A revision of the Dryinidae (Hymenoptera) Part 2

D r y i n i n ae 947 N E O T R O P I C SPEC I ES O F TRI DRYJVU5 T w e l v e spec i es are known . K E Y TO THE S P ...

Author: Massimo Olmi

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D r y i n i n ae

947

N E O T R O P I C SPEC I ES O F TRI DRYJVU5

T w e l v e spec i es are known .

K E Y TO THE S P ECI E S

F E MALES

2

H ead u n u su ally e x cav a t e d , w i t h ey es v e r y b u l gin g ( F ig . 6 4 3 A , ..... ... 2 B) . • . • • . . . . . . • • . • • • . • . . • • H ead c on v e x o r f l a t o r n o t v e r y ex cav ated, w i th ey es n o rma l l y b u lg i n g ( F i g . 6 4 3 C ) . . . . . . . . . . . . . . . . . . . . . . .. . . . . . . . . . . .4 H ead mo r e e x cavated ( F i g . 6 4 3 A I , ey es mo r e b u lg i n g ( F i g . 6 4 3 A I............................ 1. ma xiexop h t h a lmi c u s n . sp . Head less ex cav a t ed

3

4 5

6

7

8

( F i g . 643 . . . . BI . . . . . . . . . . . . . . . . . . H ead with P O L sligh t ly l o n g e r a c e I I us . . . . . . . . . . . . . . . . . . • . . .

BI ey es less b u l g i n g ( F i g . 6 4 3 .3 . . . . . . . . . . . . . . . . . . . . . . . . . . . t h an b r ead t h o f each pos t e r i o r . 8 . exophthalmicus n . sp .

H ead w i t h P O L sh o rt e r t han b r ead t h o f post e r i o r o c e l l i . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9 . que c h u a nus n . sp . .5 F o r e tarsi w i th s e gme n t 1 p r o d u c e d i n t o a h o o k . . . . .6 Fo r e tarsi with segmen ts 2 an d 3 p r o d u c e d i n t o h o oks . E n l a r ged claw with su b ap i cal t o o t h lo ca t e d v e r y n ea r apex ( F i g . 6 4 5 1; c o c o o n c i r c u la r . . . . . . . . . 2. rufic auda ( R i c h a r d s ) E n la r ged claw w i t h subap i cal to o t h mo r e d i stan t f r o m apex ( F ig . 6 4 7 1; c o c o o n ov 0 i da I . . . . . . . . . . 3 . poecilooterae ( R i c h a rd s ) H ead w i t h OL n ea rly t h r e e t i mes o r mo r e than t h r e e t i mes as l on g as P O L . . • . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 H ead w i t h O L s l i gh tly mo r e o r less l o n g e r than P O L . . . . . . . . . . . 9 Rad i al v e i n weakly c u r v i l i n ea r ( F i g . 6 4 8 A ) ; p r on o t u m almost f u l l y b l a ck . . . . . . . . . • . . . . . . . . . 4 . maximus n . sp . Rad i al v e i n s t r o n gly S - sh ap e d ( F i g . 6 4 8 B); p r on o t u m f u lly r e d d i sh - t est a c e o u s . . . . . 8 H ead ex cavate d , b l aCk , w i th c l y pe u s and man d i b l e s testac e o u s .

..... ..... ............ .5. stri a ticer)s ( K i e f f e r : - H ead almost f l at , r e d d ish . . . . . . 6 . ruficeps (Came r o n ) 9 Segmen t o f f r o n t ta r su s mo r e th an t w i c e as l o n g as segme n t 4

.

. 1 0

S e gmen t

of

f r o n t ta rsus less th an t w i c e as l o n g as segme n t 4 .

948

Mem. Amer.

•

10 Head black,

with

·

.

.

.

•

•

.

.

.

•

.

.

.

.

.

.

•

f ro n t

some t imes o rb i t s

Ent.

•

•

•

•

part

Inst.,

•

t e staceous; •

.

.

.

.

.

.

.

.

.

.

.

.

of

.

11

.

.

.

.

Disc

.

.

.

of

.

.

.

.

.

.

.

.

•

•

.

.

.

.

.

.

p r o n o t um du l l ,

.

.

•

.

•

of

1 1 .

.

•

•

•

•

•

•

•

•

•

.

.

•

.

•

clypeus,

a n dinus

n.

•

•

•

•

•

•

•

•

•

•

•

•

11

mandi b l e s and smal l ....

sp.

scutum with areolae b road......

gib b o s u s

12.

black,

p ronotum sh i ny ,

•

scutum with areolae .

n.

sp.

granulat ed and with

p icescens

around disc..................10. Disc

•

37, 1984

v e r t ex ,

Head f ully r eddish-t estaceou s ; •

•

no.

t e staceous,

not

n.

granulat ed,

n igrofla vus

st riae around disc............ 7.

some

st riae

sp.

n.

with

some

sp.

MALES

No t auli c e s visib l e and comp l e t e ; not

v i si b l e b eh i nd t h e

w i t h a median 2

ke e l

t emp l e s;

Notau l i c e s i nv i sib l e ;

occipital vei n

of

radial

f ro n s w i t h a median wi thout •

•

•

•

•

•

•

LJi stal

a median •

st rong •

•

•

•

•

•

•

•

part

wi thout

·

•

•

•

of

.

•

•

•

.

•

•

•

.

.

•

•

•

•

•

•

.

post e ri o r

•

•

•

•

v ei n

ke e l ;

keel ; •

•

.

•

•

•

.

•

•

.

•

surface

than

of

•

2.

•

.

•

•

2

part

propodeum

proximal

surface

circular .

complet e..........

l o ng as p roximal

.

p oecilopterae ( R i chards)

3.

•

l o ng e r

.

propodeum

(Ki eff e r )

ovoidal ....................

post eri o r

cocoon •

carina

nearly as

cocoon

•

radial

a median median

•

•

ke e l ;

ke e l ;

carina i n comp l e t e , surface o f

s tria ticeps

. . . . . . . . . . . . 5.

Di stal

part

o c c i p i t al post e r i o r

•

.

•

•

.

of •

•

•

part ;

frons

propodeum wi t h a •

•

•

•

•

•

•

•

•

•

•

•

•

•

•

•

•

.

rufic a u da ( R i chards)

The males of other species are u nknow n.

1.

Ft:MALE:

Tridryin u s ma xiexoph th a lmicus

fully winged;

testaceous,

with sides testaceous; abdomen black; hind coxae bulging,

shiny,

pronoturn dull,

8,75

mm;

brown;

mesothorax,

head

(Fig.

643 A)

without sculpture, invisible;

OL

sp.

head testaceous;

pro pectus black;

metathorax

abdomen black-brown;

black;

occipital carina hairy;

le ngth

with segments 2-5

n.

anten nae

pro notum

legs testaceous,

with

unusually excavated,

with

black,

and propodeum black; partly mid and with eyes very

numerous weak longitudinal keels;

more than three times as lo ng as POL;

with sides without sculpture and disc with irregular keels,

anterior transversal im pressio n and anterior collar very weak;

humped,

raised into

a carina;

posterior collar invisible;

disc

posterior transversal irnpressio n and

pro notal tubercles not reaching teguJaej

scutellum and metanotum reticulate rugose,

shiny;

scutum,

notaulices invisible;

949

Dryininae

Fig. 644

-

Chela of Tridryin us exoph t h a lmic u s n. sp.

(type)

propodeum reticulate rugose, with 2 longitudinal keels on poste"i",' "urface; fore-wing with 2 brown transversal bands and with distal part of radial vein longer' '.hem proximal part ( 2 1: 18); segment 1 of front tarsus nearly as long as segment 4; segment" 2 and 3 of front tarsus produced

Mem . A m er .

950

Fig.

645

-

Che l a

of

En!.

I n s!. ,

no.

37, 1984

Tridryinus l'uficauda

( R i chards ) ( t y p e )

951

D r y i n i n ae

i n to h oo k s ;

en l a rged c l a w

segme n t:; of front tars u s

644) w i th sub a p ic a l tooth a n d 18 l a m e l l ae;

( F i g. ( F i g.

644 )

w i th 2 r o w s of a t l e a s t 60 lame l l ae

exte n c! i n g co n ti n u o u s ly to the a p ex; MAL E :

ti b i a l

s p u r s 1 , 1 ,2 .

u n kn ow n

L O C U S TYPI C US :

Moe n ga t a p oe-Vii a v>i a

TYP I CAL IvIAT ERIAL : H OSTS:

h o l oty p e F !

( Su rin a"lf·)

i n LE .

unknown

D I STRltlU T I ON:

,'Iee>\"ur:;i c r e gi o n :

2.

SU RINAME :

Tridryin u s r u fic a u da

Moe n ga ta poe-lV i a w i a ,

( R i c h a rcJs)

/vlesodryin u s r u fic a u da R i c h a r d s 195 3 :

F E MAL E :

f u l l y w i n ge d 4, 25-6, 87 m m ;

m a n d i b les red d i s h - y e ! l ow ; black ;

h e a c b l a ck ,

abd orT,en f u lly b row n or b r ow n w i th

( F i g . 643 C )

du l l ,

h a i l'y ,

s h orter th a n P O L ; hairy,

sh i n y ,

w i th

r e a c h i n g tegu l a e ; s c u l p tu re d a s

retic u late r u gcse ;

a n terio r c o l l a r i n v i s i b le ;

scutum,

head;

i n vi s i b le,

i n c l in e d ;

ve i n

as lo n g as p ro x i ma l

f u lly

w i n ged;

tu b ercles n ot h a i ry ,

segm e n t 1

ba n d s a n d " i th d i stal

as w i th

p a r t of

p rod u c e d

i n to a h o ok ;

en I a rged c l a w

n e a r a p e x a n d w i th 20 lam e l l ae ( F i g .

( Fig.

645 ) w i th 2 row s o f a t l e a s t 80

l a m e l l ae exten d i n g c o n t i n u ou s ly to th e a p e x ; MAL E :

p o s te r i or c o l l a r very

p r on ota l

segme n t 1 of f r o n t ta r s u s s l i ghtl y

part;

located very

segme n t 5 of f r o n t ta r s u s

OL

pron otu n l

p os te r i o r s u rface w i th 2 lo n g i tu d i n a l

fore- w i n g w i th 2 b row n tr a n s ver sa l

tooth

head

p r o p o d e u m retic u l a te r u gose ,

rad i a l

s h orter th a n segm e n t 4 ;

cari na;

s c u te l l u m a n d rnetan otUrTi s h i n y ,

n ot a u l i ces

l egs red d i s h ,

occ i p i ta l c a r i n a comp l e te ;

to u c h i ng occ i p i ta l

kee l s;

6 45 ) ;

apex redd i sh;

n u mero u s str i a e arou n d d i sc ;

poste r i or s u r f a c e very

w ith s u b a p i c a l

w i th c l y p e u s a n d

thorax a n d p ro p od e u rT'

mid a n cJ h i n d c l u b s of feclOra b r ow n ;

p o s terior oce l l i

reduced a n d s h ort;

n . comb .

59 .

a n te n n a e tes taceOU5j

w i th m i d a n d h i n d c ox a e b l ack ,

LE!

l e n gth 4 m m ;

b l ack ;

ti b i a l s p u r s 1 , 1 , 2 .

m a n d i b l e s,

c l y peus a n d partl y

gen ae testaceou s; a n te n n a e testaceO U5j l egs te staceOU5, w i th h i n d cox a e p a r tly b ro w n ; a n te n n a e n o t d i s ta l ly th i c k e n e d j a n te n n a l segm e n ts i n fo l l o w i n g p ro p o r t i o n s :

9:

5:

21 :

1 3:

1 2:

1 2:

f u l l y retic u l a te r u gose ; fro n s w i th ou t med i a n c a r i n a com p l e te ; POL = 8; OL = 3; O OL = 5; s c u te l l u m a n d meta n ot u m d u ll, p ro p od e u m reti c u la te r u go s e ; med i a n

l on g i tu d i n a l

k ee l ;

1 0:

1 0:

and UPL

9:

1 0, 5;

head du l l ,

l a te r a l k ee l s ; occ i p ital = 1 ; TL = 0; s c u torTl,

f u l l y reti c u l a te r u gose ;

n o ta u l i c es i n vi s i o l e ;

p o s te r i o r s u rf a c e of p rop o d e u m w i th

b a nd b e n e a th the p terosti gm a ;

d i s ta l

p a rt of r a d i a l

ve i n

l on ger th a n

p rox i m a l p a rt ( 1 4 : 1 1 ) ; gen i ta l i a : f i g . 646; t i b i a l s p u rs 1, 1 , 2 . L O C U S T YP I C U S : S a n Ped r o d e iv\o n te s d e Oca ( C osta R i c a ) T Y P I CAL MATER I AL : HOS T S:

a stro n g

fore-w i n g h y a l i n e o r w ith a w e a k tr a n sve r s a l

h o l oty p e F !

in Trin id a d Orrnenis s p .

i n IVA. on C a c a o .

952

Mem . Amer .

F i g . 646

En!.

I n s! . ,

no.

37,

1984

M a l e ge nitalia of Tridryinus ruficauda (Richards) from S ! . Au g u s t i n e (T r i n i d a d ) (r i g h t h a l f remove d )

-

U I S T R I B U T I ON :

Neo t r o p ic reg i o n : COSTA R I CA : S a n P e d r o d e Mon t e s d e Oca,

S ! . Augu st i n e , OL! B RAZ I L : iVlanaus ( Am a zo n a s ) , BM! 50° 3D' W 1 2° 3�' 5 (Vi l a Vera), T W ! NOTES : t h e cocoon of Tr. ruficauda is a l m o s t ci rcu l a r a n d com posed of o n e o n l y lining. WA!

T R I N I DAD :

3.

il!esodryinus

Tridryinus poecilopterae ( R i ch a rd s )

p oecilop ter ae

r�

i ch a r d s 1 947:

Mesodryinus poecilopterae Ri ch a rd s : Mesodryinus sp . :

Teran

1 980 :

289 .

n . comb.

868.

Ri ch a r d s 1 9 5 3 :

59 .

D r y i n i n ae

953

F i g . 647 - C h e l a of Tridryinus p oec ii op te ra e ( R i c h a rds ) from 51. A u g u s t i n e ( Tr i n idad )

F E MA L E : f u l l y w i n g ed ; l e n g t h 6 , 0-7 , 5 m m ; head [,iceck, w i th c l yp""" errH ", a n cJ i b l e s red d i sh- tes t aceo u s ; a n t e n n a e b row n , w i t h segme n t s 1 - 2 redd i sh ; t horax a n d propodeum b l ac k ; a b domen b l ac k ; l e g s b ro w n , w i t h t i b ia e , femora a n d p a r t l y t roc h a n ters of f o r e l e g s redd i sh ; h e a d h a i r y , d u l l , ret i c u l a te ru gose, occ i p i t a l c a r i n a comp l e t e ; po s t e r i or oce l l i touch i n g occ i p i t a l ca r i n a ; O L shorter t h a n POL ; pronot u m h a i r y , sh i n y , w i t h n u merous s t r i ae a ro u n d d i sc ; pos t e r i o r co l l a r very reduced a n d s h o r t ; a n t e r i o r co l l a r i n v i s i b l e ; p ron o t a l t u b erc l e s not rea c h i n g te g u l ae ; scu t u m , sc u te l l u m a n d m e t a n o t u m sh i n y , h a i r y a n d r e t i c u l a t e ru gose ; n o t a u l i ces i n v i s i b l e ; p ropode u m re t i c u l a t e ru gose, w i t h pos t e r i or s u rf a c e v ery i n c l i n ed

954

Mem . A me r . E n ! .

I ns ! . ,

no.

37 ,

1 984

� �� . . .. . .

F i g . 648 - R a d i a l v e i n s of fema l e s of Tridryinus maXlmus n . s p . a n d striaticeps ( K i ef f e r ) ( B ) f rom Su r i n a me

( t y pe ) ( A )

a n d w i t h 2 l o n g i t u d i n a l k ee l s ; fore-w i n g w i t h 2 brown t r a n s v e r s a l b a n d s a n d w i t h d i s t a l p a r t o f r a d i a l v e i n a s l o n g a s p rox i ma l p a r t ; segme n t 1 of f ro n t t a r s u s n e a r l y as l o n g as segmen t 4 ; segmen t 1 p ro d u ced i n to a hook ; e n l a r g e d c l a w w i t h a su b a p i c a l too t h a n d 24 l a me l l ae ( F i g . 647 ) ; s e g me n t 5 of f r o n t t a rs u s ( F i g . 647 ) w i t h 2 rows of a t l e a s t 80 l a me l l ae e x t en d i n g con t i n u o u s l y to t h e a p e x ; t i b i a l s p u rs 1 , 1 , 2 . M A L E ( F i g s 634, 635 ) : f u l l y w i n g e d ; l e n g t h 4 , 0-4 , 7 m m ; h e a d b l ack , w i t h m a n d i b l e s r e d d i s h ; a n te n n a e brow n , w i t h seg men t s 1 -2 a n d 1 0 redd i s h ; some t i mes o n l y seg m e n t s 1 -2 a n d p a r t l y 3 red d i sh ; t h o r a x a n d p ropodeum b l ack ; a b domen b ro w n ; l e g s redd i s h - t e s t aceo u s , w i t h mi d a n d h i n d cox a e b row n ; some t i mes o n l y h i n d cox a e p a r t l y bro w n ; h e a d d u l l , re t i c u l a t e r u gose ; occ i p i t a l c a r i n a comp l e te ; a n t e n n a e n o t d i s t a l l y t h i c k ened ; a n t e n n a I seg men t s i n fo l l o w i n g p ropo r t i o n s : 1 2 : 9 : 2 9 : 1 5 : 1 8 : 1 5 : 1 4 : 1 2 : 1 1 : 1 4 ; f ro n t a l l i n e comp l e t e ; P O L = 1 0 ; Ol = 5 ; O O l = 8 ; O P l = 2 , 5 ; Tl = 2 ; scu t u m , sc u te l l u m , met a n o t u m a n d p ropodeum d u l l , ret i c u l a t e ru gose ; n o t a u l i c e s i n v i s i b l e ; pos t e r i o r s u rface of p ropodeum v e r y i n c l i n ed , w i t h o u t l o n g i t u d i n a l k ee l s ; fore-w i n g w i t ho u t d a rk t r a n sv e rs a l b a n d s o r w i t h a w e a k d a r k b a n d b e n e a t h t h e p t e r os t i g m a ; d i s t a l p a r t of rad i a l v e i n n e ar l y a s l o n g a s p ro x i m a l p a r t ( 1 6 : 1 5) ; t i b i a l s p u r s 1 , 1 , 2 . lOCUS TYP I C U S : S I . A u g u s t i n e ( T r i n i d ad ) T Y P I CAL MAT E R I A L : ho l o t y pe F ! a n d 1 p a ra t y p e Ivi ! i n BM . H O STS : i n T r i n i d a d Poekilloptera (=Poeciloptera) phalaenoides ( l . ) (Fl a tidae) ( R i c h a r d s 1947 ) ; i n E I Sa l v a d o r Poekilloptera completa Jac . ( R i c h a rd s 1 95 3 ) ; i n Venez u e l a Poekilloptera phalaenoides ( l . ) ( Te r a n 1 980 ) . D I ST R I B U T I O N : Neotrop i c reg i o n : T R I N I DA D : S! . Au g u s t i n e , BM ! E l SALVA D O R : S a n S a l v a d o r , O l ! PANAMA : Ba l boa ( C a n a l Zon e ) , AM ! V E N E Z U E L A : C a racas ( D . F.), WA ! O l !

4 . Tridryinus maximus n . s p .

D r y i n i n ae

955

�.",

,

\\� F i g . 649

-

.

Ch e l a of Tridryillus maxil1lus n . sp .

( type)

\\

\J

956

M e m . Amer . E n ! .

I nst . ,

no.

37 ,

1 984

F E M A L E ( F i g s 632, 63 3 ) : f u l l y w i n ged ; l e n g t h 9 , 37 m m ; head b l ac k , w i t h c l y pe u s w h i t i s h except f o r a med i a n reg i o n b l ac k ; e y e s su rrou n d e d b y a n a r row w h i t i s h s t r i p e ; a n tennae b l a c k , w i t h segme n t s 6- 1 0 l i g h t ; m ax i l l a r y p al p i w h i t i s h ; propec t u s b l ack ; p ro no t u rn b l ac k , w i t h s i d es p a r t l y red d i sh ; meso t h o r a x , m e t a t h o r a x a n d propodeum b l ac k ; a b domen b l a c k - b row n : l eg s b row n , w i t h cox a e b l ack ; dorsa I reg i o n o f s t a I k s o f fore femora a n d dorsa I reg i o n of fore t i b i ae w h i t i s h ; h e a d d u l l , g r a n u l a ted ; oce l l a r reg i o n c o n v e x ; vertex exc a v a t e d ; oce l l a r reg i o n a n d su rrou n d i n g reg i o n w i t h n u merous l o n g i t u d i n a l k ee l s ; occ i p i t a l c a r i n a comp l e t e ; t em p l es i n v i s i b l e ; poster i o r oce l l i touch i n g occ i p i t a l c a r i n a ; O L n e a r l y t h ree t i mes a s l o n g a s POL ; p ro n o t u m s h i n y , w i t h n u merous t r a n s v e rs a l k ee l s on d i sc a n d w i t h p u nc t u res a n d some k ee l s or areo l ae on s i de s a n d on a n te r i o r c o l l a r ; a n t e r i o r t r a n s v e rs a l i m p ress ion s t ron g ; a n t e r i o r co l l a r d i s t i n c t ; pos ter i or t r a n sver s a l i m press i o n s t r o n g a n d l o c a ted n e a r pos t e r i o r m a r g i n of p ro n o t u m ; pos t e r i o r co l l a r s h or t , d i s t i n c t ; d i sc h a i r y ; p rono t a l t u berc l es n o t reach i n g t e g u l a e ; scu t u m , s c u te l l u m a n d rne t an o t Unl ret i c u l a t e ru gose ; no t a u l i c e s i n v i s i b l e ; porpod e u m ret i c u l a t e r u g o s e , w i t h 2 l o n g i t u d i n a l k ee l s o n pos t e r i o r s u rf ace ; fore-w i n g w i t h 2 b row n t r asv ers a l b a n d s a n d w i t h r a d i a l v e i n n o t S- s h a ped ( F i g . 648 A ) ; d i s t a l p a r t of r a d i a l v e i n l on g e r t h a n p rox i m a l p ae t ( 24 : 1 7 ) ; seg m e n t 1 of f ro n t t a r s u s n ea r l y a s l o n g a s seg men t 4 ; seg men t 3 of f ro n t t a rs u s produ ced i n to a h ook ; en l a r ged c l aw ( F i g . 649 ) w i t h s u b a p i c a l too t h a n d 30 l a me l l a e ; segme n t 5 of fron t t a r s u s ( F i g . 649 ) w i t h 2 row s of a t l e a s t 80 l a me l l ae e x t e n d i n g c on t i n u o u s l y to the a p e x ; t i b i a l s p u rs 1 , 1 , 2 . IiIAL E : u n k now n LOCUS TYP I C U S : Z a n d e r i j ( Su r i n a m e ) TYP I CAL MAT E R I AL : h o l o t y p e F ! i n LE . HOSTS : u n k n o w n D I ST R I B UT I O N : Neo t ro p i c reg i o n : S U R I N AM E : Z a n de r i j , L E !

5.

Tridryinus striaticeps

( K i effer )

Prodryinus striaticeps K i effer 1 9 0 9 :

( K i effer ) : Tridryinus striaticeps ( K i effer ) : Tridryinus striaticeps

334 . K i effer 1 9 1 3c : 325 . K i effer 1 9 1 4b : 29 .

F E M A L E : fu l l y w i n g e d ; l e n g t h 5 , 00-7 , 50 m m ; h e a d b l ac k , w i t h c l y p e u s a n d m a n d i b l e s y e l l o w ; a n te n n a e b row n ; a n t e n n a e b row n , w i t h seg men t s 6- 1 0 t e s t aceo u s ; p ropec t u s a n d pro n o t u m redd i s h ; meso t hora x , m e t a t h orax a n d propod e u m b l a c k , abdomen b l ac k -y e l l o w ; l eg s t e s t aceou s , w i t h s t a l k s of h i n d femora a n d h i n d t i b i ae b row n ; he a d ex c a v a te d , sh i n y , w i t h n u mero u s l o n g i t u d i n a l k e e l s ; oc i p i t a l c a r i n a comp l e t e ; poste r i o r oce l l i t o u c h i n g occ i p i t a l c a r i n a ; OL more t h a n t h ree t i mes as l o n g as POL ; p ro n o t u m sh i n y ,

hairy,

w i t h n u merous w e a k s t r i ae a ro u n d d i sc ;

a n t e r i o r t r a n sv ersa l

957

Dryininae

Fig.

650

-

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of

Tridryinus striatice}ls

(Kieffer)

from

Suriname

958

Mem . Amer .

En t .

I nst . ,

no.

37,

1 984

i mpres s i on and a n te r i o r co l l a r very w e a k ; pos t e r i o r t r a n s v e r s a l i mp r e s s i o n s t rong a n d l oc a ted v e r y n e a r pos ter'ior rnal'S:j i n of p ro no t u m j p o s t e r i o r co l l a r i n v i s i b l e ; p ro no t a l t u b erc l e s n o t reach i n g t e g u l a e ; scu t u m , scu te l l u m a n d m e t a n o t u m re t i c u l a t e r u g ose ; n o t a u l i ce s i n v i s i b l e ; p ropodeum ret i c u l a t e r u g os e , w i t h 2 l on g i t u d i n a l k ee l s o n pos t e r i o r s u rface ; fore-w i n g w i t h 2 b ro w n t r a n sv e r s a l b n a d s a n d w i t h d i s t a l p a r t of r a d i a l v e i n ne a r l y a s l on g a s prox i m a l p a r t ; d i s t a l p a r t of ra d i a l v e i n s t ron g l y S-s h a ped ( F i g . 648 B ) ; segme n t 1 o f f r o n t t a r s u s l on g e r t h a n seg m e n t 4 ( 1 8 : 1 5 ) ; segme n t s 2 a n d 3 of f r o n t t a r s u s p ro d u ced i n t o hook s ; en l a rged c l a w ( F i g . 650 ) w i t h s u b a p i c a l tooth a n d 1 3 l a me l l ae ; segme n t 5 of f ro n t t a rsus ( F i g . 650 ) w i t h 2 rows of 40 l ame l l a e ex tencJ i r's; con t i n u o u s l y to t h e a p e x ; t i b i a l s p u rs 1 , 1 , 2 . M A L E : f u l l y w i n g e d ; l e n g t h 4 , 75-4 , 87 m m ; b l ac k ; m a n d i b l e s a n d c l y pe u s t e s t ac eo u s ; a n t e n n a e b row n , w i t h segmen t s 1 - 2 test aceo u s ; l eg s t e s t aceO U 5 , w i t h m i d a n d h i n d cox a e , h i n d femora a n d h i n d t i b i ae p a r t l y b l ack o r b row n ; a n t e n n a e n o t d i s t a l l y t h i c kened ; a n t en n a l segme n t s i n fo l l ow i n g p ropo r t i o n s : 8 : 7 : 23 : 1 3 : 1 4 : 1 4 : 1 2 : 1 0 : 9 : 1 1 ; h e a d d u l l , f l a t , g r a n u l a ted a n d s t r on g l y re t i c u l a t e r u gose ; f r o n s w i t h o u t med i a n a n d l a t e r a l k ee l s ; occ i p i t a l c a r i n a i ncomp l e t e , n o t v i s i b lf> bE-h i n d t h e t em p l es ; POL ; 8 ; OL ; 4 ; OOL ; 4 ; OPL ; 3 ; TL ; 0 ; scu t u m , scu t e l l u m a n d met a n o t u m d u l l , g r a n u l a ted a n d s t r on g l y re t i c u l a te r u g o se ; n o t a u l i ces comp l e t e , p o s te r i or l y sep a ra t ed ; p ropodeum re t i c u l a t e rugose, w i t h o u t t r a n s v e rs a l k ee l s ; p o s te r i o r s u rf a ce w i t h a med i a n s t r o n g l o n g i t u d i n a l k e e l ; fore-w i n g w i t h a d a r k t r a n s v ersa l b a n d b e n e a t h t h e p te ros t i g m a ; d i s t a l p a r t of r a d i a l v e i n l on g e r t h a n p ro x i m a l p a r t ( 1 8 : 1 5 ) ; t i b i a l s p u r s 1 , 1 , 2. L O C U S TYP I C U S : P a r a ( B r az i l ) TYP I CA L M A T E R I AL : ho l o t y p e F ! i n CA . H OS T S : u n k n o w n D I STR I B UT I ON : Neot rop i c reg i o n : SUR I N AME : M a R e t r a i te p l a n t a t i o n ( P a r a m a r i bo ) , L E ! BOL I V I A : At Mou t h of R i o B a u re s ( R i o I te n e z , De p t . B en i ) , AM ! B R AZ I L : P a r a , CA l Teodoro S am p a i o ( S ao P a u l o ) , TW ! OL! J a t a i ( Go i a s ) , T W !

6.

Tridryinus ruficeps ( C a m eron )

n . com b .

Dryin u s ruficeps C a meron 1 88 8 : 445 . Prodryin u s ruficeps ( C ameron ) : K i ef f e r i n K i ef f e r et M a rs h a l l 1 906 : 498 . Prodryin u s ruficeps ( C ameron ) : K i effer 1 9 1 4b : 5 1 . Les todryinus dichrous F e n t o n 1 927 : 1 ( n . sy n . ) ; locus typ ic u s C h a p a d a ( Br a z i I ) •

F E M AL E :

f u l l y w i nged ;

l e n g t h 5 , 0-7 , 1 8 mm ;

head,

a n te n n a e ,

p ropec t u s a n d

Dry i n i na e

651

F i g.

p ro n o t u m r e d d i s h ;

- C h e la of Tridryinus ruficeps

rneso t h orax ,

meta thorax

b ro w n - red d i s h o r redd i s h ;

l egs redd i sh ;

gra n u l a t e d ,

l ongi t u d i n a l

w i t h n u merous

occ i p i t a l c a rin a ve r tex ;

inc o m p l e te ,

OL more t h a n

occ i p i t a l c a r i na ;

9:'9

vis i b l e on l y

t h ree t i rnes as

p r o n o t u rn h a i r y ,

and p ro p o de u rn b lack ;

head sh i n y ,

ab d o m e n

a l m o s t fla t ,

weakl y

k ee l s on fro n s an d vertex ; beh i n d t h e oce l l i

l o n g as POL ;

sh i n y ,

( C amero n )

w i th

a n d on s i des of

p o s t e r i o r oce l l i touch i n g

n u mero u s kee l s a r o u n d d i sc ,

960

lvie m . Amer . E n t .

I n st. ,

no.

37,

1 984

A

B

Fig .

652 - Pronotum of fema l es of Tridry in u s n igrofl a vu s n . sp . from Nova Teuto n i a ( 5r a zi l ) ( A )

a n d picescen s n . sp .

( ty p e ) ( B )

with a posterior tr a n sversa l impression n e a r posterior mar gin ( th e poste r i o r co l l a r so is invis i b le ) ; t h e a n terior tra n sversa l imp ression is w e a k ; the p r o n otum so h a s a sw o l l e n disc a n d a not ve ry visible a n terior c o l l a r ; p ro n o t a l tuberc l es not r e a c h i n g tegu l a e ; scutum d u l l , reticu l a te r u gose; notau l ices i n visib l e; inste a d of t h e notau l ices there a re 2 l on gitudin a l k ee l s reachin g a p p roximate l y 0 , 5 l e n gth of scutum; scute l l um a n d met a n o tum retic u l ate rugose; p ro p odeum ret i c u l ate rugose ; posterior surface of p ro p odeum very inc l in e d , with 2 l o n gitudina l k ee l s ; fore- w i n g with a distinct b r o w n tr a n sversa l b a n d ben eath the pterostigma a n d with 2 t r a c k s of p ro x im a l b ro w n tra nsversa l b ands; dista l p a rt of r a d ial ve i n a s l on g a s p roximal p a rt ; seg me n t 1 of front ta rsus n e a r l y a s l ong a s segme n t 4; se g me n ts 2 - 3 p ro duced i n t o a hOOK; enalrged c l a w with sub a pic a l tooth and 1 3 l a me l l ae ( Fig . 6 5 1 ) ; segment 5 of front t a rsus ( F i g . 651) with 2 rows of at l e ast 50 l ame l l a e e x te n d i n g c o n tin uous l y to the a p e x ; tib i a l spurs 1 , 1 , 2 . MAL E : u n k n o w n LOC U S TYP I C U S : Bug a b a ( Pa n anla ) TYP I CAL MAT E R I AL: ho l o t y p e F! of D. ruficep s in 81'-1; ho l oty p e F! a n d 1 p a r a t y p e F! of L. dichrous i n WA . HOSTS: u n k n o w n O I STRIBU T I ON : Neotropic region: PAI�AMA: Bug a b a , BM! SUR I NAME: K w atta , L E ! P a r amaribo, TW! COLOMBIA: M a r t a ( Rio D o n Die g o , I� Sierra Neva d a d e S . ) , B M ! V E N E Z U E LA: San Esta b a n ( Puerto C a b e110), TW! E C UADOR :

Dry i n i nae

961

F i g . 653 - C h e l a o f Tridry in u s n igrofl a v u s n . s p .

( ty pe )

L i moncoc h a ( N a po ) , O L ! O T ! P E R U : T i n g o M a r i a ( Monzon V a l l ey ) , C A ! Q u i n c em i l ( M a r c a p a t a ) , UL ! TW ! BRAZ I L : N o v a T e u t o n i a ( S . t a C a t a r i n a ) , U L ! B M ! Cha p a d a ( R i o G r a n d e do Su i ) , VIA ! S a n tos ( Sao P a u l o ) , ZO ! S a n t a rem ( P a r a ) , 'ivA ! J a t a i ( Go i a s ) , OT ! Represa do R i o G r a n d e ( G u a n ab a r a ) , TW ! 7 1 ° 38 ' W 4 ° 33' S ( A mazon a s ) , TW ! OL ! ARG ENT I NA : E l do r a d o ( M i s i o n e s ) ,

AM !

7.

F E MA L E :

f u l l y w i n g ed : ;

Tridryin u s nigrofla v u s n . s p .

l e n g t h 5 , 50-8 , 62 m m ;

h e a d b l ac k ,

w i t h f ro n t p a r t

962

of ver tex

Mem . Amer .

( more a l o n g orb i t s ) ,

En! .

I n s! . ,

no.

37,

1 984

c l y p e u s a n d m a n d i b l e s y e l l ow ;

a n te n n a e

t e s t aceo u s , w i t h segmen t s 2-5 a n d 8- 1 0 b row n ; propec t u s b l ack , w i t h a med i a n dors a l reg i o n y e l l ow ; p ro n o t u rn b l ack , w i t h p os t e r i or co l l a r a n d p a r t l y pos t e r i o r l obes y e l l ow ; rnesotho ra x , rne t a t h o r a x a n d p ropodeurn b l ac k ; a b dornen b ro w n -red d i s h ; l eg s t e s t aceou s ; h e a d d u l l , ret i c u l a t e ru gose ; occ i p i t a l c a r i n a i n comp l e te , o n l y v i s i b l e beh i n d t h e oce l l i a n d o n s i des of vertex ; pos t e r i o r oce l l i n o t touch i n g occ i p i t a l c a r i n a ; O P L shorter than P O L ; O L s l i g h t l y l o n g e r t h a n P O L ; p ro n o t u rn ( F i g . 652 A) sh i n y , w i t h n urnerou s l o n g i t u d i n a I a n d t r a n sversa I k e e l s , crossed by a t r a n sversa I a n t e r i o r i rnp ress i o n ; pos t e r i o r t r a n s versa l i rn p ress i o n a n d p o s t e r i o r co l l a r i n v i s i b l ej p ron o t a l t u b e rc l e s n o t reach i n g t eg u l ae ; sc u t u rn sh i n y , re t i cu l a te ru gose ; n ot a u I i ces i nv i s i b I e ; scu te I I urn sh i n y , w i t h n u rnerous l o n g i t u d i n a I k ee l s ; m e t a n o t u rn ret i c u l a t e r u g ose ; p ropodeurn ret i c u l a t e rugo se , w i t h 2 l o n g i t u d i n a l k ee l s on pos t e r i or s u rf a ce ; fore-w i n g w i t h 2 b ro w n t r a n sversal b a n d s a n d w i t h d i s t a l p a r t of r a d i a l v e i n s h o r t t h a n prox i m a l p a r t ( 9 : 1 2 ) ; segmen t 1 of f ro n t t a rs u s nea r l y as l o n g as segmen t 4 ; segmen ts 3 a n d 2 of fron t t a rs u s p ro d uced i n t o hook s ; en l a r ged c l a w ( F i g . 653 ) w i t h s u b a p i c a l tooth a n d 8 l a me l l ae ; segmen t 5 of fro n t t a r s u s ( F i g . 653 ) w i t h 2 row s of at l e a s t 60 l a m e l l ae ex t en d i n g con t i n uo u s l y to t h e a pex ; t i b i a l s p u rs 1 , 1 , 2 . MALE : u n k n o w n L O C U S TYP I C U S : N o v a T e u t o n i a ( B r az i l ) TYP I CAL MAT E R I A L : h o l o t y p e F ! a n d 1 p a r a t y p e F ! i n B M . HOSTS : u n k n o w n D I ST R I B U T I O N : Neot rop i c reg i on : BRAZ I L : Nov a T e u t o n i a ( S . t a C a t a r i n a ) , BM !

8.

Tridryi n u s exophtha lmicus n . sp .

F E MA L E : f u l l y w i n ged ; l e n g t h 6 , 93 mm ; h e a d red d i s h - te s t aceo u s , w i t h a b ro w n s p o t on oce l l a r reg i o n : a n te n n a e t e s t aceo u s , w i t h segmen t s 1 -5 d a r k e n e d ; p ro p ec t u s b l ack ; pro n o t u m b l ack , w i t h s i des y e l l ow ; meso t h orax , m e t a t h o r a x a n d propode u m b l ac k ; a b d o m e n b row n-b l ack ; l e g s t e s t aceou s , w i t h h i n d cox a e p a r t l y b l ac k ; h e a d ( F i g . 643 B ) u n u s u a l l y exc a v a t e d , w i t h e y e s very b u l g i n g , w i t h n u merou s l o n g i t u d i n a l keels on frons; occ i p i t a l c a r i n a i n v i s i b l e ; P O L s l i g h t l y l o n g e r t h a n b r e a d t h of each poster i o r oce l l u s ; OL more than t h ree t i mes a s l o n g a s PO L ; p ron o t u rn d u l l , h a i ry , w i t h n u merous kee l s a ro u n d d i sc ; a n t e r i o r t r a n sver s a l i m p ress i o n a n d a n t er i or co l l a r weak ; pos t e r i o r t r a n s v e rs a l i m p r es s i o n a n d pos t e r i or co l l a r i n v i s i b l e ; d i sc h u m p e d , r a i sed i n to a c a r i n a ; scu t u m , scu t el l u m a n d m e t a n o t u m re t i c u l a t e ru gose ; n o t a u l i ce s i n v i s i b l e ; rne t a n o t u m r a i sed i n to a med i a n c a r i n a ; propodeu m ret i c u l a t e r u g ose w i t h 2 l o n g i t u d i n a l k ee l s ; fore-w i n g w i t h 2 b ro w n t r a n s v e rs a l b a n d s a n d w i t h d i s t a l p a r t o f ra d i a l v e i n l o n g e r t h a n p ro x i m a l p ar t ( 1 8- 1 2 ) ; segmen t 1 of fro n t t a r s u s n ea r l y a s l o n g a s segmen t 4 ;

segmen t s 2 a n d 3 o f fron t t a rs u s p rod uced i n t o

D r y i n i n ae

F i g . 654

-

C he l a o f Tr idryinus exo,ohthalmicus n . sp .

963

( type)

hook s ; e n l a r g e d c l aw ( F i g . 654) w i t h su b a p i c a l too th a n d 11 l a me l l ae ; segmen t 5 of f r o n t t a rsus ( F i g . 654) w i t h 2 rows o f 44 l a m e l l a e ex ten d i n g con t i nuou s l y t o t h e apex ; t i b i a l s p u rs 1,1,2.

964

Mem . Amer .

En!.

I ns ! . ,

no.

37,

1 984

F i g . 655 - C h e l a of Tridryin us quech u a n u s n . sp .

( ty p e )

D ry i n i n ae

MAL E : u n k n ow n LOCUS TYP I C U S : N o v a Teu t on i a ( B r az i l ) TYP I CAL MA T E R I A L : ho l o t y p e F ! i n B M . HOST S : u n k n o w n D I ST R I B U T I O N : Neo t rop i c reg i o n : BRAZ I L : BM !

9.

965

N o v a Teu t on i a

Tridryin u s quech u a n u s

n.

( S . t a Catarina ) ,

sp .

FEMAL E : f u l l y w i n g ed ; l e n g t h 6 , 56 m m ; h e a d f u l l y t e s t a ceou s ; a n t e n n a e b row n , w i t h se g men t 1 t e s t aceo u s ; t ho r a x a n d p ro p o d e u m b l ac k ; abdomen b l a ck , w i t h some reg i o n s t e s t aceo u s ; l e g s b l ac k , w i t h t a rs i , t roc h a n t e r s a n d f o r e t i b i a e p a r t l y tes t ac eo u s ; a n t e n n a e d i s t a l l y t h i c k e n e d ; a n te n n a l segmen t s i n fo l l o w i n g p ro p or t i on s : 1 6 : 8 : 3 7 : 1 5 : 1 3 : 1 1 : 9 : 9 : 7 , 5 : 1 0 ; h e a d d u l l , v e r y exca v a t e d , w i t h eyes very b u l g i n g ; f ro n s a n d v e r tex s c u l p t u red b y weak l o n g i t u d i n a l s t r i a e ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 1 ; OOL = 1 6 ; PUL shorter t h a n b read t h of each p o s t e r i o r oce l l u s ; p ro n o t u m h u mped , w i t h a s t ro n g pos t e r i o r t r a n s v e rs a l i mp r e s s i o n a n d w i t h a w e a k a n te r i o r t r a n s v e rs a l i m p re s s i o n ; po s t e r i o r co l l a r i n v i s i b l e ; p ron o t a l t u berc l e s n o t rea ch i n g t e g u l a e ; sc u t u m d u l l , f u l l y re t i c u l a t e r u gose ; n o t a u l i ces i n v i s i b l e ; sc u t e l l u m f u l l y re t i c u l a t e r u g os e ; met a n o t u m ra i sed i n to a med i a n c a r i n a , ret i c u l a t e r u g ose n e a r a n terior m a r g i n and smoo t h n e a r p o s t e r i o r m a r g i n ; propodeum ret i c u l a t e r u g o s e , w i t h o u t a t r a n s v ersa l k ee l b e t w een dor s a l a n d p o s te r i o r s u rface ; p o s t e r i o r s u rface w i t h t w o l o n g i t u d i n a l k ee l s ; fore-w i ng w i t h t w o d a r k t r a n sv e r s a l b a n d s , o n e o n t h e b a s a l c e l l s a n d a n o t h er b e n e a t h t h e p t eros t i g m a ; d i s t a l p a r t of ra d i a l v e i n l o n g e r t h a n p rox i m a l p a r t ( 1 7 : 1 4 ) ; fore t a rs a l seg men t s i n fo l l ow i n g propor t i o n s : 24 : 4 : 8 : 1 8 : 32 ; segmen t s 2 a n d 3 o f f ro n t t a r s u s produ ced i n to hook s ; en l a rged c l a w ( F i g . 655 ) w i t h a s u b a p i c a l tooth and 1 5 l a me l l a e ; se g m e n t 5 of f ro n t t a r s u s ( F i g . 655 ) w i t h 2 row s of n e a r l y 47 l a me l l ae ex t e n d i n g con t i n u ou s l y to t h e a p e x ; t i b i a l s p u r s 1 , 1 , 2 . MAL E : u n k n o w n L O C U S TYP I C U S : P u y o ( E c u ador ) TYP I CA L M A T E R I AL : ho l o t y p e F! i n O T . HOST S : u n k n o w n D I S T R I B U T I O N : N eo t rop i c reg i on : ECUADOR : P u y o ( P a s t a z a ) , O T !

10.

Tridryin u s picescens n . s p .

FEMA L E : f u l l y w i n g ed ; l e n g t h 5 , 0 mm ; h e a d b l a ck , w i t h f ro n t p a r t of v e r t e x , c l y p e u s a n d rna n d i b l e s t e s t a ceou s ; a n t e n n a e b row n , w i t h seg m e n t t e s t a c eo u s ; occ i p u t redd i s h ; tem p l es w i t h a sm a l l red d i s h s po t ; t h orax a n d p ropodeurn b l ac k , a b domen b ro w n - re d d i sh ; l e g s t e s t a ceou s , w i t h rn i d a n d

966

Mem . Am er . E n ! .

I n s! . ,

n o . 37,

1 9 84

F i g . 656 - C h e l a of Tridryinus p icescens n . sp .

( type)

h i n d cox a e b l ac k ; h e a d d u l l , h a i ry , ret i c u l a t e ru gose ; occ i p i t a l c a r i n a comp l e t e ; pos t e r i or oce l l i n o t touch i n g occ i p i t a l c a r i n a ; O P L shorter t h a n O L ; OL s h o r t e r t h a n PO L ; prono t u m ( F i g . 652 B ) d u l l , h a i ry , g r a n u l a t e d , w i t h n u m e ro u s s t r i a e a ro u n d d i sc ; a n te r i o r t r a n sver s a l i mpres s i o n a n d a n te r i o r co l l a r weak ; pos t e rior t r a nsver s a l i mp ress i o n s t ro n g a n d l o c a t e d

D r y i n i n ae

F i g . 657 - C h e l a of Tridryinus a n dinus n . sp .

967

( type)

n e a r po s t e r i o r m a r g i n o f prono t u m j pos t e r i o r co l l a r i n v i s i b l e j p ro n a t a l t u b erc l e s not reac h i n g t eg u l ae ; scu t u m , s c u t e l l u m a n d m e t a n o t u m ret i c u l a te r u gose ; not a u l i ce s i n v i s i b l e ; propode u m ret i c u l a t e r u g o s e , w i t h 2 l on g i t u d i n a l k ee l s on pos t e r i o r s u rf a c e ; fore- w i n g w i t h 2 b ro w n t r a n s v e rs a l b a n d s a n d w i t h d i s t a l p a r t of r a d i a l ve i n l o n g e r t h a n p rox i mo l p a r t ( 1 5 : 1 0 ) ; s e g m e n t 1 o f fron t t a r s u s l es s t h a n t w i ce a s l o n g a s seg men t 4 ( 1 9 : 1 2 ) ; s e g m e n t 3 of f ro n t t a r s u s p ro d u c ed i n to a hook ; e n l a r g ed c l aw ( F i g . 656 ) w i t h s u b a p i c a l too t h a n d 6 l a m e l l a e ; s e g m e n t 5 of f ron t t a r s u s ( F i g . 656 ) w i t h 1 row o f 1 3 lame l l ae ; a pex w i t h a g ro u p o f 1 3 l a m e l l a e ; t i b i a l s p u rs 1 , 1 , 2 . MALE : u n k n ow n LOCUS TYP I C US : Las Cej a s ( A r g e n t i n a ) TYP I CAL MAT E R I A L : h o l o t y p e F! i n T U . H O ST S : u n know n DISTR I B U T I O N : I�eo t rop i c reg i o n : A R G E N T I NA : L a s Cej a s ( T u c u m a n ) , TU !

11.

Tridry in u s andinus n . s p .

F E MAL E : f u l l y w i n g ed ; l e n g t h 5 , 00-5 , 62 m m ; h e a d b l ack, w i t h f ron t p a r t of ver tex , cly p e u s and m a n d i b l e s t e s t aceo u sj som e t i me s a l so orb i t s t e s t aceou sj occ i p u t b l ac k or t e s t aceou s ; an t e n n a e , p ro no t u m a n d p ropec t u s redd i s h t e s t aceou s ; mesot h o r a x , m e t a t h o r a x a n d p ropodeum b l ack ; a b domen a n d l e g s redd i s h - t e s t aceou s ; h e a d d u l l , re t i c u l a t e r u gose ; occ i p i t a l c a r i n a com p l e t e ; pos t e r i o r oce l l i n o t tou c h i n g occ i p i t a l c a r i n a ; O P L s h o r t e r t h a n OL ; O L s h o r t e r t h a n P O L ; p rono t u m d u l l , h a i r y , g r a n u l a ted , w i t h n u merous s t r i ae a rou n d d i sc ; a n te r i or t r a n sv e rs a l i m p ress i o n a n d a n t e r i o r co l l a r w e a k ; pos t e r i o r t r a n s v e rs a l i m p ress i on s t ro n g a n d l oc a ted on pos t e r i o r m a r g i n of p ro n o t u m j pos t e r i o r co l l a r i n v i s i b l e j pra na t a l t u berc l e s not reach i n g

968

Mem . Amer . E n ! .

I ns! . ,

no.

37 ,

1 984

F i g . 658 - C h e l a of Tridryl'n u s gib b o s u s n . s p .

( type)

t e g u l ae ; scu t u m , s c u t e l l u m a n d m e t a n o t u m d u l l , re t i c u l a t e r u g ose ; n o t a u l i ces i n v i s i b l e ; p ropodeum ret i c u l a t e r u g o s e , w i t h 2 l o n g i t u d i n a l k ee l s o n pos t e r i o r s u r f ac e ; fore-w i n g w i t h 2 b rown t r a n s v e rs a l b a n d s a n d w i t h d i s t a l p a r t of r ad i a l v e i n l o n g e r t h a n p rox i m a l p a r t ( 1 3 : 1 1 ) ; s e g m e n t 1 of fron t t a rs u s more t h a n t w i ce as l o n g as seg m e n t 4 ( 2 1 : 8 ) ; seg m e n t 3 of fron t t a rs u s p roduced i n t o a hook ; en l a r ged c l aw ( F i g . 657 ) w i t h s u b a p i c a l too t h a n d 5 l a m e l l a e ; segmen t 5 of f ron t t a rs u s ( F i g . 657 ) w i t h 1 row o f 1 3 l ame l l ae ; a p e x w i t h a g r o u p o f 1 2 l a me l l ae ; t i b i a l s p u r s 1 , 1 , 2. MALE : u n k n o w n L O C U S T Y P I C U S : Yacoch u y a ( A rge n t i n a ) TYP I CA L MA T E R I AL : h o l o t y p e F ! i n T U ; 1 p a r a t y p e F ! i n G V ; 1 p a r a t y p e F! in OL . HOST S : u n k n o w n

969

D ryininae

D I ST R I B U T I O N : T U!

Ol!

Neotropic region :

Cordo b a ,

ARGEI�TINA:

Yacoc h u ya ( C afayate ,

Sa l ta ) ,

GV!

1 2 . Tridryin u s gil) b o s u s n . s p .

F E r.lAlE : fu l l y wing e d : l ength 4,00-6 , 2 5 mm; head r e d d is h -testac eo u s ; sometimes oce l l ar region brown; antennae b r o w n , with seg ments 1 -2 testac eo u s; p ro p ectu s b l ac k , with dorsal re g i on testaceou s; p ro p o d e u m b l ack w i th sides l ig ht o r fu l l y r e d dish; occasiona l l y black with margins and a median stripe l i g ht or r e d dish with a b rown spot on sides; mesothorax , metathorax an d p ro p o d e u m b l ack; ab d omen reddish-b rown; l e g s testac eo u s , with m i d and hind coxae part l y b l ac k ; c l u b s o f femora p artly brown; head d u l l , retic u l ate r u g o se; occip i ta l carina comp l ete; posterior oce l l i not to u c hing occipital carina; OPl s h orter than Ol; Ol sh orter than P O L; p ronotulTI d u l l , h a i ry, w i th n u m e ro u s weak striae around disc; anterior transversa l impre ssion and ante rior co l lar weak; p osterior transversa l impres s i on strong and l ocated on posterior m ar g i n of pronotu m; p osterior co l l ar invisib l e; scutu m d u l l , h a i ry , retic u l ate r u gose; notau l ices inv i sib l e; metanotum d u l l , very s h ort and r e d u c e d ; p ro p o d e u m retic u l ate r u g os e , with 2 l ong i t u dinal k ee l s on posterior s u rface; fore-wing with 2 b rown transve rsal bands and with distal p art of radia l vein s h orter th an p roxilTIal part ( 1 2 : 1 5 ) ; s e g m ent 1 of front tar s u s more than twice as l on g as s e g m ent 4 ( 28 : 1 2 ) ; s e g m ent 3 o f front tar s u s p ro d u ce d into a h o o k ; e n l arged c l aw ( Fig . 658 ) with s ubapical tooth and 5 l am el l ae; segment 5 of fore tarsi ( Fig . 6:'8 ) with 2 r o w s of 14 l am e l l ae; apex with a g r o u p of 1 5 l ame l l ae; tibial s p u rs 1 , 1,2. MAL E : unknown lOC U S TYP I C U S : St . A u g u stine ( T rini d ad ) paraty p e TYP I CAL MAT E R I AL : h o l otype F! in BI'-I; 1 para type F! i n Ltc; p aratype F ! in F ! in WA ; 1 p araty p e F ! in C A ; 1 paratype F! i n AM ; Ol. HOSTS: unknown D I ST R I B U T IO N : f,eotro pic re gion : TR I N I D AD : St . A u g u stine ( St . Georg e ) , 81\;i! COSTA R I '=A : E s p arta, WA! SUR I N AM E : Kre p i ( C h ar l esbu r g , Paramaribo ) , lE! PER U : Tingo Maria ( Monson Va l l ey ) , CAl BOL I V I A : At m o u th of Rio Baures ( Rio I tenez, D e pt . Beni ) , AM! Rio I tenez ( Oept. [leni ) , Ol!

4 . j\;egadryin u s Richards

Mega dryin u s Ric h a r d s 1 9 5 3 :

55 .

TYPE S P E C I E S : Megadryin u s magnificus Richards 1 9 5 3 ,

orig .

d esig .

970

Mem . Am e r . E n ! .

I n st . ,

no.

37,

1 984

F i g . 659 - Fema l e of /o,egadryinu5 magnificus R i ch a rds ( t y p e )

FEMALE ( F i g s 659 , 660 ) : f u l l y w i n ged ; m ax i l l a ry p a l p i w i t h 6 segmen t s ; l ab i a l p a l p i w i t h 3 segme n t s ; en l a rged c l a w ( F i g s 659 , 660 , 661 ) more t h a n t h ree t i mes as l o n g a s fro n t t i b i a , m u c h l o n g e r t h a n a ro l i u m , w i t hou t s U b a p i c a l too t h ; no t a u l i c e s i nv i s i b l e; pron o t a l t u b e rc l e s not reac h i n g t e g u lae ; occ i p i t a l c a r i n a corn p l e te; t i b i a l s p u rs 1 , 1 , 2 . MAL E : u nKnow n DIST R I BU T I OI;J : Neot rop i c HOS T S : Flatidae SP E C I ES: 1

D r y i n i n ae

971

F i g . 6 6 0 - F em a l e of Mega dryin u s m a gn ific u s R i c h a r d s ( t y p e )

N E O T R O P I C S P E C I ES OF MEGADRYINUS

O n e spec i es i s k no w n e

IY/ega dryin u s m a gn ificus R i c h a r d s

Megadryin u s m a gn ificus R i c h a r d s 1 95 3 : 55 .

F E MA L E ( F i g s 6 5 9 , 6 60 ) : f u l l y w i n g ed; l en g t h 1 0 , 93 m m ; h e a d b l ac k , w i t h f ro n t p a r t o f v e r t e x , c l y p e u s a n d m a n d i b l es y e l l ow-red d i s h ; a n t e n n a e w i t h s e g me n t s 1 -3 red d i s h - t e s t aceou s , 4- 1 0 b row n ; p ropec t u s b l ac k ; p ro n o t u m b l a c k , w i t h a p e x e s o f t e g u l a e a n d l a tera l m a r g i n s r e d d i s h ; s c u t u m ,

972

Mem . Amer . E n ! .

I ns t . ,

n o . 37,

1 984

F i g . 6 6 1 - C h e l a of Nega dryln u s magnlhcus R i c h a r d s ( t y p e )

s c u te l l u m , p ropode u m , mesop l e u r a , m e t a p l e u r a , mesos t e r n u m a n d m e t a s t e r n u m b l ack; pet i o l e b l ack; a b d omen b row n - b l ack; fore l eg s w i t h coxae a n d troc h a n ters p a r t l y b l a ck , femora a n d t i b i ae redd i sh , t a r s i b row n i s h , en l a r ged c l a w l i g h t; m i d a n d h i n d l e g s b row n -b l ack; te g u l ae

973

D ry i n i n ae

y e l l o w ; a n t e n n a l seg men t s in fo l l ow i n g p rop o r t i on s : 22: 6: 67 : 20: 15: 9 : 7 : 7 : 6 : 1 0 ; h e a d sh i n y , h a i r y , f u l l y re t i c u l a te ru gose ; occ i p i t a l c a r i n a comp l e t e ; OPL s h o r t e r t h a n b r ea d t h o f each oce l l u s ; p ro n o t u m sh i n y , c o n v ex , w i t ho u t t r a n sversa I i m p ress i o n ; p ron o t u m w i t h n u m ero u s weak l on g i t u d i n a l k ee l s , w i t h s i des w i t h ou t scu l p t u r e ; scu t u m h a i r y , f u l l y r e t i c u l a t e ru gose ; n o t a u l i c es i n v i s i b l e ; sc u t e l l u m a n d met a n o t u m ret i c u l a te r u gose ; p ro p o d e u m f u l l y ret i c u l a t e r u g o s e , w i t h 2 l o n g i t u d i n a l k ee l s on p o s t e r i o r s u rf a ce ; poste r i o r s u rf a ce of p ro n o t u m n e a r l y as l o n g as a n t e r i o r s u rf ac e ; fore-w i n g w i t h 2 b rown t r a n sversa l b a n d s a n d w i t h d i s t a l p a r t of r a d i a l v e i n l o n ger than p rox i m a l p a r t ; en l a r ged c l a w ( F i g . 6bl) n e a r l y t h ree t i mes a s l o n g a s fore t i b i a e , w i t h ou t s u b a p i c a l too th , s p a t u l a t e a t a p e x , w i t h 1 row of nea r l y 80 p eg - l i k e h a i r s ; seg men t 5 o f fro n t t a r s u s ( F i g . 6 6 1 ) w i t h 2 row s of near l y 1 70 l a me l l a e ; a pex w i t h a g ro u p of n e a r l y 26 l a m e l l ae ; seg m e n t 1 of f r o n t t a rsu s s h o r t e r t h a n seg men t 4 ( 1 8 : 28 ) ; 1 , 1 , 2.

seg m e n t s 2 a n d 3 of fro n t t a r s u s w i t h o u t hook s ;

t i b i a l s p u rs

MALE : u n k n o w n LOC US TYP I C U S : Ar i m a ( T r i n i d ad ) TYP I C AL ''i,AT EI''AL : h o l o t y p e F ! i n WA ; 1 p a r a t y p e F ! i n B M . HOSTS : i n T r i n i d a d Ormenis s p . (Fla tidae) ( R i c h a rd s 1953 ) D I STR I B U T I ON : N eo t rop i c reg i o n : TR I N I DA D : A r i m a , \VA ! S I . A u g u s t i n e ( R i c h a rd s 1 9 53 ) ;

D i ego M a r t i n ,

tl M !

5 . A lph a dryin us n .

gen .

T Y P E S P E C I E S : A lph a dryin u s a sia ticus n. s p . F E M A L E ( F i g s 662, 663 ) : f u l l y w i n ged ; rnax i l l a ry p a l p i w i t h 6 seg men t s ; l a b i a l p a l p i w i t h 3 segme n t s ; en l a r g e d c l a w ( F i g . 666 ) a s l o n g a s or s h o r t e r t h a n front t i b i a , much l o n g e r t h a n a ro l i u m , w i t h o u t s u b a p i ca l too t h ; n o t a u l i ces com p l e t e o r i ncomp l e t e ; pron o t a l t u berc l es reac h i n g t e g u l ae ; occas i o n a l l y n o t reach i n g t eg u l a e ; occ i p i t a l c a r i n a com p l e t e or i n com p l e te ; t i b i a l s p u rs 1 , 1 , 2 . MALE : u n k n o w n D I STR I B U T I ON : P a l a e a rc t i c ( on l y fos s i l spec i e s ) , Neot rop i c , A u s t r a l i a n HOSTS : u n k n o w n SPEC I ES : 1 4

E t h i op i a n ,

PALAEARCT I C S P E C I E S CJF A LPHADRYINUS

T w o foss i l

spec i e s a r e k n ow n .

Urien t a l ,

974

Mem . Am er . E n ! .

Fig.

662

I ns! . ,

no.

37,

1 984

F e m a l e of A lph a dryin u s lamellatus n . sp .

-

( ty p e )

�EY T O THE SP E C I E S

F EMAL E S

P r on o t u n1 h u mp ed , w i t h a d i s t i n c t d i s c a n d t w o t r a n s v e r s a l i mp r e s s i o n s ( F i g . 6 6 4 B ) ; an t e r i o r a n d p o s t e r i o r c o l l a r d i s t i n c t .

.

.

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D r y i n i n ae

975

i mp r e s s i on b e t w e e n a n t e r i o r and p o s t e r i o r c o l l a r . . . • . . . . . • • . . . . . . . • . . . • • . . 2 . b altic u s n . sp . •

(F i g .

665

A)

•

•

MALES

U n k nown

Fig.

663

-

F e rn a l e of A lphadryin u s l a m el l a t u s

n .

sp .

( t y pe )

1 . A lp h a dryin u s bruesi n . s p .

F E M A L E ( F i g . 664 ) : f u l l y w i n g e d ; l e n g t h 6 , 7 rn rn ; h e a d b row n - l i g h t , w i t h t e e t h of rn a n d i b l es b row n - d a rk a n d ternp l e s b l ac k ; a n t e n n a e b l a c k , w i t h segrnen t 1 t e s t aceou s ; p ro n o t u rn black , w i t h t r a n sv e r s a l i rn press i o n a p p a ren t l y t e s t aceou s ; sc u t u m b l ac k ; s c u te l l u m a p p a r e n t l y t e s t aceou s ; rn e t a n o t u rn i n v i s i b l e ; p ropodeurn b l ac k , w i t h po s t e r i o r s u rface a p p a ren t l y

•

976

M e m . Amer .

En!.

I ns! . ,

no.

37,

F i g . 664 - F e m a l e o f A lp h a dryin u s bruesi n . s p . B : p ron o t u m . '

1 984

( t y p e )( A ) ( p a r t l y v i s i b l e ) ;

t e s t aceo u s ; ven t r a l s i de of t h o r a x b l a c k ; a b domen b l ac k ; l e g s t e s t ac e o u s , w i t h cox a e , c l u b s of femora a n d p a r t l y t i b i ae b l a c k ; a n t e n n ae 1 0seg men t e d , shor t , weak l y s l e n d e r , d i s t a l l y w e a k l y t h i ck e n e d ; d e n s e l y h a i ry ; a n t e n n a l h a i r s v e r y s h or t , m u c h s h o r t e r t h a n brea d t h o f segmen t s ; a n t en n a l seg men t s i n fo l l o w i n g propor t i o n s : 1 1 : 6 : 32 : 1 9 : 1 5 : 1 5 : 7 : 7 : 8 : 1 0 ; a n t e n n a e l es s t h a n t h ree t i me s a s l o n g a s h e a d ( l en g t h o f h e a d dorsa l l y m e a s u red from occ i p i t a l c ar i n a beh i n d t h e oce l l i to a pex of m a n d i b l e s ) : 1 22 : 50 ; c l y p e u s and a n te n n a I soc k e t s n o t v i s i b l e ; m a n d i b l es w i t h 4 teeth p r o g ress i n g l a r g e r f ro m a n t e r i o r o n e to poster i o r ; eyes a p p a re n t l y b a re ; m a l ar s p ac e w i t h a su bocu l a r su l c u s ; occ i p i t a l c a r i n a comp l e t e ; tem p l e s p rom i n en t ; eyes norma l l y b u lg i n g ; P O L � 3 , 5 ; OL � 2 ;

Dryininae

DOL

=

8;

OPL

=

3,5;

TL

=

977

6; frontal line complete; vertex and frons dull,

granulated, rugose mostly on vertex and behind the ocelli; pronoturn

(Fig.

664 B) crossed by an anterior weak transversal impression between anterior collar and disc;

disc weakly humped;

pronotum also crossed '='y a

weak posterior transversal impression between disc and posterior collar; posterior collar very short, rnuch shorter than anterior collar ( 1 ,5:5); pronotum dull, granulated, slightly shorter than head (19:26); pronotal tubercles apparently reaching or almost reaching tegulae; sides of pronotum contracted where they meet the scutum;

posterior transversal

impression and posterior collar without a distinct dorsal rnedian carina; scutum shorter than pronotum (13: 1 9), du I I,

weak I y granu I ated;

notau Iices

complete, broad, posteriorly separated, composed of numerous strong punctures; scutellum dull, weakly granulated, shorter than scutum ( 1 0: 1 3), metanotum invisible; shape of scuturr" Dryininae;

scutellum and pronotum usual for

propodeum not easily distinct;

propodeum

(dorsally viewed)

invisible because hidden under the wings; laterally viewed visible, showing numerous areolae on sides and on posterior surface; posterior surface of propodeurn apparently with two incomplete longitudinal keels; posterior surface of propodeum very step; fore-wing apparently hyaline, without dark transversal bands, with the usual venation of Dryininae; pterostigma narrow, much longer than broad open;

(38:5); radial cell apparently

distal part of radial vein much longer than proximal part (22:1 0);

radial ve',n not s-shaped;

fore wings with the usual three oasal cells

clearly enclosed (costal, median and submedian cells); hind wings apparently hyaline, without dark bands, not easily distinct;

fore-wing

very slender, with shape usual for Dryininae; front leg segments in following proportions: 1 2 (coxae): 1 4 (trochanters): 22 (femora): 2 1 (tibiae): 8 (tarsal segment 1 ): 3 (tarsal segment 2): 5 (tarsal segment 3): 6 (tarsal segment 4): 23

(tarsal segment 5):

16 (enlarged claw); fore

trochanters less than four times as long as broad: 1 4: 4;

fore trochanters

with a long proximal slender stalk, broadened after the half-way; segments 2 and 3 of front tarsus produced into hooks; rudimentary cia" not distinct;

enlarged claw much longer than arolium (22:1 0); enlarsed

claw apparently without subapical tooth with some long lanlellae;

(but apex weakly distinct) and

apex of enlarged cia" not pointed, but spatulate;

segment 5 of front tarsus with some median lame! lae and

\II,. ith

nUfT'erOU5

apical lamellae; number of lamellae on enlarged claw and on tarsal segment 5 not distinct;

mid leg segments in following proportions: 8

(coxae): 5 (trochanters): 6

15 (femora):

1 4 (tibiae):

13 (tarsal segments 1 ):

(tarsal segment 2): 4,5 (tarsal segment 3): 3 (tarsal segment 4): 4,5

,tarsal segment 5);

hind leg segments in following proportions: 1 7 (coxae):

5 (trochanters): 33

(femora):

segment 2): 5):;

7

32 (tibiae):

1 8 (tarsal segment 1 ): 8 (tarsal

(tarsal segment 3): 3 (tarsal segment 4):

tibial spurs 1 , 1 ,2;

5 (tarsal segment

abdomen without a distinct petiole;

max'lllary

Jalpi apparently with 6 segments; labial palpi apparently with 3 segments; last labial segment very long and slender; shape, length and Jreadth of wings usual for Dryininae; shape and

978

Mem . Amer.

Ent .

I n st . ,

no.

37,

1 984

morp h o l o g y of body usua l for Dryininae. M/,LE: u n k n oV\n L OC U S T YP I CUS : German a m b e r T YP I CAL �iAT E f, I AL :

h o l o t y p e F!

in CO .

H OSTS : u n k n ow n o I STR I BUT I O N : on I y k n o w n from G e rm a n a rll b e r . N O T ES : t h i s spec i e s seems show morph olog i c c h a r a c t e r i st i cs of l i v i n g ,11phaclryinus. N o spec i es o f Alphaclryinus i s l i v i n g a c t ua l l y i n Pa l a e a rc t i c r eg i o n . T h e spec i es i s n amed i n h o n o r of the l a t e spec i al i st Mr . C . T . B rues .

2 . Alphadryinus balticus n . sp .

FEI\:ALE ( F i g . 665 ) : fu l l y w i n g ed ; l e n g t h 1 , 80 mOl ; a p p a re n tly b l ac k , w i t h a n t e n n a e a n d l e gs t e s t a ceous ; a bdomen a n d a p p a re n tly ve n tr a l side o f t h o r a x b r o w n - t est a ceousj a n t e n n a e lO-se g me n t ed, s h o r t , d i st a l l y th i c k e n ed ; dense l y h a i r y ; a n t e n n a l h a i rs short, sh o r t e r t h a n b re a d t h o f segme n t s; a n t e n n a l se g m e n ts i n f ol l ow i n g p ro p o r t i o ns: 7 : 5 : 5 : 4 : 3 , 5: 3 , 5 : 3 : 3 : 3 : 6 ; a n t en n ae l e ss t h a n tw i ce a s l o n g a s h e a d ( l e n g t h o f h e a d dorsa l l y measured f rom occ i p i t a l c a r i n a be h i n d t h e oce l l i to a p e x o f mand i b l es ) : 4 2 : 25 ; clyp eus a p p a re n tly w i t h ven t r a l m a r g i n b i den t a t e ; m a n d i b l es quadr i de n t a t e , w i th tee t h p ro g re ss i n g l a r g e r f rom a n te r i o r o n e to post e r i o r ; e y es a p p a re n t l y b a r e ; ma l a r space n o t v i s i b l e ; occ i p i t a l c a r i n a comp l ete ; temp l es p romi n e n t; e y es norma l l y b u l g i n g ; POL � 5; OL � 4 ; OOL � 7 ; O PL � 5 ; : TL � 6 ; f r o n s a p p a r e n t l y w i th out med i a n k ee l ; ve r t e x a n d f ro n s du l l , g r a n u l a t ed, a p p a re n tly n o t rugose ; p ro p odeum very sh o r t , w i t h o u t d i sc, n o t humped, b u t w i th a t r a nsve rsa l i m p ress i o n b e t w ee n a n te r i o r a n d post e r i o r co l l a r ; p r o n o t a l tuberc l es a p p a re n t l y r e a c h i n g t e g u l a e; p ro n o tum much s h o r t e r t h a n h e a d ( 5 : 2 5 ) ; scutum l o n g e r t h a n p ro n o tum ( 1 3 : 5 ) ; poste r i o r co l l a r o f p r o n o tum a p p a r e n t l y ver y short; scutum d u l l , g ra n u l a ted, a p p a re n t l y n o t rugose ; n o t au l i ces comp l e t e , poster i o r l y sep a r a t ed ; scut e l l um w e a k l y v i s i b l e , a p p a r e n t l y a s sc u l p t ured a s scutum ; met a n o t um not v i s i b l e ; p ropodeum o n l y l a te ra l l y v i s i b l e , a p p a r en t l y re t i cu l a t e rug ose , w i t h dorsa l surf ace n e a r l y a s l o n g as post e r i o r sur f a c e ( 1 0 : 1 2 ) ; post e r i or surf a c e n o t v i s i b l e f rom t n e dorsa l s i de ; sh a p e of scutum, scut e l lum a n d pro podeurn usua l f o r iJryininae; s h a pe of p r o n o turn u n u sua l ; fore-w i n g h y a l i n e , w i t h out dark t r a nsversa l b a nds, w i t h t h e usua l ven a t i o n of Dryininae; p t e rost i gma n a r row, much l o n g e r t h a n b road ( 1 0 : 4 ) ; r a d i a l c e l l not f u l l y v i s i b l e ; d i st a l p a r t of r ad i a l ve i n much l o n g e r t h a n p roxim a l p a r t ( but r ad i a l p a r t o n l y p a r t l y v i s i b l e) ; r a d i a l ve i n n o t s-sh a p e d ; f o re w i n g s w i t h t h e usua l t h re e b a s a l c e l l s c l e a r l y e n c l osed ( cost a l , med i a n a n d submed i a n c e l l s ) ; h i n d w i n g s a p p a rently h y a l i n e, w i t h o u t d a r k b a n ds ; f o re-w i n g s l ende r , w i t h sh a pe usua l f o r Dryininae ; f r o n t l e g segme n t s i n fo l l ow i n g p ro p o r t i o n s: 1 1 ( co x a e ) : 5 ( t r oc h a n t e rs ) : 2 1 ( femora ) : 1 6 ( t i b i ae ) : 1 0 ( t a rsa l se gment 1 ) :

D r y i n i n ae

979

A

:" i g .

665

-

Fe m a l e of Alph a dryin u s o a l ticus n . s p . ( t y p e )( B ) ; A : p ron o t u m .

2

head and

( t a r s a l s e g m e n t 2 ) : 2 ( t a rs a l s e g m e n t 3 ) : 1 3 ( t a r s a l s e g m e n t 4 ) : 1 5 : a r s a l seg m e n t 5 ) : 1 4 ( e n l a r g e d c l a w ) ; fore t roc h a n t ers more t h a n t w i c e 2S lo n g as b ro a d ( 5 : 2 ) , w i t h a short prox i m a l s t a l k ; e n l a r ged c l a w m u c h ""lger t h a n a ro l i u m ( 1 4 : 4 ) ; en l a r g e d c l a w s p a t u l a te , w i t h apex n o t =ci n ted ; a pex o f e n l a r g e d c l a w w i t h a row of t e e t h ( 5 t e e t h v i s i b l e ) a n d "itn n u m e r o u s b r i s t l es ; s e g m e n t 5 of f ro n t t a r s u s weak l y v i s i b l e , s:o=aren t l y w i t h some med i a l l a me l l ae ( 3 l a m e l l ae v i s i b le ) ; apex w i t h a ,;-cJp of l a m e l l ae ; r u d i me n t a r y c l a w d i s t i n c t ; s e g m e n t s 2 a n d 3 of fron t :,,- s u s p roduced i n to hook s ; m i d l e g s e g m e n t s i n fo l l ow i n g p rop o r t i on s : 6 coxae ) : 2 ( troc h a n t e r s ) : 1 3 ( femora ) : 1 7 ( t i b i ae ) : 7 ( t a r s a l s e g m e n t 1 ) : 3 :srsa l se g m e n t 2 ) : 3 ( t a r sa l s e g m e n t 3 ) : 2 , 5 ( t a r s a l s e g m e n t 4 ) : 5 ( t a r s a l "",,"len t 5 ) ; h i n d l e g seg me n t s i n fo l l ow i n g p roport i o n s : 9 ( cox a e ) : 4 :-xh a n t ers ) : 1 8 ( t i b i ae ) : 7 ( t a r s a l seg men t 1 ) : 4 ( t a r s a l s e g m e n t 2 ) : 4 :5-sa l se g m e n t 3 ) : 4 ( t a r s a l s e g m e n t 4 ) : 6 ( t a r s a l seg m e n t 5 ) ; t i b i a l "=_rs 1 , 1 , 2 ;

a bdomen w i t h o u t a d i s t i n c t pe t i o l e ;

m ax i l l a ry p a l p i

980

M e m . Amer . E n ! .

I ns t . ,

no.

37,

1 984

a p p a re n t l y w i t h 6 s e g me n t s ; l a b i a l p a l p i a p p aren t l y w i t h 3 s e g me n t s ; l a s t l a b i a l seg men t not v i s i b l e ; s h a p e , l e n g t h a n d b re a d t h o f t h e w i n g s u s u a l for Dryin in a e ; s h a pe a n d mor p h o l o g y o f bod y u su a l for Dryin in a e . MALE : u n k nown LOCUS TYP I C U S : Germ a n a m b e r TYP I CAL MAT E R I AL : h o l o t y p e F ! i n CO . HOSTS : u n k n o w n D I ST R I B U T I O N : on l y k n o w n f r o m Germ a n a m b e r . N O T E S : t h i s spec i e s i s d i ffere n t from o t h e r fos s i I a n d I i v i n g spec i es of A lph a dryin u s for t h e u n u su a l s h a pe of p ro n o t u m ( ex c a v a ted , n o t h u m pe d ) . T h· i s exce p t i o n , however, i s k n o w n i n o t h e r g e n e r a ( see Dryinus irregularis n . sp . ) a n d i s not con s i dered a gen e r i c c h a ra c t e r .

E TH I OP I AN S P E C I ES OF ALPHADRYLVUS

O n e spec i es i s k now n .

A lph a dryin u s a b erra n s ( Be n o i t ) n . com b .

Dryin u s a b erra n s Be n o i t 1 95 3 b : 4 1 2 .

F E MAL E : f u l l y w i n g e d ; l e n g t h 5 , 0 m m ; he a d b l ac k , w i t h c l y p e u s a n d m a n d i b l e s t e s t a ceou s ; a n te n n a e tes t aceou s ; p ropec t u s b l ac k ; prono t u m b l ac k , w i t h m a r g i n s a n d pos t e r i o r co l l a r redd i s h ; meso t h orax , m e t a t h o r a x a n d p ropo d e u m b l ac k ; a b domen b row n ; l e g s t e s t a c eo u s , w i t h cox a e a n d c l u b s o f femora b row n ; a n t e n n ae d i s t a l l y w e a k l y t h i cken e d ; a n t en n a l s e g m e n t s i n fo l l ow i n g p ropor t i on s : 9 : 4 : 2 5 : 1 3 : 9 : 5 , 5 : 5 : 5 : 4 , 5 : 6 ; h e a d d u l l , g r a n u l a t e d ; occ i p i t a l c a r i n a comp l e te ; pos t e r i o r oce l l i t o u c h i n g occ i p i t a l c a r i n a ; OL � POL ; p ro n o t u m d u l l , g r a n u l a t e d , w i t h a n t e r i o r i m p re s s i o n a n d a n t e r i o r co l l a r weak ; p o s te r i o r t r a n s v e rs a l i m p res s i o n s t ron g ; pos t e r i o r co l l a r s h or t , d i s t i n c t ; pronot a l t u b erc l e s n o t reach i n g te g u l a e ; scu t u m s h i n y , g r a n u l a t e d ; n o t a u l i ces i n c om p l e t e , reac h i n g a p p ro x i m a te l y 0 , 65 l e n g t h of sc u t u m ; s c u t e l l u m a n d m e t a n o t u m d u l l , g r a n u l a t e d ; propod e u m re t i c u l a te r u g o s e , w i t h o u t l o n g i t u d i n a l k ee l s on pos ter i o r su rface ; fore-w i n g w i t h 3 b rown t r a n s v e r s a l b a n d s and w i t h d i s t a l p a r t o f r a d i a l ve i n n e a r l y tw i c e a s l o n g a s p rox i m al p a r t ; en l arged c l a w ( F i g . 666 ) w i t h o u t s u b a p i c a l too t h a n d w i t h 5 l ame l l ae ; segmen t 5 of f ro n t tarsus ( F i g . 666 ) w i t h 2 rows of 1 7 l a m e l l ae ; apex w i t h a g roup of 9 l a me l l ae ; s e g m e n t 1 of f ro n t t a r s u s l o n g er t h a n s e g m e n t 4 ( 1 7 : 1 0 ) ; 3 of f ro n t t a rs u s p r o d u ced i n to a hook ; t i b i a l s p u rs 1 , 1 , 2 . MALE : u n k nown LOCUS TYP I C SU : Bek i l y ( M a d a g a s c a r )

seg men t

�81

Dry i n i nae

//��-----

I

y/

i��

//

/1

,

V

',,\

'"

�

/\

F i g . 666

-

Chela of Al,)hadryinus a:oerrans

(Benoit) (type)

M em . Amer .

982

T Y P I CAL MAT E R I AL : H O STS :

En!.

holotype F !

I ns ! . ,

no.

37,

1 984

in P .

u n know n

D I STR I B U T I ON :

E t h i o p i a n reg i o n :

MADAGASCAR ;

Bek i l y ,

P!

OR I E N TAL S P E C I E S O F !l � PIIADJ( YINU.C:

T w o spec i e s are know n .

KEY T O THE SPEC I ES

F EMALES

Oc c i p ap i ca Occ i p a p i ca

i t a l c a r i n a c omp l e l p r om i n en c e ( F i g . i t a l c a r i n a i n c omp l p r om i n en c e ( F i g .

t e ; s eg m e n t 5 o f f r on t t a r s u s w i t h ou t 6 6 7 ) . . . . . . 1 . a si a ticus n . s p . l e t e ; s e g me n t 5 o f f r on t t a r su s w i t h 6 6 8 ) . . . . . . 2 . gib b o s u s n . s p .

MALES

U n know n

1 . A lp h a dryin u s a s i a ticus

n.

sp .

F E MA L E : f u l l y w i n g ed : l e n g t h 3, 75-5 , 62 m m ; h e a d b l ack, w i t h c l y p e u s a n d m a n d i b l e s y e l l o w ; some t i mes a l so ma l a r s p a c e redd i s h ; a n t e n n a e bro w n , w i t h se g men t s 1 -2 y e l l ow ; p ropec t u s b l a c k ; p ro n o t u rn b l ack, w i t h a n t e r i or m a rg i n , s i des a n d pos t e r i o r col l a r redd i sh ; rnesot h o ra x , rne t a thorax , p ropod e u m a n d p e t i o l e b l ac k ; a b domen b rown ; l e g s tes t aceou s , w i t h cox a e a n d c l u b s o f f e m o r a b row n ; somet i mes a l so t i b i ae b row n ; h e a d d u l l , g ra n u l a t ed , w i t h 1 -4 l o n g i t u d i n a l kee l s on t h e s i des of t h e oce l l i , from occ i p i t a l c a r i n a to c l y p eu s ; some t i mes a l so i r reg u l a r short kee l s are v i s i b l e on fro n s ; occ i p i t a l c a r i n a comp l e t e ; a n t e n n a e shorter t h a n bod y , w i t h seg men t 3 l o n g er t h a n seg n len t 4 ; p ron o t u m h u m p e d , w i t h a s t ron g t ra n s v e r s a l p o s t e r i o r i mp re s s i o n a n d a weak a n t e r i o r t r a n sv e rs a l i rn p ress i on ; pos t e r i or co l l a r v i s i b l e ; a n ter i or co l l a r weak l y v i s i b l e ; p ronot u m sh i n y , w e ak l y g r a n u l a t e d , w i t h n u mero u s l o n g i t u d i n a l kee l s arou n cJ d i sc , sc u t u m d u l l , g r a n u l a te d , w i t h l a t e r a l reg i o n s ret i c u l a t e r u gose ; not a u l i ces i n comp l e t e , e x t en d i n g t o b e y o n d 0 , 5 l e n g t h of sc u t u m ;

Dry i n i n ae

F i g . 667 - C h e l a of A lph adryinus a si a ticus n . sp .

983

( ty p e )

s c u t e l l u m a n d m e t a n o t u m g r a n u l a t e d ; p ropod e u m re t i c u l a t e ru gose w i t h 2 l o n g i t u d i n a l kee l s on pos t e r i or s u rf ace ; fore-w i n g w i t h 3 b ro w n t r a n s v ers a l o a n d s ( t h e t w o b a s a l b a n d s j o i n ed ) ; d i s t a l p art o f rad i a l v e i n m u c h l o n ge r t h a n p ro x i m al p a r t ; e n l a r g e d c l a w ( F i g . 667 ) not s p a t u l a t e , w i t h o u t s u b a p i c a l too t h a n d l a m e l l ae , w i t h 1 prox i m a l b i g p e g - l i ke h a i r ; se g m e n t

984

Mem . A m er . E n t .

I n st . ,

no.

37,

1 984

F i g . 668 - C h e l a of A lp h a dryin u s gib b o s u s n . s p .

( type)

5 o f f ro n t t a rs u s ( F i g . 667 ) w i t h 3-4 row s o f a t l e a s t 4 5 l ame l l ae e x t en d i n g con t i n uo u s l y to t h e a pex ; en l a rged c l a w m u c h l o n g e r t h a n a ro l i u m ; seg m e n t 1 of fron t t a r s u s s l i g h t l y l o n ger t h a n segmen t 4 ; t i b i a l spurs 1 , 1 , 2 .

Dry j n j nae

t/IAL E :

985

unknown

L O C U S TYP I C U S :

Kal abakan

T Y P I C A L �'IA T E R I A L : H OS T S :

( Sabah ,

ho l o t y p e F !

Ma l a y s i a )

in 8;

1

p a r a t y p e F" !

D I ST R I BU T I ON : V I E T I�AM :

Oriental

8 i an

(Mt.

reg i o n :

La n g ) ,

f',\A LAYS I A :

Kal abakan

F e MA L E :

f u l l y w i n g ed ;

l e n g t h 5 , 0 m rn ;

( m ore a l o n g o rb i t s ) ,

red d i s h ;

p ro p ec t u s b l ac k ;

( Tawau,

Saoah ) ,

8!

OL !

2 . A l�h a dryin u s gi 0 bo s u s n .

v er tex

i n UL .

unknown

sp .

h e a d b l a ck ,

w i t h f r o n t p a r t of

c l y pe u s and ma n d i b l es redd i sh ;

p ro n o t u m b l a c k ,

w i th anterior and

a n t e n n ae l a tera l

m a r g i n s re d d i s h ; pos t e r i o r co l l a r redd i s h ; mesoth o r a x , met a t h o r a x a n d p ro p o d e u m b l a c k , a b domen b r o w n - r e d d i s h ; l e g s t e s t a ceo u s , w i t h c l u b s o f femora a n d h i n d c o x a e b r ow n ; h e a d s w o l l e n , sh i n y , w e a k l y p u n c t a t e ; oc i p i t a l c a r i n a i n comp l e t e ; p o s t e r i or oce l I i n o t t o u c h i n g occ i p i t a l ca r i n a ; U ? L ::: P O L ;

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66SJ - C h e l a of A lph a dryin u s lamellatus n . sp .

( type)

a ro u n d co l l a rs ; p ronot a l t u b erc l e s n o t reach i n g t e g u l a e ; sc u t u m d u l l , g r a n u l a t ed a n d w e a k l y re t i c u l a t e ru gose ; n o t a u l i ces com p l e t e , s t ron g l y con verg i n g , pos t e r i o r l y sep a r a ted ; s c u te l l u m a n d m e t a n o t u m sh i n y , w ea k l y p u nc t a t e ; propode u m ret i c u l a te ru gose, w i t h 2 l on g i t u d i n a l k ee l s on pos t e r i o r s u rf a c e ; fore-w i n g w i t h 2 b ro w n t r a n s vers a l b a n d s and w i t h d i s t a l p a r t of rad i a l v e i n much l o n g e r t h a n p rox i m a l p a r t ( 28 : 1 4 ) ; seg m e n t 1 o f f ro n t t a r s u s shorter t h a n seg men t 4 ( 1 7 : 20 ) ; segme n t s 2 a n d 3

988

Mem . Amer . E n ! .

Fig.

I ns ! . ,

no.

37,

1 984

670 - C h e l a of A lp h a dryin u s p a n amensis

n .

sp.

( ty p e )

p ro d u c e d i n to hook s ; en l arged c l a w ( F i g . 669 ) w i t h o u t s u b a p i c a l too t h a n d w i t h 1 u n u s u a l v e r y l o n g l a me l l a + 4 s h o r t e r l a me l l ae ; segmen t 5 o f f ron t t a r s u s ( F i g . 669 ) w i t h 1 row of 1 1 l a me l l a e ; a pex w i t h a g roup of 1 5 l ame l l ae ; t i b i a l s p u rs 1 , 1 , 2 . MALE : u n k n o w n

Dry i n i nae

989

LOCUS TYP I C U S : Nova Teu ton i a ( B raz i l ) TYP I CA L MA T E R I AL : h o l o t y p e F ! a n d 1 p a r a t y p e F ! i n OL. HOSTS: u n k nown D I ST R I B U T I O N : Neot rop i c reg i o n : BM ! OL !

BRAZ I L :

i n B fYI ;

1

paratype F !

Nova Teuton i a ( S . t a C a t a r i na ) ,

2 . A lp h a dryi n u s p a n amensis n . s p .

F E M A L E : f u l l y w i n g e d ; l e n g t h 6 , 93 m m ; h e a d b l ac k , w i t h fron t p a r t of ver tex , c l y p e u s , m a n d i b l e s a n d orb i t s y e l l o w ; occ i p u t b row n -y e l l ow ; a n t e n n a e w i t h se g m e n t s 2-5 b row n , seg m e n t 1 p a r t l y y e l l o w , seg men t 6 w i t h d i s t a l p a r t y e l l o w ; seg men t s 7- 1 0 m i s s i n g i n h o l o t y p e ( o n l y k n o w n spec i men ) ; t h o r a x , p ropodeum a n d a b domen b l ac k ; l e g s t e s t a c eo u s j h e a d d u l l , f l a t , w i t h oce l l a r reg ion sw o l l e n , g r a n u l a te d a n d ret i c u l a t e r u g o se, h a i ry ; occ i p i t a l c a r i n a i n comp l e t e , v i s i b l e on l y beh i n d the oce l l i and on t h e s i des of v e r tex ; pos t e r i o r oce l l i not t o u c h i n g occ i p i t a l c a r i n a , O P L shorter t h a n P O L ; O L nea r l y t h ree t i mes a s l o n g as P U L ; p rono t u m sh i n y , h a i r y , w i t h n u m erou s l o n g i t u d i n a l k ee l s on co l l a r a n d w i t h n u ro erous k ee l s a ro u n d d i sc ; a n t e r i o r t r a n s versa l i m p ress ion s t r o n g ; a n ter ior co l l a r d i s t i n c t ; p os t e r i o r t r a n sver s a l i m p ress i o n s t ro n g a n d n e a r pos t e r i or m a r g i n o f pronot u m j pos t e r i o r co ! l a r i n v i s i b l e ; p ranot a l t u b erc l es not reach i n g t e g u l a e ; sc u t u m d u " , h a i r y , ret i c u l a t e ru gose ; not a u l i ces i n comp l e t e , reach i n g 0 , 3 3 l e n g t h o f sc u t u m ; sc u t e l l u m d u l l a n d ret i c u l a te r u g ose ; m e t a n o t u m sh i n y , smoo t h , w i t h o u t scu l p t u re , e x c e p t for some p u n c t u res l oc a t e d n e a r sc u te l l u m ; p ropodeum ret i c u l a t e r u gose , w i t h 2 l o n g i t u d i n a l k ee l s o n po s t e r i o r s u rface ; fore- w i n g w i t h 2 b ro w n t r a n s versa ! b a n d s a n d " i t h d i s t a l p a r t o f ra d i a l v e i n m u c h l o n g e r t h a n prox i m a l p a r t ( 25 : 1 1 ) ; r a d i a l ce l l broade l y open ; seg men t 1 of f ron t t a r s u s n e a r l y as l o n g as seg men t 4; se g m e n t s 2 and 3 of front tarsus p ro d u c e d i n to hook s ; en l a r g e d c l a w ( F i g . 670 ) w i t h o u t s u b a p i c a l too t h a n d w i t h 1 :' l a m e l l ae ; s e g rnen t 5 o f f ron t t a r s u s ( F i g . 670 ) w i t h 1 r o w of 1 0 l a m e l l ae ; a pex w i t h a g ro u p of a t l e a s t 40 l a me l l ae ; t i b i a l s p u r s 1 , 1 , 2 . MAL E : u n k n o w n L O C U S TYP I C U S : G a m boa ( P a n a m a ) TYP I C A L lviA T E R I A L : h o l o t y p e F ! i n LE . .., OSTS : u n k n o w n U I ST R I B U T I O N : Neot rop i c reg ion : PANAMA :

G a nlboa ( P i pe l i n e ['(o a d ) ,

LE !

3 . A l;,h a dryin u s parvus n . s p .

c :: MAL E : f u l l y w i n g ed ; l e n g t h 3 , 75 m m ; t e s t a ceou s , " i t h p e t i o l e b l ack ; a n t e n n ae d i s t a l l y t h i c k e n e d ; a n t e n n a i se g m e n t s i n fo l l ow i n g p roport i o n s : 8 :

990

IVlem . Amer . E n ! .

F i g . 67 1

I ns! . ,

no.

37,

1 984

- C h e l a of A l p h a dryin us p arvus n . sp .

( type )

4 , 5 : 1 9 : 1 1 : 1 0 : 9 : 7 : 6 : 6 : 7 , 5 ; h e a d d u l l , g r a n u l a te d ; frons w i t h o u t a med i a n k ee l ; occ i p i t a l c a r i n a i n com p l e t e , v i s i b l e o n l y on dors a i s i d e of o cc i p u t ; occ i p i t a l c a r i n a n o t v i s i b l e on temp l e s ; P U L � 3 ; OL � 4 ; O O L 5 , 5 ; O P L � 1 ; temp l e s i n v i s i b l e ; pro n o t u m d u l l , crossed b y a n a n t e r i o r t r a n s ve rs a l i m p ress i o n a n d a pos t e r i o r t r a n s v e rs a l i m p ress i o n ; d i sc h u m p e d ; p ro n o t a l t u b erc l e s n o t reac h i n g t eg u l a e ; pos t e i o r c o l l a r very s h or t ; pro n o t u m and scu t u m d u l l , g r a n u l a t e d ; n o t a u l i c e s i n com p l e t e , reac h i n g 0 , 3 3 l e n g t h o f sc u t u m ; sc u te l l url1 a n d m e t a n o t u m sh i n y , w e a k l y g ra n u l a t ed ; p ropodeum ret i c u l a te r u g ose ; pos t e r i o r s u rface w i t h t w o l o n g i t u d i n a l k ee l s ; fore- w i n g w i t h t w o d a r k t r a n s v e rs a l b a n d s , one o n t h e b a s a l c e l l s a n d a n o t h er b e n e a t h t h e p t e ros t i g m a ; d i s t a l p a r t of ra d i a l v e i n l o n ge r t h a n prox i m a l p a r t ( 1 3 : 8 ) ; fore t a rsa l segmen t s i n fo l l o w i n g p roport i o n s : 1 2 : 2 : 3 , 5 : 1 3 : 20 ; en l a r g e d c l a w ( F i g . 6 7 1 ) w i t h o u t s u b a p i c a l toot h , w i t h 1 r o w of 8 l a me l l ae , a m o n g w h i c h 1 l o n g a p i c a l l a me l l a ; seg m e n t 5 o f f ro n t t a rs u s ( F i g . 6 7 1 ) w i t h 2 row s o f 1 2 l a me l l ae ; a p ex w i t h a g roup of 1 0 l am e l l a e ; t i b i a l s p u rs 1 , 1 , 2 . I� ALE : u n k n o w n L O C U S TYP I C U S : C a r u a r u ( B r az i l ) TYP I CA L MAT E R I A L : ho l o t y p e F ! i n T W . H O ST S : u n k n o w n D I ST R I B UT l m, : Neot rop i c reg i o n : BRAZ I L :

C a r u a ru ,

TW!

D r y i n i n ae

991

F i g . 672 - Ch e l a of A l p h a dryin u s pegn a i n . s p .

4 . A lp h a dryin u s pegn a i n .

F E MALE :

f u l l y w i n g ed ;

l e n g t h 3 , 75-4 , 50 m m ;

( type)

sp .

h e a d b l ac k ,

w i t h fron t p a r t of

992

Mem . Amer . E n ! .

I ns! . ,

no.

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;YPE S P [ C I C: � , : Mesodryin u s n iger ( , ; ; pffpl' 1 904 ) , de s i g . by " i effer' i n K i effer e t M a rsh a l l 1 906 . F E M A L E ( F i g s 678, 679 ) : f u l l y w i n g e d ; m ax i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segme n t s ; e n l arged c l a w ( F i g . 683 ) as l o n g as f ron t t i b i a or shor t e r , m u c h l o n g er t h a n a ro l i u m , w i t hou t s u b a p i c a l too t h ; n o t a u l i ces a b s e n t ; pronot a l t u b e rc l e s n o t reac h i n g te gu l a e ; occ i p i t a l c a r i n a com p l e t e o r i n comp l e te ; t i b i a l s p u rs 1 , 1 , 2 .

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F i g . 6 7 9 - Fema l e o f Mesodryin us a e th iop icus

n .

sp.

( type)

( F i g s 680, 6 8 1 ) : f u l l y w i n ged ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; p a l p i w i t h 3 s e g m e n t s ; o t h e r g e n e r i c m o r p h o l o g i c c h a r a c t er i s t i c s : see fa m i I y d i a g n os i s . D I ST R I B U T I O N : P a l aearc t i c , E t h i o p i a n , O r i e n t a l , N e a rc t i c , I�eo t r'op i c HOSTS : Dictyop h aridae SPEC I ES : 1 5 fVI A L E

labial

PALA E A R C T I C S P E C I E S U F NESODl2 Y1NUS

T w o spec i es a r e k n ow n .

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MALES

P o s t e r i o r o c e l l i n o t t ou c h i n g o c c i p i t a l c a r i n a ; h ea d w i t h a r e o l a e a s w i d e a s a r e o l a e o f s c u t u m ; a n t en n a l s e g men t 3 s l i g h t l y l o n g e r t h a n s eg men t 4 , l e s s t h a n t h r e e t i me s a s l o n g a s 686 ) s eg me n t 2 ; g on o f o r c e p s s l i g h t l y s h o r t e r t h a n p e n i s ( F i g . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 . n iger ( K i e f f e r ) - P o s t e r i o r o c e l l i t ou c h i n g o c c i p i t a l c a r i n a ; a r e o l a e o f h e ad mu c h s ma l l e r t h a n a r e o l a e o f s c u t u m ; a n t e n n a l s e g m e n t 3 a l mo s t t w i c e a s l on g a s s eg m e n t 4 , a l mo s t f ou r t i me s a s l on g a s s eg m e n t 2 ; g on o f o r c ep s mu c h s h o r t e r t h a n p en i s ( F i g . 6 8 4 ) .

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1 . Mesodryin u s dayi n .

( p ara t y p e )

sp .

F E MA L E : f u l l y w i n ge d ; l e n g t h 3 , 87-4 , 37 m m ; head b l ack , w i t h m a n d i b l e s t e s t a ceo u s ; some t i mes w i t h a red d i s h s t r i pe a rou n d o rb i t s ; a n te n n ae b l ac k , w i t h se g m e n t s 1 -2 o r 1 -4 test aceou s : t h or a x , propodeum a n d a b domen b l ac k ; l e g s test aceo u s , w i t h cox a e , c l u b s of femora a n d somet i mes a l so t i b i ae p a r t l y b l ac k ; head sh i n y , or d u l l , s wo l l e n , g r an u l a t e d , w i t h n u merous more or l e ss s t r o n g l on g i t u d i n a l k ee l s ; a n ten n a l segmen ts i n fo l l ow i n g propor t i o n s : 1 0 : 5 : 2 1 : 1 3 : 1 2 : 1 0 : 8 : 8 : 7 : 9 ; occ i p i t a l c a r i n a comp l e te ; pos t e r i o r oce l l i n o t t o u c h i n g occ i p i t a l c a r i n a ; P O L = 6 ; O l = 4 ; OOl = 9 ; O P l = 1 ; pron o t u m ( F i g . 682 A ) s h i n y , w i t h col l a r smoo t h a n d w i t h d i sc ru gose , w i t h l o n g i t u d i n a l a n d t r a n s vers a l kee l s ; d i sc very h u m pe d ; a n t e r i o r t r a n s v ers a l i m p ress i o n s t ro n g ; a n te r i or co l l a r d i s t i n c t ; poste r i o r t r a n s vers a l i mp r es s i on s t ron g ; poster i or co l l a r i n v i s i b l e ; sc u t u m re t i c u l a te r u g os e ; not a u l i ces a b s en t ; sc u te l l u m a n d m e t a n o t u m ret i c u l a t e r u g os e ; p ropodeum ret i c u l a t e ru gose , w i t h 2 l on g i t u d i n a l k ee l s on poster i or s u rface ; fore- w i n g w i t h 2 b ro w n t r a n s v e rs a l b a n d s a n d w i t h d i s t a l p a r t of rad i a l v e i n n e a r l y as l o n g as prox i m a l p a r t ; fore t a rsa l segme n t s i n fo l l ow i n g p roport i on s : 1 4 : 3 : 5 : 8 : 1 6 ; segmen t 3 of fro n t t a r s u s produced

1 004

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I n s! . ,

no.

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1 984

A

F i g . 682

-

P r o n o t u m of fema l e s of Mesodryi n u s dayi n . s p . n iger ( K i ef fe r ) ( t y p e ) ( B )

i n to a h ook ; e n l a r ged c l a w ( F i g . 683 )

w i th subap i c a l

( ty pe ) ( A )

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tooth a n d 2 a p i c a l

s ma l l peg- l i k e l a me l l ae , i n a d d i t i o n t o 5 p rox i m a l b r i s t l es ; segmen t 5 of f ro n t t a rs u s ( F i g . 683 ) w i t h some rows of n e a r l y 34 l a me l l ae e x t e n d i n g con t i n u ou s l y t o t h e a pex ; t i b i a l s p u rs 1 , 1 , 2 . MALE ( F i g s 6 8 0 , 6 8 1 ) : f u l l y w i n ged ; l e n g t h 4 mm ; b l ack ; m a n d i b l es , c l y p e u s , a n t e r i o r reg i on o f frons ( a l so a l o n g o rb i t s ) testaceo u s ; a n t e n n a e f u l l y test aceo u s ; t e g u l ae t e s t aceo u s ; l eg s t e s t aceous, w i t h c o x a e p a rt l y b l ack , c l u b s of femora b row n ; h i n d t i b i a e b row n ; a n te n n a I seg m e n t s i n f o l l o w i n g p ro p o r t i o n s : 6 : 6 : 2 2 : 1 3 : 1 5 : 1 5 : 1 3 : 1 2 : 1 0 : 1 2 ; h e a d f u l l y ret i c u l a t e r u g ose ; a reo l ae of h e a d sma l l er t h a n a reo l ae of scu t u m ; frons w i t h o u t l o n g i t u d i n a l k ee l s ; occi p i t a l c a r i n a i n comp l e t e ; poster i o r oce l l i t o u c h i n g occ i p i t a l c a r i n a ; sc u t u m f u l l y re t i c u l a t e ru gose, d u l l ; n o t a u l i ces w e a k l y d i s t i n c t , a l most i n v i s i b l e ; sc u t e l l u m , m e t a n o t u m and p ro p o d e u m f u l l y re t i c u l a te ru gose ; fore-w i n g h y a l i ne , w i t h o u t d a rk t ra n s v e r s a l b a n d s ; d i s t a l p a r t of r ad i a l v e i n nea r l y as l o n g as p rox i m a l p a r t ( 9 : 1 0 ) ; gonoforceps ( F i g . 684 ) m u c h shorter t h a n pen i s ; t i b i a I s p u rs 1 , 1 , 2 . LOCUS TYP I C U S : Ka l a m b ak a ( G reece ) T Y P I CA L MAT E R I AL : h o l o t y p e F ! a n d 1 p a r a t y p e M ! i n BM ; 1 p a r a t y p e F ! i n MD . H OS T S : u n k n o w n D I ST R I BU T I ON : P a l a e a rc t i c reg i o n : S PA I N : M a d r i d , M D ! G R E E C E : K a l a m b a k a ( T hess a l i a ) , BM ! T U R K E Y : D i k m e n ( A n k a r a ) , BM ! NOTE :

t h e s p ec i es i s n am e d i n honor of M r . M i c h a e l D a y .

D r y i n i n ae

F i g . 683 - C he l a of Mesodryin us d a y i n . sp .

1 00 5

( type )

2 . Mesodryinus n iger ( K i effer )

Dryin u s n iger K i effer 1 904a : 3 5 2 . Mesodryin u s n iger ( K i effer ) : K i effer i n K i effer e t M a r s h a l l 1 906 : 497 . Mesodryin u s britta nicus R i c h a rds 1 93 9 : 228 ( s y n . proposed b y R i c h a r d s 1 95 3 ) ; loc u s typicus G l a n v i l l es Woo t t o n ( E n g l a n d ) . Mesodryin u s niger ( K i effer ) : R i c h a r d s 1 9 53 : 5 7 . Mesodryin u s n iger ( Kjeffer ) : C u r r a d o a n d O l m i 1 979b : 2 .

F E MAL E : f u l l y w i n ged ; l e n g t h 4 , 00-4 , 37 m m ; head b l ac k , w i t h c l y p eus a n d m a n d i b l es y e l l o w ; a n te n n a e test aceou s ; t horax a n d p ropodeum b l ac k ; abdomen b row n ; l e g s testaceo u s , w i t h c l u b s of femora a n d h i n d femora

1 006

Mem . Amer .

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I ns! . ,

no.

37,

1 984

F i g . 684 - Ma l e g en i t a l i a of Mesodryin u s dayi n . sp . ( p a r a t y pe ) from D i rk me n ( T u rk e y ) ( r i g h t h a l f removed )

b row n ; d i s t a l p a r t of m i d a n d h i n a t i b i ae b row n ; a n te n n a l segmen t s i n fo l l o w i n g p roport i o n s : 9 : 5 : 2 3 : 1 1 : 1 0 : 9 : 7 : 7 : 6 : 7 ; h e a d g ra n u l a t e d , d u I I , w i t h n u me ro u s l o n g i t u d i n a I w e a k s t r i ae o n f r o n s a n d vertex ; occ i p i t a l c a r i n a i n com p l e te , v i s i b l e o n l y b eh i n d t h e oce l l i ; poste r i o r oce l l i touch i n g occ i p i t a l c a r i n a ; p r o n o t u m ( F i g . 682 B ) d u l l , h a i r y , g ra n u l a t e d , w i th d i sc w e a k l y h u m p e d ; p ron o t u m w i t h a s t ro n g t r a n sver s a l pos t e r i o r i "' press i o n a n d a w e a k a n t e r i o r t r a n sversa I i m p ress i o n ; p o s t e r i o r co i l a r v e r y short a n d reduced ; p ro n o t a I t u b erc l es n o t reach i n g t e g u I a e ; sc u t u m d u l l , fu l l y ret i c u l a te ru gose ; n o t a u l i ces a b s e n t ; s c u te l l u m a n d m e t a n o t u rn d u l l , ret i c u l a t e rugose ; p ropodeum ret i c u l a te ru gose ; fore-w i n g w i t h 2 b row n t r a n s v e rs a l b a n d s , w i t h d i s t a l p a r t of r a d i a l v e i n s l i g h t l y l o n g er

1 00 7

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685 - C he l a of Mesodryin u s n iger ( K i ef fe r ) ( t y p e )

t h a n prox i m a l p a r t ; s e g m e n t 1 of f ro n t t a rs u s l on g e r t h a n segmen t 4 ( 1 4 : 1 1 ) ; seg m e n t s 2 a n d 3 of f ro n t t a rs u s p ro d u ced i n to hooks ; e n l a r ged c l a w ( F i g . 68 5 ) w i t h o u t s u b a p i c a l t oo t h , w i t h 6 l am e l l ae , segment 5 o f fro n t t a r s u s ( F i g . 685 ) w i t h 2 row s o f 1 0 l a me l l ae n e a r a pex ; t i b i a l s p u rs 1 , 1 ,2. MAL E : f u l l y w i n ged ; l e n g t h 4 , 37 m m ; b l ac k , w i t h m a n d i b l es b ro w n -red d i s h , a n ten n ae b row n , l e g s b ro w n - b l ack , w i t h fore t i b i a e <mel fore t a r s i y e l l ow ; a n ten n a l s e g m e n t s i n fo l l o w i n g p ropor t i on s : 5 : 5 : 1 4 : 1 2 : 1 1 : 1 1 : 1 0 : 9 : 9 : 1 0 ; head d u l l , f u l l y re t i c u l a te r u gose ; occ i p i t a l c a r i n a i n comp l e t e ; O P L shorter t h a n O L ; sc u t u m f u l l y ret i cu l a te ru gose ; n o t a u l i ces a b s e n t ; s c u te l l u m , m e t a n o t u m a n d p ropodeum ret i c u l a te r u g ose ; fore -w i n g w i t h o u t d a r k t r a n svers a l b a n d s , h y a l i n e , w i t h d i s t a l p a r t of r a d i a l v e i n a s l on g a s prox i m a l p a r t ; g en i t a l : f i g . 686 ; t i b i a l s p u rs 1 , 1 , 2 . LOCUS T Y P I C U S : V a l l o d i L u c a n i a ( I t a l y ) T Y P I CA L MAT E R I A L : h o l o t y p e F ! o f D . n iger i n GE ; ho l o t y p e F ! o f M. britt a n i c u s in OX . HOST S : u n k n o w n D I ST R I BU T I O N : P a l a e a rc t i c reg i o n : SW E D E N : E k sh a r a d ( Vm l d . ) , OT ! YUGOSLA V I A : P res p a Ge u l ( O tesevo , M a kedon i j a ) , B M ! E N G L AN D : G l a n v i l l es Woo t to n ( Dorset ) ( R i c h a r d s 1 93 9 ) ; A y no ( No r t h a n t s ) , B M ! I TALY : Geno v a , GE ! V a l l o d i L u c a n i a ( S a l er no ) , GE ! C Y P R U S : C h erkes , BM !

E T H I O P I AN S P E C I ES OF MESODR YINUS

O n e spec i e s i s k n ow n .

I 1 008

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M a l e gen i t a l i a of Mesodryinus n iger ( K i effer ) from Genova ( I t a l y ) ( ri ght h a I f removed )

Mesodryinus a ethiop i c u s n . sp .

F E M A L E ( F i g s 678 , 679 ) : f u l l y w i n ged ; l en g t h 5 , 3 1 -6 , 9 3 m m ; b l ac k , w i t h c l y p e u s a n d m a n d i b l es ferr u g i no u s ; propec t u s b l ack ; p ro n o t u m b l ac k , w i t h s i des a n d poster i o r m ar g i n redd i s h ; mesothor a x , m e t a thorax a n d propodeum b l ac k , a b domen b l ac k - b r ow n , some t i mes w i t h apex redd i s h ; l e g s testaceo u s , w i t h coxae a n d c l u b s o f femora b l ac k or b ro w n ; a n te n n a e d i s t a l l y weak l y t h i c k e ne d ; a n t e n n a l segme n t s i n fo l l ow i n g p roport i o n s : 1 6 : 6 : 2 7 : 1 9 : 1 4 : 1 1 : 9 : 9 : 8 : 1 0 ; h e a d s h i n y , fu l l y ret i c u l a te ru gose ; occ i p i t a l c a r i n a comp l e te ; O P L s horter t h a n O L ; O L shorter t h a n PO L ; p ro n o t u m d u l l , g r a n u l a ted , n o t h u mped , w i t h a weak med i a n t r a n sversa l i mp res s i on ;

I 1 009

D ry i n i n ae

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687 - C he l a of Mesodryin us a e th iopicus n .

sp.

( t y pe )

pos te r i o r t r a n s v e rs a l i m p r es s i on we a k l y v i s i b l e ; poste r i o r co l l a r very s h or t , weak l y v i s i b l e ; p rono t a l t u berc l es not rea c h i n g t e g u l ae , s c u t u m d u l l , g r a n u l a t e d a n d ret i c u l a te r u gose ; not a u l i ces a b s en t ; 2 l o n g i t u d i n a l k ee l s , reach i n g a p p r ox i m a te l y 0 , 5 l e n g t h of sc u t u m , a re s i m i l a r to not a u l i ces ( b u t t h e y a re kee l s , not f u r rows ) ; s c u te l l u n d u l l , g r a n u l a te d a n d p a r t l y re t i c u l a te ru gose ; m e t a n o t u m f u l l y re t i c u l a t e r u gose ; p ropodeum ret i c u l a te r u g os e , w i t h 2 l o n g l o n g i t u d i n a l kee l s on poste r i o r s u rface ; fore-w i n g w i t h 2 b ro w n t r n s vers a l b a n d s a n d w i t h d i s t a l p a r t of ra d i a l v e i n l o n g e r t h a n p rox i m a l p a r t ( 2 0 : 1 3 ) ; en l a rged c l a w ( F i g . 687 ) w i t h o u t s u b a p i c a l too t h a n d w i t h 3 l a me l l ae ; segmen t 5 o f fore t a r s i ( F i g . 687 ) w i t h 2 rows of 42 l a me l l ae exte n d i n g con t i n u ous l y to t h e a pex ; segme n t 3 of frorjt t a rs u s p rod uced i n to a hook ; segme n t 1 of f ro n t t a rs u s nea r l y a s l o n g a s seg m e n t 4 ; fore t roc h a n ters n e a r l y t w i ce a s l o n g a s b road ; t i b i a l s p u rs 1 , 1 , 2 . M A L E : f u l l y w i n ged ; l e n g h t 4 , 37 m m ; head b l ack , w i t h c l y p e u s a n d m a n d i b l es tes t a ceo u s ; a n t e n n a e t e s t aceo u s ; t h o r a x a n d p ropodeum b l ack ; a b domen b row n ; l e g s t e s t a c eo u s , w i t h coxae a n d c l u b s of femor a b l ac k ; a n t e n n a e not d i s t a l l y t h i c k e n e d ; a n t e n n a I s e g m e n t s i n fo l l ow i n g p ropor t i o n s : 9 : 6 : 20 : 1 3 : 1 3 : 1 4 : 1 2 : 1 1 : 9 : 1 2 ; h e a d s h i n y , r e t i c u l a te

1010

Fig.

Mem . Amer .

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no .

37 ,

1 984

M a l e ge n i t a l i a of fviesodryin u s a e th iopic u s n . s p . from Boukoko ( Ce n t r a l Afr i c a n R e p u b l i c ) ( r i g h t h a l f removed )

r u gose ; occ i p i t a l c a r i n a i n comp l e t e ; pos t e r i o r oce l l i c a r i n a ; O P L shorter t h a n O L ; O L shorter t h a n POL ; sc u t u m s h i n y , ret i c u l a t e ru gose ; not a u l i ces a b sen t ; g r a n u l a ted , w i t h some a reo l ae ; m e t a n o t u m re t i c u l a te

not touch i n g occ i p i t a l temp l es very s hor t ; s c u te l l u m d u l l , ru gose ; p ropode u m

re t i c u l a te r u gose , w i t ho u t l o n g i t u d i n a l kee l s on poste r i o r s u rf a ce ; fore w i n g w i t h o u t d a r k t r a n s v e rs a l b a n d s and w i t h d i s t a l p a r t of r a d i a l v e i n l o n ger t h a n p ro x i m a l p a r t ( 1 7 : 9 ) ; d i s t i vo l se l l a ( F i g . 688 ) l oc a t e d very n e a r a p e x of pen i s .

D ry i n i n ae

101 1

LOCUS TYP I C U S : Nj a l a ( S i e rr a Leo n e ) TYP I CA L MAT E R I AL : h o l o t y p e F ! a n d 1 p a r a t y p e F ! i n BM ; 2 p a r a t y p es ( 1 F , 1 M ) ! i n P ; 1 para type F ! i n TW. HOST S : i n Cen t r a l Afr i c a n Rep u b l i c Dic tyop h a ridae D I ST R I B U T I O N : E t h i o p i a n reg i on : S I E R R A L E O N E : Nj a l a , B M ! F reetow n , T W ! C E NT R A L A F R I CA N R E P UBL I C :

Boukoko ,

P!

OR l E N T A L SP E C I E S OF MESODR YINUS

F o u r spec i e s a re k n ow n .

K E Y T O T H E SP E C I E S

FEMALES

Fo re-wi ng hya l i n e ,

t ra n s v e r s a l b a n d s . . . . . . . . . . . . . . . . 1 . orien t a lis n . sp . F o r e -w i n g w i t h 1 - 2 d a rk t ra n s v e r s a l b a n d s 2 H e a d w e a k l y r e t i c u l a t e r u g o s e ; a r e o l a e o f h e a d sma l l ; s c u t e l l u m g r a n u I a t ed , w i t h wea k a r e o I a e n ea r I a t e ra I ma rg i n s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 . p e terseni n . sp . H e a d s t r o n g l y r e t i c u l a t e ru g o s e , ; a r e o l a e of h ea d w i d e ; s ome t i me s a r e o l a e on l y on f r on s , b u t i n t h i s c a s e v e r t ex w i t h s t r o n g l on g i t u d i n a l k e e l s ; s c u t e l l u m f u l l y s t r o n g l y r e t i c u l a t e rug o s e 3 P o s t e r i o r o c e l l i t o u c h i n g o c c i p i t a l c a r i n a ; en l a rg ed c l a w w i t h ou t l a me l l a e ( F i g . 6 9 2 ) . . . . . . . 3 . viet n . sp . P o s t e r i o r o c e l l i n o t t o u c h i n g o c c i p i t a l c a r i n a ; e n l a rg ed c l a w w i t h l ame l l a e ( F i g . 6 9 3 ) . . . . . . . . . . 4 . b a rbarus n . s p . .

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MALES

U nknown

1 . Mesodryin u s orien t a l i s n .

sp.

F E MAL E : f u l l y w i n ged ; l e n g t h 6 , 25 m m ; b l ac k ; m an d i b l es a n d p a r t l y c l y pe u s test aceou s ; a n te n n a e w i t h segmen t 1 tes t a ceo u s ( o t h e r segme n t s

1012

Mem . Amer . E n ! .

I n s t . , n o . 37 ,

1 984

A

B

c Fig.

689 - P ro n o t u m of fem a l es of Mesodryin u s b arb a ru s n . sp . ( t y p e ) ( A ) , orienta lis n . sp . ( t y pe ) ( B ) a n d petersen i n . s p . ( ty p e ) ( C ) .

m i ss i n g i n ho l o t y pe ) ; te g u l ae a n d l eg s testaceous , w i t h h i n d coxae prox i ma l l y b l ack ; head s h i n y , s t ron g l y ret i c u l ate ru gose , w i t h areo l ae w i de ; frons w i t h a med i a n l on g i t u d i n a l kee l ; occ i p i t a l ca r i n a comp l e t e ; occ i p u t smoo t h , sh i n y , w i t h o u t scu l p t u re ; P O L ; 7 ; Ol ; 4 ; OOl ; 1 0 ; poster i o r oce l l i touc h i n g occ i p i t a l car i n a ( OP l ; 0 ) ; temp l es i n v i s i b l e ( T l ; 0 ) ; p ronotum ( F i g . 689 B ) a l most f l a t , sh i n y , smoot h , mos t l y on s i de s , w i t h w e a k tra nsversa l kee l s arou n d d i sc ; p ronot a l t u berc l es not reac h i n g teg u l ae ; sc u t u m ret i c u l ate ru gose , w i t h a med i a n reg i o n not ret i cu l ate ru gose , gra n u l a ted ; notau l i ces absen t ; scute l l um ret i c u l ate ru gose , w i t h a med i a n re g i o n gran u l a t e d , not ret i c u l a te ru gose ; metanotum ret i c u l a te r u gose ; propodeum ret i c u l a te ru gose , w i t h poster i o r su rface stron g l y

Dry i n i nae

Fig.

690

-

C he l a of Mesodryin u s orienta lis n .

1013

sp .

( type)

1 01 4

M e m . A m er .

F i g . 691

-

En!.

I ns ! . ,

n o . 3 7 , 1 984

C he l a of jYjesodryin u s p e terseni n . sp .

( ty p e )

D r y i n i n ae

1 01 5

t r a n s v erse l y s t r i a te a n d w i t h 2 l on g i t u d i n a l k ee l s ; fore-w i n g h y a l i n e , w i t h o u t d a rk t r a n s v ersa I a b n d s ; d i s t a I p a r t o f r a d i a I v e i n m u c h l on ger t h a n p ro x i m a l part ( 1 8 : 1 3 ) ; fore t a rsa l segmen t s in f o l l ow i n g p ropor t i on s :

1 1 : 2 : 4 , 5 : 1 1 : 1 9 ; en l a r ged c l a w ( F i g . 690 ) sp a t u l a t e , w i t h o u t S U b a p i c a l t oo t h , w i t h a ser i es o f 6 s m a l l teet h a n d 4 l ame l l a e ; segme n t 5 o f fron t t a rs u s ( F i g . 690 ) w i t h 2 rows of n ea r l y 38 l a m e l l a e ; segme n t 3 of fron t t a rs u s p r o d u ced i n to a hook ; t i b i a l s p u rs 1 , 1 , 2 . MALE : u n k n o w n L O C U S T Y P I C US : Loh-c h a n g D i s t r i c t ( C h i n a ) TYP I CAL MAT E R I A L : h o l o t y p e F ! i n B . HOST S : u n k n o w n D I ST R I B U T I O N : O r i e n t a l reg i o n : C H I NA : Loh-c h a n g D i s t r i c t ( N K w a n g t u n g ) , B!

2 . Niesodryin u s petersen i n . sp .

F E �lA L E : f u l l y w i n ged : l e n g t h 4 , 1 6-4 , 37 m m ; head b l ack , w i t h m a n d i b l es y e l l ow or b row n ; a n te n n a e b row n , w i t h seg m e n t s 1 - 2 a n d some t i mes p a r t l y 3 y e l l ow ; t h o r a x a n d p ro p o d e u m b l ac k ; a b domen b row n ; l eg s t e s t aceou s , w i t h coxae a n d c l u b s o f femora b ro w n ; head d u l l , g r a n u l a t ed , w i t h n u mero u s w e a k s t r i ae on f ro n s ; o c c i p i t a l c a r i n a i n comp l e te , on l y v i s i b l e beh i n d t h e oce l l i ; poster i o r o c e l l i touch i n g occ i p i t a l c a r i n a ; p rono t u m ( F i g . 6 8 9 C ) d u l l , g ra n u l a ted , w i t h n u merous weak s t r i ae a ro u n d d i sc ; p ron o t u m w i t h d i sc h u m p e d ; p o s t e r i or co l l a r very s h o r t ; a n t e r i o r c o l l a r V i s i b l e ; prono t a l t u berc l es n o t reach i n g teg u l a e ; scu t u m s h i n y , w i t h l a t era l reg i o n s re t i c u l a t e r u g ose ; poster i or h a l f of med i an reg i o n ret i c u l a t e r u g ose ; a n te r i o r h a l f g r a n u l a t e d ; n o t a u l i ces a b se n t ( i n s tead of t h e not a u l i ces t here a re 2 s h o r t kee l s , n o t f u r rows ) ; scu te l l u m d u l l , g r a n u l a t e d ; m e t a n o t u m ret i c u l a te r u g ose ; propodeum ret i c u l a t e r u gose o n a n t e r i o r h a l f ; poster i or h a l f n o t ret i c u l a t e ru gose , w i t h n u merous t r a n s v e rs a l kee l s and 2 l o n g i t u d i n a l kee l s ; med i a n a rea s h i n y ; fore- w i n g w i t h 2 b ro w n t r a n s v e rs a l b a n d s , w i t h d i s t a l p a r t o f r a d i a l v e i n l o n g e r t h a n p r o x i m a l p a r t ( 1 5 : 8 ) ; seg m e n t 1 of f ro n t t a rs u s s l i g h t l y l o n g e r t h a n segmen t 4 ( 1 2 : 1 0 ) en l a rged c l a w ( F i g . 6 9 1 ) s p a t u l a t e , w i t h o u t s u b a p i c a l too t h , w i t h 9 l a me l l a e a n d a s e r i e s o f 1 2 s m a l l t ee t h ; segme n t 5 o f f ro n t t a rs u s ( F i g . 6 9 1 ) w i t h 2 rows of 24 l a m e l l ae exten d i n g co n t i n u o u s l y to t h e apex ; t i b i a l sp u rs 1 , 1 , 2 . MALE : u n k n o w n L O C U S T Y P I C U S : N of B a t u B a t u ( P h i l i p p i nes ) TYP I CAL MAT E R I AL : h o l o t y p e F ! i n CO ; 1 p a r a t y p e F ! i n B . HOST S : u n k n ow n D I S T R I B UT I O N : O r i e n t a l reg i o n : H O N G K O N G : T a i pok a u , B ! P H I L I P P I NE S : N of B a t u B a t u ( T a r a w a k a n , T a w i T a w i ) , CO ! NOT E : t h e spec i es i s n a med i n honor of M r . Borge P e terse n .

1016

Mem . A m e r .

Ent.

I ns ! . ,

n o . 37 ,

1 984

F i g . 692 - C h e l a of Mesodryinus viet n . s p .

( type)

3 . Mesodryinus viet n . sp .

F E M AL E : f u l l y w i n g ed ; l e n g t h 5 m m ; b l a ck ; m a n d i b l es tes t aceo u s ; a n t e n n ae b ro w n , w i t h segm e n t s 1 -3 a n d 1 0 t e s t aceo u s ; l eg s t e s t aceou s , w i t h cox a e , c l ub s of femora a n d t i b i ae p a r t l y b row n ; a n t e n n a e d i s t a l l y w e a k l y t h i c k en e d ; a n te n n a l seg m e n t s i n fo l l ow i n g p roport i on s : 1 3 : 7 : 2 5 : 1 2 : 1 1 : 9 : 8 : 7 : 7 : 9 ; head f u l l y ret i c u l a t e ru gose , s h i n y , fro n s w i t h a med i a n k ee l , w i t ho u t l a t e r a l kee l s ; occ i p i t a l c a r i n a comp l et e ; P O L = 4 ; O l = 3 ; OOl = 1 0 ; pos t e r i o r oce l l i touch i n g occ i p i t a I c a r i n a ; temp l es i n v i s i b I e ; p rono t u m d u l l , g r a n u l a t e d , w i t h n umerous w e a k i r re g u l a r kee l s o r areo l ae o n t h e s i de s of d i sc ; p ro n o t u m w i t h a s t o n g t r a n s ve rs a l i mpress i o n ; pos t e r i o r s u rface of p ro n o t u m h u m p e d ; p ro n o t a I t u b erc l es n o t rea c h i n g

Dry i n i nae

1017

/

Fig.

693 - C h e l a of Mesodryin u s b a rbarus n . sp .

( type)

t e g u l a e ; sc u t u m , s c u t e l l u m a n d m e t a n o t u m f u l l y ret i c u l a t e r u g o s e ; sc u t u m w i t h a n t e r i o r s u rface n e a r p ro n o t u m smoo t h a n d w i t h o u t scu l p t u r e ; n o t a u l i ces i n v i s i b l e ; propodeum ret i c u l a t e r u gose ; poste r i o r s u rface o f p ropod e u m w i t h 2 l o n g i t u d i n a l k ee l s ; fore-w i n g w i t h 1 d a r k t r a n s v e rs a l

1018

Mem . Amer .

En!.

I n s t . , n o . 37 ,

1 984

b a n d b e n e a t h t h e p t e ros t i g m a ; d i s t a l p a r t of ra d i a l v e i n l o n g e r t h a n p rox i m a l p a r t ( 1 7 : 1 1 ) ; fore t a rsa l s e g m e n t s i n fol l o w i n g p roport i on s : 1 5 : 3 : 6 : 9 : 1 9 ; e n l a r g e d c l a w ( F i g . 692 ) s p a t u l a te ; , w i t h o u t s u b a p i c a l too t h w i t h a ser i e s o f 3 sm a l l t e e t h a n d 8 b r i s t I es ; s e g m e n t 5 of f ro n t t a r s u s ( F i g . 692 ) w i t h S o m e row s of n ea r l y 42 l ame l l ae ; segmen t 3 o f f ro n t t a rs u s p roduced i n to a hoo k ; t i b i a l s p u rs 1 , 1 , 2 . M A L E : u n k no w n LOCUS TYP I C U S : D a i L a n h ( V i e t Nam ) T Y P I C A L MAT E R I A L : ho l o t y p e F ! i n B . HOST S : u n k n o w n D I S T R I BU T I O N : O r i e n t a l re g i on : V I E T N A M : D a i L a n h ( N of N h a T r a n g ) , B !

4 . Mesodryin u s b a rbarus n . s p .

F E MA L E : f u l l y w i n ged ; l e n g t h 5 , 6 2- 8 , 7 5 m m ; b l ac k ; m a n d i b l e s p a r t l y b rown and test aceo u s ; a n te n n a e b row n , w i t h segmen t s 1 -2 a n d somet i mes p a r t l y 3 test aceo u s ; p ro n o t u m w i t h m a r g i n s test aceou s ; l eg s test aceous or somet i mes w i t h h i n d cox a e , m i d and h i n d c l u b s of femora , m i d and h i n d t i b i ae p a r t l y b ro w n ; a n te n n ae not d i s t a l l y t h i c kened ; a n t e n n a I segmen t s i n fo l l ow i n g propor t i o n s : 1 8 : 6 : 5 5 : 30 : 2 2 : 1 4 : 1 2 : 1 0 : 1 0 : 1 2 ; h e a d s h i n y , s t ron g l y ret i c u l a t e ru gose , m a i n l y on frons ; vertex w i t h n u merous s t ron g l o n g i t u d i n a l kee l s ; frons w i t h a med i a n kee l ; occ i p i t a l c a r i n a comp l e te ; POL = 4 ; OL = 4 ; OOL = 1 1 ; pos t e r i or oce l l i not touch i n g occi p i t a l ca r i n a ; p ro n o t u m ( F i g . 689 A ) s h i n y , smoo t h , weak l y g ra n u l ated , w i t h d i sc w e a k l y h u m pe d , w i t h some areo l ae n e a r pos t e r i or m a rg i n ; p ron ota l t u b e rc l es not reac h i n g te g u l a e ; scutum f u l l y ret i c u l a t e r u g os e w i t h a n a n t e r i o r reg i o n n e a r prono t u m smoo t h a n d w i t h o u t scu l p t u r e ; no t a u l i ces a b se n t ; sc u te l l u m a n d m e t a n o t u m f u l l y ret i c u l a t e ru gose ; p ropodeum ret i c u l a t e ru gose , w i t h 2 l o n g i t u d i n a l kee l s on pos t e r i o r su rface ; fore- w i n g w i t h 1 d a r k t r a n s v e r s a l b a n d b e n e a t h the p t e ros t i g m a ; d i s t a l p a r t of ra d i a l v e i n n e a r l y as l o n g a s prox i m a l p a r t ( 23 : 23 ) ; fore tarsa l segmen t s i n fo l l ow i n g p roport i on s : 1 8 : 3 : 6 : 1 3 : 26 ; e n l arged c l a w ( F i g . 693 ) s p a t u l a t e , w i t h o u t s u b a p i c a l toot h , w i t h 4 a p i c a l l ame l l ae a n d some b r i s t l e s ; s e g me n t 5 of fro n t t a r s u s

( F i g . 693 ) w i t h 2 r o w s of 32 l a me l l ae e x t e n d i n g con t i n uou s l y to t h e a p e x ; t i b i a l s p u rs 1 , 1 , 2 . MALE : u n k n o w n L O C U S T Y P I C U S : W a p i k h a m t ho n g ( L aos ) TYP I CAL MAT E R I A L : ho l o t y p e F ! a n d 1 p a r a t y p e F ! i n B ; 1 p a r a t y pe F ! i n OL . HOSTS : u n k n o w n D I ST R I B U T I O N : O r i e n t a l reg i on : LAOS : W a p i k h a m t h o n g ( W a p i ) , B ! B a n V a n Eue ( V i e n t i ane ) , B! O L !

N E A R C T I C S P E C I E S O F ME::ODl2YINU::

D ry i n i nae

1019

F o u r spec i e s a re k n ow n .

KEY T O T H E S P E C I E S

F E MA L E S

S c u t u m g ra n u l a t ed , n o t r e t i c u l a t e ru g o s e ; b o d y mo s t l y 3 . solaris n . sp . t e s t a c e o u s - r ed d i s h S c u t u m f u l l y r e t i c u l a t e ru g o s e ; b od y mo s t l y b l a ck E n l a rg ed c l a w w i t h t w o s u b a p i ca l t e e t h ( F i g . 6 9 8 ) ; o c c i p i t a l ca r i n a l on g , a l mo s t r e a c h i n g ey e s 4 . fa vre a u a e n . sp . E n l a r g ed c l a w w i t h ou t s u b a p i c a l t e e t h ( F i g s 6 9 4 , 6 9 6 ) ; o c c i p i t a l c a r i n a s h o r t , v i s i b l e on l y n e a r p o s t e r i o r o c e l l i Rad i a l c e l l w i de ( F i g . 6 9 5 A ) 2 . dors a li s n . sp . R a d i a l c e l l n a r r ow ( F i g . 6 9 5 B ) . . . . . . . . . . . l . incon s u l t u s n . sp . .

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MALES

U n k no w n

1 . l'lesodryinus incon s u l t u s n .

sp.

F E MA L E : f u l l y w i n ged : l e n g t h 4 , 37-4 , 50 m m ; h e a d red d i s h - t e s t aceous , w i t h a w i d e b l ack spot on vertex a n d w i t h t w o b l ac k spots on s i des of occ i p u t ; a n t e n n a e test aceou s ; t hor a x , propodeum a n d a b domen b l ac k , except for s i des of prono t u m test aceo u s ; l eg s t e s t aceo u s , w i t h h i n d cox a e b asa l l y b l ack ; a n t e n n a e d i s t a l l y w e a k l y t h i ck e n e d ; a n t e n n a I segme n t s i n fo l l o w i n g propor t i o n s : 8 : 5 : 27 : 1 5 : 1 3 : 9 : 7 : 6 : 6 : 7 ; h e a d d u l l , f u l l y g r a n u l a t e d ; fro n s w i t h a med i a n com p l e t e l o n g i t u d i n a l k ee l ; occ i p i t a l c a r i n a i ncomp l e t e , v i s i b l e on l y be h i n d t he pos t e r i o r oce l l i ; POL = 4 ; O L = 4 , 5 ; OOL = 9 ; pos t e r i or oce l l i touc h i n g occ i p i t a l c a r i n a ; p ronot u m d u l l , g ra n u l a t e d a n d w i t h n u merous a reo l ae n e a r p o s t e r i o r l obes ; p rono t a l t u berc l es not reac h i n g teg u l a e ; p ro n o t u m w i t h a w e a k t r a n s v e rs a l a n t e r i or i m p ress i o n b e t ween a n t e r i o r co l l a r a n d d i sc ; pos ter i or co l l a r i n v i s i b l e ; d i sc l i t t l e con v e x ; s c u t u m d u l l , f u l l y ret i c u l a t e r u g o s e ; n o t a u l i ces a b s en t ; s c u te l l u m d u l l , g r a n u l a ted ; m e t a n o t u m d u l l , g r a n u l a ted ; p ropodeum ret i c u l a te r u g ose , d u l l , w i t h t w o i n comp l e t e l o n g i t u d i n a l k ee l s on pos ter i or s u rface ; fore- w i n g w i t h t w o d a r k t r a n s vers a l b a n d s , one on t h e b a s a l c e l l s a n d a n o t h e r ben e a t h t he p t e ros t i g m a ; d i s t a l p a rt of ra d i a l ve i n l on g e r t h a n p ro x i m a l p a r t ( 9 : 5 ) ( F i g . 695 B ) ; fore t a r s a l seg m e n t s i n

1 020

Mem . Amer . E n t .

I ns t . , no. 37,

1 984

F i g . 694 - C he l a of Mesodryinus inconsultus n . s p .

( type)

fo l l ow i n g proport i on s : 1 4 : 3 : 9 : 1 6 : 30 ; e n l a rged c l a w ( F i g . 694 ) w i t h o u t subap i c a l too t h a n d w i t h 1 1 l ame l l ae ; seg m e n t 5 of fro n t tarsus ( F i g . 694 ) w i t h 1 row of 1 5 l a m e l l ae exte n d i n g con t i n uo u s l y to t he apex ; segmen ts 2 a n d 3 of f ro n t tarsus prod uced i n to hooks ; t i b i a l s p u rs 1 , 1 , 2 . MALE : u n k no w n

D r y i n i n ae

1 02 1

' :::;A

"

� -. ' :; : ;-: Fig.

B �== - - - - -

695 - R a d i a l ve i n s of fem a l es of Mesodryin u s dors a li s ( t y p e ) ( A ) a n d incon s u l t u s n . sp . ( t y p e ) ( B ) .

n . sp .

LOCUS T Y P I C U S : N a tchez T r ace S . P . ( Le x i n g ton , T e n n essee , U . S . A . ) T Y P I CA L M A T E R I AL : ho l o t y p e F ! i n O T ; 1 p a r a t y p e F ! i n T W . H O ST S : u n k n o w n D I S T R I BU T I O N : Neo t rop i c reg i o n : U . S . A . : N a tchez T race S . P . ( Le x i n g ton , T e n n . ) , O T ! Moore s to w n ( N . J . ) , T W !

2 . Mesodryin u s dorsa li s n . s p .

F E MA L E : f u l l y w i n g e d ; l e n g t h 3 , 60 m m ; head b l ac k , w i t h c l y pe u s , m a n d i b l es a n d a n t e r i o r re g i on of frons tes taceou s ; a n t e n n a e test aceou s ; t ho r a x b l ac k , w i t h s i des of pronot u m test aceo u s ; p ropod e u m a n d a b domen b l a ck ; l eg s testaceo u s , w i t h prox i m a l h a l f of h i n d coxae b l ac k ; a n t e n n ae d i s t a l l y t h i c k e n e d ; a n ten n a l seg m e n t s i n fol l o w i n g p ropor t i o n s : 1 3 : 6 : 30 : 1 5 : 1 4 : 1 0 : 9 : 8 : 8 : 1 0 ; head f l a t , d u l l , g ra n u l a ted , w i t h n u m erous i rreg u l a r s t r i a e ; frons w i t h a comp l e te med i a n k ee l ; occ i p i t a l c a r i n a i ncomp l e te , v i s i b l e on l y n e a r pos t e r i o r oce l l i ; POL = 6 ; O L = 6 ; OOL 9 ', O P L = 0 ; TL = 0 ; pronot u m d u l l , w i t h a w e a k a n te r i or t r a n s v e rs a l i m p re ss i on , g r a n u l a ted a n d w i t h n u merous areo l ae on s i des of d i sc ; prono t a l tu berc l e s not reach i n g teg u l a e ; scu t u m , s c u te l l u m a n d m e t a n o t u m d u l l , f u l l y ret i c u l a t e r u g ose ; not a u l i ces a b s e n t ; p ropodeum ret i c u l a te r u gose , d u l l , w i t h o u t t r a n s versa l or l on g i t u d i n a l kee l s ; fore-w i n g w i t h two d a r k t r a n svers a l b a n d s , one on t h e b a s a l c e l l s a n d another benea t h t h e p te ros t i g m a ; ra d i a l ce l l open ; d i s t a l p a r t of r a d i a l v e i n l on g e r t h a n p ro x i m a l p a r t ( 1 5 : 1 3 ) ( F i g . 6 9 5 A ) ; fore t a r s a l segmen t s i n fo l l ow i n g propo r t i o n s : 1 8 : 3 : 6 : 1 5 : 27 ; e n l arged c l a w ( F i g . 696 ) w i t hou t s u b a p i c a l tee t h , w i t h 2 row s o f 8 l a m e l l a e ; segmen t 5 o f f ro n t t a rs u s ( F i g . 696 ) w i t h two row s of 1 8 l a me l l ae exte n d i n g con t i n uo u s l y to t h e a pe x ; t i b i a l s p u rs 1 , 1 ,2.

1 022

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1 984

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( type)

D r y i n i n ae

F i g . 697

-

1 02 3

C h e l a of Mesodryin us solaris n . sp .

MAL E : u n k n o w n L O C U S TYP I C U S : So u t h H a v e n ( N ew Yor k , T Y P I CAL MAT E R I AL : ho l o t y p e F ! i n A M . HOST S : u n k n o w n

U.

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( ty p e )

1 024

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I n s t . , n o . 37 ,

1 984

S. A . : So u t h H a v e n

( Long

I s l an d ,

N.

3 . Mesodryin u s solaris n . s p .

F E MALE : f u l l y w i n ged ; l e n g t h 4 , 68-5 , 62 m m ; red d i s h-test aceou s , w i t h p et i o l e b l ac k ; abdomen dorsa l l y darkened ; a n te n n ae w i t h l a s t t ree s e g me n t s d a r k e ne d ; a s ma l l b l ack spot i s v i s i b l e b e h i n d t h e t e g u l ae ; a n t e n n ae d i s t a l l y t h i c k e n e d ; an t en n a l seg m e n t s i n fo l l ow i n g p ropor t i o n s : 1 1 : 5 : 2 2 : 1 2 : 1 0 : 8 : 6 : 6 : 5 : 8 ; head s h i n y , smoo t h , f u l l y g r a n u l a te d ; frons w i t h a very w e a k i ncom p l e t e l o n g i t u d i n a l med i a n k ee l , v i s i b l e on l y b et ween t he a n t e n n a I soc k e t s ; occ i p i t a l c a r i n a i ncom p l e te , weak l y v i s i b l e b e h i n d t h e p o s t e r i o r oce l l i a n d on t h e s i des o f oce l l i near eyes ; POL = 5 ; O L = 3 ; OOL = 8 , 5 ; poste r i or oce l l i touc h i n g occ i p i t a l ca r i n a ; p ro n o t u m d u l l , f u l l y g r a n u l a ted , w i t h a s t r o n g a n t e r i o r t r a n sversa l i m p ress i o n ; a n t er i o r co l l a r d i s t i nc t ; d i sc h u m ped ; poste r i or c o l l a r i n v i s i b l e ; pronot a l t u be rc l es not reac h i n g t e g u l a e ; s c u t u m d u l l , f u l l y g r a n u l a ted ; not a u l i ces abse n t ; sc u te l l u m and m e t a n o t u m s h i n y , w e a k l y gra n u l a ted ; p ropod e u m ret i c u l a t e ru gose ; w i t h poster i o r s u rface w e a k l y r u gose ; pos t e r i or surface w i t h 2 comp l e t e l o n g i t u d i n a l k ee l s ; med i a n area s h i n y and a l most smoo t h ; fore-w i n g w i t h 3 d a r k t r a n s v ersa I b a n d s on t h e b a s a I c e l l s a n d b e n e a t h t he P teros t i g m a ; d i s t a l p a r t o f rad i a l v e i n l o n g e r t h a n p rox i m a l p a r t ( 1 8 : 5 ) ; fore t a rs a l seg m e n t s i n fo l l ow i n g proport i o n s : 1 8 : 3 : 5 : 1 5 : 27 ; segmen t 3 of f ro n t t a r s u s produ ced i n to a hoo k ; e n l arged c l a w ( F i g . 697 ) w i t ho u t s u b a p i c a l too t h a n d w i t h 1 2 l a me l l ae ; seg m e n t 5 of f ro n t t a r s u s ( F i g . 697 ) w i t h 2 row s of near l y 37 l ame l l ae exte n d i n g con t i n uous l y to t h e a p e x ; t i b i a l s p u rs 1 , 1 , 2 . M AL E : u n k no w n L O C U S T Y P I C U S : T a rpon S p r i n g s ( F l or i d a , U . S . A . ) T Y P I CAL MAT E R I AL : h o l o t y p e F ! i n a T ; 1 p a r a t y p e F ! i n O L . HOST S : u n k n o w n D I S T R I B U T I O N : Nearc t i c re g i o n : U . S . A . : T a rpon S p r i n g s ( F l a . ) , O T ! P u n t a Gorda ( F l a . ) , O L !

4 . Mesodryin u s favre a u a e n .

sp .

F E M A L E : f u l l y w i n ged ; l e n g t h 4 , 40 m m ; h e a d b l ac k , w i t h c l y pe u s , m a n d i b l e s , m a l a r s p ace , t e m p l e s a n d fro n t p a rt o f frons test aceou s ; a n t e n n a e test aceous ; t hor a x , propode u m a n d abdomen b l ac k ; l eg s brow n , w i t h t i b i ae a n d t a r s i l i g t h ; a n t e n n ae d i s t a l l y t h i c k e n e d ; a n t e n n a I segmen ts i n fo l l o w i n g p roport i o n s : 1 3 : 7 : 62 : 2 5 : 1 8 : 1 1 : 9 : 7 : 7 : 1 0 ; head f l a t , s h i n y , gra n u l a te d , w i t h n umerous w e a k tracks o f i r reg u l a r s t r i ae ; frons w i t h a comp l e t e med i a n k ee l ;

occ i p i t a l car i n a i ncom p l e t e ,

l a tera l l y

Dry i n i nae

F i g . 698

-

C he l a of Mesodryin u s fa vrea u a e n . sp .

1 02 5

( type)

1 02 6

Mem . Amer .

F i g . 699

-

En!.

I nst . ,

no. 37,

1 984

C he l a of Mesodryin u s foresta lis n . sp .

( type)

reac h i n g a l most pos t e r i o r e d g es of e y e s ; P O L � 7 ; Ol � 4 ; OOl � 1 2 ; OPl 1 ; T L � 4; prono t u m d u I I , crossed b y a transversa I i m p ress i o n , w i t h d i sc a n d co l l a r g ra n u l a t e d a n d w i t h some l o n g i t u d i n a l s t r i a e ; p rono t a l t u b e rc l e s n o t reac h i n g te g u l ae ; scu t u m , s c u te l l u m a n d m e t a n o t u m d u l l ,

�

f u l l y g r a n u l a te d a n d ret i c u l a t e ru gose ; n o t a u l i ces a b sen t ; p ropodeum re t i c u l a te ru gose w i t h two l o n g i t u d i n a l k ee l s on posterior s u rf a ce ; fore w i n g w i t h two d a r k t r a n sv ers a l b a n d s , one on the b a s a l ce l l s and a n o t he r benea t h the p te ros t i g m a ; d i s t a l p a r t of rad i a l v e i n l on g e r t h a n p ro x i m a l p a r t ( 1 4 : 9 ) ; r a d i a l ce l l open ; fore t arsa l segme n t s i n fo l l ow i n g propor t i o n s : 27 : s u b ap i c a l t e e t h ( F i g . 698 ) w i t h t h e apex ; t i b i a l MALE : u n k n o w n

5 : 1 2 : 2 5 : 44 ; en l arged c l a w ( F i g . 698) w i t h t w o a n d w i t h a r o w of 2 5 l a me l l ae ; s e g m e n t 5 of fro n t t a r s u s two r o w s of n e a r l y 37 l a m e l l a e e x t e n d i n g con t i n uous l y t o s p u rs 1 , 1 , 2 .

D ry i n i n ae

1 02 7

L O C U S TYP I C U S : Po rta l ( Ar i zon a , U . S . A . ) T Y P I C A L MAT E R I A L : ho l ot y p e F ! i n AM . HOS T S : u n k n o w n D I ST R I B U T I O N : Nearc t i c reg i o n : U . S . A . : 2 m i . E Por t a l ( Coch i s e C o . , Ariz . ) , AM! N O T E S : t h e spec i es i s n a m e d i n honor of M r s . M a rjo r i e F a v reau .

N E O T R O P I C S P E C I ES OF MESODRYINUS

F o u r spec i e s a re k n ow n .

KEY T O T H E SPEC I ES

F E MA L E S

H ea d w i t h OL mo r e t h a n

t w i c e as

l on g as POL ; •

2

•

3 -

en l a rg ed c l a w

w i t h o u t a p i c a l row o f t e e t h ( F i g . 6 9 9 ) 1 . forestalis n . sp . H ea d w i t h OL n ea r l y a s l on g a s o r s h o r t e r t h a n POL ; en l a rg ed c l a w w i t h an a p i ca l r ow o f t e e t h ( F i g s 7 0 0 , 7 0 1 , 7 0 2 ) 2 E n l a r g ed c l a w w i t h ou t a p i c a l l ame l l a b i g g e r t h a n o t h e r l ame l l a e ( F i g . 702 ) 4 . a terrimu s n . s p . E n l a r g ed c l a w w i t h an a p i c a l l ame l l a b i g g e r t h a n o t h e r l a me l l a e ( F i g s 700 , 701 ) 3 P r on o t u m f u l l y r e t i c u l a t e ru g o s e 2 . cara i b ic u s n . s p . P r op od e u m n o t r e t i c u l a t e ru g o s e , p a r t l y s m o o t h a n d p a r t l y 3 . r u b er n . s p . s cu l p t u r e d b y c u r v i I i n ea r k e e l s •

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MALES

U n k no w n

1 . Mesodryin u s forestalis n . s p .

F E MA L E : fu l l y w i n g ed ; l e n g t h 3 , 0-3 , 88 m m ; head red d i sh-testaceous ; somet i me s vertex d a r ke ne d ; a n t e n n ae b row n , w i th s e g m e n t s 1 -2 testaceou s ; t ho ra x , propodeum a n d a b domen b l ac k ; l eg s testaceou s , w i t h m i d a n d coxae b l ac k ; a n te n n a e d i s t a l l y t h i c k e n e d ; a n te n n a l s e g m e n t s i n fol l ow i n g proport i o n s : 7 : 5 : 2 1 : 1 2 : 9 : 7 : 5 , 5 : 5 : 4 , 5 : 5 ; head exc a v a te d , d u l l ,

1 028

Mem . Amer . E n ! .

I ns t . , no. 3 7 ,

1 984

!

!

F i g . 700 - C he l a of Mesodryin u s caraibicus n . sp .

( type)

D r y i n i nae

1 02 9

g r a n u l a te d , w i t h n u merous l o ng i t u d i n a l s t r i a e ; frons w i t h o u t med i a n kee l ; occ i p i t a l c a r i n a i ncomp l e te , v i s i b l e on l y beh i n d t h e oce l l i a n d on t h e s i d es of oce l l a r t r i a n g l e , not touch i n g eyes ; poste r i o r oce l l i touch i n g occ i p i t a l c ar i n a ; temp l es i n v i s i b l e ; POL = 2 ; O L = 5 ; OOL = 7 ; pronotum d u l l , g r a n u l a ted , w i t ho u t s t r i a e , w i t h a n a n te r i o r w e a k t r a n s v e r s a l i m p ress i on a n d w i t h a p o s te r i o r s t ro n g t r a n svers a l i mp res s i on ; d i sc h u m pe d ; pos ter i o r c o l l a r i n v i s i b l e ; p ro no t a l t u berc l es not reac h i n g t e g u l a e ; s c u t u m d u l l , f u l l y ret i c u l a te r u gose ; n o t a u l i ces absen t , sc u te l l u m d u l l , g r a n u l a te d ; m e t a n o t u m short , very red u ced , ret i c u l a te ru gose ; p ropod e u m w i t h dorsa l s urface scu l p t u re d by n u merous p a r a l l e l l on g i t u d i n a l kee l s a n d w i t h poste r i o r s u rface areo l ae a n d w i t h t wo l o n g i t u d i n a l kee l s ; fore-w i n g w i t h t wo d a r k t r a n s v e rs a l b a n d s , one on t he b as a l ce l l s a n d a n o t her benea t h t he p teros t i g m a ; d i s t a l p a r t o f r ad i a l ve i n l on ger t ha n p ro x i m a l p a r t ( 1 1 : 6 , 5 ) ; fore t a rs a l segme n t s i n fo l l ow i n g p roport i on s : 1 1 : 2 : 6 : 1 0 : 2 1 ; segme n t 3 of fro n t t a r s u s p rod uced i n to a hoo k ; e n l arged c l a w ( F i g . 699 ) w i t ho u t s u b a p i c a l too t h , w i t h 1 row of 1 2 l a m e l l ae ; segmen t 5 of f ro n t t ar s u s ( F i g . 699 ) w i t h 1 row o f 7 l a me l l ae ; a pex w i t h a g roup of 1 2 l ame l l ae ; t i b i a l s p u rs 1 , 1 , 2 . MALE : unknown LOCUS T Y P I C U S : K w a t t a ( S u r i n a m ) T Y P I CAL MAT E R I AL : ho l o t y p e F ! i n L E ; 1 p a r a t y pe F ! i n A M ; 1 p a r a t y p e i n OL . HOST S : u n k n o w n D I S T R I B U T I O N : Neotrop i c reg i o n : SUR I NA M E : K w a t t a , OL ! LE ! BOL I V I A : R i o I tenez ( De p t . Be n i ) , AM !

2 . Nesodryin u s cara i b i c u s n . s p .

F E MAL E : f u l l y w i n ged ; l e n g t h 4 , 40 mm ; y e l l ow- tes t aceou s , w i t h p et i o l e b l ac k a n d a b l ac k med i a n reg i o n o n dors a l s u rface o f p ropodeu m ; a n t e n n a e d i s t a l l y t h i c k ened ; a n te n n a l seg m e n t s i n fo l l ow i n g p roport i on s : 1 5 : 6 : 2 5 : 1 2 : 1 1 : 9 : 7 : 6 : 6 : 1 0 ; head f l a t , d u l l , g r a n u l a ted ; fro n s w i t ho u t med i a n k ee l ; occ i p i t a l ca r i n a i ncomp l e t e , v i s i b l e on l y b eh i n d t he oce l l i a n d on t h e s i d es of t he pos t e r i o r oce l l i ; POL = 4 ; O L = = 4 ; OOL = 1 0 ; O P L = 0 ; p ronot u m w i t h two sm a l l l a t e r a l b ro w n spots o n co l l a r ; p ro n o t u rn crossed b y a n a n te r i o r t r a n s vers a l i m p ress i o n , d u l l , g r a n u l a te d ; d i sc h u m p e d ; t u b erc l es not re ac h i n g teg u I a e ; scu t u m , scute I I um a n d m e t a n o t u m du I I , gra n u l a te d ; n o t a u l i ces absen t ; p ropodeum ret i c u l ate ru gose , w i t ho u t t ra n s v e rsa l a n d l o n g i t u d i n a l kee l s ; fore- w i n g w i t h t w o d a r k t r a n s vers a l b a n d s ; ra d i a l ce l l open ; rad i a l v e i n w i t h d i s t a l p a r t l o n ger t h a n p ro x i m a l p a r t ( 22 : 1 0 ) ; fore t a rsa l segme n t s i n fo l l ow i n g p ropo r t i on s : 2 1 : 4 : 5 : 1 5 : 2 3 ; e n l arged c l a w ( F i g . 700 ) w i t ho u t s u b a p i c a l too t h , w i t h a n a p i c a l row of tee t h and w i t h a row of 1 0 l a me l l ae ; segmen t 5 of fro n t t a r s u s ( F i g . 700 ) w i t h t w o ro w s of n e a r l y 30 l a me l l ae e x t e n d i n g con t i n uous l y to t h e a p ex ; t i b i a l s p u rs 1 , 1 , 2 .

1 030

M e m . Amer . E n t .

I n st . ,

no.

37 ,

1 984

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Fig.

70 1

- C he l a of Mesodryin u s ruber n . s p .

( ty p e )

D ry i n i nae

1 03 1

MALE : u n k n o w n LOCUS TYP I C U S : A r i m a V a l l e y ( T r i n i d a d ) TYP I C A L M A T E R I AL : h o l o t y p e F ! i n A M . HOST S : u n k n own D I STR I BU T I ON :

Neo t ro p i c reg i o n :

T R I N I DA D : A r i m a Va l l e y , A M !

3 . Meso dryin u s ruber n .

sp .

F E MA L E : f u l l y w i n g e d ; l e n g t h 3 , 80 m m ; red d i sh-d a r k , w i t h a n te n n ae , a p e x o f abdomen a n d a r t i c u l a t i o n s of l e g s testaceou s ; a n te n n a e d i s t a l l y t h i ck e n e d ; a n t e n n a I seg men t s i n fo l l ow i n g p ropor t i o n s : 1 2 : 6 : 2 0 : 1 1 : 9 : 8 : 7 : 6 : 5 : 8 , 5 ; head d u l l , f l a t , h a i r y , g r a n u l a t e d ; frons w i t h o u t m ed i a n k ee l ; occ i p i t a l c a r i n a i ncom p l e t e , v i s i b l e on l y beh i n d t h e oce l l i a n d on t h e s i des of t h e oce l l a r t r i a n g l e ; POL = 5 , 5 ; OL = 4 ; OOL = 9 ; O P L = 1 ; p ronot u m d u l l , h a i r y , g ra n u l a t e d , crossed b y a t r a n svers a l i mp ress i o n ; d i s c h u m p e d ; p rono t a l t u b erc l es not reach i n g te g u l a e ; sc u t u m , scu te l l u m a n d m e t a n o t u m d u l l , h a i r y , g r a n u l a t e d ; not a u l i ces a b s en t ; p ropodeum n o t ret i c u l a te r u g os e , w i t h dors a l s u rface s h i n y , scu l p t u red b y some c u rv i l i n e a r l o n g i t u d i n a l k e e l s ; pos t e r i or s u rf a c e of p ropodeum s h i n y , smoo t h , exce p t for some t r a n s v e rs a l kee l s , s i m i l a r t o areo l ae , n e a r dorsa l s u rface ; m e t a p l e u r a a n d mesop l e u r a smoo t h , h a i ry , p a r t l y gra n u l a te d ; fore w i n g s w i t h two d a r k t r a n s versa I b a n d s , One on the b a s a I ce l l s a n d a n o t h e r b e n e a t h t h e p t e ros t i g m a ; rad i a l ce l l open ; r ad i a l v e i n w i t h d i s t a l p a r t l o n ge r t h a n p rox i m a l p a r t ( 1 7 : 8 ) ; fore t a rsa l seg m e n t s i n fol l ow i n g p roport i o n s : 20 : 3 : 5 , 5 : 1 5 : 25 ; e n l arged c l a w ( F i g . 7 0 1 ) w i t ho u t s u b a p i c a l toot h , w i t h an a p i c a l row o f t e e t h a n d w i t h a row of 1 0 l a me l l ae ; seg men t 5 of fron t t a r s u s ( F i g . 701 ) w i t h tow row s of n ea r l y 40 l a me l l ae e x t e n d i n g con t i n u ou s l y to the a p e x ; t i b i a l s p u r s 1 , 1 , 2 . MALE : u n k no w n L O C U S T Y P I C U S : T he B i g h t ( C a t I s l a n d , B a h a m a s I s l a n d s ) T Y P I CA L M A T E R I AL : h o l o t y p e F ! i n A M . HOST S : u n k no w n D I ST R I B U T I O N : Neo t rop i c reg i o n : BAHAMAS I S LA N D S : T he B i g h t ( C a t

I s l an d ) ,

AM !

4 . Mesodryin u s a terrimus n .

sp .

F E MA L E : f u l l y w i n ged ; l en g t h 5 , 20 m m ; head b l ac k , w i t h c l y pe u s , m a n d i b l es , m a l a r s p ace a n d a n te r i or re g i on of frons tes t aceou s ; tem p l es p a r t l y tes t aceo u s ; t ho r a x a n d p ropodeum b l ac k ; abdomen b l ac k , w i t h a p e x b row n ; l e g s b l ac k , w i t h a rt i c u l a t i o n s a n d t a r s i l i g h t ; a n t e n n ae d i s t a l l y t h i c k e n e d ; a n t e n n a I seg m e n t s i n fo l l ow i n g p ropor t i o n s : 1 1 : 5 : 63 : 24 : 2 1 : 1 2 : 1 1 : 9 : 8 : 1 1 ; head d u l l , g ra n u l a ted a n d w i t h n u m e rous l on g i t u d i n a l

1 032

Mem . Amer . E n ! .

F i g . 702

-

I n s t . , no . 3 7 ,

1 984

Che l a of Mesodryinus a terrimus n . s p .

( t yp e )

D r y i n i n ae

1 033

kee l s ; fro n s w i t h a comp l e te med i a n kee l , occ i p i t a c a r i n a i ncom p l e te , not v i s i b l e on temp l es ; POL � 3 , 5 ; OL � 3 , 5 ; OOL � 1 1 ; O P L � 2 ; TL � 0 ; p rono t u m d u l l , crossed b y a t r a n svers a l i m p ress ion ; p ron o t a l t u b erc l es not reach i n g te g u l ae ; p ro n o t u m g ra n u l a te d , w i t h n u mero u s s t r i ae a rou n d d i sc ; s c u t u m d u l l , g r a n u l a te d , ret i c u l a te r u gose a n d w i t h n u mero u s l o n g i t u d i n a l k e e l s ; not a u l i ces a b s en t ; sc u te l l u m a s scu l p t u red a s sc u t u m ; m e t a n o t u m a s scu l p t u red a s s c u t e l l um , except for p o s t e r i o r m a r g i n smoot h ; p ropodeum d u l l , w i t h dors a l s u rf a ce scu l p t u red by n u merous l o n g i t u d i n a l k ee l s ; poster i o r s u rface ret i c u l a te ru gose , w i t h t w o l o n g i t u d i n a l keel s ; fore-w i n g w i t h t w o d a rk t r a n s vers a l b a n d s , one on the b as a l ce l l s a n d a n o t h e r benea t h the p t e ros t i g m a ; d i s t a l p a r t of r a d i a l v e i n l o n ger t h a n prox i m a l p a r t ( 1 2 : 1 0 ) ; fore t a r s a l segmen t s i n f o l l ow i n g proport i on s : 2 5 : 4 : 1 3 : 24 : 40 ; en l a rged c l a w ( F i g . 702 ) w i t h two s u b a p i c a l teeth a n d w i t h a row of 24 l a me l l ae ; s e g m e n t 5 of fro n t t a r s u s ( F i g . 702 ) w i t h two row s of nea r l y 3 8 l am e l l ae exte n d i n g con t i n uo u s l y t o the a p e x ; t i b i a l s p u r s 1 , 1 , 2 . MALE : u n k n o w n L O C U S TYP I C U S : At mou t h of R i o B a u res ( Bo l i v i a ) T Y P I CAL M A T E R I A L : ho l o t y p e F ! i n A M . HOST S : u n k no w n D I ST R I B U T I ON : Neotrop i c reg i o n : BOL I V I A : A t mo u t h o f R i o B a u res ( R i o I t enez , Dep t . Be n i ) , AM !

7 . Perodryin us R . C . L .

Perk i n s

Perodryin us R . C . L . P e r k i n s 1 907 : 4 3 . Perodryin u s R . C . L . Perk i n s : K i effer 1 9 1 4 b : 48 .

T Y P E S P E C I E S : Perodryin us amoe n u s R . C . L . Perk i n s 1 90 7 , mono t y p i c . F E M A L E ( F i g s 703 , 704 ) : f u l l y w i n g e d ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segme n t s ; en l a rged c l a w ( F i g . 7 1 1 ) s h orter t h a n fron t t i b i a , w i t h o u t s ub a p i c a l toot h , ne a r l y as l o n g a s or s l i g h t l y l on ger t h a n a ro l i u m ; not a u l i ces a b s e n t or i ncomp l e t e ; occ a s i on a l l y weak l y v i s i b l e ; p rono t a l t u berc l e s not reach i n g tegu l ae ; occ i p i t a l c a r i n a com p l e te ; t i b i a l s p u rs 1 , 1 , 2 . M A L E ( F i g s 705 , 706 ) : f u l l y w i n g e d ; m a x i I l a r y p a l p i w i t h 6 segmen ts ; l a b i a l p a l p i w i t h 3 segme n t s ; o t her gener i c morpho l o g i c c h a racter i s t i c s : see f a m i I y d i a g nos i s . D I S T R I B U T I ON : Nearct i c , HOSTS : A ca n a loniidae SPEC I ES : 5

Neotrop i c

N E A R C T I C S P E C I E S OF PERODR YL�US

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Mem . Amer .

Fig.

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( type )

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Perodry i n u 5 aliloel i U ";:, R . C . L . Perodryin u5 amoen u s R . C . L . Perodry in u s a moen us R . C . L . 1 035 . Perodryin u s a moen u s R . C . L .

Perk i n s 1 907 : 4 3 . Perk i n s : K i effer 1 9 1 4b : 4 g . Perk i n s : M u esebeck a n d W a I k l ey 1 95 1 : Perk i n s : K romb e i n 1 979 :

1 244 .

F E MA L E : f u l l y w i n ged ; l e n g t h 4 , 20-4 , 56 mm ; b l ack , w i t h a n te n n ae y e l l o w t e s t a ceou s , c l y p eus and m a n d i b l es y e l l o w , p ro no t u m y e l l o w-redd i s h , t e g u l ae a n d l e g s y e l l o w-red d i s h ; somet i mes s ta l ks of h i n d femora b l ack ; a n te n n ae very shor t , shorter t h a n b od y , d i s t a l l y t h i c k e n e d ; a n t e n n a I s e g m e n t s i n fo l l ow i n g proport i o n s : 1 0 : 5 : 2 0 : 9 : 7 , 5 : 7 : 5 : 5 : 5 : 7 ; h e a d ru gose ,

1 035

D r y i n i n ae

F i g . 704 - F e m a l e of Perodryin u s n eotrop i c u s n . s p .

h a i ry ,

f u l l y ret i c u l a t e r u gose a n d gra n u l a ted ;

fron t a l

( ty p e )

l i n e comp l e te ;

occ i p i t a l c a r i n a com p l et e ; P O L = 6 ; OL = 4 ; OL = 8 , 5 ; O P L = 2 , 5 ; T L = 4 ; O P L shorter t h a n l e n g t h of oce l l a r t r i a n g l e ; p ronot u m s h i n y , h a i ry , weak l y s t r i ate aro u n d d i sc , h u m pe d , w i t h t wo t r a n s v e rs a l i mp ress i on s ; a n t e r i o r i mp res s i o n w e a k ; poste r i o r col l a r i n v i s i b l e ; dorsa l l y t h e p ro n o t u m s h o w s a s t ro n g con s t r i c t i o n i n fro n t of t h e sc u t u m ; p ro no t a l t u b erc l es n o t reac h i n g tegu l a e ; sc u t u m s h i n y , ru gose , f u l l y ret i c u l a t e r u gose a n d g r a n u l a ted ; n o t a u l i ces i n v i s i b l e ; sc u te l l u m smoot h , gra n u l a t e d , ret i c u l a t e ru gose ; occas i o n a l l y not ret i c u l a t e ru gose ; m e t a n o t u m r u gose , ret i c u l a t e r u gose a n d g ra n u l a ted ; p ropod e u m ret i c u l a t e ru gose , w i t h t w o l o n g i t u d i n a l k ee l s o n poste r i or s u rf ace ; med i a n a rea o f pos t e r i o r s u rf ace re t i c u l a te r u g ose ; occa s i o n a l l y not ret i c u l a t e ru gose ; fore-w i n g w i t h one d a r k t r a n sversa l b a n d b e n e a t h t h e p t e ros t i g m a ; d i s t a l p a r t of r a d i a l v e i n n ea r l y as l o n g a s prox i m a l p a r t ( 1 1 : 1 0 ) ; t h e t w o p a r t s form i n g a c u r v e ; rad i a l ce l l open ; fore t a rs a l se g m e n t s i n fo l l ow i n g p ro port i on s 2 1 : 3 : 5 : 5 : 1 0 ; en l arged c l a w ( F i g . 707 ) v e r y short a n d s l en d e r , n e a r l y a s long as aro l i u m , w i t h o u t s u b a p i c a l too t h a n d l a me l l ae o r b r i s t l es ; segmen t 5 o f fron t t a r s u s ( F i g . 707 ) n e a r l y t w i ce a s l o n g a s en l a r g e d c l a w , w i t h t w o r o w s of 2 1 l a me l l ae e x t e n d i n g con t i n u ous l y to t h e

1 036

M e m . Amer. E n t .

I ns t . , n o . 3 7 ,

1 984

F i g . 705 - M a l e of Perodry inus amoen u s R . C . l . Perk i n s ( p a r a l ectotype )

apex ; t i b i a l s p u r s 1 , 1 , 2 . MALE ( F i g s 705 , 706 ) : f u l l y w i n g e d ; l e n g t h 3 , 66 mm ; b l ack ; a n tennae y e l l ow - b row n ; m a n d i b l es test aceo u s ; abdomen b row n ; l eg s y e l l ow , w i t h m i d a n d h i n d coxae brow n ; h i n d t i b i ae a n d h i n d femora brow n ; a n te n n ae not d i s t a l l y t h i c k e n e d ; a n t en n a l segme n t s in fol l ow i n g p roport i on s : 8: 7: 1 5 : 1 0 : 1 1 : 1 1 : 1 0 : 8 : 7 : 9 ; head d u l l , f u l l y s t ron g l y ret i c u l a te ru gose ; c l y pe u s weak l y emarg i n a te d j occ i p i t a l c a r i n a comp l e te ; vertex convex ; temp l e s d i s t i n ct ; fro n t a l l i n e a bsen t ; POL ; 8 ; Ol ; 4 ; OOl ; 7 ; O P l ; 3 ; T l ; 4 ; scu t u m , sc u te l l u m , metanotum a n d propodeum ret i c u l ate ru gose , d u l l ; notau l i ces i n v i s i b l e ; p ropodeum w i t h o u t tran svers a l or l o n g i t u d i n a l kee l s ; fore- w i n g h y a l i n e , w i t h o u t d a r k tran sversa l b a n d s ; d i s t a l part of rad i a l ve i n s horter t h a n p rox i m a l p a r t ( 1 1 : 1 4 ) ; rad i a l ce l l open ; g en i t a l i a : f i g . 708 ; t i b i a l s p u r s 1 , 1 , 2 . lOCUS T Y P I C U S : Nog a l es ( Ar i zo n a , U . s . A . ) T Y P I CAL M A T E R I AL : l ecto t y p e F ! a n d 1 para l ectotype M ! ( des i g n a ted b y O l m i 1 982 ) i n B .

D r y i n i n ae

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1 037

M a l e of Perodry in u s amoen u s R . C . L .

Perk i n s ( p a r a l ecto t y p e )

HOST S : i n U . S . A . A c a n a lonia (=Amp h iscep a ) b ivitta ta ( S a y ) (A cana loniidae) ( R . C . L . Per k i n s 1 907 ) D I ST R I B U T I O N : N e a rc t i c reg i o n : Concord i a ( S i n a l oa ) , OT !

U. S. A. :

Nog a l es ( A r i z . ) ,

B ! M E X I CO :

N E O T R O P I C S P E C I ES OF PERODf(YINUS

F o u r spec i e s a re k n ow n .

K E Y TO T H E S P E C I E S

F E MA L E S

F o r e w i n g s h y a l i n e , w i t h ou t d a rk t ra n s v e r s a l b a n d s • . • • • • . . • • • F o r e w i n g s w i t h d a rk t ra n s v e r s a l b an d s . . . . . . . . . . . . . . . . . . . . . . . 2 H ea d r e t i c u l a t e r u g o s e • • • • • • • • • • • • • • • • • • • • 1 . a u tunnalis n . sp .

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1 038

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D r y i n i n ae

1 039

- H ea d w i t h n um e r ou s l on g i t u d i n a l k e e l s . . . . 2 . bolivia n u s n . s p . 3 P r o n o t u m b l a c k , w i t h ma r g i n s t e s t a c e o u s 3 . neotrop icus n . sp . - P r o n o t u m t e s t a c e o u s - r ed d i s h , w i t h a n t e r i o r c o l l a r b l a ck .

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7 0 8 - Ma l e g en i t a l i a o f Perodryin u s amoen u s R . C . L . ( p a r a l ec t o t y p e ) ( r i g h t h a l f removed )

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U n k n ow n

1 . Perodryin u s a u tumn a l is n .

sp .

F E MAL E : f u l l y w i n g e d ; l e n g t h 4 , 5 6 m m ; head b l ack , w i t h m a n d i b l es a n d c I y p e u s t e s t a c eo u s ; a n te n n a e a n d l eg s test aceo u s ; t h o r a x a n d p ropode u m b l ac k , w i t h d i sc a n d p a r t o f s i des o f p ro n o t u m ye l l ow- tes t aceo u s ; a b domen b l ac k ; a n t e n n ae d i s t a l l y t h i c kened ; a n te n n a I segmen t s i n fo l l o w i n g p ropor t i o n s : 1 5 : 7 : 30 : 1 1 : 1 1 : 9 : 6 : 6 : 5 : 8 ; head d u l l , f u l l y ret i c u l a te r u gose ; frons w i t h a med i a n kee l a n d 2 l a t e r a l i n com p l e t e k ee l s ; occ i p i t a l c a r i n a comp l e te ; P O L = 3 ; O L = 3 ; O O L = 6 ; O P L = 2 ; temp l e s i n v i s i b l e ; p ro n o t u m d u l l , ha i r y , w i t h n u merous s t r i a e a ro u n d d i s c , w i t h a weak a n te r i o r t r a n s v e rs a l i m p ress i o n and a poste r i o r s t ro n g t r a n s v e rs a l i mp ress i o n ; poster i o r co l l a r i n v i s i b l e ; p ra n a t a l t u b e rc l es n o t reac h i n g t e g u l a e ; sc u t u m , s c u te l l u m a n d m e t a n o t u m d u l l , f u l l y ret i c u l a te r u gose ; not a u I i ces absen t ; p ropode u m w i t h dorsa I s u rface s c u I p t u red b y s t ro n g l on g i t u d i n a l p a ra l l e l k ee l s ; pos ter i o r s u rface w i t h 2 l on g i t u d i n a l k ee l s ; l a t e r a l a n d med i a n areas w i t h n u merous t ra n s v e rs a l k ee l s a n d w i t h some a reo l ae ; fore-w i n g h y a l i n e , w i t h o u t dark t r a n s v e r s a l b a n d s ; d i s t a l p a r t of r a d i a l v e i n l o n g e r t h a n p ro x i m a l part ( 1 6 : 1 4 ) ; fore t a rsa l seg m e n t s i n fo l l o w i n g p ropo r t i o n s : 27 : 3 , 5 : 7 : 8 : 1 4 ; en l a r g e d c l a w ( F i g . 709 ) very r e d u ced , n e a r l y a s l o n g a s a r o l i u m , w i t h o u t s u b a p i c a l tooth and l a me l l ae or peg - l i ke h a i rs ; segmen t 5 of f ro n t t a rs u s ( F i g . 709 ) w i t h some rows o f 1 8 l a me l l ae e x t e n d i n g con t i n uo u s l y to t h e a p e x ; t i b i a l s p u rs 1 , 1 , 2 . M A L E : u n k n ow n LOCUS TYP I C U S : Verd a n t V a l e ( T r i n i d a d ) TYP I CAL MAT E R I AL : ho l o t y p e F ! i n WA . HOST S : u n k n o w n D I ST R I B U T I O N : Neot rop i c reg i o n : T R I N I DA D : V e rd a n t Va l e , WA !

2 . Perodryin u s bolivia n u s n .

sp

F E M A L E : f u l l y w i n ged ; l e n g t h 5 , 0 m m ; head b l ack , w i t h m a n d i b l es t e s t a c eo u s ; a n te n n a e b r o w n - b l ack , w i t h seg m e n t s 1 -2 y e l l ow ; p ropec t u s b l ac k ; p ro n o t u m red d i s h ; meso t h o r a x , m e t a t h o r a x a n d p ropod e u m b l a ck ; abdomen b l ac k ; l e g s test aceo u s ; head sw o l l e n , d u l l , g ra n u l a t e d , w i t h n u merous l o n g i t u d i n a l kee l s ; occ i p i t a l c a r i n a comp l e te ; temp l es i n v i s i b l e ; pos t e r i o r oce l l i not touch i n g occ i p i t a l c a r i n a ; OPL s h o r t e r t h a n O L ; O L = P O L ; p r o n o t u m h a i r y , d u l l , w i t h n u merous w e a k s t r i ae a ro u n d d i s c ; a n t e r i o r t r a n s v e rs a l

i m p r e s s i o n a n d a n t e r i or co l l a r w e a k ;

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1 042

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( type)

1 043

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( type)

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no . 37 ,

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t r a n s v e r s a l i mp res s i o n stron g ; pos ter i or col I a r-' i n v i s i l; : e ; s c u t u m r-e t i c u l ate r u gose , h a i r y ; not a u l i ces w e a k l y v i s i b l e , i n comp l et e , reac h i n g a p p rox i m a te l y 0 , 5 l en g t h o f scu t u m ; s c u te l l u m a n d m e t a n o t u m ret i c u l a t e r u gose ; propodeum ret i c u l a te rugose , w i t h 2 l o n g i t u d i n a l k ee l s on pos t e r i o r s u rface ; fore- w i n g w i t ho u t d a rk t r a n sversa l b a n d s a n d w i t h d i s t a l part of radi a I v e i n l o n ger than p rox i m a I p a r t ( 1 7 : 1 4 ) ; segment 1 of f ro n t tarsus n e ar l y t h ree t i mes as l on g a s segme n t 4 ( 24 : S ) ; seg ment 3 of front tarsus p roduced i n to a hoo k ; e n l a rged c l aw ( F i g . 7 1 0 ) very reduced , s l i g h t l y l onger t h a n a r o l i u m , ap i c a l l y serra te , w i t h o u t l a me l l ae o r peg- l i ke h a i r s ; segme n t 5 of f ro n t t a rsus ( F i g . 7 1 0 ) w i t h some rows o f 2 9 l am e l l ae exten d i n g con t i n u ou s l y to t he apex ; t i b i a l s p u rs 1 , 1 , 2 . MAL E : u n k n o w n LOCUS TYP I C US : R u r e n ab a q u e ( B o l i v i a ) TYP I CAL MAT E R I AL : ho l o t y p e F ! i n BM . HOST S : u n k n o w n D I ST R I BU T I O N : Neotrop i c reg i o n : BOL I V I A : R u re n a b a q ue ( Ben i ) , B M !

3 . Perodryin u s neotropicus n . s p .

F E MALE ( F i g s 703 , 704 ) : f u l l y w i n ged ; l e n g t h 4 , 37-4 , 68 m m ; head b l ac k , w i t h c l y peus a n d m a n d i b l e s y e l l o w ; a n t e n n ae testaceou s ; propec t u s b l ac k ; p ro n o t u m b l ac k , w i t h s i des a n d pos t e r i o r co l l a r red d i s h ; mesothorax , met a t horax a n d propode u m b l ac k ; abdomen b ro w n -red d i s h ; legs test aceous , rare l y brown w i t h h i n d coxae testaceou s ; a n ten n a l segme n t s i n fo l l ow i n g propor t i o n s : 1 2 : 5 : 2 2 : 1 0 : 7 : 6 : 5 : 4 , 5 : 4 : 7 ; hea d s wo l l e n , h a i ry , d u l l , ret i c u l a t e rugose ; occ i p i t a l ca r i n a comp l et e ; pos t e r i o r oce l l i not tou c h i n g occ i p i t a l c a r i n a ; O P L = O L ; OL shorter t h a n POL ; pronot u m d u l l , h a i r y , w i t h n u merous weak s t r i a e arou n d d i sc ; a n t e r i o r t r a n svers a l i mp res s i on a n d a n t e r i o r co l l a r weak ; poster i or t r a n s v e r s a l i mp ress i o n s t rong a n d near poste r i o r m a r g i n of pron o t u m ; pos ter i o r coi l a r i nv i s ib I e ; p ro n o t a I t u berc l e s not reac h i n g tegu l ae ; scu t u m du l l , h a i r y , f u l l y re t i c u l ate ru gose ; n o ta u l i ces absen t ; s c u te l l u m ret i c u l a t e rugose ; metanotum gra n u l a ted ; p ropod eum ret i c u l a t e rugose , w i t h 2 l on g i t u d i n a l kee l s on pos terior s u rface ; fore- w i n g w i t h 3 brown t r a n svers a l b a n d s and w i t h d i s t a l part of r ad i a l ve i n l o n ger than prox i m a l p a r t ( 1 5 : 1 0 ) ; seg men t 1 of f ro n t tarsus more than t h ree t i mes as l o n g as seg m e n t 4 ( 1 8 : 5 ) ; segmen t 3 of f ro n t t arsus p roduced i n to a hook ; en l a rged c l aw ( F i g . 7 1 1 ) v e r y reduced , n ea r l y as l o n g as a ro l i u m , w i t h o u t s u b a p i c a l tooth a n d l a mel l ae or peg l i ke h a i rs ; seg men t 5 of f ro n t t a r s u s ( F i g . 7 1 1 ) w i t h some rows of 23 l ame l l ae exte n d i n g con t i n u o u s l y to the apex ; t i b i a l spurs 1 , 1 , 2 . MALE : u n k n o w n L O C U S TYP I C U S : St . A u g u s t i n e ( T r i n i d a d ) T Y P I CA L MAT E R I AL : h o l o t y pe F ! a n d 1 para t y p e F ! i n BM ; 3 p a r a t y p es F F ! i n O T ; 2 p a r a t y pes F F ! i n O L . HOST S : u n k no w n

Dry i n i nae

1 045

D I ST R I B U T I O N : Neot rop i c reg i on : M E X I C O : M u s t e ( H u i x t l a , C h i a p a s ) , O T ! T R I N I DA D : S I . A u g u s t i n e ( S I . George ) , Ol ! B M ! C u repe ( S t a . M a r g a r i t a , C i rcu l a r R d . ) , O T ! O l !

Fig.

7 1 2 - C he l a o f Perodryin u s b icolor n . s p .

4 . Perodryin u s b icolor n .

( type)

sp .

F E MA L E : f u l l y w i n ged ; l e n g t h 5 m m ; head b l ac k , w i t h c l y p e u s a n d m a n d i b I es t e s t aceous ; a n t e n n ae t e s t aceou s ; p ropec t u s b l a ck ; p ro n o t u m t e s t a ceous-re d d i s h , w i t h a n t e r i or co l l a r b l ac k ; mesot hora x , m e t a t hora x , p ropo d e u m a n d a b domen b l ack ; l eg s t e s t aceo u s , w i t h s t a l k s o f h i n d femora b l a ck ; a n t e n n a e d i s t a l l y t h i c kened ; a n t e n n a I s e g m e n t s in fo l l ow i n g propor t i o n s : 1 4 : 6 : 2 9 : 1 3 : 9 : 8 : 6 : 5 : 4 , 5 : 8 ; h e a d d u l l , h a i ry , re t i c u l a te ru gose a n d w i t h a few l on g i t u d i n a l k ee l s ; frons w i t h a med i a n comp l e t e k ee l ; occ i p i t a l c a r i n a comp l e t e ; t e m p l es i n v i s i b l e ; POL = 4 , 5 ; O l = 3 ; OOl = 5 ; O P l = 2 , 5 ; prono t u m h a i r y , s h i n y , w i t h n um e ro u s s t r i ae a ro u n d d i sc ; p rono t a l t u b erc l e s not reach i n g t e g u l ae ; pron o t u m cross(,d b y a n a n t e r i o r w e a k t r a n s v e r s a l i m p ress i on a n d b y a pos t e r i o r s t rong t ra n s v e rs a l i m p ress i o n ; d i sc h u m p ed ; pos t e r i o r co l l a r v ery shor t ; scu t u m , scu t e l l u m a n d m e t a n o t u m d u l l , ret i c u l a te ru gose ; n o t a u l i ces w e a k l y v i s i b l e , i n com p l e t e , reac h i n g a p p rox i ma te l y 0 , 5 l e n g t h o f scu t u m ; propodeum w i t h n u merous l o n g i t u d i n a l k ee l s o n dors a l s u rface ; pos t e r i o r s u rface of p ropode u m ret i c u l a t e ru gose a n d w i t h two l o n g i t u d i n a l k ee l s ; fore w i n g s

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w i t h two dark t r a n s v ersa I b a n d s , one on the b a s a I ce l l s a n d a n o t her b enea t h t he p te rost i g ma ; d i s t a l p a r t of rad i a l v e i n n ea r l y a s l on g as p rox i m a l pa r t ; fore t a r s a l segme n t s i n fo l l ow i n g p ropor t i o n s : 2 7 : 4: 9: 6 : 1 5 ; segmen t 3 o f f ro n t t a r s u s prod uced i n to a hook ; e n I a rged c l a w ( F i g . 7 1 2 ) very reduced , n ea r l y a s l o n g a s a ro l i u m , w i t h o u t s U b a p i c a l too t h a n d l ame l l ae o r peg-l i ke h a i r s ; segmen t 5 o f fron t t a r s u s ( F i g . 7 1 2 ) w i t h SOme rows of near l y 23 l a me l l ae exten d i n g con t i n u o u s l y to t he a pex ; t i b i a l spurs 1 , 1 , 2 . MALE : u n k n o w n L O C U S TYP I C US : San E s t a b a n ( Venezue l a ) TYP I CAL MAT E R I AL : h o l o t y p e F ! i n T W . HOST S : u n k n o w n D I STR I B U T I O N : Neotrop i c reg i o n : V E N E Z U E LA : San E s t a b a n ( P u erto Cab e l l o ) , TW!

8 . Cretodryin us Ponomare n k o

Cretodryin us Ponomarenko 1 975d :

1 04 .

TYPE SPEC I E S : Cretodryin us zherichini Ponomarenko 1 975d , mon o t y p i c a n d o r i g . d es i g . F EMAL E : f u l l y w i n g e d ; a n te n n a e , m ax i l l a r y a n d l ab i a l p a l p i not v i s i b l e i n the on l y k n o w n spec i me n ; prono t a l t u b erc l es a p p a re n t l y reac h i n g tegu l ae ; not a u l i ces app a ren t l y v i s i b l e ; e n l arged c l a w a p p a re n t l y w i t h o u t s u b a p i c a l toot h , w i t h l a m e l l ae , shorter t h a n fore t i b i a , much l o n ger t h a n a ro l i u m ; t i b i a l s p u r s a p p a re n t l y 1 , 1 , 2 . MAL E : u n k n o w n D I ST R I BU T I ON : P a l ae a rc t i c ( on l y foss i l spec i es ) HOST S : u n k n o w n SPEC I ES : 1

Cretodryin us zheric h in i Ponoma re n ko

Cretodryin us zherichini Ponoma r e n ko 1 975d :

1 04 .

FEMAL E : f u l l y w i n g e d ; l e n g t h 2 , 4 m m ; co l o u r of bod y s l i g h t l y v i s i b l e , apparen t l y d a r k , w i t h l e g s test aceous ; a n te n n a e , p a l p i , m a n d i b l es a n d p a r t of h e a d s l i g h t l y v i s i b l e i n on l y k n o w n spec i men ; p ro n o t u m s l i g h t l y v i s i b l e ; m a y b e p ro n o t a l t u b e rc l es reac h i n g te g u l ae ; pronotum a p p aren t l y w i t hou t d i sc a n d t r a n svers a l i m p ress i o n s ; sc u t u m a p p a re n t l y w i t h i ncomp l e te notau l i ces ; scute l l u m a n d metanotum not eas i l y v i s i b l e , bu t

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prese n t ; propodeum a p p a r en t l y w i t h areo l ae ; fore w i n g s h y a l i n e , w i t h p t e ros t i g m a l o n g a n d n a r ro w , not l a nceo l a t e , a p p a ren t l y w i t h t h ree b a s a l c e l l s e nc l osed ; rad i a l v e i n a p p a r e n t l y w i t h d i s t a l p a r t as l on g a s p ro x i m a l p a r t ; abdomen w i t ho u t a d i s t i nc t p e t i o l e ; p ropod e u m w i t h pos t e r i o r s u rface very s tep ; fore t a r s i c he l a te ; r u d i me n t a ry c l a w a p p aren t l y prese n t ; en l a r g e d c l a w a p p aren t l y w i t hou t s u b a p i c a l too t h a n d w i t h l a m e l l ae ; en l a r g e d c l a w m u c h l on g e r t h a n a ro l i u m , shorter t h a n fore t i b i a ; segme n t 5 of fron t t a rs u s app a ren t l y w i t h rows of l a me l l ae ; segmen t 4 of f ro n t t a rs u s a p p a ren t l y s l i g h t l y l o n ger t h a n segme n t 1 ; t i b i a l s p u rs apparen t l y 1 , 1 , 2 . MA L E : u n k n o w n L O C U S TYP I C U S : T a i m y r a mber ( S i b i r i a ) T Y P I CAL MAT E R I AL : h o l o t y p e F ! i n M P I . HOST S : u n k n o w n D I S T R I B U T I O N : P a l aearc t i c reg i o n : U . S . S . R . : T a i m y r a m b e r ( S i b i r i a ) , MP I ! N O T E S : Cr . zherichini i s one of o l de s t foss i l D ry i n i d s . ( n e a r l y 80 m i l l i on" years o l d v e rb a l commu n i c a t i on of Mr. A. Ponomarenko ) . T h e con d i t i o n s of t h e foss i I are too b a d to com p a r e i t w i t h o t h e r k n o w n fossi l . T he v a l i d i t y of t h e gen u s Cretodryin u s i s u n cert a i n .

F i g . 7 1 3 - Fema l e of Tra n s dryin u s notogeicus n . s p .

( t y pe )

1 04 8

Mem . Amer . E n ! .

I nst . ,

no. 37,

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8 . SU BFAM I LY TRANSDR YININAE N . SUBF .

TYPE S P E C I E S : Tran s dryin u s n . gen . F E MALE ( F i g s 1 7 , 7 1 3 ) : f u l l y w i nged ; fore-w i n g w i t h cos t a l , med i a n a n d s ubmed i a n ce l l s c l ea r l y enc l osed b y p i gmen ted v e i n s ( F i g s 1 7 , 7 1 3 ) ; m ax i l l a r y p a l p i w i t h 6 segments ( F i g . 7 1 4 D ) l a b i a l p a l p i w i t h 3 segments ( F i g . 7 1 4 C ) ; frons tarsus c he l a te ; c he l a w i t h o u t ru d i me n t a r y c l aw ( F i g . 7 1 4 A ) ; ma n d i b l es w i t h one on l y too t h ( F i g . 7 1 4 B ) ; en l a rged c l aw w i t h o u t s u b ap i c a l t o o t h or other teeth ( F i g . 7 1 4 A ) ; segmen t 5 of fro n t t a rsus w i t h l ame l l ae ( F i g . 7 1 4 A ) ; pronot a l t u berc l es ( d i rec ted towards the teg u l a e ) presen t ; occ i p i t a l c a r i n a i n v i s i b l e ; pet i o l e short ; a n te n n a e w i t h o u t t u f t s of l ong h a i r s ; t i b i a l spurs 1 , 0 , 1 . MALE : u n k n o w n D I ST R I BU T I O N ; A u s t r a l i a n HOST S : u n k n o w n GENERA: 1 SPEC I ES : 1

Tra n s dryin u s n . gen .

TYPE S P E C I E S : Tran s dryin u s notogeicus n . sp . F EMALE ( F i gs 1 7 , 7 1 3 ) : f u l l y w i n ged ; ma x i l l a r y p a l p i w i t h 6 segmen ts ; l a b i a l p a l p i w i t h 3 seg m e n t s ; pronotum crossed by a tran sversa l i m pres s i o n ( F i g . 1 7 ) ; prono t a l tuberc l es not reac h i n g te g u l a e ; n o t a u l i ces i n v i s i b l e ; occ i p i t a l car i n a i n v i s i b l e ; en l arged c l aw ( F i g . 7 1 4 A) w i t h o u t s u b ap i ca l toot h or o t h e r tee t h , w i t h o u t l ame l l ae , w i t h one b r i s t l e on a p rox i m a l prom i nence ; t i b i a l spurs 1 , 0 , 1 . MALE : u n k n o w n D I ST R I B U T I ON : A u s t r a l i a n HOST S : u n k n o w n SPEC I E S : 1

Tra n s dryin u s notogeic u s n . sp .

F E M A L E ( F i g s 1 7 , 7 1 3 ) : f u l l y w i n ged ; l e n g t h 4 , 1 8 mm ; head test aceous redd i s h , w i t h a b l ac k tra nsvers a l b a n d on vertex and 2 l a tera l b l ac k s t r i pes on f ro n s ; a n t e n n ae testaceou s , w i t h segmen ts 6-9 b l ac k a n d segme n t 1 0 w h i t i s h ; propec t u s b l ac k ; pronot u m b l ac k , w i t h l a tera l a n d poster i or m a r g i ns red d i s h ; mesothor a x , met a t horax a n d propodeum b l ack ; a b domen b l ack ; l eg s testaceo u s , w i t h coxae pa r t l y b l ack , femora pa r t l y b rown a n d m i d a n d h i n d t i b i ae brow n ; a n t e n n ae d i s t a l l y t h i ckened ; a n ten n a l segmen ts i n fo l l o w i n g propor t i o ns : 8 : 6: 1 5 : 1 0 : 9: 7 , 5 : 6: 6: 5 : 8 ; m a n d i b l es w i t h

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1 049

F i g . 7 1 4 - F em a l e of Tr a n s dryinus n o togeicus n . s p . ( t y p e ) ; A : c h e l a ; B : m a n d i b l e ; C : l ab i a l p a l p u s ; D : m ax i l l a ry p a l p u s .

one on l y toot h ; head s h i n y , g r a n u l a te d ; fron s w i t h a s h o r t med i a n k e e l v i s i b l e on l y i n fro n t of t h e a n t e r i or oce l l u s ; occ i p i t a l c a r i n a i n v i s i b l e ;

1 05 0

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En ! .

I ns ! . ,

no . 3 7 ,

1 984

POL � 4 ; Ol � 3 , 5 ; OOl � 8; p ro n o t u m d u l l , g r a n u l a te d , w i t h a s t ron g t r a n s v e rs a l i m p re s s i o n ; p ro no t a l t u b e rc l es n o t reach i n g t e g u l a e ; scu t u m d u l l , g ra n u l a t e d , w i t h c e n t r a l reg i o n r e t i c u l a t e r u gose ; n o t a u l i ces a bsen t ; sc u t e l l u m and m e t a n o t u m smoo t h , sh i n y , w i t hou t scu l p t u re ; p ropod e u m w i t h dors a l s u rf ac e s t ron g l y scu l p t u re d by n u merous p a ra l l e l l o n g i t u d i n a l k ee l s ; poster' i o r s u rf a c e smoot h , sh i n y , w i t hou t scu l p t u re or w i t h very weak t r a n s vers a l i m press i o n s , w i t h o u t areo l ae or k ee l s ; fore w i n g w i t h 2 d a r k t r a n s v e rs a l b a n d s , one on t h e b a sa l c e l l s and a n o t h e r benea t h t h e p t e ros t i g m a ; d i s t a l p a r t o f ra d i a l v e i n n ea r l y t w i c e as l on g a s prox i m a l p a r t ( 1 4 : 7 ) ; fore t a r s a l segmen t s i n fo l l ow i n g p roport i o n s : 1 3 : 3 : 5 : 1 8 : 2 5 ; e n l arged c l aw ( F i g . 7 1 4 A ) w i t h ou t t ee t h , w i t h one b r i s t l e on a p r ox i m a l p rom i nence ; segmen t 5 of fron t t a r s u s ( F i g . 7 1 4 A ) w i t h one row o f 1 4- 1 6 l a m e l l a e ; a pex w i t h a grou p o f 3 l a me l l ae ; r u d i me n t a r y c l a w a b s en t ; s e g m e n t 3 of fron t t a rs u s p ro d u ced i n to a hook ; t i b i a l s p u rs 1 , 1 , 2 . M A L E : u n k n ow n lOCUS T Y P I C U S : B r i s b a n e ( A u s t ra l i a ) TYP I CA L MAT E R I A L : h o l o t y pe F ! i n BR . H OSTS : u n k no w n D I ST R I B U T I O N : A u s tr a l i a n reg i o n : AUSTRAL I A : B r i s b a n e ( Qu een s l a n d ) , BR !

9 . SUBFAM I LY GONA TOPODINAE K I E F F E R

Gon a topodinae p artim K i effer i n K i effer e t M a r sh a l l Gona topodin a e partim K i effer 1 907b : 1 2 . Gona topodini K i effer 1 9 1 4b : 6 7 . Gona topodini R i c h a r d s 1 93 9 : 1 88 . Dryin ini p artim M u e se b eck a n d W a l k l e y 1 95 1 : 1 034 . Gona topodin a e R i c h ar d s 1 95 3 : 5 1 . Gon a topodin a e Kromb e i n 1 979 : 1 244 .

T Y P E G E N U S : Gona topu5 Lj u n g h 1 8 1 0 F E M AL E : a p terous ( F i g s 7 9 8 , 799 , 8 0 1 ,

809,

1 906 : 499 .

8 7 3 , 894 ) ; som e t i mes f u l l y

w i n g e d ( F i g s 7 1 5 , 7 1 6 , 754 , 7 5 5 , 767, 7 68 ) ; occ a s i o n a l l y m i crop t e rous ( F i g s 1 2 50 , 1 25 1 ) ; fore-w i n g ( i n f u l l y w i n ged spec i es ) w i t h cos t a l , med i a n a n d su b med i a n c e l l s c l e a r l y e n c l osed by p i g m e n t e d v e i n s ( F i g . 7 1 5 ) ; m ax i l l a ry p a l p i w i t h 2-6 segmen t s ; l a b i a l p a l p i w i t h 1 -3 segme n t s ; poss i b l e p a l p a l ra t i os : 2 / 1 , 2/2, 3/2, 4/2 , 4/3, 5/2 , 5/3, 6/2, 6/3 ( f i rs t n u m b e r � m a x i I l a ry p a l p segmen t s ; sec o n d n u m b e r = l a b i a l p a l p segmen t s ) ; m a n d i b l e s w i t h 4 t e e t h p rog ress i n g l a rger from a n t e r i or one to poster i o r ; occ i p i t a l c a r i n a a b s e n t or v i s i b l e on l y b eh i n d the oce l l i or on the s i des of the pos t e r i o r oce l l i ; occ i p i t a l c a r i n a n e v e r reac h i n g temp l es ; fore t roc h a n t e r s more t h a n tw i ce as l o n g a s b road ; segmen t 3 of fron t t a r s u s p roduced i n t o a hook ; fron t t a r s i ch e l a t e ( F i g . 708 ) ; 'ch e l a w i t h

Go n a topod i n ae

1 05 1

r u d i me n t a r y c l a w ( F i g . 7 1 9 ) ; s e g m e n t 5 o f fro n t t a rs u s a l w a y s w i t h more t h a n one p re a p i c a l l a me l l a ( F i g . 7 1 9 ) ; a n t e n n ae w i t ho u t tufts of l o n g h a i rs ; p ronot a l t u b e rc l es a b sen t ; i n w i n g e d fema l es m e t a n o t u m very reduced a n d p teros t i g m a very n a r row ; t i b i a l spurs 1 , 0 , 1 ; occa s i on a l l y 1 ,0,2. MALE ( F i gs 7 1 7 , 7 1 8 ) : f u l l y w i n ged ; fore-w i n g w i t h costa l , med i a n a n d s u b me d i a n ce l l s c l ea r l y e n c l osed b y p i gmen ted v e i n s ( F i g . 7 1 7 ) ; m a x i I l a r y p a l p i w i t h 2 - 6 s e g me n t s ; l a b i a l p a l p i w i t h 1 -3 seg m e n t s ; poss i b l e p a l p a l r a t i o s : 2 / 1 , 2/2 , 3/2 , 4/2 , 4/3 , 5/2 , 5/3 , 6/2 , 6/3 ( f i rs t n u m b e r = m ax i l l a ry p a l p segmen ts ; seco n d n u m b e r = l a b i a l p a l p segmen ts ) ; m a n d i b l es w i t h 3 tee t h p ro g res s i n g l ar g e r from a n te r i o r o n e to poster i o r ; occi p i t a l c a r i n a a b s en t ; som e t i mes i ncom p l e te a n d v i s i b l e on l y beh i n d t h e oce l l i a n d on t h e s i des o f t he pos t e r i or oce l l i ; i ncomp l e te occ i p i t a l ca r i n a n ev e r reach i n g temp l es ; occa s i on a l l y occ i p i t a l ca r i n a comp l e t e ; tem p l es u s u a l l y prese n t , occ a s i o n a l l y a b se n t ; p t e ros t i gm a a l w a y s v e r y n a rrow ; occ i p u t u s u a l l y con c a v e , occa s i on a l l y s t ra i g h t ; g e n i t a l a r m a t u re a l w a y s w i t h dors a l p rocess of gonoforcep s ; t i b i a l s p u rs 1 , 1 , 2 . D I ST R I B U T I O N : wor l d w i d e HOST S : Cic a dellidae ( exce p t f o r Idiocerin a e , Macropsinae, Typhlocyb in a e ) ,

Trop iduchida e, Lophopidae, Fl a tidae, A c a n a l o n iidae, Rica niidae, Nogodin idae, Delphacida e . G E N E RA : 1 8 S P E C I E S : 346

N O T E S ON T H E TAXONOMY O F T H E A P T E R O U S F E M A L E S

T h e c l a s s i c a l c l a ss i f i c a t i on of t h e a p t erous fema l es o f Gonatopo d i n e s , accord i n g to R . C . L . Perk i n s , K i effer a n d R i c h a r d s , w a s b ased on t h e fo l l ow i n g g e n e r i c c h aracter i s t i c s : 1 ) v e r t e x o f t h e h e a d ( ex c a v a te d , f l a t o r s w o l l en ) ; 2 ) p ro n o t u m ( w i t h or w i t h o u t t r a n sv e rs a l i m p r es s i on ) ; 3 ) en l a r g e d c l a w ( w i t h or w i t ho u t s u b a p i c a l toot h ; w i t h o r w i t h o u t s m a l l tooth a t t h e e n d of a l on g i t u d i n a l f u rrow ; w i t h l ame l l ae or b r i s t l es or p e g - I i k e h a i rs ; 4 ) n u m b e r of p a l p a l s e g m e n t s ( co n s i dered con s t a n t i n each g e n u s ) . A f t e r m y rev i s i o n of t h e wor l d spec i es a n d after a s t u d y of a b o u t 2 0 , 000 s p e c i m e n s f r o m e v e r y cou n t r y , some o f t h e a b o v e c h a ra c t e r i s t i cs are con s i dered l es s i mpor t a n t for gener i c t a xonom y . 1 ) Vertex of the hea d . Conc a v i t y a n d convex i t y are often s u b j ec t i v e , b ec a u s e t h e y d e p e n d o n t h e pos i t i o n of t h e h e a d u n der observ a t i o n . A t f i rs t I thou g h t o f fi x i n g a h e a d pos i t i on , s o t h a t t h e observ a t i on wou l d b e c o n s t a n t . I p roposed a l w
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u n de r t h e a bo v e l i ne ; s wo l l en i f t h e pos t e r i or oce l l i were abov e ; f l a t i f t h e pos t e r i o r oce l l i were o n t h e above l i n e . I observ ed , howe v e r , m a n y i n termed i a t e cases , ra n g i n g from v e r y exca v a ted t o s l i g h t l y exca v a ted , to f l a t o r w e a k l y con v e x . As t he re are i n termed i a te c a se s of conca v i t y a n d con v ex i t y , t h e v e r t e x c a n n o t b e con s i dered a g e n e r i c c h a racter i s t i c .

2 ) Pronolum . T he p resence o r a b sence o f a t r a n s versa l i m p ress i o n o n t h e p rono t u m h a s u s u a I I y b e e n con s i d e red a d e p e n d a b I e g e n e r i c c ha racter i s t i c . I , howeve r , observed n u merous spec i me n s w i t h v e r y f a i n t t r a n s v e rs a l i m p ress i on s , demo n s t r a t i n g t h e e x i s te n c e o f i n termed i a te forms . I d i d not f i n d t hese i n termed i a t e forms to be con t i n uo u s , b u t noted a def i n i t � d i s t i nc t i o n b e t w een genera w i t h s t ron g t r a n s ve rs a l i mpress i on s a n d t hose w i t hou t o r w i t h o n l y v e r y f a i n t i m p ress i o n s . T h i s d i s t i nc t i on i s u s u a l l y c l ea r l y v i s i b l e . Accor d i n g to t h e r u l es of l og i c , t h e p resence o r a bsence of a t r a n s v e rsa l i m p ress i o n on the p ro n o l u m shou l d not b e r e g a r d e d as a sou n d g e n e r i c c h a r a c te r i s t i c , s i nce t h e e x i s t ence o f i n termed i a te c a s e s l e a d s to u n cer t a i n t y i n t h e c l as s i f i c a t i on of cert a i n spec i me n s . T he exc l u s i o n o f t h i s fea t u re o f t h e p ro n o t u m a s a gener i c c h a r a c te r i s t i c wou l d l e a d to on l y t w o genera o f a p t e rous Gon a topod i nes ( Gonatop u s a n d Dicondyl u s ) , a con c l u s i o n not comp l e te l y j u s t i f i e d b y research a t t h e present t i me . 3 ) E n l arged c l a w . T h e p resence of a s u b a p i c a l t oo t h or i t s a b s e n ce a n d rep l acemen t w i t h a s ma l l tooth a t t h e e n d o f a l on g i t u d i n a l f u rrow i s t h e o n l y rea l g e n e r i c c h a r a c t er i s t i c i n G o n a topod i n es . I t i s con s i dered t h e b a s i s of t h e t a x on o m y . O n t h e coon t r a r y , t h e a b se n c e of teeth ( sm a I I o r s u b ap i c a l ) on t h e e n l arged c l a w c a n not b e con s i dered a g e n e r i c c h a r a c t er i s t i c , s i nce I o b s e r v ed spec i me n s w i t h o u t teeth a n d spec i me n s w i t h a s m a l l t o o t h i n t h e s a m e spec i e s ( fo r i n s t a nc e , Gon a top u s leptothorax C e b a I l o s ) . T h e t y p e of Trichogon a top u s raptor ( Fe n ton ) h a s a s ma l l tooth on t h e r i g h t e n l a rged c l a w bu t no teeth on t h e l ef t e n l arged c l a w . Genus E u c a mp tonyx R . C. L. Perk i n s is t herefore co n s i dered a j u n i o r s y n o n y m of Gona top u s L j u n g h . T he p resence of l am e l l ae , b r i s t l es or peg- l i ke h a i rs i s a l so a q u e s t i on a b l e c h a r a c t e r i s t i c , beca u se som e t i mes t he peg - l i ke h a i rs are very t h i c k a n d l e a v e dou b t a s to t he i r true i den t i t y - rea l l am e l l ae o r peg- l i ke h a i rs ? I n a d d i t i o n , t here are some s p ec i es ( fo r i n s t a nce Gon a top u s stria t u s K i effer ) w h i c h con t a i n both spec i m e n s w i t h b i g l a me l l ae a n d others w i t h on l y s l e n d e r h a i rs . T h i s v a r i ab i l i t y d i scou r a ges t h e use o f t hese fea t u res a s g e n e r i c c h a r a c te r i s t i cs . A crodon tochelys C u rrado seems t o b e a n excep t i on , b e c a u s e t h e p e g- l i ke h a i rs i n t h i s g e n u s c a n n e v e r be m i s t a k e n for l ame l l ae . 4 ) Number of pa l p a l segments . Both m y o w n s t u d i e s o f n u merous spec i me n s a n d t h e F re y t a g research s h o w t h a t t h e n u m b er of p a l pa l seg men t s i s v a r i a b l e i n t h e s a m e spec i e s . Genera l l y , t h e m a x i l l a ry p a l p i v a r y t h e m os t , w hereas t h e l a b i a l p a l p i seem more con s t a n t . B u t t h i s f e a t u re i s not re l i a b l e s i nce I h a v e see n , for i n s t a n ce , a spec i men of Gona top u s s tria t u s K i effer w i t h the r i g h t l a b i a l p a l p 2-segmen ted a n d t h e l ef t l ab i a l p a l p 3-seg m e n ted . T herefo re , t h e n u m b e r o f p a l p a l segmen ts shou l d not be

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c o n s i dered v a l u a b l e as g e n e r i c c h a r a c te r i s t i c . O n t h e o t h e r ha n d , have seen t h a t in n u merous spec i es t h e n u m b e r of pal pal seg men t s seems con s t a n t : for i n s t a n c e , Gon a top u s sepsoides West wood , the most common E u ropean spec i es , a l w a y s seems to have a p a l p a l r a t i o 5/2 . Cons i d e r i n g t he v a s t a mo u n t o f m a t er i a l t h a t I h a v e seen , o n e wou l d e x p e c t t o f i n d some v a r i a t i o n s . T he r e i s s t i l l d o u b t regard i n g t h e p a l p a l ra t i os , b u t I b e l i ev e t h a t t h ere a re some spec i es or g e n e r a t h a t h a v e a co n s t a n t p a l p a l r a t i o . F o r i n s t a n c e , i n t h e dozen s o f m a l e a n d fem a l e Haplogon a top u s t h a t I h a v e e xa m i n e d , each spec i men h a d a p a l p a l ra t i o of 2/1 . F o r t h i s rea son , I s t i l l b e l i e v e t h a t i n m a n y cases t h e n u m b e r o f p a l p a l segmen ts is a c h a ra c te r i s t i c of gen e r i c v a l ue , a n d my k e y s ref l ec t t h i s i n t erpret a t i on w h i c h shou l d ho l d t r u e u n l e ss f u t u re b reed i n g p ro v es o t herw i se . I n co n c l u s i on t here i s on l y one c h a r a c t e r i s t i c w h i c h h a s a d i s t i n c t l y g e n e r i c v a l ue i n the fema l es : t h e p resence on t he en l a rged c l a w of a s u b a p i c a l toot h , or i t s absence and rep l aceme n t w i t h a s m a l l tooth a t t h e e n d of a l o n g i t u d i n a l f u rrow . I f I p ropose t h a t t h i s i s t h e o n l y c h a r a c te r i s t i c f o r t h e gen er i c c l as s i f i c a t i o n , t h i s wou l d resu l t i n on l y t w o g e n e r a of a p terous fem a l es , Gon a top u s L j u n g h a n d D icondyl u s Ha l i d a y . I t h i n k , however , t h a t t h i s so l u t i o n i s t o o rad i c a l a n d not y e t s u ff i c i e n t l y j u s t i f i e d b y resea rc h . D i fferences a m o n g t h e spec i es w h i c h h a v e a more or l es s exca v a ted vertex can become more d i s t i n c t if a n i n dex of conca vity is adop ted . I f t h e spec i me n i s ob served from a fron t a l v i e w , w i t h the ou t l i n e of t he pos t e r i o r oce l l i v i s i b l e , t h e d i s t a nce b e t ween t h e poste r i o r oce l l i a n d t h e i m a g i n a r y s t r a i g h t l i n e jo i n i n g t h e u p p e r e d g e s of t h e eyes c a n b e con s i dered t h e index of con c a vity . For i n s t a n c e w i t h a d i s t a n c e of 0 . 1 80 m m , t h e i n d e x of conc av i t y i s 0 . 1 80 . T h e c a l c u l a t i on of t h e i n dex i s usefu l i n sep a ra t i n g cou p l e t s i n to c a t e g o r i e s w i t h " ve rtex more exc a v a t e d " or " v e r t e x l e s s e x c a v a t ed " .

N O T E S O N T H E TAXONOMY O F T H E W I N G E D F E M A L E S

T he c l a s s i c a l c l ass i f i c a t i o n of t h e w i n g e d fem a l es of the Gona topod i n e s , accord i n g to K i effer a n d R i c h a r d s , d i v i des t he m i n to t h ree gen era : Ech throdelp h a x R . C . L. Perk i n s , Nercetia K i effer and Neodryin u s R . C . L . Perk i n s . T he g en e r i c c l ass i f i c a t i o n w a s b ased m a i n l y on t h e n o t a u l i ces : a b s e n t i n Neodryin u s , presen t a n d poste r i or l y separa ted i n Mercetia , p rese n t a n d poster i or l y jo i n t i n Echthrodelph a x . A n o t h e r i ncomp l e te d i s t i n c t i o n w a s b a s ed on t h e p a l p a l formu l a : 4 / 2 i n Ech throdelph a x , 6/3 i n Neodryin u s , p a l p a l formu l a u n k now n i n Mercetia . I observed a w i de v a r i a b i l i t y i n t h e p a l p a l form u l a of Echthrodelp h a x : 3/2 , 4/2 , 5/2 , 5/3 , 6/3 . T h e refore , the p a l p a l form u l a can n o t b e co n s i dered a g e n e r i c c h a r a c te r i s t i c . I n Ech throdelp h a x , I fou n d t h a t t h e not a u l i ce s are poster i or l y separa t e d , not jo i n t . T h u s , t here c a n b e n o

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s e p a r a t i o n of Mercetia a n d Echthrodelp h a x b ased on t h e n o t a u I i ces . I n a dd i t i on , I fo u n d t h a t n o t a u l i ces a re n o t p re se n t , b u t a b s e n t i n Neodryin u s . T h e t i b i a l s p u r s c a n be con s i dered a good g e n e r i c c h a ra c t e r i s t i c . Con t ra r y t o t h e i n te r p re t a t i on o f R i c h a r d s ( 1 953 ) , t i b i a l s p u rs a l w a y s seem t o be i n t h e sequence 1 , 0 , 2 i n Neodryin u s , a n d 1 , 0 , 1 i n Echthrodelph ax . A n ew g en u s , A dry in us , i s p rop osed , bec a u se i t h a s t i b i a l s p u r s i n t h e s e q u e nc e 1 , 0 , 1 a n d i s w i t ho u t n o t a u l i ces .

N O T E S ON T H E TAXONOMY OF T H E MALES

T he t a xo n o m i c c l a s s i f i c a t i o n of m a l e Go n a t opod i n e s is e x t reme l y d i ff i c u l t bec a u s e o f t h e morp h o l o g i c u n i form i t y a m o n g t he v a r i o u s spec i e s , a n d t h e sex u a l d i morph i s m w i t h i n t h e v a r i o u s s p ec i e s . T he morp ho l o g i c u n i fo r m i t y see m s t o b e a resu l t o f a very s l ow e vo l u t i on ; t h e fem a l e s , on t h e o t her h a n d , h a v e evo l ve d i n t o very s pec i a l i zed , e a s i l y recog n i z a b l e form s . A l ac k o f d i s t i n g u i s h i n g fea t u re s i n t h e m a l es c a u ses d i ff i c u l t i e s w hen a t t e m p t i n g t o sep a r a te t h e v a r i ou s s pec i e s a n d g e n er a . T h e se x u a l d i morp h i s m m a k e s breed i n g exper i me n t s necess a r y for t he correct p a i r i n g of m a l es a n d fem a l e s a m o n g t h e d i fferen t spec i e s . T he l ac k of l a bora tory bred m a l e spec i me n s ( a n d t h e i r m o r p h o l o g i c u n i form i t y ) m a k e s a com p l e t e g e n er i c c l a ss i f i c a t i on i m poss i b l e ( to t h i s d a t e t here e x i s t a n u m b e r of genera i n w h i c h n o m a l e s p ec i men is k n ow n ) . T he ma l es of on I y fo u r g e n e r a (Neodryin u s , Echthrodelp h a x , Haplogon a top u s , E s a gon a top u s ) h a ve d ef i n i te gener i c c h a ra c t e r i s t i c s . T h e o t h e r m a l e Go n a topod i nes c a n n o t be g e n e r i c a l l y recog n i zed a n d m u s t b e c l a s s i f i e d u n de r s p ec i f i c k e y s ( dors a l process of gonoforcep s , s h a pe o f r a d i a l ce l l , n ot a u l i ces , e t c . ) . I n f ac t , t h ere h a s been very l i t t l e a d v a ncemen t i n t h e t a xonomy o f m a l e Gon a topod i nes s i nce t h e desc r i p t i on of t h e g e n u s L a beo H a l i d a y , w h i c h g ro u p ed a l l m a l e s u n der a s i n g l e g en u s . For t h i s r e a son , I h a ve d ec i de d t o l i s t on l y t h ree g e n e r a i n m y gen e r i c c l a s s i f i c a t i o n key s , a n d l i s t spec i e s u n der m y s p ec i f i c c l as s i f i c a t i on k e y s accord i n g t o zoogeog r a p h i c reg i o n . F r e y t a g ( 1 97 7 ) proposed a key to t h e m a l es of t h ree g e n e r a of N e a r c t i c Go n a topo d i nes : Gona top u s , Neogon atop u s , A gon a topu s . I t i s poss i b l e t h a t t h e key i s v a l i d for t h e N e a rc t i c m a l es , b u t i t i s n o t a l w a y s v a l i d i n t h e o t h e r zoogeo g r a p h i c reg i o n s . T h e c o l o u r o f t h e l eg s ( " l e g s b a n ded o r n o t " ) a n d t h e dorsa I proces s o f gonoforce p s gonoforceps r a t i o ( " dors a l p roces s of gonoforceps h a l f l e n g t h o f gonofo rceps " i n Gon a top u s or " n e a r l y s a m e l en g t h a s gonoforcep s " i n Neogona topus a n d Agon a top u s ) a r e c h a ra c te r i s t i cs n o t a l w a y s i n a g reemen t w i t h t h e gener i c d i fferences proposed i n t h e f e m a l e key . I n Gona topus bartletti n . s p . from P u e r to R i co , for i n s t a n c e , the fem a l e has l am e l l ae near l y s a m e l e n g t h ( a s i n Gona top u s ) , t he m a l e ' s l eg s a r e n o t b a n de d ( a s i n Neogo n a top u s ) a n d t h e dors a l process of gonoforceps i s n e a r l y t h e

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same l e n g t h s a s the gonoforceps ( a s in Neogon a top u s ) . I n Gon a topus zealandic u s , a new spec i e s from New Zea l a n d , the fem a l e has l a me l l ae n e a r l y the s a m e l e n g t h ( a s i n Gon a top u s ) a n d the m a l e ' s l e gs are not b a n de d ( as in Neogon a top u s ) . A no t h e r c h a racter i s t i c proposed b y F re y t a g ( 1 97 7 ) i s the p resence or absence of a n o t c h i n t h e b a s i vo l se l l a ( " w i t h n o t c h " i n Neogon a topus a n d Agona top u s ; " w i t h o u t n o t c h " i n Gona top u s ) . I n Gona top u s l u n a t u s K l u g from t h e P a l a e a rc t i c reg i o n , however , the fem a l e h a s l a me l l ae nea r l y t h e s a me l e n g t h ( a s i n Gon a topus ) a n d the m a l e h a s a d i s t i nc t n o t c h ( as i n Neogon a top u s ) I n co n c l u s i o n , a l l t h e c h a r a c te r i s t i cs p roposed b y F re y t a g ( 1 97 7 ) for t h e gener i c c l a ss i f i c a t i o n o f t h e m a l es d o n o t a l w a y s a g ree w i t h t h e gen e r i c d i fferences p roposed i n t h e fema l e k e y . A n o t her fem a l e c h aracter i s t i c proposed b y Frey t a g ( 1 97 7 ) ( " p resence o r a b sence of l on g setae " ) i s v e r y v a r i a b l e . I h a v e seen some fema l e spec i me n s o f Gona top u s l u n a t u s K l u g w i t h l o n g s e t a e a n d o t h e r fem a l e s p ec i me n s b a re . At f i rs t s i g h t a v a l i d c h a racter i s t i c ( a l re a d y p roposed b y R i c h a r d s 1 939 ) seems the l e n g t h - b rea d t h ra t i o of a n t e n n a I segmen t 3 ( " a n ten n a l seg m e n t 3 fo u r t i mes o r more t h a n fo u r t i mes a s l o n g a s b ro a d " i n m a l es b e l on g i n g to genera p a r a s i t i z i n g Fu l goroidea ; " a n te n n a I segme n t 3 t h ree t i mes or l e s s t h a n t h ree t i mes as l on g a s b ro a d " i n m a l es b e l on g i n g to genera p a r as i t i z i n g Cicadelloidea ) . T h i s c h a racter i s t i c , how ever , is on l y s p ec i f i c , b e c a u s e some excep t i o n s are k n ow n : for i n s t a n ce , the N e a rc t i c Gona top us agropyrus F e n to n , p a r as i t i z i n g Cic a delloidea , h a s a n t e n n a I segme n t 3 fo u r t i mes or more t h a n fo u r t i mes as l o n g as b roa d . •

K E Y T O T H E G E N E RA OF GONA TOPODINA E

F E MALES

Sp e c i e s w i t h r ed u c e d w i n g s ( F i g s 1 2 5 0 , 1 2 5 1 ) . . . . . . . . . . . . . . . . . . . 1 8 . Gynochelys B ru e s Sp e c i e s a p t e r ou s ( F i g s 7 9 8 , 7 9 9 , 8 0 1 , 9 8 5 ) 2 S p e c i e s f u l l y w i n g ed ( F i g s 7 1 5 , 7 1 6 , 7 5 4 , 7 5 5 , 7 6 7 , 7 6 8 ) 1 6 E n l a rg ed c l a w w i t h ou t S U b a p i c a l t o o t h , w i t h o r w i t h o u t s ma l l t o o t h a t t h e e n d o f a l on g i t u d i n a l f u r r ow ( F i g s 9 9 3 , 1 0 2 0 , 1 04 0 ) 3 E n l a r g ed c l a w w i t h s u b ap i c a l t o o t h ( F i g s 7 8 6 , 8 0 3 ) 8 P r on o t u m c r o s s ed b y a s t r on g t ra n s v e r s e i mp r e s s i on ( F i g s 1 0 3 8 , 1 066 ) 0 4 P r o n o t u m n o t c r o s s ed b y a t ra n s v e r s e i mp r e s s i on o r w e a k l y i mp r e s s ed ( F i g . 9 9 2 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 /le x i I l a ry p a l p i w i t h 2 s eg me n t s •

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Epigon a top u s R . C . L . P e rk i n s

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Max i I l a ry p a l p i w i t h 3 - 6 s egmen t s 5 Ma x i I l a ry p a l p i w i t h 3 - 5 s egmen t s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 7 . Gon a top u s L j u n g h Max i I l a ry p a l p i w i t h 6 s e g men t s 6 La b i a l p a l p i w i t h 2 s egmen t s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . • 1 6 . E s agona top u s n . g en . La b i a l p a l p i w i t h 3 s egmen t s 1 7 . Gon a top u s L j u n g h L a b i a l p a l p i w i t h 3 s egmen t s 14 Trichogon a top u s K i e f f e r L ab i a l p a l p i w i t h 2 s egmen t s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 . Tetrodontochelys R i c h a rd s P r on o t u m n o t c r o s s e d by a t ra n sv e r s e i mp r e s s i on o r weak l y i mp r e s s e d ( F i g . 9 4 8 ) 9 P r o n o t u m c r o s s ed by a s t r on g t ra n s v e r s e i mp r e s s i on ( F i g s 7 9 9 , 895 ) 12 L a b i a l p a l p i w i t h 1 s egmen t . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 0 . Haplogon a topu s R . C . L . P e r k i n s L a b i a l p a l p i w i t h 2 - 3 s egmen t s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 0 L ab i a l p a l p i w i t h 3 s egmen t s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 . Plectrogon a topoides P o n oma r e n k o La b i a l p a l p i w i t h 2 s egmen t s . . . . . . . . . . . . . . . .1 1 Max i I l a r y p a l p i w i t h 5 s e g men t s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 . Pentagon a top u s n . g e n . Ma x i I l a r y p a l p i w i t h 2 - 4 s egmen t s 9 . Dicondylus H a I i d a y E n l a rg ed c l a w w i t h o u t l a me l l a e , w i t h o r w i t h ou t b r i s t l e s o r p eg - l i k e h a i r s ( F i g . 7 8 6 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 . A crodon tochelys C u r r a d o E n l a rg ed c l a w w i t h l a me l l a e ( F i g . 8 0 3 ) 13 La b i a l p a l p i w i t h 2 s egmen t s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 4 L a b i a l p a l p i w i t h 3 s egmen t s 15 Ma x i I l a ry p a l p i w i t h 2 - 4 s egmen t s . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 . Pseu dogon a top u s R . C . L . P e r k i n s Max i I l a ry p a l p i w i t h 5 s e gmen t s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6 . Don isthorpina R i c h a rd s Max i I l a r y p a l p i w i t h 6 s egmen t s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 . Ap terodryin u s R . C . L . P e rk i n s Ma x i I l a r y p a l p i w i t h 5 s e gmen t s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 . Agon a topoides R . C . L . P e rk i n s T i b i a l s p u r s 1 , 0 , 2 . . 1 . Neodryin us R . C . L . P e rk i n s T i b i a I spu r s 1 , 0 , 1 17 N o t a u l i c e s c omp l e t e a n d d i s t i n c t ( F i g . 7 6 7 ) . . . . . . . . . . . . . . . . . . 3 . Ech throdelph a x R . C . L . P e rk i n s N o t a u l i c e s i nv i s i b l e ( F i g . 7 5 4 ) 2 . A dryinus n . g e n . •

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1 057

Gona topod i n a e

r�A L E S

2

T e mp l e s a b s en t ( F i g . 7 1 7 ) ; o c c i p u t s t ra i g h t ( F i g . 7 1 7 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 . Neodryin u s R . C . L . P e rk i n s T emp l e s p r e s e n t ( F i g s 7 7 9 , 8 3 6 ) ; o c c i p u t c o n c a v e ( F i g s 7 7 9 , 836 ) 2 H e a d w i t h a d i s t i n c t o c c i p i t a l c a r i n a f r om p o s t e r i o r o c e l l i t o t h e v en t ra l s i d e o f t h e h ea d ( F i g . 7 7 9 ) . . . . . . . . . . . . . . . . . . . 3 . E c h th rodelp h a x R . C . L . P e rk i n s H ea d w i t h o u t o c c i p i t a l c a r i n a o r w i t h o c c i p i t a l c a r i n a v e r y s h o r t , v i s i b l e on l y b eh i n d t h e o c e l l i o r on t h e s i d e s o f t h e p o s t e r i o r o c e l l i , b u t n ev e r r e a c h i n g t emp l e s ( F i g . 8 3 6 ) . . . . . . 3 L a b i a l p a l p i w i t h 1 s egmen t . . . . . . . . . . . . . . . . . . . 1 0 . Haplogona topus R . C . L . P e rk i n s . . . . . . . . . . . . 4 L a b i a l p a l p i w i t h 2 - 3 s eg m e n t s . P a l p a l ra t i 0 6 / 2 1 6 . Esagona top u s n . g e n . Pa I pa I ra t i 0 d i f f e r e n t 4 . A crodontochelys C u r rad 0 5 . Pseu dogon a top u s R . C . L . P e r k i n s 6 . Donisth orpin a R i c h a rd s 7 . A gona topoides R . C . L . P e rk i n s 8 . Ap terodryin u s R . C . L . P e rk i n s 9 . Dicondyl u s Ha I i d a y 1 3 . Tetrodon tochelys R i c h a r d s 1 7 . Gona topus L j u n g h •

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T he m a l es of Gynochelys Brues , Epigon a top u s R . C . L . Perk i n s , Trichogon a top u s K i effer , Pen tagon a top u s n . g en . , Plectrogona topoides Ponom a r e n k o , A dryin u s n . gen . a re u n k n ow n .

S P E C I F I C K EYS T O T H E KNOWN MAL E S O F GONA TOPODINAE

PALAEARCT I C R E G I ON

A n t e n n a l s egmen t 3 mo r e t h a n f o u r t i m e s a s l on g a s b r oad . . . . . 2 A n t e n n a l s eg m e n t 3 l e s s t h a n t h r e e a n d a h a l f t i me s a s l on g a s b road 15 H e a d w i t h a d i s t i n c t o c c i p i t a l c a r i n a f r om p o s t e r i o r o c e l l i t o t h e v en t r a l s i d e o f t h e h ea d ( g en u s Echthrodelph a x R . C . L . P e rk i n s ) ( F i g . 7 7 9 ) 3 H e a d w i t h ou t o c c i p i t a l ca r i n a o r w i t h o c c i p i t a l ca r i n a v e ry s h o r t , on l y v i s i b l e b eh i n d t h e o c e l l i o r on t h e s i d e s of t h e p o s t e r i o r o c e l l i , b u t n ev e r r e a c h i n g t emp i e s ( F i g . 8 3 6 ) 4 •

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1 058

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Mem .

Do rsa l

p ro c e s s . . . Do rsa l p ro c e s s (F i g . 781 ) L ab i a l p a l p i w i P e rk i n s ) Lab i a l pa l p i w i Dorsa l p ro c e s s ·

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l on g a s b r e a d t h o f t h e o c e l l i . . . . . . . . . Pseu dogona top u s fulgori ( N a k a g awa ) H e a d w i t h OOL n ea r l y t w i c e o r mo r e t h a n t w i c e a s l on g a s b r ead t h o f t h e oc e l l i . . . . . . . . . . . 10 D o r s a l p r o c e s s o f g on o f o r c ep s w i t h a p i ca l a n d i n n e r ma r g i n s e r ra t e ( F i g . 8 1 8 ) · Pseu dogonatop u s distin c t u s ( K i e f f e r ) D o r s a l p r o c e s s o f g on o f o r c e p s w i t h a p i c a l a n d i n n e r ma r g i n n o t s e r ra t e ( F i g s 9 0 1 , 9 0 3 , 9 9 5 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 D o r s a l p r o c e s s o f g on o f o r c e p s s l e n d e r a n d w i t h a p e x p o i n t ed ( F i g s 9 0 1 , 9 0 3 , 946 , 995 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 D o r s a l p r o c e s s o f g on o f o r c ep s w i t h a p ex b r o a d en e d ( F i g . 946 ) 14 D o r s a l p r o c e s s o f g on o f o r c ep s mu c h s h o r t e r t h a n g o n o f o r c ep s (F i g . 901 ) A gona topoides solidus ( H a u p t ) D o r s a l p r o c e s s of g on o f o r c ep s n ea r l y a s l on g o r s l i g h t l y s h o r t e r t h a n g on o f o r c ep s ( F i g s 9 0 3 , 9 9 5 ) 13 F o r e - w i n g w i t h ra d i a l v e i n r eg u l a r l y c u r v ed . . . . . . . . . . . . . . . . . · . A gona topoides can ariensis n . s p . .

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t h 2 - 3 s eg m e n t s o f g o n o f o r c ep s l on g a n d

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i n n e r ma rg i n e s e r ra t e ( F i g . 9 7 4 ) · !!aplogona top u s oratorius ( W e s t wood ) D o r s a l p r o c e s s o f g o n o f o r c ep s w i t h a p i c a l a n d i n n e r ma r g i n s e r ra t e ( F i g s 9 7 7 , 9 8 0 ) 6 D o r s a l p r o c e s s o f g o n o f o r c e p s b r oad en ed n e a r t h e a p e x ( F i g . 980 ) . . liaplogon a top u s apicalis R . C . L . P e rk i n s D o r s a l p r o c e s s o f g on o f o r c e p s n o t b r oad e n e d n e a r t h e a p ex ( F i g . 977 ) . Haplogon a top u s a tr a t u s E s a k i a n d Ha s h i mo t o D o r s a l p r o c e s s o f g o n o f o r c ep s v e r y s h o r t ( F i g s 8 0 8 , 8 9 3 ) ; g on o f o r c e p s mo r e t h a n t h r e e t i me s a s l on g a s d o r sa l p r o c e s s ( F i g s 808 , 893 ) 8 D o r s a l p r o c e s s o f g on o f o r c ep s v e ry l on g ( F i g s 8 0 4 , 8 1 8 ) ; g on o f o r c e p s l e s s t h a n t h r e e t i m e s a s l on g a s d o r sa l p r o c e s s ( F i g s 804 , 8 1 8 ) 9 H e a d w i t h f r o n s d u l l , w i t h ou t a med i a n d e ep s h i n y f u r r ow ; d o r s a l p r o c e s s o f g on o f o r c e p s s l e n d e r ( F i g . 8 0 8 ) . . . . Pseu dogon a top u s fl a vifem ur E sak i a n d H a s h i mo t o · . . . H e a d w i t h f r o n s sh i n y , w i t h a med i a n d e e p s h i n y f u r r ow ; d o r s a l p r o c e s s o f g o n o f o r c ep s w i t h a p ex b r oad e n ed ( F i g . 8 9 3 ) . . . . . . . . . · . . Donisthorpina p a llida ( C e b a I I o s ) •

7

1 984

o f g o n o f o r c ep s t r a n s v e r s e ( F i g . 7 7 0 ) . Echthrodelp h a x hortusen sis ( Ab d u I - N o u r ) o f g o n o f o r c ep s n ea r l y p a ra l l e l t o t h e p en i s Echthrodelp h a x fa irchildii R . C . L . P e rk i n s t h 1 s eg men t ( g en u s Haplogon a top u s R . C . L .

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G o n a topod i n a e

1 059

F o r e -w i n g w i t h rad i a l v e i n n o t r e g u l a r l y c u r v ed ; ra d i a l v e i n w i t h p r ox i ma l a n d d i s t a l p a r t s f o rm i n g an a n g l e · Tetrodon tochelys pedestris ( D a I man ) 1 4 M i n i mu m d i s t a n c e b e t w e en t h e n o t a u l i c e s n ea r l y a s l on g a s t h e b r ead t h o f t h e o c e l I i .

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Dicondyl u s bicolor ( H a I i d a y ) M i n i mu m d i s t a n c e b e t w e en t h e n o t a u l i c e s s h o r t e r t h a n t h e b r ead t h o f t h e o c e l I i · Dicondyl u s dichrom u s ( K i e f f e r ) P r op o d e u m smo o t h , s h i n y , n o t r e t i c u l a t e ru g o s e A crodontochelys bouceki C u r r a d 0 · P r op o d eu m d u l l , r e t i cu l a t e r u g o s e 16 M i n i mu m d i s t a n c e b e t w e e n t h e n o t a u l i c e s n e a r l y a s l on g a s a n t en n a l s egmen t 2 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . · Gon a top u s distingu endus K i e f f e r M i n i mu m d i s t a n c e b e t we en t h e n o t a u l i c e s mu c h s h o r t e r t h a n t h e l en g t h o f a n t en n a l s eg men t 2 17 D o r s a l p r o c e s s o f g on o f o r c ep s n ea r l y a s l on g a s g on o f o r c ep s ( F i g s 1 0 4 9 , 1 2 5 5 ) ; ra d i a l c e l l w i d e r ( F i g . 1 0 4 8 A ) 18 D o r s a l p r o c e s s o f g on o f o r c ep s mu c h s h o r t e r t h a n g on o f o r c e p s ( F i g s 1 0 7 1 , 1 0 9 5 ) ; r a d i a l c e l l s ma l l e r ( F i g . 1 0 4 8 B , C ) 19 C l y p eu s a n d f r o n s b l a c k ; d o r s a l p r o c e s s o f g on o f o r c ep s s l en d e r ( F i g . 1 04 9 ) Gona top u s stria t u s K i e f f e r C l y p eu s , a n t e r i u r a n d med i a n r e g i o n o f f r o n s , l a t e ra l r eg i on s o f f r on s a r o u n d t h e o rb i t s y e l l o w- t e s t a c e o u s ; d o r sa l p r o c e s s o f g on o f o r c ep s b r o a d e n e d a n d w i t h s ome a p i c a l pap i I l a e ( F i g . 1 2 5 5 ) . . . . . . . . . . . . . L a beo a lbosign a t u s K i e f f e r ( s e e n om i n a d u b i a ) Ba s i v o l s e l l a w i t h r i d g e ( F i g . 1 0 7 1 ) · Gona top u s l u n a t u s K I u g Ba s i v o l s e l l a w i t h o u t r i d g e ( F i g . 1 0 9 5 ) Gon a top u s sepsoides W e s t w o od ·

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ET H I OP I AN REG I ON

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Gona top u s incogn itus n . s p . N o t a u l i c e s a t l ea s t p a r t l y p r e s e n t 2 N o t a u l i c e s c omp l e t e 3 N o t a u l i e e s i n c omp l e t e 6 An t en n a l s eg men t 3 l e s s t h a n t h r e e t i m e s a s l on g a s b r o a d ; h ea d , b e t w e en p o s t e r i o r o c e l l i a n d e y e s , w i t h a smo o t h , s h i n y , v e ry w i d e a r e a , a n t e r i o r l y d e l i m i t ed b y a s t r o n g , h o r n - s h a p ed keel ·

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1 060

Mem.

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Pseu dogon a top u s corn u t u s B e n o i t ( n o m i n a d u b i a ) A n t en n a l s eg m e n t 3 mo r e t h a n t h r e e t i me s a s l on g a s b r oa d ; h ea d , b e t we en p o s t e r i o r o c e l l i a n d e y e s , w i t h o u t smo o t h , s h i n y , v e ry w i d e a r ea , a n t e r i o r l y d e l i m i t ed b y a s t r o n g h o rn - s h a p ed k e e l 4 L a b i a l p a l p i w i t h 1 s eg me n t . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . •

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Haplogon a top us k a t a ngae ( B e n o i t ) L a b i a l p a l p i w i t h 2 - 3 s eg m e n t s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . S S c u t u m w i t h n u m e r o u s , l on g i t u d i n a l , w e a k s t r i a e ; m i n i mu m d i s t a n c e b e t w e e n t h e n o t a u l i c e s l on g e r t h a n b r ead t h o f t h e o c e l l . i . . . . . . . . . . . . Neodryin u s ( 7 ) bellicosus ( 7 ) B e n o i t ( s e e A dryin u s bellicosu s ) S c u t um w i t h ou t l on g i t u d i n a l s t r i a e ; m i n i mu m d i s t a n c e b e t w e en t h e n o t a u l i c e s s h o r t e r t h a n b r ead t h o f t h e o c e l I i . . . . . . . . . . . . Pseu dogona topus ruens n ew n ame N o t a u l i c e s r e a ch i n g a p p r ox i ma t e l y 0 , 7 5 l en g t h of s c u t u m . . . . . . . . . . . . . . . . . . . . . Gon a top u s inexpectatus B e n o i t ( n o m i n a d u b i a ) N o t a u l i c e s r e a c h i n g a p p r ox i ma t e l y O , S l en g t h o f s c u t u m 7 D i s t a l p a r t o f rad i a l v e i n n ea r l y a s l on g a s p r ox i ma l p a r t ; p r op od eu m w i t h d o r s a l s u r f a c e n ea r l y a s l on g a s p o s t e r i o r s u r f a c e ; f o r e -w i n g w i t h o n l y c o s t a l c e l l c l ea r l y e n c l o s ed b y p i g m e n t ed v e i n s Aph elop u s afric a n u s B e n o i t ( n om i n a d u b i a ) D i s t a l p a r t o f ra d i a l v e i n mu c h l on g e r t h a n p r ox i ma l p a r t ; p r op od e u m w i t h d o r s a l s u r f a c e n ea r l y t w i c e a s l o n g a s p o s t e r i o r s u r f a c e ; f o r e -w i n g w i t h t h r e e c e l l s c l ea r l y e n c l o s ed b y p i g me n t ed v e i n s . A crodontochelys uga n da n us ( B en o i t ) •

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OR I E NTAL REG I ON

T emp i e s a b s en t ;

o c c i p u t s t ra i g h t . . . . . . . . . . . . . . . . . . . . . . . . . . . . Neodryin u s j a v a n u s R i c h a rd s T emp l e s p r e s en t ; o c c i p u t c o n c a v e 2 H e a d w i t h a d i s t i n c t o c c i p i t a l ca r i n a f r om p o s t e r i o r o c e l l i t o t h e v e n t ra l s i d e o f t h e h e a d . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Echthrodelphax fa irchildii R . C . L . P e rk i n s H ea d w i t h ou t o c c i p i t a l ca r i n a o r w i t h o c c i p i t a l ca r i n a v e r y s h o r t , o n l y v i s i b l e b eh i n d t h e o c e l l i o r on t h e s i d e s o f t h e p o s t e r i o r o c e l l i , b u t n ev e r r e a c h i n g t emp l e s . . . . . . . . . . . . . . . . . 3 L a b i a l p a l p i w i t h 1 s eg me n t . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Haplogo n a topus api c a l is R . C . L . P e rk i n s L a b i a l p a l p i w i t h 2 - 3 s eg m e n t s . . . 4 N o t a u l i c e s c omp l e t e a n d s ep a ra t ed 5 N o t a u l i c e s c omp l e t e a n d j o i n t ( o r v e ry n ea r , a l mo s t j o i n t ) . . . 8 D o r s a l p r o c e s s o f g o n o f o r c e p s l on g e r t h a n p en i s ( F i g s 9 1 4 , .

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Gon a topod i n ae

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91 7) 6 D o r s a l p r o c e s s o f g on o f o r c ep s s h o r t e r t h a n p en i s ( F i g s ' 8 4 1 , 878 ) 7 H ea d f u l l y y e l l ow ; a n t e n n a l s eg m e n t 3 mo r e t h a n f o u r t i me s a s l on g a s b r oad ; d o r s a l p r o c e s s o f g on o f o r c ep s w i t h a p e x s h a rp •

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(F i g . 91 4 ) A gona topoides born e a n u s n . sp . H ea d b r o wn ; a n t e n n a l s eg m e n t 3 l e s s t h a n t h r e e t i me s a s l o n g a s b r o a d ; d o r s a l p r o c e s s o f g on o f o r c e p s w i t h a p ex r o u n d ed (F i g . 9 1 7 ) Agona topoides pyrill a e ( Ma n i ) D o r s a l p r o c e s s o f g on o f o r c ep s w i t h d i s t a l h a l f b r oad en ed ; p r o x i ma l h a l f s l e n d e r ( F i g . 8 7 8 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Pseudogon a top us hospes R . C . L . P e rk i n s D o r s a l p r o c e s s o f g o n o f o r c ep s a l mo s t f u l l y r eg u l a r l y b r oa d e n ed (F i g . 841 ) T y p e B o f Pseudogona top u s m a lesiae n . sp . o r s a rawakensis n . sp . ( s e e t e x t ) P r o p o d e u m w i t h d o r s a l s u r f a c e s c u l p t u r ed by a r e o l a e mu c h s ma l l e r t h a n a r e o l a e o f t h e p o s t e r i o r su r f a c e . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Pseudogona topus s a ra ,,'aki Mo c za r P r o p o d e u m w i t h d o r s a l su r f a c e s c u l p t u r ed b y a r e o l a e n e a r l y a s w i d e a s a r e o l a e o f t h e p o s t e r i o r su r f a c e 9 D o r s a l p r o c e s s o f g o n o f o r c ep s v e r y l o n g a n d b r oad , ea s i I y •

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T y p e A o f Pseu dogo n a top u s m a lesiae n . sp . o r sara,,'akensis n . sp . ( s e e t ex t ) D o r s a l p r o c e s s o f g on o f o r c e p s v e ry r ed u c ed , a l mo s t i n v i s i b l e . . . . . . . . . . . . . . . . . . . . Dicondyl u s india n u s n . sp . •

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NEARCT I C R E G I ON

o c c i p u t s t ra i g h t . . . Neodryin u s typh locyb a e ( A s h m e a d ) T emp l e s p r e s e n t o c c i p u t c o n Ca v e An t e n n a I s e gmen t 3 l e s s t h a n t h r e e t i m e s a s l o ng a s b r oad A n t e n n a l s egmen t 3 mo r e t h a n t h r e e t i me s a s l on g a s b r oad . P r o p o d e u m smo o t h , s h i n y , n o t r e t i c u l a t e r u g o s e . . . . . . . . . . . L a beo a l b i t arsis K i e f f e r ( s e e n om i n a d u b i a . . . . P r op od e u m d u l l , r e t i c u l a t e ru g o s e . . . . . . . . . . N o t a u l i c e s c omp l e t e , p o s t e r i o r l y s ep a r a t ed . . . . . . . . . . . . . . . . T e mp I e s a b s en t ;

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N o t a u l i c e s i n c omp l e t e D o r s a l p r o c e s s o f g on o f o r c ep s s l e n d e r ( F i g . 1 1 8 8 ) . . . . . . . Gona top u s herb arum ( R . C . L . P e r k i n s ) D o r s a I p r o c e s s o f g on o f o r c ep s b r o a d en ed ( F i g . 1 0 3 6 ) . . . . . . . . . . . . . E s a gona top u s niger ( F e n t o n ) N o t a u l i c e s r e a c h i n g app r o x i ma t e l y 0 , 8 0 l e n g t h o f s c u t u m ; .

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p r o c e s s o f g o n o f o r c ep s l on g a n d s l e n d e r ( F i g . 1 2 6 5 A ) . . . Embolemus n a s u tu s ( A s h m e a d ) ( s e e n om i n a d u b i a ) .

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N o t a u l i c e s o n l y v i s i b l e o n p r ox i ma l h a l f o f s cu t u m ; d o r sa l p r o c e s s o f g on o f o r c e p s r ed u c ed a n d l ob e s h a p e d ( F i g . 1 0 1 5 ) 7 8

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Tetrodontochelys unicus ( R . C L . P e rk i n s ) P r op od e u m f u l l y smo o t h , s h i n y , n o t r e t i c u l a t e r u g o s e . . . . . . . . 8 P r op od eu m d u l I , r e t i c u l a t e r u g o s e 9 N o t a u l i c e s i n c o mp l e t e , o n l y v i s i b l e on p r ox i ma l h a l f o f s c u t u m ; s c u t u m d u l l , g ra n u l a t ed · Gona top u s cyphonotus B ra d l ey N o t a u l i c e s c omp l e t e , p o s t e r i o r l y s e p a ra t ed ; s c u t u m s h i n y , w i t h ou t s c u l p t u r e . A crodontochelys m im u s ( R . C . L . P e rk i n s ) N o t a u l i c e s c o mp l e t e a n d j o i n t . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Dicondy l u s americ a n u s ( R . C . L . P e rk i n s ) N o t a u l i c e s c o mp l e t e a n d mo r e o r l e s s s ep a r a t ed . . . . . . . . . . . 10 D o r s a l p r o c e s s o f g on o f o r c ep s mu c h s h o r t e r t h a n g o n o f o r c ep s ( F i g s 860 , 863 , 1 1 95 , 1 264 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 D o r s a l p r o c e s s o f g on o f o r c ep s n ea r l y a s l on g a s g on o f o r c ep s (F i gs 857 , 1 1 80 ) 14 D o r s a l p r o c e s s o f g on o f o r c ep s v e ry r e d u c ed ( F i g s 8 6 0 , 8 6 3 ) 12 D o r s a l p r o c e s s o f g o n o f o r c ep s l on g e r ( F i g s 1 1 9 5 , 1 2 6 4 ) . . . . . . 1 3 T h o ra x a n d p r op od eu m b l a c k . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Pseu dogo n a top u s stenocra n i R . C . L . P e rk i n s T h o ra x a n d p r o p od e u m b r own - r e d d i sh Pse u dogona top u s arizonicus R . C . L . P e rk i n s D o r s a l p r o c e s s o f g o n o f o r c ep s w i t h a p ex r o u n d ed ( F i g . 1 1 9 5 ) . . . · . . . . . . . . . . Gon a top u s a shmeadi K i e f f e r D o r s a l p r o c e s s o f g o n o f o r c ep s w i t h a p ex i r r e g u l a r l y t ru n c a t ed ( F i g . 1 2 6 4 ) . . . . . L a beo longitarsis A s h m e a d ( s e e n om i n a d u b i a ) D o r s a l p r o c e s s o f g o n o f o r c ep s w i t h med i a n a n d d i s t a l r eg i o n b r o a d e n ed ( F i g s 8 5 4 , 8 5 7 , 1 2 5 7 , 1 2 5 8 ) . . . . . . . . . . . . . . . . . . . . . . . 1 5 D o r s a l p r o c e s s o f g on o f o r c ep s f u l l y s l e n d e r o r on l y w i t h d i s t a l r eg i on b r o a d e n ed ( F i g s 1 0 1 3 , 1 1 8 0 , 1 1 9 7 , 1 2 7 0 ) . . . . . . . 1 8 D o r s a l p r o c e s s o f g on o f o r c ep s w i t h a p e x p o i n t ed ( F i g . 8 5 7 ) . · . . . . . . . . . . . . Pseu dogon a top u s sjoestedti ( K i e f f e r ) D o r s a l p r o c e s s o f g on o f o r c ep s w i t h a p e x r o u n d ed ( F i g s 8 5 4 , 1 25 7 , 1 25 8 ) 16 D o r sa l p r o c e s s o f g on o f o r c ep s w i t h d i s t a l r e g i on w i t h o u t p a p i l l a e a n d w i t h d i s t a l ma r g i n n o t s e r r a t e ( F i g . 1 2 5 7 ) . . . . . . . . . . . . . . . . . . . . . . . . . Neogona top u s ie n t om' F r ey t a g ( n om i n a d ub i a ) D o r s a I p r o c e s s o f g on o f o r c ep s w i t h d i s t a I r eg i o n c ov e r ed w i t h p ap i l l a e a n d w i t h d i s t a l ma r g i n s e r ra t e ( F i g s 8 5 4 , 1 2 5 8 ) . . . . 1 7 N o t a u l i c e s n ea r e r , a l mo s t j o i n t . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Pseu dogon a top u s c an a densis n . sp . N o t au l i ce s l es s n e a r · Neogon a top u s gra n dis F r ey t ag ( n o m i na d u b i a ) D o r s a l p r o c e s s o f g o n o f o r c ep s w i t h d i s t a l r eg i on v e r y b r o a d en ed a n d w i t h d i s t a l ma r g i n s l i g h t l y s e r ra t e ( F i g . 1 1 9 7 ) . Gona topus ombrodes ( R . C . L . P e rk i n s ) ·

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D o r s a l p r o c e s s o f g o n o f o r c ep s f u l l y s l e n d e r or a t mo s t w i t h d i s t a l r e g i on s l i g h t l y b road e n e d { F i g s 1 0 1 3 , 1 1 8 0 , 1 2 7 0 ) . . . . 1 9 1 9 R a d i a l v e i n r e g u l a r l y c u r v ed { F i g . 1 1 7 8 ) · . . . . . . . . . . Gon a top u s a gropyrus F en t o n R a d i a l v e i n w i t h d i s t a l a n d p r ox i ma l p a r t s f o rm i n g a n a n g l e {F i g . 1 269 ) 20 2 0 D o r s a l p r o c e s s o f g o n o f o r c ep s w i t h a p e x p o i n t ed { F i g . 1 2 7 0 ) .

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N E OTROP I C R E G I ON

T emp l e s a b s e n t ; o c c i p u t s t ra i g h t · . . . . . . . . . . Neodry i n u s trin ita tis R i c h a rd s T emp l e s p r e s e n t , m o r e o r l e s s p r o m i n e n t ; o c c i p u t c o n c a v e 2 A n t e n n a l s eg m e n t 3 I e s s t h a n t h r e e t i m e s a s l on g a s b r o a d · . . . . . . . . . L a beo s a n ctivincenti A sh m e a d ( n om i n a d u b i a ) A n t en n a l s eg m e n t 3 m o r e t h a n t h r e e t i me s a s l on g a s b r oad 3 P r o p o d e u m f u l l y r e t i cu l a t e ru g o s e 4 P r op od eu m a t mo s t p a r t l y r e t i c u l a t e ru g o s e , f u l l y o r p a r t l y s m o o t h , s ome t i me s r u g o s e 6 D o r s a l p r o c e s s o f g on o f o r c ep s l on g e r t h a n p en i s { F i g . 1 2 6 6 ) L a berin u s p aranen sis Og I o b l i n ( n om i n a d u b i a ) D o r s a l p r o c e s s o f g o n o f o r c e p s s h o r t e r t h a n p en i s ( F i g s 9 3 9 , 1 25 9 ) 5 D o r sa l p r o c e s s o f g on o f o r c ep s w i t h a p e x s l i g h t I y b r o a d en ed {F i g . 939) . . . . . A p terodryin u s tij u c a n u s ( A r l e ) •

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Gon a top u s bartletti n . s p . P r o p o d e u m w i t h d o r s a I s u r f a c e f u I I Y smo o t h ; p o s t e r i o r s u r f a c e r e t i cu l a t e r u g o s e ; d o r sa l p r o c e s s o f g on o f o r c ep s w i t h a p e x mo r e b r o a d e n ed { F i g . 1 2 6 8 ) · . . . . . . L a beo sim u l a n s A s h m e a d ( s e e n om i n a d u b i a ) D o r s a l p r o c e s s o f g on o f o r c ep s mu c h s h o r t e r t h a n p en i s ( F i g . 1 20 6 ) Gona top u s fern a n din a e n . s p . D o r s a l p r o c e s s o f g on o f o r c ep s s l i g h t l y s h o r t e r t h a n p en i s {F i g . 1 20 8 ) 9 L eg s b r o wn - d a rk , w i t h d i s t a l a p e x o f c ox a e , t r o c h a n t e r s a n d f emo ra y e l l ow ; p r ox i ma l a p e x o f t i b i a e a n d t a r s i y e l l ow .

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p r o c e s s o f g o n o f o r c ep s v e r y b r o a d en ed { F i g . 1 2 5 9 ) L a beo gren a den sis A s h m e a d ( s e e n om i n a d u b i a ) N o t a u l i c e s c o mp l e t e 7 N o t a u I i c e s i n c omp l e t e 8 P r o p o d e u m n o t smo o t h , f u l l y weak l y r u g o s e ; d o r sa l p r o c e s s o f g on o f o r c e p s w i t h a p e x s l i g h t I y b r o a d en ed { F i g . 1 2 2 0 ) .

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1 064

M e m . Amer .

En!.

I ns!. ,

n o . 37 ,

1 984

. . . . . . . . . . . . . . . . . . A crodontochelys cubensis R i c h a r d s L e g s t e s t a c e o u s -d a r k , w i t h d i s t a l a p ex o f c ox a e , t r o c h a n t e r s a n d f em o r a s l i g h t I y l i g h t ; p r o x i ma l a p e x o f t i b i a e a n d t a r s i s l i gh t I y l i gh t Gona top u s breviforceps K i e f f e r .

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AUSTRA L I AN REG I ON

T emp l e s a b s en t ; o c c i p u t s t ra i g h t o r a l mo s t s t ra i g h t . . . . . . . . . . . Neodryin u s koebelei R . C . L . P e rk i n s T emp l e s p r e s en t ; o c c i p u t c o n ca v e .

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H ea d w i t h a d i s t i n c t o c c i p i t a l v en t ra l s i d e o f t h e h ea d •

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Head w i shor t , pos t er i No t au l i

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ca r i n a f r om p o s t e r i o r o c e l l i •

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t h ou t o cc i p i t a l ca r i n a o r w i t h o c c i p i t a l c a r i n a v e ry o n l y v i s i b l e b eh i n d o c e l l i o r on t h e s i d e s o f t h e o r o c e l l i , b u t n ev e r r e a c h i n g t emp i e s c e s p os t e r i o r l y j o i n t •

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. . . . . . . . . . Echthrodelp h a x fairch il dii R . C . L . P e rk i n s N o t a u l i c e s p o s t e r i o r l y d i s t i n c t l y s ep a ra t ed . Ech throdelp h a x nigricollis R . C . L . P e rk i n s 4 L a b i a l p a l p i w i t h 1 s eg m e n t . . . . . . . . . . . Haplogona top u s vitie n s is R . C . L . P e rk i n s L a b i a l p a l p i w i t h 2 - 3 s e g men t s 5 5 N o t a u I i c e s i n c omp l e t e 6 N o t a u I i c e s c omp l e t e 8 6 P r op od e u m s mo o t h a n d s h i n y ; p r ox i ma l p a r t o f rad i a l v e i n mu c h s h o r t e r t ha n d i s t a I p a r t . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Gona top u s mel a n i a s ( R . C . L . P e rk i n s ) P r op od e u m r e t i c u l a t e ru g o s e , d u l l ; d i s t a l p a r t o f ra d i a l v e i n n ea r l y a s l on g a s p r ox i ma l p a r t . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 7 D o r s a l p r o c e s s o f g o n o f o r c ep s n ea r l y a s l o n g a s g o n o f o r c e p s Gona top u s zea l a n dicus n . s p . (F i g . 1 239) D o r s a l p r o c e s s o f g o n o f o r c ep s mu c h s h o r t e r t h a n g o n o f o r c ep s ( F i g . 1 246 ) 1 . . Gona top u s op t a b ilis ( R . C . L . P e rk i n s ) Gon a top u s decora t u s ( R . C . L . P e rk i n s ) 9 8 N o t a u l i c e s s e p a ra t ed 1 1 N o t au l i c e s j o i n t 9 D o r sa l s u r f a c e o f p r op od e u m smo o t h , n o t r e t i cu l a t e ru g o s e , o n l y w i t h med i a n f u r r ow . . . . . . . . . . . . . . . . Pseudogona topus s a c c h a retorum R . C L . P e rk i n s D o r s a l s u r f a c e o f p r op od eu m f u l l y r e t i c u l a t e ru g o s e 10 1 0 D o r s a l p r o c e s s o f g o n o f o r c ep s l on g , w i t h d i s t a l h a l f .

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1 T h e o n l y k n ow m a l e spec i men c a n b e l o n g t o Gona top u s op ta b ilis or decor a t u s ( see tex t ) .

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Gon a topo d i n a e

b r o a d e n ed , p a r a l l . . . . . . . . . . . . . . . . . Dorsa l p ro c e s s of 960 ) •

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1 065

e l t o t h e p en i s ( F i g . 8 7 8 ) . . . . . . . . . . . . . . . . . . . . Ps e u dogo n a top u s hospes R . C . L . P e rk i n s g on o f o r c ep s s h o r t , s l en d e r , t r a n s v e r s e ( F i g . Dicondylus perkinsi ( A s h m ea d ) •

1 1

P r op od e u m s m o o t h , n o t r e t i c u l a t e ru g o s e . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Pseu dogon a top u s dichromus R . C . L . P e rk i n s P r op o d e u m f u l l y o r a l mo s t f u l l y r e t i c u l a t e r u g o s e . . . . . . . . . . . 1 2 1 2 D o r s a l p r o c e s s o f g o n o f o r c ep s w i t h n u m e r o u s s h o r t , p o i n t ed l am e l l a e ( F i g . 9 6 9 ) ; p r o p o d e u m f u l l y r e t i c u l a t e rug o s e . . . . . . . . . . . . . . . . . . . . . . . Dicondylus alpin u s ( G o u r I a y ) D o r s a l p r o c e s s o f g on o f o r c ep s smo o t h , w i t h ou t s h o r t l a m e l l a e .

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( F i g s 8 7 5 , 8 7 6 ) ; p r o p o d e u m r e t i c u l a t e ru g o s e , b u t w i t h d o r s a l s u r f a c e s h ow i n g t w o l a t e r a l s m o o t h a r e a s . . . . . . . . . . Pseu dogon a top u s n igric a n s R . C . L . P e r k i n s .

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1 . Neodryin u s R .

C.

L Perk i n s

Neodryin u s R . C . L . Perk i ns 1 905 : 50 . Psilodryin us K i effer i n K i effer et M a r s h a l l 1 90 6 : 497 ( s y n . p ro p osed b y R i c h a r d s 1 953 ) ; t y p e spec i e s Psilodryin u s a c u ticollis ( K i effer 1 905 ) , ori g . des i g . Neodryin u s R . C . L . P e r k i n s : K i effer i n K i effer e t M a rsh a l l 1 906 : 497 . Prodry i n u s K i effer i n K i effer et M a rs h a l l 1 906 : 497 ( s y n . p roposed by R i c h a rd s 1 95 3 ) ; t y p e spec i es Prodryin u s bra chycerus ( K i effer 1 904 ) , or i g . des i g . Neodryin u s R . C . L . Perk i n s : K i effer 1 9 1 4b : 2 5 . Prodryin u s partim K i effer : K i effer 1 9 1 4b : 4 9 . Ps ilodryin u s K i effer : K i effer 1 9 1 4b : 53 . Ph a n erodryin us Roep k e 1 9 1 6 : 289 ( s y n . p roposed by R i c h a r d s 1 95 3 ) ; t y pe s p ec i e s Ph a n erodryin us ja v a n u s R o e p k e 1 9 1 6 , or i g . d es i g . Neodryin u s R . C . L . Perk i n s : R i c h a r d s 1 939 : 1 89 . Prodryin u s I< i ef fe r : R i c h a r d s 1 939 : 1 89 . Psilodryin u s K i effe r : R i c h a rd s 1 93 9 : 1 89 . L eptodryin us R i c ha rd s 1 95 3 : 67 ( n . sy n . ) ; t y p e s p ec i e s Leptodryin us leptop u s R i c h a rd s 1 95 3 , m o n o t y p i c . Neodryin u s R . C . L . Perk i n s : D l m i a n d C u rrado 1 97 6 a : 1 84 . Psilodryinus K i ef fe r : D i m i a n d C u rr a d o 1 976a : 1 84 . Prodryin u s K i ef fe r : D l m i a n d C u rrado 1 976a : 1 84 .

T Y P E SP E C I E S : Neodryin u s koebelei R . C . L . Perk i n s 1 905 , o r i g . des i g . F E MA L E ( F i g s 7 1 5 , 7 1 6 ) : f u l l y w i n ged ; m a x i I l a ry p a l p i w i t h 6 segmen t s ; l ab i a l p a l p i w i t h 3 segmen t s ; p ro no t u m crossed by a s t ro n g t r a n s v e rs a l

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1 066

Mem . Amer .

Fig.

715

-

Ent.

I ns t . ,

n o . 37 ,

1 984

F em a l e of Neodryin u s nelsoni R . C . L . Perk i n s

i m p ress i o n ( F i g . 7 1 6 ) ; n ot a u l ices i n v i s i b l e ; e n l a r g e d c l a w ( F i g . 7 1 9 ) w i t h s u b a p i c a l too t h a n d l am e l l ae or b r i s t l es or p e g - I i k e h a i rs ; t i b i a l s p u rs 1 , 0 , 2 ( F i g . 758 B ) . MALE ( F i g s 7 1 7 , 7 1 8 ) : f u l l y w i n g e d ; m a x i l l a ry p a l p i w i t h 6 s e g men t s ; l ab i a l p a l p i w i t h 3 segmen ts ; occ i p u t s t ra i g h t ; temp l es i n v i s i b l e ; o c c i p i t a l c a r i n a i n v i s i b l e o r on l y v i s i b l e b e h i n d t h e oce l l i o r o n t h e s i des o f t he pos te r i o r oce l l i , b u t n e v e r reac h i n g t e m p l es ; t i b i a l s p u rs 1 , 1 ,2.

G o n a topod i n ae

Fig. 716

-

1 067

Fem a l e of Neodryinus n e lsoni R . C . L .

D I ST R I BU T I O N : E t h i op i a n , O r i e n t a l , Nearc t i c , HOST S : Fl a tidae, Ric aniidae, Nogodinidae S P E C I E S : 26

Perk i n s

Neotrop i c , A u s t r a l i a n

E T H I O P I AN S P E C I E S O F NEODRYINUS

T w o s p ec i e s a re k n ow n .

KEY T O THE SPEC I ES

F E MALES

O c c i p i t a l c a r i n a i n v i s i b l e ; s cu t u m n o t r e t i c u l a t e ru g o s e , w i t h n u m e r ou s l on g i t u d i n a l k e e l s 1 . a n tiq u u s B en o i t O c c i p i t a l c a r i n a v i s i b l e b eh i n d t h e o c e l l i a n d on t h e s i d e s o f t h e p o s t e r i o r o c e l l i ; s c u t u m f u l l y r e t i c u l a t e ru g o s e . . . . . . . . . . •

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Mem . Amer .

1 068

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I ns!. ,

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2.

1 984

gigas C eb a l l o s

M a l e of Neodryin u s koebelei R . C . L . Perk i n s

MALES

U n k no w n

no. 37,

G o n a topod i n a e

Fig.

1 069

7 1 8 - M a l e of Neodryin u s koebelei R . C . L . Perk i n s

1 . Neodryin u s a n t i q u u s Beno i t

Neodryin u s a n ti q u u s Be n o i t 1 95 3 b : 402 .

F E MALE : f u l l y w i n g e d ; l e n g t h 4 , 37 m m ; h e a d b l ac k , w i t h m a n d i b l e s redd i s h ; a n t e n n ae b ro w n , w i t h seg m e n t 5 , d i s t a l p a r t o f seg m e n t 4 a n d p ro x i m a l p a r t of seg men t 6 ye l l o w ; p ron o t u m b l a c k , w i t h s i des p a r t l y redd i s h ; meso t h o r a x , met a t ho r a x a n d p ro p o d e u m b l a c k ; a b domen b ro w n b l ac k ; l e g s b ro w n -red d i s h ; head w ea k l y exc a v a ted , w i t h n U merous l o n g i t u d i n a l k ee l s o n fron s , v e r t e x and occ i p u t ; occ i p i t a l c a r i n a i n v i s i b l e ; p ro n o t u m crossed b y a t r a n s v e rs e i mp r es s i on , d u l l , g r a n u l a ted , w i t h n u merous l o n g i t u d i n a l kee l s on co l l a r a n d d i s c ; pos t e r i o r edges of :) rono t u m ro u n ded , not p ro d u ced i n to l ob e s ; scu t u m du I I , as scu I p t u red as ,::J ro no t u m , w i t h n u m e rous l o n g i t u d i n a l kee l s ; n o t a u l i ces i n v i s i b l e ; s c u t e l l u m a s scu I p t u red a s scu t u m ; met a n o t u m v ery reduced a n d sho r t ; p ropode u m f u l l y re t i c u l a t e ru gose , w i t ho u t l o n g i t u d i n a l k ee l s on pos ter i or s u rface ; fore- w i n g w i t h t w o b r o w n t r a n svers a l b a n d s ; d i s t a l p a r t of r a d i a l ve i n m u c h l o n ger t h a n p ro x i m a l p a r t ; fore t ro c h a n ters a p p ro x i m a te l y f o u r t i mes a s l on g a s b ro a d ; segment 1 of f ro n t t a r s u s l o n g e r t h a n seg m e n t 4 ( 20 : 1 0 ) ; e n l a r ged c l a w ( F i g . 7 1 9 ) w i t h a s U b a p i c a l [Oo t h a n d 4 p e g - l i ke h a i rs ; s e g m e n t 5 o f f ro n t t a r s u s ( F i g . 7 1 9 ) w i t h t w o - o w s of n e a r l y 9 l a m e l l ae ; a p e x w i t h a g r o u p of a t l e a s t 20 l a m e l l a e ; .,-, a x i I l a ry p a l p i w i t h 6 s e g me n t s ; l a b i a l p a l p i w i t h 3 segm e n ts ; t i b i a l s ;:) u rs 1 , 0 , 2 . '.'ALE : u n k n o w n

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LOCUS T Y P I C U S : M a d a g as c a r T Y P I CA L M AT E R I A L : ho l o t y p e F ! i n P ( b y m i s t a k e t h e ho l o t y p e w a s l ab e l l ed Neodryin u s berla n di n . sp . ) . HOSTS : u n k n o w n D I ST R I B U T I O N : E t h i o p i a n re g i o n : MADAGASCAR , P !

2 . Neodryin u s gigas Ceb a l l os

Neodryin u s gigas C eb a l l os 1 936 : 5 1 . Neodryin u s giga s C eb a l l o s : Beno i t 1 953b : 401 .

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1 072

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1 984

- C he l a of Neodryin u s gigas C eb a l l os ( t y p e )

prono t u m ro u n de d , n o t p ro d uced i n to l o b e s ; sc u t u m d u l l , f u l l y ret i c u l a t e r u gose ; n o t a u I i ce s i n v i s i b I e ; sc u t e l l u m d u I I , f u I I y re t i c u I a te r u gose ; m e t a n o t u m re t i c u l a te ru gose , v e r y red u ced a n d s h o r t ; p ropodeum f u l l y

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Mem . Amer . E n t .

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1 984

1 . Neodryin u s leptopus R i c h a r d s

Neo dryin u s (Leptodryin u s ) leptop u s R i c h a rd s 1 95 3 : 66 .

F E M A L E : f u l l y w i n ged ; l e n g t h 5 , 93 m m ; head b l ac k , w i t h f ro n t p a r t o f v e r t e x ( more a l o n g orb i t s ) , c l y p e u s a n d m a n d i b l es y e l l o w ; a n te n n ae testaceo u s , w i t h segmen ts 3-5 d a r k ; t ho r a x a n d p ropodeum b l ac k , w i t h poste r i o r s u rface o f pron o t u m red d i s h ; a b domen b row n ; l eg s b row n , w i t h t a rs i l i g h t ; hea d d u l l , g r a n u l a t e d , w i t h a s t ro n g med i a n k e e l between t he a n t E! n n a l soc k e t s ; f r o n s w i t h some short t r a n svers a l k ee l s o n t he s i d es of the med i a n k ee l ; oce l l a r t r i a n g l e e q u i l a t e ra l ; a n t e r i o r h a l f o f pronot u m

( F i g . 7 2 2 A ) c rossed b y a w e a k t r a n s v e rs a l i m p ress i on ; poste r i or h a l f w i t h a no t h e r w e a k t r a n sversa l i mp ress i o n ; c o l l a r s h i n y , w i t h o u t s c u l p t u re , s m oo t h ; d i s c s h i n y , w i t h s t r o n g I d n g i t u d i n a l k e e l s ; poster i o r ed ges o f t h e p ron o t u m n o t p ro d u ced i n to l obes ; sc u t u m s h i n y , r u gose , w i t h s t ro n g l o n g i t u d i n a l k e e l s ; n o t a u l i ces i n v i s i b l e ; s c u te l l u m s h i n y , f u l l y re t i c u l a t e r u g ose ; m e t a n o t u m i n v i s i b l e ; p ropodeum f u l l y ret i c u l a te r u gose , w i t h t w o l o n g i t u d i n a l kee l s on poste r i o r s u rf a ce ; fore t ro c h a n ters a p p ro x i m a te l y four t i mes as l o n g as b road ; fore-w i n g w i t h two b ro w n t r a n s v e r s a l b a n d s ; d i s t a l p a r t o f rad i a l ve i n m u c h l on g e r t h a n p ro x i m a l p a r t ; segmen t 1 o f f r o n t t a r s u s w e a k l y s horter t h a n segmen t 4 ( 27 : 30 ) ; e n l a rged c l a w ( F i g . 723 ) w i t h s U b a p i c a l too t h a n d 8 l a m e l l ae ; segme n t 5 of f ro n t t a r s u s ( F i g . 723 ) w i t h t w o rows of 1 8 + 1 1 l ame l l ae ; apex w i t h a g ro u p of at l ea s t 20 l a me l l ae ; m a x i l l a ry p a l p i w i t h 6 seg men t s ; l ab i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l s p u r s 1 , 0 , 2 . MALE : u n k n o w n LOCUS T Y P I C US : K uc h i n g ( Borneo , I n dones i a ) T Y P I C A L MAT E R I A L : h o l o t y p e F ! i n BM . HOST S : u n k no w n D I S T R I BU T I O N : O r i e n t a l reg i o n : I N D O N E S I A : K uc h i n g ( Borneo ) , B M ! LAOS : B a n - v a n-eue ( SE of P ho u - k o w - k u e i ) , B !

2 . Neo dryin u s retic u l a t u s ( F o u t s )

Psilodryin u s reticu l a tus F o u t s 1 92 2 : 632 . Neodryin u s retic u l a t u s ( F o u t s ) : B a l t a z a r 1 966 : 204 . F E MA L E : f u l l y w i n ged ; l e n g t h 6 , 25 m m ; head b l ac k , w i t h m a n d i b l es red i s h ; a n te n n ae b row n , w i t h segme n ts 8- 1 0 l i g h t a n d ve n t r a l face of segmen t 1 w h i te ; t ho r a x , p ropodeum a n d a b domen b l ac k ; l eg s b l ack b row n ; head v e r y e x c a v a te d , s h i n y , g r a n u l a t e d , w i t h n u merous l o n g i t u d i n a l k ee l s o n fro n s , vertex and o cc i p u t ; SOme k ee l s a re a l so v i s i b l e amon g the oce l l i ; occ i p i t a l c a r i n a i n v i s i b l e ; p ron o t u m crossed b y

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1 077

C he l a of Neodryin u s reti c u l a t u s ( F o u t s ) ( t y p e )

a t r a n s v e rs a l i m pres s i on , h a i r y , w i t h some l o n g i t u d i n a l k ee l s m a i n l y on the co l l a r ; pos t e r i o r edges of the p ro n o t u m rou n d e d , n o t p ro d u ced i n to

Mem . A m e r .

1 078

Ent.

I ns ! . ,

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l obes ; s c u t u m d u l l , g r a n u l a t ed a n d f u l l y r e t i c u l a t e r u gose , w i t h t w o k ee l s i n s t e a d of t h e no t a u l i ces ; n o t a u l i ces i n v i s i b l e ; sc u te l l u m a s scu I p t u red as sc u t u m ; m e t a n o t u m very reduced a n d s ho r t ; p ropodeum fu I I y r e t i c u l a t e r u gose , w i t h t w o l o n g i t u d i n a l k ee l s on t h e pos t e r i o r s u rface ; fore- w i n g w i t h t w o b ro w n t r a n s v e rs a l b a n d s ; d i s t a l p a r t of r a d i a l v e i n m u c h l o n g e r t h a n p rox i m a l p a r t ; fore t roc h a n t ers a p p rox i m a te l y f o u r t i me s a s l o n g a s b road ; segmen t 1 of fron t t a r s u s l o n g e r t h a n s e g m e n t 4 ( 30 : 22 ) ; en l a r g e d c l a w ( F i g . 724 ) w i t h s u b a p i c a l t o o t h a n d 5 b r i s t l es ; segme n t 5 of f r o n t t a rs u s ( F i g . 724 ) w i t h 2 row s of 1 7 + 1 1 l am e l l ae ; a p e x w i t h a g ro u p o f a t l e a s t 1 4 l am e l l ae ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l s p u r s 1 , 0 , 2 . MALE : u n known l O C U S T Y P I C US : los B a no s ( P h i I i p p i n es ) T Y P I CA L MAT E R I AL : h o l o t y p e F ! a n d 1 p a r a t y p e F ! i n VIA . HOST S : u n k no w n D I ST R I B U T I O N : O r i e n t a l reg i o n : P H I L I P P I N E S : L o s B a nos ( L u zon ) , WA !

3 . Neodryin u s chela tus n . s p .

F E MA L E : f u l l y w i n ged ; l e n g t h 6 , 25-6 , 80 m m ; h e a d b l ac k , w i t h s i des of c l y p e u s and m a n d i b l es y e l l ow ; a n te n n ae b l ac k , w i t h segme n t s 6-1 0 r ed d i s h a n d v e n t r a l f a c e of seg m e n t s 1 -2 y e l l o w ; t horax , p ropodeum a n d p e t i o l e b l ac k , w i t h s i des o f prono t u m p a r t l y redd i s h ; a b domen red d i s h ; fore l e g s b l ac k , w i t h coxae a n d t roc h a n t e r s p a r t l y y e l l ow , c l u b s of femora redd i s h , e x t e r n a l f ace of t i b i ae red d i s h , e n l a r ged c l a w y e l l ow ; m i d a n d h i n d l e g s b l a ck , w i t h coxae p a r t l y red d i s h , t roc h a n t e r s a n d c l u b s o f fe mora f u l l y red d i s h , t a r s i y e l l o w , w i t h p rox i m a l h a l f of s e g m e n t

1 b l ac k ; h e a d e xc a v a t e d , s h i n y , w i t h n u merous l o n g i t u d i n a l k ee l s on frons and v e r t e x ; occ i p i t a l c a r i n a i n comp l e t e , reach i n g a p p rox i m a te l y 0 , 5 l e n g t h of OOl ; p rono t u m ( F i g . 722 B ) crossed b y a t r a n s v e rs a l i m p ress i o n , s h i n y , a s scu l p t u re d a s h ea d , w i t h n u merous l o n g i t u d i n a l k ee l s ; p o s te r i or ed ges o f t h e p ron o t u m rou n d e d , n o t p roduced i n t o l ob e s ; scu t u m h a i r y , s h i n y , a s s c u l p t u red a s h ea d , w i t h cen t r a l reg i o n ret i c u l a t e r u gose , o n t he s i des w i t h n u merous l o n g i t u d i n a l k ee l s ; no t a u l i ce s i n v i s i b l e ; sc u te l l u m f u l l y ret i c u l a t e r u g os e , a s scu l p t u red a s h e a d ; m e t a n o t u m v ery reduced a n d s hort ; p ropodeum f u I I y re t i c u I a t e r u gose , w i t h t w o l o n g i t u d i n a l k ee l s on pos te r i or s u rface ; fore-w i n g w i t h t wo b r o w n t r a n s v e rs a l b a n d s ; d i s t a l p a r t o f rad i a l v e i n m u c h l o n g e r t h a n p rox i m a l p a r t ; segmen t 1 of f ron t t a rs u s l o n g e r t h a n seg m e n t 4 ( 35 : 2 5 ) ; e n l a r g e d c l a w ( F i g . 725 ) w i t h s u b a p i c a l t oo t h a n d 4 peg- I i k e h a i rs ; segmen t 5 of f ro n t t a rs u s ( F i g . 725 ) w i t h t w o rows of 1 0 + 8 l a me l l ae ; a p e x w i t h a g ro u p o f a t l e a s t 3 0 l ame l l a e ; m a x i l l a r y p a l p i w i t h 6 seg men t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 2 . MALE : u n k n o w n l O C U S T Y P I C U S : B a n V a n E u e ( L aos )

Mem . A m e r . E n ! .

1 080

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-

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n o . 37 ,

1 984

C he l a of Neodryinus rub escens n . s p .

( type)

T Y P I CAL M A T E R I A L : h o l o t y p e F ! i n B ; 1 p a r a t y pe F ! i n CO . HOSTS : u n k n o w n D I STR I BU T I O N : O r i e n t a l reg i o n : LAOS : B a n V a n E u e ( V i en t i a n e ) , P H I L I PP I NE S : D a l a w a n B a y ( Ba l a b ac ) , CO !

B!

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4 . Neodryin u s rubescens n .

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sp .

F E MA L E : f u l l y w i n ge d ; l e n g t h 5 , 00 m m ; h e a d b l ack , w i t h s i de s of c l y p e u s y e l l ow a n d m a n d i b l es redd i s h ; a n t e n n a e b l ac k , w i t h seg m e n t s 7- 1 0 redd i s h a n d v e n tra l face of s e g m e n t 1 y e l l o w ; t ho ra x , propode u m a n d pe t i o l e b l ac k , w i t h redd i s h n u a nces on t h e s i des o f t h e p ronot u m ; a b domen b row n - re d d i s h ; l e g s redd i s h ; head exc a v a ted , s h i n y , g r a n u l a ted , w i t h n u merous l o n g i t u d i n a l k ee l s on f ro n s a n d verte x ; occi p i t a l ca r i n a d i s t i n c t , not e x te n d i n g b e y o n d 0 , 5 l en g t h o f OOL ; p ro n o t u m crossed b y a t r a n s v e rs a l i m p re s s i o n , h a i r y , as scu l p t u red as h e a d , w i t h n u m e ro u s l o n g i t u d i n a l k ee l s on t h e c o l l a r ; poster i or e d g e s of t h e p ro n o t u m rou n ded , not produced i n to l o b e s ; s c u t u m d u l l , as sc u l p t u red as p ronot u m , ret i c u l a t e r u gose , w i t h two k ee l s i n s t e a d of t h e n o t a u l i ces ; no t a u l i ce s i n v i s i b l e ; sc u te l l u m a s scu l p t u red a s sc u t u m ; m e t a n o t u m v e r y reduced a n d s hort ; p ropod e u m f u l l y ret i c u l a t e ru gose , w i t h t w o l o n g i t u d i n a l k ee l s o n poste r i o r s u rf ace ; fore-w i n g w i t h t w o b ro w n t r a n s v e rs a l b a n d s ; d i s t a l p a r t o f ra d i a l v e i n m u c h l o n g e r t h a n p rox i m a l p a r t ; fore t roc h a n ters ap prox i m a te l y four t i mes a s l o n g a s b road ; s e g m e n t 1 of fron t t a rs u s l o n g e r t h a n s e g m e n t 4 ( 2 3 : 1 5 ) ; en l a rged c l a w ( F i g . 726 ) w i t h s u b a p i c a l tooth a n d 7 b r i s t l es ; s e g m e n t 5 of fro n t t a r s u s ( F i g . 726 ) w i t h t wo rows of 10 palpi

+

1 3 l a m e l l ae ; apex w i t h a g ro u p of a t l ea s t 1 5 l am e l l ae ; m a x i I l ary w i t h 6 s e g me n t s ; l a b i a l p a l p i w i t h 3 s e g m e n ts ; t i b i a l s p u rs 1 , 0 , 2 .

M AL E : u n k n o w n L O C U S TYP I C U S : K a n d y ( S r i L a n k a ) T Y P I CA L M A T E R I AL : h o l o t y p e F ! i n B M . HOST S : u n k no w n D I ST R I BU T I O N : O r i e n t a l reg i o n : SR I LANKA : K a n d y ,

BM !

5 . Neodryin u s dolosus n . s p .

F E M A L E : f u l l y w i n ged ; l e n g t h 4 , 68 m m ; head b l ac k , w i t h s i des of c l y p e u s a n d m a n d i b l es redd i s h ; a n te n n a e b l ac k - b ro w n , w i t h s e g m e n t s 8- 1 0 red d i s h a n d ven t r a l face of s e g m e n t 1 y e l l ow ; t hora x , p ropodeum a n d p e t i o l e b l ac k ; a b do m e n b row n ; l e g s b ro w n - b l ac k ; h e a d exca v a ted , g r a n u l a ted , s h i n y , h a i r y ; fro n s a n d v e r te x w i t h n u merous l on g i t u d i n a l k ee l s ; occ i p i t a l car i n a d i s t i n c t , not exte n d i n g beyon d 0 , 5 l en g t h of OO L ; pron o t u m crossed b y a t r a n s vers a l i m p re s s i o n , s h i n y , h a i r y , a s s c u l p t u red a s head , w i t h S o m e l o n g i t u d i n a l k ee l s on t h e co l l a r ; poste r i or edges of t h e p ro n o t u m ro u n ded , not p rod uced i n to l obes ; s c u t u m h a i r y , a s sc u l p t u red a s p ronot u m , w i t h two i n c i m p l e te k ee l s i n s tead of t h e n o t a u t i ces j no t a u l i c e s i n v i s i b l e ; s c u te l l u m a s scu l p t u red a s sc u t u m ; m e t a n o t u m v e r y reduced a n d short ; p ropod e u m f u l l y ret i c u l a t e ru gose , w i t h two l o n g i t u d i n a l k ee l s on t h e poster i o r s u rface ; fore-w i n g w i t h two b ro w n t r a n s v e rs a l b a n d s ; fore troc h a n ters a p p rox i m a te l y fo u r t i mes a s

1 082

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I nst. ,

no.

37 ,

1 984

F i g . 729 - C he l a of Neodryin us dolosus n . sp .

( ty p e )

l o n g a s b ro a d ; segmen t 1 o f f ro n t t a r s u s l o n ge r t h a n segmen t 4 ( 2 5 : 1 8 ) ; en l a r g e d c l a w ( F i g . 7 2 7 ) w i t h s u b a p i c a l too t h a n d 8 peg - I i ke h a i rs ; segmen t 5 o f f r o n t t a r s u s ( F i g . 727 ) w i t h t w o rows o f 1 3 + 1 4 l a me l l ae ; a p e x w i t h a g r o u p o f a t l ea s t 1 2 l a me l l ae ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l s p u r s 1 , 0 , 2 . I,j AL E : u n k n o w n L O C U S T Y P I C US : T a i p o k a u ( Ho n g Kon g ) TYP I CA L MAT E R I AL : ho l o t y pe F ! i n 8 . H OST S : u n k n o w n D I ST I, 1 8 U T I O N : O r i e n t a l reg i o n : HOr� G K O N G : T a i po k a u ( Ko w l oon ) , B !

6 . Neodryin u s rob ustus n .

sp .

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1 083

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728 - C h e l a of Neodryin u s rob u s t u s n . sp .

( type )

F E MA L E : f u l l y w i n g e d ; l e n g t h 5 m m ; h e a d b l ac k , w i t h m a n d i b l es redd i sh ; a n t e n n a e b l ac k , w i t h se g m e n t 1 0 redd i s h a n d v e n t r a l s i d e o f s e g m e n t 1 l i g h t ; thorax , p ropodeu m , p et i o l e , a b domen a n d l e g s b l ac k ; h e a d e x c a v a ted , d u l l , g r a n u l a t e d , w i t h n u me ro u s l o n g i t u d i n a l k ee l s o n fron s a n d verte x ; occ i p i t a l ca r i n a i n com p l et e , not exte n d i n g b e y o n d 0 , 5 l e n g t h o f OOL ; p ro n o t u m d u l l , a s scu l p t u red a s h e a d , crossed b y a t r a n svers a l i m pres s i o n , w i t h n u me ro u s l o n g i t u d i n a l k ee l s ; poster i o r e d g e s o f pron o t u m

1 084

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Mem . Amer . E n ! .

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1 984

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Gon a topod i n a e

1 085

rounded , not p rod uced i n to l ob e s ; sc u t u m d u I I , a s s c u I p t u re d as p ro no t u m , w ea k l y re t i c u l a te r u g ose , h a i r y , w i t h 2 i n comp l e te kee l s i n s t e a d of t he n o t a u l i ces ; n o t a u l i ces a b s e n t ; s c u te l l u m a s s c u l p t u red a s s c u t u m ; m e t a n o t u m very reduced , s h or t ; p ro p o d e u m f u I I y re t i c u I a te r u g ose , w i t h t w o l o n g i t u d i n a l kee l s on poste r i o r s u rface ; fore troc h a n ters approx i m a te l y fo u r t i mes a s l o n g a s b road ; fore w i n g s w i t h t w o b ro w n t r a n s v e rs a l b a n d s ; d i s t a l p a r t of rad i a l v e i n m u c h l o n g e r t h a n prox i m a l p a r t ; se g me n t 1 o f f r o n t t a r s u s l o n g e r t h a n seg m e n t 4 ( 2 7 : 1 7 ) ; en l arged c l a w ( F i g . 728 ) w i t h s u b a p i c a l too t h and 8 b r i s t l e s ; seg m e n t 5 of fro n t t a r s u s ( F i g . 728 ) w i t h t w o ro w s o f 2 1 l am e l l a e ; a p e x w i t h a g ro u p o f a t l ea s t 1 2 l a me l l ae ; m a x i I l a r y p a l p i w i t h 6 s e g me n t s ; l a b i a l p a l p i w i t h 3 s e g m e n t s ; t i b i a l s p u rs 1 , 0 , 2 . MALE : u n known L O C U S T Y P I C U S : B a n V a n E u e ( L ao s ) T Y P I CA L MAT E R I A L : h o l o t y p e F ! i n B . HOSTS : u n k n o w n D I ST R I B U T I O N : O r i e n t a l reg i o n : LAO S : B a n V a n E u e ( V i e n t i a n e ) , B !

7 . Neodryin u s j a va n us ( Roep k e )

Ph anerodryin us ja v a n u s Roepke 1 9 1 6 : 290 . Psilodryin u s thoracicus F o u t s 1 92 2 : 6 3 1 ( n . s y n . ) ; locus typicus P uerto P r i ncesa ( P h i l i p p i n es ) . Neodryin u s l a w a n a e R i c h a r d s 1 953 : 64 ( n . s y n . ) ; locus typicus K u a l a Lumpur ( Malaya ) . Neodryin u s j a v a n u s ( Roep k e ) : R i c h a r d s 1 953 : 6 1 . Neodryin us thoracic us ( F o u t s ) : B a l t az a r 1 966 : 204 .

F E MA L E : f u l l y w i n ged ; l e n g t h 5 , 00-6 , 25 m m ; h e a d b l ac k , w i t h s i de s o f c l y p e u s a n d m a n d i b l e s y e l l o w ; a n te n n ae redd i s h , w i t h v e n t ra l face of s e g m e n t 1 y e l l o w ; t hora x , p ropodeum and pe t i o l e b l ac k ; a b domen redd i s h ; l e g s redd i s h ; h e a d exca v a t e d , h a i r y ; frons a n d v e r t e x w i t h n u merous l o n g i t u d i n a l k ee l s ; occ i p i t a l ca r i n a i n comp l e t e , e x t e n d i n g beyond 0 , 5 l e n g t h o f OOL ; p ro n o t u m crossed b y a t r a n s v e rs a l i m press i o n , h a i r y , g r a n u l a t e d , w i t h some l o n g i t u d i n a l kee l s on t he s i d es of t he co l l a r ; poste r i or edges o f t h e p ro n o t u m rounded , not p rod uced i n to l ob e s ; s c u t um d u l l , h a i r y , as sc u l p t u red as p ro no t u m , w i t h t w o i n c i m p l e te k ee l s i n s t e a d of t h e not a u l i ce s ; n o t a u l i ces i n v i s i b l e ; s c u t e l l u m a s sc u l p t u red a s scu t um ; m e t a n o t u m v e r y reduced a n d s h or t ; p ropodeum f u l l y re t i cu l a te r u gose , w i t h t w o l o n g i t u d i n a l k ee l s on t h e post e r i o r s u rf ace ; fore-w i n g w i t h t w o t r a n s v e r s a I b r o w n b a n d s ; d i s t a I p a r t o f ra d i a I v e i n m u c h l on g e r t h a n p rox i m a l p a r t ; f o r e t roc h a n ters a p p rox i m a t e l y f o u r t i mes a s l on g a s b ro a d j se g m e n t 1 of fron t t a r s u s l o n g e r t h a n seg m e n t 4 ( 3 1 : 20 ) ; en I a r ged c l a w ( F i g . 729 ) w i t h s u b a p i c a l tooth a n d 8 b r i s t l e s ; seg m e n t 5 o f fron t

1 086

Mem . A mer . E n t .

I ns!. ,

n o . 37 ,

1 984

F i g . 730 - L a b i a l p a l p u s of fem a l e of !leodryin u s j a va n u s ( Roep k e ) from K ua l a Lumpur ( M a l a y a )

t a rs u s ( F i g . 72 9 ) w i t h t w o ro ws o f 1 4 + 1 5 l a m e l l ae ; apex w i t h a g ro u p o f a t l e a s t 1 5 l a me l l ae ; m a x i l I a ry p a l p i w i t h 6 s e g m e n t s ; l a b i a l p a l p i w i t h 3 seg m e n t s ( F i g . 730 ) ; t i b i a l s p u rs 1 , 0 , 2 . M AL E : f u l l y w i n ged ; l e n g t h 3 , 37-3 , 50 m m ; h e a d b l ac k , w i t h m a n d i b les y e l l o w ; t h o r a x and propod e u m b l ack ; a b do m e n b row n ; legs b row n , w i t h t a rs i a n d fore t i b i a e l i g h t ; a n te n n a l seg men t s i n fo l l o w i n g p roport i on s : 6 : 6 : 1 6 : 1 2 : 1 2 : 1 2 : 1 2 : 1 0 : 9 : 1 0 ; h e a d h a i r y , g r a n u l a ted , w i t h a m ed i a n kee l b e t w e e n t he a n t e n n a I sock e t s ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 6; O L = 1 ; OOL = 7; b re a d t h of t h e oce l l i : 5 ; te m p l es i n v i s i b l e ; s c u t u m h a i r y , d u l l , g r a n u l a ted ; n o t a u l i ces d i s t i nc t a n d com p l e t e , poster i o r l y s e p a r a t e d ; m i n i m u m d i s t a n ce b e t ween t h e n o t a u J i ces n e a r l y a s l o n g a s a n te n n a I se g m e n t 2 ( 5 : 6 ) ; sc u te l l u m a s scu l p t u red a s scu t u m ; m e t a n o t u m w i t h a n te r i or t h i rd r u g os e ; m i d a n d pos t e r i o r t h i rd smoot h , s h i n y , w i t h o u t sc u l p t u r e ; p ropodeurn fu l l y ret i c u l a t e r u gose , w i t ho u t l o n g i t u d i n a l k ee l s o n t he poste r i or s u rface ; fore-w i n g h y a l i ne , w i t ho u t d a r k t r a n s ve rs a l b a n d s ; d i s t a l p a r t of rad i a l v e i n n ea r l y a s l o n g a s p rox i m a l p a r t ; dors a l p rocess o f gon oforceps ( F i g . 7 3 1 ) s h o r t , s l en d e r , i r reg u l a r a n d po i n ted , l oc a t e d on t h e d i s t a l h a l f of t he gonoforcep s ; m a x i l l a ry p a l p i w i t h 6 s e g m e n t s ; l a b i a l p a l p i w i t h 3 s e g rn e n t s . LOCUS T Y P I C U S : Ass i n a n ( I n d o n e s i a ) T Y P I CA L �IA T [ R I AL : l ec t o t y p e F ! a n d 1 p a r a l ec t o t y p e F ! ( hereb y d es i g n a te d ) of Ph . j a va n u s i n B M ; h o l o t y pe F ! of Ps . thora cicus i n W A ; h o l o t y pe F ! a n d 4 p a r a t y p e s ( 2 F F , 2 M M ) ! o f N . l a w a n a e i n B M . HOST S : i n M a l a y a L a wa n a conspersa ( W a l k . ) ( Fl a ti da e ) a n d can dida F . ( Roepke 1 9 1 6 ) . D I STR I B U T I O N : O r i e n t a l reg i o n : I N D O I� E S I A : A s s i n a n ( Sa l a t i g a , J a v a ) , B M ! MALAYS I A : K u a l a L u m p u r ( M a l a y a ) , B M ! P H I L I P P I N E S : P u e r t o P r i ncesa ( P a l a w a n ) , VIA !

8 . Neodry in u s pseu dodiffu s u s n .

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G o n a topod i n a e

1 089

red d i s h ; a n t e n n a e b r o w n o r redd i s h , w i t h s e g m e n t s 7 - 1 0 b r ow n ; t ho r a x a n d p ropodeum b l ac k , w i t h s i des a n d pos t e r i o r m a r g i n re d d i s h ; a b domen b l a c k - b r o w n ; l e g s b row n , w i t h t a r s i l i g h t ; h e a d we a k l y e x c a v a ted , s h i n y , w i t h f ro n s a n d v e r t e x f u l l y re t i c u l a t e r u gose ; occ i p u t g r a n u l a t ed , n o t ret i c u l a t e r u gose ; occ i p i t a l c a r i n a i n comp l e te , e x te n d i n g b e y o n d 0 , 5 l e n g t h of OOL ; p ro n o t u m crossed b y a t r a n s v e rs a l i m p ress i o n , h a i ry , w i t h c o l l a r smoo t h , w i t ho u t sc u l p t u re ; d i sc w i t h some l o n g i t u d i n a l kee l s ; poste r i o r edges of t h e p ro n o t u m ro u n d ed , n o t p rod uced i n t o l obes ; sc u t u m h a i r y , f u l l y re t i c u l a t e r u gose ; n o t a u l i ces i n v i s i b l e ; s c u t e l l u m ret i c u l a t e r u gose ; met a n o t u m v e r y r e d u c e d a n d s h or t ; p ropodeum f u l l y ret i c u l a te r u gose , w i t h t w o l o n g i t u d i n a l k ee l s on t h e poste r i o r s u rf ace ; fore-w i n g w i t h t w o b r o w n t r a n s v e rs a l b a n d s ; d i s t a l p a r t o f r a d i a l v e i n m u c h l o n g e r t h a n p ro x i m a I p a r t ; fore t r oc h a n t e r s a p p ro x i m a te I y fo u r t i mes a s l o n g as b road ; s e g m e n t 1 of f ro n t t a rs u s l on g e r t h a n 4 ( 24 : 1 9 ) ; e n l a r g e d c l a w ( F i g . 732 ) w i t h s u b a p i c a l too t h a n d 1 5 peg - l i ke h a i r s ; seg men t 5 of f r o n t t a rs u s ( F i g . 732 ) w i t h t w o r o w s o f 3 + 7 l a me l l a e ; a p e x w i t h w g ro u p o f 5 l a m e l l ae ; m a x i l l a r y p a l p i w i t h 6 segm e n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 2 . MAL E : u n k n o w n lOCUS T Y P I C U S : C h i a n g d ao ( T h a i I a n d ) TYP I CAL MAT E R I A L : h o l o t y pe F ! i n B ; 2 p a r a t y p es F F ! i n WA ; 1 p a r a t y pe F!

i n OL.

HOST S : u n k n o w n D I ST R I B U T I O N : O r i e n t a l reg i o n : T HA I LAN D : C h i a n g d a o , 8 ! P H I L l P P I ;, E S : W a - w a D a m ( M t . M o n t a l b a n , R i z a l , L u z o n ) , O l ! M a n i l a ( L u zo n ) , WA ! SR I lANKA : P a l a t u p a n a ( H a m . D i s t . ) ( Ponom a r e n ko 1 9 8 1 b , u n der n a me ,Veodryin u s koebelei R . C . l . Perk i n s ) , WA ! U l h i t i y a O y a ( 1 5 m i . �m:=: of M a h i y a n g a n a , B a d . D i s t . ) , WA !

9 . Neodryin u s diffu s u s n . s p .

F E M A L E : f u l l y w i n ged ; l e n g t h 4 , 68-5 , 62 m m ; h e a d b l ac k , w i t h m a n d i b l es redd i s h ; a n t e n n ae b l a c k - b row n ; t ho r a x , p ropodeum a n d p e t i o l e b l a ck , w i t h s i des of p r o n o t u m re d d i s h ; a b domen b l ac k , w i t h a p e x re d d i s h ; l e g s b ro w n , w i t h t a r s i l i g h t ; h e a d e x c a v a ted , s h i n y , w i t h f r o n s a n d v e r t e x f u l l y re t i c u l a t e r u g ose ; occ i p u t g r a n u l a ted , n o t re t i c u l a te r u gose ; occ i p i t a l c a r i n a i n comp l e t e , e x t e n d i n g b e y o n d 0 , 5 l e n g t h of O O l ; p ro n o t u m c rossed b y a t r a n s vers a l i m p ress i o n , h a i r y , w i t h co l l a r smoot h , w i t ho u t scu l p t u r e ; d i sc w i t h some l o n g i t u d i n a l k ee l s ; poste r i o r ed ges of t h e p ro no t u m rou n de d , n o t p ro d uced i n to l o b e s ; s c u t u m h a i r y , f u l l y re t i c u l a te r u gose ; n o t a u l i ces i n v i s i b l e ; scu t e ! l u m ret i c u l a t e r u gose ; m e t a n o t u m very reduced a n d s h or t ; p ropodeum f u l l y ret i c u l a t e r u gose , w i t h two l o n g i t u d i n a l k ee l s o n the pos t e r i o r s u r f ace ; fore-w i n g w i t h two brown t r a n s v e rs a l b a n d s ; d i s t a l part o f rad i a l v e i n m u c h l o n g er than p ro x i m a l p a r t ; fore t r oc h a n ters a p p ro x i m a t e l y fo u r t i mes a s l o n g a s b ro a d ; seg men t

1 090

Mem . Amer .

Ent.

I ns t . ,

n o . 37 ,

1 984

/1 ��------ ----� ----- �

Fig.

733

-

C he l a of Neodry in u s diffu s u s n .

sp .

( ty p e )

1 of fro n t t a rs u s l o n ger t h a n seg m e n t 4 ( 2 0 : 1 4 ) ; e n l a rged c l a w ( F i g . 733 ) w i t h s u b a p i c a l too t h a n d 1 3 peg- I i ke h a i rs ; segmen t 5 of f ro n t t a rs u s ( F i g . 733 ) w i t h t w o r o w s o f 9 + 9 l a me l l ae ; a p e x w i t h a g ro u p of 1 0 l a me l l a e ; m a x i I l a r y p a l p i w i t h 6 s e g me n t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 2 . MALE : u n known L O C U S TYP I C U S : K h aoc h a n g ( T h a i l a n d ) T Y P I CA L �lAT E R I AL : h o l o t y p e F ! i n O L ; 1 p a r a t y p e F ! i n B . HOSTS : u n k n o w n D I ST R I BU T I O N : O r i e n t a l reg i o n : T HA I LAN D : K h aoch a n g ( T ra n g , K a ho p h a p p h a ) , O L ! MALAYS I A : Rej a n g D e l t a ( Sa r i k e i D i s t . , S a r a w a k ) ,

6!

G o n a topod i n ae

1 09 1

I� E A R C T I C S P E C I E S O F NEODR YINUS

T h ree s p ec i es a re k n ow n .

K E Y TO T H E S P E C I E S

F EMALES

S c u t u m g ra n u l a t ed a n d p a r t l y r e t i c u l a t e ru g o s e , n o t s c u l p t u r ed by l on g i t u d i n a l k e e l s ; p r o t h o rax f u l l y o r p a r t l y b l a c k • 1 . typ h locybae ( A s h mead ) S c u t u m f u l l y s c u l p t u r ed by p a ra l l e l l o n g i t u d i n a l k e e l s ; p r o t h o ra x b l a c k o r f u l l y r ed d i sh 2 P r o t h o rax f u l l y r ed d i sh , w i t h c o l l a r a n d d i s c smoo t h a n d s h i n y , w i t h ou t s c u l p t u r e 2 . balli n . sp . P r o t h o ra x a l mo s t f u l l y b l a ck , w i t h c o l l a r a n d d i s c d u l l , g ra n u l a t e d a n d w i t h k e e l s a r o u n d t h e d i s c 3 . incaic u s n . sp . ( s e e N e o t r o p i c s p e c i e s ) •

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MALES

O n I y t h e ma I e of N.

typh locyb a e i s k n ow n .

1 . Neodryin u s typ h locyb a e ( As h m e a d ) n . com b .

L a beo typ h locyb a e A s h m e a d 1 893 : 89 . Dryin u s ormenidis A s h me a d 1 903 : 1 92 ( s y n . p roposed b y M u esebeck a n d W a l k l e y 1 95 1 ) ; locus typ i c u s S a n d u s k y ( O h i o , U . S. A . ) . Dryin u s n igrellus B r u e s 1 90 4 : 1 1 7 ( sy n . p roposed b y M u esebec k a n d Wa l k l e y 1 95 1 ) ; locus typicus P a r k v i l l e ( N . Y . , U . S . A . ) . Dryin u s ormenidis A s h mea d : R . C . L . Perk i n s 1 906a : 4 9 2 . Neodryin u s arizonic u s R . C . L . Perk i n s 1 90 7 : 38 ( n . s y n . ) ; locus typ ic u s Noga l e s ( A r i z . , U. S. A . ) . Neodryin u s d u b iosus R . C . L . Perk i n s 1 907.: 39 ( n . s y n . ) ; locus typ ic u s Nog a l e s ( A r i z . , U . S. A . ) . Dryinus ormenidis A s h mead : K i effer 1 90 7 b : 7 . Psilodryin u s n igrell u s ( B r u es ) : K i effer 1 907b : 8 . L a b eo typ h locyb a e A s h m e a d : K i effer 1 907b : 1 6 .

1 092

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Ent.

I n st . ,

no . 3 7 ,

1 984

F i g . 734 - C h e l a of Neodryin u s typ h locyb a e ( As h mead ) from Sa n d u sk y (Ohio)

Gon a topod i n ae

1 093

--- --------------

F i g . 735 - Ma l e gen i t a l i a of Neodryin u s typh locyb a e ( A shmead } ( l ec t o t y p e )

Prodryin u s ormenidis ( As h m e a d ) : K i effer 1 9 1 4b : 5 1 . Psilodryinus arizon icus ( R . C . L . Perk i n s ) : K i effer 1 9 1 4b : 5 5 . Psilodryinus dubiosus ( R . C . L . Perk i n s ) : K i effer 1 9 1 4b : 55 . L a b erius typhlocyb a e ( As h me a d ) : K i effer 1 9 1 4b : 65 . Gon a top u s typ h locy b a e ( As h me a d ) : B r u e s 1 9 1 6 : 6 1 4 . Psilodryin u s carin a t u s Fou t s 1 924 : 1 60 ( n . s y n . ) ; locus typicus C a r l i s l e ( P a . , U. S. A . ) . Psilodryin u s arizon icus ( R . C . L . Perk i n s ) : M u e sebeck a n d Wa l k l ey 1 95 1 : 1 03 5 . Psilodryinus carin a tus Fou t s : M u esebeck a n d Wa l k l e y 1 95 1 : 1 03 5 .

1 094

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n o . 37,

1 984

Psilodryin u s dubiosus ( R . C . l . P e rk i n s ) : Mu esebec k and W a l k l ey 1 9 5 1 : 1 03 5 . Psilodryin u s typ h locyb a e ( As h m ea d ) : M u e sebeck a n d W a I k l ey 1 95 1 : 1 0 35 . Neodryin u s ormenidis ( As h m e a d ) : R i c h a rd s 1 95 3 : 68 . Neodryin u s (Prodryin u s ) arizon ic u s R . C . l . Perk i n s : Krombe i n 1 97 9 : 1 244 . Neodryin u s (Prodryin u s ) c a rin a t u s ( Fou t s ) : K romb e i n 1 979 : 1 244 . Neodryin u s (Prodryin u s ) du b io s u s R . C . l . Perk i n s : Krom b e i n 1 979 : 1 244 . Neodryin u s (Prodryin u s ) typ h locyb a e ( As h me a d ) : Krom b e i n 1 97 9 : 1 24 4 .

F E M A L E : fu l l y w i n g e d ; l e n g t h 4 , 00-5 , 1 2 m m ; h e a d b l ac k o r b row n , w i t h c l y p e u s a n d m a n d i b l es y e l l o w ; a n te n n a e u s u a l l y y e l l o w , w i t h segmen t s 6-8 b row n , occas i o n a l l y fu l l y y e l l ow ; p r o n o t u m b l ac k , w i t h s i de s a n d poster i o r m a r g i n red d i sh ; mesot h o ra x , m e t a t h orax a n d propodeum b l ac k ; abdomen b ro w n - b l ac k ; l e g s u s u a l l y t e s t a ceou s , occ a s i on a l l y w i t h cox a e , c l u b s o f femora a n d t i b i a e d a r k e n e d o r b l ac k i sh ; h e a d a l mo s t f l a t , s h i n y , g r a n u l a t e d a n d w e a k l y rug ose ; occ i p i t a l c a r i n a w e a k l y v i s i b l e on t he s i des of the oce l l i , not e x t e n d i n g b e y o n d 0 , 5 l e n g t h o f OOl ; p r o n o t u m c rossed b y a t r a n s v e rs a l i m p ress i o n , s h i n y , g r a n u l a t e d , w e a k l y s t r i a t e ; pos t e r i o r e d g e s of p ro n o t u m rou n ded , n o t p rodu ced i n t o l obes ; sc u t u m s h i n y , g r a n u l a t e d , w i t h t r a c k s o f a reo l ae , m a i n l y a l o n g i n v i s i b l e n o t a u l i ces ; sc u te l l u m sh i n y , w i t h a n t e r i or h a l f g r a n u l a ted a n d poster i or h a l f smooth a n d w i t h o u t sc u l p t u re ; m e t a n o t u m very reduced a n d short ; p ropodeum fu l l y ret i c u l a t e r u gose , w i t h l o n g i t u d i n a l kee l s on poster i o r s u rf a ce ; i n s o m e spec i me n s a t r a n svers a l f o l d i s v i s i b l e b e t ween a n t er i or a n d p o s t e r i o r s u rf a c e of p ropodeu m ; fore-w i n g w i t h two b ro w n t r a n s versa l b a n d s ; d i s t a l p a r t of ra d i a l v e i n as l o n g a s , or s l i g h t l y l o n ger t h a n prox i m a l p a r t ; fore troch a n ters a p p rox i m a te l y f o u r t i mes as l o n g as b road ; seg m e n t 1 of fro n t tarsus l on g e r than segme n t 4 ( 20 : 1 1 ) ; en l arged c l a w ( F i g . 734 ) w i t h s u b a p i c a l tooth a n d 4 b r i s t l e s ; seg men t 5 of fron t t a rs u s ( F i g . 7 3 4 ) w i t h 1 row of 23 l a m e l l a e ; a p ex w i t h a group of n ea r l y 1 0 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 6 segmen ts ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 2 . MAL E : f u l l y w i n g e d ; l e n g t h 2 , 8 1 m m ; head b l a c k - b row n , w i t h m a n d i b l es y e l l o w ; a n ten n ae b ro w n ; t h o r a x a n d p ropodeum b l ac k ; a b domen b ro w n t e s t a ceo u s ; l e g s t e s t aceou s ; a n t e n n a l seg men t s i n f o l l ow i n g proport i on s : 7 : 6 : 1 0 : 8 : 8 : 8 : 7 : 7 : 7 : 1 0 ; head sh i n y , g ra n u l a t ed , w i t h f r o n s a n d vertex re t i c u l a t e ru gose , w i t h a med i a n kee l b e t w een t jo e a n te n n a l sock e t s ; POL = 8 ; O L = 3 ; OOl = 4 ; b read th o f t h e oce l l i : 4 ; occ i p i t a l c a r i n a i n v i s i b l e ; te m p l es i n v i s i b l e ; oce l l a r t r i a n g l e w i t h b a s e v ery l o n g ; s c u t u m d u l l , g ra n u l a t e d ; n o t a u l i ces i n comp l e t e , reach i n g a p p rox i ma te l y 0 , 75 l e n g t h o f scu t u m ; scu te l l u m a n d m e t a n o t u m g r a n u l a t e d ; p ropodeum ret i c u l a te rugose, w i t hou t l o n g i t u d i n a l k ee l s o n pos ter i or

1 09 5

Gona topod i n a e

s u rf ac e ; fore-w i n g h y a l i n e ,

w i t h o u t d a rk t r a n s v ers a l b a n d s ;

d i s t a l part

o f ra d i a l v e i n n e a r l y as l o n g as p rox i m a l p a r t ; dor s a l p rocess of gonoforceps ( F i g . 735 ) shor t , n a rrow a n d po i n ted ; t i b ; a I s p u rs 1 , 1 , 2 . L O C U S T Y P I C US : Wa s h i n g ton ( D . C . , U . s . A . ) T Y P I CAL M A T E R I A L : l ec t o t y p e M ! a n d 1 p a r a l ec to t y p e M ! ( h ereby des i g n a ted ) of L. typ h locyb a e in WA ; l ec to t y p e F ! and 2 para l ec t o t y pes F F ! ( h e r e b y des i g n a t ed ) of D . ormenidis i n WA ; l ec t o t y pe F ! ( d es i g n a ted b y O l m ; 1 982 ) of N. arizon icus in B ; h o l o t y p e F ! of N. dubiosus in B ; h o l o t y pe F ! of Ps . carin a t u s i n WA ; h o l o t y p e F ! of D . n igrellus i n A M . H O S T S : i n U . S . A . Metca lfa (� Ormenis) pruinosa ( Sa y ) a n d Ormenis septentrionalis ( Sp i n . ) ; Typ hlocy b a s p . a I so accord i ng to A s h m e a d 1 893 ( i nform a t i o n p ro b a b l y w ron g :

the

l ec t o t y p e i n fact

is l a b e l l ed as f o l l ow s :

" P a r : o n Homop t e r a o n C e l t i s + E l m , J u l y 23 . 83" ; n o h o s t i s g l u ed toge t h er w i t h the t y pe spec i men ; the i n for m a t i on so is o n l y in o r i g i n a l desc r i p t i on a n d not o n l a b e l s ) . PARAS I T E S : i n U . S . A . Cheiloneurus swezey; A s h m e a d ( Sw ez e y 1 903 ; R . C . L . Perk i n s 1 906b ) . D I S T R I B U T I O N : Nearc t i c reg i o n : CANADA : V i n e l a n d ( O n t . ) , a T ! S h e t l a n d ( On t . ) , O T ! U . S . A . : Nog a l es ( Ar i z . ) , B ! K i r k w ood ( Mo n t . ) , WA ! Koppe ' s B r i d g e ( We l l bo r n , Brazos Co . , T ex . ) , T E ! H e a r n e ( Rober t so n Co . , T ex . ) , OL ! Mercedes ( Tex . ) , WA ! C M ! W i c h i t a ( K a n s . ) , OL ! A n n Arbor ( M i ch . ) , BM ! I ow a ( M uesebeck a n d W a l k l ey 1 95 1 ) ; U r b a n a ( I I I . ) , WA ! Terrace P a r k ( Oh i o ) , B ! C o l u m b u s ( O h i o ) , B ! S a n d u s k y ( Oh i o ) , WA ! B ! Ben t on v i l l e ( Ark . ) , WA ! A i k en ( S . C . ) , O l ! a T ! Wash i n g t on ( D . C . ) , WA ! C a r l i s l e ( P a . ) , WA ! N . C u m b er l a n d ( P a . ) , Ol ! Moores t o w n ( N . J . ) , T W ! I n d l ow v i l l e ( N . Y . ) , I T ! N e w b u r g h ( N . Y . ) , ZO ! S i x M i l e C reek ( I t h a c a , N . Y . ) , O T ! Pou g h k e� p s i e ( N . Y . ) , O L ! T W ! P a r k v i l l e ( lo n g I s l a n d , N . Y . ) , A M ! Sou t h H a ven ( lo n g I s l a n d , N . Y . ) , OL ! H a rbor H i l l ( N . Y . ) , C M !

2 . Neodryin u s b a l l i n .

sp .

F E M A L E : f u l l y w i n ged ; l en g t h 4 m m ; h e a d b l ac k , w i t h c l y p e u s a n d m a n d i b l es t e s t aceo u s ; a n t e n n ae t e s t aceou s ; p ro t horax redd i s h ; meso t h o r a x , m e t a t h orax a n d p ropodeu m b l ac k ; a b domen b l ac k ; fore l e g s t e s t a ceou s ; m i d a n d h i n d l e g s b row n , w i t h t a r s i l i g h t ; a n te n n a e d i s t a l l y t h i ck e n e d ; a n t e n n a I segmen t s i n fo l l ow i n g p roport i on s : 1 4 : 6 : 24 : 1 1 : 1 0 : 7 : ( l a s t four a n t en n a l

seg m e n t s m i s s i n g

i n t h e on l y k n o w n spec i men ) ;

head f l a t ,

w i t h oce l l a r reg i o n s w o l l en , d u l l , g r a n u l a t e d a n d f u l l y ret i c u l a t e rugose ; frons w i t h a m ed i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 4 ; OL = 2 ; OOL = 1 0 , 5 ; p ro n o t u m sh i n y , smoo t h , w i t ho u t p o s t e r i or l obes d i rected t o w a r d s the t eg u l ae , w i t h a s t r o n g a n t e r i o r t r a n sv e rs a l i m p res s i on ; d i sc h u mped ; p ron o t u m w i t h o u t scu l p t u re , w i t h on l y w e a k s t r i ae a r o u n d t h e d i sc ; scu t u m d u l l , f u l l y sc u l p t u red b y l o n g i t u d i n a l p a ra l l e l k ee l s ; n ot a u l i ces i n v i s i b l e ; sc u te l l u m sh i n y , w i t h p os t e r i o r h a l f smoo t h a n d w i t hou t scu l p t u re ; a n t e r i o r h a l f p a r t l y ret i c u l a t e ru gose a n d w ea k l y

1 096

Mem . A m e r . E n t .

I n st . ,

n o . 37,

1 984

F i g . 736 - C h e l a of Neodryinus b a ll i n . sp .

( ty p e )

1 097

G o n a topod i n a e

g r a n u l a ted ; m e t a n o t u m very reduced and f u l l y re t i c u l a t e ru gose ; p ropodeum f u l l y ret i c u l a t e ru gose a n d w i t h t w o i n comp l e t e l o n g i t u d i n a l kee l s o n poster i or s u rf ace ; fore-w i n g w i t h t w o d a r k t ra n sversa I b a n d s ; d i s t a l p a r t o f ra d i a l v e i n l on g e r t h a n p ro x i m a l p a r t ; fore t a r s a l segmen t s i n fo l l ow i n g p roport i on s : 1 8 : 3 : 8 : 1 7 : 3 2 ; en l arged c l a w ( F i g . 7 3 6 ) w i t h s u b a p i c a l t o o t h a n d 6 b r i s t l es ; segmen t 5 o f f r o n t t a r s u s ( F i g . 7 3 6 ) w i t h 1 ro w of n e a r l y 2 2 l am e l l a e ; apex w i t h a g rou p of a t l ea s t 30 l a m e l l ae ; m a x i I l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 2 . MALE : u n k nown L O C U S T Y P I C U S : T i n aj a s A l t a s ( Ar i z . , U . S . A . ) TYP I CA L M A T E R I AL : h o l o t y p e F ! i n WA . HOSTS : u n k n o w n D I ST R I B U T I ON : Nearc t i c reg i on : U . S . A . : T i n a j a s A l t a s ( Ar i z . ) , NOT E S : t h e s p ec i es i s n a med i n h onor of M r . E . D . B a l l .

WA !

N E O T R O P I C S P E C I E S OF NEODR YINUS

N i n e s p ec i es a re k n o w n .

K E Y TO T H E S P E C I E S

F E I\IA L E S

S eg m e n t

2

3

4

5

5 of

f r on t

t a r s u s a l mo s t

t w i c e as

l on g a s

en l a rg ed

c I a w ( F i g . 7 3 7 ) • . . . • 1 . a c u t i co 11 i s ( K i e f f e r ) S eg m e n t 5 o f f r on t t a r s u s s l i g h t l y l o n g e r t h a n en l a rg ed c l a w ( F i g s 739 , 740 , 742 , 743, 744, 745, 746 , 747 ) . . . . . . . . . . . . . . . . 2 F o r e -w i n g w i t h ra d i a l v e i n s t r o n g l y S - s h a p ed ( F i g . 7 3 8 B , D , E ; F i g . 741 C ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .3 F o r e - w i n g w i t h ra d i a l v e i n n o t o r w e a k l y S - s h a p e d ( F i g . 7 3 8 A , C) • • • • • . • • • • • • • • • • • • • • • • • . • • • • . • . • . . • • • • . . . . . • . . • • • • • . • . • . • • • 7 D i s t a l p a r t o f ra d i a l v e i n l on g e r t h a n p r o x i ma l p a r t ( F i g . 7 3 8 D) • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • . • • • . • • . • • • • • . • • .4 D i s t a l p a r t o f ra d i a l v e i n a s l on g a s o r sh o r t e r t h a n p r ox i ma l pa r t ( F i g . 738 B , E ; F i g . 7 4 1 C ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 H ea d a n d s c u t u m g ra n u l a t ed , n o t r e t i c u l a t e r u g o s e ; b od y r ed d i sh - t e s t a c e ou s , w i t h i r r e g u l a r b r own s p o t s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 . dom inicd )l I J ,-, ( E v a n s ) H ea d a n d s c u t u m g r a n u l a t ed a n d r e t i c u l a t e r u g o s e ; b od y b l a c k . . . . . . . . . . . . . . . . . . . . . . 3 . albosign a tu s n . sp . D i s t a l p a r t o f ra d i a l v e i n a s l on g a s p r ox i ma l pa r t ( F i g . 7 3 8

Mem . A m e r . E n ! .

1 098

E )

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(Fig.

747 )

( F i gs 744 , . .8 745 ) S c u t u m f u l l y w e a k l y r e t i c u l a t e r u g o s e , w i t h ou t l o n g i t u d i n a l 6 . m a c u licornis ( C a m e r o n ) . k ee I s S c u t u m w i t h n u me r o u s I ong i t u d i n a l k e e l s , p a r t I y w i t h s i d e s s t r o n g l y r e t i cu l a t e rug o s e . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . · .. .. . . . . . . .. . .. . .. . . . .. . .. .. 7 . i n c a i c u s n . sp . •

8

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no.

D i s t a l pa r t of 738 B, 741 C ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . P r on o t u m w i t h d i s c mo r e p r o t r u d i n g ( F i g . 7 4 1 B ) . . . . . . . . . . . . . . . . . . . . . 4 . r a di a l i s n . s p . P r on o t u m w i t h d i s c l e s s p r o t r u d i n g ( F i g . 7 4 1 A ) . 8 . bra c hycerus ( K i e f f e r ) S eg m e n t 5 o f f r on t t a r s u s w i t h t w o r o w s o f l a m e l . . 9 . trin i t a tis R i c h a rd s r ow o f l a me l l a e S e g m e n t 5 o f f r on t t a r su s w i t h .

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MALES

O n l y t h e m a l e of N.

trin i t a tis R i c h a r d s

i s k n ow n .

1 . Neo dryin u s a c u ticollis ( K i ef fe r )

Dryi n u s a c u ticollis K i effer 1 90 5 b : Neodryin u s

e t Marsha I I

9.

(Psilodryi n u s ) a c u ticollis

( K i effe r ) :

K i effer i n K i ef f e r

1 906 : 497 .

Neodryin u s a c u ticollis

( K i effe r ) : R i ch a r d s 1 953 : 6 2 . ( K i effe r ) : O l m i a n d C u r rado 1 976a :

Psilodryi n u s a c u ticollis

1 85 .

F E i'<', A L E : f u l l y w i n g ed ; l e n g t h 4 , 20 m m ; h e a d b l ac k , w i t h c l y p e u s a n d m a n d i b l e s y e l l o w ; a n t e n n a e w i t h segmen t s 1 -3 y e l l ow ; o t h e r seg men ts m i s s i n g in the on l y k n o w n spec i men ; pron o t u rn b l a c k , w i t h redd i s h n u a n ces on s i des a n d pos t e r i or m a rg i n ; meso t h orax , m e t a t h o r a x a n d p ropod e u m b l a c k ; a b domen b l a c k ; l eg s y e l l ow -redd i sh , w i t h cox a e d a r k e n e d ; a n te n n a l segmen t s i n fo l l o w i n g p roport i on s : 1 6 : 8 : 3 2 ( ot h er segme n t s m i ss i n g i n t h e on l y k n o w n s p ec i me n ) ; h e a d f l a t , h a i r l es s , ret i c u l a t e r u gose ; occ i p i t a l c a r i n a i n comp l e t e , on l y v i s i b l e beh i n d t h e ocel l i ; POL � 5 ; O L � 3 ; OOL � 1 1 ; O P L � 1 ; p ro n o t u m d u l l , c rossed b y a t r a n s v e r s a l i m p res s i on , h a i r l es s , i rreg u l a r l y s t r i a t e ; pos t e r i or edges rou n de d , not p ro d uced i n t o l obes ; scu t u m h a i ry , scu l p t u red b y l on g i t u d i n a l k ee l s ; scu te l l u m f l a t , g r a n u l a t ed a n d ret i c u l a t e r u gose ;

G o n a topod i n a e

\ \

\(

F i g . 737

-

C he l a of Neodryin u s a c u ticollis ( K i effer ) ( t y p e )

p ropod e u m ret i c u l a t e ru gose a n d w i t h dors a l s u rface scu l p t u red b y l o n g i t u d i n a l k ee l s ; fore-w i n g w i t h t w o d a r k t r a n svers a l b a n d s ; d i s t a l

1 099

1 1 00

Mem . Amer . E n t .

I n st . ,

no . 37 ,

1 984

F i g . 738 - R a d i a l v e i n s of fem a l es of Neodryin u s trin ita tis R i c h a rd s ( t y p e ) ( A ) , radialis n . sp . ( t y pe ) ( B ) , incaicus n . sp . ( t y pe ) ( C ) , dom in i c a n u s ( E v a n s ) ( ty p e ) ( D ) a n d peru vianus n . sp . ( t y pe ) ( E )

1 1 01

G o n a topod i n a e

p a r t of ra d i a l v e i n m u c h l o n ge r t h a n p ro x i m a l p a r t ( 1 6 : 8 ) ; r ad i a l v e i n ( F i g . 737 ) w ea k l y S-s h a ped ; en l a r g e d c l a w w i t h s u b a p i c a l tooth a n d 6 b r i s t l es ; seg men t 5 of f ro n t t a rs u s ( F i g . 737 ) a l most t w i ce as l o n g a s en l a rged c l a w , w i t h a row o f 27 l a m e l l ae ; a p e x w i t h a g r o u p of a p p rox i m a t e l y 1 4 l a m e l l ae ; m a x i I l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 2 . MAL E : u n k n o w n LOCUS TYP I C US: M a n a g u a ( N icaragua ) HOST S : u n k n o w n D I ST R I B U T I O N : Neot rop i c reg i on : N I CARAG U A :

2 . Neodryin u s domin ica n u s

Prodryin u s domin ica n u s E v a n s 1 969 :

Managua,

GE !

( E v a n s ) n . com b .

12.

F E M A L E : f u l l y w i n g e d ; l en g t h 3 , 75 m m ; h e a d t es t aceo u s , w i t h or w i t h o u t b l ac k oce l l a r s po t ; a n t e n n ae fu scou s , w i t h seg men t 1 test aceou s ; p ron o t u m f u l l y t e s t aceo u s or w i t h t wo b l ac k spots o n t h e s i des o f t h e d i sc ; scu t u m b ro w n , w i t h a n U-shaped t es t aceo u s spot o n med i a n a re a ; sc u t e l l u m t e s t a ceo u s or b ro w n i s h ; m e t a n o t u m b row n ; p ro p o d e u m fu I I y t e s t aceo u s o r w i t h b r o w n i r reg u l a r spo t s ; a b domen a n d l e g s tes t a ceou s ; h e a d d u l l , w e a k l y e x c a v a t ed , g r a n u l a ted ; occ i p i t a l c a r i n a i n comp l e t e , o n l y v i s i b l e o n t h e s i des of t h e ace l l i ; p rono t u m c r o s s e d b y a t r a n s v e rs a l i m p ress i o n , d u l l , a s scu l p t u red a s h e a d , w i t h w e a k l o n g i t u d i n a l k ee l s ; poster i o r e d g e s of p ron o t u m rou n d e d , n o t p ro d u ced i n to l ob e s ; scu t u m h a i ry , g r a n u l a ted ; n ot a u l i ce s i n v i s i b l e ; scu te l l u m a s scu l p t u red a s scu t u m ; m e t a no t u m v e r y reduced a n d s ho r t ; p ropodeu m ret i c u l a t e rugose, w i t h s i des a n d p o s te r i o r s u rf ace w ea k l y ru gose ; p o s t e r i or s u rf ace w i t h o u t l on g i t u d i n a l k ee l s ; fore-w i n g w i t h t w o b ro w n t r a n sver s a l b a n d s ; r a d i a l v e i n ( F i g . 738 D ) s t ro n g l y S-sh a p e d , w i t h d i s t a l p a r t m u c h l o n g er t h a n p rox i m a l p a r t ; fore t roch a n ters a p p ro x i m a t e l y fou r t i mes as l o n g a s b ro a d ; en l a rged c l a w ( F i g . 739 ) w i t h s u b a p i c a l t o o t h a n d 6 p eg - l i k e h a i rs ; segmen t 5 of f ro n t t a rs u s ( F i g . 739 ) w i t h t w o rows of 1 0 + 1 0 l a m e l l ae ; a p ex w i t h a group of a t l ea s t 1 5 l a m e l l ae ; seg men t 1 of f ro n t t a r s u s l on g e r t h a n seg m e n t 4 ( 2 2 : 1 2 ) ; m ax i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 seg men t s ; t i b i a l s p u r s 1 , 0 , 2 . MALE : u n k nown LOCUS TYP I C US : ho l o t y p e F ! a n d 1 p a r a t y p e F ! i n IVA . HOST S : u n k n o w n D I ST R I BU T I O N : Neo t rop i c reg i o n : DOM I N I CA : C l a r k e H a l l ( L a y o u V a l l ey ) , WA !

3 . Neodryin u s a l bosign a tu s n .

sp .

1 1 02

Mem . Amer .

En\.

I nst . ,

n o . 37,

1 984

F i g . 739 - C h e l a of Neodryin us domin ic a n u s ( E v a n s ) from Dom i n i c a

F E MA L E : f u l l y w i n g e d ; l e n g t h 3 , 75 m m ; b l ac k ; m a n d i b l es t e s t aceo u s ; a n t e n n a e t e s t aceo u s , w i t h seg men t s 6-9 b l ac k a n d segmen t 1 0 w h i t i s h ; l eg s t e s t aceo u s , w i t h c l u b s of fore femora b l ack a n d m i d a n d h i n d cox ae p a r t l y b l ac k ; a n te n n ae d i s t a l l y t h i c k e n ed ; a n t e n n a l seg men t s in fo l l o w i n g

1 1 03

Gon a topod i n ae

Fig.

p ro p o r t i o n s :

740 - C h e l a of Neodryin u s a lbosign a t u s n . sp .

14:

5:

30 :

15:

13:

9:

7:

6:

5:

9;

head d u l l ,

( type)

exc a v a ted ,

g r a n u l a t ed a n d re t i c u l a t e r u g ose ; f ro n s w i t h a comp l e t e m ed i a n k ee l ; occ i p i t a l c a r i n a i ncomp l et e ,

v i s i b l e beh i n d t h e oce l l i

a n d on t h e s i des of

t h e pos t e r i or oce l l i , reach i n g a p p rox i ma te l y 0 , 5 l e n g t h of OOl ; POL = 3 ; O l = 2 , 5 ; OOl = 1 0 ; pos t e r i o r oce l l i tou c h i n g t h e o c c i p i t a l c a r i n a ; t e m p l es i n v i s i b l e ; p ron o t u m cros sed by a s t ro n g t ra n s v ers a l i m p re ss i o n , d u l l , g r a n u l a ted ; p ron o t u m n o t p ro d u ced i n t o l o bes d i rec t ed t o w a r d s t h e t e g u l ae ; d i sc h u mped ; scu t u m d u l l , g r a n u l a t ed a n d ret i c u l a t e r u g ose ; n o t a u l i ces i n v i S i b l e ; s c u t e l l u m as scu l p t u red as scu t u m ; m e t a n o t u m very s h o r t , reduced , d u l l ; p ro p o d e u m w i t h dors a l s u rface scu l p t u red b y n u me ro u s , p a ra l l e l , l o n g i t u d i n a l k ee l s , i n t e r r u p ted b y t r a n s v e rs a l k ee l s ; po s t er i or s u rface ret i c u l a t e ru gose a n d w i t h t wo l on g i t u d i n a l k ee l s ; fore w i n g w i t h t w o d a rk t r a n s v e rs a l b a n d s ; d i s t a l p a r t of rad i a l v e i n l on g er t h a n p ro x i m a l p a r t ( 1 8 : 1 1 ) ; rad i a l v e i n s t ron g l y S-s haped ; fore t a r s a l s e g m e n t s i n fo l l o w i n g p ropor t i o n s : 2 2 : 3 : 5 : 1 0 : 1 8 ; en l arged c l a w ( F i g . 740 ) w i t h s u b a p i c a l tooth a n d 6 b r i s t l es ; s e g m e n t 5 of f ro n t t a rs u s ( F i g . 740 ) w i t h t w o rows of n e a r l y 2 5 l a me l l ae e x t en d i n g c on t i n u o u s l y t o t h e apex ; m a x i I l a ry p a l p i w i t h 6 s e g m e n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 2 . MA L E : u n k n ow n lOCUS T Y P I C U S : 50° 30 ' W , 1 2 ° 30 ' S ( V i l a V e r a , T Y P I C A L M A T E R I A L : ho l o t y p e F ! i n T W . HOSTS : u n k n o w n D I ST R I BU T I O N : TW !

Neot rop i c reg i o n :

BRAZ i l :

Braz i l )

50° 30 ' W 1 2 ° 30 ' S ( V i l a Vera ) ,

1 1 04

Mem . Amer . E n ! .

I ns! . ,

n o . 37 ,

1 984

A

B

v

c

F i g . 741 - P ro n o t u m ( l a tera l l y v i ewed ) ( A ) a n d rad i a l v e i n ( C ) of fema l e o f Neodryin u s brachycerus ( K i effe r ) ( t y p e ) ; p ron o t u m ( l a t e ra l l y v i ewed ) of f em a l e of Neodryin u s radialis n . sp . ( t y p e ) ( B )

4 . Neodryin u s radia l is n . sp .

F E M A L E : f u l l y w i n ged ; l e n g t h 4 , 50-6 , 00 m m ; h e a d b l ac k , w i t h m a n d i b l e s y e l l o w ; a n te n n a e testaceo u s , w i t h segmen t s 1 -2 a n d 7- 1 0 b ro w n ; t h o r a x a n d p ropodeum b l ack ; a b domen b ro w n ; l e g s b row n , w i t h t a r s i l i g h t ; h e a d d u l l , ret i c u l a t e ru gose, h a i ry ; occ i p i t a l c a r i n a i n com p l e t e , o n l y v i s i b l e b eh i n d t h e oce l l i a n d o n t h e s i des of t h e pos t e r i o r oce l l i ; pronot u m d u l l , g r a n u l a t e d , scu l p t u red b y n u merous l o n g i t u d i n a l k ee l s ( more n u merous o n t h e s i des of t h e co l l a r ) , c rossed b y a s t ron g t r a n s v e r s a l i m p res s i o n ; d i sc r a i se d i n to a c a r i n a ; s c u t u m d u l l , g r a n u l a t e d , w i t h n u merous l o n g i t u d i n a l k ee l s ; n o t a u l i ce s i n v i s i b l e ; sc u t e l l u m d u l l , g r a n u l a t e d a n d ret i c u l a te ru gose ; m e t a n o t u m short a n d red u c ed ; propodeum re t i c u l a t e ru gose , w i t h t w o l on g i t u d i n a l k ee l s o n poster i o r su rf ace ; fore-w i n g w i t h t w o b ro w n t r a n sv e rs a l b a n d s ; d i s t a l p a r t of r a d i a l v e i n ( F i g . 738 B ) s t ron g l y S-s h a ped , shorter t h a n p rox i m a l p a r t ; segmen t 1 o f fron t t a r s u s n e a r l y t w i c e a s l o n g a s s e g m e n t 4 ( 22 : 1 0 ) ; e n l a r ged c l aw ( F i g . 742 ) w i t h

1 1 05

Gon a topod i n ae

F i g . 742 - C h e l a of Neodryin u s r a d i a l i s n .

suba p i c a l 742 ) w i t h m a x i l l a ry s p u rs 1 , 0 ,

sp .

( type)

tooth a n d 5 peg - l i k e h a i r s ; seg m e n t 5 of fron t t a r s u s ( F i g . two rows of 1 9 l a m e l l a e ; apex w i t h a g ro u p of 1 4 l a me l l ae ; p a l p i w i t h 6 segmen ts ; l a b i a l p a l p i w i t h 3 segmen ts ; t i b i a l 2.

1 1 06

Mem . Am e r . E n ! .

I n s! . ,

no . 37,

1 984

F i g . 743 - C h e l a o f Neodryin u s peruvianus n . s p ,

MALE : u n k no w n L O C U S T Y P I C U S : Nov a T e u t o n i a ( B raz i l ) TYP I C AL MAT E R I AL : h o l o t y p e F ! i n B M ;

1

paratype F !

( type)

i n TW .

Gon a topod i n a e

HOSTS : u n k n o w n D I ST R I B U T I O N : Neot rop i c reg i o n : BRAZ I L : BM ! Teodoro S a m p a i o ( Sao P a u l o ) , TW !

1 1 07

Nov a Teuton i a ( S . t a C a t a r i n a ) ,

5 . Neodryin u s peruvia n u s n . s p .

F E M AL E : f u l l y w i n ged ; l e n g t h 5 , 3 1 m m ; h e a d b l ac k , w i t h m a n d i b l es tes t aceo u s ; a n te n n ae test aceou s , w i t h seg m e n t s 6-1 0 b row n ; t h ora x , propode u m a n d a b domen b l ac k ; l e g s test aceo u s , w i t h coxae p a r t l y b l ack ; c l u b s of femora a n d t i b i ae p a r t l y b row n ; h e a d d u l l , r e t i c u l a te r u gose, h a i ry , exc a v a ted ; occi p i t a l c a r i n a on l y v i s i b l e beh i n d the oce l l i a n d o n t h e s i d e s of t h e poster i or oce l l i ; p ro n o t u m d u l l , c r o s s e d b y a s t ron g t r a n s versa l i m p re s s i o n , gra n u l a t e d , w i t h n u me ro u s l o n g i t u d i n a l k ee l s o n t h e s i de s of t h e c o l l a r ; d i sc ra i sed i n to a c a r i n a ; s c u t u m d u l l , g r a n u l a ted a n d ret i c u l a t e ru gose ; n o t a u l i c e s i n v i s i b l e ; s c u t e l l u m d u l l , ret i c u l a t e r u gose ; m e t a n o t u m very reduced a n d short ; p ropodeum ret i c u l a t e ru gose, w i t h t w o l o n g i t u d i n a l k ee l s on p o s te r i or s u rface ; fore-w i n g w i t h t w o b ro w n t r a n s v e rs a l b a n d s ; d i s t a l p a r t of rad i a l v e i n ( F i g . 738 E ) stron g l y S-sh a p e d , a s l o n g a s p ro x i m a l p a r t ; seg men t 1 of fron t t a r s u s l o n ger t h a n seg men t 4 ( 1 7 : 1 0 ) ; en l arged c l a w ( F i g . 743 ) w i t h s u b a p i ca l tooth a n d 8 b r i s t l es ; seg m e n t 5 of f ro n t t a r s u s ( F i g . 743 ) w i t h t w o row s of 20 l a me l l a e ; a p e x w i t h a g ro u p of at l ea s t 20 l a me l l ae ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 seg men t s ; t i b i a l s p u r s 1 , 0 , 2 . MALE : u n k n o w n L O C U S TYP I C U S : P u c a l l p a ( Peru ) TYP I CAL M A T E R I AL : h o l o t y p e F ! i n C A . HOSTS : u n k n own D I ST R I BU T I O N : Neot rop i c reg i on : P E R U : P u c a l l p a , C A l

6 . Neodryin u s m a c u l icornis ( C a m eron )

Dryin u s m a c ulicornis C a meron 1 888 : 444 . Dryin us a l ticola C a meron 1 888 : 444 ( n . s y n . ) ; locus typicus V o l c a n de C h i r i q u i ( P a n a m a ) . Prodryin u s a l ticola ( C ameron ) : K i effer 1 9 1 4b : 5 2 . Prodryin u s m a c ulicornis ( C a meron ) : K i effer 1 9 1 4b : 5 2 . Neodryin u s a lticola ( C ameron ) : R i c h a r d s 1 953 : 6 8 . Neodryin u s m a c ulicornis ( C amero n ) : R i c h a r d s 1 953 : 69 .

F E M AL E : f u l l y w i n g e d ; l en g t h 4 , 68-5 , 00 m m ; h e a d b l ac k , w i t h m a n d i b l es y e l l ow ; c l y pe u s y e l l ow or b l ack ; a n te n n a e b row n , w i t h segmen t 5 y e l l o w ; occ a s i o n a l l y a n t e n n ae y e l l o w ,

w i t h seg m e n t s 5- 1 0 b ro w n ; p ro no t u m b l ac k ,

Mem . A me r . E n t .

1 1 08

F i g . 744

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1 1 09

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w i t h s i d es r e d d i s h ; meso t h o r a x , m e t a t ho r a x a n d p ro p o d e u m b l ac k ; a b domen b r o w n - b l ack ; l e g s b ro w n , w i t h t a r s i l i g h t ; h e a d w ea k l y exc a v a t ed , d u l l , f u l l y ret i c u l a t e r u gose ; occ i p i t a l c a r i n a i n com p l e t e , o n l y v i s i b l e o n t h e s i de s of t h e oce l l i ; p ro no t u m d u l l , h a i ry , c rossed b y a t r a n s v e rs a l i m press i o n , g r a n u l a t ed ; p o s t e r i or edges of t h e p ro n o t u m rou n d e d , n o t p ro d u ced i n t o l obes ; s c u t u m h a i ry , g r a n u l a t ed , f u l l y ret i c u l a t e r u gose ; n o t a u l i ces i n v i s i b l e ; s c u t e l l u m as scu l p t u red as sc u t u m ; m e t a n o t u m v e r y reduced a n d s h o r t ; p ropodeu m f u l l y re t i c u l a t e ru gose, w i t h t w o l o n g i t u d i n a l k ee l s o n p o s t e r i o r s u rface ; fore-w i n g w i t h two b ro w n t r a n s v e rs a l b a n d s ; ra d i a l v e i n n o t o r weak l y S-sh a p e d , w i t h d i s t a l p a r t m u c h l on g e r t h a n p rox i m a l p a r t ; fore t r oc h a n ters a p p rox i m a t e l y f o u r t i mes a s l o n g a s b road ; seg men t 1 of f r o n t t a r s u s l on g e r t h a n seg m e n t 4 ( 25 : 1 7 ) ; e n l a rged c l a w ( F i g . 744 ) w i t h s u b a p i c a l t o o t h a n d 8 b r i s t l es ; seg m e n t 5 of f r o n t t a r s u s ( F i g . 744 ) w i t h 1 row of 22 l a m e l l ae ; a p ex w i t h a g r o u p of a t l ea s t 1 5 l a m e l l ae ; m a x i l l a r y p a l p i w i t h 6 s e g me n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 2 . MALE : u n k n o w n L O C U S T Y P I C U S : M i ra n d i l l a ( P a n a m a ) T Y P I CA L M A T E R I A L : h o l o t y p e F ! of D. D. a l ticola i n B M . H OST S : u n k n o w n D I ST R I BU T I O N : Neot rop i c reg i on : V o l c a n de C h i r i q u i , BM !

m a c u l icornis i n BM ; h o l o t y p e F ! of

PANAI,IA :

M i ran d i l l a ,

7 . Neodryin u s inca icus

n.

BM !

Bugaba,

BM !

sp .

F E M A L E : f u l l y w i n g ed ; l e n g t h 3 , 56-5 , 30 m m ; h e a d b l ack , w i t h m a n d i b l es t e s t aceou s ; a n t e n n a e t e s t aceo u s , w i t h seg men t s 6-1 0 b row n ; t h o r a x a n d p ro p o d e u m b l a c k ; a b domen b row n ; l e g s t e s t a ceo u s , w i t h cox a e p a r t I y b l ack ; c l u b s of femora p a r t l y b row n ; some t i me s a l so t i b i ae b r ow n ; h e a d w e a k l y e x ca v a ted , d u l l , f u l l y ret i c u l a t e r u gose, h a i r y ; occ i p i t a l c a r i n a on l y v i s i b l e beh i n d t h e oce l l i a n d o n t h e s i des o f t h e p o s t e r i o r oce l l i ; p ro no t u m d u l l , c rossed b y a t r a n s v e rs a l i m p ress i on , h a i r y , g r a n u l a t ed ; p o s t e r i o r e d g e s of t h e p r o n o t u m rou n d e d , n o t p ro d u ced i n to l obes ; scu t u m h a i ry , g r a n u l a ted , w i t h n u merous l on g i t u d i n a l k ee l s , p a r t l y w i t h s i des ret i c u l a t e r u go s e ; n o t a u l i ces i n v i s i b l e ; s c u t e l l u m d u l l , g r a n u l a t ed , ret i c u l a t e r u gose n e a r a n t e r i o r m a rg i n ; m e t a n o t u m v e r y red uced a n d s h o r t ; p ro p od e u m f u l l y re t i c u l a t e r u g o s e , w i t h t w o l o n g i t u d i n a l k ee l s o n pos ter i o r s u rf ace ; fore-w i n g w i t h t w o b r o w n t ra n s v e rs a l b a n d s ; r a d i a l v e i n ( F i g . 738 C ) w e a k l y S-sh a ped , w i t h d i s t a l p a r t m u c h l on g e r t h a n p r ox i m a l p a r t ; fore t roc h a n t e r s a p p ro x i m a te l y f o u r t i mes a s l o n g a s b ro a d ( 27 : 6 ) ; s e g m e n t 1 of f ro n t t a r s u s a s l o n g a s 4 ; en l a r g e d c l a w ( F i g . 745 ) w i t h s u b a p i c a l t o o t h a n d 7-8 b r i s t l es ; s e g m e n t 5 of f r o n t t a r s u s ( F i g . 745 ) w i t h 1 row of 1 5-20 l a m e l l a e ; a pex w i t h a g r o u p of a t l ea s t 30 l a m e l l a e ; m a x i l l a r y p a l p i w i t h 6 s e g me n t s ; l a b i a l p a l p i w i t h 3 s e g me n t s ;

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paratype F ! in OL . F ! i n OT ; 1 p a r a t y p e F ! i n AM ; H O ST S : u n k n o w n D I ST R I BU T I O N : N e a rc t i c reg i on : U . S . A . : Bear W a l l ow ( S ! . C a t a l i n a M t s . , Ar i z . ) , A M ! Neot rop i c reg i o n : M E X I C O : E I P a l m i to ( S i n a l oa ) , OT ! V E N E Z U E LA : S a n E s t e b a n , O L ! V e n ez u e l a , WA ! P E R U : S a n J o r g e , C A l

8 . Neod""yin u s brachycerus ( K i effer )

Dryi n u s brachycerus K i effer 1 904a : 3 5 1 . Prodryin u s brachycerus ( K i effer ) : K i effer i n K i effer et M a r s h a I I 1 906 : 497 . Prodryin u s brachycerus ( K i effer ) : B r u c h 1 9 1 5 : 443 . Neodryin u s brachycerus ( K i effer ) : R i c h a r d s 1 953 : 6 1 . Prodryin u s brachycer u s ( K i effer ) : O l m i a n d C u rr a d o 1 976a : 1 88 .

F E M AL E : f u l l y w i n ged ;

l e n g t h 4 , 07 m m ; h e a d b l ac k ,

y e l l ow-brow n ; c l y p e u s b l ac k ; a n t e n n ae b row n , t horax ,

w i t h m a n d i b l es

w i t h v e n t ra l f ac e of

segmen t 1

wh i t i s h ;

w i th t a r s i

l i g h t ; a n t en n a l segmen t s i n fo l l o w i n g p ropor t i on s :

p ropodeum a n d a b domen b l ac k ;

l eg s b l ac k , 1 5 : 6 : 26 :

1 5 : 1 2 : 9 : 7 : 6 : 6 : 9 ; h e a d w e a k l y exca v a ted , h a i r l es s , d u l l , ret i c u l a t e ru gose ; occ i p i t a l c a r i n a i n com p l e t e , o n l y v i s i b l e b eh i n d t h e o ce l l i ; P O L = 3 ; OL = 3 ; OOL = 1 3 ; pronot u m ( F i g . 7 4 1 A ) g r a n u l a ted , w e a k l y s t r i a t e , crossed b y a t r a n sversa l i m p re s s i on ; pos t e r i o r e d g e s o f pron o t u m rou n de d , n o t p ro d uced i n to l obes ; sc u t u m h a i ry , s c u l p t u red b y l o n g i t u d i n a l k ee l s ; scu t e l l u m f l a t , a l u t aceou s ; met a n o t u m v e r y redu ced a n d s h o r t ; p ropod e u m ret i c u l a t e ru gose , w i t h dors a l s u rface scu l p t u red b y l on g i t u d i n a l k ee l s ; fore-w i n g w i t h t w o d a r k t ra n s v e rs a l b a n d s ; ra d i a l v e i n ( F i g . 74 1 C ) s t ron g l y S-s h a p e d , w i t h d i s t a l p a r t m u c h shorter t h a n p rox i m a l p a r t ; en l a r ged c l a w ( F i g . 746 ) w i t h s u b a p i c a l tooth a n d 5 b r i s t l es ; segmen t 5 of fron t t a rs u s ( F i g . 746 ) w i t h t w o rows of 4 + 9 l a m e l l a e ; apex w i t h a g ro u p of 25-30 l a m e l l ae ; m ax i l l a ry p a l p i w i t h 6 s e g me n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 2 . M A L E : u n k no w n L O C U S TYP I C U S : T i g r e ( A r g e n t i n a ) TYP I CA L M A T E R I AL : h o l o t y p e F ! i n G E . HOSTS : u n k n o w n D I STR I BU T I O N : Neot rop i c reg i o n : A R G E N T I NA : T i g r e , G E !

9 . Neodryin u s trim"ta tis R i c h a rd s

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1 984

F i g . 746 - C h e l a of Neodryin u s bra chycerus ( K i effer ) ( t y p e )

Neodryin u s trin ita tis R i ch a r d s 1 95 1 : 8 1 6 .

Gon a topod i n a e

Fig.

1113

747 - C h e l a of i'ieo dryin u s trin ita tis R i c h a rds ( t y p e )

F E M AL E : f u l l y w i n g e d ; l e n g t h 4 , 68-6 , 00 m m ; h e a d b l ac k , w i t h m a n d i b l es redd i s h ; a n t e n n a e b row n , w i t h segmen t 5 , d i s t a l p a r t of segmen t 4 a n d ven t r a l f a c e o f segme n t 1 y e l l ow ; p ro n o t u m b l ac k , w i t h s i des p a r t l y

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F i g . 748 - M a l e g en i t a l i a of Neodryin u s trin ita tis R i ch a rds ( a l l o t y p e )

redd i sh ; mesothora x , met a t h ora x , propodeum a n d a b domen b l ac k ; l eg s redd i s h , w i t h cox a e p a r t l y b l ac k ; h e a d w e a k l y exc a v a t e d , w i t h f ro n s a n d v ertex s t ron g l y or w e a k l y re t i c u l a te r u gose, somet i me s w i t h w e a k l o n g i t u d i n a l k ee l s ; occ i p i t a l c a r i n a i n comp l e t e , e x t e n d i n g b e y o n d 0 , 5 l e n g t h of OOL ; p ro n o t u m crossed b y a t r a n svers a l i m p ress i o n , h a i r y , d u l l , g r a n u l a t e d ; d i sc r a i sed i n t o a c a r i n a ; pos t er i or edges o f p ro no t u m rou n d e d , n o t p rod u ced i n t o l obes ; sc u t u m d u l l , a s scu l p t u red a s prono t u m , w i t h n u me ro u s , p a ra l l e l a n d l on g i t u d i n a l k ee l s ; n o t a u l i ce s

Gon a topo d i n ae

1115

i n v i s i b l e ; sc u te l l u m as scu l p t u red as sc u t u m , b u t w i t h o u t k ee l s ; m e t a n o t u m v e r y red uced a n d short ; p ropodeum f u I I y ret i c u I a t e ru gose, w i t h two l o n g i t u d i n a l k ee l s on t h e post e r i or s u rf a c e ; fore-w i n g w i t h two b ro w n t r a n svers a l b a n d s ; r a d i a l v e i n ( F i g . 738 A) not or w e a k l y S s h a p e d , w i t h d i s t a l p a r t m u c h l on g e r t h a n prox i m a l p a r t ; fore troc h a n ters a p p rox i m a t e l y four t i mes a s l o n g a s b roa d ; segmen t 1 of f ro n t t a rs u s l on g e r t h a n seg m e n t 4 ( 1 9 : 1 4 ) ; e n l a rged c l a w ( F i g . 747 ) w i t h s u b a p i c a l too t h a n d n u me ro u s b r i s t l es ; seg m e n t 5 of fron t t a rs u s ( F i g . 747 ) w i t h two rows of 1 0 + 7 l a m e l l ae ; a p ex w i t h a group o f a t l ea s t 1 5 l a m e l l ae ; m a x i I l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 2 . MA L E : f u l l y w i n g e d ; l e n g t h 2 , 37 m m ; h e a d b l ac k , w i t h m a n d i b l es red d i s h ; a n te n n a e b row n ; t horax a n d p ropod e u m b l ack ; a b domen b ro w n ; l eg s b row n , w i t h t a r s i l i g h t ; a n te n n a l segme n t s i n fo l l ow i n g p ropor t i on s : 6 : 5 : 1 2 , 5 : 9 : 9 : 1 0 : 8 : 8 : 7 : 9 ; a n te n n a l seg m e n t 3 more t h a n t h ree t i me s a s l o n g a s b road ( 1 2 , 5 : 2 , 5 ) ; h e a d s h i n y , g r a n u l a ted , w i t ho u t med i a n k e e l b e t w ee n t h e a n t en n a l soc k e t s ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L � 8 ; O l 3 ; OOl � 6 ; b r e a d t h of t h e oce l l i s h o r t e r t h a n OOl ( 3 : 6 ) ; temp l es i n v i s i b l e ; sc u t u m d u l l , g r a n u l a ted ; no t a u l i c e s i ncom p l e t e , reach i n g a p p rox i m a te l y 0 , 5 l e n g t h of scu t u m ; scu t e l l u m a n d met a n o t u m a s scu I p t u red a s scu t u m ; p ropodeum w i t h a med i a n f u rrow o n a n te r i or s u r f ace ; two l a tera l , s i des of t h i s f u rrow ;

smooth reg i o n s w i t h o u t sc u l p t u r e a re v i s i b l e on t h e poster i or s u rf a c e r u g ose , w i t h o u t areo l ae a n d

l o n g i t u d i n a l k ee l s ; fore-w i n g h y a l i n e , w i t h o u t d a r k t r a n s v e r s a l b a n d s ; d i s t a l p a r t of rad i a l v e i n n e a r l y as l o n g as p rox i m a l p a r t ; g en i t a l i a w i t h dor s a l p rocess of gonoforceps ( F i g . 748 ) l o n g , s l en d e r a n d po i n t ed ; m a x i l l a ry p a l p i w i t h 6 segme n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 2 . lOCUS T Y P I C U S : St . A u g u s t i n e ( T r i n i d a d ) TYP I C AL M A T E R I AL : ho l ot y p e F ! , a l l ot y p e M ! a n d 4 p a ra t y pes ( 3 M M , 1 F ) ! i n BM . HOSTS : i n T r i n i d a d Ormenis sp . on c a c a o . D I ST R I B U T I O N : N eo t rop i c reg i o n : T R I N I DA D : St . A u g u s t i n e , BM ! O L ! WA ! BRAZ i l : S . t a R i t a do C e d ro ( M i n a s ) , M D ! E C UA D O R : L i moncoc h a ( N a po ) , OT !

AUST RAL I AN S P E C I E S OF NEODR YINUS

F o u r spec i e s a re k now n .

K E Y TO T H E S P E C I E S

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P o s t e r i o r su r f a c e o f l ong i t u d i na I k e e I s • • • . . • • . • • • • • • . • • . • • • • • • • • • • • • • • • • • • • • • • • • • 2 O c c i p i t a l c a r i n a i n v i s i b l e o r i n c omp l e t e ( i n t h i s c a s e o n l y v i s i b l e b eh i n d t h e o c e l l i o r on t h e s i d e s o f t h e p o s t e r i o r o c e l I i , b u t n ev e r r e a ch i n g 0 , 5 l e n g t h o f OOL ) . . . • . • • • • . . . • . • • • . . . • . . . . . . . . . . . . . . . . . . . . . . 2 . nelsoni R . C . L . P e rk i n s O c c i p i t a l c a r i n a r ea c h i n g a p p r ox i ma t e l y 0 , 5 l en g t h o f OOL . . . . 3 S c u t u m f u l l y r e t i c u l a t e ru g o s e ; en l a rg ed c l a w mo r e t a p e r i n g , w i t h s u b a p i c a l t oo t h s ma l l e r ( F i g . 7 5 1 A ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 . koebelei R . C . L . P e rk i n s S c u t u m n o t r e t i c u l a t e ru g o s e , w i t h n u me r o u s l on g i t u d i n a l k e e l s ; en l a rg ed c l a w l e s s t a p e r i n g , w i t h s u b a p i ca l t o o t h b i g ( F i g . 7 5 3 ) . . . . . . . . . . . . . . . . 4 . a m b on i n . sp .

MALES

O n l y t h e m a l e of N.

koebelei i s k n ow n .

R . C . L . P e r k i n s ( 1 90 5 ) desc r i bed a l so t h e m a l e of N. raptor R . C . L . P e r k i n s ( � n e lsoni R . C . L . Perk i n s ) . I h a v e s t u d i ed a m a l e determ i n ed a s

raptor b y t h e s a m e R . C . L . P e rk i n s : nigrica n s ( R . C . L . Perk i n s ) .

i t ' s rea l l y a m a l e o f Pseudogona top u s

1 . Neodryin u s cheesmani n .

sp .

F E M AL E : fu l l y w i n g e d ; l e n g t h 5 , 0 m m ; h e a d b l ac k , w i t h s i de s of c l y pe u s a n d m a n d i b l es y e l l ow ; a n t e n n a e w i t h segme n t s 1 -4 y e l l ow ; o t h er segmen t s m i ss i n g i n t h e on l y k no w n s p ec i me n ; t ho r a x a n d p ropodeu m b l ac k , w i t h red d i sh n u a nces o n p o s t e r i o r m a r g i n o f d i sc o f p ro n o t u m ; red d i s h n u a n ce s a l so a re v i s i b l e o n s i des of p ro n o t u m ; a b domen b l ac k ; l e g s y e l l o w , w i t h m i d a n d h i n d cox a e b row n ; c l u b s of femora b row n ; h e a d s h i n y , exca v a t e d , w i t h o u t scu l p t u re , w i t h fron t p a r t o f v ertex a n d occ i p u t g r a n u l a t e d ; occ i p i t a l c a r i n a i n com p l e t e , reac h i n g a p p rox i m a t e l y 0 , 5 l e n g t h o f OOL ; p ro n o t u m crossed b y a t r a n sv e rs a l i m p ress i o n , w i t h n u merous l on g i t u d i n a l k ee l s , w ea k l y g r a n u l a t e d ; poster i or e d g e s of p ron o t u m rou n d e d , not p ro d u ced i n to l obes ; scu t u m d u l l , g ra n u l a ted , w i t h n u merous l o n g i t u d i n a l k e e l s ; n ot a u l i ces i n v i s i b l e ; sc u t e l l u m g r a n u l a t ed , h a i ry ; m e t a n o t u m v e r y red uced a n d s h o r t ; p ropodeum f u l l y ret i c u l a t e ru gose , w i t h o u t l o n g i t u d i n a l k ee l s o n p o s t e r i o r s u rf ac e ; a n te r i or s u rface n ea r l y 1 , 5 t i me s a s long a s poster i o r s u rf ace ; fore-w i n g w i t h t w o b ro w n t r a n sv e r s a l b a n d s ; f o r e t roc h a n ters a p p rox i m a t e l y fou r t i mes a s l o n g a s b road ; s e g m en t 1 of fron t t a r s u s s l i g h t l y l on g e r t h a n segmen t 4 ( 2 1 : 20 ) ;

1 1 17

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en l a rged c l a w ( F i g . 749 ) w i t h su b a p i ca l t oo t h a n d 8 l a me l l ae ; segmen t 5 of f ro n t t a rs u s ( F i g . 749 ) w i t h t w o ro w s of 1 4 + 1 0 l a m e l l ae ; a pex w i t h a g ro u p of a t l e a s t 1 2 l a m e l l ae ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 2 .

l ab i a l

M A L E : u n k no w n LOCUS T Y P I C US : h o l o t y p e F ! i n B M . HOSTS : u n k n ow n D I ST R I B U T I ON : A u s t r a l i a n reg i on : NEW G U I N E A : M t . B a d u r i ( J a p en I s l a n d ) , BM ! N O T E : t h e spec i es i s n a me d i n honor of M r . L . E . C hees m a n .

2 . Neodryin u s nelsoni R . C . L . Perk i n s

Neodryin u s nelsoni R . C . L . P e rk i n s 1 90 5 : 5 2 . Neodryin u s rap tor R . C . L . Perk i n s 1 90 5 : 5 2 ( n . s y n . ) ; locus typicus B u n d a berg ( Au s t r a I i a ) • Neodryinus raptor R . C . L . Perk i n s v a r . umbratus R . C . L . P e r k i n s 1 905 : 5 2 ( n . s y n . ) ; locus typicus Queens l a n d ( Au s t r a l i a ) . n e c Neodryin u s nelsoni R . C . L . Per k i n s : Ponomarenko 1 98 1 b : 881 .

F E M A L E ( F i g s 7 1 5 , 7 1 6 ) : f u l l y w i n ged ; l e n g t h 3 , 66-5 , 49 m m ; h e a d b l ac k , w i t h s i d es o f c l y pe u s a n d m a n d i b l e s y e l l o w ; a n t en n ae y e l l ow ; t h o r a x a n d p ropode u m b l ac k , w i t h red d i sh n u a nces on s i des o f co l l a r ; red d i sh n u a nces a re a l so v i s i b l e on p os t e r i or m a rg i n of d i sc ; a b d omen b row n ; l eg s b ro w n , w i t h t a r s i l i g h t ; h e a d w e a k l y exca v a t e d , sh i n y ; ver tex w i t h n u mero u s l on g i t u d i n a l k ee l s , g r a n u l a ted a mon g k ee l s ; occ i p i t a l c a r i n a i n v i s i b l e o r w e a k l y v i s i b l e on s i des of oce l l i ; p ro n o t u m s h i n y , crossed b y a t r a n svers a l i m p ress i on , w i t h n u merous l o n g i t u d i n a l k ee l s o n co l l a r a n d d i sc ; d i sc a n d p a r t l y co l l a r g r a n u l a ted ; poster i o r edges o f p ronot u m rou n d e d , n o t p ro d u ced i n to l obes ; sc u t u m f u l l y ret i c u l a te r u gose, w i t h t w o i ncomp l e te l o n g i t u d i n a l k ee l s on pos ter i o r su rface ; a n t e r i o r s u rface of p ropod e u m n ea r l y a s long a s posterior s u r face ( 25 : 22 ) ; fore-w i n g w i t h t w o b row n t r a n sversa l b a n d s ; d i s t a l p a r t o f r a d i a l v e i n m u c h l on ger t h a n p rox i m a l p a r t ; fore t roc h a n t e r s a p p rox i m a te l y f o u r t i mes a s l o n g a s b road ; segmen t 1 of fron t t a r s u s s l i g h t l y shor ter t h a n segmen t 4 ( 1 5 : 1 6 ) ; e n l a rged c l a w ( F i g . 750 ) w i t h s u b a p i c a l too t h a n d 8 l a me l l a e ; segmen t 5 of f ro n t t a r s u s ( F i g . 750 ) w i t h t w o rows of 25 l a me l l ae ; a pex w i t h a g rou p of a t l ea s t 1 7 l a m e l l a e ; m ax i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 2 . M A L E : u n k n ow n ( see p a ge 1 1 1 6 ) . L O C U S T Y P I C U S : C a i r n s ( A u s t ra l i a ) T Y P I C AL MAT E R I AL :

l ec t o t y p e F ! a n d 1

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O l m i 1 982 ) of tv.. nelsoni in B ; l ec t o t y p e F ! and 4 p a ra l ec to t y pes ( 2 F F , 2 MM ) ! ( de s i g n a ted b y O l m i 1 98 2 ) of N . raptor i n B ; t y p e F ! o f N . raptor v a r . umbratus p e r h a p s l o s t ( O l m i 1 982 ) . HOSTS : i n A u s t ra I i a Fl a tid a e : Colgar pera c u t u s , Sip h a n ta sp . , Massila sp . , Sephena sp . , Ph a n tiopsis (� A p h a n oph a n t i a ) s p . ; Ricaniida e : Privesa s p . , Scolypopa s p . ; Nogodinida e : Gaetulia sp . ( R . C . L . P e r k i n s 1 906c ) , Privesa aphrophoroides W a I k . ( T hompson 1 9 54 ) . D I ST R I BU T I O N : A u s t ra l i a n reg i o n : AUSTRAL I A : C a i r n s ( Q u ee n s l a n d ) , B ! B u n d a b e r g ( Qu e e n s l a n d ) , OL ! B ! B r i s b a n e ( Qu e e n s l a n d ) , Q U ! Jo l i mo n t C reek ( M a ck a y , Queen s l a n d ) , B M ! Sa r i n a R a n g e ( 1 1 K m S S a r i n a , N Queen s l a n d ) , B R ! N OT E : N. nelsoni w a s recorded by m i s t a k e from Sri L a n k a by P o n o m a r e n k o ( 1 98 1 b ) . T h e spec i me n s e e n b y Mrs . P o n o m a r e n k o i s rea l l y a n o t h er spec i es of Neodryin u s n o t recog n i za b l e bec a u se t h e h e a d i s m i ss i n g . i s n o t a nelson i .

I t,

however,

3 . Neodryin u s koebelei R . C . L . Perk i n s

Neodryin u s koebelei R . C . L . Perk i n s 1 90 5 : 5 1 . Neodryin u s koebelei R . C . L . P e rk i n s : Dodd 1 9 1 4c : 2 . nec Neo dryin u s koeb e l i R . C . L . P e rk i n s : P o n o m a r e n k o 1 98 1 b : 882 .

F E MAL E : f u l l y w i n g e d ; l en g t h 4 , 39-5 , 85 m m ; h e a d b l ack , w i t h s i de s of c l y pe u s a n d m a n d i b l es y e l l ow ; a n t e n n a e y e l l ow ; t h o r a x a n d p ropodeum b l ac k , excep t for redd i sh n u a n ces o n s i des of p ro n o t u m ; red d i s h n u a n ces a r e v i s i b l e a l so o n p os t e r i o r m a rg i n of p ro n o t u m ; abdomen brow n ; l eg s brow n , w i t h t a r s i l i g h t ; h e a d e x c a v a t ed , sh i n y ; ver tex w i t h n u merous l o n g i t u d i n a l k ee l s and w i t h o u t scu l p t u re among k ee l s ; f ro n t part of v e r t e x g r a n u l a te d a n d r u gose ; occ i p i t a l c a r i n a i n com p l e t e , reac h i n g a p p ro x i ma t e l y 0 , 5 l e n g t h o f OOL ; p ron o t u m s h i n y , crossed b y a t ra n s v e rs a l i m p re s s i on , w i t h n u merous l on g i t u d i n a l k ee l s on co l l a r a n d d i sc ; p o s t e r i o r e d g e s o f p r o n o t u m rou n ded , n o t p ro d u ced i n t o l obes ; sc u t u m f u l l y re t i c u l a t e r u g o s e , s h i n y ; n o t a u l i ce s i n v i s i b l e ; i n spec i me n s j u s t e m e r g ed n o t a u l i ces d i s t i n c t a n d i n comp l e t e ; scu t e l l u m sh i n y a n d ret i c u l a t e ru gose ; m e t a n o t u m v e r y reduced a n d s h o r t ; propod e u m f u l l y ret i cu l a t e ru gose , w i t h t w o i n com p l e t e l on g i t u d i n a l k ee l s o n p o s te r i o r s u rf ac e ; a n te r i or s u rf a c e n e a r l y as l o n g as pos t e r i or s u rf ace ( 25 : 22 ) ; fore-w i n g w i th t w o b ro w n t ra n sv e rs a l b a n d s ; d i s t a l p a r t of r a d i a l v e i n m uch l o n g e r t h a n p rox i m a l p a r t ; fore t roch a n ters a p p rox i m a te l y four t i mes a s l o n g a s broad ; segme n t 1 of f ro n t t a rs u s s l i g h t I y shorter t h a n segmen t 4 ( 22 : 23 ) ; e n l a rged c l a w ( F i g . 7 5 1 A ) w i t h s u b a p i c a l t oo t h a n d 1 2 l a m e l l ae ; segm e n t 5 of f ro n t t a rs u s ( F i g . 7 5 1 A ) w i t h t w o rows of 37 l a m e l l ae ; apex w i t h a g ro u p of at l ea s t 20 l a me l l ae ; m ax i l l a r y p a l p i w i t h

1 1 21

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a s l o n g a s b road ( 4 , 66 ) ; head d u l l , f u l l y ret i c u l a t e ru gose ; f ro n s w i t h a med i a n k ee l b e t w een t h e a n t en n a l soc k e t s ; occ i p i t a l c a r i n a i ncom p l e t e , on l y v i s i b l e beh i n d t h e oce l l i ; P O L = 8 ; OL = 3 ; O O L = 6 ; b re a d t h of t h e oce l l i : 4 ; temp l es i n v i s i b l e ; scu t u m f u l l y ret i c u l a t e ru gose , d u l l ; n o t a u l i ce s com p l e t e , poster i or l y s e p a r a t e d ; m i n i m u m d i s t a nce b e t w e n t h e n o t a u l i ce s a s l o n g a s b r ea d t h of t h e oce l l i ; sc u t e l l u m d u l l , ret i c u l a t e ru gose ; me t a n o t u m s h i n y , w i t h a n te r i o r h a l f ret i c u l a t e r u g os e ; poster i o r h a l f smoo t h , w i t h o u t s c u l p t u re ; p ropode u m f u l l y ret i c u l a te ru gose, w i t hou t l o n g i t u d i n a l k ee l s on pos t e r i or s u rf a c e ; fore-w i n g h y a l i n e , w i t hou t d a rk t r a n sv ers a l b a n d s ; d i s t a l p a r t of r ad i a l v e i n n e a r l y as l on g as prox i m a l p a r t ; dors a l p rocess o f gonoforc e p s ( F i g . 7 5 2 ) short a n d s l en d e r ; m a x i I l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l spurs 1 , 0 , 2 . LOCUS T Y P I C US : B u n d a b e r g ( A u s t r a l i a ) TYP I CAL M A T E R I A L : l ec t o t y p e F ! a n d 4 p a r a l ec to t y pes ( 2 F F , 2 MM ) ! ( de s i g n a t e d b y O l m i 1 982 ) i n B . H O S T S : i n A u s t r a l i a Colgar per a c u t u s ( R . C . L . Perk i n s 1 906c ) a n d Massila sp . ( T h o m p so n 1 954 ) . D I ST R I B U T I ON : A u s t ra l i a n reg i on : AUSTRAL I A : B u n d a b erg ( Qu ee n s l a n d ) , B ! OL l Tow n s v i l l e ( C a i r n s , Queens l a n d ) , B ! BM ! N e l son ( C a i rn s , N Q u een s l a n d ) ( Do d d 1 9 1 4c ) ; C a i rn s ( Qu een s l a n d ) , B ! B r i s b a n e ( Queen s l a n d ) , QU ! Como ( New Sou t h W a l es ) , RY ! Bo t a n y B a y ( Ne w Sou t h W a l es ) , WA ! N O T E : N. koebelei ( u n d e r n a me koebeli ) w a s recorded b y m i s t a k e f rom S r i L a n k a b y Ponomarenko ( 1 9 8 1 b ) . T h e spec i me n seen b y Mrs . Ponomarenko is rea l l y a Neodryin u s pseudodiffu s u s n . sp . Neodryi n u s ) .

( see O r i e n t a l spec i es of

4 . Neodryin u s a m boni n .

sp .

F E MA L E : f u l l y w i n ged ; l en g t h 6 , 25 m m ; h e a d b l ac k , w i t h s i des of c l y pe u s a n d m a n d i b l es y e l l o w ; a n t e n n a e b row n , w i t h segmen t s 9- 1 0 red d i sh a n d v e n t r a l f ace o f segmen t 1 w h i te ; t ho r a x a n d p ropod e u m b l ac k , w i t h s i des of p ron o t u m red d i sh ; d i sc of p ro n o t u m w i t h pos t er i or m a r g i n redd i sh ; a b domen b ro w n - b l ac k ; l eg s brow n , w i t h t a r s i a n d fore t roc h a n ters l i g h t ; h e a d d u l l , ex c a v a te d , ru gose, w i t h n u merous i rreg u l a r k ee l s ; occ i p i t a l c a r i n a i n com p l e t e , reach i n g a p p rox i m a t e l y 0 , 5 l e n g t h of OOL ; prono t u m crossed b y a t r a n sv e rs a l i m press i o n , h a i r y , w i t h n u merous l on g i t u d i n a l k ee l s ; poster i o r edges o f p rono t u m not p ro d u c e d i n to l obes ; sc u t u m h a i ry , w i t h n u me ro u s l on g i t u d i n a l k ee l s , n o t ret i c u l a t e r u g os e ; n o t a u l i ces i n v i s i b l e ; sc u t e l l u m h a i ry and a s scu l p t u red a s p rono.t u m ; m e t a n o t u m v e r y red uced a n d short ; p ropodeum re t i c u l a t e r u g os e , w i t h t wo l o n g i t u d i n a l k ee l s o n poster i o r s u rf a c e ; a n t l-' r' i l H ::- U f Li t n : y 1 , :',3 t i mes a s l o n g a s pos t e r i or s u rf ac e ; fore-w i n g w i t h t w o b ro w n t r a n s v er s a l b a n d s ; h i nd-w i n g w i t h a p e x b ro w n ; fore t r'oc h a n l r�T."-, d P � :., 1 ';< ; 1 11 ., 1. i : ,I I 11 �

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1 1 26

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T Y P E S P E C I E S : A dryin u s cerrutii ( Beno i t 1 95 1 ) F E M A L E ( F i g s 754 , 75 5 ) : f u l l y w i n ged ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l ab i a l p a l p i w i t h 3 segmen t s ; p ron o t u m crossed b y a s t ro n g t r a n s vers a l i m p ress i o n ; not a u l i c e s i n v i s i b l e ( F i g . 756 ) ; en l a r g e d c l a w ( F i g . 757 ) w i t h s u b a p i c a l t oo t h a n d b r i s t l es o r p e g- l i k e h a i rs ; ·t i b i a l s p u rs 1 , 0 , 1 ( F i g . 758 A ) . MALE : u n k no w n D I ST R I B U T I O N : E t h i op i a n , HOST S : u n k no w n S P E C I ES : 7

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E T H I O P I AN S P E C I E S OF ADR YINUS

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P r on o t u m w i t h d i s c v e ry p r o m i n e n t a n d p o i n t ed ( F i g s 7 5 5 , 7 5 6 A ) ; me s op l eu ra v e ry p r om i n en t ( F i g . 7 5 4 ) . . . . • • . . • • . • • . . • • . . • • • cerrutii ( B en o i t ) . . . . . . . . . . . . . . . . . . . . . . . 1 . P r on o t u m w i t h d i s c p r o m i n en t , b u t n o t p o i n t ed ( F i g . 7 5 6 B ) ; me s op l eu ra l e s s p r om i n en t • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • 2 F o r e -w i n g w i t h d i s t a l p a r t o f r a d i a l v e i n a s l on g a s p r ox i ma l pa r t • • • • • • • • • • • • . • • • • • • • • • • • . . • • • • • • • • • • • • • • • • • • • • • • • • • . • • • . • 3 F o r e -w i n g w i t h d i s t a l p a r t of r a d i a l v e i n a t l ea s t 1 , 4 t i me s a s l on g a s p r ox i ma l p a r t . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 P r on o t u m f u l l y b l a c k ; l eg s a n d a n t en n a e b l a c k . . . . . . . . . . . . . . . . . . . . . . . . . • . . . • . . • • . . . . • . 4 . oweni n . s p . P r on o t u m b l a c k , w i t h s i d e s r ed d i s h ; l eg s a n d a n t en n a e f u l l y r ed d i sh - t e s t a c e o u s • • • • • 5 . rugidorsa lis ( T u r n e r ) O c c i p u t g ra n u l a t ed , w i t h ou t p a ra l l e l k e e l s ; o c c i p i t a l c a r i n a d i s t i n c t a n d i n c omp l e t e • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • . . . . . . . . . . . . . . . . . . . . . . . 2 . nigerian u s n . sp .

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1 . A dryi n u s cerrutii ( Be n o i t )

Neodryinus cerrutii Beno i t 1 95 1 d :

n . com b .

17.

F E MA L E ( F i g s 7 5 4 , 7 55 ) : f u l l y w i n ged ; l en g t h 6 , 0 mm ; h e a d b l ac k , w i t h m a n d i b l es p a r t l y redd i s h ; a n te n n a e b l ac k , w i t h ven t r a l f a c e of s e g me n t y e l l o w ; t ho r a x , p ropodeu m , a b domen a n d l eg s b l ac k , w i t h t a r s i l i g h t ; h e a d exc a v a t e d , d u l l , w i t h t e m p l es p rom i n e n t , h a i r y , g r a n u l a te d , w i t h n u merous p a ra l l e l kee l s on f ro n s , v er t ex a n d occ i p u t ; occ i p i t a l c a r i n a i n v i s i b l e ; p roo n o t u m ( F i g . 756 A ) crossed b y a t r a n s v e rs a l i mp ress i o n , g r a n u l a t e d , w i t h n u mero u s k ee l s a rou n d d i sc a n d co l l a r ; d i sc ( F i g . 756 A) very p rom i n e n t a n d po i n t e d ; p os t e r i o r edges of p ro n o t u m rou n d e d , n o t p ro d u ced i n to l obes ; meso p l eu r a v e r y p rom i n en t ; sc u t u m sh i n y , f u l l y

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B F i g . 758 - H i n d t i b i a l s p u rs of f e m a l es of A dryinus cerr u t i i ( Be n o i t ) ( t y p e ) ( A ) a n d Neodryin u s domin icanus ( E v a n s ) ( t y p e ) ( B ) .

ret i c u l a t e ru gose ; n o t a u l i ces i n v i s i b l e ; s c u t e l l u m f u l l y re t i c u l a t e ru gose , w i t h o u t s c u l p t ure a m o n g a r eo l a e ; m e t a n o t u m very reduced a n d s h o r t ; p ro p o d e u m f u l l y ret i c u l a t e ru gose , w i t h o u t l on g i t u d i n a l k ee l s on pos ter i or s u r f ace ; fore-w i n g w i t h t w o b ro w n t r a n s v e rs a l b a n d s ; d i s t a l p a r t of r a d i a l v e i n s l i g h t l y l on g e r t h a n p rox i m a l p a r t ( 1 0 : 9 ) ; fore t r oc h a n ters a p p ro x i m a t e l y f o u r t i mes a s long a s b ro a d ; segment 1 of f ron t t a r s u s l on g e r t h a n segmen t 4 ( 30 : 1 5 ) ; en l a rged c l a w ( F i g . 757 ) w i t h s u b a p i c a l tooth a n d 3 b r i s t l es ; segmen t 5 o f f ro n t t a rs u s ( F i g . 757 ) w i t h t w o row s of 1 0 + 3 l a me l l ae ; a pex w i t h a g r o u p o f 1 5 l a me l l ae ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 ( F i g . 758 A ) . MA L E : u n k n o w n lOC U S TYP I C US : N a i rob i ( Ken y a ) T Y P I CAL M A T E R I AL : h o l o t y p e F ! i n TV . HOSTS : u n k n o w n D I ST R I B U T I O N : E t h i op i a n reg i o n : K E NYA : N a i rob i , T V ! Z I MBABW E : V i c t o r i a Fa l l s , BM !

2 . A dryin u s n igeri a n u s n . s p .

F E MAL E : f u l l y w i n ge d ; l en g t h 4 , 1 2 m m ; h e a d b l ac k , w i t h m a n d i b l es red d i sh ; a n t e n n a e testaceo u s ; t h o r a x a n d p ropodeum b l ac k , w i t h s i des of p ronot u m redd i s h ; a b domen b l a c k - b ro w n ; l eg s b l a c k - b row n , w i t h t a r s i l i g h t ; h e a d w ea k l y ex c a v a ted , d u l l , g r a n u l a ted , f u l l y ret i c u l a t e ru gose ; occ i p i t a l c a r i n a i nc o m p l e t e , reach i n g a p p ro x i m a t e l y 0 , 5 l en g t h of OOl ; occ i p u t g r a n u l a t ed , w i t h o u t a reo l a e or k ee l s ; p ron o t u m c rossed b y a t r a n s v e rs a l i m p re ss i o n , d u l l , g r a n u l a ted , w i t h some s t r i a e on c o l l a r

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a ro u n d t h e d i sc ; p o s te r i o r e d g e s of p ro n o t u m rou n d e d , not p ro d u c e d i n to l obes ; sc u t u m f u l l y r e t i c u l a t e ru gose ; n o t a u l i ces i n v i s i b l e ; scu t e l l u m fu l l y r e t i cu l a t e r u g os e ; m e t a n o t u m very reduced a n d s h or t ; p ro p o d e u m ret i c u l a t e ru gose, w i t h t w o l o n g i t u d i n a l k ee l s on p os t e r i o r surface ; fore- w i n g w i t h t wo d a r k t ra n s v e rs a l b a n d s ; d i s t a l p a r t o f ra d i a l v e i n a p p rox i m a t e l y 1 , 66 a s l on g a s p rox i ma l p a r t ; fore t roc h a n ters a p p rox i m a t e l y f o u r t i mes a s l o n g a s b ro a d ; segmen t 1 of f ron t t a r s u s l on g e r t h a n s e g me n t 4 ( 22 : 1 4 ) ; e n l arged c l a w ( F i g . 759 ) w i t h s u b a p i c a l t oo t h a n d 1 2 p e g - l i k e h a i rs ; s e g m e n t 5 of fron t t a r s u s ( F i g . 759 ) w i t h t w o row s of 5 + 4 l a me l l a e ; a pex w i t h a g rou p of a t l ea s t 6 l a me l l a e ; max i l l a ry p a l p i w i t h 6 seg m'en t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k no w n L O C U S TYP I C U S :

Z u g u rma ( N i ge r i a )

TYP I CA L MATER I AL : h o l o t y p e F ! i n BM . HOST S : u n k n o w n D I STR I B U T I ON : E t h i op i an reg i o n : N I G E R I A : Z u g u r m a ( Mok w a , BM !

3 . A dryin u s b ellieosus

( Be n o i t )

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F E M A L E : f u l l y w i n ged ; l en g t h 5 , 00- 5 , 62 m m ; h ead b l ack , w i t h m a n d i b l es red d i sh ; an t e n n a e t es t aceou s , w i t h seg m e n t s 7- 1 0 b l ack ; occa s i on a l l y an t e n n a e f u l l y b l ac k or b ro w n , w i t h ven t r a l face of segmen t 1 w h i t i sh ; t horax a n d p ro p o d e u m b l ac k ; p rono t u m somet i me s w i t h s i de s a n d poster i or m a r g i n red d i sh ; a b domen b r o w n or b row n-red d i sh ; l eg s b l ac k , w i t h t a r s i l i g h t ; h e a d exc a v a t e d , s h i n y , g r a n u l a ted , w i t h n u merous p a ra l l e l k ee l s o n f ron s , ver tex a n d occ i p u t ; occ i p i t a l car i n a i n v i s i b l e ; p ron o t u m crossed b y a t r a n s v e rs a l i m p ress i o n , d u l l , g r a n u l a t e d , w i t h n u merous s t r i ae on c o i l ar a n d a r o u n d t h e d i s c ; pos ter i or ed ges o f p ron o t u m r o u n d ed , n o t p ro d u c e d i n t o l obes ; sc u t u m d u l l , g r a n u l a t e d , re t i cu l a t e ru gose a n d w i t h l on g i t u d i n a l k ee l s ; n o t a u l i ce s i n v i s i b l e ; sc u t e l l u m as scu l p t u red as sc u t u m ; m e t a n o t u m v ery red uced a n d shor t ; p ropodeum ret i c u I a t e ru gose , w i t h t w o l o n g i t u d i n a l k ee l s on pos t e r i o r s u rfac e ; fore- w i n g w i t h t wo d a r k t r a n s v e r s a l b a n d s ; d i s t a l p a r t of rad i a l v e i n a p p rox i m a te l y 1 , 4 - 1 , 5 t i mes a s l o n g a s p rox i m a l p a r t ; fore t ro c h a n ters a p p rox i m a t e l y fou r t i mes as l o n g a s b road ; segmen t 1 of fron t t a rs u s l on g e r t h a n segmen t 4 ( 24 : 1 4 ) ; e n l a rged c l a w ( F i g . 760 ) w i t h S U b a p i c a l too t h a n d 1 1 p e g - l i k e h a i rs ; s e g me n t 5 of f ro n t t a r s u s ( F i g . 760 ) w i t h t w o rows of 1 0 + 6 l a me l l a e ; a p ex w i t h a g r o u p of a t l ea s t 1 5 l a me l l ae ; ma x i l l ary p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k n ow n . Ben o i t ( 1 95 1 c ) descr i be d t h e a l l o t y p e M ; t h i s s p ec i men

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re a l l y b e l on g s to a n other gen u s , b e c a u s e i t h a s p a l p a l ra t i o 4/2 . L O C U S T Y P I CUS : Loa s h i V a l l ey ( Z a i re ) T Y P I CAL MAT E R I AL :

holotype F !

i n TV .

HOST S : u n k n o w n D I ST R I B UT I ON : E t h i o p i a n reg i on : Z A I R E : Loash i V a l l ey ( K i w u ) , T V ! K a z i b a ( U p e m b a N a t . P a r k ) , O L ! TV ! UGA N D A : R u z u m b u s a , BM ! N a m u t a m b a , OL ! BM ! Z I M BA BWE : C h i s h a w a s h a ( Sa l i sb u ry ) , BM !

4 . A dryin u s oweni n . sp .

F E M AL E : f u l l y w i n ged ;

l e n g t h 6 mm ;

b l ac k ,

w i t h m a n d i b l es d i s t a l l y

t e s t a ceou s ; m i d a n d h i n d t a r s i l i g h t ; a n te n n a e d i s t a l l y t h i ck e n e d ; a n t en n a l s e g m e n t s i n fo l l ow i n g p ropor t i o n s : 2 2 : 7 : 29 : 1 7 : 1 3 : 1 0 : 7 : 5 : 5 : 8 ; h e a d exc a v a t e d , s h i n y , g r a n u l a ted , w i t h n u m erous areo l ae a n d l o n g i t u d i n a l kee l s ; occ i p u t s c u I p t u red b y l o n g i t u d i n a I k e e l S ; occ i p i t a I c a r i n a i n v i s i b l e ; frons w i t h a med i a n k ee l ; P O L = 7 ; OL = 3 ; OOL = 1 6 ; temp l es v ery p ro m i n en t ; p ro n o t u m crossed b y a t r a n s v e rsa I i m p re s s i on , d u l l , g r a n u l a t e d , w i t h i r reg u l a r k ee l s ; co l l a r w i t h a n t e r i or m a r g i n s t ron g l y e x c a v a t e d ; d i sc h u m p ed , b u t not p o i n t ed ; pronot u m w i t h o u t poster i or l obes d i rected tow a r d s t h e t e g u l ae ; sc u t u m s h i n y , w i t h n u mero u s l on g i t u d i n a l a n d t ra n sv e rs a l k ee l s ; n o t a u l i ces i n v i s i b l e ; s c u te l l u m ret i c u l a t e ru gose ; m e t a n o t u m very sho r t ,

ret i c u l a t e r u g os e ;

p ropodeum

ret i c u l a t e ru gose, w i t h o u t l o n g i t u d i n a l or t r a n svers a l k ee l s ; fore-w i n g w i t h two d ar k t r a n svers a l b a n d s ; d i s t a l p a r t of ra d i a l v e i r. as l o n g a s p rox i m a l p a r t ; f o r e t a rsa l segme n t s i n fo l l ow i n g p roport i o n s : 29 : 4 : 8 : 1 3 : 26 ; en l a rg ed c l a w ( F i g . 761 ) w i t h s u b a p i c a l too t h a n d 5 b r i s t l es ; seg m e n t 5 of front t a rs u s ( F i g . 761 ) w i t h t w o row s o f 3 2 l a m e l l a e e x t e n d i n g con t i n u o u s l y to t h e apex ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n w o n LOCUS TYP I CU S : Kamp a l a ( U gan d a ) T Y P I CAL M A T E R I A L : ho l ot y pe F ! i n T W . HOST S : u n k n o w n D I ST R I B UT I ON : E t h i o p i a n re g i on : UGANDA : K a m p a l a , T W ! N OT E : t h e spec i es i s n a med i n honor of M r . Den i s F . Owen .

5 . A dryin u s rugidors a l i s ( T u r n e r ) n . com b .

Psilodryi n u s rugidors a lis T u rner 1 9 28 : 1 50 . Neodryin u s rugidors a l i s ( T u rn er ) : R i c h a rd s 1 953 : 69 .

F E M A L E : f u l l y w i n ged ;

l en g t h 4 , 68 m m ;

h e a d b l ac k ,

w i t h c l y pe u s a n d

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1 1 36

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1 984

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m a n d i b l es redd i s h ; a n t e n n a e red d i s h - t e s t aceou s ; p rono t u m b l ac k , w i t h s i des a n d pos t er i or m a r g i n redd i sh ; meso t h o ra x , m e t a t ho r a x a n d p ropode u m b l ac k ; a b domen t e s t aceo u s , w i t h t e r g i t e 1 p a r t I y b l ac k ; l eg s red d i s h - t es t aceo u s ; h e a d w ea k l y exc a v a ted , s h i n y , ret i c u l a te r u gose o n f ro n s a n d vertex ; occ i p u t w i t h n u merous k ee l s ; occ i p i t a l c a r i n a i n v i s i b l e ; p ro n o t u m ( F i g . 756 B ) crossed b y a t r a n sversa l i m p ress i o n , d u l l , g r a n u l a ted , s t r i a te on co l l a r a n d a rou n d t h e d i sc ; p o s te r i or e d g e s of p ro n o t u m rou n d e d , not p rod uced i n to l obes ; sc u t u m s h i n y , ret i c u l a t e

1 1 37

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ru gose ; n ot au l i ces a b sen t ; sc u te l l u m re t i c u l a t e r u gose ; pron o t u m , scu t u m a n d s c u t e l l u m g ra n u l a t e d a m o n g areo l ae ; m e t a n o t u m v e r y reduced a n d shor t ; p ropod e u m ret i c u l a t e r u gose , w i t ho u t l on g i t u d i n a l k ee l s on pos t e r i o r s u rf a c e ; fore-w i n g w i t h two dark t r a n sversa l b a n d s ; d i s t a l p a r t o f ra d i a l v e i n a p prox i ma te l y a s l o n g as p rox i m a l p a r t ; f o r e t roch a n t e r s a p p ro x i m a t e l y f o u r t i me s a s l o n g a s b road ; segmen t 1 of f ro n t t a r s u s l o n g e r t h a n s e g me n t 4 ( 2 1 : 8 ) ; e n l a r g e d c l a w ( F i g . 762 ) w i t h s u b a p i c a l t oo t h a n d 5 b r i s t l es ; seg m e n t 5 o f f r o n t t a rs u s ( F i g . 762 ) w i t h t w o rows of 7 + 7 l a me l l a e ; a pex w i t h a g ro u p of a t l ea s t 1 5 l a m e l l ae ; m ax i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M AL E : u n k n o w n L O C U S T Y P I C U S : P o r t S t . J o h n s ( Sou t h A f r i c a ) HOSTS : u n k no w n D I ST R I B U T I ON : E t h iop i a n reg i on : S O U T H A F R I CA : P o r t S! . J o h n s

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T w o spec i es are k n ow n .

KEY T O T H E S P E C I E S

FEMALES

Scu t u m pu n c t a t e , wi t h ou t I ong i t ud i n a l k e e l s • . • • • • • . . . • . . . . . . . . . . . . . . . . . . . . . . . . . . 1 . gracilis ( F e n t on ) S c u t u m w i t h n u m e r ou s l on g i t u d i n a l k e e l s , n o t p u n c t a t e , g r a n u I a t ed among k e e I s • • • • 2 . meridiana lis n . sp .

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a n d m a n d i b l es y e l l o w ; a n te n n a e y e l l ow , w i t h segmen t s 6-9 b row n ; a p e x of segmen t 5 d a r k ; p ro no t u m b l ac k , w i t h po s t e r i o r m a r g i n and s i des redd i s h ; mesothora x , meta thorax and p ropodeum b l ack ; a b domen b row n ; l eg s t e s t aceou s ; h e a d s h i n y ,

a l mo s t f l a t ,

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occ i p i t a l c a r i n a v i s i b l e o n l y beh i n d t h e oce l l i ; pro n o t u m crossed b y a t r a n sv e rs a l i m p ress i on , sh i n y , w i t hou t scu l p t u re or w ea k l y g r a n u l a t e d , w i t h n u merous s t r i a e ; p o s t e r i or edges of p ro n o t u m rou n d e d , n o t p roduced i n to l ob e s ; sc u t u m sh i n y , w i t h o u t scu l p t u re , excep t for n u me ro u s p u n c t u res a n d some a reo l ae m a i n l y a l o n g t h e i n v i s i b l e n o t a u l i ces ; scu t e l l u m sh i n y , w i t h a n t e r i o r h a l f g r a n u l a t e d ; poster i o r h a l f w i t hou t scu l p t u re ; m e t a n o t u m very reduced a n d shor t ; p ropodeum ret i c u l a t e r u go s e , w i t h t w o l o n g i t u d i n a l kee l s o n pos t e r i o r s u rf ac e ; fore-w i n g w i t h t w o b r o w n t r a n s v e r s a l b a n d s ; d i s t a l p a rt of r a d i a l v e i n a p p rox i m a t e l y a s l o n g a s prox i m a l p a r t ; fore t roc h a n ters a p p rox i m a te l y fo u r t i mes a s l on g a s b ro a d ; c h e l ae m i s s i n g i n t h e o n l y k n o w n spec i m en ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M AL E : u n k no w n L O C U S TYP I C U S : Berm u d a TYP I CAL M A T E R I AL : ho l ot y p e F ! i n WA . H OST S : u n k n o w n D I S T R I B U T I O N : N e a rc t i c reg i o n : B E R M U D A , WA !

2 . A dryin u s meridion a lis n . sp .

F E MA L E : f u l l y w i n ged ; l e n g t h 4 , 0 m m ; h e a d b l ac k , w i t h c l y pe u s a n d m a n d i b I es t e s t aceou s ; a n te n n a e t e s t aceou s , w i t h segmen t s 7- 1 0 b row n ; t hora x , p ropodeu m a n d a b domen b l ack ; l eg s b l ac k , w i t h t a r s i t e s t aceo u s ; h e a d sh i n y , g r a n u l a te d , ret i cu l a t e r u gose , a l mo s t f l a t ; occ i p i t a l c a r i n a o n l y v i s i b l e beh i n d t h e oce l l i a n d o n t h e s i des o f t h e poster i o r oce l l i ; p rono t u m crossed b y a t r a n sv ersa I i mp ress i on , w i t h o u t scu I p t u re or w e a k l y g r a n u l a t e d , sh i n y , w i t h some l o n g i t u d i n a l k ee l s o n t he s i des of t h e c o l l a r ; poster i or edges of p ro no t u m ro u n ded , not p ro d u ced i n t o l obes ; scu t u m h a i ry , s h i n y , w i t h n u me ro u s l on g i t u d i n a l k ee l s , w e a k l y g r a n u l a ted ; n o t a u l i ces i n v i s i b l e ; s c u t e l l u m d u l l , w i t h a n ter i o r h a l f ret i c u l a t e ru gose ; post e r i o r h a l f smoot h ; m e t a n o t u m very reduced a n d s h o r t ; p ropode u m ret i c u l a t e r u gose, w i t h t w o l on g i t u d i n a l k ee l s o n post e r i o r s u rf ace ; fore-w i n g w i t h t w o d a rk t ra n sversa l b a n d s ; d i s t a l p a r t of r a d i a l v e i n n e a r l y t w i ce a s l o n g a s p rox i m a l p a r t ; fore t roc h a n ters a p p ro x i m a t e l y f i v e t i mes a s long a s b roa d ; en l a r g e d c l a w ( F i g . 763 ) w i t h s u b a p i c a l t oo t h a n d 7 b r i s t l es ; seg m e n t 5 of f ro n t t a r s u s ( F i g . 763 ) w i t h 1 row of 26 l a me l l ae ; a pex w i t h a g ro u p of n e a r l y 1 5 l a m e l l ae ; m ax i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; M AL E : u n k n o w n LOCUS T Y P I C U S : S a n Lore n zo I s l a n d ( Mex i co ) TYP I CA L M A T E R I AL :

h o l o t y pe F !

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HOSTS : u n k n o w n D I ST R I BU T I ON : N e a rc t i c reg i on : M E X I C O : S a n Lorenzo I s l an d ( G u l f of C a l i fo rn i a ) , C A l

3 . Ech throdelph a x R . C . L . Perk i n s

Echthrodelp h a x R . C . L . Perk i n s 1 90 3 : 36 . Echthrodelp h a x R . C . L . Perk i n s : K i effer i n K i effer et M a r sh a l l .1 906 : 489, 493 . Mercetia K i effer i n K i effer et M a r s h a l l 1 906 : 495 ( n . sy n . ) ; t y p e spec i es Mercetia longicollis ( K i effer 1 905 ) , or i g . d es i g . ( a t f i rs t s i g h t Mercetia seems v ery d i ffere n t f r o m Ech throdelp h a x ; rea l l y t h e d i fferences are o n I y s p ec i f i c ) . Echthrodelp h a x R . C . L . Perk i n s : K i effer 1 9 1 4b : 1 6 . Mercetia K i ef fe r : K i effer 1 9 1 4b : 1 9 . Echthrodelp h a x R . C . L . Perk i n s : R i c h a r d s 1 939 : 1 88 . Pterogona top u s Ab d u l -Nour 1 976 : 275 ( n . sy n . ) ; t y pe s p ec i es Pterogon a top u s hortusensis A b d u l -Nour 1 97 6 , monot y p i c .

TYPE S P E C I E S : Echthrodelp h a x fa irchildii R . C . L . Perk i n s 1 90 3 , monot y p i c . F E M A L E ( F i g s 767, 768 ) : f u l l y w i n ged ; m a x i I l a ry p a l p i w i t h 3-6 segmen t s ; l a b i a l p a l p i w i t h 2-3 segmen t s ; p ron o t u m ( F i g . 768 ) crossed b y a s t ro n g t r a n s v e r s a l i m p ress i on ; n o t a u l i c e s ( F i g . 767 ) comp l e t e a n d d i s t i nc t ; e n l a rged c l a w ( F i g . 769 ) w i t h s u b a p i c a l t o o t h a n d l a me l l ae ; t i b i a l s p u rs 1 , 0, 1 . M A L E ( F i g s 7 7 9 , 7 80 ) : f u l l y w i n ged ; m ax i l l a ry p a l p i w i t h 4 segme n t s ; l a b i a l p a l p i w i t h 2 seg m e n t s ( t h e i n form a t i on s a b o u t t h e n u mber of p a l p a l seg men t s i n m a l es a re c e r t a i n l y i n com p l e t e ; p ro b a b l y t he p a l p a l ra t i o i s not d i ff e r e n t i n b o t h sexes ) ; occ i p u t concave ; temp l e s p rese n t ; h e a d w i t h a d i s t i n c t occ i p i t a l car i n a f ro m p o s te r i or oce l l i t o ven t r a l s i de of t h e h ea d ; t i b i a l s p u rs 1 , 1 , 2 . D I ST R I BU T I ON : Pa l ae a rc t i c , E t h i op i a n , O r i en t a l , Nearc t i c , . Au s t ra l i a n HOSTS : Delph a cidae SPEC I ES : 1 4

PALAEARC T I C S P E C I E S O F ECHTHRODELPHAX

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F o r e -w i n g h y a l i n e , w i t h ou t d a rk t r a n sv e r s a l b a n d s . . . . . . . . . . . . . . . . . . . . . . . 1 . fairchildii R . C . L . P e rk i n s ( s e e A u s t r a l i a n s p e c i e s ) 2

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F o r e -w i n g w i t h a t l ea s t on e d r a k t r a n sv e r s a l b a n d . . . . . . . . . . . . 2 S c u t u m d u l l , f u l l y r e t i c u l a t e ru g o s e ; p r op od e u m w i t h t w o l o n g i t u d i n a l , med i a n a n d p a ra l l e l k e e l s on d o r s a l a n d p o s t e r i or su r f a c e • . • • • . • . . . • • • . . • • • • • • . . . . . . . . . . 2 . b a eticus ( C eb a l l o s ) S c u t u m sh i n y , smo o t h , w e a k l y p u n c t a t e , w i t h o u t s c u l p t u r e amon g p u n c t u r e s , n o t r e t i c u l a t e ru g o s e ; p r op od eu m w i t h o u t l on g i t u d i n a l , p a r a l l e l a n d med i a n k e e l s on d o r sa l a n d p o s t e r i o r su r f a c e , a t mo s t w : t h t w o i n c omp l e t e k e e l s on p o s t e r i or s u r f a c e • . • • • . . • • • • • • • • • • • • . • • • • • . . . • . • • • • • • . • . . • • . . 3 P o s t e r i o r su r f a c e o f p r op o d e u m w i t h t wo i n c omp l e t e I ong i t ud i n a l k e e l s ; o c e l l a r t r i an g l e equ i l a t e ra l o r s l i g h t I y i s o s c e l e s ( F i g . 7 6 4 D ) ; f o r e -w i n g w i t h t wo d a rk t r a n sv e r s a l b a n d s . . . . . 3 . longicollis ( K i e f f e r ) P o s t e r i o r su r f a c e o f p r op o d eu m w i t h o u t l o n g i t u d i n a l k e e l s ; o c e l l a r t r i a n g l e s t r on g l y i s o s c e l e s ( F i g . 7 6 4 A ) ; f o r e - w i n g w i t h on e d a rk t ra n sv e r s a I b a n d ; s ome t i m e s w i t h a t s e c on d b a n d • • • • • • • • . • • • • • • • • • • • . . • • • • • . • . . . • • • . • • • S c u t u m t e s t a c e o u s o r t e s t a c e o u s - r ed d i sh ; h ea d w i t h t h a n t h r e e t i me s a s l o n g a s POL ; n o t a u l i c e s j o i n t . . . . . . . . . . . 4 . h ortusensis ( Ab d u I - N o u r ) S c u t u m b l a c k ; h e a d w i t h O L a p p r ox i ma t e l y o n e a n d a t i me s a s l on g a s P O L ; n o t a u l i c e s s ep a r a t ed . . . . . . . . . . . . . . . . . . 5 . italicus n . sp .

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2 . Echthrodelp h a x b a eticus

( C e b a I l os ) n . com b .

Mercetia b a etica Ceb a l l os 1 927 : 1 00 . Mercetia ten u icornis K i effer 1 924 : 4 2 ( n . sy n . ) ; locus typicus A l geri a . Dryin u s victorovi Ponomarenko 1 970 : 4 3 1 ( n . s y n . ) ; locus typicus K l y u c h i ( U . S. S. R . )

F E M A L E : f u l l y w i n ged ; l en g t h 5 , 75-6 , 37 m m ; h e a d redd i s h , w i t h a w i d e b l ack s p o t on ver tex a n d f ron s ; occ a s i on a l l y h e a d b l ac k , w i t h occ i p u t a n d t e m p l e s red d i sh ; somet i mes a l so frons a l on g orb i t s y e l l ow ; a n t e n n a e f u l l y t e s t aceo u s ; c l y peu s , m a n d i b l es a n d m a l a r s p ac e a l w a y s t e s t aceo u s ; p rono t u m red d i sh , w i t h b l ack s p o t s on t ra n s v e r s a l i mp re s s i on a n d on s i d e s ; occas i on a l l y p rono t u m b l ac k , w i t h m a r g i n s redd i s h ; sc u t u m , s c u te l l u m , met a n o t u m , p l e u r a , p ropodeum a n d p e t i o l e b l ac k ; a b domen b l ack , w i t h red d i s h n u a n ces ; l eg s t es t aceou s , w i t h c l u b s of femora a n d cox ae p a r t l y b l ac k ; h e a d ( F i g . 764 C ) d u l l , g r a n u l a te d , ret i c u l a t e ru gose , w i t h i rreg u l a r k ee l s o n frons a n d v e r t e x , w e a k l y h a i ry ; f ro n s w i t h a med i a n l on g i t u d i n a l k e e l ; oce l l a r t r i a n g l e e q u i l a t e ra l ; p rono t u m crossed b y a t r a n s v e rs a l i m p re ss i o n , s h i n y , h a i r y , w i t h i r reg u l a r k ee l s ; pos t er i or edges o f p ron o t u m n o t p ro d u ce d i n to l obes d i rected t o w a r d s t h e t e g u l ae ; scu t u m d u l l , ret i c u l a t e ru gose, h a i ry ; not a u l i c e s d i s t i n c t a n d com p l e t e , poster i or l y sep a r a ted ; s c u te l l u m ret i c u l a t e r u g os e ; m e t a n o t u m

1 1 43

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Fema l e o f Ech throdelp h a x baeticus ( Ce b a l l o s ) from G a v a r n i e ( F r a n ce ) ; A : c h e l a ; B : l a b i a l p a l p u s .

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1 984

redu ced t o a s h o r t t r a n s v e rs a l f u r ro w ; p ro p o d e u m f u l l y ret i c u l a t e r u go s e , w i t h t w o com p l et e l on g i t u d i n a l kee l s on dors a l a n d p o s ter i or s u rface ; fore-w i ng w i th t w o d a r k t ra n s versa I b a n d s ; fore t roc h a n t e r s a p p rox i ma te l y f i ve t i mes a s l on g as b road ; segme n t 1 o f fron t t a r s u s s l i g h t l y s h o r t e r t h a n segme n t 4 ( 24 : 30 o r 24 : 26 ) ; en l arged c l a w ( F i g . 765 A ) w i t h s u b a p i c a l t oo t h and 6-8 l ame l l ae ; segment 5 of f ro n t t a r s u s ( F i g . 765 A ) w i t h t w o row s of 4 2 l ame l l ae ex t en d i n g con t i n u ou s l y t o t h e apex ; max i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i ( F i g . 765 B ) w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . M A L E : u n k no w n LOCUS TVI- I C U S : A l g ec i ras ( Sp a i n ) TYP I CA L M A T E R I AL : h o l o t y p e F ! o f N . b a etica i n M D ; h o l o t y p e F !

and victorovi i n MO ; h o l o t y pe F of N. tenu icornis l os t ; n e o t y p e F ! ( h ereby des i g n a t e d ) o f N. tenu icornis i n P . H O ST S : u n k no w n D I STR I B U T I ON : P a l aearc t i c reg i on : U . S . S . R . : K l y u ch i ( Sa r a t o v P rov i n ce ) , MO ! B u k h ara ( Uzbek i s t a n ) , L N ! K y n a k ( u n der n a me Nercetia turanica n . s p . G u ss a k o v s k j i : u n desc r i bed s p ec i es ) , L N ! A l m a ( C r i mea ) ( Ponomarenko 1 970 ) ; F RAN C E : G a v a r n i e ( H a u tes-p y re ne es ) , L O ! SPA I N : A l g ec i ra s , M D ! ALG E R I A : La S t i d i a ( O ran ) , P ! N O T E S : t h e spec i me n s from A l ger i a seem s l i g h t l y d i ffere n t f rom t h e E u ropean a n d As i a t i c p o p u l a t i on s . T h e head a n d p ro n o t u m are a l most f u l l y smoo t h and w i t hou t scu l p t u re ( i n E u ropean a n d A s i a t i c spec i mens head a n d p rono t u m a re f u l l y ru gose ) , t h e n o t a u l i ces are w eak l y v i s i b l e ( s t ron g l y v i s i b l e i n E u ropean a n d A s i a t i c fema l es ) , t h e sc u t u m i s mos t l y scu l p t u red b y i rreg u l a r k ee l s ( mo s t l y ret i c u l a t e r u gose i n E u ropean and As i a t i c spec i men s ) , t he t w o p a ra l l e l l on g i t u d i n a l k ee l s of t h e p ropode u m a re s l i g h t l y d i s t i nc t ( s t ron g l y v i s i b l e i n E u ropean a n d A s i a t i c p o p u l a t i on s ) . Perh a p s t h e North Afr i c an p o p u l a t i on i s a s u b spec i es o r a s e p a r a t e d spec i es ( ten u icornis K i effer ) . I h a v e seen on l y a fem a l e from A l g er i a ; o t h e r s p ec i mens need . p aratype F of D .

3 . Echthrodelp h a x longicollis ( K i effer ) n . com b .

Dryin u s Mercetia 495 . Mercetia Mercetia

longicollis K i effer 1 905a : 75 . longicollis ( K i effer ) : K i effer i n K i effer et M a r s h a I I 1 906 : longicollis ( K i effer ) : longicollis ( K i effer ) :

K i effer 1 9 1 4b : 1 9 . Ceb a l l os 1 927 : 9 9 .

F E M AL E : f u l l y w i n g e d ; l e n g t h 4 , 37-4 , 56 m m ; head tes taceous-red d i sh ; occas i on a l l y p a r t l y red d i s h , w i t h a w i d e b l ac k spot on v e r te x a n d f ro n s ; m a n d i b l es , c l y pe u s a n d a n t e n n a e test aceou s ; p ronot u m f u l l y redd i s h ; occ a s i on a l l y red d i s h , w i t h b l ack s p o t s o n t ra n sversa l i mp ress i on a n d o n

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1 1 46

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s i de s ; sc u t u m a n d sc u t e l l u m red d i s h ; m e t a n o t u m , mesop l eu r a , me t a p l eu r a a n d p ropodeu m b l ack ; occ a s i on a l l y met a n o t u m red d i s h ; a b domen b l ack , w i t h red d i sh n u a n ces ; l e g s f u l l y or p a r t l y t e s t aceo u s ; occ a s i o n a l l y w i t h c o x a e , c l u b s of femora a n d t i b i ae p a r t l y b l ac k or b row n ; a n t en n a l segmen t s i n f o l l ow i n g p ropo r t i o n s : 9 : 5 : 20 : 1 6 : 1 3 : 1 0 : 7 : 5 : 5 : 7 ; h e a d ( F i g . 7 6 4 D ) d u l l o r sh i n y , g r a n u l a te d , w i t h w e a k i r regu l a r k ee l s ; occ i p i t a l c a r i n a i n comp l e t e , s l i g h t l y v i s i b l e beh i n d t h e oce l l i ; O L = 4 ; OOL = 3 ; p ro n o t u m crossed b y a t ra n s vers a l i m p ress i o n , sh i n y ; d i sc w i t h o u t scu l p t u re ; s i des p u n c t a t e a n d w i t h w e a k k ee l s ; p rono t u m occ a s i on a l l y f u l l y w i t h ou t scu l p t u re ; n o pos t e r i o r l ob e s d i rected tow a rd s

1 1 47

Gon a topod i n ae

F i g . 768 - F em a l e of Ech throdelp h a x h ortusensis ( Ab d u l -Nou r ) from Sc i o l ze ( I taly)

t h e t e g u l ae ; scu t u m s h i n y , smoo t h , w e a k l y p u n c t a t e , w i t h o u t scu l p t u re a m o n g p u n c t u res ; n o t a u l i ces com p l e t e , pos t er i or l y sep a ra t e d ; scu t e l l u m du I I , s t ro n g I y p u n c t a t e ; m e t a no t u m red u ce d to a s h o r t t r a n sv ersa I f u rrow ; propo d e u m d u l l , f u l l y ret i c u l a t e r u gose, w i t h i ncomp l et e l o n g i t u d i n a l k ee l s o n p o s te r i or s u rf a c e ; fore-w i n g w i t h t w o d a r k t ra n s v ers a l b a n d s ; p t erost i g m a very n a rrow ; d i s t a l p a r t o f r a d i a l v e i n l on ger t h a n prox i ma l p a r t ; fore t a rs a l seg men t s i n fo l l o w i n g p roport i on s : 2 1 : 2 : 5 : 20 : 3 0 ; en l a r ged c l a w ( F i g . 766 ) w i t h s u b a p i c a l t oo t h a n d 8-9 l am e l l ae ; seg m e n t 5 of f ro n t t a r s u s ( F i g . 766 ) w i t h t wo row s o f 2 1 -26 l a me l l ae ; a p e x w i t h a g ro u p of 1 3-20 I "- me l l ae ; m a x i l l a r y p a l p i w i t h 6 seg me n t s ; l a b i a l p a l p i w i th 3 segmen t s ; t i b i a l spurs 1 , 0 , 1 . MALE : u n k no w n L O C U S T Y P I C US : Mon t a rco ( Sp a i n ) TYP I CAL M A T E R I A L : h o l o t y p e F ! i n M D . H O ST S : u n k n o w n D I ST R I BU T I ON : P a l a e a rc t i c reg ion : S PA I N : Mon t a rco, M D ! S i erra d e E s p u n a ( To t a n a , M u rc i a ) , BM ! P a r a c u e l l os de J a r am a , M D ! E I P a rdo ( M a d r i d ) ( Ce b a l l os 1 92 7 ) ; I TALY : B a g n o l o ( Me l d o l a , For l ) ) ( Z a n g h e r i P . , 1 969 ) •

4 . Echthrodelp h a x hortu sensis ( Ab d u l -Nou r ) n . com b .

Pterogon a top u s hortusensis A b d u I -Nou r 1 97 6 : 276 . FEMALE ( F i g s 767,

768 ) : f u l l y w i n ge d ;

l en g t h 2 , 62-3 , 1 2 mm ;

head

1 1 48

Mem . Amer . E n ! .

I n s! . ,

n o . 37,

1 984

F i g . 769 - C h e l a o f Echthrodelp h a x hortusen sis ( Ab d u l -No u r ) ( I ta l y )

from Sc i o l ze

1 1 49

Gon a topod i n a e

Fig.

770 - M a l e ge n i t a l i a of Echthrodelp h a x hortu sensis Torrev i ej a ( Sp a i n )

( Ab d u l -No u r ) from

t e s t a c eou s , u s u a l l y w i t h a b ro w n spot o n v e r t e x ; a n te n n a e , p ro n o t u m a n d scu t u m t e s t aceou s ; s c u te l l u m , m e t a n o t u m a n d p ropod e u m b l ac k ( excep t for a p e x of p ropod e u m y e l l ow ) ; pet i o l e b l ack ; a b domen y e l l o w , w i t h terg i te 1 p a r t l y b l ack a n d t r a n s v e r s a l b ro w n b a n d s ; l eg s t e s t a ceou s ; h e a d ( F i g . 764 A ) sh i n y , smooth , w i t h o u t sc u l p t u re ; oce l l a r t r i a n g l e s t ron g l y i sosce l es ( F i g . 764 A ) ; POL = 1 ; O L = 4 ; OOL = 7 ; f ro n s w i t h med i a n k ee l , v i s i b l e a l so amon g oce l l i ; p ro n o t u m crossed b y a t r a n s v e r s a l i m p ress i o n , w i t h co l l a r a n d s i d es sh i n y a n d w i t h o u t scu l p t u re ; d i sc d u l l a n d g r a n u l a t e d ; pos t er i or edges of prono t u m n o t produced i n to l o b e s ; scu t u m sh i n y , w e a k l y p u n c t a t e ; n o t a u l i ces d i s t i n c t a n d co m p l e t e , j o i n t a t t h e pos t e r i o r m a r g i n o f sc u t u m ; s c u t e l l u m sh i n y , smoo t h , w e ak l y p u n c t a t e ; m e t a n o t u m very reduced , shor t ; p ropode u m f u l l y ret i c u l a te ru gose , w i t h o u t l on g i t u d i n a l k ee l s ; fore-w i n g w i t h one b row n t r a n sv e rs a l b a n d ; fore troch a n ters a p p rox i m a t e l y fou r t i mes a s l o n g as b road ; en l a r ged c l a w ( F i g . 769 ) w i t h s u b a p i c a l toot h a n d 4-5 l a me l l ae ; seg m e n t 5 of f ro n t t a rs u s ( F i g . 769 ) w i t h 1 r o w o f 7- 1 1 l a me l l ae ; a p e x w i t h a g ro u p o f 6 - 1 1 l a me l l ae ; m ax i l l a r y p a l p i w i t h 5 seg men t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l spurs 1 , 0 , 1 . MALE : f u l l y w i n g e d ;

l en g t h

1 , 37 m m ;

h e a d b l ac k ,

w i t h m a n d i b l es y e l l ow ;

Mem . A me r . E n ! .

1 1 50

I nst . ,

n o . 37,

1 984

a n te n n a e b row n ; t h orax a n d propod e u m b l ack ; a b domen red d i s h - b row n ; l eg s t e s t a ceou s ; a n t e n n a l seg men t 3 f o u r or more t h a n f o u r t i me s as l on g a s b ro a d ; h e a d s h i n y ,

w i t h o u t scu l p t u r e ; fron t a l

l i n e a b s e n t ; occ i p i t a l

c a r i n a comp l e t e ; scu t u m h a i r y , w e a k l y g r a n u l a ted ; n o t a u l i ces com p l e t e a n d d i s t i n c t , pos t er i o r l y s e p a r a t e d ; scu t e l l u m a n d met a n o t u m sh i n y , w i t hou t scu l p t u re ; p ropod e u m ret i c u l a t e ru gose , w i t h o u t l on g i t u d i n a l k ee l s o n p o s t e r i or s u rface ; fore-w i n g h y a l i n e , w i t ho u t b ro w n t r a n sversa l b a n d s ; dors a l p rocess o f gon oforceps ( F i g . 770 ) v e r y s h or t , po i n ted a n d t r a n sverse ; m a x i l l a ry p a l p i w i t h 4 s e g men t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . L O C U S T Y P I C US : H o r t u s ( F rance ) TYP I CA L M A T E R I AL : h o l o t y p e F ! a n d 2 p a r a t y p es F F ! i n P . H O ST S : i n F r a n c e Rib a u todelp h a x imit a n s R i b a u t ( Ab d u l -N o u r 1 976 ) ; i n I t a I y L a odelp h a x s triatellus ( F a l i e n ) a n d Megadelp h a x sordidulus ( St a I ) D I ST R I B U T I OI� : P a l a e a rc t i c reg i o n : F R AN C E : H o r t u s ( He ra u l t ) , P ! M o n t a r n a u d ( He ra u l t ) , P ! S PA I N : Torrev i ej a ( A l i c a n te ) , B M ! I TALY : Sc i o l ze ( Tor i n o ) , O L ! Port acomaro ( As t i ) , E N ! V o l a n o ( Fe r r a r a ) , V E ! C a se l l a ( G enova ) , G E ! CANARY I SLANDS : G r a n C a n a r i a , T N ! E G Y P T : Ben i You ssej , WA ! F a y ed , WA ! G i z a , WA !

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5 . Ech throdelp h a x i t a l icus n . s p .

F E MA L E : f u l l y w i n g e d ; l e n g t h 2 , 56-2 , 80 m m ; head t e s t aceou s , w i t h a b ro w n oce l l a r s p o t ; a n t e n n a e , l eg s a n d p ro t horax t e s t a ceOU 5 j rneso t horax , m e t a t horax a n d p ropodeum b l ac k ; a b domen b ro w n , w i t h tes t ac eo u s reg i on s ; a n t en n a l s e g me n t s i n f o l l o w i n g p ropor t i on s : 6 : 1 2 : 8 : 6 : 5 : 6 : 5 : 5 : 6 : 9 ; head s h i n y , w e a k l y g ra n u l a t e d ; frons w i t h a com p l e t e med i a n k ee l ; P O L = 2 ; OL = 3 ; OOL = 9 ; occ i p i t a l c a r i n a s l i g h t l y v i s i b l e b eh i n d t h e oce l l i a n d on t h e s i des o f t h e pos t e r i or oce l l i ; pron o t u m d u l l , c rossed b y a t r a n s v e r s a l i m p re s s i o n ; a n t e r i o r co l l a r s h i n y a n d weak l y g r a n u l a t ed ; d i sc d u l l a n d s t ron g l y g ra n u l a t e d ; p o s te r i o r e d g e s of pron o t u m n o t p roduced i n to l obes ; scu t u m sh i n y , w i t h o u t sc u l p t u r e , smoo t h ; m e t a n o t u m ru gose , d u l l ; propodeum d u l l , re t i c u l a t e ru gose ; p o s t e r i or a n d a n t e r i or s u rface w i t h o u t l o n g i t u d i n a l k ee l s ; fore-w i n g w i t h o n e d a r k t r a n s v e r s a l b a n d benea t h t h e p t e ro s t i g m a ; rad i a l ce l l open ; r a d i a l ve i n w i t h d i s t a l p a r t m u c h l on g e r t h a n p rox i m a l p a r t ( 25 : 8 ) ; fore t a rs a l seg men t s i n f o l l ow i n g p ropor t i o n s : 1 7 : 4 : 5 : 1 4 : 20 ; segme n t 3 o f f ro n t t a r s u s p ro d u ced i n to a hook ; e n l arged c l a w ( F i g . 7 7 1 ) w i t h s u b a p i c a l tooth a n d 4 l a me l l a e ; seg m e n t 5 o f f r o n t t a rs u s ( F i g . 771 ) w i t h o n e row o f 1 3 l a me l l ae ; a p ex w i t h a g r o u p o f 6 l ame l l ae ; m a x i l l a r y p a l p i w i t h 5 segme n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MA LE : u n k n o w n L O C U S T Y P I C US :

Pa l o ( I ta l y )

T Y P I CA L M A T E R I AL : HOSTS: u n k nown

ho l o t y p e F !

i n GE ;

1 p a r a t y pe F !

in P.

1 1 51

Gon a topod i n a e

F i g . 771

-

C h e l a of Echthrodelp h a x italicus n . sp .

( ty p e )

Mem . Amer .

1 1 52

D I ST R I BU T I ON :

En ! .

P a l ae a rc t i c reg i o n :

A n d a l ou se s ( O r a n ) ,

I ns ! . ,

I TALY :

n o . 37,

1 984

P a l o ( Roma ) ,

G E ! ALG E R I A :

P!

E T H I OP I AN S P E C I E S O F E CHTHRODELPHA X T h ree s p ec i es a re k n ow n .

KEY T O THE SPEC I ES

F E MA L E S

2

P r o p o d e u m w i t h t wo m ed i a n , l on g i t u d i n a l a n d p a ra l l e d o r sa l a n d p o s t e r i o r s u r f a c e ; sp e c i e s b i g . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 . obsc ul'uS ( C eb a l l o s ) P r op od e u m w i t h ou t l on g i t u d i n a l k e e l s ; s p e c i e s s ma l l S c u t u m b l a c k ; p r on o t u m f u l l y o r p a r t l y b l a ck , s o me t d i s c t e s t a c e ou s o r r ed d i s h . . . . . . . . . . . . . . . . . . . . . . . . . 1 . migl'a tol'ius B en o i t S c u t u m a n d p r on o t u m t e s t a c e ou s 2 . a iel' n . s p . .

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l k ee l s on . . . . . . . . . . . . . . . . . . .2 i me s w i t h . . . . . . . . . . •

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MALES

U n k no w n

1 . Ech thl'odelp h a x m igl'a tOl'ius Benoi t

Ech thl'odelp h a x migl'a tol'iu s Beno i t 1 953b : 397 .

F E MA L E : f u l l y w i n ge d ; l e n g t h 2 , 25 m m ; head test aceo u s , w i t h b ro w n t r a n s v e rs a l b a n d on vertex ; a n te n n ae t e s t a c eo u s ; prono t u m f u l l y b l ac k ; occ a s i o n a l l y c o l l a r b l ack a n d d i sc test aceo u s or redd i s h ; p ropec t u s b l ac k ; scu t u m , s c u te l l u m , met a n o t u m a n d propo d e u m b l ac k ; a b domen b l ack ; l eg s t e s t aceo u s ; h e a d s h i n y , w i t h o u t s c u l p t u re , except for occ i p u t a n d f ro n t p a r t o f v ertex g ra n u l a t ed ; ocel l a r t r i a n g l e s t ron g l y i sosce l e s , w i t h med i a n k ee l ; O L m u c h l on ger t h a n POL ; p ron o t u m cros sed b y a t r a n s v e r s a l i mp ress i o n , w i t h co l l a r smooth a n d w i t h ou t scu l p t u re ;

d i sc

g r a n u I a te d ; pos t e r i o r edges of prono t u m not p rod uced i n to l obes ; scu t u m

1 1 53

G o n a t opod i n ae

Fi g . 772 - C h e l a of Echthrodelp h a x m igratorius Be n o i t

( type)

sh i n y , w i t h o u t sc u l p t u re ; n o t a u l i ces d i s t i n c t a n d com p l e t e , j o i n t a t t h e pos t e r i o r m a r g i n of sc u t u m ; sc u t e l l u m sh i n y , w i t h o u t sc u l p t u re ; m e t a n o t u m v e r y redu ced a n d s h o r t ; p ropodeum r e t i c u I a t e ru gose, w i t h t r a n s versa I k ee l s , w i t h o u t l on g i t u d i n a l kee l s on p os t e r i o r s u rface ; fore-w i n g h y a l i n e , w i t h o u t d a rk t r a n svers a l b a n d s ; fore t roc h a n ters a p p ro x i m a t e l y f o u r t i mes as l o n g a s b road ; en l arged c l a w ( F i g . 772 ) w i t h s u b a p i c a l too t h a n d 4 l a me l l a e ; segmen t 5 of f ro n t t a r s u s ( F i g . 772 ) w i t h 1 row of 1 1 l a me l l ae ; apex w i t h a g ro u p of 1 0 l a me l l a e ; m a x i l l a r y p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : Bek i l y ( M a d a g asc a r ) TYP I'CAL M A T E R I A L : h o l o t y p e F ! i n P . HOST S : u n k no w n D I ST R I BU T I O N : E t h i op i a n reg i o n : SOMAL I A : A f g o i ( Be n a d i r ) , OL ! ZA I R E : G a r a m b a r\ & t . P a r k , TV ! E d w ard L a k e , T V ! Z I M BAB WE : S a l i s b u ry , O T ! S O U T H A F R I C A : Derdepoor t , B M ! Port S t . Joh n s ( Po n do l a n d ) , B M ! BO ! MADAGASCAR : Bek i I y , P ! T V !

M e m . Ame r . E n t .

1 1 54

Fig.

773

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I nst . ,

no. 37,

1 984

C h e l a of Ech throdelphax a fer n . sp . from Af g o i

( Som a l i a )

Gon a topod i n a e

1 1 55

2 . Ech throdelph a x a fer n . s p .

F E MALE : f u l l y w i n g e d ; l en g t h 2 , 37-2 , 50 m m ; h e a d , a n t e n n a e , p ro no t u m a n d sc u t u m t es t aceou s ; occa s i o n a l l y sc u t u m red d i s h - b row n ; s c u te l l u m , m e t a n o t u m a n d p ropodeum b l ac k ; a b domen t e s t aceo u s , w i t h t e r g i te 1 p a r t l y b l ac k ; l eg s tes t aceou s ; h e a d sh i n y , w i t h o u t sc u l p t u re , except for occ i put and f r o n t part of v ertex g ra n u I a ted ; oce l l a r tri a n g I e s t rong I y i sosce l es , w i t h med i a n k ee l ; OL m u c h l o n ge r t h a n P O L. ; p rono t u m crossed by a t r a n s v e rs a l i m p ress i o n , w i t h c o l l a r sh i n y and w i t h o u t scu l p t u re ; d i sc d u l l a n d g r a n u l a t e d ; pos t e r i o r e d g e s of pron o t u m not produce d i n t o l ob e s ; s c u t u m s h i n y , w i t h o u t sc u l p t u r e ; n o t a u l i ce s d i s t i n c t a n d com p l et e , j o i n t a t t h e pos t er i or m a rg i n o f sc u t u m ; s c u t e l l u m s h i n y , w i t h o u t sc u I p t u r e ; met a n o t u m v er y red uced a n d s h o r t ; p ropodeum ret i c u I a t e r u go s e , w i t h t r a n s v e rs a l k ee l s , w i t h o u t l o n g i t u d i n a l k ee l s o n pos t er i or s u rf a c e ; fore-w i n g w i t h o n e b ro w n t r a n sv e rs a l b a n d ; fore t r oc h a n ters a p p rox i m a t e l y fou r t i me s a s long a s b ro a d ; en l a rged c l a w ( F i g . 773 ) w i t h s u b a p i c a l t oo t h a n d 4 l a me l l ae ; segmen t 5 of fron t t a rs u s ( F i g . 773 ) w i t h 1 row of 8 l a me l l ae ; a p e x w i t h a g ro u p of 7-9 l a m e l l ae ; m a x i l l a r y p a l p i w i t h 5 s e g me n t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : S a l i s b u ry ( Z i m b a b w e ) TYP I CAL M A T E R I AL : h o l o t y p e F ! a n d 3 p a r a t y pes F F ! i n BM ; 4 p a r a t y pe s F F ! i n O L ; 2 p a r a t y pes F F ! i n T V . H OSTS : u n k n o w n D I ST R I BU T I ON : E t h i op i a n reg i o n : SOMAL I A : Af g o i ( Ben a d i r ) O L ! Z A I R E : G a r a m b a N a t . P a rk , OL ! TV ! Z I M BA BWE : S a l i sb u r y , BM ! C h i sh a w a s h a ( Sa l i s b u r y ) , BM ! SOU T H A F R I CA : K a t berg ( E C a pe P rov i n ce ) , OL ! BM ! P o r t St . J o h n s ( Po n do l a n d ) , BM !

3 . Echthrodelp h a x o b s curus ( C e b a I l os ) n . com b .

Mercetia o b s c ur a C e b a I l os 1 936 : 45 . Mercetia o b scura C e b a I l os : Ben o i t 1 953b : 399 .

F E M A L E : f u l l y w i n g e d ; l e n g t h 6 , 93 m m ; h e a d red d i sh -ferru g i no u s , w i t h occ i p u t a n d oce l l a r reg i on b l ac k ; a n te n n a e w i t h segmen t s 1 -8 t e s t aceo u s ( ot h er seg men t s m i ss i n g i n t h e o n l y k n o w n spec i me n ) ; t h o r a x , p ropod e u m a n d a b domen b l ack ; l e g s testaceo u s , w i t h c l u b s o f femora d a rkened ; h e a d d u l l , h a i r l es s , g r a n u l a t e d ; occ i p i t a l c a r i n a i n com p l et e , o n l y v i s i b l e beh i n d t h e oce l l i a n d o n t h e s i des of t h e pos t e r i or oce l l i ; oce l l a r t r i a n g l e eq u i l a tera l ; p rono t u m crossed b y a t r a n svers a l i m p ress i o n , d u l l , h a i r l es s , smoo t h , a l u t a ceou s , w i t h some p u n c t u res o n d i sc ; d i sc w i t h a short t r a n s v e rs a l i m p ress i o n b eh i n d t h e a n t e r i o r t r a n svers a l i m p ress i on ;

Mem . A me r . E n t .

1 1 56

F i g . 774

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I nst . ,

n o . 37,

1 984

C h e l a of Echthrodelp h a x ob scurus ( C e b a l l os ) ( t y p e )

poster i o r edges o f p rono t u m n o t p rod uced i n to l obes ; sc u t u m d u l l , g ra n u l a t e d , h a i r l es s ; not a u l i ce s v i s i b l e a n d com p l e t e , poster i or l y sep a r a t e d , b u t v e r y n e a r , s t ro n g l y con verg i n g ; s c u t e l l u m sh i n y , smoo t h , w i t h o u t sc u l p t u r e ; m e t a no t u m very s h o r t a n d reduced , s t ron g l y re t i c u l a t e r u gose ; p ropod e u m ret i c u l a t e r u gose , w i t h t w o l o n g i t u d i n a l k ee l s o n p o s t e r i o r s u rf ac e ; dorsa l s u rface a l so w i t h t w o med i a n l o n g i t u d i n a l k ee l s ; fore- w i n g w i th t w o b ro w n t r a n s vers a l b a n d s ; d i s t a l p a r t o f r ad i a l v e i n m u c h I c C' g e r t h a n p rox i m a l p a r t ( 3 1 : 1 0 ) ; h i n d -w i n g w i t h d i s t a l reg ion d a rk ; s e g me n t 1 of f ron t t a r s u s s h o r t e r t h a n segmen t 4 ( 1 5 : 1 9 ) ; e n l a rged c l a w ( F i g . 774 ) w i t h s u b a p i c a l t oo t h and 10 l a me l l ae ; segmen t 5 of f ro n t t a rs u s ( F i g . 774 ) w i t h t w o rows of 2 9 l a me l l ae ; a p e x w i t h a g ro u p o f 2 5 l a m e l l ae ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k no w n L O C U S T Y P I C U S : Pe r i n e t ( M a d a g a s ca r ) T Y P I CAL M A T E R I AL : h o l o t y p e F ! i n M D . HOSTS : u n k n ow n D I ST R I BU T I O N : E t h i op i a n reg i on :

MADAGASCAR :

Pe r i n e t ,

MD !

1 1 57

Gona topod i n a e

OR I E N TAL S P E C I E S OF ECHTHRODELPHAX

T h ree sp e c i es a re k n o w n .

K E Y TO T H E S P E C I E S

F E MA L E S

F o r e -w i n g w i t h t wo b r own t ra n s v e r s a I b a n d s ; b od y f u I I y t e s t a c e ou s , w i t h p e t i o l e b l a ck a n d a b d omen d a rk e n ed •

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1 . rufus n . sp . F o r e -w i n g h y a l i n e , w i t h ou t d a rk t ra n s v e r s a l b an d s ; t h o r ax a n d p r op od eu m p a r t l y b l a c k 2 S c u t u m b l a ck ; o c e l l a r t r i a n g l e w i t h ou t med i a n k e e l . . . . . . . . . . . . . . . . . . . . . . 2 . fairchildii R . C . L . P e rk i n s ( s e e A u s t ra l i a n sp e c i e s ) S c u t u m r ed d i sh ; o c e l l a r t r i a n g l e w i t h d i s t i n c t med i a n k e e l . . . . . . . . . . . . . . 3 . laotian u s n . sp . •

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MALES

O n l y the m a l e of E .

fairchildii is k no w n .

1 . Ech throdelp h a x rufus n . sp .

F E MAL E : fu l l y w i n ged ; l en g t h 2 , 5 m m ; t e s t aceo u s , w i t h pet i o l e b l ack a n d a b domen d a rkened ; h e a d sh i n y , w i t h o u t sc u l p t u re , excep t for occ i p u t a n d p a r t of v e rt ex g r a n u l a t ed ; oce l l a r t r i a n g l e s t ro n g l y i sosce l e s , w i t h med i a n k e e l ; OL m u c h l on g e r t h a n POL ; p ro n o t u m crossed b y a t ra n s v e rs a l i m p ress i o n , w i t h co l l a r s h i n y a n d w i t hou t sc u l p t u r e ; d i sc d u l l a n d g r a n u l a ted ; pos t e r i or e d g e s of p rono t u m n o t produ ced i n to l o b e s ; scu t u m s h i n y , w i t h ou t sc u l p t u re ; no t a u l i ce s d i s t i nc t a n d comp l e t e , j o i n t a t t h e pos t e r i or m a r g i n of scu t u m ; scu t e l l u m sh i n y , w i t h ou t scu l p t u re ; m e t a no t u m v e r y reduced a n d shor t ; p ropodeu m ret i c u I a t e ru gose , w i t h s t rong t ra n svers a l s t r i ae m a i n l y on pos t e r i or s u rf ace ; pos t e r i or s u rface w i t h ou t l o n g i t u d i n a l k ee l s ; fore-w i n g w i t h t w o b ro w n t r a n s v e rs a l b a n d s ; fore t roc h a n t ers a p p rox i m a te l y fo u r t i mes a s l on g a s b road ; en l a rged c l a w ( F i g . 775 ) w i t h s u b a p i c a l too t h and 4 l a m e l l ae ; segme n t 5 of fron t t a rs u s ( F i g . 775 ) w i t h 1 row of 7-9 l a me l l ae ; a p ex w i t h a g ro u p of 7-8 l a me l l ae ; m a x i I l a r y p a l p i w i t h 5-6 segmen t s ; l a b i a l p a l p i w i t h 3

1 1 58

Mem . Amer . E n ! .

I ns! . ,

n o . 37,

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F i g . 775 - C h e l a of Ech throdelp h a x rufus n . s p . from K h aoc h a n g ( Th a i l a n d )

s e g me n t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k no w n L O C U S T Y P I C U S : De T h a N g o n e ( L aos ) T Y P I CAL M A T E R I AL : h o l o t y p e F ! i n B ; 1 p a ra t y p e F ! i n OL . HOSTS: u nknown D I ST R I BU T I O N : O r i en t a l reg i o n : LAOS : D e T h a N gone ( V i en t i a n e , B ! T HA I LA N D : K h aoc h a n g ( T r a n g , K h a op h a p p h a ) , OL !

C i S i on ) ,

2 . Ech throdelp h a x fa irchildii R . C . L . Perk i n s ( se e A u s t r a l i a n reg i o n )

3 . Echthrodelp h a x la otia n us n . s p .

F E MA L E : fu I I y w i n g ed ; l en g t h 2 , 5 m m ; h e a d , a n te n n a e , p rono t u m a n d scu t u m t e s t aceou s ; sc u t e l l u m , m e t a no t u m , p ropodeum a n d p e t i o l e b l ack , excep t for a p ex of p ropod e u m y e l l ow j a b domen t e s t aceo u s , w i t h terg i t e p a r t l y b row n ; h e a d sh i n y , w i t hou t sc u l p t u r e , excep t for occ i p u t a n d p a r t of v e r t e x g r a n u l a ted ; o c e l l a r t r i a n g l e s t ro n g l y i sosce l es , w i t h d i s t i n c t

1 1 59

Gon a topod i n a e

F i g . 776

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C h e l a of Echthrodelp h a x l a o tia n u s n . s p .

( type)

med i a n k ee l ; OL m u c h l on g er t h a n POL ; p r o n o t u m cros sed b y a t ra n s v er s a l i m p re ss i o n , w i t h co l l a r w i t h o u t scu l p t u re ; d i sc g r a n u l a t e d ; p o s te r i o r e d g e s of p ro n o t u m n o t p roduced i n t o l ob e s ; sc u t u m s h i n y , w i t ho u t sc u l p t u re ; n o t a u l i ce s d i s t i n c t a n d comp l et e , poster i or l y s e p a r a t e d ; sc u t e l l u m s h i n y , w e a k l y p u nc t a t e ; met a n o t u m very redu ced a n d s h o r t ; p ro p o d e u m ret i c u l a t e r u g o s e , w i t h s t ro n g t ra n svers a l k ee l s , w i t h o u t l on g i t u d i n a l k ee l s on p o s te r i o r s u rf a ce ; fore- w i n g h y a l i n e , w i t h o u t d a rk t ra n s v e r s a I b a n d s ; fore t roc h a n t e r s a p p rox i m a t e l y f o u r t i mes a s l o n g as b roa d ; e n l a rged c l a w ( F i g . 776 ) w i t h s u b a p i c a l t oo t h a n d 5 l a m e l l ae ; seg m e n t 5 of fron t t a r s u s ( F i g . 776 ) w i t h 1 row of 9 l a m e l l ae ; a pex w i t h a g ro u p o f 9 l a m e l l ae ; m a x i l l a r y p a l p i w i t h 5 segmen t s ; w i t h 2 s e g me n t s ; t i b i a l s p u r s 1 , 0 , 1 . MAL E : u n k n o w n L O C U S TYP I C U S : V i e n t i a n e ( L aos ) TYP I CAL M AT E R I AL : h o l o t y p e F ! i n B . H OS T S : u n k n o w n D I ST R I B U T I O N : O r i e n t a l reg i o n : LAOS : V i e n t i a n e , B !

labial

palpi

Mem . Amer . E n t .

1 1 60

F i g . 777

I nst . ,

n o . 37,

1 984

C h e l a of Echthrodelp h a x krom bein i n .

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( ty p e )

N E A R C T I C S P E C I ES O F ECHTHRODELPHAX

O n e spec i es i s k no w n .

Ec h th ro d elph a x kromb eini n . s p .

F E MA L E : f u l l y w i n ged ; l e n g t h 1 , 9 3 m m ; h e a d , a n t e n n a e , prothorax , s c u t u m , abdomen a n d l e g s y e l l ow - t e s t aceou s ; s c u te l l u m , m e t a n ot u m , mesop l e u r a , m e t a p l eu ra a n d p ropode u m b l ac k ; a n te n n ae d i s t a I I y t h i ck e n e d ; a n t en n a l seg m e n t s i n fo l l o w i n g p ropor t i on s : 6 : 3 : 4 : 3 : 3 : 3 : 3 : 3 , 5 : 5 ; h e a d sh i n y , a l mo s t f l a t , smoo t h , w i t h o u t scu l p t u re ;

3:

occ i p i t a l c a r i n a i n v i s i b l e ; frons w i t h m ed i a n k ee l ; P O L = 0 , 5 ; Ol = 3 ; OOl = 5 ; p ronot u m s h i n y , c rossed b y a t r a n s v ers a l i m p re s s i o n , w e a k l y g r a n u l a ted ; pos t e r i o r e d g e s n o t p ro d u ce d i n to l obes ; scu t u m , sc u t e l l u m a n d m e t a n o t u m smoo t h , sh i n y , w i t h o u t scu l p t u re ; not a u l i c e s comp l e t e , poster i or l y sep a r a t ed ; p ropodeum d u l l , f u l l y ret i c u l a t e ru gose ; fore-w i n g h y a l i n e , w i t hou t d a r k t r a n s v e r s a l b an d s ; d i s t a l p a r t o f r a d i a l v e i n m u c h l on g e r t h a n prox i m a l p a r t ( 1 2 : 4 ) ; fore t a r s a l seg men t s i n fo l l o w i n g p ropor t i o n s : 7 : 1 , 5 : 2 : 8 : 1 1 ; en l a rged c l a w ( F i g . 777 ) w i t h s u b a p i c a l tooth a n d 4 l a me l l ae ; s e g m e n t 5 o f fron t t a r s u s ( F i g . 777 ) w i t h 1 row of 6 l a me l l ae ; a p e x w i t h a g ro u p of 7 l a me l l ae ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k no w n l O C U S T Y P I C U S : Cora l G a b l es ( F l o r i d a , U . S . A . ) T Y P I CA L M A T E R I AL : HOST S : u n k no w n

holotype F !

i n WA .

Gon a topod i n ae

1 1 61

D I ST R I BU T I O N : N e a rc t i c re g i on : U . S . A . : Cor a l G a b l es ( F l a . ) , WA ! NOT E : t h e spec i es i s n a med i n honor of M r . K a r l V . K rombe i n .

A U S T R A L I A N S P E C I E S OF ECiITH:70JJELPJJAX

F o u r s p ec i es a re k n ow n .

K E Y TO T H E S P E C I E S

F E MA L E S

F o r e - w i n g w i t h t wo d a rk t ra n s v e r sa I b a n d s F o r e - w i n g h y a l i n e , w i t h ou t d a r k t ra n s v e r s a T h o r a x a n d p r op o d e u m f u l l y t e s t a c e o u s 3 . 'o ifa s c i a t u s R . C . L . T h o ra x a n d p r op o d e u m p a r t l y b l a c k . . . . . . . . . . . . . . . . . . . . . . . 4 . pelop s n . s p . P r o n o t u m t e s t a c e ou s l . fairch ildii R . C . L . P r on o t u m b l a c k 2 . n igricol1is !: L C . L . .

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P e rk i n s P e rk i n s

MALES

N o t au l i c e s p o s t e r i o r l y

j o i nt 3 . fairchildii R . C . L . P e rk i n s N o t a u l i c e s p o s t e r i o r l y d i s t i n c t l y s ep a ra t e d 2 . nigricollis I' . C . L . P e rk i n s .

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pelops a n d b ifa s c i a t u s a re u n ko w n .

1 . Ech throdelp h a x fairchildii R . C . L .

Perk i n s

Echthrodelp h a x fairchildii R . C . L . Perk i n s 1 903 : 37 . Echthrodelp h a x fa irchildii R . C . L . P e r k i n s : K i effer 1 9 1 4b : 1 7 . Echthrodelp h a x bicolor E s a k i a n d H a s h i moto 1 93 1 : 29 ( n . s y n . ) ; locus typic u s O i t a ( K y u sh u , J a p a n ) . Echthrodelp h a x bicolor E s a k i a n d H a s h i m o t o : E s a k i a n d H a s h i moto 1 93 3 : 1 9 . Ech throdelp h a x .'Jicolor E s a k i a n d H a s h i moto : E s a k i a n d H a s h i moto

Mem . Amer . E n t .

1 1 62

I ns t . ,

no.

37,

1 984

\

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778

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C h e l a of Echthrodelp h a x fa irchildii R . C . L . N a n goku ( J a p a n )

Perk i n s from

Gon a topod i n ae

1 1 63

1 9 35 : 1 8 . Ech throdelp h a x b icolor E s a k i a n d H a s h i mo t o : E s a k i a n d H a s h i moto 1 9 36 : 1 8 . Ech throdelp h a x b i color E s a k i a n d H a s h i moto : E s a k i a n d S a m e s h i m a 1 939 : 42 ; Ech throdelp h a x b i color E s a k i a n d H a s h i moto : E s a k i a n d S a m e s h i m a 1 940 : 27 . nec Echthrodelp h a x fairch ildii R . C . L . Perk i n s : N a g y 1 967b : 332 .

F E MALE : f u l l y w i n ge d ; l en g t h 2 , 20-2 , 92 m m ; h e a d t e s t aceou s , u s u a l l y w i t h t r a n s v e r s a I b ro w n b a n d on v ertex ; a n t e n n a e a n d p ro n o t u m t e s t aceou s ; scu t u m , sc u t e l l u m , m e t a no t u m , propod e u m a n d p e t i o l e b l ac k ( except for

F i g . 779 - M a l e of Ech throdelp h a x fairchildii R . C . L . P e rk i n s from E w a ( Oa h u ,

H aw a i i an

I s l ands )

1 1 64

Fig.

Mem . Amer .

En! .

I ns! . ,

n o . 37 ,

1 984

780 - M a l e of Ech throdelp h a x fa irchildii R . C . L . ( O a h u , H aw a i i a n I s l a n d s )

Perk i n s f rom E w a

a pex o f p ropodeum y e l l ow ) ; a b domen t e s t aceo u s , w i t h terg i tes 1 , 2 a n d 4 p a r t l y b ro w n ; l eg s test aceo u s ; h e a d ( F i g . 764 B ) s h i n y , w i t hou t s c u l p t u re ; oce l l a r t r i a n g l e s t r on g l y i sosce l e s ( F i g . 764 B ) , w i t h o u t med i a n keel ; OL m u c h l on ger t h a n P O L ; pron o t u m crossed by a t r a n svers a l i mpres s i o n , s h i n y ; poster i o r e d g e s o f p ro n o t u m n o t p ro d u ced i n to l obes ; scu t u m smoo t h , sh i n y , w e a k l y p u nc t a t e ; n ot a u l i ce s d i s t i nc t a n d comp l e t e , p o s t er i o r l y s e p a r a ted ; sc u te l l u m s moo t h , s h i n y , w e a k l y p u n c t a t e ; m e t a n o t u m very red u ced a n d s h o r t ; p ropodeum ret i cu I a te ru gose , w i t h t ra n s v e r s a l kee l s ; poster i or s u rf ace w i t h o u t l o n g i t u d i n a l k ee l s ; fore-w i n g h y a l i ne , w i t h o u t d a rk t r a n sversa l b a n d s ; fore t roc h a n ters a p p rox i m a t e l y f o u r t i mes as l o n g a s b ro a d ; segmen t 4 of f r o n t t a r s u s as l on g a s segmen t 1 ; en l a rged c l a w ( F i g . 778 ) w i t h s u b a p i c a l tooth a n d 4-5 l a m e l l ae ; seg men t 5 of f ro n t t a r s u s ( F i g . 778 ) w i t h 1 row o f 9- 1 2 l a m e l l ae ; a p ex w i t h a g r o u p of 6-1 1 l a me l l ae ; m a x i l l a r y p a l p i w i t h 3-4 segmen t s ; l ab i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE ( F i gs 780 , 7 8 1 ) : f u l l y w i n g e d ; l e n g t h 1 , 68-1 , 90 m m ; h e a d b ro w n or b l ac k , w i t h m a n d i b l es y e l l ow ; a n t en n ae b row n ; thorax and p ropodeum b l ac k ; a b domen b row n ; l eg s y e l l o w ; a n ten n a l segmen t s in f o l l o w i n g p roport i on s : 5 : 5 : 9 : 1 0 : 1 0 : 9 : 1 0 : 9 : 9 : 1 1 ; a n t en n a l segmen t 3 f o u r o r

Gon a topod i n a e

F i g . 781

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M a l e ge n i t a l i a of Ech throdelph a x fa irchildii R . C . l . Perk i n s f rom los B a n o s ( P h i I i p p i n es )

more t h a n fou r t i mes as l o n g a s b ro a d ; h e a d s h i n y , smoo t h , w i t hou t scu l p t u re , w e a k l y h a i ry ; t e m p l es p rom i n e n t , n e a r l y a s l o n g a s e y e s ; occ i p i t a l c a r i n a comp l e t e ; POL = 5 ; Ol = 3 ; OOl = 3 ; b rea d t h of t h e oce l l i : 1 ; oce l l a r t r i a n g l e w i t h o u t med i a n k e e l ; f r o n s w i t h med i a n k e e l b e t w ee n t h e a n t en n a l sock e t s ; sc u t u m s h i n y , w i t h o u t scu l p t u re o n p o s t e r i or h a l f ; a n te r ior h a l f g r a n u l a ted ; n o t a u l i c e s d i s t i n c t , comp l e t e , j o i n t a t t h e p o s t e r i o r m a rg i n o f sc u t u m ; scu t e l l u m s h i n y , smoo t h , w i t h o u t scu l p t u re ; m e t a no t u m s h or t , v e r y red uced , s h i n y ; p ro p o d e u m ret i c u l a t e r u gose , w i t h o u t l on g i t u d i n a l k ee l s on poster i or s u rf ace ; fore-w i n g h y a l i n e , w i t h o u t d a r k t r a n sv e r s a I b a n d s ; d o r s a I p rocess o f g o n oforceps ( F i g . 7 8 1 ) v e r y short and s l en d e r ; m a x i l l a ry p a l p i w i t h 4 seg me n t s ; l a b i a l p a l p i w i t h 2 s e g men t s ; t i b i a l s p u rs 1 , 1 , 2 . lOCUS T Y P I C U S : E w a P l a n t a t i on ( O a h u , H a w a i i a n I s l a n d s ) TY P I CAL M A T E R I AL : l ec t o t y p e F ! ( d es i g n a ted b y O l m i 1 982 ) of E . fa irchildii i n B ; h o l o t y p e F ! a n d a l l o t y p e M ! of E . b icolor i n KY . H O S T S : i n t h e H a w a i i a n I s l a n d s Perkin siella s a cch aricida K i rk . a n d A lo h a ipom e a e ( R . C . l .

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( Ho rv a t h ) ( E s a k i and H a s h i moto 1 9 33 ) , L a odelp h a x s tria tell u s F a l l e n a n d Nilap a rva ta l ugens ( S t a l ) ( = oryzae M a t s . ) ( T hompson 1 954 ) ; i n I n d i a Nilap a rva ta s p . ; i n T a i w a n Nilaparva ta l u gens ( S t a l ) ( C h u a n d H i ra s h i m a 1 98 1 ) ; i n t h e P h i l i p p i n e s Nilap arva t a l u gens ( S t a l ) ( C h a n d ra 1 980b ) a n d Soga tella furcifera ( Ho rv a t h ) ; accord i n g t o C h a n d ra ( 1 980b ) i n t h e P h i l i p p i n e s E . fairchildii c a n p a r a s i t i z e a l so t h e C i c a d e l l i d Nephotettix sp . : t h i s i n forma t i on m u s t b e conf i rmed . PARAS I T E S : i n H a w a i i a n I s l a n d s Hypergon a top u s h a l<'aiien s is R . C . L . Perk i n s ( T i m b er l a k e 1 92 2 ) . D I ST R I B U T I O N : A u s t ra l i a n reg i o n : HAWA I I AN I SL A N D S : E w a P l a n t a t i on ( O a h u ) , B ! O L ! K a h u k u ( O a h u ) , B ! Hon o l u l u ( O a h u ) , BM ! K a u a i ( R . C . L . Perk i n s 1 905 ) ; e s t a b l i sh e d o n t h e o t h e r H a w a i i a n I s l a n d s for b i o l og i c a l con t ro l ( R . C . L . Perk i n s 1 90 5 ) . O r i en t a l reg i o n : TA I WA N : G a k k o , KY ! K i j i n , O L ! T a i n a n H s i e n ( K w a n tze l i n g ) , B ! I N D O N E S I A : P ek a l on g a n ( J a v a ) , B ! I N D I A : H o l a l u ( M a n d y a ) , BM ! P H I L I P P I N E I SLAN DS : J a nopo l ( T a n a u a n , B a t a n g a s ) , BM ! Los B a n o s ( L a g u n a ) , BM ! P a l aearc t i c reg i on : JAPAN : K a m i -g u n ( Koch i P r ef . ) , O L ! N I ! Nan g o k u ( Koc h i P ref . ) , N I ! O i t a ( K y u s h u ) , KY ! NOT E : accord i n g to N a g y ( 1 967b ) E . fairchildii occ u r s a l so i n Roman i a . h a v e not seen t h e on l y fem a l e s p ec i men fou n d b y M r . N a g y n e a r A g i gea . Accord i n g to t h e descr i p t i on , h o w e v e r , t he h e a d i s b l ac k , w hereas i n E . fairchildii i t ' s t e s t aceo u s , u s u a I I y w i t h a b ro w n t r a n sversa I b a n d o n vertex . I n a d d i t i on t he fore-w i n g s h o w s a d a rk t r a n svers a l b a n d , t h i n k t h a t t h i s fema l e spec i me n w h ereas i n E . fairch ildii i t ' s h y a l i n e . from Rom a n i a i s a n e w s p ec i es , d i ffere n t a l so from t h e o t h e r P a l a e a rc t i c spec i es .

2 . Echthrodelp h a x n igricollis R . C . L . Perk i n s

Ech throdelp h a x n igricollis R . C . L . P e rk i n s 1 90 5 : 49 . Ech throdelp h a x nigricollis R . C . L . Perk i n s : K i effer 1 9 1 4b :

1 7.

F E MAL E : f u l l y w i n g ed ; l e n g t h 1 , 64-2 , 1 9 m m ; h e a d t e s t aceou s , w i t h b ro w n t r a n svers a l b a n d on vertex ; a n ten n ae t e s t aceou s ; t horax a n d p ropodeum b l ac k ; a b domen b ro w n , w i t h a n i n t ermed i a t e terg i t e t e s t aceou s ; l eg s t e s t aceo u s ; h e a d s h i n y , w i t h o u t sc u l p t u re , except for occ i p u t a n d f ro n t p a r t o f vert ex gra n u l a t e d ; oce l l a r t r i a n g l e s t ron g l y i sosce l es , w i t h med i a n k ee l ; OL m u c h l o n g e r t h a n POL ; p ro n o t u m s h i n y , w i t h co l l a r w i t h o u t s c u l p t u re ; d i sc g r a n u l a ted ; poster i o r ed ges of pronot u m n o t p ro d u ced i n to l obes ; scu t u m s h i n y , w i t hou t sc u l p t u re on poster i o r h a l f ; a n te r i o r h a l f gra n u l a t e d ; n o t a u l i c e s d i s t i n c t a n d com p l e t e , poster i or l y s e p a ra t ed ; sc u te l l u m s h i n y , w i t hou t scu l p t u re ; m e t a n o t u m very reduced and s h or t ; p ropodeu m ret i c u l a t e ru gose ,

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post e r i o r s u rf ac e ; pos ter i or s u rf ace w i t h o u t l on g i t u d i n a l k ee l s ; fore- w i n g h y a l i n e , w i t h o u t d a r k t r a n svers a l b a n d s ; fore t roc h a n t e r s a p p rox i m a te l y f o u r t i me s a s l o n g a s b ro a d ; segmen t 4 o f f r o n t t a r s u s a s l o n g a s seg m e n t 1 ; en l a r ged c l a w ( F i q . 7;:;2 ) w i t h s u b a p i c a l tooth a n d 5 l a me l l ae ; seg m e n t 5 of f ro n t t a rs u s ( F i g . 782 ) w i t h 1 row of 8 l a m e l l ae ; a p ex w i t h a g ro u p o f 7 l a me l l ae ; m a x i l l a r y t p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i

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F i g . 783 - M a l e g en i t a l i a of Echthrodelp h a x n igricollis R . C . L . P e rk i n s f rom C a i rn s ( Queen s l a n d )

w i t h 2 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : f u l l y w i n ged ; l e n g t h 1 , 46 m m ; red d i s h -b row n , w i t h l e g s y e l l o w ; a b domen y e l l ow - b row n ; a n t e n n a e s l en d e r , l on g e r t h a n b od y , not d i s t a l l y t h i ck e n e d ; a n t en n a l seg m e n t s i n fo l l o w i n g p ropor t i on s : 4 : 4 : 9 : 9 : 9 : 1 0 : 9 : 9 : 9 : 1 1 ; a n t en n a l seg men t 3 more t h a n f i ve t i me s a s l o n g a s b road ( 5 , 33 ) ; h e a d s h i n y , smoo t h , f i n e l y p u n c t a t e , w i t h o u t scu l p t u re a mon g p u n c t u r e s ; f ro n t a l l i n e a b se n t ; occ i p u t conc a v e ; temp l es d i s t i n c t ; occ i p i t a l ca r i n a com p l et e , w e a k l y d i s t i n c t ; P O L = 3 ; OL = 2 ; OOL = 4 ; sc u t u m f i ne l y p u n c t a te , w i t h o u t s c u l p t u re a mon g p u n c t u res , smoo t h , s h i n y ; n ot a u l i ce s com p l e t e , poste r i or l y s e p a r a t e d ; m i n i m u m d i s t a nce b e t ween t h e not a u I i ce s l o n g e r t h a n b r e a d t h of t h e oce l l i ( 1 , 5 : 1 ) ; sc u te l l u m a n d m e t a n o t u m smoo t h , s h i n y , w i t h o u t sc u l p t u re ; p ropodeum ret i c u l a te ru gose, d u l l , w i t h o u t k ee l s ; fore-w i n g h y a l i n e , w i t hou t d a r k

1 1 69

Gon a topod i n ae

t r a n sv ersa l b a n d s ; d i s t a l p a r t o f r a d i a l v e i n l on ge r t h a n p rox i m a l p a r t ( 1 7 : 5 ) ; dorsa I process o f gonoforceps ( F i g . 783 ) l ob e - s h a p ed ; m a x i l l a r y p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 2 seg men t s ; t i b i a l s p u r s 1 , 1 , 2 . LOCUS T Y P I C U S : C a i r n s ( A u s t ra l i a ) TYP I CA L M A T E R I A L : l ec t o t y p e F ! a n d 2 p a r a l e c t o t y pes ( 1 F , 1 M ) ! ( d es i g n a t ed b y O l m i 1 982 ) i n B . HOST S : i n A u s t r a I i a Hadeodelp h a x sp . a n d forms a I I i e d to Lib urn ia ( R . C . L . P erk i n s 1 906c ) . D I ST R I B U T I O N : A u s t ra l i a n reg i o n : AUSTRAL I A : C a i r n s ( Q ueens l a n d ) , B !

3 . Echthrodelp h a x b ifa s c i a t u s R . C . L .

OL !

Perk i n s

Ech throdelp h a x b ifa s c i a t u s R . C . L . Perk i n s 1 905 : 49 . Ech throdelph a x b ifa scia tus R . C . L . P erk i n s : K i effer 1 9 1 4b :

17.

F E M A L E : f u l l y w i n ged ; l e n g t h 2 , 1 9- 2 , 57 m m ; t e s t aceou s , w i t h pe t i o l e b l ac k a n d a b d omen d a r k e n ed ; h e a d sh i n y , w i t h o u t scu l p t u r e , except for occ i p u t a n d f r o n t p a r t o f ver t ex g r a n u l a t ed ; oce l l a r t r i a n g l e s t ron g l y i sosce l e s , w i t h med i a n k ee l ; O L m u c h l on g e r t h a n POL ; p ro n o t u m crossed b y a t r a n sver s a l i m p ress i o n , sh i n y ; co l l a r w i t h o u t scu l p t u re ; d i sc g r a n u l a t e d ; poster i o r edges of p ron o t u m n o t p roduced i n t o l o bes ; sc u t u m sh i n y , w i t h o u t scu l p t u re ; n o t a u l i ces d i s t i n c t a n d com p l e t e , j o i n t a t t h e pos t e r i o r m a r g i n of scu t u m ; s c u te l l u m sh i n y , w i t h o u t sc u l p t u r e ; m e t a n o t u m very reduced and short ; propod e u m ret i c u l a t e ru gose , w i t h s t ro n g t r a n s v e rs a l k ee l s ; pos t e r i o r s u rface o f p ropodeum w i t h o u t l o n g i t u d i n a l k ee l s ; fore w i n g w i t h two b r o w n t ra n s v ersa l b a n d s ; fore t roc h a n ters a p p rox i m a te l y f o u r t i mes a s l on g a s b ro a d ; seg men t 4 of f ro n t t a r s u s a s l o n g as seg m e n t 1 ; en l arged c l a w ( F i g . 784 ) w i t h s u b a p i c a l t o o t h a n d 4 l a m e l l a e ; seg men t 5 o f f r o n t t a r'SUS ( F i g . 784 ) w i t h 1 row o f 9 l a m e l l a e ; a p ex w i t h a g ro u p of 7 l a me l l ae ; m a x i l l a r y p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 2 s e g m e n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k nown L O C U S T Y P I C U S : B u n d a b e r g ( Au s t r a I i a ) TYP I CAL M A T E R I AL : l ec t o t y p e F ! a n d 2 p a r a l ec t o t y p e s F F ! ( d es i g n a t ed b y O l m i 1 982 ) i n B . H O ST S : i n A u s t r a l i a L i b u rn i a sp . a n d a l l i ed forms ( R . C . L . Perk i n s 1 906c ) • D I ST R I B U T I ON : A u s t ra l i a n reg i on : AUSTRAL I A : OL !

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h e a d a l mo s t f l a t , s h i n y , w e a k l y g r a n u l a t ed ; f rons w i t h a med i a n k ee l ; occ i p i t a l c a r i n a i ncom p l e t e , on l y v i s i b l e on t h e s i des of t h e pos t e r i or oce l l i ; POL = 2 ; Ol = 3 ; OOl = 5 , 5 ; p rono t u m crossed by a t ra n svers a l i m p res s i on ; c o l l a r s h i n y , smoo t h , w i t h ou t scu l p t u re ; d i sc d u l l , g r a n u l a te d ; scu t u m , sc u te l l u m a n d m e t a n o t u m s h i n y , smoo t h , w i t hou t

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sc u l p t u re ; n o t a u l i ces com p l et e , poste r i or l y sep a ra t e d , b u t very nea r ; p ropodeu m d u l l , f u l l y re t i c u l a t e ru gos e , w i t ho u t l on g i t u d i n a l o r t r a n s v e r s a l kee l s ; f o r e w i n g s w i t h t w o d a r k t r a n s vers a l b a n d s , one on t he b a s a l c e l l s and a n o t h e r b e ne a t h t he p t e ros t i g m a ; d i s t a l p a r t of r a d i a l v e i n much l on ger t h a n p rox i m a l part ( 9 : 4 , 5 ) ; fore t a rs a l segme n t s i n fo l l o w i n g p ro p o r t i on s : 8 : 1 , 5 : 2 : 8 : 1 2 ; en l a r g e d c l a w ( F i g . 785 ) w i t h a s u b a p i c a l t oo t h a n d 5 l a me l l ae ; segment 5 of f ro n t t a rs u s ( F i g . 785 ) w i t h one row of 6 l a me l l a e ; apex w i t h a g ro u p 0 7 l a me l l ae ; t i b i a l s p u rs 1 ,0, 1 . MALE : u n k no w n L O C U S TYP I C U S : E u r i m b u l a C k . ( A u s t r a l i a ) TYP I CAL M A T E R I A L : h o l o t y pe F ! i n U Q . HOSTS : u n k n o w n D I STR I BU T I ON : A u s t ra l i a n reg i o n : AUSTRAL I A : H i l l Hea d ,

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A crodontochelys C u r r a d o 1 976 :

13.

TYPE S P E C I ES : A crodontochelys b o uceki C u rrado 1 976 , o r i g . d es i g . F E M A L E ( F i g s 798 , 7 99 ) : a p terou s ; m a x i I I a ry p a l p i w i t h 2-4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; p ro n o t u m ( F i g . 799 ) c rossed b y a s t ro n g t ra n s v e r s a l i m p res s i on ; e n l a rged c l a w ( F i g . 7 9 5 ) w i t h s u b a p i c a l toot h , w i t h o u t l a me l l ae , w i t h b r i s t l es o r p e g - I i ke h a i rs ; t i b i a l s p u rs 1 , 0 , 1 . M A L E ( F i g s 789 , 7 90 ) : f u l l y w i n g e d ; m a x i I I a ry p a l p i w i t h 3-4 segmen t s ; l ab i a l p a l p i w i t h 2 segmen t s ; occ i p u t con c a v e ; t em p l es p resen t ; occ i p i t a l c a r i n a i n v i s i b l e o r on l y v i s i b l e beh i n d t h e oce l l i o r o n t he s i des o f t he p os t e r i or oce l l i , b u t never reach i n g t em p l es ; t i b i a l s p u r s 1 , 1 , 2 . D I STR I BU T I ON : w or l d w i de HOSTS : Cicadellida e , exce p t for I diocerinae, Macropsinae, Typh locy b in a e . S P E C I ES : 7 NO T E : t he spec i e s of A crodontochelys are very u n i form : t h e fema l es h a v e h e a d f l a t a n d b o d y mos t l y b l ac k or b row n ; t he m a l es h a v e p ropodeum smoo t h and w i t ho u t s c u l p t u r e . T h e on l y spec ies s l i g h t l y d i ffere n t is the N e a rc t i c A . mimus ( R . C . L . Perk i n s ) , recog n i z a b l e for t he h e a d w e a k l y exc a v a t e d a n d for t h e b o d y redd i sh - t e s t aceou s . T h e o t her spec i es h a v e f em a l es so u n i form t h a t t h ey a re n o t recog n i z a b l e . T h e m a l es , on t he con t r a r y , are e as i l y recog n i za b l e for t he i r d i ffere n t g en i t a l i a . T h e m a l e o f A . mimus i s d i fferent a l so for t h e comp l e te n o t a u l i ces ( i n comp l e t e i n o t her s pec i es ) .

K E Y TO T H E W O R L D S P E C I E S OF A CRODONTOCHE[ YS

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N o t a u I i c e s i n c omp l e t e 2 D o r s a l p r o c e s s o f g on o f o r c ep s d i s t a l l y b r oa d en ed ( F i g . 7 9 1 A ) . . . . . . . uganda n u s ( B en o i t ) ( E t h i op i a n ) D o r s a l p r o c e s s o f g on o f o r c ep s n o t d i s t a l l y b r o a d en e d ( F i g . 787 ) 3 .

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ex t e n d i n g b ey on d

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( F i g . 7 8 7 ) . . . bouceki C u r ra d o ( P a l a ea r c t i c ) D i s t i v o l s e l l a e x t e n d i n g b e y on d t h e a p e x o f . • . . c u bensis ( R i c h a rd s ) ( N e o t r op i c ) .

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T he ma l es of A .

vitiensis,

americ a n u s a n d sinensis a re u n k now n .

PALAEARCT I C S P E C I ES OF A CRODONTOCHE L YS

O n e spec i es i s k n ow n .

A crodontoch elys l:Jouceki C u rrado

A crodontochelys bouceki C u rrado 1 976 :

14.

F E MA L E : a p t e rou s ; l en g t h 2 , 8-4 , 0 mm ; h e a d b l ac k , w i t h m a n d i b l e s , c l y p e u s a n d front p a r t of vertex y e l l ow ; a n te n n a e b l ac k , w i t h segments 1 -3 y e l l o w ; t h o ra x , propodeu m , pet i o l e and a b domen b l ac k , except for apex of propodeu m redd i s h ; l eg s ye l l ow , w i t h coxae a n d c l u b s of femora a l most f u l l y b l ac k ; fore t i b i ae w i t h e x t ern a l s i de b l ack ; m i d a n d h i n d t i b i ae w i t h b o t h ext rem i t i e s b l ac k ; h e a d f l a t , d u l l , g ra n u l a t ed ; fron t a l l i ne comp l ete ; p ro n o t u m crossed b y a s t ro n g t r a n s v e rs a l i m p ress i on , a s scu l p t u red as h e a d , s h i n y ; s c u t u m as scu l p t u red as head ; me t a n o t u m t r a n s verse l y s t r i a t e , as scu I p t u re d a s scu t u m ; s i d e s of m e t a n o t u m rounded ; m e t a t h o r a x + p ropodeum d u l l , a s scu l p t u re d as m e t a n o t u m , w i t h poster i o r s u rface t r a n s v erse l y s t r i a te ; mesop l e u ra a n d metap l e u r a w i t h o u t s t r i ae ; meso-met a p l e u r a l s u t u re ob so l e t e ; s e g m e n t 1 of f r o n t t a r s u s near l y t w i ce a s l on g a s segment 4 ; en l a r g ed c l a w ( F i g . 786 ) w i t h s u b a p i c a l too t h a n d 5-6 peg- l i k e h a i rs ; segment 5 o f fron t t a r s u s ( F i g . 786 ) w i t h 2 rows of 1 1 -22 l a me l l ae ; a pex w i t h a g ro u p of 7-9 l a me l l a e ; m ax i l l a ry p a l p i w i t h 4 seg men t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E : f u l l y w i n ged ; l e n g t h 2 , 7-2 , 8 m m ; b l ac k or b row n , w i t h m a n d i b l es ye l l ow ; l eg s b a n d e d ; a n t e n n a l segment 3 l ess t h a n t h ree a n d a h a l f t i mes a s l on g as b ro a d ( 3 , 33 ) ; head s h i n y , a l u taceo u s , w i t h med i a n kee l between the a n t e n n a I soc k e t s ; temp l es d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e ; s c u t u m s h i n y , a l u t aceou s ; n ot a u l i ces i ncom p l e t e , rea c h i n g a p p rox i m a te l y

Gon a topod i n ae

Fig.

787

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1 1 75

M a l e gen i ta I i a of A crodontochelys b ouceki from La Herr ad u ra ( S p a i n ) ; r i g t h h a l f removed .

0 , 5 l e n g t h of scu t u m ; s c u t e l l um a n d m e t a n o t u m s h i n y , smoo t h , weak I y p u n c t a t e ; p ropodeum smoo t h , s h i n y , not ret i c u l a te ru gose , w i t ho u t scu l p t u r e ; dorsa l s u rface w i t h a med i a n f u r ro w ; fore- w i n g h y a l i ne , w i t h o u t d a r k t ra n svers a l b a n d s ; dors a l p rocess of gonoforcep s ( F i g . 787 ) n o t d i s t a l l y b roadened , cu rved , w i t h a p i c a l t h i rd p a ra l l e l to t he pen i s ; d i s t i v o l se l l a n o t e x t e n d i n g beyon d t h e a pex of the gonoforce p s ; m a x i I I a r y p a l p i w i t h 4 s e g me n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . LOCUS T Y P I C U S : Nerj a ( S p a i n ) TYP I CA L M A T E R I AL : h o l o t y p e F ! ; a l l ot y pe M ! a n d 1 3 p a r a t y pes ( 1 2 F F , 1

Mem . Amer . E n t .

1 1 76

Fig.

788

-

I n s t . , n o . 37 ,

1 984

C he l a of A crodontochelys uga n d a n u s ( Be n o i t ) from Serere ( Ug a n d a )

Gon a topod i n a e

M ) ! i n B M ; 4 p a r a t y p es F F ! i n WA. HOST S : u n k n o w n

i n EN ;

1

1 1 77

paratype F !

i n TO ;

1

paratype F !

D I ST R I BU T I O N : P a l a e a rc t i c reg i o n : SPA I N : Nerj a ( M a l a g a ) , BM ! E N ! B e t e r a ( V a l e n c i a ) , M D ! L a Herr a d u ra ( G r a n a d a ) , B M ! C a l a h o n d a ( G r a n a d a ) , B M ! C a l e l l a d . C o s t a ( B a rce l o n a ) , B M ! Ben i c a s i m ( C a s t e l l on ) , B M ! C YP R U S : Asom a t o s , O L ! CANARY I SLAN D S : S a n Seb a s t i a n ( Gomer a ) , O L ! H E ! B a rra nco de Los J u ncos ( Te n e r i fe ) , M D ! E I Socorro ( G u i m a r , T e n er i fe ) , M D ! E I M e d a n o ( T e n e r i fe ) , T N ! B . co d e L a L e n a ( T acoro n t e , T e n er i f e ) , O L !

F i g . 789

-

( Te n e r i fe ) ,

M a l e o f A crodontochelys uga n d a n u s

TN!

C u m b re F .

Fria

( Be no i t ) from U g a n d a

E T H I O P I A N S P E C I E S O F A CRODONTOCHEL YS

A crodon tochelys uga n da n u s ( Beno i t ) n . com b .

1 1 78

Mem . Amer .

En!.

I ns ! . ,

no . 3 7 ,

1 984

F i g . 790 - M a l e o f A C l'odontochelys uga n da n u s ( Beno i t ) from U g a n d a

Pla tygo n a topus uga n da n u s Beno i t 1 95 1 a : 300 . Pla tygona top u s uga n da n u s Beno i t : Beno i t 1 95 1 d : 1 9 . Rhynchogona top u s u ga n d a n u s ( Ben o i t ) : O l m i a n d C u r r a d o 1 976a : 1 73 .

F E M AL E : a p terou s ; l e n g t h 2 , 50-3 , 43 m m ; h e a d b l ac k , w i t h m a n d i b l es , c I y p e u s a n d f ro n t p a r t o f vertex ye l l o w ; a n te n n ae b r ow n , w i t h s e g me n t s 1 -3 y e l l o w ; h e a d f l a t , s h i n y , g r a n u l a ted ; pron o t u m crossed b y a s t ro n g t ra n sversa l i m p res s i o n , s h i n y , h a i r l es s , g ra n u l a ted ; scu t u m g r a n u l a ted ; m e t a n o t u m g r a n u l a t e d a n d w i t h some t r a n s v e r s a l k ee l s ; m e t a t h o r a x + p ropodeum s h i n y , g r a n u l a ted ; meso-me t a p l e u r a l s u t u re o b so l e t e ; p os t e r i o r s u rface of p ropodeum t r a n s verse l y s t r i a t e ; fore t a r s a l segmen t s i n fo l l ow i n g p roport i on s : 1 5 : 2 : 3 : 7 : 1 2 ; e n l a rged c l a w ( F i g . 788 ) w i t h s u b a p i c a l t oo t h a n d 4-6 peg- l i ke h a i rs ; segmen t 5 of fron t t a rs u s ( F i g . 788 ) w i t h t w o rows o f 1 4-20 l a m e l l ae ; a pex w i t h a g ro u p of 7-9 l am e l l ae ; m a x i I l a r y p a l p i w i t h 2-4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE ( F i g s 789 , 790 ) : f u l l y w i n ge d ; l en g t h 1 , 56-2 , 37 m m ; b l ac k or b ro w n , w i t h m a n d i b l es y e l l ow a n d t a rs i l i g h t ; a n ten n a l segme n t s i n fo l l ow i n g propor t i on s : 5 : 5 : 9 : 9 : 7 : 8 : 8 : 7 ( l a s t t w o s e g m e n t s m i ss i n g i n t h e o n l y seen rna l e s pec i men ) ; a n t en n a l s e g m e n t 3 more t h a n t h ree t i rnes a s l on g as b ro a d ( 4 , 0 ) ; head s h i n y , h a i r y , w e a k l y p u n c t a t e , weak l y a l u t aceo u s , w i t h a rned i a n kee l between t h e a n te n n a l sock e t s ; t e m p l es d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e ; POL � 6 ; Ol � 4 ; OOl � 2 ; b re a d t h of t h e oce l l i : 3 ; s c u t u m h a i ry , a l u t ac eo u s , d u l l ; no t a u l i ces

Gon a topod i n a e

1 1 79

F i g . 79 1 - M a l e of A crodontochelys uga n d a n u s ( Be n o i t ) f rom Serere ( U g a n d a ) ; A : g e n i t a l i a ; B : m a x i l l ary p a l p u s ; C : l a b i a l p a l p u s .

i ncomp l et e , reach i n g a p p rox i m a t e l y 0 , 5 l e n g t h of s c u t u m ; s c u te l l u m a n d m e t a n o t u m s h i n y , h a i ry , as scu l p t u red as scu t u m ; p ropod e u m smoot h , not ret i c u l a t e r u gose , s h i n y , w i t h o u t scu l p t u re , w i t h weak med i a n f u rrow ; fore-w i n g not crossed by d a rk b a n d s ; dors a l p rocess of gonoforceps ( F i g . 791 A ) d i s t a l l y broadened ; m a x i l l a ry p a l p i ( F i g . 7 9 1 B ) w i t h 4 segmen t s ; l a b i a l p a l p i ( F i g . 7 9 1 C ) w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 1 , 2 . LOCUS TYP I C U S : B u l u l o ( U g a n d a ) T Y P I CAL M A T E R I A L : h o l o t y pe F ! i n G E . HOST S :

u n k no w n

Mem . Amer . E n t .

1 1 80

I nst . ,

n o . 37,

1 984

D I ST R I BU T I O N : E t h i o p i a n reg i o n : UGAN D A : Bu l u l o , G E ! K a w a n d a , O l ! BM ! Serere , BM ! Z I MBABW E : S a l i sb u ry , B M ! K E NY A : N a i rob i , T R ! O l g a s a l i c , TV ! SO U T H A F R I C A : K l oof ( N a t a l ) , BM ! BO ! D r a k e n s berg ( V a n Reenen , N a t a l ) , O l ! BM ! H a r r i sm i t h ( O r a n g e S t a t e ) , BM ! Port St . Joh n s ( Po n do l a n d ) , ( Zu l u l a n d ) ,

Ol !

BM ! G i n g i n d h l o v u

( Z u l u l an d ) ,

BM !

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BM !

OR I E N T A L S P E C I ES OF A CRODONTOCHE L YS

O n e spec i es i s k now n .

A crodontochelys sinensis n . sp .

F E MAL E : a p terou s ; l en g t h 2 , 00-3, 1 2 m m ; h e a d b l ack or b row n , w i t h m a n d i b l es , c l y p e u s a n d fron t p a r t of vertex y e l l o w ; t h o r a x , p ropod e u m , p e t i o l e a n d a b d omen b l ack or b row n , e x c e p t f o r a p ex of p ropodeum redd i sh ; l eg s y e l l o w , w i t h coxae a n d c l u b s of femora p a r t l y b l a ck or b row n ; fore t i b i ae w i t h e x tern a l s i d e b l ac k ; m i d and h i n d t i b i ae w i t h b o t h e x t r em i t i es b l a ck ; h e a d f l a t , sh i n y , h a i r l es s , weak l y g r a n u l a t e d ; prono t u m crossed b y a s t ro n g t r a n s ve rsa l i m pres s i o n , s h i n y , h a i r l es s , a s scu l p t u red a s h ea d ; scu t u m s t ron g l y g r a n u l a ted ; meso-me t a p l eu r a l s u t ure o b so l e te ; m e t a n o t u m w i t h s i des rou n d e d , weak l y t r a n s v e rse l y s t r i a t e ; m e t a t h o r a x + p ropodeum a s scu l p t u red a s p ron o t u m , exce p t for dors a l s u rface smooth a n d w e a k l y a l u t a ceou s ; mesop l e u r a , m e t a p l e u r a a n d poster i or s u rface o f p ropodeu m t r a n sverse l y s t r i a te ; fore t a rs a l segmen t s i n fo l l ow i n g p roport i on s : 1 5 : 2 : 3 : 7 : 1 3 ; en l a rged c l a w ( F i g . 792 ) w i t h s u b a p i c a l too t h a n d 3-5 peg- l i k e h a i rs ; seg m e n t 5 of fron t t a r s u s ( F i g . 792 ) w i t h two rows of 1 0- 1 9 l a me l l ae ; a p ex w i t h a g ro u p of 6-9 l a me l l ae ; m a x i l l a r y p a l p i w i t h 3-4 segmen t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l spurs 1 , 0 , 1 . MAL E : u n k no w n lOCUS T Y P I C U S : M a c a u T Y P I CAL M A T E R I AL : h o l o t y p e F ! a n d 2 p a r a t y pes F F ! i n B ; 3 p a r a t y pes F F ! in Ol . H OST S : u n k n o w n D I ST R I BU T I O N : O r i e n t a l reg i o n : CH I NA : NAM : Dj i r i n g , Ol !

Macau ,

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N E A R C T I C SPEC I E S OF A CRODONTOCHEL YS

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1 1 81

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K E Y TO T H E S P E C I E S

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( ty p e )

Mem . A me r .

1 1 82

En!.

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no.

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1 984

FEMALES

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MALES

O n l y t he m a l e o f A .

mimus i s k n o w n .

1 . A crodon tochelys america n us

( F e n ton ) n . com b .

Epigon a top u s america n us F e n ton 1 92 1 : 70 . Pa chygon a top u s nearcticus F e n ton 1 927 : 6 ( n . s y n . ) ; locus typ icus S i ou x C i t y ( I ow a , U. S. A . ) .

F E MA L E :

a p te ro u s ;

l en g t h 2 , 68-3 , 37 m m ; h e a d b ro w n or b l ac k , w i t h a n d f r o n t p a rt o f v e r te x y e l l ow ; a n te n n a e b row n , w i t h segme n t s 1 -3 y e l l o w ; t hora x , p ropode u m , pet i o l e a n d a b domen b l ac k or b rown , except for a pex of p ropodeum red d i s h ; l eg s y e l l ow , w i t h coxae and c l u b s of femora a l mo s t f u l l y b l ac k ; fore t i b i a e w i t h e x t e r n a l s i d e b l ac k ; m i d a n d h i n d t i b i ae w i t h b o t h ex trem i t i es b l ac k ; head f l a t , s h i n y , h a i r l es s , g ra n u l a t e d ; p ronot u m c rossed b y a s t ron g t r a n svers a l i m p re s s i on , s h i n y , h a i r l es s , g r a n u l a ted ; s c u t u m d u l l , g r a n u l a ted ; m e t a n o t u m w i t h s i des ro u n ded , s h i n y , smoo t h , n o t s t r i a t e ; mesometap I e u ra I s u t u re o b so l e t e ; meta t horax + p ropode u m g r a n u I a t e d , excep t for a n te r i o r s u rface smooth a n d w i t hou t sc u l p t u r e ; mesop l e u r a , me t a p l e u r a a n d pos t e r i or s u rface of p ropode u m t r a n sverse l y s t r i a te ; fore t a rs a l segme n t s i n fo l l ow i n g p ropor t i o n s : 1 2 : 2 : 3 : 7 : 8 ; e n l arged c l a w ( F i g . 793 ) w i t h s u b ap i c a l tooth a n d 4-6 p e g - I i k e h a i rs ; segmen t 5 of f ro n t t a r s u s ( F i g . 793 ) w i t h t w o rows o f 6- 1 3 l a me l l ae ; a p e x w i t h a g ro u p of 7-8 l a me l l ae ; m a x i I l a ry p a l p i w i t h 2-3 segme n t s ; l a b i a l p a l p i w i t h 2 seg m e n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n own LOCUS T Y P I C U S : Ames ( I ow a , U. S. A . ) T Y P I CAL M A T E R I AL : h o l o t y p e F ! a n d 4 p a r a t y pes F F ! of E . america n u s i n W A ; h o l o t y p e F ! of P . n earcticus i n WA . HOST S : i n U . S . A . Balc1 u t h a imp icta V a n D . ( T ho m p son 1 954 ) . D I ST R I BU T I O N : N e a rc t i c reg i o n : U . S . A . : W aco ( Mc Len n a n C o . , T e x . ) , ma n d i b l e s ,

c l ypeus

Gon a topod i n a e

F i g . 793

-

C he l a of A crodantachelys america n u s ( Fe n ton ) ( t y p e )

BY ! O L ! W e l der Preserve ( S i n ton , S . P a t r i c i o Ca . , Tex . ) , O L ! Ames ( I ow a ) , WA ! S i a u x C i t y ( I o w a ) , WA ! A l an g S t . Marys ( F l a . ) , WA !

2 . A cradantachelys mimus ( R . C . L . Perk i n s ) n . camb .

1 1 83

Mem . Amer . E n ! .

1 1 84

Fig.

794

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P ro n o t u m ( A )

I n st . ,

no.

37 ,

1 984

a n d h e a d ( B ) of fema l e of A crodontochelys m im u s ( R . C . L . Perk i n s )

Neogon a top u s m im u s R . C . L .

Perk i n s 1 90 7 : 3 3 .

F E MA L E : a p terou s ; l e n g t h 2 , 37-3 , 00 m m ; t e s t aceou s-red d i s h , w i t h a n te n n a e b ro w n i s h ( w i t h seg men t s 1 -3 t e s t aceo u s ) ; pet i o l e b l ac k ; b o d y s h i n y ; head and p ro n o t u m u n sc u l p t u re d ; h e a d ( F i g . 794 B) weak l y e x c a v a t e d ; p ronot u m ( F i g . 794 A ) crossed b y a s t ro n g t ra n svers a l i m p ress i o n ; m e t a t h o r a x + p ropodeum w i t h pos t e r i o r s u rf a c e s t ro n g I y t r a n s verse I y s t r i a t e ; meso-me t a p l e u r a l s u t u re ob so l e t e ; segment 4 of f r o n t t a rs u s s horter t h a n segment 1 ( 1 8 : 23 ) ; e n l a r g e d c l aw ( F i g . 795 ) w i t h s U b a p i c a l tooth a n d 4-5 p e g - I i k e h a i rs ; segmen t 5 o f fron t t a rs u s ( F i g . 795 ) w i t h t w o rows of 1 3 l a me l l ae ; apex w i t h a g ro u p of 9 l ame l l ae ; m a x i l l a r y p a l p i w i t h 3-4 seg men t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u r s 1 ,0, 1 . MALE : f u l l y w i n ged ; l en g t h 2 , 37 m m ; b row n ; l eg s n o t b a n ded ; a n t en n a l segme n t s i n fo l l ow i n g p ro po r t i o n s : 6 : 5 : 9 , 5 : 1 0 : 9 : 9 : 8 : 9 : 7 : 1 0 ; a n ten n a I segment 3 more t h a n t h ree t i mes a s l o n g as b ro a d ( 9 , 5 : 2 , 5 ) ; head s h i n y , weak l y p u n c t a t e , w i t h o u t med i a n kee l b e t ween t h e a n t en n a l socke t s ; temp l es d i s t i nc t ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 7 ; O L = 3 ; OOL = 5 ; scu t u m s h i n y , smoo t h , w i t h o u t scu l p t u re ; n o t a u l i ces com p l e t e , poster i or l y sepa r a t e d ; m i n i m u m d i s t ance bet ween t h e no t a u l i ces n ea r l y as l on g a s b re a d t h of t h e oce l l i ( 2 ) ; s c u t e l l u m a n d m e t a n o t u m s h i n y , n o t ret i c u l a te r u gose , w i t h o u t scu l p t u re ; d o rs a l s u rface w i t h a med i a n f u r row ; fore-w i n g h y a l i n e , w i t h o u t d a rk t ra n s v e rs a l b a n d s ; dors a l p rocess of gonoforceps ( F i g . 7 9 6 ) n o t d i s t a l l y b roadened , c u rved ; b as i vo l se l l a w i t h r i d g e ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 s e g men t s ; t i b i a l s p u rs 1 , 0 , 1 .

1 1 85

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lOCUS T Y P I C U S : Nog a l es ( A r i z . , U . S . A . ) T Y P I CA L MAT E R I A L : l ectotype F ! a n d 3 p a r a l ec to t y pes F F ! ( d es i g n a ted b y O l m i 1 982 ) i n B . HOST S : u n k n o w n D I ST R I BU T I O N : N e a r c t i c reg i on : U . S . A . : Nog a l es ( A r i z . ) , B ! O l !

N E O T R O P I C S P E C I ES O F A CRODONTOCHE L YS

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Cyrtogo n a top u s c u b en s is R i c h a rd s 1 969 : 8 0 .

F E MAL E : a p terou s ; l en g t h 2 , 62-3 , 75 m m ; h e a d b l ac k , w i t h m a n d i b l es , c l y p e u s a n d fron t p a r t o f vertex y e l l ow ; a n t e n n ae b row n , w i t h segmen t s 1 -3 y e l l o w ; t h o r a x , propo d eu m , p e t i o l e a n d a b domen b l ac k , except for apex of p ropode u m redd i s h ; l eg s y e l l ow , w i t h coxae and c l u b s of femora a l most f u l l y b l ac k ; fore t i b i ae w i t h e x t e rn a l s i de b l ac k ; m i d and h i n d t i b i ae w i th b o t h e x t rem i t i es b l ac k ; h e a d f l a t , sh i n y , h a i r l ess , g r a n u l a te d ; p ron o t u m crossed b y a s t ron g t r a n svers a l i m p res s i o n , s h i n y , h a i r l es s , g r a n u l a t e d ; sc u t u m g ra n u l a te d ; m e t a n o t u m w i t h s i de s rou n ded , t r a n sverse l y s t r i a t e ; m e t a t ho r a x + p ropodeum s h i n y , g ra n u l a t e d ; mesop l eu r a a n d m e t a p l e u r a n o t s t r i a te ; pos t e r i or s u rface of p ropodeum t r a n s v e rs e l y s t r i a t e ; meso-me t a p l eu r a l s u t u re o b so l e te ; fore t a rs a l

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1 1 89

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segmen t s i n f o l l ow i n g p ropo r t i on s : 1 3 : 2 : 3 : 6 , 5 : 1 2 ; e n l arged c l a w ( F i g . 797 ) w i t h s u b a p i ca l too t h a n d 5 peg- I i ke h a i rs ; segmen t 5 of f r o n t t a rs u s ( F i g . 797 ) w i t h t w o row s of 1 0- 1 8 l a m e l l a e ; apex w i t h a g r o u p o f 7 l am e l l ae ; m a x i I l a r y p a l p i w i t h 3-4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : f u l l y w i n ged ; l en g t h 2 m m ; b l ac k , w i t h l e g s b a n ded a n d t a r s i l i g h t ; a n t en n a l segmen t s i n fo l l ow i n g propo r t i o n s : 5 : 5 : 8 , 5 : 8 : 8 : 8 : 7 , 5 : 7 , 5 : 7 , 5 : 9 ; a n t en n a l segmen t 3 more t h a n t h ree t i mes as l o n g a s b ro a d ( 3 , 2 1 ) ; h e a d s h i n y , weak l y a l u t aceou s ; t e m p l e s d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e ; scu t u m s h i n y , h a i r y , a l u t aceou s ; n o t a u l i c es i ncomp l e te , reach i n g a p p ro x i m a t e l y O , S I e n g t h of sc u t u m ; s c u t e l l u m a n d m e t a n o t u m s h i n y , a l u t aceo u s ; p ro p o d e u m s moo t h , s h i n y , w i t ho u t sc u l p t u re ; dors a l s u rface w i t h med i a n f u rrow ; fore-w i n g h y a l i n e , w i t h o u t d a r k t r a n svers a l b a n d s ; dors a l p rocess o f g o noforceps n o t d i s t a l l y b roadened , c u rved , w i t h a p i c a l t h i rd p a ra l l e l to t he pen i s ; d i s t i vo l se l l a e x t en d i n g b e y o n d t h e apex of t he gonoforceps ; m a x i I l a r y p a l p i w i t h 4 segmen t s ; w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 .

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Neogona top u s vitien sis R . C . L . Perk i n s 1 906a : 490 . Gon a top u s vitien sis ( R . C . L . P e rk i n s ) : T hom p son 1 95 4 : 2 2 1 .

F E MA L E ( F i g s 798 , 799 ) : a p terou s ; l e n g t h 2 , 00-3 , 2 5 m m ; head b ro w n o r b l ac k , w i t h m a n d i b l e s , c l y p e u s a n d f ro n t p a rt o f v e r tex y e l l o w ; a n t en n ae b ro w n , w i t h s e g m e n t s 1 -3 y e l l o w ; t h ora x , propod eu m , p e t i o l e a n d a b domen b l ac k , except for a pe x of p ropo d e u m red d i s h ; occ a s iona I I y a b domen b row n ; l eg s y e l l o w , w i t h coxae and c l u b s of femora a l most f u l l y b l ac k ; fore t i b i ae w i t h e x t e rn a l s i de b l ac k ; m i d a n d h i n d t i b i ae w i t h b o t h e x t r em i t i es b l ac k ; h e a d f l a t , h a i r l es s , s h i n y , g r a n u l a ted ; p ro n o t u m crossed b y a t r a n s ve rs a l i m p ress i on , g r a n u l a te d , s h i n y ; scu t u m g r a n u l a te d ; m e t a n o t u m w i t h s i des rou n d e d , g r a n u l a t e d , t r a n s v e rs e l y s t r i a t e ; m e t a t horax + p ropode u m g ra n u l a t e d ; mesop l eu r a a n d m e t a p l eu r a not t r a n svers e l y s t r i a t e ; pos t e r i or s u rface of p ropod e u m t r a n s v erse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re o b so l e t e ; fore t a r s a l s e g m e n t s i n fo l l o w i n g propor t i on s : 1 3 : 2 : 3 : 7 : 1 2 ; e n l a r g e d c l a w ( F i g . 800 A ) w i t h s u b ap i c a l tooth a n d 3-7 peg - l i ke h a i rs ; s e g m e n t 5 of f ro n t t a r s u s ( F i g . 800 A ) w i t h t w o rows of 1 3- 2 1 l a me l l ae ; a p e x w i t h a g ro u p of 8- 1 0 l am e l l ae ; m a x i I l a ry p a l p i ( F i g . 800 B ) w i t h 3-4 segmen t s ; l ab i a l p a l p i ( F i g . 800 C ) w i t h 2 s e g m e n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n LOCUS T Y P I CUS : Suva ( F i j i I s l an d s ) T Y P I C A L M A T E R I AL : l ec t o t y p e F ! ( de s i g n a t e d b y O l m i 1 98 2 ) i n B . HOST S : i n F i j i I s l a n d s E u leimonios sp . a n d Deltocep h a l u s sp . ( R . C . L . Perk i n s 1 906c ) . D I ST R I B U T I ON : A u s t ra l i a n reg i on : F I J I I SL A N D S : S u v a ( V i t i L e v u ) , B ! B M ! Pemb u t u , B ! TONGA I SLAN D S : N u k u a l of a ( To n g a t a p u ) , B ! O L ! N e i afu ( V a v a u ) , B! AUST RAL I A : B a l fo u r R a . ( v i a Ben a rk i n , S E Queens l a n d ) , O L ! Y a r ra m a n ( SE Q u e e n s l a n d ) , U Q ! Forest S t a t i on ( B u l b u r i n S t a t e F o re s t , v i a M a n y P ea k s , Q u e e n s l a n d ) , U Q ! B r i sb a ne ( Queen s l a n d ) , U Q !

5 . Pseu dogon a top u s R . C . L .

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Pseu dogon a top u s R . C . L . Perk i n s 1 90 5 : 3 4 . Pa rago n a top u s R . C . L . Perk i n s 1 905 : 40 ( n . s y n . ) ; t y pe s p ec i es Paragon a top u s n igricans R . C . L . Perk i n s 1 90 5 , mono t y p i c . Pse u dogon a top u s R . C . L . P e rk i n s : K i effer 1 906 : 488 . Paragon a top u s R . C . L . Perk i n s : K i effer 1 906 : 488 .

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1 1 92

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Pristogon a top u s K i effer 1 9 1 3c : 325 ( s y n . p roposed b y O l m i a n d C u rrado 1 974 ) ; t y pe s p ec i es Pristogon a topus den tatiforceps ( K i effer 1 904 ) , o r i g . d es i g . P r i s t o go n a t o p u s K i effe r : K i effer 1 9 1 4 b : 7 1 . Pseudogon a top u s R . C . L . P e rk i n s : K i effer 1 9 1 4b : 79 . Paragon a top u s R . C . L . P e rk i n s : K i effer 1 9 1 4b : 87 . Paragon a top u s R . C . L . P e rk i n s : R i c h a r d s 1 939 : 1 95 . Pseu dogon a top u s R . C . L . P e rk i n s : R i c h a r d s 1 939 : 1 95 . Pristogon a top u s K i effer : R i c h a rds 1 939 : 1 95 . Pseu dogon a topoides W i I I i a m s 1 956 : 90 ( n . s y n . ) ; t y pe spec i es Pseudogon a topoides m a uriti a n u s W i l l i a m s 1 956 , o r i g . des i g . Pseu dogon a top u s R . C . L . P e rk i n s : J . F . P e rk i n s 1 976 : 32 . Paragon a top u s R . C . L . P e rk i n s : O l m i 1 979 : 2 1 8 .

T Y P E S P E C I E S : Pse u dogon a top u s kuranda e R . C . L . desi g .

P e rk i n s 1 90 5 , or i g .

F E MA L E ( F i g s 801 , 809 ) : a p terou s ; m a x i l l a r y p a I p i w i t h 2-4 s e g me n t s ; l a b i a l p a l p i w i t h 2 segme n t s ; p rono t u m ( F i g . 8 0 1 ) crossed b y a s t ro n g t r a n s ve rs a l i m p ress i on ; e n l a rged c l a w ( F i g . 803 ) w i t h s u b a p i ca l too t h a n d l a me l l a e ; t i b i a l s p u rs 1 , 0 , 1 . MALE ( F i g s 836 , 837 ) : f u l l y w i n ge d ; m a x i l l a r y p a l p i w i t h 2-4 segmen t s ; l a b i a l p a l p i w i t h 2 s e g m e n t s ; occ i p u t conca v e ; t em p l es p rese n t ; occ i p i t a l c a r i n a i n v i s i b I e ; occ as i o n a I I y o n I y v i s i b I e b e h i n d t h e oce l l i o r o n t he s i des of t h e pos t e r i or oce l l i , b u t never reac h i n g tem p l es ; t i b i a l s p u rs 1 ,1 ,2.

1 1 93

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s eg m e n t 4 . . . . . . . . . . . . . 7 . dromedarius ( A . C o s t a ) A n t e n n a e b r own , w i t h s e gmen t s 1 - 2 a n d 1 0 y e l l ow o r wh i t e ; s eg m e n t 1 o f f r on t t a r s u s n ea r l y a s l on g a s o r l on g e r t h a n s eg m e n t 4 . • • • • • . • . . . • • 8 . a l b osign a tu s ( K i e f f e r ) 9 S c u t u m b l a c k o r b r own . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 0 S c u t u m y e l l ow ; o c ca s i on a l l y w i t h a n t e r i o r s u r f a c e b l a c k a n d p o s t e r i o r s u r f a c e y e l l ow o r r ed d i s h . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 1 0 P r on o t u m w i t h o u t s c u l p t u r e ; a n t e r i o r s u r f a c e o f me t a t h o ra x +

1 1

p r o p od eu m w i t h o u t t ra ck o f med i a n f u r r o w • • • • • . • . • • • • • • • • • • • . . . . . . . . . . . . . . . . . . . . . . 9 . a ug u s t a e C u r ra d o a n d O l m i P r o n o t u m w e a k l y o r s t r o n g l y g ra n u l a t ed ; a n t e r i o r s u r f a c e o f me t a t h o ra x + p r op o d e u m w i t h t ra ck o f med i a n f u r r o w • • • • • . . • • S eg m e n t 1 o f f r on t t a r s u s l on g e r t h a n s eg m e n t 4 ; me t a n o t u m wi t h

s i d e s mo r e r ou n d ed ( F i g . 8 1 6 A ) . . . . . . . . . . • . . • • • • • • • . • • . • • 1 0 . distin c t u s ( K i e f f e r ) S e gmen t o f f r on t t a r s u s s h o r t e r t h a n s eg me n t w i t h s i d e s l e s s r o u n d ed . . . . . . . . . . . . . . . . . . . . ·

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1 3 H ea d w i . . . . . . . H ea d w i i r regu l

th . . th ar

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4 ; me t a n o t u m

. . • . • • 1 1 . priesneri n . s p . l a t e ra l p r o t r u s i o n s r ou n d ed ( F i g . 8 2 0 A ) • • • • . . . . . 1 2 . septemden t a t u s ( J . S a h I b e rg ) l a t e r a l p r o t r u s i on s p o i n t ed ( F i g . 8 2 0 8 ) . . . . . 1 3

v e r t ex s m o o t h a n d w i t h o u t s c u l p t u r e . . . . . . . . . . . . . . . . . . . . . . . . . 8 . a l bosign a tu s ( K i e f f e r ) v e r t e x d u l l , r u g o s e , g ra n U l a t ed , w i t h s h o r t k e e l s . . . . . . 1 3 . ligusticus C u r r a d o a n d O l m i

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MALES

2

D o r sa 808) . D o r sa D o r sa 818) . D o r sa (Fig.

l

p r o c e s s o f g o n o f o r c e p s v e ry s h o r t a n d p o i n t ed ( F i g . • . . • • . • . . . • • • • • 4 . fl a vifumur E s a k i a n d H a s h i mo t o l p r o c e s s o f g on o f o r c ep s v e ry l o n g ( F i g s 8 0 4 , 8 1 8 ) . . . . . . 2 l p r o c e s s o f g o n o f o r c e p s w i t h i n t e rn a l s i d e s e r r a t e ( F i g . . . . . • . . • . • • . . • • . 1 0 . distinc t u s ( K i e f f e r ) l p r o c e s s o f g on o f o r c ep s w i t h i n t e r n a l s i d e n o t s e r ra t e 8 0 4 ) • • • • • • • • • • • • 1 . fu lgori ( N a k a g a wa ) .

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T h e m a l es of t h e o t h e r spec i es a re u n k n o w n .

1 . Pseudogona top u s fulgori ( N a k a g a w a )

n . comb .

Gon a top u s fulgori N a k a g a w a 1 906 : 1 63 . Haplogona top u s fulgori ( N a k a g a w a ) : E s a k i a n d H a s h i m o t a

1 1 95

Gona topo d i n a e

A

B

F i g . 802 - Meso t h o r a x , m e t a t ho r a x a n d p ropodeum of fem a l es of Pse u dogon a top u s foca rilei n . s p . ( A ) a n d fulgori ( N a k a g a w a ) ( B )

1 930 : 2 1 . Paragon a top u s fulgori ( N a k a g a w a ) : Moch i z u k i

1 943 : 9 9 .

F E MA L E ( F i g . 8 0 1 ) : a p t erou s ; l en g t h 3 , 1 2-3 , 37 m m ; h e a d brow n-b l ac k , w i t h fro n t p a r t of v er t e x , c l y p e u s a n d m a n d i b l es y e l l o w ; a n t e n n a e b row n , w i t h s e g m e n t s 1 -2 y e l l o w ; t ho r a x a n d p ropode u m brow n-b l ac k , occa s i o n a l l y w i t h b ro w n -red d i s h n u a nces on p ropode u m ; pet i o l e a n d abdomen b l ac k ; l eg s y e l l o w , w i t h c l u b s o f fore femora a n d fore t i b i ae brow n ; m i d a n d h i n d coxae p a r t l y brown ; h e a d e x c a v a t e d , s h i n y , smoo t h , w i t ho u t scu I p t u r e , exce p t for occ i p u t g r a n u I a t e d ; p rono t u m crossed b y a s t ro n g t r a n s v e rsa l i m press i o n , s h i n y ; co l l a r w i t ho u t scu l p t u r e ; d i s c g r a n u l a t e d ; s i des of p ro n o t u m g r a n u l a ted ; scu t u m d u l l , g r a n u l a ted , w i t h s t ron g t r a n svers a l k ee l s ; scu t u m very l on g , n e a r l y t w i ce or more t h a n t w i ce as l o n g as sc u te l l u m ; m e t a n o t u m w i t h s i de s rou n ded ; m e t a t horax + p ro p o d e u m ( F i g . 802 B ) s h i n y , w e a k l y g r a n u l a t ed ; meso p l e u r a , m e t a p l e u r a a n d pos t er i or s u rface of p ropod e u m t r a n s v e rse l y s t r i a t e ; fore t a rsa l seg m e n t s i n fo l l o w i n g p ropo r t i o n s : 20 : 3 : 6 : 1 6 : 25 ; en l a r g e d c l aw ( F i g . 803 ) w i t h s u b a p i c a l t oo t h a n d 4-5 l a me l l ae ; segme n t 5 o f fro n t t a r s u s ( F i g . 803 ) w i t h t w o row s of 1 0 + 1 2 l a m e l l ae ; a pe x w i t h a g ro u p o f 5 l a m e l l ae ; m a x i I l a ry p a l p i w i t h 2 segmen t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : f u l l y w i n ged ; l en g t h 2 , 1 2-2 , 50 m m ; h e a d b l ac k , w i t h m a n d i b l es y e l l ow ; a n ten n ae b ro w n - y e l l o w ; t ho r a x a n d p ropode u m b l ack ; a b domen b ro w n ; I e g s y e l l o w ; a n te n n a l s e g m e n t 3 four t i mes or more t h a n four t i mes as l o n g a s b roa d ; h e a d s h i n y , h a i ry , g r a n u l a ted , w i t ho u t med i a n k ee l b e t w e e n t h e a n te n n a l soc k e t s ; scu t u m h a i ry , s h i n y , g r a n u l a t ed ; n o t a u l i ces comp l et e , p o s t er i or l y j o i n t ; sc u te l l u m a n d m e t a n o t u m s h i n y , h a i ry ,

weak l y p unctate ;

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dul l ,

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1 1 96

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M e m . A me r . E n ! .

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no. 37,

1 984

803 - C he l a of Pseu dogon a top u s fulgori ( N a k a g a w a ) from K a m i - g u n ( J ap a n )

on pos t e r i or s u rface ; occ i p i t a l c a r i n a i n v i s i b l e ; p rocess o f gonoforceps ( F i g . 804 )

l on g ,

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t em p l es p r om i n en t ;

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w i t h a l atera l poi n t ;

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1 1 97

n

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F i g . 804 - M a l e g en i t a l i a of Pseudogona top u s fulgori K a m i -g u n ( J a p a n )

( N a k a g a w a ) from

m a x i l l a r y p a l p i w i t h 3 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l spurs 1 , 1 , 2 . LOCUS T Y P I C U S : J a p a n T Y P I C A L M A T E R I AL : p rob a b l y l o s t . HOST S : i n J a p a n Soga tella furcifera ( Ho rv a t h ) a n d L a odelp h a x striatellus ( F a l l en ) ( Moch i z u k i 1 943 ) . D I ST R I BU T I O N : P a l a e a rc t i c reg i o n : JAPA N : K a m i -g u n ( Koc h i Pref . ) , OL ! N I ! N a n g o k u ( Koch i P r e f . ) , OL ! N I ! S u g a t a -cho ( M a t s u e C . , Sh i m a n e Pref . ) , O L ! M U ! M a t u z a k i ( F u k u o k a ) ( Moc h i z u k i 1 943 ) , K Y ! O L !

2 . Pseu dogona topus c a melin u s

( K i ef fe r )

Gon a topus camelin u s K i effer 1 904a : 361 . Gon a topus den ta tiforceps K i effer 1 904a : 362 ( s y n . p roposed by O l m i and C u rr a d o 1 974 ) ; locus typ i c u s G i g l i o I s l a n d

1 1 98

Fig.

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n o . 37 ,

1 984

805 - C he l a of Pseudogon a top u s camelinus ( K i effer ) l a P a l m a ( C a n a ry I s l a n d s )

from S . C r u z d e

( I ta l y ) . Pris togon a top u s den t a tiforceps ( K i effer ) : K i effer 1 9 1 3c : 325 .

Gona topo d i n a e

1 1 99

Dicondylys camelin u s ( K i effer ) : K i effer 1 9 1 4 b : 7 5 . Pseu dogona top u s c a melin u s ( K i effer ) : O l m i a n d C u r r a d o 1 974 : 1 8 1 .

F E MA L E : a p t e ro u s ; l en g t h 2 , 68-4 , 00 m m ; testaceou s-red d i s h , w i t h p et i o l e b l ac k ; occas i on a l l y v e r te x , p ron o t u m , m e t a n o t u m a n d m e t a t h o r a x + p ro p o d e u m red d i s h - d a r k ; h e a d exc a v a ted , s h i n y , w i t h o u t scu l p t ure , h a i r l es s ; p ro n o t u m crossed b y a t r a n s v e rs a l i m p re s s i on , s h i n y , h a i r l es s , w e a k l y g r a n u l a ted ; s c u t u m d u l l , g r a n u l a t e d ; s c u te l l u m s h i n y , smoot h , w i t ho u t scu l p t u re ; met a n o t u m v e r y short , g ra n u l a ted , w i t h s i des ro u n de d ; m e t a t h o r a x + p ropode u m sh i n y ; a n t e r i o r s u rface smoo t h , w i t h o u t scu I p t u re , w i t h or w i t h o u t t r a c k of l on g i t u d i n a l f u r ro w ; poster i o r s u rface , mesop l eu ra a n d met a p l e u ra t r a n s v erse l y s t r i a t e ; meso-m e t a p l e u r a l s u t u re obso l e t e or w e a k l y d i s t i n c t ; fore t a r s a l segmen t s i n fo l l o w i n g p roport i on s : 1 4 : 2 : 4 : 1 1 : 1 7 ; e n l a rg e d c l a w ( F i g . 805 ) w i t h s u b a p i c a l too t h a n d 5 l am e l l ae ; s e g m e n t 5 of front t a rs u s ( F i g . 805 ) w i t h t w o row s of 1 5- 1 8 l a m e l l ae ; apex w i t h a g ro u p of 8 l a me l l ae ; occ a s i on a l l y rows o f l ame l l ae exten d i n g co n t i n uo u s l y to t h e a pex ; m a x i l l a ry p a l p i w i t h 4 segmen t s ; l ab i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k n o w n LOCUS TYP I CU S : G i g l i o I s l a nd ( I t a l y ) T Y P I CAL M A T E R I AL : h o l o t y p e F ! of G . camelinus i n G E ; h o l ot y pe F ! of G . denta tiforceps i n G E . HOST S : u n k no w n D I ST R I BU T I O N : P a l a e a rc t i c reg i o n : S PA I N : G i b ra l t a r , WA ! A l m u ne c a r ( G r a n a d a ) , OL ! OT ! I TALY : G i g l i o I s l a n d ( Grosseto ) , G E ! Sc i c l i ( R a g u s a ) , P L ! CANARY I SLAN D S : S . C r u z de l a P a l m a ( L a P a l m a ) , H E ! O L !

3 . Pseu dogona topus rosell a e C u rrado a n d O l m i

Pseu dogon a top u s rosellae C u rrado a n d O l m i

1 974a : 234 .

F E M A L E : a p t e ro u s ; l e n g t h 3 , 48-3 , 80 m m ; b l ac k , w i t h fro n t p a r t of v ertex , c l y p e u s a n d m a n d i b l es y e l l o w ; a n t en n ae b l ac k , w i t h segmen t s 1 -3 y e l l o w ; l eg s y e l l o w , w i t h c l u b s of femora p a r t l y b row n ; m i d a n d h i n d coxae p a r t l y b row n ; h e a d exca v a ted , s h i n y , g r a n u l a ted ; p ron o t u m crossed b y a s t ro n g t r a n s v e rs a l i m p ress i o n , s h i n y , g r a n u l a ted ; s c u t u m d u l l , g ra n u l a ted ; m e t a n o t u m v e r y short , n ea r l y as l on g a s scu te l l u m , t ra n s verse l y s t r i a t e ; m e t a t horax + p ropodeu m d u l l , g ra n u l a ted ; poste r i o r s u rface of propo d e u m t r a n s v e rse l y s t r i a t e ; a n t e r i o r s u rface w i t h m ed i a n f u r ro w ; p l e u r a not t r a n s v erse l y s t r i a t e ; fore t a rs a l s e g m e n t s i n fo l l o w i n g propor t i o n s : 20 : 4 : 5 , 5 : 1 3 : 20 ; e n l arged c l a w ( F i g . 806 ) w i t h s u b a p i c a l t oo t h a n d 4-6 l a me l l ae ; s e g m e n t 5 o f fro n t t a rs u s ( F i g . 806 ) w i t h 2-3

1 200

Mem . A m e r . E n ! .

I n s! . ,

no.

37 ,

1 984

F i g . 806 - C he l a of Pse u dogon a top u s rosel l a e C u rrado a n d O l m i

( paratype )

rows of 23-29 l a me l l a e ; a p e x w i t h a g rou p of 5-8 l a m e l l a e ; m a x i I l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k no w n

Gon a topod i n a e

F i g . 807

-

1 20 1

C he l a o f Pseu dogona top u s fla vifemur E s a k i a n d H as h i moto f ro m K a m i - g u n ( Ja p a n )

Mem . A me r .

1 202

Ent.

I n st . ,

no .

37 ,

1 9 84

L O C U S T YP I CU S : P i a n d e l l ' A l p e ( I t a l y ) TYP I C A L M A T E R I A L : ho l o t y pe F ' a n d 2 p a r a t y p es F F ! i n E N ; 2 p a r a t y pe s F F ! i n OL . HOSTS : i n I t a I y Megadelp h a x sordidulus ( St a I ) , Dicra notropis h a m a ta ( Bo h . ) a n d divergens K i r s c h b a u m ( C u r rado a n d O l m i 1 974a ) . D I ST R I BU T I ON : P a l a e a r c t i c reg i o n : I TALY : P i a n de l l ' A l p e ( F e n e s t re l l e , Tori no) ,

EN! OL!

4 . Pseu dogon a top u s fl a vifemur E s a k i a n d H a s h i moto

Pseu dogon a topus Pseu dogon a top u s H as h i mo t o 1 933 : Pseu dogon a top u s H as h i mo t o 1 936 : Pseu dogo n a top u s Moch i z u k i 1 94 1 :

fla vifemur fl a v ifemur 20. fla vifemur 20. fla vifemur 26 .

Esak i

a n d H a s h i moto 1 9 3 2 : 28 . a n d H a s h i mo t o : E s a k i a n d

Esak i

a n d H a s h i moto : E s a k i

and

E sa k i

a n d H a s h i moto :

Esak i

and

Esak i

F E MAL E : a p t e ro u s ; l en g t h 3 , 1 2- 3 , 75 m m ; head b row n , w i t h occ i p u t , c l y p eu s , f ro n t p a r t o f v e r t e x a n d m a n d i b l es y e l l ow ; a n t e n n a e b row n , w i t h seg m e n t 1 y e l l ow ; t h o ra x , p ropodeu m , p e t i o l e a n d abdomen b l ac k ; l eg s y e l l o w , w i t h c l u bs of fore femora b ro w n ; h e a d exc a v a ted , s h i n y , g r a n u l a t e d , w i t h l on g i t u d i n a l k ee l s o n v e r t e x ; p rono t u m c rossed b y a s t rong t r a n sv e rs a l i mp ress i o n , s h i n y , g r a n u l a t e d ; scu t u m s h i n y , g r a n u l a ted ; sc u t e l l u m s h i n y ; m e t a n o t u m n e a r l y a s l o n g a s sc u te l l u m , t r a n s v e rse l y s t r i a t e , w i t h s i des rou n d e d ; m e t a t horax + p ropodeum d u l l , g r a n u l a t e d ; mesop l e u r a , m e t a p l e u ra a n d p o s t e r i o r s u rface of p ropodeum t r a n s verse l y s t r i a t e ; fore t a r s a l seg m e n t s i n fo l l ow i n g p ropo r t i o n s : 2 1 : 3 , 5 : 6 : 1 7 : 2 5 ; en l a r g e d c l a w ( F i g . 807 ) w i t h s u b a p i c a l too t h a n d 5 l am e l l a e ; s e g m e n t 5 of f r o n t t a r s u s ( F i g . 8 0 7 ) w i t h 2 ro w s of 1 3 l a m e l l ae ; a pex w i t h a g r o u p of 7 l a me l l ae ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l sp u rs 1 , 0 , 1 . M AL E : f u l l y w i n ge d ; l e n g t h 2 , 56-3 , 25 m m ; h e a d b l ac k ; a b domen b ro w n b l ac k ; l eg s y e l l ow ; a n t en n a l segmen t 3 f o u r o r m o r e t h a n four t i mes a s l o n g a s b ro a d ; h e a d d u l l , h a i r y , g r a n u l a ted ; temp l e s p rom i n en t ; occ i p i t a l c a r i n a i n v i s i b l e ; f ro n s w i t hou t med i a n kee l b e t ween t h e a n t en n a l soc k e t s ; sc u t u m d u l l , g r a n u l a ted , h a i ry ; n o t a u l i ces comp l e t e , pos t e r i o r l y j o i n t ; s c u t e l l u m a n d m e t a n o t u m s h i n y , we a k l y p u n c t a t e ; p ropodeum ret i c u l a t e r u gose , w i t h o u t kee l s on poste r i or s u rf ace ; dors a l p rocess of gonoforceps ( F i g . 808 ) sho r t , po i n t ed ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l ab i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u r s 1 , 1 , 2 . LOCUS T Y P I C U S : J a p a n T Y P I CA L MAT E R I AL : probab l y l o s t HOSTS :

i n J a p a n Nil ap a rvata lugens ( S t a l ) ( � oryzae M a t s . ) ( E s a k i a n d

G o n a topod i n ae

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H a s h i moto 1 933 ; G r i s t a n d Lever 1 969 ) . D I ST R I B U T I O N : P a l a e a rc t i c reg i o n : JAPAN : K a rn i - g u n ( K oc h i P ref . ) , OL ! N I ! N a n go k u ( K oc h i P ref . ) , N I ! M U ! F u k uok a , KY ! O r i en t a l reg i o n : the reco rds from P h i I i p p i nes ( C h a n d r a 1 978 a ; C h u a a n d D y c k 1 982 ; I . R . R . I . E n to mo l og y D e p a r t m e n t 1 980 , 1 98 1 ) a n d T a i w a n ( C h u a n d H i r a s h i m a 1 98 1 ) m u s t b e c h ecked .

5 . Pseudogona top u s orth ola b is ( K i effer )

Gon a topus orth olabis K i effer i n K i effer et M a r s h a l l 1 906 : 506 . Dicondyl u s ortb olabis ( K i effer ) : K i effer 1 9 1 4b : 7 4 . Dicondylus ortbolabis ( K i effer ) : Ber l a n d 1 9 28 : 1 46 .

1 204

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1 206

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a n d C u r r a d o 1 974 : ( I ta l y ) .

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F E M A L E ( F i g . 809 ) : a p t e rou s ; l en g t h 3 , 00-4 , 40 m m ; h e a d t e s t aceo u s or b l ac k , w i t h m a n d i b l es , c l y pe u s and f ro n t part of vertex y e l l o w ; a n t en n a e b ro w n - b l ac k , w i t h segm e n t s 1 -2 a n d p a r t l y 3 y e l l o w ; p ro p ec t u s b l ack o r redd i s h ; p,rono t u m b l ac k ; scu t u m b ro w n -redd i s h or ye l l o w , w i t h a n t e r i or s u rface b l ac k ; s c u t e l l u m a n d m e t a t ho r a x + p ropodeum b l ac k , w i t h a p e x of p ropode u m ye l l o w ; p e t i o l e b l ac k ; a b domen b ro w n - b l ac k ; l eg s y e l l o w , w i t h coxae , c l u b s of femora a n d t i b i a e p a r t l y b r ow n ; head excav a ted , s h i n y or d u l l , g r a n u l a t e d ; p ro n o t u m crossed by a t ra n s v e rs a l i m p res s i o n , s h i n y , g r a n u l a t e d ; scu t u m d u l l , g ra n u l a t e d ; sc u t e l l u m s h i n y , w i t h o u t scu l p t u r e , smoo t h ; m e t a n o t u m g r a n u l a t e d , s h i n y or d u l l , w i t h some k ee l s a n d w i t h s i des r o u n d e d ; a n te r i or s u rface o f p ropodeum s h i n y o r d u l l , smoo t h , w i t h o u t sc u l p t u re or g r a n u l a t e d , w i t h t rack of med i a n f u r ro w ; mesop l eu r a , m e t a p l e u r a a n d p os t e r i o r s u rface of p ro p o d e u m t r a n s verse l y s t r i a t e ; p o s te r i o r s u rf a c e o f p ro p o d e u m g r a n u I a ted a m o n g s t r i ae ; meso m e t a p l eu r a l s u t u re o b so l e t e ; e n l arged c l a w ( F i g . 8 1 0 ) w i t h s u b a p i c a l t o o t h a n d 3-6 l a me l l ae ; segmen t 5 o f f ro n t t a rs u s ( F i g . 8 1 0 ) w i t h 2 rows of 20-29 l a me l l ae ; a p e x w i t h a group of 5-9 l am e l l ae ; m a x i l l a ry p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k no w n L O C U S TYP I C U S : Ucc i a n i ( Corse , F ra n ce ) T Y P I CA L MAT E R I AL : ho l o t y pe F ' of G. orth ola b is i n B U ; ho l o t y p e F ! of

Ps . igilii in G E . HOST S : u n k n o w n D I ST R I BU T I ON : P a l aearc t i c reg i o n : F R A N C E : Ucc i a n i ( Corse ) , BU ! I TALY : G i g l i o I s l a n d ( G rosseto ) , G E ! OL ! Sc i c l i ( R a g u s a ) , P L ' EGYPT : F a y u m , WA !

6 . Pseu dogon a top u s foc arilei n .

sp .

F E M AL E : a p t e ro u s ; l en g t h 4 , 8 1 m m ; h e a d y e l l ow-red d i s h , w i t h b r o w n s p o t on v e r t e x ; a n t e n n ae b row n , w i t h segmen t s 1 -2 a n d 1 0 y e l l ow ; p ro p ec t u s redd i s h ; p ro n o t u m redd i s h- t e s t ac eo u s ; scu t u m ye l l o w , w i t h a n a r ro w a n t er i or b ro w n s t r i p e ; sc u t e l l u m t e s t a c eo u s ; m e t a n o t u m a n d m e t a t ho r a x + p ropodeum red d i s h - t e s t aceou s ; p et i o l e b l ac k ; a b domen redd i s h ; l eg s red d i s h - t es t ac eo u s , w i t h c o x a e a n d c l u b s of femora p a r t l y b r ow n ; h e a d e xc a v a t e d , d u l l , g r a n u l a t e d , w i t h s o m e l o n g i t u d i n a l k ee l s on v e r t e x ; p ron o t u m s h i n y , c rossed by a s t ro n g t ra n s v ersa l i m p ress i o n , w i t ho u t scu l p t u r e ; sc u t u m s h i n y , a l u t aceo u s , w i t h some l on g i t u d i n a l k ee l s ;

1 207

Gon a t opod i n ae

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1 209

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2 s e g m e n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n known lOCUS TYP I C U S : Ucce l l i n a N a t . P a r k ( I t a l y ) TYP I CAL M A T E R I AL : h o l o t y pe F ! i n Ol . HOS T S : u n k no w n D I ST R I B U T I ON : P a l aearc t i c reg i o n : I TALY : Ucce l l i n a N a t . Park ( G ros seto ) , Ol ! NOT E : t he spec i es i s n a med i n honor of M r . A l e s s a n dro Foc a r i I e .

7 . Pseudogona top u s dromedarius ( A . C o s t a )

Dicondylus dromedarius A . Cos t a 1 882 : 38 . Gon a topus dromeda rius ( A . Cos t a ) : D a l l a Torre 1 898 : 545 . Gon a top u s dromedarius ( A . Cos t a ) : K i effer i n K i effer e t M a r s h a l l 1 90 5 : 9 7 . Gon a top u s dromedarius ( A . Cos t a ) : K i effer 1 9 1 4b : 1 09 . Pseudogona top u s dromedari u s ( A . Cos t a ) : C u rrado a n d O l m i 1 974a : 227 .

F E M A L E : a p t e rou s ; l e n g t h 2 , 88 m m ; y e l l ow - t e s t aceo u s , w i t h p e t i o l e b l ack ; body s h i n y , smoo t h , h a i r l es s , w i t ho u t scu l p t u re ; head exca v a t e d ; prono t u m crossed b y a s t ro n g t r a n sversa I i m p ress i on ; m e t a no t u m w i t h some t r a n s vers a l k ee l s and w i t h s i des p ro t r u d i n g ; p rot r u s i o n s po i n t ed ; meso met a p l e u r a l s u t u re d i s t i n c t a n d comp l e t e ; mesop l eu r a , m e t a p l e u r a a n d pos t e r i o r s u rface o f p ropodeum t ra n s verse l y s t r i a t e ; s e g m e n t 1 o f fro n t t a rs u s s l i g h t l y shorter t h a n seg m e n t 4 ( 8 : 1 0 ) ; en l arged c l a w ( F i g . 8 1 2 ) w i t h s u b a p i c a l t oo t h a n d 7 l a m e l l a e ; seg m e n t 5 of f ron t t a r s u s ( F i g . 8 1 2 ) w i t h 2 rows of 6 + 1 0 l a me l l a e ; a p e x w i t h a g ro u p of 20 l a me l l ae ; m a x i l l a ry p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n known L O C U S TYP I C U S : A l g h ero ( I t a l y ) TYP I CA L MAT E R I A L : h o l o t y p e F l os t ( or i g i n a l l y i n NA ) ; ( d es i g n a ted b y C u rrado a n d O l m i 1 974a ) i n E N . HOST S : u n k n o w n D I ST R I BU T I O N : P a l a e a rc t i c reg i on :

8.

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EN !

Pseu dogon a top u s a l bosign a tus ( K i effe r )

Gon a top us a l b osign a tu s K i effer 1 904a : 358 . Pseu dogon a top u s s p . R i c h a rd s 1 972 : 543 . Pse u dogona top u s a l b osign a tu s ( K i effer ) : O l m i a n d C u rrado 1 974 :

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1 86 . Pseu dogona topus separa t u s R i c h a r d s partim : A b d u I -N o u r 1 97 6 : 274 .

F E M A L E : a p tero u s ; l en g t h 2 , 73-3 , 75 m m ; h e a d t e s t aceou s-red d i s h or red d i s h , occ a s i o n a l l y p a r t l y b l ack ; a n t e n n a e b row n , w i t h s e g me n t s 1 -2 a n d 1 0 w h i t e or y e l l ow ; t h orax a n d propo d e u m redd i s h or b row n , w i t h s c u t e l l u m a n d a p e x of p ropod e u m y e l l ow ; p e t i o l e b l ac k ; a b domen b ro w n o r b l ac k , w i t h terg i t e 1 red d i s h ; l e g s y e l l o w , w i t h f e m o r a b row n i s h ; h e a d e xc a v a ted , s h i n y , smoo t h , w i t h o u t scu l p t u re ; p ro n o t u m crossed b y a s t ro n g t r a n s v e rs a l i m press i o n , s h i n y , smoot h , w i t h o u t scu l p t u re ; scu t u m ru gose , d u l l , w i t h some l o n g i t u d i n a l k ee l s ; scu te l l u m smoo t h , s h i n y ; m e t a n o t u m t ra n sverse I y s t r i a t e , w i t h s i des p ro t r u d i n g ; p ro t r u s i o n s p o i n t ed ( formed by a t r a n s vers a l kee l on mesop l e u r a ) ; m e sop l e u ra , m e t a p l eu r a a n d pos t e r i o r s u rf ace o f propo d e u m t r a n s v e rse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re d i s t i n c t a n d comp l e t e ; a n t e r i o r s u rf ace of m e t a t horax + p ropode u m s h i n y , smoo t h , w i t h t r a c k of m ed i a n f u r row ; e n l a r g e d c l a w ( F i g . 8 1 3 ) w i t h s U b a p i c a l t oo t h a n d 7-8 l a m e l l ae ; s e g m e n t 5 o f f r o n t t a r s u s ( F i g . 8 1 3 ) w i t h 2 rows o f 1 2- 1 6 l a m e l l a e ; a p e x w i t h a g ro u p of 8 - 1 1 l a me l l ae ; m a x i l l a r y p a l p i w i t h 3-4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , l . MALE : u n k n own LOCUS TYP I C U S : G i g l i o I s l a n d ( I t a l y ) TYP I CA L MAT E R I A L : ho l o t y pe F ! i n GE . HOSTS : i n F r a nce Rib a u todelph a x imitans R i b . ( A b d u I -Nour 1 97 6 u n der n a me Ps e udogon a topus separatus R i c h a rd s partim ) ; i n I srael p e r h a p s L a odelp h a x stria tellus ( F a l l e n ) ( R i c h a rds 1 972 ) . D I ST R I BU T I O N : P a l a e a rc t i c reg i o n : F RA N C E : S t . -Ge l y - d u -Fesc ( He r a u l t ) , P ' S PA I N : A l m u n ecar ( G r a n a d a ) , O T ! I TA L Y : G i g l i o I s l a n d ( G ros seto ) , G E ! I SR AE L : Rehovo t , GV ' CANARY I SLANDS : T aod i o ( T e n e r i fe ) , M D ! E G Y P T : F a y e d , WA r O L !

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F E M A L E : a p terou s ; l en g t h 2 , 60 m m ; b l ack ; d i s t a l a p e x of cox ae , m i d a n d h i n d t r oc h a n t e r s , s t a l k of femor a , p a r t o f t i b i ae a n d t a r s i y e l l o w ; b o d y s h i n y , h a i r l es s , w i t ho u t s c u l p t u re ; h e a d e x c a v a t e d ; pron o t u m ( F i g . 8 1 4 ) crossed b y a s t ro n g t r a n s v e rs a l i m p ress i o n ; s c u t u m w i t h some l o n g i t u d i n a l k ee l s ; m e t a n o t u m w i t h s i des p ro t ru d i n g ; p r o t ru s i o n s p o i n ted ; meso m e t a p l eu r a l s u t u re d i s t i n c t ; mesop l e u r a , m e t a p l e u r a a n d pos t e r i o r s u rface of propodeum t r a n s v erse l y s t r i a t e ; fore t a r s a l segmen t s in fo l l o w i n g propor t i o n s : 1 3 : 2 , 5 : 3 , 5 : 1 0 : 1 6 ; en l a rged c l a w ( F i g . 8 1 5 ) w i t h s u b a p i c a l toot h a n d 7-8 l am e l l ae ; seg m e n t 5 o f fron t t a r s u s ( F i g . 8 1 5 ) w i t h 2 rows of 1 1 l a me l l a e ; a pex w i t h a g ro u p of 7-8 l a m e l l ae ; m a x i l l a r y p a l p i w i t h 4 s e g m e n t s ; l a b i a l p a l p i w i t h 2 s e g m e n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u nknown L O C U S T Y P I C U S : Port aco m a r o ( I t a l y ) TYP I CAL M A T E R I AL : ho l o t y pe F ! i n E N . HOSTS : i n I t a l y Megadelp h a x sp . ( C u rr a d o a n d O l m i 1 974a ) . D I STR I BU T I O N : P a l aearc t i c reg i o n : I TA L Y : Port acomaro ( A s t i ) , E N !

1 0 . Pseudogo n a top u s distin c tu s ( K i effe r )

Gona topod i n ae

1213

F i g . 8 1 6 - F em a l e of Pseu dogon a top us dis tin c t u s ( K i effer ) : p r o t h o r a x a n d p ropodeum ( A , B ) ; h e a d ( C ) .

Gona top us distin c t u s K i effer i n K i effer et M a rs h a I I 1 906 : 509 . Dicon dylus distin ctus ( K i effe r ) : K i effer 1 9 1 4b : 75 . Dicondy I u s rob u s t u s C e b a I l os 1 927 : 1 05 ( n . s y n . ) ; locus typicus C e rc ed i l l a ( Spa i n ) . Pseu dogon a top u s distin c t u s ( K i effer ) : R i c h a r d s 1 939 : 205 . Pseu dogon a top u s distin c t u s ( K i effer ) : Beh r i sc h 1 95 2 : 33 . Pseu dogon a top u s dis tinctus ( K i effer ) : N a g y 1 967a : 1 24 . Pseudogo n a top u s distinctus ( K i effe r ) : C u rrado a n d D l m i 1 974a : 224 . Pseu dogon a top u s distin ctus ( K i effer ) : W a l off 1 974 : 1 0 3 . Pseu dogona topus distin c t u s ( K i effer ) : W a l off 1 975 : 664 . Pseu dogon a top u s distinctus ( K i effer ) : J . F . P e r k i n s 1 97 6 : 32 . nec Pseu dogon a topus distin c t u s ( K i effe r ) : P o n o m a r e n k o 1 978 : 23 . Pseu dogon a top u s dis tin c t u s ( K i effer ) : Moczar 1 083b : 207 .

F E M AL E : a p t e ro u s ; l e n g t h 3 , 5-4 , 0 m m ; h e a d b l ac k , w i t h m a n d i b l es , c l y pe u s a n d fron t p a r t o f v e r t e x y e l l ow ; a n t e n n a e b l ac k , w i t h segmen t s 1 -2 y e l l o w ; t hora x , p ro po d e u m , p et i o l e a n d a b do m e n b l ac k ; l eg s y e l l o w , w i t h c l u b s of fore femora b row n i s h ; h e a d ( F i g . 8 1 6 C ) excav a t e d , s h i n y , g r a n u l a t ed ; p r o no t u m ( F i g . 8 1 6 A , B ) crossed b y a s t ro n g t r a n svers a l

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i m p res s i on , h a i r l es s , w e a k l y a l u t a c eo u s ; scu t u m d u l l , r u gose ; met a n o t u m t r a n s verse l y s t r i a t e , w i t h s i des p r o t r u d i n g ; p r o t r u s i o n s p o i n te d ; meso met a p l e u r a l s u t u re d i s t i n c t and comp l e t e ; mesop l e u r a , me t a p l eu r a a n d pos t e r i o r s u rface o f p ropodeum t ra n s v e rse l y s t r i a t e ; a n t e r i o r s u rface of p ro p o d e u m w e a k l y r u g ose , w i t h t rack of med i a n fu rrow ; fore t a rsa l se g m e n t s i n fo l l ow i n g p ro p o r t i o n s : 1 5 : 2 : 4 : 1 1 : 1 8 ; en l a r g e d c l a w ( F i g . 8 1 7 ) w i t h s u b a p i c a l t oo t h a n d 5-9 l a m e l l ae ; seg m e n t 5 of fron t t a rs u s ( F i g . 8 1 7 ) w i t h 2 r o w s o f 2 1 -27 l a m e l l ae ; a p e x w i t h a g r o u p of 5- 1 0 l a me l l a e ; m a x i I l a ry p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : f u l l y w i n g ed ; l en g t h 2 , 5 m m ; head b l ac k , w i t h m a n d i b l e s redd i s h ; a n te n n a e b row n ; t ho r a x a n d p ropodeum b l ac k ; a b domen b ro w n - r e d d i s h ; l e g s b row n , w i t h t a r s i l i g h t ; a n t e n n a I segmen t 3 fo u r t i mes or more t h a n four t i mes as l o n g as b ro a d ; h e a d s h i n y , g r a n u l a t ed , w i t h o u t med i a n kee l b e t ween a n t e n n a l socke t s ; t em p l es p r om i n en t ; occ i p i t a l c a r i n a i n v i s i b l e ; s c u t u m s h i n y , g r a n u l a ted ; n o t a u l i ces comp l e t e , p o s t e r i o r l y sep a ra t e d ; m i n i m u m d i s t a n c e b e t w een t h e n o t a u l i ces s h o r t e r t h a n b re a d t h o f oce l l i ; scu t e l l u m a n d m e t a n o t u m s h i n y , smoo t h , w e a k l y p u n c t a t e ; p ropodeum ret i c u l a t e r u g o s e , sh i n y , w i t h o u t l on g i t L1 d i n a i k ee l s o n pos t e r i o r s u rface ; dors a l p rocess of gonoforceps ( F i g . 8 1 8 ) l o n g , w i t h i n n e r s i d e serr a t e ; m a x i I l a r y p a l p i w i t h 4 seg men t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . L O C U S TYP I C U S : N e w F o r e s t ( E n g l a n d ) TYP I CAL MAT E R I A L : h o l o t y p e F ! o f C . distin ctus i n BU ; ho l o t y p e F ! o f D . rob u s t u s i n M D . HOST S : i n E n g l a n d Dicra n o tropis h a m a ta ( Bo h . ) , Gra vestein iella boldi ( Sco t t ) , Ditropis pteridis S p i no l a , Hyledelp h a x elega n t u l u s ( Boh . ) ( � L a odelp h a x elega n t u l u s ) , la vesella pellucida (F . ) , la vesella discolor ( Boh . ) , /{ossl"igi anella exigua ( Boh . ) ( � N u irodelp h a x exigu u s ) , Rib a u todelp h a x a ngulosus ( R i b a u t ) , Para lib urn ia dalei ( Scot t ) ( R i c h a rds 1 9 39 , 1 948 ; W a l off 1 974 ) ; in G e r m a n y Dicra n o tropis h a m a t a ( Boh . ) ( S t ru b i n g 1 956 ) ; i n F i n l a n d la vesella pellu c ida ( F . ) ( He i k i n h e i mo 1 9 5 7 : t h e a l l ot y pe M ! of Dicondylus lindbergi H e i k . i s re a l l y a m a l e of Pseu dogon a topus distin ctus ) . D I ST R I BU T I O N : P a l a e a rc t i c reg i o n : M O N GO L I A : 1 1 Km S from P a s s Zos i j n d a v a a ( 90 K m S f rom U l a n - B a a t o r , Ce n t r a l a i m a k ) , B U ' ( t h e o n l y seen s p ec i men i s a m a l e : i t has n o t a u l i c e s s l i g h t l Y d i ffere n t from t y p i c a l m a l es o f Ps . distin c t u s ; t he m i n i m u m d i s t a n ce b e tween t h e n o t a u l i ces i n f a c t i s n e a r l y a s l o n g a s b re a d t h o f oce l l i ; v a r i ab i l i t y o r a n o t h e r s p ec i es ? ) ; F I N L AN D : H a n k o ( U u de n m a ) , H E ! L a i h i a ( V a a s a n ) ( a l l o t y pe M ! of Dicondylus lindbergi He i k i n h e i mo ) , HE ! S W E D E N ( He l l e n 1 953 ) ; R O M M� I A UJ a g y 1 967a ) ; G E R MANY ( S t r u b i n g 1 956 ) ; B E L G I U M ( P i c a r d 1 93 2 ) ; F R AN C E : Fon t a i n e b l ea u ( Se i n e - e t - M a r n e ) , O L I E N G LA N D : New Forest ( H a n t s So u t h ) , BU ! S i l wood P a rk ( Be r k s h i re ) , B M ! L i c h f i e l d D i s t r . ( C a r r 1 93 1 ) ; n u merous l oc a l i t i es l i s t ed by R i c h a r d s 1 93 9 and J . F . P e r k i n s 1 976 from E n g l a n d a n d W i g h t I s l a n d ; S PA I N : Cerced i l l a ( M ad r i d ) , M D ! M a d r i d , P ! I TA L Y : Pecc i a ( R i v a V a l do b b i a , Vercel l i ) , O L '

Mem . A me r . E n ! .

1 21 6

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n o . 37 ,

1 984

M a l e g en i t a l i a of Pseu dogon a top u s distinctus L a i h i a ( F i n l a n d ) ( r i g h t h a l f remo ved )

( K i effer ) from

N OT E : accord i n g to Ponomarenko ( 1 978 ) Ps . distin c t u s occurs i n U . S . S . R . ( C r i mea ) ; I h a ve s t u d i e d t h e spec i me n s seen b y M r s . Pono m a ren k o : re a l l y t he y b e l o n g t o a n o t h e r s p ec i e s , Ps . sep temden t a t u s ( J . Sa h l b e rg ) .

1 1 . Pseudogona top u s priesneri n . sp .

Gon a topod i n a e

1217

F E MA L E : a p t e r ou s ; l e n g t h 2 , 4- 3 , 2 mm ; b ro w n or b ro w n-red d i s h , w i t h t a r s i a n d a n t e n n a I s e g me n t 1 t e s t a c eo u s ; a n t e n n ae d i s t a l l y t h i c k e n e d ; a n t en n a l s e g me n t s i n fo l l o w i n g proport i o n s : 1 0 : 6 : 1 0 : 7 : 6 : 6 : 5 , 5 : 5 , 5 : 5 : 1 0 ; h e a d exc a v a te d , d u l l , we a k l y g r a n u l a ted ; fro n s w i t h a com p l e t e med i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 2 ; O L = 3 ; DOL = 1 0 ; t e m p l e s p r o t r u d i n g ; p ro no t u m crossed b y a s t ro n g t ra n svers a l i m p res s i o n , d u l l , w e a k l y g r a n u l a t e d ; s c u t u m d u l l , g r a n u l a t e d ; met a n o t u m w i t h n u merous t ra n s v e rs a l k ee l s , h o l l o w b eh i n d t h e sc u te l l u m , w i t h s i des p r o t r u d i n g ; p r o t r u s i on s p o i n te d ; meso-me t a p l e u r a l s u t u re d i s t i n c t a n d comp l e t e ; m e t a t h o r a x + p ro p o d e u m g r a n u I a ted , excep t for a n t e r i or s u rface s h i n y a n d w i t ho u t sc u l p t u re ; mesop l e u r a , m e t a p l e u r a a n d pos t e r i o r s u rface of p ro p o d e u m t r a n s v e rse l y s t r i a t e ; d i sc of m e t a t horax + p ropodeum w i t h a t r a c k of med i a n fu rrow ; fore t a rsa l segmen t s i n fo l l ow i n g p ropor t i on s : 1 3 : 3 : 5 : 1 6 : 25 ; en l arged c l a w ( F i g . 8 1 9 ) w i t h s u b a p i c a l t o o t h a n d 1 1 l a me ' i ae ; seg m e n t 5 of fron t t a r s u s ( F i g . 8 1 9 ) w i t h 2 rows of n e a r l y 1 6 l a m e l l a e ; a p ex w i t h a g ro u p of n e a r l y 9 l a me l l ae ; m a x i l l a ry p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 seg men t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k no w n L O C U S TYP I C U S : M e a d i ( E g y p t ) paratype F ! i n OL . TYP I CAL MAT E R I AL : h o l o t y p e F ! i n WA ; HOSTS : u n k n o w n D I ST R I B U T I O N : P a l a e a rc t i c reg i o n : E GYPT : M e a d i , WA ! O L ! NOT E : t h e s pec i es i s n a med i n honor o f M r . H . P r ", es n e r .

1 2 . Pseu dogona top u s septemden t a t u s ( J . Sa h l berg )

n.

com b .

Gon a top u s septemden t a t u s J . Sa h l berg 1 9 1 0 : 1 1 . Pseu dogon a top u s sep a ratus R i c h a r d s 1 939 : 207 ( n . s y n . ) ; locus typicus Port hc a w l ( E n g l a n d ) . nec Pseu dogona topus sep a ra tus R i c h a rds : Ab d u l - N o u r 1 976 : 274 ( see Pseu dogon a top u s a lbosign a tus K . a n d ligu s ticus C u rrado and O l m i ) . Pseu dogon a top u s distinctus ( K i effer ) : Ponomarenko 1 97 8 : 2 3 .

F E MA L E : a p t e ro u s ; l e n g t h 3 , 0-3 , 5 m m ; head b l ac k , w i t h occ i p u t , m a n d i b l e s , c l y pe u s a n d f ro n t p a r t of v e r t e x y e l l o w ; t horax a n d p ropod e u m b l ac k , w i t h p o s t e r i o r h a l f o f sc u t u m ye l l o w ; a p ex of p ropodeum y e l l o w ; occ a s i o n a l l y pron o t u m b row n , w i t h re d d i sh n u a n c es ; a n t e n n ae b row n , w i t h s e g me n t s 1 -2 a n d 1 0 ye l l o w ; a b domen b rown-b l a c k , w i t h t e rg i t e 1 p a r t l y red d i s h ; l e g s y e l l ow , w i t h c l u b s of fore femora b ro w n i s h ; occas i o n a l l y c l u b s o f m i d a n d h i n d femora a n d m i d a n d h i n d coxae b ro w n i s h ; h e a d e xc a v a t e d , s h i n y , a l u t aceou s , wea k l y gra n u l a t e d ; p ron o t u m crossed by a s t ro n g t r a n sversa l i m p res s i o n , s h i n y , w i t ho u t scu l p t u re or w e a k l y g ra n u l a t e d ; m e t a n o t u m t r a n sv e r se l y s t r i a t e , w i t h

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F i g . 820 - Mesotho ra x , m e t a t h o r a x a n d p ropodeum of fem a l es of Pseu dogon a top u s septemde n t a t u s ( J . Sa h l berg ) ( A ) a n d ligu sticus C u rr a d o and O l m i ( 8 )

s i des p r o t ru d i n g ; p ro t r u s i o n s rou n ded ( F i g . 820 A ) ; meso- m e t a p l eu r a l s u t u re d i s t i n c t a n d com p l e t e ; a n t e r i o r s u rface o f m e t a t ho r a x + p ropod e u m w i t h t ra c k o f m ed i a n f u rrow ; mesop l eu r a , m e t a p l e u r a a n d pos t e r i or s u rface of propode u m t r a n s verse I y s t r i a t e ; fore t a rs a I s e g m e n t s i n fo l l o w i n g p ropo r t i o n s : 1 5 : 2 : 4 : 1 0 : 1 8 ; en l arged c l a w ( F i g . 8 2 1 ) w i t h s u b a p i c a l t o o t h a n d 8- 1 3 l a m e l l a e ; seg m e n t 5 of fron t t a rsus ( F i g . 8 2 1 ) w i t h 2 rows of 1 6- 2 1 l a m e l l ae ; a p e x w i t h a g roup of 6-25 l a m e l l a e ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k no w n LOCUS TYP I CU S : Juustjarv i

( F i n l an d )

TYP I CAL M A T E R I AL : h o l o t y pe F ! o f G . septemden t a t u s i n H E ; h o l o t y p e F ! a n d 2 p a r a t y pes F F ' of Ps . sep a r a t u s i n 8 M . HOSTS : i n U . S . S . R . Meta delp h a x prop i n q u a F i e b . ( Po n o m a r e n k o 1 978 , u n der n a me Ps . distin c tu s ) ; Aphrodes nervo s u s by m i s t a k e l i s t e d b y L i n d b e r g ( 1 950 ) A . D I ST R I BU T I ON : P a l aearc t i c reg i o n : U . S . S . R . : C r i m ea ( Po n o m a r e n k o 1 978 , u n der n a m e Pseu dogon a top u s dis tin ctus K . ) , P K ! F I NLA N D : J u u s t j a rv i ( K a re I i a ) , H E ! ROMAN I A ( N a g y 1 9 72 ) ; G E R M A NY : Hopfenberg ( Se h l i s , L e i p z i g ) , O L ! Sch l osspark ( L e i p z i g-K l zsc h . ) , HA ! SW I T Z E R L A N D : N y o n , G V ! E N G LA N D : P o r t h c a w l ( G l a m o r g a n ) , B M ! S t . M e r r y n ( C o r n w a l l E a s t ) , B M ! I TA L Y : C a s a l e R o l e t t o ( Mo n g r a n do , Verce l l i ) , OL ! S Z '

1 3 . Pseudogona top u s ligusticus C u r rado a n d

0I

mi

Pseu dogon a top u s ligu s ticus C u rrado a n d 0 I m i 1 974a : 229 . Pseu dogon a top u s separa t u s R i c h a r d s partim : A b d u I -No u r 1 976 :

Mem .

1 2 20

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821

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274 .

F E MAL E : a p tero u s ; l en g t h 2 , 60-3 , 7 5 m m ; h e a d b ro w n - b l ac k , w i t h c l y p eu s , occ i p u t , m a n d i b l es a n d f r o n t p a r t o f v e r t e x y e l l ow ; occa s i on a l l y occ i p u t b row n ; a n te n n a e b ro w n - b l ac k , w i t h seg men t s 1 - 2 y e l l o w ; a n te n n a l s e g me n t 1 0 l i g h t or b ro w n ; p ropec t u s b ro w n - b l ack ; p ro n o t u m b l ac k ; occa s i o n a l l y d i sc re d d i s h , co l l a r a n d s i des b ro w n - r e d d i s h ; sc u t u m y e l l ow , w i t h a n ter i or s u rface b row n - b l a c k ; sc u te l l u m a n d m e t a n o t u m redd i s h ; m e t a t h o r a x + propodeum b ro w n - b l ac k , occ a s i on a l l y w i t h red d i s h n u a n ces o n a n t e r i or s u rf a ce ; apex of p ro p o d e u m y e l l ow ; a b domen b ro w n - b l ac k ; l eg s red d i s h , w i t h c l u b s of femora b row n i s h ; h e a d s h i n y , e xc a v a t ed , g r a n u l a ted ; p r o n o t u m s h i n y , w i t ho u t sc u l p t u r e , c rossed b y a s t ro n g t r a n sv e rs a l i m p ress i o n ; s c u t u m a n d s c u te l l u m smoot h ; m e t a n o t u m h o l l ow b e h i n d t he s c u t e l l u m , w i t h s i des p r o t ru d i n g ; p r o t r u s i on s p o i n ted ( F i g . 820 B ) ; meso-me t a p l eu r a l s u t u re d i s t i nc t and comp l e t e ; meso p l eu ra ,

Gon a topod i n a e

Fig.

1 22 1

822 - Che I a of Pseu dogon a top u s ligusticus C u rrado a n d 0 I m i from A l t a re ( I t a l y )

met a p l e u ra a n d p o s t e r i o r s u rface of p ropodeum t r a n sverse l y s t r i a t e ; met a t ho r a x + propodeum s h i n y , w i t h o u t scu l p t u re a m o n g s t r i a e ; fore t a r s a l s e g me n t s i n fo l l o w i n g p ropor t i o n s : 1 4 : 3 : 6 : 1 7 : 2 6 ; e n l arged c l a w ( F i g . 822 ) w i t h s u b a p i c a l tooth a n d 9- 1 0 l a me l l ae ; seg m e n t 5 of fro n t t a rs u s ( F i g . 8 2 2 ) w i t h t w o rows o f 1 7-23 l a me l l ae ; a p e x w i t h a g ro u p of 1 5-23 l a me l l ae ; m a x i l l a ry p a l p i w i t h 4 s e g me n t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k no w n L O C U S T Y P I C U S : S a n Lorenzo d i C a s a n ov a ( I t a I y ) T Y P I CAL MAT E R I A L : ho l o t y p e F ! i n GE . HOST S : i n F r a nce Rib a u todelp h a x imi tans R i b a u t ( A b d u l - N o u r 1 976 , u n der n a m e Pseu dogona topus s ep a r a t u s R i c h a rds p a rt im ) . D I S T R I BU T I O N : P a l a e a rc t i c reg i on : H U N GARY : K rs k u n h a l a s ( Bo g arzo ) , B U ! Tomp a , B U ! V a cz , B U ! B O ! F R A N C E : S t . Ge l y -d u-Fesc ( He r a u l t ) , P ! SPA I N : Torrev i ej a ( A l i c a n t e ) , OL ! B M ! P a l amos ( B a rce l o n a ) , SZ ! I TA L Y : S a n B e n e d e t t o B e l bo ( C u n eo ) , O L ! S a n Lore n zo d i C a s a n o v a ( Ge n ov a ) , G E ! A l t a re ( Sa v on a ) , P L !

E T H I O P I A N S P E C I E S O F PSEUDOGONA TOPUS

1 22 2

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KEY T O T H E SPEC I ES

F E MA L E S

2 3

4

5

6

M e t a n o t u m w i t h s i d e s r o u n d ed ( F i g . 8 2 4 ) ; me s o -m e t a p l eu ra l s u t u r e ob s o l e t e • • • • • . . • . • . . • . . . . • . . . . • . . . • . . • • • . . • . . . • • • • . . • • 2 M e t a n o t u m w i t h s i d e s p r o t r u d i n g ( p r o t ru s i o n s r o u n d e d o r p o i n t ed ) ( F i g s 8 3 1 A , 8 3 3 A ) ; me s o -me t a p l e u ra l s u t u r e d i s t i n c t • . . . . • . • • . . . . • . • . . . . • • • • . . . . . • • . . . . . . . . . . . . . . . . . . . . . . . . . . . . . • 6 T h o ra x a n d p r op od e u m a t l ea s t p a r t l y b l a c k • • • • • • • • • • • . . • • . . . . 3 T h o ra x a n d p r o p o d e u m y e l l ow - t e s t a c e ou s o r r ed d i s h -y e l l ow • . . . • 4 P r on o t u m a n d s c u t u m b l a c k . . . . 1 . pilo s u s B en o i t P r o n o t u m w i t h c o l l a r b l a ck a n d d i s c y e l l ow ; s c u t u m y e l l ow , w i t h a n t e r i o r s u r f a c e b l a c k . , 2 . perkinsiella vorus B e n o i t B od y w i t h ou t s c u l p t u r e , v e ry s h i n y . . . . . . . . . . . . . . . . . . . . . . . . . . . . • • • • . • . • • • • • • . . . • • • . . • • • • • • • • 3 . seyrigi ( B e n o i t ) B od y s t r o n g l y o r w e a k l y g ra n u l a t ed . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 H e a d f u l 'l y y e l l ow - t e s t a c e o u s . 4 . polli B en o i t H ea d b r own - b l a c k , w i t h o c c i p u t , c l y p eu s , man d i b l e s a n d f r on t p a r t o f v e r t e x y e l l ow . . . . . . . . 5 . ruens n ew n a m e B od y r ed d i s h -d a rk o r a t I e a s t p a r t I y b l a ck o r b r own . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6 . a c u t u s n . sp . B o d y y e l l ow - t e s t a c e ou s o r r ed d i s h - l i g h t , w i t h p e t i o l e b l a c k ; o c c a s i o n a l l y a n t en n a e o r a b d omen b r own o r p a r t l y b l a ck . . . . . .

7

P r o n o t um w i t h (F i g . 831 B ) . . P r on o t u m w i t h (F i g . 833 B ) . .

t . t .

ra n s v e r s a . . . . . . . . . ra n s v e r s a . . . . . . . . .

l i mp r e s s i on s t r o n g l y d e ep a n d b r oad . . . . . . 7 . spira c u laris B en o i t l i mp r e s s i o n n a r r ow a n d l e s s d e ep . . . . . . 8 . similis ( B r u e s )

MALES

O n l y t h e m a l e of Ps .

ruens i s k no w n .

1 . Pse u dogona top u s pilosus B e n o i t

Pseu dogo n a top u s p il o s u s Beno i t 1 9 53b : 390 .

.

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F i g . 823 - C h e l a of Ps e u dago n a t op u s pilasus B e n a i t

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( type )

F E MA L E : a p terou s ; l en g t h 4 , 5 m m ; h e a d b l ac k , w i t h m a n d i b l es a n d c l y p e u s y e l l ow ; a n t e n n a e b r o w n - b l ac k , w i t h seg men t s 1 a n d 1 0 y e l l ow o r w h i t e ; t ho r a x a n d p ropodeum b l ac k , w i t h a p e x o f p ropode u m red d i s h ; pet i o l e a n d a b do m e n b l ac k ; l eg s b ro w n - b l a c k ; h e a d exc a v a t ed , s h i n y , smoo t h , h a i r l es s , w i t h o u t s c u l p t u re ; frons w i t h 2 med i a n k ee l s con n ec t i n g c l y p e u s to oce l l a r reg i on ; oce l l a r reg i o n r u g os e ;

p rono t u m crossed b y a

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1 224

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I n s! . ,

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1 984

F i g . 824 - T ho r a x a n d p ropodeum of fema l e of Pseu dogon a top u s perkinsiella vorus Be n o i t ( t y p e )

s t ro n g t ra n s vers a l i m p ress i o n , smoo t h , w i t h o u t s c u I p t u re , e x c e p t for some k ee l s o n s i d es ; scu t u m r u gose , w i t h some l on g i t u d i n a l k ee l s ; s c u t e l l u m s h i n y , smoo t h ; m e t a n o t u m t r a n s v e rse l y s t r i a t e , w i t h s i des rou n d e d ; meso m e t a p l eu r a l s u t u re obso l e t e ; m e t a t ho r a x + p ro p o d e u m w i t h t r a c k of med i a n f u r ro w ; p l e u r a a n d pos t e r i o r s u rface o f p ro p o d e u m t r a n s v e rse I y s t r i a t e ; a n te r i o r s u rface o f m e t a t horax + p ropodeum smoo t h , w i t ho u t scu I p t u r e ; segme n t 1 o f f ro n t t a rs u s a s l o n g a s s e g m e n t 4 ; en l a rged c l a w ( F i g . 8 2 3 ) w i t h s u b a p i c a l too t h a n d 1 2 l a me l l a e ; s e g me n t 5 o f fron t t a rs u s ( F i g . 8 2 3 ) w i t h 2 rows of 2 2 l a me l l ae ; apex w i t h a g ro u p of a t l e a s t 1 2 l a me l l ae ; m a x i l l a r y p a l p i w i t h 4 segme n ts ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E : u n k no w n LOCUS TYP I CUS :

I soana I a

( M a d a g asc a r )

T Y P I CAL M A T E R I AL : h o l o t y pe F ! i n P . HOSTS : u n k n o w n D I ST R I BU T I O N : E t h i op i a n reg i on : MADAGASCAR :

2.

I so a n a l a ,

P!

Pseudogo n a top u s perkinsiella vorus B e n o i t

Pseu dogon a top u s perkinsiella vorus Ben o i t 1 96 3 : 341 .

1 22 5

Gona topod i n a e

F E M A L E : a p terou s ; l e n g t h 3 , 5 m m ; h e a d b l ac k ( accord i n g t o t he or i g i n a l desc r i p t i o n : h e a d m i ss i n g i n t h e o n l y k n o w n s p ec i m e n ) ; a n t e n n ae y e l l o w , w i t h s e g m e n t 1 0 w h i t e ( accord i n g to t he o r i g i n a l desc r i p t i o n ) ) ; p rono t u m y e l l o w , w i t h co l l a r b l ac k ; sc u t u m y e l l o w , w i t h a n t e r i or s u rf a c e b l ac k ; s c u t e l l u m ye l l ow ; m e t a n o t u m red d i sh ; m e t a t ho r a x + p ro p o de u m b l ac k , w i t h a pe x of p ro p o d e u m ye l l o w ; pe t i o l e b l ac k ; a b domen y e l l o w - r e d d i s h ; l eg s y e l l ow ; h e a d m i s s i n g i n t he o n l y k n o w n s p ec i men ; a n t en n a l seg m e n t 1 more t h a n 7 t i mes as l o n g as b roa d , s h o r t e r t h a n 2 + 3 ( 4 3 : 50 ) ; seg m e n t 3 t h ree t i mes as l o n g as b ro a d ( accord i n g to t he or i g i n a l descr i p t i on ) ; p ron o t u m ( F i g . 824 ) smoo t h , s h i n y , w i t ho u t sc u l p t u r e , crossed b y a s t ro n g t r a n s v e rs a l i m p re ss i o n ; s c u t u m a n d scu te l l u m s h i n y , h a i r l ess , w i t ho u t sc u l p t ure ; m e t a n o t u m t r a n s v erse l y s t r i a t e , w i t h s i des r o u n ded ; met a t ho r a x + p ropodeum ( F i g . 824 ) s h i n y , s m oo t h , h a i r l ess ; p l e u ra a n d pos t e r i o r s u rface o f p ropode u m t r a n sv erse l y s t r i a t e ; fore l e g s m i s s i n g i n t he o n l y k n ow n spec i me n ; c he l a not descr i bed i n t h e o r i g i n a l desc r i p t i o n ; m a x i I l a r y p a l p i w i t h 4 s e g m e n t s ( accord i n g to t h e or i g i n a l descr i p t i on ) ; l ab i a l p a l p i w i t h 2 se g m e n t s ( accord i n g to t he o r i g i n a l d e s c r i p t i on ) ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k no w n L O C U S T Y P I C U S : B r i c k a v i l l e ( M a d a g as c a r ) TYP I CAL M A T E R I AL : h o l o t y pe F ' i n GV . HOSTS : i n M a d a g a scar Perk insiella s a c ch aricida K i r k . ( Be n o i t 1 96 3 : 341 ) . D I ST R I BU T I O N : E t h i o p i a n reg i o n : MADAGASC A R : B r i c k a v i l l e , GV !

3 . Pseu dogo n a top u s seyrigi

( Be n o i t )

n . comb .

A go n a topoides seyrigi B e n o i t 1 953 : 388 .

F E M A L E : a p terou s ; l en g t h 2 , 75 m m ; y e l l ow - t e s t aceo u s , w i t h p e t i o l e b l ac k ; a n t e n n a e w i t h seg m e n t s 3- 1 0 we a k l y b row n i s h ; a b domen p a r t l y b row n i s h ; b o d y s h i n y , smoot h , h a i r l es s , w i t ho u t sc u l p t u re ; h e a d excav a te d ; p ro no t u m crossed b y a s t ro n g t r a n s versa I i m p ress i o n ; meso-me t a p l eu r a I s u t u re o b s o l e t e ; p o s te r i o r s u rface of p ropodeum t r a n s v e rse l y s t r i a t e ; seg m e n t 1 o f fron t t a r s u s l on ge r t h a n segmen t 4 ( 1 3 : 1 0 ) ; en l a r g e d c l a w ( F i g . 8 2 5 ) w i t h s u b a p i c a l t o o t h a n d 4-5 l a me l l ae ; s e g m e n t 5 of f ro n t t a r s u s ( F i g . 825 ) w i t h 2 rows o f 1 9-30 l a me l l ae e x t e n d i n g con t i n uo u s l y t o t he a p e x ; a pex n o t p ro m i n e n t ; m a x i I l a ry p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : F o r t -D a u p h i n ( M a d a g a s ca r ) T Y P I CAL M A T E R I A L : ho l o t y p e F ! i n P ; 1 p a r a t y p e F ! i n T V . HOSTS : u n k n o w n D I S T R I B U T I O N : E t h i o p i a n reg i o n : MADAGASCA R : F o r t - D a u p h i n , P ' T V '

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C he l a of Pseu dogon a top u s seyrigi ( Ben o i t ) ( t y p e )

4 . Pseu dogon a top u s polli B e n o i t

Pseu dogona topus polli Ben o i t 1 95 1 c : 8 .

1 227

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826 - C h e l a of Pseudo go n a tonus polli Beno i t

( t y pe )

F E MA L E : a p terou s ; l en g t h 3 , 00-3 , 62 mm ; y e l l ow - red d i s h , w i t h p e t i o l e b l ac k ; h e a d s h i n y , exc a v a ted , wea k l y g r a n u l a t ed ; f ro n s smoo t h , w i t h o u t s c u I p t u re ; p r o n o t u m crossed b y a s t ro n g t r a n sve rsa I i m p ress i on , s h i n y , g r a n u l a ted ; scu t u m , sc u t e l l u m a n d m e t a t ho r a x + p ropode u m g r a n u l a t e d ; m e t a n o t u m w i t h s i des ro u n ded ; meso-met a p l e u r a l s u t u re o b so l e t e ; a n te r i o r

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s u rface of me t a t ho r a x + p ropodeum w i t h t r a c k of med i a n f u rrow ; p o s te r i o r s u rf a c e o f p ro p o d e u m t r a n s verse l y s t r i a t e ; mesop l e u r a a n d m e t a p l e u ra n o t t r a n s v e rs e l y s t r i a t e ; fore t a r s a l segm e n t s i n fo l l o w i n g p roport i o n s : 1 3 : 2 : 4 : 1 1 : 1 8 ; en l a r g e d c l a w ( F i g . 826 ) w i t h s u b a p i c a l toot h a n d 4-5 I a m e l i a e ; s e g m e n t 5 of fron t t a r s u s (F i g . 8 2 6 ) w i t h 2 rows of 20-37 l a m e l l ae e x t e n d i n g cont i n u ou s l y to t he a p e x ; apex prom i n e n t ; m a x i l l a ry p a l pi w i t h 4 segmen t s ; l ab i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k no w n L O C U S TYP I C US :

K a l om b o ( Z a i re )

TYP I CAL M A T E R I AL : ho l o t y pe F ! i n T V . HOST S : u n k no w n D I ST R I B U T I O N : E t h i op i a n reg i o n : ZA I R E : K a l om b o , T V ! SO U T H A F R I CA : Mossel B a y ( C a p e P rov i n c e ) , B M ! Ceres ( C a p e Prov i nc e ) , B M ! Somerset E a s t ( C a p e Prov i n ce ) , OL !

5 . Pseu dogon a top u s ruens n e w n a me

n ec Pseudogona topus r u fu s F o u t s 1 935 : Parago n a top us rufus D l m i 1 97 9 : 2 1 9 .

1 58 .

F E MAL E : a p terou s ; l e n g t h 2 , 8 1 -3 , 1 2 m m ; h e a d red d i s h , w i t h reg i on n e a r oce l l i b row n i s h ; f ro n t p a rt of v e r t e x y e l l ow-re d d i s h ; a n te n n ae b ro w n , w i t h segmen t s 1 -3 y e l l o w ; t h o r a x a n d p ropodeum redd i s h ; pe t i o l e a n d a b do m e n b l ac k , except for a b d om i n a l terg i te 1 red d i s h ; l e g s ye l l ow redd i s h , w i t h c l u b s of f o r e f e m o r a b row n ; h e a d e x c a v a ted , s h i n y , smoo t h , w i t h o u t scu l p t u r e i n fro n t of t h e oce l l i ; occ i p u t a n d fron t p a r t of v e r t e x g ra n u l a ted ; p ron o t u m d u l l , g r an u l a t e d , crossed b y a s t rong t r a n s v e rs a l i m p ress i o n ; sc u t u m d u l l , w i t h some l on g i t u d i n a l s t r i ae ; m e t a n o t u m f l a t , t r a n s verse I y s t r i a t e , a s l o n g a s sc u t u m ; m e t a t h o r a x + p ro p o d e u m d u I I ; occas i o n a l l y a n t er i or su rface w i t h t r a c k of med i a n f u r ro w ; poster i o r s u rf a c e , mesop l e u r a a n d me t a p l e u r a t r a n s v erse l y s t r i a t e ; meso-me t a p l eu r a l s u t u re obso l e t e ; fore t a rs a l seg men t s i n fo l l o w i n g p ropor t i o n s : 9 : 2 : 3 : 6 : 1 0 ; e n l a r ged c l a w ( F i g . 827 ) w i t h s u b a p i c a l tooth a n d 5 l a me l l ae ; s e g m e n t 5 of f r o n t t a rs u s ( F i g . 8 2 7 ) w i t h 2 rows o f 1 5- 1 7 l a me l l ae ; a p e x w i t h a g r o u p of 7 - 8 l a me l l ae ; m a x i I l a ry p a l p i w i t h 2 - 4 segmen t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE ( F i g . 828 ) : f u l l y w i n ged ; l e n g t h 2 , 5 m m ; he a d b l ac k , w i t h m a n d i b l es y e l l ow ; a n t en n ae b row n ; t h o r a x a n d p ropodeum b l ac k ; a b domen b row n ; teg u l ae and l e g s y e l l o w ; a n t e n n a I s e g m e n t s i n fo l l o w i n g p roport i o n s : 5 , 5 : 5 , 5 : 1 2 : 1 1 : 1 0 : 1 0 : 1 0 : 1 0 : 9 : 1 3 ; a n t e n n a I segmen t 3 more t h a n t h ree t i mes as l on g as b ro a d ( 4 , 8 ) ; h e a d h a i ry , d u l l , g r a n u l a t ed , w i t ho u t med i a n k e e l b e t ween t he a n te n n a l soc k e t s ; t e m p l es d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 8 ; DL = 4 ; DOL = 3 ; b r ea d t h of t h e oce l l i : 3 ; s c u t u m h a i r y , g r a n u l a ted ; n o t a u l i ces comp l e t e ,

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1 229

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poste r i o r l y s e p a r a t e d ; m i n i m u m d i s t a nce b e t ween t h e n o t a u l i ces shorter t h a n b re a d t h of t h e oce l l i ( 2 : 3 ) ; sc u t e l l u m and m e t a n o t u m h a i ry , g r a n u l a ted ; p ropodeum d u l l ; pos t e r i o r s u rface ret i c u l a te ru gose ; d i sc w i t h med i a n f u rrow a n d w i t h s h o r t i r reg u l a r k ee l s ; fore-w i n g h y a l i n e , w i t ho u t

1 230

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F i g . 828 - M a l e of Pseu dogona top u s ruens n .

1 984

name ( p aratype)

d a r k t r a n svers a l b a n d s ; dors a l p rocess of g o noforceps ( F i g . 829 ) d i s t a l l y s h a rp ; m a x i l l a r y p a l p i w i t h 2 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . lOCUS T Y P I C U S : S w a k op m u n d ( So u t h W e s t Afr i ca ) T Y P I CAL M A T E R I AL : h o l o t y p e F ! a n d 8 p a r a t y pes ( 7 F F , 1 M ) ' i n B M ; 2 p a r a t y pes F F ! i n O l . HOST S : u n k n o w n D I ST R I BU T I O N : E t h i o p i a n reg i o n : SO U T H W E S T A F R I CA : S w a k o p m u n d , B M ! Ol !

S O U T H A F R I CA : G r a h a m s tow n

( C ape Prov i nce ) ,

BM !

6 . Pseu dogon a top u s a c u t u s n .

sp .

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829 - M a l e g e n i t a l i a o f

Pseudogona to,o u s

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n.

n a me ( p a r a t y pe )

F E M A L E : a p terou s ; l e n g t h 2 , 5-3 , 5 mm ; h e a d b l a c k or b row n , w i t h m a n d i b l es , c l y p e u s a n d f ro n t p a r t of v e r t e x y e l l ow ; a n t e n n a e b row n , seg m e n t s 1 -2 y e l l ow ;

a n te n n a l seg m e n t 1 0

w i th

I i g h t ; t h o r a x a n d p ropodeum

b l ac k o r b rown-redd i s h , w i t h poster i or h a l f of scu t u m a n d a p e x o f propode u m y e l l ow ; pet i o l e b l ac k ; a b domen b l ac k o r b ro w n - b l ac k ; l e g s y e l l o w , w i t h cox a e , c l u b s of femora a n d t i b i a e b row n i s h ; h e a d s h i n y , g ra n u l a te d , exc a v a t ed ; p r o n o t u m crossed b y a s t r o n g t r a n s v e rs a l i m p ress i o n , d u l l , g r a n u l a ted ; m e t a n o t u m t r a n s v erse l y s t r i a t e , w i t h s i d es p r o t r u d i n g ; p ro t r u s i o n s p o i n t ed ; m e t a t h o r a x + p ropod e u m s h i n y , smoot h , w i t h o u t sc u l p t ure ; meso p l e u r a , m e t a p l e u r a a n d p o s te r i o r s u rface o f

1 2 32

Mem . Amer.

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F i g . 830 - C h e l a of Pse u dogon a to,o u s a c u tu s n .

sp .

( type)

p r'opod e u m t r a n sverse l y s t r i a t e ; d i sc w i t h t r ack o f med i a n f u r row ; meso m e t a p l e u r a l s u t u re d i s t i n c t a n d comp l e t e ; fore t a rs a l segmen t s i n fo l l o w i n g p ropor t i o n s : 1 3 : 2 : 4 : 1 3 : 20 ; en l a rged c l a w ( F i g . 830 ) w i t h s u b a p i c a l t oo t h a n d 6 - 1 1 l a m e l l a e ; seg men t 5 of f ro n t t a r s u s ( F i g . 830 ) w i t h t wo rows of 1 0- 2 1 l a m e l l a e ( oc c a s i on a l l y s h o r t e s t ro w o n l y l oc a t e d on

1 233

Gon a topod i n a e

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831

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F em a l e of Pseu dogon a top u s spira c u l a ri s Beno i t B : t ho r a x a n d p ropodeu m ; C : h e a d .

( p a r a t y pe ) : A ,

t h e p r ox i m a l h a l f ) ; a p e x w i t h a g ro u p of 8-20 l a me l l ae ; m a x i I l a r y p a l p i w i t h 4 s e g m e n t s ; l a b i a l p a l p i w i t h 2 s e g m e n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k nown l O C U S TYP I C US : Mosse l B a y ( So u t h Af r i c a ) T Y P I CAL M A T E R I A L : ho l o t y pe F ! i n BM ; 2 p a r a t y pes F F ! i n O l . HOST S : u n k no w n D I ST R I B U T I O N : E t h i o p i a n reg i o n : S O U T H A F R I CA : Mosse l B a y ( C a p e P rov i n ce ) , BM ! O l !

7 . Pseu dogon a top u s spir a c u l a ris Beno i t

Pseu dogona top u s spira c u l aris Beno i t 1 96 3 : 339 .

F E M A L E : a p t e ro u s ; l en g t h 3 , 00-3 , 43 m m ; y e l l o w - t e s t a c eo u s , w i t h p e t i o l e b l ac k ; bod y s h i n y , h a i r l es s , w i t h o u t sc u l p t u r e , smoo t h ; h e a d ( F i g . 8 3 1 C ) e x c a v a t ed ; p ro n o t u m ( F i g . 8 3 1 A , B ) crossed b y a s t ro n g t r a n s v ersa l i m p ress i o n ; met a n o t u m w i t h s i des p ro t r u d i n g ; p ro t r u s i o n s r o u n d e d ; meso met a p l e u ra l s u t u re d i s t i n c t a n d comp l e t e ; m e t a t h orax + p ro p o d e u m ( F i g . 8 3 1 A , B ) w i t h med i a n f u rrow ; meso p l e u r a , m et a p l e u ra a n d p o s t e r i o r s u rf a c e of p ropodeum t r a n s v e rse l y s t r i a t e ; s e g m e n t 1 of f ro n t t a rs u s s l i g h t l y s h o r t e r t h a n segmen t 4 ( 1 3 : 1 4 ) ; en l a r g e d c l a w ( F i g . 8 3 2 ) w i t h

1 234

Mem . A me r .

F i g . 832

-

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I n s! . ,

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C he l a of Pseudogo n a top u s spira c ularis Ben o i t ( p a r a t y p e )

s u b a p i c a l too t h a n d 7 l a me l l ae ; s e g m e n t 5 of fron t t a r s u s ( F i g . 832 ) w i t h two rows of 20 l a me l l ae ; a p e x w i t h a g rou p of 1 7 l a me l l a e ; m a x i l l a r y p a l p i w i t h 4 s e g me n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E : u n k no w n LOCUS TYP I CU S : B r i c k a v i l i e ( Mad agascar ) T Y P I C A L MAT E R I AL : ho l o t y pe F ! a n d 1 p a r a t y pe F ! i n G V ; 2 p a r a t y pes FF ! i n TV. HOS T S : i n M a d a g a s c a r Perkinsiella s a ccharicida K i r k . ( B e n o i t 1 963 ) . D I ST R I BU T I ON : E t h i op i a n reg i o n : MADAGASCA R : B r i c k a v i l l e , G V ! T V !

8 . Pseu dogon a top u s similis ( B r u e s )

n . com b .

Gon a top u s similis B r u e s 1 906a : 1 07 . Pseudogon a topoides m a uritia n u s W i I I i a m s 1 956 : 9 0 ( n . s y n . ) ; locus typic u s Pa I m y re ( M a u r i t i u s ) . nee Pseu dogon a top u s similis F re y t a g 1 978 : 1 39 .

F E MA L E : a p terou s ; l e n g t h 2 , 50-2 , 8 1 m m ; y e l l ow - t e s t aceou s , w i t h p et i o l e b l ac k ; a n te n n a e b ro w n , w i t h s e g m e n t s 1 -2 a n d 1 0 w h i te o r y e l l o w ; occ a s i on a l l y segmen t 1 0 b ro w n ; a b domen b row n i s h or y e l l ow - t e s t aceou s ; body s h i n y , smoot h , w i t h o u t scu l p t u re ; h e a d ( F i g . 833 C ) exc a v a t e d ; p ro n o t u m ( F i g .

833 A ,

B ) crossed b y a s t ro n g t r a n s v e rs a l

i m p ress i o n ;

1 235

Gon a topod i n a e

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833 - F e m a l e of Pseu dogon a top u s similis ( B r u e s ) ( t y p e ) : A , a n d p ropod e u m ; C : h e a d .

B:

thorax

m e t a n o t u m t r a n s v e r se l y s t r i a t e , w i t h s i des p ro t r u d i n g ; p r o t r u s i o n s ro u n de d ; meso-me t a p l e u r a l s u t u re d i s t i n c t a n d comp l e t e ; mesop l e u r a , met a p l e u r a a n d pos t e r i o r s u rface of p ro p o d e u m t ra n s verse l y s t r i a t e ; segmen t 1 of f r o n t t a r s u s s l i g h t l y l on g e r t h a n segmen t 4 ( 1 2 : 1 1 ) ; e n l a r g e d c l a w ( F i g . 834 A ) w i t h s u b a p i c a l t oo t h a n d 8 - 1 1 l a me l l ae ; seg m e n t 5 o f f r o n t t a r s u s ( F i g . 834 A ) w i t h t w o rows o f 1 6- 1 9 l a me l l ae ; a p ex w i t h a g r o u p o f 9 - 1 3 l ame l l a e ; m a x i l l a ry p a l p i ( F i g . 834 B ) w i t h 3-4 segmen t s ; l a b i a l p a l p i ( F i g . 834 C ) w i t h 2 segme n t s ; t i b i a l s p u r s 1 , 0, 1 . MALE : u n k nown L O C U S TYP I C U S : A l goa B a y ( So u t h A f r i c a ) TYP I CAL MAT E R I A L : ho l o t y p e F ' o f C . similis i n M L ; p a r a t y pe s F F ' of Ps . m a uritia n u s i n B M .

ho l o t y p e F '

and 2

HOSTS : i n M a u r i t i u s Perkinsiella s a cch aricida K i rk . a n d Dicran otropis m u iri K i r k . ( W i I I i a ms 1 956 ) . D I S T R I B U T I O N : E t h i o p i a n reg i o n : M A U R I T I U S : P a l m y r e , B M ! SOU T H A F R I C A : A l goa B a y ( C a p e P rov i n ce ) , M L ! Mosse l B a y ( C a pe P ro v i n ce ) , B M ' Port S t . Johns ( Po n do l a n d ) , B M ! L E SO T H O : M a m a t h es , O l ' PARAS I T E S : i n M a u r i t i u s Ch eiloneurus gona topodis R . C . l . ( Encyr't ida e ) ( W i l l i a m s 1 95 6 ) .

Perk i n s

N O T E S : Ps . s imilis a n d m a uritia n u s p rov i s i o n a l l y a re co n s i dered s y n o n y m s ; o n l y the s t u d y o f m a l es wi I I con f i rm the s y n o n y m y .

OR I E N T A l S P E C I E S OF

PSEUDOCONA TOPUS

Mem . Amer .

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En!.

I ns! . ,

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1 984

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Fig.

834 - F e m a l e of Ps eu dogon a top u s similis ( B r u es ) ( t y p e ) : A : c h e l a ; max i l l a ry pa l p u s ; C : l a b i a l p a l p u s .

Ten spec i es are k n ow n .

KEY T O T H E S P E C I ES

FEMALES

B:

1 237

G o n a topod i n a e

Th o ra x and p r op o d e u m f u l l y b l a ck or b r o wn - d a rk ( a t mo s t a p ex o f p r op o d e u m l i g h t ) . . . . . . . . . . . . . 8 . hospes R . C . L . P e r k i n s ( s e e A u s t r a l i a n s p e c i e s ) •

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keeI s M e s o -m e t a p l e u r a l s u t u r e ob s o l e t e . . . . . . . . . . . . . 3 . n u du s R . C . L . P e r k i n s M e s o -m e t a p l e u r a l s u t u r e mo r e o r l e s s d i s t i n c t . . . . . . . . . . . . . . . M e t a t h o ra x + p r o p o d e u m w i t h o u t med i a n f u r r o w . . . . . . . . . . . . . . . . M e t a t h o r a x + p r op o d e u m w i t h a w e a k o r s t r o n g m ed i a n f u r r o w M e t a n o t u m w i t h l a t e r a l p r o t ru s i o n s mo r e p r om i n en t . . . . . . . . . . . · . . 4 . m a lesiae n . s p . M e t a n o t u m w i t h l a t e r a l p r o t r u s i o n s l e s s p r om i n en t ( F i g . 8 4 4 A) 5 . validus n . sp . Ivie t a n o t u m w i t h l a t e ra l p r o t r u s i o n s mo r e p r om i n en t ( F i g . 8 4 6 B ) ; m e t a t h o ra x + p r o p o d e u m w i t h m ed i a n f u r r o w s t r o n g e r ; m e s o -me t a p l e u r a l s u t u r e v e ry s t r o n g a n d d i s t i n c t ( F i g . 8 4 6 B ) . . . . . . . . . . . . . 6 . l a n k a e P o n oma r en k o M e t a n o t u m w i t h l a t e r a l p r o t ru s i on s l e s s p r o m i n en t ( F i g . 8 4 6 A ) ; m e t a t h o ra x + p r op o d e u m w i t h med i a n f u r r o w w e a k ; m e s o -me t a p l eu ra l s u t u r e w e a k l y d i s t i n c t , on l y v i s i b l e n ea r t h e l a t e r a l p r o t ru s i o n s o f m e t a n o t u m ( F i g . 8 4 6 A ) . . . . . . . . . . . . . . . . . . 7 . m u iri n . sp . •

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1 . sarawaki Mo c z a r A n t e r i o r s u r f a c e o f p r op o d e u m d u l l , g r a n u I a t ed M e t a t h o ra x + p r op od e u m b r own , ex c ep t f o r a p ex o f p r op o d e u m l i gh t 9 . soga t a e R o h w e r Me t a t h o r a x + p r op od e u m r ed d i sh - t e s t a c e o u s 1 0 . p u s a n u s n . sp . A n t e r i o r s u r f a c e o f m e t a t h o r a x + p r o p o d e u m w i t h n u m e r ou s I ong i t ud i n a l k e e l s 2 . s a ra wa kensis n . sp . An t e r i o r s u r f a c e o f me t a t h o ra x + p r o p o d e u m s mo o t h , w i t h o u t ·

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M e t a n o t u m w i t h s i d e s p r o t r u d i n g ; p r o t ru s i on s r o u n d ed o r p o i n t ed ( F i g s 8 4 4 , 8 4 6 ) ; m e s o -m e t a p l e u r a l s u t u r e d i s t i n c t obso l et e. A n t e r i o r s u r f a c e o f p r op od e u m sh i n y , w i t h ou t s c u l p t u r e . . . .

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MALES

N o t au l i c es p o s t e r i o r l y j o i n t N o t a u l i c e s p o s t e r i o r l y s ep a ra t ed •

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P r o p o d e u m w i t h d o r s a l s u r f a c e s c u l p t u r ed by a r e o l a e mu c h s ma l l e r t h a n a r e o l a e o n p o s t e r i o r s u r f a c e •

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1 . s a ra waki Mo c za r P r o p od eum w i t h d o r sa l s u r f a c e s cu l p t u r ed b y a r e o l a e n ea r l y a s w i d e a s a r eo l ae o n p o s t e r i o r s u r f a c e Ma l e A ( Pseudogon a top u s m a lesiae o r sarawakensis : s e e n o t e o n p a g e 1 2 4 1 D o r s a l p r o c e s s o f g on o f o r c ep s w i t h d i s t a l h a l f b r o a d e n ed ( F i g . 8 7 8 ) ; p r o x i ma l ha l f s l e n d e r ( F i g . 8 7 8 ) . . . . . . . . . 8 . hospes R . C . l . P e rk i n s ( s e e A u s t ra l i a n s p e c i e s ) D o r sa l p r o c e s s o f g on o f o r c ep s a l mo s t f u l l y r eg u l a r l y b r oad en ed (F i g . 841 ) Ma l e B ( Pseu dogon a top u s m a lesiae o r sara w a kensis : s e e n o t e o n p a g e 1 2 4 1 ) •

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T h e m a l es of Ps . n u du s , m u iri, soga tae, p u s a n u s , validus a n d l a n k a e are u n k no w n . M a l es of m a lesiae a n d sara"lakensis : s e e n o t e o n p a g e 1 24 1 .

1 . Pseudogona top u s sarawaki Mocz a r

Pseudogona topus sara waki Moczar 1 979 : 7 7 . Pseu dogon a top u s ponomarenkoi Moczar 1 979 : 80 ( n . s y n . ) ; locus typicus P a y a h P a l oh ( Sa r a w a k ) . Pseu dogon a top u s ponomarenkoi Moc z a r : Ponomarenko 1 98 1 b : 880 .

F E M A L E : a p t e rou s ; l en g t h 3 , 00-4 , 1 6 m m ; head red d i s h - b ro w n , w i t h m a n d i b l es , c l y pe u s a n d f ro n t p a r t o f v e r t e x y e l l o w ; occ a s i o n a l l y head mos t l y d a rk o r b l ac k ; a n t e n n a e b ro w n , w i t h segme n t s 1 -2 y e l l o w ; t horax and p ropodeum red d i s h- t e s t a c eou s ; occ a s i on a l l y scu t u m y e l l ow ; p e t i o l e b l ac k ; a b domen b l ac k , w i t h red d i s h n u a n ces ; l eg s t e s taceou s ; a n t en n a l segmen t s i n f o l l ow i n g p roport i on s : 1 0 : 7 , 5 : 27 : 1 1 : 1 0 : 1 0 : 9 : 8 : 8 : 1 2 ; h e a d e xc a v a ted , s h i n y , smoo t h , w e a k l y a l u t aceou s ; P O L = 7 ; O l = 4 ; OOl = 1 2 ; p ro n o t u m crossed by a strong t ra n sversa I i mp ress i on , s h i n y , w e a k I y g ra n u l a t e d ; m e t a n o t u m s h i n y , t r a n sverse l y s t r i a t e , w i t h s i des roun ded ; meso-me t a p l eu r a l s u t u re o b so l et e ; m e t a t horax + p ropodeum s h i n y , smoo t h , occ a s i o n a l l y w i t h w e a k t r a c k of med i a n f u r ro w ; mesop l e u r a , met a p l eu r a a n d p os t e r i or s u rface o f p ropodeum t r a n s verse l y s t r i a t e , w i t ho u t scu l p t u re among s t r i a e ; fore t a rs a l segmen t s i n fo l l o w i n g p roport i on s : 1 5 : 3 : 4 : 1 5 : 23 ; e n l arged c l aw ( F i g . 835 ) w i t h s u b a p i c a l tooth a n d 6-7 l a me l l a e ; s e g me n t 5 of fron t t a r s u s ( F i g . 835 ) w i t h t w o rows o f 1 9-20 l a me l l ae ; a p e x w i t h a g ro u p of 5-8 l a m e l l ae ; m a x i l l a r y p a l p i w i t h 2-4 segmen t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE ( F i gs 836 , 837 ) : f u l l y w i n g ed ; l en g t h 2 , 25 m m ; b l ac k or b row n ;

Gona topod i n a e

F i g . 835

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1 239

C h e l a of Pseudogona topus sarawaki Mocz a r ( t y p e )

a n t e n n a e brow n ; m a n d i b l es a n d l eg s y e l l o w ; a b do m e n b row n ; a n t e n n a l segme n t s i n fo l l o w i n g p ropor t i on s : 6 : 6 : 1 1 : 9 : 9 : 9 : 9 : 9 : 9 : 1 2 ; a n t en n a l segmen t 3 more t h a n t h ree t i mes a s l on g as b road ( 4 , 4 ) ; h e a d s h i n y , h a i r y , g r a n u l a t e d ; frons w i t h o u t med i a n k e e l b e t ween t he a n t e n n a l socke t s ; P O L = 7 ; O L = 3 ; OOL = 3 ; b re a d t h o f t h e oce l l i : 4 ; temp l es d i s t i nc t ; occ i p i t a l c a r i n a i n v i s i b l e ; scu t u m g r a n u l a t ed , h a i r y , d u l l ; n o t a u l i ces d i s t i n c t a n d comp l e t e , poster i or l y j o i n t ; sc u t e l l u m a n d

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F i g . 836 - M a l e o f Pseu dogon a to[J u s sara w a k i Moczar f rom S r i L a n k a

met a n o t u m s h i n y , smoo t h , we a k l y p u n c t a t e ; p ropodeum d u l l , ret i c u l a t c r u gose ; fore- w i n g h y a l i n e , w i t h o u t d a rk t r a n s v ersa l b a n d s ; dors a l p rocess o f gonoforceps ( F i g . 838 ) w i t h a p e x po i n ted a n d w i t h l a t e r a l s u b a p i c a l po i n t ; m a x i l l a r y p a l p i w i t h 3-4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . L O C U S TYP I C US : P a y a h P a l o h ( S a r a w a k , M a l a y s i a ) TYP I CA L MAT E F< l A L : ho l o t y p e F ! o f Ps . s a ra vJ a k i i n BU ; h o l o t y pe F ! a n d p a r a t y p (� F ! o f Ps . [Jonomarenkoi i n BU . HOST S : i n S r i L a n k a a n d M a l a y s i a Nilapa rva t a lugen s S t a l a n d Soga tella s p . ; in t he P h i l i p p i nes Nilaparva ta lugen s SI a l a n d Toya a ttenua t a D i s ! . D I ST R I B U T I O N : O r i e n t a l reg i on : SR I LAN KA : K a l u g om u w a , O L ! G a n n oru w a ( K a n d y 0 i s t r . ) ( Po n o m a re n k o 1 98 1 b , u n d e r n a me Pseudogo n a topus ponomarenkoi Mocza r ) , WA ! T i sm a d a ( K a n . d i s ! . ) ( Po n o m a ren k o 1 98 1 b , u n d e r n a me Ps e u dogo n a t op us ponomarenkoi Mocz a r ) , WA ! I N O I A : M a d ras ( T a m i z h a g a n ) , ZO ! MALAYS I A : P a y a h P a l o h ( S a r a w a k ) , BU ! M a l a y s i a , O L !

G o n a topod i n a e

1 24 1

F i g . 8 3 7 - M a l e o f Pseu dogon a top u s sara1'la k i Moc z a r from S r i L a n k a

TA I WA N : Kose n , O L I P H I L I P P I N E S : Los B a nos ( L a g u n a ) , ( Be n g u e t , L u zo n ) , WA ! M a n i l a , KY !

BM !

Bag u i

2 . Pseudo gona top u s sarawa kensis n . s p .

F E MA L E : a p tero u s ; l e n g t h 5 , 00-5 , 3 1 m m ; h e a d b r o w n - b l ac k , w i t h fron t p a r t of v e r t e x ( more a l o n g o rb i t s ) , c l y p e u s a n d m a n d i b l es t e s t aceou s ; a n te n n a e brow n , w i t h seg m e n t s 8- 1 0 y e l l o w ; t h o r a x a n d p ro p o d e u m ferr u g i n o u s ; p et i o l e a n d a b domen b l ac k ; l eg s b row n ; head exc a v a ted , smoo t h , s h i n y , w i t ho u t scu l p t u re ; sc u t u m s h i n y , w i t h some w e a k l on g i t u d i n a l k ee l s ; s c u t e l l u m smoo t h , w i t ho u t sc u l p t u r e ; m e t a n o t u m t ra n sverse l y s t r i a t e , l a t e ra l l y p r o t r u d i n g ; l a t e r a l p r o t r u s i o n s r o u n d e d ; a n t e r i o r s u rf ace of m e t a t ho r a x + p ropode u m w i t h n u merous l on g i t u d i n a l k ee l s ; m e t a t h o r a x + p ropodeum s h i n y , w i t ho u t scu l p t u r e ; p o s te r i o r s u rf a c e of p ropodeum a n d p l e u r a t r a n sverse I y s t r i a t e ; meso- m e t a p I e u r a I s u t u re comp l e t e , wea k l y d i s t i n c t ; segme n t 1 of fron t t a r s u s s h o r t e r t h a n seg m e n t 4 ( 1 0 : 1 5 ) ; en l a r g e d c l a w ( F i g . 839 ) w i t h s u b a p i c a l t oo t h a n d 1 1 l a m e l l ae ; segmen t 5 of f ro n t t a rs u s ( F i g . 839 ) w i t h 2 rows of 1 7 + 5 l a me l l ae ; a p e x w i t h a g r o u p of a t l ea s t 20 l a m e l l ae ; m a x i I l a ry p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . M A L E : u n k no w n ( see notes ) L O C U S TYP I C U S : G u n o n g M u l u ( S a ra w a k , M a l a y s i a ) TYP I CA L MAT E R I A L : ho l o t y p e F ! a n d 1 p a r a t y p e F ! i n B M . HOST S : u n k n o w n D I ST R I BU T I O N : O r i e n t a l reg i o n : MA LAYS I A : G u no n g M u l u ( Sa r a w a k ) , NOTES :

BM !

i n BM t here i s a ser i es of m a l e s p ec i me n s of G o n a topod i n es fou n d

Mem . Amer .

1 242

F i g . 838

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I nst . ,

no.

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M a l e g e n i t a l i a of Pseu dogon a top u s sarawaki Moczar from S r i Lanka

b y m a l a i se t r a p s i n G u n o n g M u l u N a t i o n a l P a r k ( S a r a w a k ) . T h ese spec i me n s p a r t l y are m a l es of Agon a topoides bengalensis n . sp . ( w i t h m a x i l l a r y p a l p i w i t h 5 segmen t s a n d l a b i a l p a l p i w i t h 3 segmen t s ) . O t h e r spec i m e n s are m a l es of t wo new s p ec i es of Pseu dogon a top u s : m a lesiae a n d sara w a ken sis . T h e y h a v e m a x i l l a r y p a l p i w i t h 3-4 s e g me n t s a n d l a b i a l p a l p i w i t h 2 s e g me n t s . A ser i es o f t hese m a l es h a s n ot a u l i ce s p o s t er i or l y

1 243

Gon a topod i n ae

F i g . 839 - C he l a of Pseudogo n a topus sara"/a kensis n .

sp .

( type)

j o i n t a n d dors a l p rocess of gon oforceps w i t h a pex r o u n ded ( m a l e A ) ; a seco n d ser i es h a s not a u l i ces poster i o r l y s e p a r a ted a n d dors a l process of gonoforceps w i t h apex p o i n ted ( m a l e B ) . I t i s i m poss i b l e , w i t ho u t b reed i n g s , t o refer t hese m a l es t o m a lesiae o r sarawa kensis . Bo t h m a l es show t h e f o l l ow i n g morpho l og y : f u l l y w i n g ed ; l e n g t h 1 , 37-2 , 50 m m ; bod y b ro w n or b l ac k , w i t h a n te n n ae b ro w n or p a r t l y t e s t aceo u s ; l e g s y e l l o w ; a n t e n n a I segmen t s i n fo l l o w i n g p ropor t i o n s : 5 : 5 : 9 : 7 : 7 : 7 : 8 : 7 : 6 : 1 0 ( m a l e A ) ; 5 : 6 : 9 : 9 : 8 : 8 : 8 : 8 : 8 : 1 1 ( m a l e B ) ; a n t en n a l segmen t 3 i n b o t h m a l es more t h a n t h ree t i mes as l o n g a s b road

( m a l e A : 4 , 0 ; ma l e B :

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s p . or

3 , 66 ) ; h e a d d u l l , h a i r y , r u gose ; occ i p i t a l c a r i n a i n v i s i b l e ; t emp l es short , d i s t i n c t ; P O L = 5 ; Ol = 2 ; OOl = 2 , 5 ; b r e a d t h of t he oce l l i : 3 ; sc u t u m d u l l , h a i r y , p u nc t a t e ; n o t a u l i ce s com p l e t e , poste r i o r l y j o i n t ( m a l e A ) o r sep a r a ted ( m a l e B ) ; i n m a l e B m i n i m u m d i s t a n c e b e t ween t he

1 245

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sp . or

n o t a u l i ces s h o r t e r t h a n b re a d t h of t h e oce l l i ( 2 : 3 ) ; s c u t e l l u m a n d met a n o t u m s h i n y , h a i r y , p u n c t a t e , smoo t h ; p ropodeum re t i c u l a t e r u gose ; fore-w i n g h y a l i n e , w i t h o u t d a r k t ra n s ver s a l b a n d s ; dors a l p rocess of gonoforceps l o n g , b r o a d , w i t ; , a p e x rou n d e d ( m a l e A : f i g . 840 ) or po i n ted ( m a l e B : f i g . 841 ) ; m a x i I l a ;' y p a l p i w i t h 3-4 segmen t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u r s 1 , 1 , 2 .

3 . Pseu dogon a top u s n u du s R . C . L . P e r k i n s

Ps e u dogona topus n u dus R . C . L . P e r k i n s 1 9 1 2 :

12.

F E MAL E : a p terou s ; l en g t h 2 , 74 m m ; red d i s h - t e s t aceo u s , w i t h pet i o l e b l ac k a n d a b domen b ro w n - re d d i s h ; a n t e n n a e b r ow n , w i t h seg men t s 1 -2 y e l l ow ; b o d y h a i r l es s , s h i n y , g r a n u l a t ed ; h e a d e x c a v a ted ; p ro n o t u m c rossed b y a s t ro n g t r a n s v e rs a l i m p re s s i o n ; m e t a n o t u m w i t h s i des p r o t r u d i n g ; p ro t r u s i o n s rou n d e d ; meso-me t a p l e u r a l s u t u re o b so l e t e ; met a t ho r a x + p ropod e u m w i t h o u t med i a n f u rrow ; mesop l e u r a , met a p l e u r a a n d pos t e r i o r s u rface o f p ropodeum t r a n s verse l y s t r i a t e ; se g m e n t 4 of f ro n t t a r s u s a s l o n g a s segmen t 1 ; e n l a r g e d c l a w ( F i g . 842 ) w i t h s u b a p i c a l too t h a n d 6 l a me l l a e ; s e g m e n t 5 of f ro n t t a r s u s ( F i g . 842 ) w i t h 2 rows of 1 4 l a m e l l ae ; apex w i t h a g r o u p o f 6 l a me l l ae ; m a x i I l a r y p a l p i w i t h 4 seg men t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 .

1 246

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I n st . ,

no.

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F i g . 842 - C he l a of Pseudogo n a topus nudus R . C . L . Perk i n s ( t y p e )

M AL E : u n k n o w n L O C U S T Y P I C U S : P ek a l o n g a n ( J a v a , I n d o n es i a ) T Y P I CA L M A T E R I A L : l ec t o t y p e F ! ( d es i g n a ted b y O l m i 1 982 ) i n B . HOST S : i n I n dones i a Dicran otropis s p . ( R . C . L . Perk i n s 1 980 ) ; i n t h e P h i l i p p i nes Soga tella furcifera ( Ho rv a t h ) ( C h a n d r a 1 980b ; I . R . R . I . E n to mo l o g y D e p a r t m e n t 1 980 ) , Nil aparvata l u gens ( St a l ) , Stenocra n u s s p .

1 247

Gon a topod i n ae

F i g . 843

-

C he l a o f Pseu dogon a top u s ma lesiae n .

sp .

( type)

1 248

Mem . A m e r . E n ! .

I ns ! . ,

n o . 37 ,

1 984

( C h a n d r a 1 980b ) . Accord i n g to C h a n d r a ( 1 980b ) i n t he P h i l i p p i nes occ a s i o n a l l y Ps . n u dus c a n p a r a s i t i ze a l so C i ca de l l i d s : Nephotettix n igropictus ( S t a l ) , Nephotettix virescens ( D i s t a n t ) and Recilia dors a lis ( Mo t s c h u l s k y ) . T h i s l a s t i n forma t i o n s m u s t b e c h eck e d . D I ST R I BU T I O N : O r i e n t a l reg i on : I N D O N E S I A : P ek a l on g a n ( J a v a ) , B ! P H I L I P P I N E S ( C h a n d r a 1 980b ; I . R . R . I . E n tomo l o g y D e p a r t m e n t 1 980 ) .

4 . Pseu dogon a top u s m a lesiae n . sp .

F E M A L E : a p t e ro u s ; l e n g t h 3 , 0 m m ; t e s t a c eo u s , w i t h p e t i o l e b l ac k ; a n t e n n a e t e s t aceou s , w i t h segmen t 1 0 w h i t i s h ; head v e r y e xc a v a ted , s h i n y , smoo t h , w i t h o u t scu l p t u re , h a i r l ess ; p rono t u m crossed b y a s t ro n g t ra n s v ersa l i m p ress i o n , smoo t h , h a i r l es s , s h i n y ; s c u t u m a n d s c u t e l l u m s h i n y , smoo t h a n d w i t h o u t sc u l p t u re ; m e t a n o t u m smoo t h , s h i n y , w i t ho u t sc u l p t u re , w i t h s i des p ro t r u d i n g ; p ro t r u s i o n s po i n ted ; me t a t ho r a x + p ropode u m s h i n y , smoo t h , w i t h o u t sc u l p t u re , except for n u me ro u s t r a n s v e rs a l s t r i a e o n pos t e r i o r s u rface ; p l eu r a w i t ho u t t ra n sv e rs a l k ee l s ; meso- me t a p l eu r a l s u t u re d i s t i nc t a n d comp l e t e ; segmen t 1 of fron t t a rs u s s ho r t e r t h a n segmen t 4 ( 1 3 : 1 6 ) ; en l a r g e d c l a w ( F i g . 843 ) w i t h s U b a p i c a l tooth a n d 7 l a me l l ae ; seg m e n t 5 of fron t t a rs u s ( F i g . 843 ) w i t h 2 row s of 1 9 l a me l l a e ; a p e x w i t h a g ro u p of 1 5 l a m e l l a e ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k no w n ( see notes o n p a g e 1 24 1 ) . L O C U S T Y P I C U S : G u no n g M u l u ( Sa r a w a k , M a l a y s i a ) T Y P I CAL M A T E R I AL : ho l o t y p e F ! i n BM . HOST S : u n k n o w n D I ST R I B U T I O N : O r i e n t a l reg i on : M A LAYS I A : G u n o n g M u l u ( Sa r a w a k ) , BM !

5 . Pseu dogon a top u s validus n . sp .

F E MA L E : a p terou s ; l e n g t h 2 , 5 m m ; head t e s t aceo u s-redd i s h or b ro w n redd i s h , w i t h m a n d i b l e s , c l y p e u s a n d fron t p a r t of v e r t e x y e l l o w ; a n te n n a e b row n , w i t h s e g m e n t s 1 -2 y e l l o w ; t horax a n d p ropod e u m red d i s h or b ro w n-re d d i s h ; p e t i o l e b l ac k ; a b domen b ro w n-red d i s h ; l eg s y e l l ow ; head ( F i g . 844 C ) exca v a t e d , smoo t h , s h i n y , w i t ho u t sc u l p t u re ; p ron o t u m ( F i g . 8 4 4 B ) crossed b y a s t ro n g t r a n svers a l i m p ress i o n , smoo t h , s h i n y , w i t ho u t scu l p t u re ; sc u t u m a n d sc u te l l u m smoo t h , s h i n y , w i t h o u t scu l p t u re ; m e t a n o t u m n o t t r a n s verse I y s t r i a t e , w i t h s i des p r o t r u d i n g ; p ro t r u s i on s p o i n ted ; meso-me t a p l eu r a l s u t u re o n l y d i s t i nc t on pos t e r i o r h a l f ; m e t a t ho r a x + p ropodeum ( F i g . 844 A ) smoo t h , s h i n y , w i t ho u t sc u l p t u re ; mesop l e u r a , met a p l eu r a a n d pos t e r i o r s u rface of p ropodeum t r a n sverse l y s t r i a t e ; segmen t 1 of fro n t t a r s u s a s l o n g a s segme n t 4 ; en l arged c l a w ( F i g . 845 ) w i t h s u b a p i c a l t oo t h a n d 5 l a m e l l ae ; segmen t 5 of f ro n t t a rs u s

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c

Fig.

844 - F em a l e of Pse u dogon a top us validus n . s p . t h o r a x a n d p ropode u m ; C : he a d .

( type) : A , B :

( F i g . 845 ) w i t h t w o rows of 1 3 l a me l l a e ; apex w i t h a group of 9 l a m e l l ae ; m a x i I l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E : u n k no w n L O C U S TYP I C U S : Spooner R o a d ( M a l a y a , M a l a y s i a ) T Y P I CA L MAT E R I A L : h o l o t y p e F ! i n BM ; 1 p a r a t y p e F ! i n OL . HOST S : u n k n o w n D I ST R I B U T I O N : O r i en t a l reg i o n : SR I L A N KA : Y a k k a l a , O L ! MALAYS I A : Spooner R o a d ( K u a l a L u m p u r , M a l a y a ) , BM !

6 . Pseu dogon a top us l a n k a e P o n o m a r e n k o

Pseu dogon a top us lankae P o n o m a r e n k o 1 98 1 b : 8 8 1 .

F E MA L E : a p t e ro u s ; l e n g t h 3 , 32 m m ; r e d d i s h - b r o w n , w i t h a n t e n n a e tes t aceou s ; l eg s w i t h t roc h a n ters a n d fore c o x a e p a r t l y w h i t i s h ; h i n d s i d e of h i n d femora w h i t i s h ; a p e x of p ropodeum w h i t i s h ; h e a d d u l l , a l u t aceou s ; o c c i p i t a l c a r i n a i n v i s i b l e ; f ro n s w i t h a com p l e t e med i a n k ee l ; POL = 2 ; OL = 3 ; OOL = 1 0 ; head e x c a v a ted ; temp l es p rom i n e n t ; a n ten n a e d i s t a l l y t h i c k e n e d ; a n t e n n a I segmen t s i n fo l l o w i n g p ro po r t i o n s : 1 4 : 8 : 1 4 : 9 : 9 : 6 : 6 : 6 : 6 : 1 1 ; p r o n o t u m c rossed b y a s t ro n g t ra n s v ersa l i m p ress i o n , s h i n y , smoo t h , w i t ho u t scu l p t u r e ; scu t u m smoo t h , s h i n y , w i t ho u t scu l p t u r e , w i t h som" t r a n s v e r s a l kee l s o n s i des ; s c u te l l u m s h i n y , smoo t h , w i t ho u t scu l p t u re ; t ho r a x h a i r l es s ; met a n o t u m f i a t , wea k l y ho l l o w

Mem . Amer . E n ! .

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C h e l a of Pseudogon a top u s v a l idus n . sp .

( t y pe )

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F i g . 846 - T h o r a x a n d p ropodeum o f fem a l es o f Pseudogona topus mUlrl n . s p . ( t y pe ) ( A ) a n d l a n k a e Ponomarenko ( t y p e ) ( B )

b e h i n d t h e s c u te l l u m , smoot h , s h i n y , w i t h o u t scu l p t u re , w i t h s i de s protru d i n g ; p ro t r u s i o n s rou n ded ; meso-me t a p ! e u r a l s u t u re d i s t i nc t a n d comp l e te ; m e t a t horax + propode u m ( F i g . 8 4 6 B ) smoot h , s h i n y , w i t h o u t sc u l p t u re , excep t for n u merous t r a n s vers a l s t r i a e on poster i or s u rf ace of p ropode u m ; p l e u r a w i t h o u t t r a n s versa l s t r i a e ; d i sc of m e t a t horax + propodeum w i t h a s t ro n g med i a n f u r row ; fore t a r s a l s e g me n t s i n fo l l ow i n g proport i on s : 1 5 : 4 : 6 : 20 : 3 1 ; en l arged c l a w ( F i g . 847 ) w i t h s U b a p i c a l too t h a n d 8 l ame l l a e ; segment 5 of fron t t a rs u s ( F i g . 847 ) w i t h t w o row s of 1 1 + 1 0 l ame l l ae ; a p e x w i t h a g ro u p of n e a r l y 20 l a me l l a e ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E : u n k no w n LOCUS TYP I CUS : G i l i m a l e ( S r i Lan k a ) TYP I CAL M A T E R I A L : h o l o t y p e F ! i n W A . HOST S : u n k no w n D I ST R I B U T I O N : O r i en t a l reg i on : SR I LAN K A : G i l i m a l e ( I n d u r u w a J u n g l e , R a t . D i s t . ) , WA ! W a t u r a w a Forest ( 5 i n h a r a j a J u n g l e , R a t . D i s t . ) , OL ! N OT E S : Ps . l a n k a e i s very s i m i l a r to Ps . ma lesia e : t h e on l y d i fference i s t h e a b sence o f med i a n f u rrow o n m e t a t horax + propo d e u m .

7 . Pseu dogon a top u s m u iri n .

sp .

F E M AL E : a p t e ro u s ; l e n g t h 2 , 5 m m ; red d i s h - t e s t a ceou s , w i t h pe t i o l e b l ac k a n d a b domen b ro w n -red d i s h ; a n te n n a e b row n , w i t h segmen t s 1 -2 y e l l ow ; head du I I , e x c a v a ted , weak I y g r a n u I a ted ; p ro n o t u m crossed b y a s t ro n g t r a n s v e r s a l i m p ress i o n , s h i n y , w i t h o u t sc u l p t u r e ; m e t a n o t u m s h i n y , w i t h s i des p r o t r u d i n g ; p ro t r u s i o n s rou n ded ( F i g . 846 A ) ; meso-met a p l eu r a l s u t u re w e a k l y d i s t i nc t ; p o s t e r i o r s u rface of p ropod e u m t r a n s verse l y

Mem . Amer.

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F i g . 847 - C h e l a of Pseu dogon a top u s l a n k a e P o n o m a r e n k o ( t y pe )

striate;

segme n t 1 of fron t t a r s u s as l o n g as s e g m e n t 4 ; en l a r ged c l a w ( F i g . 848 ) w i t h s u b a p i c a l too t h a n d 6-7 l a m e l l ae ; seg m e n t 5 of fro n t

1 2 53

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F i g . 848

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C h e l a of Pse u dogoll a top u s m u iri n . s p .

( type )

1 254

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I ns ! . ,

n o . 37 ,

1 984

F i g . 849 - C h e l a of Pseudogo n a topus soga t a e R o h w e r ( t y p e )

t a r s u s ( F i g . 848 ) w i t h 2 rows o f 1 5- 1 7 l a m e l l ae ; a p e x w i t h a g ro u p of 1 1 - 1 2 l a me l l ae ; m a x i l l a r y p a l p i w i t h 3 segme n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E : u n k no w n L O C U S TYP I CU S : Pek a l on g a n ( J av a , I n do n es i a ) TYP I CA L M A T E R I A L : ho l ot y pe F ! i n OL . HOST S : u n k n o w n D I S T R I B UT I O N : O r i e n t a l reg i on : I N D O N E S I A : Pek a l on g a n ( Ja v a ) , OL ! N O T E : t h e spec i es i s n a m e d i n honor of M r . F . M u i r .

8 . Pseu dogona topus hospes R . C . L .

Perk i n s

See A u s t ra l i a n spec i es

9 . Pseudogo n a topus soga t a e R o h w e r n . eme n d .

Pseu dogon a top u s soga tea Rohwer 1 920 : 1 60 ( soga tea i s p r i n t i n g -error : t h e t y p e i s l a b e l l ed soga tae ) .

1 25 5

Gona topod i n a e

F E MA L E : a p terou s ; l e n g t h 3 , 0 m m ; head b row n , w i t h c l y pe u s a n d m a n d i b l e s y e l l ow ; a n t e n n a e b row n , w i t h segmen t 1 y e l l ow a n d 2 - 3 l i g h t ; t horax a n d p ropodeum b ro w n , w i t h sc u t u m redd i s h a n d a p e x of p ropodeum y e l l o w ; p e t i o l e b l ac k ; a b domen b ro w n - d a r k ; l eg s y e l l ow - b row n ; b o d y ha i r l es s ; h e a d e xc a v a ted , s h i n y , g r a n u l a ted ; p ro n o t u m crossed b y a s t r o n g t r a n sversa l i m p ress i on , weak l y g r a n u l a ted ; s c u t u m d u l l , g r a n u l a ted ; m e t a n o t u m f l a t , t r a n sverse l y s t r i a t e , w i t h s i des ro u n ded ; m e t a t h o r a x + p ropodeum d u l l , as scu l p t u red as sc u t u m ; mesop l e u r a , m e t a p l e u r a a n d poste r i or s u rface of p ropodeum t r a n sverse l y s t r i a te ; meso-me t a p l eu r a l s u t u re obso l e t e ; segmen t 1 of fro n t t a r s u s s l i g h t l y l on ge r t h a n s e g me n t 4 ( 1 0 : 8 ) ; en l arged c l a w ( F i g . 849 ) w i t h s u b a p i ca l too t h a n d 4 l a me l l a e ; seg m e n t 5 of fro n t t a r s u s ( F i g . 849 ) w i t h 2 ro w s of 23 l a me l l ae exten d i n g con t i n uo u s l y to the a p e x ; m a x i I l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E : u n k n o w n ( t h e m a l e a l l ot y p e desc r i b e d b y Rohwer 1 920 i s rea l l y a m a l e of Haplogon a top u s orien t a l i s Rohwer : i t h a s m a x i I l a r y p a l p i w i t h 2 seg m e n t s a n d l a b i a l p a l p i w i t h 1 segme n t ) . LOCUS TYP I CU S : Pusa ( I n d i a ) T Y P I CAL MAT E R I AL : ho l o t y p e F ! i n WA ( to g e t h e r w i t h t h e a l l o t y p e M ! n o t b e l o n g i n g to t h i s s p ec i es ) . HOST S : i n I nd i a Soga ta s p . ( Ro h w e r 1 920 ) . D I ST R I BU T I O N : O r i en t a l reg i on : I N D I A : P u s a ( B i h a r ) , WA ! C u t t ack ( O r i ss,a ) ( K u r i a n 1 954b ) .

1 0 . Pseu dogon a top u s p u s a n u s n .

F E MA L E :

a p terou s ;

vertex b row n ;

l en g t h 3 , 0-3 , 5 m m ;

abdomen b r ow n ;

sp .

test aceous-red d i s h ;

head w i t h

a n t e n n a I segmen ts 3- 1 0 b row n ;

s i des of

sc u t u m a n d m e t a n o t u m d a r k ; a n t e n n a e d i s t a l l y t h i c k e n e d ; a n t en n a l s e g m e n t s i n fo l l ow i n g p roport i on s : 8 : 7 : 1 5 : 8 : 7 : 7 : 7 : 7 : 6 : 9 ; head exc a v a t e d , d u l l , g r a n u l a t e d ; frons w i t h a com p l et e med i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L � 3 ; O L � 4 ; OOL � 9 ; p ron o t u m crossed b y a s t ro n g t r a n sv e r s a l i m p ress i on , d u l l , g r a n u l a ted ; scu t u m d u l l , g r a n u l a ted ; met a n o t u m f l a t , t r a n s verse l y s t r i a t e , w i t h s i des r o u n d e d ; m e t a t horax + p ro p o d e u m d u l l , g r a n u l a ted , w i t h poster i or s u rface a n d p l e u ra t r a n sverse l y s t r i a t e ; meso-meta p l e u r a l s u t u re obso l e te ; fore t a rsa l seg m e n t s i n fo l l o w i n g p ropor t i on s : 1 9 : 2 : 6 : 1 4 : 23 ; e n l a rged c l a w ( F i g . 850 ) w i t h s U b a p i c a l tooth a n d 4 l a me l l a e ; s e g m e n t 5 of fron t t a rs u s ( F i g . 850 ) w i t h 3 rows of 1 2 l a me l l ae ; a pex w i t h a g ro u p of 6 l a me l l a e ; m a x i I l a r y p a l p i w i t h 3 s e g me n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n LOCUS TYP I CU S : Pusa ( I n d i a ) T Y P I C A L M A T e R I AL : h o l o t y p e F ! a n d 1 p a r a t y p e F ! i n N O . HOS T S :

u n k no w n

M e m . Amer . E n t .

1 256

D I STR I BU T I O N : O r i e n t a l reg i o n :

F i g . 850

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I nst. ,

n o . 37 ,

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I ND I A : Pusa ( B i h a r , NO !

C h e l a of Pseudogona top u s p u s a n us n . s p .

( p ar a t y p e )

Go n a topod i n a e

1 257

NEARCT I C SPEC I E S O F PSEUDOGONA TOPUS

E i g h t s p ec i es are k n ow n .

KEY T O T H E SPEC I ES

F EMALES

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B o d y r ed d i s h ; a n t e r i o r s u r f a c e o f me t a t h o ra x + p r op o d e u m mo r e i n c l i n ed ( F i g . 8 5 5 B ) ; s e gmen t 4 o f f r o n t t a r s u s a s l on g a s o r s l i g h t I y s h o r t e r t h a n s eg m e n t 1 ( mo r e t h a n 8 5 % ) . . . . . . . . . . . . . . • • • . . . • • . . . . • • • • • . . • • • • . . • • • • • 3 . sjoes tedti ( K i e f f e r ) B od y y e l l o w - r e d d i s h ; a n t e r i o r su r f a c e o f me t a t h o r a x + p r op o d e u m l e s s t a r s u s sh o r t e r

5

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. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 . freytagi n e w n a me Me s o - me t a p l eu ra l su t u r e ob s o l e t e . . . . . . • • . • • . • . . . . • • . • • • . • • • . . 2 T h o r a x a n d p r op od e u m f u l l y b l a c k ( a t m o s t a p ex o f p r o p od eu m I i g h t ) • • • • • • • • . • • • • • • • • • • • • • • • 2 . c a n a densis n . s p . T h o r a x a n d p r op od e u m f u I I Y o r p a r t I y r edd i s h o r t e s t a c e ou s . • . 3 B o d y y e l l ow - t e s t a c e ou s o r r e d d i s h , w i t h p e t i o l e b l a c k ; o c c a s i on a l l y a b d omen b r own o r p a r t l y b l a c k ; o c c a s i on a l l y h e a d w i t h v e r t ex b r own ; o c c a s i on a l l y a n t e n n a e p a r t l y b r own o r b l a c k

8 5 8 8 ) ; s eg m e n t 4 o f f r on t ( l e s s t han 8 0 % ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 . arizonicus R . C . L . P e r k i n s Me t a t h o r a x + p r op od e u m r e d d i s h , w i t h b r own i r r eg u l a r s p o t s . . . . . . . . . . . . . . . . . . . . . . . . . 5 . ontarianus n . sp . Me t a t h o r a x + p r op od e u m b l a ck , a t mo s t w i t h a p ex o f p r op od e u m I i g h t a n d w i t h me t a n o t u m p a r t l y r ed d i s h • . • . . . . . • • • • . • • . . . • . . 6 S e g me n t 4 o f f r o n t t a r su s l on g e r t h a n 1 • • • • . . • • • • • • . • • . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6 . s tenocra n i R . C . L . P e rk i n s S e g m e n t 4 o f f r o n t t a r s u s s h o r t e r t h a n s e gmen t 1 • • • • • . . . ' " • • 7 Scu t um b l a ck ; 0 , 7 t i me s a s l . . . . . . . . . . S c u t u m y e l l ow ; 0 , 7 5 t i me s a s . . . . . . . . . . . . . .

i n c l i n ed ( F i g . t h a n s eg m e n t 1

s eg m e n t 4 o f f r o n t t a r s u s s h o r t e r ( a p p r ox i ma t e l y on g a s s eg m e n t 1 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 . a u toxen o b i u s R . C . L . P e rk i n s s eg m e n t 4 o f f r o n t t a r s u s l o n g e r ( mo r e t h a n l on g a s s e gmen t 1 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 . m a gn u s B r own

Mem.

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I n s! . ,

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MALES

2

3

D o r s a l p r o c e s s o f g on o f o r c e p s mu c h s h o r t e r t h a n g o n o f o r c ep s ( F i g s 860 , 863 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 D o r s a l p r o c e s s o f g on o f o r c ep s n ea r l y a s l o n g a s g on o f o r c ep s ( F i g s 854 , 857 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 T h o r a x a n d p r op od eu m b l a c k • • • • 6 . sten ocra n i R . C . L . P e rk i n s T h o rax a n d p r op od e u m b r own - r ed d i s h . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 . arizonicus R . C . L . P e rk i n s D o r s a l p r o c e s s o f g o n o f o r c ep s w i t h a p ex p o i n t ed ( F i g . 8 5 7 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 . sjoestedti ( K i e f f e r ) D o r sa l p r o c e s s o f g on o f o r c ep s w i t h a p ex r o u n d ed ( F i g . •

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T h e ma l es of t he o t h e r spec i es are u n k no w n .

1 . Pseu dogon a top us freytagi n e w n a me

Pseu dogona top u s similis F r e y t a g 1 978 : Pseu dogon a top u s similis ( B r u e s 1 906 ) .

1 39 ;

p reocc u p i ed b y

F E MA L E : a p terou s ; l e n g t h 1 , 0-4 , 0 m m ; b ro w n or b ro w n - y e l l ow , w i t h p e t i o l e b l ac k ; a n t e n n ae b ro w n - b l ac k , w i t h seg m e n t 1 -2 a n d 1 0 y e l l ow o r w h i te ; m a n d i b l es a n d l eg s y e l l o w ; h e a d s h i n y , exc a v a t ed , smoo t h , g r a n u I a t ed ; prono t u m crossed b y a s t ro n g t r a n sversa I i m p ress i o n , smoo t h , s h i n y , w i t ho u t sc u l p t u re ; s c u t u m s h i n y , smoo t h , w i t ho u t scu l p t u re , w i t h some t ra c k s of k ee l s ; sc u t e l l u m s h i n y , smoo t h , w i t ho u t sc u l p t u r e ; met a no t u m w i t ho u t t ra n s v e r s a l s t r i a e , w i t h s i des p r o t r u d i n g ; meso m e t a p l e u r a l s u t u re d i s t i n c t ; p l eu r a and p o s te r i o r s u rface of p ropodeum t r a n sverse l y s t r i a t e ; seg m e n t 1 of fron t t a rs u s a s l o n g as segmen t 4 ; en l arged c l a w ( F i g . 8 5 1 ) w i t h s u b a p i c a l t oo t h a n d 7-9 l a me l l ae ; segme n t 5 o f f ro n t t a r s u s ( F i g . 85 1 ) w i t h t w o rows o f 8 + 1 0 l a me l l a e ; apex w i t h a g ro u p of 1 3 l a me l l a e ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : Lex i n g to n ( K y . , U . S . A . ) TYP I C A L M A T E R I AL : h o l o t y p e F a n d 1 0 p a r a t y p e s F F i n WA ; 7 p a r a t y pes FF i n K E . HOST S : i n U . S . A . Delp h a codes l u tulenta ( V a n D u zee ) ( F re y t a g 1 978 ) D I ST R I BU T I O N : Nearct i c reg i o n : U . S . A . : 29 . 1 m i . N of M i n era l W e l l s ( J a c k C o . , Tex . ) , BY ! 2 . 8 m i . 5 o f Somerv i l i e ( W a s h i n g ton Co . , Tex . ) , O L ! Lex i n g to n ( K y . ) , O L ! C M ! WA ! KE !

•

Gon a topod i n a e

Fig.

851

1 259

- C he l a of Pseu dogon a top u s freytagi n . n a me f rom L e x i n g t o n ( Ken t u c k y )

N O T E S : Ps . freytagi w a s reare d b y F re y t a g ( 1 978 ) con t i n u o u s l y i n l a bo r a t o r y for o n e y e a r . T he l y to k u s rep ro d uc t i on occ u r s i n t h i s s p ec i e s .

1 260

Mem . Amer . E n ! .

I ns ! . ,

no . 37 ,

1 984

F i g . 852 - T horax and p ropodeum of fem a l es of Pseudogo n a top u s s te n ocra n i R . C . L . Perk i n s from P r i nce A l bert ( Sa s k a t c h e w a n ) ( A ) a n d c a n a densis n . s p . ( t y p e ) ( B ) .

A p p ro x i m a te l y 30 d a y s a re n eeded to com p l e t e a I i f e c y c l e from e g g to a d u l t . R e a r i n g con d i t i o n s w e re u s u a l l y m a i n t a i n ed at 3 2 ° C a n d a 1 6-hour day. T h e spec i es i s n a m e d i n honor o f M r . P a u l H . Frey t a g .

2 . Pseudogona top u s c a n a densis n . s p .

F E M A L E : a p tero u s ; l en g t h 2 , 8 1 -3 , 00 m m ; b l ac k ; head w i t h m a n d i b l es , c l y pe u s a n d a n te r i or reg i o n of frons ( more a l o n g orb i t s ) tes t aceou s ; a n t en n ae brown , w i t h s e g m e n t s 1 -'7. testaceo u s ; l eg s test aceou s , w i t h fore c l u b s of femora a n d fore t i b i ae brow n ; a n te n n ae d i s t a l l y t h i ck e n e d ; a n ten n a l segmen t s i n fo l l ow i n g proport i o n s : 7 : 6 : 1 1 : 8 : 7 : 7 : 6 , 5 : 6 : 5 , 5 : 9 ; h e a d d u l l , g r a n u l a t e d , e x c a v a ted ; fro n s w i t h med i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L ; 2 ; OL ; 3 ; OOL ; 9 ; p ro n o t u m crossed by a s t ron g t r a n s vers a l i mp ress i on , d u l l , w e a k l y g ra n u l a ted ; sc u t u m d u l l , g r a n u l a t ed ; m e t a n o t u m t r a n s verse l y s t r i a t e , w i t h s i des p rotr u d i n g ; p ro t r u s i on s rou n ded ( F i g . 852 B ) ; me t a t horax + propodeum d u l l , g r a n u l a t e d , w i t h n u merous s t ron g t r a n s versa l s t r i a e on poster i or s u rface ; p l e ura w i t h weak t r a n sversa l s t r i ae ; d i sc a n d a n ter i or s u rface of m e t a t horax + p ropodeum w i t h a s t ron g track of med i a n f u r row ; meso-

Gon a topod i n a e

F i g . 8 5 3 - C h e l a of Pseu dogon a top u s c a n a densis n .

1 26 1

sp .

( type)

m e t a p l e u r a l s u t u re o b so l e t e ; fore t a rs a l seg m e n t s i n fo l l ow i n g p roport i o n s : 1 5 : 3 : 5 : 1 1 : 1 7 ; e n l a r ged c l a w ( F i g . 853 ) w i t h s u b a p i c a l t oo t h a n d 4-5 l a m e l l ae ; segme n t 5 of fron t t a rs u s ( F i g . 853 ) w i t h t wo row s of 1 5- 1 7 l a m e l l ae ; a p e x w i t h a g ro u p o f 6-1 0 l a me l l ae ; m a x i I l a r y p a l p i w i t h 4

Mem . Amer . E n ! .

1 262

Fig.

I n s! . ,

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1 984

854 - M a l e g en i t a l ia of Pseu dogon a top u s c a n a densis n. C a l dor ( C a l i for n i a )

sp . from

segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : f u l l y w i n g e d ; l e n g t h 2 , 4 m m ; b l ac k ; m a n d i b l es y e l l o w , w i t h t ee t h b ro w n ; a n t e n n a e a n d l e g s b row n ; a n t e n n a e n o t d i s t a l l y t h i ck e n e d ; a n te n n a l s e g me n t s i n fo l l o w i n g p ropor t i on s : 7 : 5 : 1 1 : 1 2 : 1 2 : 1 2 : 1 0 : 1 1 : 1 1 : 1 2 ; a n t en n a l segmen t 3 more t h a n t h ree t i mes a s l o n g a s b ro a d ( 1 2 : 3 ) ; h e a d s h i n y , g r a n u l a t ed ; f r o n s w i t ho u t m ed i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 9 ; O l = 5 ; OOl = 4 ; temp l es d i s t i nc t ; occ i p u t conca v e ; s c u t u m d u l l , g ra n u l a ted ; n o t a u l i ce s comp l e t e , p o s t er i or l y sep a r a t e d a n d very n ea r ; scu t e l l u m a n d m e t a n o t u m s h i n y , smoo t h , w i t ho u t scu l p t u re or v e r y w e a k l y p u n c t a t e ; p ropodeum d u l l , ret i c u l a t e r u g o s e ; fore-w i n g h y a l i n e , w i t ho u t d a r k t ra n s v e rs a l b a n d s ; d i s t a l p a r t of ra d i a l v e i n m u c h l on g e r t h a n prox i m a l p a r t ; r a d i a l c e l l open ; dorsa l p rocess of g o n oforceps ( F i g . 854 ) n ea r l y a s l o n g as gonoforc ep s , very b roa d , w i t h d i s t a l reg i on s h o w i n g p a p i I l a e ; d i s t a l m a rg i n serra t e ; m a x i I l a r y p a l p i w i t h 2 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . l O C U S T Y P I C U S : Red Deer ( A l be rt a , C a n a d a ) T Y P I CAL MAT E R I A L :

h o l ot y p e F !

and 1

p a r a t y pe F !

i n OT ;

3 p a r a t ypes

1 26 3

Gon a topod i n ae

F i g . 855 - F e m a l e of Pseu dogo n a topus sjoestedti ( K i effer ) ( t y p e ) : A , t ho r a x a n d p ropod eu m ; C : hea d .

B:

F F ! i n T E ; 1 parat y p e F ! i n AM ; 2 parat ypes ( 1 F , 1 M ) ! i n O L . HOST S : u n k n o w n D I ST R I BU T I O N : Nearc t i c reg i on : CANADA : N o r m a n W e l l s ( N . W . T . ) , a T ! R e d Deer ( A l t a . ) , O T ! U . S . A . : C a l dor ( E I Dora do Co . , C a l i f . ) , OL l T E ! S a n S i meon ( Sa n L u i s Ob i spo C o . , C a l i f . ) , A M !

3 . Pseudogo n a topus sjoestedti ( K i effer )

n . com b .

Gon a top u s sjoestedti K i effer 1 904b : 525 . Pseu dogon a top u s american us R . C . L . Perk i n s 1 905 : 37 ( n . s y n . ) ; locus typicus C o l u m b u s ( O h i o , U . S . A . ) .

F E M A L E : a p t e rou s ; l e n g t h 3 , 47-3 , 56 m m ; red d i s h , w i t h pe t i o l e b l ac k ; a b domen p a r t l y b ro w n i s h ; head ( F i g . 855 C ) exca v a ted , s h i n y , g ra n u l a t e d ; p rono t u m ( F i g . 855 A , B ) s h i n y , crossed b y a s t ro n g t r a n svers a l i m p ress i o n , g r a n u l a ted ; sc u t u m a n d s c u te l l u m s h i n y , smoot h ; m e t a n o t u m t r a n sverse l y s t r i a te , w i t h s i des rou n d e d ; meso-me t a p l e u r a l s u t u re obso l et e ; m e t a t ho r a x + propode u m ( F i g . 855 A , B ) g r an u l a ted , w i t h

Mem . Amer . E n t .

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Gon a topod i n a e

1 265

F i g . 857 - M a l e gen i ta I i a of Pseu dogon a top u s sjoestedti ( K i effer ) from L e x i n g ton ( K e n t u c k y ) ( r i g h t h a l f removed )

t r a c k of med i a n f u r row ; a n t e r i or s u rf ace v e r y i n c l i n ed ; p l eu r a not t r a n s v erse l y s t r i a t e ; pos t e r i o r s u rf a c e of p ropodeum t ra n s v erse l y s t r i a t e ; segmen t 4 o f fron t t a r s u s a s l o n g a s o r s l i g h t I y shorter t h a n segmen t 1 ( 1 4 : 1 6 i n h o l o t y p e of Ps . sjoes tedti ; 29 : 34 i n h o l o t y p e of Ps . america n u s ) ; e n l arged c l a w ( F i g . 856 ) w i t h s u b a p i c a l t oo t h a n d 4-7 l a me l l ae ; segmen t 5 of fron t t a rs u s ( F i g . 856 ) w i t h t wo rows of 1 1 -24 l a me l l ae ; a p e x w i t h a g ro u p of 6-1 1 l a me l l ae ; m a x i l l a r y p a l p i w i t h 4 s e g me n t s ; l ab i a l p a l p i w i t h 2 s e g m e n t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E : f u l l y w i n ged ; l e n g t h 1 , 8 m m ; b l ac k ; c l y p e u s t e s t a ceou s ; m a n d i b l es y e l l ow ; a n ten n a e , a b domen a n d l eg s b row n ; t a r s i y e l l o w ; h e a d s h i n y , w e a k l y g ra n u l a ted ; frons w i t hou t med i a n k ee l ; occ i p u t conc a v e ; occ i p i t a l c a r i n a i n v i s i b l e ; tem p l es d i s t i n c t ; POL � 7 ; O L = 3 ; OOL � 3 ; sc u t u m d u l l , g ra n u l a ted ; not a u l i ces com p l e t e , pos t e r i or l y s e p a r a t e d ; sc u t e l l u m a n d met a n o t u m s h i n y , smoo t h , w i t h o u t sc u l p t u r e ; p ropodeum w e a k l y ret i c u l a t e ru gose , w i t ho u t t r a n s v e rsa l or l o n g i t u d i n a l k ee l s ; fore- w i n g h y a l i n e , w i t ho u t d a rk t r a n sv e rs a l b a n d s ; r a d i a l c e l l open ; d i s t a l p a r t of rad i a l v e i n l on g e r t h a n p rox i m a l p a r t ; dors a l p rocess of gonoforceps ( F i g . 857 )

w i t h a p e x p o i n ted ; m a x i I l a ry p a l p i w i t h 4 segmen t s ;

l ab i a l

1 266

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c

Q;J}

F i g . 858 - F e m a l e of Ps e u do go n a t op u s arizonicus R . C . L . P e r k i n s from Mon t gomery ( Georg i a ) : A , B : t horax a n d propode u m ; C : h ea d .

p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u r s 1 , 1 , 2 . L O C U S TYP I C U S : T e x a s ( U . S . A . ) TYP I CA L MAT E R I AL : h o l o t y p e F ! of G . sjoestedti i n ST ; l ec t o t y pe F ! ( d es i g n a t e d b y O l m i 1 982 ) of Ps . cunerica n u s i n 3 . HOST S : i n U . S . A . Liburnia sp . ( R . C . L . P e r k i n s 1 906c ) D I ST R I B U T I ON : N e a rc t i c reg ion : U . S . A . : Tex a s , ST ! Waco ( Mc L e n n a n C o . , T e x . ) , O L ! BY ! Co l u m b u s ( O h i o ) , B ! L e x : n g ton ( K y . ) , K E ! H i g h l a n d s ( N . C . ) , O T ! Sea C l i ff ( Lo n g I s l a n d , N . Y . ) , C M !

4 . Pseu dogon a top u s arizon icus R . C . L . P e r k i n s

Pse u dogon a top u s arizonicus R . C . L . P e r k i n s 1 907 : 3 6 . F E MA L E : a p t e ro u s ; l en g t h 2 , 1 9-2 , 40 m m ; y e l l o w - t e s t aceo u s , w i t h p e t i o l e b l ac k ; occa s i on a l l y a b domen b row n ; h e a d ( F i g . 858 C ) exc a v a t e d , s h i n y , g r a n u l a ted ; p ron o t u m ( F i g . 858 A , B ) crossed b y a s t ro n g t ra n sversa l i m p re ss i on , s h i n y , g r a n u l a t e d ; m e t a n o t u m w i t h s i d es rou n d e d ; m e t a t h o r a x + propo d e u m ( F i g . 858 A , B ) s h i n y , g r a n u l a ted ; a n t er i o r s u rface of m e t a t ho r a x + p ropod e u m v e r y i n c l i ne d ; p l e u r a not t r a n svers e l y s t r i a t e ; pos t e r i o r s u rface o f p ropod e u m t r a n s v e rse l y s t r i a t e ; meso-me t a p l eu r a l s u t ure o b so l e t e ; seg m e n t 4 of fron t t a rs u s shorter t h a n segmen t 1 ( 20 : 25 i n h o l o t y p e ; 9 : 1 2 i n a s p e c i men from Mon t g omery ) ; e n l a rg ed c l a w ( F i g . 859 ) w i t h s u b a p i c a l too t h a n d 4-5 l a m e l l ae ; segmen t 5 of f r o n t t a r s u s ( F i g . 859 ) w i t h t wo row s of 1 0- 1 7 l a me l l ae ; a p e x w i t h a g ro u p o f 5- 1 0 l a m e l l ae ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ;

Gon a topod i n ae

F i g . 859

-

C he l a of Pseudogona topus arizonicus R . C . L . Mon t gomery ( Georg i a )

1 267

Perk i n s from

1 268

Mem . Amer . E n t .

I n st . ,

no .

37 ,

1 984

F i g . 860 - M a l e g en i t a l i a o f Pseu dogona topus arizon icus R . C . L . from N og a l es ( A r i zo n a )

Perk i n s

t i b i a l s p u rs 1 , 0 , 1 . MALE : f u l l y w i n g e d ; l e n g t h 2 , 3 m m ; b ro w n-red d i s h , w i t h l eg s ye l l ow ; a n t e n n a e n o t d i s t a l l y t h i ck e n ed ; a n t e n n a I segme n t s i n f o l l o w i n g propor t i o n s : 5 : 6 : 1 0 : 1 0 : 1 0 : 1 0 : 9 : 8 : 8 : 1 0 ; a n ten n a l s e g m e n t 3 n e a r l y f i ve t i mes a s l o n g a s b roa d ; head s h i n y , weak l y g r a n u l a ted ; t em p l es s hor t , b u t d i s t i nc t ; occ i p i t a l ca r i n a i n v i s i b l e ; fron t a l l i n e weak , b u t d i s t i nc t ; P O L = 7 ; OL = 2 ; O O L = 3 , 5 ; s c u t u m weak l y g r a n u l a ted ; not a u l i ces com p l et e , poster i or l y s e p a r a ted ; m i n i m u m d i s t a n c e b e t ween t h e n o t a u l i ces s h o r t e r t h a n b rea d t h of t h e oce l l i ( 2 : 3 ) ; s c u te l l u m a n d m e t a no t u m s h i n y , smoo t h , w i t h o u t scu l p t u r e ; p ropode u m ret i c u l a t e ru gose ; a n t e r i or s u rface w i t h m ed i a n f u rrow ; fore- w i n g h y a l i n e , w i t ho u t d a r k t r a n svers a l b a n d s ; d i s t a l p a r t of r a d i a l ve i n l o n g e r t h a n p rox i ma l p a r t ( 20 : 1 1 ) ; dors a l process of gonoforceps ( F i g . 860 ) very shor t , m u c h shorter t h a n g o noforce p s ; m a x i l l a r y p a l p i w i t h 4 s e g me n t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 1 , 2 . LOCUS T Y P I C U S : Nog a l es ( A r i z . , U . S . A . )

1 26 9

G o n a topod i n a e

F i g . 861

-

C he l a of Pseu dogon a top u s ontari a n u s n .

sp.

( type)

T Y P I CAL M A T E R I A L : l ec t o t y p e F ! a n d 2 p a ra l ec t o t y pes ( 1 F , 1 M ) ! ( d es i g n a te d b y O l m i 1 98 2 ) i n B . HOST S : i n U . S . A . Stoba era s p . ? ( R . C . L . P e r k i n s 1 90 7 ) a n d Sa cch arosydne s a ccharivora . D I S T R I BU T I ON : N e a rc t i c reg i o n : U . S . A . : Nog a l es ( A r i z . ) , B ! F e l l sm e re ( F l a . ) , O L ! WA ! S a p e l o I s . ( G a . ) , O L !. WA ! M o n t g o m e r y ( G a . ) , I T !

5 . Pseu dogon a top u s on taria n u s n .

sp .

F E MAL E : a p t e rou s ; l e n g t h 3 , 1 8-3 , 62 m m ; h e a d b ro w n , w i t h occ i p u t , c l y p e u s , m a n d i b l es a n d fro n t p a r t of v e r t e x ( more a l o n g o rb i t s ) t es t aceous or redd i sh ; a n ten n ae b row n , w i t h seg m e n t s 1 -2 t e s t a c eo u s ; p rono t u m red d i s h - d a r k ; sc u t u m w i t h pos t e r i o r h a l f redd i s h ; a n te r i or h a l f brow n ; s c u t e l l u m b l ac k ; m e t a t ho r a x + p ro p o d e u m red d i s h-d a rk o r red d i s h - b ro w n , w i t h a p e x t e s t aceo u s ; p e t i o l e b l ac k ; a b domen red d i s h d a rk ; l e g s t e s t aceo u s-red d i sh , w i t h c o x a e and c l u b s of femora p a r t l y b ro w n ; a n te n n a e d i s t a l l y t h i ckened ; a n t en n a l seg men t s i n f o l l ow i n g p ro p o r t i on s : 9 : 5 : 1 4 : 8 : 7 : 7 : 6 : 6 : 5 : 9 ; h e a d e x c a v a ted , g r a n u l a ted , d u l l ; f r o n s w i t h a med i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 1 , 5 ; OL = 3; O O L = 8; p ro no t u m c rossed b y a s t ro n g t r a n s v e r s a I i m p res s i on , d u I I , g r a n u l a te d ; sc u t u m d u l l , g r a n u l a t ed ; scu t e l l u m s h i n y , smoo t h ; m e t a n o t u m r u g ose , w i t h s i des p r o t ru d i n g ; p ro t r u s i on s rou n ded ; m e t a t h o r a x + p ro p o d e u m d u l l , g r a n u l a t ed , w i t h n u merou s t r a n s v e rs a l s t r i ae on p l eu r a a n d p o s t e r i o r s u rf a c e ; meso-me t a p l eu r a l s u t u r e obso l e t e ; m e t a t ho r a x +

1 270

Fig.

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n o . 37 ,

1 984

862 - C h e l a of Pseudogona topus stenocra n i R . C . L .

Perk i n s from O h i o

propo d e u m v e r y h u mped , a s i n P a l aearc t i c Gon a top u s l u n a tus K l u g ; fore t a rs a l seg men t s in fo l l ow i n g p ropor t i o n s : 1 5 : 3 : 5 : 1 5 : 2 1 ; en l arged c l a w w i t h s u b a p i c a l t oo t h a n d 6 l a me l l a e ; segme n t 5 o f fron t t a rs u s w i t h t w o r o w s of n ea r l y 30 l a me l l ae e x t en d i n g con t i n uo u s l y t o t h e a p e x ; m a x i l l a r y p a l p i w i t h 4 s e g me n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n

G o n a topod i n a e

L O C U S TYP I C U S : G o l den L . ( O n t a r i o , C a n a d a ) T Y P I CAL MAT E R I AL : ho l ot y p e F ! a n d 1 p a r a t y p e F ! i n AM ;

1

paratype F !

1 27 1

i n OT ;

paratype F !

i n OL .

HOST S : u n k n o w n D I ST R I BU T I O N : N e a rc t i c reg i on : CANAD A : G o l d e n L . ( O n t . ) , OT ! O L ! U . S . A . : K a l b f l e i sc h F i e l d S t a t i on ( H u n t i n g ton , L o n g I s l a n d , N . Y . ) , A M !

6 . Pseudogona topus stenocra n i R . C . L .

Perk i n s

Pseu dogon a top u s sten ocra n i R . C . L . P e r k i n s 1 905 : 38 . Pseu dogona topus stenocra n i v a r . d u b io s u s R . C . L . P e r k i n s 1 905 : 39 ( s y n . p roposed b y F re y t a g 1 978 ) ; locus typ i c u s C o l u m b u s ( Oh i o , U . S . A . ) . Pseudogon a top u s iowensis F e n to n 1 924 : 1 90 ( s y n . p roposed b y F re y t a g 1 978 ) ; locus typ icus Ames ( I o w a , U . S . A . ) .

F E MAL E : a p t e rou s ; l e n g t h 2 , 62-3 , 29 m m ; h e a d t e s t a ceous or y e l l o w-b row n ; occ a s i o n a l l y r e d d i s h - b r ow n ; u s u a l l y w i t h v e r t ex b row n i s h ; f r o n t p a r t of v e r t e x , c l y p e u s and m a n d i b l es y e l l o w ; a n t e n n a e u s u a l l y b ro w n , w i t h s e g " l e n t s 1 -3 a n d 1 0 y e l l ow ; occas i o n a l l y s e g m e n t 1 0 w h i t e ; occ a s i o n a l l y a n t e n n a e fu l l y t e s t a c eo u s ; p ron o t u m redd i s h , occas i o n a l l y w i t h b r o w n spots o n i d es a n d b e t ween c o l l a r a n d d i sc ; s c u t u m redd i s h , o cc a s i o n a l l y w i t h s i de s b l a ck ; occ a s i on a l l y sc u t u m f u l l y b l ack ; sc u t e l l u m a n d m e t a t ho r a x + p ro p o d e u m b l a c k ; p e t i o l e b l ac k ; a b domen red d i s h o r b ro w n ; l e g s y e l l o w ; h e a d excav a te d , s h i n y or d u l l , g r a n u l a t ed ; p ro n o t u m c rossed by a s t ro n g t r a n s v e rs a l i m p ress i on , s h i n y , w e a k l y g ra n u l a t ed ; sc u t u m d u l l , g r a n u l a t e d ; m e t a n o t u m w i t h s i des rou n d e d ( F i g . 852 A ) ; m e t a t ho r a x + p ro p o d e u m d u l l , g r a n u l a te d ; p l e u r a a n d pos t e r i o r s u rface of p ropod e u m t ra n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re o b so l e t e ; seg men t 1 of f ro n t t a r s u s s h o r t e r t h a n seg m e n t 4 ( 25 : 26 i n l ec t o t y p e o f Ps . s tenocra n i ; 1 1 : 1 3 i n ho l o t y pe of Ps . iO\
1 272

Mem . Amer . E n ! .

I n s! . ,

no .

37 ,

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F i g . 863 - M a l e g en i t a l i a of Pseu dogona topus stenocra n i R . C . l . P e rk i n s from lex i n g to n ( Ke n t uc k y ) ( r i g h t h a l f removed )

t ra n sversa l or l o n g i t u d i n a l k ee l s ; fore- w i n g h y a l i n e , w i t h o u t d a r k t r a n s v e rs a l b a n d s ; ra d i a l ce l l open ; d i s t a l p a rt of ra d i a l v e i n m u c h s horter t h a n prox i m a l p a r t ; dors a l p rocess of gonoforceps ( F i g . 863 ) very red u ced ; b as i vo l se l l a w i t h o u t r i d g e ; m a x i I l a r y p a l p i w i t h 4 segme n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . lOCUS T Y P I C US : O h i o ( U . S . A . ) TYP I C A L M A T E R I A L : l ec t o t y p e F ! a n d 2 p a r a l ec t o t y p e s F F ! ( des i g n a ted b y O l m i 1 98 2 ) of Ps . s tenocra n i i n B ; l ec to t y pe F ! ( des i g n a ted b y O l m i 1 98 2 ) of Ps . stenocrani v a r . d u b iosus i n B ; h o l o t y p e F ! of Ps . iowensis i n WA . HOST S : i n U . S . A . Stenocran u s dorsa l i s a n d Delp h a codes l u tu l en ta ( V a n D u zee ) ( R . C . l . Perk i n s 1 906c ; F re y t a g 1 978 ; O l m i 1 982 ) . D I ST R I BU T I ON : Nearc t i c reg i on : CANADA : P r i nce A l b e r t ( Sa s k . ) , O T ! K enosee ( Sa sk . ) , OT ! loc keport ( N . 5 . ) , O T ! u . S . A . : l i n co l n ( A u s t i n , T ex . ) , O l ! O T ! M i d l a n d Co . ( M i c h . ) ( F rey t a g 1 978 ) ; Ames ( I o w a ) , WA ! O h i o , B ! lex i n g t o n ( K y . ) , K E ! Sea C l i ff ( N . Y . ) , C M ! O n eon t a ( N . Y . ) ,

1 273

G o n a t o po d i n ae

F i g . 864 - C he l a of Pseu dogon a top u s a u toxen o b i u s R . C . L . Moores t o w n ( New Jersey )

Perk i n s from

TW!

7 . Pseu dogon a top u s a u toxeno b i u s R . C . L .

Perk i n s

Pseu dogon a top u s a u toxenob i u s R . C . L . P e r k i n s 1 907 : 3 7 .

F E M A L E : a p terou s ;

l e n g t h 3 , 0-4 , 2 m m ;

h e a d b ro w n ,

w i t h m a n d i b l es ,

c l y p e u s a n d f ro n t p a r t o f v e r t e x y e l l o w ; occ a s i o n a l l y occ i p u t a n d ven t r a l s i de t e s t ac eo u s ; a n t e n n ae b row n , w i t h s e g m e n t s 1 -2 y e l l ow ; p rono t u m redd i s h , w i t h b ro w n s po t s o n s i des a n d b e t ween d i sc a n d co l l a r ; occ a s i o n a l l y p ron o t u m mos t l y b row n ; scu t u m b l ack , w i t h o r w i t ho u t b ro w n n u a nces ; scu t e l l u m a n d met a t horax + p ro p o d e u m b l ac k ( w i t h a p e x y e l l o w ) ; a b domen b ro w n-b l ac k ; l eg s y e l l o w - re d d i s h , w i t h m i d a n d h i n d coxae

1 274

Mem . Amer . E n t .

I n st . ,

no . 37 ,

1 984

F i g . 865 - C he l a of Pseu dogon a top u s magnus Brown f rom B r i g h ton ( F l or i d a )

Gon a topod i n ae

1 275

b ro w n - b l ac k ; c l u b s of femora b row n ; head e x c a v a t e d , s h i n y , g ra n u l a t ed ; p ron o t u m crossed by a s t ro n g t ra n s v ersa l i m p res s i on , d u l l , g ra n u l a t e d ; m e t a n o t u m w i t h s i des roun d e d ; meso-me t a p l e u r a l s u t u re o b so l e t e ; m e t a t ho r a x + p ro p o d e u m d u l l , g r a n u l a ted ; p os t e r i o r s u rface of p ropodeum t ra n s v e rse l y s t r i a te ; segment 4 o f f r o n t t a rs u s shorter than s e g m en t 1 ( 28 : 40 i n t h e ho l ot y p e ) ; en l a rged c l a w ( F i g . 864 ) w i t h s u b a p i c a l t oo t h a n d 4 l am e l l ae ; segmen t 5 o f fron t t a rs u s ( F i g . 864 ) w i t h t w o row s of 1 9-22 l a m e l l ae ; apex w i t h a group o f 6-9 l a me l l a e ; occa s i o n a l l y segme n t 5 w i t h rows of l a me l l ae e x t e n d i n g con t i n uo u s l y to t he a pe x ; m a x i I l ary p a l p i w i t h 4 s e g me n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n known L O C U S T Y P I C U S : N o g a l es ( A r i z . , U . S . A . ) T Y P I CAL M A T E R I A L : l ec to t y p e F ! ( des i g n a te d by O l m i

1 98 2 )

in B.

HOST S : i n U . S . A . Stob a era sp . ( R . C . L . P e r k i n s 1 907 ; O l m i 1 982 ) . D I ST R I B U T I O N : N e a rc t i c reg i on : U . S . A . : No g a l es ( A r i z . ) , B ! Moores t o w n ( N . J . ) , TW !

8 . Pseu dogo n a topus m a gn u s Brow n

Pseu dogon a top u s m a gn u s B ro w n 1 940 :

11.

F E M A L E : a p t e ro u s ; l e n g t h 3 , 75-5 , 00 m m ; h e a d b row n , w i t h m a n d i b l es , c l y pe u s a n d fron t p a r t o f v e r t e x y e l l o w ; a n t en n ae b row n , w i t h s e g me n t s 1 -2 a n d p a r t l y 3 y e l l ow ; p ro n o t u m a n d sc u t u m y e l l o w or redd i s h ; s c u te l l u m red d i s h o r y e l l ow ; occ a s i o n a l l y s c u t e l l u m b ro w n ; m e t a n o t u m b l ac k o r brow n ; occ a s i on a l l y p a r t l y red d i s h ; m e t a t h o r a x + p ropodeum b l ac k , w i t h a p e x ye l l o w ; p e t i o l e b l ac k ; a b domen y e l l ow - re d d i s h , w i t h a p i c a l t e r g i tes b ro w n ; l eg s y e l l o w ; h e a d s h i n y , exc a v a ted , w e a k l y g ra n u l a te d ; p rono t u m crossed b y a s t ro n g t ra n svers a l i m p ress i on , s h i n y , smoo t h , w e a k l y g r a n u l a t e d ; scu t u m d u l l , g r a n u l a t e d ; m e t a n o t u m t ra n sverse I y s t r i a t e , w i t h s i d e s r o u n ded ; meso-me t a p I e u r a I s u t u re obso l e t e ; m e t a t ho r a x + propode u m s h i n y , g ra n u l a t e d ; p l eu ra a n d p o s t e r i or s u rface of p ropodeum t ra n s v erse l y s t r i a t e ; a n t e r i or s u rface of m e t a t ho r a x + p ropodeum w i t h t rack o f med i a n f u rrow ; segmen t 1 o f f ro n t t a rs u s l o n g e r t h a n seg m e n t 4 ( 1 2 : 1 0 ) ; e n l arged c l a w ( F i g . 865 ) w i t h s U b a p i c a l too t h a n d 5-7 l a me l l a e ; segmen t 5 o f fron t t a rs u s ( F i g . 865 ) w i t h 2 rows of 1 6-22 l a me l l ae ; apex w i t h a g ro u p of 6 l a me l l a e ; m a x i l l a r y p a l pi w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : C rom w e l l ( I n d . , U . S . A . ) T Y P I CAL M A T E R I AL : ho l o t y p e F ! i n WA . HOST S : u n k n o w n D I ST R I B U T I O N : Nearc t i c reg i o n : U . S . A . : C ro m we l l ( W a w asee L ak e , I n d . ) , iVA ! L a k e I s t o k p o g a ( F l a . ) , OL ! B r i g h to n ( F l a . ) , G V ! G a i n s v i l i e ( F l a . ) ,

Mem . Amer . E n t .

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I n st . ,

n o . 37 ,

1 984

WA!

N E O T R O P I C S PE C I E S O F PSEUDOGONA TOPUS

F i ve spec i es a re k n ow n .

KEY T O T H E SPEC I ES

F E MA L E S

Me s o -me t a p l e u r a l

s u t u r e d i s t i n c t a n d c omp l e t e • • • • • • • • • • . • • • • • • • • • • 5 . morenoi n . s p . M e s o -me t a p l e u ra l s u t u r e ob s o l e t e • • • • • • • • • • . • • • • • • • • • • • H ea d m o r e e x c a v a t ed ; b od y p a r t l y b l a c k . • • • • • • • • • • • . . • • H ea d w e a k l y e x c a v a t ed ( F i g s 8 6 6 C , 8 6 8 C ) ; b od y mo s t l y •

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t e s t a c e ou s • • • • • • • • • • • • • • • • • • • • • S c u t u m y e l l ow ; p r o n o t u m r ed d i s h

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3 . delp h a cidis n . sp . S c u t u m b l a c k ; p r o n o t u m mo s t l y b l a c k . • • • • • • • • • • • • • . • • • • • • · . • . . . . . • . • . . . . . • . . . . . . . • . . . . 4 . m u esebecki n . sp . A n t e n n a l s eg m e n t s 1 - 9 y e l l ow ; s eg me n t 4 o f f r on t t a r s u s s h o r t e r t h a n s egmen t 1 ( n ea r l y 8 5% - 9 1 % ) • • • . • • • • • . . • . • • • • . . . . . • • . . . . • • . • . . . . . . • . . . . 1 . variistri a t u s F en t on A n t en n a l s eg m e n t s 3 - 9 b r own or b l a c k ; s egmen t 4 of f r on t t a r s u s a s l on g a s o r s l i g h t l y l on g e r t h a n s eg me n t 1 • • • • • · 2 . fla v u s n . sp . •

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MALES

U n k no w n

1 . Pseu dogon a top u s variistriatus Fen ton

Pseu dogon a top u s variistri a t u s F e n ton 1 92 7 : 5 . Pseu dogon a top u s s a ccharivorae R i c h a r d s 1 972 : 544 ( n . s y n . ) ; locus typ icus B e r n a r d Lodg e ( Ja m a i c a ) .

1 27 7

G o n a topod i n a e

F i g . 8 6 6 - F e m a l e of Pseudogona topus variistria t u s F e n ton t h o r a x a n d p ropod e u m ; C : h e a d .

( t y pe ) ;

A,

B:

F E MA L E : a p t ero u s ; l e n g t h 2 , 1 8-3 , 00 m m ; y e l l o w - t e s t aceo u s , w i t h pe t i o l e b l ac k ; a n t e n n ae y e l l o w - t e s t a c eo u s ; h e a d ( F i g . 866 C ) s h i n y , e x c a v a ted , smoo t h , h a i r l ess , wea k l y a l u t aceou s ; p r o n o t u m ( F i g . 866 A , B ) c rossed b y a s t ro n g t r a n sv e rs a l i mp ress i o n , smoot h , s h i n y , wea k l y g r a n u l a t ed ; sc u t u m smoo t h , s h i n y , w ea k l y g r a n u l a ted ; m e t a n o t u m t r a n s v e r se l y s t r i a t e , w i t h s i des rou n d e d ; m e t a t ho r a x + p ropodeum ( F i g . 866 A , B ) s h i n y , smoo t h ; a n t e r i o r s u rf ace w e a k l y i nc l i n ed ; p l e u r a a n d p os t e r i or s u rface of p ro p o d e u m t ra n s v erse l y s t r i a t e ; meso-met a p l e u ra l s u t u re obso l e t e ; seg men t 1 of f ro n t t a r s u s l on g e r t h a n segmen t 4 ( 1 4 : 1 2 ) ; e n l a r g e d c l a w ( F i g . 867 ) w i t h s u b a p i c a l too t h a n d 4-5 l a m e l l ae ; s e g m e n t 5 of f ro n t t a rs u s ( F i g . 867 ) w i t h t w o rows of 8- 1 3 l a m e l l ae ; a p e x w i t h a g r o u p of 4-9 l a m e l l ae ; m a x i I l a r y p a l p i w i t h 4 s e g me n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E : u n k no w n LOCUS T Y P I C U S : R i o P i e d r a s ( P u e r t o R i c o ) TYP I CA L MAT E R I A L : ho l o t y p e F ! o f Ps . variis tria tus i n WA ; ho l ot y p e F ! a n d 3 p a r a t y pes F F ! of Ps . saccharivorae i n B M . HOST S : i n J a m a i c a Sa ccharosydne s a cch arivora ( We s t w . ) . D I ST R I BU T I O N : Neot rop i c reg i o n : BAHAMA S : A b a c o , BM ! P U E R T O R I CO : R i o P i e d r a s , WA ! JAMA I CA :

Ber n a r d Lo dge , B M !

2 . Pseu dogon a top us fla vus n . sp .

Mem . A m e r . E n ! .

1 278

Fig.

I n s! . ,

no.

37 ,

1 984

867 - C h e l a of Pseu dogon a top u s variis tria tus F e n t o n

( t y pe )

F E M A L E : a p terou s ; l e n g t h 2 , 3 1 -2 , 37 m m ; y e l l o w - t e s t aceou s , w i t h b l ac k ; a n t e n n a e b ro w n , w i t h segmen t s 1 , 2 a n d 1 0 w h i te ; h e a d C ) e x c a v a ted , s h i n y , wea k l y a l u t a c eo u s ; p ro n o t u m ( F i g . 868 A, crossed b y a s t ro n g t r a n s v e rs a l i m p ress i o n , s h i n y , g r a n u l a t e d ; g r a n u l a ted ; m e t a n o t u m t r a n s verse l y s t r i a t e , w i t h s i de s rou n d e d ;

pet i o l e ( F i g . 868 B) sc u t u m meso-

1 279

Gona topod i n a e

F i g . 868 - F em a l e of Pseu dogona top u s fla vus n . sp . a n d p ropod e u m ; C : h e a d .

m e t a p l e u r a l s u t u re o b so l et e ;

( ty pe ) ;

m e t a t h o r a x + p ropodeum ( F i g .

A,

B:

868 A ,

t ho r a x

B)

w i th

a n t er i or s u rf ace w e a k l y i n c l i ned ; fore t a rs a l s e g m e n t s i n fo l l ow i n g propor t i on s : 8 : 2 : 2 , 5 : 9 : 1 4 ; s e g me n t 1 of front t a rs u s as l o n g a s s e g m e n t 4 i n a p a r a t y pe ; en l arged c l a w ( F i g . 869 ) w i t h s u b a p i c a l t oo t h a n d 4 - 5 l ame l l a e ; segmen t 5 o f fron t t a rs u s ( F i g . 869 ) w i t h t wo rows of 8-1 7 l a me l l ae ; a p e x w i t h a g ro u p of 8-1 1 l a me l l ae ; m a x i I l a r y p a l p i w i t h 3 segmen t s ; l ab i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k no w n L O C U S T Y P I C U S : G u a d a l ou p e T Y P I CAL M A T E R I A L : ho l o t y pe F ! a n d 2 p a r a t y pes F F ! i n B M ; 1 p a ra t y pe F ! i n WA ; 1 p a r a t y p e F ! i n CA . HOST S : u n k n o w n D I ST R I B U T I ON : Neot rop i c reg i o n : G R E N A D A : G r a n d E t a n g , BM ! W E S T I N D I E S : S a i n t L uc i a , WA ! GUADALOU P E , BM ! C H I L E : A n g o l , C A l BRAZ I L : Nova T e u ton i a ( S . t a C a t a r i n a ) , B M !

3 . Pseu dogona top u s delp h a c idis n .

sp .

F E M AL E : a p terou s ; l e n g t h 3 m m ; h e a d t e s t a c eo u s , w i t h v e r t e x d a r k ; a n t e n n a e b row n , w i t h s e g m e n t s 1 -2 b row n ; p ro t h o r a x redd i s h ; sc u t u m y e l l o w ; m e t a t ho r a x + p ropodeum b l ac k , w i t h poster i or reg i on of t he propodeu m t e s t a c eous ; a b domen red d i s h - t e s t a ceou s ; p e t i o l e b l ac k ; l e g s t e s t a c eo u s , w i t h h i n d coxae p a r t l y b l ac k ; s t a l k o f h i n d femora brown ; a n t e n n a e d i st a l l y t h i ck e n e d ; a n t en n a l s e g m e n t s i n fo l l ow i n g p ropor t i on s : 8 : 4 : 1 1 : 6 : 5 : 5 : 5 : 5 : 5 : 8 ; h e a d e x c a v a t e d , s h i n y , w i t ho u t sc u l p t u re ;

Mem . A m e r . E n ! .

1 280

Fig.

869

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I n s ! . , n o . 37 ,

1 984

C h e l a of Pseudogona top u s fl a vu s n .

sp .

( type )

frons w i t h med i a n k e e l ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 1 , 5 ; OL = 1 , 5 ; OOL = 8 ; p ronot u m crossed b y a s t ro n g t ra n sversa l i mp ress i on , s h i n y , w i t ho u t scu l p t u r e ; scu t u m s h i n y , w i t h some w e a k l on g i t u d i n a l k ee l s ; m e t a n o t u m smoot h , s h i n y , w i t hou t scu l p t u re , w i t h s i de s rounded ; m e t a t h o r a x + p ropodeum smoo t h , s h i n y , w i t h o u t scu l p t u re , except for p l eu ra a n d poste r i or s u rface of p ropodeum t r a n s verse l y s t r i a t e ; meso m e t a p l eu r a l s u t u re obso l e te ; fore t a rs a l s e g me n t s i n fol l ow i n g p roport i on s : 1 4 : 2 : 4 : 1 1 : 1 6 ; en l a r g e d c l a w ( F i g . 870) w i t h a s U b a p i c a l tooth a n d 6 l a me l l ae ; s e g m e n t 5 of fro n t t a r s u s ( F i g . 870 ) w i t h 2 rows of 1 4 l a m e l l ae ; a pe x w i t h a g ro u p o f 7 l a me l l ae ; m a x i l l a ry p a l p i w i t h 4 s e g me n t s ; l a b i a l p a l p i w i t h 2 s e g m en t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n LOCUS T Y P I C US : N o v a Teu ton i a ( B r az i l ) T Y P I CA L MAT E R I A L : ho l o t y p e F ! i n O T . HOST S : u n k n o w n D I ST R I B U T I ON : Neotrop i c reg i on : BRAZ I L : N o v a T e u ton i a ( S . t a C a t a r i n a ) , OT !

4 . Pseu dogon a top u s m u esebecki n . sp .

F E MAL E : a p t e ro u s ; l en g t h 3 , 62 m m ; h e a d b l ack , w i t h a n te r i o r reg i on of fron s , c l y pe u s a n d m a n d i b l es testaceo u s ; a n t e n n a e b row n , w i t h s e g m e n t s

1 , 2 a n d 1 0 test aceou s ; t h o r a x a n d p ropodeum b l ack , except f o r s i des o f p ro n o t u m red d i s h ; a b domen b rown ; l eg s b row n , w i t h t a r s i l i g h t ; a n ten n a e d i s t a l l y t h i ck ened ; a n t e n n a I segme n t s i n fo l l ow i n g p ropor t i o n s : 1 0 : 5 : 1 3 : 6 : 5 : 5 : 5 : 5 : 5 : 9 ; hea d d u l l , v e r y exc a v a t e d , g r a n u l a t e d ; fro n s w i t h med i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 1 ; O L = 1 , 5 ; OOL = 6 , 5 ; p ro n o t u m crossed b y a s t ro n g t r a n s v e r s a l i m p ress i o n , s h i n y , w i t h o u t scu l p t u re on s i des a n d on co l l a r ; d i sc g r a n u l a t e d ; s c u t u m d u l l , w i t h some l on g i t u d i n a l k ee l s ; s c u te l l u m wea k l y v i s i b l e ; m e t a n o t u m t r a n sverse l y s t r i a t e ; s i de s of m e t a not u m p rot r u d i n g ( t w o po i n ted p rot r u s i on s per s i de ) ;

Gon a t opod i n a e

F i g . B70

-

C h e l a o f Pseu dogon a top u s delp h a cidis n .

1 2B1

sp .

( type)

met a t ho r a x + p ropodeum s h i n y , t ra n s v ers e l y s t r i a t e o n p l eu r a a n d poster i or s u rf ace ; a n t e r i or s u rf ace smoo t h , w i t ho u t scu l p t u re ; meso m e t a p l e u r a l s u t u re obso l et e ; fore t a rs a l seg men t s in f o l l o w i n g p roport i on s : 1 3 : 2 , 5 : 6 : 1 4 : 24 ; en l a r g e d c l a w ( F i g . B 7 1 ) w i t h s u b a p i c a l tooth a n d 5

Mem . A m e r . E n ! .

1 282

\

\

F i g . 87 1

I n s! . ,

n o . 37 ,

1 984

i

\�

- C he l a of Pseu dogon a top u s m u esebecki n .

sp.

( type)

l ame l l ae ; s e g m e n t 5 of f ro n t t a r s u s ( F i g . 8 7 1 ) w i t h t w o rows of 1 2 l a me l l ae ; a p e x w i t h a g r o u p of a t l ea s t 1 5 l a me l l ae ; m a x i l l a r y p a l p i 4 segmen t s ; l a b i a l p a l p i w i t h 2 seg men t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k no w n L O C U S T Y P I C U S : G u a t em a l a

with

1 283

Go n a t o p o d i n ae

F i g . 872

-

C h e l a of Pseudogona top u s morenoi n .

sp.

( paratype)

1 284

Mem . Amer . E n t .

I n st . ,

no.

37 ,

1 984

T Y P I CAL M A T E R I A L : h o l o t y p e F ! i n W A . HOST S : u n k no w n D I ST R I B U T I ON : Neot rop i c reg i on : G U A T E MALA , WA ! N OT E : t h e spec i es i s n amed i n honor of M r . C a r l F . W . M u esebec k .

5 . Pseu dogona topus morenoi n . sp .

F E MA L E : a p t e ro u s ; l en g t h 2 , 4-3 , 0 m m ; h e a d b l ack , w i t h c l y pe u s a n d m a n d i b l es t e s t aceo u s ; a n te n n ae b l ac k , w i t h segmen t s 1 -2 t e s t aceo u s ; seg m e n t 1 0 w h i t e ; t h o ra x , p ropode u m a n d a b domen b l a c k - b ro w n , w i t h scu t u m a n d a p e x of p ropodeum y e l l o w ; l eg s b row n , w i t h t a r s i a n d t roc h a n t ers y e l l ow ; i n i mm a t u re s p ec i me n s b o d y b ro w n - l i g h t ; a n t e n n ae d i s t a l l y t h i ck e n e d ; a n t en n a l seg m e n t s i n fo l l ow i n g p ropor t i on s : 7 : 6 : 8 : 6 : 6 : 6 : 5 : 5 : 5 : 8 ; h e a d exc a v a ted , d u l l , g r a n u l a ted ; fron t a l l i n e comp l et e ; occ i p i t a l c a r i n a i n v i s i b l e ; POL � 1 ; OL � 2 ; OOL � 7; p ro n o t u m c rossed b y a s t ro n g t r a n s v e rs a l i mp ress i o n , s h i n y , w e a k l y gra n u l a t e d ; s c u t u m a n d s c u t e l l u m d u l l , g r a n u l a t e d ; meso-me t a p l eu r a l s u t u re comp l et e ; me t a n o t u m weak l y ho l l ow b e h i n d t h e sc u t e l l u m , w i t h s i des p ro t ru d i n g ; p ro t r u s i on s r o u n d e d ; m e t a t ho r a x + p ropode u m d u l l , g r a n u l a ted ; p l eu r a a n d pos ter i or s u rface of p ropod e u m t r a n s verse l y s t r i a t e ; fore t a r s a l s e g m e n t s i n fo l l o w i n g p ropo r t i o n s :

1 2 : 3 : 4 : 1 2 : 1 9 ; e n l a rged c l a w ( F i g . 872 ) w i t h s U b a p i c a l t oo t h a n d 9 l a m e l l ae ; s e g me n t 5 of fro n t t a r s u s ( F i g . 872 ) w i t h t wo rows o f n e a r l y 1 5 l a me l l ae ; a p e x w i t h a g ro u p o f n e a r l y 1 0 l a me l l a e ; m a x i I l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : I n gen i o C a u c a ( Co l om b i a ) T Y P I C A L M A T E R I A L : h o l o t y p e F ! a n d 2 p a r a t y p es F F ! i n O L . H O ST S : i n C o l o m b i a Perkinsiella s a cch aricida (Delph a c i da e ) . D I S T R I B U T I O N : Neot rop i c reg i on : COLO M B I A : I n gen i o C a u c a ( V a l l e ) , O L ! NOT E S : t h e s p ec i es i s n a m e d i n honor of t h e f i n d e r M r . L u i s E n r i q u e Moreno A rc e .

A U ST RAL I A N S P E C I ES OF PSEUDOGONA TOPUS

E i g h t s p ec i es are k n o w n .

K E Y TO T H E S P E C I E S

F E MA L E S

Gona topod i n ae

1 285

IAe t a n o t u m w i t h s i d e s r o u n d ed ( F i g . 3 7 3 ) ; rne s o -rne t a p l e u r a l s u t u r e ob s o l e t e . . • • . . • • • . . . . . . . .2 1'v1. e t a n o t u m w i t h s i d e s p r o t r u d i n g ( F i g s 8 7 9 , 8 8 2 , 3 8 5 ) ;

2

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p r o t r u s i on s r o u n d e d o r p o i n t ed ; me s o -rne t a p l e u r a l s u t u r e ob s o l e t e o r d i s t i n c t . . . . . • • . . . . . . . . . . . . . . . . . . . . • . . . . . • . . . • . • • 4 T h o ra x a n d p r o p o d e u m b l a c k , w i t il a p e x o f p r o p o d e u m y e I l ow • • . • . . . . . . . . . . . . . . . . 1 . n igri c a n s ( R . C . L . P e rk i n s ) ! 2 . hosp es K . C . L . P e r k i n s ! T h o r a x a n d p r op od e u rn t e s t a c e o u s o r b r own - r e d d i s h . .. .. 3 M e t a n o t u m , me s op l e u r a , me t a p l e u r a a n d p r o p o d e u m s t r o n g l y t r a n s v e r s e l y s t r i a t e , ex c e p t f o r t h e a n t e r i o r s u r f a c e o f me t a t h o r a x + p r op ocJ e u m l on g i t u d i n a l l y s t r i a t e . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 . pl uris u l c a tus n . s p . Me t a t h o r a x + p r op od e u m w i t h w e a k t r a n sv e r s a l s t r i a e on p l e u ra and p o s t e r i o r s u r f a c e ; no l o n g i t u d i n a l s t r i a e . . . . . . . . . . . . . . . . . •

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Paragon a top u s n igri c a n s R . C . L . P e r k i n s 1 90 5 : 4 1 . Pseudogona top u s mel a n a crias R . C . L . P e r k i n s 1 906a : 487 ( s y n . p roposed b y R . C . L . Perk i n s 1 9 1 2 ) ; locus typ icus S u v a ( F i j i I s l ands ) . Paragon a top u s n igri c a n s R . C . L . P e r k i n s : K i effer 1 9 1 4b : 8 7 . Paragon a top u s n igri c a n s R . C . L . P e r k i n s : O l m i 1 979 : 2 1 8 .

1 288

Mem . Amer . E n ! .

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1 984

F i g . 875 - M a l e gen i t a l i a of Pseu dogon a top u s n igricans ( R . C . L . Perk i n s ) from B u n da b e r g ( Qu e e n s l an d ) : spec i me n w i t h a p e x of dors a l p rocess of gonoforceps l es s rou n d e d .

F E M A L E ( F i g . 873 A ) : a p t e rou s ; l en g t h 2 , 37-2 , 92 m m ; b row n - red d i s h or b row n ; occ a s i on a l l y b l ac k ; a n te n n a e b row n , w i t h segmen t s 1 -2 or 1 -3 y e l l ow ; m a n d i b l es , c l y pe u s a n d fron t p a r t of vertex y e l l ow ; occ a s i o n a l l y p ro no t u m b l ac k ; m e t a t ho r a x + p ropode u m b ro w n - b I a c k or b l ac k ( e xcep t for apex of p ropod e u m y e l l ow ) ; pe t i o l e b l ac k ; l eg s y e l l ow , occ a s i on a l l y w i t h c l u b s o f femora b row n ; h e a d ( F i g . 873 B ) exc a v a t e d , s h i n y , h a i r y , g ra n u l a t e d ; p rono t u m ( F i g . 873 C ) g r a n u l a ted , crossed by a s t ro n g t r a n sversa I i m p ress i on ; m e t a n o t u m a s l o n g a s s c u t u m , t r a n s verse I y

Gon a topod i n a e

1 289

\ F i g . 876 - M a l e g en i t a l i a of Pseu dogona top u s nigricans ( R . C . L . Perk i n s ) from B u n d a be r g ( Qu een s l a n d ) : spec i men w i t h a p e x o f d o rs a l p rocess of gonoforceps more rou n d e d ( pen i s a n d r i g h t h a l f removed ) .

s t r i a t e , meso- m e t a p l e u r a l s u t u re obso l e t e ; p o s t e r i or s u rface of p ropod e u m t ra n s v e rse l y s t r i a te ; seg men t 4 of f ro n t t a r s u s s h o r t e r t h a n se g m e n t 1 ( 20 : 27 ) ; en l a r g e d c l a w ( F i g . 874 ) w i t h s u b a p i c a l tooth a n d 3-4 l a me l l ae ; segmen t 5 o f f ro n t t a r s u s ( F i g . 874 ) w i t h t w o rows o f 9-2 1 l a m e l l ae ; a pex w i t h a g r o u p of 4-9 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 2 segmen t s ; l a b i a l p a l p i w i t h 2 segmen ts ; t i b i a l s p u r s 1 , 0 , 1 . MALE : fu l l y w i n ged ; l e n g t h 1 , 90-2 , 1 9 m m ; b row n ; a n te n n a l seg m e n t s i n fo l l o w i n g p ro p o r t i o n s : 5 : 5 : 9 : 8 : 8 : 9 : 7 : 8 : 7 : 1 0 ; a n t e n n a I seg men t 3 more t h a n t h ree t i mes as l o n g as b r o a d ( 5 , 0 ) ; h e a d s h i n y , g ra n u l a te d ;

1 290

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877 - C h e l a of Pseu dogonatopus hospes R . C . L . Perk i n s from M ac a u

fron t a l l i ne a b s en t ; occ i p i t a l c a r i n a i n v i s i b l e ; t em p l es d i s t i n c t ; P O L = 7 ; O L = 3 ; OOL = 3 , 5 ; b re a d t h of t h e oce l l i : 3 , 5 ; s c u t u m s h i n y , g r a n u l a ted ; not a u l i ces com p l e t e , p o s t e r i o r l y jo i n t ; s c u t e l l u m a n d m e t a n o t u m s h i n y , i n d i s t i n c t l y p u n c t a te ; p ropodeu m w i t h pos t e r i o r su rface ret i c u l a t e r u gose ; dors a l s u rface w i t h a med i a n l o n g i t u d i n a l f u rrow a n d w i t h t w o l a tera l smoo t h areas ; fore-w i n g h y a l i n e , n o t crossed b y d a r k b a n d s ; dors a l p rocess of gonoforceps ( F i g s 875 , 87G ) d i s t a l l y rou n d e d ; m a x i l l a ry p a l p i w i t h 2 s e g m e n t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 1 , 2 . L O C U S T Y P I C U S : B u n d a b er g ( Queen s l a n d , A u s t r a l i a ) TYP I C A L MAT E R I A L : l ec t o t y p e F ! a n d 2 p a r a l ec t o t y p es F F ! ( des i g n a ted b y O l m i 1 98 2 ) o f P . nigricans i n B ; l ec t o t y p e F ! a n d 2 p a r a l e c t o t y p es F F ! ( d e i s g n a t e d b y O l m i 1 98 2 ) of Ps . mela n a cria s i n B . Stenocran u 5 a n d L i b urnia ( R . C . L . I IOST S : i n I; u s t r a I i a Delpha cidae n e a r Perk i n s 1 9J5c ) . :) I ST '� I B U T I ,] ' I : ". u s t r c1 1 i a n reg i on : A U S T R A L I A : B u n d a berg ( Queens l a n d ) , Ol ! B ! F I J I I SLANDS : S u v a ( V i t i levu ) , B !

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F E MAL E : a p tero u s ; l e n g t h 3 , 62 -4 , 37 m m ; h e a d b row n , w i t h m a n d i b l e s , c l y p e u s a n d f ro n t p a r t o f v e r t e x y e l l o w ; a n t e n n a e b row n , w i t h segme n t s 1 -2 y e l l o w j p ro n o t u m , sc u t u m , s c u te I I u rn , m e t a n o t u m a n d m e t a t ho r a x + p ropode u m b l a c k ; a p e x of propo d e u m y e l l ow ; p et i o l e b l ac k ; a b domen b ro w n - b l a c k ; l e g s y e l l ow , w i t h p a r t of cox a e a n d cl ubs of femora b ro w n i s h ; h e a d exc a v a t e d , s h i n y , w i t ho u t sc u l p t u re , exce p t for occ i p u t a n d f ro n t p a r t o f vertex g ra n u l a ted ; prono t u m g ra n u l a t e d , crossed b y a s t ro n g t r a n s v e rs a l i m press i o n , s h i n y ; s c u t u m g r a n u l a te d ; s c u t e l l u m d u l l , w i t ho u t scu l p t u re ; met a n o t u m v e r y l on g , t r a n s verse l y s t r i a t e , n e a r l y a s l o n g a s sc u t u m , w i t h s i d e s ro u n ded ; p l e u r a a n d pos t e r i o r s u rface of

1 292

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I ns! . ,

n o . 37 ,

1 984

p ropode u m t r a n sverse l y s t r i a te ; meso-me t a p l e u r a l s u t u re o b so l e t e ; m e t a t ho r a x + p ropodeum s h i n y , w i t h t r a c k of med i a n f u r row , i n d i s t i n c t l y scu l p t u re d ; s e g me n t 4 of fro n t t a rs u s shorter t h a n segm e n t 1 ( 40 : 47 ) ; e n l arged c l a w

( F i g . 877 ) w i t h s u b a p i c a l

too t h a n d 4-9 l a me l l ae ;

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5 o f f r o n t t a r s u s ( F i g . 877 ) w i t h t w o r o w s o f 20-25 l ame l l ae ; a p e x w i t h a g ro u p of 7- 1 0 segmen t s ; m a x i I l a r y p a l p i w i t h 2-4 segme n t s ; l ab i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E : f u l l y w i n g e d ; l e n g t h 2 , 30-2 , 43 m m ; b ro w n - b l ack , w i t h frons fu l l y o r a l most f u l l y t e s t aceou s ; a n te n n a e n o t d i s t a l l y t h i ckened ; a n te n n a l seg m e n t s i n fo l l o w i n g propor t i on s : 6 : 5 , 5 : 1 0 : 9 : 8 : 9 : 8 : 8 : 7 : 1 0 ; a n t e n n a I s e g me n t 3 more t h a n t h ree t i mes as l o n g as b ro a d ( 4 , 3 3 ) ; h e a d s h i n y , h a i r y , g r a n u l a t e d ; occ i p i t a l c a r i n a a b se n t ; POL � 8 ; O L � 3 ; O O L 3 ; b r ea d t h of t h e oce l l i : 4 ; t e m p l e s d i s t i n c t ; s c u t u m h a i ry , g r a n u l a t ed ; n o t a u l i ces d i s t i nc t a n d comp l e t e , p o s t er i or l y s e p a r a ted ; m i n i m u m d i s t a n c e b e t ween t h e n o t a u l i c e s s ho r t e r t h a n b re a d t h o f t h e oce l l i ( 2 , 5 : 4 ) ; s c u te l l u m a n d met a n o t u m w e a k l y p u n c t a t e , smoo t h , w i t h o u t s c u l p t u re a m o n g p u nc t u re s ; p ropod e u m d u l l , re t i c u l a t e ru gose ; fore-w i n g h y a l i n e , w i t ho u t d a r k t r a n s v e rs a l b a n d s ; dors a l p rocess of g o n oforcep s ( F i g . 878 ) b ro a d a n d po i n ted ; m a x i l l a r y p a l p i w i t h " "egmen t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u rs 1 , 1 , 2 . LOCUS TYP I CU S : C h i n a T Y P I CA L M A T E R I A L : l ec t o t y pe F ! a n d 3 p a ra l ec t o t y pes F F ! ( de s i g n ated b y �

Olmi

1 982 ) o f Ps . hospes i n B ;

ho l o t y p e F ! o f G .

s a u teri i n E B .

HOST S : i n C h i n a a n d H a w a i i a n I s l a n d s Perk insiella s a ccharicida K i rk . ( Delph a c idae ) ; i n M a l a y a Soga tella furcifera ( Hor v . ) ( T ho m p so n 1 954 ; G r i s t a n d L e v e r 1 969 ) . PARAS I T E S : i n H a w a i i a n I s l a n d s ffelegon a top u s pseu doph anes R . C . L . Perk i n s ( Swezey 1 9 1 9 ; T i m b e r l a k e 1 9 1 9 ) . D I S T R I BU T I O N : O r i e n t a l reg i o n : MACAU , B ! HONG K O N G : T a i L u n g F arm , B M ! I ND O N E S I A : Pek a l on g a n ( J a v a ) , B ! D i e n g P l a t e a u ( J a v a ) , B ! P a l o h ( K a l i m a n t a n , Borneo ) , B M ! MALAYS I A : Se l a n gor ( M a l a y a ) , B M ! K u a l a L u m p u r ( Se l a n go r , M a l a y a ) , VA ! T u a ra n ( S a b a h ) , VA ! Pa y a h P a l oh ( 1 s t D i v . , S a r a w a k ) , B U ! THA I LA N D : H a d d Y a i ( So n g k h l a ) , V A ! Na K a m H a i ( No n B u a L a m p oo ) , VA ! T u n g Saew , VA ! T u n g F ab a d , VA ! C h a i n a t , K Y ! O L ! B a n m akok , VA ! M a e T a n g , V A ! L a m p a n g , K Y ! M ae-Th e n g , K Y ! H o t , K Y ! C h i en g D ao , K Y ! TA I WA N : T a i n a n , E B ! Kose n , O L ! KY ! G a k k o , K Y ! S h i nsh i , KY ! A u s t r a l i a n reg i o n : HAWA I I AN I SL A N D S : Hon o l u l u ( O a h u ) , B ! W a i a l a u ( O a h u ) , OL ! B ! B M ! S u g a r C o . ( O a h u ) , B ! E w a ( O a h u ) , B ! B M ! C A ! B a rb e r ' s P I . ( O a h u ) , B ! M a noa S u b s ta t i o n ( O a h u ) , B ! Oa h u , OL ! WA ! A h u k i n i ( K a u a i ) , B ! P u u nene ( Mau i ) , B ! NO T E S : spec i e s e s t a b l i s h e d i n t h e H a w a i i a n I s l a n d s from cocoo n s sen t f rom C h i n a ( R . C . L . Perk i n s 1 9 1 2 ) . P ro b a b l y t he s p ec i es is e x t i n c t in t he H a w a i i a n I s l a n ds b e c a u s e I h a v e seen i n w or l d c o l l ec t i on s no spec i me n co l l ec ted more rece n t l y t h a n 1 928 .

G o n a topod i n a e

F i g . 879

-

1 293

F e m a l e of Pseu dogon a top us dichrom u s R . C . L . Per k i n s f rom B u n d a b e r g ( Queen s l a n d ) : t h o r a x and p ropod e u m .

3 . PseudGgona top u s dichromus R . C . L .

Perk i n s

Pseudogo n a topus dichromus R . C . L . P er k i n s 1 905 : 37 .

F E MA L E : a p t e ro u s ; l e n g t h 2 , 34-2 , 60 m m ; y e l l o w - t e s t aceo u s , w i t h pe t i o l e b l ac k a n d a b domen b row n ; a n t e n n a e y e l l ow - b rown , w i t h seg men t s 1 -2 y e l l ow ; se g m e n t 1 0 p a r t l y y e l l o w ; b o d y sh i n y , h a i r l es s , w i t ho u t s c u l p t u r e ; h e a d e x c a v a t e d ; p ro n o t u m ( F i g . 879 ) crossed by a s t ro n g t r a n s v e r s a l i m p ress i o n ; meso-m e t a p l e u r a l s u t u re d i s t i n c t , comp l e t e ; met a n o t u m w i t h s i d e s protr u d i n g ; p ro t r u s i o n s po i n t e d ( F i g . 879 ) ; p l e u r a a n d pos t e r i o r s u rface t r a n s v erse l y s t r i a te ; seg m e n t 4 of f ro n t t a r s u s s l i g h t l y s ho r t e r t h a n se gmen t 1 ( 2 1 : 2 2 ) ; en l a r g e d c l a w ( F i g . 880 ) w i t h s U b a p i c a l too t h a n d 6-8 l a m e l l a e ; s e g m e n t 5 of f ro n t t a r s u s ( F i g . 880 ) w i t h t w o row s of 1 0-20 l a m e l l a e ; a p e x w i t h a group of a t l ea s t 9-1 4 l a me l l ae ; m a x i l l a ry p a l p i w i t h 4 segme n ts ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l sp u rs 1 , 0 , 1 . MALE : f u l l y w i n ge d ; l e n g t h 1 , 9 m m ; b row n , w i t h c l y pe u s a n d m a n d i b l e s y e l l ow ;. a n t e n n a l seg m e n t s i n fo l l ow i n g p ropor t i on s : 5 : 4 , 5 : 6 : 5 , 5 : 5 , 5 : 5 , 5 : 5 , 5 : 6 : 4 : 8 ; a n t e n n a I s e g m e n t 3 more t h a n t h ree t i mes a s l o n g a s b ro a d ( 3 , 52 ) ; h e a d g r a n u l a te d ; fron t a l l i n e a b s en t ; occ i p i t a l c a r i n a a b se n t ; temp l es d i s t i n c t ; P O L = 6 ; O L = 3 ; O O L = 4 ; b rea d t h of t h e oce l l i : 2 ; s c u t u m g r a n u l a t e d ; n o t a u l i ces comp l e t e , we a k l y v i s i o l e i n t h e p o s t e r i o r t h i r d ; s c u t e l l u m g r a n u l a t e d ; met a n o t u m smoot h ; p ro p o d e u m smoot h , n o t re t i c u l a te r u gose ; fore s u rface w i t h med i a n f u rrow ; fore-w i n g h y a l i n e , w i t h o u t d a rk t r a n s ver s a l b a n d s ; dors a l ;o rocess of gonoforce!.)s

1 2 94

Mem . A me r . E n ! .

I n st . ,

no.

37 ,

1 984

F i g . 880 - C he l a of Pse u clogon a top u s clichrom us R . C . L . B u n d a be r g ( Quee n s l a n d )

( F i g . 881 )

lon g ,

s l e n d e r a n d po i n t e d ;

max i l l a ry p a l p i

Perk i n s f rom

w i t h 3-4 segme n t s ;

l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u r s 1 , 0 , 1 . L O C U S TYP I C U S : B u n d a b e r g ( Queens I a n d , A u s t r a I i a ) T Y P I CAL MAT E R I A L : l ec t o t y pe F ' a n d 4 p a r a l ec t o t y pes ( 3 F F , M ) ! ( de s i g n a ted b y O l m i 1 98 2 ) i n El ; 5 p a r a l ec t o t y pes ( 4 F F , 1 M ) ! i n El M . HOST S : i n A u s t r a l i a Lib urnia s p . ( R . C . L . P er k i n s 1 906c ) ; b red i n t h e H a w a i i a n I s I a n d s from Perkinsiella s a cch aricicla K i rk . ( Delph a c idae ) . D I ST R I B U T l m.J : A u s t ra l i a n reg i on : AUST RAL I A : B u n d a berg ( Queens l a n d ) , O L ! B ! B M ! Red l y n c h ( C a i r n s , Q u een s l a n d ) ( R . C . L . Perk i n s 1 905 ) .

4 . Pseu dogon a top u s saccharetorUln R . C . L . P e r k i n s

Pseu dogon a topus s a c c h a retorum R . C . L . Perk i n s 1 905 : 36 . Pseu dogon a topus s a c c h a retorum r' . C . L . P e rk i n s : I< i ef f e r 1 9 1 4 b : 84 .

F E MAL E :

a p tero u s ;

l e n g t h 2 , 92-3 , 25 m m ;

t e s t aceous-red d i s h

( I i g h t or

d a rk ) , w i t h p e t i o l e b l a c k and a b domen b ro w n - r ed d i s h ; l a s t a n t e n n a I seg men t w h i t e ; b o d y s h i n y , h a i r l e ss , w i t ho u t sc u l p t u re ; head ( F i g . 882 C ) e x c a v a ted ; p ro n o t u m ( F i g . 882 A, B ) c rossed b y a s t ro n g t r a n s v e r s a l i m press i on ; m e t a n o t u m w i t h s i des p ro t r u d i n g ; p ro t r u s i o n s r o u n d e d ; mesome t a p l e u r a l s u t u re o-J s u i f..; t ,_, ; rT ! C t 2 t , /r a x T ) " ( c> U • 882 A , B ) w i t h deep , med i a n f u rrow ; p l e u ra a n d pos te r i o r s u rface of p ropode u m

G o n a topod i n ae

1 295

F i g . 8 8 1 - M a l e g e n i t a l i a of Pseu dogon a top u s dichrom u s R . C . L . Perk i n s from B u n d a b erg ( Queen s l a n d ) : d i fferen t s h a pes o f dors a l p rocess of gonoforce p s .

t r a n s v e r s e l y s t r i a te ; seg m e n t 4 o f fron t t a r s u s l on g er t h a n segmen t 1 ( 30 : 24 ) ; en l a r g e d c l a w ( F i g . 883 ) w i t h s u b a p i c a l toot h a n d 7-8 l a me l l a e ; segmen t 5 o f f ro n t t ar s u s ( F i g . 883 ) w i t h t w o rows o f 1 4- 1 6 l a me l l ae ; a pex w i t h a g ro u p of 1 1 - 1 4 l a me l l a e ; m a x i l l a r y p a l p i w i t h 4 seg men t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MA L E : f u l l y w i n g e d ; l e n g t h 2 , 1 9 m m ; b row n - t e s t aceo u s , w i t h a n te n n ae , l e g s , a b domen a n d f ro n t p a r t of h e a d l i g h t ; a n t e n n a e not d i s t a l l y t h i c k ened ; a n t e n n a l segmen t s i n f o l l o w i n g proport i o n s : 6 : 5 : 1 0 : 9 : 9 : 8 : 8 : 8 : 8 : 1 0 ; a n t en n a l segme n t 3 n e a r l y f i v e t i mes a s l o n g a s b road ; h e a d d u l l , w e a k l y g r a n u l a ted ; t em p l e s shor t , b u t d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L � 6 ; O L � 2 , 5 ; O O L � 4 ; sc u t u m d u l l , g r a n u l a ted ; n o t a u l i ces co m p l e t e , pos t er i o r l y s e p a r a ted ; m i n i m u m d i s t a nce b e t ween t h e n o t a u l i ces shorter t h a n b re a d t h of t h e oce l l i ( 1 , 5 : 4 ) ; scu t e l l u m a n d m et a n o t u m s h i n y , smoo t h ; p ropod e u m s h i n y , w i t h dorsa l s u r face smoo t h , not ret i c u l a t e r u gose , w i t h med i a n f u r row ; pos t e r i o r s u rf a c e ret i c u l a t e r u g os e ; fore-w i n g h y a l i n e , IV i t ho u t d a rk t r a n s v e rs a I b a n d s ; dorsa I p rocess of gonoforceps ( F i g . 884 ) l o n g , d i s t a l l y poi n t ed ; m a x i l l a ry p a l p i w i t h 4 s e g m e n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . L O C U S TYP I C U S : H a m b l edon ( A u s t ra l i a )

1 296

Fig.

Mem . Amer . E n t .

I nst . ,

no . 37 ,

1 984

882 - F e m a l e of Pseudogona top u s s a ccha retorum R . C . L . P e r k i n s f rom H a l i fa x ( Queens l a n d ) ; A , 8 : t ho r a x a n d p ropodeu m ; C : h e a d .

T Y P I CA L MAT E R I AL : l ec to t y p e F ! a n d 1 p a r a l ec t o t y p e M ! ( d es i g n a ted b y O l m i 1 982 ) i n B . HOST S : i n A u s t r a l i a Perkin siella s a c ch aricida K i rk . ( R . C . L . P e r k i n s 1 906c ) • D I S T R I B U T I O N : A u s t r a l i a n reg i on : A U S T R A L I A : H a m b l edon ( C a i r n s , Queen s l a n d ) , B ! H a l i fa x ( Queens l a n d ) , B ! B u n d a b erg ( Q ueens l a n d ) , BR ! S a n d h i l l ( Queen s l a n d ) , OL !

5 . Pseu dogon a top u s kuranda e R . C . L .

Perk i n s

Pseu dogona topus k ur a n da e R . C . L . Perk i n s 1 905 : 35 . Pseu dogon a top u s kura n dae R . C . L . Perk i n s : K i effer 1 9 1 4b : 85 .

F E M A L E : a p t e rou s ; l e n g t h 2 , 74-2 , 92 m m ; y e l l o w - t e s t aceou s , w i t h p e t i o l e b l ac k a n d a b domen b row n ; a n te n n ae b row n , w i t h segme n t s 1 -2 a n d 1 0 y e l l o w ; m e t a t ho r a x + p ropodeum w i t h some b ro w n s p o t s ; body h a i r l es s , s h i n y , w i t ho u t sc u l p t u re ; h e a d e xc a v a t ed ; p ro n o t u m crossed b y a s t r o n g t r a n s v e rs a l i m p ress i on ; m e t a not u m w i t h s i des p r o t r u d i n g ; p ro t r u s i on s rou n d e d ( F i g . 8 8 5 A ) ; meso-m e t a p l e u r a l su t u re obso l e te ; met a t ho r a x + p ropode u m w i t ho u t med i a n f u r ro w ; p l eu r a a n d pos t e r i or s u rface of p ropode u m t r a n s v erse l y s t r i a te ; s e g me n t 4 of f ron t t a rs u s as l on g a s

Gon a topod i n a e

Fig.

883

-

C h e l a of Ps e u Jogon a top us s a c c h aretorum R . C . L . H a l i f a x ( Qu een s l a n d )

1 297

Perk i n s f rom

segmen t 1 ; e n l a r ged c l a w ( F i g . 886 ) w i t h s u b a p i c a l t o o t h a n d 7 l ame l l ae ; segmen t 5 o f f ro n t t a r s u s ( F i g . 886 ) w i t h t w o row s of 2 2 l a me l l ae ; a p e x w i t h a g ro u p of a t l ea s t 1 1 l a me l l ae ; m a x i I l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MA L E : u n k n o w n

1 2 98

M em . A mer .

En!.

I ns! . ,

n o . 37 ,

1 984

F i g . 884 - M a l e g en i t a l i a of Pseu dogon a topus s a cch aretorum R . C . L . P e r k i n s ( p a r a t y pe ) ( l e f t h a l f remov ed )

LOCUS TYP I C U S : Kuranda ( Austra I i a ) T Y P I CA L MAT E R I AL : l e c t o t y p e F ! a n d 1 p a r a l ec t o t y p e F ! ( d es i g n a ted b y O l m i 1 982 ) i n B . HOST S : i n A u s t r a l i a Lib urnia sp . ( R . C . L . Perk i n s 1 906c ) . D I S T R I B U T I O N : A u s t r a l i a n reg i o n : AUST RAL I A : K u r a n d a ( C a i rn s , Queens l a n d ) , B !

6 . Pseudogo n a top u s jun cetorum R . C . L . Perk i n s

Pseu dogon a topus juncetorum R . C . L . Perk i n s 1 90 5 : 36 . Ps eu dogon a top u s p a l u s tris R . C . L . Perk i n s 1 90 5 : 36 ( n . sy n . ) ; locus typicus Bu n d a b erg ( A u s t r a I i a ) • Pseu dogon a topus juncetorum R . C . L . P e rk i n s : K i effer 1 9 1 4b : 8 4 .

Gona topod i n a e

1 299

F i g . 885 - l a t e r a l prot r u s i o n s of m e t a n o t U nl i n fem a l es of Pseu dogon a top u s kurandae R . C . L . Perk i n s ( t y p e ) ( A ) a n d juncetorum R . C . L . Perk i n s ( type ) ( B ) .

Pseudogon a top us p a l u s tris R . C . l . P e rk i n s : K i effer 1 9 1 4 b : 85 .

FEMALE :

a p t e rou s ;

l e n g t h 2 , 37-3 , 47 m m ;

t e s t a ceous-red d i s h ,

w i t h p et i o l e

b l ac k a n d a b domen b ro w n o r b r o w n - b l ac k ; a n te n n a e b row n , w i t h seg m e n t s 1 -2 a n d 1 0 y e l l o w ; b o d y s h i n y , h a i r l es s , w i t ho u t scu l p t u r e ; h e a d exc a v a te d ; p ro no t u m c r o s s e d b y a s t ro n g t r a n s v e rs a l i m p re s s i o n ; m e t a n o t u m w i t h s i d e s p r o t ru d i n g ; prot r u s i o n s r o u n d e d ( F i g . 885 B ) ; meso-m e t a p l e u r a l s u t u re o b so l e t e ; m e t a t h o r a x + p ropodeum w i t ho u t med i a n f u r row o r w i t h t r ack o f med i a n f u r ro w ; p l e u r a a n d pos t e r i or s u rface of propode u m t r a n s v e rse l y s t r i a t e ; se g m e n t 4 of f ron t t a rs u s as l o n g as seg men t 1 ( i n l ec t o t y p e of Ps . p a l u s tris ) or s h o r t e r t h a n s e g m e n t 1 ( 25 : 30 : i n l ec t o t y p e of Ps . juncetorum ) ; e n l a rged c l a w ( F i g . 887 ) w i t h s u b a p i c a l too t h a n d 6 l a m e l l ae ; seg m e n t 5 of f ro n t t a r s u s ( F i g . 887 ) w i t h t wo ro w s of 1 4 l a m e l l ae ; a p e x w i t h a g ro u p of 8- 1 5 l a m eJ l ae ; m a x i l l a ry p a l p i w i t h 4 s e g me n t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n l O C U S T Y P I C U S : C h i l ders ( A u s t ra l i a ) T Y P I CAL M A T E R I AL : l e c to t y p e F ! ( de s i g n a t e d b y O l m i 1 982 ) of Ps . juncetorum i n B ; l ec t o t y p e F ! a n d 2 p a r a l ec to t y p e s F F ! ( des i g n a ted b y O l m i 1 982 ) o f Ps . p a l u s tris i n B . HOST S : i n A u s t ra l i a Lib urnia s p . ( R . C . L . Perk i n s 1 906c ) . D I ST R I B U T I O N : A u s t r a l i a n reg i o n : A U S T R A L I A : C h i l ders ( Qu e e n s l a n d ) , B ! B u n d a b e r g ( Q ueens l a n d ) , B ! B r i s b a n e ( Qu e e n s l a n d ) , B !

7 . Pseudogona topus pluri s u l c a t u s n . s p .

F E M AL E : a p t e ro u s ; l e n g t h 2 , 68 m m ; t e s t a ceous-re d d i s h , w i t h p e t i o l e b l ac k ; a n te n n a e d i s t a l l y t h i c k e n e d ; a n t e n n a l s e g me n t s i n fo l l ow i n g p ropor t i on s : 9 : 5 : 1 8 : 8 : 7 : 6 , 5 : 5 , 5 : 6 : 5 : 9 ; h e a d v e r y e xc a v a ted , s h i n y , s moot h , w i t ho u t sc u l p t u re ; frons w i t h m ed i a n , com p l e t e k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 1 ; Ol = 1 ; O O l = 7 ; p ro n o t u m s h i n y , smoo t h , w i t ho u t s c u I p t u r e , crossed b y a s t r o n g t r a n s versa I i m p ress i on ; sc u t u m s h i n y , w i t hou t sc u l p t u r e ; m e t a n o t u m t r a n s ve r se l y s t r i a te , w i t h s i des ro u n ded ; meso-m e t a p l e u r a l s u t u re o b so l e t e ; p l e u r a a n d pos t e r i o r s u rface of p ropod e u m s t ro n g I y , t r a n s v erse I y s t r i a t e ; a n t e r i or s u rface of p ropod e u m l o n g i t u d i n a l l y s t r i a te ; fore t a r s a l s e g men t s i n fo l l o w i n g propor t i on s : 1 3 : 2 : 4 : 1 5 : 23 ; e n l a rged c l a w ( F i g . 888 ) w i t h s u b a p i c a l too t h a n d 6

1 300

Fig.

Mem . A m e r . E n ! .

886

-

I ns! . ,

no.

37 ,

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C he l a of Pse u dogona top u s kurandae R . C . L .

Perk i n s ( t y p e )

l ame l l ae ; se g m e n t 5 o f f ro n t t a r s u s ( F i g . 8 8 8 ) w i t h t w o rows o f 1 9 l am e l l ae ; a p e x w i t h a g r o u p of a t l e a s t 1 5 l a m e l l ae ; m a x i l l ary p a l p i 4 segmen t s ; l ab i a l p a l p i w i t h 2 seg men ts ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : I rod R a n g e ( Queens l a n d , A u s t r a l i a ) T Y P I CAL MAT E R I AL :

holotype F !

i n UO.

with

Gon a topod i n ae

F i g . 887

-

1 30 1

C he l a o f Pse u dogo n a top u s j u n cetorum R . C . L .

HOST S : u n k no w n D I ST R I B U T I O N : A u s t r a l i a n reg i o n : A Li S T R A L l A : Pe n i n s u l a , N O u e e n s l a n d ) , 'Y' !

Perk i n s ( t y p e )

I rod R a n g e ( C a pe York

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C he l a o f Pseu dogon a top us pl uris u l c a tus n . sp .

( type)

8. Pseu dogon a topus op a c u s R . C . L . Perk i n s

Pseu dogon a topus op a e u s R . C . L . Perk i n s 1 905 ( o n l y fem a l e ) : 3 8 . nee Pseudogon a topus op a e u s R . C . L . Perk i n s 1 905 m a l e : 38 .

Gon a t opod i n ae

F i g . 889 - C h e l a of Pseu dogon a top u s op a c u s R . C . L .

1 303

Perk i n s ( t y p e )

F E MA L E : a p t e ro u s ; l e n g t h 2 , 1 9-3 , 29 m m ; h e a d b ro w n -red d i s h ; a n te n n a e b ro w n-re d d i s h , w i t h segmen t s 1 -2 y e l l ow ; t h o r a x a n d propodeu m b ro w n re d d i s h ( exce p t f o r a p e x o f p ropodeum y e l l ow ) ; p e t i o l e b l ack ; a b domen b row n-red d i s h ; l e g s b ro w n - red d i s h , w i t h t a r s i a n d fore t i b i ae y e l l ow ; head e x c a v a t e d , d u l l , g r a n u l a t ed ; p r o n o t u m d u l l , g r a n u l a t ed , crossed b y a s t r o n g t r a n s v e r s a I i m p ress i o n ; meso-m e t a p I e u ra I s u t u re o b so l e t e ; m e t a n o t u m w i t h o u t t r a n s v e rs a l k ee l s , w i t h s i des ro u n ded ; p l eu r a a n d p o s te r i or s u rface o f p ro p o d e u m i n d i s t i nc t l y , t r a n s v erse l y s t r i a t e ; s e g m e n t

1 304

Mem . A m e r .

Ent .

I nst . ,

no.

37 ,

1 9 84

4 of f ro n t t a r s u s s l i g h t l y l on g e r t h a n seg men t 1 ( 26 : 25 ) ; en l a r g e d c l a w ( F i g . 889 ) w i t h s u b a p i c a l too t h a n d 7 - 1 1 l a m e l l ae ; segmen t 5 o f fron t t a rs u s ( F i g . 889 ) w i t h t w o rows of 1 3- 2 1 l am e l l ae ; a p e x w i t h a g ro u p of a t l e a s t 1 2 l a m e l l ae ; m a x i I l a ry p a l p i w i t h 4 s e g me n t s ;

l ab i a l p a l p i w i t h

2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n ow n ; t h e on l y k n o w n m a l e ( p a r a l ec t o t y p e ) i s re a l l y a m a l e s p ec i m e n of !/aplogo n a top u s a p ic a lis R . C . L . P e r k i n s ( p a l p a l ra t i o 2/ 1 ) . L O C U S TYP I CU S : B u n d a b e r g ( A u s t r a l i a ) T Y P I CA L M A T E R I AL : l ec t o t y pe F ! a n d 2 p a r a l ec t o t y p e s ( 1 F , M)! ( des i g n a t e d b y O l m i 1 982 ) i n B ( t h e p a r a l ec t o t y pe M i s re a l l y a m a l e spec i m e n ' o f fla plogona topus a,�ica lis R . C . L . I-er'k i n s ) . HOST S : i n A u s t r a l i a Liburnia sp . ( ne a r ,-'h 1orion a ) ( R . C . L . Perk i n s 1 906c ) . D I S T R I B U T I O N : A u s t r a l i a n reg i on : A U S T R A L I A : B u n d aberg ( U ueen s l a n d ) , B ! B r i s b a n e ( Queens l a n d ) , B !

6 . Donisthorp ina R i c h a r d s

Donisthorpina R i c h a r d s 1 939 : 2 0 1 . T Y P E S P E C I E S : Donisthorp i n a form icicola R i c h a r d s 1 939 � 1) . p a l l ida ( C e b a l l os 1 927 ) , o r i g . des i g . F E MA L E ( F i g s 894 , 895 ) : a p terou s ; m a x i I l a r y p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; p r o n o t u m ( F i g . 895 ) crossed b y a s t ro n g t ra n s v e rs a l i m p ress i o n ; en l a rged c l a w ( F i g . 890 ) w i t h s u b a p i c a l too t h a n d l am e l l ae ; t i b i a l s p u rs 1 , 0 , 1 . M A L E ( F i g s 8 9 1 , 892 ) : f u l l y w i n g ed ; m a x i l l a ry p a l p i w i t h 5 s e g me n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; occ i p u t con c a v e ; occ i p i t a l c a r i n a i n v i s i b l e o r o n l y v i s i b l e b e h i n d t h e oce l l i o r o n t he s i des o f t h e pos t er i o r oce l l i , b u t n e v e r reach i n g t e m p l e s ; t e m p l es u i s t i n c t ; t i b i a l s p u r s 1 , 1 , 2 . D I ST R I B U T I O N : P a l a e a rc t i c , O r i e n t a l HOST S : Delp h a cidae SPEC I ES : 2

K E Y TO T H E SP E C I E S

F EMALES

B od y s l e n d e r ; s c u t u m l on g t a r s u s w i t h 1 r o w o f l am e . . . . . . . . . . . . . . . . . . . . . . . . . Body rob u s t ; s c u t um sh o r t w i t h 2 r o w s o f l am e l l a e •

•

a n d n a r r ow ; s eg m e n t 5 o f f r o n t l l ae . . . . a s ia tica n . sp . ( O r i en t a l ) a n d b r o a d ; s eg m e n t 5 o f f r o n t t a r s u s p a llida ( C eb a l l o s ) ( P a l a ea r c t i c ) .

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Go n a topod i n ae

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1 306

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I ns! . ,

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1 984

F i g . 89 1 - M a l e o f Don isthor<'Jina p a llida ( Ce b a l l o s ) P a r t ec i p a n z a ( I t a l y )

Donisthorpina p a lhda

( C e b a 1 1 05 )

from Bosco d e l l a

n . com b .

Dicondylus p a llidus Ceb a l l os 1 9 27 : 1 07 . Donisthorpin a form icicola R i c h a r d s 1 939 : 2 0 1 ( n . sy n . ) ; locus typ icus M a t l e y Gog ( E n g l a n d ) . Donisth ori'J i n a form icicola !� i c h a r d s : C u rrado a n d 0 1 m i 1 974b : 3 9 .

F E MA L E : a p tero u s ; l e n g t h 3 , 00 -3 , 1 8 m m ; t e s t a c eo u s ; occa s i on a l l y v e r t e x , o c c i p u t , t em p l es , p l e u r a , s c u te l l u m , m e t a no t u rn a n d p a r t o f p ro p o d e u m b row n i s ll ; a n t e n n a e b ro w n , w i t h segmen t s 1 -3 y e l l o w ; p e t i o l e b l ac k ; p a rt

Go n a topod i n ae

1 30 7

F i g . 892 - !,! a l e o f iJon isth ol','J inD. ,c; a lliJa ( Ce b a l l o s ) f rom Bosco d e l l a P a r t ec i p a n z a ( I t a l y )

of a b domen b ro w n -b l ac k ; a n t en n a l seg men t s i n fo l l o w i n g propo r t i o n s : 7 : 5 : 1 1 : 7 : 5 : 5 : 4 : 4 : 4 : 7 , 5 ; h e a d s h i n y , exc a v a ted , w i t h o u t scu l p t u re , excep t for f ro n t p a r t of v e r t e x a n d occ i p u t g r a n u l a t e d ; p ro n o t u m sh i n y , a l u t ac eo u s , crossed b y a s t ron g t ra n s v e rsa l i m p ress i on ; p l eu r a a n d p os t e r i o r s u rface of p ropode u m t ra n sv e r se l y s t r i a te ; m e t a t h orax + p r o p o d e u m s h i n y , w i t ho u t sc u l p t u re ; meso-me t a p l e u ra l s u t u re o b so l e t e ; fore t a rsa l se g m e n t s i n fo l l o w i n g p ropor t i on s : 1 2 : 2 : 4 : 1 2 : 1 8 ; e n l a r g e d c l a w ( F i g . 890 ) w i t h s u b a p i c a l too t h a n d 4-6 l a m e l l ae ; segmen t 5 of f r o n t t a rs u s ( F i g . 890 ) w i t h t w o r o w s of 1 7-22 l a me l l ae ; a pex w i t h a g r o u p of 6- 1 2 l a m e l l a e ; m a x i l l ary p a l p i w i t h 5 s e g me n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l S p U i'S 1 , 0 , 1 . ;,1 A L E ( F i g s 8 9 1 , 89 2 ) : f u l l y w i n g e d ; l e n g t h 2 , 62 m m ; b l ac k , w i t h m a n d i b l e s , t e g u l a e , l eg s a n d a n te n n a l seg m e n t s 1 -2 ye l l ow ; a n t e n n a l seg m e n t 3 more t h a n four t i me s a s l o n g a s b ro a d ( 4 , 33 ) ; h e a d s h i n y , h a i r y , w i t ho u t SCU l p t u r e or w e a k l y p u n c t a t e ; fron t a l l i n e a b s e n t ; t e m p l es d i s t i n c t ; OCC i p i t a l ca r i n a i n v i s i b l e ; sc u t u m h a i ry , g r a n u l a t e d ; n o t a u l i ce s com p l e t e , pos t e r i or l y j o i n t ; sc u t e l l u m a n d m e t a n o t u m s h i n y , w i t ho u t scu l p t u re or w e a k l y p u n c t a t e ; p ropod e u m re t i c u l a te ru gose , w i t ho u t k ee l s ; dorsa I p rocess of gonoforceps ( F i g . 893 ) very s hor t , d i s t a I I y b ro a d e n e d ; m a x i l l a r y p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l spurs 1 , 1 , 2 . LOCUS TYP I C U S : E I P a rdo ( Sp a i n ) T Y P I CAL M A T E f� I A L : h o l o t y p e F ! of iJ . pallidus i n M D ; h o l o t y p e F ! of D .

forw icicola i n B M .

1 308

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I n s! . ,

no.

37 ,

1 984

F i g . 893 - M a l e g en i t a l i a of Don isthorp in a pall ida ( Ce b a l l os ) from Beme l e n ( Ho l l a n d ) ( r i g h t h a l f removed )

HOST S : i n I t a I y Xa n thodel p h a x stramineus ( St a I ) ( Delph a cida e ) ( C u r r a d o a n d O l m i 1 97 4 b ) ; i n Ho l l a n d Rib a u todelp h a x pungens ( R i b a u t ) a n d Muellerianella fairmairei P e rr i s ( = extr u s a Sco t t ) ( C . F . M . d e n B i em a n leg. ) • D I ST R I BU T I ON : P a l a e a rc t i c reg i o n : JAPAN : H i rosh i m a , K Y ! H U N GA R Y : O . Sebes h e l y , BU ! H O l lA N D : Beme l en , B E ! O l ! E N G l A ['>J D : M a t l e y Bog ( New Forest , H a n t s Sou t h ) , BM ! SPA I N : E I P a r d o ( M a d r i d ) , M D ! I T ALY : C a n t a l u p a ( To r i no ) , O L l Mon d a g no l a ( F rabosa Sop ra n a , C u neo ) , O l ! Bosco d e l l a P a r t ec i p a n z a ( T r i no Verce l l ese , Verc e l l i ) , O l ! A L G E R I A : Bon a ( A b d u l - N o u r 1 974 ) .

O R I E N T A L S P E C I E S O F DONISTHOi2PINA

O ne s p ec i es i s k n ow n .

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1 309

894 - Fem a l e of UonisthoriJ i n a a s i a ti c a n . s p .

Don i s thorp i n a a s i a t i c a n .

( t y pe )

sp .

F E MA L E ( F i g s 894 , 895 ) : a p t e rou s ; l en g t h 3 m m ; t e s t aceou s , w i t h p e t i o l e b l ac k j head exc a v a ted , s h i n y , a l u t aceo u s ; p ronot u m sh i n y , h a i r l ess , a l u t aceo u s , c rossed by a s t ron g t r a n s v e r s a l i m p res s ion ; scu t u m very s l e n d e r , s h i n y , w i t ho u t sc u l p t u re ; m e t a n o t u m s h i n y , w i t h o u t sc u l p t u re , not st r i a t e ; m e t a t ho r a x + p ropod e u m s h i n y , w i t ho u t scu l p t u re ; p l e u r a a n d poste r i o r s u rf a c e o f propo d e u m t r a n s verse l y s t r i a t e ; meso-me t ap l e u r a l s u t u re o b so l e t e ; fore t a r s a l se g m e n t s i n fo l l o w i n g p roport i o n s : 1 3 : 2 : 4 : 1 3 : 1 9 ; e n l arged c l a w ( F i g . 896 ) w i t h s u b a p i c a l too t h a n d 6 l a m e l l a e ;

1310

Mem . Amer.

Ent.

I ns t . ,

no . 37 ,

1 984

F i g . 895 - F em a l e of Don isthorpina a s ia tica n . sp .

( type)

segme n t 5 o f f ron t t a rs u s ( F i g . 896 ) w i t h 1 row o f 1 2 l a m e l l ae ; a p e x w i t h a g ro u p of 8 l a me l l a e ; m a x i I l a r y p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k no w n L O C U S TYP I C U S : Gom b a k ( M a l a y a , M a l a y s i a ) TYP I C A L M A T E R I A L : h o l o t y p e F ! i n B M . HOST S : u n k n o w n D I STR I B U T I O N : O r i e n t a l reg i o n : MALAYS I A : Gom b a k

7 . A gona to,rJoides R . C . L .

( Ma l ay a ) ,

B fv\ !

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A gona topoides R . C . L . P e r k i n s 1 907 : 33 .

T Y P E S P E C I E S : Agona topoides synchroIilus R . C . L . P e r k i n s 1 907 , mono t y p i c . F E M A L E ( F i g s 906 , 907 ) : a p terou s ; m a x i l l a ry p a l p i w i t h 5 s e g me n t s ; l a b i a l p a l p i w i t h 3 segmen ts ; p ro n o t u m ( F i g . 907 ) c rossed b y a s t ron g t r a n s v e rsa l i mp ress i on ; e n l a r g e d c l a w ( F i g . 897 ) w i t h s u b a p i c a l too t h a n d l a m e l l a e ; t i b i a l s p u rs 1 , 0 , 1 . MALE ( F i g s 899 , 900 ) : f u l l y w i n ge d ; m a x i l l a r y p a l p i w i t h 5 seg m en t s ; l a b i a l p a l p i w i t h 3 s e g m e n ts ; occ i p u t conc a v e ; occ i p i t a l c a r i n a i n v i s i b l e o r v i s i b l e on l y b e h i n d t h e oce l l i o r o n t h e s i de s o f t h e pos te r i or oce l l i , b u t n e v e r r e a c h i n g temp l e s ; temp l es d i s t i nc t ; t i b i a l s p u r s 1 , 1 , 2 . D I ST R I B U T I O N : P a l a e a rc t i c , E t h i o p i a n , O r i en t a l , N e a r c t i c , Neo t rop i c HOST S : Delp h a cidae

Go n a topod i n a e

F i g . 896

-

C he l a of Don isthorpina a s i a tica n . sp .

1 31 1

( type)

1 31 2

Mem . Amer. E n ! .

SPEC I ES :

I ns! . ,

n o . 37 ,

1 984

15

PALAEA R C T I C S P E C I E S OF A GONA TOPOIDES

T h ree spec i es are k n ow n .

K E Y TO T H E SPEC I E S

F E MA L E S

2

M e t a n o t u m h o l l ow b e h i n d t h e s c u t e l l u m ; m e s o -m e t a p d i s t i n c t , s t r on g . . . . . . . . . . . 1 . felix n . sp . M e t a n o t u m n o t h o l l ow b eh i n d t h e s cu t e l l u m , a l mo s t -me t ap l eu r a l s u t u r e ob s o l e t e o r i n d i s t i n c t . . . . . . . B od y a l mo s t f u l l y b l a ck 2 . solidus ( H a u p t ) B od y a l mo s t f u l l y r ed d i s h - t e s t a c e ou s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 . can a riensis n . sp . •

•

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D o r sa l p r o c e s s of g on o f o r c ep s ( F i g . · . . . . . . . . . . . . . . . . D o r sa l p r o c e s s o f g on o f o r c ep s ( F i g . . . . . . . . . . . . . . . . . ·

T h e m a l e of A .

g on o f o r c e p s 9 0 1 ) ; n o t au . . . . . . . . . .2. g on o f o r c e p s 90 3 ) ; n o t au . . . . . . . . . . 3.

v e ry s h o r t , mu ch s h o r t e r t h a n l i c e s j o i n t ; ra d i a l c e l l s ma l I solidus ( H a u p t ) v e r y l on g , n ea r l y a s l on g a s l i c e s s ep a ra t ed ; ra d i a l c e l l w i d e can ariensis n . sp .

felix i s u n k n ow n .

1 . A gona topoides felix n .

sp.

F E M A L E : a p te rou s ; l e n g t h 2 , 1 8- 2 , 68 m m ; red d i s h - t e s t a ceou s , w i t h p et i o l e b l ac k a n d a b domen b row n ; a n te n n a e w e ak l y , d i s t a l l y t h i c k e n e d ; a n t e n n a I seg m e n ts i n fo l l o w i n g p ropor t i o n s : 8 : 5 : 1 2 : 6 : 6 : 6 : 5 : 4 : 4 : 6 ; head d u l l , f u l l y g r a n u l a t e d , w e a k l y e x c a v a t e d ; occ i p i t a l c a r i n a i n v i s i b l e ; fron t a l l i ne a b se n t ; POL = 2 ; O L = 3 ; OOL = 8 ; p ro n o t u m s h i n y , crossed by a s t ron g t r a n s v e rs a l i m p ress i o n , w i t h i n d i s t i nc t l o n g i t u d i n a l s t r i ae ;

1 31 3

Go n a topod i n ae

F i g . 897

-

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n .

sp .

( t y pe )

1 31 4

Fig.

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I nst . ,

no . 3 7 ,

1 984

898 - C he l a of Agon a topoides solidus ( H a u p t ) from S a n i co d i A l f i a n o N a t t a ( I ta l y )

Gon a topod i n a e

F i g . 899

-

1315

M a l e o f ilgon D. to , ) oides sol i d u s ( H a u p t ) from M a sserano ( I t a l y )

s c u t u m g r a n u l a ted , d u l l ; s c u t e l l u m d i s t i n c t ; m e t a n o t u m t r a n s verse l y s t r i a t e , h o l l o w be h i n d t h e sc u te l l u m , w i t h s i des p r o t r u d i n g ; protru s i on s pO i n t ed ; meso-me t a p l eu r a l s u t u r e d i s t i nc t , com p l e t e ; m e t a t horax + p ropode u m s h i n y ; a n t e r i o r s u rf a c e ',. i t ll l on g i t u d i n a l , i n d i s t i nc t s t r i ae ; pos ter i o r s u rface tra n s v e rse l y s t r i a t e ; fore t a rsa l seg men ts i n fo l l ow i n g proport i o n s : 1 2 : 2 : 5 : 1 3 : 20 ; en l arged c l a w ( F i g . 897 ) w i t h s u b a p i c a l too t h a n d 6 l a me l l ae ( i n a d d i t i o n t o 1 b r i s t l e ) ; seg men t 5 of fron t t a rs u s ( F i g . 897 ) w i t h 2 row s o f 1 0 l a m e l l a e ; a p e x w i t h a g ro u p o f 1 2 l am e l l a e ; m a x i l l a r y p a l p i w i t h 5 s e g me n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : I� i o C a b r a ( Ca n ary I s l a n d s ) TYP I CAL MAT E R I A L : h o l o t y p e F ! a n d 2 p a r a t y p es F F ! i n M D ; 2 p a r a t y pes F F ! in O L . HOST S : u n k no w n D I ST R I B U T I O N : Pa l ae a rc t i c reg i o n : C A I"ARY I SL A N D S : R i o C a b r a ( F u e r te v e n t u r a ) , M D ! O L !

1316

Fig.

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En!.

I ns! . ,

no . 37 ,

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900 - M a l e of A gona to,'Joides solidus ( H a u p t ) from M a sserano ( I t a l y )

2 . Ilgon a to[Joides solidus ( H a u p t ) n . com b .

A llogon a topus Agona topoides locus typicus Agona topoides

solidus H a u p t 1 938 : 2 7 . joh a n n a e C u r r a do a n d O l m i 1 975 : 28 ( n . sy n . ) ; San i co d i A l f i a no N a t t a ( I t a l y ) . joh a n n a e C u rr a d o a n d O l m i : O l m i 1 98 1 : 1 1 6 .

F E MALE : a p t e rou s ; l en g t h 3 , 4 m m ; h e a d b ro w n , w i t h m a n d i b l es , c l y p e u s a n d f ron t p a r t of v ertex y e l l ow ; a n t e n n a e b row n-b l ac k , w i t h s e g m e n t s 1 -3 y e l l ow ; t ho r a x a n d propode u m b l ac k ; a b domen b ro w n -b l ac k ; l e g s y e l l o w , w i t h p a r t of coxae a n d p a r t of c l u b s of femora b l ac k ; h e a d e x c a v a ted , s h i n y , gra n u l a ted , h a i r l ess ; prono t u m h a i r l es s , g r a n u l ated , crossed b y a s t ron g t r a n s v e rsa l i m press i o n ; s c u t u m s h i n y , g r a n u l a ted , w i t h some l o n g i t u d i n a l k ee l s ; sc u t e l l u m s h i n y , g ra n u l a te d ; m e t a n o t u m tra n s verse l y s t r i a t e ; m e t a t horax + propodeum s h i n y , g r a n u l a ted ; a n te r i or s u rface of m e t a t ho r a x + p ropodeum w i t h track of med i a n f u rrow ; p l e u ra a n d pos t e r i o r s u rf ace o f p ropod e u m t r a n s v erse l y s t r i a t e ; meso-me t a p l e u ra l s u t u re o b so l e t e ; fore t a rsa l seg m e n t s i n fo l l o w i n g p ropor t i on s : 1 4 : 2 : 3 : 9 : 1 4 ; en l a r g e d c l a w ( F i g . 898 ) w i t h s u b a p i c a l too t h a n d 6-7 l a me l l a e ; seg m e n t 5 o f f ro n t t a r s u s ( F i g . 898 ) w i t h t wo rows o f 1 9 l a me l l ae ; a p e x w i t h a g ro u p of 4-5 l am e l l ae ; m a x i l l a ry p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 . r,1ALE ( F i g s 899 , 900 ) : f u l l y w i n g e d ; l en g t h 3 m m ; head b l ac k , w i t h

Gonatopod i n ae

1 31 7

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F i g . 90 1

-

M a l e gen i t a l i a of A gona topoides sohdus C a s te l l a l fero ( I t a l y ) ( r i g h t h a l f removed )

( Ha u p t )

from

1318

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I nst . ,

no . 37 ,

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m a n d i b l es test aceo u s ; a n t e n n a e b l a c k - b row n ; t horax a n d propod e u m b l ack ; a b do m e n b row n ; l e g s y e l l ow - b row n , w i t h t a r s i l i g h t ; a n t en n a l seg men t 3 more t h a n four t i mes as l o n g as b ro a d ( 5 , 6 ) ; h e a d s h i n y , h a i r y , g ra n u l a ted ; f ro n t a l l i n e a b s en t ; temp l es d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e ; scu t u m s h i n y , g r a n u l a ted ; not a u l i ce s comp l e te , poste r i o r l y j o i n t ; s c u t e l l u m a n d m e t a n o t u m s h i n y , wea k l y p u n c t a te ; p ropod e u m w i t h pos t e r i or s u rface ret i c u l a te ru gose ; a n te r i o r s u rface w i t h areo l ae sma l l a n d w i t h med i a n f u rrow ; poster i or s u rface w i t h areo l ae w i de ; dorsa l p rocess of gonoforceps ( F i g . 90 1 ) short , d i s t a l l y s l e n d e r ; m a x i I l a ry p a l p i w i t h 5 segmen t s ; l a � i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 1 , 2 . LOCUS TYP I CU S : Bel l i nchen ( German y ) TYP I CA L M A T E R I AL : h o l o t y p e F ! of A . solidus i n HA ; h o l o t y p e F ! o f A . joh a n n a e i n E N . HOST S : i n I t a I y Mega delp h a x sp . a n d L a odelp h a x s tria tell u s ( F a l i e n ) Welp h a cida e ) ; i n Germ a n y Muirodelph a x a u bei ( P e r r i s ) ( = L ib urnia a u b e i ) ( H a u p t 1 938 ) . D I ST R I BU T I O N : P a l a e a rc t i c reg i o n : G E R M AN Y : B e l l i nc h e n ( O d e r ) , HA ! I TALY : M asserano ( Verce l l i ) , O L ! C a s te l l a l fero ( As t i ) , E N ! Sa n i co d i A l f i a n o N a t t a ( A l es s a n d r i a ) , E N !

3 . Agona tofJoides c a n ariensis n .

sp .

F E MA L E : a p t erou s ; l e n g t h 3 , 56 m m ; red d i s h - t e s t aceou s , w i t h pos ter i or s u rface of p ropod e u m d a r k ; pe t i o l e b l ack ; a b domen b row n ; a n t e n n ae b ro w n , w i t h s e g m e n t s 1 -2 test aceou s ; a n t e n n a e w e a k l y d i s t a l l y t h i c k e n e d ; a n t e n n a I seg m e n t s i n fo l l o w i n g p ropor t i on s : 7 : 5 : 1 3 : 7 : 7 : 6 : 5 : 5 : 5 : 6 , 5 ; h e a d sh i n y , exca v a ted , wea k l y g r a n u l a ted ; fron t a l l i n e presen t ; occ i p i t a l ca r i n a i n v i s i b l e ; POL = 1 ; O L = 2 ; O O L = 7 ; p ro n o t u m crossed b y a s t ro n g t r a n s v e rs a l i mp ress i o n , d u l l , g r a n u l a ted ; s c u t u m a n d scu te l l u m d u l l , g r a n u l a ted ; m e t a n o t u m n o t h o l l o w b e h i n d t h e sc u te l l u m , a l most f l a t , t r a n s v erse l y s t r i a t e , w i t h s i des rou n d e d ; m e t a t horax + p ropo d e u m d u l l , g r a n u l a t e d , w i t h pos ter i or s u rface a n d p l e u r a t ra n s v e rse l y s t r i a te ; meso-me t a p l eu r a l s u t u r e i n d i s t i n ct ; fore t a r s a l seg men t s i n fo l l ow i n g proport i on s : 1 4 : 2 : 4 : 1 0 : 1 6 ; en l a rged c l a w ( F i g . 902 ) w i t h s u b a p i c a l toot h a n d 7 l am e l l ae ; seg m e n t 5 of f ro n t t a r s u s ( F i g . 902 ) w i t h t w o row s of 20 l a m e l l ae ; a p e x w i t h a g ro u p of 9 l a m e l l ae ; t i b i a l s p u rs 1 , 0 , 1 . MALE : f u l l y w i n g e d ; l e n g t h 2 , 5 m m ; b l ac k ; m a n d i b l es b l ac k , w i t h tee t h test aceou s ; a n t e n n ae b row n ; l eg s b ro w n , w i t h fore t i b i ae test aceo u s ; a n te n n ae n o t d i s t a l l y t h i c k ened ; a n t e n n a I se g m e n t s i n fo l l ow i n g proport i on s : 6 : 4 , 5 : 9 : 9 : 9 : 8 : 8 : 8 : 7 : 9 ; a n t e n n a I se g m e n t 3 four t i mes a s long as b road ; h e a d d u l l , g r a n u l a t e d ; occ i p i t a l c a r i n a i n v i s i b l e ; f ro n s w i t ho u t k ee l s ; POL = 7 ; O L = 3 ; OOL = 4 ; scu t u m d u l l , g r a n u l a ted ; n o t a u l i ces comp l e te , pos ter i or l y s e p a r a t e d ; sc u t e l l u m w e a k l y g r a n u l a ted ; m e t a n o t u m s h i n y , w i t ho u t scu l p t u re ; p ropodeum ret i c u l a t e

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r u gose ; dorsa l s u r face w i t h med i a n f u rrow a n d w i t h v e r y s m a l l a reo l ae ; poster i o r s u rface w i t h w i de areo l ae ; fore- w i n g h y a l i n e , w i t ho u t d a rk t r a n s v e rs a l b a n d s ; gonoforceps w i t h dors a l p rocess ( F i g . 903 ) l on g , n a rrow , n e a r l y a s l on g a s gonoforce p s ; t i b i a l s p u rs 1 , 1 , 2 . L O C U S TYP I C U S : B a rra nco d e l os J u ncos ( C a n a r y I s l a n d s ) TYP I CA L M AT E R I AL : h o l o t y p e F ! i n M D ; 1 p a r a t y p e M ! i n O L .

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F i g . 903 - M a l e g en i t a l i a of A gona to,'Joides c a n a riensis n . s p . f ro m C . no de l a s M e rcedes ( C a n a r y I s l a n d s ) ( r i g h t h a l f remov e d )

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HOST S : u n k n o w n 0 1 S T R I B U T I O N : P a I a e a r c t i c r e g i on : C A N A R Y I SLANOS : B a rr a n co de l o s J u ncos ( T e n e r i fe ) , M O ! C . n o d e l as Mercedes ( T e n e r i fe ) , O L !

E T H I O P I A N SP E C I E S O F AGONA TOPOIDES

F i ve spec i e s are k n ow n .

K E Y T O T H E SPEC I E S

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E n l a rg ed c l aw w i t h a p ex v e ry b r oad

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(F i g . 905 ) 1 fu s c u s n . sp . E n l a rg ed c l aw w i t h a p e x t a p e r i n g , n o t b r o a d ( F i g s 9 0 8 , 9 0 9 , 91 0 , 91 1 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .2 Me t a t h o ra x + p r o p o d e u m w i t h a s t r o n g med i a n f u r r ow ( F i g . 9 0 6 ) .

3

2 . a c u m in a tu s n . sp . Me t a t h o ra x + p r o p o d e u m w i t h o u t s t r o n g med i a n f u r r ow Me t a n o t u m w i t h s i d e s r o u n d ed ; me t a t h o ra x + p r o p o d e u m b l a ck

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. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 . c a fer n . sp . Me t a n o t u m w i t h s i d e s p r o t r u d i n g ; p r o t r u s i o n s r o u n d ed o r s h a r p ; b od y f u l l y r ed d i s h , w i t h p e t i o l e b l a c k 4 H e a d s h i n y , w i t h ou t s c u l p t u r e , e x c ep t f o r o c c i p u t g ra n u l a t ed . . . . . . , . 4 . r u b er n . sp . H ea d d u l l , f u l l y g r a n u l a t ed S . ceres n . sp . ·

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MALES U n k nown 1 . A go n a topoides fu s c u s n . sp .

F E M A L E ( F i g . 904 A ) : a p t e r ou s ; l en g t h 3 , 1 2-4 , 00 m m ; head , a n t e n n a e , t hora x , p ropod e u m , a b domen a n d l eg s tes t aceous-re d d i s h or redd i s h - d a r k ; p et i o l e b l ac k ; occa s i o n a l l y a b do m e n b rown i s h ; h e a d f l a t , d u l l , h a i r y , g r a n u l a t e d ; p ron o t u m d u l l , h a i ry , w i t h n u merous w e a k l o n g i t u d i n a l k ee l s , crossed b y a s t ro n g t r a n s v ersa I i m pres s i on ; s c u t u m d u I I , w i t h some l o n g i t u d i n a l k ee l s ; m e t a n o t u m d u l l , t r a n s v erse l y s t r i a t e , ho l l o w b e h i n d t h e sc u te l l um , w i t h s i des p r o t r u d i n g ; p ro t r u s i o n s po i n ted ; met a t ho r a x + p ropodeum d u l l ; a n te r i o r s u rface w i t h n u merous l o n g i t u d i n a l k ee l s ; p l e u r a

1 322

Mem . Amer .

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I n s! . ,

no.

37 ,

1 984

F i g . 904 - F e m a l e of Agona topoides fu scus n . s p . from Van R e e n e n ( So u t h A f r i c a ) ( A ) ; B : he a d ; C : t h o r a x a n d p ropodeum l a tera l l y v i ewed .

a n d pos t e r i o r s u rf a ce n o t or w e a k l y t ra n s v e rse l y s t r i a t e ; meso- m e t a p l e u r a l s u t u re comp l e t e , d i s t i nc t ; fore t a rsa l segmen t 1 s l i g h t l y s h o r t e r t h a n seg m e n t 4 ( 1 4 : 1 5 ) ; e n l a r g e d c l a w ( F i g . 905 ) w i t h a pe x v e r y b ro a d , w i t h s u b a p i c a l too t h a n d 5-7 l am e l l a e ; segmen t 5 of fro n t t a r s u s ( F i g . 905 ) w i t h t w o ro w s of 1 2- 1 7 l a me l l ae ( s ho r t e s t row exte n d i n g o r not e x t en d i n g b e y o n d 0 , 5 l en g t h of segmen t ) ; a p e x w i t h a g ro u p of 1 0-20 l a m e l l a e ; m a x i I l a ry p a l p i w i t h 5 s e g m e n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k no w n L O C U S TYP I C U S : Mosse l B a y ( So u t h Afr i c a ) TYP I CA L MAT E R I AL : ho l o t y p e F ! a n d 3 p a r a t y pe s F F ! i n B M ; 1 p a r a t y p e F! in OL .

Gon a topod i n ae

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-

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1 323

( t y pe )

1 324

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F em a l e of Agona topoides a c um i n a t u s n .

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( type)

HOST S : u n k n o w n D I ST R I B U T I O N : E t h i o p i a n reg i on : SOU T H A F R I CA : Mosse l B a y ( C a p e P rov i n ce ) , B M ! K a t b e r g ( E C a p e P ro v i n ce ) , OL l BM ! V a n Reenen ( D r a k e n s Berg , N a t a l ) , B M !

2 . A gona topoides a c umin a tu s n . s p .

F E MALE

( F i g s 906 , 907 ) :

a p tero u s ;

l e n g t h 3 , 1 2 mm ;

red d i s h- b ro w n ,

with

1 325

Gon a topod i n ae

F i g . 907

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( type )

m a n d i b l e s , c l y p e u s a n d front p a r t of vertex y e l l ow ; a n ten n ae y e l l ow , w i t h segmen ts 8- 1 0 brow n ; p e t i o l e b l ac k ; t a r s i a n d a p e x of p ropodeum y e l l o w ; h e a d d u l l , exc a v a t ed , g r a n u l a ted ; p ro n o t u m crossed b y a s t ro n g t r a n s v e rs a l i m p ress i on , s h i n y , w i t h s h o r t , i rreg u l a r kee l s ; sc u t u m s c u l p t u red b y l o n g i t u d i n a l k ee l s ; sc u te l l u m smoo t h a n d s h i n y ; m e t a n o t u m w i t h s i des p ro t r u d i n g ; p r o t r u s i o n s p o i n t ed ; m e t a t ho r a x + p ropodeum s h i n y , as scu I p t u re d as p ro n o t u m , wi th a s t ro n g med i a n f u rrow ; meso m e t a p l e u r a l s u t u re d i s t i n c t ; poste r i or s u rf ace of p ropode u m t r a n s verse l y s t r i a t e ; p l e u r a not t r a n sv e rse l y s t r i a t e ; fore t a rsa l seg men ts i n fo l l ow i n g p ropor t i on s : 9 : 2 : 4 : 1 0 : 1 6 ; en l arged c l a w ( F i g . 908 ) w i t h s u b a p i c a l t oo t h a n d 7 l a m e l l ae ; segme n t 5 o f fron t t a r s u s ( F i g . 908 ) w i t h t w o rows of 1 7 l a me l l ae ; apex w i t h a g ro u p of 1 6 l a m e l l ae ; m a x i I l a ry p a l p i w i t h 5 segme n t s ; l ab i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E : u n k no w n L O C U S T Y P I C U S : U p e m b a N a t i on a l P a r k ( Z a i re ) T Y P I CAL M A T E R I A L : ho l o t y p e F ! i n T V . HOST S : u n k n o w n D I ST R I BU T I ON : E t h i op i a n reg i on : Z A I R E : U p e m b a N a t i o n a l P a r k , T V !

3 . A gona topoides cafer n . s p .

1 32 6

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1 984

C he l a of Agon a topoides acumin a t u s n . sp .

( t ype )

G o n a topod i n ae

F i g . 909 - C he l a of A go n a to;)oides c a fer n . s p .

1 327

( type)

F E MAL E : a p terou s ; l e n g t h 3 , 62 m m ; h e a d a n d a n te n n ae redd i s h - d a r k ; a n t e n n a I s e g m e n t 1 0 w h i t e ; p ron o t u m a n d scu t u m t e s t a c eo u s ; sc u te l l u m a n d m e t a t ho r a x + propodeum b l ac k , e x c e p t f o r a p e x of p ro p o d e u m r e d d i s h ; p e t i o l e b l ac k ; a b d o m e n red d i s h - d a r k ; l e g s t e s t aceo u s ; h e a d exc a v ated , d u I I , g r a n u I a t e d ; p ro no t u m crossed b y a s t ro n g t r a n sversa I i m p re s s i o n , s h i n y , smoot h , w i t h some l o n g i t u d i n a l kee l s ; m e t a n o t u m s h o r t , h o l l o w b e h i n d t h e s c u te l l u m , w i t h s i des rou n d e d ; m e t a t h o r a x + p ropodeurn s h i n y , w i t h o u t scu l p t u re ; pos t e r i or s u rf ace of propode u m a n d p l eu r a t r a nsverse l y s t r i a te ; meso- m e t a p l e u r a l s u t u re d i s t i n c t , com p l e t e ; fore t a r s a l segmen t 1 shorter t h a n seg m e n t 4 ( 1 5 : 1 7 ) ; e n l arged c l a w ( F i g . 909 ) w i t h a p e x t a p er i n g , n o t b ro a d , w i t h s u b a p i c a l t oo t h a n d 8 l a m e l l ae ; s e g m e n t 5 o f f ro n t t a r s u s ( F i g . 909 ) w i t h t w o rows of 30 l a me l l ae ; a p e x w i t h a g ro u p of 25 l a me l l ae ; m a x i l l a ry p a l p i w i t h 5 segmen ts ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : M a tjesfon t e i n ( So u t h Afr i c a ) T Y P I CAL M A T E R I A L : ho l o t y p e F ! i n B M . HOSTS : u n k no w n D I ST R I B U T I O N : E t h i o p i a n reg i o n : SOU T H A F R I C A : M a t jesfon t e i n P rov i n ce ) , B M !

( Cape

4 . A gona topoides ruber n . s p .

F E MA L E : a p terou s ; l e n g t h 3 , 43 m m ; red d i s h - b ro w n , w i t h p et i o l e d a r k ; a n te n n a e a n d t a r s i l i g h t ; h e a d e x c a v a t e d , s h i n y , smoo t h , h a i r l ess , w i t h o u t sc u l p t u r e ; occ i p u t h a i r y ; p ron o t u m crossed by a s t ro n g

1 328

Mem . A m er . E n t .

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9 1 0 - C h e l a of A gona topoiJes I'u J)eI' n . s p .

( type)

t r a n s v e rs a l i m p ress i o n , s h i n y , smoot h , h a i r l es s , w i t h o u t scu l p t u re ; s c u t u m sh i n y , w i t h some l o n g i t u d i n a l k ee l s ; m e t a n o t u m smoo t h , n o t t r a n sve rse l y s t r i a t e ; m e t a t h o r a x + p ro p o d e u m s h i n y , smoo t h , w i t h o u t s c u l p t u re ; p l e u r a a n d pos t e r i o r s u r'face of p ropode u m t r a n sverse I y s t r i a t e ; meso-me tap l e u ra I s u t u re obso l e te ; fore t a r s a l segmen ts i n fo l l o w i n g p ropo r t i o n s : 1 4 : 2 : 4 : 1 1 : 1 7 ; e n l a r g e d c l a w ( F i g . 9 1 0 ) w i t h s U b a p i c a l too t h a n d 5 l a me l l ae ; s e g m e n t 5 of f ro n t t a r s u s ( F i g . 9 1 0 ) w i t h two rows of 1 3 l a me l l ae ; a p e x w i t h a g ro u p of 1 3 l a me l l a e ; m a x i I l a r y p a l p i w i t h 5 segme n t s ; l a b i a l p a l p i " i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MA L E : u n k n o w n L O C U S TYP I C U S : M i l nerton ( Sou t h A f r i c a ) T Y P I CAL MA T E R I AL : h o l o t y p e F ! i n B lv\ . HOSTS : u n k n o w n , D I STR I I3 U T I O N : E t h i o p i a n reg i o n : S O U T H A F R I C A : ''' d l n erton , 13 M !

5 . A gon a toi)oides ceres n . s p .

F E M A L E : a p terou s ; l e n g t h 3 , 75 mrn ; red d i s h - te s t aceo u s , w i t h p e t i o l e b l ac k ; head exca v a t e d , d u " , g r a n u l a te d ; p ro no t u m d u " , g ra n u l a t e d , crossed b y a s t r o n g t r a n s v e rs a l i m p re ss i o n ; scu t u m d u l l , g ra n u l a t e d ; m e t a n o t u m t r a n s v e rse l y s t r i a t e , h o l l o w b e h i n d t h e sc u t e l l u m , w i t h s i d e s p r o t r u d i n g ; p ro t r u s i on s ro u n d e d ; rne t a t ho r a x + p ropode u m d u l l , g ra n u l a t e d ; p o s t e r i o r s u rface of p ropodeurn a n d p l eu r a t r a n s v erse l y s t r i a t e ; a n te r i o r s u rface of propo d e u m w i t h n u me ro u s weak

l o n g i t u d i n a l k ee l s ;

meso-me t a p l e u r a l s u t u r e

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1 984

d i s t i n c t , com p l e t e ; fore t a r s a l s e g m e n t 1 shorter t h a n segmen t 4 ( 1 3 : 1 6 ) ; e n l a r g e d c l a w ( F i g . 9 1 1 ) w i t h a pex t a p er i n g , not b road , w i t h s u b a p i c a l too t h a n d 6- 1 2 l a m e l l ae ; segment 5 o f front t a r s u s ( F i g . 9 1 1 ) w i t h t w o r o w s of 1 3-23 l ame l l ae ; a p e x w i t h a g r o u p o f 1 2- 1 8 l a me l l ae ; m a x i l l a r y p a l p i w i t h 5 segmen t s ; l ab i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : Ceres ( So u t h A fr i c a ) TYP I CA L MAT E R I AL : ho l o t y p e F ! a n d 1 p a ra t y pe F ! i n B M . HOST S : u n k n o w n D I ST R I BU T I O N : E t h i op i an reg i o n : SO U T H A F R I CA : Ceres ( C a p e P rov i nce ) , BM ! Port S t . Johns ( Po n do l a n d ) , BM !

O R I E NT A L S P E C I E S OF AGONA TOPOIDES

F o u r s pec i es a re k n ow n .

K E Y TO T H E S P E C I E S

F E MA L E S

+

D i s c o f m e t a t h or a x

2

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3

p r op od eu m w i t h s t r o n g

t ra n s v e r s a l

kee l s

•

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1 . benga lensis n . s p . D i s c o f m e t a t h o ra x + p r op od eu m w i t h o u t t ra n s v e r s a l k e e l s ; p o s t e r i o r k e e l s p r e s en t on l y on p o s t e r i o r s u r f a c e o f p r op od eu m ; o c c a s i o n a l l y s h o r t k e e l s a r e v i s i b l e o n t h e g r o u n d o f t h e m e d i a n f u r r ow 2 V e r t ex o f h ea d , me t a n o t u m a n d me t a t h o ra x + p r o p o d e u m v e ry s h i n y , smo o t h , w i t h o u t s c u l p t u r e . . . . . . . . . . . . . 2 . born e a n u s n . s p . V e r t ex o f h e a d , me t a n o t u m a n d me t a t h o ra x + p r o p o d e u m d u l I , g r a n u I a t ed 3 A n t en n a e s h o r t , mo r e d i s t a l l y t h i c k en ed ; a n t en n a l s e g men t 9 •

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s l i g h t l y l on g e r t h a n b r o a d . . 3 . insularis n . s p . A n t en n a e l on g , l e s s d i s t a l l y t h i c k e n ed ; a n t en n a l a p p r o x i ma t e l y t h r e e t i me s a s l on g a s b r o a d 4 . pyrill a e ( Ma n i ) •

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s eg men t •

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MALES

H ea d f u l l y y e l l ow ;

a n t en n a l

s e g men t

3 mo r e

t han

f o u r t i me s a s

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G o n a topod i n a e

1 33 1

l on g a s b r o a d ; d o r sa l p r o c e s s o f g on o f o r c ep s w i t h a p ex s h a rp ( F i g . 9 1 4 ) . . . . . . . . . . . . . . . . . . 2 . born e a n u s n . s p . H e a d b r own ; a n t en n a l s eg m e n t 3 l e s s t h a n t h r e e t i me s a s l on g a s b r o a d ; d o r s a l p r o c e s s o f g o n o f o r c e p s w i t h a p ex r o u n d ed ( F i g . 9 1 7 ) . . . . . . . . . . . . . . . . . . 4 . pyri l l a e ( Ma n i )

T he m a l es of A .

F i g . 91 2

-

insularis a n d bengalen sis a re u n k n ow n .

C he l a of Agona topoides benga lensis n .

sp.

( type)

1 332

Mem . Amer .

Ent.

I ns! . ,

n o . 37 ,

1 984

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C he i a of !l gona topoides b orn e a n u s n . s p .

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Gon a topod i n ae

1 33 3

1 . Agona topoides benga len sis n . sp .

F E MA L E :

a p tero u s ;

l e n g t h 3 , 50-5 , 3 1

mm ;

y e l l o w - t e s t a ceou s ,

w i t h t horax

a n d p ropode u m red d i s h- b row n ; p e t i o l e b l ac k ; h e a d s h i n y , exc a v a ted , h a i r l es s , w i t h o u t scu l p t u re ; prono t u m crossed b y a s t rong t r a n svers a l i mp res s i on , s h i n y , smoot h , h a i r l es s , w i t h o u t scu l p t u re , w i t h some l a tera l k ee l s ; s c u t u m s h i n y , smoo t h , w i t h some l on g i t u d i n a l kee l s ; sc u t e l l u m s h i n y , smoo t h , rec t a n g u l a r ; m e t a n o t u m t r a n s verse l y s t r i a t e , w i t h s i des p ro t r u d i n g ; p ro t r u s i on s ro u n ded ; m e t a t h o r a x + propodeum s h i n y , w i t h a s t ro n g m e d i a n f u rrow ; poster i o r s u rface a n d d i sc of m e t a t h o r a x + propodeum t r a n s verse l y s t r i a t e ; p l e u r a not t r a nsverse l y s t r i a t e ; meso m e t a p l e u r a l s u t u re d i s t i n c t ; fore t a r s a l segme n t s i n fo l l ow i n g p ropor t i on s : 1 6 : 4 : 5 : 20 : 3 3 ; e n l a rged c l a w ( F i g . 9 1 2 ) w i t h s u b a p i c a l too t h a n d 8 l a m e l l ae ; segmen t 5 of f ro n t t a r s u s ( F i g . 9 1 2 ) w i t h 1 row of 1 3 l a m e l l a e ; apex w i t h a g ro u p o f 2 0 l a m e l l ae ; m a x i l l ary p a l p i w i t h 5 segmen ts ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M AL E : u n k n o w n L O C U S T Y P I C U S : M i s h m i H i l l s ( A ss a m , I n d i a ) T Y P I CA L M A T E R I AL : h o l o t y p e F ! i n BM . HOST S : u n k n o w n D I ST R I B U T I O N : O r i e n t a l reg i on : I N D I A : M i s h m i H i l l s ( D e l a i V a l l ey , Assam ) , B M !

2 . Agona topoides born e a n u s n .

sp .

F E MA L E : a p tero u s ; l e n g t h 3 , 52 m m ; t e s t a ceou s - b row n ; a n te n n ae d i s t a l l y weak l y t h i c kened ; a n ten n a l segme n t s i n fo l l ow i n g p ropor t i on s : 1 2 : 7 : 2 2 : 1 2 : 1 1 : 9 : 7 : 7 : 7 : 1 1 ; h e a d very exca v a ted , s h i n y , smoo t h , w i t h o u t scu l p t u re ; occ i p i t a l c a r i n a i n v i s i b l e ; fron t a l l i n e com p l e te ; POL = 1 ; O L 3 ; OOL = 1 0 ; p ro n o t u m crossed b y a s t ro n g t ra n svers a l i mp ress i o n , w i t h o u t scu l p t u re , h a i r l es s ; scu t u m a n d s c u te l l u m s h i n y , smoot h , w i t ho u t scu l p t u re ; s i d es of scu t u m w i t h t wo l a t e r a l poi n t s ; m e t a n o t u m smoot h , s h i n y , w i t h o u t sc u l p t u r e , ho l l ow b e h i n d t h e s c u t e l l u m ; meso-me t a p l e u r a l s u t u re d i s t i n c t , com p l e t e ; m e t a t ho r a x + p ropodeum s h i n y , smoot h , w i t h o u t scu l p t u re , w i t h a s t r o n g med i a n f u r row ; pos t e r i o r s u rface o f p ropode u m a n d p l e u ra t r a n sverse l y s t r i a t e ; fore t a r s a l segmen t s i n fo l l ow i n g proport i o n s : 1 9 : 5 : 8 : 22 : 3 3 ; en l a rged c l a w ( F i g . 9 1 3 ) w i t h s u b a p i c a l too t h a n d 8 l a me l l ae ; segment 5 o f f ron t t a r s u s ( F i g . 9 1 3 ) w i t h t w o rows of 1 1 + 2 ( p ro x i m a l ) l am e l l a e ; a pex w i t h a g ro u p of n e a r l y 1 8 l ame l l a e ; m a x i l l a r y p a l p i w i t h 5 segmen t s ; l ab i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : f u l l y w i n g e d ; l e n g t h 1 , 87 m m ; head , a n t e n n a e a n d l eg s test aceou s ; t h o r a x , propode u m a n d a b domen b ro w n ; a n ten n a l segme n t s i n fo l l ow i n g proport i o n s : 6 : 6 : 1 2 : 8 : 8 : 8 : 8 : 8 : 7 : 1 0 ; a n t e n n a I segme n t 3 more

1 334

Mem . Amer . E n t .

I n s! . ,

no . 3 7 ,

1 984

F i g . 9 1 4 - M a l e gen i t a l i a of Agon a topoides born e a n u s n . s p .

( paratype)

t h a n f o u r t i mes a s l on g a s b ro a d ( 4 , 8 ) ; h e a d h a i ry , s h i n y , a l u t aceo u s ; fron t a l l i ne a b sen t ; frons w e a k l y g r a n u l ated ; occ i p i t a l ca r i n a a b s en t ; temp les very s h o r t , red uced ; POL = 6 ; O L = 2 , 5 ; OOL = 2 ; b re a d t h o f t h e oce l l i : 3 ; s c u t u m h a i r y , s h i n y , g r a n u l a t ed ; n o t a u l i ces com p l e t e , poster i or l y sep a r a ted ; m i n i m u m d i s t a n ce b e t w ee n t h e n o t a u l i ces s l i g h t l y s horter t h a n b rea d t h of t h e oce l l i ( 2 , 5 : 3 ) ; s c u te l l u m a n d m e t a n o t u m s h i n y , smoot h , w i t h o u t scu l p t u re ; p ropode u m ret i c u l a te r u gose , w i t ho u t l o n g i t u d i n a l kee l s on pos ter i or s u rface ; fore-w i n g h y a l i n e , w i t ho u t d a r k t ra n s v e r s a l b a n d s ; dors a l p rocess of gonoforceps ( F i g . 9 1 4 ) l o n g , b roa d , p o i n te d ; m a x i l l a r y p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 3 seg men t s ; t i b i a l spurs 1 , 1 , 2 . LOCUS TYP I C U S : G u no n g M u l u ( Sa r a w a k , M a l a y s i a ) T Y P I C A L M A T E R I A L : h o l o t y p e F ! a n d 1 p a ra t y pe M ! i n SM . HOSTS : u n k n o w n D I STR I BU T I O N : O r i e n t a l reg i o n : MALAYS I A : G u no n g M u l u ( Sa r a w a k ) , BM !

3 . Agona topoides in s u l a ris n . s p .

1 335

Go n a topod i n a e

Fig.

9 1 5 - C he l a of Agon a topoides insul aris n . sp .

( t y pe )

F E MA L E : a p terou s ; l e n g t h 3 , 2 m m ; t e s t a ceou 5 , w i t h p e t i o l e b l ack ; a n te n n a e b row n , w i t h seg m e n t s 1 -3 a n d 1 0 t e s t aceo u s ; a n t en n ae d i s t a I I y t h i ck e n e d ; a n te n n a l s e g me n t s i n fo l l ow i n g p roport i on s : 1 0 : 7 : 1 8 : 1 0 : 9 :

1 336

Mem . A m e r . E n t .

Fig. 916

-

I n st . ,

no. 37,

1 984

C he l a of Agon a topoides pyrillae ( M a n i ) ( t y p e )

8 : 8 : 6 : 5 , 5 : 1 0 ; a n te n n a l segmen t 9 s l i g h t l y l on ge r t h a n b ro a d ( 5 , 5 : 4 ) ; head d u l l , very e x c a v a t e d , g r a n u l a ted ; fron t a l l i n e com p l e t e ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L � 1 ; O L � 3 ; OOL � 1 2 ; p ron o t u m c rossed b y a s t ro n g t r a n svers a l i m p ress i o n , s h i n y , weak l y g r a n u l a ted ; sc u t u m s h i n y , w i t hou t sc u l p t u r e ; m e t a n o t u m h o l l ow b e h i n d t h e sc u t e l l u m , w e a k l y t r a n s verse l y s t r i a t e ; m e t a n o t u m w i t h s i des p ro t ru d i n g ; p ro t r u s i o n s p o i n ted ; fore t a rs a l segmen t s i n fo l l ow i n g p ropo r t i on s : 1 6 : 4 : 6 : 1 9 : 30 ; e n l a r g e d c l a w ( F i g . 9 1 5 ) w i t h s u b a p i c a l too t h a n d 8 l a me l l a e ; segmen t 5 of fron t t a r s u s ( F i g . 9 1 5 ) w i t h t w o rows of 3 ( p rox i m a l ) + 1 2 l a m e l l ae ; a p e x w i t h a g ro u p of 1 2 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 5 segme n t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n LOCUS T Y P I C U S : U l h i t i y a O y a ( S r i L a n k a ) T Y P I CA L MAT E R I A L : h o l ot y pe F ! i n WA . HOST S : u n k n o w n D I ST R I B U T I ON : O r i en t a l reg i o n : S R I LANKA : U l h i t i y a O y a ( 1 5 m i . N N E of M a h i y a n g a n a , B a d . D i s t . ) , WA !

4 . Agon a topoides pyril l a e ( M a n i )

n .

com b .

Gon a topod i n a e

1 337

F i g . 9 1 7 - M a l e g en i t a l i a of Agon a topoides pyrill a e ( M a n i ) ( a l l o t y p e )

Ps e u dogon a top u s pyril l a e M a n i 1 942 : 1 6 1 . Pseu dogon a top u s pyrillae M a n i : K u r i a n 1 954a : 2 5 8 .

F E MA L E : a p tero u s ; l e n g t h 4 , 2 m m ; t e s t aceo u s-red d i s h , w i t h a b domen b l ac k ; a n te n n a e d i s t a l l y t h i ck e n ed ; a n t e n n a I segme n t s i n fo l l o w i n g p ro p or t i on s : 1 1 : 8 : 30 : 20 : 1 5 : 1 3 : 1 0 : 1 0 : 9 : 1 1 ; a n te n n a l segmen t 3 a p p ro x i m a te l y t h ree t i mes as l on g as b ro a d ( 9 : 3 ) ; h e a d e x c a v a ted , d u l l , g r a n u l a te d ; f ro n t a l l i n e comp l e t e ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 1 ; O L = 3 ; O O L = 1 4 ; p ro n o t u m crossed b y a s t ro n g t r a n sversa l i mp res s i on , s h i n y , w e a k l y g r a n u l a t e d ; sc u t u m s h i n y , w e a k l y g ra n u l a t e d ; s c u te l l u m s h i n y , i n d i s t i nc t ; m e t a n o t u m ho l l o w b eh i n d t h e sc u te l l u m , w i t h s i des p ro t r u d i n g ; p ro t r u s i o n s po i n ted ; m e t a t horax + p ropodeum d u l l , g r a n u l a ted , w i t h a s t ro n g med i a n f u r ro w , w i t h s t ro n g t r a n sversa l s t r i ae on pos t e r i o r s u rface ; d i sc ru gose , n o t t r a n sv erse I y s t r i a t e ; meso-

1 338

Mem . A m e r . E n t .

I n st . ,

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1 984

m e t a p l e u r a l s u t u re d i s t i n c t , comp l e t e ; mesop l eu r a not t r a n s verse l y s t r i a t e ; me t a p l e u r a t r a n s v e rs e l y s t r i a t e ; fore t a r s a l segme n t s i n fo l l ow i n g p roport i on s : 2 2 : 5 : 7 : 25 : 40 ; en l a rged c l a w ( F i g . 9 1 6 ) w i t h s u b a p i c a l tooth a n d 1 1 l a me l l ae ; seg men t 5 of fron t t a rs u s ( F i g . 9 1 6 ) w i t h 1 row of 1 5 l a m e l l a e ; a p e x w i t h a g ro u p of a p prox i m a te l y 9 l a m e l l ae ; seg men t 3 of front t a r s u s p ro d u c e d i n to a hook ; m a x i l l a r y p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : f u l l y w i n ged ; l e n g t h 2 , 4 m m ; b row n , w i t h a n te n n a e , m a n d i b l es a n d l e g s t e s t aceou s ; a n t e n n ae n o t d i s t a l l y t h i c k e n ed ; a n te n n a l s e g m e n t s i n fo l l ow i n g p roport i on s : 7 : 6 : 8 : 7 : 7 : 6 : 6 : 6 : 6 : 9 ; a n t en n a I s e g m e n t 3 l es s t h a n t h re e t i me s as l o n g as b ro a d ( 8 : 3 ) ; h e a d d u l l , gra n u l ated , w i t hou t fron t a l l i n e ; occ i p u t conc a v e ; occ i p i t a l c a r i n a i n v i s i b l e ; temp l e s d i s t i nc t ; P O L = 8 ; O L = 4 ; O O L = 4 ; sc u t u m s h i n y , g ra n u l a ted ; not a u l i ces com p l e t e , poste r i o r l y sep a r a te d ; m i n i m u m d i s t a nce b e t w een t h e notau l i ces s horter t h a n b re a d t h of the ocel l i ( 1 , 5 : 3 ) ; s c u te l l u m a n d m e t a n o t u m s h i n y , smoo t h , p u n c t a te , w i t h o u t s c u l p t u re a mong p u n c t u res ; propode u m d u l l , ret i c u l a t e r u g o s e , w i t h o u t t r a n s versa l o r l on g i t u d i n a l k ee l s ; dors a l s u rface w i t h a m ed i a n n a rrow f u r ro w ; fore-w i n g h y a l i ne , w i t h o u t d a r k t r a n s v e rs a l b a n d s ; r a d i a l c e l l open ; d i s t a l p a r t of r ad i a l v e i n l on g e r t h a n p rox i m a l p a r t ( 2 3 : 1 5 ) ; d o r s a I p rocess of gonoforceps ( F i g . 9 1 7 ) l o n g , d i s tB l l y b roadened ; m a x i l l a r y p a l p i w i t h 5 segme n t s ; l a b i a l p a l p i w i t h 3 s e g me n ts ; t i b i a l s p u r s 1 , 1 , 2 . LOCUS TYP I CU S : K a r n a l ( I n d i a ) T Y P I CA L M A T E R I AL : h o l o t y pe F ! a n d a l l o t y p e M ! i n N O . HOST S : i n I n d i a Pyril l a s p . ( M a n i 1 942 ) . D I ST R I BU T I O N : O r i e n t a l reg i on : I N D I A : K a rn a l ( P u n j a b ) , N O ! D a u ra l a F a rm ( Meeru t ) , N O !

N E A RC T I C S P E C I E S O F AGONA TOPOIDES

T w o s p ec i es a re k n ow n .

KEY T O THE SPEC I ES

F E MA L E S

S e g m e n t 4 o f f r o n t t a r s u s s h o r t e r t h a n s eg me n t 1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 . synchromus R . C . L . P e r k i n s S e gm e n t 4 o f f r o n t t a r s u s l on g e r t h a n s eg m e n t 1 2 . c a lidus n . s p . •

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Go n a topod i n ae

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MALES

U n k no w n

1 . Agona topoides synchromus R . C . L .

Perk i n s

A gona topoides synchromus R . C . L . Perk i n s 1 907 :

33 .

F E MAL E : a p terou s ; l e n g t h 4 , 0 m m ; redd i s h - t e s t aceou s , w i t h pet i o l e b l ac k ; body n e a r l y h a i r l es s ; h e a d exc a v a ted ; p ron o t u m crossed b y a s t ron g t r a n s v e rs a l i m p ress i on ; p o s t e r i or s u rface of p ropodeum t ra n sverse I y s t r i a t e ; segme n t 5 of f ro n t t a r s u s l on g e r t h a n s e g m e n t 4 ( 76 : 50 ) ; se g m e n t 4 of fron t t a r s u s shorter t h a n segmen t 1 ( 1 04 : 1 40 ) ; en l a rged c l a w w i t h s u b a p i c a l too t h a n d 5 l ame l l a e ; segmen t 5 of fron t t a r s u s w i t h two row s o f 8 + 9 l a m e l l ae ; apex w i t h a g ro u p o f 2 l a me l l a e ; m a x i I l a ry p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MAL E : u n k n o w n L O C U S T Y P I C U S : Nog a l e s ( A r i z . , U . S . A . ) T Y P I CAL MAT E R I A L : l ec t o t y pe F ! a n d 2 p a r a l ec t o t y pes F F ! ( d e s i g n a ted b y O l m i 1 98 2 ) i n B . HOSTS : i n U . S . A . Bosta era ( � Cochise ) a p a c h e a n u s K i rk . ( Delph a cida e ) ( R . C . L . Perk i n s 1 907 ) ; Bosta era n a s u ta B a l l ( T ho m p son 1 95 4 ) D I ST R I BU T I O N : N e a rc t i c reg i on : U . S . A . : Nog a l e s ( A r i z . ) , 8 ! •

2 . Agona topoides ca lidus

n .

sp .

F E MAL E : a p terou s ; l e n g t h 3 , 06 m m ; tes t aceo u s , w i t h pe t i o l e b l ac k ; a n te n n ae b ro w n , w i t h seg m e n t s 1 -3 a n d p a r t of 4 t e s t aceo u s ; segmen t 1 0 a n d p a r t of 9 w h i te ; a n t e n n a e d i s t a l l y t h i c kened ; a n t en n a l s e g me n t s i n fo l l ow i n g p ropor t i o n s : 8 : 5 : 1 5 : 8 : 8 : 6 : 6 : 6 : 6 : 9 ; head exc a v a ted , s h i n y , g ra n u l a ted ; fron t a l l i n e com p l e t e ; occ i p i t a l c a r i n a i n v i s i b l e ; POL � 1 ; OL � 2; OOL � 9; p ro n o t u m crossed by a s t ro n g t r a n svers a l i m p res s i on , s h i n y , smoo t h , w i t ho u t scu l p t u re ; scu t u m d u l l , g ra n u l a t e d , w i t h o u t l a tera l po i n ted p ro t r u s i on s ; sc u te l l u m i n v i s i b l e ; m e t a n o t u m n o t f l a t , t r a n s verse l y w e a k l y s t r i a te , w i t h s i des p ro t ru d i n g ; p rot r u s i on s po i n te d ; meso-met a p l e u r a l s u t u r e obso l e t e ; m e t a t ho r a x + p ropodeum s h i n y , w i t h o u t sc u l p t u re among s t r i ae a n d on a n te r i o r s u r f ace o f p ropode u m ; p o s te r i or s u rface o f p ropodeum a n d p l e u r a t r a n sv erse I y s t r i a t e ; fore t a rs a l segmen t s i n fo l l ow i n g p roport i on s : 1 0 : 3 : 6 : 1 5 : 24 ; e n l a r ged c l aw ( F i g . 9 1 8 ) w i t h s u b a p i c a l too t h a n d 1 5 l a me l l a e ; seg men t 5 of f ron t

1 340

Mem . Amer . E n ! .

Fig.

I ns! . ,

no.

37 ,

1 984

9 1 8 - C he l a of Agona topoides c a lidus n . s p .

( type )

t a rs u s ( F i g . 9 1 8 ) w i t h t wo row s of n e a r l y 50 l ame l l ae e x t en d i n g co n t i n uo u s l y t o t h e apex ; m a x i I l a ry p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l s p u r s 1 , 0 , 1 .

Gona topod i n a e

F i g . 91 9

-

1 34 1

C he l a of Agon a topoides longichel a t u s ( F e n ton ) { t y p e )

MALE : u n k n o w n L O C U S T Y P I C U S : W a s h i n g ton ( D . C . , U . S . A . ) TYP I CA L MAT E R I AL : ho l o t y pe F ! i n WA . HOST S : u n k n o w n D I ST R I BU T I O N : N e a rc t i c reg i o n :

U.

S . A . : W a s h i n g ton

(D.

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N E O T R O P I C SP E C I E S OF A GONA TOPOIDES

O n e s p ec i es i s k n ow n .

A gon a top o id e s l ongich e l a t u s ( Fe n ton ) n . com b .

Dicondylus l o ngich e l a tu s F e n ton 1 927 : 4 .

F E MAL E : a p terou s ; l e n g t h 3 , 3 1 m m ; h e a d red d i sh - b row n , w i t h m a n d i b l es a n d c l y p e u s y e l l ow ; fron t p a r t of vertex l i g h t ; a n te n n ae ye l l ow ; p ro n o t u m redd i s h ; co l l a r w i t h b ro w n spo t s ; s c u t u m , s c u t e l l u m , m et a t horax + p ropodeum a n d pet i o l e b l ac k ; a b domen redd i s h- b row n ; l eg s y e l l o w redd i s h ; h e a d e x c a v a t e d , d u l l , g r a n u l a t e d , w i t h s h o r t t r a n s v e rs a l kee l s n e a r med i a n fron t a l l i ne ; p rono t u m crossed b y a s t ro n g t ra n s vers a l i m p re s s i on , s h i n y , smoo t h , w i t ho u t scu l p t u r e , w i t h some l a tera l kee l s ; scu t u m d u l l , w i t h l on g i t u d i n a l k e e l s ; s c u te l l u m s h i n y , smoot h ; met a t horax + propodeum d u l l , w i t h o u t scu l p t u re ; m e t a no t u m , p l e u r a and pos ter i or s u rface of p ropodeum t r a n s v erse l y s t r i a te ; meso-me t a p l eu r a l s u t u re d i s t i n c t ; fore t a rs a l seg m e n t s i n fo l l ow i n g p roport i on s : 1 4 : 2 : 5 : 1 5 : 2 5 ; e n l a r g e d c l a w ( F i g . 9 1 9 ) w i t h s u b a p i c a l too t h a n d 6 l a me l l a e ; segmen t 5 of f ro n t t a rs u s ( F i g . 9 1 9 ) w i t h 2 row s of 1 0 + 1 3 l a me l l ae ; a p e x w i t h a g ro u p of a t l e a s t 1 5 l ame l l ae ; m a x i l l a ry p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k no w n L O C U S T Y P I C U S : P e r n a m b uco ( B r a z i I ) TYP I CAL MAT E R I AL : ho l o t y p e F ! i n WA . HOSTS : u n k no w n D I S T R I B U T I O N : Neot rop i c reg i o n : BRAZ I L : Pern a m b uco ( Bo n i t o ) , WA !

8 . Ap terodryin u s R . C . L .

Perk i n s

Ap terodryin u s R . C . L . Perk i n s 1 907 : 1 4 . Dolichochelys O l m i a n d C u rrado 1 976a : 1 77 ( n . s y n . ) ; t y p e s p ec i es Dolichochelys longicornis ( K i effer 1 904 ) , o r i g . d es i g . Noga top u s Ponomarenko 1 979a : 357 ( n . s y n . ) ; t y p e s p ec i es Noga top u s gob iensis Ponom a renko 1 97 9 , o r i g . d e s i g .

T Y P E S P E C I E S : Apterodryin u s torvus R . C . L . Perk i n s 1 907 , monot y p i c . F E MA L E ( F i g s 926 , 927 ) : a p terou s ; m a x i I l a ry p a l p i w i t h 6 segmen t s ; labial

p a l p i w i t h 3 segme n t s ; e n l a rged c l a w ( F i g . 920 )

w i th subap ical

G o n a to pod i n a e

F i g . 920

-

1 343

C he l a of Ap terodryinus gob iensis ( Ponom a r e n ko ) ( t y p e )

too t h a n d l a m e l l ae ; p ro n o t u m ( F i g . 927 ) crossed b y a s t ro n g t r a n sv e r s a l i m p ress i o n ; t i b i a l s p u rs 1 , 0 , 1 . MALE : f u l l y w i n g e d ; m a x i I l a ry p a l p i w i t h 6 segme n t s ; l a b i a l p a l p i w i t h 3

1 344

Mem . Amer .

En! .

I ns! . ,

no . 37 ,

1 984

segme n t s ; tem p l es d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e or on l y v i s i b l e beh i n d t he oce l l i o r o n t h e s i des o f t h e pos t e r i or oce l l i , b u t never reac h i n g tem p l es ; occ i p u t concave ; t i b i a l s p u rs 1 , 1 , 2 . D I ST R I B U T I O N : wor l d w i de HOST S : A c a n a loniida e , Fl a tidae SPEC I ES : 1 7

PALAEA R C T I C S P E C I E S O F APTERODR YINUS

O n e spec i es i s k n o w n .

Ap terodryin u s gobiensis ( Ponom a ren k o ) n . com b .

Noga top u s gobiensis Ponom arenko 1 979a : 357 .

F E MAL E : a p t e ro u s ; l e n g t h 3 , 5 m m ; tes t aceou s , w i t h pe t i o l e b l ac k ; a n t e n n ae d i s t a l l y t h i c kened ; a n t e n n a l segme n t s i n fo l l ow i n g p ropor t i o n s : 7 : 4 : 1 1 : 7 : 6 : 6 : 5 : 5 : 6 : 7 ; head d u l l , exca v a ted , g ra n u l a t ed ; occ i p i t a l ca r i n a i n v i s i b l e ; fron t a l l i n e com p l e t e ; P O L � 1 ; OL � 2 ; OOL � 7 ; p ro n o t u m crossed by a s t ron g t r a n s versa l i m p ress i on , s h i n y , w i t h o u t scu l p t u re ; m e t a no t u m h o l l o w beh i n d t he scu t e l l u m ; meso-me t a p l e u r a l s u t u re d i s t i n c t , com p l e t e ; m e t a t horax + p ropodeum s h i n y , w i t hou t scu l p t u re , excep t for pos t e r i o r s u rface of p ropodeum a n d p l e u r a t r a n s v erse I y s t r i a t e ; fore t a rs a l segmen t s i n fo l l ow i n g p roport i on s : 1 4 : 3 : 4 : 1 6 : 2 3 ; en l arged c l a w ( F i g . 920 ) w i t h s u b a p i c a l t oo t h and 1 0 l a m e l l ae ; segme n t 5 of f ron t t a rs u s ( F i g . 920 ) w i t h 2 row s of 26 l ame l l ae ; a pe x w i t h a g ro u p of n e a r l y 1 1 l a me l l ae ; m ax i l l a r y p a l p i w i t h 6 segme n t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : h o l o t ype F ! i n L N . H OS T S : u n k no w n D I S T R I B U T I O N : P a l aearc t i c reg i o n : M O N GO L I A : S h a ra-K h u l s n y - B u l a g ( B a j a n - K ho n g o r a y m a g , C e n t r a l Mongo l i a ) , L N ! NO T E S : A . go b iensis i s v e r y s i m i l a r t o Agon a topoides felix n . s p . from C a n a r y I s l a n d s : the on l y d i fferences concern t he n u m ber of m a x i l l a r y p a l p a l segmen t s , t h e h e a d ( more e x c a v a t e d i n A . gobien sis ) a n d t he a n t e r i o r s u rf ace of met a t ho r a x + p ropodeum ( w i t ho u t scu l p t ure i n A . go b iens is , w i t h w e a k l o n g i t u d i n a l s t r i a e i n Ag. felix ) .

E T H I O P I AN S P E C I E S O F APTERODR YINUS

Gon a topod i n ae

1 345

T h ree s p ec i es a re k n o w n .

K E Y TO T H E S P E C I E S

F E MA L E S

M e s o - me t a p l e u ra l s u t u r e ob s o l e t e 1 . somerseti n . s p . Me s o -me t a p l eu ra l s u t u r e d i s t i n c t , c omp l e t e 2 A n t e r i o r s u r f a c e o f m e t a t h o ra x + p r op o d e u m l e s s i n c l i n ed ( F i g . 9 2 2 A ) ; me t a t h o ra x + p r op o d e u m a l mo s t f u l l y b l a c k ; e n l a rg ed . . . . . . . . . . . . c l a w w i t h a p ex v e r y b en t ( F i g . 9 2 3 ) . . . . . . . . . . • . . . 2 . richardsi n . s p . A n t e r i o r s u r f a c e o f me t a t h o ra x + p r op o d e u m m o r e i n c l i n ed ( F i g . 9 2 2 8 ) ; me t a t h o ra x + p r op o d e u m f u l l y f e r ru g i n ou s ; e n l a r g ed c l a w w i t h a p ex I e s s b en t ( F i g . 9 2 4 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 . rufu l u s n . s p . •

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MALES

Unknown

1 . A p terodryin u s somerseti n .

sp .

F E M AL E : a p tero u s ; l e n g t h 4 , 37 m m ; head b row n , w i t h front p a r t o f ver te x , c l y p e u s a n d m a n d i b l e s y e l l o w ; a n t e n n ae test aceo u s ; p r o n o t u m w i t h co l l a r b l ack ; d i sc b ro w n - re d d i s h ; scu t u m y e l l o w , w i t h a n te r i o r m a rg i n b l ac k ; sc u te l l u m a n d met a t ho r a x + p ropodeum b l ac k ; a b domen b ro w n b l ac k ; l eg s tes taceous , w i t h p a r t o f c o x a e a n d p a r t o f c l u b s o f femora b row n ; head exca v a ted , s h i n y , w i t ho u t scu l p t u re , except for occ i p u t a n d f r o n t p a rt o f v e r t e x gra n u l a t e d ; p ro n o t u m c rossed b y a s t ron g t ra n s v e rs a l i m p ress i o n , d u l l , weak l y g r a n u l a t e d ; sc u t u m d u l l , g r a n u l a t ed , l a tera l l y w i t h t w o po i n t ed p ro t r u s i o n s ; m e t a n o t u m t r a n s v erse l y s t r i a t e ; meso m e t a p l e u r a l s u t u re obso l e te ; m e t a t h o r a x + p ropodeum s h i n y , w i t h a n t e r i o r s u rface w e a k l y a l u t aceou s ; pos t er i or s u rface of p ropodeum t r a n s verse l y s t r i a t e ; p l e u r a g r a n u l a t e d , w i t h some t r a n s v e rsa l k ee l s ; fore t a r s a l s e g m e n t 1 as l o n g as seg m e n t 4 ; en l arged c l a w ( F i g . 9 2 1 ) w i t h s u b a p i c a l t oo t h a n d 9 l ame l l a e ; segmen t 5 o f f ro n t t a r s u s ( F i g . 92 1 ) w i t h t w o r o w s o f 3 ( p ro x i m a l ) + 1 6 l am e l l ae ; a p e x w i t h a g ro u p o f 1 4 l a me l l ae ; m a x i I l a r y p a l p i w i t h 6 s e g me n t s ; l a b i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 .

1 346

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Fig.

921

I nst . ,

n o . 37 ,

1 984

- C he l a of Apterodryin u s somerseti n . s p .

( type)

MALE : u n k n o w n LOCUS T Y P I C U S : Somerset E a s t ( So u t h A f r i ca ) T Y P I CA L M AT E R I A L : ho l o t y pe F ! i n B M . HOSTS : u n k no w n D I ST R I BU T I O N : E t h i o p i a n reg i o n : SOU T H A F R I CA : Somerset E a s t P rov i n ce ) , B M !

( Cape

2 . Ap terodryinus rich ardsi n . s p .

F E MAL E : a p te ro u s : l e n g t h 4 , 8 1 m m ; head , p rono t u m , scu t u m a n d s c u te l l u m redd i s h- d a r k ; a n t e n n ae t e s t a ceou s , w i t h segmen t 1 0 w h i te ; m e t a t horax + p ropodeu m b l ac k - b row n , excep t for m e t a n o t u m a n d a p e x of p ropodeum redd i s h ; p e t i o l e b l ac k ; a b domen and l eg s t e s t aceo u s ; head excav a ted , d u I I , g r a n u I a ted ; p ro n o t u m crossed b y a s t ro n g t r a n sversa I i m p ress i on , d u l l , g ra n u l a ted ; s c u t u m s h i n y , w i t h some l on g i t u d i n a l k ee l s ; m e t a n o t u m h o l l ow b eh i n d t h e s c u te l l u m , t r a n s v erse l y s t r i ate ; m e t a t horax + p ropod e u m ( F i g . 922 A ) d u l l , g r an u l a t e d ; a n t e r i o r s u rface w i t h s o m e w e a k l o n g i t u d i n a l kee l s ; p l e u r a a n d pos ter i or s u rf a ce of p ropodeum t r a n s verse l y s t r i a te ; meso-me t a p l e u ra l s u t u re d i s t i n c t , com p l e t e ; fore t a rsa l segmen t 1 shorter t h a n segmen t 4 ( 9 : 1 3 ) ; en l a rged c l aw ( F i g . 9 2 3 ) s t rong l y c u r ve d , w i t h s u b a p i ca l too t h , 4 l a me l l ae a n d , i n a d d i t i on , a tooth l oc a te d on t h e p rox i m a l t h i rd of t h e i n tern a l s i d e ; segme n t 5 of front t a r s u s ( F i g . 923) w i t h t w o rows of 32 l a m e l l ae ; a p e x w i t h a g r o u p of n e a r l y 40 l ame l l ae ; m a x i I l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : M o s s e l B a y ( So u t h A f r i c a ) T Y P I CA L M A T E R I A L : h o l o t y p e F ! i n B M . HOST S : u n k n o w n D I ST R I BU T I ON : E t h i o p i a n reg i on : SOUTH A F R I CA : Mos s e l B a y ( C a p e P rov i nce ) , B M !

Go n a t opod i n ae

1 347

A

F i g . 922 - Meso t hora x , m e t a t horax a n d p ropodeum ( l a t e ra l l y v i ewed ) o f fema l e s of Ap terodryinus richa rdsi n . sp . ( t y p e ) ( A ) a n d rufu l u s n . s p . ( type) ( 8 )

NOTE :

t he s p ec i es i s n amed i n honor of M r . O w a i n W .

3 . Ap terodryinus rufu l u s n .

R i c h a rds .

sp .

F E MAL E : a p terou s ; l e n g t h 3 , 7 5-4 , 8 1 m m ; ferr u g i nou s , w i t h p e t i o l e b l a ck ; head e x c a v a t e d , d u l l , g r a n u l a t ed ; p ron o t u m crossed by a s t ron g t r a n s ve rs a l i m p res s i o n ; co l l a r s h i n y , weak l y g r a n u l a ted ; d i sc d u l l , s t ro n g l y g r a n u l a ted ; scu t u m d u l l , g r a n u l a t e d ; m e t a n o t u m d u l l , t r a n s v erse l y s t r i a t e , h o l l o w b e h i n d t h e s c u t e l l u m ; m e t a t horax + p ropodeu m ( F i g . 9 2 2 B ) d u l l , gra n u l a ted ; a n t e r i o r s u rface of m e t a t horax + p ropode u m v e r y i nc I i ned ; pos t er i or s u rf ace of p ropodeum t ra n s versel y s t r i a t e ; p l eu r a n o t t r a n sverse l y s t r i a te ; meso-me t a p l e u r a l s u t u re d i s t i n c t , comp l e t e ; fore t a r s a l segme n t 1 s horter t h a n segmen t 4 ( 1 4 : 1 6 ) ; en l a rged c l a w ( F i g . 924 ) w i t h s u b a p i c a l too t h and 8 l a m e l l ae ; segmen t 5 of f ro n t t a r s u s ( F i g . 924 ) w i t h t w o row s of 2 4 l a m e l l a e ; a p e x w i t h a g ro u p of 2 0 l a m e l l ae ; m a x i I l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : P o r t SI . Jo h n s ( Sou t h Afr i c a ) TYP I CAL M A T E R I A L : HOSTS : u n k n o w n

holotype F !

and 1

p a ra t y pe F !

i n BM .

1 348

Mem . Ame r . E n t .

I nst . ,

no.

37 ,

1 984

F i g . 923 - C he l a o f A p terodryin u s richd rdsi n . s p .

D I S T R I BU T I O N : E t h i o p i a n reg i o n : ( Po n do l a n d ) , B M !

SOU T H A F R I CA :

( type)

Port St . Johns

O F, I E N T A L S P E C I E S O F A PTERODR YINUS

T wo s p ec i es are k n ow n .

K E Y TO T H E S P E C I E S

F E MA L E S

A n t e n n a e v e ry as

l on g ,

l on g a s s e gmen t

s l end e r , w i t h 2

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s egmen t .

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Fig.

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1 349

( t y pe )

1 350

Mem . Amer . E n ! .

I ns! . ,

no.

37 ,

1 984

1 . a s i a ticus n . s p . A n t e n n a e s h o r t , w i t h s eg men t 4 s l i g h t I y l on g e r t h a n s e g m e n t 2 . . . . . . . . . . . . . . . . . . . 2 . insul aris n . s p . ( s e e A u s t ra l i a n s p e c i e s ) .

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MALES

U n k no w n

1 . A p terodryin u s a siaticus n .

sp.

F E MAL E : a p t e ro u s ; l e n g t h 6 m m ; y e l l ow-ferr u g i nou s , w i t h pe t i o l e b l ac k ; a n t en n a l segmen t s i n fol l ow i n g p ropor t i o n s : 9 : 8 : 40 : 32 : 2 5 : 1 7 : 1 1 : 8 : 7 : 1 0 ; head e x c a v a te d , s h i n y , w i t ho u t sc u l p t u re , e x c e p t for occ i p u t a n d fron t p a r t o f v e r t e x g r a n u I a t e d ; p ro n o t u m crossed b y a s t ro n g t r a n sversa I i mp ress i o n , s h i n y , h a i r l es s , g ra n u l a ted ; scu t u m g ra n u l a ted , w i t h t w o l a tera l p o i n te d p ro t ru s i on s ; sc u te l l u m s h i n y , d i s t i nc t ; met a n o t u m t ra n s v e r se l y s t r i a t e , n o t h o l l ow beh i n d t h e s c u te l l u m ; m e t a t horax + p ropode u m s h i n y , w i t ho u t sc u l p t u re ; p l e u r a , poster i or a n d a n t e r ior s u rface of m e t a t ho r a x + p ropod e u m t r a n sverse I y s t r i a te ; meso-me t a p I e u r a I s u t u re d i s t i nc t , corc p l e te ; fore t a r s a l segmen t 1 shorter t h a n segmen t 4 ( 1 5 : 20 ) ; en l a rged c l a w ( F i g . 925 ) w i t h s u b a p i c a l too t h a n d 8 l a m e l l ae ; segme n t 5 o f fron t t a r s u s ( F i g . 925 ) w i t h 2 rows of 2 2 + 1 9 l a me l l ae ; a p e x w i t h a g ro u p o f a t l e a s t 3 0 l am e l l a e ; m a x i l l a r y p a l p i w i t h 6 segme n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M AL E ; u n k n ow n L O C U S T Y P I C U S : T o n d a n o ( S u l a w e s i , I n dones i a ) T Y P I CA L MAT E R I AL : h o l o t y p e F ! i n B M . H O ST S ; u n k n o w n D I ST R I BU T I ON : O r i e n t a l reg ion : I N DO N ES I A : T o n d a no ( Su l awes i ) , BM !

2 . Ap terodryinus ins ularis n . ( see A u s t r a l i a n reg i on )

sp .

N E A R C T I C S P E C I E S OF APTERODR YINUS

T h ree s p ec i es are k n ow n .

K E Y TO T H E S P E C I E S

Gon a topod i n ae

F i g . 925

-

C he l a o f Apterodryin u s a s i a ticus n .

1 35 1

sp .

( type)

1 35 2

Mem . A m e r .

Ent.

I nst . ,

n o . 37 ,

1 984

F E MA L E S

D i s c o f p r on o t u m p o s t e r i o r l y s t r o n g l y c omp r e s s ed i n t o a med i a n , a cu t e , ra i s ed ca r i n a . 1 . torvus R . C . L . P e rk i n s D i s c o f p r on o t u m r ou n d ed , n o t c omp r e s s ed i n t o a med i a n , a c u t e , . . . . . . .2 ra i s ed c a r i n a Me t a n o t u m n o t h o l l ow b e h i n d t h e s c u t e l l u m . . . . . . . . . 2 . secundus ( F en t on ) Me t a n o t u m h o l l ow b e h i n d t h e s c u t e l l u m . . . . . . . . . . . . . . . . • • 3 . c a liforn icus ( A s h m e a d ) .

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MALES

F i g . 926

-

F e m a l e of Apterodryin u s torvus R . C . L .

( A r i zo n a )

Perk i n s f r o m Nog a l es

Gon a topod i n ae

F i g . 927

-

1 353

F e m a l e of A p terodryin u s torvus R . C . L . Perk i n s from Nog a l es ( A r i zo n a )

U nknown

1 . A p terodryinus torvus R . C . L . Perk i n s

A p terodryin u s torvus R . C . L . Perk i n s 1 907 :

14.

F E MA L E ( F i g s 926 , 9 27 ) : a p te ro u s : l e n g t h 3 , 43-4 , 68 m m ; head b l ac k , w i t h m a n d i b l e s , c l y pe u s a n d m a l a r s p ace y e l l ow ; occ a s i on a l l y head f u l l y f e r r u g i n o u s ; a n ten n ae f u I I y redd i s h ; occ a s i o n a I I y a n te n n ae b ro w n , w i t h segmen t s 9- 1 0 o r 8- 1 0 w h i t i s h ; t ho r a x a n d p ropodeum b l a c k ; p ro n o t u m w i t h w e a k red d i s h n u a nces on t h e s i des ; occ a s i o n a l l y a l so on d i sc a n d co l l a r ; a b domen red d i s h o r b l ac k ; l eg s ferru g i nous o r y e l l ow , w i t h coxae , t roc h a n ters , c l u b s of femora a n d t i b i ae b ro w n i s h ; head e x c a v a t e d , d u l l , h a i ry , g r a n u l a t e d ; occ i p i t a l ca r i n a on l y v i s i b l e b e h i n d t he oce l l i ; p ro n o t u m g r a n u l a te d , crossed b y a s t ro n g t r a n s v e rs a l i m p ress i on , s h i n y , h a i r l es s , w i t h d i sc poster i or l y s t ron g l y com p ressed i n to a med i a n , a c u t e , r a i sed c a r i n a ; sc u t u m s h i n y , h a i r l es s , g r a n u l a ted ; m e t a n o t u m t r a n s v e r se l y s t r i a t e ; meta t h o r a x + p ropode u m s h i n y , w i t h a n te r i o r s u rf ace smoot h , w i t ho u t scu l p t u re ; p l eu ra a n d pos t e r i o r s u r f ace of p ropodeum

1 354

Mem . Amer .

Ent.

I nst . ,

no . 37 ,

1 984

F i g . 928 - C he l a of Apterodryin u5 torvu5 R . C . L . Perk i n s from Nog a l es ( A r i zon a )

Gon a topod i n ae

F i g . 929

-

1 355

F ore t i b i a of fem a l e of Ap terodryinus secundus

( F e n ton ) ( t y p e )

t ra n s v e r se l y s t r i a te ; meso-me t a p l eu r a l s u t u re d i s t i n c t ; mesop l e u ra a n d m e t a p l e u r a l oc a te d o n d i fferen t p l a nes ; fore t i b i a e n o t c u rved ; fore t a r s a l segmen t s i n fo l l o w i n g proport i o n s : 1 6 : 2 : 5 : 1 9 : 2 9 ; en l a rg e d c l a w ( F i g . 928 ) w i t h s u b a p i c a l too t h a n d 1 2 l am e l l ae ; segmen t 5 of fron t t a r s u s ( F i g . 928 ) w i t h t w o row s o f 27 l a m e l l a e ; a p e x w i t h a g ro u p of a t l ea s t 2 5 l a me l l ae ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : Nog a l e s ( A r i z . , U . S . A . ) TYP I C A L M A T E R I AL : l ec t o t y pe F ! a n d 2 p a r a l e c t o t y pes F F ! ( des i g n a t e d b y O l m i 1 982 ) i n B ; 1 p a r a l e c t o t y p e F ! i n BM . HOST S : i n U . S . A . Mistharnop h a n tia son ora n a (Fl a tidae) a n d Amphiscep a imm a c u l a ta (A c a n a lon iida e ) ( R . C . L . Perk i n s 1 90 7 ) . D I ST R I B U T I O N : Nearc t i c re g i o n : U . S . A . : No g a l e s ( A r i z . ) , B ! BM ! OL ! Rock C reek ( Gordo n v i l l e , G r a y so n Co . , Tex . ) , T E ! F a r m i n g d a l e ( N . Y . ) , TW !

2 . Ap terodryi n u s secundus ( F e n to n )

n . com b .

E u c a mptonyx secundus F e n t o n 1 927 : 5 .

F E MA L E : a p t e ro u s ; l e n g t h 3 , 7 5-5 , 40 m m ; fer r u g i n o u s - y e l l ow , w i t h pe t i o l e b l ac k ; occa s i o n a l l y a b do m e n b l ac k ; a n te n n a e y e l l ow , w i t h seg men t s 3-6 d a r k e n e d ; h e a d more or l ess e x c a v a t e d , s h i n y , g r a n u l a t e d ; occ i p i t a l c a r i n a i n comp l e t e ; p ron o t u m crossed b y a s t ro n g t r a n s v e rs a l i m p res s i o n , s h i n y , g ra n u l a t e d , w i t h d i sc rou n de d ; scu t u m d u l l , g r a n u l a t ed ; scu t e l l u m invisible;

meta thorax

+

p ro p o d e u m d u l l ,

g ra n u l a te d ;

metanotu m ,

p l eura ,

1 356

Mem . Amer . E n ! .

Fig.

I ns ! . ,

no.

37 ,

1 984

930 - C h e l a o f Apterodryin u s secundus ( F en ton ) ( t y p e )

G o n a topod i n a e

1 357

pos ter i o r and a n t e r i o r s u r f ace of m e t a t ho r a x + propodeum t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re d i s t i n c t ; meso p l e u r a a n d met a p l e u r a l o c a t e d o n t h e s a m e p l a n e ; fore t i b i ae ( F i g . 929 ) s t ron g l y c u r v e d ; fore t a rs a l segme n t s i n fo l l ow i n g proport i o n s : 20 : 4 : 1 0 : 3 3 : 50 ; e n l a rged c l a w ( F i g . 930 ) w i t h s u b a p i c a l too t h a n d 7- 1 3 l a m e l l a e ; segment 5 of f ro n t t a r s u s ( F i g . 930 ) w i t h t w o rows o f 36-47 l a m e l l ae ; a pex w i t h a g ro u p o f a t l ea s t 20-30 l ame l l ae ; m a x i l l a r y p a l p i w i t h 6 segme n t s ; l ab i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k n o w n L O C U S T Y P I C U S : Western O h i o ( U . S . A . ) T Y P I CA L MAT E R I A L : h o l o t y p e F ! a n d 1 p a r a t y p e F ! i n WA . HOST S : i n U . S . A . A c a n a lonia con ica ( Sa y ) ( A c a n a loniida e ) . D I S T R I B U T I O N : Nea rc t i c reg i on : U . S . A . : P resco t t ( A r i z . ) , WA ! P i nery C a n . ( C h i r . M ts . , A r i z . ) , OT ! P or t a l ( Coc h i se C o . , A r i z . ) , A M ! S a n t a A n a R e f u g e ( A l a mo , Tex . ) , WA ! L a w re n ce ( Do u g l a s C o . , K a n s . ) , W A ! Western O h i o , Wi, ! A rc h bo l d B i o i . S t a . ( L a k e P l ac i d , H i g h l a n d s C o . , F l a . ) , O L ! P l u m mers I s . ( M d . ) , WA ! P o u g h k eeps i e ( N . Y . ) , T W !

3 . Ap terodryin u s c a l iforn ic u s ( A shmea d )

n . com b .

Gona top u s c a liforn i c u s A s h m e a d 1 893 : 85 .

F E MALE : a p t e ro u s ; l e n g t h 3 , 00-3 , 75 mm ; head f u l l y test aceo u s ; occ a s i o n a l l y b row n , w i t h f ro n t p ar t of vertex a l o n g orb i t s , c l y p e u s a n d m a n d i b l e s tes t a ceou s ; a n t en n ae f u l l y testaceo u s ; occas i o n a l l y b row n , w i t h v e n t r a l face of segmen t 1 y e l l o w ; t h o r a x a n d propodeum f u l l y b ro w n or re d d i s h-test aceo u s ; occ a s i o n a l l y red d i sh-test aceou s , w i t h p ropodeum b row n ; a b domen b row n ; l eg s test aceo u s or b row n ; a n te n n a e d i s t a l l y t h i c k e ne d ; a n t en n a l segme n t s i n f o l l ow i n g p ropo r t i on s : 9 : 6 : 1 3 : 8 : 7 : 5 : 5 : 5 : 5 : 7 ; head sh i n y , exca v a ted , h a i r y , g r a n u l a ted ; p ron o t u m crossed b y a s t r o n g t r a n svers a l i m press i o n , ro u n ded , w i t h d i sc a l u t aceou s ; co l l a r smoo t h , w i t h o u t scu l p t u r e ; sc u te l l u m d i s t i n c t ; m e t a n o t u m ho l l o w b eh i n d t h e sc u te l l u m ; m e t a t h o r a x + p ropodeum g r a n u l a t e d ; p l e u ra a n d p os t e r i o r s u rface of propodeum t r a n s v e rse l y s t r i a te ; meso-me t a p l eu r a l s u t u re d i s t i n c t , com p l e t e ; segme n t 1 of fron t t a rs u s shorter t h a n seg men t 4 ( 2 1 : 27 ) ; en l a rged c l a w ( F i g . 9 3 1 ) w i t h s u b a p i c a l tooth a n d 7 - 1 1 l a m e l l ae ; segme n t 5 o f f ro n t t a r s u s ( F i g . 9 3 1 ) w i t h t w o rows o f 1 6- 1 9 l a me l l ae ; a pex w i t h a g ro u p of a t l e a s t 1 4- 1 8 l a me l l a e ; m a x i l l a r y p a l p i w i t h 6 segme n t s ; l ab i a l p a l p i w i t h 3 segmen ts ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u nknown L O C U S TYP I C U S : Los A n g e l e s ( C a l i f . , U . S . A . ) T Y P I CAL MAT E R I A L : ho l o t y p e F ! i n WA . HOST S : u n k n o w n D I S T R I B U T I O N : N e a rc t i c reg i o n : U . S . A . : L o s A n g e l es ( C a l i f . ) , WA ! M a r s h

1 358

Mem . Amer . E n ! .

F i g . 931

O r . Sp r .

-

I n s! . ,

no . 37 ,

C he l a of Ap terodryin u s c a l ifornicus

( Ca l i f . ) , C A !

P rescott

(Ari z . ) , WA!

1 984

( A s hmead ) ( t y p e )

OL !

N E O T R O P I C S P E C I ES OF APTERODR YINUS

Gon a topod i n ae

1 359

Seven s p ec i es are k n ow n .

K E Y TO T H E S P E C I E S

F E MA L E S

2

3

4

5

6

S eg m e n t 5 o f f r o n t t a r s u s w i t h d i s t a l p a r t w e a k I y b e n t ( F i g s 932 , 933 ) • • • • • • • • • • . • . . . . • • • • • • . • • . . • 2 S e g m e n t 5 o f f r o n t t a r s u s w i t h d i s t a l p a r t s t r on g l y b en t ( F i g s 934 , 9 3 5 , 936 , 940 , 9 4 1 ) . . . . . . . . . . . . . . • . . . . . . . . . . . . . . . . . . . . . . 3 Me s o -me t a p l eu r a l s u t u r e o b s o l e t e ; b od y mo s t l y y e l l o w - f e r ru g i n o u s . . . . . . . . . . . . . . . . . 1 . citri n u s n . s p . Me s o -me t a p l e u ra l s u t u r e d i s t i n c t , c omp l e t e ; b o d y mo s t l y b l a ck • • . . • • . . . • • • • • • • • . . . • • . • 2 . arn a udi n . s p . Me s o -me t a p l eu r a l s u t u r e o b s o l e t e . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6 . longicornis ( K i e f f e r ) Me s o -me t a p l eu r a l s u t u r e d i s t i n c t , c omp l e t e • . . • • . . • • • • • • . • • • . • 4 Me t a t h o r a x + p r op od e u m w i t h a n t e r i o r s u r f a c e f u l l y s m o o t h , w i t h o u t s c u l p t u r e ; s eg m e n t 4 o f f r o n t t a r s u s s l i g h t I y s h o r t e r t h a n s eg m e n t 1 . . . . . . . . . . . . . . . 7 . bellottii n . s p . Me t a t h o ra x + p r o p o d e u m w i t h a n t e r i o r s u r f a c e a t I e a s t p a r t I y g ra n u l a t ed o r t ra n s v e r s e l y s t r i a t e ; s egmen t 4 o f f r o n t t a r s u s l on g e r t h a n s e g men t 1 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .5 B o d y mo s t l y b l a ck . . . . . . . . . . . . 5 . t ij u c a n u s ( A r l e ) B od y f u l l y o r m o s t l y t e s t a c e o u s o r r ed d i s h ; o c c a s i on a l l y f e r rug i n ou s • • • • . • • • • • • • • • • • • • • • • • • • . • • . . • . • • . • • . . • • . • • • • • • • • • 6 S e g m e n t 5 o f f r on t t a r s u s w i t h t w o r ow s o f l a me l l a e ( F i g . 9 3 4 ) ; b od y d u l l . . . . . . . . . . . . . . 3 . tes taceus ( C a m e r o n ) S eg m e n t 5 o f f r o n t t a r s u s w i t h 1 r ow o f l a me l l a e ( F i g . 9 3 5 ) b od y sh i n y • • • . • • • . • . . • • . • • . . • 4 . menkei n . s p .

MALES

O n l y t he m a l e o f A.

tij u c a n u s is k n ow n .

1 . Ap terodryinus citrin u s n .

sp.

F E MALE : a p t e r o u s ; l e n g t h 3 , 37-4 , 37 m m ; y e l l ow-fer r u g i no u s , w i t h p e t i o l e b l ac k ; a n t e n n ae l o n g , s l e n d e r ; h e a d exc a v a te d , s h i n y , g r a n u l a t e d ; p ro n o t u m d u l l , crossed b y a s t ro n g t r a n s v e r s a l i m p re s s i on , g r a n u l a t ed ;

1 360

Mem . Amer .

En!.

I n s! . ,

no. 37,

1 984

I

F i g . 932 - C he l a of Ap terodryin u s citI'in u s n . s p .

( type )

s c u t u m d u l l , g ra n u l a ted , l a ter a l l y w i t h t w o po i n t ed p ro t r u s i on s ; sc u t e l l u m i n v i s i b l e ; met a n o t u m not h o l l o w b e h i n d t h e sc u t e l l u m , t r a n s v erse l y s t r i a t e ; met a t ho r a x + p ro p o d e u m d u l l , g r a n u l a t e d ; p l e u ra , pos ter i or

Gon a topod i n ae

1 36 1

I

,

F i g . 933 - C he l a of A p terodryin u s a rn a udi n .

sp .

( type)

s u rf a ce a n d p a r t o f a n t e r i o r s u rface o f me t a t ho r a x + p ropodeum t r a n s v e r se l y s t r i a t e ; meso-me t a p l e u ra l s u t u re o b so l e te ; fore t a rs a l se g m e n t 1 shorter than s u b a p i ca l too t h w i t h 2 rows of l a me l l ae ; m a x i I

segmen t 4 ( 1 5 : 25 ) ; en l a rged c l a w ( F i g . 932 ) w i t h a n d 6-7 l am e l l ae ; segmen t 5 o f f ro n t t a r s u s ( F i g . 932 ) 1 2- 1 9 l am e l l ae ; a p e x w i t h a g ro u p of a t l ea s t 20-30 l a ry p a l p i w i t h 6 s e g men t s ; l a b i a l p a l p i w i t h 3 s e g me n t s ;

1 362

Mem . A m e r . E n ! .

I ns ! . ,

no.

37 ,

1 984

t i b i a l sp urs 1 , 0 , 1 . M A L E : u n k no w n LOCUS T Y P I C U S : S. Jose ( Co s t a R i c a ) T Y P I CAL MAT E R I AL : h o l o t y p e F ! i n O L ; I p a r a t y p e F ! i n W A . HOST S : u n k no w n D I ST R I B U T I ON : Neo t rop i c reg i o n : COSTA R I CA : S . Jos e , O L ! C U B A : S . Rangel , WA!

2 . Ap terodryin u s arn a udi n . s p .

F E MA L E : a p te ro u s ; l e n g t h 3 , 75-4 , 37 m m ; head b l ack , w i t h c l y p e u s , ma l a r s p ace a n d m a n d i b l es y e l l ow ; a n t e n n a e b ro w n , w i t h s e g m e n t s 1 -2 a n d 6- 1 0 t e s taceo u s ; t ho r a x a n d p ropodeum b l ack ; a b d omen b l ack-b row n ; l eg s test aceo u s , w i t h coxae b l ack i s h ; h e a d exc a v a t e d , d u l l , g r a n u l a t e d ; p rono t u m d u l l , g r a n u l a ted , crossed b y a s t ro n g t r a n s v e rs a l i m p res s i o n ; s c u t u m s h i n y , w i t h some l o n g i t u d i n a l k ee l s ; scu te l l u m i n v i s i b l e ; met a t ho r a x + p ropodeum d u l l , g ra n u l a t e d , w i t h n u merous l on g i t u d i n a l k ee l s o n pos t e r i o r s u rface a n d p l e u r a ; meso-me t a p l e u r a l s u t u r e d i s t i nc t , comp l e t e ; seg m e n t 1 o f f ro n t t a r s u s s horter t h a n s e g me n t 4 ( 1 0 : 1 5 ) ; e n l a r g e d c l a w ( F i g . 9 3 3 ) w i t h s u b a p i c a l too t h a n d 8 l ame l l a e ; seg m e n t 5 of f ro n t t a r s u s ( F i g . 933 ) w i t h 2 row s of 1 4 + 1 4 l a m e l l ae ; apex w i t h a g ro u p of at l ea s t 40 l ame l l ae ; m a x i l l a ry p a l p i w i t h 6 s e g me n t s ; l ab i a l p a l p i w i t h 3 s e g m e n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : Academy B a y ( Sa n t a C r u z I s l a n d , Ga l a p agos I s l a n d s ) TYP I CAL MAT E R I AL : H OST S : u n k n o w n

holotype F!

i n CA ;

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p aratype F !

i n OL .

D I S T R I B U T I O N : Neotro p i c reg i o n : GALAPAGOS I SLANDS : Academy B a y C ru z I s l a n d ) , CA ! OL ! N O T E S : t h e s p ec i e s i s n a m e d i n honor of M r . P a u l A r n a u d j u n i o r .

3 . A p terodryinus tes ta ce u s ( C ameron )

(Santa

n . com b .

Gona top u s testaceus C a meron 1 888 : 440 . A go n a topoides jujuyen sis O g l ob l i n 1 95 1 : 488 ( n o s d y n . ) ; locus typ ic u s Cerro Pera l e s ( Argen t i n a ) .

F E MA L E : a p terou s ; l e n g t h 4 , 62-7 , 00 m m ; ferr u g i nou s , w i t h p e t i o l e b l ack ; head exc a v a t e d , d u l l , g r a n u l a t e d ; p ronot u m crossed b y a s t ro n g t r a n s v e r s a l i mp re s s i o n , s h i n y , h a i r l es s , g r a n u l a t e d ; s c u t u m d u l l , g ra n u l a te d , l a t e ra l l y w i t h t w o p o i n ted p ro t r u s i on s ; scu te l l u m w e a k l y v i s i b l e , d u l l ; m e t a n o t u m t r a n sv erse l y s t r i a t e , n o t h o l l ow b eh i n d t h e

Gon a topod i n a e

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scu t e l l u m ; m e t a t horax + propodeu m s h i n y , g r a n u l a t e d ; p l e u r a , pos ter i or s u rface of p ropode u m a n d p a r t of a n t e r i or s u rface t r a n s v erse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re d i s t i nc t , comp l e t e ; fore t a rs a l seg men t 1 shorter t h a n segment 4 ( 24 : 3 1 ) ; en l a rged c l a w ( F i g . 934 ) w i t h s u b a p i c a l t oo t h a n d 6-1 1 l a m e l l ae ; segmen t 5 of f ro n t t a r s u s ( F i g . 934 ) w i t h 2 rows of 1 9-37 l a m e l l ae ; apex w i t h a group of 20-40 l a m e l l ae ; m a x i I l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k n o w n L O C U S T Y P I C US : Vo l c a n de C h i r i q u i ( P a n a m a )

1 364

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( type)

TYP I CAL M A T E R I AL : ho l o t y p e F ! of G . te s t a c e u s i n B M ; h o l o t y p e F ! o f A . juju ye n sis i n T U . HOST S : u n k no w n D I ST R I B U T I O N : Neo t rop i c reg i o n : PANAMA : Vo l c a n de C h i r i q u i , B M ! G a m boa ( C a n a l Zone ) , L E ! Taboga I s l a n d , WA ! B u g a b a , B M ! Per a l e s ( J u j u y ) , T U ! B u enos A i res , O L l

4 . Ap terodl'yin us menkei n .

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sp .

F E MA L E : a p te ro u s ; l e n g t h 4 , 25 m m ; t e s t aceou s-red d i s h , w i t h some i r reg u l a r d a r k s p o t s ; p e t i o l e b l ac k ; s c u t u m y e l l ow ; l eg s t e s t aceou s , w i t h

Gon a topod i n a e

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1 365

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coxae , c l u b s o f femora a n d fore t i b i ae b row n ; a n t e n n a e d i s t a l l y t h i c k e n e d ; a n ten n a l segme n t s i n fo l l ow i n g p roport i o n s : 9 : 6 : 2 3 :

15: 13:

1 0 : 8 : 5 : 5 : 9 ; h e a d s h i n y , exca v a ted , g ra n u l a ted ; fron t a l l i n e com p l e te ; occ i p i t a l ca r i n a i n comp l e t e , on l y v i s i b l e be h i n d t h e oce l l i a n d on t h e s i des of t he poste r i or oce l l i , b u t n e v e r reac h i n g t e m p l es ; POL � 2 ; O L 1 , 5 ; OOL � 1 0 ; p ro n o t u m crossed b y a s t ro n g t r a n s v e rsa l i m p ress i on , s h i n y , w e a k l y g r a n u l a ted ; scu t u m d u l l , g r a n u l a ted ; s c u te l l u m s hort , t r a n sverse ; m e t a n o t u m t r a n s v erse l y s t r i a t e , w i t h s i des p ro t r u d i n g ; p ro t r u s i o n s p o i n te d ; m e t a t horax + p ropode u m s h i n y , w i t h p l e u r a a n d post e r i o r s u rf ace t r a n sverse l y s t r i a t e ; a n te r i or s u rface a l most w i t h o u t sc u l p t u r e ; meso-met a p l eu r a l s u t u re d i s t i nc t , com p l e t e ; fore t a r s a l segments in fo l l ow i n g p ropor t i o n s : 1 7 : 3 : 6: 20 : 30 ; e n l a rged c l a w ( F i g . 935 ) w i t h s U b a p i c a l too t h a n d 6 l am e l l ae ; segme n t 5 of front t a rs u s ( F i g . 935 ) w i t h 1 row of 1 3 l a m e l l ae ; a p e x w i t h a g ro u p of at l ea s t 20 l a me l l ae ; m a x i l l a r y p a l p i w i t h 6 segme n t s ; l ab i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k n o w n L O C U S T Y P I C U S : L a Lo l a ( C o s t a R i c a ) T Y P I CA L MAT E R I A L : h o l o t y p e F ! i n WA . H O ST S : u n k n o w n D I ST R I BU T I O N : Neo t rop i c reg i on : C O S T A R I CA : L a Lo l a , W A ! N OT E : t h e s p ec i e s i s n a med i n honor o f M r . A r no l d M e n k e .

5 . A p terodryi n u s tij u c a n u s

( A r l e ) n . com b .

Pseu dogon a top u s tij u c a n u s A r l e 1 935 : 5 2 .

F E MA L E : a p terou s ; l e n g t h 6 m m ; h e a d b l ac k , w i t h m a n d i b l es tes t aceou s ; c l y p e u s l a ter a l l y test aceou s , med i a l l y b row n ; a n te n n ae t e s t aceo u s ; t ho r a x a n d p ropodeum b l ac k , w i t h sc u t u m y e l l o w ; a b domen b ro w n - t e s t aceou s ; l eg s testaceou s , w i t h cox ae , troc h a n ters a n d femora p a r t I y b row n ; a n t e n n a e

1 366

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1 984

937 - M a l e of Apterodryin u s tij u c a n u s ( A r l e ) from F l ores t a de T i j uc a ( B r'a z i I )

d i s t a l l y t h i c kened ; a n t e n n a I segmen t s i n fo l l ow i n g p ropor t i o n s : 1 0 : 8 : 32 : 20 : 1 7 : 1 2 : 8 : 7 : 5 . 5 : 9 ; head e x c a v a t e d , d u l l , g r a n u l a t e d , l o n g i t u d i n a l l y weak l y s t r i a t e ; f ro n t a l l i n e com p l e te ; occ i p i t a l c a r i n a i n com p l e te , on l y v i s i b l e b e h i n d t h e ocel l i a n d o n t h e s i des of t h e pos ter i or oce l l i , b u t never reac h i n g tem p l es ; pron o t u m s h i n y , crossed b y a s t ron g t r a n sversa I i m p ress i on , a s scu I p t u red a s head ; s c u t u m d u I I , g r a n u l a ted ; sc u te l l u m very sho r t , t r a n s v erse ; m e t a n o t u m n o t f l a t , n o t t r a n sverse l y s t r i a t e , w i t h t wo l a tera l po i n t s ; met a t ho r a x + p t�opodeum d u l l , g ra n u l a t e d , w i t h pos t e r i or s u rface s t ron g l y t r a n sverse l y s t r i a t e ; p l eu r a n o t t r a n s v erse l y s tr i a te ; meso - m e t a p l e u r a l s u t u re d i s t i n c t , com p l e te ; fore t a r s a l segmen t s i n fo l l o w i n g p ropor t i on s : 2 2 : 3 : 6 : 27 : 37 ; e n l arged c l a w ( F i g . 936 ) w i t h s u b a p i c a l too t h a n d 9-1 2 l a m e l l ae ; segme n t 5 of fron t t a r s u s ( F i g . 936 ) w i t h t w o row s o f nea r l y 3 6 l a me l l a e ; a p e x w i t h a g ro u p of n e a r l y 1 2 l a me l l ae ; m a x i l l a l'y p a l p i w i t h 6 segme n ts ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E ( F i g s 937 , 938 ) : f u l l y w i n ged ; l e n g t h 2 , 68 m m ; b l ac k ; m a n d i b l es t e s t aceo u s ; l eg s b row n , w i t h t a r s i l i g h t ; a n t e n n ae n o t d i s t a l l y t h i ck e n e d ; a n ten n a l segmen t s i n fol l ow i n g p roport i on s : 6 : 5 : 1 1 : 1 1 : 1 0 : 1 0 : 9 : 9 : 8 :

Go n a topod i n ae

F i g . 938 - M a l e of Ap terociryirllls

1 367

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8 ; a n t e n n a I s e g m e n t 3 more t h a n t h ree t i mes as l o n g as b ro a d ( 1 6 : 3 , 5 ) ; h e a d d u l l , g .' a n u l a t ed ; f l'o n t a l l i n e a b s e n t ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 6 , 5 ; O L = 2 , 5 ; OOL = 4 ; t em p l e s w e a k l y d i s t i n c t ; sc u t u m d u l l , g r a n u l a te d ; n o t a u l i ces com p l e te , p o s t e r i o r l y separ�a t e cJ ; m i n i m u m d i s t a n c e b e t w e e n t h e n o t a u l i c es as l o n g as b r e a d t h of t h e oc e l l i ; sc u t e l l u m a n d m e t a n o t u m s h i n y , smoo t h , weak l y p u n c t a t e , w i t ho u t scu l p t u re a m o n g p u n c t u re s ; p ro p o d e u m d u l l , f u l l y ret i c u l a t e r u gose ; fore-w i n g h y a l i n e , w i t ho u t d a rk t ra n s v e rsa l b a n d s ; r a d i a l c e l l c l osed : d i s t a l p a r t of r a d i a l v e i n t w i ce a s l o n g a s p ro x i m a l p a r t ( 24 : 1 2 ) ; gonoforceps w i t h a s h o r t dors a l p .'ocess ( F i g . 939 ) , w i t h a p e x b ro a d ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l s p u rs 1 , 1 , 2 . LOCUS TYP I C U S : P i co d a T i j u c a ( 3 r az i l ) ( ho l o t y p e ) ; F l o res t a d a T i j u c a ( B r a z i I ) ( n eo t y p e ) TYP I CA L MAT E R I AL : ho l o t y p e F l o s t ( p erson a l com m u n i c a t i on o f M r . A r l e ) ; n eo t y ;:> e F ' ( h ereby d es i g n a te d ) i n T W . HOST S : u n k n o w n D I STR I B U T I O N : Neo t rop i c reg i o n : BliAZ I L : F l ores t a d a T i j u c a ( G u a n a b a .'a ) , T W ' P i co d a T i j uc a ( G u a n a b a r a ) ( A r l e 1 935 ) .

6 . A p terodryin u s longicornis

( K i effer ) n . comb .

Gon a topus longicornis K i effer 1 904a : 357 . Dicondylus longicol'n is ( K i effer ) : K i ef f e r 1 9 1 4 b : 78 . Dolichochelys longicornis ( K i effer ) : O l m i a n d C u r-rado 1 976a :

1 78 .

1 368

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939 - M a l e g e n i t a l i a of Ap terodry inlls tljll c a n ll s ( A r l e ) from F l o re s t a de T i j uc a ( B r a z i I )

F E MALE : ap terou s ; l e n g t h 5 m m ; y e l i ow - t e s t a c eo u s , w i t h p e t i o l e b l ac k ; a n te n n a l segme n t s i n fo l l o w i n g p ro p o r t i o n s : 8 : 8 : 3 1 : 23 : 1 8 : 1 4 : 9 : 8 : 7 : 1 0 ; h e a d s h i n y or d u l l , smoo t h , h a i r l es s , w i t ho u t scu l p t u r e , excep t for occ i p u t and p a r t o f v e r t e x g ra n u l a te d ; POL � 1 , 5 ; OL � 2 , 5 ; OOL � 1 3 ; p rono t u m crossed b y a s t ro n g t r a n s vers a l i m p ress i o n , s h i n y , h a i r l e ss , g r a n u l a t e d ; sc u t u m d u l l , g r a n u l a te d , w i t h t w o l a t e ra l , p o i n te d protrus i o n s ; scute l l um weak l y v i s i b l e ; met anotum s h i n y or d u l l , not h o l l o w b e h i n d t h e sc u t e l l u m , t r a n sverse l y s t r i a t e ; m e t a t h or a x + p ro p o d e u m s h i n y o r d u l l , w i t h o u t s c u l p t u re ; p l e u ra , pos t e r i o r s u rface of m e t a t h o r a x + propo d e u m a n d p a r' t of a n t e r i o r s u rface t r a n sverse l y s t r i a t e ; meso m e t a p l e u ra l s u t u re o b so l e t e ; fore t a r s a l segmen t s i n fo l l o w i n g p roport i o n s : 20 : 3 : 5 : 25 : 37 ; e n l a r g e d c l a w ( F i g . 940 ) w i t h s u b a p i c a l t oo t h a n d 7 l a me l l a e ; se g m e n t 5 of fro n t t a r' s u s ( F i g . 940 ) w i t h t w o rows of 29 l a me l l ae ; a pex w i t h a g r o u p of 25 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 6 segme n t s ; l a b i a l p a l p i w i t h 3 seg men t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u nk nown L O C U S T Y P I C U S : Coro i co a n d C h i l u m a n i ( Bo l i v i a )

Gon a topod i n ae

F i g . 940

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C h e l a of A p terodryin u s longicorn i s ( Br a z i I )

TYP I CAL MAT E R I AL :

ho l o t y p e F !

1 369

( K i effer ) f rom Nov a Teu ton i a

i n GE .

H O ST S : u n k n o w n D I ST R I B U T I O N : Neot rop i c reg i o n : BRAZ I L : Nov a T e u t o n i a ( S . t a C a t a r i n a ) , BM ' B OL I V I A : Coro i co a n d C h i l u m a n i , G E !

7 . A p terodryin u s bellottii n . s p .

1 370

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F E MAL E : a p terou s ; l e n g t h 4 , 8 m m ; head b row n , w i t h m a n d i b l e s , c l y p e u s a n d a n te r i or re g i o n of f ro n s t e s t a c eo u s ; i n a d d i t i o n l a te r a l reg i o n s o f f r o n s ( more a l o n g orb i t s ) testaceo u s ; a n te n n a e b row n , w i t h seg m e n t s 1 a n d 8- 1 0 y e l l ow ; propec t u s redd i s h - t e s t aceo u s ; pro n o t u m redd i s h testaceo u s , w i t h t r a n sv e rs a l i m p ress i o n d a r k ; scu t u m red d i s h - t e s t aceo u s ; sc u te l l u m , m e t a t ho r a x + propo de u m , mesop l e u r a a n d m e t a p l eu r a b l ac k ; a b domen t e s t a c eo u s ; l eg s t e s t aceo u s , w i t h p a r t of coxae a n d p a r t of c l u b s o f femora b l ac k ; a n t e n n a e d i s t a l l y t h i ck e n e d ; a n t en n a l segmen t s i n f o l l ow i n g propor t i o n s : 1 4 : 8 : 24 : 1 5 : 1 1 : 1 1 : 9 : 9 : 9 : 1 3 ; h e a d v ery exc a v a t ed , s h i n y , smoo t h , w i t ho u t scu l p t u re , w i t h t r a c k s o f t r a n sversa l , short k ee l s o n a n te r i or h a l f of f r o n s a n d o n t h e s i des o f fron t a l l i n e ; fron t a l l i n e comp l e te ; occ i p i t a l c a r i n a i ncomp l e t e , o n l y v i s i b l e b eh i n d t h e oce l l i ; P O L = 2 ; O l = 4 ; O O l = 1 2 ; p os t e r i o r oce l l i touched b y t h e t r a c k o f occ i p i t a l c a r i n a ; p ron o t u m w i t h a s t ron g t r a n s v e rs a l i m pres s i o n , s h i n y , smoo t h , w i t ho u t sc u l p t u re ; sc u t u m a n d sc u te l l u m s h i n y , smoo t h , w i t h o u t scu l p t u re ; s c u t u m w i t h a l a t e r a l s t r o n g p o i n t ; meso-m e t a p l eu r a l s u t u re d i s t i n c t , comp l e t e ; m e t a n o t u m reduced to a short t r a n s v e rs a l k ee l , wea k l y v i s i b l e b e h i n d t he sc u te l l u m ; m e t a t h o r a x + p ropode u m s h i n y , smoot h , w i t h o u t sc u l p t u re , except for t r a n svers a l st r i ae o n p l e u ra a n d pos t e r i o r s u r f a c e ; fore t a r s a l se g m e n t s i n f o l l o w i n g p ropor t i on s : 25 : 5 : 8 : 24 : 38 ; e n l a rged c l a w ( F i g . 9/+ 1 ) w i t h s u b a p i c a l too t h a n d 7 l a m e l l ae ; seg m e n t 5 o f f r o n t t a rs u s ( F i g . 941 ) w i t h t w o rows o f n e a r l y 1 8 l a me l l ae ; apex w i t h a g r o u p o f n e a r' l y 9 l am e l l ae ; m a x i l l a ry p a l p i w i t h 6 segme n t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k no w n l O C U S T Y P I C U S : R o a d F ac a t a t i v a -Ano l a i m a ( C o l omb i a ) TYP I CAL M A T E R I AL : h o l o t y p e F ! i n A M . HOST S : u n k n o w n D I ST R I B U T I O N : Neot rop i c reg i o n : ( C u n d i n a m a rc a ) , AM ! NOTES:

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t h e spec i es i s n a med i n honor of M r .

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K E Y TO T H E S P E C I E S

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1 37 2

Mem . Amer . E n t .

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Me s o -me t a p l e u r a l s u t u r e v e r y s t r o n g and d i s t i n c t h o l l ow b e h i n d t h e s c u t e l l u m . 1 . n o togeicus n . sp .

me t a n o t u m

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U n k no w n

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sp.

F E MA L E : a p t e ro u s ; l en g t h 4 , 00-4 , 25 m m ; red d i s h - t e s t aceou s , w i t h p a r t of t he a b domen b ro w n ; a n t e n n a e wea k l y d i s t a l l y t h i c k e n e d ; a n te n n a I segmen t s i n fo l l o w i n g propor t i o n s : 9 : 7 : 1 7 : 1 0 : 9 : 7 : 5 : 4 , 5 : 4 , 5 : 6 , 5 ; h e a d e x c a v a ted , s h i n y , w i t ho u t s c u l p t u re , excep t for occ i p u t a n d a n te r i o r

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942 - C he l a of ,1p ter'oJryin u s no togeic u s n .

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re g i on of f ro n s g r a n u l a t e d ; fron t a l l i n e com p l e t e ; POL = 2 ; OL = 3 ; OOL 9; occ i p i t a l c a r i n a i n v i s i b l e ; p ro n o t u m sh i n y , crossed b y a s t ro n g t ra n s ve rs a l i m p ress i o n , w e a k l y t r a n sverse l y s t r i a t e ; scu t u m s h i n y , w i t h w e a k l on g i t u d i n a l k ee l s ; sc u te l l u m smoo t h , s h i n y , w i t ho u t sc u l p t u r e ; m e t a no t u m t r a n s v erse l y s t r i a t e , ho l l o w b e h i n d t he s c u te l l u m ; meso m e t a p l e u r a l s u t u re d i s t i n c t , com p l e te ; m e t a t ho r a x + p ropodeum s h i n y , w i t ho u t sc u l p t u re , w i t h p os t e r i o r s u rface t r a n sverse l y s t r i a t e ; a n te r i or s u rface w i t h a w e a k med i a n f u r ro w ; mesop l e u r a n o t t r a n s verse l y s t r i a te ; m e t a p l e u r a t r a n sverse l y s t r i a te ; fore t a rs a l s e g me n t s i n fo l l ow i n g p ropor t i o n s : 1 8 : 3 : 6 : 1 6 : 2 7 ; e n l a r ged c l a w ( F i g . 942 ) w i t h s U b a p i c a l too t h a n d 9 l a m e l l ae ; segmen t 5 o f fro n t t a rs u s ( F i g . 942 ) w i t h 1 row of 1 3 l a m e l l ae ; apex w i t h a g ro u p of 1 7 l a m e l l a e ; m a x i I l a r y p a l p i w i t h 6 segme n t s ; l ab i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n LOCUS TYP I C U S : Perth ( A u s t ra l i a ) T Y P I CA L MAT E R I AL : ho l o t y pe F ! i n B M ; 1 p a r a t y p e F ! i n O L . HOSTS : u n k n o w n D I ST R I BU T I O N : A u s t r a l i a n reg i o n : AUSTRAL I A : P e r t h ( Western A u s t r a l i a ) , BM !

2 . A p terodryin u s insul aris n . s p .

F E MAL E : a p terou s ; l e n g t h 2 , 5 m m ; y e l l o w - t e s t aceou s , w i t h p e t i o l e b l ac k a n d a b domen b row n i s h ; a n te n n a l segme n t s i n fo l l ow i n g p roport i o n s : 8 : 5 :

1 1 : 6 , 5 : 5 : 4 : 5 ( l a s t t h ree segmen t s m i s s i n g i n t he o n l y k n o w n spec i me n ) ; h e a d s h i n y , exca v a ted , h a i r l ess , w i t h o u t scu l p t u re , exce p t for o cc i p u t a n d f ro n t p a r t of v e r t e x weak I y g r a n u I a ted ; pron o t u m c rossed b y a s t r o n g t r a n s v e rs a l i mp ress i o n , s h i n y , w i t ho u t s c u l p t u re ; scu t u m s h i n y , w i t ho u t sc u l p t u re , l a tera l l y w i t ho u t po i n t s ; scu t e l l u m w e a k l y v i s i b l e ; m e t a n o t u m n o t ho l l o w beh i n d t he sc u te l l u m ; m e t a t ho r a x + propodeum s h i n y , w i t ho u t sc u l p t u re ; p l eu r a and p os t e r i o r s u rf ace of propodeum t r a n s v e r se l y s t r i a t e ; meso-me t a p l eu r a l s u t u re obso l e t e ; segmen t 1 of f ro n t t a rs u s a s l o n g a s segmen t 4 ; e n l a rged c l a w ( F i g . 943 ) w i t h s u b a p i c a l too t h a n d 8 l a m e l l ae ; seg m e n t 5 o f f ro n t t a rs u s ( F i g . 943 ) w i t h t wo ro w s of 2 + 1 3 l a m e l l ae ; a pex w i t h a g ro u p o f a t l ea s t 1 6 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l ab i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k no w n L O C U S T Y P I C U S : L a r a t I s l a n d ( I n dones i a ) TYP I C A L M A T E R I A L : ho l o t y p e F ! i n B . HOST S : u n k n o w n D I ST R I B U T I O N : A u s t r a l i a n reg i o n : I N DONES I A : L a r a t I s l a n d ( T a n i m b a r I s l an d s ) , B !

9 . Dicondylus H a l i d a y

Gon a topod i n a e

1 3 75

nec L a beo C u v i e r 1 8 1 7 : 1 94 . Dicondyl u s H a l i d a y i n C u r t i s 1 8 29-30 : 1 1 0 . Dicondyl u s H a l i d a y 1 833 : 2 73 . L a beo H a l i d a y 1 83 3 : 273 ( s y n . p roposed by R i c h a r d s 1 937 ) ; t y p e s p ec i e s L a beo v i tripennis H a l i d a y 1 83 3 , monot y p i c . n ec L a bea B l a n c h a r d 1 840 : 430 . L a beola H a l d e m a n 1 84 2 : 1 92 ( s y n . p roposed b y K ro m b e i n 1 979 ) , new name. La bea B l a n c h a r d : D a l l a Torre 1 898 : 542 . L a beo H a l i d a y : K i effer i n K i ef f e r et M a r' s h a l l 1 905 : 220 . Trigo n a topus K i effer 1 9 1 3c : 325 ( n . sy n . ) ; t y p e spec i es Trigon a top u s b ifarius ( K i ef f e r 1 906 ) , o r i ,) . d es i ,] . L a � erius I( i e ffer 1 9 1 4 b : 59 ( s y n . p roposed b y R i c h a rd s 1 9 37 ) ; new name. Trigon a topus !< i effer : K i ef f e r 1 9 1 4b : 7 2 . Dicondyl u s H a l i d a y : K i ef f e r 1 9 1 4b : 7 2 . L a berinus O g l ob l i n 1 932 : 266 ( m i s p e l l i n g o r l a p s u s ) . Dicondylus H a l i d a y : R i c h a r d s 1 935 : 1 47 . L a berius K i effer : R i c h a r ds 1 9 36a : 1 69 . Dicon dyl u s H a l i d a y : R i c h a r d s 1 937 : 1 1 8 . Idologon a topus O g l ob l i n 1 95 3 : 1 1 1 ( n . s y n . ) ; t y p e spec i es Idologon a top u s nigrithorax O g l o b l i n 1 95 3 , o r i g . des i g .

TYPE S P E C I E S : Dicondylus bicolor ( H a l i d a y 1 828 ) , d e s i g . b y K i effer ( 1 9 1 4b ) • F E MA L E ( F i g s 947 , 948 ) : a p terou s ; m a x i l l a r y p a l p i w i t h 2-4 segmen t s ; l a b i a l p a l p i w i t h 2 segme n t s ; p r o n o t u m ( F i g . 948 ) n o t crossed b y a s t r o n g t r a n s v e rs a l i m p ress i o n or w ea k l y i m p ressed ; e n l a rged c l a w ( F i g . 944 ) w i t h s u b a p i c a l too t h a n d l a m e l l ae ; t i b i a l s p u r s 1 , 0 , 1 . MALE ( F i g s 952 , 953 ) : f u l l y w i n g ed ; m a x i l l a r y p a l p i w i t h 2-4 s e g m e n t s ; l a b i a l p a l p i w i t h 2 segmen ts ; tem p l es d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e o r on l y v i s i b l e b e h i n d the oce l l i o r o n t h e s i des o f t h e poste r i o r oce l l i , b u t n e v e r reac h i n g temp l e s ; occ i p u t con c a ve ; t i b i a l s p u rs 1 , 1 , 2 . D I ST R I B U T I O N : wor l d w i de HOST S : Delp h a cidae SPEC I ES : 1 3

PALAEARCT I C S P E C I E S O F D ICOND YLUS

T w o s p ec i e s are k n ow n .

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F E MA L E S

Me t a t h o ra x + p r o p od e u m , s c u t e l l u m a n d p a r t o f t h e f i r s t a b d om i n a l t e rg i t e r e d d i s h o r y e l l ow - r ed d i s h . . . . . . . . . . . . . . . . . . . dich rom u s ( K i e f f e r ) · • • • • . • • . • . . . . . • • . . . . • . . • • • • • 1 . Me t a t h o r a x + p r o p od e u m , s c u t e l l u m a n d f i r s t a b d om i n a l t e rg i t e b l a c k ( e x c e p t f o r t h e r ed d i s h a p ex o f p r o p o d eu m ) • • . . • . • • • . • . • • ·

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b icolor ( H a I i d a y )

MALES

M i n i mu m d i s t a n c e b e t w e e n t h e n o t a u l i c e s n e a r l y a s l o n g a s b r ead t h o f t h e o c e l l i ( F i g . 9 4 5 B ) . . • • • • • . . • • • • • . . • • • • • • • • • . · • . . . . . • • . . . • . . , . . . . . . . • . . . . . 2 . b icolor ( Ha I i d a y ) M i n i mu m d i s t a n c e b e t w e e n t h e n o t a u l i c e s s h o r t e r t h a n b ,' ea d t h o f t h e o c e l l i ( F i g . 9 4 5 A ) • • • l . dichrom u s ( K i e f f e r )

1 . Dicondyl u s dichrom u s

( K i ef f e r )

Gon a to p u s dichrom u s K i effer i n K i e f f e r et M a rs h a l l 1 906 : 505 . Dicondylus dichrom u s ( K i ef f er ) : Ber l a n d 1 928 : 1 46 . Gon a to p u s rufescens H e l l e n 1 93 5 : 8 ( s y n . p roposed b y He l l en

1 95 3 ) ; nomen n u d u m . D i c ondyl u s helleni R a a t i k a i n e n

1 96 1 : 1 26 ( n . s y n . ) ; locus ( F i n l and ) . Dicondy l u s dichrom u s ( K i effe r ) : R a a t i k a i n en 1 96 1 : 1 3 1 . typ i c u s L a i h i a

D i c on dy l u s s i b iric u s Ponomarenko 1 965 : typ i c u s R o m a n o v k a ( S i b i r i a , U . S . S . R . ) .

624

( n . sy n . ) ; locus

( K i ef f er ) : Moc z a r 1 965 : 393 . R a a t i k a i ne n 1 97 0 : 3 2 1 . ffaplogon a top u s p a c e i C u rr a d o a n d O l m i 1 97 7 : 3 1 7 ( n . s y n . ) ; l o c u s typ i c u s G r i so l i a ( I t a l y ) . Pseu dogon a topus rufu s Ponomaren k o 1 97 9 a : 355 ( n . s y n . ) ; l o c u s typ i c u s 2 5 K m N of B u l g a n ( Mo n g o l i a ) . Dicon dyl u s dichrom u s

Dicondylus helleni R a a t i k a i n en :

( K i ef f er ) : Mocza r 1 983b : 206 . ( R a a t i k a i n e n ) : Moc z a r 1 983b : 206 .

Dicondyl u s dichrom u s Dicondy l u s helleni

F E MAL E : a p t e rou s ; l e n g t h 2 , 50-3 , 25 m m ; red d i s h or y e l l ow - t e s t a ceou s , w i t h a n te n n a I se g m e n t s 3-9 b row n ; occ a s i on a l l y ver tex o f head b row n ; p e t i o l e b l ac k ; a b domen red d i s h - y e l l ow o r b ro w n - b l ac k , occ a s i o n a l l y w i t h f i rs t t e r g i t e red d i s h ;

h e a d d u l l , exc a v a ted ,

g r a n u l a ted ;

p ron o t u m n o t

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/ //// /

F i g . 944 - C he l a o f Diconclyl u s dichrom u s

( K i e f fe r ) f r o m L a i h i a ( F i n l a n d )

c r'osseu

b y a s t ro n g t r a n s v e rs a l i m p ress i o n o r we a k l y i m p ressed , d u l l , g r a n u l a t e d ; s c u te l l u m d u l l , w i t ho u t scu l p t u re ; met a n o t u m w i t h s i de s p r o t r u d i n g ; p ro t r u s i on s po i n ted ; met a t h o r a x + p ropodeum d u l l , g ra n u l a t ed , w i t h pos t er i or s u rface a n d p l e u ra t r a n s verse l y s t r i a t e ; meso me t a p l e u ra l s u t u re obso l e t e ; seg m e n t 1 o f f ro n t t a rs u s a p p rox i m a te l y t w i ce as l o n g as segmen t 4 ( 1 2 : 6 ) ; en l a r g e d c l a w ( F i g . 944) w i t h s u b a p i c a l too t h a n d 4-5 l ame l l a e ; segmen t 5 of f ron t t a rs u s ( F i g . 944 ) w i t h 2 rows o f 6-9 l a me l l ae ; a p e x w i t h a g ro u p o f 3-6 l a me l l ae ; m a x i l l a ry p a l p i w i t h 2-4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : f u l l y w i n ged ; l e n g t h 2 , 37-2 , 62 mm ; b l ac k ; m a n d i b l e s y e l l ow ; a n te n n a e a n d t e g u l ae b ro w n ; l e g s b ro w n - y e l l ow ; a n t e n n a I s e g me n t 3 f o u r t i mes or m o r e t h a n f o u r t i mes a s l on g a s b ro a d ( 5 , 25 ) ; h e a d s h i n y , h a i r y , g r a n u l a te d ; f ro n t a l l i n e a b s e n t ; t em p l e s d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e ; occ i p u t conc a v e ; sc u t u m s h i n y , g r a n u l a ted ; n o t a u l i ces d i s t i n c t , comp l e te , pos t e r i or l y sep a r a ted ; m i n i m u m d i s t a nce b e t ween t h e n o t a u l i ces ( F i g . 945 A) m u c h s ho r t e r t h a n b r e a d t h o f the oce l l i ; s c u t e l l u m a n d m e t a n o t u m s h i n y , smoo t h , p u n c t a t e ; p ropodeum ret i c u l a t e ru gose , w i t h o u t k ee l s ; fore- w i n g h y a l i n e , w i t ho u t d a rk t r a n s v e rs a l b a n d s ; dors a l p rocess o f gonoforceps ( F i g . 946 A ) w i t h p a d d l e- s h a p ed a p e x ; m a x i I l a r y p a l p i w i t h 4 s e g m e n t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u rs 1 , 1 , 2 . L O C U S TYP I C U S :

Corse ( F r a n c e )

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F i g . 945 - N o t a u l i ces of m a l es o f Dicondylus dich rom u s ( K i ef f e r ) f rom Csevharaszt ( H u ngary ) ( A )

a n d b icolor ( H a l i d a y ) f rom Germ a n y

(B).

T Y P I C A L M A T E R I AL : h o l o t y p e F ! o f G . dichromus i n B U ; h o l o t y p e F ! a n d 3 p a r a t y p e s ( 1 F , 2 M M ) ! of D. helleni i n H E ; o t h e r p a r a t y pes of D . hellen i i n H E a n d i n R a a t i k a i n e n ' s col l ec t i o n ; ho l o t y pe F ! a n d a l l o t y p e M ! of D . sibiric u s i n MO ; 2 p a ra t y pes ( 1 F , 1 M ) ! o f D . s i b iric u s i n B�� ; h o l o t y p e F ! of H. p a ce i i n V R ; h o l o t y p e F ! of Ps . rufus i n L N . HOST S : i n F i n I a n d Megadelp h a x sordidulus ( S t a I ) , Un k a nodes excisa ( Me l . ) ( = Delp h a x excisu s ) ( Delp h a c ida e ) ( R a a t i k a i ne n 1 960 , 1 96 1 , 1 970 ) . B I OLOGY : e g g , l a rv a , cocoon , p u p a a n d l i fe cy c l e desc r i b e d b y R a a t i k a i nen ( 1 96 1 ) . N A T U RAL E N E M I ES : i n F i n l a n d A c hol'olop h u s gracilipes ( K ramer ) ( A carin a , Erythraeida e ) ( R a a t i k a i n e n 1 96 1 ) . D I ST R I B U T I ON : P a l a e a rc t i c reg i on : M O N GOL I A : 2 5 K m N of Bu l g a n ( R i ve r U I j a s t a y n-Go l , H o v d a y m a g , W e s t Mon g o l i a ) , L N ! 1 1 K m W f r o m Somon Baj a n n u u r ( Bu l g a n a i m a k ) , B U ! 3 K m N from Somon Uenc i n T a l U e n c ( C hovd a i m a k ) , B U ! Somon Baj a n zo g t ( C en t r a l a i m a k ) , B U ! U l a n - B a a to r ( C en t r a l a i m ak ) , B U ! U . S . S . R . : Romanov k a ( Ye r a v n e s n k i y D i s t r i c t , B u r y a t A . S . S . R . , S i b i r i a ) , B M ! WA ! M O ! F I NLA N D : L a i h i a ( V a a s a n ) , H E ! T v a rm i n n e , H E ! H a n go ( U u de n ma ) , H E ! H e n r i k sb e r g ( R a a t i k a i ne n 1 96 1 ) ; K a u n i a i ne n ( R a a t i k a i n en 1 96 1 ) ; T a i p a l sa a r i ( R a a t i k a i n en 1 96 1 ) ; SWEDE N : A n ge l ho l m ( K r i s t i a n s t a d ) , H E ! H U NGAR Y : M a t ra sz t i m re , B U ! Csev h a r a sz t , B U ! V a l ko , B U ! B u k k-h g s . , B U ! Hosszu herc ( B u k k ) , B U ! G E RMANY : B er l i n , E B ! A U ST R I A : Vos l a u , P ! D E N M A R K , C O ! HOLLA N D : T e rsche l l i n g , L E ! SW I T Z E R L AN D : Vercor i n ( V a l a i s ) , LO ! Les P o n t s ( N e u c h a te l ) , LO ! F R A N C E : Corse , B U ! SPA I N : Sa l ou ( T a rr a g on a ) , BM ! O L ! I TALY : Bu sso l en o ( T or i n o ) , O L ! G r i so l i a ( Cosenza ) , V R !

2 . Dicondylus b icolor ( H a l i d a y )

Dryin u s b icolor H a l i d a y 1 828 i n C u r t i s 1 828 : 206 . Dicondylus bicolor H a l i d a y i n C u rt i s 1 829-30 : 1 1 0 . L a beo vitripennis H a l i d a y 1 833 : 273 ( s y n . p roposed b y R i c h a rds 1 939 ) ; locus typicus I r e i a n d . A n t aeon excis u s Wes t wood 1 833b : 497 ( s y n . proposed by R i c h a rd s 1 939 ) ; locus typicus C a m b r i d g e ( E n g l a n d ) .

Gon a topod i n a e

Fig.

1 379

946 - M a l e g e n i t a l i a of Dicondylus dichromus ( K i effe r ) from L a i h i a

( F i n l an d ) ( A )

a n d b icolor ( H a l i d a y ) from Born t a l removed )

( Ge r m a n y ) ( 3 ) ( r i g h t h a l f

Gon a topus conjunctus K i effer i n K i effer et M a r s h a l l 1 90 5 : 1 1 9 ( n . s y n . ) ; locus typ i c u s S t a v a n ger ( Norw a y ) . Gon a top u s b ifarius K i effer i n K i effer e t M a r s h a l l 1 906 : 502 ( n. s y n . ) ; locus typ i c u s Corse ( F r a n ce ) . Gon a top u s b icolor ( H a l i d a y ) : K i effer i n K i effer e t M a r s h a l l 1 906 : 50S . C a b eo excisus ( We s t wood ) : K i effer 1 907b : 1 5 . Pla tygo n a topus b ifa rius ( K i effer ) : K i effer 1 907b : 1 9 . Trigon a top u s b ifarius ( K i effer ) : K i effer 1 9 1 3c : 325 . Dicondylus b icolor ( H a l i d a y ) : K i effer 1 9 1 4b : 7 3 . Gon a topus decretorius H a u p t 1 9 1 6 a : 4 6 ( n . s y n . ) ; loc u s typ ic u s S w i n e m u n d e ( Ge r m a n y ) . L a berius excis u s ( We s t wood ) : B er l a n d 1 925 : 1 45 . Trigon a top u s b ifa rius ( K i e ffer ) : Ber l a n d 1 92 5 : 1 45 . L a berius vitripennis ( H a l i d a y ) : R i c h a rd s 1 935 : 1 45 . Dicondylus b icolor ( Ha l i d a y ) : R i ch a rds 1 9 35 : 1 47 . Dicondylus b icolor ( H a l i d a y ) : B e h r i sch 1 95 2 : 34. D icondylus lindbergi H e i k i n he i mo 1 957 : 78 ( n . sy n . ) ; locus typicus L a i h i a ( F i n l a nd ) . Dicondyl us lindbergi H e i k i n h e i m o : R a a t i k a i n en 1 96 1 : 1 3 1 . Dicondylus b icolor ( H a l i d a y ) : R a a t i k a i n e n 1 96 1 : 1 3 1 . Haplogon a top u s b ifa ri u s ( K i effer ) : Moc z a r 1 965 : 391 .

1 380

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F i g . 947 - F e m a l e of Dicondyl u s bicolor ( H a l i d a y ) from L a i h i a

(Fin land)

Dicondyl u s lindb ergi H e i k i n h e i mo : R a a t i k a i n e n 1 967 : 88 . Gon a topus conjunctus K i effe r : R a a t i k a i n e n 1 967 : 88 . Dicondylus b icolor ( H a l i d a y ) : N a g y 1 9 67 a : 1 24 . Haplogon a top u s b ifarius ( K i effer ) : C u r r a d o a n d O l m i 1 974b : 4 1 . Dicondyl u s b icolor ( H a l i d a y ) : W a l off 1 974 : 1 04 . Dicondyl u s b icolor ( H a l i d a y ) : W a l of f 1 97 5 : 662 . Dicondyl u s b icolor ( H a l i d a y ) : J . F . P e r k i n s 1 976 : 3 1 .

F E MALE ( F i g s 947 , 948 ) : a p t e ro u s ; l e n g t h 2 , 75-3 , 00 m m ; h e a d b ro w n b l ac k , w i t h m a n d i b l e s , c l y pe u s a n d f ro n t p a r t of v e r t e x ( more a l o n g o rb i t s ) y e l l o w ; a n te n n a e b row n , w i t h segme n t s 1 -2 a n d occ a s i o n a l l y 1 0 y e l l o w o r w h i t i s h ; p rono t u m f u l l y y e l l ow-tes t a ceou s ; occ a s i o n a l l y co l l a r b l ac k a n d d i sc y e l l o w ; r a re l y pro n o t u m a l mo s t f u l l y b l ac k ; sc u t u m y e l l o w , w i t h s i des b l ac k ; sc u te l l u m a n d m e t a t ho r a x + p ropod e u m b l ac k , w i t h a p e x of p ropode u m y e l l o w ; a b domen b l ac k ; l eg s y e l l o w or y e l l ow b row n , occ a s i o n a l l y w i t h c l u b s o f femora f u l l y o r p a r t l y b row n i s h ; head e xc a v a ted , g r a n u I a t e d ; p ro no t u m n o t crossed by a s t ro n g t ra n s versa I i m p res s i o n or w e a k l y i m p ressed , s h i n y , g r a n u l a te d ; scu t u m g r a n u l a t e d ;

Gon a topod i n ae

1 38 1

F i g . 9 4 8 - F em a l e of Dicondyl u s bicolor ( H a l i d a y ) f rom L a i h i a

met a n o t u m t r a n s v e rs e l y s t r i a te ,

w i t h s i des p r o t r u d i n g ;

( F i n l an d )

p ro t r u s i o n s

p o i n t ed ; pos t e r i o r s u rface of p ropodeum a n d p l eu ra t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re o b so l e t e ; occ a s i o n a l l y d i s t i n c t , b u t v e r y w e a k ; fore t a rs a l s e g m e n t 1 a p p ro x i m a te l y t w i ce as l o n g as segmen t ' 4 ; en l a r g e d c l a w ( F i g . 949 ) w i t h s u b a p i c a l too t h a n d 4 - 5 l a m e l l ae ; segme n t 5 of f ro n t t a rs u s ( F i g . 949 ) w i t h t w o row s o f 6-9 l ame l l ae ; a p e x w i t h a g ro u p o f 3-6 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 2-4 segmen t s ; l ab i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . Ivt" L ,, : f u l l y w i n ged ; l e n g t h 1 , 57-2 , 00 mm ; h e a d b l ac k , w i t h m a n d i b l es y e l l o w ; a n te n n ae b l a c k - b ro w n , w i t h p a r t of seg men t s 1 -2 y e l l ow ; t ho r a x a n d p ro p o d e u m b l ack ; a b domen b l a c k - b ro w n ; l eg s y e l l o w , w i t h p a r t o f c o x a e a n d p a r t of femora b row n ; a n te n n a e n o t d i s t a l l y t h i ck e n e d ; a n t e n n a l se g m e n t s i n fo l l o w i n g p ropor t i o n s : 5 : 4 , 5 : 7 : 8 : 7 , 5 : 7 , 5 : 6 , 5 : 6 , 5 : 6 : 8 , 5 ; a n t e n n a l s e g m e n t 3 f o u r t i mes o r more t h a n fou r t i mes a s l o n g a s b ro a d ( 4 , 0 ) ; h e a d s h i n y , g r a n u l a t ed ; fron t a l l i n e a b sen t ; t em p l es d i s t i n c t ; no s h i n y , smoo t h a re a i s v i s i b l e on t h e s i des of t he poster i o r oce l l i ; occ i p i t a l ca r i n a i n v i s i b l e ; scu t u m s h i n y ; n o t a u l i ces com p l e t e , poste r i o r l y sep a r a ted ; m i n i m u m d i s t a n c e b e t ween t he n o t a u l i ce s ( F i g . 945 B ) n e a r l y a s l o n g a s b r ea d t h of the oce l l i ; s c u te l l u m a n d met a n o t u m s h i n y , w ea k l y p u n c t a t e ; p ropodeum re t i c u l a t e r u gose ; fore-w i n g h y a l i ne , w i t h o u t d a rk t r a n s v e rs a l b a n d s ; dors a l p rocess of g o n oforceps ( F i g . 946 G ) w i t h p a d d l e- s h a p e d a p e x ; m a x i I l a r y p a l p i w i t h 3 segme n t s ; l ab i a l p a l p i w i t h 2 segmen ts ; t i b i a l s p u r s 1 , 1 , 2 . L O C U S T Y P I C US : u n c er t a i n ( D e a l i n E n g l a n d accord i n g t o Mocz a r 1 965 ; D a ren t h Wood i n E n g l a n d accord i n g to R i c h a r d s 1 939 ) . T Y P I C A L MAT E R I AL : ho l o t y p e F ! a n d 1 p a r a t y p e F ! of D . b icolor i n D U ; ho l o t y p e F ! of G. conjunctus i n P ; h o l o t y p e F ! a n d 1 p a ra t y pe F ! of D . lindb ergi i n HE ; t he a l l ot y p e M ! o f D . lindb ergi i n H E i s rea l l y a m a l e

1 382

F i g . 949

M e m . Amer .

-

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Go n a topo d i n a e

1 383

o f Pseu dogon a top us dis tin c t u s ( K i effe r ) ; ho l o t y p e F ! of G . b ifarius i n BU ; h o l o t y p e M ! of L . vi tripe n n is i n D U ; h o l o t y p e M of A . excisus i n B M . HOST S : i n F i n l a n d lavesella pellucida ( F . ) ( H ek i n h e i mo 1 957 ; R a a t i k a i n en 1 967 ) , lavesella o b s c urella ( Boh . ) , Megadelp h a x sordidu l u s ( S t a l ) , Dicra n o tropis h a m a t a ( Boh . ) , Stiroma 'J icarina ta ( H . S . ) , Criom orp h u s a l b om argin a tu s C u r t . ( R a a t i k a i n e n 1 967 ) , Un k a n odes excis a ( M e l . ) ( = Delph a x excis u s ) ( L i n db e r g 1 950 ) ; i n I ta I y D icranotropis h a m a t a ( Bo h . ) ( C u rrado a n d O l m i 1 974b ) ; i n E n g l a n d lavesella pellucida ( F . ) , Criomorph u s albomargin a t u s C u r t i s , Hyledelp h a x elega n t u l u s ( Boh . ) ( W a I off 1 975 ) ; Dicra n o tropis h a m a t a ( Bo h . ) ( H a s s a n 1 93 9 ; W a l of f 1 975 ) ; D i tropis p teridis ( Sp i n . ) ( R i c h a rd s 1 933 ) ; Rib a u todelp h a x s p . ( = Delp h a codes coll inus Bo h . ) ( R i c h a rd s 1 93 9 ) ; Gra vesteiniella boldi ( Scot t ) ( R i c h a r d s 1 948 ) ; i n F r a nce Delp h a x collin a e Bo h . ( M a n ev a l 1 93 5 ) a n d Stiroma sp . ; i n Germ a n y Criomorp h u s b ic arin a tu s ( H . S . ) ( S t r u b i n g 1 95 6 ) , Un k a n odes excisa ( Me l . ) ( = Delp h a x excis u s ) ( H a u p t 1 9 1 6a ) ; i n Be l g i u m I2ib a u todelph a x p u n gens ( R i b a u t ) ( C . F . M . d e n B i e m a n l e g . ) ; i n Ho l l a n d Dicra n o tropis h a m a ta ( Bo h . ) , Rib a u todelp h a x collin u s ( Boh . ) , Delp h a cinus mesomelas ( Bo h . ) ( C . F . M . den B i e m a n l e g . ) . B I OLOGY : e g g , l a r v a , cocoo n , p u p a a n d l i fe c y c l e desc r i bed b y R a a t i k a i n e n ( 1 96 7 ) . U n i v o l t i n e i n F i n l a n d ( R a a t i k a i n en 1 967 ) . D I S T R I B U T I O N : P a l a e a rc t i c reg i o n : F I N L A N D : L a i h i a ( V a a s a n ) , H E ! WA ! K o k k o l a ( V a a sa n ) , H E ! Y l i s t a ro ( He i k i n h e i mo 1 957 ) ; U u s i k i rk k o ( He i k i n he i mo 1 957 ) ; Ec kero ( A l a n d ) , H E ! NORWAY : S t a v a n g e r , P ! H U NGARY : B u k k s z e n t l l e k , B U ! H u n g a r y ( N a g y 1 967a ) ; f
E T H I OP I AN S P E C I E S OF D lcn,vD YLUS

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F i g . 950 - C h e l a of Dicondy l u s p a l l idior n .

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F E MAL E : a p t e rou s ; l en g t h 3 , 37 m m ; h e a d b row n , w i t h f ro n t p a r t of ver tex , c l y p e u s a n d m a n d i b l es redd i s h ; a n te n n ae t e s t a c eo u s ; p r o n o t u m redd i s h -d a r k ; scu t u m , s c u t e l l u m , m e t a t ho r a x + p ropodeum red d i s h t e s t aceou s ; pe t i o l e b l ac k ; a b domen re d d i s h - t e s t aceou s ; l e g s y e l l ow ; h e a d exc a v a ted , s h i n y , w i t ho u t scu l p t u re or we a k l y g r a n u l a ted ; p ro n o t u m n o t crossed b y a s t r o n g t r a n s v e rs a l i m pres s i o n or w e a k l y i m p ressed , s h i n y , g r a n u l a ted ; scu t u m d u l l , w i t h some l o n g i t u d i n a l kee l s ; m e t a n o t u m t r a n s v e r se l y s t r i a te , w i t h s i d es r o u n d e d ; m e t a t ho r a x + p ropode u m s h i n y , w i t ho u t scu l p t u re ; p l e u r a a n d pos t e r i o r s u rf a ce o f p ropodeum t r a n s v e r se l y s t r i a t e ; meso-me t a p l e u r a l s u t u re obso i e t e ; fore t a rs a l seg men t 1 a p p rox i m a t e l y t w i ce as l o n g as seg men t 4 ; e n l a r ged c l a w ( F i g . 950 ) w i t h s u b a p i c a l too t h a n d 6 l a m e l l a e ; seg men t 5 of fro n t t a r s u s ( F i g . 950 ) w i t h t w o ro w s o f 1 1 l am e l l ae ; a p e x w i t h a group o f 8 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 3 s e g m e n t s ; l a b i a l p a l p i w i t h 2 s e g m e n t s ; t i b i a l s p u rs 1 , D , 1 . MAL E : u n k n o w n L O C U S T Y P I C U S : K a t b erg ( Sou t h A f r i ca ) TYP I C A L MAT E R I A L : h o l o t y p e F ! i n BM . HOST S : u n k no w n D I ST R I B U T I O N : E t h i o p i a n reg i o n : SOU T H A F R I CA : K a t b erg ( C a p e P rov i n c e ) , BM !

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O n e spec i es i s k n ow n .

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1 385

( type )

F E MA L E : a p t e ro u s ; l e n g t h 3 , 00-3 , 75 m m ; h e a d b l ack , w i t h occ i p u t , f ro n t p a r t o f v e r t e x , c l y p eu s a n d m a n d i b l es y e l l o w ; a n te n n a e b l ac k , w i t h v e n t r a l s i de of segmen t 1 y e l l o w ; p a r t of segmen t 1 0 y e l l o w ; t ho r a x , p ropode u m a n d a b domen b l ac k ; l eg s y e l l ow , w i t h p a r t of fore femora b row n ; i n a d d i t i o n part of fore and m i d t i b i a e b row n ; head e x c a v a ted , s h i n y , g r a n u l a te d ; f ro n t p a r t of v e r t e x w i t h some k ee l s ; p r o n o t u m d u l l ,

1 386

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g r a n u l a te d , w i t ho u t k ee l s , not crossed b y a s t ro n g t ra n svers a l i m p res s i on or w e a k l y i m presse d ; sc u t u m d u l l , w i t h some l on g i t u d i n a l k ee l s ; m e t a n o t u m a n d met a t ho r a x + p ropodeum f u l l y t r a n sverse l y s tr i a t e ; m e t a no t u m w i t h s i d e s ro u n d e d ; meso-me t a p l eu r a l s u t u re o b so l e te ; fore t a rs a l s e g me n t s i n fo l l ow i n g p ropor t i o n s : 1 7 : 2 , 5 : 5 : 1 4 : 2 1 ; en l a rged c l aw ( F i g . 9 5 1 ) w i t h su b a p i c a l t oo t h a n d 5-6 l a me l l ae ; s e g m e n t 5 of fron t t a rs u s ( F i g . 9 5 1 ) w i t h t w o rows of 1 8-23 l ame l l ae ; a pex w i t h a g ro u p of 8-1 0 l a me l l ae ; m a x i l l a ry p a l p i w i t h 4 segmen t s ; l ab i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE ( F i g s 952 , 953 ) : f u l l y w i n g e d ; l e n g t h 2 , 50-2 , 8 1 m m ; b l ac k , w i t h m a n d i b l es a n d t e g u l ae y e l l ow ; a n te n n ae b row n ; l eg s y e l l ow , w i t h h i n d coxae b ro w n ; a n te n n a l s e g m e n t s i n fo l l o w i n g p roport i on s : 1 0 : 7 : 1 4 : 1 5 : 1 4 : 1 3 : 1 3 : 1 3 : 1 2 : 1 3 ; a n te n n a l seg m e n t 3 more t h a n t h re e t i me s a s l o n g a s b ro a d ( 4 , 33 ) ; h e a d s h i n y , h a i r y , g r a n u l a t e d ; fron t a l l i ne a b se n t ; temp l es d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L � 1 0 ; OL � 3 , 5 ; OOL � 2 ;

Gon a topod i n a e

F i g . 953 - H a l e of :J iconuy l u s illcJ i o. n u s

1 387

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s p . fronc H o l a l u

( I nd i a )

b re a d t h of t h e oce l l i : 5 , 5 ; sc u t u m g r a n u l a ted ; n o t a u l i c e s d i s t i n c t , com p l e t e , pos t er i o r l y jo i n t o r v e r y n e a r , a l most j o i n t ; sc u t e l l u m s h i n y , smoo t h , we a k l y p u n c t a t e ; met a n o t u m d u l l , w i t h a n t e r i o r s u rface p u n c t a te ; p os t e r i o r s u rf a c e smoo t h ; p ropod e u m re t i c u l a t e ru gose , w i t h m ed i a n s u rface de l i rn i ted by s t r o n g kee l s ; fore-w i n g h y a l i n e , w i t h o u t d a r k t r a n s v e rs a l b a n d s ; dors a l p rocess o f gonoforceps ( F i g . 954 ) v e r y red uced , short , i n v i s i b l e i f t h e pen i s i s n o t sep a r a t e d from t h e g o n oforc e p s ; m a x i l l a r y p a l p i w i t h 4 s e g m e n t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 1 , 2 . LOCUS T Y P I C U S : Ho l a l u ( I n d i a ) T Y P I CAL MAT E R I A L : h o l o t y p e F ! a n d 7 p a r a t y p e s ( 4 F F , 3 MM ) ! i n 8 M ; 2 p a r a t y pes F F ! i n OL ; 2 p a r a t y p e s ( 1 F , 1 M ) ! i n K Y . H O ST S : i n I n d i a Nilaparvata sp . : i n t h e P h i i i p p i n e s Nil aparvata lugen s Sta l . D I S T R I B U T I O N : O r i e n t a l reg i o n : I N D I A : H o l a l u ( Ma n d y a ) , 8M ! OL ! C u t t a c k ( O r i ss a ) , BM ! P H I L I P P I N E S : K Y ! S h i s h i , KY !

Los B a n o s ( L a g u n a ) , 8 M !

N C: A R C T I C S P E C I ES OF DI COND YLUS

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954 - M a l e g en i t a l i a of Dicondyl u s indi a n u s n . s p . from Ho l a l u ( I n d i a ) ( r i g h t h a l f removed )

Dicondyl u s american us ( R . C . L . P e r k i n s )

Haplogon a top u s amerl c a n u s R . C . L . P e r k i n s 1 905 : 40 . Haplogon a topus americ a n u s R . C . L . Perk i n s : M u esebeck a n d W a i k l e y 1 95 1 : 1 036 . IIaplogon a topus americ a n u s R . C . L . Perk i n s : K ro m b e i n 1 979 : 1 246 . Dicondyl u s americ a n u s ( R . C . L . P e rk i n s ) : G i r i a n d F re y t a g 1 98 2 : 1 21 .

F E MAL E : a p terou s ; l e n g t h 1 , 62-2 , 87 m m ; y e l l o w - t e s t aceou s ; h e a d w i t h v e r t e x b ro w n ; a n t e n n a l s e g m e n t s 3-9 a n d a b domen b ro w n j occ a s i on a l l y p ron o t u m a n d m e t a t ho r a x + p ro p o d e u rn b rown or b ro w n - b l ac k ; a n te n n a I seg men t s i n fo l l o w i n g p ro p or t i on s : 7 : 4 : 7 : 5 : 4 : 4 : 4 : 4 : 4 : 7 ; h e a d

Go n a topod i n a e

1 389

F i g . 955 - C h e l a o f Dicondyl u s americ a n u s ( R . C . l . P e rk i n s ) from Lex i n g ton ( K e n t uc k y )

exc a v a ted , s h i n y , s moot h , w e a k l y g r a n u l a t ed ; p ron o t u m s h i n y , smoot h , w i t ho u t sc u l p t u re , n o t crossed b y a t r a n s ver s a l i m p res s i o n ; scu t u m s h i n y , w i t h n u merous l o n g i t u d i n a l k ee l s ; sc u te l l u m s h i n y , smoo t h , w i t ho u t scu l p t u r e ; m e t a t ho r a x + p ropod e u m s h i n y , g ra n u l a t ed , w i t h n u merous t r a n s v e rs a l s t r i a e on pos t e r i or s u rface ; p l e u r a t r a n svers e l y s t r i a t e ; m eso-me t a p l e u r a l s u t u re o b so l et e ; fore t a r s a l se g m e n t s i n fol l ow i n g p ropo r t i o n s : 1 1 : 3 : 4 : B : 1 4 ; en l a r ged c l a w ( F i g . 955 ) w i t h s u b a p i c a l t oo t h a n d 4-7 l am e l l ae ; s e g m e n t 5 o f fron t t a rs u s ( F i g . 955 ) w i t h t w o rows of 5- 1 0 l a me l l a e ; a p e x w i t h a g ro u p of a t l e a s t 5 - 6 l a me l l ae ; m a x i l l a r y p a l p i w i t h 2-3 seg men t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : f u l l y w i n ged ; l e n g t h 2 , 0(;- 2 , 56 m m ; b l ac k ;· m a n d i b l es a n d t eg u l ae testaceo u s ; a n te n n a e b row n ; l eg s t e s t aceo u s , w i t h p a r t of cox a e b l ac k ; c l u bs of femora b ro w n i s h ; a n te n n a I s e g me n t s i n fo l l ow i n g p ro p o r t i o n s : 5 : 4 : 6 : 7 : 7 : B : 7 : 6 : 6 : B ; h e a d s h i n y , smoo t h , w i t ho u t scu l p t u re or w e a k l y g r a n u l a ted , m a i n l y on t e m p l es ; P O L = 1 0 ; O l = 4; O O l = 3 ; b re a d t h of t h e oce l l i : 3 ; occ i p i t a l c a r i n a i n v i s i b l e ; sc u t u m s h i n y , h a i ry , smoot h , w i t ho u t scu l p t u re or weak l y g ra n u l a ted ; n ot a u l i ces comp l e t e , poste r i or l y j o i n t ; sc u te l l u m a n d m e t a n o t u m s moo t h , s h i n y , w i t ho u t sc u l p t u r e ; p ropod e u m ret i c u l a t e r u gose ; fore-w i n g h y a l i n e , w i t ho u t d a rk t r a n sv e rs a l b a n d s ; d i s t a l p a r t of ra d i a l ve i n m u c h l o n g e r t h a n p rox i m a l p a r t ; dors a l p rocess o f gonoforceps ( F i g . 9 5 6 ) l on g , s l e n d e r , s i n u a t e ,

1 390

Mem . A m e r .

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1 984

F i g . 956 - M a l e g e n i t a l i a of Dicondyl u s amer'ic a n u s ( R . C . L . Perk i n s ) f rom Lex i n g to n ( Ke n t u ck y ) ( r i g h t h a l f removed ) .

w i t h a p e x rou n d e d ; m a x i I l a r y p a l p i w i t h 2 s e g me n t s ; l ab i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . LOCUS TYP I C U S : C o l u m b u s ( O h i o , U . S . A . ) T Y P I CAL MAT E R I AL : l ec t o t y p e F ! a n d 2 p a r a l ec t o t y pe s F F ! ( de s i g n a t e d b y O l m i 1 982 ) i n B ; 1 p a r a l ec t o t y p e F ! i n BM . HOST S : i n U . S . A . Liburnia s p . ( R . C . L . P e r k i n s 1 906c ) , Delp h a codes campestris V a n D . a n d l u t u len ta Van D . ( T hompson 1 954 ) . D I STR I B U T I ON : N e a rc t i c reg i on : A LASKA : A n to n L a rsen B a y ( Ko d i a k ) , OT ! CANAD A : O t t a w a ( O n ! . ) , OT ! U . S . A . : C o l u m b u s ( O h i o ) , B ! BM ! O h i o ( G i r i a n d F re y t a g 1 982 ) ; Lex i n g t o n ( K y . ) , OL ! K E ! Sape l o I s . ( G a . ) , WA ! OL !

N E O T R O P I C S P E C I ES OF D I COND YUfS

O n e spec i e s i s k n ow n .

Dicondylus nigrithorax ( O g l ob l i n ) n . com b .

Idologon a top u s n igrithorax O g l ob l i n 1 95 3 :

1 1 2.

Go n a topod i n a e

F i g . 957

-

1 39 1

C h e l a o f Dicondy l u s n igrithorax ( O g l o::> i i n ) ( p a r a t y p e )

F E M AL E : a p t e rou s ; l e n g t h 4 , 75 m m ; h e a d b ro w n , w i t h a n t e r i o r h a l f of fron s , c I y p e u s a n d m a n d i b l e s t e s t aceo u s ; an t e n n a e test aceou s ; t h o r a x a n d p ropodeum b ro w n ; p e t i o l e b row n ; a b domen a n d l e g s test aceou s ; a n t en n a l segme n t s i n fo l l o w i n g p roport i o n s : 1 2 : 1 0 : 26 : 1 5 : 1 2 : 1 1 : 1 0 : 1 0 : 9 : 1 5 ; h e a d d u l l , g ra n u l a ted , w i t h v e r t e x excav a t e d ; frons w i t h t r ack of i ncom p l e t e fron t a l l i n e ; POL = 3 ; OL = 4 , 5 ; OOL = 1 3 ; occ i p i t a l c a r i n a a l mos t f u l l y i n v i s i b l e , o n l y v i s i b l e n e a r t he pos t e r i o r oce l l i ; t e m p l es d i s t i n c t ; p r o n o t u m w e a k l y t r a n sverse l y i m p ressed ; co l l a r d u l l , g ra n u l a ted ; d i sc s h i n y , w e a k l y p u n c t a te ; s c u t u m d u l l , ru gose ; sc u t e l l u m s hort , d u l l ; m e t a no t u m d u l l , n o t ho l l ow b e h i n d t h e s c u t e l l u m , ru gose ; m e t a t h o r a x + p ropodeum d u l l , g r a n u l a t ed ; p l e u r a , a n t e r i o r a n d pos t e r i o r s u rface of m e t a t ho r a x + p ropod e u m t r a n sverse l y s t r i a t e ; meso-m e t a p l e u ra l s u t u r e obso l e t e ; fore t a rs a l seg men t s i n fo l l o w i n g p ropo r t i o n s : 2 4 : 5 : 7 :

1 392

M e m . A me r .

En!.

I n s! . ,

no.

37 ,

1 984

1 8 : 33 ; en l a r g e d c l a w ( F i g . 957 ) w i t h s u b a p i c a l too t h a n d 6 l a m e l l ae ; seg men t 5 o f f r o n t t a rs u s ( F i g . 957 ) w i t h t w o rows o f ne a r l y 9 - 1 1 l a m e l l ae ; a p e x w i t h a g r o u p of n e a r l y 7-8 l a m e l l ae ; seg men t s 2 a n d 3 o f f r o n t t a rs u s p ro d uced i n t o a hook ; m a x i l l a r y p a l p i w i t h 3 s e g me n t s ; l ab i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M AL E : u n k n o w n L O C U S TYP I C U S : M a s a f u e r a I s l a n d ( J . F e r n a n dez I s l a n d s , C h i l e ) TYP I CAL Iv:AT E R I A L : ho l o t y p e F p rob a b l y l os t ( i n SW o n l y a s l i de o f t h e m o u t h - p a r t s i s v i s i b l e ! ) ; 1 p a r a t y p e F ' i n PT . HOSTS : u n k n o w n D I ST R I B UT I O N : Neo t rop i c reg i o n : C H I L E : M a s a f u e r a I s l a n d ( J . F e r n a n de z I s l a n d s ) , SW ! PT !

AUSTRAL I A N S P E C I E S OF D I COND YL US

Seven s p ec i es are k n ow n .

KEY TO T H E SP E C I E S

F E MA L E S

Me t a n o t u m w i t h s i d e s f' o u n d e d ( F i g . 9 5 8 A ) . . . . . . . . . . . . . . . Me t a n o t u m w i t h s i d e s p r o t r u d i n g ; p r o t r u s i o n s r ou n d ed o r 2

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. . . . 1 . perkinsi ( A s h m e a d ) t a r s u s l on g e r t h a n s eg men t rufus ( F o u t s ) . . • • • . . . . 2 . s cu l p t u r ed b y i r r e g u l a r , l o n g i P r on o t um d u l l . . . . . . . . . . . . . . . . . . . . . . 3 . dui]iu5 n . s p . P r o n o t u m s h i n y , smo o t h , w i t h o u t s cu l p t u r e . . . . Me t a n o t u m w i t h I a t e r a I p r o t I' U S i on s r o u n d ed ( F Me t a n o t u m w i t h l a t e r a l p r o t r u s i on s p o i n t ed ( F ·

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M e t a t h o ra x + p r op o d e u m d u l l , f u l l y s c u l p t u l' ed by s t r o n g t r a n s v e r s a l k e e l s . • . . . . 6 . ocean icus n . s p . Me t a t h o r a x + p r op od e u m s h i n y , w i t h a n t e r i o r s u r f a c e w i t h o u t s c u l p t u r e ; p l e u ra a n d p o s t e r i o r s u r f a c e t r a n sv e r s e l y s t r i a t e . . . . . . . . . . . . . . . . . . . . . . . 7 . a lp inus ( G o u r l a y ) T h o r a x a n d p r op o d e u m r ed d i s h - t e s t a c e o u s . • • . . . • • . • • . . . . . . . . . . . . . . . . . . . . . . . 4 . ,'< iefferi ( R . C . L . P e rk i n s ) T h o r ax a n d p r op o d e u m b l a c k . • • • • • • • • • • • . . . . . . . . . . . . . . . . . . . . . . 5 . prim itivus n . s p . ·

Go n a t opod i n ae

1 39 3

F i g . 9 5 8 - F e m a l e of Dicondylus perkinsi ( As h m e a d ) { A ) : B : h ea d ; C : t h o r a x a n d p ropodeum ( l a t e ra l l y v i ew ed ) ; D : fore t a rsa l seg m e n t s 1 -4 .

1 394

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MALES

N o t a u l i c e s j o i n t ; d o r s a l p r o c e s s o f g o n o f o r c ep s w i t h a p e x s e r ra t e ( F i g . 9 6 9 ) 7 . a lp in u s ( G o u r l a y ) N o t a u l i c e s s ep a ra t ed ; d o r sa l p r o c e s s o f g on o f o r c ep s w i t h a p ex n o t s e r ra t e ( F i g . 9 6 0 ) . 1 . perkinsi ( A s h me a d ) •

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T h e m a l es of D. primitivus, rufu s , a re u n k n ow n .

du b i u s ,

oceanic u s a n d kiefferi

1 . Dicon dy I u s perkinsi ( As h m e a d ) n . com b .

Gon a top u s perkin s i A s h m e a d 1 90 1 : 293 . Gon a top u s h a le a k a l a e A s h m e a d 1 90 1 : 293 ( n . s y n . ) ; locus typicus H a l e a k a l a ( M a u i , H a w a i i a n I s l a n d s ) . L a beo h a w a iiensis A s h m e a d 1 90 1 : 294 ( n . sy n . ) ; locus typicus Mo l o k a i Mts ( Ha w a i i a n I s l a n d s ) . Pseu dogo n a top u s perkinsi ( As h m e a d ) : R . C . L . Perk i n s 1 9 1 2 : 487 .

F E MA L E ( F i g . 958 A ) : a p t e ro u s ; l e n g t h 2 , 00-3 , 75 m m ; co l o u r v a r i a b l e ; re d d i s h - te s t a c eo u s , w i t h p e t i o l e b l ac k ; occ a s i on a l l y b row n i s h o r b row n b l ack ; h e a d ( F i g . 958 B ) exc a v a t e d , s h i n y , g r a n u l a ted ; p ron o t u m ( F i g . 958 C ) n o t crossed b y a s t ro n g t r a n s v e rs a l i m p res s i on or w e a k l y i mp ressed , s h i n y , w i t h o u t scu l p t u re ; scu t u m g r a n u l a ted ; m e t a t ho r a x + p ropodeum f u l l y t r a n sv e r se l y s t r i a t e ; segmen t 1 of fron t t a rs u s ( F i g . 958 D ) shorter t h a n segmen t 4 ( 35 : 38 ) ; en l a rged c l a w w i t h s u b a p i c a l tooth and 4-6 l a m e l l ae ; segmen t 5 of f r o n t t a rs u s w i t h t w o rows of 1 0-21 l a m e l l a e ; a p e x w i t h a g r o u p of 4-9 l a m e l l ae ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l ab i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M AL E : f u l l y w i n g e d ; l e n g t h 2 m m ; b ro w n-redd i s h , w i t h a b domen l i g h t ; a n te n n a e y e l l ow - b row n ; fron t p a r t o f verte x , c l y pe u s , m a n d i b l es a n d t e g u l a e y e l l o w ; l e g s ye l l o w , w i t h coxae b ro w n i s h ; a n ten n a l seg m e n t s i n fo l l o w i n g proport i o n s : 6 : 5 : 1 0 : 9 : 9 : 9 : 8 : 8 : 8 : 1 2 ; a n t e n n a I segme n t 3 more t h a n t h ree t i mes as

l o n g as b roa d ( 3 , 6 ) ;

hea d s h i n y ,

h a i ry ;

t e m p l es

d i s t i nc t ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 9 ; OL = 2 ; OOL = 1 ; b rea d t h of t h e oce l l i : 4 ; fron t a l l i n e b a sen t ; scu t u m s h i n y , g r a n u l a t ed ; n o t a u l i ces comp l e t e , poster i o r l y sep a ra ted ; m i n i m u m d i s t a nce b e t w een the n o t a u l i ces as l o n g a s b rea d t h of t h e oce l l i ; scu te l l u m a n d m e t a n o t u m smoo t h , s h i n y , w ea k l y p u n c t a te ; p ropodeum ret i c u l a te ru gose , w i t ho u t l on g i t u d i n a l kee l s ; fore-w i n g h y a l i n e , w i t ho u t d a r k t r a n svers a l b a n d s ; dors a l p rocess of g o n oforceps ( F i g . 9 5 9 ) s l en der , d i s t a l l y w e a k l y b ro a dened ; m a x i l l a ry p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 s e g me n ts ; t i b i a l s p u rs 1 , 1 , 2 .

Gon a toood i n a e

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1 395

( As h m e a d )

LOCUS T Y P I C U S : Mo l o k a i M t s ( H a w a i i a n I s l a n d s ) T Y P I CAL MAT E R I AL : h o l o t y p e F ! of G . perk i n s i i n B M ; ho l o t y p e F ! o f G . h a l e a k a l a e i n B,�I ; h o l o t y p e F ! of '- . h a l,'a iiensis i n B M . HOST S : i n t h e H a w a i i a n I s l a n d s f{eli s i a svezeyi ( B r i d we l l 1 920 ) , I l b urnia coprosmicola M u i r

( T hompson 1 954 ) , l l b urnia !w ae K i rk .

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a n d o t he r Delp h a cidae a l l i ed to L i b urnia ( R . C . L . P erk i n s 1 906c ) . PARAS I T E S : i n t h e H a w a i i a n I s l a n d s Hypergo n a topus h a w a iiensis ( R . C . L . Perk i n s ) , vulcan u s T i mb e r l ak e , brunneipes T i m b e r l a k e , flavipes T i m b e r l a k e ( T i m b e r l a k e 1 922 ) . D I ST R I B U T I O N : A u s t r a l i a n reg i o n : HAWA I I A N I SLAN D S : W a i a n a e ( O a h u ) , B !

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F i g . 9 6 1 - F em a l e of Dicondylus d u b i u s n . s p . ( t y p e ) : A : h e a d ( f ron t a l l y v i e w ed ) ; B : h e a d ( do r s a l l y v i e w ed ) ; C : meso t h o ra x , m e t a t h o r a x a n d p ropod e u m ( do r s a I I y v i e w e d ) ; 0 : t h o r a x a n d p ropode u m ( I a te r a l l y v i e w ed ) ;

E : fore t a rs a l segmen t s 1 -4 .

P o a moho ( O a h u ) , B ! M t . K a a l a ( O a h u ) , B ! O L ! O l y m p u s ( O a h u ) , WA ! K on a h u a h u i ( Oa h u ) , B ! WA ! M . T a n t a I u s ( O a h u ) , B ! O L ! P u u K a n e h o a ( O a h u ) , B! W a i a u ( O a h u ) , B! C a s t l e T r i a l ( O a hu ) , B! Ka umuohoua

1 398

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1 984

F i g . 962 - C h e l a of Dicondylus d u b i u s n . s p .

( type )

( O a h u ) , WA ! B ! W a i k a ne ( O a h u ) , B ! L a n i h u l i ( O a h u ) , B ! P u u K a l en a ( O a h u ) , B ! P a l o l o ( O a h u ) , B ! Hono l u l u ( O a h u ) , B ! K a i n a l u ( Mo l ok a i ) , B ! W a i ko l u ( Mo l o k a i ) , B ! K a m i l o l o a ( M o l ok a i ) , B ! M o l ok a i M t s ( M o l o k a i ) , B M ! K a l a u p a p a Looko u t ( M o l ok a i ) , B ! K a w e l a ( Mo l ok a i ) , B ! H a e l a a u ( M a u i ) , B ! P u u N i u n i a u ( M a u i ) , B ! K a u a i k i n a n a ( K a u a i ) , B ! Kokee ( K a u a i ) , B ! Moh i h i R i d g e ( K a u a i ) , B ! M t s . W a i me a ( K a u a i ) , B ! H a l em a n u ( K a u a i ) , B ! M t . H u a l a l a i ( H a w a i i ) , B ! H a w a i i N a t i o n a l P a rk ( H a w a i i ) , C A l H u m u u l a ( Hawa i i ) , B ! U-Hamak ua ( Hawa i i ) , B ! K i l a uea ( H aw a i i ) , B ! O l aa ( H a w a i i ) , B ! L a n a i h a l e ( L a n a i ) , B ! H a l ep a a k a i ( L a n a i ) , B !

2 . Dicondy lu s rufus ( F o u t s ) n . com b .

Pseu dogon a top u s rufus F o u t s 1 935 : 1 58 . Pseudogona top u s rugo s u s F o u t s 1 935 : 1 59 ( n . s y n . ) ; locus typicus Ooumu ( M a rq ue s a s l s i a n d s ) .

F E MAL E : a p terou s ; l e n g t h 4 , 2 m m ; red d i s h- t e s t aceo u s , w i t h pe t i o l e b l ac k ; h e a d exca v a ted , s h i n y , g ra n u l a te d ; p ron o t u m s h i n y , w ea k l y i m p ressed b y a t r a n s vers a l f u rrow , w i t ho u t sc u l p t u re ; sc u t u m g ra n u l a ted ; m e t a t ho r a x + p ropode u m f u l l y t r a n s verse l y s t r i a t e ; m e t a n o t u m w i t h s i des roun ded ; meso-me t a p l e u ra l s u t u re o b so l e t e ; seg men t 1 of front t a rs u s l on g er t h a n segmen t 4 ( 38 : 35 ) ; en l a rged c l a w w i t h s u b a p i c a l too t h a n d 6 l a me l l ae ; segmen t 5 o f fron t t a r s u s w i t h t w o rows o f 1 9- 2 1 l a m e l l ae ; apex w i t h a g ro u p of 8-9 l am e l l a e ;

m a x i l l a r y p a l p i w i t h 4 segme n t s ;

l ab i a l

pa l p i w i th

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1 399

s p u rs 1 , 0 , 1 .

MALE : u n k no w n LOCUS TYP I C U S : T e i v i p a k e k a ( N u k u H i v a , M a r q u e s a s I s l a n d s ) TYP I C A L MAT E R I AL : h o l o t y p e F ! o f Ps . rufus i n B ; h o l o t y p e F ! o f Ps . rugos u s i n B . HOST S : u n k n o w n D I ST R I B U T I O N : A u s t r a l i a n reg i o n : MARQU E SAS I SLM, D S : T e i v i p a k e k a ( N u k u H i v a ) , B ! Oo u m u ( N u k u H i v a ) , B '

3 . Dicondylus du b i u s n .

sp.

F E M,<\L E : ap terou s ; l e n g t h 5 , 3 1 m m ; red d i s h - t e s t a c eo u s , w i t h a n t e n n a I segme n t s 5-9 b row n ; pet i o l e b l ac k ; a b domen b row n ; h e a d ( F i g . 9 6 1 A , B ) d u l l , exc a v a t e d , g ra n u l a t e d ; pron o t u m ( F i g . 9 6 1 D ) d u l l , wea k l y i m p ressed b y a t r a n sversa I f u rrow , w i t h n u merous i rreg u I a r k e e l s ; s c u t u m s h i n y , w i t h some l o n g i t u d i n a l k ee l s ; m e t a t ho r a x + p ropodeum ( F i g . 9 6 1 C ) a n d m e t a no t u m ret i c u l a t e ru gose ; m e t a n o t u m w i t h s i d es p r o t r u d i n g ; p ro t r u s i o n s p o i n ted ; meso-me t a p l e u r a l s u t u re o b so l e te ; segmen t 1 of f ro n t t a r s u s l on g e r t h a n segmen t 4 ( 26 : 2 1 ) ( F i g . 9 6 1 E ) ; en l a rged c l a w ( F i g . 962 ) w i t h s u b a p i c a l t oo t h a n d 7 l a m e l l ae ; seg m e n t 5 of fron t t a rs u s ( F i g . 962 ) w i t h t w o row s of 2 3 l a m e l l a e ; apex w i t h a g r o u p of 1 8 l a m e l l ae ; m a x i l l a r y p a l p i w i t h 4 seg men t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MA L E : u n k no w n LOCUS T Y P I C U S : N a n d a r i v a t u ( V i t i Lev u , F i j i I s l a n d s ) HOST S : u n k n o w n D I S T R I B U T I O N : A u s t r2 , i a n reg i on : F I J I I SLAN D S : N a n d a r i v a t u ( V i t i Le v u ) , B!

4 . Dicondyl u s k iefferi ( R . C . L . Perk i n s )

n . com b .

Pseu dogon a topus kiefferi R . C . L . P e r k i n s 1 906a : 487 .

FEMALE :

a p terou s ;

l e n g t h 3 , 70-4 , 20 m m ;

red d i s h - tes t aceo u s ,

w i t h antenna I

s e g m e n t s 6-9 b ro w n i s h ; o t her a n t e n n a l s e g me n t s y e l l ow ; head b ro w n b l ac k ; m a n d i b l e s , c l y pe u s a n d f ro n t p a r t of v e r t e x ( more a l on g orb i t s ) y e l l o w ; p e t i o l e b l ac k ; a b domen b ro w n - b l ac k ; body s h i n y ; head exc a v a ted , g ra n u l a te d ; p ro n o t u m s h i n y , w i t ho u t sc u l p t u re , w e a k l y i m pressed by a t r a n s v e rs a l f u rrow ; scu t u m sh i n y , w i t hou t s c u l p t u re , w i t h some l o n g i t u d i n a l k ee l s ; m e t a n o t u m w i t h s i des p ro t r u d i n g ; pro t r u s i o n s p o i n ted ; m e t a t horax + p ro p o d e u m q u i te i r regu l a r l y s t r i a t e ; segmen t 1 of f ro n t t a r s u s l o n g e r t h a n segmen t 4 ( 49 : 36 ) ; en l a r g e d c l a w ( F i g . 963 ) w i t h

1 400

M e m . Amer . E n ! .

I nst. ,

no.

37 ,

1 984

F i g . 963 - C he l a of Dicondyl u s k iefferi ( R . C . L . Perk i n s ) ( t y p e )

s u ba p i c a l too t h a n d 7 l am e l l ae ; s e g m e n t 5 o f fro n t t ar s u s ( F i g . 963 ) w i t h t wo row s of 1 7 l a m e l l ae ; a p e x w i t h a g ro u p of 1 0 l a m e l l ae ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : S u v a ( V i t i Lev u , F i j i I s l a n d s ) TYP I C A L MAT E R I AL : l ec t o t y p e F ! a n d 1 p a r a l ec t o t y p e F ! ( de s i g n a t ed b y

G o n a topod i nae

1 40 1

F i g . 964 - Fem a l e of Dicondylus primitivus Il . s p . ( t y p e ) : A : meso t ho ra x , m e t a t h o r a x a n d p ropod e u m ( dorsa I I y v i ew ed ) ; B : h e a d ( dorsa I I y v i ewed ) ; C : thorax a n d p ropodeum ( l a tera l l y v i ewed ) ; D : head ( front a l l y v i ew ed ) .

O l m i 1 982 ) i n B . HOS T S : i n F i j i I s l a n d s p e r h a p s Va n u a vitien s i s ( R . C . L . Perk i n s 1 906a ) . D I ST R I BU T I O N : A u s t r a l i a n r eg i on : F I J I I SLAN D S : S u v a ( V i t i L e v u ) , B !

5 . Dicondylus primitivus n . s p .

F E M A L E : a p t erou s ; l en g t h 3 , 75 m m ; b l ac k , w i t h m a n d i b l es , c l y p e u s , f ron t p a r t of vertex a n d a n te n n ae ye l l ow ; l e g s b rown , w i t h p a r t of coxae , troch a n ters , s t a l k s of femora , t a r s i y e l l ow ; head ( F i g . 964 B , D ) excava ted , s h i n y , smoot h , w i t h o u t scu l p t u re ; p ro n o t u m ( F i g . 964 C ) s h i n y , smoo t h , w i t ho u t scu l p t u re , w ea k l y i m p ressed b y a t r a n s v erse f u r row ; sc u t u m s h i n y , w i t h some l on g i t u d i n a l kee l s ; met a n o t u m re t i c u l a t e r u gose , w i t h s i des p ro t ru d i n g ; p r o t r u s i ons p o i n ted ; meta t ho r a x + p ro p o d e u m ( F i g . 964 A ) ret i c u l a t e ru gose ; a n te r i o r s u rface smoo t h , w i t h a l on g i t u d i n a l f u rrow ; poster i or s u rface ret i c u l a t e r u g ose ; meso-m e t a p l eu r a l s u t u re d i s t i nc t ; segmen t 1 o f f r o n t t a r s u s l on ge r t h a n segme n t 4 ( 1 7 : 1 5 ) ; e n l a rged c l a w ( F i g . 965 ) w i t h s u b ap i c a l too t h a n d 8 l am e l l ae ; seg m e n t 5 of f ron t t a r s u s ( F i g . 965 ) w i t h two rows of 1 7 l a me l l ae ; a p e x w i t h a g ro u p of 1 0 l am e l l ae ; m a x i I l a r y p a l p i w i t h 4 segme n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n

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L O C U S TYP I C U S : V i l a ( E f a te , Ne w H eb r i de s ) TYP I CAL M A T E R I AL : ho l o t y p e F ! i n B . HOST S : u n k n ow n D I ST R I B U T I O N : A u s t ra l i a n reg i o n : N E W H E B R I D E S : V i l a ( E f a t e ) , B ! ! !

6 . Dicon dylus oceanicus n .

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Mem . Amer . E n ! .

I ns t . ,

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1 984

spot o n v e r te x ; a n te n n ae t e s t aceou s ; p ropec t u s red d i s h ; p ron o t u m b ro w n , w i t h s i des re d d i s h ; sc u t u m brown ; m e t a t h o r a x + p ropodeum a n d s c u te l l u m b l ac k ; abdomen b ro w n ; l e g s test aceou s , w i th p a r t of coxae b ro w n ; a n ten n ae d i s t a l l y t h i ckened ; a n t e n n a I segments i n fo l l o w i n g p roport i on s : 1 0 : 9 : 3 1 : 2 3 : 1 4 : 1 0 : 1 0 : 8 : 8 : 1 1 ; head s h i n y , w e a k l y g r a n u l a ted ; f ron t a l l i ne i n com p l e te , s u r r o u n d e d by a l on g i t u d i n a l f u r ro w ; occ i p i t a l c a r i n a s hort , o n l y v i s i b l e o n the s i des of t h e pos ter i or ocel l i ; P O L = 1 ; O L = 2 ; OOL = 1 0 ; p ronot u m s h i n y , smoot h , w i t ho u t scu l p t u re , w e a k l y crossed b y a t r a n s vers a l i m p ress i o n ; scu t u m d u l l , ru gose , w i t h some l o n g i t u d i n a l k ee l s ; scu te l l u m d u l l , p u n c t a te ; met a n o t u m w i t h s i des p rot r u d i n g ; p ro t r u s i on s r o u n ded ; m e t a t ho r a x + p ropodeum f u I l y scu I p t u re d b y s t ron g tra n s v ersa l kee l s ; d i sc ret i c u l a t e r u gose ; meso-me t a p l eu r a l s u t u re d i s t i n c t , com p l ete , i r regu l a r ; fore t a rs a l segments i n f o l l o w i n g p roport i o n s : 24 : 4 : 7 : 2 1 : 34 ; e n l arged c l aw ( F i g . 966 ) w i t h s u b a p i ca l too t h a n d 1 2 l a m e l l ae ; s e g m e n t 5 o f front t a r s u s ( F i g . 966 ) w i t h t w o rows o f 39 l a me l l ae ; a pe x w i t h a g ro u p of n e a r l y 1 3 l ame l l ae ; m a x i l l a r y p a l p i w i t h 4 segmen ts ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : L o r d H o w e I s l a n d ( So l omon I s l a n d s ) TYP I CA L MAT E R I AL : ho l o t y pe F ! i n A D . HOST S : u n k n o w n D I ST R I B U T I ON : A u s t r a l i a n reg i o n : AD !

SOLOMON

I SLAN D S : L o r d How e

7 . Dicondyl u s alpin u s ( Go u r l a y )

I s l an d ,

n . com b .

Neogon a top u s a lpinus Gou r l ay 1 954 : 4 .

F E MA L E ( F i g . 967 A ) : a p terou s ; l e n g t h 2 , 1 8-4 , 06 m m ; h e a d t e s t aceous redd i s h , occa s i o n a l l y d a r k ; a n t e n n ae f u l l y test aceo u s or brow n , w i t h segments 1 -2 a n d 1 0 tes taceo u s ; occ as i o n a l l y i n a d d i t i on segmen t 3 f u l l y o r p a r t I y test aceo u s ; t h o r a x a n d p ropodeum f u I I y redd i s h-test aceo u s ; occa s i o n a l l y d a rkened ; abdomen red d i s h-testaceous or b row n ; l eg s tes taceou s , occ a s i o n a l l y w i t h coxae a n d c l u b s of femora d a r k e n ed ; h e a d ( F i g . 967 B ) e xc a v a ted , s h i n y , s m oo t h , w i t h o u t scu l p t u re , h a i r l ess ; p ron o t u m ( F i g . 967 C ) not crossed by a t r a n s versa l i m p res s i o n or w e a k l y i m p ressed , smoo t h , w i t ho u t scu l p t u re , h a i r l es s ; sc u t u m s h i n y , w i t h w e a k l on g i t u d i n a l k ee l s ; met a n o t u m s m oo t h , not t r a n s v e r se l y s t r i a te o r weak l y s t r i a t e , w i t h s i des p ro t r u d i n g ; p ro t r u s i o n s p o i n te d ; met a t ho r a x + p ropodeum smoo t h , w i t ho u t sc u l p t u r e , w i t h p l eu ra a n d p o s te r i or s u rf ace t r a n s v e r se l y s t r i a t e ; occ a s i on a l l y p l e u r a a n d post e r i or s u rface not t r a n s v erse l y s t r i a te ; meso-me t a p l eu r a l s u t u re obso l e te ; fore t a rs a l segme n t s i n fo l l o w i n g p roport i on s : 1 3 : 2 : 3 : 8 : 1 5 ; e n l a rged c l aw ( F i g . 968 ) w i t h s u b a p i c a l toot h a n d 4-5 l a me l l ae ; seg m e n t 5 o f fron t t a rs u s

G o n a topod i n a e

F i g . 967

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1 405

F em a l e of Dicondyl u s a lp n u s ( Go u r l a y ) ( A ) ; B: h e a d ( f ron t a l l y v i ewed ) ; C : t h o r a x a n d p ropode u m ( l a te ra l l y v i ew ed ) .

-

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1 406

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1 984

from Beeby ' s K n o b ( New

1 407

Go n a t o pod i n a e

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from Nervou s

( F i g . 968 ) w i t h t w o rows o f 8-9 l a me l l a e ; a p e x w i t h a g ro u p of 5-9 l a m e l l a e ; m a x i I l a r y a n d l a b i a l p a l p i w i t h 2 seg men t s ; t i b i a l sp u rs 1 , 0 , 1 . MALE : f u l l y w i n ged ; l e n g t h 2 , 00-3 , 43 m m ; h e a d b l ac k or b row n ; occa s i o n a l l y redd i s h ; a n t e n n ae b row n , occ a s i o n a l l y t e s t aceo u s ; t h o r a x a n d propod e u m b l a ck , occ a s i o n a l l y red d i s h ; a b domen b l a ck or b row n , occa s i on a l l y redd i s h ; l e g s t e s t aceo u s , w i t h h i n d coxae d a r k e n e d ; a n t en n a l seg men t s i n fo l l ow i n g p ropor t i o n s : 8 : 6 : 1 2 : 1 2 : 1 2 : 1 2 : 1 1 : 1 1 : 1 0 : 1 2 ; a n te n n a l se g m e n t 3 more t h a n t h ree t i mes a s l o n g a s b ro a d ( 4 , 8 ) ; h e a d s h i n y , h a i r y , s t ro n g l y p u n c t a t e , occ a s i o n a l l y smoo t h , w i t h o u t s c u l p t u re ; temp l es d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 7 ; OL = 4 ; OOl = 4 ; b re a d t h o f t h e oce l l i : 3 ; scu t u m s h i n y , h a i r y , g r a n u l a ted ; n o t a u l i ces comp l e t e , poster i or l y jo i n t ; sc u t e l l u m and m e t a no t u m s h i n y , smoot h , w e a k l y p u n c t a t e ; p ropod e u m f u l l y ret i c u l a t e r u gose ; dors a l p rocess o f gon oforceps ( F i g , 969 ) l o n g , p a r a l l e l to t h e gonofo rceps ; a p i c a l reg i o n of dors a l p rocess w i t h n u m e ro u s h y a l i ne po i n t s and w i t h s e r r a t e m a r g i n ; m a x i I l a r y a n d l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . lOCUS T Y P I C U S : Beeb y ' s K n o b ( Ne w Z e a l a n d ) T Y P I CA L MAT E R I A L : ho l o t y p e F a n d 3 p a r a t y p es F F ! i n N Z . HOST S : i n New Z ea l a n d Nil a p a r v a t a myers i .

1 408

Mem . Amer . E n ! .

I ns t . ,

n o . 37 ,

1 984

D I ST R I BU T I O N : A u s t r a l i a n reg ion : NEW Z EALAN D : A p p l eb y ( Ne l son ) , N Z ! Beeby ' s K n o b ( Ne l son ) , N Z ! W a t t l e B a y ( Au ck l a n d ) , O L ! N Z ! L y n f i e l d ( W a t t l e B a y , A u ck l a n d ) , N Z ! M a k e t u S t re a m ( A u c k l a n d ) , N Z ! T i t i ra n g i ( A u ck l a n d ) , O L ! P a n dora ( Nor t h l a n d ) , N Z ! U n u w h ao ( No r t h l a n d ) , N Z ! R i b bon wood ( M i d C a n te r b u r y ) , N Z ! Cra i g i e b u rn Forest ( Nervou s K n ob , C a n te r b u r y ) , N Z ! L i n d i s P a s s ( C e n t ra l O t a go ) , N Z ! S t e p h e n s I s l a n d ( M a r l borou g h SOU Cl CS ) , N Z !

1 0 . Haplogon a top u s R . C . L .

Mid

Perk i ns

Haplogon a top u s R . C . L . P e r k i n s 1 905 : 39 . Monogo n a top u s R i c h a r d s 1 939 : 200 ( s y n . p roposed by O l m i a n d C u rrado 1 979b ) ; t y p e spec i es Mon ogon a top u s oratorius ( We s twood 1 833 ) , mono t y p i c . Congodryin u s Beno i t 1 950 : 223 ( n . sy n . ) ; t y p e spec i es Congodryin u s k a tangae Ben o i t 1 950 , or i g . d es i g .

T Y P E S P E C I E S : Haplogon a topus apica lis R . C . L .

Perk i n s 1 905 , o r i g . d es i g .

F E MA L E ( F i g s 970 , 975 ) : a p terou s ; m a x i l l a r y p a l p i w i t h 2 segme n t s ; l a b i a l p a l p i w i t h 1 segmen t ; prono t u m ( F i g . 975 ) not crossed b y a t r a n s v e rs a l i m p ress i o n or w e a k l y i mp ressed ; e n l a rged c l a w ( F i g . 97 1 ) w i t h s u b a p i c a l too t h a n d l a me l l ae ; t i b i a l s p u rs 1 , 0 , 1 . MALE ( F i g s 972 , 973 ) : f u l l y w i n ged ; m a x i l l a r y p a l p i w i t h 2 s e g me n t s ; I a b i a I p a I p i w i t h 1 seg m e n t ; temp l es d i st i n c t ; occ i p u t con c a v e ; occ i p i t a l c a r i n a i n v i s i b l e o r on l y v i s i b l e b e h i n d t h e oce l l i o r o n t he s i des o f t h e poster i o r oce l l i , b u t n ev e r reac h i n g temp l es ; t i b i a l s p u r s 1 , 1 , 2 . D I ST R I BU T I O N : P a l a e a rc t i c , E t h i op i a n , O r i e n t a l , Neotrop i c , A u s t r a l i a n HOST S : Delph a c idae SPEC I ES : 7

PALAEAR C T I C S P E C I E S OF HA PLOGONA TOPUS

T h ree spec i e s a re k n o w n .

K E Y TO T H E S P E C I E S

FEMALES

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D o r sa l p r o c e s s of g on o f o r c ep s l on g , s l en d e r , w i t h o u t a p i c a l , s e r ra t e ma r g i n ( F i g . 9 7 4 A ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 . ora torius ( We s t wood ) D o r s a l p r o c e s s o f g on o f o r c e p s w i t h a p i c a l , s e r ra t e ma r g i n (F i gs 977 , 980 ) 2 D o r sa l p r o c e s s o f g o n o f o r c e p s d i s t a l l y b r o a d en ed ( F i g . 9 8 0 ) . . . . . . . . . . . . . 1 . apic a l is R . C . L . P e rk i n s ( s e e O r i e n t a l sp e c i e s ) D o r sa l p r o c e s s o f g on o f o r c ep s n o t d i s t a l l y b r o a d en ed ( F i g . 9 7 7 ) . . . . . . . . . . 3 . a tr a t u s E s a k i a n d H a s h i mo t o •

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1 . Haplogona top u s apic a l is R . C . L .

P e rk i n s

( see O r i en t a l spec i e s )

2 . Hap logon a top u s oratorius

( We s t wood )

Gona top u s ora torius W e s t w oo d 1 833b : 496 . Gon a top u s mayeti K i effer i n K i effer et M a rs h a l l 1 905 : 1 03 ( s y n . p roposed b y A b d u I - N o u r 1 974 ) ; locus typicus A u s t r i a . Dicondyl u s ora tori u s ( We s t w oo d ) : K i effer 1 907b : 2 2 . Gon a top u s oratorius W e s t wood : R i c h a r d s 1 935 : 1 45 . Monogo n a top u s ora torius ( We s t wood ) : R i c h a r d s 1 939 : 200 . Monogona top u s ora torius ( We s t wood ) : R a a t i k a i n en 1 96 1 : 1 3 1 . Monogo n a top u s ora torius ( We s t wood ) : C u rrado a n d 0 I m i 1 972 : 1 43 . Monogo n a top u s ora torius ( We s t wood ) : A b d u l -N o u r 1 974 : 1 0 1 5 . Haplogo n a top u s ora torius ( We s t wood ) : O l m i a n d C u rr a d o 1 97 9 b : 39 .

F E MA L E ( F i g . 970 ) : a p t e rou s ; l en g t h 2-3 m m ; h e a d b ro w n or b l a c k , w i t h m a n d i b l es , c l y pe u s a n d f ro n t p a r t of vertex y e l l ow ; a n te n n a e b row n , w i t h s e g m e n t s 1 -2 a n d 1 0 y e l l o w ; t ho r a x a n d p ropodeum red d i s h ; p e t i o l e a n d

1 T h e fem a l es o f H. oratorius a n d a tr a t u s a re n o t i d en t i f i a b l e ; t h e m a l e s , on t h e con t r a r y , a re e a s i l y recog n i za b l e for t he d i ffere n t p rocesses o f t h e gonoforcep s .

Mem . A m e r . E n ! .

1410

I ns! . ,

no.

37 ,

1 984

F i g . 970 - F e m a l e of lIaplogo n a top u s ora tori u s ( Westwood )

a b domen b l ac k ;

l e g s y e l l ow - t e s t a ceou s ; occ a s i o n a l l y bod y f u l l y b l ac k ;

h e a d excav a te d , s h i n y , w e a k l y g ra n u l a ted ; p ronot u m not crossed b y a t r a n s v e rs a l i m p ress i o n , s h i n y , smoo t h , h a i r l es s , w i t ho u t scu l p t u re ; scu t u m

Gon a topod i n a e

F i g . 971

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1 41 1

( Vv e s t w ood ) from C a s t e l l a l fero

( I taly )

d u l l , h a i r l e s s , w i t h some l o n g i t u d i n a l kee l s ; m e t a n o t u m , p l e u ra a n d pos t e r i or s u rf ace o f p ropode u m t r a n s v erse l y s t r i a t e ; m e t a t h o r a x + p ropode u m w i t ho u t scu I p t u re a m o n g s t r i ae a n d on a n te r i o r s u rface ; m e so m e t a p l e u r a l s u t u re obso l e t e ; fore t a rs a l s e g me n t s i n fo l l ow i n g p roport i o n s : 1 3 : 2 ; 3 ; 8 ; 1 3 ; e n l a rged c l a w ( F i g . 9 7 1 ) w i t h s u b a p i c a l too t h a n d 4-5 l a me l l ae ; s e g m e n t 5 of fron t t a rs u s ( F i g . 9 7 1 ) w i t h two ro w s of 6- 1 2 l am e l l ae ; a p e x w i t h a g r o u p of 3-9 l a m e l l ae ; m a x i l l a r y p a l p i w i t h 2 segmen t s ; l a b i a l p a l p i w i t h 1 seg men t ; t i b i a l s p u rs 1 , 0 , 1 . MALE ( F i g s 9 7 2 , 97 3 ) : f u l l y w i n g e d ; l e n g t h 2 , 1 8-2 , 40 m m ; b l ac k , w i t h a n t en n ae b ro w n ; m a n d i b l e s y e l l ow - b row n ; l e g s b row n , w i t h t a r s i l i g h t ;

1412

Mem . A m e r . E n ! .

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972 - M a l e of Haplo go n a top u s ora tori us

head

( We s t w ood ) f r o m T o n co ( I t a l y )

s h i n y , h a i r y , g r a n u l a t e d ; f ro n t a l l i n e a b se n t ; t e m p l es d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e ; s c u t u m d u l l , g r a n u l a ted ; n o t a u l i ce s d i s t i nc t , com p l e t e , s t ron g l y c o n v e r g i n g , b u t r u n n i n g p a r a l l e l n e a r pos t e r i o r m a r g i n o f sc u t u m ; s c u t e l l u m a n d met a n o t u m d u l l , h a i ry , weak l y p u n c t a t e ; p ropodeum r e t i c u l a t e ru gose , w i t ho u t kee l s ; fore- w i n g h y a l i n e , w i t ho u t d a rk t r a n sversa l b a n d s ; do" sa l p rocess of g o n oforceps ( F i g . 974 ) l o n g , s l en d e r , n o t d i s t a l l y b ro a d en e d ; m a x i I l a r y p a l p i w i t h 2 segmen t s ; l a b i a l p a l p i w i t h 1 s e g m e n t ; t i b i a l s p u rs 1 , 1 , 2 . LOCUS TYP I C U S : R i p l e y ( E n g l a n d ) TYP I CA L MAT E R I AL : ho l o t y p e F ! o f G . oratorius i n OX ; ho l o t y p e F ! of G .

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F i g . 973 - M a l e of Haplogon a top us oratorius

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i n MT .

HOSTS : i n I t a l y L a odelp h a x stria tel l u s ( F a l l en ) a n d Mega delphax sordidulus ( St a l ) ( C u rrado a n d O l m i 1 972 ) . D I ST R I B U T I O N : P a l a e a rc t i c re g i on : U . S . S . R . : K u o l emaj a r v i ( I s t h m u s K a re l i c u s ) , H E ! C Z E C HOSLOVAK I A : R u m u n s k o J u l om i t a , P G ! AUSTR I A , M T ! E N G LA N D : R i p l ey ( Su rrey ) , OX ! I T ALY : Port acomaro ( As t i ) , E N ! To nco ( As t i ) , E N ! C as te l l a l fero ( A s t i ) , E N ! San i co di A l f i a n o N a t t a ( A l es s a n d r i a ) , E N ! Pever a g no ( C u neo ) , SZ ! M a g n a n o ( Verce l l i ) , E N !

3 . Haplogona top u s a tratus E s a k i a n d H a s h i moto

Haplogo n a topus a tr a t u s E sa k i and Hash i moto 1 9 32 : 2 5 . Haplogon a top u s s u chovi Ponomarenko 1 970 : 429 ( n . s y n . ) ; locus typicus O m s k ( S i b i r i a , U. S. S. R . ) .

F E MALE ( F i g . 975 ) : a p t erou s ; l e n g t h 2 , 37-3 , 1 2 m m ; head tes taceo u s , w i t h vertex b l ack ; a n te n n ae b ro w n , w i t h segme n t s 1 -3 a n d 1 0 y e l l ow ; t ho r a x a n d p ropode u m test aceou s ; pet i o l e a n d a b domen b l ac k ; l eg s testaceou s ; h e a d exc a v a t e d , s h i n y , g r a n u l a t e d ; p ro n o t u m not c rossed b y a t r a nsversa l i mp ress i o n , s h i n y , w i t ho u t scu l p t u re ; s c u t u m s h i n y , w i t h w e a k l o n g i t u d i n a l k ee l s ; metano t u m , p l eu r a a n d p o s t e r i o r su rface of p ropodeu m t r a n s verse l y s t r i a t e ; met a t ho r a x + p ropodeum s h i n y , w i t h o u t scu l p t u re on a n ter i or s u rface a n d a m o n g s t r i a e ; s e g m e n t 1 o f fron t t a r s u s l o n ger t h a n segme n t 4 ( 1 2 : 8 ) ; e n l a rged c l aw ( F i g . 976) w i t h s u b a p i c a l

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974 - M a l e g en i t a l i a of [!a plogo n a top u s ora torius ( We s t w oo d ) from Tonco ( I t a l y ) ( A ) ; B : m a x i l l a ry p a l p u s ; C : l ab i a l p a l p u s .

t o o t h a n d 4-5 l a me l l ae ; seg m e n t 5 o f fron t t a rs u s ( F i g . 976 ) w i t h t wo rows of 7-9 l a m e l l ae ; a p e x w i t h a g ro u p of 3-4 l a me l l ae ; m a x i I l a r y p a l p i w i t h 2 segmen t s ; l a b i a l p a l p i w i t h 1 segmen t ; t i b i a l s p u rs 1 , 0 , 1 . MALE : f u l l y w i n ged ; l e n g t h 2 , 25-2 , 50 m m ; b l ack , w i t h a n te n n ae b row n ; l e gs , m a n d i b l e s a n d te g u l ae y e l l o w ; a n te n n a l s e g m e n t 3 more t h a n four t i mes a s long a s b ro a d ( 4 , 4 ) ; h e a d h a i ry , w e a k l y p u n c t a t e , s h i n y ; t e m p l es d i s t i n c t ; occ i p u t co n c a v e ; occ i p i t a l c a r i n a i n v i s i b l e ; fron t a l l i n e a b sen t ; sc u t u m g r a n u l a t e d ; n o t a u l i ce s d i s t i n c t , com p l e t e , poster i or l y jo i n t ; s c u te l l u m a n d m e t a n o t u m s h i n y , w e a k l y p u n c t a t e ; p ropod e u m

Go n a topod i n ae

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1415

F e m a l e of JJaplogon a top us a tr a t u s c s a k i Japan

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re t i c u l a te r u g ose , w i t h o u t kee l s ; dors a l p rocess o f gonoforcep s ( F i g . 977 ) l o n g , n o t d i s t a l l y b roadened , w i t h i n ner m a r g i n serra t e ; m a x i l l a ry p a l p i w i t h 2 segmen t s ; l a b i a l p a l p i w i t h 1 segmen t ; t i b i a l s p u rs 1 , 1 , 2 . L O C U S T Y P I C U S : U t s u nom i y a ( J a p a n ) TYP I CA L M A T E R I A L : l ec to t y p e F ! a n d 1 p a ra l ecto : y p e M ! ( h ereby des i g n a t e d ) of H. a tr a t u s in K Y ; ho l o t y p e F ! a n d 1 7 p a r a t y p e s ( 1 2 F F , 5 M M ) of H. s u c h o v i i n MO . HOST S : i n J a p a n L a odelp h a x stri a tellus ( F a l l e n ) ( E s a k i a n d H a s h i moto 1 93 2 ) D I ST R I B U T I O N : P a l a e a rc t i c reg i o n : JAPAN : K a m i -g u n ( Koc h i P r ef . ) , O L ! N I ! U t s u n om i y a ( Ho n s h u ) , K Y ! S u g a t a-cho ( M a t s u e C . , Sh i m a n e P ref . ) , OL ! •

M U ! F u k uo k a ( K y u s h u ) ( E s a k i a n d H a s h i moto 1 9 32 ) ; A m a k u s a ( T o m i ok a , K y u s h u ) { E s a k i a n d H a s h i moto 1 93 2 ) ; S O U T H K O R E A : N aj u , O L ! Su weon , O L ! U . S . S . R . : O m s k ( We s t e r n S i b i r i a ) , M O !

E T H I O P I AN S P E C I E S O F HA PLOGONA TOPUS

O n e spec i e s i s k n ow n .

Haplogon a top u s k a tangae ( Be n o i t )

Congodryinus ka tangae Ben o i t 1 950 : 223 .

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F E MA L E : u n k n o w n MALE : f u l l y w i n g e d ; l en g t h 2 , 3 1 m m ; h e a d b l ac k , w i t h m a n d i b l e s y e l l o w ; a n te n n ae brow n ; t h o r a x a n d p ropodeu m b l ac k ; a b domen b row n ; l eg s t e s t ac eo u s ; a n t en n a l segmen t s i n fo l l ow i n g p roport i o n s : 7 : 5 : 1 0 : 1 0 : 9 : 1 0 : 9 : 9 : 9 : 1 1 ; h e a d sh i n y , h a i r y , w ea k l y g r a n u l a t ed ; POL = 8 ; O l = 3 ; OOl = 5 ; b re a d t h o f t h e oce l l i : 3 , 5 ; occ i p i t a l c a r i n a i n v i s i b l e ; t e m p l es d i s t i nc t ; occ i p u t conca v e ; fron t a l l i ne a b sen t ; scu t u m h a i r y , s h i n y , w e a k l y g r a n u l a t ed ; n o t a u l i ce s com p l e t e , p o s t e r i or l y j o i n t ; sc u t e l l u m a n d m e t a n o t u m sh i n y , w e a k l y p u n c t a t e ; propod e u m re t i c u l a t e r u g o s e , w i t h o u t l o n g i t u d i n a l kee l s on p o s te r i or s u rface ; fore-w i n g h y a l i n e , w i t ho u t d a rk t ra n sver s a l b a n d s ; d i s t a l p a r t of r a d i a l v e i n m u c h l on g e r t h a n p rox i m a l p a r t ; dors a l p rocess of g o noforc e p s ( F i g . 978 ) l o n g , s l en de r , w i t h apex serr a t e , n o t d i s t a l l y b ro a d e n ed ; m a x i I l a ry p a l p i w i t h 2 segmen t s ; l a b i a l p a l p i w i t h 1 segmen t ; t i b i a l s p u rs 1 , 1 , 2 . l O C U S T Y P I C U S : E l i s a b e t h v i l l e ( Z a i re ) TYP I CAL MAT E R I AL : h o l o t y p e M ! i n T V . HOST S : u n k n o w n D I ST R I B U T I O N : E t h i o p i a n reg i o n : Z A I R E : E l i s a be t hv i l l e , TV !

OR I E N T A L S P E C I E S O F HAPWGONA TOPUS

G o n a topod i n ae

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Haplogon a top u s apicalis R . C . L . Perk i n s 1 90 5 : 39 . Haplogon a top u s moestus R . C . L . Perk i n s 1 90 5 : 40 ( n . s y n . ) ; locus typ icus C a i r n s ( A u s t r a l i a ) . Haplogon a topus brevicornis R . C . L . P e rk i n s 1 906a : 489 ( n . sy n . ) ; locus typ icus T weed R i ver ( Au s t r a l i a ) . Haplogon a topus orien talis R o h w e r 1 920 : 1 59 ( n . s y n . ) ; locus typicus B i l a s p u r ( I n d i a ) . Haplogon a top u s japon icus E s ak i a n d H a s h i moto 1 93 1 : 25 ( n . s y n . ) ; locus typ icus F u k u o k a ( J a p a n ) . l-/aplogon a top u s orienta lis R o h w e r : O t a k e , Som a s u n d a ra m a n d A b e y koon 1 97 6 : 289 . l-/ap logon a topus orienta lis R o h w e r : O t a k e 1 97 7 : 4 6 . Monogon a top u s orien t a l is ( Ro h w e r ) : C u rr a d o a n d O l m i 1 979 a : 1 8 1 . l-/aplogon a top u s apica lis R . C . L . P e rk i n s p a rtim : O l m i a n d C u rr a do 1 97 9 b : 42 . Haplogon a top u s orien t a li s R o h we r : O l m i a n d C u rrado 1 979b : 4 3 .

Gon a topo d i n a e

1419

F i g . 9 7 9 - C he l a o f Haplogo n a top u s a p i c a l i s R . C . L . P e r k i n s from S r i Lanka

F E MALE : a p t e ro u s ; l e n g t h 2 , 1 2-3 , 75 m m ; h e a d y e I l ow - t e s t aceou s ; occas i o n a l l y b l ac k o r b row n , w i t h m a n d i b l es , c l y p e u s a n d f ro n t p a r t of v er t e x y e l l ow ; a n te n n ae b row n , w i t h s e g me n t s 1 -2 a n d 1 0 y e l l ow ; t h o r a x , p ropodeum a n d l e g s y e l l ow - t e s t aceou s ; occa s i on a l l y s i des of p ro n o t u m a n d m e t a t ho r a x + propodeum b ro w n o r b l ac k ; p e t i o l e b l ac k ; a b domen redd i s h ; occas i on a l l y more or l es s b row n ; h e a d exc a v a ted , s h i n y , h a i r l es s ,

1 420

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F i g . 980 - M a l e gen i t a l i a of Haplogona topus apicalis R . C . L . Perk i n s f rom Sri L a n k a

g r a n u l a t e d ; p ro n o t u m n o t crossed b y a t r a n svers a l i m p ress i o n , s h i n y , smoo t h , h a i r l es s , w i t ho u t sc u l p t u re ; scu t u m s h i n y , w i t h some l o n g i t u d i n a l kee l s ; m e t a n o t u m , p l e u r a a n d p o s te r i or s u rface o f p ropodeum t r a n sverse l y s t r i a t e ; m e t a t horax + p ropodeu m s h i n y , w i t ho u t s c u l p t u re o n a n t e r i o r s u rface a n d a mo n g s t r i a e ; meso-met a p l e u r a l s u t re o b so l e t e ; fore t a rs a l segmen t s i n fo l l o w i n g p ro po r t i on s : 1 0 : 2 : 3 : 7 : 1 1 ; e n l arged c l a w ( F i g . 979 ) w i t h s u b a p i c a l too t h a n d 3-6 l am e l l ae ; segme n t 5 of fron t t a rs u s ( F i g . 979 ) w i t h t w o r o w s o f 7- 1 0 l a me l l ae ; a pe x w i t h a g ro u p o f 2-7 l a m e l l ae ; m a x i l l a r y p a l p i w i t h 2 segmen t s ; l a b i a l p a l p i w i t h 1 segme n t ; t i b i a l s p u rs 1 , 0 , 1 . M AL E : f u l l y w i n g e d ; l e n g t h 2 , 0-2 , 5 mm ; b l a c k or b ro w n , w i t h m a n d i b l es ye l l o w ; a n t e n n ae b ro w n a n d l e g s y e l l ow ; occ a s i on a l l y a n ten n a l segme n ts 1 -2 y e l l o w ; a n t en n a l segme n t s i n fo l l ow i n g p ropor t i on s : 7 : 7 : 1 2 : 1 1 : 1 1 : 1 0 : 9 : 9 : 9 : 1 2 ; a n t e n n a l segme n t 3 more t h a n t h ree t i mes a s l o n g a s b ro a d ( 4 , 0-4 , 5 ) ; h e a d s h i n y , h a i ry , weak l y g ra n u l a ted ; P O L = 9 ; O L = 3 ; OOL = 3 ; b re a d t h o f the o c e l l i : 4 ; fron t a l l i ne a b sen t ; tem p l es d i s t i nc t ; occ i p i t a l c a r i n a i n v i s i b l e ; occ i p u t concave ; s c u t u m h a i ry , d u l l or s h i n y ,

G o n a topo d i n a e

1 42 1

g r a n u l a ted ; n o t a u l i ce s comp l e te , s t ron g l y co n v erg i n g , j o i n t or a l most j o i n t ; s c u te l l u m and m e t a n o t u m s h i n y , smoo t h , f i n e l y p u n c t a t e , h a i r y ; propode u m re t i c u l a t e ru gose , w i t ho u t k e e l s ; fore-w i n g h y a l i n e , w i t h o u t d a r k t r a n svers a l b a n d s ; dorsa l p rocess of gon oforceps ( F i g . 980 ) l o n g , d i s t a l l y b ro a d e n ed ; m a x i I l a ry p a l p i w i t h 2 segmen t s ; l a b i a l p a l p i w i t h segmen t ; t i b i a l s p u rs 1 , 1 , 2 . LOCUS TYP I C U S : B u n d a berg ( A u s t r a l i a ) TYP I CAL M A T E R I A L : l ec t o t y p e F ! a n d 3 p a r a l ec t o t y pes ( 2 F F , 1 M ) ! ( de s i g n a ted b y O l m i 1 98 2 ) of H. apicalis i n B ; l ec t o t y p e F ! ( de s i g n a t e d b y O l m i 1 982 ) o f H. moes tus i n B ; h o l o t y p e F ! of H. brevicornis in B; h o l o t y p e F ! and 3 p a r a t y p es (2 F F , 1 M ) ! of H. orien talis in W A ; h o l o t y p e F ! and a l l o t y p e M ! of H. japon icus in KY . HOST S : i n I n d i a Soga t a s p . ( Ro h w e r 1 920 ) ; i n Sri L a n k a Soga tella furcifera ( Ho rv a t h ) and Nil aparva t a lugens ( S t a l ) ( O t a k e , Som a s u n d a r a m a n d A b e y koon 1 976 ) ; i n t h e P h i l i p p i nes I s l a n d s Nilaparvata lugens ( S t a l ) ; i n M a l a y s i a Sog a t a s p . ; i n J a p a n Soga tella furcifera ( Ho rv a t h ) ( E s a k i a n d H a s h i mo t o 1 93 1 ) a n d L a odelp h a x striatellus F a I I . ( T hompson 1 954 ) ; i n A u s t r a l i a L i b urn ia sp . ( n e a r Chlorion a ) a n d Hadeodelp h a x s p . ( R . C . L . Perk i n s 1 906c ) . D I ST R I B U T I O N : O r i e n t a l reg i o n : I N D I A : B i l a s p u r ( Ja n i g i r ) , WA ! O r i s s a ( K u r i a n 1 954a ) ; Ja n i g i r , N O ! K a n k e y F a r m , N O ! P u s a ( B i h a r ) ( erroneou s l y a l l o t y p e M ! o f Pseu dogon a topus soga t a e R o h w er ) , WA ! S R I LANK A : T i s m a d a ( K a n . D i s t . ) , W A ! K a l u g o m u w a ( K a n d y D i s t r i c t ) ( s pec i me n s desc r i bed b y D t a ke , Som a s u n d a r a m a n d A b e y koon 1 97 6 , now i n OL ) , DL ! C H I N A : S h e k T u n g ( K u a n g T u n g ) , B ! MALAYS I A : K u a l a L u m p u r ( M a l a y a ) , B M ! T a r a t ( K u ch i n g , S a r a w a k ) , Y A ! P H I L I P P I N E S : L o s B a n o s ( L a g u n a ) , B M ! TA I W AN : K i j i n , K Y ! T H A I LA N D : T u n g S ae w , YA ! P a s a n g , YA ! T u n g F a b a d , YA ! B a n m a kok , YA ! C hom-Thon g , K Y ! P a l ae a rc t i c reg i on : JAPA N : K a m i - g u n ( Koc h i P ref . ) , OL ! N I ! F u k u o k a ( Ky u s h u ) , K Y ! N i s h i k a w a t s u -c h o ( M a ts u e C . , S h i m a n e Pref . ) , O U M U ! A u s t ra l i a n re g i on : AUSTRAL I A : B u n d a b e r g ( Queen s l a n d ) , B ! C h i l d ers ( Qu e e n s l a n d ) , B ! C a i rn s ( Queens l a n d ) , B ! B r i s b a n e ( Qu een s l a n d ) , B ! S h e p p a r t o n ( V i c t or i a ) , U Q ! T weed R i v e r ( Co n d o n g , New So u t h W a l es ) , B !

N E O T R O P I C SP E C I ES OF HA PLOGONA TOPUS

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1 . Haplogon a top us insul aris Og l ob l i n

Haplogo n a top u s insul aris O g l ob l i n 1 95 3 :

1 1 3.

F E MA L E : ap terou s ; l e n g t h 4 , 6 m m ; t e s t aceou s , w i t h oce l l a r t r i a n g l e , p ropec t u s , sc u t u m , sc u te l l u m , a p e x o f p ropode u m , pe t i o l e a n d p a r t of a b domen b row n ; m e t a t h o r a x + p ro p o d e u m red d i s h ; h e a d d u l l , g r a n u l a t e d ; occ i p i t a l c a r i n a i n v i s i b l e ; POL � 2 ; OL � 3 ; OOL � 1 0 ; fron t a l l i n e a b se ;o � ; a n t e n n a e m i s s i n g i n t h e on l y k n o w n spec i men ; p ron o t u m sh i n y , w e a k l y g ra n u l a te d , not crossed b y a t r a n s v e rs a l i m p ress i o n ; s c u t u m d u l l , g r a n u l a t e d , w i t h some l o n g i t u d i n a l k ee l s ; s c u t e l l u m d i s t i n ct , short ; m e t a n o t u m t r a n s v e rse l y s t r i a t e ; m e t a t h o r a x + propod e u m d u l l ; p l e u r a , d i sc , a n te r i o r a n d pos t e r i or s u rfaces t r a n s v erse l y s t r i a t e ; meso me t a p l e u r a l s u t u re obso l e te ; p a r t of fore l e g s m i s s i n g i n the on l y k n o w n spec i m en ; fore t a r s i fu l l y m i s s i n g i n t h e on l y k n o w n spec i men { accord i n g to O g l ob l i n - 1 953- e n l a r g e d c l a w w i t h s u b a p i c a l tooth a n d 5 l a me l l a e ; segmen t 5 of fro n t t a rsus w i t h t w o ro w s of 8 l a m e l l a e ; a p e x w i t h a g r o u p of 1 0 l a m e l l a e } ; m a x i l l a ry p a l p i w i t h 2 segmen ts ; l a b i a l p a l p i w i t h 1 segmen t ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n w o n L O C U S T Y P I C U S : B a h i a C u m b er l a n d ( M a s a t i e r r a I s l a n d , J . F e r n a n dez I s l ands , C h i I e ) TYP I CAL MA T E R I A L : h o l o t y p e F ! { l a b e l l ed Hap l ogon a top u s kuscheli a n u s } P T ( s l i d e of mou t h p a r t s o f t h e ho l o t y pe i n SW ! ) .

in

HOST S : u n k n o w n D I ST R I B U T I O N : Neot rop i c reg i o n : C H I L E : B a h i a C u m b e r l a n d ( M a s a t i erra I s l a n d , J. F e r n a n dez I s l a n d s ) , P T !

2 . Haplogon a topus hern a n dezae n .

sp.

F E MA L E : a p t e ro u s ; l e n g t h 1 , 6-3 , 2 m m ; head b l ac k , w i t h c l y p e u s a n d m a n d i b l es t e s t aceo u s ; a n te n n ae b l ac k , w i t h segmen ts 1 -2 t e s t aceo u s ; segmen t 1 0 l i g h t ; p ronot u m a n d sc u t u m b l ac k or b row n ; occ a s i o n a l l y sc u t u m test aceou s ; s c u t e l l u m a n d m e t a t ho r a x + p ropode u m b l ack , w i t h a p e x o f propo d e u m t e s t a ceou s ; a b domen b l ac k ; l eg s tes t aceou s , w i t h c l u b s of femora a n d t i b i a e b ro w n ; a n t e n n a e d i s t a l l y t h i c k e ned ; a n t e n n a l segmen t s i n fo l l ow i n g p ropor t i on s : 9 : 5 , 5 : 1 0 : 6 : 6 : 5 : 5 : 5 : 6 : 1 0 ; h e a d

1 424

Mem . A m e r .

En!.

I n s! . ,

n o . 37 ,

1 984

excav a te d , d u l l , g r a n u l a ted ; fron t a l l i ne com p l e t e ; occ i p i t a l c a r i n a o n l y v i s i b l e n e a r t he pos t e r i o r oce l l i ; POL � 1 , 5 ; O L � 1 , 5 ; DOL � 8 ; p ro n o t u m n o t crossed b y a t r a n s v e rs a l i m p ress i o n o r w e a k l y i m p ressed , s h i n y , weak l y gra n u l a t e d ; s c u t u m d u l l , g r a n u l a t e d ; scu t e l l u m s h i n y , smoo t h ; m e t a n o t u m t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re o b so l e t e ; p l eu r a a n d p o s t e r i o r s u rf ac e o f m e t a t ho r a x + p ropode u m t r a nsverse l y s t r i a t e ; a n t er i or s u rf ace o f m e t a t ho r a x + p ro p o d e u m s h i n y , smoo t h , n o t t r a n sv erse l y s t r i a t e ; fore t a rs a l seg men t s i n f o l l o w i n g p ropor t i on s : 1 4 : 3 : 4 : 1 2 : 1 9 ; e n l a r g e d c l a w ( F i g . 9 8 1 ) w i t h s u b a p i c a l tooth a n d 6 l ame l l ae ; segmen t 5 o f f r o n t t a rs u s ( F i g . 9 8 1 ) w i t h t w o rows of n e a r l y 8 l a me l l ae ; a p e x w i t h a g ro u p of 8 l a me l l ae ; m a x i l l a ry p a l p i w i t h 2 s e g me n t s ; l a b i a l p a l p i w i t h 1 segmen t ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : C . I . A . T . ( P a l m i r a , C o l o m b i a ) TYP I CA L MA T E R I A L : ho l o t y p e F ! a n d 40 p a ra t y p es F F ! i n O L ; 1 p a r a t y p e F ! i n G V ; 1 p a r a t y p e F ! i n BU ; 2 p a r a t y pe s F F ! i n B M . HOST S : i n C o l o m b i a a n d Costa R i ca Soga todes oryzicol a . D I S T R I B U T I ON : Neo t ro p i c reg i on : COSTA R I CA : G u a n ac os t a , O L ! COLOMB I A : C . I . A . T . ( P a l m i r a , V a l l e ) , OL ! BM ! BU ! G V ! NO T E S : t h e s p ec i es i s n a m e d i n honor o f t he f i n d e r M i s s M a r t h a d e l P i l a r Hern a n d e z .

AUSTRAL I A N S P E C I E S O F HA PLOGONA TOPUS

T w o s p ec i es are k n ow n .

KEY T O THE SPEC I ES

FEMALES

T h e fem a l es o f H. apica lis a n d vitiensis are n o t i de n t i f i ab l e . T h e m a l e s , on t h e con t r a r y , are ea s i l y recog n i za b l e for t he d i ffere n t p rocesses o f g o n oforceps

MALES

D o r sa . . . . . D o r sa . . . . .

l . l .

p r o c e s s o f g on o f o r c ep s s l e n d e r ( F i g . 9 8 3 ) . . . . . . . . . . . . . . . . . . . . . . 1 . vitiensis R . C . L . P e rk i n s p r o c e s s o f g on o f o r c e p s b r oad ( F i g . 9 8 0 ) . . . . . . . . . . . . . . . . . . . . . . . . 2 . apica lis R . C . L . P e r k i n s ( s e e O r i en t a l s p e c i e s )

Gon a topod i n a e

Fig.

1 42 5

9 8 2 - C h e l a o f Haplogon a top u s vitiensis R . C . L . T a l ofofo ( G u a m )

1 . Haplogon a top u s vitiensis R . C . L .

Haplogo n a top u s Haplogo n a topus Haplogon a top u s Haplogo n a top u s C u rrado 1 97 9 b :

vitiensis vitien s is vitiensis apica lis 42 .

P e r k i n s from

Perk i n s

R . C . L . Perk i n s 1 906a : 488 . R . C . L . P e r k i n s : Swezey 1 928 : 670 . R . C . L . P e r k i n s : W i l l i a m s 1 93 1 : 247 . R . C . L . P e r k i n s partim : O l m i a n d

F E MAL E : a p t e ro u s ; l e n g t h 2 , 5-3 , 0 m m ; h e a d b ro w n , w i t h m a n d i b l es , c l y p e u s a n d fron t p a rt o f v e r t e x y e l l ow ; a n te n n a e b ro w n , w i t h s e g m e n t s 1 -2 a n d 1 0 y e l l o w ; t ho r a x a n d p ropod e u m redd i s h ; p e t i o l e b l ac k ; a b domen b row n ; l e g s y e l l ow - t e s t aceo u s ; head exc a v a t e d , s h i n y , h a i r l es s , g r a n u l a ted ; p ro no t u m n o t crossed by a t r a n sversa l i m p r e s s i o n , s h i n y , smoo t h , h a i r l es s , w i t ho u t scu l p t u re ; scu t u m s h i n y , w i t h some l o n g i t u d i n a l k ee l s ; m e t a n ot u m , p l eu ra a n d p os t e r i or s u rf ace o f p ropodeum t r a n s v e r se l y

Mem . A m e r .

1 426

Fig.

983

-

Ent.

I nst . ,

no. 37,

1 984

M a l e ge n i t a l i a of !!a plogon a top us viti e nsis R . C . L . F i j i I s l ands

Perk i n s from

s t r i a te ; meso-me t a p l e u r a l s u t u re o b so l e te ; m e t a t horax + p ropod e u m w i t hou t scu l p t u re on a n te r i o r s u rface a n d amon g s t r i ae ; fore t a rsa l s e g m e n t s i n fo l l ow i n g p roport i o n s : 1 2 : 2 : 3 : 7 : 1 2 ; e n l arged c l a w ( F i g . 982 ) w i t h s u b a p i c a l too t h a n d 3-4 l a m e l l ae ; segmen t 5 of fron t t a rs u s ( F i g . 982 ) w i t h two ro w s of 7-9 l ame l l ae ; a p e x w i t h a g ro u p of 2-5 l a m e l l a e ; m a x i l l a r y p a l p i w i t h 2 segmen t s ; l a b i a l p a l p i w i t h 1 segmen t ; t i b i a l s p u rs 1 , 0 , 1 . M A L E : f u l l y w i n ged ; l e n g t h 2 m m ; b ro w n or b l ac k , w i t h m a n d i b l es a n d l eg s y e l l ow ; a n te n n a l s e g m e n t s i n fo l l ow i n g p roport i on s : 7 : 6 : 8 : 8 , 5 : 9 :

Gon a topod i n a e

1 427

8: 7: 8: 7 : 1 1 ; a n t e n n a l segmen t 3 more t h a n t h ree t i mes a s l o n g as b road ; head s h i n y , h a i r y , w e a k l y p u n c t a t e , n o t g r a n u l a t e d ; fron t a l l i n e a b s en t ; t em p l es d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 9 ; O L = 3 ; O O L = 3 ; b r ea d t h of t h e ocel l i : 4 ; sc u t u m ha i ry , d u l l , g ra n u l a t ed ; n o t a u l i ces d i s t i nc t , c o m p l e t e , s t ron g l y c o n v e r g i n g , p o s t e r i o r l y j o i n t ; s c u t e l l um a n d m e t a n o t u m s h i n y , h a i r y , w e a k l y p u n c t a t e ; p ropodeum ret i c u l a t e r u gose , w i t h o u t k ee l s ; fore- w i n g h y a l i n e , w i t ho u t d a rk t r a nsvers a l b a n d s ; dors a l p rocess o f gonoforceps ( F i g . 983 ) l o n g , s l en d e r , d i s t a l l y b roade n e d ; m a x i l l a r y p a l p i w i t h 2 segme n t s ; l a b i a l p a l p i w i t h 1 segmen t ; t i b i a l s p u rs 1 , 1 , 2 . L O C U S TYP I C U S : S u v a ( F i j i I s l a n d s ) T Y P I CAL MAT E R I A L : ho l o t y p e F ! o f H. vitien sis i n B . H O ST S : i n t h e F i j i I s l a n d s Perkinsiella vitiensis K i r k a l d y ; i n t h e H a w a i i a n l s i a n d s Peregrin u s ma idis A s h m . a n d Perk insiella s a ccharicida K i r k . ( T ho m p s o n 1 954 ) . PARAS I T E S : i n t h e H a w a i i a n I s l a n d s Ceraphron a bn ormis R . C . L . Perk i n s ( u n d e r n a m e Haplogon a topus vitien sis R . C . L . Perk i n s ) a n d Cheiloneurus americ a n u s ( R . C . L . Perk i n s 1 906b : u n de r n a me Haplogon a top u s vitien sis ) . D I ST R I B U T I O N : A u s t r a l i a n reg i o n : MAR I AN A I SLANDS : T a l ofofo ( G u am ) , BN ! F I J I I SLAN D S : S u v a ( V i t i L e v u ) , B M ! N a v u a ( V i t i L e v u ) , B ! La u to k a ( V i t i L e v u ) , B M ! HAWA I I AN I SL A N D S : O a h u , B ! E w a ( O a h u ) , B M ! NOT E S : I h a v e seen m a l es from t h e F i j i I s l a n ds a n d from A u s t r a l i a ; t h e y h a v e g en i t a l i a very d i fferen t ; t he g en i t a l i a o f m a l es f r o m A u s t r a l i a a re very s i m i l a r to g e n i t a l i a of H. orien talis ; I t h i n k so t h a t p o p u l a t i on s of A u s t r a l i a are b e l on g i n g to orie n t a lis ( p reocc u p i e d b y apicalis ) , w h ere a s p o p u l a t i on s f r o m t h e M a r i a n a , F i j i a n d H a w a i i a n I s l a n d s a re vitien sis . H. vitiensis w a s i n t ro d u c e d i n t o H a w a i i a n I s l a n d s from F i j i i n 1 906 . T he p resence of h y p e r p a r a s i t e s ( E n c y r t i d s ) h a s g re a t l y reduced t h e eff i c i en c y of t h i s s p ec i es a g a i n s t Perkinsiella s a ccharicida K i rk . ( W i l l i a m s 1 93 1 ) . P r o b a b l y t h e i n t rod u c t i on h a d no s uccess .

2 . Haplogo n a top u s apicalis C . L . ( see O r i e n t a l s p ec i e s )

1 1 . Pen tagon a topus n .

Perk i n s

gen .

TYPE S P E C I E S : Pen tagon a top u s mon t a n u s n . s p . F E MA L E ( F i g s 985 , 986 ) : a p t e rou s ; m a x i I l a r y p a l p i w i t h 5 s e g me n t s ; l a b i a l p a l p i w i t h 2 segme n t s ; e n l arged c l a w ( F i g . 984 ) w i t h s U b a p i c a l t o o t h a n d l a me l l ae ; p ro n o t u m ( F i g . 986 ) n o t crossed b y a t r a n s vers a l i mp res s i on or w e a k l y i m p ressed ; t i b i a l s p u rs 1 , 0 , 1 . M AL E : u n k n o w n D I ST R I B U T I ON : N e a rc t i c , A u s t r a l i a n H O ST S : u n k no w n

1 428

Mem . Amer .

En!.

I n s! . ,

no.

37 ,

1 984

F i g . 984 - C he l a of Pen tago n a topus b orealis n . s p .

SPEC I ES :

( type)

2

N E A R C T I C S P E C I E S OF PENTA GONA TOPUS

O n e s p ec i es i s k n ow n .

Pen tagon a top u s bore a l i s n . s p .

F E M A L E : a p t e r o u s ; l en g t h 3 , 1 8 m m ; b l a c k ; m a n d i b l es , c l y p e u s a n d f ro n t p a r t o f v e r t e x ( more a l o n g o rb i t s ) t e s t aceou s ; a n t e n n ae t e s t aceous-d a r k ; l eg s b l ac k , w i t h t roc h a n ters , t i b i ae a n d t a r s i t e s t aceo u s ; a n t e n n a e d i s t a l l y t h i c k e n e d ; a n ten n a l segmen t s i n f o l l ow i n g p roport i on s : 8 : 5 : 1 3 : 1 0 : 8 : 8 : 6 : 6 : 6 : 1 0 ; h e a d s h i n y ; frons smoo t h , w i t h o u t scu l p t u re ; t em p l es a n d occ i p u t g r a n u l a t ed ; f ron t a l l i n e comp l e t e ; occ i p i t a l c a r i n a o n l y v i s i b l e b e h i n d t h e oce l l i a n d o n t he s i de s o f t he p o s t e r i o r ocel l i ; POL � 2; O L � 2 , 5 ; OOL � 9; p ro no t u m v e r y w ea k l y crossed by a t ra n sversa l i m p re s s i o n , g ra n u l a t e d ; scu t u m s h i n y , w i t h some l on g i t u d i n a l k ee l s ; met a n o t u m f l a t , d u l l , t ra n s v erse l y s t r i a t e ; m e t a t h o r a x + p ro p o d e u m d u l l , gra n u l a ted , w i t h p l eu ra a n d pos t e r i o r s u rf ac e t r a n s v e rse l y s t r i a te ; a n t e r i o r s u rf ace of p ropodeum w i t h a n i n com p l e t e t r ack of med i a n f u r row ; meso-me t a p l e u r a l s u t u re o b so l e t e ; fore t a r s a l segmen t i n fo l l ow i n g p ropor t i o n s : 1 7 : 3 , 5 : 5 : 1 3 : 22 ; en l a rged c l a w ( F i g . 984 ) w i t h s u b a p i c a l t o o t h a n d 6 l a me l l ae ; seg m e n t 5 o f f r o n t t a rs u s ( F i g . 984 ) w i t h t wo row s

Gon a topod i n ae

F i g . 985 - Fema l e of Pen t a gon a top u s m on ta n u s

1 429

n.

sp .

( type)

o f 9 + 1 2 l am e l l ae ; apex w i t h a g ro u p o f 1 4 l a m e l l a e ; m a x i l l a r y p a l p i w i t h 5 s e g m e n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M AL E : u n k n o w n LOCUS TYP I C U S : P a r k e Reserve ( Qu ebec , C a n a d a ) TYP I CAL M A T E R I A L : h o l o t y pe F ! i n QT . HOST S : u n k n o w n D I ST R I BU T I ON : Nea rc t i c reg i o n : CANAD A : P a r k e Reserve ( Qu e . ) , O T !

AUSTRAL I AN S P E C I E S OF PENTA GONA TOPUS

1 430

Mem . A m e r . E n t .

I ns t . ,

no.

37,

1 984

F i g . 986 - F e m a l e of Pen tagon a top u s mon t a n u s n . s p .

( type)

O n e spec i es i s k n ow n .

Pen tagon a top u s m on t a n u s n . s p .

F E M A L E ( F i g s 985 , 986 ) : a p t e ro u s ; l e n g t h 3 , 2 5 m m ; b l ack , w i t h m a n d i b l es , c l y p e u s a n d fron t p a r t o f v e r t e x y e l l o w ; a n te n n ae b row n ; l eg s b ro w n b l ack , w i t h p ar t of troc h a n t e rs a n d p a rt of t a r s i y e l l ow ; head e x c a v a t e d , d u l l , w i t ho u t scu l p t u re ; f ron t a l l i n e comp l e t e , weak ; p ro n o t u m w e a k l y t r a n s v e r se l y i m p ressed , s h i n y ; co l l a r smoo t h , w i t h o u t scu l p t u re ; d i sc w i t h n umero u s , w e a k , l o n g i t u d i n a l kee l s ; m e t a n o t u m d u l l , t r a n s v erse l y s t r i a t e ; m e t a t horax + p ro p o d e u m d u l l ; a n te r i o r s u rf ace w i t h s t r o n g l on g i t u d i n a l k ee l s ; p l e u r a a n d pos t e r i o r s u rf ace o f p ropodeum t r a n sverse l y s t r i a t e ; meso-m e t a p l e u r a l s u t u re d i s t i nc t ; seg m e n t 1 o f f r o n t t a rs u s l on g e r t h a n segm e n t 4 ( 1 5 : 1 3 ) ; en l a r g e d c l a w ( F i g . 987 ) w i t h s u b a p i c a l t oo t h a n d 6 l a me l l ae ; se g me n t 5 of fro n t t a r s u s ( F i g . 987 ) w i t h t w o rows of 6 + 1 2 l a me l l a e ; a p e x w i t h a g r o u p of 8 l a me l l ae ; m a x i l l a ry p a l p i w i t h 5 segme n t s ; l ab i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k no w n LOCUS TYP I C U S : M t . P i o r a ( New G u i n e a ) T Y P I CA L M A T E R I A L : ho l o t y p e F ! i n B . HOST S : u n k n o w n D I S T R I B U T I O N : A u s t r a l i a n reg i on : N E W GU I N EA : M t . P i o ra ( N E N e w G u i n ea ) , B!

G o n a t o po d i n a e

Fig.

987 - C h e l a o f Pen tagon a topus mon t a n u s n .

1 43 1

sp .

( type)

1 2 . Plectrogona topoides Ponomarenko

Plec trogona topoides Ponomarenko 1 975a : 3 1 8 .

T Y P E S P E C I E S : Plectrogon a topoides chortinis Ponom a r e n ko 1 97 5 , o r i g . des i g . F E M A L E ( F i g s 988 , 989 ) : a p tero u s ; m a x i I l a r y p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 3 segme n t s ; pron o t u m ( F i g . 989 ) n o t crossed b y a t r a n s versa l i m p ress i o n or weak l y i m p ressed ; e n l a rged c l a w ( F i g . 990 ) w i t h s u b a p i c a l too t h a n d l a me l l a e ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n D I S T R I B U T I O N : P a l a e a rc t i c , Nearc t i c H O ST S : u n k n o w n SPEC I ES : 2

PALAEAR C T I C S P E C I E S OF PLECTROGONA TOPOIDES

1 432

Mem . Amer . E n t .

I n st . ,

no.

37,

1 984

F i g . 988 - F e m a l e of Plectrogon a top u s nearcticus n.

sp .

( type)

O n e s p ec i es i s k now n .

Plectrogona topoides chortin is Ponomarenko

Plectrogon a topoides chortin i s P o n o m a r e n k o 1 97 5 a : 3 1 8 .

F E M A L E : I h a v e n o t seen t h e on l y k n o w n s p e c i men . See t he o r i g i n a l descri p t i on . MALE : u nknown L O C U S T Y P I C U S : U s c i u g h u n-Obo ( A n d a-X u d u k , U v e r-X a n g a i a i m a k , Mo n g o l i a )

Gona topod i n ae

1 433

F i g . 989 - F e m a l e of Plec trogon a top u s nearcticus n . s p .

T Y P I CAL MAT E R I A L : HOSTS : u n k n o w n

( t y pe )

t h e t y p e seems l os t .

D I STR I BU T I O N : P a l ae a rc t i c reg i o n : MONGOL I A : U ver-X a n g a i a i mak ) ( Ponom a re n k o 1 975a ) .

U sc i u g h u n -Obo ( A n d a-X u d u k ,

N E A R C T I C S P E C I E S OF PLECTROGONA TOPOIDES

O n e s p ec i es i s k n ow n .

Plectrogona topoides nearcticus n . s p .

F EM A L E ( F i g s 988 , 989 ) : a p terou s ; l e n g t h 2 , 5 m m ; head t e s t aceo u s , w i t h a b l ac k b a n d on verte x ; a n te n n ae b row n , w i t h segmen t s 1 -2 a n d p a rt of 3 t e s t a ceou s ; p rope c t u s t e s t aceou s ; prono t u m b l ac k , w i t h s i des red d i s h ; mesothora x , m e t a t ho r a x a n d p ropodeum b l ac k ; abdomen t e s t aceo u s ; l e g s t e s t aceo u s ; a n t e n n ae d i s t a l l y t h i c k e n e d ; a n t en n a l seg m e n t s i n fo l l ow i n g propor t i o n s : 9 : 5 : 1 1 : 5 : 5 : 4 , 5 : 5 : 4 : 4 : 6 ; head e x c a v a ted , d u l l , g r a n u l a ted ; f ro n t a l l i n e comp l e t e ; occ i p i t a l ca r i n a i n v i s i b l e ; POL = 2 ; O L = 2 ; OOL = 6 ; p ro n o t u m weak l y crossed b y a t r a n s v e rs a l i m p re s s i o n , s h i n y , w i t h o u t s c u l p t ure ; scu t u m s h i n y , w i t h o u t scu l p t u re ; m e t a n o t u m w i t h t w o l a t e r a l p o i n t s ; m e t a t ho r a x + p ropodeum s h i n y , a l mo s t w i t ho u t scu l p t u re or weak l y g r a n u l a ted ; mesop l eu r a n o t t r a n sverse l y s t r i a t e ; me t a p l e u r a a n d poster i o r s u rface of prop o d e u m t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re obso l e t e ; fore t a rs a l seg m e n t s i n fo l l ow i n g p ropor t i on s : 1 0 : 2 : 4 :

Mem . Amer . E n t .

1 434

F i g . 990

-

I ns t . ,

n o . 37 ,

1 984

C h e l a of Plectrogon a top u s n e arcticus n .

sp .

( type)

1 2 : 2 0 ; e n l a r g e d c l a w ( F i g . 990 ) w i t h s u b a p i c a l too t h a n d 8 l a m e l l a e ; segmen t 5 o f f r o n t t a r s u s ( F i g . 990 ) w i t h t w o row s o f n e a r l y 4 0 l a me l l ae e x t e n d i n g con t i n uo u s l y to t h e a pe x ; m a x i I l a r y p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n known L O C U S TYP I CU S : F a i rf a x C o . ( V i r g i n i a , U . S . A . ) T Y P I CA L MAT E R I AL : HOST S : u n k no w n D I ST R I B U T I ON :

holotype F !

N e a rc t i c reg i o n :

i n WA . U.

S . A . : F a i rfax C o .

( V a . ) , WA !

1 3 . Tetrodontochelys R i c h a rds

Tetrodontochelys R i c h a rds 1 939 : 2 1 7 . Epigon a topoides R i c h a r d s 1 93 9 : 2 2 1 ( n . s y n . ) ; t y p e spec i es Epigon a topoides p u licarius ( K l u g 1 8 1 0 ) , or i g . d es i g . Tetra don tochelys J . F . P erk i n s 1 976 ( j a n u a ry ) : 35 ( emen d . i n va l i d ) . Neogon a topoides A b d u l -Nou r 1 976 ( se p te m b e r ) : 274 ( n . s y n . ) ; t y p e s p ec i es Neogon a topoides cla udiae A b d u l -N o u r 1 97 6 , monot y p i c .

TYPE SPEC I E S :

Tetrodon tochelys ljunghii ( We s t wood 1 833 ) ( = T . pedestris ) ,

or i g . d es i g . F E MA L E ( F i g s 9 9 1 , 992 , 9 97 ) : a p te ro u s ; m a x i I l a r y p a l p i w i t h 2-5 seg m e n t s ; l ab i a l p a l p i w i t h 2 segme n t s ; p ron o t u m ( F i g . 992 ) n o t crossed b y a t r a n s v e r s a l i m p ress i o n or w e a k l y i m p resse d ; e n l arged c l a w ( F i g . 9 9 3 ) w i t ho u t s u b a p i c a l too t h , w i t h a s m a l l t oo t h a t the e n d of a l on g i t u d i n a l f u rrow a n d w i t h b r i s t l es o r l a me l l ae ; t i b i a l s p u rs 1 , 0 , 1 .

Gon a topod i n ae

F i g . 991

-

F e m a l e of Tetrodon tochelys pedes tris ( I ta l y )

1 435

( D a l m a n ) from Verce l l i

1 436

Mem . Amer . E n t .

I n st . ,

no. 37,

1 984

F i g . 992 - F e m a l e of Tetrodon tochelys pedestris ( D a l m a n ) from Verce l l i ( I ta l y )

M A L E ( F i g s 995 , 1 0 1 2 ) : f u l l y w i n g e d ; m a x i l l a r y p a l p i w i t h 2-4 segmen t s ; l a b i a l p a l p i w i t h 2 segme n t s ; occ i p u t con c a v e ; t em p l es d i s t i nc t ; occ i p i t a l c a r i n a i n v i s i b l e o r on l y v i s i b l e b e h i n d t h e oce l l i o r o n t h e s i de s o f t h e poste r i o r oce l l i , b u t n e v e r reac h i n g temp l es ; t i b i a l s p u rs 1 , 1 , 2 . D I ST R I B U T I ON : P a l a earct i c , E t h i op i a n , O r i e n t a l , N e a rc t i c , A u s t r a l i a n HOST S : Cic a delli da e , exc e p t for Typhlocyb in a e , Idiocerin a e , Macrop s in a e SPEC I ES: 1 8

PALAEAR C T I C S P E C I E S O F TETRODONTOCHEL YS

S i x spec i es a re k n ow n .

K E Y T O T H E SPEC I E S

F E MA L E S

S eg m e n t

5 of

f r on t

t a r su s w i t h

2-7 m i n u t e

l a me l l a e o n

the

Gon a topod i n ae

di stal

2

3

ha l f

(F i gs 993 ,

pedes tris ( D a l ma n ) !

2 . s a k a ii ( E s a k i a n d Ha s h i mo t 0 ) 1 S e g m e n t 5 o f f r on t t a r s u s w i t h a l on g r o w o f b i g l a me l l a e o n d i s t a l a n d p r ox i ma l h a l f ( F i g s 9 9 8 , 9 9 9 , 1 0 0 0 , 1 0 0 1 ) ; o c ca s i on a l l y o n l y o n t h e d i s t a l h a l f , b u t i n t h i s c a s e l ame l l a e v e ry b i g , n o t m i n u t e . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 B od y y e l l ow - t e s t a c e o u s o r r ed d i s h , w i t h p e t i o l e b l a c k , a b d omen b r o wn i sh and o c c a s i on a l l y p a r t of a n t e n n a e b r own . . . . . . . . . . . . . 3 B o d y a t l ea s t p a r t l y b l a ck . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 B o d y y e l l ow - r ed d i sh ( w i t h p e t i o l e b l a c k a n d o c ca s i on a l l y a b d o m e n b r own i sh ) 3 . p u licarius ( K I u g ) B o d y r ed d i sh ( w i t h p e t i o l e b l a ck , a b d omen a n d o c ca s i o n a l l y •

4

996 ) . 1 .

1 437

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p a r t o f a n t e n n a e b r o wn i sh ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 . pseudo a u da x n . s p . · . . . . . . . . . . . . . . . H ea d f l a t , w i t h t emp l e s v e ry l o n g . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 . aegyptiacus n . s p . · H ea d e x c a v a t ed , w i t h t emp l e s s h o r t . . . . . . . . . . . . . . . . . . . . . . . . . . . 6 . a u da x n . s p . · .

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MALES

O n l y t h e m a l e of T.

pedestris i s k n ow n .

1 . Tetrodon tochelys pedestris ( D a I m a n )

Gon a top u s pedes tris D a l m a n 1 8 1 8 : 86 . nee Dicondylus pedestris ( D a l m a n ) : H a l i d a y i n C u r t i s 1 829-30 . Gon a top u s ljunghii W e s t wood 1 83 3 b : 496 ( s y n . p ro p osed by O l m i a n d C u rr a d o 1 977 ) ; locus typ ic u s W i m b l e don Common ( E n g l a n d ) . Pachygon a top u s pedestris ( D a l m a n ) : K i effer i n K i effer e t M a r s h a l l 1 906 : 500 . Dicondylus ljungh ii ( We s t wood ) : K i effer 1 907b : 22 . Gon a top u s leu costom u s J . S a h l b e r g 1 9 1 0 : 1 4 ( s y n . p roposed b y O l m i 1 97 9c ) ; locus typ ic u s J a a k k i m a ( F i n l a n d ) . Pl a tygona top u s p e dest-ris ( D a l m a n ) : K i effer 1 9 1 4b : 9 2 . Pla tygon a top u s pedestris ( D a l m a n ) : Ber l a n d 1 9 28 : 1 47 . Gon a top u s Ijunghii W e s t w oo d : R c i h a r d s 1 935 : 1 46 . Tetrodon toch e lys Ijunghii ( We s t wood ) : R i c h a rds 1 939 : 2 1 7 .

! T h e f em a l e s o f T . pedestris ( from Wes t e r n P a l ae a rc t i c reg i o n ) a n d s a k a ii ( from J a p a n ) seem i n d i s t i n g u i s h a b l e . O n l y t h e s t u d y of m a l e s from J a p a n w i l l c l e a r t h i s p ro b l em .

1 438

Mem . Amer . E n t .

I n st . ,

no . 37 ,

1 984

F i g . 993 - C he l a of Tetrodon tochelys pedestris ( D a l ma n ) from Verce l l i ( I tal y )

Pachygo n a top u s arnoldii typ ic u s D e rk u l ( U . s . s . Pla tygon a top u s pedestris Tetrodon tochelys ljunghii Pachygon a top u s arnoldii 1 60 .

P o nomar e n k o 1 96 6 : 1 578 ( n . sy n . ) ; loc u s R.). ( D a l ma n ) : N a g y 1 967 a : 1 23 . ( We s t wood ) : N a g y 1 967a : 1 23 . Ponomare n k o : C u rr a d o a n d 0 I m i 1 972 :

Gon a topod i n ae

1 439

F i g . 994 - M a Ie of Tetrodontochelys pedestris ( D a I m a n ) from Verce l l i ( I ta l y )

Tetra dontochelys Iju n gh i i ( West wood ) : J . F . P e rk i n s 1 976 : 3 5 . Tetrodon tochelys pedestris ( D a l m a n ) : O l m i a n d C u rrado 1 97 7 : 76 . Pachygon a top u s arnoldii Ponoma ren ko : Ponomarenko 1 978 : 2 5 .

FEMALE

( F i gs 9 9 1 , 992 ) :

a p t e rou s ;

l en g t h 2 , 82-3 , 00 m m ;

head b l ac k or

b row n , w i t h m a n d i b l e s , c l y p e u s and fron t part of v e r t e x y e l l ow o r y e l l ow - b row n ; a n t e n n a e y e l l ow o r y e l l ow - b row n ; t ho r a x , p ropodeum a n d a b domen b l ac k ; l e g s y e l l ow , occ a s i on a l l y w i t h p a r t of coxae a n d p a r t o f c l ub s of femora b ro w n or b l ac k ; h e a d ( F i g . 1 006 G ) w e a k l y e x c a v a ted , s h i n y , g r a n u l a t e d ; temp l es d i s t i n c t ; occ i p i t a l ca r i n a i n v i s i b l e ; p ro no t u m

( F i g . 1 006 G ) s h i n y , n o t crossed b y a t r a n sversa l i mp re s s i o n o r w ea k l y i m p ressed , w e a k l y g r a n u l a ted ; scu t u m d u l l , g r a n u l a ted ; m e t a n o t u m show i n g a sem i c i rc l e- s h a p e d kee l ; m e t a t h o r a x + p ro p o d e u m ( F i g . 1 006 G ) g ran u l a te d , w i t h t r a c k of med i a n f u rrow ; poster i or s u rface of p ropod e u m t r a n s verse l y s t r i a t e ; meso-me t a p l eu r a l s u t u r e o b so l e t e ; en l arged c l a w ( F i g . 993 ) w i t ho u t s u b a p i c a l too t h , w i t h a sm a l l tooth a t t h e e n d o f a l o n g i t u d i n a l f u r row a n d w i t h some b r i s t l es ; s e g m e n t 5 of f ro n t t a rs u s ( F i g . 993 ) w i t h 1 row of 2-7 m i n u t e l a me l l ae o n t h e d i s t a l h a l f ; a p e x w i t h a

1 440

Fig.

Mem . A m e r . E n ! .

I ns t . ,

n o . 37 ,

1 984

995 - M a I e gen i ta I i a of Tetrodontochelys pedestris ( D a I m a n ) from Verce l l i ( I t a l y )

g ro u p of 4-9 l ame l l ae ; m a x i I l ary p a l p i w i t h 2-4 s e g me n t s ; l ab i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E ( F i g . 994 ) : fu l l y w i n ged ; l e n g t h 2 , 4 m m ; head b l ac k , w i t h m a n d i b l es y e l l ow ; a n te n n a e b l ack-b row n , w i t h p a r t of s e g me n t s 1 -2 y e l l ow ; t ho r a x a n d propodeu m b l ac k ; a b domen b rown-b l ac k ; l eg s b ro w n -b l ac k , w i t h s t a l k o f fore femora a n d fore t i b i a e y e l l ow ; a n t e n n a I segmen t 3 more t h a n f o u r t i mes a s l o n g a s b road ( 5 , 6 ) ; tem p l es d i s t i n c t ; occ i p u t conca ve ; occ i p i t a l cari n a i n v i s i b l e ; scutum d u l l ,

g ra n u l a t e d ; n o t a u l i ce s comp l e t e , d i s t i nc t ,

poste r i o r l y s e p a r a t e d ; sc u te l l u m a n d m e t a n o t u m g r a n u l a t e d ; p ropodeum ret i c u l a t e r u gose ; fore-w i n g h y a l i n e , w i t h o u t d a rk t r a n sversa l b a n d s ; dorsa l process of gonoforceps ( F i g . 995 ) l on g , s l e n d e r , w i t h a p e x po i n t e d ; m a x i I l a ry p a l p i w i t h 3-4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . LOCUS TYP I C U S : V a s tergot l a n d ( Sweden ) T Y P I CAL MAT E R I AL : l ec t o t y p e F ! ( de s i g n a t e d by O l m i a n d C u rrado 1 977 ) of G. pedestris i n ST ; h o l o t y p e F ! of G. lju nghii i n O X ; ho l o t y p e F ! o f G . l e u costomus i n H E ; h o l o t y p e F ! a n d 1 p a r a t y pe F ! of P . a rnoldii i n M O . HOST S : i n I t a l y Macrosteles l a evis ( R i b a u t ) ( C u rrado a n d O l m i 1 972 , u n der n ame Pachygon atop u s arnoldii ) ; in F i n l a n d b y m i s t a k e Calligypona o b scurella Bo h . , accord i n g to L i n d b e r g ( 1 95 0 ) ; i n U . S. S. R. by m i s t a ke Delph a c i da e . accord i ng to Ponom a re n k o ( 1 97 8 ) .

Gona topod i n a e

1 44 1

F i g . 996 - C he l a o f Tetrodontochelys s a k a ii ( E s a k i a n d H a s h i moto ) from Kami -gun ( Japan )

D I STR I B U T I O N : P a l a e a rc t i c reg i o n : U . S . S . R . : D e rk u l ( l u g a n s k , U k ra i ne ) , M O ! K a m e n k a ( N aro Fom i n s k , Moscow ) ( Ponomarenko 1 966 ) ; F I NLAN D : J a a k k i ma ( K a re l i a ) , H E ! P a r g a s ( Po r i n ) , HE ! Y l a n e ( Por i n ) , H E ! n umerous o t h e r l oc a l i t i es ( He l so , lojo , E k e n a s , T e r i jok i , Sot t u n g a , H a i l u o t o , Jou t s e n o , P a r i k k a l a ) l i s ted i n He l l e n ( 1 953 ) ; SWE D E N : Vastergot l a n d , ST ! HUNGARY : l i l l af U red , B U ! ROMAN I A ( N a g y 1 967a ; 1 972 ) ; G E R MANY : B e l l i nc h e n ( O d er ) , HA ! AUSTR I A : Vos l a u ( N i eder-Osterre i c h ) , P ! F RA N C E : R a m i l l o n s , P ! E N GLA N D : W i m b l edon Common ( Su rrey ) , OX ! l i c h f i e l d D i s t r . ( C a r r 1 93 1 ) ; I TALY : Ros t a ( To r i n o ) , O l ! C h i v asso ( T o r i n o ) , Ol ! Verce l l i , O l !

2.

Tetrodontochelys s a k a ii ( E sak i a n d H a s h i moto )

n.

com b .

Mem . Amer .

1 442

En!.

I nst . ,

no .

37 ,

1 984

Epigon a top u s s a k a i i E s a k i a n d H a s h i moto 1 933 : 20 . Epigon a top u s s a k a ii E s a k i a n d H a s h i mo to : E s ak i a n d Moch i z u k i 1 94 1 : 2 7 .

F E MA L E : a p terou s ; l en g t h 2 , 50-2 , 75 m m ; h e a d b l ack , w i t h occ i p u t l i g h t ; fron t p a r t of v e r t e x , c l y p e u s a n d m a n d i b I e s y e l l ow ; a n t e n n a e t e s t aceo u s ; t hora x , p ropode u m , pet i o l e a n d abdomen b l ack ; l eg s y e l l ow ; h e a d w e a k l y exca v a ted , a l most f l a t , s h i n y , w i t h o u t scu l p t u re , except for occ i p u t g ra n u l a te d ; p ro n o t u m w e a k l y c rossed b y a t r a n s v e rs a l i m p res s i on , s h i n y , w ea k l y g r a n u l a ted ; scu t u m s h i n y , g r a n u l a t e d ; m e t a n o t u m t ra n s verse l y s t r i a t e ; m e t a t horax + propodeum s h i n y , g r a n u l a t e d ; p l e u r a w e a k l y t r a n s v e rs e l y s t r i a te ; pos t e r i o r s u rface of p ropod e u m t ra n s v e rs e l y s t r i a te on l y on s i des a n d on pos t e r i o r t h i rd ; segme n t 1 of fron t t a r s u s l on ge r t h a n segmen t 4 ( 1 1 : 9 ) ; e n l arged c l a w ( F i g . 996 ) w i t h o u t s u b a p i c a l too t h , w i t h a sm a l l too t h a t t h e e n d of a l on g i t u d i n a l f u r row a n d w i t h 3 b r i s t l es ; s e g m e n t 5 of fron t t a r s u s ( F'i g . 996 ) w i t h 1 row of 6 m i n u te l ame l l ae l o c a t e d on t h e d i s t a l h a l f ; a p e x w i t h a g roup of 6 l a me l l ae ; m ax i l l a r y p a l p i w i t h 2 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l spurs 1 , 0 , 1 . MALE : u n k n o w n LOCUS T Y P I C U S : O i t a ( K y u s h u , J a p a n ) ( ho l o t y p e ) ; S h i i d a-cho ( J a p a n ) ( n eo t y p e ) TYP I CA L MAT E R I A L : h o l o t y p e l o s t ; n e o t y p e F ! ( he r e b y des i g n a t ed )

in OL .

HOST S : i n J a p a n Nep hotettix b ip u n c t a t u s cincticeps U h l e r ( E sak i a n d H a s h i moto 1 93 3 ) a n d Nep h otettix apicalis ( Mo t s . ) ( G r i s t a n d L e v e r 1 969 ) . D I ST R I B UT I O N : P a l a e a rc t i c reg i on : JAPA N : K a m i - g u n ( Koch i P r ef . ) , N I ! O i t a ( K y u s h u ) ( E s a k i a n d H a s h i moto 1 933 ) ; Sh i i d a-cho ( C h i k u j o u - g u n , F u k uo k a Pref . ) , O L ! M U !

3.

Tetrodontochelys p u licarius ( K l u g ) n . com b .

Gon a top u s pu licarius K l u g 1 8 1 0 : 1 65 . Gona top u s bolivari C e b a l l os 1 927 : 1 03 ( n . sy n . ) ; locus typ icus S a n Rafael ( Sp a i n ) . Epigon a topoides p u l ic arius ( K l u g ) : R i c h a rd s 1 93 9 : 2 2 1 . Epigona topoides n o t a t u s Ponom a r e n ko 1 972 : 676 ( n . s y n . ) ; locus typicus M o u n t-Zamry n-U I a ( Mo n g o l i a) • Epl'gon a topoides p ulica rius ( K l u g ) : O l m i a n d C u rr a d o 1 97 5 : 6 1 . Neogona topoides c1a udiae A b d u l - N o u r 1 976 : 274 ( n . sy n . ) ; locus typ ic u s S I . Ge l y -d u - F e sc ( F r a n ce ) . Epigona topoides p u licarius ( K l u g ) : Ponomarenko 1 97 8 : 26 .

F E MA L E

( F i g . 9 97 ) :

a p te rou s ;

l e n g t h 2 , 00-2 , 25 m m ;

y e l l ow - te s t a c eo u s ,

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F e m a l e of Tetrodon tochelys p u licarius ( K l u g ) ( t y p e )

1 443

1 444

Fig.

Mem . Amer .

Ent.

I nst . ,

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998 -. C he l a of Tetrodon tochelys pulicarius ( K l u g ) F esc ( F r a n c e )

from 5 1 . Ge l y - d u

p e t i o l e b l ac k a n d a b domen b ro w n i s h ; a n te n n a l s e g m e n t s i n fo l l o w i n g p roport i o n s : 5 : 3 : 3 , 5 : 3 : 2 : 3 : 2 , 5 : 3 : 3 : 4 ; head s h i n y , smoo t h , exca v a t e d , w i t h o u t sc u l p t u re ; p ro n o t u m s h i n y , smoo t h , w i t h o u t scu l p t u re ,

Gon a topod i n ae

F i g . 999 - C he l a of Tetrodontochelys p se u doa udax n . sp .

1 445

( t y pe )

n o t crossed by a t r a n s v e rs a l i m p ress i on or weak l y i m p resse d ; s c u t u m s h i n y , smoo t h ; m e t a t horax + p ropodeum s h i n y , smoo t h ; pos t e r i or s u rface weak l y t r a n sverse l y s t r i a te ; segment 1 of f ro n t tarsus a s l on g as segment 4; e n l a rged c l a w ( F i g . 998 ) w i t h o u t s u b a p i c a l toot h , w i t h a s m a l l too t h at the e n d of a l on g i t u d i n a l f u r row and w i t h 3-4 b r i s t l es ; s e g me n t 5 of f ro n t t a rs u s ( F i g . 998 ) w i t h 1 r o w o f 9- 1 4 l ame l l a e ; a p e x w i t h a g rou p of 4-9 l am e l l ae ; m a x i l l a r y p a l p i w i t h 3-4 s e g me n t s ; l ab i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 .

Mem . Amer.

1 446

Ent.

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1 984

MALE : u n k no w n L O C U S TYP I C U S : Ger m a n y TYP I CAL MATER I AL : h o l o t y p e F ! o f G . pulicari u s i n HU ;

holotype F !

and 1

p a r a t y pe F ! of N. cla u di a e i n P ; h o l o t y p e F ! of G . bolivari i n M D ; ho l o t y p e F ! o f E . notatus i n LN . HOST S : i n F ra n ce A darrus taurus R i b a u t ( A b d u l -N o u r 1 976 ) . D I STR I B U T I O N : P a l a e a rc t i c reg i on : MONGOL I A : Mo u n t - Z a m r y n - U l a ( C e n t r a l a y m a g , Cen t r a l Mo n g o l i a ) , LN ! U . S. S . R . ( Ponom a re n k o 1 97 8 ) ; G E R M ANY : B e r l i n , H U ! F R A N C E : S ! . G e l y - d u - F esc ( He r a u l t ) , P ! SPA I N : S a n R a f a e l ( Segov i a ) , M D !

4 . Tetrodon toch elys pseudo a u da x n . sp .

F E MA L E : a p t e rou s ; l e n g t h 2 , 62 m m ; red d i s h - t e s t aceou s , w i t h pet i o l e b l ac k a n d a b domen b ro w n ; a n t e n n ae b ro w n , w i t h s e g m e n t s 1 -2 a n d prox i m a I reg i o n of s e g m e n t 3 test aceou s ; a n t e n n ae d i s t a l l y t h i ckened ; a n t en n a l segme n t s i n fo l l o w i n g p roport i on s : 6 : 2 : 4 : 6 : 4 : 4 : 4 : 3 , 5 : 3 , 5 : 4 ; head exca v a ted , s h i n y ; frons w i t h o u t sc u l p t u r e ; vertex and occ i p u t w e a k l y g r a n u l a t e d ; occ i p i t a l ca r i n a i n v i s i b l e ; POL = 1 ; OL = 1 ; OOL = 6 ; temp l e s d i s t i nc t ; fron t a l l i n e com p l e te ; p ro n o t u m n o t crossed b y a t r a n s v e rs a l i m p reSS i o n , d u l l , w e a k l y g r a n u l a t e d ; sc u t u m w ea k l y scu l p t u red b y l o n g i t u d i n a l s t r i a e , d u l l ; m e t a t horax + p ropodeum d u l l , w e a k l y g r a n u l a ted ; p l e u ra a n d pos t e r i o r s u rface of p ropodeum very w e a k I y t r a n sverse I y s t r i a te ; meso-m e t a p l eu r a l s u t u re o b so l e te ; fore t y a rs a l segmen t s i n fo l l ow i n g p roport i o n s : 9 : 2 : 3 : 1 0 : 1 5 ; e n l a rged c l a w ( F i g . 999 ) w i t h o u t s u b a p i c a l too t h , w i t h a sm a l l too t h a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d w i t h 5 peg- l i ke h a i rs ; s e g m e n t 5 of f ro n t t ar s u s ( F i g . 999 ) w i t h 1 row of 1 8 l a m e l l ae ; apex w i t h a grou p of 6 l a me l l ae ; m a x i l l a ry p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . M A L E : u n k no w n L O C U S TYP I C U S : B a j a m a r ( C a n a r y I s l a n d s ) TYP I CA L MAT E R I A L : ho l o t y p e F ! i n M D . HOST S : u n k n o w n D I S T R I B U T I O N : P a l aearc t i c reg i o n : CANARY I SLAN D S : B a j a m a r ( T e n e r i fe ) , MD!

5.

Tetrodon tochelys a egyp ti a c u s n . sp .

F E MA L E : a p t e ro u s ; l e n g t h 2 , 48-2 , 80 m m ; head b ro w n , w i t h c l y p e u s a n d m a n d i b l es l i g h t ; a n te n n a e b row n , w i t h segmen t s 1 -4 test aceo u s ; t ho r a x a n d propo d e u m b l ac k ; a b domen b row n ; l e g s b row n , w i t h fore t i b i ae a n d fore t a r s i tes t aceo u s ; a n te n n ae d i s t a l l y t h i c k e n e d ; a n t e n n a I segmen ts i n fol l ow i n g propor t i o n s : 7 : 5 : 8 : 6 : 4 , 5 : 4 , 5 : 4 : 4 : 4 : 5 ; head f l a t , d u l l ,

G o n a topod i n a e

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1 000 - C h e l a of Tetrodontochelys a egyp tiacus n . s p .

g r a n u l a ted ; 2 ; OOl = i m p ress i o n ; g r a n u l a ted ; t ra n s v e rsa l

1 447

( t y pe )

fron t a l I i n e comp l e te ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 1 ; O l 8 ; T l = 7 ; p ronot u m s h i n y , g r a n u l a t e d , w i t h o u t t r a n sversa l m e t a n o t u m t r a n s verse l y s t r i a t e ; m e t a thorax + p ropodeum d u l l , pos t er i or su rface of p ropodeum d i s t a l l y w i t h some t ra c k s of s t r i a e ; meso-me t a p l eu r a l s u t u re obso l e t e ; mesop l e u r a a n d m e t a p l e u ra w i t h some t r a c k s o f t r a n s v e rs a l s t r i ae ; fore t a rsa l segme n t s i n fo l l ow i n g p roport i o n s : 1 1 : 3 : 3 , 5 : 7 : 1 3 ; e n l a rged c l aw ( F i g . 1 000 ) w i t h o u t s u b a p i c a l toot h , w i t h a s m a l l too t h a t the e n d o f a l o n g i t ud i n a l f u r row and w i t h 1 row of 1 br i s t l e and 3 long l a m e l l ae ; s e g me n t 5 of f ro n t t a rs u s ( F i g . 1 00 0 ) w i t h 1 r o w of 6 l o n g l a me l l ae ; a p e x w i t h a g roup of 6 l o n g l a me l l ae ; m a x i l l a ry p a l p i w i t h 4 segmen ts ; l ab i a l p a l p i w i t h 2 segmen t s ; t i b i a l spurs 1 , 0 , 1 . MALE : u n k no w n l O C U S TYP I C U S : W a d i H o f ( E g y p t ) TYP I CAL MAT E R I A I_ : ho l ot y p e F ! i n W A ; 1 p a r a t y p e F ! i n O L . HOST S : u n k n o w n D I STR I B U T I ON : P a l a e a rc t i c reg i o n : E GY P T : W a d i Hof , WA ! W a d i D i g l a , OL ! =

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F E MAL E : a p t e ro u s ; l en g t h 2 , 37 m m ; head b row n , w i t h c l y p e u s , m a n d i b l es a n d a n t e r i o r reg i on of frons test aceou s ; a n t e n n a e b row n , w i t h s e g me n t s 1 -3 tes taceou s ; p ro t ho r a x b ro w n , w i t h p ro n o t u m l a tera l l y redd i sh ; mesothor a x , m e t a t ho r a x , p ropode u m , pet i o l e a n d a b domen b l ack ; l e g s b row n , w i t h t roch a n ters , t i b i ae a n d t a r s i test aceou s ; a n t e n n a e not d i s t a l l y t h i ck e n e d ; a n t e n n a I s e g m e n t s i n fo l l o w i n g p roport i o n s : 6 : 4 : 7 : 4 : 4 : 4 : 3 , 5 : 3 , 5 : 3 : 5 ; head s h i n y , e x c a v a te d , w i t h frons smoo t h a n d w i t h o u t scu l p t u re ; occ i p u t a n d vertex g r an u l a ted ; occ i p i t a l ca r i n a i n v i s i b l e ; p ro n o t u m d u l l , g r an u l a ted , not crossed b y a t r a n s v e rs a l i m p res s i o n ; sc u t u m w i t h l o n g i t u d i n a l k ee l s , w i t h o u t l a te r a l p o i n t s ; m e t a t horax + p ropode u m d u l l , g r a n u I a t e d ; poster i o r s u r f ac e of p ropodeum a n d p l e u r a not t r a n s v erse I y s t r i a t e or w i t h v e r y weak s t r i ae ; meso-met a p l e u r a l s u t u r e obso l et e ; fore t a rs a l s e g m e n t s i n fo l l ow i n g p ropor t i o n s : 9 : 1 , 5 : 3: 9 : 1 4 ; en l a r ged c l a w ( F i g . 1 00 1 ) w i t ho u t s u b a p i c a l too t h , w i t h a sm a l l too t h at the e n d of a l o n g i t u d i n a l f u r row a n d w i t h 5 p e g - l i k e h a i r s ; s e g m e n t 5 of fron t t a r s u s ( F i g . 1 00 1 ) w i t h 1 r o w of 2 1 l a me l l ae ; apex w i t h a g ro u p o f 8 l ame l l ae ; p rox i m a l reg i o n of segme n t 5 w i t h a shor t , i n n e r , serrate m a rg i n ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l spurs 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : T a ho d i o ( C a n a r y I s l a n d s ) TYP I CAL MAT E R I AL : h o l o t y p e F ! i n M D . HOST S : u n k n o w n D I ST R I B U T I ON : P a l a e a rc t i c reg i o n : C A NARY I SL A N D S : T a hod i o ( T e n er i fe ) , MD!

E T H I O P I AN S P E C I E S O F TETRODONTOCHEL YS

Seven s p ec i e s a re k n ow n .

K E Y T O T HE SP E C I E S

FEMALES

B od y y e l l ow- t e s t a c e ou s o r r ed d i s h , w i t h p e t i o l e b l a ck , a b d omen b r o wn i s h and a n t en n a e o c c a s i on a l l y p a r t l y b r own i s h . . . . . . . . . . . 2 B od y a t l ea s t p a r t l y b l a c k . . . . . . . . . . . . . . . . . . . . .4 B od y y e l l ow- t e s t a c e ou s , w i t h p e t i o l e b l a ck a n d o c c a s i on a l l y a b d omen b r own i s h l . ochre u s n . sp . B od y r e d d i sh , w i t h p e t i o l e b l a c k ; a b d omen a n d o c c a s i on a l l y p a r t l y a n t en n a e b r o wn i s h 3 A n t e r i o r s u r f a c e o f p r o p o d e u m s h i n y , smo o t h , w i t h ou t s c u l p t u r e .

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1 006 0 , E , F ) 5 5 Me t a t h o r a x + p r o p o d e u m s h i n y , w i t h ou t s cu l p t u r e 6 7 Me t a t h o ra x + p r op od e u m d u l I , g ra n u l a t ed 6 S c u t u m y e l l ow ; t h o ra x a n d p r op o d e u m r ed d i s h - b r own o r r ed d i s h -b l a ck 2 . obscurus n . s p . S c u t u m b l a ck ; t h o ra x a n d p r op od e u m b l a ck · . . . . . . • • • . . . . . . . • . . • . . . . • 5 . n igrior n . sp . 7 B od y mo r e r o b u s t ( F i g . 1 0 0 6 D ) ; s eg me n t 5 of f r on t t a r s u s l e s s s l end e r ( F i g . 1 00 8 ) 6 . a fer n . sp . B od y mo r e s l en d e r ( F i g . 1 0 0 6 F ) ; s eg m e n t 5 o f f r on t t a r s u s mo r e s l end e r ( F i g . 1 00 9 ) 7 . varipes ( B r u e s ) •

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MALES

Un known

1 . Tetrodontochelys ochreu s n . sp .

F E MA L E : a p t e ro u s ; l e n g t h 2 , 1 8 m m ; y e l l ow - t e s t aceo u s , w i t h pe t i o l e b l ac k ; head a l most f l a t , s h i n y , w i t h o u t scu l p t u re or weak l y g ra n u l a t e d ; p ro n o t u m v e r y w e a k l y t r a n sve rse l y i m p ressed , s h i n y , g r a n u l a ted ; m e t a t ho r a x + propo d e u m s h i n y , w i t hou t scu l p t u re ; p l e u r a a n d p o s t e r i o r s u rface of propode u m t r a n s verse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re o b so l e t e or s l i g h t l y v i s i b l e on pos t e r i o r h a l f ; s e g m e n t 1 of f ro n t t a r s u s as l on g a s segme n t 4 ; e n l arged c l a w ( F i g . 1 002 ) w i t ho u t s u b a p i c a l toot h , w i t h a s m a l l toot h a t the e n d of a l o n g i t u d i n a l fu rrow a n d w i t h some b r i s t l es ; s e g m e n t 5 of fron t t a r s u s ( F i g . 1 00 2 ) w i t h 1 r o w of 1 ( p rox i m a l ) + 1 4 l ame l l ae ( ex t e n d i n g b e y o n d 0 , 5 l e n g t h of segmen t ) ; a p e x w i t h a g ro u p of 1 1 l ame l l ae ; prox i m a l reg i on of s e g m e n t 5 w i t h a n i n ner , serrate m a rg i n ; max i l l ary p a l p i w i t h 3 s e g me n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k no w n L O C U S TYP I C U S : O k a h a n d j a ( Sou t h W e s t A f r i c a ) T Y P I CAL MAT E R I AL : h o l o t y pe F ! i n BM . HOSTS : u n k n o w n D I STR I B UT I O N : E t h i o p i a n reg i o n : S O U T H W E S T A F R I CA : O k a h a n d j a , B M !

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1 003 - C h e l a of Tetrodon tochelys obscurus n .

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F E MA L E :

Ent.

sp .

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Tetrodon tochelys obscurus n . sp .

a p terou s ; l e n g t h 2 , 1 2- 2 , 37 m m ; h e a d red d i s h - b ro w n , w i t h fron t p a r t of ver t e x , c l y p e u s a n d m a n d i b l es y e l l ow ; a n t e n n a e y e l l ow - b row n ; p ronot u m red d i sh-b ro w n ; scu t u m ye l l ow ; s c u t e l l u m a n d m e t a t ho r a x + p ropode u m re d d i s h - b row n or b l ac k ; a p e x of p ropodeum y e l l ow ; p e t i o l e b l ac k ; abdomen red d i s h - b ro w n ; l eg s y e l l ow , w i t h c l u b s o f fore femora

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1 454

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37 ,

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meso-m e t a p l e u r a l s u t u re obso l e t e ; fore t a r s a l seg me n t s i n fo l l ow i n g p ropor t i o n s : 8 : 2 : 3 : 7 : 1 3 ; en l a r ged c l a w ( F i g . 1 003 ) w i t h o u t s u b a p i c a l toot h , w i t h a sm a l l too t h a t t h e e n d o f a l o n g i t u d i n a l f u r row a n d w i t h 4-6 peg- l i k e h a i r s ; segment 5 of f ro n t t a r s u s ( F i g . 1 00 3 ) w i t h 1 row of 1 2- 1 5 l a m e l l a e ; apex w i t h a g r o u p o f 6-8 l a me l l ae ; m a x i I l a r y p a l p i w i t h 3 segme n t s ; l ab i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k n o w n LOCUS TYP I C U S : Mosse l B a y ( Sou t h Af r i ca ) TYP I CAL MAT E R I A L : ho l o t y p e F ! a n d 37 p a r a t y p es F F ! i n BM ; 7 p a r a t y p es F F ! i n OL . HOST S : u n k n o w n D I ST R I B U T I O N : E t h i o p i a n P rov i n ce ) , BM ! OL !

reg i o n : SO U T H A F R I CA :

Mosse l B a y

( C a pe

3 . Tetrodontochelys rubripes n . sp .

F E MA L E : a p te ro u s ; l e n g t h 3 , 56 m m ; redd i s h , w i t h p e t i o l e b l ack a n d a b domen b ro w n i s h ; head exca v a t ed , s h i n y , g r a n u l a ted ; p rono t u m not crossed b y a t r a n s v e rs a l i m p res s i on or weak l y i m p ressed , d u l l , w i t h o u t scu l p t u re or w e a k l y a l u t ac eo u s ; meso-me t a p l e u r a l s u t u re obso l e t e ; m e t a t ho r a x + p ropode u m d u l l , g r a n u l a t e d ; p l e u r a n o t t r a n s v e rse l y s t r i a t e ; pos t e r i o r s u rface o f p ropodeum w e a k l y t r a n s verse l y s t r i a te ; seg m e n t 1 of fron t t a rs u s as l o n g as s e g m e n t 4 ; en l a r g e d c l a w ( F i g . 1 004 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d o f a l on g i t u d i n a l f u r row and w i t h 5 peg- l i ke ha i rs ; segmen t 5 of fron t t a rs u s ( F i g . 1 004 ) w i t h 1 row of 1 6 l a me l l ae ; a pe x w i t h a g ro u p of 1 0 l am e l l ae ; m a x i I l a r y p a l p i w i t h 4 segme n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : L r . U m h l a t u z i R . ( Sou t h Afr i c a ) TYP I C A L MAT E R I A L : h o l o t y p e F ! i n B M . HOST S : u n k n o w n D I ST R I B U T I O N : E t h i o p i a n reg i on : SOU T H A F R I CA : ( Z u l u l a n d ) , BM !

Lr . Umh l atuzi R .

4 . Tetrodon toch elys z a irensis n . sp .

F E MAL E : a p tero u s ; l e n g t h 2 , 25 m m ; h e a d b row n , w i t h m a n d i b l es , c l y p e u s a n d f r o n t p a r t o f v e r t e x y e l l ow ; a n te n n a e b ro w n , w i t h s e g me n t s 1 -2 a n d p a r t o f segmen t 3 y e l l ow ; t ho ra x , p ropodeum a n d a b domen b l a ck ; l eg s y e l l o w , w i t h c l u b s o f femora a n d coxae b row n i s h ; h e a d ( F i g . 1 006 C ) s h i n y , exca v a te d , g r a n u l a ted ; p ronot u m ( F i g . 1 006 C ) not crossed b y a t r a n s v e rsa l i m p ress i on or w e a k l y i m p ressed , g r a n u l a ted ; scu t u m , sc u te l l u m a n d m e t a t ho r a x + p ropod e u m ( F i g . 1 006 C ) g r a n u l a t e d ; s c u t u m w i t h some

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k ee l s ; p l e u r a a n d pos t e r i o r s u rface o f propod e u m n o t t r a n s v e rse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re obso l e t e ; a n t e r i o r s u rface o f m e t a t horax + p ropode u m v e r y i nc l i ned ; fore t a rs a l segmen ts i n fo l l o w i n g p ropor t i o n s : 7 : 2 : 3 , 5 : 8 : 1 4 ; e n l a r g e d c l a w ( F i g . 1 005 ) w i t h o u t s u b a p i c a l too t h , w i t h a sm a l l tooth at t h e e n d of a l o n g i t u d i n a l f u rrow a n d w i t h 6 peg- l i k e h a i rs ; s e g m e n t 5 o f f ro n t t a rs u s ( F i g . 1 005 ) w i t h 1 row of 1 9 l a m e l l ae ; a p e x w i t h a g ro u p of at l e a s t 9 l a me l l ae ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C US : G a r a m b a N a t i o n a l P a r k TYP I CA L MAT E R I A L : h o l o t y p e F ! i n T V . HOST S : u n k n o w n

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Park , T V !

Tetrodon tochelys n igrior n . s p .

F E MA L E : a p terou s ; l e n g t h 2 , 1 8 m m ; h e a d b r o w n - b l ac k , w i t h c l y p e u s , m a n d i b l es a n d frons a l o n g orb i t s y e l l ow ; a n t e n n a e b row n , w i t h seg m e n t s 1 -2 y e l l o w ; t h o r a x , p ropod e u m , p e t i o l e a n d a b domen b l ac k ; l e g s t e s t a c eo u s , w i t h m i d a n d h i n d c o x a e b row n ; c l u b s of femora a l so b ro w n ; h e a d exca v a t e d , s h i n y , h a i r l e s s , w i t h o u t s c u l p t u r e ; p r o n o t u m s h i n y , n o t crossed b y a t r a n s v e rs a l i m press i on , w i t h o u t scu l p t u re , h a i r y ; sc u t U ITl s h i n y , w i t h some l o n g i t u d i n a l k ee l s ; m e t a t h o r a x + p ropodeum s h i n y , w i t h o u t sc u l p t u r e ; p l e u r a a n d p os t e r i o r s u rface of propode u m w i t h w e a k t r a n s v e rs a l s t r i a e ; meso-me t a p l e u r a l s u t u re o b so l e t e ; s e g m e n t 1 of fron t t a rs u s s l i g h t l y l on g e r t h a n segmen t 4 ( 8 : 7 ) ; e n l a r g e d c l a w ( F i g . 1 007 ) w i t h o u t s u b a p i c a l too t h , w i t h a sma l l too t h a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d w i t h 4 l ame l l ae ; s e g m e n t 5 of fro n t t a rs u s ( F i g . 1 007 ) w i t h 1 row of 1 8 l a me l l ae ; apex w i t h a g ro u p of 5 l a me l l a e ; p rox i m a l reg i o n of segmen t 5 w i t h an i n n e r , serrate m a rg i n ; m a x i l l a ry p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n known L O C U S T Y P I C U S : V a n R e e n e n ( So u t h Afr i c a ) T Y P I CA L MAT E R I AL : h o l o t y p e F ! a n d 1 p a r a t y pe F ! i n BM . HOST S : u n k n o w n D I ST R I B U T I O N : E t h i o p i a n reg i o n : SOU T H AF R I CA : V a n Reenen ( D r a k e n s Berg , N a t a l ) , BM !

6.

Tetrodo n tochelys a fer n .

sp.

F EM AL E : a p terou s ; l e n g t h 2 , 50-2 , 62 m m ; h e a d b ro w n - b l ac k , w i t h m a n d i b l es , c l y p e u s a n d f r o n t p a r t of vertex y e l l o w ; a n t e n n a e b row n , w i t h segme n t s 1 -2 y e l l o w ; pronot u m b ro w n - b l ac k , p a r t l y re d d i s h ; sc u t u m , sc u te l l u m a n d m e t a t ho r a x + propode u m b l ac k ; a b domen brow n - b l ac k o r b l ac k ; l eg s y e l l o w or y e l l ow - b ro w n , w i t h cox a e , c l u b s of femora a n d t i b i ae b row n i s h ; h e a d ( F i g . 1 006 D ) exca v a t e d , s h i n y , g r a n u l a t e d ; p ro n o t u m ( F i g . 1 006 D ) n o t crossed b y a t r a n s v ersa l i mp ress i on or w e a k l y i m p ressed , s h i n y , g r a n u l a t e d ; s c u t u m g ra n u l a t e d , w i t h some k ee l s ; s c u te l l u m smoo t h , sh i n y , w i t h o u t sc u l p t u re ; m e t a n o t u m not s t r i a t e ; m e t a t ho r a x + p ropodeum ( F i g . 1 006 D ) d u l l , g r a n u l a t e d , w i t h t rack of med i a n f u r row ; p l e u r a a n d p o s te r i o r s u rface o f propod e u m n o t t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re obso l e te or w e a k l y d i s t i n c t ; fore t a rs a l segmen ts i n fo l l ow i n g p ropor t i o n s : 9 : 2 : 3 : 9 : 1 6 ; en l a rged c l a w ( F i g . 1 008 ) w i t h o u t s U b a p i c a l toot h , w i t h a sma l l too t h a t t h e e n d o f a l o n g i t u d i n a l f u r ro w a n d w i t h 4-6 peg- l i k e h a i r s ; segmen t 5 of fron t t a r s u s ( F i g . 1 00 8 ) w i t h 1 row of 1 7-20

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7 . Tetrodontochelys varipes ( B r u e s ) n . com b .

Gon a top u s varipes B r ues 1 906a :

1 06 .

F E MAL E : a p tero u s ; l e n g t h 2 , 8 1 m m ; head a n d a n t e n n ae y e l l ow-red d i s h ; p ron o t u m b l ac k , w i t h s i des red d i s h ; s c u t u m a n d m e t a t ho r a x + p ropodeum

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b l ac k ; a b domen b row n ; l e g s y e l l o w ; h e a d ( F i g . 1 006 F ) e x c a v a t e d , s h i n y ; occ i p u t g r a n u l a t e d ; f ro n s w i t h o u t scu l p t u re ; p ro n o t u m ( F i g . 1 006 F ) s h i n y , g ra n u l a ted , n o t crossed b y a t r a n sversa l i m p ress i on ; scu t u m , s c u te l l u m a n d m e t a t ho r a x + p ropo d e u m ( F i g . 1 006 F ) d u l l , g r a n u l a ted ; p l eu r a n o t t r a n s verse l y s t r i a t e ; pos t e r i o r s u rface o f propodeum t r a n sverse l y s t r i a t e ; meso-m et a p l e u r a l s u t u re obso l et e ; fore t a rs a l seg m e n t s i n fo l l ow i n g p ropor t i o n s : 9 : 2 : 3 : 1 0 : 1 5 ; en l a r g e d c l aw ( F i g . 1 009 ) w i t h o u t s ub a p i c a l t oo t h , w i t h a s m a l l too t h a t t h e e n d o f a l o n g i t u d i n a l f u rrow a n d w i t h some b r i s t l es ; s e g m e n t 5 of fron t t a r s u s ( F i g . 1 009 ) w i t h 1 row of 26 l am e l l ae ; apex w i t h a g rou p of 8 l a me l l ae ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 s e g m e n t s ; t i b i a l s p u r s 1 , 0 , 1 . M A L E : u n k no w n L O C U S T Y P I C U S : M a s h on a l a n d ( Z i m b a b w e ) T YP I C A L MAT E R I AL : h o l o t y p e F ! i n M L . HOST S : u n k n o w n D I S T R I BU T I ON : E t h i o p i a n reg i o n : Z I MBABW E : M a shon a l a n d ( S a l i sb u ry ) , M L !

O R I E N T A L S P E C I ES OF TETRODONTOCHE L YS

O n e spec i e s

is k n ow n .

Tetrodontochelys l u cens n . s p .

Neogon a top u s lun a tu s ( K l ug ) : Mocz a r 1 979 : 83 .

F E M A L E : a p tero u s ; l e n g t h 2 , 37-3 , 1 2 m m ; red d i s h - t e s t aceous ( d a r k or l i g h t ) , w i t h p et i o l e b l ac k a n d occ a s i on a l l y abdomen a n d v e r t e x b ro w n i s h ; a n te n n a I segmen t s i n fo l l o w i n g proport i on s : 6 : 4 : 7 : 5 : 5 : 5 : 5 : 4 : 4 : 6 ; h e a d e xc a v a t e d , s h i n y , smoo t h , w i t h o u t scu l p t u re , e x c e p t for occ i p u t g ra n u I a te d ; p ro n o t u m not crossed b y a t r a n s v e rs a I i m press i o n , s h i n y , smoot h , w i t h o u t sc u l p t u re ; sc u t u m s h i n y , w i t h some l o n g i t u d i n a l kee l s ; metathorax + p ropodeum s h i n y , w i t h o u t scu l p t u re ; a n t e r i or s u rface w i t h a pos t e r i or s t ro n g med i a n f u rrow ; occ a s i on a l l y w i t h t r a c k of m ed i a n f u r row ; s u rface t r a n s v erse l y s t r i a t e ; p l e u r a n o t t r a n s verse l y s t r i a t e or weak l y s t r i a te ; meso-me t a p l e u r a l s u t u re obso l e t e ; fore t a rs a l seg m e n t s i n fol l ow i n g propor t i o n s : 1 0 : 2 : 3 : 1 0 : 1 4 ; e n l a r g e d c l a w ( F i g . 1 0 1 0 ) w i t h o u t su b a p i c a l too t h , w i t h a s ma l l too t h a t t h e e n d o f a l o n g i t u d i n a l f u rrow a n d w i t h 5 peg- l i ke h a i rs ; seg m e n t 5 of fron t t a r s u s ( F i g . 1 0 1 0 ) w i t h 1 row of 1 6-20 l a m e l l ae ; apex w i t h a g ro u p of a t l e a s t 8- 1 0 l ame l l ae ; prox i m a l reg i on of segmen t 5 w i t h an i n n e r , serrate m a rg i n ; m a x i l l a r y p a l p i w i t h 3-4 segmen t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE :

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LOCUS TYP I C U S : Pek a l o n g a n ( J a v a , I n dones i a ) TYP I C A L M AT E R I AL : ho l ot y p e F ! i n B ; 4 p a r a t y p es F F ! i n VA ; 2 p a r a t y p es F F ! in B U ; 1 p a ra t y p e F ! in O L . HOST S : i n S a r a w a k Nep h o tettix virescens ( D i s t a n t )' , Nephotettix n igrop ic t u s ( S t a l ) a n d Recilia dorsa li s ( Mo t s c h u l sk y ) . D I ST R I B U T I O N : O r i e n t a l reg i o n : I N D O N E S I A : Pek a l o n g a n ( J a v a ) , B ! O L ! MALAYS I A : P a p a r ( Sa b a h ) , YA ! T u a r a n ( Sa b a h ) , VA ! P a y a h ( P a l o h , 1 s t D i v i s i o n , S a ra w a k ) ( Mocz a r 1 979 , u n der n a me Neogo n a topus l u n a t u s K l u g ) , B U ! P H I L I P P I N E S : P i l i ( C a m a r i n e s Su r ) , YA !

N E A R C T I C S P E C I ES OF TETRODONTOCHEL YS

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t a r s u s w i t h a sh o r t row of 2 - 7 m i n u t e ha l f ( F i g . 1 0 1 1 ) • • • • • • • • . • • • • • • . • • • • • • • • • • • • • • • • • • • . . . • . • • . . • . . . . . . . . . . 1 . plesius ( F e n t on ) S eg m e n t 5 o f f r o n t t a r s u s w i t h 1 - 2 l o n g r o w s o f l on g l a me l l a e (F i gs 1 01 4 , 1 0 1 6 ) • • • . . • • . • . . • • • . . • • • • . • • • • • . . • • • • • • • . • . • • • • • • • 2 Me s o s oma mo s t l y o'r f u l l y b l a c k . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . • • . • • . • • • • • • • • • • • • • • . • • • • • • • 3 . peculiaris ( B r u e s ) Me s o s oma f u l l y o r mo s t I y t e s t a c e o u s o r r ed d i s h • • • • • • . • . • • • • • • • . . • • . . • • • • . • • . . • • • . • • • • • . • • • 2 . unicus ( R . C . L . P e rk i n s )

MALES

N o t a u l i c e s c omp l e t e ; a n t en n a l s e g m e n t 3 mo r e t h a n t w i c e a s l on g a s s eg me n t 2 ; p r o p o d e u m r e t i cu l a t e r u g o s e ; d o r s a l p r o c e s s o f g on o f o r c ep s l on g a n d s l e n d e r ( F i g . 1 0 1 3 ) • • • • • • • • . • • • • • • • • • • • • • • • . • • . . • • . • • • • • • • • • • • • • • • 1 . plesius ( F en t on ) N o t a u l i c e s i n c omp l e t e ; a n t en n a l s eg m e n t 3 l e s s t h a n t w i c e a s l on g a s s eg m e n t 2 ; p r o p o d e u m g ra n u l a t e d , n o t r e t i c u l a t e ru g o s e ; d o r s a l p r o c e s s o f g on o f o r c ep s d i f f e r en t I y s h a p ed ( F i g . 1 0 1 5 ) . . . . . . . . . . . . . . . . . . . . . . . 2 . u n i c u s ( R . C . L . P e rk i n s )

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n . eme n d .

Epigon a topus plesuis Fen ton 1 92 7 : 9 (plesuis i s p r i n t i n g-error for pl e s ius : plesius i s w r i t te n o n t h e type l oc a l i t y l ab e l ) .

F E MAL E : a p terou s ; l en g t h 2 , 00-2 , 8 1 m m ; h e a d b ro w n or b l ack , w i t h m a n d i b l es , c l y p e u s a n d occa s i on a l l y front p a r t of vertex y e l l o w ; a n t e n n ae y e l l o w , occ a s i o n a l l y w i t h segme n t s 4- 1 0 b row n ; p ro n o t u m b l ac k or b row n ; s c u t u m , scu te l l u m , m e t a t ho r a x + p ro p o d e u m a n d p et i o l e b l ack ; a b domen b row n - b l ack o r b l ac k ; l eg s y e l l o w , w i t h c l u b s of femora , m i d and h i n d coxae b row n i s h ; h e a d w e a k l y exca v a ted , s h i n y , w e a k l y p u n c t a te , w e a k l y g r an u l a ted ; p ro n o t u m not crossed b y a t r a n s v e rs a l i m p re s s i o n or weak l y i mp re s se d , s h i n y , g r a n u l a ted ; s c u t u m d u l l , s t r on g l y g r a n u l a ted ; m e t a n o t u m l a te ra l l y w i t h two p o i n ts , dorsa l l y w i t h a sem i c i rc l e- s h a ped k ee l ; m e t a t horax + p ropodeum d u l l , g r a n u l a ted ; p l e ura not t r a n sverse l y s t r i a t e ; p os te r i or s u rface o f p ro p o d e u m tra n sverse l y s t r i a t e ; occ as i on a l l y not s t r i a t e ; meso-me t a p l eu r a l s u t u re o b so l e t e ; segment 1 of fron t tarsus as l o n g a s segment 4; en l a r g e d c l a w ( F i g . 1 0 1 1 ) w i t ho u t s u b a p i c a l toot h , w i t h a s m a l l too t h a t t h e e n d o f a l o n g i t u d i n a l fu rrow a n d w i t h 3 p e g- l i k e h a i rs ; s e g m e n t 5 of f ro n t t a r s u s ( F i g . 1 0 1 1 ) w i t h 1 row of 5-7 m i n u te l a me l l ae on d i s t a l h a l f ; a p e x w i t h a g ro u p of 7-9 l ame l l ae ; m a x i l l a r y p a l p i w i t h 2-4 seg m e n ts ;

l a b i a l p a l p i w i t h 2 segmen t s ;

t i b i a l s p u rs 1 , 0 , 1 .

1 466

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1 0 1 3 - M a l e g en i t a l i a of Tetrodon toch elys plesius C h a t h a m ( O n t a r i o ) ( r i g h t h a l f remov ed )

( F e n ton ) from

M A L E ( F i g . 1 0 1 2 ) : f u l l y w i n ged ; l e n g t h 2 , 0-2 , 5 m m ; b l ac k ; m a n d i b l es test aceo u s ; a n te n n ae a n d a b domen b row n ; l eg s b ro w n , n o t b a n de d ; a n t e n n a e n o t d i s t a l l y t h i c k ened ; a n ten n a l s e g m e n t s i n fo l l o w i n g p ropor t i o n s : 4 : 4 : 1 0 : 1 0 : 9 : 8 : 8 : 7 : 6 : 1 0 ; a n te n n a l s e g m e n t 3 more t h a n t h ree t i mes a s l o n g a s b ro a d ( 4 , 4 ) ; h e a d s h i n y , g r a n u l a t e d ; fron t a l l i ne a b s en t ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L � 6 ; OL � 3 ; O O l � 3 ; b r e a d t h of t h e oce l l i : 3 ; temp l es d i s t i nc t ; s c u t u m , sc u te l l u m a n d m e t a n o t u m s h i n y , g ra n u l a t e d ; n o t a u l i ces com p l e t e , p o s t er i or l y s e p a r a te d ; m i n i m u m d i s t a n c e b et ween t h e n o t a u I i ces s h o r t e r t h a n b re a d t h of t he oce l l i

(2:3) ;

p ropodeum

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g ran u l a ted , re t i c u l a t e r u g ose ; dors a l s u rface w i t h o u t med i an f u rrow ; fore w i n g h y a l i n e , w i t h o u t d a r k t r a n s v e rs a l b a n d s ; d i s t a l p a r t of r a d i a l v e i n l on g e r t h a n p rox i m a l p a r t ; dorsa l p rocess o f gon oforce p s ( F i g . 1 0 1 3 ) l o n g a n d s l e n d e r ; b a s i vo l se l l a w i t ho u t r i d g e ; m a x i I l a r y p a l p i w i t h 3-4 segme n t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u r s 1 , 1 , 2 . LOCUS T Y P I C U S : E l k Po i n t ( 5 . D . , U . S . A . ) T Y P I CAL MAT E R I AL : h o l o t y p e F ! a n d 2 p a r a t y p es F F ! i n WA .

Mem . Amer . E n t .

1 468

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n o . 37 ,

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HOSTS : in C a n a d a and U . S. A. Macrosteles fa scifrons St a l ; in C a n a d a a l so Macrosteles divis u s . D I ST R I B U T I ON : N e a rc t i c reg i o n : CANADA : R e i n deer Depot ( M acke n z i e De l t a , N . W . T . ) , O T I a L I V a ncou ver ( B . C . ) , a T ! V i n e l a n d ( a n t . ) , a T ! O L ! O t t a w a ( O n t . ) , aL l OT ! C h a t h a m ( O n t . ) , a L I W A ! U . S . A . : E l k Po i n t ( 5 . D . ) , W A ! U n i vers i t y F a rm ( R amsey C o . , M i n n . ) , W A I Read ( D . C . ) , a L I a T ! Oswego C o . ( N . Y . ) , W A !

2.

Tetrodontochelys unicus ( R . C . L . Perk i n s ) n . com b .

Cha lcogon a top u s unicus R . C . L . Perk i n s 1 90 7 : 25 . Gon a top u s haploth ora x R . C . L . P e r k i n s 1 90 7 : 35 ( n . s y n . ) ; locus typicus N og a l es ( A r i z . , U . S . A . ) . Pachygona top u s m in im u s F e n ton 1 92 7 : 6 ( n . s y n . ) ; locus typicus E l k Po i n t ( S . D . , U . S . A . ) . Pachygona top u s min imus F e n t o n : B a rret t , W e s t d a I a n d R i c h a rd �o n 1 965 : 2 1 6 .

F E MAL E :

a p terou s ;

l en g t h 2 , 00-4 , 02 m m j redd i s h- test aceou s ,

b l ac k ; occ a s i on a l l y a b do m e n b row n ;

w i th pet i o l e

a n t e n n a e testaceous , occas i o n a l l y w i t h

segmen t s 4-1 0 or 5-1 0 b row n ; some t i mes a n ten n a l segme n t s 6-1 0 b row n ; a n te n n ae d i s t a l l y t h i ckened ; a n t en n a I segmen t s i n fo l l ow i n g p ropor t i o n s : 6 : 4 : 8 : 4 : 4 : 3 , 5 : 3 , 5 : 3 , 5 : 3 , 5 : 5 , 5 ; h e a d ( F i g . 1 006 A ) f l a t o r w e a k l y exc a v a t e d , s h i n y , w i t h o u t scu l p t u re or weak l y g ra n u l a ted ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 1 , 5 ; O L = 1 , 5 ; O O L = 6 , 5 ; p ro n o t u m ( F i g . 1 00 6 A ) s h i n y , not crossed b y a transvers a l i m p ress i o n o r w e a k l y i m pressed , v e r y w e a k l y g ra n u l a ted ; sc u t u m g r a n u l a t e d , l a tera l l y w i t ho u t p o i n t s ; m e t a n o t u m , met a t horax + p ro p o d e u m ( F i g . 1 00 6 A ) a n d p l e u r a w e a k l y g r a n u l a ted ; post e r i o r s u rface of p ropodeum s t ron g l y , f u l l y or p a rt l y t r a n s v erse l y s t r i a te ; p l e ura not t r a n sverse l y s t r i a t e ; meso- me t a p l eu r a l s u t u re obso l e te ; fore t a rsa l segments i n fo l l o w i n g p roport i o n s : 9 : 2 : 3 : 1 0 : 1 5 ; en l a r g e d c l aw ( F i g . 1 0 1 4 ) w i t ho u t s u b a p i c a l toot h , w i t h a sm a l l too t h a t t h e e n d of a l o n g i t u d i n a l fu rrow and w i t h 2-6 peg- l i ke h a i rs ; segment 5 o f fron t t a r s u s ( F i g . 1 0 1 4 ) w i t h 1 row of 1 4- 2 1 l a m e l l ae or w i t h 2 rows B.f 3- 1 6 l ame l l ae ; a p e x w i t h a g r o u p of 7 - 1 4 l a me l l a e ; m a x i l l a ry p a l p i w i t h 2-5 segme n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : f u l l y w i n ge d ; l e n g t h 1 , 5-2 , 0 m m ; b l ac k ; m a n d i b l es tes taceo u s ; a n ten n ae b ro w n ; l e g s b a n ded ; a n ten n ae not d i s t a l l y t h i ckened ; a n t e n n a I segmen t s i n fo l l o w i n g p roport i o n s : 4 : 3 , 5 : 5 : 4 , 5 : 4 , 5 : 4 , 5 : 4 : 4 : 3 , 5 : 5 , 5 ; a n t e n n a I segment 3 l es s t h a n t h ree t i mes a s l o n g a s b ro a d ( 2 , 5 ) ; h e a d s h i n y or d u l l , w e a k l y g r a n u l a ted ; f ro n t a l l i n e a b sen t ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 4 ; O L = 3 ; OOL = 4 , 5 ; b re a d t h of t h e oce l l i : 3 ; scu t u m , scute l l um a n d met a n o t u m s h i n y , g r a n u l a te d ; n o t a u l i ces i n com p l ete , d i s t i nc t or w e a k l y v i s i b l e , reac h i n g a p p rox i m a t e l y 0 , 5 l e n g t h of scu t u m ;

Gon a topod i n ae

Fig.

1 0 1 5 - M a l e gen i t a l i a of Tetrodon tochelys u n ic u s ( R . C . L . from P a s a d e n a ( I d a h o ) ( l e f t h a l f removed )

1 469

Perk i n s )

p ropod e u m d u l l , f u l l y g r a n u l a t e d ; dors a l s u rface w i t h a s t ro n g med i a n f u rrow ; fore- w i n g h y a l i ne , w i t h o u t d a r k t r a n s ve rs a l b a n d s ; d i s t a l p a r t o f r ad i a l ve i n l on ger t h a n p ro x i m a l p a r t ; gonoforceps ( F i g . 1 0 1 5 ) w i t h a b a n d- s h a p e d dorsa l mem b r a n o u s p rocess ; b a s i v o l se l l a w i t h o u t r i d ge ; m a x i l l a r y p a l p i w i t h 3 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . L O C U S TYP I C US : Nog a l es ( A r i z . , U . S . A . ) TYP I CA L M A T E R I A L : h o l o t y p e F ! of Ch . unicus i n B ; l ec t o t y p e F ! a n d 3 p a r a l ec t o t y p e s F F ! ( des i g n a t e d b y O l m i 1 982 ) of G. haplothorax i n B ;

1 470

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Ent .

I n st . ,

no .

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h o l o t y p e F ! of P. minim u s i n WA . HOST S : i n U . S . A . E u tettix tenella ; i n C a n a d a Nacrosteles fascifrons ( S t a l ) ( B a r re t t , Wes t d a l a n d R i c h a r d son 1 965 , u n der n ame Pachygona top u s

m in im u s ) . D I STR I BU T I ON : Nearct i c reg i o n : C A N AD A : So d a C reek ( B . C . ) , O T ! L a n d i n g ( Sas k . ) , OT ! B i rc h H i l l s ( S as k . ) , aT ! OL ! F i scher B r a n c h ( M a n . , B a r r e t t l e g . ) , O T ! W i n n i pe g ( M a n . , B a r r e t t l e g . ) , a T ! O L ! Ste . A g a r h e ( M a n . , B a rr e t t l e g . ) , O T ! M a rmora ( O n t . ) , OT ! U . S . A . : B a t t l es n a k e R i d g e ( Be n to n C o . , W a sh . ) , WA ! P a s a d e n a ( I d a ho ) , WA r No g a l es ( A r i z . ) , B ! Mon t a n a ( B a r re t t , Wes t d a l a n d R i c h a rdson 1 965 ) ; B i g B e n d N . P . ( T e x . ) , O T ! 29 . 1 m i . N of M i nera l W e l l s ( J ack Co . , T e x . ) , BY ! 9 . 9 m i . N of S tephen v i l i e ( E r a t h Co . , T e x . ) , O L ! North D a ko t a ( B a rre t t , Wes t d a l a n d R i c h a r d son 1 965 ) ; F a rgo ( N . D . ) , W A r E l k Po i n t ( S . D . ) , VIA ! M i n n eso t a ( B a rret t , Wes t d a l a n d R i c h a rdson 1 965 ) .

3.

Tetrodontochelys peculiaris ( B r u e s )

Gona top u s peculiaris B r u e s 1 903 : 1 25 . A gona top u s heterothorax R . C . L . Perk i n s 1 907 : 3 2 ( n . s y n . ) ; loc u s typ icus Nog a l es ( A r i z . , U . S . A . ) . Gona top u s peculiaris Brues : M u esebeck a n d W a l k l e y 1 95 1 : 1 038 .

F E MA L E : a p te rou s ; l e n g t h 3 , 1 2-3 , 66 m m ; head red d i s h - t e s t aceo u s , occ a s i o n a l l y w i t h v e r t e x d a r k e n e d ; a n te n n a e b row n , w i t h segme n t s 1 -3 test aceo u s ; p ronot u m b l ac k , w i t h s i des red d i s h ; occ a s i o n a l l y poster i o r m a r g i n re d d i s h ; mesot ho r a x , m e t a t ho r a x a n d p ropodeum b l ack , w i t h a p e x o f propode u m redd i s h ; a b d o m e n b l ac k , occ as i o n a l l y p a rt l y red d i s h ; l e g s test aceo u s ; h e a d weak l y e x c a v a t e d , a l most f l a t , s h i n y , weak l y g r a n u l a ted ; prono t u m not crossed b y a t r a n svers a l i m p ress i on o r weak l y i m pressed , s h i n y , g r a n u l a t e d ; scu t u m s h i n y , w i t h SOme l o n g i t u d i n a l k ee l s ; m e t a n o t u m s h i n y , t r a n sverse l y s t r i a t e ; m e t a t horax + propode u m s h i n y , g ra n u I a t e d ; p l e u ra a n d pos t e r i o r s u rf a ce of p ropodeum t r a n sverse I y s t r i a t e ; meso-m e t a p l e u r a l s u t u re obso l ete ; seg m e n t 4 of fron t t a r s u s a p p rox i m a t e l y as l o n g as s e g me n t 1 ; e n l a r g e d c l a w ( F i g . 1 0 1 6 ) w i t h o u t s u b a p i c a l toot h , w i t h a s m a l l too t h a t t h e e n d o f a l on g i t u d i n a l f u rrow and w i t h 5-7 peg- l i k e h a i rs ; seg men t 5 of f ro n t t a rs u s ( F i g . 1 0 1 6 ) w i t h 2 rows of 22-30 l a me l l ae or w i t h 1 row of 9-1 5 l a m e l l ae ; a p ex w i t h a g ro u p of 6 - 1 2 l a m e l l a e ; occ as i on a l l y segmen t 5 w i t h l a m e l l ae on l y on d i s t a l h a l f ; m a x i I l a ry p a l p i w i t h 4-5 s e g me n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n LOCUS T Y P I C US : A u s t i n ( T e x . , U . S . A . ) TYP I CAL MAT E R I AL : l ec t o t y p e F ! ( h ereby de s i g n a t e d ) of G . peculiaris i n C M ; 1 p a r a l ec to t y p e F ! of G . peculi aris i n W A ; h o l o t y p e F ! of A .

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1 47 1

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1 0 1 7 - C he l a of Tetrodon tochelys anom a l u s ( R . C . L . P e rk i n s ) from O a h u ( Ha wa i i a n

I s l ands)

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1 473

heterothorax in B . HOST S : i n U . S . A . u n k n o w n Cieadellida e . D I STR I BU T I ON : N e a rc t i c reg i o n : CANADA : P e l i c a n l a k e ( N i n e t t e , M a n . ) , OT ! F i n l a n d ( O n t . ) , O l ! U . S . A . : S t a n U ( C a l i f . ) , Ol ! Nog a l es ( A r i z . ) , B ! A u s t i n ( T ex . ) , WA ! C M ! 1 1 . 2 m i . S of lev e l l a n d ( Hock l e y Co . , T ex . ) , BY ! S i o u x C i t y ( I o w a ) , WA ! C l emson ( S . C . ) , WA ! accord i n g to F o u t s ( 1 93 6 ) t h i s s p ec i es occ u rs a l so i n H a w a i i a n I s l a n d s ( Ho n o l u l u ) ( u n der n ame A gon a top u s heterothora x ) : record certa i n I y erroneo u s .

AUSTRAL I AN S P E C I E S O F TETRODONTOCHEL YS

O n e s p ec i es i s k n ow n .

Tetrodontoehelys a nomalus ( R . C . l .

Gon a top u s anomala R . C . l . Perk i n s 1 9 1 2 :

Perk i n s ) n . com b .

14.

F E MALE : a p terou s ; l en g t h 2 , 1 2-2 , 8 1 m m ; red d i s h , w i t h p e t i o l e b l ac k ; occa s i o n a l l y a b do m e n b row n ; head ( F i g . 1 006 B ) s h i n y , w i t h o u t sc u l p t u re , excav a t e d ; p ro n o t u m ( F i g . 1 006 B ) not crossed b y a t r a n s v e rs a l i m p res s i o n or w e a k l y i m p ressed , s h i n y , w i t ho u t sc u l p t u r e ; sc u t u m s h i n y , w i t h some k e e l s ; meta thorax + p ro p o d e u m ( F i g . 1 006 B) s h i n y , w i t h o u t s c u l p t ure ; p l eu r a a n d p o s te r i o r s u rface of p ropodeum t r a n s v e rse I y s t r i a t e ; s e g m e n t of f ro n t t a r s u s s l i g h t l y l o n g e r t h a n s e g m e n t 4 ( 1 0 : 9 ) ; e n l arged c l a w ( F i g . 1 0 1 7 ) w i t ho u t s u b a p i c a l toot h , w i t h a s m a l l too t h a t the e n d of a l o n g i t u d i n a l f u rrow a n d w i t h 3-5 b r i s t l es ; s e g m e n t 5 of f ro n t t a rs u s ( F i g . 1 0 1 7 ) w i t h 1 ro w o f 1 5- 2 1 l a m e l l ae ; a p e x w i t h a group o f 8- 1 0 l a me l l ae ; m a x i l l a r y p a l p i w i t h 3-4 segmen ts ; l ab i a l p a l p i w i t h 2 seg men t s ; t i b i a l spurs 1 , 0 , 1 . MAL E : u n k n o w n lOCUS TYP I CU S : F i j i I s l a n d s T Y P I CAL MAT E R I AL : l ec t o t y p e F ! ( des i g n a t e d b y O l m i 1 982 ) i n B . HOSTS : u n k n o w n D I S T R I BU T I O N : A u s t r a l i a n reg i o n : F I J I I SLAN D S , B ! HAWA I I A N I SLAN D S : O a h u , WA ! B ! O l ! P a l o l o ( O a h u ) , WA ! Hopa u n a u ( Ha w a i i ) , B !

14.

Trichogo n a top u s K i effer

Triehogon a top u s K i effer 1 909 : 336 . nee Triehogona top u s H e l l e n 1 930 : 2 .

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Perk i n s ) from

T Y P E S P E C I E S : Tric hogona topus rubriceps K i effer 1 909 , hereby d es i g n a t ed . F E MA L E ( F i g s 20 , 1 02 7 ) : a p tero u s ; m a x i I l ar y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 s e g me n t s ; p ro n o t u m ( F i g . 1 02 1 C ) not c rossed b y a t r a ns v e rs a l i m p ress i o n or w e a k l y i m p ressed ; en l a r g e d c l a w ( F i g . 1 0 1 8 ) w i t ho u t s u b a p i c a l too t h , w i t h o r w i t ho u t a sm a l l too t h a t t he e n d o f a l o n g i t u d i n a l f u r row a n d w i t h or w i t ho u t b r i s t l es or l ame l l ae or p e g - I i ke h a i rs ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n D I ST R I B U T I O N : N e a rc t i c , Neotro p i c HOSTS : u n k n o w n SPEC I ES : 8

Gona topod i n ae

1 47 5

N EA R C T I C SP E C I E S OF TRICHOGONA TOPUS

O n e spec i e s i s k n ow n .

Trichogon a top u s simula tor ( R . C . L . Perk i n s ) n . com b .

Chalcogon a top u s s im u l a tor R . C . L . Perk i n s 1 907 : 2 6 . Cha lcogon a top u s koe b elei R . C . L . Perk i n s 1 907 : 26 ( n . s y n . ) ; locus typicus Nog a l es ( Ar i z . , U . S. A . ) .

F E MALE : a p t e ro u s ; l e n g t h 4 , 94-5 , 87 m m ; fer r u g i n o u s-red d i s h , w i t h a n t e n n a l segmen t s 6-8 o r 7- 1 0 brow n i s h ; occ a s i o n a l l y a n t en n a l segmen t 1 0 w h i t i s h ; pe t i o l e b l ack ; occ a s i o n a l l y a b domen p a r t l y b l ac k ; a n te n n a I s e g me n t s i n f o l l ow i n g p roport i o n s : 1 1 : 7 : 29 : 1 6 : 1 2 : 1 1 : 9 : 8 : 7 : 9 ; head excav a t e d , s h i n y , g r a n u l a t e d ; p ro no t u m not crossed by a t r a n s v e rsa l i m p res s i o n or w e a k l y i m p ressed , g r a n u l a t e d ; sc u t u m , sc u t e l l u m and m e t a t ho r a x + propode u m g r a n u l a ted ; pos t e r i or s u rface of propodeum t r a n s verse l y s t r i a t e ; meso-me t a p l e u ra l s u t u re o b so l e te ; seg m e n t 4 o f f ro n t t a r s u s as l on g a s segme n t 1 ; en l arged c l a w ( F i g . 1 0 1 8 ) w i t h o u t s u b a p i c a l toot h , w i t h a sm a l l tooth a t t h e e n d of a l o n g i t u d i n a l fu rrow a n d w i t h 3-5 l a m e l l ae ; seg m e n t 5 of f ro n t t a r s u s ( F i g . 1 0 1 8 ) w i t h some ro w s of 2 1 -39 l a me l l ae ; a p ex b e n t a t r i g h t a n g l e , w i t h a group o f a t l e a s t 1 5-20 l a me l l ae ; m a x i l l ary p a l p i w i t h 6 segme n t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MA L E : u n k n o w n LOCUS T Y P I C US : Nog a l es ( A r i zo n a , U . S . A . ) TYF' I CAL MATE R I AL : l ec t o t y p e F ! a n d 1 p a r a l ec t o t y p e F ! ( des i g n a ted b y O l m i 1 982 ) o f Ch . koebelei i n B ; l ec t o t y p e F ! a n d 1 p a r a l ecto t y p e F ! ( des i g n a te d b y O l m i 1 982 ) of Ch . simulator i n B . HOST S : u n k n o w n D I ST R I B UT I O N : Nearc t i c reg i o n : U . S . A . : N og a l es ( Ar i z . ) , B ! WA ! S . A n to n i o ( Tex . ) , W A ! M E X I C O : T a m a z u n c h a l e ( S a n L u i s Potos] ) , CA l

N E O T RO P I C SP E C I E S OF TRICHOGONA TOPUS

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S t a l k b e t we en p r on o t u m a n d me t a t h o ra x + p r op od e u m v e ry l on g , s l e n d e r , c omp o s ed o f s c u t u m , s c u t e l l u m a n d me t a n o t u m , a s l on g a s me t a t h o ra x + p r op od eum ( F i g . 1 0 2 3 A ) . . . . 3 . p a lliditarsis ( C a m e r o n ) S t a l k b e t w e en p r on o t u m a n d me t a t h o r a x + p r op o d e u m v e ry s h o r t a n d b r oa d ( F i g . 1 0 2 7 ) ; me t a t h o ra x + p r op od e u m n ea r l y t wo - t h r e e ·

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1 . richardsi n . sp . Me s o -me t a p l e u ra l s u t u r e ob s o l e t e ; s eg m e n t 5 of f r on t t a r s u s w i t h a p e x b en t a t r i g h t a n g l e ( F i g s 1 0 2 2 , 1 0 2 4 , 1 0 2 5 , 1 0 2 6 , 1 02 8 , 1 02 9 ) E n l a r g ed c l a w w i t h o u t l a me l l a e o r b r i s t l e s o r p eg - l i k e h a i r s , w i t h a r o w o f t e e t h on p r o x i ma l r eg i o n o f i n n e r s i d e ( F i g . 1 0 2 2 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 . hispidus n . sp . E n l a r g ed c l a w w i t h o u t r o w s o f t e e t h on p r o x i ma l r eg i on o f i n n e r s i d e , w i t h 1 r o w o f l a me l l a e o r p eg - I i k e h a i r s o r •

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s e gmen t 5 o f f r on t t a r s u s (F i g . 1 020 ) . . . . . . . . . . . . . .

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1 477

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p ropode u m a n d a bdomen b l ac k ; l eg s red d i s h , w i t h coxae p a r t l y b l ac k a n d c l ub s of femora p a r t l y b ro w n i s h ; a n te n n a l s e g me n t 3 n e a r l y a s l o n g a s s e g m e n t 1 ; h e a d ( F i g . 1 0 1 9 A ) e x c a v a t e d , d u l l , h a i ry , g r a n u l a ted ; p ro n o t u m ( F i g . 1 0 1 9 B , C ) not crossed b y a t r a n s v e r s a l i m p ress i on o r w e a k l y i mp ressed , d u l l , h a i r y , g r a n u l a t e d ; s c u t u m s h i n y , g r a n u l a t ed , w i t h some l on g i t u d i n a l k ee l s ; s c u te l l u m d u l l , g r a n u l a ted ; m e t a n o t u m n a r row ; m e t a t ho r a x + p ropodeum ( F i g . 1 0 1 9 B , C ) d u l l , g r a n u l a ted ; p l e u r a a n d poste r i o r s u rf ac e o f propodeum n o t t r a n sv erse l y s t r i a te ; meso-me t a p l e u r a l s u t u re d i s t i n c t ; fore t a rs a l s e g m e n t s i n fo l l ow i n g p roport i on s : 1 5 : 3 : 5 : 2 0 : 30 ; e n l a r g e d c l a w ( F i g . 1 02 0 ) w i t hou t s u b a p i c a l too t h , w i t h a s m a l l tooth a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d w i t h 1 0 p e g- l i k e h a i rs ; s e g m e n t 5 of front t a rs u s ( F i g . 1 02 0 ) w i t h 2 rows of 40 l a me l l ae ; a p e x not bent at r i g h t a n g l e , w i t h a g ro u p of 1 5 l a me l l ae ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k no w n L O C U S TYP I C U S : 1 2 0 50 ' S 5 1 0 47 ' W ( B r a z i l ) T Y P I CA L M A T E R I A L : ho l o t y p e F ! i n BM . HOST S : u n k no w n D I S T R I BU T I O N : Neotrop i c reg i o n : BRAZ I L : 1 2 0 50 ' S 5 1 0 47 ' W ( M a to

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F EMALE ( F i g . 1 02 1 A ) : a p t e ro u s ; l en g t h 6 , 56 m m ; h e a d b l ack , w i t h m a n d i b l es y e l l o w ; c l y p e u s b l ac k or y e l l ow , w i t h c en t r a l spot b l ac k ; a n te n n a e b row n , w i t h segmen ts 1 -3 red d i s h or b row n - l i g h t ; thor a x , p ropod e u m a n d a b domen b l ac k ; l eg s redd i s h , w i t h b l ac k spots on coxae a n d c l u b s of femor a ; occa s i on a l l y l eg s b l ack , w i t h tarsi l i g h t ; a n te n n a l segme n t 3 n e a r l y t h ree t i mes a s l o n g a s seg m e n t 1 ; head ( F i g . 1 02 1 B ) exca v a t e d , d u l l , g ra n u l a t e d , w i t h some short k e e l s ; p ro n o t u m ( F i g . 1 02 1 C ) not crossed b y a t r a n s versa l i m p re s s i on o r w e a k l y i m p ressed ; sc u t u m d u l l , h a i ry or h a i r l ess , g ra n u l a ted ; s c u te l l u m b roa d , s h or t , h a i ry o r h a i r l es s , g r a n u l a t e d ; m e t a t horax + p ropodeum d u l l , h a i r y , g ra n u l a ted ; m e t a n o t u m a n d p l eu r a not s t r i a t e ; posterior surface of p ropod e u m on l y l a tera l l y t r a n s verse l y s t r i a te ; med i a n reg i o n smoot h ; meso-me t a p l eura l s u t u re o b so l e t e ; a b domen h a i r y ; fore t a rsa l segmen t s i n fo l l ow i n g proport i o n s : 1 7 : 4 : 5 : 1 6 : 2 5 ; en l arged c l a w ( F i g . 1 02 2 ) w i t h o u t s u b a p i c a l toot h , w i t h a sma l l too t h a t t h e e n d of a l on g i t u d i n a l f u r row a n d w i t h 1 row of tee t h , w i t h o u t b r i s t l es or peg- l i ke h a i rs or l ame l l a e ; s e g m e n t 5 of f ro n t t a r s u s ( F i g . 1 022 ) w i t h 2 rows of 76-78 l ame l l ae e x t en d i n g con t i n u o u s l y t o t h e apex ; apex b e n t a t r i g h t a n g l e ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l spurs 1 , 0 , 1 . MAL E : u n k n o w n LOCUS TYP I C U S : B a rb acoas ( Co l om b i a ) T Y P I CAL MAT E R I A L : h o l o t y p e F ! i n BM ; 1 p a r a t y pe F ! i n E B . HOST S : u n k n o w n D I ST R I B U T I ON : COLOMB I A : B a rb acoas ( N a r i no ) , BM ! BRAZ I L : C a mpo de Jordao ( Sao P a u l o ) , E B !

3.

Trichogona top u s p a lliditarsis ( C a meron ) n . com b .

Gon a top u s p a l liditarsis C a meron 1 88 8 : 441 .

F E MALE ( F i g . 1 023 A ) : a p terou s ; l e n g t h 7-9 m m ; red d i s h , w i t h t hor ax , p ropodeum a n d p et i o l e b l ac k ( except for s i des of p ro n o t u m p a r t l y redd i s h ) ; occa s i on a l l y h e a d most l y b l ac k ; a n te n n ae v e r y l on g , s l e n de r , w i t h s e g m e n t 3 t h ree t i me s a s l o n g a s segmen t 1 ; head ( F i g . 1 023 B ) e x c a v a t e d , s h i n y , h a i r l es s , g r a n u l a ted ; pronotum ( F i g . 1 02 3 C ) weak l y t r a n sve rse l y i m pressed , h a i r y , s h i n y , w e a k l y g r a n u l a ted ; s t a l k ( F i g . 1 023 A ) b e t w ee n p rono t u m a n d m e t a t horax + p ropode u m v e r y l o n g a n d s l en de r , composed o f sc u t u m , scu te l l u m a n d m e t a n o t u m , as l o n g a s m e t a t horax + p ropode um ; u s u a l l y s c u t u m , s c u t e l l u m a n d m e t a n o t u m t r a n sv erse l y s t r i a t e ; s t a l k w i t h s i des p ro t r u d i n g ; p ro t r u s i on s po i n te d ; scu te l l u m w e a k l y v i s i b l e ; m e t a t horax + propodeum s h i n y , h a i r y , w i t ho u t scu l p t u re ; poster i o r s u rface of p ropodeum f u l l y o r most l y t ra n sverse l y s t r i a t e ; p l e u r a g r a n u l a t ed , w i t h i r regu l a r s t r i a e ; s e g m e n t 1 of fron t t a rs u s s horter t h a n s e g me n t 4 ( 1 9 : 22 ) ; e n l a r g e d c l a w ( F i g . 1 024 ) w i t ho u t su b a p i ca l toot h , w i t h a sm a l l too t h a t

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t h e e n d o f a l o n g i t u d i n a l f u rrow a n d w i t h 9 p e g - l i k e h a i rs ; segmen t 5 of fro n t t a r s u s ( F i g . 1 024 ) w i t h 2 rows of 4 + 24 l am e l l ae ; a p e x b e n t a t r i g h t a n g l e , w i t h a g ro u p of a t l e a s t 1 5 l a me l l ae ; m a x i l l a ry p a l p i w i t h 6 segme n t s ; l a b i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k n o w n L O C U S T Y P I C U S : V o l c a n de C h i r i q u i ( P a n a m a )

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TYP I C A L M A T E R I A L : h o l o t y p e F ! i n BM . H O ST S : u n k n o w n D I ST R I B U T I O N : Neo t rop i c reg i o n : PANAMA : Vo l c a n de C h i r i q u i , 1 2 ° 3 1 ' S 5 5 ° 3 7 ' W ( S i n o p , M a t o Grosso ) , T W !

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Cha lcogon a top u s rap tor F e n ton 1 927 : 8 .

F E MALE : a p tero u s ; l e n g t h 7 , 0-7 , 5 m m ; head b l ac k , w i t h m a n d i b l es b ro w n redd i s h ; a n t e n n a e b row n , w i t h s e g m e n t s 1 -3 o r 1 -4 a n d 8- 1 0 y e l l ow ; t ho r a x a n d p ro p o d e u m b l ac k ; a b domen b ro w n - b l ac k , w i t h s i des redd i s h ; l eg s y e l l ow-brow n ; h e a d exca v a ted , s h i n y , g ra n u l a ted ; occ i p u t ha i r y ; p ro n o t u m n o t crossed b y a t r a n s v e rs a l i m p res s i on or w e a k l y i m p ressed , h a i r y , g r an u l a ted ; sc u t u m a n d m e t a t h o r a x + p ropodeum h a i r y , g ra n u l a t ed ; poste r i o r s u rf a c e of p ro p o d e u m not t r a n s v erse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re obso l e t e ; s e g m e n t 4 o f fro n t t a r s u s a s l o n g a s seg m e n t 1 ; e n l a rg e d c l a w ( F i g . 1 02 5 ) w i t h o u t s u b a p i c a l toot h , w i t h a s m a l l too t h a t t h e e n d o f a l o n g i t u d i n a l f u rrow a n d w i t h 5 b r i s t l es ; sm a l l t o o t h occ a s i on a l l y a b s e n t ; s e g m e n t 5 of f ro n t t a r s u s ( F i g . 1 02 5 ) w i t h 2 rows of 1 5- 1 9 l ame l l ae ; a p e x b e n t a t r i g h t a n g l e , w i t h a g ro u p o f a t l e a s t 50 l a me l l a e ; m a x i l l a r y p a l p i w i t h 6 seg m e n t s ; l ab i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k n o w n L O C U S TYP I C U S : P o r t o Be l l o ( P a n ama ) T Y P I CAL MAT E R I AL : h o l o t y p e F ! i n W A . HOST S : u n k n o w n D I ST R I BU T I O N : N eo t ro p i c reg i o n : PANAMA : Porto B e l l o , WA ! P E R U : T i n g o M a r i a ( Mo n zo n V a l l e y ) , CA ! 1 1 ° 3 ' 5 7 5 ° 1 7 ' W , MO ! E C U A DO R , WA !

5.

Trichogon a top u s a l b omargin a t u s ( C a m eron ) n . com b .

Gon a top u s albomargin a tu s C a meron 1 88 8 : 441 . Gon a top u s dromedarius C ameron 1 888 : 443 ( n . s y n . ) ; loc u s typ i c u s V o l c a n de C h i r i q u i ( P a n a m a ) . Gon a top u s camelus D a l l a Torre 1 898 : 545 ( n . s y n . ) ; new n a me . Trichogo n a top u s bogotensis K i effer 1 909 : 337 ( n . s y n . ) ; loc u s typ ic u s Bogota ( C o l o mb i a ) .

F EMAL E : a p terous ; l e n g t h 3 , 75 -5 , 3 1 m m ; b l ac k , occ a s i on a l l y w i t h c l y p e u s p a r t l y redd i s h ; m a n d i b l e s y e l l o w ; a n te n n ae fu l l y t e s t aceous or b l ac k ; occas i o n a l l y b ro w n , w i t h s e g m e n t s 1 -3 a n d 1 0 red d i s h ; l eg s redd i s h , w i t h coxae b ro w n i s h o r b l ac k ; t a r s i testaceo u s ; h e a d exca v a ted , d u l l , ha i r l es s , g r a n u l a t e d , w i t h s hort t r a n svers a l k ee l s a l o n g o r b i ts ; p ro n o t u m not crossed by a t r a n s v e rs a l i m p ress i o n o r wea k l y i m p ressed , d u l l , h a i r y , g r a n u l a ted ; s c u t u m g ra n u l a t e d , w i t h some l o n g i t u d i n a l k ee l s ; sc u te l l u m w e a k l y v i s i b l e ; m e t a t h o r a x + p ropodeum d u l l , g r a n u l a ted ; p l e u r a not s t r i a te ; p os te r i o r s u rface of p ropode u m u s u a l l y p a rt l y t ra n s v e rs e l y s t r i a te ,

1 486

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occas i o n a l l y n o t s t r i a t e ; meso-me t a p l eu r a l s u t u re o b so l e t e ; seg m e n t 1 o f f ro n t t a rs u s s l i g h t l y s ho r t e r t h a n segmen t 4 ( 1 3 : 1 4 ) ; en l a r ged c l a w ( F i g . 1 026 ) w i t ho u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d o f a l o n g i t u d i n a l f u rrow a n d w i t h 5-7 p e g- l i ke h a i r s ; segme n t 5 of fron t t a r s u s ( F i g . 1 026 ) w i t h 2 row s of 1 4-29 l a m e l l ae ; a p e x b e n t a t r i g h t a n g l e , w i t r, a g ro u p of at l e a s t 9-29 l a m e l l a e ; m a x i l l a r y p a l p i w i t h 6 segm e n ts ; l a b i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k n o w n L O C U S TYP I C U S : V o l c a n de C h i r i q u i ( P a n a m a ) TYP I CAL MAT E R I A L : h o l o t y p e F ! of G. albomargin a t u s i n B M ; h o l o t y pe F ! of G . dromedarius i n BM ; h o l o t y p e F ! o f T. bogoten s is i n H U . HOST S : u n k n o w n D I ST R I BU T I O N : Neotrop i c reg i o n : C O S T A R I CA , WA ! PANAMA : Vo l c a n d e C h i r i q u i , B M ! BU g a b a , BM ! V E N E Z U E LA : C a r a c a s , WA ! C O ! C O L O M B I A :

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F E MA L E ( F i g s 20 , 1 027 ) : a p t e ro u s ; l e n g t h 3 , 2 5-4 , 1 8 m m ; head redd i s h , occas i o n a l l y w i t h oce l l a r reg i o n b ro w n i s h ; a n t e n n a e y e l l ow-red d i s h , occas i o n a l l y w i t h s e g m e n t s 5- 1 0 b ro w n ; thorax a n d p ropodeum b l ack ; a b domen b ro w n - b l a c k , w i t h a p e x a n d terg i t e 1 redd i s h ; l e g s y e l l ow redd i s h ; head e x c a v ated , d u l l , h a i r l es s , g r a n u l a ted , w i t h some shor t , t r a n s v e r s a I k e e l s a l o n g o r b i t s ; p ro no t u m n o t crossed b y a t r a n s versa I i m pres s i o n or weak l y i m p ressed , d u l l , h a i r l es s , g r a n u l a ted ; sc u t u m d u l l , g r a n u l a ted ; m e t a n o t u m w i t h s i des rounded ; m e t a t ho r a x + p ropodeum d u l l , g r an u l a ted ; p l e u r a a n d p os t e r i o r s u rface of p ropodeum not t r a n s verse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re o b so l e te ; fore t a r s a l s e g m e n t s i n fo l l ow i n g p ropor t i on s : 1 7 : 3 : 5 : 1 5 : 2 2 ; en l arged c l a w ( F i g . 1 028 ) w i t h o u t s u b a p i c a l too t h , w i t h a s ma l l too t h a t t h e e n d o f a l o n g i t u d i n a l f u r row a n d w i t h 5

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7.

Trichogon a top u s

rubriceps K i effer

Trichogon a top us rubriceps K i effer 1 909 : 3 3 6 . Trichogon a top u s rubriceps K i effer : K i effer 1 9 1 4b : 86 .

Gon a topod i n ae

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1 489

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F E MA L E : a p terou s ; l e n g t h 5 , 62 m m ; h e a d fu l l y redd i s h ; a n t e n n a e b row n , w i t h s e g m e n t s 1 -4 a n d 1 0 red d i s h - t e s t aceou s ; t ho r a x , p ropodeum a n d a b domen b l a ck ; l eg s b l ac k o r b ro w n , w i t h t roc h a n ters , t a r s i , s t a l k s of fore femora a n d fore t i b i a e test aceo u s ; a n t e n n a e d i s t a l l y t h i c k e n e d ; a n te n n a l se g m e n t s i n fo l l o w i n g p ro p or t i on s : 1 2 : 7 : 3 1 : 1 6 : 1 4 : 1 2 : 8 : 8 : 7 : 1 0 ; h e a d exca v a te d , d u l l , g ra n u l a t e d ; occ i p i t a l ca r i n a i n v i s i b l e ; fron t a l l i ne comp l e te ; P O L = 3 ; O L = 3 ; OOL = 1 1 ; p ro n o t u m d u l l , not crossed b y a t r a n s v e rs a l i m p re s s i o n , g r a n u l a ted ; s c u t u m d u l l , g r a n u l ated , w i t h some l o n g i t u d i n a l k ee l s ; sc u te l l u m i n v i s i b l e ; m e t a t h o r a x + p ropodeum d u l l , g r a n u l a ted ; poste r i o r s u rf a ce of p ropodeum o n l y l a tera l l y t r a n s v erse l y s t r i a t e ; cen t r a l reg i o n smoot h ; p l e u r a n o t t r a n s v e rse l y s t r i z t e ; meso-

1 490

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m e t a p l e u r a l s u t u re obso l e t e ; fore t a r s a l segmen t s i n fo l l o w i n g p ropor t i o n s : 20 : 5 : 8 : 24 : 36 ; en l a r g e d c l a w ( F i g . 1 02 9 ) w i t ho u t s u b a p i c a l toot h , w i t h a sma l l too t h at t h e e n d of a l o n g i t u d i n a l fu rrow a n d w i t h 7 p e g - l i k e h a i rs ; segmen t 5 of f ro n t t a rs u s ( F i g . 1 029 ) w i t h 2 rows of n e a r l y 2 3 l a m e l l ae ; a p e x w i t h a g r o u p of n e a r l y 8 l a me l l ae ; m a x i l l a r y segme n t s ; l ab i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M AL E : u n k n o w n L O C U S T Y P I C U S : Bo g o t a ( C o l o mb i a ) T Y P I CA L MAT E R I AL : h o l o t y p e F ! i n H U . HOST S : u n k no w n D I ST R I B U T I O N : Neotrop i c reg i o n : C O L O M B I A :

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Epigon a top u s R . C . L . Perk i n s 1 90 5 : 45 .

T Y P E S P E C I E S : Epigon a top u s solitarius R . C . L . P e r k i n s 1 90 5 , o r i g . des i g . F E MALE : a p t erou s ; m a x i l l a r y p a l p i w i t h 2 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; p r o n o t u m crossed by a s t r o n g t r a n s v e r s a I i m p ress i on ; en I a r ged c l a w ( F i g . 1 030 ) w i t ho u t s u b a p i c a l too t h , w i t h a s m a l l tooth a t the end of a l o n g i t u d i n a l furrow and w i t h sorne b r i s t l es o r p e g- l i ke h a i rs ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n D I ST R I B U T I O N : A u s t r a l i a n H OSTS : u n k n o w n SPEC I E S : 2

AUSTRAL I AN S P E C I E S OF EPIGONA TOPUS

T w o s p ec i es are k n ow n .

KEY T O T H E S P E C I E S

F E MA L E S

B o d y s l e n d e r ; me t a t h o ra x + p r op o d e u rn b l a c k , w i t h a p e x y e l l ow ; s eg m e n t 5 o f f r o n t t a r s u s w i t h o u t a r e g u l a r r ow o f l a me l l a e , b u t o n l y w i t h a f ew i r r e g u l a r l a me l l a e ( F i g . 1 0 3 0 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 . fa l l a x R . C . L . P e rk i n s

Gon a t o po d i n ae

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B o d y r o b u s t ; me t a t h o ra x + p r o p o d eum r ed d i s h ; s eg men t 5 o f f r o n t t a r s u s w i t h a r eg u l a r r o w o f l a me l l a e ( F i g . 1 0 3 1 ) •

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1 . Epigon a top u s fa l l a x R . C . L . P e rk i n s

Epigon a top u s fa llax R . C . L . Perk i n s 1 90 6 a : 490 .

F E MA L E : a p tero u s ; l e n g t h 3 , 1 5 m m ; b ro w n - b l ac k , w i t h m a n d i b l es a n d c l y p e u s y e l l o w ; a n t e n n a e b row n i s h , w i t h s e g m e n t s 1 - 3 y e l l ow ; p rono t u m b row n ; s c u t u m a n d a b domen b row n ; a pex of propodeum y e l l ow ; l eg s y e l l o w , w i t h coxae a n d c l u b s of femora b ro w n ; head exca v a t e d , s h i n y , smoo t h , w i t ho u t sc u I p t u re ; prono t u m crossed b y a s t ro n g t r a n svers a l i m p ress i o n , s h i n y , smoot h , w i t ho u t sc u l p t u re ; s c u t u m a n d m e t a t h o r a x + p ropodeum d u l l , a l u t aceou s ; pos ter i or s u rf a c e of p ropodeum t r a n s verse l y s t r i a te ; seg m e n t 4 of f r o n t t a rs u s s horter t h a n segmen t 1

( 20 : 30 ) ;

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c l a w ( F i g . 1 03 0 ) w i t ho u t s u b a p i c a l t oo t h , w i t h a sm a l l too t h a l on g i t u d i n a l f u rro w , w i t ho u t b r i s t l es o r p e g - l i k e h a i rs or s e g me n t 5 o f f ro n t t a r s u s ( F i g . 1 03 0 ) w i t h o u t a r e g u l a r row w i t h o n l y 2 l o n g l a m e l l ae ; apex w i t h a g ro u p of 9 l a m e l l ae ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u r s 1 , 0 , 1 . MAL E : u n k n o w n L O C U S TYP I C U S : M i t t a g o n g

a t t he e n d o f l a me l l ae ; o f l a m e l l ae , m a x i l l a ry a n d

(Austra l i a )

T Y P I CAL MAT E R I AL : l ectot y p e F ! ( des i g n a t e d b y O l m i 1 982 ) i n B . HOST S : u n k n o w n D I ST R I BU T I O N : A u s t r a l i a n reg i o n : AUSTRAL I A : M i t t a g o n g ( N ew So u t h W a l es ) , B!

2 . Epigo n a topus solitarius R . C . L . Perk i n s

Epigon a top u s solitariu s R . C . L . Perk i n s 1 90 5 : 4 5 .

F E MAL E : a p terou s ; l en g t h 3 , 4 m m ; redd i s h , w i t h m a n d i b l es , c l y p e u s a n d f r o n t p a rt of vertex y e l l ow ; a n t e n n a e b ro w n i s h , w i t h segmen t s 1 -2 y e l l ow ; p et i o l e b l ac k ; a bdomen b ro w n - b l a c k , w i t h t e r g i t e 1 p a r t l y redd i s h ; l e g s y e l l o w , w i t h f o r e f e m o r a b row n i s h ; h e a d exc a v a te d , s h i n y , smoo t h , w i t h o u t s c u l p t u re ; p ro n o t u m c rossed b y a s t ron g t r a n svers a l i m p re s s i o n , s h i n y , smoo t h , w i t ho u t scu I p t u re ; sc u t u m a n d m e t a t ho r a x + p ropodeum d u I I , a l u t ac eo u s ; poste r i or s u r f ace of p ropodeum t r a n sverse l y s t r i ate ; seg men t 4 of f ro n t t a r s u s shorter t h a n s e g m e n t 1 ( 26 : 27 ) ; e n l a rged c l a w ( F i g . 1 03 1 ) w i t ho u t s u b a p i c a l too t h , w i t h a sm a l l t o o t h a t t h e e n d of a l o n g i t u d i n a l f u rrow , w i t ho u t b r i s t l es o r peg - l i ke h a i rs o r l a me l l ae ; seg men t 5 o f f r o n t t a rs u s ( F i g . 1 03 1 ) w i t h 1 row of 1 6 l a me l l ae ; a p e x w i t h a g ro u p of 3 l a me l l a e ; m a x i l l a r y a n d l a b i a l p a l p i w i t h 2 segmen ts ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k no w n LOCUS T Y P I C U S : B u n d ab e r g ( A u s t r a l i a ) T Y P I CAL MATE R I A L : l ecto t y p e F ! ( d es i g n ated b y O l m i 1 982 ) i n B . HOSTS : u n k n o w n D I STR I BU T I O N : A u s t r a l i a n reg i o n : AUSTRAL I A : B u n d a berg ( Queens l a n d ) , B !

1 6 . Es agona top u s n .

gen .

T Y P E S P E C I E S : Es a go n a top u s niger ( F e n ton 1 92 4 ) F E MALE ( F i g s 1 037 , 1 038 ) : a p t e ro u s ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l ab i a l p a l p i w i t h 2 segme n t s ; pronotum ( F i g . 1 03 8 ) c rossed b y a s t r o n g t r a n s v e rs a l i m p ress i o n ; e n l a rg ed c l a w ( F i g . 1 03 4 ) w i t h o u t s u b a p i c a l toot h , w i t h a s m a l l tooth a t t h e e n d of a l o n g i t u d i n a l f u r row a n d w i t h b r i s t l es or peg- l i ke h a i rs ; t i b i a l s p u rs 1 , 0 , 1 .

1 494

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MALE ( F i g s 27 , 1 0 35 ) : f u l l y w i n g e d ; m a x i l l a r y p a l p i w i t h 6 segme n ts ; l a b i a l p a l p i w i t h 2 segmen t s ; t e m p l es d i s t i n c t ; occ i p u t con c a v e ; occ i p i t a l c a r i n a i n v i s i b l e o r on l y v i s i b l e b e h i n d t h e oce l l i o r o n t h e s i des of t h e poste r i o r oce l l i , b u t n e v e r reach i n g t em p l e s ; t i b i a l s p u r s 1 , 1 , 2 . D I ST R I B U T I O N : Nearc t i c HOST S : Cic a dellida e , exce p t for Typhlocyb in a e , SPEC I E S : 2

I diocerin a e , Ma cl'opsin a e .

N E A R C T I C S P E C I ES OF ESA GONA TOPUS

T w o s p ec i es a re k n ow n .

KEY T O T H E SPEC I ES

F E MA L E S

Me t a t h o r a x + p r o p o d e u m b r oad e r b eh i n d t h e s c u t e l l u m ( F i g . 1 0 3 3 B ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 . n iger ( F e n t on ) Me t a t h o r a x + p r o p o d e u m l e s s b r oad b e h i n d t h e s cu t e l l u m ( F i g . 1 0 3 3 A ) . . . . . . . . . . . . . . . . . . . . . . . 2 . perdebilis ( R . C . L . P e rk i n s )

MALES

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n iger is k n ow n .

1 . Esagona top u s n iger ( F e n t o n ) n . com b .

Chalcogon a top u s nigrus Fen t o n 1 924 : 1 9 3 . Chalcogon a top u s niger Fen ton : K romb e i n 1 958 : 1 00 . Neogon a topus n iger ( F e n t o n ) : F re y t a g 1 977 : 574 . Neogon a topus niger ( F en t o n ) : F re y t a g 1 980 : 1 45 .

F E M AL E : a p t e ro u s ; l e n g t h 2 , 37-3 , 00 m m ; h e a d ferru g i n o u s , w i t h fron t p a rt of v e r te x , c l y p e u s a n d m a n d i b l e s y e l l o w ; occ a s i o n a l l y v e r t e x w i t h a b l ac k t r a n s v e rs a l b a n d ; a n te n n a e y e l l o w , w i t h s e g m e n t s 8- 1 0 d a reken ed ; t ho r a x a n d p ropodeum ferr u g i n o u s , w i t h apex of p ro p o d e u m y e l l ow ; occ a s i o n a l l y t horax a n d p ro p o d e u m w i t h d a r k s po t s ; p et i o l e b l ac k ; a b domen b r o w n ferru g i no u s ; l e g s y e l l ow-redd i s h ; bod y some t i mes f u l l y y e l l ow , w i t h p e t i o l e b l ac k ; h e a d ( F i g . 1 03 2 B ) e xc a v a t e d , sh i n y ; v e r t e x w i t ho u t sc u l p t u re ; f ro n s a n d occ i p u t gra n u l a t e d ; p ro n o t u m ( F i g . 1 03 2 A ) crossed b y a s t ron g t ra n s v ers a l i m p re ss i on , s h i n y ; co l l a r smoo t h , w i t h o u t sc u l p t u re ; d i sc g ra n u l a t e d ; s i des of p ro n o t u m w i t h l on g i t u d i n a l k ee l s ; scu t u m s h i n y , w i t h l on g i t u d i n a l k ee l s ; met a n o t u m t r a n s v e rse l y s t r i a t e ; met a t horax + p ropodeum ( F i g . 1 03 2 A) s h i n y , g r a n u l a t e d ; l a t e r a l l y v i e wed t h i c k n ess of m e t a t horax + p ro p o d e u m greater t h a n t h i c k n ess of scu t u m ( F i g . 1 03 3 B ) ; poster i or s u rface of p ro p o d e u m a n d p l e u r a t r a n sve rse I y s t r i a t e ; meso-me t a p l eu r a I s u t u re obso l e te ; segmen t 4 of f ro n t t a rs u s s l i g h t l y l on g e r t h a n segmen t ( 1 3 : 1 2 ) ; en l a r g e d c l a w ( F i g . 1 03 4 ) w i t ho u t s u b a p i c a l too t h , w i t h a s m a l l t o o t h a t t h e e n d of a l o n g i t u d i n a l f u r ro w a n d w i t h 7-9 p e g - I i k e h a i rs ;

1 496

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C h e l a of E s a gon a topus n iger ( F en t o n ) ( t y p e )

segme n t 5 o f f ro n t t a rs u s ( F i g . 1 034 ) w i t h 2 rows o f 1 7 l a me l l ae o r w i t h row of 1 3- 1 5 l a m e l l a e ( p rox i m a l l a me l l a e l o n g er t h a n med i a l a n d d i s t a l l a m e l l a e ) ; a p e x w i t h a g r o u p o f 7- 1 8 l a me l l ae ; m a x i I l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 0 , . MALE ( F i g s 27 , 1 035 ) : f u l l y w i n g e d ; l e n g t h 2 m m ; b l ac k , w i t h m a n d i b l es y e l l o w ; a n t e n n a e , l e g s a n d a b domen b row n ; a n te n n a e n o t d i s t a l l y t h i c k e n e d ; a n t e n n a l seg m e n t s i n fo l l o w i n g p roport i on s : 4 : 5 : 7 : 6 : 6 : 6 : 6 : 5 , 5 : 5 , 5 : 1 0 ; a n te n n a I s e g m e n t 3 l ess t h a n t h ree t i mes a s l on g a s b ro a d ( 2 , 8 ) ; h e a d s h i n y , g r a n u l a t e d ; occ i p u t con c a v e ; o cc i p i t a l c a r i n a i n v i s i b l e ; t e m p l es d i s t i n c t ; f ro n t a l l i n e a b s e n t ; POL = 6 ; O L = 2 ; DOL = 4 ; s c u t u m s h i n y , g r a n u l a t ed ; n o t a u l i ces com p l e t e , poster i or l y sep a r a ted ; m i n i m u m d i s t a nc e b e t ween t h e n o t a u l i ces s horter t h a n b re a d t h o f t h e oce l l i ( 2 : 3 ) ; s c u te l l u m a n d met a n o t u m s h i n y , smoo t h , w i t h o u t s c u l p t u re ; p ro p o d e u m f u l l y ret i c u l a t e ru gose ; fore-w i n g h y a l i n e , w i t ho u t d a r k t ra n s v e rs a l b a n d s ; r a d i a l ce l l o p e n ; d i s t a l p a r t of r a d i a l ve i n l on g er t h a n p ro x i m a l p a r t ( 1 7 : 8 ) ; dors a l p rocess o f gonoforceps ( F i g . 1 03 6 ) b ro a d e n e d ; m a x i l l a r y p a l p i w i t h 6 s e g m e n t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u rs 1 , 1 , 2 . L O C U S T Y P I C U S : Ames ( I o w a , U . S . A . ) T Y P I CA L MAT E R I AL : h o l o t y p e F ! i n WA . H O ST S : i n U . S . A . Scap hoideus p a ludosus B a l l ( F re y t a g 1 97 7 , 1 980 ) . D I ST R I B U T I O N : N e a rc t i c reg i o n : C AN AD A : Marmora ( On t . ) , O T ! S . M i n o n i c o ( O n t . ) , O T ! S t . D a v i d s ( O n t . ) , O T ! OL ! U . S . A . : R a msey C o . ( N . D . ) , OL !

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( Ke n t u ck y )

Ames ( I o w a ) , W A ! L e x i n g to n ( K y . ) , K E ! R ob e r t so n C o . ( K y . ) ( F r e y t a g 1 977 ) ; Harr i s b u rg ( P a . ) , W A ! M E X I CO : C u e r n a v aca ( Mo re l os ) , WA !

2 . Esagona topus perdebilis ( R . C . L .

Perk i n s ) n . com b .

Ch a lcogon a top u s perdebilis R . C . L . P er k i n s 1 907 : 27 .

F E MA L E ( F i g s 1 03 7 , 1 038 ) : a p t e rou s ; l en g t h 2 , 92-3 , 1 8 m m ; y e l l ow t e s t aceo u s , w i t h p e t i o l e b l ac k ; a n t e n n a e y e l l o w , w i t h seg men t s 8- 1 0 w e a k l y b row n ; h e a d exca v a ted , s h i n y ; v e r t e x w i t h o u t scu l p t u re ; f ro n s a n d occ i p u t g ra n u l a te d ; p ro n o t u m crossed b y a s t r o n g t r a n svers a l i m p ress i o n , s h i n y , w i t ho u t sc u l p t u re ; sc u t u m s h i n y , w i t ho u t scu l p t u re ; m e t a n o t u m t r a n s verse l y s t r i a te ; m e t a t horax + p ro p o d e u m s h i n y , w i t ho u t scu l p t u re ; l a t e ra l l y v i ewed t h i c k n es s of m e t a t horax + p ro p o d e u m s m a l l e r t h a n t h i ck n e s s of s c u t u m ( F i g . 1 033 A ) ; pos t e r i o r s u rface of p ropodeu m t r a n s verse l y s t r i a t e ; meso m e t a p l e u r a l s u t u re o b so l e t e ; segmen t 4 of f ro n t t a r s u s s l i g h t l y l o n g e r t h a n s e g me n t 1 ( 25 : 22 ) ; e n l arged c l a w ( F i g . 1 039 ) w i t h o u t s u b a p i c a l too t h , w i t h a sma l l t oo t h a t t h e e n d of a l on g i t u d i n a l f u r ro w a n d w i t h 7 p e g- l i k e h a i rs ; segmen t 5 of fro n t t a r s u s ( F i g . 1 03 9 ) w i t h 1 row of 1 3 l a me l l ae ;

1 498

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a p e x w i t h a g r o u p of a t l e a s t 9 l a me l l ae ; m a x i I l a ry p a l p i w i t h 6 segme n t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : No g a l e s ( A r i z . , U . S . A . ) T Y P I CAL M A T E R I AL : l ec t o t y p e F ! a n d 1 p a r a l ec t o t y p e F ! ( d es i g n a t e d b y O l m i 1 982 ) i n B . HOST S : u n k n o w n D I ST R I B U T I O N : N e a rc t i c reg i on : U . S . A . : N og a l es ( A r i z . ) , B ! OL ! W i ndow T ra i l ( B i g B e n d N . P . , T ex . ) , O T !

1 7 . Gon a topus L j u n g h

Gon a top u s Lj u n g h 1 8 1 0 : 1 6 1 . Neogon a top u s R . C . L . Perk i n s 1 90 5 : 42 ( n . s y n . ) ; t y p e s p ec i es Neogon a top u s ombrodes R . C . L . P e r k i n s 1 905 , o r i g . des i g .

Gon a topod i n a e

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1 499

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Pa chygon a top u s R . C . L . Perk i n s 1 90 5 : 4 5 ( n . s y n . ) ; t y p e s p ec i es Pachygon a top u s mel a n i a s R . C . L . Perk i n s 1 905 , or i g . des i g . Ch alcogona topus R . C . L . P e r k i n s 1 90 5 : 46 ( n . s y n . ) ; t y p e s p ec i es Cha lcogon a top u s gigas R . C . L . Perk i n s 1 905 , o r i g . des i g . Eugon a top u s R . C . L . Perk i n s 1 905 : 46 ( n . s y n . ) ; t y p e s p ec i es Cha lcogon a topu5 (Eugon a top u s ) pseudochromus R . C . L . Perk i n s 1 90 5 , ori g . d es i g . Pla tygon a top u s K i effer i n K i effer e t M a r s h a l l 1 906 : 500 ( n . s y n . ) ; t y p e s pec i es Pla tygona topus plan iceps ( K i effer 1 904 ) , des i g n a te d by R i c h a r d s 1 93 9 . Eucamp tonyx R . C . L . Perk i n s 1 907 : 2 8 ( n . s y n . ) ; t y p e spec i e s Eucamp tonyx tes taceus R . C . L . P e rk i n s 1 907 ,

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monot y p i c . A gona top u s R . C . L . P e r k i n s 1 907 : 29 ( n . s y n . ) ; t y p e spec i es A gona top u s p a l l idieornis R . C . L . P erk i n s 1 90 7 , d es i g n a t e d b y R i c h a rd s 1 93 9 . Cyrtogona topus K i effer 1 907b : 1 9 ( n . s y n . ) ; t y p e s p ec i e s Cyrtogon a top u s breviforeeps ( K i effer 1 904 ) , monot y p i c . nee Triehogona topus K i effer 1 909 : 3 3 6 . Digon a top u s K i effer 1 9 1 3c : 325 ( s y n . p roposed b y R i c h a r d s 1 939 ) ; t y p e spec i e s Digon a top u s j a v a n u s ( R . C . L . P e r k i n s 1 9 1 2 ) , monot y p i c . Triehogon a top u s H e l l en 1 930 : 2 ( n . s y n . ) ; t y p e spec i es Triehogo n a top u s lindbergi H e l l en 1 930 , o r i g . des i g . Me tagon a topus O g l o b l i n 1 93 2 : 264 ( n . s y n . ) ; t y p e s p ec i es Me tagon a top u s brasiliensis O g l ob l i n 1 9 3 2 , ori g . des i g . Gona top u s Lj u n g h : R i c h a rd s 1 937 : 1 1 8 . A llogon a topus H a u p t 1 938 : 2 7 ( n . s y n . ) ; t y p e s pec i es A llogon a top u s procerus H a u p t 1 938 , o r i g . des i g . Ple e trogon a top u s R i c h a r d s 1 939 : 207 ( n . s y n . ) ; t y p e s p ec i e s Pleetrogon a top u s stri a t u s ( K i effer 1 905 ) , o r i g . des i g . Rhynehogon a top us Ben o i t 1 953b : 390 ( n . s y n . ) ; t y p e spec i e s Rhynchogon a top u s meridion a l is Beno i t 1 95 3 , or i g . d es i g . M a decago n a top u s B e n o i t 1 95 3b : 392 ( n . s y n . ) ; t y p e s pec i es Ma deeagon a top u s a n k ara trus ( Ce b a l l os 1 936 ) , or i g . d es i g . Cyrtogona topoides P o n o m a ren k o 1 96 6 : 1 579 ( n . s y n . ) ; t y p e s pec i es Cyrtogona topoides borealis P o n o m a r e n k o 1 966 , o r i g . d es i g .

Gon a t opod i n ae

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1 50 1

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T Y P E S P E C I E S : Gona top u 5 formicarius L j u n g h 1 8 1 0 , mono t y p i c . F E MA L E ( F i g s 1 065 , 1 066 , 1 090 ) : a p t e ro u s ; m a x i l l a ry p a l p i w i t h 3-6 segmen t s ; l a b i a l p a l p i w i t h 2-3 segmen t s ; p a l p a l r a t i o s : 3/2 , 4/2 , 4/3 , 5/2 , 5/3 , 6/3 ; p ro n o t u m ( F i g . 1 06 6 ) crossed b y a s t ro n g t r a n s v e r s a l i m p re s s i o n ; en l a rged c l a w ( F i g . 1 04 0 ) w i t h o u t s u b a p i c a l too t h , w i t h or w i t ho u t a s m a l l too t h at the e n d of a l o n g i t u d i n a l f u rrow a n d w i t h b r i s t l e s o r p e g- l i k e h a i rs or , r a re l y , l a me l l ae ; t i b i a l s p u rs 1 , 0 , 1 . MALE ( F i g s 1 1 78 , 1 1 79 ) : fu l l y w i n g e d ; m a x i I l a r y p a l p i w i t h 3-6 segme n t s ; l a b i a l p a l p i w i t h 2-3 segmen t s ; p a l p a l r a t i o s : 3/2 , 4/2 , 4/3 , 5/2 , 5/3 , 6/3 ; t em p l es d i s t i n c t ; occ i p u t co n c a v e ; occ i p i t a l c a r i n a i n v i s i b l e or o n l y v i s i b l e b e h i n d t h e oce l l i or o n t h e s i des of t h e pos t e r i o r oce l l i , b u t n e v e r reac h i n g t e m p l e s ; t i b i a l s p u rs 1 , 1 , 2 . D I ST R I B U T I O N :

wo r l d w i de

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PALAEARCT I C S P E C I E S OF GONA TOPUS

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S eg m e n t 1 o f f r on t t a r s u s a p p r o x i ma t e l y t w i c e a s l on g a s s egmen t 4 . . • . • • • • • . • • • • . • • • • • • • • • • . • • • • • • • • • • . • • • . • • • . . . . • . . . . 2 S egmen t o f f r on t t a r s u s s h o r t e r , a s l ong a s , o r s l i g h t I y l o n g e r t h a n s e g men t 4 . . . . . . . • • • . . • . . . . • . . . . . . . . . . . • . . . . . . . . . . • 3 H ea d , t h o ra x a n d p r o p o d e u m f e r ru g i n ou s • • • • . • . . • • • . . • • . • • • • • • • • • • • • . . . . . . . . . . . . • • . . . . • • 1 . vistos u s n . s p . H ea d mo s t l y b l a c k ; t h o r a x a n d p r o p o d e u m b l a c k . . . . . . . . . . . . . . . . . . •

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(F i g s 1 050 , 1 074 ) • • • • • • . • • • • • . • • • . . • • • • • • • • • • . • • • . • • • . • • • • . • • • 4 S e g m e n t 5 o f f r on t t a r s u s w i t h i n n e r s i d e p r ox i ma l l y n o t s e r ra t e ( F i g s 1 0 8 6 , 1 0 9 2 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 8 S eg m e n t 5 o f f r on t t a r s u s w i t h 2 - 3 r o w s o f l a me l l a e e x t en d i n g b ey o n d 0 , 5 l en g t h o f s eg m e n t ( F i g . 1 0 4 2 ) . . . . . . . . . . . . . . . . . . . . . . 5 S eg m e n t 5 o f f r o n t t a r s u s w i t h 1 r o w o f l a me l l a e ( F i g . 1 0 5 9 ) ; o c ca s i o n a l l y w i t h 2 r o w s , b u t i n t h i s c a s e o n e o n l y r o w e x t e n d i n g b ey o n d 0 , 5 l en g t h o f s eg m e n t ( F i g s 1 0 5 4 , 1 0 5 8 ) . . . . . 1 1 S c u t u m a t l ea s t p a r t l y r ed d i s h - t e s t a c e ou s ; s eg m e n t 5 o f f r on t

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t a r s u s w i t h m ed i a l l a m e l l a e n o t l o c a t ed o n a d i s t i n c t p r om i n en c e ( F i g . 1 0 4 2 ) . 3 . doderoi ( O l m i a n d C u r r a d o ) S c u t u m b l a c k ; o c c a s i on a l l y r ed d i s h o r y e l l o w , b u t i n t h i s c a s e s eg m e n t 5 o f f r o n t t a r s u s w i t h me d i a l l a m e l l a e l o c a t ed o n a d i s t i n c t p r om i n e n c e ( F i g . 1 0 4 4 ) . . . . . . . . . . . . . . . . . . . . . . . . .6 P r o n o t u m p a r t I y r ed d i s h - t e s t a c e ou s ; s eg m e n t 5 o f f r o n t t a r s u s w i t h l a m e l l a e b eg i n n i n g i n p r ox i ma l t h i rd ( F i g . 1 0 4 3 ) 4 . b ilinea tus K i e f f e r P r on o t u m b l a c k ; o c c a s i on a l l y p a r t l y o r f u l l y r e d d i s h o r y e l l ow , b u t i n t h i s c a s e s eg m e n t 5 o f f r o n t t a r s u s w i t h l am e l l a e b e g i n n i n g i n d i s t a l h a l f ( F i g . 1 0 4 4 ) . . . . . . . . . . . . . . . . . 7 S eg m e n t 5 o f f r o n t t a r s u s w i t h l a m e l l a e b e g i n n i n g i n d i s t a l ha l f ( F i g s 1 044 , 1 08 1 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 S e g me n t 5 o f f r o n t t a r s u s w i t h l a m e l l a e b eg i n n i n g i n p r ox i ma l .

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st r i at e . . . . . . . . . . . . . . . . . . . . . 8 . lran l C U S n . s p . �e t a t h o ra x + p r o p o d e u m w i t h a s t r o n g med i a n f u r r o w . . . 12 Me t a t h o ra x + p r op od e u m w i t h o u t med i a n f u r r o w o r o n l y w i t h t ra c k o f med i a n f u r r o w . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 4 M e t a t h o r a x + p r op od e u m b l a c k , w i t h a p ex r e d d i s h . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 . vollenhoveni ( O l m i a n d C u r ra d o ) Me t a t h o ra x + p r o p o d e u m r ed d i s h 1 3 H ea d f l a t 1 0 . a l b oli n e a t u s K i e f f e r H ea d ex c a v a t ed . . . . . . . . 1 1 . s u arezi n . sp . M e s o -m e t a p l eu ra l s u t u r e d i s t i n c t a n d c omp l e t e . . . . . . . . . . . . . . . . 1 5 M e s o -m e t a p l eu ra l s u t u r e ob s o l e t e o r w e a k l y v i s i b l e o n l y i n .21 p r o x i ma I h a I f . . H e a d f u I I y r ed d i s h - t e s t a c e o u s . . . . . . . . . " . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 . dimidi a t u s S a h I b e rg H e a d a t l e a s t p a r t l y b l a ck o r b r own . . 16 Me t a n o t u m w i t h l a t e r a l p o i n t s . . . . . . . . . . 1 7 M e t a n o t u m w i t h o u t l a t e ra l p o i n t s . . . . . . . . . . . . . . . .18 P r o t h o ra x a n d scu t u m f u l l y b l a ck . . . . . . . . . . · 1 3 . a tl a n ticus n . sp . P r o t h o r a x a n d s c u t u m p a r t l y r ed d i s h . . . . . . . . . . ·

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1 504

Mem . Amer . E n ! .

I ns! . ,

no.

37 ,

1 984

1 4 . u n ilineatus K i e f f e r 1 8 T h o r a x a n d p r op o d e u m f u I I y r ed d i s h - t e s t a c e ou s 1 5 . aegyptiacus n . s p . T h o ra x a n d p r op o d e u m p a r t I y o r f u I I y b l a ck . . . . . . . . . . ·

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. . . . .1 9

1 9 Me t a n o t u m h o l l ow b e h i n d s c u t e l l u m . . . . . . . . . . . . 1 7 . b ar b a tellus R i c h a rd s Me t a n o t u m n o t h o i l o w b eh i n d s c u t e l l u m . . . . . . . . . . . . . . . . . . . . . .

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2 0 P o s t e r i o r s u r f a c e o f p r o p o d e u m t ra n s v e r s e l y s t r i a t e ; me t a t h o r a x + p r op od e u m s h i n y , w i t h o u t s c u l p t u r e , e x c ep t f o r s t r i a e on a n t e r i o r a n d p o s t e r i o r s u r f a c e . . . . . . . . . . . . . . . . . . . . . .

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1 6 . plumbeus n . s p . P o s t e r i o r s u r f a c e o f p r op od e u m n o t t r a n s v e r s e l y s t r i a t e ; me t a t h o ra x + p r o p o d e u m du l l , g ra n u l a t e d . . . . . . . . . . . . . . . . . . •

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4 0 . gra ecus n . s p . T h o ra x a n d p r op o d e u m f u l l y b l a c k , ex c ep t f o r a p ex o f p r op od e u m I i gh t 22 T h o ra x a n d p r op o d eu m a t i ea s t p a r t l y y e l l ow o r r ed d i s h . . . . . . . 2 4 2 2 H e a d a l mo s t f l a t ( F i g . 1 0 5 6 B ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 8 . a ter n ew n a me H ea d v e r y ex c a v a t ed ( F i g . 1 0 5 6 A ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 3 2 3 S eg m e n t 5 o f f r on t t a r s u s w i t h a p i c a l l a m e l l a e n ea r l y s a m e l e n g t h ( F i g . 1 0 6 4 ) . . . . 1 9 . lin dbergi H e l l en S e g m e n t 5 o f f r o n t t a r s u s w i t h s ome a p i c a l l ame l l a e mu c h •

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l o n g e r t h a n o t h e r a p i c a l l a me l l a e ( F i g . 1 0 5 7 ) . . . . . . . . . . . . 1 3 . a tl a n ticus n . s p . 2 4 Me t a n o t u m f I a t 25 Me t a n o t u m i n c l i n ed , n o t f l a t . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 6 2 5 S c u t e l l u m d i s t i n c t , l a t e r a l l y w i t h t wo p o i n t s ; s t a l k b e t we en p r o t h o ra x a n d me t a t h o r a x + p r o p od e u m s h o r t ; a n t e r i o r s u r f a c e o f me t a t h o ra x + p r o p o d e u m u s u a l l y d u l l , g r a n u l a t ed . . . . . . . . . . . . . 2 0 . l u n a tus K I u g S c u t e l l u m i n d i s t i n c t , l a t e ra l l y w i t h o u t p o i n t s ; s t a l k b e t w e en p r o t h o ra x a n d me t a t h o r� x + p r op o d e u m v e r y l on g a n d s l en d e r ; a n t e r i o r s u r f a c e o f me t a t h o ra x + p r op o d e u m s h i n y , w i t h o u t s c u l p t u r e . . . . . . . . . . . . . 3 9 . schenklingi S t r a n d ( O r i e n t a l s p e c i e s ) 2 6 S c u t e l l u m a n d me t a t h o ra x + p r op o d e u m b l a ck , e x c ep t f o r a p ex o f p r op o d e u m l i g h t . . . . . . . 2 1 . tenerifei n . s p . S c u t e l l u m a n d me t a t h o ra x + p r op o d e u m r ed d i s h - t e s t a c e o u s . . . . . . 2 7 2 7 Me t a t h o ra x + p r op od e u m a n d h e a d s h i n y , w i t h o u t s c u l p t u r e , smo o t h ( e x c e p t f o r t ra n s v e r s a l s t r i a e o n p l eu ra a n d p o s t e r i o r s u r f a c e o f p r op o d e u m ) . 2 2 . can ariensis n . s p . Me t a t h o r a x + p r op o d e u m a n d h e a d d u l l , g ra n u l a t ed ( i n a d d i t i on t o t ra n s v e r s a l s t r i a e on p l eu ra a n d p o s t e r i o r s u r f a c e o f p ropodeum) 2 3 . wittmeri n . s p . 2 8 Me t a t h o r a x + p r o p o d e u m w i t h a mo r e o r l e s s s t r o n g med i a n f u r r ow 29 Me t a t h o r a x + p r op od e u m w i t h ou t med i a n f u r r o w . . . . . . . . . . . . . . . . . 3 5 •

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Go n a to po d i n a e

1 505

29 S e g m e n t 5 of f r o n t t a r s u s w i t h l a m e l l a e b eg i n n i n g i n d i s t a l ha l f ( F i g . 1 07 6 ) 2 4 . n igrosign a tu s S a h l b e rg S e g me n t 5 o f f r o n t t a r s u s w i t h l a me l l a e b eg i n n i n g i n p r ox i ma l t h i rd ( F i g s 1 0 7 7 , 1 0 7 8 ) 30 3 0 S e g men t 5 o f f r on t t a r s u s w i t h ap i c a l l a m e l l a e n ea r l y s a m e l en g t h ( F i g . 1 0 7 7 ) 2 5 . p l a n iceps K i e f f e r S eg m e n t 5 o f f r o n t t a r s u s w i t h 1 - 3 a p i c a l l am e l l a e mu c h l on g e r t h a n o t h e r a p i c a l l a me l l a e ( F i g s 1 0 7 8 , 1 0 8 3 ) 31 3 1 S e g men t 1 o f f r on t t a r s u s s l i g h t l y s h o r t e r t h a n s e g m e n t 4 32 S e g me n t 1 o f f r o n t t a r s u s l o n g e r t h a n s eg men t 4 33 3 2 S eg m e n t 5 o f f r on t t a r s u s w i t h 2 r o w s o f l a me l l a e ex t en d i n g b e y on d 0 , 5 l en g t h o f s eg m e n t ( F i g . 1 0 9 6 ) . . . . . . . . . . . . . . . . . . . . . . 3 7 . rhaensis P o n oma r e n k o S e g me n t 5 o f f r on t t a r s u s w i t h 1 r ow o f l a m e l l a e ; o c ca s i o n a l l y w i t h 2 r o w s , b u t i n t h i s c a s e o n e o n l y r o w e x t e n d i n g b ey on d 0 , 5 l e n g t h o f s eg m e n t ( F i g . 1 0 7 8 ) 2 6 . campestris P o n oma r e n k o 3 3 T h o ra x , p r o p o d e u m a n d mo s t p a r t o f h e a d r e d d i s h . . . . . . . . . . . . . . . . . . . . . . 1 0 . a lbolin e a t u s K i e f f e r T h o ra x , p r op o d eu m a n d mo s t p a r t o f h ea d b r own -b l a c k 34 3 4 P o s t e r i o r s u r f a c e o f p r op od eu m s t r o n g l y t ra n s v e r s e l y s t r i a t e 2 9 . bernardi ( P i c a rd ) P o s t e r i o r s u r f a c e o f p r o p o d e u m n o t t ra n s v e r s e l y s t r i a t e 3 0 . form icarius L j u n g h 3 5 Me s o -m e t a p l e u r a l s u t u r e ob s o l e t e 36 Me s o -me t a p l e u ra l s u t u r e d i s t i n c t a n d c omp l e t e ( sepsoides g roup ) 42 3 6 T emp l e s v e r y l on g , a p p r ox i ma t e l y 0 , 5 - 0 , 8 a s l o n g a s e y e s ( F i g . 1 079 A ) 2 7 . s u b tilis n . s p . T emp l e s s h o r t , l e s s t h a n 0 , 5 t i me s a s l on g a s e y e s 37 3 7 B o d y y e l l o w - t e s t a c e o u s , w i t h p e t i o l e b l a c k a n d a b d omen b r o wn i s h ; o c ca s i on a l l y t wo b r own s p o t s o n s i d e s of p r op od e u m a n d s cu t u m 2 8 . spectrum ( Sn e l l e n v a n V o l l e n h ov en l B o d y d i f f e r e n t l y c o l o r ed ; h ea d , t h o ra x a n d p r o p o d e u m r ed d i s h o r a t l e a s t p a r t I y b r own o r b l a ck 38 3 8 S c u t e l l u m a n d m e t a t h o ra x + p r op od eu m b l a c k ( ex c e p t f o r a p e x o f p r op od e u m l i g h t ) S c u t e l l u m a n d m e t a t h o ra x + p r op od e u m r ed d i s h o r r e d d i s h -b r own . . .41 3 9 A n t e r i o r s u r f a c e o f m e t a t h o ra x + p r op o d e u m s h i n y , s m o o t h , w i t h ou t s c u l p t u r e 3 1 . mongolieus P o n oma r e n k o A n t e r i o r s u r f a c e o f m e t a t h o ra x + p r o p o d e u m d u l l , g ra n u l a t e d 40 4 0 S e g m e n t 5 o f f r on t t a r s u s w i t h 2 r o w s o f l a m e l l a e l o e a t ed o n a d i s t i n c t p r o m i n en c e ( F i g . 1 0 4 4 ) • • 5 . distinguendus K i e f f e r S eg men t 5 o f f r o n t t a r s u s w i t h 1 r ow o f l a me l l a e ; l a m e l l a e n o t l o c a t ed o n a d i s t i n c t p r om i n e n c e ( F i g . 1 0 8 6 ) •

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1 506

41

Mem .

Amer.

En!.

I ns! . ,

n o . 37 ,

1 984

3 2 . longus ( P o n oma r e n k o ) T h o r a x a n d p r o p o d e u m d u l I , g ra n u l a t ed 3 3 . n ivos u s n . s p . T h o r a x a n d p r op o d e u m s h i n y , w i t h o u t s c u l p t u r e , e x c ep t s t r i a e on p l e u r a a n d p o s t e r i o r s u r f a c e o f p r o p o d e u m .

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ex t en d i n g b e y o n d 0 , 5 l en g t h o f s e g m e n t ( F i g . 1 0 8 5 ) 1. . . . . . . . . . . 4 3 Segmen t 5 of f ron t t a r s u s wi t h row o f l ame l l a e ; o c ca s i on a l l y w i t h 2 r ow s , b u t i n t h i s c a s e o n l y o n e r ow ex t e n d i n g b e y on d 0 , 5 l en g t h o f s eg me n t ( F i g . 1 0 6 2 ) � . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 6 A n t e r i o r s u r f a c e o f m e t a t h o ra x + p r op od e u m w i t h s ome d i s t i n c t l on g i t u d i n a l k e e l s . . . . 3 5 . cherson esius P o n oma r e n k o A n t e r i o r s u r f a c e o f m e t a t h o ra x + p r op o d e u m w i t h o u t d i s t i n c t . . 44 l on g i t u d i n a l k e e l s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . A n t e r i o r s u r f a c e o f m e t a t h o ra x + p r op o d e u m smo o t h , n o t g ra n u l a t e d ( F i g . 1 0 9 1 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 6 . s ep s o id e s W e s t w o o d A n t e r i o r s u r f a c e o f m e t a t h o ra x + p r op o d e u m g ra n u l a t ed . . .45 P o s t e r i o r s u r f a c e o f m e t a t h o ra x + p r o p o d e u m s t r o n g l y t r a n s v e r s e l y s t r i a t e . . 3 1 . mongolicus P o n oma r en k o P o s t e r i o r s u r f a c e o f m e t a t h o ra x + p r o p o d e u m n o t t r a n s v e r s e l y st r i a t e 3 0 . form icarius L j u n g h A n t e r i o r s u r f a c e o f me t a t h o r a x + p r op o d e u m smo o t h , w i t h o u t s c u l p t u r e ( F i g . 1 0 9 1 ) . 3 6 . sepsoides W e s t w o o d A n t e r i o r s u r f a c e o f m e t a t h o ra x + p r op o d e u m g r a n u l a t ed 47 S c u t u m a l mo s t f u l l y y e l l o w . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 7 . b ar .� a t e l l u s R i c h a r d s Scu t um b l a ck . . . 48 P o s t e r i o r s u r f a c e o f p r op o d e u m s t r o n g l y t ra n s v e r s e l y s t r i a t e . . . . . . . . . . . . . . . . . . . . . . 3 8 . b a rb ieri n . sp . P o s t e r i o r s u r f a c e o f m e t a t h o ra x + p r op od e u m n o t t r a n s v e r s e l y 5t r i at e j ra r e l y v e ry weak l y s t r i a t e . . . . . . . . . . . . . . . 3 0 . formicarius L j u n g h •

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M i n i mu m d i s t a n c e b e t we en t h e n o t a u l i c e s n ea r l y a s l on g a s a n t e n n a I s egmen t 2 5 . distingu e n d u s K i e f f e r M i n i mu m d i s t a n c e b e t w e e n t h e n o t a u l i c e s mu c h s h o r t e r t h a n •

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1 T he f o I l o w i n g spec ies ( sep soides, form icarius, b a rb a te l I u s , chersonesius, mongolic u s , b arbier i ) are b e l o n g i n g to s ep soi de s g ro u p , i n c l u d i n g very s i m i l a r s p ec i e s .

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G o n a t opod i n a e

Fig.

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C he l a of Gon a top u 5 v i s t o s u s n . sp .

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( type)

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no .

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l en g t h of a n t en n a l s eg m e n t 2 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 D o r s a l p r o c e s s o f g o n o f o r c e p s n ea r l y a s l o n g a s g o n o f o r c ep s ( F i g . 1 0 4 9 ) ; rad i a l c e l l w i d e r ( F i g . 1 0 4 8 A ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6 . stri a t u s K i e f f e r D o r sa l p r o c e s s o f g o n o f o r c ep s mu ch s h o r t e r t h a n g on o f o r c e p s ( F i g s 1 0 7 1 , 1 0 7 2 , 1 0 9 5 ) ; r a d i a l c e l l s ma l l e r ( F i g . 1 0 4 8 B , C ) . . . . . . . . . . . . . .3 B a s i v o l s e l l a w i t h n o t ch ( F i g . 1 0 7 1 ) . . . . . 2 0 . l u n a tus 1< I u g G a s i v o l s e l l a w i t h ou t n o t c h ( F i g . 1 0 9 5 ) 3 6 . sepsoides W e s t w o o d •

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T h e m a l es of t h e o t h e r s p ec i e s of Gon a top u s a r e u n k no w n .

1 . Gon a topus vistosus n .

sp .

F E MALE : a p te ro u s ; l e n g t h 2 , 3 1 -3 , 1 2 m m ; h e a d ferr u g i n o u s , w i t h m a n d i b l e s a n d fron t p a r t o f v e r t e x y e l l o w ; a n t e n n a e b row n , w i t h segmen t s 1 -3 y e l l ow ; t ho r a x a n d p ropode u m fer r u g i no u s ; p e t i o l e b l ac k ; a b domen b l ack , w i t h segmen t 1 ferru g i n o u s ; l e g s y e l l ow - t e s t aceo u s , w i t h b o t h e x t rem i t i es of m i d a n d h i n d t i b i ae b row n ; h e a d w e a k l y excav a ted , s h i n y , h a i r l ess , g r a n u l a ted ; te m p l e s l es s t h a n 0 , 5 as l o n g as eyes ; p ro n o t u m crossed by a s t rong t r a n s v e rs a l i m p re ss i o n , s h i n y , h a i r l es s , w i t h o u t scu l p t u r e ; scu t u m s h i n y , g r a n u l a ted ; met a n o t u m n o t t r a n s verse l y s t r i a t e ; m e t a t ho r a x + p ropode u m sh i n y , w e a k l y a l u t aceo u s ; pos t e r i o r s u rface of propodeum g r a n u l a te d ; p l e u r a and p o s te r i o r s u rf ace of p ro p o d e u m not or w e a k l y t ra n s v erse l y s t r i a t e ; meso- m e t a p l eu r a l s u t u re o b so l e t e ; seg m e n t 1 of f ro n t t a r s u s t w i ce a s l o n g a s s e g m e n t 4 ( 1 4 : 7 ) ; e n l a rged c l aw ( F i g . 1 040 ) too t h , w i t h a s m a l l t oo t h a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d w i t h 3-6 peg- I i k e h a i rs ; seg men t 5 o f fron t t a rs u s ( F i g . 1 040 ) w i t ho u t s u b a p i c a l

w i t h i n ner s i d e not serra t e , w i t h 2 rows of 1 4- 1 6 v e r y s m a l l l a m e l l a e ; a pex b ro a d e n e d , w i t h a g rou p of 6-1 1 l a m e l l ae ; m a x i I l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n lOCUS TYP I CU S : Guenav i s t a ( Ca n a r y I s l an d s ) TYP I CAL MAT E R I AL : ho l o t y p e F ! i n H E ; 1 p a r a t y p e F ! i n SU ; 3 p a r a t y p es F F ! i n M D ; 1 p a r a t y p e F ! i n T N ; 2 p a ra t y pe s F F ! i n Ol . HOST S : u n k no w n D I ST R I B U T I O N : P a l a e a rc t i c reg i on : CANARY I SLANDS : B u e n a v i s t a ( T ener i fe ) , H E ! B a r r a n co de T a hod i o ( T e n er i fe ) , O l ! Los O r g a nos ( A g u a M a n s a , T e ner i fe ) , O l ! B a r r a nco H o n do ( T e n er i fe ) , M D ! l a d e r a de G i U m a r ( T e n er i fe ) , M D ! C os t a de S . t a C r u z ( T e n e r i fe ) , M O ! T e n er i fe , T t" ! l a P a l ma ,

SU i

Gon a t o p o d i n ae

Fig.

1 509

1 04 1 - C he l a of Gonatop u s horvathi K i effer from N i k o l a e v s k - n a-A m u re ( U . S . S . R . ) ( A ) ; B : m a x i l l a ry p a l p u s .

2 . Gonatopus horvathi K i effer

Gon a topus horvathi K i effer i n K i effer et M a r s h a l l

1 90 6 : 503 . Moczar 1 965 : 387 . Cyrtogona topoides borealis Ponomare n k o 1 966 : 1 579 ( n . s y n . ) ; locus typ ic u s K a m j e n k a ( U . S . 5 . R . ) . Cyrtogon a topoides borealis Ponomare n k o : Ponomarenko 1 978 : 2 6 .

Pla tygon a topus horva th i ( K i effer ) :

F E MAL E : a p t e ro u s i l en g t h 3 , 00-3 , 1 2 m m i b l a c k , w i t h m a n d i b l e s , c l ypeus and fro n t part of vertex y e l l ow ; a n te n n ae b row n , wi t h segmen t s 1 -3 y e l l ow ; l eg s y e l l o w , w i t h coxa e a n d c l u b s of femora brow n ; m i d t i b i ae w i t h i n ne r s i de of a p i c a l ex trem i t i es brow n ; head f l a t , s h i n y , h a i r l es s , w i t h o u t scu l p t ur e , exce p t for fro n t p a r t o f vertex a n d occ i p u t g r an u l a t e d ; pronot u m crossed b y a s t rong t r a n svers a l i m press i on , s h i n y , h a i r l es s , g r an u l a te d ; sc u t u m g ra n u l a ted , w i t h some l on g i t u d i n a l k ee l s ; m e t a n o t u m t ra n s verse l y s t r i a t e ; meta t horax + propod e u m d u l l , g ra n u l a te d ; poste r i or s u rface of propodeum a n d p l e u r a s t ro n g l y t r a n sverse l y s t r i a te ; meso met a p l eu r a l s u t u re obso l e t e ; seg m e n t 1 of fro n t t a r s u s t w i ce as l on g a s

1 51 0

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C he l a of Gon a topu5 doderoi ( O l m i a n d C u rr a d o ) from T a u r i r t ( Morocco)

Gona topod i n a e

151 1

segme n t 4 ( 1 4 : 7 ) ; e n l a r g e d c l aw ( F i g . 1 04 1 A ) w i t h o u t s u b a p i ca l toot h , w i t h a sm a l l tooth a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d w i t h 6 peg- l i ke h a i rs ; segme n t 5 of f ro n t t a r s u s ( F i g . 1 04 1 A ) w i t h i n n e r s i de n o t serra t e , w i t h 2 row s of near l y 1 7 l a me l l ae ; apex b roadened , w i t h a g ro u p of 1 0 l a mel l ae ; m a x i l l a r y p a l p i w i t h 4 segme n t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l spurs 1 , 0 , 1 . MALE : u n k no w n LOCUS TYP I C U S : K rasova ( Hu n g a ry ) TYP I CAL MAT E R I AL : h o l o t y pe F ! of G. horva t h i i n B U ; h o l ot y pe F ! a n d 3 p a r a t y pes F F of C. bore a l i s i n MO . HOST S : i n U . S . S . R . Psammotettix stria t u s L . ( Ponoma ren ko 1 97 8 : u n der n a me Cyrtogon a topoides borealis Pon . ) . D I ST R I BU T I ON : P a l aearc t i c reg i o n : U . S . S . R . : N i k o l aevsk-n a -A m u re ( Hab arovsk ) , H E ! K a m j e n k a ( L j s v i e n , Perm ) , MO ! T a t a r k a ( Lj s v i e n , Perm ) ( Ponomarenko 1 966 ) ; S a k h a l i n ( Ponomarenko 1 966 ) ; HUNGARY : K r a sov a , B U ! ROMAN I A : M i l c o i u ( Arges ) ( Teodorescu 1 967 ) .

3 . Gon a topus doderoi ( O l m i a n d C u rrado ) n . com b .

Gon a top u s unilin e a t u s v a r i e t e K i effer 1 904a : 359 . Neogona top u s doderoi O l m i a n d C u r r a d o 1 97 4 : 1 95 .

F E MALE : a p terou s ; l e n g t h 3 , 43-4 , 1 2 m m ; head test aceou s , w i t h b ro w n spot o n ver t e x ; a n te n n ae b l ac k , w i t h segmen t s 1 -3 and part of 4 t e s t aceou s ; p ronot u m test aceo u s , w i t h pos t e r i or s u rface o f d i sc b l ack ; scu t u m test aceou s , w i t h s i des b l ac k ; occa s i on a l l y posterior h a l f b l ack ; scut e l l um test aceous or b l ack ; meta t horax + p ropodeum b l ack ; occ as i on a l l y m e t a n o t u m b ro w n - t e s t aceou s ; a b domen b l ack ; l eg s test aceou s ; head exc a v a te d , h a i r y , weak l y g r a n u l a te d ; prono t u m crossed by a s t ro n g t r a nsvers a l i m press i o n , s h i n y , smoo t h , w i t ho u t scu l p t u re ; sc u t u m s h i n y , w i t h some l on g i t u d i n a l k ee l s ; m e t a n o t u m f l a t ; met a t horax + p ropod e u m s h i n y , w i t ho u t sc u l p t u re ; p l e u r a a n d pos t e r i o r s u rface of p ropodeum t r a n sverse l y s t r i a t e ; meso met a p l e u r a l s u t u re d i s t i n c t a n d comp l e t e ; en l a rged c l a w ( F i g . 1 04 2 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d of a l o n g i t u d i n a l fu rrow a n d w i t h 2-4 peg- l i k e h a i r s o r b r i s t l e s ; seg m e n t 5 of fro n t t a r s u s ( F i g . 1 04 2 ) w i t h 2 row s o f 22-40 l ame l l ae exten d i n g beyon d 0 , 5 l e n g t h of segme n t ; apex w i t h a g ro u p of 1 4-20 l a me l l ae ; m a x i l I a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MAL E : u n k n o w n L O C U S TYP I C U S : Go l fo A r a n c i ( I t a l y ) TYP I CAL MAT E R I AL : h o l o t y p e F ! i n GE . HOSTS : u n k n o w n D I STR I BU T I O N : P a l aearc t i c reg i o n : I TALY : Go l fo A r a n c i ( Sa s s a r i ) , G E ! MOROCCO : T a u r i r t , SU ! S . Lorenzo ( Me l i l i a ) , MD !

1512

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! I \

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Fig.

1 04 3 - C h e l a of Gonatopus bilin e a t u s K i effer from G i g l i o I s l a n d ( I !al y )

Gon a t opod i n a e

1513

4 . Gon a topus b ilin e a t u s K i effer

Gonatopus bilineatus K i effer 1 904a :

359 .

Gonatopus glaber K i effer i n K i effer et M a r s h a l l

1 905 : 1 1 1 ( n . s y n . ) ; loc u s typicus Mon t-de-M a r s a n ( F rance ) . Gon a topus glaber K i effe r : Ber l a n d 1 928 : 1 5 1 . Neogon a topus glaber ( K i effer ) : A b d u l -Nour 1 974 : 1 0 1 2 . Neogon a topus bilin e a t u s ( K i effer ) : O l m i a n d C u rrado 1 97 4 : 1 98 . Neogon a topus glaber ( K i effer ) : A b d u l -No u r 1 97 6 : 273 .

F E MAL E : a p t e ro u s ; l e n g t h 4 , 20-4 , 25 m m ; head red d i s h-testaceou s , w i t h a b l ack b a n d on v er t e x ; a n te n n ae b l ac k , w i t h segmen t s 1 -3 test aceou s ; occ i p u t redd i s h - t e s t aceou s ; occas i on a l l y occ i p u t b l ack o r red d i s h w i t h temp l e s b l ac k ; pronotum redd i s h-test aceou s , w i t h t r a n svers a l i m press i o n a n d m o s t p a r t o f d i sc b l ack ; scu t u m , s c u te l l u m , m e t a t horax + p ropodeum a n d pe t i o l e b l ack ; a bdomen b l ack , w i t h red d i s h n u a nces ; l eg s red d i s h t e s t aceo u s , occa s i o n a l l y w i t h coxae a n d c l u b s o f femora b row n i s h ; head exc a v a te d , s h i n y , g r a n u l a t e d ; p ro n o t u m crossed b y a s t ro n g t r an svers a l i mp ress i o n , s h i n y , g r an u l a te d ; scu t um d u l l , w i t h some l o n g i t u d i n a l k ee l s ; m e t a n o t u m s h i n y , w i t h o u t scu l p t u re , w i t h o r w i t h o u t t r a n svers a l s t r i ae ; met a t horax + p ropodeum d u l l , w i t ho u t sc u l p t u re , except for p l e u r a a n d poster i or s u rface o f propodeum t r a n s verse l y s t r i a t e ; meso-me t a p l eura l s u t u re d i s t i nc t a n d comp l e t e ; fore t a r s a l segmen t s i n fol l ow i n g p roport i o n s : 1 2 : 2 : 4 : 1 2 : 2 1 ; e n l a rged c l aw ( F i g . 1 04 3 ) w i t h o u t s ub a p i c a l t oo t h , w i t h a s ma l l tooth at t h e e n d of a l o n g i t u d i n a l f u rrow a n d w i t h 2-4 b r i s t l es or peg- l i ke h a i r s ; seg ment 5 of fro n t t a r s u s ( F i g . 1 043 ) w i t h i n n er s i d e p rox i ma l l y serrate a n d w i t h 2 row s of 28-39 l am e l l ae exten d i n g beyond 0 , 5 l e n g t h of segme n t ; apex w i t h a g ro u p of 9-33 l a me l l ae ; m a x i l l a ry p a l p i w i t h 4 segme n t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k no w n L O C U S TYP I C U S : G i g l i o I s l a n d ( I t a l y ) TYP I CA L MAT E R I AL : h o l o t y pe F ! of G . bilin e a tus i n G E ; h o l o t y p e F of G . glaber l os t ( Ab d u l -Nour 1 974 ) . HOSTS : i n F r a nce Aphrodes bicin e tu s Sc h r k . a n d Thamnotettix dil utior K b m . ( Ab d u l -Nour 1 976 ) . D I S T R I BU T I O N : P a l aearc t i c reg i o n : F RANC E : Mon t-de-Marsan ( L a n d es ) , M T ! La F i g a rede ( Hera u l t ) , P ! Cette ( He r a u l t ) , P ! M a n d u e l ( Ga rd ) , P ! I TALY ; G i g l i o I s l a n d ( Grosseto ) , G E ! Samb i a se ( C a t a n z a ro ) , G E !

5 . Gona top u s distinguendus K i effer

nee Gonatopus flavieornis Da I m a n

1 8 1 8 : 83.

Gona top u s fla vieornis C . G . T homson 1 860 :

1 81

( s y n . proposed b y

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1 044 - C h e l a of Gona top us distinguendus K i effer from E I P a rdo ( Spa i n )

Gon a topod i n ae

Fig.

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1 04 5 - R a d i a l v e i n of ma l e of Gonatop u s distinguendus K i effer from V e l e t a ( Sp a i n )

D l m i 1 977 ) ; loc us typ ic u s S k a n e ( Swede n ) . Gonatopus distinguendus K i effer i n K i effer et M a rsh a l l 1 90 5 : 1 1 6 . Gonatopus lu teicornis K i effer i n K i effer e t M a r s h a l l 1 905 : 1 1 7 ( s y n . p roposed by D l m i a n d C u rrado 1 976b ) ; locus typicus T r i es t e ( I ta l y ) • Gonatopus exc a vatus J . S a h l berg 1 9 1 0 : 9 ( n . s y n . ) ; loc us typicus Mets a p i r t t i ( U . S. S. R . ) . Gonatopus distinguen dus K i effer : O g l ob l i n 1 924b : 1 2 1 . Gonatopus lichtensteini P i c a r d 1 93 2 : 30 ( n . s y n . ) ; locus typicus of h o l o t y pe B o l quere ( F rance ) ; locus typicus of n eo t y pe T u rk e n Sc h a n tze ( A u s t r i a ) . A llogon a topus procerus H a u p t 1 938 : 27 ( n . sy n . ) ; locus typicus B e l l i nchen ( Germ a n y ) . Neogon a topus distinguen dus ( K i effer ) : R i c h a r d s 1 93 9 : 2 1 4 . Gonatopus thomsoni H e l l <§n 1 953 : 9 4 ( n . s y n . ; n e w n a me i n s tead of G. fla vicornis C . G . T homson p reocc u p i e d by G. fla vicornis Dalman) • Gonatopus rossicus Ponom arenko 1 965 : 627 ( n . sy n . ) ; locus typicus K i n e l ( U . S. S. R . ) . Neogona topus distinguen dus ( K i effer ) : D l m i a n d C u rr a do 1 97 4 : 2 1 2 . Neogona topus distinguendus ( K i effer ) : Ab d u l -Nour 1 97 6 : 273 . Neogona top u s distinguendus ( K i effer ) : D l m i a n d C u rrado 1 976b : 72 .

F E MAL E : a p t erou s ; l e n g t h 2 , 68-3 , 75 m m ; co l ou r v a r i a b l e from b l ack to redd i s h-b l ack , w i t h i n termed i a te co l ou rs ; t h e b l ack v ar i e t y i s t he t y p i c a l distinguendu s ; t h e redd i s h v a r i e t y i s t h e t y p i c a l exca vatus ( = rossic u s , proceru s, lich tenstein i ) ; head redd i sh or red d i sh-testaceo u s ( occas i on a l l y y e l l ow ) , w i t h b ro w n or b l ac k b a n d o n vertex ; occ a s i on a l l y w i t h b l ack Sp0t beh i n d temp l e s ; occ a s i on a l l y b a n d beh i n d tem p l es j o i n t to vertex spo t ; a n te n n ae y e l l ow-red d i s h , w i t h segmen t 1 0 b ro w n i sh ; p ro n o t u m f u l l y b l ack ( v a r i e t y distinguen dus ) o r redd i s h w i t h s i des and t r a n sversa l i mp ress i o n b ro w n ( v a r i e t y exc a vatus ) ; occ a s i on a l l y p ro n o t u m redd i s h , w i t h d i sc p a r t l y o r fu l l y brow n or b l ack ; scu t u m b l ac k ( v a r i e t y distinguendus ) or y e l l o w o r red d i s h ( v a r i et y exc a v a t u s ) ; occas i o n a l l y sc u t u m b row n ; scute l l u m a n d meta t horax + propodeum b l ac k , w i t h apex o f p ropodeum l i g h t ; a b domen b l ack ( v a r i et y distinguen dus ) or red d i s h w i t h f i rs t and l a s t te rg i tes b l ack

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F i g . 1 04 7 - C he l a 0" ( F rance ) ( B : e n l a r g e d c l e ,>. h a i rs ) e "

Fig.

1 046 - M a l e gen i t a l i a of Gonatopus distinguendus K i effer from V e l e t a ( Sp a i n ) ( l eft h a l f removed )

( v a r i e t y exc a va tu s ) ; head exc a v a te d , s h i n y , h a i r l es s , weak l y g r a n u l a ted ; pronot u m crossed b y a s t r o n g t r a n s v e rs a l i m p ress i o n , d u l l , g r a n u l ated ; scu t u m s h i n y , w i t ho u t scu l p t u re or g ra n u l a ted ; met a n o t u m t r a n s v erse l y s t r i a t e ; meso-met a p l eu r a l s u t u re obso l e te ; me t a t horax + p ropod e u m d u l l , g r a n u l a t e d ; p l e u ra not or weak l y t r a n sverse l y s t r i a te ; poster i o r s u rface of p ropode u m weak l y t r a n s verse l y s t r i a t e ; seg m e n t 1 of f ro n t t a r s u s as l o n g a s segment 4 ; e n l a rged c l a w ( F i g . 1 04 4 ) w i t ho u t s u b a p i c a l tooth , w i t h a sm a l l tooth at t h e e n d of a l on g i t u d i n a l f u r row a n d w i t h 4-6 peg- l i k e

•

=

h a i r s ; segm e n t 5 of fro n : serrate ( occ a s i o n a l l y no: - l oc a t e d on a d i s t i n c t p r e maxi l l ary pa l p i w i t h 4 v a r i e t y exc a va tu s ) ; l a b · _ MAL E : fu l l y w i n ged ; l e r :: ' a n t e n n ae a n d l eg s b a n c e � 3 , 5 : 4 , 5 : 7 , 5 : 7: 6 , 5 : 7 : : t h ree t i mes as l o n g as = - - - absen t ; occ i p i t a l cari n e scu te l l u m a n d metanotu- sep a r a t e d ; m i n i m u m d i s : =. - : OO a n ten n a l segmen t 2 ; p r c = : : � . med i a n fu rrow o n dorsa : -

....................................--------�

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F i g . 1 04 7 - C he l a of Gon atop u s stria tus K i effer from F on t a i neb l ea u ( F rance ) ( B : e n l arged c l a w w i t h l a me l l ae ) ; A : en l a r ged c l a w ( w i t h peg- l i ke h a i r s ) of a spec i me n from Lojo ( F i n l a n d ) .

h a i r s ; segment 5 of front t a r s u s ( F i g . 1 044 ) w i t h i n n er s i de proxi m a l l y serrate ( oc ca s i o n a l l y not serrate ) a n d w i t h 2 row s of 1 2- 2 1 l ame l l ae l oc a te d on a d i s t i nc t prom i nence ; apex w i t h a group of 3 - 1 1 l am e l l ae ; m a x i l l a r y p a l p i w i t h 4 ( i n v a r i e t y distinguen dus ) o r 5 segme n t s ( i n v a r i e t y exc a v a t u s ) ; l ab i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : f u l l y w i n g e d ; l e n g t h 2 , 3 1 mm ; b l ac k ; m an d i b l es test aceou s ; a n te n n ae a n d l e g s b a n de d ; a n t e n n a I segmen t s i n fo l l ow i n g p roport i o n s : 3 , 5 : 4 , 5 : 7 , 5 : 7 : 6 , 5 : 7 : 6 , 5 : 6 , 5 : 6 , 5 : 9 ; a n ten n a l seg m e n t 3 l e s s t h a n t h ree t i mes a s l on g a s broad ( 2 , 86 ) ; head d u l l , g ra n u l a t e d ; f ro n t a l l i ne absen t ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 5 ; OL = 2 , 5 ; OOL = 4 ; scu t u m , scute l l u m a n d met a n o t u m d u l l , g r a n u l a ted ; n o t a u l i ces com p l e t e , poster i o r l y separa ted ; m i n i m um d i s t a n c e bet ween t h e n o t a u l i ces near l y a s l on g a s a n te n n a I segme n t 2 ; propodeum ret i c u l a t e r u gose , w i t h a l on g i t u d i n a l m ed i a n f u rrow o n dorsa l s u rf a ce ; fore-w i n g h y a l i n e , w i t h o u t d a r k

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t ra n s vers a l b a n d s ; r a d i a l ve i n : f i g . 1 045 ; dors a l p rocess of gonoforceps ( F i g . 1 04 6 ) s ho r t ; b as i v o l se l l a wi t ho u ! r l d g e ; max i l l ary p a l p I w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 1 , 2 . LOCUS T Y P I C U S : Wey mou t h ( E n g l a n d ) TYP I CAL MAT E R I AL : lecto t y pe F ! a n d 1 p a r a l ec to t y pe F ( d es i g n ated by Moczar 1 96 5 ) of G . dis tinguen dus In B U ; ho l ot y pe F ! of G . luteicornis i n V I ; l ec t o t y p e F ! ( des i g n a ted b y O l m i 1 97 7 ) o f G . flavicornis C . G . T homson I n L U ; h o l o t y pe F ! of G. exc a v a t u s in HE ; h o l o t y p e F of G . lichtensteini l o s t , neo t y p e F ! ( hereby d es i g n a ted ) I n Pi h o l ot y pe F ! of A. procerus I n HA ; 1 p a r a t y p e F ! of A . procerus i n B M ; ho l ot y pe F ! of G . ros sicus i n M O ; 1 p a r a t y p e F ! of G . ros sicus i n B M . HOSTS : I n F r a nce Circ ulifer dubiosus ( M a t s u m u ra ) ( = Deltocep h a l u s ruscinonensis R i b a u t ) ( P i c a rd 1 932 ) , P a l u d a elong a t a ( W a g n e r ) ( = Rhopa lopyx elonga tu s ) , A darrus taurus R i b . , Araldus propin q u u s K b m . ( A b d u l -Nour 1 976 ) ; In U. S. S . R . Ps ammotettix stria t u s L. ( Ponomare n k o 1 965 , 1 978 ) a n d Mocuellus collin u s Bah . ( Ponomarenko 1 978 ) ; i n Germ a n y b y m i s t a k e Eurysa lurida F i eb . ( Ha u p t 1 938 ) . D I ST R I BU T I O N : P a l aearc t i c reg i on : MONGOL I A ( Ponomare n k o 1 979a ) ; U . S . S . R . : M e t s a p i rt t l ( So v i e t K a r e l i a ) , HE ! K i ne l ( K u y b y s h e v ) P rov i n ce ) , MO ! BM ! SWE D E N : Sk a n e , L U ! H o l a n d , L U ! HUNGARY : K e l eb l a , B U ! Domsod ( A p a j p u szta ) , B U ! B u d a pes t , B U ! ROMAN I A : Top l e t ( Ba n a t ) ( Teodorescu 1 967 ) ; C Z E C HOSLOVAK I A ( O g l ob l i n 1 924b ) ; GE RMANY : B e l l i nchen ( O der ) , HA ! B M ! D Li n e n R a n t u m ( Sy l t , Sch l esw i g-Ho l s t e i n ) , HA ! AUSTR I A : T u rken Schantze ( W i en ) , P! A u s t r i a , CO! D E N MA RK , CO! SW I T Z E R LA ND : Lens ( V a l a l s ) , GV! FRAN C E : B o l q u ere ( P y ren ees O r i e n t a l e s ) ( P i c a rd 1 932 ) ; Corse ( R i c h a r d s 1 939 ) ; E N GLAN D : Weymou t h ( Dorset ) , B U i o t her l oca l i t i es I n R i c h a r d s ( 1 93 9 ) ; I R E LA N D ( R i c h a r d s 1 939 ) ; SPA I N : E I P a rdo ( M a d r i d ) , BM ! Pozue l o de C a l a t r a v a ( C l u d a d Rea l ) ( ca l l . by La F u e n te , det . Gonatopus pedestris D a l m a n ) , M D i V i l l atobas ( To l e do ) , M D i V e l e t a ( S i erra Neva d a ) , O L ! I TALY : T r i es te , V I ! Cec l n a ( U vorno ) , GE !

6 . Gon a top u s stria tus K i effer

Dicondyl u s p e destris ( D a l m a n ) : H a l i d a y i n C u r t i s 1 829-30 : 1 1 0 . Gona topus stria tu s K i effer i n K i effer e t M a r s h a l l 1 905 : 9 2 . Gona top u s stria tus K i effer : K i effer i n K i effer et M a r s h a l l 1 906 : 500 . Plectrogon a top u s stria tus ( K i effer ) : R i c h a r d s 1 939 : 208. Agona topoides stria t u s ( K i effer ) : Mocz a r 1 96 5 : 396. Plec trogo n a top u s rich a rdsi Moczar 1 965 : 396 ( n . sy n . ) ; locus typ ic u s D e a l ( E n g l a n d ) . A gona topoides tauricus Ponomarenko 1 965 : 623 ( sy n . proposed b y C u rrado a n d O l m i 1 975 ) ; l o c u s typicus G u r s u f ( U . S . S . R . ) . Plectrogon a top u s richardsi Mocza r : C u rrado a n d O l m i 1 976 : 1 . n e c Agon a topoides stria t u s ( K i effer ) : J . F . Perk i n s 1 976 : 32 .

Gon a topod i n ae

1519

� /�"' . .. � A � .

Fig.

1 048 - R a d i a l v e i n s of m a l es of Gona topus stria t u s K i effer ( A ) , l u n a tus K l u g ( B ) a n d sepsoides West wood ( C ) .

Plectrogon a topus richardsi Mocza r : J . F . Perk i n s 1 976 : 3 2 . A gona topoides stria tus ( K i effer ) : Ponom arenko 1 978 : 2 3 .

F E MALE : a p t e rou s ; l e n g t h 3-4 m m ; h e a d b ro w n or b l ack , w i t h m a n d i b l es , c l ypeu s a n d fro n t part of vertex red d i s h-test aceou s ; occ i p u t red d i s h ; a n t e n n ae b l ack , w i t h segme n ts 1 - 3 ye l l ow ; t hora x , p ropodeum , p e t i o l e a n d a b domen b l ac k ; l eg s red d i sh-ye l l ow , occ a s i on a l l y w i t h c l u b s o f fore femora b row n i s h ; head excavated , s h i n y , g r a n u l a ted ; pronotum crossed b y a s t ron g t r a n svers a l i m press i on , s h i n y , w i t h o u t scu l p t u re , f i n e l y h a i ry ; scu t u m s h i n y , w i t h some l on g i t u d i n a l k ee l s ; m e t a t ho r a x + p ropodeum s h i n y , w i t ho u t scu l p t u re , excep t for p l eu r a a n d pos t e r i or s u rf a ce o f p ropodeum t r a n s verse l y s t r i a te ; met a no t u m smoo t h , n o t t r a n sverse l y s t r i a t e ; meso met a p l e u r a l s u t u re on l y p rox i m a l l y d i s t i nc t ; segmen t 1 of fron t t a r s u s s l i g h t l y shorter t h a n segmen t 4 ( 1 0 : 1 2 ) ; e n l arged c l a w ( F i g . 1 047 ) w i t ho u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d of a l o n g i t u d i n a l f u rrow and w i t h 5-7 peg- l i ke h a i r s or l a mel l ae ; segme n t 5 of fron t t a r s u s ( F i g . 1 047 ) w i t h i n n er s i d e prox i m a l l y serr a t e , w i t h 2-3 rows of 24-40 l ame l l ae e x t en d i n g beyond 0 , 5 l e n g t h of seg men t ; apex w i t h a g roup of 6-40 l ame l l a e ; m ax i l l ary p a l p i w i t h 4-5 seg men t s ; l a b i a l p a l p i w i t h 2-3 segme n ts ; p a l p a l r a t i o s : 4/3 , 5/3 , 5/2 ( i n a fema l e speci men from Orop a , I t a l y , m ax i l l ary p a l p i w i t h 5 segme n t s , r i g h t l a b i a l p a l p i 3-segmen ted , l e f t l a b i a l p a l p i 2-seg men ted ) ; t i b i a l s p u r s 1 , 0 , 1 . MALE : f u l l y w i n g e d ; l e n g t h 1 , 69-2 , 37 mm ; b l ac k ; m a n d i b l es b l ac k , w i t h d i st a l h a l f test aceo u s ; l eg s brow n , w i t h fore t i b i a e test aceous ; a n te n n ae not d i s t a l l y t h i c k e ne d ; a n ten n a l segme n t s i n fol l ow i n g p roport i on s : 6 : 4 , 5 : 6 , 5 : 7 : 7 : 7 : 7 : 7 : 7 : 1 0 ; a n t e n n a I segmen t 3 l ess t h a n 3 , 5 t i mes a s l o n g a s broad ( 2 , 25 ) ; h e a d d u l l , g r a n u l a t e d a n d ret i c u l a t e ru gose ; vertex w i t h

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1 049 - M a l e g e n i t a l ia of Gon a top u s stria t u s K i effer ( l eft h a l f removed )

a s h i n y area between oce l l a r t r i a n g l e a n d eyes ; s h i n y a rea not exten d i n g t o eyes ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 6 ; O l = 3 ; OOl = 5 ; temp l es d i s t i n c t ; occ i p u t conca ve ; sc u t u m d u l l , fu l l y g ra n u l a te d ; not a u l i ces comp l e te , poste r i or l y separa ted ; scute l l u m d u l l , fu l l y g ra n u l a t e d ; met a n o t u m s h i n y , smoot h , w i t h o u t scu l p t u re ; p ropodeum s t ron g l y ret i c u l a t e ru gose , w i t h o u t l o n g i t u d i n a l kee l s on poster i o r s u rface ; fore- w i n g h y a l i n e , w i t h o u t d a r k t r a n s vers a l b a n ds ; r a d i a l v e i n : f i g . 1 048 A ; dorsa l p rocess of gonoforceps ( F i g . 1 049 ) very l on g , s l e n d e r ; b a s i vo l se l l a w i t h o u t r i d g e ; m a x i l l a r y p a l p i w i t h 5 segmen t s ; l ab i a l p a l p i w i t h 2-3 segme n t s ; t i b i a l spurs 1 , 1 , 2 . lOCUS TYP I C US : T r i este ( I t a l y ) ( ho l o t y p e ) ; Kort v e l yes ( H u n g a ry ) ( neot y pe ) . TYP I CA L MAT E R I AL : h o l o t y pe F of G. stri a tus l os t ( Mocze a r 1 965 ) ; n eo t y pe F ! ( des i g n a te d b y Mocz a r 1 96 5 ) i n B U ; h o l o t y pe F ! a n d 1 p a r a t y pe F ! of Pl . ric hardsi i n B M ; h o l o t y p e F ! of A . ta uric u s i n M O . HOST S : i n E n g l a n d Aphrodes b icinctus ( Sc h r k . ) ( W a l off 1 975 : u n der n a m e Plectrogon a top u s richardsi Mocz a r ) . D I ST R I B U T I O N : P a l aearc t i c reg i o n : MONGO L I A ( Po n o m a re n k o 1 978 ) ; U . S . S . R . : Gu rsuf , MO ! F I N LAN D : lojo ( U u d e n m a ) , H E ! POLAND : U j he l y i , B U !

G o n a topod i n a e

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F i g . 1 050 - Che l a of Gon a topus vollenhoveni ( O l m i a n d C u rrado ) ( t y p e )

HUNGARY : SzSzt M i k l o s , B U ! M a r i a b es n y o , B U ! B a kon y s z . l a sz l o , B U ! F o t ( B u dapest ) , B U ! Mohacs i -Sz i ge t , B U ! Godo l l o , B U ! R a v . , B U ! Be i kos ( Bospor u s ) , B U ! Nogradsza k a l , B U ! Csob a n k a , BU! Z amard i -f . , B U ! T a p o l c afo , B U ! B a l a toncs i c s o , B U ! ROMAN I A ( N a gy 1 972 ; Teodorescu 1 967 ) ; C Z E C HOSLOVAK I A : C a r l s b a d , B U ! G E R MANY : T h u r i n g e n , G E ! K y ff h a u ser ( T h u r i n ge n ) , OL ! Be l l i nc h e n ( O der ) , HA ! AUSTR I A : P ra te r ( W i en ) , P ! Aust r i a , CO ! DENMAR K , CO ! SW I T Z E RLAN D : C h u r ( Gr i g i on i ) , ZO ! F RANC E : F on t a i n e b l e a u ( Se i ne-et-Marne ) , H E ! T honon - I es-Ba i n s ( H a u te-Sa vo i e ) , GV ! Cornu s-L ' Hosp i t a l e t du L a rzac (Aveyro n ) , P ! EN GLAN D : St u d l a n d ( Dorset ) , B M ! Shotover ( a xon ) , B M ! Jersey , B M ! D e a l ( Ke n t E a s t ) , B M ! P l y mo u t h ( from n e s t o f Formica fus c a , accord i n g t o K i effer i n K i effer e t M a r s h a l l 1 906 ) , B U ! I R E LAN D : some l oc a l i t i e s i n R i c h a rd s ( 1 93 9 ) ; I TALY : S . Benedetto B e l bo ( C u n eo ) , P L ! Oropa ( Verce l l i ) , BO ! T r i es te , B U ! Sp i l amberto ( Moden a ) , E B ! L ' Aq u i l a , GE ! G R E E C E : Monemv a s i a ( Pe l op o n n esos ) , B U ! T U R K E Y : O rd u , G V !

7 . Gonatop u s vollenhoveni ( O l m i a n d C u rrado)

Neogon a topus vollenhoven i O l m i a n d C u rrado 1 974 :

n . com b .

207 .

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F i g . 1 05 1 - C h e l a of Gon a top u 5 ira n i c u s n . s p .

( t y pe )

F E M AL E : a p te ro u s ; l e n g t h 3 , 1 2 m m ; head b ro w n , w i t h fro n t p a r t of vertex , c l y p e u s a n d m a n d i b l e s y e l l ow-red d i s h ; a n t e n n a e b ro w n , w i t h segmen t s 1 -2 y e l l ow ; prono t u m redd i s h , w i t h b ro w n spots on transversa l i mpress i on , s i des a n d d i sc ; sc u t u m y e l l ow , w i t h a n te r i or s u rface b row n ; scu t e l l urn a n d met a t horax + propodeum b l ac k , w i t h a p ex o f propodeum I i g h t ; a b domen brown -red d i s h , w i t h b row n -b l ac k n u a nces ; l e g s y e l l ow-red d i s h , w i t h coxae and c l u b s of femora brow n i s h ; head exc a v a t e d ; p ro n o t u m s h i n y , crossed by a s t ro n g t r a n sversa I i mpress i o n ; m e t a n o t u m fl a t , s h i n y , t r a n s v ersel y s t r i a t e ; met a t ho r a x + propodeum w i t h s t rong med i a n f u rrow ; pos t e r i o r s u rface of p ropodeum t r a n s verse l y s t r i a t e ; e n l a rged c l a w ( F i g . 1 050 ) w i t ho u t sub a p i c a l too t h , w i t h a s m a l l t oo t h a t t h e e n d of a l on g i t u d i n a l f u rrow a n d w i t h 6 b r i s t l es ; segme n t 5 o f fro n t t a r s u s ( F i g . 1 05 0 ) w i t h 2

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row s of 2 + 1 5 l a m e l l ae ( shortest rOw not extend i n g b e y o n d 0 , 5 l e n g t h of segmen t ) ; apex w i t h a g roup of 16 l am e l l ae ; maxi I l a ry p a l p i w i t h 4 segme n t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k no w n LOCUS T Y P I C US : Ho l l a n d T Y P I CAL MATER I A L : h o l o t y pe F ! i n GE . HOSTS : u n k n o w n D I STR I BU T I O N : P a l aearc t i c reg i o n : HOLLAN D , G E !

8 . Gon a topus iranicus n .

sp.

F E MAL E : a p t e ro u s ; l e n g t h 5 , 62 m m ; h e a d b l ack , w i t h c l y p e u s , m a n d i b l es a n d m a l a r sp ace y e l l ow ; a n te n n ae b l ack , w i t h seg men ts 1 -2 pa r t l y ye l l ow ; t h ora x , p ropodeum a n d a b domen b l ack ; l e g s b l ack , w i t h che l ae , m i d t i b i ae , m i d t a r s i a n d h i n d t a r s i y e l l ow-bro w n ; head exca v a ted , s h i n y , g r a n u l a t ed ; p ro n o t u m crossed b y a s t ro n g t r a n s v e rs a l i mp ress i on , s h i n y ; d i sc smoot h : co l l a r a n d s i des g r a n u l a te d ; scu t u m s h i n y , w i t h s t ro n g l on g i t u d i n a l k ee l s ; scute l l u m s h i n y , smoo t h ; m e t a n o t u m s h i n y , w i t h a t r a n s v e rs a l e l l i p t i c kee l ; sc u t u m l at e ra l l y w i t h 2 po i n t s ; met a t horax + p ropodeum s h i n y , smoot h , weak l y g ra n u l a ted ; pos t e r i or s u rface of p ropod e u m a n d p l e u r a t r a n s verse l y s t r i a t e ; meso-me t a p l eu r a l s u t u re d i s t i nc t a n d com p l e t e ; e n l a rged c l a w ( F i g . 1 05 1 ) w i t ho u t s u b a p i c a l toot h , w i t h a sma l l toot h a t t h e e n d of a l on g i t u d i n a l fu rrow a n d w i t h 4 b r i s t l e s ; segme n t 5 of fron t t a r s u s ( F i g . 1 05 1 ) w i t h i n n e r s i d e p rox i m a l l y serra t e , w i t h 2 rows of 32 l a me l l ae exte n d i n g beyon d 0 , 5 l e n g t h of segme n t ; apex w i t h a g ro u p of at l ea s t 1 7 l a m e l l ae ; ma x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segme n ts ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k no w n LOCUS T Y P I C US : Z i a r a n ( I ra n ) T Y P I CAL MATER I AL : h o l o t y p e F ! i n P G . HOST S : u n k no w n D I ST R I B U T I ON : P a l aearc t i c reg i on : I RAN : Z i a r a n ( Northern I ra n ) , PG !

9 . Gonatopus popovi Ponomaren k o

Gonatopus popovi Ponomarenko 1 965 : 629 . Gonatop u s popovi Ponomarenko :

Ponoma renko 1 97 8 : 2 6 .

F E MALE : a p t erou s ; l en g t h 4 , 5 mm ; h e a d m i s s i n g i n t h e o n l y k n o w n spec i me n ; accord i n g to t h e o r i g i n a l descri p t i on head b l ack , w i t h c l y p e u s , m a nd i b l es a n d a n ten n a l segmen t 1 y e l l o w ; a n te n n a I segme n ts 2- 1 0 b l ack , accord i n g to t h e o r i g i n a l descr i p t i on ; t hora x , propodeum a n d a b domen

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b l ack ; l eg s testaceo u s , w i t h coxae a n d c l u b s of femora b row n -b l ack ; accord i n g to t he o r i g i n a l descr i p t i o n head exc a v a te d ; p ronotum crossed b y a s t ro n g t ra n svers a l i mpress i o n , d u l l , gra n u l ated ; sc u t u m a n d s c u te l l um d u l l , g ra n u l a ted ; sc u t um l a tera l l y w i t h 2 p o i n t s ; m e t a n o t u m h o l l ow beh i n d scute l l um , r u gose ; meta t horax + propodeum d u l l , g ra n u l a ted , w i t h pos ter i or s u rface t r a n s v erse l y s t r i a t e ; p l e u r a not t r a n s verse l y s t r i a t e ; d i sc w i t h a track of med i a n f u rrow ; meso-me t a p l eura l s u t u re d i s t i n c t a n d comp l e te ; fore t a rs a l segmen ts i n fo l l ow i n g propor t i o n s : 2 3 : 5 : 8 : 2 3 : 36 ; e n l arged c l aw ( F i g . 1 052 ) w i t ho u t s u b a p i c a l t oo t h , w i t h a s ma l l tooth at the e n d of a l on g i t u d i n a l f u rrow and w i t h 4 b r i s t l e s ; segme n t 5 of f ro n t tarsus ( F i g . 1 05 2 ) w i t h 2 row s of 9 + 1 8 l ame l l ae extend i n g beyond 0 , 5 l e n g t h of segme n t ; apex w i t h a gro u p of n e ar l y 1 7 l a m e l l ae ; m a x i l l ar y p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k no w n L O C U S TYP I C U S : K i ne l ( U . S . S . R . ) TYP I CA L MATE R I AL : h o l o t y p e F ! i n MO . HOSTS : i n U . S . S . R . Thamnotettix sp . ( Ponomare n k o 1 965 , 1 97 8 ) D I ST R I BU T I ON : P a l a e a rct i c reg i o n : U . S . S . R . : K i ne l ( K u y b y shev Prov i nc e ) , MO !

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1 0 . Gon a top u s a l b ol in e a t u s K i effer

Gon a top u s a l b ol in e a tus K i effer i n K i effer et M a rs h a l l 1 905 : 95 . Gon a top u s a l b oline a t u s K i effer v a r . interruptus K i effer i n K i effer e t M a r s h a l l 1 905 : 95 ( s y n . p roposed b y A b d u l -N o u r 1 974 ) ; locus typ ic u s Bone ( A l ger i a ) . Gon a top u s polychrom u s Mars h a l l i n K i effer et M a rs h a l l 1 90 5 : 1 05 ( n . s y n . ) ; locus typ i c u s C e t t e ( F rance ) . Pachygona top u s (Pla tygon a topus) polychrom u s ( M a rs h a l l ) : K i effer i n K i effer et M a rs h a l l 1 906 : 500 . Pla tygon a top u s a lb olineatus ( K i effer ) : K i effer 1 9 1 4b : 9 1 . Pla tygona topus in terrup t u s ( K i effer ) : K i effer 1 9 1 4b : 92 . Pla tygon a top u s polychromus ( M a rs h a l l ) : K i effer 1 9 1 4b : 93 . Pla tygon a top u s polychrom u s ( M a r s h a l l ) : B e r l a n d 1 92 8 : 1 49 . Pla tygon a top u s polychromus ( M arsha I I ) : Bern a rd 1 935b : 5 3 . Pla tygon a top u s a l b olinea t u s ( K i effe r ) v a r . deserticus Ben o i t 1 95 1 a : 299 ( s y n . proposed by O l m i a n d C u rrado 1 976a ) ; locus typ ic u s G i a l o oas i s ( L i b y a ) . Chalcogon a topus polychro m u s ( M arsh a l l ) : Mocz a r 1 96 5 : 388 . Chalcogon a top u s a l b olineatus ( K i effer ) : Abd u l -Nour 1 974 : 1 0 1 3 . Ma deca gon a top u s a lbolinea t u s ( K i effer ) : 0 1 m i a n d C u rrado 1 976a : 1 66 .

F E MALE : ap tero u s ; l e n g t h 3 , 28-4 , 1 2 m m ; head redd i s h , w i t h f ro n t p a r t of vertex , c l y p e u s and m a n d i b l es w h i t i s h ; a n te n n ae w h i t i s h , w i t h segmen t s 7- 1 0 d a r k ; t horax a n d propodeum red d i s h ; pet i o l e b l ac k ; a b domen b ro w n b l ac k , w i t h ter g i t e 1 p a r t l y red d i s h ; l e g s y e l l ow - w h i t i s h , w i t h ex tern a l faces of t i b i ae a n d femora b ro w n i sh ; occa s i on a l l y ( v a r . deserticus Beno i t ) body a l most f u l l y w h i t i s h ; head weak l y exc a v ated , s h i n y , h a i r l es s , w i t h o u t scu l p t u r e , w i t h some l o n g i t u d i n a l kee l s ; pronotum crossed b y a s t ron g t r a n s vers a l i m press i o n , smoo t h , h a i r l es s , w i t h o u t scu l p t u re , w i t h some l o n g i t u d i n a l k ee l s on the s i d es ; s c u t u m s h i n y , w i t h some l on g i t u d i n a l k ee l s ; sc u te l l u m smooth , s h i n y , very i nc l i ne d ; m e t a n o t u m h o l l o w beh i n d sc u te l l u m , w i t h some t r a n s v ersa l kee l s ; me t a t horax + propodeum w i t h a s t ron g med i a n fu rrow ; pos t e r i o r s u rface of p ropodeum tran s verse l y s t r i a t e ; meso-me t a p l e ura l s u t u re d i s t i n c t a n d comp l e te ; segmen t 1 o f fron t t a r s u s l o n ge r t h a n segmen t 4 ( 1 4 : 1 1 ) ; en l a rged c l a w ( F i g . 1 05 3 ) w i t ho u t s U b a p i c a l toot h , w i t h a sm a l l too t h a t t h e e n d o f a l o n g i t u d i n a l f u rrow a n d w i t h 2 peg- l i k e h a i r s ; segme n t 5 of fro n t t a r s u s ( F i g . 1 05 3 ) w i t h i n n er s i de prox i m a l l y not serr a t e ( occas i o n a l l y serrate ) , w i t h 2 rows of 2 ( p rox i m a l ) + 1 6- 1 9 l a me l l ae beg i n n i n g i n prox i m a l t h i rd ; apex w i t h a group of 7- 1 0 l ame l l ae ( 1 -3 l a me l l ae are m u c h l on ger t h a n o t he r l ame l l ae ) ; ma x i l l a ry pa l p i w i t h 6 segme n t s ; l a b i a l p a l p i w i t h 3 segmen ts ; t i b i a l spurs 1 , 0 , 1 . MALE : u n k no w n LOCUS T Y P I C US : Bone ( A l ge r i a )

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TYP I CAL MATE R I AL : h o l o t y pe F ! of G . a lbolinea t u s i n M T ; h o l ot y pe F ! of G. in terrup t u s i n M T ; h o l o t y pe F ! of G . polychromus i n B U ; h o l o t y pe F ! a n d 4 p a r a t y p es F F ! of Pl . deserticus i n GE ; 2 p a r a t ypes F F ! of Pl. deserticus in T V . HOST S : u n k n o w n D I ST R I B U T I ON : P a l aearc t i c reg i o n : F RANC E : C e t t e ( He r a u l t ) , B U ! V i l l epey ( Bernard 1 935b ) ; L a R i ne l l e ( Corse ) , P ! I TALY : Mess i n a , CO ! ALGER I A : Bon e , M T ! L I BYA : G i a l o oas i s ( C i re n a i c a ) , GE ! T V ! EGYP T : W a d i ' O f , W A !

1 1 . Gon a topus s u arezi n . sp .

F E MALE : aptero u s ; l e n g t h 4 mm ; redd i s h , w i t h a n te n n a l segmen t s 9- 1 0 b row n ; sc u t e I I u m a n d meso-metap l e u r a l s u t u re brow n ; ab domen p a r t I y b row n ; m i d a n d h i n d coxae p a r t l y brow n ; head exc a v a te d , sh i n y , smoo t h , w i t hou t sc u l p t u re ; pronot u m crossed b y a strong tra nsvers a l i mp ress i on , s h i n y , smoo t h , w i t ho u t sc u l p t u re ; sc u t u m s h i n y , w i t h some l o n g i t u d i n a l k ee l s ; scute l l u m sh i n y , smoo t h , w i t hou t scu l p t u re ; m e t a n o t u m s h i n y , very l on g , t r a n s verse l y st r i a t e ; meso-metap l e u ra l s u t u re d i s t i n c t ; meta t horax + p ropode u m sh i n y , smoot h , w i t ho u t scu l p t u re , w i t h poster i or s u rface t r a n sverse l y s t r i a t e ; d i sc w i t h a weak med i a n fu rrow ; p l eu r a n o t t r a n s verse l y s t r i a te ; seg m e n t 1 o f fro n t t a r s u s s l i g h t l y shorter t h a n segme n t 4 ( 1 8 : 1 9 ) ; e n l arged c l a w ( F i g . 1 05 4 ) w i t h o u t s u b a p i c a l toot h , w i t h a sm a l l tooth at t h e end of a l on g i t u d i n a l f u rrow a n d w i t h 1 0 peg- l i k e h a i rs or b r i s t l es ; segme n t 5 of front t a rs u s ( F i g . 1 054 ) w i t h i n ne r s i d e prox i m a l l y serra te , w i t h 2 row s of 25 l ame l l ae ( s hortest row not exten d i n g beyond 0 , 5 l e n g t h of segmen t ) ; apex w i t h a g ro u p of 1 0 l a me l l ae ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i th 3 segmen ts ; t i b i a l spurs 1 , 0 , 1 . MAL E : u n k n o w n LOCUS TYP I C U S : Ameskrou ( Morocco ) T Y P I CAL MAT E R I AL : h o l o t y pe F ! i n SU o H05T S : u n k no w n D I ST R I B U T I ON : P a l aearc t i c reg i o n : MOROCCO : Amesk ro u ( A g ad i r ) , SU I NOTES : t h e spec i es i s n a med i n honor of M r . F r a n c i sco J . S u arez .

1 2 . Gonatopus dimidi a t u s S a h l berg

Gon a topus dimidia t u s Sa h l berg 1 9 1 0 : Neogon a topus dimidia t u s ( S a h l berg ) :

17. R i c h a rd s 1 97 2 : 541 .

F E MALE : a p te ro u s ; l en g t h 4 mm ; head , pro n o t u m a n d sc u t u m fu I I y testaceou s-re d d i s h ; a n t e n n ae brown , w i t h segments 1 -3 y e l l ow ; meta t horax

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p ropodeu m , p e t i o l e a n d a b domen b l ac k ( excep t for apex of p ropode u m l i g h t ) ; l eg s y e l l ow-redd i s h ; h e a d exca v a t e d , d u l l , h a i r l es s , a l u t aceo u s ; p ronot u m crossed by a s t ro n g t r a n svers a l i m p ress i on , s h i n y , w i t h o u t scu l p t u re ; s c u t u m s h i n y , w i t h some w e a k l o n g i t u d i n a l k ee l s ; m e t a n o t u m n o t t ra n s v e rse l y s t r i a t e ; m e t a t horax + p ropodeum s h i n y , smoo t h , w i t h o u t scu l p t u r e , excep t f o r p l e u r a a n d poste r i or s u rface of p ropodeum t ra n s verse l y s t r i a t e ; meso-me t a p l e u r a l s u t u r e d i s t i nc t a n d com p l e te ; segment 1 of front t a r s u s s l i g h t l y shorter t h a n segment 4 ( 1 5 : 1 7 ) ; e n l arged c l a w ( F i g . 1 05 5 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l tooth a t t h e e n d of a l o n g i t u d i n a l f u r row a n d w i t h 4 p e g - l i k e h a i rs ; s e g me n t 5 o f front t a rs u s ( F i g . 1 05 5 ) w i t h i n n e r s i d e p rox i m a l l y serr a t e , w i t h 2 r o w s of 3 + 24 l a m e l l ae ( shortest row not e x t e n d i n g b e y o n d 0 , 5 l en g t h of segmen t ) ; apex w i t h a g ro u p of 9 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 4 segme n t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k no w n L O C U S T Y P I C U S : Jer i c ho ( I sr a e l ) T Y P I CA L M A T E R I AL : H O ST S : u n k no w n D I S T R I B U T I ON :

holotype F !

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F E MAL E : a p terou s ; l e n g t h 2 , 68-3 , 37 m m ; b l ac k , w i t h m a n d i b l es , c l y pe u s a n d fron t p a r t of v e r t e x y e l l ow ; a n t en n ae b l ac k , w i t h segments 1 -2 y e l l ow ; l eg s b row n , w i t h t roc h a n ters , s t a l k s of femora a n d t a r s i l i g h t ; h e a d ( F i g . 1 056 A ) very exca v a ted , s h i n y , g r a n u l a ted ; p ro n o t u m crossed b y a s t ro n g t r a n s v e rs a l i m p re s s i on , s h i n y , g r a n u l a ted ; sc u t u m d u l l , g r a n u l a ted ; m e t a n o t u m t r a n s v e rs e l y s t r i a te ; m e t a t ho r a x + p ropodeum g r a n u l a t e d , w i t h p l e u r a a n d pos t e r i o r su rf ace of p ropodeum t r a n s v e r se l y s t r i a t e ; a n t e r i o r s u rface of m e t a t ho r a x + p ropode u m w i t h some l on g i t u d i n a l k ee l s ; meso m e t a p l eu r a l s u t u re o b so l e t e ; occa s i on a l l y f u l l y v i s i b l e , b u t i n d i s t i n c t ; occas i on a l l y on l y v i s i b l e n e a r m e t a n o t u m ; fore t a rsa l segmen t s i n fo l l ow i n g

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p ropo r t i on s : 1 4 : 3 : 5 : 1 3 : 20 ; en l arged c l a w ( F i g . 1 05 7 ) w i t h o u t s u b a p i c a l too t h , w i t h a s ma l l toot h a t t h e e n d of a l on g i t u d i n a l f u rrow a n d w i t h 2 peg- l i ke h a i rs or b r i s t l es ; segmen t 5 of front t a r s u s ( F i g . 1 057 ) w i t h i n ner s i de p rox i m a l l y serra t e , w i t h 1 row of 20-25 l ame l l a e ; occas i o n a l l y w i t h 2 row s a n d i n t h i s case shortest row n o t exten d i n g beyond 0 , 5 l en g t h of segmen t ; apex w i t h a g roup o f 1 0- 1 3 l ame l l a e ; m a x i I l a r y p a l p i w i t h 5 segme n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k n ow n LOCUS TYP I C U S : B e t a n c u r i a ( C a n a r y I s l a n d s ) TYP I CAL MATER I A L : h o l o t y p e F ! i n HE ; 1 p a r a t y pe F ! i n M D ; 1 p a r a t y p e F! in OL. HOST S : u n k no w n D I ST R I BU T I O N : P a l aearc t i c reg i o n : SPA I N : Poz u e l o , M D ! CANARY I SLAN D S : Betancu r i a ( F uerte ven t u r a ) , HE ! MOROCCO : M e l i l l a , O L !

1 4 . Gon a top u s unilinea tus K i effer

Gon a top u s unilin e a t u s K i effer 1 904a : 358 . nee Gon a top u s unilin e a t u s v a r i e t e K i effer 1 904a : 359 . Neogon a top u s unilin e a tus ( K i effe r ) : O l m i a n d C u rrado 1 974 : 191 .

F E MALE : a p t e ro u s ; l e n g t h 3 , 80-4 , 87 mm ; h e a d test aceou s , w i t h a b ro w n s p "' , on vertex ; occ a s i o n a l l y t wo l a tera l b ro w n s p o t s a re prese n t n e a r temp l e s ; a n t e n n a e b l ack , w i t h seg men ts 1 -2 ( occa s i o n a l l y a l so p a r t o f segmen t 3 ) y e l l ow ; p ron o t u m w i t h co l l a r a l mo s t fu l l y test aceous o r b l aCk w i t h a n t e r i o r m a rg i n test aceo u s ; d i sc test aceous w i t h a n te r i or m a r g i n b l ack ; scu t u m b l a ck ( i n h o l o t y p e ) or red d i s h w i t h marg i n s b l ack ( i n a s p ec i men from Empon a , Greece ) ; scut e l l u m , meta t horax + p ropodeu m , p e t i o l e a n d a b domen b l ack ( e xcep t for a pex o f p ropodeum l i g h t ) ; l e g s y e l l ow redd i s h , w i t h b ro w n s p o t s on co xae ; occ a s i on a l l y c l ub s of fore femora b row n i s h ; head excava ted , s h i n y , w i t h o u t scu l p t u re , f i ne l y h a i r y ; p ro n o t u m crossed b y a s t ro n g t r a n s v e r s a l i m press i o n , d u I I , w i t hou t scu l p t u re , f i n e l y h a i r y ; sc u t u m d u l l , w i t h some l on g i t u d i n a l k ee l s ; m e t a n o t u m s h i n y , smoot h , w i t h o u t scu l p t u r e , w i t h some short , t r a n sversa l s t r i a e ; met a t horax + propodeum ( F i g . 1 060 A ) s h i n y , w i t ho u t scu l p t u re , w i t h p l e u r a a n d pos ter i o r su rface tran s verse l y s t r i a t e ; meso-me t a p l e ura l s u t ure d i s t i n c t a n d com p l e te ; segme n t 1 of fron t t a r s u s s horter t h a n segm e n t 4 ( 1 6 : 2 0 ) ; en l a rg e d c l a w ( F i g . 1 05 8 ) w i t h o u t s u b a p i c a l toot h , w i t h a sm a l l too t h at t h e end of a l on g i t u d i n a l fu rrow a n d w i t h 3-5 peg- l i ke h a i rs ; segmen t 5 of fron t t a rs u s ( F i g . 1 05 8 ) w i t h i n n er s i de p ro x i ma l l y serra t e , w i t h 2 rows of 20-24 + 2-5 l a m e l l ae ( s hortest row not exten d i n g beyon d 0 , 5 l e n g t h o f seg me n t ) ; apex w i t h a g ro u p o f 1 0- 1 4 l am e l l a e ; m a x i l l a ry p a l p i w i t h 4 segme n t s ; l ab i a l p a l p i w i t h 2 segmen t s ; t i b i a l

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spurs 1 , 0 , 1 . MAL E : u n k n o w n L O C U S TYP I C U S : Temp i o P a u s a n i a ( I t a l y ) TYP I CA L MAT E R I A L : h o l o t y p e F ! i n GE . HOSTS : u n k no w n D I ST R I BU T I ON : P a l aearc t i c reg i o n : I T ALY : Tem p i o P a u s a n i a ( Sa s s a r i ) , GE ! G R E E C E : E m p o n a ( R hodes ) , GV !

1 5 . Gon a top u s a egyp t i a c u s n . s p .

F EMALE : a p terou s ; l e n g t h 2 , 8 1 m m ; head b ro w n , w i t h fro n t p a r t o f verte x , c l ypeus a n d m a n d i b l es test aceou s ; a n te n n ae b row n , w i t h seg m e n t s 1 -2 test aceou s ; t horax a n d propodeum test aceou s-re d d i s h ; pet i o l e b l ack ; abdomen brow n ; l e g s testaceous ; a n t e n n ae d i s t a l l y t h i ckened ; a n te n n a l segme n t s i n fol l ow i n g propo r t i on s : 6 : 4 : 1 0 : 6 : 5 : 5 : 4 , 5 : 4 : 4 : 6 ; head weak l y exca v a te d , d u l l , g ra n u l ated ; f ron t a l l i ne com p l ete ; occ i p i t a l c a r i n a i n v i s i b l e ; POL ; 1 ; O L ; 2 ; OOL ; 5 ; pronotum crossed b y a s t ro n g t r a n svers a l i m press i on , d u l l , weak l y g r a n u l a te d ; sc u t u m d u l l , g r a n u l a ted ; m e t a n o t u m w i thout l a ter a l po i n t s , ho l l ow beh i n d scute l l u m ; meso meta p l e u r a l s u t u re d i s t i n c t a n d com p l e te ; m e t a t horax + p ropodeum weak l y g r a n u l a te d , w i t h p l e ura a n d pos t e r i o r su rface t r a n sve rse l y s t r i a t e ; fore t a r s a l seg m e n t s i n fo l l ow i n g proport i o n s : 1 2 : 2 : 3 : 1 0 : 1 5 ; en l arged c l a w ( F i g . 1 059 ) w i t ho u t s u b a p i c a l too t h , w i t h a s m a l l too t h at the e n d of a l o n g i t u d i n a l fu rrow a n d w i t h 7 peg- l i ke h a i rs ; seg m e n t 5 of front t a r s u s

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( F i g . 1 0 59 ) w i t h 1 row of 1 2 l a me l l ae ; apex w i t h a g ro u p of 7 l a me l l ae ; m a x i l l a r y p a l p i w i t h 5 s e g m e n t s ; l a b i a l p a l p i w i t h 2 segm e n t s ; t i b i a l spurs 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : So l o u m ( E g y p t ) TYP I CA L MAT E R I AL : h o l o t y p e F ! i n WA . HOST S : u n k n o w n D I ST R I B UT I O N :

P a l a e a rc t i c reg i o n :

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1 6 . Gon a topus plumbeus n . s p .

F E MAL E : a p t e ro u s ; l e n g t h 4 , 37 m m ; head b l a ck , w i t h frons a l o n g orb i t s , c l y p e u s a n d m a n d i b l e s y e l l ow ; a n t e n n a e , t ho r a x , p ropod e u m , p e t i o l e a n d a b domen b l ac k ; l e g s t e s t a c eo u s , w i t h coxae a n d c l u b s o f femora p a r t l y b l ac k ; head e x c a v a t e d , s h i n y , w i t h o u t sc u l p t u re , excep t for occ i p u t g ran u l a t e d ; p ro no t u m sh i n y , crossed b y a s t ro n g t r a n sv e rs a l i m pres s i on , w i t h co l l a r g ra n u l a t ed ; d i sc w i t ho u t sc u l p t u re ; scu t u m s h i n y , smoo t h , w i t ho u t sc u l p t u re ; met a n o t u m s h i n y , t r a n sv e rse l y s t r i a te ; m e t a t ho r a x + p ropod e u m ( F i g . 1 060 B ) s h i n y , w i t ho u t scu l p t u re , excep t for p l e u r a a n d poster i o r s u rf a ce t r a n sverse l y s t r i a t e ; d i sc a n d a n te r i o r s u rface w i t h w e a k l o n g i t u d i n a l kee l s ; meso-me t a p l e u ra l s u t u re d i s t i n c t a n d comp l e t e ; se g m e n t 1 o f f ro n t t a r s u s s l i g h t l y l o n ge r t h a n seg m e n t 4 ( 1 5 : 1 3 ) ; en l a rged c l a w (Fig.

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l o n g i t u d i n a l f u r ro w a n d w i t h 5 b r i s t l es ; s e g m e n t 5 o f f ro n t t a r s u s ( F i g . 1 06 1 ) w i t h 1 row of 20 l a me l l ae ; apex w i t h a g ro u p of 9 l am e l l ae ; m a x i l l a r y p a l p i w i t h 4 seg men t s ; l ab i a l p a l p i w i t h 2 seg men t s ; t i b i a l sp urs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : P a l m ac h i m ( I s r ae l )

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TYP I CAL MA T E R I AL : h o l o t y pe F ! i n B M . HOST S : u n k n o w n D I ST R I BU T I O N : P a l aearc t i c reg i o n : I SRA E L : P a l m a c h i m , BM !

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Gon a top u s b ar b a tellus R i c h a rd s

Gon a top u s krogeri H e l l en 1 935 : 8 ( n omen n u d um ) . Gon a topus b ar b a tellus R i c h a rds 1 9 39 : 2 1 3 . Gon a topus b arb a tellus R i c h a rds : H e l l en 1 9 53 : 95 .

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c l y p e u s a n d m a n d i b l es y e l l o w ; a n t e n n ae b l ack , w i t h seg men t s 1 - 2 p ro n o t u m b l ack ; sc u t u m y e l l o w , w i t h a n t e r i o r s u rface b l ack ;

s c u t e l l u m , m e t a t ho r a x + p ro p o d e u m a n d p e t i o l e b l ack , e x c e p t for a p e x of p ro p o d e u m l i g h t ; a b domen b l a c k , w i t h red d i s h n u ances ; l eg s y e l l ow , w i t h coxae p a r t l y b l ac k a n d c l u b s of femora b row n i s h ; h e a d w e a k l y exca v a ted , du I I , g ra n u I a t e d , h a i ry ; p ro n o t u m crossed b y a s t r o n g t r a n s v e r s a I i m p ress i o n , d u l l , g r a n u l a ted ; sc u t u m d u l l , w i t h some l o n g i t u d i n a l k ee l s ; m e t a no t u m s t ron g l y ho l l o w b e h i n d sc u te l l u m ; meso-me t a p l e u ra l s u t u re d i s t i n c t a n d comp l e t e ; m e t a t ho r a x + p ro p o d e u m d u l l , g r a n u l a ted , w i t h t rack of med i a n f u rrow , w i t h p l e u r a t r a n s verse l y s t r i a te ; pos t e r i o r s u rf a ce of propod e u m a l m o s t fu l l y n o t s t r i a t e , o n l y w i t h some s t r i ae on pos t e r i o r h a l f ; s e g m e n t 1 o f fron t t a r s u s s l i g h t l y s h o r t e r t h a n seg m e n t 4 ( 1 1 : 1 2 ) ; en l a r g e d c l a w ( F i g . 1 06 2 ) w i t ho u t su b a p i c a l too t h , w i t h a sm a l l too t h a t t h e e n d o f a l o n g i t u d i n a l f u rrow a n d w i t h 1 -2 b r i s t l es ; occ a s i o n a l l y w i t ho u t b r i s t l e s ; seg men t 5 of f ro n t t a r s u s ( F i g . 1 062 ) w i t h i n n e r s i d e u s u a l l y p rox i m a l l y serra t e ( oc c a s i o n a l l y n o t s e r r a t e ) , w i t h 1 row of 1 6-23 l a m e l l ae ; a p e x w i t h a g ro u p o f 7- 1 1 l a me l l a e ; m a x i l l a ry p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 2 seg men t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : I c k l i n g h a m ( E ng l a n d ) ( ho l o t y p e w i t h no l oc a l i t y l a b e l : a m p ro p os i n g to des i g n a t e as locus typ ic u s t h e l oc a l i t y on p a r a t y p e l ab e l ) . TYP I CA L MAT E R I AL : h o l o t y p e F ! i n OX ; 1 p a r a t y p e F i n M R . HOST S : u n k n o w n D I ST R I B U T I O N : P a l a e a rc t i c reg i o n : F I N LAN D : Sei s k a r i ( R . K . ) , H E ! T e r i j o k i ( W . -H . ) , H E ! GE RMANY : l e t t i n-Hohen ( H a l l e-S ) , O l ! He i de b r i n k ( W o l l i n ) , HA ! D E N MAR K , C O ! HOLLAN D : Me i j en d e l , LE ! T e rsche l l i n g , LE ! B E L G I U M ( J . F . Perk i n s 1 976 ) ; ENGLA N D : I c k l i n g h a m ( S uffo l k E a s t ) ( R i c h a r d s 1 93 9 ) .

1 8 . Gon a topus a ter n e w n a m e

nec Gon a topus plan iceps K i effer 1 904a : 355 . Gon a top u s hispanicus K i effer : H e l l en 1 930 : 4 . Pla tygon a top u s p l a n iceps ( K i e ffer ) : R i c h a r d s 1 939 : 2 1 9 . nec Plectrogon a top u s ric h ardsi Moczar 1 965 : 396 . Pla tygon a top u s ric h a rdsi O l m i a n d C u rrado 1 974 : 1 77 ( n .

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s y n . : p reocc u p i ed b y Plectrogon a top u s richardsi Moczar 1 965 , now Gon a top u s rich ards i ) .

F E MAL E : a p t e rou s ; l e n g t h 2 , 5 m m ; b l ac k , w i t h m a n d i b l es , c l y p e u s a n d fron t p a r t of v e r t e x ( more a l o n g orb i t s ) y e l l o w ; a n te n n ae b l a ck , w i t h segme n t s 1 -2 y e l l ow ; l eg s b ro w n , w i t h t i b i ae a n d t a r s i l i g h t ; h e a d ( F i g . 1 056 B ) a l mos t f l a t , weak l y exc a v a t e d , d u l l , g ra n u l a t ed ; p ro n o t u m crossed by a s t r o n g t r a n s v e rs a l i m p ress i on , s h i n y , g r a n u l a ted ; scu t u m d u l l , g ra n u I a ted ; m e t a n o t u m d u I I , t r a n s v erse I y s t r i a te ; m e t a t h o r a x + p ropode u m d u l l , g r a n c: l a ted , w i t h p l e u r a a n d pos t e r i o r s u rface t r a n sverse l y s t r i a t e ; meso-m e t a p l eu r a l s u t u re o b so l e t e ; fore t a r s a l se g m e n t s i n fo l l ow i n g p ropor t i o n s : 1 1 : 2 : 3 : 1 1 : 1 7 ; e n l a rged c l a w ( F i g . 1 063 ) w i t ho u t s u b a p i c a l t oo t h , w i t h a s ma l l too t h a t t he e n d o f a l o n g i t u d i n a l f u rrow a n d w i t h 9 peg- l i k e h a i rs ; se g m e n t 5 of fro n t t a r s u s ( F i g . 1 063 ) w i t h i n n e r s i de p rox i m a l l y serra te , w i t h 1 row of 1 7 l a me l l ae ; a p e x w i t h a g ro u p of 4-8

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1 064 - C h e l a of Gon a top u 5 lindbergi He l l e n

l a me l l a e ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k n o n w L O C U S TYP I C U S : Corse ( F r a n c e ) TYP I CA L MAT E R I A L : ho l o t y p e F ! i n B M .

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1 9 . Gon a topus lindb ergi He l l en

Gon a topus { Trichogona top u s } lindbergi H e l l e n 1 930 : 2 .

F E MA L E : a p te ro u s ; l e n g t h 6 , 75 m m ; b l ack , w i t h f ron t p a r t o f vertex ( more a l o n g orb i t s ) , c l y pe u s a n d m a n d i b l es y e l l o w ; a n t en n ae b row n , w i t h segme n t s 1 -2 y e l l o w ; l e g s b l ack , w i t h t a r s i b row n ; head excav a ted , d u l l , g ra n u l a te d ; frons w i t h i rreg u l a r kee l s ; occ i p i t a l c a r i n a i n v i s i b l e ; p ronot u m crossed b y a s t ron g t r a n s v e rs a l i m p ress i o n , g r a n u l a ted , d u l l , w i t h n u merous l on g i t u d i n a l kee l s ; sc u t u m d u l l , g r a n u l a ted , w i t h some l on g i t u d i n a l kee l s , l a tera l l y w i t h 2 p o i n t s ; s c u te l l u m sma l l , smoo t h , s h i n y ; meta t horax + propodeum d u l l , g r a n u l a ted , w i t h p l e u ra a n d pos t e r i o r s u rface t r a n s verse l y st r i a t e ; a n te r i o r s u rf ace scu l p t u red b y l o n g i t u d i n a l s t r i ae ; meso-me t a p l e u r a l s u t u re w i t h on l y t h e prox i m a l t h i rd d i s t i nc t ; d i s t a l p a r t obso l e t e ; segment 1 o f fro n t t a r s u s l o n g e r t h a n segmen t 4 ( 22 : 1 7 ) ; e n l a r g e d c l a w ( F i g . 1 064 ) w i t h o u t s u b ap i c a l too t h , w i t h a sm a l l tooth a t t h e e n d of a l on g i t u d i n a l f u rrow a n d w i t h 7 m i n u te peg- l i ke h a i rs ; segmen t 5 of fro n t t a r s u s ( F i g . 1 064 ) w i t h 1 row of 1 1 l a me l l a e ; a p ex w i t h a g ro u p o f a t l ea s t 1 4 l am e l l ae ; m a x i l l ary p a l p i w i t h 6 seg men t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MAL E : u n k n o w n LOCUS TYP I C US : M a l a g a ( Sp a i n ) TYP I CA L MAT E R I AL : h o l o t y pe F ! i n H E . HOST S : u n k no w n D I ST R I BU T I O N : P a l aearc t i c reg i on : SPA I N : M a l a g a ( A n d a l uc i a ) , HE !

2 0 . Gon a top u s l u n a t u s K l u g

Gon a top u s l u n a tus K l u g 1 8 1 0 : 1 64 . Gon a top u s b ifa s c i a t u s K i effer 1 904a : 360 ( n . sy n . ) ; locus typ i c u s G i g l i o I s l a n d ( I t a l y ) . Gon a top u s gracilicornis K i effer 1 904a : 361 ( n . sy n . ) ; locus typicus G i g l i o I s l a n d ( I t a l y ) . Gon a top u s filicornis K i effer i n K i effer et M a r s h a l l 1 905 : 1 1 2 ( s y n . proposed by A b d u l -Nour 1 974 ) ; locus typicus M o n t - de-Marsan ( F r a n ce ) . Gon a top u s myrmecop h i l u s K i effer i n K i effer et M a r s h a l l 1 90 5 : 1 1 3 ( s y n . proposed b y R i c h a r d s 1 939 ) ; locus typ i c u s

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Fig.

1 066 - F em a l e of Gona top u s l u n a tus K l u g

L u xem b u r g . Gona top us gracilis K i effer i n K i effer et M a rs h a l l 1 90 5 : 1 1 5 ( s y n . proposed b y A b d u I -N o u r 1 974 ) ; locus typicus M o n t - d e - M a r s a n ( F r a n ce ) . Gona top u s marsh a l l i K i effer i n K i e ffer et M a r s h a I I 1 905 : 1 2 1 ( s y n . p roposed b y Moc z a r 1 965 ) ; locus typic us S w a n a g e ( Engl and ) Gon a topus gracilipes K i effer i n K i effer et M a r s h a I I 1 906 : 508 ( s y n . proposed b y Moczar 1 965 ) ; locus typ icus Szomo tor ( Hu n gary ) Gona top u s raptoripes S t r a n d 1 9 1 9 : 202 ( n . s y n . ) ; locus typ ic u s R a t z d orf ( Ge r m a n y ) . Gon a topus marshalli K i e ffe r : L i c h t e n s te i n a n d P i c a r d 1 920 : 54 . Gona top us filicornis K i effe r : Ber l a n d 1 928 : 1 52 . Gon a topus gracilis K i effer : Ber l a n d 1 92 8 : 1 52 . Gon a top u s myrmecop h i l u s K i effer : B er l a n d 1 928 : 1 52 . Gona top u s marshalli K i effe r : Ber l a n d 1 9 28 : 1 53 . Gon a topus myrmecoph il u s K i e ffer : G i ord a n i So i k a 1 932 : 1 26 . Gon a top u5 l u n a tus K I u g : B e r n a r d 1 939 : 1 1 2 . Neogo n a topus l u n a tus ( K l u g ) : R i c h a r d s 1 939 : 2 1 6 . Neogon a top u s l u n a tus ( K l u g ) : N a g y 1 967a : 1 24 . Neogo n a top us s p . 2 R i c h a r d s 1 97 2 : 543 ( n . sy n . ) . Neogon a top u s b ifa s c i a t u s ( K i e ffer ) : O l m i a n d C u rr a d o 1 974 : 202. Neogo n a top u s l u n a tus ( K l u g ) : O l m i a n d C u rr a d o 1 974 : 207 . Neogon a top u s l u n a t u s ( K I u g ) : A b d u I -N o u r 1 974 : 1 0 1 2 . •

•

1 543

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8

Fig.

1 067 - A n t e n n ae of fem a l es of Gon a top u s c a n a riensis n . s p . a n d l u n a t u s K l u g from Fon t a i n eb l ea u ( F ra n ce ) ( B ) .

( t ype ) ( A )

Neogon a top u s l u n a t u 5 ( K l u g ) : Ab d u l -Nour 1 976 : 273 . Neogon a top u s l u n a t u s ( K l u g ) : Ponomarenko 1 978 : 27 . Gon a top u 5 l u n a t u s K I u g : Moczar 1 983a : 1 94 .

F E MA L E ( F i g s 1 065 , 1 066 ) : ap tero u s ; l e n g t h 2 , 3 1 -3 , 56 m m ; head y e l l ow redd i s h , w i t h o r w i t ho u t b ro w n b a n d on vert e x ; a n t e n n ae b row n , w i t h segme n t s 1 -2 y e l l o w ; prono t u m red d i s h - te s t aceou s , occas i on a l l y w i t h t r a n s v e rs a l i m pres s i o n a n d most p a r t of d i sc b ro w n ; sc u tu m y e l l o w , occ as i o n a I I y w i t h a n te r i o r a n d pos t e r i o r m a r g i n s b row n ; sc u te l l um a n d m e t a t horax + p ropodeum b l ac k ( w i t h a p e x of p ropod e u m l i g h t ) o r occas i o n a l l y y e l l o w - redd i s h ; a b domen b ro w n or b l ack , w i t h re d d i s h n u ances ; l e g s y e l l o w -redd i s h , occas i o n a l l y w i t h coxae a n d c l u b s o f femora b row n i s h ; head exca v a t e d , s h i n y , w i t ho u t scu l p t u re , excep t for occ i p u t a n d f ro n t p a r t of vertex g r a n u l a ted ; a n t e n n ae ( F i g . 1 067 B ) s l e n d e r , more t h a n 3 , 5 t i mes as l o n g a s hea d ; pron o t u m crossed by a s t ro n g t r a n sversa l i m p ress i on , s h i n y , g r a n u l a te d ; sc u t u m s h i n y , w i t h some l on g i t u d i n a l k ee l s ; m e t a n o t u m f l a t , weak l y t r a n s v erse l y s t r i a t e ; m e t a t ho r a x + propodeum d u l l ( occa s i on a l l y s h i n y ) , s t r on g l y or weak l y g ra n u l a t e d , w i t h t r a c k of m e d i a n f u r row ; p l eu r a not t r a n s ve rse l y s t r i a te ; pos t e r i o r s u rface o f propode u m t r a n s verse l y s t r i a t e ; meso-me t a p l eu r a l s u t u re obso l e t e ; fore t a rs a l segme n t s i n fol l o w i n g p roport i o n s : 1 3 : 2 : 4 : 1 3 : 2 1 ; en l a r ged c l a w ( F i g . 1 068 ) w i t ho u t s u b a p i c a l too t h , w i t h a sm a l l too t h a t the e n d of a l on g i t u d i n a l f u rrow a n d w i t h 4-7 peg- I i ke h a i rs o r b r i s t l es ; segmen t 5 of fro n t t a r s u s

Gon a topod i n ae

Fig.

1 068

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1 545

C he l a of Gon a top u s l u n a tus K l u g from F on t a i neb l e a u ( F rance )

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1 546

Fig.

1 069

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Ent.

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no.

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1 984

M a l e of Gon a topus l u n a tus K l u g from L e u t r a Va l l e y

( Germ a n y )

( F i g . 1 068 ) w i t h i n ner s i d e p ro x i m a l l y serra t e , w i t h 2 row s of 1 -3 + 1 2 1 7 l a m e l l a e ( s h o r t e s t row n o t e x t en d i n g b e y o n d 0 , 5 l en g t h of segme n t ) ; m a x i l l a r y p a l p i w i t h a g r o u p of 9- 1 4 l am e l l ae ; m a x i l l a ry p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 s e g m e n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE ( F i g . 1 069 ) : f u l l y w i n ged ; l e n g t h 1 , 7 5 - 1 , 87 m m ; b l ack , w i t h m a n d i b l es red d i s h ; t eg u l ae y e l l o w ; l e g s b a n ded ; a n te n n a e n o t d i s t a l l y t h i ck e n e d ; a n t e n n a I se g m e n t s i n fo l l o w i n g p ropo r t i on s : 4 , 5 : 4 : 6 : 5 , 5 : 5 , 5 : 5 , 5 : 5 , 5 : 5 , 5 : 5 , 5 : 7 , 5 ; a n t e n n a l segmen t 3 l ess t h a n 3 , 5 t i mes a s l o n g as b ro a d ( 2 , 6 ) ; head d u l l , h a i r y , g ra n u l a t e d , w i t h a sh i n y , smoo t h area between eyes and pos t e r i o r ace l I i i occ i p i t a l c a r i n a i n v i s i b l e ; t em p l es d i s t i n c t ; scu t u m d u l l , h a i r y , g r a n u l a ted ; n o t a u l i ces com p l e t e , p o s t er i or l y sep a r a t e d ( F i g . 1 070 B ) ; sc u t e l l u m d u l l , g ra n u l a ted ; m e t a n o t u m s h i n y , w i t ho u t scu l p t u r e ; p ro p o d e u m w e a k l y ret i c u l a t e ru gose , w i t ho u t kee l s ; dors a l s u rface w i t h a w e a k m ed i a n f u r ro w ; ra d i a l v e i n : f i g . 1 048 B ; dorsa l p rocess of gonoforceps n a rrow a n d s i n u a t e , d i s t a l l y r o u n ded p a r a l l e l to t he pen i s ( F i g . 1 07 1 ) or rou n de d ( F i g . 1 072 B ) ; b a s i v o l se l l a w i t h n o t c h ( F i g . 1 07 1 ) ; m a x i l l a ry p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . LOCUS TYP I C U S : Ber l i n ( Ge r m a n y ) TYP I CAL MAT E R I AL : l ec t o t y p e F ! of G . l u n a tu s ( d es i g n a t e d b y Moczar 1 96 5 ) i n H U ; l ec t o t y p e F ! a n d 3 p a r a l ec t o t y pes F F ! ( de s i g n a t e d b y Moczar 1 965 )

1 547

G o n a t o po d i n ae

F i g . 1 070 - Not a u l i ce s of m a l es of Gon a top u s sepsoides Wes t w ood from M a sserano ( I t a l y ) ( A ) a n d l u n a t u s K l u g from L e u t r a Va l l e y ( Ge r m a n y )

of G . m a rs h a ll i i n B U ; l ec t o t y p e F ! a n d 4 p a ra l ec t o t y pes F F ! ( d es i g n a ted by Moc z a r 1 965 ) of G. gracilipes i n BU ; h o l o t y p e F of G. filicornis in M T ; h o l o t y pe F of G. gracilis i n M T ; h o l o t y p e F ! of G. myrmecop h i l u s i n W S ; h o l o t y p e F ! o f G . b ifa scia tus i n G E ; h o l o t y p e F ! o f G . gracilicornis i n GE ; h o l o t y p e F ! a n d 1 p a r a t y p e F ! of G . raptoripes i n E B . H O ST S : i n F r a nce A d arrus ta uru s R i b a u t ( A b d u l -Nour 1 976 ) ; i n Germ a n y En a n tiocep h a l u s corn utus ( H . - S . ) , E u scelis incis u s , Erra s t u n u s s p . ( S a n der i n I i t t . ) , Deltocep h a l u s p a sc uorum ( H a u p t l eg . ) , Psammotettix stri a tus ( Ha u p t l e g . ) ; in F i n l a n d Arth a l deus p a s cuellus ( F a l l . ) ( L i n d b e r g 1 950 ) ; i n I t a I y Psammotettix a lien u s ( D a h l b . ) . D I S T R I BU T I O N : P a l aearc t i c reg i o n : K O R E A : N u n g-ra do I s l a n d ( P y o n g y a n , P ro v o So u t h P y o n g a n ) , B U ! M O N GO L I A ( Po n o m a r e n k o 1 979a ) ; F I N L A N D ( He l l e n 1 9 53 ) ; SWE D E N : S k a n e , L U ! Ho l a n d , L U ! Lj u n g b y ( Sk a n e ) , L U ! H U N GARY : SZomo t o r , BU ! Kecsem e t , BU ! K i s k u n h a l a s ( Bo g a r zo ) , BU ! S z i kra ( K ecse m e t ) ( Moczar 1 965 ) ; F a r k a s d , BU ! G y o n , BU ! C sepe l . , BU ! K o r t O l t e s , BU ! N y i re g y h a z a , B U ! Vacz ( T u dosdom b ) , BU ! B u d apest , BU ! Hortob a g y , B U ! D i as ( K i s b a l a ton ) , B U ! OrSz . M i k l os , BU ! Bon y , BU ! Mezo t u r , BU ! K . N . P . ( B u g ac ) , BU ! K u t a to h a z ( B u g ac ) , S U ! C s e v h a r a sz t , B U ! ROMAN I A ( N a g y 1 967a ; 1 972 ; Decu B u r g he l e 1 96 1 ; Teodo rescu 1 967 ) ; C Z E C H OSLOVAK I A : M i k u l o v ( P a w l ov s k e ) , SA ! G E R MANY : A n g e r ( Go r a u e r , B a y er n ) , E B ! H e i d e ( Rostoc k e r , Sch l e s w i g - Ho l s t e i n ) ,

EB!

F i now ( B randen burg ) ,

EB!

Bel l i nchen

( O der ) , HA ! Hohe n p r i e s s n i t z ( D u b e n a . M . ) , HA ! B e r l i n , H U ! R a t zdorf ( B r a n d e n b u r g ) , E S ! W u rz b u r g , G V ! L e u t r a V a l l e y ( Je n a ) , SA ! J u l c h e n doff ( S t e rn b er g , Meck l e n b u r g ) , E B ! AUST R I A : I g l s ( T i ro l ) , O T ! T u rken sch a n z e ( W i e n ) , P ! D E N M AR K , CO ! H O L LA N D : D e n H a a g , L E ! L U X E M B U R G , W S ! SW I T Z E R LA N D : L e n s ( V a l a i s ) , L O ! F R A N C E : B o l q u ere , P ! C h ambon-s u r-L . , P ! Corse ( Mocz a r 1 96 5 ) ; S t e . E n i m i e ( Lozere ) , B M ! Mon t-de-Marsan ( K i e ffer i n K i effer et M a rs h a l l 1 90 5 ) ; Fon t a i n e b l e a u ( Se i ne-e t - M a r n e ) , H E ! S I . A y g u l f ( Va r ) , P ! S I . P a u l -e t - V a l m a l l e ( Hera u l t ) , P ! P a r i s ( K i effer i n K i effer e t M a rs h a l l 1 905 ) ; E N G L A N D : S w a n a ge ( P u r b eck I s l a n d ) , B U ! B u r n h a m D u nes ( Norfo l k E a s t ) , HA ! Dea l ( K e n t E a s t ) , OX ! Si I wood P a rk ( Berk s . ) , B M ! S t u d l a n d ( Dorset ) , B M ! S p u r n H e a d ( York s ) , B M ! Quernera i s ( Jersey ) , B M ! S PA I N : A l g ec i ras ( He l l en 1 9 30 ) ; I TA LY : S a r re ( Aos t a ) , 8 M ! S t a z z a n o ( A l es s a n d r i a ) , GE ! Mon ca l v o ( A s t i ) , O L ! S a v i g l i a n o ( C u n eo ) , P L ! S . B e n e d e t t o Be l b o ( C u n eo ) , P L ! Sc i o l ze ( To r i n o ) , O L ! C h i a n occo ( T ori n o ) , O L ! Pece t to ( To r i n o ) , O L ! P u n t a Sab b i o n i ( Ve n ez i a ) ( u n d e r n a m e Gon a top u s myrmecop h i l u s , s e e G i o rd a n i So i k a 1 932 ) , M I ! V i se r b a ( F or i ) ) , BO ! G i o g o d i B a r d i n e t o ( S a v o n a ) , G E ! G i g l i o I s l a n d ( Grosse t o ) , G E ! V a s t o M a r i n a

1 548

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F i g . 1 07 1 - M a l e gen i t a l i a of Gon a top u s l u n a t u s K l u g from L e u t r a V a l l e y ( Germa n y ) ( dorsa I process o f gonoforceps p a ra I l e l t o pen i s ) ; N : notc h .

( C h i e t i ) , P L ! I sc h i a ( N a p o l i ) , HA ! P aes t u m ( S a l er no ) , E N ! T U R K EY : S m y rn a , WA ! I SRAE L : Rehovot , G V ! MADE I RA : Porto S a n to , HE !

2 1 . Gon a top u s tenerifei n . sp .

F E MAL E : a p t e ro u s ;

l en g t h 3 , 62-4 , 00 mm ; head , a n t e n n ae a n d p ro n o t u m

Gona topod i n a e

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1 549

1 072 - Dors a l process of g o noforceps of Gon a top u s sepsoides West wood from M a s�erano ( I t a l y ) ( A ) a n d l u n a tus K l u g from M i k u l o v ( C zechos l o v a k i a ) ( B ) ( dors a l process t r a n sverse )

tes t aceous-red d i sh ; occ as i o n a l l y a n t e n n a l se g me n t s 6- 1 0 b row n ; sc u t u m b row n - re d d i s h ; scu t e l l u m b l a c k ; m e t a t ho r a x + p ropodeum b l ac k , e xc e p t for apex of p ropod e u m l i g h t ; pet i o l e a n d a b domen b l ac k ; l e g s test aceous redd i s h , w i t h coxae a n d c l u b s of femora b row n ; a n t e n n a e l e ss t h a n 3 , 5 t i mes a s l o n g a s h e a d ; head exc a v a t e d , sh i n y , w i t ho u t scu l p t u re ; pron o t u m s h i n y , w i t ho u t scu l p t u re , h a i r l es s ; sc u t u m s h i n y , w i t h some l o n g i t u d i n a l k ee l s ; m e t a no t u m s moo t h , n o t s t r i a te t ra n s v e r se l y ; m e t a t h o r a x + p ropodeum smoo t h , s h i n y , w i t ho u t sc u l p t ure ; p l eu r a t r a n s v erse l y n o t s t r i a t e ; pos ter i or s urface of p ropode u m t r a n s verse l y s t r i a t e ; meso-me t a p l e u r a l su t u re o b so l e te ; fore t a r s a l se g m e n t s i n fo l l o w i n g p roport i o n s : 1 4 : 3 : 4 : 1 4 : 2 2 ; e n l a r g e d c l a w ( F i g . 1 07 3 ) w i t ho u t s u b a p i c a l too t h a n d l a me l l ae ; w i t h s ma l l too t h a t t h e e n d of a l on g i t u d i n a l fu rrow a n d 3 peg- l i ke h a i rs ; se g m e n t 5 of fro n t t a rs u s ( F i g . 1 07 3 ) w i t h i n ner s i d e p rox i m a l l y serr a t e a n d w i t h 2 rows of 1 ( p rox i m a l ) + 22 l am e l l ae ; a p e x w i t h a g ro u p of 6 l a me l l ae ; m a x i l l a r y p a l p i w i t h 5 seg men t s ; l a b i a l p a l p i w i t h 3 seg men t s ; t i b i a l spurs 1 , 0 , 1 . MALE : u n k no w n L O C U S T Y P I C U S : P u e r t o d e l a C r u z ( C a n a ry I s l a n d s ) paratype F ! TYP I C A L MAT E R I A L : h o l o t y p e F ! i n H E ; 1 p a r a t y p e F ! i n P ; i n OL. HOSTS : u n k no w n D I S T R I B U T I ON : P a l aearc t i c reg i on : CANARY I SLAN D S : P u e r t o d e l a C r u z

1 550

Mem .

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1 073 - C h e l a of Gon a top u5 tenerlfei n . sp .

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sp .

F E MA L E : a p tero us ; l e n g t h 3 , 25 m m ; t e s t aceous-red d i s h , w i t h pet i o l e b l a ck ; a b domen b l ac k w i t h red d i s h n u ances ; head e x c a v a ted , s h i n y , w i t h o u t sc u l p t u r e ; a n te n n a e ( F i g . 1 067 A ) l ess t h a n 3 , 50 t i mes a s l o n g as h e a d ; p ro n o t u m crossed by a t r a n s versa l i m p res s i o n , sh i n i n g , w i t ho u t scu l p t u re , w i t h some l o n g i t u d i n a l k ee l s ; m e t a n o t u m smoo t h , n o t t r a n s verse l y s t r i a te ; m e t a t h o r a x + p ropodeum s h i n y , w i t h o u t scu l p t u re ; p l e u ra w e a k l y t r a n s verse l y s t r i a t e ; pos t e r i o r s u rface o b so l e t e ; fore t a r s a l seg men t s i n fo l l ow i n g propor t i o n s : 1 2 : 3 : 4 : 1 2 : 2 1 ; e n l a r ged c l a w ( F i g . 1 074 ) w i t ho u t s u b a p i c a l too t h a n d l a me l l ae , w i t h a s m a l l too t h a t t h e e n d o f a l o n g i t u d i n a l f u r row a n d 4 peg - I i k e h a i r s ; seg men t 5 of fron t t a rs u s ( F i g . 1 074 ) w i t h i n ner s i d e pro x i m a l l y serra t e a n d w i t h 2 row s of 2 + 1 7 l a m e l l ae ( s h o r t e s t row n o t e x t e n d i n g b e y o n d 0 , 5 l en g t h of segmen t ) ; a p ex w i t h a g ro u p of 1 0 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 5 segme n t s ; l a b i a l p a l p i w i t h 3 s e g m e n t s ; l a b i a l p a l p i w i t h 3 seg men t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n l O C U S T Y P I C U S : M o n t e Ag u i rre ( C a n ary I s l a n d s ) TYP I CAL MAT E R I A L : h o l o t y p e F ! i n H E . HOST S : u n k n o w n D I STR I B U T I O N : P a l a e a rc t i c reg i o n : CANARY I SL A N D S :

Mon t e A g u i rre

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1 074 - C h e l a o f Gon a top u s c a n a rien sis n .

1 55 1

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23 . Gona top u s wit tmeri n . sp .

F E MAL E : a p t e rou s ; l en g t h 2 , 84 m m ; t e s t aceo u s-red d i s h , w i t h p e t i o l e b l ack , a bdomen p a r t l y b ro w n , a n t e n n a I seg m e n t s 4- 1 0 b row n ; a n te n n a e d i s t a l l y t h i ck e n e d ; a n te n n a I se g m e n t s i n fo l l o w i n g p ropo r t i on s : 1 0 : 6 : 1 0 : 6 : 6 : 7 : 6 : 6 : 5 : 8 ; h e a d a l m o s t f l a t , d u l l , g r a n u l a te d ; fron s w i t ho u t m ed i an k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; POL � 2 ; OL � 2 , 5 ; O O L � 9 ; p r o n o t u m crossed by a s t r o n g t r a n s v e r s a I i m p ress i o n , d u I I , g r a n u I a t ed ; m e t a n o t u m d u I I , t r a nsverse l y s t r i a t e , not h o l l o w b e h i n d t h e sc u t e l l u m ; m e t a t h o r a x + p ro p o d e u m d u l l , g r a n u l a ted , w i t h pos t e r i o r s u rface t r a n s v erse l y s t r i a t e ;

1 55 2

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1 07 5 - C h e l a of Gon a top u s wittmeri n . sp .

( type)

meso-me t a p l e u r a l s u t u re o b so l e t e ; d i sc o f m e t a t ho r a x + p ro p o d e u m w i t h a very w e a k t r ack of med i an fu rrow ; fore t a r s e I segmen t s i n fol l o w i n g p ropor t i on s : 1 4 : 4 : 5 : 1 8 : 25 ; e n l a rged c l a w ( F i g . 1 075 ) w i t h o u t s u b a p i ca l too t h , w i t h a s m a l l too t h a t the e n d of a l on g i t u d i n a l f u r row a n d w i t h a row of 8 p e g- l i k e l a m e l l a e ; seg men t 5 of fron t t a rs u s ( F i g . 1 075 ) w i t h row of 1 9 l a m e l l ae ; apex w i t h a g ro u p of n e a r l y 9 l a me l l ae ; m a x i l l a ry

G o n a topod i n ae

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1 076

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C h e l a of Gon a topus n igrosign a tu s J . S a h l berg

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p a l p i w i t h 4 s e g m e n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l MALE : u n k n o w n LOCUS TYP I C U S : S a k k a ra ( E g y p t ) T Y P I CA L MA T E R I AL : h o l o t y p e F ! i n WA . HOST S : u n k n o w n D I ST R I B U T I O N : P a l aearc t i c reg i o n : E GYPT : S a k k a r a , WA ! NOT E S : t he s p ec i e s is n a m e d i n honor of M r . W . W i t t mer .

s p u rs 1 , 0 , 1 .

24 . Gon a top u s nigrosign a t u s J . S a h I berg Gon a top u s n igrosign a tu s J . S a h l b e r g 1 9 1 0 :

15.

F E MA L E : a p t e rou s ; l e n g t h 3 , 1 2 m m ; y e l l o w - t e s t aceo u s , w i t h p et i o l e b l ac k ; head e x c a v a t e d , d u l l , g r a n u l a t e d on occ i p u t ; v e r t e x w i t h o u t scu l p t u re , w i t h fron t p a r t g ra n u l a ted ; p rono t u m crossed b y a t r a n s v e rs a l i m p ress i o n , s h i n y , w i t ho u t scu l p t u re or w e a k l y g ra n u l a t ed ; scu t u m s h i n y , g r a n u l a ted ; m e t a n o t u m w e a k l y tra n s v e rse l y s t r i a te ; m e t a t horax + p ropodeum w i t h s t ron g med i a n f u r row , w i t hou t scu l p t u r e ; p l e u r a a n d pos t e r i or s u rface of p ropod e u m t r a n s v erse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re obso l e t e ; segmen t 1 of fron t t a r s u s s l i g h t l y l o n ger t h a n segmen t 4 ( 1 2 : 1 1 ) ; e n l arged c l a w ( F i g .

1 076 ) w i t ho u t s U b a p i c a l too t h a n d l a me l l ae , w i t h a sma l l too t h a t t h e e n d of a l o n g i t u d i n a l fu rrow a n d 3 p e g- l i k e h a i rs ; segmen t 5 of fro n t t a r s u s ( F i g . 1 076 ) w i t h i n ne r s i de p ro x i m a l l y not serra t e , w i t h 2 r o w s of 4 + 1 1 l a me l l a e l o c a t ed on d i s t a l ha l f of seg me n t 5 on a d i s t i n c t p rom i n ence ; apex w i t h a g ro u p of 9 l a me l l a e ( some l a me l l a e a r e m u c h l o n g er t h a n o t h e r l a me l l ae ) ; m a x i I l a r y p a l p i w i t h 4 segmen t s ; l ab i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : Dsc h i l a r i k ( T u rk i s t a n ) TYP I CA L MAT E R I AL : h o l o t y p e F ! i n H E . HOST S : u n k no w n D I ST R I BU T I O N : P a l aearc t i c reg i o n : T U R K I STAN : Dsch i l a r i k , H E !

25 . Gon a topus plan iceps K i effer

Gon a topus p l a n iceps K i effer 1 904 a : 355 . Pla tygon a topus pla n iceps ( K i effer ) : K i effer i n K i effer e t M a r s h a l l 1 90 6 : 500 . Pla tygon a top u s b uysson i Ber l a n d 1 928 : 1 47 ( n . s y n . ) ; loc u s typ icus St-Rem y - I a - V a r e n n e ( F ra n ce ) . Gon a top u s ruficollis H e l l en 1 9 30 : 3 ( n . s y n . ) ; locus typ i c u s Mogador ( Morocco ) . n ec Ph tygon a top u s p l a n iceps ( K i e ffer ) : R i c h a r ds 1 939 :

G o n a topod i n a e

Fig.

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C h e l a of Gona topus p l a n iceps K i effer from Mogador ( Morocco )

1 556

Mem . Amer .

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I ns ! . ,

no.

37 ,

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219. Pl a tygo n a top u s plan iceps ( K i effer ) : O l m i a n d C u rrado 1 974 : 1 75 .

F E MA L E : a p t e ro u s ; l en g t h 3 , 0-3 , 5 m m ; h e a d b l ac k o r b r o w n - b l ac k ; occa s i on a l l y b row n-redd i s h , w i t h fro n t p a r t of v e r t e x , c l y p eu s , m a l a r s p ace a n d m a n d i b l e s y e l l o w ; a n ten n ae b row n , w i t h seg m e n t s 1 -3 , 1 -4 o r

1 -6 y e l l o w ; p rono t u m redd i s h or b ro w n ; occa s i on a l l y b ro w n w i t h red d i s h n u a nces ; sc u t u m y e l l ow - re d d i s h or red d i sh ; occa s i on a l l y b l ac k w i t h b ro w n n u a nces ; sc u t e l l u m a n d m e t a t horax + p ro p o d e u m b ro w n-red d i s h or red d i s h ; occa s i o n a l l y b l ac k w i t h b ro w n n u a n ces ; p e t i o l e a n d a b domen b l a ck , i n some s p ec i m e n s w i t h b r o w n n u a n ces ; l e g s y e l l ow-red d i s h , w i t h coxae a n d c l u b s o f femora b ro w n i s h ; h e a d w e a k l y e x c a v a ted , d u l l , g r a n u l a t e d , i n some s p ec i m e n s w i t h short t r a n sv e rs a l k ee l s a l o n g orb i t s ; occ i p i t a l c a r i n a n o t com p l e te , v i s i b l e on l y b e h i n d t h e oce l l i ; p ron o t u m crossed b y a t r a n s v e rs a l i m p re ss i o n , s h i n y ; d i sc w i t ho u t scu l p t u re ; co l l ar ( mos t l y o n s i d es ) g ra n u l a ted ; sc u t u m s h i n y , w i t h s o m e w e a k l o n g i t u d i n a l k e e l s ; s c u te l l u m v e r y i n c l i n ed ; m e t a n o t u m s h o r t , h o l l o w b e h i n d t h e scu te l l u m ; m e t a t h o r a x + p ropodeum w i t ho u t scu l p t u re , w i t h s t ro n g med i a n fu rrow ; p l e u r a a n d pos t e r i o r su rface of p ropodeum t r a n sverse l y s t r i a t e ; meso m e t a p l e u r a l s u t u re d i s t i n c t a n d comp l e t e ; se g m e n t 1 of f ro n t t a r s u s s l i g h t l y s h o r t e r t h a n se g m e n t 4 ( 1 0 : 1 1 ) ; en l a rged c l a w ( F i g . 1 077 ) w i t ho u t s u b a p i c a l too t h a n d l a m e l l ae , w i t h a s ma l l t oo t h a t t h e e n d of a l o n g i t u d i n a l f u r ro w a n d 2-8 p e g - I i ke h a i r s ; seg m e n t 5 of f ro n t t a r s u s ( F i g . 1 077 ) w i t h i n n e r s i d e p ro x i m a l l y n o t serra t e , w i t h 2 r o w s of 1 3-25 l a me l l ae b eg i n n i n g o n p ro x i m a l t h i rd of segmen t 5 ( s hortes t row not e x t en d i n g b e y o n d 0 , 5 l e n g t h of segmen t ) ; apex w i t h a g r o u p of 8- 1 2 l a me l l ae ; m a x i I l a ry p a l p i w i t h 4-5 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k n o w n L O C U S TYP I C U S : G i g l i o I s l a n d ( I t a l y ) TYP I C A L M A T E R I AL : ho l o t y p e F ! of G . p l a n iceps i n G E ; h o l o t y p e F ! of Pl . b uyssoni i n P ; h o l o t y p e F ! of G. ruficollis i n H E . HOST S : u n k n o w n D I ST R I BU T I O N : P a l a e a rc t i c reg i o n : H U N GARY : V a c z T u dosdomb , BU ! C se p e l , B U ! R OMAN I A ( N a g y 1 972 ) ; G E R M ANY : H e i deb r i n k ( W o l l i n ) , HA ! F R A N C E : Mon t pe l l i e r ( He r a u l t ) , P ! S t-Rem y - I a-Varen n e ( M a i ne-et-Loi r e ) , P ! I T ALY : G i g l i o I s l a n d ( G rosseto ) , G E ! C i rceo ( L a t i n a ) , BO ! CANARY I SL A N D S : G a l d a r ( G ran C a n a r i a ) , H E ! M e d a n o ( T e n er i fe ) , M D ! M O R OC C O : M o g a do r , H E ! E GYPT : A b o u l a i r , WA ! W a d i U m a s s a d , WA !

Wadi

EI

T i h , WA !

26 . Gon a topus campes tris Ponomarenko

Gon a topus campestris P o n o m a ren k o 1 965 : 628 .

OL !

Gon a topod i n ae

Fig.

1 5 57

1 078 - C h e l a of Gon a top u s camp estris Ponomarenko ( t y p e )

Gon a topu s c a mpestris Ponomarenko 1 978 : 2 6 . Gon a top u s c a mpestris Ponomaren k o : Moczar 1 983b : 207 .

F E MAL E : a p t e rou s ; l e n g t h 2 , 7 m m ; h e a d m i s s i n g i n h o l o t y p e ( o n l y seen s p ec i me n ) : accord i n g to t h e o r i g i n a l desc r i p t i on head y e l l ow , w i t h v e r t e x b l ac k a n d a n t e n n a e b l ac k w i t h s e g m e n t 1 y e l l ow ; t h o r a x and p ropod e u m b l ack , w i t h red d i s h n u a nces on p r o t ho ra x ; occas i on a l l y b ro w n -re d d i s h , w i t h sc u t u m a l most f u l l y y e l l ow ; a b domen b ro w n - t e s t aceo u s ; l eg s t e s t aceou s ; p ro n o t u m w e a k l y g r a n u l a t ed , s h i n y , crossed b y a s t ro n g t r a n s v ersa l i m press i on ; sc u t u m d u l l , g ra n u l a ted ; m e t a n o t u m ru gose , h o l l ow b e h i n d t he s c u te l l u m ; meso-me t a p l e u r a l s u t u r e d i s t i n c t a n d comp l e t e ; m e t a t horax + propode u m d u l l , g r a n u l a t e d ; p l e u r a n o t t r a n sverse l y s t r i a t e ; pos t e r i or s u rface o f p ropod e u m c e n t r a l l y n o t s t r i a t e , o n s i de s t r a n s v erse l y s t r i a te ; d i sc o f me t a t horax + p ropode u m w i t h a more o r l e ss s t ro n g med i a n f u rrow ; i n some spec i me n s on l y w i t h a t rack o f med i a n f u rrow ; fore tarsa l s e g me n t s i n fo l l ow i n g p ropor t i o n s : 9 : 2 : 3 : 1 0 : 1 6 ; en l arged c l a w ( F i g . 1 07 8 ) w i t ho u t su b a p i c a l t oo t h , w i t h a s m a l l too t h a t t h e e n d of a l on g i t u d i n a l f u rrow a n d w i t h one b r i s t l e ; s e g m e n t 5 of fron t t a rs u s ( F i g . 1 078 ) w i t ho u t p rox i m a l serra t e p a r t , w i t h 1 row o f 1 0 l a me l l ae or w i t h 2 rows of 2 ( p ro x i m a l ) + 1 6 l a m e l l a e ; a p e x w i t h a g ro u p of 7 l a me l l a e ( i n t h i s g ro u p 3 l am e l l ae are m u c h l o n g e r t h a n o t h e r l a me l l ae ) ; m a x i l I a r y p a l p i w i t h 5 s e g m e n t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k no w n

1 5 58

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B

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Heads of f e m a l es of Gona top u s s u b tilis n . s p . from C a n a r y dorsa l l y v i e wed ) a n d cla vicornis ( F e n ton ) ( B : dorsa l l y v i ewed ; C : fron t a l l y v i ew e d ) .

-

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paratype F !

i n MO.

HOST S : i n U . S . S . R . Ps ammotettix stri a t u s L . ( Ponomarenko 1 965 , 1 978 ) . D I STR I B U T I ON : P a l aearc t i c reg i o n : MONGOL I A : S b a n k o f Orog n u u r l ak e ( U v s a i ma k ) , BU ! U . S . S . R . : K i ne l ( K u y b y s hev P ro v i n ce ) , M O !

27 . Gon a top u s s u b tilis n .

sp .

F E MAL E : a p terou s ; l e n g t h 3 , 1 2-3 , 43 m m ; head ferr u g i n o u s or b row n , w i t h m a n d i b l es a n d c l y pe u s y e l l o w ; a n ten n ae fu l l y test aceous or b ro w n , w i t h seg m e n t s 1 -2 y e l l ow ; t h o r a x a n d p ropodeum ferr u g i n o u s ; i n some spec i m e n s p ro t h o r a x a n d scu t u m b ro w n - red d i s h a n d m e t a t h o r a x + p ropod e u m b l ac k ; p et i o l e a n d a b domen b l a c k - b row n ; l e g s b row n , w i t h fore t a rs i , e x t e r n a l f ace of fore t i b i ae a n d m i d p a r t o f h i n d t i b i ae y e l l ow ; i n some spec i m e n s t i b i ae y e l l o w , w i t h apexes b row n ; h e a d f l a t , s h i n y , h a i r l e s s , g r a n u l a t e d ; t e mp l es very l o n g , appro x i m a te l y 0 , 5 as l o n g as e y e s ; p rono t u m h a i r l es s , crossed b y a t r a n s v ersa l i m press i o n , s h i n y , w e a k l y a l u t aceou s ; scu t u m s h i n y , h a i r l es s , w i t ho u t scu l p t u re , w i t h some l on g i t u d i n a l k ee l s ; m e t a n o t u m t r a n s v e r se l y s t r i a t e ; m e t a t horax + p ropodeum s h i n y , h a i r l e ss , w i t h o u t scu l p t u re ; p l e u r a a n d pos t e r i o r s u r face of propodeum t ra n s v e rse l y s t r i a te ; meso-met a p l eu r a l s u t u re obso l e t e ; se g m e n t 1 o f fron t t a r s u s l on g e r t h a n seg men t 4 ( 1 2 : 7 ) ; e n l a rged c l a w ( F i g . 1 080 ) w i t ho u t su b a p i c a l t o o t h a n d l am e l l ae , w i t h a s m a l l too t h a t t h e e n d o f a l on g i t u d i n a l f u rrow a n d 3-4

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p e g - l i k e h a i rs ; seg m e n t 5 of fro n t t a r s u s ( F i g . 1 080 ) w i t h i n n e r s i de p r o x i m a l l y n o t serra t e , w i t h 2 ro w s of 8- 1 4 v e r y l o n g l a m e l l ae ; apex n o t b roa d e n e d , w i t h a g ro u p of 6- 1 3 l a m e l l ae ; m a x i I l a r y p a l p i w i t h 4 - 5 s e g m e n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n known L O C U S TYP I C U S : S a n Seb a s t i a n ( C a n a r y I s l a n d s ) TYP I CA L MAT E R I A L : ho l o t y p e F ! i n H E ; 3 p a r a t y pes F F ! i n WA ; 1 p a r a t y p e F ! i n OL . HOST S : u n k n o w n D I S T R I B UT I O N : P a l aearc t i c reg i o n : CANARY I SLAN D S : S a n S e b a s t i a n ( Gomera ) , OL ! H E ! E GYP T : W a d i Hoff , WA ! W a d i E I T i h , WA !

28

Gon a top u s spectrum ( Sn e l l e n v a n Vo l l en hoven )

Dryin us spectrum S n e l l e n v a n Vo l l e n h o v e n 1 874 : 1 59 . Gon a topus spectrum ( S ne l l en v a n Vo l l e n h o v e n ) : K i effer i n K i effer e t M a r s h a l l 1 90 5 : 9 9 . Gon a top u s spectrum ( S ne l l en v a n Vo l l e n h o v e n ) v a r . a l b i d u s K i effer in K i effer e t Marsha l l 1 906 : 504 ( sy n . p rop osed b y O l m i a n d C u rrado 1 975 ) ; locus typ ic u s S v e t o B r d o ( H u n g ary ) . Gon a top u s fla v us Hed i c k e 1 92 6 : 243 ( n . s y n . ) ; locus typ i c u s B e l l i n c h e n ( Ge r m a n y ) . Gon a top u s drom edarius ( A . C os t a ) : Bern a r d 1 939 : 1 1 1 . Neogon a top u s spectrum ( Sne l l en v a n Vo l l e n h o v e n ) : O l m i a n d C u rr a d o 1 975 : 55 . Neogon a topus a lb idus ( K i e ffer ) : Ponom a r e n k o 1 978 : 2 6 .

F E MA L E : a p t e ro u s ; l en g t h 2 , 80-3 , 1 2 m m ; y e l l o w - t e s t aceou s , w i t h p e t i o l e b l ac k a n d ab domen b ro w n i s h ; rare l y m e t a t ho r a x + p ro p o d e u m d a r k ; SOme s p ec i m e n s w i t h two b ro w n spots o n s i des of p ropode u m ; head e xc a v a ted , w i t ho u t sc u l p t u re , s h i n y , h a i r l es s ; p ron o t u m crossed b y a t r a n s v e rs a l i m p ress i o n , s h i n y , smoo t h ; m e t a n o t u m s h i n y , t r a n s v e r se l y s t r i a t e ; m e t a t ho r a x + p ro p o d e u m s h i n y , w i t ho u t sc u l p t u re ; p l e u r a a n d p o s t e r i o r s u rface o f p ro p o d e u m t r a n s ve rse I y s t r i a t e ; meso-me t a p l e u r a I s u t u r e o b so l e t e ; s e g m e n t 1 of fron t t a r s u s a s l o n g a s s e g m e n t 4 ; en l a r g e d c l a w ( F i g . 1 08 1 ) w i t h o u t s u b a p i c a l too t h a n d l a me l l ae , w i t h a sma l l too t h a t t h e e n d of a l o n g i t u d i n a l f u r row a n d 4-5 p e g - l i k e h a i rs o r b r i s t l es ; se g m e n t 5 of f r o n t t a r s u s ( F i g . 1 08 1 ) w i t h i n n e r s i de p ro x i m a l l y n o t serra t e , w i t h 2 rows of 1 5- 1 7 l a m e l l a e l oc a t e d on d i s t a l h a l f of seg m e n t on a d i s t i n c t p rom i ne n c e ; apex w i t h a group of 7- 1 0 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 4-5 segmen t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U 5 TYP I C U S : Vo g e l en z a n g ( Ho l l a n d ) TYP I C A L MA T E R I AL :

ho l o t y pe F ! o f D .

spectrum i n L E ;

ho l o t y p e F !

of G.

Gon a topod i n ae

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C he l a of Gon a topus spectrum ( S ne l l e n v a n V o l l en ho v e n ) from Be l l i n c he n ( Ge rm a n y )

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1 082 - Head of fema l e of Gon a top u s bernardi ( P i c a rd ) ( t y pe )

a lb idus i n B U ; h o l o t y p e F ! of G. fla vu s i n H U . HOST S : i n F r a n ce A darrus ta urus R i b . , Araldus prop in q u u s K b m . , P a l u da elonga t a ( W g n . ) ( = Rhop a lopyx elonga t u s ) . D I ST R I B U T I O N : P a l aearc t i c reg i on : U . S . S . R . ( Ponomarenko 1 978 , u n der n ame Neogon a topus a lbidus K . ) ; HUNGARY : N a g y s ze n a s , B U ! Tompa , B U ! Sveto B r do , B U ! V a c z . T u dosdomb , B U ! K e l eb i a , B U ! Berh i d a , B U ! B U LGAR I A : S a n d a n s k i , B M ! G E R MANY : B e l l i nchen ( Od er ) , HU ! H A ! AUST R I A : Vos l o w ( u n der name Gon a top u s dromeda rius Cos t a : Bernard 1 939 ) , P ! HOLLAN D : Voge l en z a n g , L E ! SW I T Z E R LAND : Peney , G V ! F RANCE : L a F i g arede ( He r a u l t ) , P! S t P a u l -et-V a l m a l l e ( Herau l t ) , P ! F rej u s ( V a r ) ( u n der n a m e Gon a topus dromedarius Cos t a : B e r n a r d 1 939 ) , P ! G R E E C E : S a k u l evo ( M akedon i a ) . P ! M t . H o l omo n d a s ( H a l k i d i k i ) , OL !

29 . Gon a top u s bernardi ( P i c a r d ) n . com b .

Pla tygo n a top u s polychromus ( M a r s h a l l ) v a r . bernardi P i c a r d 1 932 : 29 . Pla tygon a top u s polychromus ( M a r s h a l l ) v a r . bernardi P i c a r d : Bern a rd 1 932 : 7 1 . Pla tygon a top u s polychrom us ( M a r s h a l l ) v a r . bern ardi P i c a rd : Bernard 1 935b : 53.

F E MALE : a p te ro u s ; l e n g t h 3 , 00-3 , 4 3 mm ; brown-b l ac k , w i t h m a n d i b l es , c l y p e u s a n d fro n t p a r t of vertex y e l l ow ; a spec i me n from H u n g a ry w i t h body ( most l y hea d ) b row n ; a n ten n ae b row n , w i t h segme n ts 1 -2 p a r t l y y e l l ow ; l e g s b ro w n , w i t h w h i te spots on prox i m a l exter n a l p a r t of fore t i b i ae , on extern a l s i d e of m i d a n d h i n d troc h a n ters , on e x tern a l face of m i d a n d h i n d t i b i a e ; hea d ( F i g . 1 08 2 ) d u l l , weak l y exc a v a te d , w i t h n u mero u s l o n g i t u d i n a l kee l s on fron s ; occ i p i t a l car i n a not com p l e te , v i s i b l e on l y b e h i n d oce l l i ; pron o t u m crossed b y a t r a n s v e r s a l i mp ress i o n , smoo t h , h a i r l e s s , w i t h o u t scu l p t u re , w i t h Some l on g i t u d i n a l kee l s on s i des ; sc u t u m

Gon a topod i n ae

Fig.

1 083

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1 563

1 564

Mem . Amer . E n t .

I n st . ,

no. 37,

1 984

s h i n y , w i t h some l o n g i t u d i n a l k ee l s ; s c u t e l l u m smoo t h , s h i n y , v e r y i n c l i n ed ; met a no t u m h o l l o w b e h i n d t h e scu t e l l u m , t r a n s verse l y s t r i a t e ; m e t a t horax + p ropodeum w i t h s t ro n g m ed i an f u rrow ; pos t e r i or s u rface of p ropod e u m t r a n s v erse l y s t r i a te ; meso-me t a p l eu r a l s u t u r e d i s t i nc t a n d comp l e t e ; seg m e n t 1 o f fron t t a r s u s l o n g e r t h a n s e g me n t 4 ( 1 4 : 1 1 ) ; e n l arged c l a w ( F i g . 1 083 ) w i t ho u t su b a p i c a l too t h a n d l am e l l ae , w i t h a s m a l l too t h a t t h e e n d of a l on g i t u d i n a l fu rrow a n d 2 b r i s t l es ; segmen t 5 of fro n t t a rs u s ( F i g . 1 083 ) w i t h i n ne r s i de pro x i m a l l y serra t e , w i t h 2 row s o f 2-3 + 1 6 l a me l l ae b eg i n n i n g o n p ro x i m a l t h i rd of segmen t 5 ; a pex w i t h a group of 9 l a me l l a e ( on e l ame l l a i s m u c h l on ge r t h a n o t he r l a m e l l ae ) ; m a x i l l ary p a l p i w i t h 6 segmen ts ; spurs 1 , 0 , 1 .

l a b i a l p a l p i w i t h 3 segmen t s ;

tibial

MAL E : u n k no w n L O C U S T Y P I C U S : Sa i n t -A y g u l f ( F r a n ce ) TYP I CA L M A T E R I A L : l ec t o t y p e F ! ( hereb y d es i g n a te d ) i n P . HOST S : i n F r a nce A thys a n u s heydeni o n Tamarix ( P i c ard 1 332 ) ( p rob a b l y Opsius sta etoga l u s F i eb . ) D I ST R I B U T I O N : P a l a e a rc t i c reg i o n : HUNGAR Y : V a c z . T u dosdomb ( i n n e s t of a n t ) , B U ! F R A NC E : Sa i n t-Ay g u l f ( V a r ) , P i Vi I l e p e y ( Be r n a r d 1 932 ; 1 935b ) ; I TALY : M u s e i ( C a g l i a r i ) , Ol !

30 . Gon a top u s form iearius L j u n g h

Gon a top u s form iearius lj u n g h 1 8 1 0 : 1 62 . Gon a top u s formiearius L j u n g h : K i effer i n K i effer e t M a r s h a l l 1 905 : 93 . Gon a top u s form iearius Lj u n g h : K i effer 1 9 1 4 b : 1 09 . n e e Gon a top u s form iearius L j u n g h : L i c h te n s t e i n a n d P i c a r d 1 920 : 54 ( se e Gon a top u s sepsoides W e s t wood ) . n e e Gon a top u s form iearius L j u n g h : B er l a n d 1 928 : 1 5 1 ( see Gona topus sepsoides Wes t w ood ) . Gona topus formiein u s K ro g e r u s 1 932 : 1 24 ( n omen n u dum ) . Gon a top u s form iearius lj u n g h : R i c h a r d s 1 93 9 : 2 1 3 . n ee Gon a top u s form iearius L j u n g h : Ber l a n d 1 940 : 1 2 1 . Gon a topus formiearius Lj u n g h : H e l l en 1 953 : 95 . Gona topus formiearius Lj u n g h : 0 I m i a n d C u rr a d o 1 97 5 : 49 . Gon a top u s formie arius Lj u n g h : P o n o m a r e n k o 1 978 : 2 6 .

F E MAL E : a p t e rou s ; l e n g t h 2 , 24-3 , 40 m m ; h e a d b l ac k , w i t h occ i p u t , fro n t p a r t o f v e r te x , c l y p e u s a n d m a n d i b l e s y e l l ow - red d i s h ; a n te n n a e b l a ck w i t h segm e n t s 1 -2 y e l l o w ; pro n o t u m fu l l y b l ac k or b row n , w i t h d i sc a n d a n te r i o r m a r g i n o f co l l a r redd i s h ; scu t u m , s c u te l l u m a n d met a t horax + p ropodeum b l a c k , except for apex of p ropode u m red d i s h ; p e t i o l e a n d a b domen b l ac k w i t h terg i t e 1 y e l l ow - red d i s h ; l e g s y e l l o w w i t h coxae a n d

Gon atopod i n ae

Fig.

1 56 5

1 084 - C he l a of Gon a top u s form icarius L j u n g h from Portacomaro ( I t a l y )

c l u b s of femora b row n i s h ; a n t e n n a I segmen ts i n fol l o w i n g p ropor t i on s : 6 : 5 : 1 0 : 5 : 4 , 5 : 5 : 4 , 5 : 4 , 5 : 4 : 6 ; head more or l ess e xca v a ted , d u l l , g r a n u I ated ; POL = 2 ; OL = 2 ; OOL = 7 ; p ro n o t u m crossed b y a strong t r a n sversa l i m p ress i o n , s h i n y , g ra n u l a ted ; sc u t u m d u l l , w i t h some l o n g i t u d i n a l k ee l s ; met a n o t u m t r a n sverse l y not str i a te ; m e t a t h o r a x + p ropodeum d u l l , g ra n u l a t e d ; rare l y d i sc w i t h a strong med i a n f u r row ; p l e u r a a n d poster i or s u rface of p ropodeum t r a n sverse l y not s t r i a te or weak l y s t r i a t ? ; meso-me t a p l e u r a l s u t u re d i s t i n c t and com p l e te ; segme n t of f ro n t t a r s u s s l i g h t l y l o n ger t h a n segment 4 ( 1 2 : 1 1 ) ; e n l arged c l a w ( F i g . 1 084 ) w i t h o u t s u b a p i c a l too t h , w i t h a sm a l l too t h a t the e n d of a l o n g i t u d i n a l f u r ro w a n d w i t h some b r i s t l es or peg- l i ke h a i rs ; segment 5 of f ro n t t a r s u s ( F i g . 1 084 ) w i t h i n ner s i de p ro x i m a l l y not serra te , w i t h one row of 1 0-20 l a me l l ae be g i n n i n g on p ro x i m a l t h i rd of segment 5 or w i t h t w o row s ( on e row not exten d i n g b e y o n d 0 , 5 l en g t h of segme n t ) ; r a re l y w i t h t w o row s o f l a me l l ae e x t e n d i n g beyond 0 , 5 l e n g t h o f segme n t ; apex of segmen t 5 w i t h a g ro u p of 7-1 4 l a me l l ae ; m a x i l l a ry pa l p i w i t h 5 segmen ts ; l a b i a l p a l p i w i t h 2 segme n ts ; t i b i a l spurs 1 , 0 , 1 . MAL E : u n k no w n L O C U S TYP I C U S : Sk a rs i o ( Sweden ) TYP I CA L MATER I AL : h o l o t y pe F ! i n LU . HOST S : i n I ta I y Diplocolen u s a b domin a lis ( F . ) , Diplocolen u s n igrifrons ( K b m . ) , Ps ammotettix a lien u s ( D a h l b . ) ( O l m i a n d C u rrado 1 974 ) ; i n F r a nce Psammotettix confinis ( D a h l b . ) ( M a n ev a l 1 932 ) . D I ST R I BU T I ON : P a l aearc t i c reg i o n : U . S . S . R . ( Ponomarenko 1 978 ) ; F I N LAN D : Sei s k a r i ( R . J . ) , H E ! H a t t u l a ( R . K . ) , H E ! SW EDEN : S k a rs i o ( Sma l a n d ) , L U ! Ofrabo ( Lj u n g h 1 8 1 0 ) ; HUNGARY : B u d apes t , B U ! BO !

1 566

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1 085 - C he l a of Gon a top u s mongol icus Ponomarenko ( t y p e )

N o g r a d s z ak a l , B U ! P a ra d , BU ! C s i k Szep v i z , BU ! T a h i t o t fo l u , B U ! Vacz ( T u do s d om b ) , B U ! Vacz-Szod , B U ! T a s n a d , B U ! S i mo n torn y a , B U ! D i osd , B U ! C Z E C H O SLOVAK I A : P i e s t i n g , V I ! G E R MA N Y : Be l l i n c h e n ( O d er ) , HA ! K y f f h a u ser ( T h u r i n gen ) , E B ! T h Li r i n ge n , P ! A U ST R I A , C O ! H O l lA N D : M i t l e r O dergeg . , l E ! B E L G I U M ( P i c a r d 1 932 ) ; l U X E M B U R G ( K i e ffer i n K i effer 1 905 ) ; F RA N C E : C he n e re i l l es ( M a n e v a l 1 932 ) ; I T ALY : S a n i co ( A l es sa n d r i a ) , E N ! P o r t acomaro ( A s t i ) , E N ! Mon ca l vo ( A s t i ) ( A s t i ) , E N ! P i a n d e l l ' A l p e ( F e n e s t re l l e , To r i n o ) , E N ! C o l l e T h u i I e , Aos t a ) , E N ! T r i e s t e ( K i effer i n K i effer e t M a rs h a l l

e t M a r sh a l l d i A l f i ano N a t t a , Ol ! C a n e l l i S . C a r l o ( la 1 905 ) .

1 567

Gonatopo d i nae

3 1 . Gon a torJUs mongolicus P o n o rr,are n k o

Gon a topus Gon a topus Gon a top u s Con a top u s

F E t" A L E :

mongolicus mongol icus mongolicus mon gol icus

aptero u s : le n gth

Pono mare n k o 1 97 2 : 678 . P o n omare n k: o 1 97 5 a : 3 1 5 . P o n omare n k o 1 97 9 a : 355 . P o n omare n k o : �/'toczar 1 983b :

2 , 5-2 , 7 mm :

blac k :

207 .

i n some s pec i me n s h ead an d

proth o r ax partl y testaceo u s : l e gs t estaceou s , w ith clu b s of femora b ro w n i s h ; ante n n ae w i th s e gm e nts 1 - 2 t e staceo u s , segment 3 b ro w n , s e gments 4- 1 0 m i s s i n g i n h oloty pe ( o n l y s e e n spec i m en ) : h ead o f holoty pe i n sli de ( colou r n o t v i s i ble ) ; ante n n al s e gments 1 - 3 i n f ollo w i n g proporti o n s : 8 : 5 : 1 0 ; pro n otu m c rossed by a s t r o n g tran s v ersal i m pre s s i o n , s h i n y , w ith o ut sc ulptu re or v e r y w eakly gran ulate d ; sc utu m d ull , lo n gi tu d i n ally s t r i at e ; metan o t u m s h o r t , hollow b e h i n d th e scu t ellu m ; rnetath o r ax + propo d e u m s h i n y , w i t h an t e r i or s u r face w i th o u t sculptu r e , smooth ( i n h oloty pe ) ; occas i o n ally ( i n spe c i m e n s i n B U ) ante r i o r s u rface gran ula t e d; pos t e r i o r s u rface and ple u ra tran s v e r s ely s t r i ate ; meso metapl e u ral s u t u r e o b s olete ( i n h oloty pe ) o r w eakly d i sti n c t an d com plete ( i n spec i me n s i n B U ) ; fore tar s al segm e nts i n fo l l o w i n g proporti o n s : 1 2 , 5 : 2 , 5 : 3 : 1 1 : 2 0 ; enlarged claw ( Fi g . 1 08 5 ) w i th o ut s u bapi cal t ooth , w ith a small tooth at th e e n d of a lon gi tu d i n al f u r r o w and w i th 3 b r i s tle s; segment 5 of f ront tar s u s ( Fi g . 1 085 ) w i th i n n e r s i d e p r o x i mally n o t s e r r at e , w i t h t w o r o w s o f 3 + 1 7 lam e l lae e x t e n d i n g b e y o n d 0 , 5 len g t h o f segment; ape x w i th a gro u p of nearly 9 lamellae ; max i llary palpi w i th 5 segm e nts ; lab i al pa l pi w i th 2 segments ; ti b i al spu r s 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : So n g i n o ( Mo n goli a ) T Y P I CAL M A T E R I A L : holo t y pe F ' i n L N . HOSTS: u n known D I ST R I BU T I O N : P alaearc t i c regi o n : M O N G O L I A : So n g i n o ( Ulaan Baatar ) , L N ! 20 Km S f rom Somon O e i ge rzogt ( M i ttelgob i a i mak ) , B U ! 26 Km SIN from Somon Sc h a r ga ( See T u n ama I , C h o v s gol a i mak ) , B U ! 1 1 K m OSO f rom Somon Baj an zo gt ( C entral a i mak ) , B U ' N O T E S : t h e spec i me n s k e pt i n BU seem sli ghtl y d i fferent f rom h o l oty pe , b ecause th e y h av e meso-metapl eu r al s u t u re w eak , b ut d i sti n c t , and ante r i o r s u r face o f propo d e u m d ull an d gran ulated .

32.

Gon a top u s longus ( P o n omare n k o ) n . comb . )

Neogon a top u s longus P o nornar e n k o 1 97 2 : 679 . Neogona top u s longus P o n omare n k o 1 975a : 3 1 6 . Neogo n a t o p u s longus Pon omare n k o 1 9 79a : 354 .

Mem . Ame r .

1 568

F E MAL E :

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f u l l y t e s t aceo u s ; a n te n n ae w i t h s e g me n t s 1 - 3 t e s t aceou s , o t her se g m e n t s m i s s i n g i n h o l o t y pe ( on l y seen spec i me n ) ; m e t a t h o r a x + propod e u m a n d p e t i o l e b l ac k ; a b domen b row n - te s t aceou s ; a n t e n n a l seg men t s i n fo l l ow i n g p ropor t i o n s : 6 : 4 , 5 : 6 ( seg me n t s 4- 1 0 m i ss i n g i n h o l o t y pe ) ; h e a d v e r y d a m aged i n h o l o t y p e , n o t d i s t i nc t l y v i s i b l e ; pron o t u m crossed b y a s t r o n g t r a n sv e r s a l i m p ress i o n , d u l l , g r a n u l a t e d ; m e t a n o t u m n o t f l a t , d u l l , g r a n u l a t ed a n d w e a k l y t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l su t u r e o b so l e t e ; m e t a t h o r a x + propodeum d u l l , f u l l y g r a n u l a t ed , w i t h pos t e r i o r s u rf ace t r a n sverse l y s t r i a t e ; p l e u r a n o t s t r i a t e t r a n sverse l y ; fore t a rs a l seg men t s i n fo l l ow i n g propor t i o n s : 1 2 , 5 : 2 : 3 : 1 0 : 1 6 ; e n l a rg e d c l a w ( F i g . 1 086 ) w i t h o u t s u b a p i c a l too t h , w i t h a sma l l too t h a t t h e e n d of a l o n g i t u d i n a l f u r row a n d w i t h 4 b r i s t l es ; seg m e n t 5 of fro n t t a r s u s ( F i g .

1 569

Gona topod i n ae

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1 087 - C h e l a of Gon a top u5 n ivos u s n . sp .

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L O C U S T Y P I C U S : 4 0 K m N o f M a n d a i -Gob i T Y P I CAL MAT E R I AL : h o l o t y p e F ! i n L N . H O ST S : u n k n o w n D I S T R I BU T I O N : ( M i d d l e-Gob i ) ,

P a l a e a rc t i c reg i o n : LN !

33.

( Mo n g o l i a )

MONGOL I A : 4 0 K m N o f M a n d a i -Gob i

Gon a top u s n i vos u s n . sp .

F E MAL E : a p terou s ; l e n g t h 3 , 0 m m ; y e l l o w-red d i s h , w i t h p e t i o l e b l ac k a n d a bdomen b ro w n ; h e a d exca v a ted , s h i n y , h a i r l es s , w i t h o u t scu l p t ure o n most p a r t of v e r t e x , g r a n u l a te d o n occ i p u t a n d fro n t p a r t o f v e r te x ; p ron o t u m crossed b y a t r a n s v e rs a l i m p re ss i o n , s h i n y , w e a k l y g ra n u l a t e d ; scu t u m s h i n y , g ra n u l a ted ; m e t a n o t u m s h o r t , w e a k l y t r a n s v erse l y s t r i a t e ; m e t a t h o r a x + p ropodeum du I I , g r a n u I a ted ; p l e u r a a n d pos t e r i o r s u rface of p ropod e u m t r a n s v e rse l y s t r i a te ; meso-m e t a p l eu r a l s u t u re o b so l e t e ; seg men t of fro n t t a r s u s s l i g h t l y s ho r t e r t h a n segmen t 4 ( 1 1 : 1 2 ) ; en l arged c l a w ( F i g . 1 087 ) w i t ho u t s u b a p i c a l too t h a n d l a m e l l a e , w i t h a s m a l l t oo t h a t t he e n d o f a l on g i t u d i n a l fu rrow a n d w i t h 4 p e g - l i k e h a i r s ; s e g m e n t 5 of f ro n t t a rs u s ( F i g .

1 08 7 ) w i t h i n n e r s i d e pro x i m a l l y not serra t e , w i t h 2

row s of 4 + 1 3 l a m e l l ae b eg i n n i n g on d i s t a l h a l f of seg m e n t 5 ; a p e x w i t h a g ro u p of 5 l a m e l l ae ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C US : S r i n a g a r ( K a s h m i r ) T Y P I CAL MAT E R I A L : ho l o t y p e F ! i n O L . HOST S : u n k n o w n D I ST R I BU T I O N :

P a l a e a rc t i c reg i o n :

I ND I A : S r i n agar ( Kash m i r ) , OL !

34 . Gona top u s brunn eicollis ( R i c h a r d s ) n . com b .

Neogon a topus brunneicollis R i c h a rd s 1 97 2 : 539 . Neogon a topus sp . 1 R i c h a rd s 1 972 : 543 .

F E MALE : a p t e ro u s ; l e n g t h 2 , 50-3 , 1 2 m m ; head red d i s h or b row n ; occas i on a l l y b ro w n-redd i s h , w i t h m a n d i b l es a n d c l y p e u s l i g h t ; a n t e n n a e y e l l ow ; t h o r a x a n d p ropode u m red d i s h o r b ro w n - r ed d i s h ; scu t u m somet i mes y e l l ow ; p e t i o l e b l ac k ; a b domen b l ac k or b row n ; occ a s i on a l l y red d i s h ; l eg s y e l l ow , w i t h coxae a n d c l u b s o f femora b row n i s h ; h e a d e x c a v a ted , h a i r l es s , s h i n y , smoo t h , w i t ho u t scu l p t u re ; scu t u m s h i n y , w i t h some l on g i t u d i n a l k ee l s ; m e t a n o t u m t r a n s v erse l y s t r i a te ; m e t a t ho r a x + p ropodeum t ra n s v erse l y s t r i a te ; meso-me t a p l e u r a l s u t u re o b so l e te ; seg men t 1 of fro n t

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t a rs u s as l o n g as s e g m e n t 4 ; en l arged c l aw ( F i g . 1 08 8 ) w i t h o u t s u b a p i c a l t oo t h a n d l a m e l l ae , w i t h a sm a l l toot h a t t h e e n d o f a l o n g i t u d i n a l fu rrow a n d w i t h 4-7 peg- l i ke h a i rs ; segmen t 5 of fro n t t a r s u s ( F i g . 1 088 ) w i t h i n n er s i de p ro x i m a l l y not serra t e , w i t h 2 rows of 1 5-25 l a me l l a e ( so m e t i mes b eg i n n i n g o n i s t a l h a l f of segme n t 5 ) ; apex w i t h a g ro u p of 6-9 l ame l l ae ( some l a m e l l a e are m u c h l o n ge r t h a n o t h e r l a me l l ae ) ; m a x i l l a ry p a l p i w i t h 4-5 s e g me n t s ; l ab i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k n o w n L O C U S TYP I C U S : R e hovot ( I s r a e l ) TYP I CA L M A T E R I AL : h o l o t y p e F ! i n G V ; 1 p a r a t y p e F ! i n B M ; 1 p a ra t y pe F ! i n H a r p a z co l l ec t i on . HOS T S : i n I s ra e l u n k no w n Cic a dellidae D I ST R I B U T I O N : P a l aearc t i c reg i o n : U . S. S. R . : T a s h k e n t ( U z b e k i s t a n ) , WA ! S PA I N : C a l e l l a d . C o s t a ( B a rce l on a ) , B M ! I SR A E L : Rehovo t , GV ! CANARY I SLA N D S : G r a n T a raj a l ( F u e r t ev e n t u r a ) , O l ! H E ! B a rranco de T a ho d i o ( T e n er i fe ) , O L ! M O ! Med a n o ( Te n er i fe ) , M O ! L o s C a l derones ( M e d a n o , T e n e r i fe ) , M O !

35 . Gon a topus cherson esius Ponomarenko

Gon a top u s chersonesius Ponomarenko 1 970 : 2 5 7 . Gon a top u s chersonesius P o n o m a re n ko : Ponomarenko 1 978 : 2 6 .

F E MALE : a p te ro u s ; l e n g t h 4 , 37 m m ; h e a d t e s t aceous-re d d i sh ; a n te n n a e b row n , w i t h s e g me n t s 1 -2 tes t aceou s ; t h o r a x a n d p ropode u m b ro w n -red d i s h , excep t for p rono t u m y e l l ow - t e s t a ceou s ; a b domen b row n - re d d i s h ; l eg s t e s t a c eo u s ; a n te n n ae d i s t a l l y t h i ck e n e d ; a n t e n n a l segmen t s i n fo l l ow i n g p ropo r t i o n s : 1 3 : 8 : 1 9 : 1 2 : 1 2 : 1 0 : 8 : 9 : 7 : 1 2 ; h e a d e x c a v a te d , g ra n u l a te d , s h i n y ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 4 ; OL = 4 ; OOL = 1 3 ; p rono t u m crossed b y a s t r o n g t r a n s v ersa l i m p re ss i o n , sh i n y , g r a n u l a ted ; s c u t u m d u l l , w i t h some l on g i t u d i n a l k ee l s ; s c u t e l l u m b roa d , smoo t h ; m e t a n o t u m r u g o s e , weak l y ho l l o w b e h i n d t h e s c u te l l u m ; meso-me t ap l e u r a l s u t u r e com p l e t e a n d d i s t i n c t ; m e t a t horax + p ropode u m d u l l , w i t h n u merous s t ro n g s t r i ae o n p l e u ra and pos t e r i or s u rface of p ropodeu m ; a n te r i or s u rface of p ropodeum smoo t h , except for some d i s t i n c t a n d l o n g l on g i t u d i n a l k ee l s ; fore tarsa l s e g m e n t s i n fo l l ow i n g p roport i o n s : 2 2 : 5 : 7 : 2 2 : 33 ; e n l a r g e d c l a w ( F i g . 1 089 ) w i t ho u t s u b a p i c a l toot h , w i t h a s m a l l tooth a t t h e e n d of a l on g i t u d i n a l fu rrow a n d w i t h 2 b r i s t l es ; segmen t 5 of f ro n t t a r s u s ( F i g . 1 089 ) w i t h 2 rows of 4 + 1 8 l a me l l ae e x t e n d i n g b e y o n d 0 , 5 l e n g t h o f seg m e n t 5 ; a p e x w i t h a g ro u p o f 1 0 l a me l l a e ; m a x i I l a ry p a l p i w i t h 5 segmen ts ; l a b i a l p a l p i w i t h 2 s e g m e n t s ; t i b i a l spurs 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S :

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Gon a top u s sepsoides Wes t wood 1 633b : 496 . Gon a top u s p il o s u s T homson 1 860 : 1 80 ( s y n . p roposed by R i c h a r d s 1 939 ) ; locus typ ic u s Sma I a n d ( Sweden ) . L a beo nigerrimus Forster 1 86 1 : XXX I X ( n . sy n . ) ; locus typ icus Rosegg t h a l ( S w i tzer l a n d ) . L a b eo pusillus Szep l i ge t i 1 90 1 : 1 56 ( n . sy n . ) ; locus typ i c u s K a z a n ( U . S. S . R . ) . Gon a top u s h ispanicus K i effer i n K i effer e t M a r s h a l l 1 90 5 : 1 1 8 ( n .

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sy n . ) ; locus typ ic us Pozue l o de C a l a t r a v a ( S p a i n ) . p roposed b y R i c h a r d s 1 939 ) ; locus typicus C a mber ( E n g l a n d ) . Dicondylus sepsoides ( West wood ) : K i effer 1 907b : 2 2 . Gonatop u s borealis J . S a h l berg 1 9 1 0 : 1 2 ( s y n . p roposed by He l l en 1 9 53 ) ; locus typicus K u u s a mo ( U . S . S. R . , near K u u samo p rov i nce i n F i n l a n d ) : accor d i n g t o D r . An ders A l b rec h t , Z oo l o g i c a l M u seum o f t h e U n i v ers i t y , He l s i n k i , ( person a l comm u n i c a t i o n ) p rob ab l y S a h l berg ' s f i n d i s from t h e R u s s i a n s i de of t he border ) . L a b erius p usill us ( Szep l i ge t i ) : K i effer 1 9 1 4b : 6 2 . Gon a top u s wagneri S t r c n d 1 9 1 9 : 20 1 ( n . sy n . ) ; locus typicus V e l ten ( German y ) Gon a top u s formicari us Lj u n g h : L i c h t e n s t e i n a n d P i c a r d 1 920 : 5 4 . Gon a top u s pilos us T homson : Ber l a n d 1 928 : 1 50 . Gonatopus formicari u s L j u n g h : Berl a n d 1 928 : 1 5 1 . Gon a top u s sepsoides West wood : R i c h a rd s 1 935 : 1 46 . Gonatopus sociabilis K i effer : R i c h a r d S 1 935 : 1 47 . Gonatopus sepsoides Wes t w ood : Behri sch 1 95 2 : 3 3 . Dicondylus p u s illus ( Szep l i ge t i ) : Mocz a r 1 96 5 : 394 . Gon a top u s sepsoides Wes t w oo d : N a g y 1 967a : 1 24 . Gona top us sepsoides West wood : C u rrado a n d O l m i 1 972 : 1 52 . Gon a topus sepsoides West wood : O l m i a n d C u rrado 1 974 : 2 1 5 . Gon a top u s sepsoides West wood : Abdu l -N o u r 1 976 : 266 . Gonatopus sepsoides West wood : O l m i 1 976 : f i g s 6-9 . Gonatop u s sepsoides Westwood : Ponom a re n k o 1 97 8 : 2 6 . Gonatop u s sepsoides West wood : S a n der 1 980 : 5 8 . Gon a top us sepsoides West wood : O l m i 1 98 1 : 1 1 6 . Dicon dyl us p usillus ( Szep l i ge t i ) : Moczar 1 983a : 1 94 . Dicondylus p u sillus ( Szep l i ge t i ) : Mocz a r 1 983b : 206 . Gonatop u s sociabilis K i effer 1 907 a : 7 ( s y n .

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F E MA L E ( F i g . 1 090 ) : ap terou s ; l e n g t h 2 , 50-3 , 75 mm ; co l o u r very v a r i a b l e ; u s u a l l y b l a c k , w i t h m a n d i b l e s , c l y p e u s a n d fron t p a r t o f vertex y e l l ow ; a n te n n ae b l ack , w i t h segmen ts 1 -2 a n d some t i mes p a r t l y 3 ye l l ow ; ab domen b l ac k , w i t h red d i s h n u a n ces ; l e g s y e l l ow-red d i s h , w i t h c l u b s of femora b row n i s h ; in some spec i m e n s p ro n o t u m a l mo s t fu l l y redd i s h o r w i t h s i de s redd i s h a n d t r a n s v e r s a l i m p ress i o n b l ack , sc u t u m y e l l ow or b l ac k redd i s h , sc u te l l u m a n d m e t a t hora x + p ropodeum b l ack-redd i s h , ab domen a l most fu l l y re d d i s h ; hea d exca v a ted , s h i n y or d u l l , a l u t aceo u s or g r a n u I a te d ; p ro n o t u m crossed by a t r a n sversa I i m press i o n , s h i ny or du I I , g r an u l a ted , h a i r y or h a i r l ess ; s c u t u m g r a n u l a t e d , w i t h some l on g i t u d i n a l kee l s ; met a n o t u m t r a n sv erse l y s t r i a t e ; me t a t horax + p ropodeum ( F i g . 1 09 1 ) s h i n y or d u l l , h a i r y , g r a n u l a te d ; p l e u r a a n d. poste r i o r s u rface of p ropode u m t r a n sverse l y s t r i ate ( F i g . 1 09 1 ) ; meso-met a p l e ura l s u t u re d i s t i nc t a n d comp l e t e ; fore t a r s a l segme n t s i n fo l l ow i n g proport i o n s : 1 2 : 2 : 4 : 1 4 : 2 1 ; e n l a r g ed c l a w ( F i g s 1 092 , 1 093 ) w i t h o u t s u b a p i c a l tooth a n d l a mel l ae , w i t h a sm a l l too t h a t t h e e n d o f a l o n g i t u d i n a l fu rrow a n d w i t h

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some b r i s t l es o r peg - l i k e h a i rs ; seg men t 5 of fron t t a rs u s ( F i g s 1 092 , 1 094 ) w i t h i n n e r s i d e p r o x i m a l l y n o t serra t e , w i t h 1 row of 1 1 - 1 2 l a m e l l ae b eg i n n i n g on p ro x i m a l t h i rd of s e g m e n t 5 ; a p e x w i t h a g ro u p of 7-1 7 l a m e l l ae ; m a x i l l a r y p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l spurs 1 , 0 , 1 . M AL E : fu l l y w i n ged ; l e n g t h 1 , 68-2 , 75 m m ; b l ac k , w i t h m a n d i b l es a n d t e g u l ae y e l l o w ; a n t e n n ae b ro w n - b l ac k ; l eg s y e l l o w - b ro w n , w i t h coxae d a r k ; a n t e n n a l se g m e n t s i n fo l l o w i n g p ropor t i on s : 4 : 4 : 6 : 5 , 5 : 5 , 5 : 5 : 5 : 5 : 5 , 5 : 8 ; a n t e n n a l seg m e n t 3 l es s t h a n 3 , 5 t i mes a s l o n g as b ro a d ( 3 , 2 ) ; a n t e n n ae not d i s t a l l y t h i c k e n e d ; h e a d d u l l , h a i r y , g r a n u l a t e d , w i t h a s h i n y area b e t w e e n oce l l a r t r i a n g l e a n d eyes , w i t h s h o r t i r reg u l a r k ee l s o n v e r t e x a n d fro n s ; tem p l es d i s t i n c t ; occ i p i t a l ca r i n a i n v i s i b l e ; scu t u m d u l l , g r a n u l a t e d ; n o t a u l i ce s com p l e t e , p o s t e r i o r l y s e p a r a ted ( F i g . 1 070 A ) ; m e t a n o t u m s h i n y , w i t ho u t scu l p t u re ; p ropod e u m d u l l , s t ron g l y ret i c u l a t e ru gose , w i t h a med i a n fu rrow o n a n t e r i o r s u rface ; r ad i a l v e i n : f i g . 1 048 C ; dors a l process of g o noforceps ( F i g s 1 072 A, 1 09 5 ) s h or t , t r a n s verse , w i t h a m e m b r a n o u s p a r t on dors a l s i d e ; b a s i v o l se l l a w i t ho u t n o t c h ; m a x i l l a r y p a l p i w i t h 5 s e g m e n t s , l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . L O C U S TYP I C US : B l ac k g a n g C h i n e ( I s l e of W i g h t , E n g l a n d ) TYP I CA L MAT E R I A L : ho l o t y p e F ! o f G . sepsoides i n OX ; l ec to t y p e F ! ( d e s i g n a te d b y O l m i 1 977 ) of G . pilos u s i n ST ; h o l o t y p e F ! of G .

h isp a n ic u s i n P ; ho l o t y pe F ! o f G . soc i a b ilis i n B M ; l ec t o t y p e F ! ( hereby des i g n a t ed ) of G . borealis in H E ; h o l o t y p e M ! of L. nigerrim u s in V I ; h o l o t y p e M ! of L . pusillus i n BU . HOST S : i n I t a I y Diplocolen u s a b domin a l i s ( F . ) , Psammotettix stria t u s L . ( = a lien u s D a h l b . ) ( C u rrado a n d O l m i 1 972 ) ; i n B u l g a r i a Ja ssargus repletu s ; i n F r a nce Eu scelis plebejus F a l l e n , Ps ammotettix stria tus L . , Ps ammotettix n ota tus M e l . , Psammotettix p u toni T h en . , Ja s s a rgus obtu sivalvis K b m . , A darrus ta urus R i b . , A darrus gen icula tus R i b . , A darrus prop i n q u u s F i eb . , A rocep h a l u s s a gittarius R i b . , Golde u s h arpago R i b . , Rhopa lopyx elonga tus W g n . ( A b d u l -N o u r 1 976 ) ; in G e r m a n y E u scelis incis u s ( K b m . ) ( Sa n der 1 980 ) , Arocep h a l u s l a n guidus ( F l . ) ( Sa n der 1 980 ) ; i n A u s t r i a Deltocep h a l u s a ssimilis F a l l . ( T hompson 1 954 ) a n d Sorhoa n u s xan thoneurus ( F i eb . ) ( M i k 1 882 ) ; i n U . S . S . R . Turr u t u s socia lis ( F l o r ) ( Po n o m a r e n k o 1 968 ) ; i n E n g I a n d Ps ammotettix confinis D a h l b . , Ja s s a rgus pseudocellaris F l or , Ja s s argus flori F i eb e r , Erra s t u n u s ocellaris F a l l . , Cic a d u l a sp . ( W a l off 1 974 ) ; Arth a l de u s p a scuellus ( F a l l . ) ( R i c h ar d s 1 939 ; W a l off 1 974 ) ; Psammotettix n o dosus R i b . ( W a l off 1 975 ) ; Ps ammotettix s tr i a t u s L . ( R i c h a r d s 1 939 ; W a l off 1 975 ) ; i n F i n l a n d Diplocolenus a b domin alis ( F . ) ( L i n d berg 1 950 ) • B I OLOGY : i n forma t i o n s i n A b d u l - N o u r ( 1 976 ) . D I ST R I B U T I O N : P a l a e a rc t i c reg i o n : K O R E A : N u n g-ra do I s l a n d ( P y o n g y a n , P ro v o Sou t h P y o n g a n ) , B U ! M O N G O L I A : 2 0 K m S from Somon -zo g t ( M i t te l gob i a i m ak ) , B U ! M o n go l i a ( P o n o m a re n k o 1 979 a ) ; U . S . S . R . : K a z a n , B U ! K u u s a m o , H E ! K a n t a l a h t i ( So v i e t L ap l a n d ) , H E ! K r a s n o j a r s k ( S i b i r i a ) , H E ! SWE D E N : S k a n e , L U ! Sma l a n d . ST ! R a f t e n ( Sk a n e ) , L U ! G a n to f t a ( Sk a ne ) ,

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L U ! POLAN D : K U h n a u , HA ! S i l es i a , V I ! H U N GA R Y : U j - F e h er t o , BU ! T i h a n y , B U ! N . E n yed , BU ! Zemp l e n i h g s . , B U ! Boros Benede k , BU ! R ev . , B U ! Vacz ( T u dosdomb ) , B U ! Sz . -F e h e r to , B U ! O r s z t M i k l os , BU ! B u d a p es t , B U ! M t s . Borzso n y , BU ! N a g y v a zso n y ( K a b h e g y ) , BU ! R O M A N I A : A g i gea ( Do b rogea ) ( N agy 1 967 a ) ; B U L GA R I A : K U s t e n d i l . , B U ! B u l g a r i a , SA ! O L ! C Z E C H O SLOVAK I A : D o n ov a l i ( Z v o l a n ) , BU ! G E R MA N Y : V e l t e n ( Z i e g e l e i en ) , E B ! K y ff h a u se r ( T h U r i n g e n ) , E B ! S t i f t s b a u m Se h l i s ( L e i p z i g ) , E B ! L e i pz i g , E B ! HA ! H a l l e ( S . ) , E B ! Aachen ( No r d r h e i n-Westfa l e n ) , L E ! W a l l e n d orf ( R he i n l a n d ) , E B ! B e l l i n c h e n ( O der ) , HA ! K refe l d ( Nordrhe i n -W e s t f a l e n ) , HA ! D U nen R a n t u m ( S y l t , Sch l e sw i g -Ho l s t e i n ) , HA ! W a l d k i sc h ( B a d e n ) , B ! T h U r i n g en , G E ! A U S T R I A : P ra t e r ( W i e n ) , P ! T u rk e n-Sc h a n ze ( W i e n ) , P ! K a l k s b u r g ( N i eder-Osterre i c h ) , V I ! B i b erw i e r ( T i ro l ) , E B ! D E N M A R K : R u de r h e g e n , WA ! D e n m a r k , C O ! SW I T Z E R LA N D : N y o n , G V ! Gorn e r g r a t ( V a l a i s ) , G V ! Roseg g t h a l , V I ! HOLLA N D : D e n H a a g , L E ! Noord w i j k , L E ! D r i e b e r g , L E ! F R A N C E : H y eres ( V a r ) , P ! O r l ea n s ( Lo i re t ) , P ! T e n ce ( H a u te-Lo i re ) , P ! D i jon ( Co te-d ' O r ) , P ! Pot henee ( C o te-d ' O r ) , P ! M a n d u e l ( G ard ) , P ! S t Ge l y -d u - F esc ( He r a u l t ) , P ! L a t t es ( Hera u l t ) , P ! L a F i g a rede ( Hera u l t ) , P ! Mon t p e l l i e r ( Herau l t ) , P ! S t P a u l -e t - V a l m a l l e ( He r a u l t ) , P ! Mon t a r n a u d ( He r a u l t ) , P ! Combe de M a r t i es ( He r a u l t ) , P ! Le P u y , P ! E NGLAN D : C r o y d e ( N D e v o n ) , B M ! S t u d l a n d ( D orset ) , HA ! B a r n e s ( Lo n d o n ) , O X ! D u l w i c h , O X ! B l ac k g a n g C h i n e ( I s l e of W i g h t ) , OX ! n u merou s l oc a l i t i es i n R i c h a rd s ( 1 939 ) ; I R E L A N D ( R i c h a r ds 1 939 ) ; S PA I N : Poz u e l o de C a l a t ra v a , P ! I T ALY : S t a zz a n o ( A l e s s a n d r i a ) , G E ! S a n i co d i A l f i a n o N a t t a ( A l es s a n d r i a ) , E N ! Portacomaro ( A s t i ) , E N ! Monca l vo ( A s t i ) , O L ! Tonco ( A s t i ) , E N ! Ses s a n t ( A s t i ) , E N ! C a n e l l i ( A s t i ) , E N ! Sa v i g l i a n o ( C u n eo ) , SZ ! S . Benede t to B e l bo ( C u n eo ) , O L ! P L ! Sc i o l ze ( T o r i no ) , O L ! La C a s s a ( T or i n o ) , O L ! P i a n de l l ' A l p e ( Fe n e s t re l l e , Tor i n o ) , E N ! F e n e s t re l l e ( To r i n o ) , O L ! C h a m p i a s - d u - C o l ( Se s t r i e r e , To r i no ) , OL ! L a g o de l Monce n i s i o ( T or i n o ) , O L ! M a sserano ( Verce l l i ) , O L ! B a l m ucc i a ( Verce l l i ) , O L ! Pecc i a ( R i v a V a l d o b b i a , Verce l l i ) , OL ! R i fu g i o Pas tore ( A l a g n a Va l s es i a , Vercel l i ) , O L ! A l p e P i a n a l u n g a ( A l a g n a Va l ses i a , Verce l l i ) , O L ! V a l F erret ( Co u r m a y eu r , Aos t a ) , E N ! C o l l e S. C a r l o ( L a T h u i I e , Aos t a ) , E N ! Po R i ver , C O ! l eso l o ( Ve n e z i a ) , VR ! C a s t e l v e t ro ( Mode n a ) , BO ! Sp i l a m b e r t o ( Mode n a ) , G E ! A l b i s so l a ( Sa v on a ) , O L ! Cec i n a ( L i vorn o ) , GE ! P i a n G r a n d e ( C a s te l l ucc i o d i Norc i a , Per u g i a ) , OL ! O r t e ( V i t e r bo ) , OL ! R i e t i , B O ! Sa sso-F u r b a ra ( C e r v e t er i , R o ma ) , T R ! R a n d azzo ( C a t a n i a ) , E N ! G R E E C E : M t . H o l o m o n d a s ( H a l k i d i k i ) , OL ! T U R K E Y : A n k a r a , B U !

3 7 . Gona topus rhaen sis P o n o m a r e n k o

Gon a top u s rhaensis P o n o m a re n k o 1 970 : 4 3 1 . Gon a top u s rhaensis Ponorn a re n ko : Pon o m a re n k o 1 978 : 2 6 .

F E MAL E :

a p terou s ;

l en g t h 3-3 , 5 m m ; h e a d b row n ,

w i t h occ i p u t , c l y p e u s

Go n a topod i n a e

1 583

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Fig.

1 096 - C h e l a of Gon a topus rhaensis Ponomarenko ( t y p e )

a n d m a n d i b l e s t e s t acou s ; a n t en n ae b row n , w i t h seg m e n t s 1 -2 t e s t a ceou s ; t h orax a n d p ropode u m b l ac k ; a b d omen b ro w n - te s t aceou s ; l eg s t e s t a ceou s , w i t h coxae a n d c l u b s of femora b row n ; a n te n n a e d i s t a l l y t h i c k e n e d ; a n t e n n a I se g m e n t s i n fo l l o w i n g p roport i o n s : 1 1 : 6 : 1 5 : 9 : 9 : 8 : 6 : 6 , 5 : 6 : 9 ; h e a d s l i g h t l y exca v a ted , d u l l , g ra n u l a t e d ; frons w i t h a com p l e t e m ed i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 3 ; O L = 3 ; O O L = 1 3 ; p rono t u m crossed b y a t r a n s v e r s a I i m p ress i o n , d u I I , g ra n u l a ted ; scu t u m

1 584

Mem . A m e r .

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En! .

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n o . 37 ,

1 984

1 097 - C h e l a of Gon a top u 5 b a rb ieri n . sp .

w i t h some l on g i t u d i n a l

k ee l s ;

sc u t e l l u m d u l l ,

gran u l ated ;

( type)

met a n o t u m w e a k l y

h o l l ow b e h i n d the s c u t e l l u m ; meso-me t a p l e u r a l s u t u re d i s t i nc t a n d comp l e te ; p o s te r i o r s u rface of p ro p o d e u m t ra n s verse l y s t r i a t e ; p l eu ra t r a n sverse l y not s t r i a t e ; m e t a t h o r a x + p ropodeum d u l l , g r a n u l a te d , w i t h a t r a k of

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1 585

med i a n l o n g i t u d i n a l furrow ; fore t a rs a l seg ments i n fo l l ow i n g p ropor t i o n s : 2 0 : 5 : 6 : 2 2 : 35 ; en l a rged c l a w ( F i g . 1 096 ) w i t h o u t s u b a p i c a l toot h , w i t h a sma l l too t h at t h e end of a l o g i t u d i n a l fu rrow a n d w i t h 3 b r i s t l es ; segme n t 5 of front t a r s u s ( F i g . 1 096 ) w i t h 2 row s of 1 6 + 6 l a me l l ae exte n d i n g b e y o n d 0 , 5 l e n g t h of segmen t ; apex w i t h a group of 1 3 l a me l l ae ; m a x i l l a r y p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : K i n e l ( U . S . S . R . ) T Y P I CAL MAT E R I AL : h o l o t y p e F ! a n d 6 p a r a t y pes F F ! i n L N ; 28 p a r a t y pes FF! i n PK. HOST S : i n U . S . S . R . Psammotettix stri a t u s ( Ponomarenko 1 978 ) D I ST R I BU T I O N : P a l aearc t i c reg i o n : U . S . S . R . : K i ne l ( K u y b y s h e v P ro v i nce ) , L N ! P K !

38 .

Gon a top u s barbieri n . s p .

F E MALE : aptero u s ; l e n g t h 3 , 60 mm ; head b l ac k , w i t h fro n t p a r t o f vertex ( more a l o n g orb i t s ) , c l y p e u s and m a n d i b l es test aceous ; a n te n n ae b row n , w i t h segments 1 -2 test aceo u s ; t h o ra x , propod e u m a n d a b domen b l ac k ; l e gs b a n de d , b ro w n , w i t h tars i , troc h a n ters a n d s t a l k s of femora test aceo u s ; a n te n n ae d i s t a l l y t h i ckened ; a n t e n n a I seg ments i n fo l l ow i n g proport i o n s : 1 0 : 7 : 1 9 : 1 0 : 9 : 9 : 7 : 7 : 7 : 9 ; head d u l l , g ra n u l ated ; f ron t a l l i ne comp l e te ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 3 ; OL = 2 ; OOL = 8 ; pron o t u m crossed b y a s t r o n g t r a n s vers a l i m pres s i o n , s h i n y , g r a n u l a ted ; sc u t u m s h i n y , w i t h l o n g i t u d i n a l k ee l s , l a tera l l y w i t h two po i n t s ; m e t a n o t u m not h o l l o w b e h i n d the scute l l u m , s t ron g l y r u gose a n d scu I p t u red b y t r a n s v ersa I k ee l s ; meso-met a p l e u r a l s u t u re comp l e te , weak , b u t d i s t i n c t ; mesop l e u r a t r a ns verse l y n o t s t r i a te ; m e t a t horax + p ropodeum s h i n y , w i t h a n ter i o r s u rface g ra n u l ated a n d w i t h some l o n g i t u d i n a l kee l s ; poster i or s u rface a n d metap l e u ra s t ron g l y t r a n s verse l y s t r i a t e ; fore tarsa l seg m e t n s i n fo l l ow i n g propor t i on s : 1 9 : 3 : 5 : 1 8 : 27 ; en l a rged c l a w ( F i g . 1 097 ) w i t h o u t s u b a p i c a l too t h , w i t h a sma l l too t h a t t h e e n d o f a l o n g i t u d i n a l fu rrow a n d w i t h two b r i s t l es ; seg ment 5 of front t a r s u s ( F i g . 1 097 ) w i t h a row of 1 9 l a me l l a e ; a pex w i t h a group o f near l y 1 3 l am e l l ae ; ma x i l l a ry p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k no w n L O C U S T Y P I C US : L a St i d i a ( A l g er i a ) TYP I CAL MAT E R I AL : ho l ot y p e F ! i n P . HOST S : u n k no w n D I STR I BU T I O N : P a l aearc t i c reg i o n : ALGER I A : L a St i d i a ( Oran ) , P ! NOT ES : the spec i es i s n a med i n honor of M r . J . B a r b i e r .

Mem . A m e r . E n t .

1 586

Fig.

1 098

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1 984

C he l a of Gon a topus gra ec u s n . sp .

( type)

G o n a topod i n a e

1 587

3 9 . Gon a topus schenklingi S t r a n d ( see O r i en t a l spec i e s )

40 . Gon a top us graecus n .

sp .

F E MAL E : a p te rou s ; l e n g t h 2 , 48-2 , 80 m m ; b l ac k , w i t h c l y p e u s , m a n d i b l es , a n te n n a l se g m e n t 1 , t a rs i , t i b i ae a n d troc h a n ters t e s t aceou s ; occ a s i on a l l y s c u t u m p a r t l y re d d i s h ; a n te n n ae d i s t a l l y t h i c k e n e d ; a n t e n n a I segmen t s i n fo l l o w i n g propor t i o n s : 1 5 : 8 : 1 7 : 1 5 : 1 2 : 1 2 : 9 : 9 : 7 : 1 1 ; h e a d exc a v a t e d , d u l l , g ra n u l a te d ; fron t a l l i n e comp l e t e ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 1 ; Ol = 3 ; O O l = 1 0 ; prono t u m crossed by a s t ro n g t r a n s vers a l i m p ress i o n , d u l l , g r a n u l a ted ; , scu t u m d u l l , f u l l y g r a n u l a ted ; s c u t e l l u m s l i g h t l y v i s i b l e , g ra n u l a t e d , d u l l ; m e t a no t u m f l a t , not h o l l o w b e h i n d t h e s c u t e l l u m , d u l l , g ra n u l a t e d , n o t s t r i a t e ; m e t a t horax + p ropodeum d u l l , g ra n u l a t e d , w i t ho u t s t r i ae on p l e u ra a n d pos t e r i o r s u rface ; occ a s i o n a l l y d i sc w i t h a t r ack of med i a n f u r row ; meso-me t a p l e u r a l s u t u re d i s t i n c t a n d comp l e t e ; fore t a rs a l s e g me n t s i n fo l l ow i n g propor t i o n s : 1 5 : 3 : 6 : 1 4 : 2 3 ; segme n t 3 of fron t t a r s u s p r o d u ced i n t o a hook ; en l a r g e d c l a w ( F i g . 1 098 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d o f a l o n g i t u d i n a l fu rrow and w i t h 5 peg- l i k e h a i rs ; segmen t 5 of fro n t t a rs u s ( F i g . 1 098 ) w i t h i n ner s i de pro x i m a l l y serra t e a n d w i t h one row of 1 6 l a me l l ae ; a p e x w i t h a group of n e a r l y 1 1 l am e l l ae ; m a x i l l a ry p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k nown l O C U S T Y P I C U S : K a l e n d z i ( A h a i a , Greece ) T Y P I CAL M A T E R I A L : ho l o t y p e F ! a n d 1 p a ra t y p e F ! i n O l . HOST S : i n Greece Doratura s p . D I ST R I B UT I O N : P a l a e a rc t i c reg i o n : G R E E C E : K a l e n d z i ( M t , E r y m a n t hos , A h a i a ) , Ol !

E T H I O P I A N S P E C I ES OF GONA TOPUS

T h i r t y spec i es are k n o w n .

K E Y TO T H E S P E C I E S

F E MALES

S eg men t s eg men t

1 o f f r on t t a r s u s app r o x i ma t e l y t w i c e a s l on g a s 4 . . . . . . . . . . . . . . . . . . . . 1 . meridion alis ( B e n o i t )

Mem . A me r . E n ! .

1 588

I n s! . ,

S eg m e n t 1 of f r on t t a r s u s as l on g e r t h a n s eg me n t 4 .

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1 984

or sh o r t e r or s l i g h t l y .

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o f f r on t t a r su s w i b e y on d 0 , 5 l e n g t h . . . . . . . . . . . . . . . . . . . o f f r on t t a r s u s w i

3

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S eg m e n t 5 e x t en d i n g . . . . . . . . . S eg m e n t 5

2

. 18

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S cu t u m v e ry l on g , n e a r l y t h r e e t i me s a s l on g a s b r o a d ( F i g . 1 1 22 B ) . . . 2 . longi u s c u l u s n . s p . S c u t u m s h o r t e r , t w i c e o r l e s s t h a n t w i c e a s l on g a s b r oad S c u t u m l a t e r a l l y w i t h 2 p o i n t ed p r o m i n en c e s . . . S c u t u m l a t e ra l l y w i t h ou t 2 p o i n t ed p r om i n en c e s . . .

4

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Me s o -me t a p l eu ra s u t u r e o b s o l e t e o r s l i g h t l y d i s t i n c t a n d i n c omp l e t e M e s o -me t a p l e u ra l s u t u r e d i s t i n c t a n d c o mp l e t e . . . .

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4 5

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t h 2 r o w s o f l a me l l a e ; s h o r t e s t r o w o f s egmen t ( F i g . 1 1 0 1 ) . . . . . . . . . . . . . . . . . 3 . a us tr a l is ( C e b a l l o s ) t h 1 r o w o f l ame l l a e ( F i g . 1 1 0 6 ) o r

w i t h 2 r o w s ( i n t h i s c a s e s h o r t e s t r o w n o t ex t e n d i n g b ey o n d 0 , 5 l en g t h o f s eg m e n t ) ( F i g . 1 1 0 7 ) 6 6 F r on s w i t h ou t l on g i t u d i n a l k e e l s . . . 7 F r on s w i t h n u me r o u s l on g i t u d i n a l k e e l s . . . . . . . 8 7 H ea d f I a t . . . . 7 . a n k a r a trus ( C eb a I l o s ) H ea d s t r o n g I y e x c a v a t ed 8 . capensis B ru e s 8 H e a d l e s s e x c a v a t ed ( F i g . 1 1 0 2 A ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . · . . 4 . pilosipes B ru e s H ea d mo r e e x c a v a t ed ( F i g . 1 1 0 2 B ) 9 9 An t e r i o r s u r f a c e o f p r o p o d e u m s h i ny , smoo t h , w i t h ou t s cu l p t u r e . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 . guigliae ( B en o i t ) A n t e r i o r s u r f a c e o f p r o p o d e u m d u l l , w i t h n u m e r o u s l on g i t u d i n a l kee I s 6 . s eyrigi ( C eb a I l o s ) 1 0 T h o r a x a n d p r op od e u m y e l l ow - t e s t a c e ou s o r t e s t a c e o u s - r ed d i s h , s ome t i me s w i t h b r own sp 0 t s 1 1 T h o ra x a n d p r o p o d e u m b l a c k . . . . . . 16 1 1 T emp l e s mo r e p r om i n en t ( F i g . 1 1 0 9 A , C ) . . . . . . . 12 •

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T emp l e s

l e s s p r om i n en t ( F i g s 1 1 0 9 B , 1 1 1 3 ) . . . . . . . . . . . . . . f lat . . . . 9 . mosseli n . sp . V e r t ex v e ry e x c a v a t ed . 1 0 . rubrithorax ( B e n o i t ) B od y f u l l y y e l l ow - t e s t a c e ou s , w i t h p e t i o l e b l a ck B od y a t l e a s t p a r t l y b l a ck o r b r own o r d a rk A n t e r i o r su r f a c e o f p r op od e u m mo r e i n c l i n ed ( F i g . 1 1 1 1 B 3 0 . ericeti n . sp . A n t e r i o r su r f a c e o f p r o p o d e u m l e s s i n c l i n ed ( F i g . 1 1 1 1 A · 1 1 . hya l in u s n . sp . S c u t e l l u m a n d me t a t h o r ax + p r op od eu m b l a ck , w i t h a p ex o f p r op od eu m r ed d i s h . . . 1 2 . ridens n . sp . S c u t e l l u m a n d me t a t h o rax + p r op od e u m r ed d i sh , w i t h b l a c k b r own n u a n c e s o n p l e u ra 1 3 . comm u n is n . sp . P o s t e r i o r s u r f a c e o f p r op od e u m t r a n s v e r s e l y n o t s t r i a t e . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 4 . l u teipes ( B en o i t )

1 2 V e r t ex a l mo s t

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P o s t e r i o r s u r f a c e o f p r op od eu m t r a n s v e r s e l y

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1 589

G o n a topod i n ae

1 7 An t e r i o r s u r f a c e o f

p r op od eu m s h i n y , w i t h ou t s c u l p t u r e • • • . . . • • . . . . . . . . . . . . . . . 1 5 . bekilya n u s ( B e n o i t ) An t e r i o r s u r f a c e o f p r op od eu m d u l l , w i t h n u me r o u s weak l on g i t u d i n a l s t r i a e . . . • • . . . . • • • 1 6 . incognitus n . sp . S c u t u m v e ry l on g , n ea r l y t h r e e t i me s a s l on g a s b r oa d • • • • • . . . . . • . . . . • • • • . • • • . • . • . • • • . . . . • . . . . . 1 7 . leptothorax C eb a I l o s S c u t u m sh o r t e r , l e s s t h a n t w i c e a s l on g a s b r oa d . . . . . . • • • . . . . 1 9 Me t a t h o r ax + p r op od e u m w i t h s t r ong c en t ra l f u r r ow • • • • • • • • . • • • 2 0 M e t a t h o r ax + p r op od e u m w i t h ou t s t r o n g c en t ra l f u r r ow o r on l y w i t h w e a k t ra c k • • • • • . • • • • . . • • • • • • • • • • • • • . . • • • . . . . • . • . . • • • . . . . 2 4 Me t a n o t u m w i t h s i d e s r o u n d ed ( F i g . 1 1 2 4 A ) • • • . • • • . . . • • . . . • • . • . • • . . . . • • • . . . . • • • • . • • • • • • . • . • • . . • 1 8 . a e thiopicus n . sp . Me t a n o t u m w i t h s i d e s r i s i n g t o a r o u n d ed o r s h a rp l a t e ra l p o i n t ( F i g . 1 1 24 B ) • • • . . . . • • • . . . . . • • . . . • . • . . . • . • • . . • . . . • . . . . • • . • • • . 2 1 Me t a n o t u m n o t h o l l ow b eh i n d t h e s c u t e l l u m ( F i g . 1 1 2 6 B ) . . • . . • • . • • • • • • . . • • • • • • • . • • • • . • . • • • • • . . • 1 9 . t a n a n arivei n . sp . M e t a n o t u m v e ry h o l l ow b eh i n d t h e s c u t e l l u m ( F i g . 1 1 2 6 C ) . • . • . 2 2 S egmen t 5 o f f r on t t a r s u s w i t h l a me l l a e l o c a t ed o n d i s t a l h a l f ( F i g . 1 1 2 8 ) • • • • • • . • . . • • • . . . • • • • 2 0 . a rm a tus n . sp . S egmen t 5 o f f r on t t a r su s w i t h l a me l l a e a l s o on p r ox i ma l t h i rd ( F i g s 1 1 29 , 1 1 30 ) • • • • • • • • • • • • • • • • • • • • • • • . • • • • • • • • • • • • • • • • • • • • 23 S egmen t 5 o f f r on t t a r s u s w i t h on e r o w o f l a me l l a e ( F i g . 1 1 2 9 ) ; t h o ra x a n d p r op od e u m b r own -d a rk . • • • • • • • . . • • • • • . . . • . • . . . . . • • • • • • • • • • • • • . . • . • . . . • • • • • • . . • • . . • • • . 2 1 . o k a h a ndjae n . sp . S e g men t 5 o f f r on t t a r s u s w i t h 2 r o w s o f l a me l l a e ( F i g . 1 1 3 0 ) ; t h o ra x a n d p r op od eu m b l a ck • • • • • 2 2 . john s i n . sp . M e s o p l eu ra w i t h a l a t e ra l p o i n t ( F i g . 1 1 3 1 A ) . . . . . . . . . • . . • . • • • • . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 3 . taylori n . sp . M e s op l eu ra w i t h ou t l a t e ra l p o i n t ( F i g . 1 1 3 1 B ) . • . • . . • • . . . • • • • 2 5 S c u t u m w i t h o u t t wo l a t e ra l p o i n t ed p r o m i n en c e s ( F i g . 1 1 3 1 B ) • • . • • • • • . • • • • • • • • • . • • • • • • • • • • • • • • • 2 4 . upem b a n u s n . sp . S c u t u m w i t h t wo l a t e ra l p o i n t ed p r o m i n en c e s ( F i g . 1 1 0 3 A , F , G ) .

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t h e s cu t e l l u m ( F i g . 1 1 2 6 0 ) • • • • • . • • • • . . . . . • • • • • . . • . • • • • • • . . . • • • • . . 2 5 . benoiti n . sp . Me t a n o t u m v e ry h o l l ow b eh i n d t h e s c u t e l l u m ( F i g . 1 1 2 6 A ) . . . . . 2 7 2 7 T h o rax a n d p r op o d eu m f u l l y b l a c k ; i n s ome s p e c i men s a p ex o f p r op od eu m r ed d i sh • • • • • . • . . . • • . . • • • • • • • • • • . . . . • • • • . . • • • . . • • • • • 2 8 T h o ra x a n d p r op od eu m r ed d i s h -d a rk o r r ed d i s h - l i g h t . . . . . . . . . . . 2 9 2 8 T h o ra x a n d p r op od eu m w i t h ou t s c u l p t u r e , sh i n y , smo o t h . . . . . . . . . . • • • • • • • • • • • . • • • • • • • • • • • • • • • • • • . 2 6 . troch a n tericus ( B en 0 i t ) T h o ra x a n d p r o p od e u m g ra n u l a t ed , d u l I • • • . • . • • • . . . . . • . . • • • • . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 7 . op a c u s n . sp . 2 9 T h o ra x a n d p r o p o d e u m r e d d i s h - I i g h t ; I eg s f u l l y t e s t a c e o u s r ed d i sh . . . . . . • • • • • • • . • • • • • • . . . . 2 8 . p a trizii B e n o i t T h o rax a n d p r o p o d e u m r e d d i s h -d a rk ; i n s ome s p e c i men s r ed d i sh l i g h t , b u t i n t h i s ca s e m i d a n d h i n d t i �:- I a e a n d f emo ra w i t h 2 6 Me t a n o t u m n o t

h o l l ow b eh i n d

1 590

Mem . A me r . E n ! .

b o t h a p ex e s b r own and

I ns! . ,

c en t ra l •

pa r t . 2 9 .

no.

37,

1 984

y e l l ow • . . . . . . . . turneri ( B e n o i t )

MALES

On l y the m a l e of G .

in cogn i t u s n . sp .

1 . Gon a top u s meridion a l is

i s k n ow n .

( Be n o i t ) n . com b .

Rhyn chogon a top u s meridion alis Benoi t 1 9 53b : 3 9 1 .

F E �. AL E : a p t e ro u s ; l en g t h 2 , 25-2 , 7 5 m m ; b l ac k , w i t h c l y p e u s a n d m a n d i b l es y e l l ow ; a n te n n a e a n d t a r s i b row n ; h e a d f l a t , s h i n y , h a i r l es s , g r a n u l a t e d ; p ro no t u m crossed b y a t r a n s v ers a l i m p ress i o n , sh i n y , w e a k l y g r a n u l a ted ; sc u t u m d u l l , g r a n u l a t e d , w i t h some l o n g i t u d i n a l k ee l s ; m e t a n o t u m d u l l , w e a k l y t ra n s v e rse l y s t r i a t e ; m e t a t h o r a x + p ropodeum s h i n y , g r a n u l a t ed ; p l eu r a a n d pos t e r i or s u rface of p ropod e u m n o t or w ea k l y t r a n sv erse l y s t r i a t e ; segmen t 1 of f ro n t t a rs u s a p p ro x i m a t e l y t w i c e a s l on g a s seg m e n t 4 ( 1 3 : 6 ) ; en l arged c l a w ( F i g . 1 099 ) w i t h o u t s u b a p i c a l t oo t h a n d l a me l l ae , w i t h a sm a l l t o o t h a t t h e e n d o f a l o n g i t u d i n a l f u rrow a n d 3-5 peg- l i k e h a i rs ; segmen t 5 o f fron t t a rs u s ( F i g . 1 099 ) w i t h i n n e r s i d e prox i m a l l y n o t serra t e , w i t h 2 r o w s of 1 3- 1 5 u s u a l l y v e r y sma l l l a me l l a e ; a p e x b ro a d e n e d , w i t h a g roup of 4 - 9 l a m e l l ae ; m a x i I l a r y p a l p i w i t h 3 - 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M AL E : u n k n o w n L O C U S T Y P I C U S : E s i r a ( M a d a g ascar ) T Y P I CAL MAT E R I AL : h o l o t y p e F ! a n d 1 p a r a t y pe F ! i n P ; 1 p a r a t y p e F ! i n TV . H OST S : u n k no w n D I ST R I B U T I ON : E t h i o p i a n reg i on : MADAGASCA R : E s i r a , P ! Rogez , T V ! Bek i l y , P ! T V ! B e h a r a , TV ! RUAN DA : K i se n y i , T V ! O L ! Z A I R E : R u t s h u r u ( K i w u ) , T V ! S O U T H A F R I C A : Somerset E a s t ( C a p e P ro v i nce ) , BO ! B M ! E s h o w e ( Z u l u l a n d ) , B M ! T r a n s v a a l N a t . Bo t a n i c a l G a rd e n s ( P reto r i a ) , AM '

2 . Gon a top u s longi u s c u l u s n .

sp.

F E MAL E : a p terou s ; l e n g t h 4 , 37 m m ; h e a d t e s t aceous-red d i s h ; a n t e n n a e b row n , w i t h segmen t s 1 -2 t e s t aceou s ; p ron o t u m a n d s c u t u m t e s t aceous red d i s h , w i t h s i des d a r k e n e d ; s c u t e l l u m not v i s i b l e ; meta t h o r a x + p ropodeum b l ac k , w i t h a p e x red d i s h a n d redd i s h n u a n ces on m e t a n o t u m ;

G o n a topod i n a e

Fig.

1 099

-

C h e l a o f Gon a top u s meridionalis ( Be n o i t ) ( t y p e )

1 59 1

Mem . A me r . E n t .

1 592

Fig.

I nst . ,

n o . 37,

1 984

1 1 00 - C he l a of Gon a top u s longi u s c u l u s n .

sp .

( ty p e )

p et i o l e b l ac k ; a b domen b ro w n-redd i sh ; l e g s tes taceous-red d i sh ; h e a d exca v a t e d , d u l l , g r a n u l a ted ; p rono t u m c rossed b y a t r a n s v e rs a l i m p ress i o n , d u l l , g r a n u l a ted ; scu t u m v e r y l o n g ( F i g . 1 1 22 B ) , n e a r l y 3-4

1 593

Gon a topod i n a e

t i mes a s l on g as broa d , d u l l , g r a n u l a ted ; scu te l l u m n o t v i s i b l e ; meso m e t a p l eu r a l s u t u re obso l e te ; meta thorax + p ropodeum d u l l , g r a n u l a ted ; p l e u r a a n d post e r i or su rface of p ropod eum t r a n sverse l y not s t r i a t e ; segmen t 1 of fron t t a r s u s s l i g h t l y shorter t h a n seg men t 4 ( 1 5 : 1 7 ) ; en l arged c l a w ( F i g . 1 1 00 ) w i t h o u t s u b a p i c a l too t h , w i t h a sma l l too t h a t t h e e n d of a l on g i t u d i n a l f u rrow a n d w i t h 7 peg- l i k e h a i r s ; segme n t 5 of f ro n t t a rsus ( F i g . 1 1 00 ) w i t h i n ner s i de prox i m a l l y ser r a t e , w i t h 1 row of 1 7 l a m e l l ae ; apex w i t h a g ro u p of 1 2 l ame l l ae ; m ax i l l a ry p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k no w n l O C U S T Y P I C US : P o r t S t . Joh n s ( Sou t h Af r i c a ) TYP I CAL MATER I AL : ho l ot y p e F ! i n BM HOSTS : u n k no w n D I STR I BU T I O N : E t h i op i a n reg i o n : S O U T H A F R I CA : Port St . Joh n s ( Pondo l a n d ) , B M !

3 . Gona top u s a u s tralis

( Ceba l l os ) n . com b .

Eucamp tonyx a u stralis Ceba I l os 1 936 :

57 .

F EMAL E : a p t e ro u s ; l en g t h 4 , 30-5 , 00 mm ; b l ack , w i t h m a n d i b l es , c l y p e u s a n d m a l a r sp ace ( more a l on g orb i t s ) y e l l ow ; a n te n n ae brow n , w i t h segme n t s 1 -2 y e l l ow ; l eg s y e l l ow , w i t h cox a e p a r t l y b l ack a n d c l u b s of femora p a r t l y b row n ; head exca v a ted , sh i n y , w i th weak or s t ro n g l o n g i t u d i n a I kee l s on fron s , g ra n u I a ted ; p ro n o t u m crossed b y a s t rong t ra n sv e rs a l i m p ress i o n , sh i n y , smoo t h , g r a n u l a t ed ; in some spec i mens d i sc w i t h o u t scu l p t u re ; scu t u m g r an u l a t e d , w i t h some l o n g i t u d i n a l k ee l s a n d l a t e ra l l y w i t h t w o po i n t s ; me t a t horax + p ropodeum smoo t h , g r a n u l a ted ; a n t e r i or s u rf ace w i t h o u t l on g i t u d i n a l k ee l s ; m e t a n ot u m , p l e u r a a n d pos ter i o r s u rface o f propodeum t r a n sverse l y s t r i a t e ; a b domen smoo t h , w i t hou t scu l p t u re ; segmen t 1 o f f ro n t t a rs u s shorter t h a n segme n t 4 ( 1 5 ; 1 9 ) ; en l arged c l a w ( F i g . 1 1 0 1 ) w i t h a sm a l l too t h at t h e e n d of a l on g i t u d i n a l f u rrow a n d w i t h 5-8 l a m e l l a e ; too t h l oc a ted very near a pex ; en l arged c l a w shorter ( 80% ) t h a n segme n t 5 of f ro n t t a rs u s ; segment 5 of front t a r s u s ( F i g . 1 1 0 1 ) w i t h i n ne r s i de p rox i m a l l y serra t e , w i t h t wo row s of a t l ea s t 1 7-22 l a me l l ae exten d i n g beyon d 0 , 5 l e n g t h of segmen t ; a pex w i t h a g ro u p of at l ea s t 1 3- 1 7 l a m e l l ae ; ma x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k no w n L O C U S TYP I CUS : Bek i I y ( M a d a g a scar ) T Y P I CAL MAT E R I AL : ho l ot y p e F ! i n P . HOSTS : u n k no w n D I ST R I B U T I O N : E t h i o p i a n reg i on : MADAGASCAR : Bek i l y , P ! TV ! L E SOTHO : M a m a t hes, BM ! SOUT H AFR I CA : M i l nerton ( Cape P ro v i n ce ) , Ol ! BM ! CONGO :

Mem . Amer .

1 594

Fig.

1 101

En! .

I n s! . ,

n o . 37 ,

1 984

- C h e l a of Gon a top u s a u s tralis ( Ce b a l l os ) ( t y p e )

1 595

Gon a t o pod i n a e

Fig.

1 1 0 2 - H e a d s of fem a l es of Gon a top u s pilosipes Brues

( ty pe ) ( A )

and

guigliae ( Ben o i t ) ( t y p e ) ( B )

M i twaba,

CA !

4 . Gona t op u s pilosipes Brues

Gon apotus pilosipes Brues 1 906a :

1 04 .

F E M AL E : a p terou s ; l en g t h 4 , 50-7 , 43 m m ; h e a d b row n ; a n t en n ae w i t h seg men t s 1 - 3 red d i s h ( o t h e r seg men t s m i s s i n g i n ex a m i n e d spec i me n s ) ;

body

b l ack , w i t h s i des a n d pos t e r i or s u rface of p rono t u m red ; t a r s i l i g h t ; h e a d ( F i g . 1 1 02 A ) e x ca v a t e d , sh i n y g r a n u l a t ed , w i t h s o m e k ee l s on f ro n s ; occ i p u t h a i ry ; p rono t u m crossed b y a t ra n s v e rsa l i m p re ss i on , sh i n y , w i t h o u t sc u l p t u r e , w i t h some k ee l s on s i d e s , w i t h l o n g h a i rs ; scu t u m s h i n y , w i t h l o n g h a i rs , w i t h 2 l a t era l a n d poi n ted k ee l s ( F i g . 1 1 03 B ) ; a n t e r i or s u rface of p ropodeum w i t h n u merous l o n ,::' l t u d i n a l k ee l s ; p l e u r a a n d pos t e r i o r s u rf a c e o f p ropod e u m t r a n s v erse I y s t r i a t e ; m e t a t horax + p ropod e u m w i t h l o n g h a i rs ; meso-p l e u r a l s u t u re obso l e t e ; fore t a rs a l segmen t s i n fo l l ow i n g p roport i on s : 24 : 4 : 8 : 23 : 3 5 ; en l arged c l aw ( F i g . 1 1 04 A ) w i t h a sm a l l too t h a t t h e e n d of a l on g i t u d i n a l f u rrow a n d w i t h 8 l a me l l a e ; tooth l oc a t e d v e r y near apex ; e n l a rged c l a w s h o r t e r ( 8 5% ) t h a n seg men t 5 o f f ron t t a rs u s ; seg m e n t 5 o f fron t t a rs u s ( F i g . 1 1 04 A ) w i t h i n ner s i de p rox i m a l l y serr a t e , w i t h t w o rows of 1 7 l a me l l ae ( s hortest row not e x t en d i n g b e y on d 0 , 5 l e n g t h of segmen t ) ; a p ex not b e n t at r i g h t a n g l e , w i t h a g ro u p o f 1 9 l a me l l a e ; m a x i l l ary p a l p i w i t h 6 seg men t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k now n LOCUS T Y P I CUS :

Bo t h a v i l l e ( So u t h Af r i c a )

1 596

Mem . Amer . E n t .

I n st . ,

no. 37,

1 984

� B M� /\

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F i g . 1 1 03 - Scu t u m , scu te l l u m a n d met a n o t u m of fem a l es of Gon a top u s benoiti n . sp . ( A ) , pilosipes B rues ( C ) , communis n . sp . ( D ) , l u teipes ( Be n o i t ) ( E ) , p a trizii Beno i t ( F ) , troch a n teri c u s ( Be n o i t ) ( G ) a n d upem b a n u s n . sp . ( H ) .

TYP I CAL MAT E R I A L : h o l o t y p e F ! i n M L . HOSTS : u n k n o w n D I ST R I B U T I O N : E t h i op i a n reg ion : SOU T H A F R I CA : Both a v i l l e ( Or a n g e ) , Z I MBAB W E : B u l a w a y o , B M !

ML!

5 . Gon a top u s guigl i a e ( Be n o i t ) n . com b .

Digo n a top u s guigliae Benoi t 1 95 1 a : 298 . Ch a lcogon a top u s guigliae ( Be n o i t ) : O l m i a n d C u rrado 1 976a :

1 69 .

F E M AL E : a p terou s ; l e n g t h 5 , 26 mm ; head b l ack , w i t h c l y p e u s , f ro n t p a r t o f vertex a n d ma n d i b l es y e l l ow ; a n te n n ae b l ack , w i t h segme n t s 1 -2 y e l l ow ; t horax , p ropodeu m a n d a b domen b l ack , except for w e a k b ro w n n u a nces on abdome n ; I egs y e l l ow , w i t h brow n spots o n cox a e , fore t roc h a n t ers, c l u bs of femor a , exte rn a l face of fore t i b i ae ; a n ten n a l seg men ts in fol l ow i n g p roport i on s : 1 2 : 1 2 : 3 3 : 2 7 : 2 1 : 1 9 : 1 5 : 1 5 : 1 4 : 1 8 ; head ( F i g . 1 1 02 B ) exc a v a ted ; frons w i t h n u merous l on g i t u d i n a l k ee l s ; POL = 3 ; O l = 4 ; OOl = 1 5 ; p ro n o t u m crossed b y a t r a n s versa l i m p ress i o n , sh i n y , smoo t h , h a i ry ; scu t u m a l u t aceou s , l a tera l l y w i t h 2 p o i n t s ; me t a t h orax + propodeum sh i n y , smoot h ; metanot u m , p l e u ra a n d poster ior s u rface of p ropodeum t r a n sverse l y s t r i a t e ; e n l a rged c l a w ( F i g . 1 1 05 ) w i t h o u t s u b a p i c a l too t h , w i t h a sma l l

Gon a topod i n a e

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Che l a of Gon a topus pilosipes Brues ( t y p e ) ( A ) ; m a x i l l a ry and l a b i a l ( C ) pa l p i .

1 597

(B)

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1 1 05 - C h e l a of Gon a top u s guigliae

1 984

( Beno i t ) ( t y p e )

tooth a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d 7 p e g - l i k e h a i rs ; seg m e n t 5 of f ro n t t a rs u s ( F i g . 1 1 05 ) w i t h i n n e r s i d e p rox i m a l l y serr a t e , w i t h t w o rows of 2 ( p rox i m a l ) + 26 l a m e l l ae ; apex w i t h a grou p of 1 6- 1 8 l a me l l ae ; m a x i I l a r y p a l p i w i t h 6 segmen t s ; s p u rs 1 , 0 , l .

l a b i a l p a l p i w i t h 3 s e g me n t s ;

tibial

MALE : u n k nown L O C U S T Y P I C US : B u s s u B u s o g a ( U g a n d a ) p aratype F! i n TV . T Y P I CA L M A T E R I AL : h o l o t y p e F ! i n G E ; H O ST S : u n k no w n D I ST R I B U T I ON : E t h i o p i a n reg i on : UGAN D A : B u s s u - B u so g a , G E ' TV !

6 . Gon ap o tus seyrigi ( C eb a l l os )

n . com b .

Trichogon a top u s seyrigi Ceb a l l os 1 936 : 59 . E u c a mp tonyx seyrigi ( C e b a l l os ) : Beno i t 1 953b : 386 .

F E MAL E :

a p terous ;

l en g t h 6 , 0 m m ;

b l ac k ,

w i t h m a n d i b l es a n d c l y p e u s

y e l l ow ; a n t e n n a e b l ac k ; l eg s b l ac k , w i t h m i d a n d h i n d t i b i ae , t roch a n te r s , m i d a n d h i n d t a r s i b row n i s h ; a n t e n n a l seg m e n t s i n fo l l o w i n g p ropor t i on s : 1 1 : 9 : 26 : 2 3 : 1 5 : 1 5 : 1 2 : 1 2 : 9 : 1 4 ; h e a d exc a v a t e d , d u l l ; f ro n s h a i r l es s ; occ i p u t h a i r y ; frons a n d v e r t e x w i t h som e l o n g i t u d i n a l k ee l s ; p r o n o t u m crossed b y a t r a n svers a l i m p ress i o n , s h i n y , smoot h ; a n te r i o r s u rface of d i sc a l u t aceou s ; c o l l ar a n d s i d e s of p ron o t u m h a i r y ; scu t u m w i t h some k ee l s ; sc u t e l l u m s l i g h t l y v i s i b l e ; m e t a t h orax + p ro p o d e u m h a i ry ; p l e u r a a n d poster i o r su rface of p ropodeum t ra n s v e rse l y s t r i a t e ;

Gon a topod i n a e

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1 599

C h e l a of Gon a topus seyrigi ( C eb a l l os ) ( t y pe )

m e t a n o t u m a n d a n te r i o r s u rf a ce of p ropodeum w i t h l on g i t u d i n a l k ee l s ; a b domen a n d I e g s v e r y h a i r y ; seg m e n t 1 of f ro n t t a r s u s m u c h s h o r t e r t h a n seg men t 4 ( 20 : 32 ) ; en l a r g e d c l a w ( F i g . 1 1 06 ) w i t h 1 0 l a me l l ae , w i t h o u t s u b a p i c a l too t h a n d s m a l l t oo t h a t t he e n d of a l on g i t u d i n a l f u r row ; seg men t 5 of f ro n t t a rs u s ( F i g . 1 1 06 ) w i t h 1 row of 25 l a me l l a e , apex w i t h a g r o u p of a t l ea s t 1 6 l a me l l ae ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 seg men t s ( accord i n g t o C e b a l l os 1 936 ) ; i n h o l o t y p e t h e p a l p i are m i s s i n g ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n known

M e m . A me r . E n ! .

1 600

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n o . 37,

1 984

1 1 07 - C h e l a of Gon a top u s a n k ara trus ( Ce b a l l os ) ( t y p e )

LOCUS T Y P I C U S : M a d a g a s c a r T Y P I CAL M A T E R I AL : h o l o t y p e F ! i n P . H O ST S : u n k n o w n D I ST R I BU T I ON : E t h i op i a n reg i on : MADAGASC AR ,

P!

1 60 1

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7 . Gon a t op u s ankara trus ( C eb a l l o s ) n . com b .

Chalcogon a top u s a n kara trus C e b a I l os 1 936 : 55 . Ma decagon a top u s a n k a r a trus ( C e b a l l os ) : Beno i t 1 953 b : 392 .

F E M AL E : a p terou s ; l e n g t h 3 , 2 5 m m ; b l ack , w i t h m a n d i b l es , c l y pe u s , t a r s i a n d a n t en n a l seg men t s 5 - 6 b ro w n ; a n ten n a l segmen t s i n fo l l o w i n g p ro p or t i on s : 8 : 4 : 1 1 : 5 : 5 : 5 : 4 : 4 : 4 : 6 ; h e a d f l a t , d u l l , g r a n u l a ted ; t e m p l es p rom i n en t ; p ro n o t u m crossed by a t ra n sversa I s t ro n g i m p ress i o n , s h i n y , w i t h o u t sc u l p t u re , w i t h some weak l o n g i t u d i n a l s t r i ae on d i sc ; sc u t u m s h i n y , w i t h some weak l o n g i t u d i n a l k ee l s ; s c u t e l l u m smoo t h , sm a l l , s l i g h t l y v i s i b l e ; scu t u m w i t h 2 l a tera l po i n t s ; met a n o t u m very redu ced , smoo t h , n o t h o l l ow beh i n d t h e sc u te l l u m ; m e t a t h orax + p ropodeum d u l l , w i t h n u merous w e a k l on g i t u d i n a l s t r i ae on a n t er i o r s u rf a ce , d i sc a n d p o s t e r i or s u rf a ce ; p o s t e r i o r su rface w i t h a f e w s t ro n g t r a n svers a l k ee l s n e a r p e t i o l e ; meso-me t a p l e u r a l s u t u re obso l e t e ; meso p l eu r a a n d me t a p l e u ra w i t h n u merous weak t r a n s v ers a l s t r i a e ; fore t a rs a l segme n t s i n fo l l ow i n g p roport i o n s : 1 0 : 2 : 5 : 1 3 : 2 1 ; seg men t 3 o f fron t t a r s u s p ro d u ced i n to a h ook ; e n l arged c l a w ( F i g . 1 1 07 ) w i t h o u t s u b a p i c a l t oo t h , w i t h a s m a l l t oo t h a t t h e e n d of a l on g i t u d i n a l fu rrow a n d 5 p e g - l i ke h a i rs ; segmen t 5 of f ro n t t a rs u s ( F i g . 1 1 07 ) w i t h i n ner s i ci � p rox i m a l l y n o t ser r a t e , w i t h 2 rows of 2 ( p rox i m a l ) + 22 l a me l l a e ; a p e x w i t h a g rou p of 6 l a me l l ae ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 seg men t s ( accord i n g to Be n o i t 1 953b l a b i a l p a l p i w i t h 3 segme n t s ; accord i n g t o C e b a l l os 1 936 m a x i l l a r y p a l p i 6-se g m e n t e d ) ; in h o l o t y p e t h e p a l p i are m i s s i n g ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : A n k a r a t ra ( M a d a g a s c a r ) TYP I CA L M A T E R I A L : h o l o t y p e F ! i n M D . HOSTS : u n k no w n D I ST R I B U T I ON :

E t h i op i a n reg i o n :

MADAGASCAR :

Ankaratra,

MD !

8 . Gon a top u s capen sis Brues

Gon a top u s capen sis B r u es 1 906a :

1 05 .

F E MA L E : a p terou s ; l e n g t h 5 , 1 2- 5 , 62 m m ; head b row n ; a n te n n ae redd i s h ; t h ora x , p e t i o l e a n d a b domen b l ac k ; l e g s red d i s h , w i t h Coxae b l ack a n d c l u b s of femora b row n ; h e a d ex c a v a t e d , sh i n y , g r a n u l a t e d , h a i r l ess ; p ro n o t u m crossed b y a t r a n sversa l i m p ress i on , g r a n u l a ted , h a i r l es s ; scu t u m h a i r l es s , w i t h 2 l a tera l po i n t ed kee l s ; m e t a n o t u m , p l e u r a a n d poster i o r su rface o f p ropodeum t r a n s v erse l y s t r i a t e ; a n t e r i o r su rface of

Mem . A m e r .

1 602

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1 9 84

C h e l a of Gon a top u s capen s i s B r u e s ( t y p e )

propo d e u m w i t h l on g i t u d i n a l s t r i a e ; meso-me t a p l eu r a l s u t u re o b so l e t e ; fore t a r s a l seg men t s in fo l l o w i n g p rop o r t i on s : 1 7 : 4 : 7: 30 : 42 ; en l a rg ed c l a w ( F i g . 1 1 08 ) w i t h a s m a l l tooth at t h e e n d of a l on g i t u d i n a l f u rrow a n d w i t h 8 s m a l l l a m e l l ae , i n a d d i t i o n to 1 b r i s t l e ; too t h very f a r f rom t h e apex ; a p e x v e r y s l en d e r ; segme n t 5 of f ron t t a r s u s ( F i g . 1 1 08 ) w i t h i n n e r s i d e p rox i m a l l y serra t e , w i t h t w o r o w s of 1 ( p rox i m a l ) + 23-25 l a me l l ae ;

1 603

Gon a topod i n a e

F i g . 1 1 09 - H e a d s of f e m a l es of Gon a top u s mosseli n . s p . ( t y p e ) ( A ) , comm unis n . s p . ( p a r a t y pe ) ( B ) a n d ru brithora x ( Be n o i t ) ( t y p e ) ( C ) .

a p ex very p rom i n e n t a n d b e n t a t r i g h t a n g l e , 1 0 l a me l l ae ; segmen t s ;

m a x i l l a r y p a l p i w i t h 6 seg men t s ;

w i t h a g ro u p of a t l ea s t 6l a b i a l p a l p i w i th 3

t i b i a l spurs 1 , 0, 1 .

MALE : u n k n o w n L O C U S TYP I C U S : W i l l owmore ( So u t h A f r i c a ) T Y P I CAL M A T E R I AL : h o l o t y p e F ! i n ML . HOSTS : u n k n o w n D I STR I B U T I ON : E t h i op i a n reg i on : SO U T H A F R I CA : Worcester ( C a p e P ro v i n ce ) , BM ! L i t t l e K a rroo ( Ce re s , C a pe P rov i n ce ) , BM ! W i l l o w more ( C a p e P rov i nce ) , ML !

9 . Gona top u s mosseli n .

sp.

F E MALE : a p terou s ; l en g t h 3 , 1 2 mm ; t e s t aceo u s - red d i s h , w i t h pe t i o l e b l ack ; h e a d ( F i g . 1 1 09 A ) a l mo s t f l a t , s h i n y , w i t h o u t scu l p t u re or w e a k l y g r a n u l a ted ; temp l es very p ro m i n e n t ; pron o t u m crossed b y a t r a n s v e rs a l i m p re s s i o n , s h i n y , w i t h o u t sc u l p t u re ; scu t u m s h i n y , w i t h some l on g i t u d i n a l k ee l s ; m e t a n o t u m s h i n y , not h o l l ow beh i n d t h e sc u te l l u m , t r a n sverse l y s t r i a t e ; m e t a t h o r a x + p ropodeum s h i n y , smoo t h , w i t h o u t scu l p t u re ; mesop l e u ra t r a n s v e rse l y not s t r i a t e ; pos ter i o r s u rf ace of p ropode u m a n d m e t a p l e u ra t r a n s v e r se l y s t r i a t e ; meso-me t a p l e u r a l s u t u re o b so l e te ; seg m e n t 1 of f ro n t t a r s u s s l i g h t l y s h o r t e r t h a n segme n t 4 ( 1 0 : 1 1 ) ; en l a r g e d c l a w ( F i g . 1 1 1 0 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l tooth a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d w i t h 6 l a m e l l a e , i n a d d i t i o n to 1 b r i s t l e ; seg m e n t 5 o f f ro n t t a rs u s ( F i g . 1 1 1 0 ) w i t h i n n e r s i de p rox i m a l l y serra t e , w i t h 1

1 604

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1 984

C h e l a of Gon a top u s mosseli n . sp .

( ty p e )

1 605

G o n a topod i n ae

A

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- Meso t h o r a x , m e t a t horax a n d p ropodeum of fema l es of Gon a top u s hya linus n . s p . ( t y p e ) ( A ) a n d ericeti n . s p . ( t y pe ) ( B ) .

row of 1 6 l a m e l l ae ; a p ex w i t h a g ro u p of 1 5 l a me l l ae ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k no w n LOCUS T Y P I C U S : Mosse l B a y ( So u t h Af r i c a ) T Y P I C A L M A T E R I A L : h o l o t y pe F ! i n B M . H O ST S : u n k no w n D I ST R I B U T I ON : E t h i o p i a n reg i o n : S O U T H A F R I CA : P rov i nce ) , B M !

Mosse l

1 0 . Gon a top u s rubrithorax ( Be n o i t )

Bay

( Cape

n . comb .

Neogon a topus ru brithorax Beno i t 1 953b : 393 .

F E M A L E : a p t erou s ; l e n g t h 3 , 0 m m ; y e l l ow - red d i s h , w i t h head weak l y b ro w n i s h , a n t e n n a l segmen t s 3- 1 0 b row n i sh , p e t i o l e a n d a b domen b l ack b row n , c l u b s of fore femora b row n ; a n t en n a l segme n t s in fo l l o w i n g p ropor t i o n s : 4 : 4 : 1 1 : 6 : 4 , 5 : 5 : 4 : 4 : 4 : 5 , 5 ; head ( F i g . 1 1 09 C ) smoo t h , exc a v a t e d , s h i n y , h a i r l es s , w i t h o u t sc u l p t u re ; p ro n o t u m d u l l , crossed b y a t r a n s v e rs a l i mp ress i o n ; sc u t u m d u l l , w i t h some l on g i t u d i n a l k ee l s ; m e t a n o t u m , p l e u ra a n d p o s t e r i o r s u rf a ce of p ropodeum t ra n sve rse l y s t r i a t e ; a n t e r i or s u rface o f p ropode u m smoo t h ; c h e l ae m i s s i n g i n h o l o t y p e ( on l y k no w n s p ec i men ) ; m a x i l l a r y p a l p i w i t h 4 s e g men t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n ow n LOCUS T Y P I C U S : Bek i l y ( M a d a g asca r ) T Y P I C AL M A T E R I AL : ho l o t y pe F ! i n P . HOST S : u n k n ow n

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E t h i o p i a n reg i on :

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( type)

P!

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F E M AL E : a p t e ro u s ; l e n g t h 2 , 6 2 m m ; y e l l o w - t e s t aceo u s , very l i g h t , w i t h p e t i o l e b l ac k ; h e a d exc a v a t ed , sh i n y , smoo t h , w i t h o u t scu l p t u r e , except for occ i p u t g r a n u l a t ed ; p ron o t u m c rossed b y a t r a n svers a l i m p ress i o n , sh i n y , w e a k l y g r a n u l a ted ; m e t a n o t u m t r a n sverse l y s t r i a t e , n o t h o l l ow beh i n d t h e sc u te l l u m ; meso-m e t a p l eu ra l s u t u re o b so l et e ; m e t a t h o r a x + p ropod e u m ( F i g . 1 1 1 1 A ) w i t h o u t scu l p t u r e , s h i n y , smoo t h ; p l e u ra a n d poster i o r s u rf a ce of p ro p o d e u m w e a k l y t r a n s v e r se l y s t r i a t e ; s i de s o f m e t a no t u m rou n d ed ; segm e n t 1 o f fro n t t a rs u s a s l o n g as segmen t 4 ; en l a r ged c l a w ( F i g . 1 1 1 2 ) w i t h o u t s u b a p i c a l t oo t h , w i t h a sm a l l too t h a t t h e e n d of a l on g i t u d i n a l f u rrow a n d w i t h 4 b r i s t l es ; seg m e n t 5 of f ro n t t a rsus ( F i g . 1 1 1 2 ) w i t h i n n e r s i d e p rox i m a l l y n o t serr a t e , w i t h 2 rows of 2 ( p rox i m a l ) + 1 4 l a m e l l ae ; apex w i t h a g r o u p of 1 2 l a me l l ae ; m a x i I l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 seg men t s ; t i b i a l s p u r s 1 , 0 , 1 . M AL E : u n k n o w n L O C U S T Y P I C U S : P o r t S t . Joh n s ( So u t h Af r i c a ) TYP I CAL M A T E R I AL : h o l o t y p e F ! i n B M . H O ST S : u n k no w n D I ST R I B U T I O N : E t h i op i a n reg i o n : S O U T H A F R I CA : P o r t St . Joh n s ( Po n do l a n d ) , B M !

1 2 . Gon a top u s ridens n . s p .

F E M A L E : a p t e ro u s ; l e n g t h 2 , 50 m m ; h e a d t e s t aceous-red d i s h ; a n t e n n ae b row n , w i t h seg men t s 1 -2 a n d p a r t l y 3 y e l l o w , s e g me n t s 9- 1 0 a n d p a r t l y 8 w h i t e ; p rono t u m b row n-redd i s h ; scu t u m y e l l ow , w i t h a n te r i o r m a r g i n b row n ; scu t e l l u m a n d m e t a t h o r a x + p ro p o d e u m b l ac k , w i t h a p ex of p ro p o d e u m redd i sh ; p e t i o l e b l ack ; a b domen b row n-red d i s h ; l e g s test aceou s ;

Mem . Am e r .

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s l i g h t l y p rom i n e n t ; p rono t u m crossed by a t ra n s vers a l i m p ress i o n , sh i n y , smoo t h , h a i ry , w i t h o u t scu l p t u re ; scu t u m s h i n y , w i t h som e l o n g i t u d i n a l k ee l s ; m e t a n o t u m , p l e u r a a n d p o s te r i or s u rface o f p ropodeum t r a n s v e rse l y s t r i a t e ; m e t a t horax + propod e u m w i t ho u t sc u l p t u re , smoo t h , s h i n y ; meso m et a p l e u r a l s u t u re o b so l e t e ; segmen t 1 of f ro n t t a rs u s s l i g h t l y shorter t h a n seg m e n t 4 ( 1 0 : 1 1 ) ; en l a r g e d c l a w ( F i g . 1 1 1 4 ) w i t h o u t s u b a p i c a l t oo t h , w i t h a s m a l l tooth a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d w i t h 7 peg - l i k e h a i rs ; segmen t 5 of f ro n t t a rs u s w i t h i n n e r s i de p rox i m a l l y n o t serra t e , w i t h 1 row o f 1 9 l a me l l ae ; apex w i t h a g ro u p o f 1 2 l a me l l a e ; m ax i l l a r y

1 609

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1 1 1 5 - C he l a o f Gon a top u s comm unis n .

p a l p i w i t h 5 segmen t s ;

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sp .

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t i b i a l s p u rs 1 , 0 , 1 .

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1 984

1 1 1 6 - H e a d s of fe m a l es of Gon a top u s i u teipes ( Be n o i t ) ( t y p e ) ( A ) a n d incogn itus n . sp . ( t y pe ) ( B ) .

L O C U S T Y P I C U S : Mosse l B a y ( So u t h A f r i c a ) TYP I C AL M A T E R I AL : h o l o t y p e F ! i n BM . HOSTS : u n k n o w n D I ST R I B U T I O N : E t h iop i a n reg i o n : S O U T H A F R I CA : P ro v i nce ) , B M !

Mosse l

Bay

( Ca p e

1 3 . Gon a top us comm u n i s n . sp .

F E M AL E : a p tero u s ; l e n g t h 2 , 87-3 , 1 2 m m ; h e a d f u l l y t e s t aceous or brow n , w i t h occ i p u t , f ro n t p a r t o f v e r t e x , c l y p e u s a n d m a n d i b l e s y e l l ow ; a n t en n ae y e l l o w , d i s t a l l y w e a k l y b r o w n i s h ; p ro no t u m y e l l o w or brow n ; scu t u m y e l l ow , w i t h a n te r i o r m a rg i n b l ack o r b row n ; s c u te l l u m a n d m e t a t ho r a x + p ro p o d e u m y e l l o w or redd i s h ; i n some s p ec i m e n s w i t h b l ack or b r o w n n.u a n ce s o n p l e u r a a n d m e t a not u m ; p e t i o l e b l ack ; a b domen b r o w n - r ed d i s h o r y e l l ow ; l eg s y e l l o w ; i n some spec i m e n s c l u b s of fore femora b row n i s h ; h e a d ( F i g . 1 1 09 B ) exc a v a t ed , sh i n y , smoo t h , w i t h o u t scu l p t u re ; p rono t u m crossed b y a t r a n s v e rs a l i m p res s i on , sh i n y , s m oo t h , h a i r l es s , w i t h o u t scu l p t u re ; sc u t u m d u l l , g r a n u l a t ed , w i t h o u t l a t e r a l po i n t s ( F i g . 1 1 03 D ) ; m e t a n o t u m w e a k l y r u gose ; m e t a t ho r a x + p ro p o d e u m s h i n y , smoo t h , w i t h o u t scu l p t u re ; p l e u r a a n d p o s t e r i or s u rface of p ro p o d e u m t ra n s verse l y s t r i a t e ; meso-me t a p l eu r a l s u t u re o b so l e t e ; fore t a r s a l seg men t s i n fo l l o w i n g p ro p o r t i o n s : 1 2 : 2 : 4 : 1 2 : 1 8 ; en l a r g e d c l a w ( F i g . 1 1 1 5 ) w i t ho u t s u b a p i c a l too t h , w i t h a sm a l l tooth a t t h e e n d o f a l o n g i t u d i n a l f u rrow a n d w i t h 3-5 p e g - l i k e h a i rs or b r i s t l es ; seg men t 5 of f ro n t t a r s u s ( F i g . 1 1 1 5 ) w i t h i n n e r s i de p rox i m a l l y s e r r a t e or n o t s e r r a t e , w i t h 1 row of 1 8-26 l a me l l a e ; a p e x n o t ben t a t r i g h t a n g l e , w i t h a g r o u p of 9- 1 0 l a m e l l ae ; m a x i l l a r y p a l p i w i t h 4- 5 segmen t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k no w n L O C U S T Y P I C U S : K i r i b a t a ( Z a i re ) TYP I CAL M A T E R I A L : ho l o t y p e F ! i n T V ; 3 p a r a t y p es F F ! p a r a t y p es F F ! i n O L . HOST S : u n k n o w n

i n BM ;

2

1 61 1

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1 1 1 7 - C h e l a of Gon a top u s l u teipes ( Ben o i t ) ( t y p e )

D I ST R I B U T I O N : E t h i o p i a n reg i o n : Z A I R E : K i r i b a t a ( M . R u w enzo r i ) , T V ! S O U T H A F R I C A : Mos s e l B a y ( C a p e Prov i n ce ) , O L ! B M ! Somerset E a s t ( C a p e P rov i n ce ) ,

BM !

TANZAN I A :

14.

Z a n z i b a r ( Z anz i b a r

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Gon a topus l u teipes ( Be n o i t )

Pla tygon a top u s l u teipes Ben o i t 1 9 5 1 d :

n .

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19.

F E M A L E : a p terou s ; l en g t h 3 , 1 2 m m ; h e a d b ro w n , w i t h f ro n t p a r t of v e r t e x , c l y p e u s a n d m a n d i b l es y e l l ow ; a n te n n a e y e l l ow , w i t h se g m e n t s 4 - 1 0 b ro w n ; t ho r a x a n d p ropodeu m b l ac k ; a b domen b l ac k , w i t h some b ro w n n u ances ; l eg s y e l l ow , w i t h c l u bs of fore femora b row n i s h a n d m i d a n d h i n d t i b i ae w i t h b o t h a pexes b row n i s h ( a s i n A crodontochelys C u r r a d o a n d i n some Gon a top u s ) ; head ( F i g . 1 1 1 6 A ) s h i n y , a l mo s t f l a t , w ea k l y e x c a v a t e d , g r a n u l a t ed ; pron o t u m c rossed b y a t r a n s versa l i m p ress i o n , g ra n u l a t e d , s h i n y ; sc u t u m s t ron g l y g r a n u l a t e d , d u l l , w i t h o u t l a t e r a l po i n t s ( F i g . 1 1 03 E ) ; m e t a n o t u m d u l l , g r a n u l a t e d , w i t h some t r a n s v e r s a l s t r i a e ; m e t a t h o r a x + p ro p o d e u m d u l l , g r a n u l a te d ; p l e u r a a n d p o s te r i o r s u rf ace of p ropode u m t r a n s v e r se l y n o t s t r i a t e ; meso-met a p l eu r a l s u t u re o b so l e t e ; fore t a r s a l segmen t s i n fo l l ow i n g p ropor t i o n s : 1 3 : 2 : 2 , 5 : 1 0 : 1 5 ; en l a r ged c l a w ( F i g . 1 1 1 7 ) w i t hou t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d of a l on g i t u d i n a l f u rrow a n d w i t hou t or w i t h some b r i s t l es or p eg - I i k e h a i r s or l a m e l l ae ; segmen t 5 of f ron t t a r s u s w i t h i n n e r s i d e p rox i m a l l y s e r r a t e o r not se r r a t e , w i t h 1 on l y l o n g l a me l l a , i n a d d i t i on to 4 v e r y s h o r t l a me l l ae or peg - l i k e h a i rs a n d 1 row of b r i s t l es ( i n h o l o t y p e : f i g . 1 1 1 7 ) ; w i t h 1

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1612

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1 1 1 8 - Segm e n t 5 o f fron t t a rs u s o f f e m a l e o f Gon a top u s l u teipes ( Be n o i t ) from O k a h a n d j a ( So u t h West Af r i c a )

row of 20 l a me l l ae ( F i g . 1 1 1 8 ) i n a spec i men f ro m O k a h a n dj a ; a pex p rom i n e n t , w i t h a g ro u p o f 4-8 l a m e l l a e ; m ax i l l a ry p a l p i w i t h 4-5 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l spurs 1 , 0 , 1 . M A L E : u n k no w n L O C U S T Y P I C U S : O l g a s a l i c ( Ken y a ) TYP I CAL MAT E R I AL : ho l o t y pe F ! i n T V . HOST S : u n k no w n D I ST R I B U T I ON : E t h i o p i a n reg i o n : A F R I CA : O k a h a n dj a , BM !

K E NYA : O l g a s a l i c ,

T V ! SOU T H W E S T

1 5 . Gon a top u s bekilya n u s ( Be n o i t ) n . c om b .

Neogon a top u s bekily a n u s Ben o i t 1 95 3 b : 394 .

F E M A L E : a p t e ro u s ; l e n g t h 3 , 0 m m ; b l ac k , w i t h c l y pe u s a n d m a n d i b l es y e l l o w ; a n t e n n a e b row n , w i t h segmen t s 1 -2 or o n l y 1 y e l l o w ; l eg s y e l l o w , w i t h c o x a e a n d c l u b s of femora b l ac k -b ro w n ; head exca v a t e d , s h i n y , h a i r l es s , w i t ho u t scu l p t u re , excep t for f r o n s a n d occ i p u t g r a n u l a t e d ; p ro n o t u r, -crossed by a t r a n svers a l i m p ress i o n , s h i n y , smoo t h , w i thou t scu l p t u r e ; sc u t u m s h i n y , w i t h some l o n g i t u d i n a l k ee l s , w i t h o u t l a t era l po i n t s ( F i g . 1 1 0 3 C ) ; m e t a n o t u m t ra n sv erse l y s t r i a t e ; met a t horax + p ro p o d e u m sh i n y , w i t h o u t sc u l p t u r e ; p l e u ra a n d pos t e r i o r s u rf a c e of p ro p o d e u m t r a n s v e rse l y s t r i a t e ; meso-me t a p l eu r a l su t u r e o b so l e t e ; se g m e n t of f ro n t t a rs u s s h o r t e r t h a n segmen t 4 ( 1 1 : 1 4 ) ; en l a r g e d c l a w ( F i g . 1 1 1 9 ) w i t hou t s u b a p i c a l toot h , w i t h a s m a l l t oo t h a t t he e n d of a l o n g i t u d i n a l f u rrow a n d 7-9 peg - l i k e h a i rs ; s e g me n t 5 o f fron t t a rs u s ( F i g . 1 1 1 9 ) w i t h i n n e r s i de p rox i m a l l y serr a t e , w i t h 1 row of 1 7-24 l a me l l a e ; a pe x not b e n t a t r i g h t a n g l e , w i t h a g ro u p of 8- 1 0 l a m e l l a e ; m ax i l l a ry p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 .

1613

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1 1 1 9 - C h e l a of Gon a top u s bekilya n u s

( Ben o i t ) ( t y p e )

M A L E : u n k no w n L O C U S T Y P I C U S : Bek i l y ( M a d a g a s c a r ) T Y P I CA L M A T E R I AL : h o l o t y pe F ! a n d 7 p a r a t y p es F F ! i n P ; 4 p a r a t y pes i n TV . H O ST S : u n k n o w n D I ST R I B U T I ON : E t h i op i a n reg i o n : MADAGASCAR : Bek i l y , P ! T V ! Rogez , T V !

1 6 . Gon a top u s incogn i t u s n . sp .

F E M AL E : a p terou s ; l e n g t h 2 , 0 m m ; h e a d b row n - b l ac k , w i t h c l y p e u s , m a l a r space a n d m a n d i b l e s y e l l ow ; a n t e n n a e b ro w n , w i t h segmen t s 1 -3 y e l l ow ; t h o r a x , p ropodeum a n d a b domen b l ac k ; l e g s y e l l ow - b row n , w i t h c o x a e , c l u b s o f femora a n d t i b i a e b row n i sh ; a n t e n n ae d i s t a l l y t h i ch ened ; a n t e n n a l

Mem . A me r .

1 61 4

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1 1 20 - C h e l a of Gon a top us incogn itus n . sp .

( type )

seg men t s i n fo l l ow i n g proport i o n s : 6 : 4 , 5 : 8 , 5 : 5 : 5 : 5 : 4 : 4 : 4 : 6 ; h e a d ( F i g . 1 1 1 6 B ) very exc av a t e d , s h i n y , w i t h o u t scu l p t u re , excep t f o r f r o n s a n d occ i p u t g r a n u l a ted ; p r o n o t u m c r o s s e d b y a t r a n s vers a l i m p ress i o n , s h i n y , h a i r l es s , g r a n u l a t ed ; sc u t u m d u l l , w i t h some l on g i t u d i n a l k ee l s ; m e t a n o t u m t r a n sve rse l y s t r i a t e ; m e t a t h or a x + p ropodeum d u I I , g r a n u I a ted ; a n t e r i or s u rface w i t h w e a k l o n g i t u d i n a l s t r i ae ; p l e u r a a n d p o s t e r i o r s u rface of p ro p o d e u m t ra n sv erse l y s t r i a t e ; meso-met a p l e u r a l s u t u re o b so l e t e ; r a re l y v i s i b l e o n l y a t r a c k of s u t u re ; fore t a r s a l s e g me n t s i n f o l l ow i n g p ro p or t i o n s : 9 : 2 : 3 : 9 : 1 5 ; en l arged c l a w ( F i g . 1 1 20 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l t oo t h n e a r a p e x a t t h e e n d o f a l on g i t u d i n a l f u rrow a n d 5-6 peg - l i k e h a i rs ; s e g m e n t 5 o f f ro n t t a r s u s ( F i g . 1 1 20 ) w i t h i n n e r s i d e p rox i m a l l y serra t e , w i t h 1 row of 1 5- 1 6 l a m e l l ae ; apex very p ro m i n e n t a n d ben t a t r i g h t a n g l e , w i t h a g r o u p of 7-9 l a m e l l a e ; m ax i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MA L E : f u l l y w i n ged ; l e n g t h 1 , 6 2 m m ; b ro w n ; m a n d i b l e s t e s t aceou s ; an t e n n ae a n d l e g s n o t b a n ded , b row n ; a n t e n n a e n o t d i s t a l l y t h i ck e n e d ; a n t en n a l s e g me n t s i n fo l l o w i n g p ropor t i o n s : 3 : 3 , 5 : 4 : 4 : 4 : 4 : 3 , 5 : 3 , 5 : 3 , 5 : 4 , 5 ; a n t e n n a I s e g m e n t 3 l e s s t h a n t h ree t i mes a s l o n g a s b road ( 2 , 7 5 ) ; h e a d s h i n y , h a i ry , w e a k l y g r a n u l a t ed ; fron t a l l i n e a b s e n t ; occ i p i t a l c a r i n a i n v i s i b l e ; reg i o n b e t w een e y e s a n d p o s te r i o r ace l l i w i t h a smoo t h s h i n y a rea ; POL = 4 ; Ol = 1 , 5 ; OOl = 2 , 5 ; b r ea d t h of t h e oce l l i : 2 , 5 ; s c u t u m s h i n y , h a i ry , weak l y g r a n u l a t ed ; n o t a u l i ces i n v i s i b l e ; s c u t e l l u m a n d m e t a n o t u m h a i ry , s h i n y , w i t h o u t scu l p t u re ; p ropode u m s h i n y , smoo t h , w i t h o u t scu l p t u r e , w i t h a m ed i a n fu rrow o n a n t e r i or s u rf ace ; fore-w i n g h y a l i n e , w i t h ou t d a r k t r a n s v ers a l b a n d s , on l y w i t h cos t a l ce l l c l ea r l y enc l o sed ; b a s i v o l se l l a w i t h n o tc h ; gonoforce p s ( F i g . 1 1 2 1 ) w i t h a dors a l m e m b r a n o u s p rocess l o n g a n d n a r row ;

m ax i l l a ry p a l p i w i t h 4

1 61 5

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segmen t s ;

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M a l e g en i t a l i a of Gon a top u s incogn i tus n . ( T a n z a n i a ) ( l ef t h a l f removed )

l a b i a l p a l p i w i t h 2 segmen t s ;

LOCUS TYP I CU S : Kawanda ( U g a n d a ) TYP I C A L M A T E R I A L : h o l o t y p e F ! a n d 1 i n OL .

s p . from Pon g w e

t i b i a l s p u rs 1 , 1 , 2 .

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H O ST S : i n T a n z a n i a Cic a d u l i n a zea e . D I ST R I B U T I O N : E t h i o p i a n reg i o n : UGAN D A : K a w a n d a , B M ! TANZAN I A : Pon g w e , B M ! O L ! N O T E S : t h e p a r a t y p e M ! w a s o b t a i ned p a r t h enogenet i ca I I y f rom a n e g g I a i d b y t h e p a r a t y p e F ! i n T a n z a n i a b y rVI r . K i r k p a t r i ck .

1 7 . Gon a top u s lep tothorax C e b a I l os

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F i g . 1 1 22 - Fema l e of Gon a top u s l ep tothorax C eb a l l os f rom M a m a t h es ( Le so t h o ) ( A ) ; S : meso t h orax , m e t a t h o r a x a n d p ropodeum of fem a l e of Gon a topus longi u s c u l u s n . s p . ( t y p e ) .

F E M A L E ( F i g . 1 1 2 2 A ) : ap t erou s ; c l y p e u s a n d f r o n t p a r t of v e r tex

l en g t h 4 , 50 m m ; b l ac k , w i t h m a n d i b l e s , ( more a l o n g o r b i t s ) redd i s h ; a n t e n n a e

b ro w n , w i t h segmen t s 1 - 2 y e l l ow ; a n t en n a l segmen t s i n fo l l o w i n g p ro p or t i on s : 1 0 : 1 0 : 2 2 : 2 1 : 1 7 : 1 3 : 1 1 : 1 0 : 1 0 : 1 1 ; p rono t u m b l ack w i t h red d i sh n u a n ces ; a p e x o f p ropodeum redd i s h ; l eg s y e l l o w , w i t h m i d a n d h i n d coxae b r o w n i s h , c l u b s of fore a n d h i n d femora b ro w n i s h ; h e a d exca v a t e d , sh i n y , h a i r l e s s , smoo t h , w i t h s o m e s h o r t k ee l s n e a r oce l l i ; s i des of occi p u t r u gose,

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1 1 24

Mesothorax , m e t a t ho r a x a n d p ropodeum of f em a l es of Gon a topus a e th iop i c u s n . sp . ( t y p e ) ( A ) and o k a h a n djae n . s p . ( t y p e ) ( B ) .

i m p re ss i o n , g r a n u l a t ed ; sc u t u m v e r y l o n g , n e a r l y t h ree t i me s as l on g a s b road ( 1 5 : 5 ) , d u l l , g r a n u l a t e d ; s c u t e l l u m sh i n y a n d smoo t h , c i rc u l a r ; s t a l k b e t w een p ron o t u m a n d m e t a n o t u m composed of scu t u m a n d scu t e l l u m ( u s u a l l y i n o t h e r Gona topod i n e s t h e s t a l k i s composed on l y of t h e scu t u m ) ; scu t u m + s c u t e l l u m ( F i g . 1 1 2 2 A ) n e a r l y f o u r t i mes as l on g as b road ( 2 0 : 5 ) ; m e t a t h orax + p ropod e u rn d u l l ; m e t a n o t u m , p l eu r a and pos t e r i or s u rf ace of p ropod e u m t r a n s v e r se l y s t r i a t e ; a n t e r i o r s u rf ace of p ropod e u m sh i n y , w i t h 4 med i a n k ee l s a n d , i n s o m e s p ec i m en s , w i t h t r a c k of med i a n f u r row ; m e t a n o t u m v ery l on g ; seg men t 1 of f ron t t a r s u s s l i g h t l y l o n g e r t h a n s e g m e n t 4 ( 1 7 : 1 6 ) ; en l arged c l a w ( F i g . 1 1 23 ) w i t h o u t su b a p i c a l toot h , w i t h o r w i t hou t a sm a l l too t h a t t h e e n d of a l o n g i t u d i n a l fu rrow a n d w i t h 6 l a me l l ae or p e g - l i k e h a i rs ; seg m e n t 5 of fron t t a r s u s ( F i g . 1 1 23 ) w i t h row of 1 8-23 l a m e l l a e ; a pex w i t h a g ro u p of a t l e a s t 1 1 -20 l a me l l ae ; m ax i l l a ry p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 2 seg m e n t s ; t i b i a l s p u rs 1 , 0 , 1 . M AL E : u n k no w n L O C U S T Y P I C U S ; Bek i l y ( M a d a g a sc a r ) T Y P I CA L M A T E R I A L : h o l o t y pe F ! i n P . HOS TS : u n k n o w n D I ST R I B U T I O N : E t h i o p i a n reg ion : MADAGASCAR : Bek i l y , P ! L E S O T H O ; M a m a t h es , B M ! O L !

Gon a topod i n ae

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1 1 25 - C h e l a of Gon a top u s a e th iop ic u s n . s p .

( ty p e )

1 8 . Gon a topus a e t h iopic u s n . s p .

F E M A L E : a p t e ro u s ; l en g t h 4 , 8 1 m m ; h e a d b l ac k , w i t h m a n d i b l es y e l l o w ; a n te n n a e brow n , w i t h seg m e n t 1 y e l l ow ; t h o r a x a n d p ropodeu m b l ac k , w i t h 2 red d i sh s p o t s o n s i des of co l l a r o f p ron o t u m ; a b domen b l ac k ; l eg s t e s t aceous-red d i sh , w i t h c o x a e a n d c l u b s o f f e m o r a b row n i sh ; h e a d exc a v a t e d , sh i n y , w i t h o u t scu l p t u re , e x c e p t for occ i p u t a n d f ro n t p a r t of v e r t ex g r a n u l a t e d ; p ro n o t u m sh i n y , crossed b y a t r a n sv e rs a l i m p ress i o n , h a i r y , w i t h o u t scu l p t u re , excep t for s i d es g r a n u l a t e d ; sc u t u m sh i n y , w i t h some l o n g i t u d i n a l k ee l s ; met a n o t u m ho l l ow b e h i n d t h e sc u t e l l u m ; m e t a t h o r a x + p ro p o d e u m ( F i g . 1 1 24 A ) sh i n y , w i t h o u t scu l p t u re , w i t h med i a n fu rrow ; a n t e r i o r s u rface of p ropode u m w i t h l o n g i t u d i n a l k ee l s ; me t a p l e u ra a n d poster i or s u rface o f p ropodeum t ra n sverse l y s t r i a t e ; mesop l e u ra t r a n sverse l y n o t s t r i a te ; meso-me t a p l e u r a l

s u t u re d i s t i n c t a n d comp l e t e ;

1 620

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F i g . 1 1 26 - Meso t ho ra x , m e t a t h o r a x a n d p ro p o d e u m ( l a t e r a l l y v i ewed ) of fem a l e s of Gon a top u s tro c h a n tericus ( Ben o i t ) ( p a r a t y p e ) ( A ) , tanan arivei n . s p . ( t y p e ) ( B ) , o k a h a n dj a e n . s p . ( t y pe ) ( C ) a n d benoiti n . s p . ( t y p e ) ( D ) .

fore t a r s a l seg men t s i n fo l l ow i n g p ropo r t i on s : 2 0 : 4 : 6 : 26 : 38 ; e n l a rged c l a w ( F i g . 1 1 25 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l too t h at the end o f a l o n g i t u d i n a l f u rrow a n d 3 p e g - I i k e h a i rs ; a p ex of en l a r g e d c l a w t a p e r i n g ; seg men t 5 of f ron t t a r s u s ( F i g . 1 1 25 ) w i t h i n n e r s i d e p rox i m a l l y n o t serra t e , w i t h 2 rows of 3 + 2 7 l a m e l l ae ; a p e x w i t h a g ro u p o f a t l ea s t 1 0 l a m e l l a e ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : F i c h ( E t h i op i a ) TYP I C A L M A T E R I A L : ho l o t y p e F ! i n B M . HOST S : u n k n o w n D I ST R I B U T I ON : E t h i o p i a n reg i o n : E TH I O P I A : F i ch , B M !

1 9 . Gona top u s t a n a n arivei n . s p .

F E MAL E : a p t e rou s ; l en g t h 3 , 37 m m ; h e a d b r o w n -b l ac k , w i t h f r o n t p a r t o f v e r t e x , c l y p e u s a n d m a n d i b l es y e l l ow ; a n t en n ae y e l l o w ; t h o r a x , p ro p o d e u m a n d a b domen b l ac k ; l eg s y e l l o w , w i t h m i d a n d h i n d cox a e a n d c l u b s of femora b row n ; head exc a v a t e d , sh i n y , w i t h o u t scu l p t u re , exce p t for f r o n s a n d occ i p u t g r a n u l a t ed ; p ro n o t u m crossed b y a t ra n s v e rs a l i m p ress i on , s h i n y , w i t h o u t scu l p t u re o r w e a k l y g r a n u l a ted ; scu t u m s h i n y , w i t h some l on g i t u d i n a l k ee l s ; m e t a no t u m sh i n y , t r a n s v e rse l y s t r i a t e ; m e t a t h o r a x + p ro p o d e u m ( F i g . 1 1 2 6 B ) s h i n y , w i t h o u t scu l p t u r e , w i t h a w e a k

G o n a topod i n a e

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1 1 2 7 - C h e l a o f Gon a top u s t a n a n arivei n .

1 62 1

sp .

( type)

l o n g i t u d i n a l a n d med i a n k e e l con n e c t i n g m e t a n o t u m t o d i sc ; p l eu r a a n d p o s t e r i o r s u rf ace o f p ro p o d e u m t r a n s v erse l y s t r i a t e ; meso-me t a p l eu r a l s u t u re d i s t i nc t a n d com p l e t e ; seg men t 1 o f f ro n t t a rs u s as l o n g as se g m e n t 4 ; en l a rged c l aw ( F i g . 1 1 27 ) w i t h o u t s u b a p i c a l toot h , w i t h a s m a l l t oo t h a t t h e e n d o f a l o n g i t u d i n a l f u r ro w a n d 7 p e g - l i k e h a i rs ; s e g m e n t 5 o f f ro n t t a r s u s ( F i g . 1 1 2 7 ) w i t h i n n e r s i d e p rox i m a l l y serra t e , w i t h 1 row o f 1 6 l a m e l l ae ; a p ex w i t h a g ro u p o f 8 l a m e l l a e ; m a x i I l ary p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n LOCUS TYP I CU S :

T a n a n a r i v e ( M a d a g a sca r )

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1 62 2

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1 1 28 - C h e l a of Gon a topus arm a t u s n .

sp .

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1 62 3

G o n a topo d i n a e

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holotype F !

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i n OL . MA DAGASCA R :

Tananarive,

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20 . Gon a top u s arm a t u s n . s p .

F E MAL E : a p t e rou s ; l e n g t h 2 , 8 1 m m ; h e a d b row n - b l ac k , w i t h fro n t p a r t o f v er t e x a l o n g orb i t s , c l y p e u s a n d m a n d i b l e s y e l l ow ; a n t e n n ae b row n , w i t h seg men ts 1 -2 a n d p a r t I y 3 y e l l o w ; t h o r a x a n d p ro p o d e u m b row n - redd i sh ; p e t i o l e b l ac k ; a b domen b r o w n - b l a c k ; l eg s t e s t aceou s , w i t h c l u b s of fore femora d a r k ; head exc a v a te d , s h i n y , w i t h o u t scu l p t u re ; p ron o t u m crossed by a t ra n sv e rs a l i m p ress i o n , sh i n y , w i t h o u t scu l p t u re ; sc u t u m d u l l ; m e t a n o t u m h o l l ow b e h i n d t h e sc u t e l l u m , t r a n sverse l y s t r i a t e ; m e t a t h orax + p ro p o d e u m d u l l , w i t h o u t scu l p t u re ; a n t e r i o r s u rf ace of p ro p o d e u m w i t h m ed i a n f u rrow ; p l eu ra a n d p o s t e r i o r s u rf ace of p ro p o d e u m t r a n sverse l y s t r i a t e ; meso-met a p l e u ra l s u t u re d i s t i n c t a n d com p l e t e ; seg m e n t 1 o f f ro n t t a r s u s s h o r t e r t h a n seg men t 4 ( 1 0 : 1 2 ) ; en l a r g e d c l a w ( F i g . 1 1 28 ) w i t h o u t s u b a p i c a l t oo t h , w i t h a s m a l l t o o t h a t t h e e n d of a l o n g i t u d i n a l f u rrow and w i t h a ser i es of t ee t h , i n a d d i t i o n to 8 p e g - l i k e h a i rs ; seg men t 5 of f ron t t a r s u s ( F i g . 1 1 28 ) w i t h i n ner s i d e p rox i m a l l y not se r r a t e , w i t h 2 rows of 4 + 1 3 l a m e l l a e ; a p e x w i t h a g ro u p of 4 l a m e l l ae ; m a x i l l ary p a l p i w i t h 5 seg men t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l sp u rs 1 , 0 , 1 . MALE : u n k nown L O C U S T Y P I C U S : M a m a t h es ( L esot h o ) TYP I CAL MAT E R I A L : h o l o t y p e F ! i n ElM . H OST S : u n k n o w n D I STR I B U T I O N : E t h i o p i a n r e g i o n : L E S O T HO :

Mama thes,

BM !

2 1 . Gon a top u s ok a h a n djae n . s p .

F EM A L E : a p te ro u s ; l en g t h 3 , 1 2 m m ; h e a d b r o w n - b l ac k , w i t h fron t p a r t of v er t e x ( more a l o n g or b i t s ) , c l y p e u s a n d m a n d i b l es y e l l ow ; a n te n n a e b ro w n , w i t h segme n t s 1 -2 a n d p a r t I y 3 y e l l ow ; t h o r a x , p ro p o d e u m a n d a b domen b ro w n - b l ac k ; l eg s t e s t aceou s , w i t h cox a e and c l u b s of femora brow n ; h e a d a l mo s t f l a t , s h i n y , g r a n u l a t ed ; occ i p i t a l c a r i n a v i s i b l e o n l y beh i n d t h e oce l l i ; p ro n o t u m crossed b y a t r a n s v e rs a l i m p re ss i o n , sh i n y , w i t ho u t scu l p t u re ; m e t a n o t u m ( F i g . 1 1 26 C ) s h or t , t r a n sv erse l y s t r i a t e , h o l l ow b e h i n d t he sc u te l l u m ; meso-me t a p l e u ra l s u t u re d i s t i n c t a n d com p l e t e ; m e t a t h or a x + p ro p o d e u m ( F i g . 1 1 24 B ) sh i n y , w i t h o u t scu l p t u re ; p l e u r a a n d p o s te r i o r s u rf ace of p ro p o d e u m t r a n s verse l y s t r i a t e ; a n t e r i or s u rface o f p ro p o d e u m w i t h s t ro n g m ed i a n f u rrow ; seg men t 1 of f r o n t t a rs u s a s l on g a s seg men t 4 ; en l a r ged c l a w ( F i g . 1 1 29 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l t oo t h a t t h e of a l o n g i t u d i n a l f u r row a n d w i t h 7 p e g - I i k e h a i rs ; seg men t 5

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1 624

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1 62 5

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o f f ro n t t a rs u s w i t h i n n e r s i de p r'ox i m a l l y n o t serra t e , w i t h 1 row o f 1 4 l a me l l ae ; apoex w i t h a g r o u p of a t l ea s t 7 l a m e l l ae ; m a x i l l a �y p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u rs : 1 , 0 , 1 . M A L E : u n k no w n L O C U S T Y P I C U S : O k a h a n d j a ( So u t h W e s t A f r i c a )

M e m . Amer . E n ! .

1 626

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Meso t h o r a x , m e t a thorax and p ropodeum of fema I es of Gon a top u s taylori n . s p . ( p a r a t y p e ) ( A ) a n d upemb a n u s n . s p . ( t y p e ) ( B ) .

TYP I CA L M A T E R I AL : ho l o t y p e F ! i n B M . H O ST S : u n k n o w n D I ST R I B U T I O N : E t h i o p i a n reg i o n : S O U T H W E S T A F R I CA :

22 . Gon a top u s joh n s i n .

O k a h a n dj a ,

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F E MA L E : a p t ero u s ; l en g t h 2 , 52-3 , 43 m m ; h e a d b l ack , w i t h c l y p eus a n d m a n d i b l es y e l l ow ; a n t e n n a e b ro w n , w i t h s e g me n t s 1 -2 a n d p a r t l y 3 y e l l ow ; t h o r a x a n d p ropodeum b l ac k , w i t h a pex of p ro p o d e u m redd i s h ; a b domen b l ac k ; l eg s t e s t aceo u s , w i t h coxae and c l u b s of femora b ro w n o r b l ac k ; h e a d e x c a v a ted , s h i n y , smoo t h , w i t ho u t scu l p t u r e ; p ro n o t u m crossed by a t r a n sv e r s a l i m p ress i o n , s h i n y , w i t h o u t scu l p t u re , exce p t for s i des g r a n u l a t ed ; scu t u m l a t e r a l l y w i t h 2 p o i n ted p rom i n ences ; m e t a n o t u m h o l l ow beh i n d t h e scu t e l l u m , t ra n sverse l y s t r i a t e ; s i de s of m e t a n o t u m p r o t ru d i n g ; p r o t ru s i on s ro u n ded ; m e t a t h o r a x + p ro p o d e u m s h i n y , w i t h o u t scu l p t u re or w ea k l y a l u t aceou s ; a n t e r i o r s u rf ace of p ro p o d e u m w i t h s t ro n g cen t r a l f u r ro w ; p l eu r a a n d poster i or s u rf a ce of p ro p o d e u m t r a n s verse l y s t r i a t e ; meso-me t a p l eu r a l su t u re d i s t i nc t a n d com p l e t e ; seg men t 1 o f f ro n t t a r s u s shorter t h a n s e g m e n t 4 ( 1 3 : 1 5 ) ; e n l a rged c l a w ( F i g . 1 1 30 ) w i t h o u t subap i c a l

too t h ,

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a n d 8 p e g- l i k e h a i r s ; s e g m e n t 5 o f f ro n t t a rs u s ( F i g . 1 1 3 0 ) w i t h 2 ro w s o f 4 ( p rox i m a l ) + 2 1 l am e l l a e ; a p e x w i t h a g ro u p of 1 4 l a m e l l ae ; m a x i I l a ry p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 2 seg men t s ; t i b i a l s p u r s 1 , 0 , 1 . M A L E : u n k no w n L O C U S T Y P I C U S : P o r t S I . J o h n s ( Sou t h Afr i c a ) T Y P I CAL M A T E R I AL : HOST S :

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1 628

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D I ST R I B U T I ON : E t h i op i a n reg i on : S O U T H A F R I CA : P o r t S t . J o h n s ( P o n do l a n d ) , B M ! S O U T H W E S T A F R I CA : 30 M i . SE Kee t m a n s h oo p ,

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2 3 . Gona top u 5 taylori n . s p .

F E MA L E : a p terou s ; l e n g t h 4 , 06-4 , 37 m m ; h e a d b l ack , w i t h m a l a r s p a c e , c l y pe u s a n d m a n d i b l es y e l l ow ; a n t e n n a e b ro w n , w i t h seg men t s 4 - 7 y e l l ow ; t h o r a x a n d p ropodeum b l ac k , w i t h red d i s h n u a nces on d i sc a n d s i des of p rono t u m ; a b domen b l ac k ; l eg s y e l l ow - b row n ; head excav a te d , s h i n y , w i t h o u t scu l p t u re , excep t for f ro n t p a r t o f v e r t e x a n d occ i p u t w e a k l y r u gose a n d g r a n u I a t e d ; p r o n o t u m crossed b y a t ra n sv e r s a I i m p ress i o n , s h i n y , w i t h ou t scu l p t u re , excep t f o r s i des o f d i sc a n d co l l a r show i n g l o n g i t u d i n a l k ee l s a n d g r a n u l a t e d ; scu t u m sh i n y , w i t h some l on g i t u d i n a l k ee l s , l a t e r a l l y w i t h o u t 2 po i n t s ; sc u t e l l u m sh i n y , smoo t h , w i t h o u t scu l p t u r e ; met a n o t u m s h i n y , t ra n s v e rse l y n o t s t r i a t e , ho l l ow beh i n d t h e sc u t e l l u m ; meso p l eu ra l a te r a l l y w i t h 2 po i n t s n e a r m e t a n o t u m ( F i g . 1 1 3 1 A ) ; m e t a t ho r a x + p ropode u m s h i n y , w i t ho u t scu l p t u re ; p l e u ra a n d p o s te r i o r s u rface o f p ropodeum s t ron g l y t ra n sverse l y s t r i a t e ; meso-me t a p l eu ra l s u t u re d i s t i n c t a n d comp l e t e ; se g m e n t 1 of f r o n t t a r s u s s h o r t e r t h a n segmen t 4 ( 1 9 : 2 1 ) ; en l a r g e d c l a w ( F i g . 1 1 3 2 A ) w i t h o u t s u b a p i c a l t oo t h , w i t h a sm a l l tooth a t t h e e n d of a l on g i t u d i n a l f u r ro w a n d 2-6 p e g - I i k e h a i rs o r b r i s t l es ; seg m e n t 5 of f r o n t t a rs u s ( F i g . 1 1 32 A ) w i t h i n n e r s i de p rox i m a l l y n o t se r r a t e , w i t h 1 row of 1 7 l a m e l l ae or 2 rows of 3 ( p r ox i m a l ) + 25-30 l a me l l ae ; a p e x w i t h a g r o u p of a t l ea s t 8-20 l a me l l ae ; m a x i I l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i ( F i g . 1 1 3 2 B ) w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k no w n lOCUS TYP I CU S : Bugusa g a ( U g a n d a ) TYP I CAL M A T E R I AL : h o l o t y pe F ! i n CA ; 1 p a r a t y p e F ! i n O l . H OSTS : u n k no w n

a n d 2 p a ra t y pe s F F !

i n BM ;

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paratype F !

D I S T R I B U T I O N : E t h i o p i a n reg i o n : U G AN D A : B u g u s a g a , B M ! N a m u t a m b a , B M ! B u go t a , O l ! B M ! S O U T H A F R I C A : P o r t S t . J oh n s ( P o n do l a n d ) , B M ! K E N I A : K i l b w ez i , C A l I� O T E S : t h e s p ec i es i s n a med i n honor o f M r . H . C . T a y l or .

24 . Gon a top u s upem b a n u s n . s p .

F E MA L E : a p terou s ; l e n g t h 4 , 37 m m ; b l ack , w i t h c l y pe u s , m a n d i b l es , v en t r a l f ace of a n t en n a l segmen t 1 y e l l ow ; l e g s redd i sh , w i t h cox ae a n d c l u b s of fore femora b l ac k o r b ro w n ; h e a d exca v a t e d , d u l l , g r a n u l a t ed and w i t h s h o r t i r reg u l a r k ee l s ; p rono t u m crossed b y a t r a n s v e rs a l i m p ress i o n , d u l l , g r a n u l a t e d ; sc u t u m g r a n u l a t e d , w i t h o u t l a t e r a l po i n t s

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w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l t oo t h a t t h e e n d of a l on g i t u d i n a l f u rrow a n d 4 p e g - l i k e h a i rs ; a pex o f e n l a rged c l a w t a p er i n g ; seg m e n t 5 of f ro n t t a r s u s ( F i g . 1 1 3 3 ) w i t h i n n e r s i d e p rox i m a l l y not serr a t e , w i t h 2 rows of 4 + 23 l a me l l a e ; a pex w i t h a g r o u p of a t l e a s t 1 0 l a m e l l a e ; m a x i l l a r y p a l p i w i t h 6 segme n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MA L E : u n k no w n L O C U S T Y P I C U S : U p e m b a N a t i o n a l P a rk ( Z a i re ) TYP I CAL M A T E R I A L : h o l o t y p e F ! i n T V . HOSTS : u n k n o w n D I S T R I B U T I ON : E t h i o p i a n reg i o n : Z A I R E : U p e m b a N a t .

Park ,

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25 . Gon a topus benoiti n . s p .

F E M AL E : a p t e ro u s ; l en g t h 3 , 1 2 m m ; y e l l ow -redd i s h , w i t h a n t e n n a l segme n t s 7-9 b row n i s h , p e t i o l e b l a ck a n d a b domen b ro w n ; h e a d exc a v a t e d , sh i n y , w i t h o u t scu l p t u re , w i t h t r a c k o f w e a k l on g i t u d i n a l s t r i a e o n fron s ; p ron o t u m crossed b y a t ra n sve rs a l i m p res s i o n , sh i n y , smoo t h , w i t h o u t scu l p t u re ; sc u t u m sh i n y , w i t h o u t scu l p t u re , w i t h some l o n g i t u d i n a l w e a k k ee l s a n d l a te r a l l y w i t h 2 po i n t s ( F i g . 1 1 03 D ) ; met a n o t u m ( F i g . 1 1 26 D ) smoo t h , n o t or s l i g h t l y ho l l ow beh i n d t h e scu t e l l u m , t r a n s verse l y n o t s t r i a t e ; m e t a t h o r a x + p ropodeu m sh i n y , smoo t h , w i t h o u t scu l p t u re ; me t a p l eu ra a n d poster i or s u rface of p ropod e u m t r a n s verse l y s t r i a t e ; mesop l e u r a t r a n sverse l y n o t s t r i a te ; meso-me t a p l eu r a l s u t u re d i s t i n c t a n ,' comp l e t e ; fore t a rsa l segmen t s i n fo l l ow i n g p ropo r t i on s : 1 2 : 2 : 4 : 1 6 : 23 ; en l a rged c l a w ( F i g . 1 1 34 ) w i t h o u t s u b a p i c a l toot h , w i t h a s m a l l too t h a t t h e e n d o f a l on g i t u d i n a l f u rrow a n d 7 p eg - l i k e h a i rs ; s e g m e n t 5 o f fron t t a rs u s ( F i g . 1 1 34 ) w i t h i n n e r s i d e p rox i m a l l y not serra t e , w i t h 2 ro w s of 4 + 23 l a me l l ae ; a p ex w i t h a g ro u p of 1 1 l a me l l ae ; a pex of e n l a r ged c l a w t a p e r i n g ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l spurs 1 , 0, 1 . MALE : u n k n o w n L O C U S T Y P I C U S : I v o n d ro ( M a d a g a sc a r ) TYP I CAL M A T E R I A L : ho l o t y p e F ! i n T V . HOSTS : u n k no w n D I S T R I B U T I O N : E t h i op i a n reg i o n : MADAGASCAR : I vo n d r o , T V ! N O T E S : t h e spec i e s i s n a med i n honor of M r . P . L . G . Ben o i t .

26 .

Gon a top u s tro c h a n teri c u s ( Ben o i t )

n . comb .

Eucamp tonyx troch a n tericus B e n o i t 1 95 3 b : 387 . Gon a top u s scu tellaris Beno i t 1 9 53b : 396 ( n . s y n . ) ; locu s typ icus Bek i I y ( M a d a g a sc a r ) .

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a pe x of en l a r g e d c l aw t a p e r i n g ; m a x i l l a r y p a l p i w i t h 5-6 segmen t s ; l a b i a l p a l p i w i t h 2-3 segmen t s ; possi b l e p a l p a l r a t i os 5/2 o r 6/3 ; t i b i a l s p u r s 1 , 0, 1 . MALE : u n k nown L O C U S T Y P I C U S : Bek i l y ( M a d a g a sc a r ) TYP I CA L M A T E R I A L : h o l o t y pe F ! o f E . tro c h a n tericus i n P ; paratype F ! of E . troch a n teric u s i n TV ; h o l o t y p e F ! a n d 3 p a r a t y p es F F ' of G . s c u tella ris i n P ; 2 p a r a t y pes F F ! of G . s c u te l l a ri s i n TV . HOST S : u n k n o w n D I ST R I BU T I O N : E t h i o p i a n reg i on : MADAGASCAR : A F R I C A : M a tj esfo n t e i n ( C a p e Prov i n ce ) , B M !

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27 . Gon a top u s op a c u s n . s p .

F E M A L E : a p t e ro u s ; l e n g t h 5 , 6 2 mm ; h e a d b l ac k , w i t h m a n d i b l es y e l l ow ; a n t en n ae b l ac k , w i t h v e n t r a l f ace of seg m e n t 1 y e l l ow ; t h o r a x , p ropodeu m , p e t i o l e a n d a b d omen b l ac k ; l eg s b r o w n - b l a c k , w i t h t a r s i l i g h t ; h e a d exc a v a t e d , d u l l , g r a n u l a t e d ; p ropode u m c rossed b y a t r a n s v e rs a l i m p res s i o n , d u l l , g r a n u l a t e d ; scu t u m d u l l , g r a n u l a t ed , w i t h some l on g i t u d i n a l k ee l s ; sc u t e l l u m d u l l , g r a n u l a t e d ; m e t a n o t u m h o l l ow beh i n d t h e sc u t e l l u m , t r a n sverse l y n o t s t r i a t e ; m e t a t h o r a x + p ropodeum d u l l , g r a n u l a t e d , w i t h some t ra n s v e rs a l s t r i ae on p l e u ra a n d on s i d e s o f pos t e r i o r s u r f a ce of p ropod eu m ; med i a n p a r t of p o s t e r i or s u r f a c e t r a n s v e r se l y n o t s t r i a t e ; segmen t 1 of f ron t t a r s u s s l i g h t l y l on ge r t h a n segmen t 4 ( 20 : 1 9 ) ; en l a r g e d c l a w ( F i g . 1 1 3 6 ) w i t h o u t s u b a p i c a l toot h , w i t h a sm a l l too t h at t h e e n d of a l on g i t u d i n a l f u r row a n d 2 b r i s t l es ; seg m e n t 5 o f f r o n t t a r s u s ( F i g . 1 1 3 6 ) w i t h 2 rows o f 1 ( p rox i m a l ) + 23 l a m e l l a e ; a p e x w i t h a g r o u p o f 1 6 l a me l l ae ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E : u n k no w n LOCUS TYP I C U S : Kyagur ( U g a n d a ) T Y P I CAL MAT E R I AL : h o l o t y p e F ! i n B M . H O ST S : u n k no w n D I ST R I B U T I O N : E t h i o p i a n reg i on :

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s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d o f a l o n g i t u d i n a l f u rrow and 4-5 pe g - l i k e h a i rs ; se g m �rl t 5 of f r o n t t a r s u s ( F i g . 1 1 37 ) w i t h i n n e r s i d e p rox i m a l l y n o t ser r a t e , w i t h 2 r o w s of 3 - 4 + 2 1 -28 l a me l l ae ; a p ex w i t h a g r o u p of 9- 1 3 l a me l l a e ; a p e x of en l a r g e d c l a w t a p e r i n g ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MA L E :

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F E M A L E : a p t e ro u s ; l e n g t h 3 , 1 2- 3 , 75 m m ; y e l l ow - red d i s h o r b ro w n -red d i sh , w i t h p e t i o l e b l ac k a n d a b domen b row n i s h ; l e g s w i t h m i d a n d h i n d t i b i ae b ro w n ; coxae w i t h b o t h a pe x es b row n a n d cen t r a l reg i o n y e l l ow ; h e a d exc a v a t e d , s h i n y , w i t hou t scu l p t u re , excep t for f r o n t p a r t of v e r t e x , s i des o f f ro n s and occ i p u t g r a n u I a ted ; p ro n o t u m crossed b y a t r a n sversa I i m p ress i on , s h i n y , w i t h o u t scu l p t u re ; scu t u m d u l l ; met a n o t u m f l a t , ho l l ow beh i n d t h e s c u t e l l u m , t r a n sv erse l y s t r i a t e ; meta t h orax + p ro p o d e u m sh i n y , smoo t h , w i t ho u t scu l p t u re ; p l eu r a a n d poster i o r su rface of p ro p o d e u m t ra n s v e rs e l y s t r i a te ; meso-me t a p l eu ra l s u t u re d i s t i n c t a n d comp l e t e ; seg men t 1 o f fron t t a rs u s s l i g h t l y shorter t h a n se g m e n t 4 ( 1 4 : 1 6 ) ; en l a rged c l a w ( F i g . 1 1 38 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l tooth at t h e e n d of a l o n g i t u d i n a l f u rrow and 5-6 p eg - I i k e h a i rs ; seg men t 5 of f ro n t t a r s u s ( F i g . 1 1 38 ) w i t h i n n e r s i d e p rox i m a l l y serr a t e or n o t serra t e , w i t h t w o row s of 3 - 5 ( p rox i m a l ) + 1 7-20 l a m e l l ae ; a p ex w i t h a g ro u p of a t l ea s t 1 0- 1 7 l a m e l l ae ; m a x i I l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 seg men t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k no w n L O C U S T Y P I C U S : P o r t S t . J o h n s ( So u t h Afr i c a ) T Y P I C A L M A T E R I AL : h o l o t y pe F ! a n d 3 p a r a t y p es F F ! i n B M ; 3 p a r a t y pes F F ! i n TV . H O ST S : u n k no w n D I ST R I B U T I ON : E t h i o p i a n reg i on : SO U T H A F R I CA : Port S I . J o h n s ( Po n do l a n d ) , O L ! B M ! K l oof ( N a t a l ) , B M ! K a t b erg ( E C a p e P rov i n ce ) , B M ! Worc e s t e r ( C a p e Prov i nce ) , B M ! M i l n e r t o n ( C a p e Tow n ) , ( C a p e P ro v i n ce ) , B M ! C e res ( C a p e P rov i n ce ) , B M ! OL !

8M !

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30 . Gon a top u s e ric e ti n . s p .

F E MA L E : a p t e rou s ; l en g t h 3 , 52 m m ; y e l l o w - redd i sh , w i t h p e t i o l e b l ac k ; a n t e n n a e d i s t a l l y t h i c k ened ; a n t e n n a I segmen t s i n f o l l o w i n g p roport i on s : 8 : 6 : 1 8 : 1 3 : 1 0 : 9 , 5 : 8 : 8 : 8 : 8 ; h e a d sh i n y , w e a k l y g r a n u l a ted , s l i g h t l y e x c a v a t e d ; f ro n s w i t h a com p l e t e m ed i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ;

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i m p ress i o n , sh i n y , w i t h o u t scu l p t u r e ; scu t u m s h i n y , w e a k l y r u gose ; sc u t e l l u m i n v i s i b l e ; m e t a n o t u m sh i n y , n o t ho l l o w beh i n d t h e sc u t e l l u m , t r a n s v e r se l y n o t s t r i a t e ; scu t u m w i t hou t l a t er a l po i n t s ; meso-me t a p l eu r a l su t u re o b so l e t e ; a n te r i o r s u r f a c e o f p ropodeum ( F i g . 1 1 1 1 B ) w i t h o u t scu l p t u re , s h i n y ; p o s t e r i o r s u r f ace of p ropode u m a n d p l e u r a t r a n sverse l y s t r i a t e ; fore t a rsa l segmen t s i n f o l l ow i n g p ropo r t i o n s : 1 5 : 3 : 4 : 1 8 : 28 ; en l a r g e d c l a w ( F i g . 1 1 39 ) w i t hou t s u b a p i c a l toot h , w i t h a sma l l too t h a t t h e e n d o f a l o n g i t u d i n a l f u rrow a n d w i t h 6 b r i s t l e s ; segme n t 5 o f f ro n t t a r s u s ( F i g . 1 1 39 ) w i t h 1 row o f 2 1 l a me l l ae ; a p e x w i t h a g r o u p o f 9 l a m e l l a e ; m a x i I l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l spurs 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : S h a m u I a m b a ( K i v u , Z a i re ) TYP I CA L MAT E R I AL : h o l o t y p e F ! i n OL . HOSTS : u n k n o w n D I ST R I B U T I O N : E t h i o p i a n reg i o n : Z A I R E : S h a m u l a m b a ( K i v u ) ,

OL !

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Segmen t o f f r o n t t a r s u s a p p r o x i ma t e l y t w i c e a s l on g a s s e g m e n t 4 . . . . . . . . . . . . . . . . . . . . . 1 . y a s u m a t s u i n . sp . Segmen t o f f r o n t t a r s u s sh o r t e r , a s l o n g a s , o r s l i g h t I y l on g e r t h a n s eg m e n t 4 . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .2 P r ox i ma l p a r t of f o r e t a r s a l s e g m e n t 5 s e r r a t e on i n n e r ed g e • . • . . . . . • • • • • • . • • • • 3 ( F i g . 1 1 42 ) . . . . . . . . . . . . • • . . . . . • • . . . . P r ox i ma l p a r t o f f o r e t a r s a l s eg m e n t 5 n o t s e r r a t e on i n n e r .8 ed g e ( F i g . 1 1 5 1 ) Me s o - me t a p l e u r a l

su t u r e d i s t i n c t a n d c omp l e t e . . . . . . . . . . . . . . 1 8 . t h a i n . sp . Me s o - me t a p l e u r a l su t u r e ob s o l e t e . • • • • . • . • • • . • • • . . . . . . . . . . . P o s t e r i o r su r f a c e o f p r o p od e u m n o t t r a n sv e r s e l y s t r i a t e . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 . a tten u a t u s n . sp . P o s t e r i o r su r f a c e o f p r op o d e u m t r a n s v e r s e l y s t r i a t e • • . . • • . Me t a n o t u m v e ry f l a t , mu ch l on g e r t h a n l e n g t h o f s c u t e l l u m . .

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. . . .3. a p p r ox i ma t e l y

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1 640

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1 984

t h a n s cu t e l l u m 6 Me t a t h o r a x + p r op o d e u m w i t h a n t e r i o r su r f a c e d u l I , g r a n u l a t e d , w i t h a t r a k o f med i a n f u r r ow 4 . mysoren sis n . sp . Me t a t h o r a x + p r op od e u m w i t h a n t e r i o r su r f a c e sh i n y , s m o o t h , .

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p r op o d eu m t e s t a c e o u s - r ed d i s h 5 . p l e b ei u s ( R . C . L . P e rk i n s ) H ea d l e s s ex c a v a t ed ( F i g . 1 1 4 4 B ) ; S '- ' J t u m a n d me t a t h o r a x + p r op o d e u m b l a c k o r b r own -b l a c k 6 . perp o l i t u s ( R . C . L . P e rk i n s ) .9 Me s o - me t a p l eu ra l s u t u r e ob s o l e t e . . . . . . . . . . . . . . 1 1 Me s o - me t a p l e u r a l su t u r e d i s t i n c t a n d c o mp l e t e . . . H ead f I a t . . . . . . . . . . . . . . . . . . . . 9 . a bjectus n . s p . H e a d ex c a v a t ed . . 10 .

8

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w i t h o u t s c u l p t u r e a n d w i t h ou t t r a k o f med i a n f u r r ow H ea d mo r e ex c a v a t ed ( F i g . 1 1 4 4 A ) ; s c u t u m a n d me t a t h o ra x + .

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1 0 S eg m e n t 5 o f f r o n t t a r su s w i t h t w o r o w s o f l a me l l a e on d i s t a l ha l f ( F i g . 1 1 47 ) 7 . j a v a n u s ( R . C . L . P e rk i n s ) S eg m e n t 5 o f f r on t t a r s u s w i t h t w o r o w s o f l a me l l a e b eg i n n i n g a t p r ox i ma l t h i rd ( F i g . 1 1 4 8 ) . . . 8 . crista t u s ( K i e f f e r ) 1 1 M e t a t h o ra x + p r o p o d e u m w i t h a mo r e o r l e s s s t r o n g med i a n f u r r ow ; i n s o m e sp e c i men s f a i n t I y v i s i b l e a n d s i m i l a r t o a n a r row k e e l 1 2 .

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Me t a t h o ra x + p r op o d e u m w i t h ou t med i a n f u r r ow 1 5 1 2 S c u t u m , s c u t e l l u m a n d me t a t h o r a x + p r op o d e u m t e s t a c e ou s - r ed d i sh . . . . . . . . 13 S c u t u m , s c u t e l l u m a n d me t a t h o ra x + p r o p o d e u m b l a ck 14 1 3 O c c i p u t mo r e e x c a v a t ed ( F i g . 1 1 5 0 C ) ; a n t e r i o r su r f a c e o f me t a t h o r a x + p r o p o d e u m mo r e i n c l i n ed ( F i g . 1 1 5 0 A ) ; me t a t h o ra x .

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+ p r o p od e u m w i t h med i a n f u r r o w d e ep e r ; h e a d a l mo s t f l a t ( F i g . 1 1 50 B ) 1 0 . daunus n . sp . O c c i p u t l e s s ex c a v a t ed ( F i g . 1 1 5 2 C ) ; a n t e r i o r su r f a c e o f me t a t h o ra x + p r op o d e u m l e s s i n c l i n ed ( F i g . 1 1 5 2 A ) ; me t a t h o r a x + p r op o d e u m w i t h med i a n f u r r o w l e s s d e e p ; h e a d ex c a v a t ed ( F i g . 1 1 . p h i l ippin u s n . sp . 1 1 52 B ) S p e c i e s sma l l ; me t a t h o r a x + p r o p o d e u m w i t h a n a r r ow med i a n f u r r ow s i m i l a r t o a k e e l ; v e r t ex w e a k l y ex c av a t ed . . 1 2 . m a ur u s K i e f f e r Sp e c i e s b i g ; me t a t h o ra x + p r op od e u m w i t h a b r o a d med i a n f u r r ow ; v e r t ex v e ry ex c a v a t ed 1 3 . medius n . sp . T h o r a x a n d p r op o d e u m b l a c k . . . . . . . . . . . . . . . . . 16 T h o r a x a n d p r op o d e u m f u l l y o r mo s t l y r ed d i s h 17 Me t a n o t u m s m o o t h , n o t h o l l ow b eh i n d t h e s c u t e l l u m ( F i g . 1 1 4 4 D ) 1 4 . l u c i d u s ( R oh w e r ) M e t a n o t u m t r a n sv e r s e l y s t r i a t e , h o l l ow b eh i n d t h e s c u t e l l u m ( F i g . 1 1 44 C ) 1 5 . m a gn u s n . s p . P r o n o t u m , s c u t u m a n d a n t e r i o r su r f a c e o f me t a t h o r a x + p r o p o d e u m s t r o n g l y s c u l p t u r ed b y l on g i t u d i n a l s t r i a e ; me t a n o t u m n o t v i s i b l e ( t h e i n c l i n ed a n t e r i o r su r f a c e o f t h e me t a t h o ra x + p r op odeum i s d e l i m i t ed d i r e c t l y b y t h e s cu t e l l u m ) ( F i g . 1 1 5 8 B ) .

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1 64 1

Gon a topod i n a e

b od y h a i r y ( F i g . 1 1 5 8 B ) 1 6 . sitae P o n oma r e n k o P r o n o t u m , s c u t u m a n d a n t e r i o r s u r f a c e o f me t a t h o ra x + p r o p o d e u m smo o t h , n o t s c u l p t u r ed b y l on g i t u d i n a l s t r i a e ; me t a n o t u m v i s i b l e , f l a t , h o l l ow b eh i n d s c u t e l l u m ( F i g . 1 1 5 8 A ) ; b od y .

a l mo s t

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1 1 58 A ) . . 1 7 .

in dianus n .

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MALES

Un known

1 . Gon a top u s yasum a ts u i n . s p .

F E MAL E : a p t ero u s ; l en g t h 2 , 1 8- 2 , 50 m m ; b l ac k ; h e a d w i t h m a l a r s p a c e , c l y p e u s , m a n d i b l es , f r o n t p a r t of v e r t ex ( p a r t i c u l a r l y a l o n g orb i t s ) y e l l ow ; a n t e n n a e b row n , w i t h seg men t s 1 or 1 - 2 a n d p a r t l y 3 t e s t aceou s ; l eg s b ro w n , w i t h d i s t a l p a r t of cox a e , t roc h a n t e r s , a r t i c u l a t i o n s a n d t a r s i y e l l o w ; a n t e n n a I segme n t s i n fo l l o w i n g p roport i o n s : 6 : 3 : 7 : 4 : 3 : 3 : 3 : 3 : 3 : 5 ; h e a d w e a k l y co n v e x , s h i n y , g r a n u l a ted , excep t for v e r t e x w h i c h i s smooth a n d w i t h o u t sc u l p t u re ; occi p i t a l c a r i n a i n v i s i b l e ; p ron o t u m crossed b y a t r a n s versa l i m p ress i o n , s h i n y , w i t h o u t scu l p t u re , in some a re a s w ea k l y g r a n u l a t e ; d s c u t u m d u l l , g r a n u l a ted , l a t e ra l l y w i t h o u t po i n t s ; s c u t e l l u m n o t v i s i b l e ; m e t a n o t u m w i t h some t ra n sv e rs a l k ee l s ; rne t a t h o r a x + p ro p o de u m s h i n y , g r a n u l a ted ; p o s t e r i or s u rf ace w i t h on l y a few t ra n s v e r s a l s t r i ae near a p ex ; p l eu r a n o t s t r i a t e ; meso-me t a p l e u ra l s u t u re o b s o l e t e ; fore t a r s a l segmen t s i n fo l l ow i n g p ropo r t i on s : 1 0 : 2 : 2 , 5 : 5 : 9 ; e n l a rged c l a w ( F i g . 1 1 40 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d o f a l o n g i t u d i n a l f u rrow a n d w i t h 1 b r i s t l e a n d 3 p e g - l i k e h a i r s ; seg men t 5 o f f ro n t t a r s u s ( F i g . 1 1 40 ) w i t h i n ner s i d e p rox i m a l l y n o t ser r a t e , w i t h t w o rows o f 1 2 l a me l l ae ; a p e x w i t h a g rou p of 6 l a m e l l a e ; m a x i I l a r y p a l p i w i t h 3 seg me n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MA L E : u n k n o w n LOCUS TYP I C U S : P h a t t h a l u n g ( Th a i l an d ) T Y P I CA L M A T E R I AL : h o l o t y pe F ! i n YA ; 1 p a r a t y p e F ! i n O L ; 2 p a r a t y p e s F F ! i n KY . HOST S : u n k no w n D I ST R I BU T I O N : O r i en t a l reg i o n : T H A I LA N D : P h a t t h a l u n g , YA ! C h a i n a t , K Y ! M A L AYS I A : N o r t h e r n reg i o n , OL ! N O T E S : t h e spec i es i s n a med i n honor of t h e l a te M r . K e i zo Y a s u m a t s u .

2 . Gon a topus a tten u a t u s n . s p .

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1 1 40 - C he l a of Gon a top u s yasum a ts u i n . s p .

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F E MALE : a p t e rou s ; l en g t h 4 , 62-5 , 1 8 m m ; b l ac k ; m a n d i b l es , c l y p eu s , m a l a r space a n d a n a rrow f ron t a l s t r i p e a l o n g orb i t s t e s t aceou s ; a n te n n a e b ro w n , w i t h segmen t s 1 -2 tes t ac eo u s ; l e g s b r o w n - b l ac k ; a n t e n n ae d i s t a l l y t h i c k e n e d ; a n te n n a l segmen t s i n f o l l ow i n g p ropo r t i o n s : 1 4 : 7 : 24 : 1 6 : 1 3 : 11 :

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g r a n u l a ted ; fron t a l l i n e comp l e t e ; occ i p i t a l c a r i n a i n v i s i b l e ( on l y t w o t r a c k s v i s i b l e n e a r p o s t e r i o .- oce l l i ) ; POL = 1 , 5 ; OL = 3 ; OOL = 1 0 ; p ron o t u m w i t h a s t ro n g t ra n sv e rs a l i m p re s s i on ; scu t u m w i t h a l on g i t u d i n a l med i a n k ee l a n d w i t h t r a c k s o f o t h er s h o r t l a t e r a l k ee l s ; m e t a n o t u m n o t f l a t ; rn e t a t h o r a x + propod e u m w i t h s i d es rou n d e d , w i t h o u t t r a n s v e rs a l s t r i ae on p l eu r a a n d p o s t e r i o r or a n t e r i o r s u rf ace ; rneso-met a p l eu r a l s u t u re o b so l e t e ; fore t a rs a l segme n t s i n fo l l o w i n g p roport i o n s : 1 6 : 4 : 5 : 2 3 : 34 ;

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l a b i a l p a l p i w i t h 3 segmen t s ;

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3 . Gon a top u s schen z(lingi S t r a n d

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s y n . ) ; loc u s

F E M A L E : a p t e rou s ; l en g t h 3 , 00-4 , 00 m m ; h e a d t e s t aceo u s , some t i mes w i t h v er t ex d a rk e n e d ; a n t en n ae t e s t aceo u s or b row n , w i t h seg men ts 1 -2 t e s t aceo u s j p ropec t u 5 , pro n o t u m , scu t u m a n d sc u t e l l u m test aceO U 5 , some t i mes w i t h w e a k d a rk s p o t s o n s i des o f p rono t u m ; m e t a n o t u m b l ack o r t e s t aceo u s w i t h s i des b l ac k ; m e t a t h o r a x + p ropodeu m , mesop l e u r a a n d m e t a p l eu r a b r o w n or b l ack , w i t h a p e x o f p ropodeu m t e s t aceo u s ; p e t i o l e b l ack ; a b d o m e n t es t aceou s - d a r k o r b row n , w i t h seg men t 1 p a r t l y t e s t aceo u s ; l eg s f u l l y t e s t aceo u s ; a n t e n n ae d i s t a l l y s l i g h t l y t h i c k e n ed ; a n t en n a l seg men t s i n f o l l o w i n g p ropor t i o n s : 9 : 6 : 1 3 : 1 0 : 8 : 8 : 6 : 6 : 6 : 7 ; h e a d e xc a v a ted , s h i n y , smoo t h , w i t h o u t scu l p t u re or w e a k l y g ra n u l a t e d : P O L = 1 ; OL = 1 , 5 ; t em p l es p rom i n en t ; occ i p i t a l c a r i n a i n v i s i b l e ; p ro n o t u m crossed b y a s t ro n g t r a n s versa l i m p res s i o n , smoo t h , sh i n y , w e a k l y g r a n u I a t ed ; sc u t u m very n a rrow a n d l o n g ; m e t a n o t u m f I a t , t r a n s v erse I y s t r i a t e , m u c h l on g e r t h a n scu t e l l u m ; m e t a t h o r a x + p ro p o d e u m sh i n y , s m oo t h , very p rom i n e n t ; p l eu ra t r a n s v erse l y s t r i a t e ; pos t e r i o r s u rf a ce t r an s v e rse l y s t r i a t e ; a n t e r i o r s u rf ace smoo t h a n d w i t h o u t scu l p t u re ; meso m e t a p l e u ra l s u t u re o b so l e t e ; fore t a rs a l seg m e n t s in fo l l o w i n g p roport i o n s : 1 5 : 2 : 4 : 1 3 : 20 ; en l a rg ed c l a w ( F i g . 1 1 4 2 ) w i t h o u t s u b a p i c a l t oo t h , w i t h a sm a l l t o o t h a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d w i t h 5-7 p eg - I i k e h a i r s ; s e g m e n t 5 of f ro n t t a r s u s ( F i g . 1 1 4 2 ) w i t h i n n er s i d e p ro x i m a l l y serr a t e , w i t h o n e row of 1 4 l a me l l a e ; a p ex w i t h a g ro u p of 9- 1 2 l a me l l ae ; m a x i I l a r y p a l p i w i t h 5 s e g men t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M AL E :

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P a l a e a rc t i c reg i o n : J A P A 'l : F u k u o k a ( K y u s h u ) , KY ! f" O T E S . t h i s spec i es i s re l a t e d to G. l u n a t u s K l u g f ro m t h e P a l a e a rc t i c reg i o n . T h e d i fference i s f ou n d i n t h e m u c h l o n ger s t a l k b e t ween p ron o t u m a n d m e t a t h o ra x + p ropodeu m ( v e r y s h o r t i n l u n a t u s ) , a n d t h e a b sence o f scu l p t u re on t h e a n t e r i or su rface a n d d i sc o f m e t a t horax + p ropodeu m ( i n l u n a tu s t h i s reg i o n i s w e a k l y o r , more f re q u e n t l y , s t ro n g l y g r a n u l a te d ) .

4 . Gon a to[l u s mysoren s i s n .

sp .

F E IYI AL E : a p terou s ; l en g t h 3 , 1 8 m m ; t e s t aceous-re d d i s h , w i t h p e t i o l e b l a ck ; a n t e n n a e very d a m a g ed i n h o l o t y p e ( o n l y t h e f i rs t s e g m e n t p resen t ) ; h e a d excav a t e d , d u l l , g r a n u l a te d ; f ro n s w i t h a med i a n k ee l ; occi p i t a l c a r i n a i n v i s i b l e ; P O L = 1 , 5 ; O L = 2 , 5 ; O O L = 7 , 5 ; p ro n o t u m crossed b y a t r a n s v e rs a l f u rrow , sh i n y , w e a k l y g r a n u l a t e d ; scu t u m sh i n y , w i t h some l on g i t u d i n a l k ee l s ; m e t a n o t u m shor t , n e a r l y a s l o n g a s scu te l l u m , not h o l l o w beh i n d the sc u t e l l u m ; meso-me t a p l e u r a l s u t u re obso l e t e ; meta t h o r a x + p ro p od e u m d u l l , f u l l y g r a n u l a ted ; p l eu r a a n d post e r i o r s u r f a c e t r a n sverse l y s t r i a te ; a n t e r i o r s u rf a ce w i t h a t r ack of med i a n f u rrow ; f o re t a rsa l se g me n t s i n f o l l ow i n g p ro p or t i o n s : 1 3 : 2 , 5 : 3 , 5 : 1 3 : 2 1 ; en l arged c l a w ( F i g . 1 1 43 ) w i t h o u t s u b a p i c a l t oo t h , w i t h a sm a l l too t h at t h e en d o f a

l o n g i t u d i n a l f u rrow a n d w i t h 8 peg - l i k e h a i r s ; s e g m e n t 5 o f f r o n t t a r s u s ( F i g . 1 1 4 3 ) w i t h i n n e r s i de p ro x i m a l l y se r r a t e , w i t h 1 row o f 2 1 l a me l l a e ; a p ex w i t h a g r o u p o f 9 l a m e l l ae ; m a x i l l a r y p a l p i w i t h a t l e a s t 2 segmen t s ( o t h e r segmen t s m i s s i n g i n h o l o t y p e ) ; l a b i a l p a l p i w i t h 2 segm e n t s ; t i b i a l s p u rs 1 , 0 , 1 . M AL E : u n k n o w n LOCU S T Y P I C U S : B a n g a l ore ( I n d i a ) T Y P I CA L MAT E R I A L : h o l o t y p e F ! i n S M . H O ST S : i n I n d i a A thys a n u s s p . D I ST R I B U T I O N : O r i en t a l reg i o n : I N D I A : B a n g a l ore ( til y sore ) , B M !

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a n d perpolitus ( R . C . L . P e r k i n s ) f ro m P e k a l on g a n ( J a v a ) ( B ) ; meso t h o r a x a n d m e t a t h o r a x ( l a ter a l l y v i ew ed ) of fem a l es o f Gon a top u s m agn u s n . s p . ( t y p e ) ( C ) a n d l u cidus ( Ro h w e r ) ( t y p e ) ( D ) .

segrnen t s 3 - 1 0 b ro w n , p e t i o l e b l ac k a n d a b do rnen b row n - b l ack ; h e a d ( F i g . 1 1 44 A ) exc a v a ted , sh i n y , srnoo t h , h a i r l e s s , w i t h o u t scu l p t u re o r w e a k l y scu l p t u red ; p r o n o t u rn c rossed b y a t r a n s v e rs a l i rnp ress i o n , s h i n y , h a i r l es s , w i t h o u t scu l p t u re ; scu t u m w i t hou t scu l p t u r e ; rn e t a t h o r a x + p ro p o d e u m smoo t h , sh i n y , w i t h o u t scu l p t u re ; p o s t e r i o r s u rf ace of p ropod e u m t r a n s v e r se l y s t r i a t e ; p l e u r a n o t t r a n s v e r se l y s t r i a t e ; meso-rne t a p l e u r a l s u t u re o b so l e t e ; se g m e n t 1 of f r o n t t a r s u s s l i g h t l y s h o r t e r t h a n seg m e n t 4 ( 9 : 1 1 ) ; en l a r g e d c l a w ( F i g . 1 1 45 ) w i t h ou t s u b a p i c a l t oo t h , w i t h a sm a l l too t h a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d 9 p e g - l i k e h a i r s ; segmen t 5 of f r o n t t a r s u s ( F i g . 1 1 4 5 ) w i t h i n n e r s i d e p rox i rn a l l y serra t e , w i t h 1 row of 1 5 l a rn e l l ae ; apex w i t h a g r o u p of 1 0 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 2 segrnen t s ; t i b i a l s p u rs 1 , 0 , 1 . M AL E : u n k n o w n L O C U S T Y P I C U 5 : Pek a l o n g a n ( J a v a , I n d o n es i a ) T Y P I CAL MAT E R I A L : l ec t o t y p e F ! a n d 2 p a r a l ec t o t y pes F F ' ( d es i g n a t ed b y O l m i 1 98 2 ) i n B . HOST S : i n I n do n es i a n o t i d en t i f i e d Cica delliclae D I ST R I B U T I O[� : O r i en t a l reg i on : I N D O N E S I A : P ek a l on g a n ( J a v a ) , B ! R o b a n

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F E MA L E : a p t e rou s ; l en g t h 2 , 92 m m ; h e a d b ro w n - red d i s h or b row n - b l ac k , w i t h f r o n t p a r t o f v e rt e x , c l y p e u s a n d m a n d i b l es y e l l ow ; a n t e n n ae b ro w n , w i t h s e g me n t s 1 -3 y e l l ow ; p rono t u m b ro w n - red d i s h or b ro w n - b l ac k ; s c u t u m , s c u t e l l u m a n d me t a t ho r a x + p ro p o d e u m b l ac k o r b row n - b l ac k ; a b domen b row n ; l e g s t e s t aceo u s ; head sl i g h t l y ex c a v a ted , s h i n y , smoo t h , w i t h o u t scu l p t u re ; p ro n o t u m crossed b y a t ra n sv e rs a l i m p re s s i o n , s h i n y , smoo t h , w i t ho u t scu l p t u r e ; scu t u m s h i n y , w i t h some l o n g i t u d i n a l k ee l s ; m e t a t h o r a x + p ropodeum s h i n y , w i t h o u t scu l p t u re ; me t a p l e u r a a n d pos t e r i o r s u rface of p ro p o d e u m t r a n s v e r s e l y s t r i a t e ; mesop l e u r a not t r a n sv erse l y s t r i a t e ; meso m e t a p l e u ra l s u t u re o b so l e t e ; se g m e n t 1 o f f ro n t t a r s u s s l i g h t l y s h o r t e r t h a n seg m e n t 4 ( 9 : 1 0 ) ; en l a rged c l a w ( F i g . 1 1 46 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l t oo t h at t h e e n d of a l o n g i t u d i n a l f u rrow a n d 3-6 peg - l i k e h a i rs ; seg m e n t 5 of f r o n t t a r s u s ( F i g . 1 1 4 6 ) w i t h i n n e r s i d e p rox i m a l l y serr a t e , w i t h 1 r o w o f 1 2- 1 3 l a me l l a e ; a p e x w i t h a g roup of 5 - 7 l a me l l ae ; m a x i l l a ry p a l p i w i t h 5 seg men t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l

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spurs 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : Pek a l on g a n ( J a v a , I n d o n es i a ) TYP I CA L M A T E R I AL : l ec to t y p e F ! a n d 2 p a r a l ec t o t y pes F F ! O l m i 1 982 ) i n B .

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a p t e rou s ;

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p a r t of v e r tex , c l y p e u s a n d m a n d i b l es y e l l ow ; a n t e n n ae y e l l o w , w i t h s e g me n t s 8- 1 0 or 5- 1 0 b row n i sh or b l ac k i sh ; prono t u m re d d i s h - b row n ; scu t u m , sc u t e l l u m a n d met a t horax + propo d e u m b l ac k ; a b domen b row n ; l eg s t e s t aceou s , w i t h cox ae a n d c l u b s o f femora b ro w n i s h ; h e a d exc a v a t ed , s h i n y , g r a n u l a t e d ; pron o t u m crossed b y a t ra n sv e rs a l i m p ress i o n , sh i n y , g r a n u l a ted ; scu t u m sh i n y , w i t h some l on g i t u d i n a l k ee l s ; met a t ho r a x + p ropodeu m sh i n y , g r a n u I a t e d ; met a n o t u m , m e t a p I e u r a a n d pos t e r i o r s u rf ace of propo d e u m t ra n sversel y s t r i a t e ; mesop l eu r a not t r a n sv erse I y s t r i a t e ; meso-me t a p l eu r a l s u t u re o b so l e t e ; segmen t 1 of f ron t t a r s u s l on g e r t h an seg m e n t 4 ( 1 2 : 1 0 ) ; e n l a rged c l a w ( F i g . 1 1 47 ) w i t h o u t s u b a p i c a l too t h , w i t h a sm a l l tooth a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d 2-8 p e g - I i k e h a i rs , i n a d d i t i on t o 1 -4 b r i s t l es ; seg m e n t 5 of f ron t t a r s u s ( F i g . 1 1 47 ) w i t h i n n e r s i de prox i m a l l y not serr a t e , w i t h t w o rows of 1 1 - 1 6 l a me l l ae on d i s t a l h a l f ; a p e x w i t h a g ro u p of 5- 1 0 l a m e l l ae ; m ax i l l a ry p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M AL E : u n k no w n L O C U S T Y P I C U S : Pek a l on g a n ( J a v a , I n dones i a ) TYP I CAL M A T E R I A L : l ec t o t y p e F ! a n d 4 p a r a l ec t o t y pes F F ! ( de s i g n a te d by O l m i 1 98 2 ) in B . H O ST S : i n I n dones i a Deltocep h a l u s sp . ( R . C . L . Perk i n s 1 9 1 2 ; O l m i 1 982 ) . D I ST R I BU T I O N : O r i en t a l reg i o n : I N D O N E S I A : Pek a l on g an ( J a v a ) , B ! OL ! R o b a n ( J a v a ) , B ! Serd a n g ( Su m a t r a ) , BM !

8 . Gon a top u s cri s t a t u s

( K i e f fe r ) n .

com b .

ilaplogon a top u s crista t u s K i effer 1 92 2 : 67 . Mega gon a top u s b irm a n u s O l m i a n d C u rrado 1 97 6 a : 1 78 ( n . sy n . ) ; locus typicus P a l on ( B u r m a ) . Ch a lcogona topus cris t a t u s ( K i ef f er ) : C u rrado a n d O l m i 1 979a : 1 80 .

F E M AL E : a p t e rou s : l en g t h 6 , 25-7 , 46 m m ; b l ac k , w i t h m a n d i b l es a n d s i des of c l y p e u s y e l l ow ; a n t e n n a e b l ac k , in some spec i m e n s w i t h seg m e n t

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b ro w n -y e l l ow ; t a r s i l i g h t ; i n some s p ec i m e n s cox a e , c l u b s of femora a n d t i b i ae redd i sh ; a n t e n n a I segme n t s i n f o l l ow i n g proport i on s : 1 7 : 1 0 : 36 : 30 : 2 2 : 2 2 : 20 : 1 6 : 1 4 ; 1 5 ; head exca v a t ed , s h i n y , smoot h , w i t h o u t scu l p t u re ; p ro n o t u m crossed b y a t r a n sv ersa l i m press i o n , sh i n y , smoo t h , w i t h o u t scu l p t u re ; d i sc n o t reg u l a r l y rou n ded , b u t s t ron g l y compressed poster i or l y i n to a med i a n ca r i n a ; sc u t u m d u l l , w i t h some l on g i t u d i n a l k e e l s ; m e t a n o t u m t r a n sv erse l y s t r i a t e ; m e t a t horax + p ropodeum sh i n y , w i t h o u t scu l p t u re ; p l e u r a a n d pos ter i or su rface o f propo d e u m t r a n s verse l y s t r i a t e ;

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sp .

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meso-me t a p l eu r a l s u t u re obso l e t e ; m e t a n o t u m ho l l o w beh i n d sc u t e l l u m ; fore t a rsa l s e g m e n t s i n f o l l ow i n g p ropor t i o n s : 2 6 : 4 : 1 0 : 32 : 46 ; en l arged c l a w ( F i g . 1 1 48 ) w i t h o u t s u b a p i c a l too t h , w i t h a sm a l l tooth a t t h e e n d of a l on g i t u d i n a l f u rrow a n d 8 p e g - l i ke h a i r s ; seg m e n t 5 of f ron t t a r s u s ( F i g . 1 1 48 ) w i t h i n n e r s i de p rox i m a l l y n o t serra t e , w i t h t w o rows of 36-46 l a me l l ae b e g i n n i n g at p ro x i m a l t h i rd of the segmen t ; apex w i t h a g r o u p of

1 6 55

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F i g . 1 1 50 - F em a l e of Gon a top us d a u n u s n . sp . ( t y p e ) : A: t h o r a x a n d p ro p o d e u m ( l a t e r a l l y v i ew ed ) ; B , C : h e a d ( f ro n t a l l y a n d dor s a l l y v i ew e d ) .

at l e a s t 1 5- 1 7 ! a me l l a e ; m ax i l l a r y p a l p i 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MA L E : u n k n o w n

w i t h 6 segmen t s ;

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l O C U S T Y P I C U S : C u e r n o s Mou n t a i n s ( P h i l i p p i n e I s l a n d s ) TYP I CA L M A T E R I A L : ho l o t y p e F ! of H. crista t u s i n WA ; h o l o t y pe F ' of M . b irm a n u s i n G E . HOST S : u n k n o w n O I ST R I BU T I O N : O r i e n t a l reg i o n : P H i l i P P I N E S : C u e r n o s Mo u n t a i n s ( Neg ros ) , WA ! B U R M A : P a l o n ( P e g u ) , G E !

9 . Gonatop u s a bjectus n .

sp .

F E MAL E : a p t e ro u s ; l en g t h 2 , 1 2 m m ; red d i s h - t e s t aceo u s , w i t h pet i o l e b l a c k ; a n t e n n a e d i s t a l l y t h i ck e n e d ; a n t en n a l s e g m e n t s i n fo l l ow i n g p ropor t i o n s : 5 : 4 : 4 , 5 : 3 , 5 : 2 , 5 : 3 , 5 : 3 , 5 : 3 , 5 : 4 : 6 ; h e a d f l a t , sh i n y , w i t ho u t scu l p t u re , excep t f o r occ i p u t a n d a n t er i or s u rf a ce of f r o n s w h i c h a r e w e a k l y g ra n u l a ted ; f r o n s w i t h a med i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 2 ; O l = 2 ; O O l = 5 , 5 ; p ro n o t u m c rossed b y a s t ro n g t r a n s v e rs a l i m p ress i o n , d u l l , w e a k l y g r a n u l a ted ; sc u t u m d u l l , w i t h w e a k l on g i t u d i n a l s t r i a e ; met a n o t u m w i t h t r a n s versa l s t r i a e ;

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1 65 6

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m e t a t h o r a x + p ro p o d e u m s h i n y , w i t h o u t scu l p t u r e , w i t h p o s t e r i o r s u rf ac e a n d p l e u ra t ra n sverse l y s t r i a t e ; d i sc of p ro p o d e u m w i t h a w e a k m ed i a n i m p re ss i on ; fore t a r s a l s e g me n t s i n fo l l o w i n g p ro p or t i o n s : 8 : 2 : 3 : 6 , 5 : 1 2 ; en l a r g e d c l a w ( F i g . 1 1 49 ) w i t h o u t t ee t h , w i t h 4 peg - l i k e h a i rs ; seg men t 5 of f ro n t t a r s u s ( F i g . 1 1 49 ) w i t h some b r i s t l es a n d o n e on l y l a m e l l a ; a p ex w i t h a g ro u p of n ea r l y 4 l a m e l l ae ; t i b i a l s p u r s 1 , 0 , 1 ; m a x i l l a r y p a l p i w i t h 4 seg men t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k no w n LOCUS TYP I C U S : Ma l l eswaram ( I n d i a ) T Y P I CAL M A T E R I AL : h o l o t y p e F ! i n BM . HOST S : u n k no w n D I ST R I B U T I ON : O r i en t a l reg i on : I N D I A : M a l l es w a r a m ( M y sore ) ,

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F E MA L E : a p tero u s ; l en g t h 2 , 75-3 , 20 m m ; red d i s h -t e s t aceou s , w i t h p e t i o l e b l ack ; a n t e n n ae d i s t a l l y t h i ck ened ; a n te n n a l seg m e n t s i n fo l l o w i n g p ro p o r t i o n s : 9 : 4 : 1 1 : 7 : 6 , 5 : 6 : 6 : 5 : 5 : 6 ; h e a d ( F i g . 1 1 50 B , C ) d u l l , g r a n u l a ted , w i t h t e m p l es v e r y prom i n e n t a n d occ i p u t v e r y exca v a te d ; f ro n s w i t h a n i n com p l e t e m ed i a n k ee l ; occ i p i t a l c a r i n a i n com p l et e , v i s i b l e b e h i n d pos ter i o r oce l l i a n d o n s i des o f post e r i o r oce l l i ; POL � 1 ; O L � 3 ; OOL � 6 , 5 ; p ro n o t u m s h i n y , crossed by a s t r o n g t r a n sv ersa l f u r row , w i t h o u t scu l p t u re , exce p t for t h e s i des w h i c h a r e l on g i t u d i n a l l y s t r i a t e ; scu t u m s h i n y , w i t h some l o n g i t u d i n a l s t r i ae ; s c u te l l u m s h i n y , i n c l i n ed ; m e t a no t u m h o l l o w b e h i n d sc u te l l u m , t r a n s v e rse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re s t ron g , d i s t i nc t a n d comp l e t e ; m e t a t ho r a x + p ro p o d e u m ( F i g . 1 1 50 A ) w i t h o u t scu l p t u re , except for p o s te r i or su rface a n d p l eu ra w h i ch a r e t r a n sv erse l y s t r i a t e ; a n t er i or s u rface w i t h a m ed i a n s t ro n g f u rrow ; a n t e r i o r s u rf ace v e r y i n c l i ned ; fore t a r s a l segmen t s i n fo l l o w i n g p ropor t i o n s : 1 2 : 3 : 4 : 1 3 : 2 1 ; en l a r g e d c l aw ( F i g . 1 1 5 1 ) w i t h o u t s u b a p i c a l toot h , w i t h a s m a l l t oo t h a t t h e e n d o f a l on g i t u d i n a l f u r row a n d w i t h 7 p e g - l i k e h a i rs ; se g m e n t 5 of f ro n t t a r s u s ( F i g . 1 1 5 1 ) w i t h i n n e r s i d e p r ox i m a l l y n o t se r r a t e , w i t h 2 rows of 4 ( o n p r ox i m a l h a l f ) + 1 8 l a m e l l ae ; a p e x w i t h a g r o u p of 1 3 l a me l l ae ; m a x i l l a r y p a l p i w i t h 5 segmen t ; l a b i a l p a l p i w i t h 2 s e g me n t s ; MALE : u n k nown

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F E MAL E : a p t e ro u s ; l en g t h 4 , 00-4 , 1 8 m m ; y e l l ow -red d i s h , w i t h a n t en n a l segme n t s 3- 1 0 b r ow n , p e t i o l e b l ac k a n d a b domen b row n - red d i s h ; a spec i men f ro m V i e t N a m h a s b ro w n spots o n ver tex , p ropec t u s , p ro n o t u m a n d p ropodeu m ; h e a d ( F i g . 1 1 5 2 B , C ) exca v a t e d , sh i n y , w i t h o u t scu l p t u re ; p rono t u m crossed b y a t r a n s v e r s a l i m p ress i o n , s h i n y , smoo t h , w i t hou t scu l p t u re ; sc u t u m d u l l , w i t h some l on g i t u d i n a l k ee l s , l a t e r a l l y w i t h 2 po i n t s ; m e t a t h o r a x + p ropodeum ( F i g . 1 1 52 A ) s h i n y , w i t h o u t scu l p t u re a n d w i t h med i a n f u rrow ; p l e u r a a n d p o s t e r i o r s u r f ace of p ropodeu m t r a n s v e rse l y s t r i a t e ; meso-me t a p l eu r a l s u t u re i d s t i n c t a n d com p l e t e ; seg m e n t 1 o f f ron t t a r s u s a s l o n g a s se g men t 4 ; en l arged c l a w ( F i g . 1 1 5 3 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l tooth a t t h e e n d o f a l o n g i t u d i n a l f u rrow a n d 9- 1 3 pe g - l i k e h a i rs ; segmen t 5 o f f r o n t t a r s u s ( F i g . 1 1 53 ) w i t h i n n er s i d e p r ox i m a l l y n o t serr a t e , w i t h 2 r o w s o f 32 l a me l l ae ; a pex w i t h a g r o u p of 9-30 l a me l l a e ; m a x i I l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : IH, ,, n o w n L O C U S T Y P I C U S : Los B a n o s ( P h i l i p p i n e I s l a n d ) T Y P I C A L MAT E R I A L : h o l o t y p e F ! a n d 1 p a r a t y pe F ! i n OL . H O ST S : u n k n o w n D I ST R I BU T I O N : Or i e n t a l reg i o n : P H I L I P P I N E S : Los Banos ( L u zo n ) , O L ! V I E T NAM : Dj i r i n g , OL !

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F E MAL E : ap terou s ; l e n g t h 5 , 43 m m ; h e a d b r ow n , w i t h occ i p u t , c l y p e u s , m a n d i b l es a n d fron t p a r t of v e r t e x redd i s h ; a n t e n n a e b r o w n - b l ack , w i t h segmen t s 1 -2 y e l l ow-redd i s h ; t h o ra x , p ropodeu m , p e t i o l e a n d a b domen b l a ck ( w i t h red d i sh n u a n ces on d i sc of p r o n o t u m ) ; l eg s red d i s h , w i t h cox a e , c l u b s o f femora a n d t i b i ae f u l l y o r p a r t l y b row n ; h e a d exc a v a t e d , sh i n y , smoo t h , w i t hou t scu l p t u re ; p ron o t u m crossed b y a t r a n sversa l i m p res s i o n , h a i ry , s h i n y , w i t h o u t scu l p t u r e ; scu t u m d u l l , w i t h some l on g i t u d i n a l k ee l s , l a te ra l l y w i t h 2 po i n t s ; m e t a n o t u m h o l l ow beh i n d sc u t e l l u m , t r a n sv e rse l y s t r i a t e ; m e t a t h orax + p ro p o d e u m sh i n y , w i t h o u t scu l p t u re , w i t h a s t ro n g med i a n f u r row ; p l eu r a a n d p o s t e r i o r s u rface o f p ropodeu m t ra n sverse l y s t r i a t e ; meso- m e t a p l e u ra l su t u r e d i s t i n c t a n d com p l e t e ; seg men t 1 of f ro n t t a r s u s s h o r t e r t h a n seg men t 4 ( 1 3 : 2 1 ) ; en l a r ged c l a w ( F i g . 1 1 5 5 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l tooth a t t h e e n d of a l o n g i t u d i n a l fu rrow a n d 1 1 peg - l i k e h a i r s ; seg men t 5 of fron t t a r s u s ( F i g . 1 1 55 ) w i t h i n n e r s i d e p rox i m a l l y n o t serra t e , w i t h 2 rows of 40 l a m e l l a e ; apex w i t h a g ro u p of a t l e a s t 30 l a me l l ae ; m ax i l l a r y p a l p i w i t h 6 segme n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . M A L E : u n k no w n L O C U S TYP I C U S : K i r i R o m ( C a m b o d i a ) T Y P I CA L M A T E R I AL : h o l o t y p e F ' i n B . HOSTS : u n k no w n D I S T R I B U T I O N : O r i en t a l reg i o n : CAMBOD I A : K i r i Rom , B !

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l o n g i t u d i n a I kee l s on f ro n s a n d vertex ; p ro n o t u m c rossed b y a t ra n s v e r s a I i m p ress i on , h a i r y , sh i n y , w i t h o u t scu I p t u re ; d i sc of p r o n o t u m reg u I a r l y ro u n de d , n o t comp ressed i n to a med i a n c a r i n a pos t e r i o r l y ; sc u t u m sh i n y , w i t h l on g i t u d i n a l s t r i a e ; m e t a n o t u m n o t h o l l ow beh i n d sc u t e l l u m ( F i g . 1 1 44 D ) ; m e t a t h o r a x + p ropodeum sh i n y , w i t h o u t scu l p t u r e ; p l eu r a a n d pos t e r i o r s u rface o f p ropodeu m t ra n s v erse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re d i s t i n c t , b u t i rreg u l a r , n o t con t i n uo u s ; fore t a rsa l segme n t s i n fo l l ow i n g p roport i o n s : 1 2 : 3 : 4 : 1 9 : 28 ; e n l a rged c l a w ( F i g . 1 1 56 ) w i t h o u t s u b a p i c a l toot h , w i t h a s m a l l tooth a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d 6 peg l i k e h a i rs ; s e g me n t 5 of front t a r s u s ( F i g . 1 1 56 ) w i t h i n ner s i d e p ro� i m a l l y not ser r a t e , w i t h t w o rows of 1 8- 1 9 l a me l l ae ; a p ex w i t h a g ro u p of 1 4 l a m e l l ae ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k n o w n LOC U S T Y P I C U S : P u s a ( I n d i a ) TYP I CA L M A T E R I A L : h o l o t y p e F !

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F E MA L E : a � t erous j l en g t h 6 , 87 m m ; h e a d t e s t aceou s ; a n t e n n a e b ro w n , w i t h s e g me n t s 1 -2 red d i sh ; t h o r a x a n d p ropod e u m b l ack , w i t h redd i s h n u a n ces on m e t a n o t u m ; a b domen b l ack ; l e g s redd i s h , w i t h sta I k s of femora a n d t a r s i b l ac k i s h ; h e a d sh i n y , smoo t h , exc a v a t e d , w i t h o u t scu l p t u re ; p ro no t u m crossed b y a t ra n sv e rs a l i m p ress i o n , sh i n y , smoo t h , w i t h o u t scu l p t u re ; sc u t u m a l u t aceo u s , comp ressed i n t o a ra i sed med i a n c a r i n a , l a t e ra l l y w i t h 2 po i n t s ( o n e p e r s i de ) ; met a n o t u m h o l l o w b e h i n d s c u t e l l u m ( F i g . 1 1 44 e ) , t r a n s v e rse l y s t r i a t e ; met a t horax + p ropodeum sh i n y , w i t h o u t scu l p t u re ; p l eu ra a n d pos t e r i o r s u rface of p ropodeum t r a n sv e rse l y s t r i a te ( s t r i ae a l so a r o u n d p ropodea l s p i ra c l es ) ; meso-me t a p l eu r a l s u t u re d i s t i n c t a n d comp l e t e ; segmen t 1 of f ro n t t a r s u s s h o r t e r t h a n segmen t 4 ( 27 : 3 3 ) ; en l a r g e d c l a w ( F i g . 1 1 57 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d o f a l on g i t u d i n a l f u rrow a n d 1 0 pe g - I i k e h a i rs ; segmen t 5 o f f ro n t t a r s u s ( F i g . 1 1 5 7 ) w i t h i n n e r s i d e p rox i m a l l y n o t serr a t e , w i t h t w o rows o f 1 4 l a m e l l ae ; a p e x w i t h a g ro u p of at l ea s t 20 l a me l l ae ; m a x i I l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . M A L E : u n k no w n LOC U S T Y P I C U S : F y a n ( V i e t N a m ) TY P I CAL M A T E R I AL : h o l o t y pe F ! i n B . HOST S : u n k n o w n D I ST R I B U T I ON : O r i en t a l reg i o n : V I E T NAM : F y a n , B !

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s u t u re d i s t i n c t a n d comp l e t e ; a b domen a n d l e g s h a i r y ; fore t a rs a l seg men t s i n fo l l o w i n g p ropor t i o n s : 1 8 : 3 : 4 : 1 2 : 1 8 ; en l a r g e d c l a w ( F i g . 1 1 59 ) w i t h o u t s u b a p i c a l t oo t h , w i t h a s m a l l too t h a t the e n d o f a l on g i t u d i n a l f u rrow a n d w i t h 5 b r i s t l e s ; seg m e n t 5 o f f r o n t t a rs u s ( F i g . 1 1 5 9 ) w i t h 2 row s of 20 l a me l l ae ; a pex w i t h a g r o u p of n e a r l y 1 5 l a me l l ae ; m a x i I l a r y p a l p i w i ! h 5 s e g me n t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k n o w n L O C U S T Y P I C U S : C o l o m b o ( Sr i L a n k a ) TYP I CAL M A T E R I AL : h o l o t y pe F ! i n W A . H O ST S : u n k n o w n D I ST R I B U T I O N : O r i en t a l reg i o n : SR I L A N K A : D i s ! . ) , WA !

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r o w s , t h e s h o r t e s t r o w d o e s n o t ex t e n d b ey o n d 0 , 5 l e n g t h o f s e g men t ( F i g . 1 1 7 7 ) . . . . . . . . . . . . . . . 22 7 L a b i a l p a l p i w i t h 2 s e g men t s . . . . . . 8 L a b i a l p a l p i w i t h 3 s eg m e n t s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 8 T h o r a x a n d p r o p od e u m mo s t l y b l a ck . . . . . . . . . . . . . . . . . . . . . . . . .9 Th o r ax and p r op od e u m f u l l y f e r ru g i n ou s o r t e s t a c e ou s . . . . .10 9 H e ad ex c a v a t ed 9 . c on t ortulus P a t t o n H ea d f l a t 1 0 . p l a n ip e s n . sp . 1 0 Me t a t h o r a x + p r o p od eu m u s u a l l y w i t h o u t s c u l p t u r e , ex c ep t f o r s t r i a e on p l e u r a a n d p o s t e r i o r s u r f a c e ; s e gmen t 5 o f f r on t t a r s u s w i t h l a me l l a e a t d i s t a l h a l f ( F i g . 1 1 7 7 ) . . 1 2 . agropyrus F e n t on Me t a t h o r ax + p r op od e u m u s u a l l y g ra n u l a t ed , ex c e p t f o r s t r i a e on p l e u r a a n d p o s t e r i o r su r f a c e ; s e g m e n t 5 of f r' on t t a r s u s w i t h l a me l l a e b eg i n n i n g a t p r ox i ma l h a l f ( F i g . 1 1 7 6 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 1 . mimoides ( R . C . L . P e r k i n s ) 1 1 P o s t e r i o r s u r f a c e o f p r op od e u m n o t t r a n s v e r s e l y s t r i a t e o r s t r i a t e on l y o n p o s t e r i o r h a l f . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 2 P o s t e r i o r su r f a c e o f p r op o d e u m f u l l y t r a n sv e r s e l y s t r i a t e . . . . . . 14 1 2 B od y p a r t I y b l a ck . . . 6 . echo ( R . C . L . P e r k i n s ) .

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B o d y f u l l y t e s t a c e ou s o r r ed d i s h -d a rk , w i t h p e t i o l e b l a c k a n d a b d omen i n s ome sp e c i me n s b r own . . . . . . . . . . . . . 1 3 M e t a n o t u m f l a t ( F i g . 1 1 7 0 A ) . . 7 . elongatus n . sp . Me t a n o t u m n o t f l a t ( F i g . 1 1 7 0 B ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 . fre yta gi n . sp . H e a d , t h o r a x a n d p r op od e u m f u l l y t e s t a c e o u s o r y e l l ow t e s t a c e ou s o r r e d d i s h - t e s t a c e ou s o r f e r r u g i n ou s o r r ed d i s h 1 5 H ea d , t h o r a x a n d p r o p od e u m a t l ea s t p a r t l y b l a ck o r b r own . . . . 1 9 E n l a r g ed c l a w mo r e c u rv ed ( F i g . 1 1 8 1 ) ; s e g men t 5 o f f r on t t a r s u s w i t h t w o sh o r t e r r o w s of l a me l l a e ( F i g . 1 1 8 1 ) . . . . . 1 3 . fre quens ( R . C . L . P e r k i n s ) E n l a rg e d c l a w l e s s c u rv ed ( F i g s 1 1 8 3 , 1 1 8 4 ) ; s e g men t 5 o f f r on t t a r s u s w i t h t w o l on g e r r o w s of l a me l l a e ( F i g s 1 1 8 3 , 1 1 8 4 ) . . 16 H ea d mo r e e x c a v a t ed ( F i g . 1 1 8 2 A , 0 ) 17 .

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M e t a t h o ra x + p r o p o d e u m w i t h a n t e r i o r s u r f a c e l e s s i n c l i n ed ( F i g . 1 2 7 1 6 ) . . . . . . . . . . . . . . . . 1 9 . pseu dochrom u s ( R . C . L . P e rk i n s ) P r o p o d e u m f u l l y y e l l ow - r ed d i s h . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

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1 9 . pseudochrom u s ( R . C . L . P e r k i n s ) P r o n o t u m a t l ea s t p a r t l y b l a ck . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 0 . flavi&ons A s h me a d 2 2 P o s t e r i o r s u r f a c e o f p r op o d e u m w i t h ou t t ra n s v e r s a l s t r i a e o r wi t h s t r i ae o n I y o n p ost er i or ha I f 23 P o s t e r i o r s u r f a c e o f p r op o d e u m f u l l y a n d s t r on g l y t ra n s v e r s e l y st r i a t e 27 2 3 H e a d l e s s e x c a v a t ed , a l mo s t f l a t ( F i g . 1 1 8 6 A ) ; m e s o m e t a p l eu r a l s u t u r e d i s t i n c t a n d c o mp l e t e . . . . . . . . . . . . . . . . . . . . . 2 4 H e a d mo r e e x c a v a t ed ( F i g . 1 1 8 6 B , 0 ) ; m e s o -m e t a p l eu r a l s u t u r e o b s o l e t e o r on l y w i t h p r ox i ma l t h i rd d i s t i n c t . . . . . . . . . . . . . . . . 2 5 .

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2 4 Me t an o t um f l a t , p r o p o d e u m r ed d i Me t a n o t u m n o t f i n c l i n ed d own t .

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2 5 L ab i a l Lab i a l

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pa l p i pa l p i 2 6 S eg men t 5 o f l en g t h ( F i g . S eg me n t 5 o f 1 1 96 ) . . . . . . . 2 7 L ab i a l p a l p i Lab i a l pa l p i 2 8 Me t a t h o rax +

.

m o r e t h a n 0 , 5 a s l on g a s s c u t e l l u m ; t h o r a x a n d s h - t e s t a c e ou s . 2 7 . innitidus ( R . C . L . P e rk i n s ) l a t ; a n t e r i o r s u r f a c e o f m e t a t h o r a x + p r op od e u m o s c u t e l l u m ; t h o ra x a n d p r op od e u m b l a ck . . . . . . . . 2 8 . brooks; n . s p . w i t h 3 s eg m en t s . 2 1 . rapax ( R . C . L . P e rk i n s ) w i t h 2 s eg m e n t s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 6 .

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f r on t t a r s u s w i t h l a me l l a e o f n ea r l y t h e s a m e 1 1 9 4 ) . . . . . . . . . . . 2 2 . a sh m e a di K i e f f e r f r o n t t a r s u s w i t h l a m e l l a e l on g e r n ea r b a s e ( F . . . . . . . . . . . . . . . . . 2 3 . ombrodes ( R . C . L . P e rk i n s ) w i t h 3 s eg men t s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . w i t h 2 s eg me n t s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . p r op o d e u m b l a c k . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

ig . . . 28 . . 31 . . 29

M e t a t h o ra x + p r op o d e u m r e d d i s h -y e l l o w . . . . . . . . . . . . . . . . . . . . . . . . 3 0 2 9 M e s o -m e t a p l e u r a I s u t u r e o b s o l e t e . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

1 673

Gon a topod i n ae

p araleptias ( R . C . L . P e rk i n s ) a n d c o mp l e t e monilis n . s p . 30 S e g men t 4 of I y s h o r t e r t h a n s e g me n t 1 argyrias ( R . C . L . P e rk i n s ) S eg men t 4 o f t h a n s eg m e n t 1 . . . . . . . . 2 6 . curriei K r omb e i n 3 1 H ea d l e s s e x c a v a t ed ( F i g . 1 1 8 6 E ) ; b od y y e l l o w- t e s t a c e ou s , w i t h p e t i 0 I e b l a ck , a b d omen a n d a n t e n n a e p a r t I y b r own i s h 1 2 . agropyrus F en t o n H ea d mo r e ex c a v a t ed ; a t l ea s t s c u t u m , s c u t e l l u m a n d m e t a t h o ra x + p r o p od e u m p a r t l y b l a ck 2 9 . p a l lidiceps ( R . C . L . P e rk i n s ) •

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18. su t u r e d i s t i n c t 24. f r on t t a r s u s s l i g h t 25 . f r o n t t a r s u s l on g e r •

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Me s o -m e t a p l eu r a l .

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MALES

A n t en n a l

2

s eg m e n t 3 l e s s . . . . . . . . . . . . . . . . . . . . . . . A n t en n a l s eg m e n t 3 mo r e P r o p o d e u m f u l l y smo o t h ,

l o n g a s b r oad 1 7 . herb arum ( R . C . L . P e rk i n s ) t h r e e t i me s a s l on g a s b r oad 2 , n o t r e t i cu l a t e r u g o s e 2 . cyphonotus B ra d I e y P r o p od eu m d u I I , r e t i c u I a t e r u g o s e 3 D o r s a l p r o c e s s o f g on o f o r c e p s mu c h sh o r t e r t h a n g on o f o r c ep s ( F i g . 1 1 9 5 ) ; b a s i v o l s e l l a w i t h ou t n o t ch ( F i g . 1 1 9 5 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 2 . a shmeadi K i e f f e r D o r s a l p r o c e s s o f g o n o f o r c ep s n ea r l y a s l on g a s g o n o f o r c e p s ( F i g s 1 1 8 0 , 1 1 9 7 ) ; b a s i v o l s e l l a w i t h n o t ch ( F i g s 1 1 8 0 , 1 1 9 7 ) 4 D o r s a l p r o c e s s o f g on o f o r c ep s w i t h v e ry b r oad d i s t a l r e g i on a n d w i t h d i s t a l ma r g i n s l i g h t l y s e r r a t e ( F i g . 1 1 9 7 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 3 . ombrodes ( R . C . L . P e rk i n s ) D o r s a l p r o c e s s o f g o n o f o r c ep s f u l l y s l e n d e r o r a t mo s t w i t h s l i g h t l y b r oa d e n ed d i s t a l r e g i o n ( F ' g . 1 1 8 0 ) . . . . . . . . . . . . . . . . . . 1 2 . agropyrus F en t o n •

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t h an . . . . . . t han sh i n y

t h r e e t i me s a s

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T h e m a l es of t h e o t h e r s p ec i es of Gon a top u s a re u n k now n . F re y t a g ( 1 977 ) desc r i bed' t w o new s p ec i es of Neogon a topu s ( fentoni and grandis ) . On I y the ma l es a r e k n ow n . They a re t r e a t e d as nomina incerta e sedis because I fee l the m o r p h o l og i c c h a r a c t e r i s t i c s d o not s u ff i c i en t l y i n d i c a t e t h a t the two s p ec i es a r e of the gen u s Gon a top u s .

1 . Gon a top u s cla vicornis ( F e n ton )

Cyrtogo n a topus cla vicornis F e n ton 1 9 27 : 3 .

n.

comb .

Mem . A mer .

1 674

Fig.

F E MA L E :

En\.

I n s\ . ,

no.

37,

1 984

1 1 62 - C h e l a of Gon a top u s c 1 a vicornis ( F e n t on ) ( t y pe )

a p t e rou s ;

l e n g t h 3 , 30 m m ;

h e a d b row n - b l ac k ,

w i t h f ron t p a r t of

v e r t e x , c l y p e u s and m a n d i b l es y e l l ow ; a n t en n ae b ro w n , w i t h se g m e n t s 1 -2 y e l l ow ; t ho r a x a n d p ro p o d e u m b l ac k ; a b d omen b row n - b l ac k ; l e g s y e l l o w red d i s h ; head ( F i g . 1 079 B, C ) f l a t , s h i n y , g r a n u l a t ed ; t em p l es v e r y s h or t , l ess t h a n 0 , 5 a s l o n g as eyes ; p ro n o t u m crossed b y a s t ro n g t r a n s v e rs a l i m p ress i o n , sh i n y , smoo t h , w i t h o u t scu l p t u r e ; scu t u m s h i n y , w i t h some l on g i t u d i n a l k ee l s ; m e t a n o t u m n o t v i s i b l e ; a n t e r i o r s u rf a ce of m e t a t ho r a x + p ropodeum very i n c l i n e d ; m e t a t h o r a x + p ro p o d e u m sh i n y , w i t h ou t scu l p t u r e ; pos t e r i o r su rface o f p ro p o d e u m t r a n sverse l y s t r i a t e ; p l eu r a n o t t r a n sverse l y s t r i a t e ; meso-me t a p l eu r a l s u t u re obso l e t e ; m e t a t h o r a x + p ropod e u m w i t h a s t ro n g med i a n f u rrow ;

fore t a r s a l s e g me n t s

Gon a topod i n a e

Fig.

1 1 63

-

C h e l a of Gon a top u s cyphonotus Br a d l ey from C h i I I i w a c k Co l u m b i a )

1 675

( Br i t i sh

Mem . A me r .

1 676

Fig.

1 1 64

-

Ent.

I nst . ,

n o . 37,

1 984

R a d i a l v e i n of m a l e of Gon a top u s cyphonotus B r a d l ey f rom Ye l l o w k n i fe ( N . W. T . , C a n a d a )

i n f o l l ow i n g p ro p or t i on s : 1 3 : 2 : 3 : 5 : 8 ; fore t roch a n t e r s n ea r l y as l o n g a s fore c o x a e , a b o u t t w i c e a s l o n g a s b ro a d ( 1 2 : 5 ) ; e n l a rged c l a w ( F i g . 1 1 62 ) w i t ho u t s u b a p i c a l t o o t h a n d l a me l l ae , w i t h a s m a l l too t h a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d 4 b r i s t l es ; seg m e n t 5 o f f ro n t t a r s u s ( F i g . 1 1 62 ) w i t h 1 row of v e r y l o n g 7 l a m e l l ae e x t en d i n g con t i n u ou s l y t o apex w h i c h has 5 l a m e l l ae ; a p e x n o t b ro a dened ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . M A L E : u n k no w n L O C U S T Y P I C U S : B ro w n sv i l l e ( Tex . , U . S . A . ) TYP I CAL M A T E R I AL : h o l o t y p e F ! a n d 1 p a r a t y p e F ! HOST S : u n k no w n D I ST R I B U T I ON : N e a rc t i c reg i o n :

U.

S. A . :

i n WA .

B ro w n s v i l l e ( T ex . ) ,

WA !

2 . Gona top u s cyphonotus B r a d l e y

Gon a top u s cyphonotus B r a d l e y 1 906 : 380 . Gon a topu5 fou tsi W h i t t a k e r 1 9 28 : 388 ( n . s y n . ) ; locus typ ic u s C h i l l i w ack ( B . C . , C a n ada ) .

F E MAL E : a p terou s ; l e n g t h 1 , 68-3 , 25 m m ; h e a d b ro w n o r t e s t aceo u s - b ro w n , w i t h f ro n t p a r t of v e r t e x , c l y pe u s a n d m a n d i b l es y e l l o w ; a n t en n ae b ro w n , w i t h s e g me n t s 1 -2 o r 1 -5 y e l l ow ; t ho r a x , p ro p o d e u m a n d a b domen b l ac k ; r a re l y t ho r a x a n d p ropodeum or on I y p ro t h o r a x t e s t aceo u s-d a r k ; l eg s y e l l o w , w i t h cox a e a n d , i n some spec i me n s , c l u b s o f femora b row n ; h e a d f l a t , g r a n u l a t ed , s h i n y ; p ro no t u m crossed b y a t ra n s v e rs a l i m p re s s i o n , s h i n y , h a i r l es s , g r a n u l a ted ; sc u t u m g r an u l a te d ; met a t ho r a x + p ropodeum d u l l , g r a n u l a t ed ; m e t a n o t u m t r a n s v erse l y s t r i a t e ; p o s t e r i o r s u rf ac e of p ropodeu m and p l e u r a more o r l es s w e a k l y t ra n sverse l y s t r i a t e ; meso m e t a p l eu ra l s u t u re o b so l e t e ; segmen t 1 of f r o n t t a r s u s t w i ce a s l o n g as segmen t 4 ( 1 2 : 6 ) ; e n l a r ge d c l aw ( F i g . 1 1 63 ) w i t h o u t su b a p i c a l t oo t h a n d l a m e l l ae , w i t h a s m a l l t oo t h a t t h e e n d of a l on g i t u d i n a l f u r row a n d 6 peg - l i k e h g a i rs ; seg men t 5 of f ro n t t a r s u s ( F i g . 1 1 63 ) w i t h 2 rows of 1 5- 1 9 l a m e l l ae ; a p ex b ro a d e n ed , w i t h a grou p o f 8- 1 0 l a m e l l ae ; m a x i l l a r y p a l p i

Gon a topod i n ae

Fig.

1 67 7

1 1 65 - M a l e g e n i t a l i a of Gon a top u s cyp hon o t u s B r a d l ey f rom Y e l l ow k n i fe ( N . W . T . , C a n a d a ) ( r i g h t h a l f removed )

w i t h 3-4 segmen t s ; l ab i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . M AL E : f u l l y w i n ged ; l en g t h 2 , 3 1 -2 , 8 1 mm ; b l ac k ; a b domen , p ropodeum a n d l eg s b a n d e d ; a n te n n a e n o t d i s t a l l y t h i ck e n e d ; a n t e n n a I segmen t s i n fo l l ow i n g p ropor t i o n s : 7 : 5 : 1 1 : 1 0 : 1 0 : 9 : 9 : 9 : 8 : 1 0 ; a n t e n n a I se g m e n t 3 more t h a n t h ree t i me s as l o n g as b road ( 5 , 6 ) ; h e a d g ra n u l a t e d , d u l l ; f ro n s w i t h a med i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 8 ; OL = 3 ;

1 678

Mem .

Fig.

Ame r .

Ent.

I nst . ,

no.

37,

1 1 66 - C h e l a of Gon a topus mayori n .

1 984

sp .

( type)

Gon a topod i n ae

1 679

OOl = 2; b re a d t h of the oce l l i : 4; scu t u m d u l l , g r a n u l a te d ; n o t a u l i ces i n com p l e t e , reach i n g a p p rox i m a t e l y 0 , 5 l en g t h of scu t u m ; sc u t e l l u m a n d m e t a n o t u m sh i n y , w i t h o u t scu l p t u re o r v e r y w e a k l y g r a n u l a t e d ; p ro p o d e u m sh i n y , smoot h , w i t ho u t scu l p t u re ; fore-w i n g h y a l i n e , w i t h o u t d a r k t r a n sv e r s a l b a n d s ; r a d i a l v e i n : f i g . 1 1 64 ; gonoforceps ( F i g . 1 1 65 ) w i t h a d o rs a l p rocess s h o r t a n d n a rrow ; b a s i v o l se l l a w i t h o u t n o t c h ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u r s 1 , 1 , 2 . l O C U S TYP I C U S : Dow n i e C reek ( B . C . , C a n a d a ) TYP I CAL MAT E R I AL : h o l o t y p e F ! of G. cyphonotus i n I T ; h o l o t y p e F ! a n d 2 p a r a t y pes F F ! of G. fo u t s i i n BM ; 3 p a r a t y p es F F ! of G . fou tsi i n LA . HOST S : i n C a n a d a j\:'a crosteles fascifron s ( S t a l ) . D I ST R I B U T I O N : N e a rc t i c reg i on : CANA D A : Ye l l o w k n i fe ( N . W . T . ) , O T ' O l ' C h i l l i w a c k ( B . C . ) , B ty', ! lA � Dow n i e C reek ( Se l k i r k Mo u n t a i n s , B . C . ) , I T ! M i ss i on C i t y ( B . C . ) , O l ! O T ' S l a v e l a k e ( A l t a . ) , W A ! C a s t o r ( A l t a . ) , O T ! W a i n w r i g h t ( A l t a . ) , O l ' O T ' l a n c a s t e r P a rk ( A l t a . ) , O T ! A l d e r F l a t s ( A l t a . ) , OT � F i s h e r B r a n c h ( M a n . ) , O T ! W i n n i p eg ( M a n . ) , O T ! F l o F r a n c e s ( On l o ) , OT ! U . S. A . : Homestead ( F l a . ) , O T !

3 . Gon a top u s mayori n .

sp .

F E M AL E : a p t e rou s ; l en g t h 3 , 00 m m ; f u l l y t e s t aceo u s , w i t h p e t i o l e b l a c k ; h e a d f l a t , sh i n y , g r a n u l a t e d ; p ro n o t u m crossed by a t r a n s v e r s a l i m p ress i o n , s h i n y , smoo t h , w i t h o u t scu l p t u re ; scu t u m sh i n y , w i t h some l o n g i t u d i n a l k ee l s ; sc u te l l u m sh i n y and smoo t h , w i t h o u t scu l p t u re ; m e t a n o t u m ( F i g . 1 1 86 C ) s h i n y a n d smoo t h , w i t h o u t K �e l s ; m e t a t ho r a x + p ro p o d e u m smooth a n d sh i n y , w i t ho u t sc u l p t u re , w i t h n u merous t r a n s v e rs a l k ee l s o n p os t e r i o r s u rf ace a n d p l e u r a ; viSible;

meso-met a p l eu r a l s u t u re w e a k l y

seg m e n t 1 of f ron t t a rsu s a p p rox i m a t e l y

t w i ce as

l o n g a s seg men t

4 ( 1 2 : 6 ) ; e n l a r g e d c l a w ( F i g . 1 1 66 ) w i t h o u t s u b a p i c a l too t h , w i t h a sm a l l too t h a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d 4 b r i s t l es ; segmen t 5 of front t a r s u s ( F i g . 1 1 66 ) w i t h i n n e r s i d e p ro x i m a l l y not se r r a t e , w i t h 2 rows of 1 0 l a me l l ae ; a p e x w i t h a g r o u p of 9 l a m e l l a e ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n l O C U S TYP I C U S : E I M a y o r ( C a l i f . U . S . A . ) T Y P I CAL M A T E R I AL : h o l o t y p e F � i n C A . HOS T S : u n k no w n D I ST R I B U T I ON : N e a rc t i c reg i o n : U . S . A . : E I M a y o r ( C a l i f . ) , C A '

4 . Gon a to o u s portalen sis n .

sp .

F E MA L E : a p t e ro u s ; l e n g t h. 1 , 60 - 2 , 65 m m ; h e a d b l ac k or b row n , w i t h a n t er i or h a l f o f f r o n s , c l y p e u s a n d m a n d i b l es t e s t aceou s ; a n t e n n a e b row n ,

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( type)

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1 68 1

n a me ( t y p e )

w i t h se g me n t s 1 -3 tes t aceo u s ; t h o r a x , p ropod e u m , a b domen , l e g s u su a l l y t e s t aceou s ; i n some spec i me n s mesosoma d a r k ened ; p e t i o l e b l ac k ; a n t e n n ae d i s t a l l y t h i ck e n e d ; a n t en n a l segmen t s i n f o l l ow i n g p ropor t i o n s : 6 : 4 : 9 : 5 : 4 , 5 : 5 : 4 : 4 : 4 : 6 ; h e a d f l a t , d u l l , g r a n u l a t e d ; frons w i t h a s h o r t i n comp l e t e me d i a n k ee l ; occ i p i t a l ca r i n a i n v i s i b l e ; p r o n o t u m crossed b y a t r an s v e rs a l i m p re s s i on , d u l l , w e a k l y g r a n u l a ted ; sc u t u m a n d m e t a t h o r a x +

1 682

Me m . Amer .

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p ropode u m d u I I , w e a k I y g r a n u I a ted ; poste r i o r s u r f ace of propod e u m a n d p l e u r a n o t t r a n s v e r se l y s t r i a te ; meso-me t a p l e u r a l s u t u re obso l e t e ; fore t a r s a l se gmen t s in fo l l o w i n g propor t i o n s : 1 1 : 2: 3: 5 , 5 : 1 0 ; P O L = 3 ; OL 2 ; OOL = 6 ; e n l a r g e d c l a w ( F i g . 1 1 67 ) w i t h o u t s u b a p i c a l too t h , w i t h a sm a l l too t h a t t h e e n d of a l on g i t u d i n a l f u r row a n d w i t h 5-6 peg - l i k e h a i r s ; s e g me n t 5 of fron t t a r s u s ( F i g . 1 1 67 ) w i t h i n n e r s i d e p rox i m a l l y n o t ser r a t e , w i t h 2 rows of 1 1 - 1 2 l a me l l a e ; apex w i t h a g r o u p of 6-7 l a me l l ae ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l spurs 1 , 0, 1 . MAL E : u n k no w n LOCUS T Y P I C U S : Por t a l ( A r i z . U . S . A . ) TYP I C AL MAT E R I AL : ho l o t y p e F ! a n d 2 p a r a t y pes F F ! i n O T ; 1 p a r a t y p e F ! i n T E ; 1 p a r a t y p e F ! i n A M ; 3 p a r a t y p es F F ! i n OL . HOST S : u n k n o w n D I S T R I B U T I O N : N e a rc t i c reg i o n : U . S . A . : S a g e h e n C reek ( N e v a d a C o . , C a l i f . ) , OL ! Om i r a ( L a s se n C o . , C a l i f . ) , T E ! Port a l ( C a v e C reek C y n . , Ar i z . ) , OL l O T ! W a r d ( C o l o . ) , AM ! K e r rv i l l e ( Tex . ) , O T ! OL !

5 . Gon a top u s translueidus n .

n a me

n e e Gon a top u s testaeeus C a meron 1 888 : 440 . E u e a mptonyx testaeeus R . C . L . Perk i n s 1 907 : 28 .

F E M AL E : a p t e rou s ; l e n g t h 2 , 87-4 , 02 m m ; y e l l ow - t e s t aceo u s or red d i s h - d a r k , w i t h pe t i o l e b l ack , head , i n some spec i me n s , b r ow n i sh ; a n t e n n ae f u l l y t e s t aceous or w i t h segme n t s 1 -4 y e l l ow - t e s t ac eo u s a n d 5- 1 0 b row n i sh ; h e a d exc a v a t ed or a l mo s t f l a t , d u l l or s h i n y , s t r o n g l y or w e a k l y g r a n u l a ted ; p ro n o t u m crossed by a s t r o n g t r a n sversa I i m p re s s i o n , g r a n u I a ted ; m e t a t h o r a x + p ropodeum g r a n u l a t e d ; p os t e r i o r su rf ace a n d p l e u r a t r a n s v e r se l y s t r i a t e ; a n t e r i or su rf ace w i t h l o n g i t u d i n a l k ee l s ; meso - met a p l e u r a l s u t u re obso l e t e ; segmen t 4 of fron t t a rs u s as l o n g as seg m e n t 1 ; en l a r g e d c l a w ( F i g . 1 1 68 ) w i t h 4 - 9 l a me l l a e , w i t h o u t s u b a p i c a l too t h a n d a sm a l l t o o t h a t t h e e n d o f a l o n g i t u d i n a l f u rrow ; a p e x n o t po i n t ed and bent a t r i g h t a n g l e ; segmen t 5 of f ro n t t a rs u s ( F i g . 1 1 68 ) w i t h t w o rows of 1 5 - 1 6 l a me l l a e ; a p e x w i t h a group of a t l e a s t 20-40 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen ts ; t i b i a l spurs 1 , 0, 1 . MALE : u n k no w n LOC U S T Y P I C U S : Nog a l e s ( A r i z . , U . S . A . ) T Y P I C A L MAT E R I AL : ho l o t y pe F ! i n B . H O ST S : u n k no w n D I ST R I B U T I O N : N e a rc t i c reg i o n : U . S . A . : D a n v i l l e ( C a l i f . ) , O L ! Nog a l e s ( A r i z . ) , B ! Sab i n a C a n y on ( Ar. i z . ) , WA ! Bowmer R a n c h ( Be l l C o . , Tex . ) , O T ! F a rm i n g d a l e ( N . Y . ) , T W !

1 68 3

Gon a topod i n a e

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1 1 69

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C h e l a o f Gon a top u s e c h o ( R . C . L . Perk i n s ) ( t y p e )

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Chalcogon a top u s echo R . C . L . Perk i n s 1 90 7 : 23 .

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1 1 70 - Thorax sp .

and p ropodeum of f e m a l es of Gona top u s elonga tus n . ( t y pe ) ( A ) a n d freyt a gi n . sp . ( t y p e ) ( B ) .

F E M A L E : a p t e ro u s ; l e n g t h 4 , 2 1 m m ; h e a d a n d p rono t u m y e l l ow - redd i sh ; a n te n n a e y e l l ow-red d i s h , w i t h seg men t s 5 - 1 0 d a r k e n e d ; scu t u m , scu t e l l u m , me t a t h o r a x + p ropod e u m , p e t i o l e a n d a b domen b l ac k ; l e g s y e l l ow - redd i sh ; h e a d e x c a v a t e d , sh i n y , g r a n u l a ted j p r o no t u m crossed by a t r a n s v e r s a l i m p ress i o n , g r a n u l a t ed ; m e t a t h o r a x + p ropodeum g r a n u l a t ed ; pos ter i o r s u r f a c e of p ropodeum t r a n sverse l y s t r i a t e o n l y o n poste r i o r h a l f ; segmen t 4 of f r o n t t a r s u s l on g e r t h a n segmen t 1 ( 40 : 30 ) ; e n l a rged c l a w ( F i g . 1 1 69 ) w i t h o u t s u b a p i c a l t oo t h , w i t h a sm a l l too t h a t t h e e n d o f a l on g i t u d i n a l f u rrow a n d 1 1 peg - l i k e h a i rs ; segmen t 5 of f r o n t t a r s u s ( F i g . 1 1 69 ) w i t h 2 rows of 7 + 1 6 l a me l l a e ex t en d i n g b e y o n d 0 , 5 l e n g t h of segmen t ; a p ex w i t h a g ro u p o f a t l ea s t 1 0 l a me l l ae ; m ax i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : Nog a l es ( A r i z . , U . S . A . ) TYP I CAL MAT E R I AL : l ec to t y p e F ! ( des i g n a t ed b y O l m i 1 982 ) i n B . HOSTS : u n k no w n D I ST R I B U T I O N : N e a rc t i c reg i o n : U . S . A . : No g a l es ( A r i z . ) , B !

7 . Gon a top u s elonga t u s n . sp .

F E MAL E :

a p terou s ;

l en g t h 4 , 37-4 , 75 m m ;

red d i s h -d a rk or f e rr u g i n o u s ,

w i th

1 68 5

G o n a to pod i n ae

p e t i o l e b l ac k , a n te n n ae b l ac k , w i t h seg men t s 1 -4 b r ow n ; a n t en n ae d i s t a l l y t h i ck ened ; a n t e n n a I segme n t s i n fo l l o w i n g p ropor t i on : 1 1 : 7 : 26 : 1 6 : 1 0 : 1 0 : 8 : 8 : 7 : 1 0 ; h e a d s h i n y , w e a k l y g ra n u l a t ed , exc a v a t ed ; f ron s w i t h a med i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; P O ' � 2 ; OL = 2 ; OOL = 1 0 ; p ro n o t u m c rossed b y a t ra n s v e r s a l i m p ress i o n , d u l l , w e a k l y g r a n u l a t ed ; m e t a n o t u m ( F i g . 1 1 70 A ) f l a t , g r a n u l a t ed , t r a n s v e r se l y s t r i a te ; m e t a t h o r a x + p ro p o d e u m d u l l , g r a n u l a ted ; p l eu ra n o t s t r i a te t r a n s v e r se l y ; p o s t e r i o r s u rf ac e o f p ro p o d e u m not s t r i a t e t r a n s v erse I y , on I y w i t h a f e w s t r i a e n e a r a pex ; meso-met a p l e u r a l s u t u re o b so l e t e ; fore t a r s a l seg men t s i n fo l l ow i n g p roport i o n s : 1 5 : 3 : 5 : 20 : 32 ; en l a r g e d c l a w ( F i g . 1 1 7 1 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l t oo t h a t t h e e n d o f a l o n g i t u d i n a l f u r row a n d w i t h 1 0- 1 7 s m a l l l a me l l a e , p l u s 1 b r i s t l e ; s e g m e n t 5 of f r o n t t a r s u s ( F i g . 1 1 7 1 ) w i t h 2 rows of 29-4 l am e l l ae ; a pex w i t h a g ro u p of a t l e a s t 1 8-30 l a m e l l a e ; m a x i l l a ry p a l p i w i t h 6 s e g me n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u nknown L O C U S T Y P I C U S : T a l l a h a ssee ( F l o r i d a , U . S . A . ) T YP I CAL M A T E R I AL : h o l o t y p e F ! i n WA ; 1 p a r a t y p e F ! i n O L ; 1 p a r a t y p e F ! i n O T ; 1 p a r a t y p e F ! i n AM . HOSTS: u nknown D I ST R I B U T I O N : N e a rc t i c reg i o n :

( H i l l sb o ro u g h C o . ,

Fla. ),

OL !

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S. A . :

T a l l a h a s see ( F l a . ) ,

E l fers ( F l a . ) ,

WA !

Tampa

O T ! j a c k s on v i l l e ( F l a . ) ,

8 . Gon a top u 5 freytagi n .

AM '

sp .

F E MAL E : a p t ero u s ; l en g t h 2 , 50-3 , 37 m m ; tes t a ceou s , w i t h p e t i o l e b l ack ; some t i mes a b domen b row n ; a n te n n a e d i s t a l l y t h i c k e n e d ; a n t en n a l seg men t s i n fo l l ow i n g p ropor t i o n s : 7 : 4 : 1 0 : 8 : 6 : 6 : 5 : 5 : 4 , 5 : 6 ; a n t en n ae tes t ac eo u s , w i t h s e g me n t s 4- 1 0 or 6- 1 0 b row n ; h e a d a l mo s t f l a t , d u l l , g r a n u l a ted ; f r o n s w i t h a med i a n k ee l , occ i p i t a l c a r i n a i n v i s i b l e ; P O L 2; O L = 1 , 5 ; O O L = 8 ; p r o n o t u m cros sed b y a t r a n s v e r s a I i m p ress i o n , d u I I , g r a n u l a ted , h a i r l es s ; sc u t u m , s c u t e l l u m , m e t a n o t u m ( F i g . 1 1 70 B ) , m e t a t h o r a x + p ropodeum a n d p l eu r a d u l l , f u l l y g r a n u l a ted , w i t h o u t k ee l s o r o t h e r sc u l p t u r e , e x c e p t for p o s t e r i o r s u rface o f p ropodeum w h e re there are some t ra n sv e rs a l k ee l s on s i des of pos t e r i or h a l f ; meso-me t a p l e u r a l s u t u re o b so l e t e ; fore t a rs a l seg men t s i n fo l l o w i n g p ropo r t i on s : 1 1 : 2 : 3 , 5 : 9 : 1 5 ; en l a r g e d c l a w ( F i g . 1 1 72 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d of a l on g i t u d i n a l f u rrow a n d w i t h 5 peg - l i k e h a i r s ; seg m e n t 5 o f fron t t a r s u s ( F i g . 1 1 72 ) w i t h 2 rows o f 2+6 o r 1 +3 o r 2+5 l a m e l l ae ( u s u a l l y l oc a ted on a d i s t i n c t p rom i n e n ce ) e x t e n d i n g b e y o n d 0 , 5 l en g t h of segmen t ; a p ex w i t h a g r o u p of 9- 1 5 l a m e l l ae ; m a x i I l a r y p a l p i w i t h 5-6 segmen t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n known L O C U S T Y P I C U S : C h a n n i n g ( T ex a s , U . S . A . ) T Y P I C A L M A T E R I AL :

holotype F !

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in O T ; 1 p a r a t y p e F ! in C M . HOSTS : u n k nown D I S T R I B U T I ON : N e a r c t i c reg i on : U . S . A . : M i l n es a n d ( Roosev e l t C o . , N . M ex . ) , WA ! C h a n n i n g ( O l d h a m C o . , Tex . ) , \VA ! L i m p i a C n . ( F t . D a v i s , T ex . ) , O T ! G r a n i t e M t . ( Tex . ) , X C M ! N O T E S : t h e spec i e s i s n a med i n h o n o r of M r . P a u l H . F rey t a g .

9 . Gon a topus contortu l u s P a t to n

Gona top u s con tort u l u s P a t t o n 1 879 : 65 . Neogon a top u s contor t u l u s ( P a t ton ) : F re y t a g 1 97 7 : 5 7 2 .

F E M AL E : a p terou s ; l en g t h 2 , 37-3 , 62 mm ; h e a d b l ac k or b ro w n - r e d d i s h , u s u a l l y w i t h occ i p u t , f r o n t p a r t o f v e r t e x , c l y p e u s a n d m a n d i b l es y e l l ow ; a n te n n ae b row n , w i t h seg men t s 1 -2 y e l l ow ; p ro n o t u m b ro w n o r b l ac k or t e s t aceo u s , u s u a l l y w i t h d i sc b row n i sh o r b l ac k ; scu t u m b r o w n o r b l ac k o r y e l l o w ; s c u t e l l u m a n d m e t a t h or a x + p ro p o d e u m b l ac k o r b l a c k - b row n , w i t h a p ex of p ro p o d e u m r e d d i s h ; a b domen b row n - b I ack or b r o w n -redd i s h ; l e g s t e s t aceou s ; h e a d exc a v a ted , s h i n y , w e a k l y g r a n u l a t ed ; p ro n o t u m c rossed by a t r a n s v e rs a l i m p re s s i o n , s h i n y , w e a k l y g r a n u l a t ed ; scu t u m s h i n y , l a ter a l l y w i t h 2 p o i n t s ( F i g . 1 1 86 F ) , w i t h some l o n g i t u d i n a l k ee l s ; met a n o t u m t r a n s v e r se l y s t r i a t e ; m e t a t h o r a x + p ro p o d e u m s h i n y , w i t h o u t scu l p t u r e ; p l e u r a a n d p os t e r i or s u rf a c e of p ro p o d e u m t r a n s v e rse l y s t r i a t e ; meso-me t a p l eu r a l s u t u re o b so l e t e ; seg men t 1 o f f ron t t a r s u s a s l o n g a s seg men t 4 ; en l a r g e d c l a w ( F i g s 1 1 73 , 1 1 74 ) w i t ho u t s u b a p i c a l toot h , w i t h a s m a l l t oo t h a t t h e e n d of a l on g i t u d i n a l f u rrow a n d 5-6 p eg - l i k e h a i r s ; seg men t 5 o f f r o n t t a r s u s ( F i g . 1 1 73 ) w i t h 2-3 row s o f 2 1 -26 l a me l l ae e x t e n d i n g b e y o n d 0 , 5 l en g t h of segmen t ; a p ex w i t h a g ro u p of 7 - 1 6 l a me l l a e ; m a x i I l a ry p a l p i w i t h 4-5 segme n t s ; l a b i a l p a l p i w i t h 2 s e g m e n t s ; t i b i a l spurs 1 , 0, 1 . MAL E : u n k n o w n L O C U S TYP I C U S : W a t e r b u ry ( U . S . A . ) TYP I CA L M A T E R I A L : h o l o t y p e F ! i n PH . HOSTS : i n U . S . A . � a ta ll1s sayi ( Fe n ton 1 9 1 8 a , 1 9 1 8 b ) a n d E u tettix ten ella Baker ( Se v er i n 1 924 ) ; in C a n a d a Endria inimica S a y . D I ST R I B U T I O N : N e a rc t i c reg i on : C ANADA : B l a n c h a r d R . ( m i . 93 H a y n e s H w a y , B . C . ) , O T ! C r i s t o p h e r L a k e ( Sa s k . ) , O T ! W i n n i p eg ( M a n . ) , OT ! O L ! P i l o t M o u n d ( M a n . ) , O T ! U . S . A . : M i l V a l l ey ( M a r i n Co . , C a l i f . ) , C A ! P i n e V a l l ey ( Sa n D i ego C o . , C a l i f . ) , C A ! S a g e h e n C reek ( Ne v a d a Co . , C a l i f . ) , T E ! D r y de n ( T e r re l l Co . , T ex . ) , W A ! 1 1 , 2 m i . S of Leve l l a n d ( H ock l e y C o . , Tex . ) , BY ! A l f a l fa C o . ( Ok l a . ) , WA ! A n n A r b o r ( M i c h . ) , T W ! S i ou x C i t y ( I ow a ) , WA ! C o l u m b u s ( Oh i o ) , OL ! S w a n n a n o s ( N . C . ) , A M ! W a t e r b u ry ( Co n n . ) , P H ! S a l i s b u ry C e n t er ( N . Y . ) , OL ! TW ! M E X I C O : C a r r ( C u e r n a v ac a , More l o s ) , O T ! O L !

1 689

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F E MAL E : a p t e ro u s ; l en g t h 3 , 75 m m ; h e a d b r ow n , w i t h m a n d i b l es , c l y peu s , a n t e r i o r reg i on o f fron s , occ i p u t a n d t e m p l es tes taceo u s ; a n te n n a e t e s t ac eo u s , w i t h seg men t s 6-1 0 b row n ; t h o r a x a n d p ro p o d e u m b l ac k , w i t h p ron o t u m p a r t l y red d d i s h ; a b domen b row n -red d i s h ; l e g s t e s t aceou s , w i t h coxae p a r t l y b l ac k , c l u b s of femora b row n , h i n d t i b i ae w i t h b o t h a pexes b ro w n ; a n t e n n a e d i s t a l l y t h i ck e n e d ; a n t en n a l s e g m e n t s i n fo l l o w i n g p ropor t i o n s : 1 0 : 5 : 1 6 : 1 0 : 8 : 7 : 6 : 6 : 6 : 8 ; h e a d f l a t , d u l l , gra n u l a t ed ; f ro n s w i t h an i ncom p l e t e m e c i a n k ee l ; occ i p i t a l c a r i n a 5 1 i g h t l y v i s i b l e b e h i n d oce l l i ; POL = 2 , 5 ; O l = 2 , 5 ; OOl = 1 0 ; O P l = 1 , 5 ; p ro n o t u m crossed b y a t ra n s v e rs a l i m p ress i o n , sh i n y , smooth or w e a k l y g r a n u l a t e d , scu t u m w i t h n u mero u s l o n g i t u d i n a l k ee l s ; s c u t e l l u m s h i n y a n d smoo t h ; m e t a n o t u m scu l p t u red b y t r a n sv ersa l k ee l s ; m e t a t h o r a x + p ropode u m s h i n y , w e a k l y g r a n u l a t e d , w i t h p o s te r i o r s u rf a ce f u l l y t r a n s v e rse l y s t r i a t e ; p l eu ra

1 690

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t r a n sverse l y s t r i a te : meso-me t a p l e u r a l su t u re obso l e t e : fore t a r s a l seg men t s i n f o l l o w i n g p roport i on s : 1 5 : 3 : 8 : 1 6 : 28 : e n l a r g e d c l aw ( F i g . 1 1 75 ) w i t hou t s u b a p i c a l toot h , w i t h a s m a l l too t h a t t h e e n d a l o n g i t u d i n a l f u rrow a n d w i t h 8 peg- l i k e h a i r s : segmen t 5 o f f ro n t t a rs u s ( F i g . 1 1 75 ) w i t h i n n e r s i d e prox i m a l l y serra t e , w i t h 2 row s of 7 + 1 7 l a me l l ae : a p e x w i t h a g r o u p of a t l e a s t 25 l a m e l l ae ; m a x i I l a r y p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u r s 1 , 0 , 1 .

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1 1 78 - M a l e of Gon a topus a gropyrus F e n ton from Lex i n g to n

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F E MAL E : a p t er'o u s ; l e n g t h 2 , 3 1 -4 , 37 mm ; h e a d f u l l y tes t aceo u s or b row n , w i t h occ i p u t , f ro n t p a r t o f v ertex , c l y p e u s a n d m a n d i b l e s y e l l ow tes t aceo u s ; t h o r a x a n d p rop odeu m t e s t aceou s or ferr u g i n o u s ; p e t i o l e b l ac k ; a b domen ferru g i n o u s- b row n i s h ; l e g s tes t aceo u s ; a n t en n a e b row n , w i t h segme n t s 1 -2 y e l l ow ; h e a d very s l i g h t l y exca v a t e d , a l mo s t f l a t , s h i n y , h a i r l es s , g r a n u l a t e d ; p r o n o t u m c rossed b y a t r a n svers a l i m p ress i o n , s h i n y , w i t h o u t scu l p t u re or w e a k l y scu l p t u red o n s i d es ; scu t u m d u l l , g r a n u l a t ed ; s c u t e l l u m l o n g , smoo t h ; p l eu r a , m e t a n o t u m a n d p o s t e r i or s u rf ace of p ropode u m s t r o n g I y t ra n s v e rse I y s t r i a t e ; m e t a t h o r a x + p ropode u m du I I , g r a n u l a ted ; meso-me t a p l e u ra l s u t u re o b so l e t e ; segme n t 1 of f r o n t t a r s u s s l i g h t l y l o n g er t h a n segme n t 4 ( 1 1 : 1 0 ) ; e n l a r g e d c l a w ( F i g . 1 1 76 ) w i t h ou t s u b a p i c a l t oo t h , w i t h a s m a l l too t h a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d 4-8 b r i s t l es or p eg - l i k e h a i r s ; seg m e n t 5 of fron t t a r s u s ( F i g . 1 1 76 ) w i t h t w o rows of 25-36 l a m e l l ae b e g i n n i n g a t p rox i m a l t h i r d a n d e x te n d i n g b e y o n d 0 , 5 l e n g t h o f segmen t ; a p e x w i t h a g rou p o f 1 0- 1 5 l a me l l ae ( 2 of t h ese very l o n g ) ; m ax i l l a r y p a l p i w i t h 4-5 segm e n t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k n o w n

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l O C U S T Y P I C U S : Nog a l es ( A r i z . , U . S . A . ) TYP I CAL M A T E R I AL : l ec t o t y pe F ! ( d es i g n a t ed b y O l m i 1 98 2 ) i n B . HOSTS : u n k no w n D I S T R I B U T I O N : N e a rc t i c reg i o n : U . S . A . : U t a h ( M u esebeck a n d W a l k l e y 1 95 1 ) ; N og a l es ( A r i z . ) , B ! P a l o D u ro ( R a n d a l C o . , T ex . ) , I T ! F t . D a v i s ( P t . R oc k s , Tex . ) , O T ! C a n u t i l l o S t a . ( T ex . ) , WA ! O l ! 1 1 , 6 m i . N of So u r l a k e ( H a r d i n C o . , Tex . ) , BY ' You n g s p o r t Bocomes R a n c h ( Be l l C o . , Tex . ) , Ol !

1 2 . Gon a top u s agropyrus Fen ton

Gon a topus agropyrus Fen ton 1 92 1 : 7 1 . Neogon a topus a gropyrus ( F e n t o n ) : F re y t a g 1 977 : 572 .

F E MA L E : a p t e ro u s ; l en g t h 2 , 0- 3 , 0 m m ; y e l l ow - t e s t a c eo u s , w i t h p e t i o l e b l ac k a n d a b do m e n some t i mes b r o w n i sh ; a n t e n n a e y e l l ow w i t h seg men t s 4- 1 0 b row n i s h ; h e a d ( F i g . 1 1 86 E ) very s l i g h t l y exc a v a t ed , sh i n y , w i t h o u t scu l p t u r e , except f r o n t p a r t of v e r t e x a n d occ i p u t w h i c h a r e w e a k l y g r a n u l a t ed ; p ron o t u m c rossed b y a t ra n s v e rs a l i m p res s i o n , sh i n y , w i t h o u t scu l p t u re ; sc u t u m sh i n y , w e a k l y g r a n u l a t ed , l a ter a l l y w i t h 2 p o i n t s ; m e t a n o t u m w i t h some t r a n s versa l s t r i a e ; m e t a t h o r a x + p ropodeum sh i n y ,

1 696

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F i g . 1 1 80 - M a l e g en i t a l i a of Gon a top u s agropyrus F e n t o n from Lex i n g to n ( K e n t u c k y ) ( A ) ; B : d i ffere n t s h a pes of dors a l p rocess of gonoforcep s ; N : notch .

u s u a l l y w i t h o u t sc u l p t u re ; p l e u r a n o t t ra n s verse l y s t r i a t e ; pos t e r i o r su rface of propode u m t r a n s v erse I y s t r i a t e ; meso-me t a p I e u r a I s u t u re obso l e t e ; segmen t 1 of fro n t t a rs u s s l i g h t l y l on ge r t h a n s e g m e n t 4 ( 1 0 : 9 ) ; e n l a r g e d c l a w ( F i g . 1 1 77 ) w i t ho u t s U b a p i c a l too t h , w i t h a s m a l l too t h a t t he e n d o f a l on g i t u d i n a l f u r ro w a n d 4-6 peg - l i k e h a i rs ; seg men t 5 of f ro n t t a r s u s ( F i g . 1 1 77 ) w i t h 1 row of 9- 1 2 l a m e l l a e or t w o rows of 4 ( p rox i m a l or d i s t a l ) + 2 1 l a m e l l ae ; a p e x w i t h a g ro u p of 7 l a m e l l ae ( 2 of

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1 697

t hese very l o n g ) ; m a x i l l a r y p a l p i w i t h 3-5 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE ( F i g s 1 1 78 , 1 1 79 ) : fu l l y w i n g e d ; l e n g t h 2 , 0-2 , 5 m m ; b l a c k ; m a n d i b l es y e l l ow ; a n te n n ae b row n ; teg u l ae y e l l o w ; l e g s b row n , n o t b a n ded , w i t h t i b i ae a n d t a r s i l i g h t ; a n te n n a l segme n t s i n fo l l o w i n g p ropor t i o n s : 6 : 6 : 1 4 : 1 4 : 1 4 : 1 4 : 1 2 : 1 1 : 1 0 : 1 3 ; a n ten n a l s e g me n t 3 more t h a n f i ve t i mes a s l o n g a s b r o a d ( 6 , 5 ) ; h e a d s h i n y , we a k l y p u n c t a t e , h a i r y , w i t ho u t sc u l p t u r e , except for p u n c t u res ; fro n t a l I i n e a b se n t ; POL � 1 0 ; OL � 3 ; OOL � 3 ; b rea d t h of t h e oce l l i : 4 ; temp l e s d i s t i nc t ; occ i p i t a l c a r i n a i n v i s i b l e ; scu t u m d u l l , g r a n u l a ted , h a i r y ; n o t a u l i ces comp l e t e , poster i o r l y s e p a r a t e d ; m i n i m u m d i s t a nce b e t w een t h e n o t a u l i ces shorter t h a n b rea d t h of oce l l i ( 2 : 4 ) ; s c u te l l u m a n d m e t a n o t u m s h i n y , smoot h , w e a k l y p u n c t a t e ; p ropodeum re t i c u l a te r u g ose , w i t ho u t k e e l s ; dors a l p rocess o f gonoforceps ( F i g . 1 1 80 ) n ea r l y a s l o n g a s gonoforceps , not b roadened a p i c a l l y , s l ender ; b a s i vo l se l l a w i t h n o tc h ; m a x i l l a r y p a l p i w i t h 3-4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . LOCUS TYP I C U S : Ames ( I ow a , U . S . A . ) T Y P I CAL M A T E R I A L : h o l o t y p e F ! , a l l o t y p e M ' a n d 2 p a r a t y p es F F ! of G . a gropyru s i n W A . HOST S : i n U . S . A . Ps ammotettix affin is ( G . a n d B . ) ( F en t o n 1 9 2 1 ) ; Ps ammotettix stria t u s , Gra minella n igrifron s ( F orbes ) , L a ta l u s sayi ( F re y i a g 1 97 7 ) ; i n C a n a d a Endria inimica . O I ST R I BUT I O N : N e a rc t i c reg i on : CANADA : S t . D a v i d s ( O n t . ) , OT ' O t t a w a ( O n t . ) , O T ! U . S . A . : S u n n y s i de ( H u a c h u c a M t s . , A r i z . ) , OT ! 25 , 1 m i . �J of A n g l e t o n ( B razor i a C o . , T e x . ) , OL ! 5 , 3 m i . N of La G r a n g e ( F a y e t t e Co . , T e x . ) , BY ! O L ! B i g B e n d N . P . ( T ex . ) , OT ! 2 , 8 m i . S of Somer v i l l e ( W a s h i n g to n Co . , Te x . ) , BY ! W a co ( Me Len n a n Co . , Te x . ) , B Y ! 25 , 4 m i . W of Refu g i o ( Bee C o . , Te x . ) , Te x . ) , BY ! W i scon s i n ( F rey t a g 1 977 ) ; Ames ( I o w a ) , WA ! Le x i n g ton ( K y . ) , OL ! P t . V a l l e y ( G a . ) , WA ! I t h a c a ( N . Y . ) , TW !

1 3 . Gon a topus fre quens

(R.C.L.

Perk i n s ) n . co m b .

Chalcogon a to o u s fre q uens R . C . L . Perk i n s 1 907 : 2 1 . A gon a top u s p a llidicornis R . C . L . Perk i n s 1 90 7 : 29 ( n . s y n . ) ; locus typ i c u s N og a l es ( A r i z . , U. S . A . ) . A gona top u s p a ll idicornis R . C . L . P e r k i n s v a r . d u 6 iosus R . C . L . P e r k i n s 1 90 7 : 30 ( n . s y n . ) ; loc u s typ icus Nog a l es ( A r i z . , U . S . A . ) .

F E MALE : a p terou s ; l en g t h 2 , 70-4 , 02 m m ; y e l l o w - t e s t aceo u s , w i t h pe t i o l e b l ac k a n d a b domen b row n i s h ; a n t en n ae t e s t a ceou s , w i t h segmen t s 4- 1 0 d a rk e n e d ; h e a d e xc a v a ted , d u l l , g r a n u l a t e d ; pron o t u m crossed b y a t r a n svers a l i m p ress i o n , s h i n y , w i t ho u t sc u l p t u re or w e a k l y g r a n u l a ted ; s c u t u m g ra n u l a te d ;

m e t a n o t u m we a k l y t r a n sve rse l y s t r i a t e ;

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1 698

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1 700

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1 984

p ropodeum s h i n y , w i t h o u t sc u l p t u re ; p l e u r a a n d poster i or s u rface of p ropodeum t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t ure o b so l e t e ; seg m e n t of fro n t t a rs u s a s l on g a s seg m e n t 4 ; en l a rged c l a w ( F i g . 1 1 8 1 ) w i t h o u t s u b a p i c a l toot h , w i t h a s m a l l too t h a t t h e e n d of a l o n g i t u d i n a l fu rro w a n d 2-5 p e g - I i k e h a i rs ; seg men t 5 of fro n t t a r s u s ( F i g . 1 1 8 1 ) w i t h 2 r o w s of 7 - 1 3 l a me l l ae ( e x t en d i n g b e y o n d 0 , 5 l e n g t h of segmen t ) ; a p e x w i t h a g r o u p of 7- 1 3 l a m e l l ae ; m a x i l l a r y p a l p i w i t h 5-6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n knonw L O C U S T Y P I CU S : No g a l es ( A r i z . U . S . A . ) TYP I CAL M A T E R I A L : l ec t o t y pe F ! a n d 3 p a r a l ec t o t y pes F F ! ( d es i g n a ted b y O l m i 1 98 2 ) o f CU. fre quens i n B ; l ec t o t y p e F ! a n d 1 p a r a l ec t o t y pe F ! ( des i g n a t e d b y O l m i 1 982 ) o f A . pa llidicornis i n B ; t y p i c a l m a te r i a l o f A .

pall idicornis v a r . d u b io s u s l os t . HOST S : i n U . S . A . n o t i d en t i f i e d C i c a d e l l i d s . D I S T R I B U T I O N : N e a rc t i c roeg i o n : U . S . A . : T a ss a j a r a H o t ( C a l i f . ) , CA l N og a l es ( A r i z . ) , O L ! B ! A r i zo n a , B M !

1 4 . Gon a top u s areol a t u s ( F en ton )

n . com b .

Chalcogon a top u s areol a tu s F e n ton 1 927 : 7 .

F E M AL E : a p terou s ; l en g t h 3 , 4 3-4 , 1 0 m m ; ferr u g i n ou s , w i t h p e t i o l e b l a ck ; h e a d ( F i g . 1 1 82 A ) exc a v a t e d , d u l l , h a i r l e s s , gra n u l a t e d ; p ron o t u m crossed b y a t r a n s vers a l i m p res s i on , d u l l , g r a n u l a ted ; s c u t u m d u l l , g r a n u l a ted , l a t e ra l l y w i t h 2 po i n t s ( F i g . 1 1 82 B ) ; s c u te l l u m d u l l a n d g r a n u l a ted ; m e t a n o t u m t r a n s verse l y s t r i a t e ; m e t a t ho r a x + propodeum ( F i g . 1 1 82 C ) d u l l , gra n u l a t e d ; p l e u r a a n d pos t e r i or s u rface of p ropodeum t r a n s v e rse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re o b so l e t e ; fore t a rs a l se g m e n t s i n f o l l o w i n g p ropor t i on s : 1 4 : 3 : 4 : 20 : 30 ; e n l arged c l a w ( F i g . 1 1 83 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l t oo t h a t t h e e n d of a l o n g i t u d i n a l f u r row a n d 1 1 - 1 2 peg - l i k e h a i rs ; segme n t 5 o f fro n t t a rs u s ( F i g . 1 1 83 ) w i t h 2 rows of 26-36 l a me l l ae e x t en d i n g b e y o n d 0 , 5 l e n g t h of segmen t ; apex w i t h a g r o u p o f 1 3-20 l a me l l a e ( some of t h ese very l o n g ) ; m a x i l l a r y p a l p i w i t h 6 s e g me n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : F a l l s C h u rc h ( V a . , U . S . A . ) TYP I CAL MAT E R I A L : h o l o t y p e F ! a n d 1 p a r a t y p e F ! i n IVA . HOST S : u n k no w n D I ST R I B UT I O N : N e a rc t i c reg i o n : U . S . A . : U n i ve r s i t y of I l l i n o i s ( Woods , I I I . ) , O L ' C o l u m b i a ( S . C . ) , T W ! F a l l s C h u rc h ( V a . ) , ( F e n ton 1 927 ) ; G r a s s y me a d ( V a . ) , WA ! W a s h i n g to n ( D . C . ) , WA ! C h e v y ( C h a s e L a k e , M d . ) , WA '

Gon a topod i n a e

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C h e l a of Gon a top u s h a rp ax ( K rombe i n ) ( t y p e )

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Chalcogon a top u s h a rpax Krombe i n 1 9 56 :

1 58 .

F E MALE : a p t e ro u s ; l en g t h 3 , 25-3 , 70 m m ; tes t aceous o r redd i s h - d a r k , w i t h p e t i o l e b l a ck a n d a b domen b ro w n - red d i s h ; h e a d ( F i g . 1 1 82 D ) exca v a ted , s h i n y , w i t h o u t sc u l p t ure , e xc e p t for occ i p u t a n d fron t p a r t of v e r t e x w h i c h a re g ra n u l a te d ; pron o t u m crossed b y a t r a n s v e rs a l i m p ress i o n , v e r y s h i n y , s moo t h , h a i r l ess , w i t h o u t scu l p t u re ; scu t u m s h i n y , w i t ho u t scu l p t u re ; m e t a n o t u m s h i n y , w i t h some t r a n s v e r a I s t r i a e ; m e t a t h o r a x + p ro p o d e u m ( F i g . 1 1 82 F ) s h i n y , w i t h o u t sc u l p t u re ; s c u t u m l a ter a l l y w i t h 2 po i n t s ( F i g . 1 1 82 E ) ; m e t a n o t u m a l so l a te ra l l y w i t h 2 poi n t s ( F i g . 1 1 8 2 E ) ; meso-met a p l eu r a l s u t u re obso l e t e ; p l e u r a a n d p o s t e r i or s u r face of p ropode u m t r a n s v ers e l y s t r i a te ; seg m e n t 4 of fron t t a rs u s n e a r l y a s l o n g a s s e g m e n t 1 ; e n l a r ged c l a w ( F i g . 1 1 84 ) w i t ho u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d of a l on g i t u d i n a l fu rrow a n d 7 peg - l i k e h a i rs ; seg m e n t 5 of fron t t a rs u s ( F i g . 1 1 84 ) w i t h 2 rows of 5-6 + 24 l a m e l l a e e x t e n d i n g b e y o n d 0 , 5 l e n g t h of s e g m e n t ; a p e x w i t h a g ro u p of 7-20 l a me l l ae ; m a x i I l a ry p a l p i w i t h 6 s e g m e n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k nown L O C U S TYP I C U S : L o s t R i ver S t a te P a rk ( W . V a . , U . S . A . ) TYP I C AL MAT E R I AL : h o l o t y p e F ! i n WA ! HOSTS : u n k no w n D I ST R I BU T I O N : N e a rc t i c reg i o n : C A NA D A : P r o E d w ard Co . ( On ! . ) , U . S . A . : Lost R i v e r S t a t e P a rk ( IV . V a . ) , WA !

1 6 . Gon a top u s solitarius

(R.C.L.

Perk i n s )

OT !

n . com b .

Ch alcogon a topus solitarius R . C . L . Perk i n s 1 90 7 : 2 2 . Ch alcogon a topus xestocep h a l u s R . C . L . Perk i n s 1 907 : 24 ; loc u s typ icus N og a l es ( A r i z . , U. S. A . ) Agona top u s ferrugin e u s R . C . L . P e rk i n s 1 907 : 30 ( n . s y n . ) ; loc u s typ i c u s Nog a l e s ( A r i z . , U . S . A . ) A gona top u s picescens R . C . L . Perk i n s 1 907 : 30 ( n . s y n . ) ; locus typicus Nog a l es ( A r i z . , U. S. A . ) Chalcogon a top u s xestocep h a l u s R . C . L . P e rk i n s : K i effer 1 9 1 4 b : 1 0 1 . A gona top u s ferrugin e u s R . C . L . P e rk i n s : K i effer 1 9 1 4 b : 1 32 . A gon a top u s p icescen s R . C . L . Perk i n s : K i effer 1 9 1 4 b : 1 3 3 .

F E MA L E : a p t e rou s ; l e n g t h 3 , 6-5 , 1 m m ; redd i s h-ferru g i n ou s , w i t h p e t i o l e b l ac k ; a n t e n n ae red d i sh-ferru g i no u s , w i t h s e g m e n t s 4 - 1 0 o r 6- 1 0 o r 7- 1 0

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b r ow n ; a b domen occ a s i on a l l y b r o w n i s h ; a n t e n n a e d i s t a l l y t h i c k e n e d ; a n t e n n a l s e g m e n t s i n fo l l o w i n g p roport i o n s : 1 4 : 6 : 1 7 : 1 0 : 8 : 9 : 7 : 6 : 6 : 9 ; h e a d f l a t , d u l l , g r a n u l a te d ; fron t a l l i n e comp l et e ; tem p l es d i s t i n ct ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 1 , 5 ; OL = 2 ; O O L = 1 0 ; p ron o t u m s h i n y , smoot h , w i t h o u t scu l p t u re , crossed b y a s t ro n g t r a n s ver s a l i m p res s i o n ; scu t u m w i t h l on g i t u d i n a l k ee l s ; m e t a n o t u m t r a n sverse l y s t r i a t e ; m e t a t ho r a x + p ro p o d e u m ( F i g . 1 1 85 A ) d u l l , g r a n u l a ted , w i t h p l e u ra a n d p o s te r i or s u rface t r a n sve rse l y s t r i a t e ; meso-me t a p l eu r a l s u t u re o b so l ete ; seg m e n t 4 of fro n t t a r s u s as l on g as or s l i g h t l y l on ge r t h a n seg m e n t 1 ( 37 : 3 5 ) ; en l a r g e d c l a w ( F i g . 1 1 85 B ) w i t h o u t s U b a p i c a l toot h , w i t h a sm a l l too t h a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d w i t h 5-8 peg - l i k e h a i rs ; s e g m e n t 5 o f fron t t a rs u s ( F i g . 1 1 85 B ) w i t h 2 rows o f 20-28 l a me l l ae e x t en d i n g b e y o n d 0 , 5 l e n g t h of s e g m e n t ; a p e x w i t h a g r o u p of a t l ea s t 1 2- 1 6 l a m e l l ae ; m a x i I l a r y p a l p i w i t h 5-6 segmen t s ; l ab i a l p a l p i w i t h 3 segme n t s ; t i b i a l spurs 1 , 0 , 1 . MALE : u n k nown L O C U S TYP I C U S : No g a l e s ( A r i z . , U . S . A . ) TYP I CAL MA T E R I A L : h o l o t y p e F ! of Ch . solitarius i n B ; ho l o t y pe F ! of Ch . xestocep h a l u s i n B ; l ec t o t y p e F ! a n d 3 p a r a l ec t o t y pes F F ! ( de s i g n a ted b y O l m i 1 9 82 ) of A . £errugin e u s i n B ; l ec t o t y p e F ! ( des i g n a ted b y O l m i 1 98 2 ) o f A . p icescens i n B . HOST S : u n k n o w n D I ST R I B U T I O N : N e a rc t i c reg i o n : U . S . A . : No g a l es ( A r i z . ) , B !

1 7 . Gon a top u s herb arum ( R . C . L .

P er k i n s )

n . com b .

1 705

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�F F i g . 1 1 86 - H e a d s of fem a l es of Gon a topus innitidus ( R . C . L . P e rk i n s ) ( A ) , ombrodes ( R . C . L . P e rk i n s ) ( B ) , a shmeadi K i effer ( D ) , a gropyru s F e n t o n ( E ) a n d herb arum ( R . C . L . P e rk i n s ) ( G ) ; meso t ho r a x a n d m e t a t ho r a x of fema l es of Gon a top u s mayori n . s p . ( C ) a n d contortulus P a \ ton ( F ) ; t h o r a x a n d p ro p o d e u m of fem a l e of Gon a top u s herb arum ( R . C . L . P e r k i n s ) ( H ) .

Cha lcogon a top u s herb arum R . C . L . P e r k i n s 1 907 : 23 .

F E MAL E : a p terou s ; l e n g t h 3 , 4 7-5 , 4 9 m m ; ferr u g i no u s , w i t h pet i o l e b l ac k a n d a n t en n a l s e g m e n t s 6- 1 0 b row n ; h e a d ( F i g . 1 1 86 G ) v e r y s l i g h t l y e x c a v a ted ,

a l most f l a t ,

g r a n u l a ted ; p rono t u m crossed b y a t ra n s vers a l

i m p ress i o n , s h i n y , w i t h o u t sc u l p t u re ; sc u t u m g r a n u l a te d , l a te ra l l y w i t h 2 p o i n t s ; m e t a n o t u m t r a n s v e rse l y s t r i a t e ; m e t a t h o r a x + p ropodeum ( F i g . 1 1 86 H ) w ea k l y g ra n u l a ted ; p l eu r a a n d pos t e r i o r s u rface of p ropodeum t r a n s verse l y s t r i a t e ; meso-m e t a p l e u r a l s u t u re o b so l e t e ; fore t a rsa l seg men t s 1 -4 i n f o l l ow i n g p ropor t i o n s : 1 4 : 3 : 5 : 1 6 ; en l a r ged c l a w ( F i g . 1 1 87 ) w i t ho u t s u b a p i c a l t oo t h , w i t h o r w i t ho u t a s m a l l too t h a t t h e e n d o f a

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1 1 88 - M a l e g e n i t a l i a of Gon a top u s h e r ba r u m ( R . C . L . Perk i n s ) from Somerset ( C a l i for n i a ) ( r i g h t v o l se l l a removed )

l o n g i t u d i n a l f u rrow a n d w i t h 5-8 peg- l i k e h a i r s ; seg men t S of f ro n t t a rs u s ( F i g . 1 1 87 ) w i t h i n ne r s i de p ro x i m a l l y n o t serra t e , w i t h t wo rows o f 1 7-34 l a m e l l ae e x t en d i n g b e y o n d 0 , 5 l e n g t h of s e g m e n t ; a p e x w i t h a g ro u p of 1 0- 1 7 l a m e l l ae ; m a x i I l a r y p a l p i w i t h 5-6 seg men t s ; l a b i a l p a l p i w i t h 3 seg men t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : f u l l y w i n g e d ; l e n g t h 1 , 8-2 , 0 m m ; b l ac k ; m a n d i b l e s p a r t l y y e l l o w ; l eg s n o t b a n ded ; a n t e n n ae n o t d i s t a l l y t h i c k e n e d ; a n t en n a I segmen t s i n fo l l o w i n g p roport i on s : 4 : 4 : 7 : 6 : 6 : 6 : 6 : 5 : 5 : 8 ; a n t e n n a I s e g m e n t 3 l e ss t h a n t h ree t i mes as l o n g as b ro a d ( 7 : 3 ) ; h e a d d u l l , g r a n u l a ted ; f r o n s w i t h a com p l e t e med i a n k ee l ; occ i p i t a l c a r i n a a b sen t ; occ i p u t conca v e ; P O L 5 ; O L = 1 , 5 ; OOL = 5 ; t e m p l es p ro m i ne n t ; s c u t u m d u l l , g r a n u l a ted ; n o t a u l i ces comp l e t e , poster i o r l y sep a r a ted ; s m a l l es t d i s t a n c e b e t w e e n n o t a u l i ce s n ea r l y a s l o n g a s b re a d t h of oce l l i ( 2 : 2 ) ; sc u t e l l u m a n d m e t a no t u m smoo t h , s h i n y , w i t h o u t sc u l p t u re , f i n e l y p u n c t a t e ; propode u m ret i c u l a t e r u gose ; fore-w i n g h y a l i n e , w i t ho u t d a r k t r a n s vers a l b a n d s ; r a d i a l v e i n w i t h d i s t a l p a r t m u c h l on ge r t h a n prox i m a l p a r t ; r a d i a l ce l l

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o p e n ; dors a l p rocess of gonoforceps ( F i g . 1 1 88 ) v e r y l o n g a n d n a rrow ; b a s i vo l se l l a w i t h n o t c h ; m a x i l l a r y p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 3 seg men t s ; t i b i a l s p u r s 1 , 1 , 2 .

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HOSTS : u n k no w n O I ST R I BUT I O N : N e a rc t i c reg i o n : U . S . A . : S . Berd i n o C o . ( C a l i f . ) , IVA ! A p p l e V a l l e y ( C a l i f . ) , O L ! OT ! Somerset ( E I Dorado C o . , C a l i f . ) , T E ' OL ! N og a l es ( A r i z . ) , B ! B M ! O L ! C a s a G r a n d e ( A r i z . ) , BM ' W a co ( M c Len n a n C o . , Te x . ) , BY ! 5 , 3 m i . N o f L a G r a n g e ( F a y e t te Co . , T e x . ) , OL !

18.

Gon a top us paralep ti a s U ' . C . L .

Perk i n s )

n.

com b .

Cha lcogon a top u s paralep tias R . C . L . Perk i n s 1 907 : 1 8 . Cha lcogon a top u s euprepes R . C . L . Perk i n s 1 907 : 1 8 ( n . sy n . ) ; locus typ i c u s Nog a l es ( A r i z . , U . S. A . ) . Cha lcogon a top u s euprepoides R . C . L . Perk i n s 1 907 : 1 9 (n . s y n . ) ; locus typicus Nog a l e s ( A r i z . , U . S . A . ) . Cha lcogo n a top u s cuprepes ( e rro r ) R . C . L . Perk i n s : K i effer 1 9 1 4 b : 99 . Cha lcogon a top u s cuprepoides ( e rror ) R . C . L . Per k i n s : K i effer 1 9 1 4b : 99.

F E MA L E : a p tero u s ; l en g t h 3 , 66- 5 , 1 2 m m ; head b r o w n or re d d i s h , w i t h m a n d i b l es , fron t p a r t of vertex a n d s i des of c l y pe u s y e l l o w ; a n t e n n ae b r o w n w i t h seg m e n t s 5-7 y e l l ow , or y e l l o w w i t h seg m e n t s 8- 1 0 or on l y 1 0 b row n ; p ro n o t u m t e s t aceo u s or b ro w n w i t h s i des l i g h t or redd i s h ; scu t u m b ro w n -y e l l o w or b l a ck o r redd i s h ; occ a s i on a l l y re d d i s h w i t h s i des b l ac k ; sc u te l l u m , m e t a t ho r a x + p ropodeu m , p e t i o l e a n d a b domen b l a c k ; l e g s fu l l y t e s t aceo u s or y e l l o w , w i t h coxae , femora a n d t i b i a e p a r t l y b row n ; head e xc a v a t e d , s h i n y , g r a n u l a ted ; p ron o t u m crossed b y a s t ro n g t r a n s versa l i m p ress i o n , g ra n u l a t ed ; m e t a t ho r a x + p ropodeum s h i n y , w i t h o u t sc u l p t u re ; r a re l y g r a n u l a t ed a n d w i t h a t r a c k of m ed i a n f u rrow ; p o s t e r i o r s u r face of p ropodeum a n d p l e u r a t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re ob so l e t e ; seg m e n t 4 of fro n t t a r s u s l o n ge r t h a n s e g m e n t 1 ( 4 5 : 40 or 46 : 3 5 or 44 : 39 ) ; e n l a rged c l aw ( F i g s 1 1 89 , 1 1 90 ) w i t ho u t s u b a p i c a l too t h , w i t h a s m a l l t oo t h a t t he e n d of a l on g i t u d i n a l f u r ro w a n d 7- 1 2 peg - l i k e h a i rs ; s e g me n t 5 of fro n t t a r s u s w i t h 1 ( F i g . 1 1 90 ) o r 2 row s ( F i g . 1 1 89 ) of 1 6-36 l a m e l l ae ; apex w i t h a g rou p of at l e a s t 1 2- 1 6 l a m e l l a e ; m a x i l l a ry p a l p i w i t h 6 s e g me n t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C US : Nog a l e s ( A r i z . , U . S . A . ) TYP I CAL MA T E R I A L : l ec t o t y p e F ! a n d 1 p a ra l ec t o t y p e F ! ( d es i g n a t ed b y O l m i 1 982 ) of Ch . p ara leptias i n B ; ho l o t y p e F I o f Ch . e u pre p es i n B ; ho l o t y p e F ! of Ch . euprepoides i n B . HOSTS : u n k no w n D I S TR I BU T I O N : N e a rc t i c re g i o n : U . S . A . : S a n J ac i n to M t s . ( I d y l l w i l d ,

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F E MA L E : ap terou s ; l e n g t h 3 , 9 1 -4 , 1 8 m m ; h e a d r e d d i s h or y e l l ow-redd i s h , w i t h b l ac k s p o t o n v e r t e x ; a n te n n ae y e l l o w-red d i s h o r b r o w n , w i t h segmen t s 1 -2 t e s t aceo u s ; p r o n o t u m y e l l o w-redd i s h ; sc u t u m , scu te l l u m , m e t a t h o r a x + p ropodeum a n d pe t i o l e b l a c k ; ab domen b l ac k o r t e s t aceo u s ; l e g s y e l l o w-redd i s h ; a n te n n ae d i s t a l l y t h i c k e n e d ; a n t e n n a I seg men t s i n fo l l ow i n g p ro p or t i o n s : 1 3 : 9 : 35 : 20 : 1 5 : 1 2 : 1 1 : 1 0 : 9 : 1 2 ; h e a d exca v a ted , s h i n y , g r a n u l a t ed ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 2 , 5 ; O L 2 ; O O L = 1 1 ; f ro n t a l I i n e com p l e t e ; p r o no t u m c rossed b y a s t ro n g t r a n s v e rs a l i m p ress i o n , g r a n u l a ted ; scu t u m d u l l , g r a n u l a t ed ; meso m e t a p l e u r a l s u t u re obso l e t e ; m e t a t h o r a x + p ro p o d e u m g r a n u l a ted ; p o s t e r i o r s u rface t r a n s v e rse l y s t r i a t e ; seg m e n t 4 o f f ron t t a rs u s l on g e r t h a n segmen t 1 ( 37 : 3 1 ) ; en l a r ged c l a w ( F i g . 1 1 9 1 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d o f a l o n g i t u d i n a l f u rrow a n d w i t h 1 1 - 1 4 p e g - I i k e h a i rs ; segme n t 5 o f f r o n t t a r s u s ( F i g . 1 1 9 1 ) w i t h 2-3 rows o f 32-56 l a m e l l ae e x t en d i n g b e y o n d 0 , 5 l e n g t h of segmen t ; a p e x w i t h a g r o u p of a t l ea s t 1 1 -30 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 6 segme n t s ; , l a b i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 . M AL E : u n k n o w n L O C U S T Y P I C U S : Co l u m b u s ( O h i o , U . S . A . ) TYP I C A L MAT E R I A L : l ec t o t y p e F ' ( d es i g n a t ed b y O l m i 1 982 ) i n B . HOSTS : i n U . S . A . Ph lepsius sp . ( R . C . L . Per k i n s 1 906c ; O l m i 1 982 ) . D I ST R I B U T I O N : N e a rc t i c reg i o n : U . S . A . : 4 m i . S of W a r ren ( T y l e r C o . , Tex . ) , O L ! 1 0 , 5 m i . N of H a l l et s v i l l e ( L a v ac a C o . , Tex . ) , B Y ! R e i s e l ( F a l l s Co . , T e x . ) , WA ! 3 , 8 m i . S o f Sea l y ( W a l l er C o . , Te x . ) , B Y ! 2 5 , 1 m i . N of A n g l e t o n ( B razo r i a Co . , Te x . ) , B Y ! C o l u m b u s ( O h i o ) , B ! B a r r e n I s . ( Lo n g I s l a n d , N . Y . ) , AM !

20 . Gon a top u s fl a vifrons A s h m e a d

Gon a topus fl a vifrons A s h m e a d 1 89 3 : 84 . Cha lcogon a top u s lep t i a s R . C . L . P e r k i n s 1 907 : 1 7 ( n . sy n . ) ; locus typicus Nog a l es ( A r i z . , U . S . A . ) . Cha lcogon a top u s fl a vifrons ( A s h m e a d ) : K ro m b e i n 1 9 56 : 1 59 .

F E MA L E : a p t e rou s ; l en g t h 3 , 37-4 , 50 m m ; h e a d b l ac k , w i t h c l y p e u s , m a n d i b l es a n d f ro n t p a r t of v e r t e x y e l l ow ; occ i p u t b l ac k or y e l l o w - b ro w n ; a n t e n n a e b r o w n w i t h s e g m e n t 1 y e l l ow , or y e l l o w w i t h segme n t s 8- 1 0 d a rk ; t horax a n d p ropodeum b l a c k ; i n some s p ec i m e n s p rono t u m p a r t l y red d i s h ;

Gon a topod i n a e

Fig.

1 71 3

1 1 92 - C he l a o f Gon a topus flavifrons A s h m e a d ( t y p e )

abdomen b rown-b l ac k ; l eg s y e l l ow - b ro w n , w i t h coxae , femora a n d t i b i a e brown i s h ; a n te n n a e d i s t a l l y t h i c k e n e d ; a n t en n a l se g m e n t s i n f o l l o w i n g p ropor t i on s : 8 : 7 : 1 6 : 1 4 : 1 0 : 8 : 6 , 5 : 6 : 5 : 7 , 5 ; h e a d s h i n y , e xc a v a ted , h a i r l es s , g ra n u l a ted ; occ i p i t a l c a r i n a i n v i s i b l e ; fro n t a l I i n e comp l e t e ; P O L = 2 ; OL = 2 ; OOL = 8 ; p rono t u m crossed b y a s t ro n g t r a n s vers a l i m press i on , w ea k l y g r a n u l a ted ; m e t a n o t u m t r a n s verse l y s t r i a t e ; m e t a t h o r a x + p ropod e u m d u l l , g r an u l a ted ; pos t er i or s u rface f u l l y t r a n sverse l y s t r i a t e ;

Mem . A me r . E n ! .

1714

Fig.

I n s ! . , n o . 37 ,

1 984

1 1 93 - C he l a of Con a to p u s r a p a x ( R . C . L .

Perk i ns ) ( type)

p l e u ra p a r t l y t r a n s v erse l y s t r i a t e ; meso-met a p l eu r a l s u t u r e obso l e t e ; se g m e n t 4 of fro n t t a r s u s l on g e r t h a n s e g m e n t 1 ( 30 : 26 ) ; en l a r g e d c l a w ( F i g . 1 1 92 ) w i t ho u t s u b a p i c a l too t h , w i t h a s m a l l t o o t h a t t he e n d of a l o n g i t u d i n a l f u r row a n d 8- 1 0 peg- l i k e h a i rs ; seg m e n t 5 of f ro n t t a r s u s

1 71 5

Gon a topod i n a e

( F i g . 1 1 9 2 ) w i t h 2 rows of 26-3 l a m e l l ae e i t h e r i n t e r r u p t ed or e x t en d i n g co n t i n uo u s l y t o a p e x ; a p e x w i t h a g rou p o f a t l e a s t 1 3-30 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 6 seg men t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MA L E : u n k no w n LOCUS TYP I CU S : A l bany ( N . Y . , T Y P I CA L M A T E R I A L : ho l o t y p e F ! p a r a l ec t o t y pes F F ! ( des i g n a t ed HOST S : u n k n o w n D I ST R I B U T I O N : N e a rc t i c reg i o n : ( A r i z . ) , O L ! F l a g s t a f f ( Ar i z . ) , ( A s h m e a d 1 893 ) .

U. S. A . ) of G . fl a vifron s i n WA ; b y O l m i 1 982 ) i n B .

l ec t o t y p e F !

and 2

U . S . A . : L a s Vegas ( N e v . ) , WA ! W i l l i a m s O L ! Nog a l es ( Ar i z . ) , B ' A l b a n y ( N . Y . )

2 1 . Gon a top u s rapax ( R . C . L .

Perk i n s )

n . comb .

Ch a lcogon a top u 5 rap a x R . C . L . Perk i n s 1 907 : 20 .

F E MA L E : a p terou s ; l en g t h 5 , 30 m m ; h e a d re d d i s h - y e l l ow , w i t h s m a l l b r o w n o c e l l a r s po t ; a n te n n a e red d i sh-ye l l o w , w i t h s e g me n t s 3- 1 0 w e a k l y d a r k e n e d ; p ron o t u m re d d i s h - y e l l o w ; scu t u m re d d i s h - y e l l ow , w i t h s i des b row n ; scu t e l l u m b ro w n ; m e t a t ho r a x + p ro p o d e u m b l a c k , w i t h a n t e r i o r s u rf a ce red d i s h ; d i sc w i t h a b ro w n s p o t ; a p e x of p ro p o d e u m redd i s h ; p e t i o l e a n d a b domen b l ac k ; l eg s redd i s h - y e l l o w ; h e a d exc a v a ted , s h i n y , g r a n u l a t ed ; p r o n o t u m crossed by a t r a n s vers a l i m p ress i o n , g r a n u l a t ed ; m e t a t h o r a x + p ro p o d e u m g r a n u I a ted ; p o s t e r i o r s u rface of p ropodeum t r a n s v e rs e l y s t r i a te on l y on p o s te r i or h a l f ; seg m e n t 4 of fore t a r s i s l i g h t l y l on g e r t h a n seg men t 1 ( 50 : 4 8 ) ; e n l a rged c l a w ( F i g . 1 1 9 3 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l t oo t h a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d 1 2 p e g l i k e h a i rs ; seg men t 5 of f r o n t t a rs u s ( F i g . 1 1 93 ) w i t h 2 rows of 3 + 21 l am e l l ae ( s h o r t e s t r o w n o t e x t e n d i n g b e y o n d 0 , 5 l e n g t h of s e g m e n t ) ; a p e x w i t h a g r o u p of a t l e a s t 20 l a me l l ae ; m a x i l l a r y p a l p i w i t h 6 seg men t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : N og a l es ( Ar i z . , U . S . A . ) TYP I CAL M A T E R I A L : l ec t o t y p e F ! ( des i g n a ted by O l m i 1 982 ) i n B . HOST S : i n U . S . A . n o t i de n t i f i e d C i c a d e l l i d s . D I S T R I BU T I O N : N e a rc t i c reg i o n : U . S . A . : No g a l e s ( A r i z . ) , B !

2 2 . Gon a topu5 a shme a di K i effer

nee Dryin u 5 � ieolor H a l i d a y 1 828 i n C u r t i s 1 8 28 : 206 . Gon a top u s ,"ieolor A s h m e a d 1 893 : 8 5 .

Mem . A m e r . E n ! .

1 71 6

Fig.

1 1 94

-

I n s ! . , n o . 37 ,

1 984

C he l a of Gon a top u s ashme a di K i effer from Sp r i n g C reek ( Georg i a )

1 71 7

Gon a topod i n a e

Gon a top u s ashmea di K i e ffer i n K i e ffer e t M a r s h a l l 1 905 : 1 08 ( ne w n am e ) . 1 Neogon a top u s erythrodes R . C . L . P e r k i n s 1 905 : 4 3 ( s y n . p roposed b y F re y t a g a n d B ac k 1 977 ) ; locus typicus C o l u m b u s ( U . S . A . ) . Neogon a top u s obscurissimus R . C . L . P e r k i n s 1 905 : 43 ( s y n . p roposed by F re y t a g a n d Back 1 977 ) ; locus typicus C o l u m b u s ( U . S. A . ) . Neogon a top u s brunnescens R . C . L . P e r k i n s 1 90 5 : 44 ( s y n . p roposed b y F re y t a g a n d Back 1 977 ) ; locus typicus O h i o ( U . S . A . ) . Gon a top u s ashmeadi K i effer : K i effer 1 907b : 2 1 . Gon a top u s b icolor A s h m e a d : K i effer 1 9 1 4 b : 1 22 . Gon a topus affinis F e n ton 1 9 1 8b : 266 ( s y n . p roposed b y F re y t a g a n d B a c k 1 977 ) ; l o c u s typ icus B a y V i ew ( O h i o , U . S . A . ) . Gona topus inimicis F e n ton 1 9 1 8b : 270 ( s y n . p roposed b y F rey t a g a n d B a c k 1 977 ) ; l o c u s typicus B a y V i ew ( O h i o , U . S . A . ) . Gon a top u s similis F e n ton 1 9 2 1 : 7 1 ( s y n . p roposed b y F rey t a g a n d B a c k 1 977 ) ; l o c u s typ icus A m es ( I ow a , U . S . A . ) . Epigon a top u s ten uis F e n ton 1 927 : 8 ( n . s y n . ) ; locus typicus L a f a y e t t e ( I n d . , U . S. A . ) . Gon a top u s b icolor A s h m ea d : F re y t a g a n d Back 1 977 : 222 .

F EMALE : a p t e rou s ; t e s t aceou s ,

l en g t h 1 -4 m m ; co l o u r v e r y v a r i a b l e ;

f u l l y red d i s h

w i t h p e t i o l e b l a c k a n d a n t e n n a I seg men t s 1 -2 y e l l ow ,

3- 1 0

b row n ; i n some s pec i me n s w i t h b ro w n oce l l a r spot ; somet i mes co l o u r redd i s h- b r ow n , w i t h head b ro w n ( w i t h y e l l o w m a n d i b l es a n d c l y pe u s ) , scu t u m red d i s h - b ro w n , scu t e l l u m a n d m e t a t h o r a x + p ropodeu m b l a c k ; head ( F i g . 1 1 86 D) exc a v a ted , s h i n y o r d u l l , w e a k l y g r a n u l a t e d ; p ron o t u m c rossed b y a t r a n s v e rs a l i m p ress i o n , w i t hou t sc u l p t u re or w e a k l y g r a n u I a te d ; s c u t u m d u I I , g r a n u I a t e d ; m e t a n o t u m a n d m e t a t h o r a x + p ropod e u m w i t h o u t scu l p t u r e or w e a k l y g r a n u l a t ed ( sC U l p t u re more v i s i b l e on p o s te r i or s u rface ) ; p l eu r a a n d pos t er i or s u r f ace of p ropode u m not t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re o b so l e t e ; segme n t 1 of fron t t a r s u s s l i g h t l y s h o r t e r t h a n segm e n t 4 ( 1 1 : 1 3 ) ; en l a r g e d c l a w ( F i g . 1 1 94 ) w i t hou t s u b a p i c a l t oo t h , w i t h a s m a l l too t h a t t he e n d of a l o n g i t u d i n a l

1 K i effer ( 1 90 5 ) con s i de red Gon a top u s b icolor A s h m e a d 1 893 p reoc c u p i e d b y Gon a top u s b icolor ( H a I i d a y 1 828 ) a n d p roposed t he n e w n a me Gona topus a shmeadi . T h i s i s , howev e r , a Case of seco n d a r y homonomy , b e c a u s e H a l i d a y desc r i bed h i s b icolor as Dryin u s a n d a c t u a l l y h i s /:; icolor i s b e l o n g i n g t o t h e g e n u s D icondyl u s . T h e A s h m e a d s p ec i es i s , o n t h e con t r a r y , i n t h e g e n u s Gon a top u s . Bicolor A s h m e a d s h o u l d h a v e s o p r i or i t y o v e r a s hmeadi K i effer . T h i s p rob l em shou l d h a v e , however , a n o t he r s o l u t i on on t he b as i s o f t h e r u l es o f zoo l og i c a l nomenc l a t u re . A s hmeadi K i effer and b icolor Ashmead a re i n fac t s y n on y m s . T h e s a m e K i effer ( 1 907b ) was t he f i rs t rev i se r and p l aced b icolor A s h m e a d as sy non y m of a sh me a di . G . a shmeadi so a c t u a I I y has p r i or i t y O V e r G . b icolor .

1 71 8

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Mem . A me r .

Ent.

I nst. ,

no.

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1 984

1 1 9 5 - M a l e ge n i t a l i a of Gon a top u s il s hmeadi K i e ffer from L e x i n g t on ( Kentuck y )

f u rrow a n d 4-7 p e g - l i k e h a i rs ; s e g m e n t 5 of fron t t a rs u s ( F i g . 1 1 94 ) w i t h 1 row o f 8-20 l a m e l l ae ( n ea r l y eq u a l i n l en g t h ) ; a p e x w i t h a g r o u p o f 6-8 l a m e l l a e ( 2-3 of t h ese v e r y l o n g ) ; m a x i l l a r y p a l p i w i t h 4-5 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E : f u l l y w i n ged ; l e n g t h 1 , 0-4 , 0 m m ; b l ac k or b r o w n i s h-b l ac k , w i t h t e g u l ae y e l l ow , m a n d i b l es y e l l ow , l e g s b a n ded ; a n t e n n a I s e g me n t s i n fo l l o w i n g p ropor t i o n s : 5 : 5 : 9 : 9 : 8 : 8 : 8 : 8 : 8 : 1 1 ; a n t e n n a I seg men t 3 more t h a n t h ree t i mes CIS l o n g as b road ( 4 ) ; h e a d h a i r y , s h i n y , g r a n u l a ted ; D O L � 8 ; D L � 3 ; DOL � 3 ; b r e a d t h o f t he oce l l i : 3 ; t e m p l es d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e ; scu t u m d u l l , g r a n u l a ted , h a i r y ; n o t a u l i ces comp l e t e , p o s t er i o r l y sep a r a t ed ; m i n i m u m d i s t a n c e b e t ween n o t a u l i ces shorter t h a n b re a d t h o f oce l l i ( 2 : 3 ) ; sc u t e l l u m and m e t a n o t u m s h i n y , smoot h , w e a k l y p u n c t a t e ; p ro p o d e u m ret i c u l a t e r u g ose , w i t h o u t k ee l s ; dorsa l p rocess of gonoforceps ( F i g . 1 1 9 5 ) m u c h s h o r t e r t h a n gonoforce p s , d i s t a l l y b ro a d en e d ; b a s i vo l se l l a w i t h o u t n o t c h ; ' m a x i l l a r y p a l p i w i t h 4 - 5 segmen t s ; l ab i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 1 , 2 . L O C U S T Y P I C US : Se l m a ( A l a b a m a , U . S . A . ) TYP I C A L MAT E R I AL : h o l o t y p e F ! of G . b icolor A s h m e a d i n \V A ; l ec t o t y pe F ' ( de s i g n a t ed b y O l m i 1 982 ) o f N . erythrodes i n B ; l ec t o t y p e F ' ( de s i g n a t ed b y D l m i 1 98 2 ) o f N . oh sc urissimus in B ; l ec t o t y p e F ! ( de s i g n a ted by D l m i 1 982 ) o f N. brunnescens i n B ; h o l o t y p e F ! a n d a l l o t y p e M ! o f G . a ffin is i n WA ; h o l o t y p e F ' o f G . inimicis i n \VA ; h o l o t y p e F ! o f G . similis i n WA ;

1 71 9

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�f \

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C h e l a of Gon a top u s ombrodes ( R . C . L . (Ohio)

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1 720

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1 984

h o l o t y pe F ! of E. ten u is i n WA . HOST S : i n U . S . A . Amblysell u s curtisii ( F i tc h ) ( F rey t a g a n d B a c k 1 977 ) A thys a n u s curtisii ( R . C . L . P er k i n s 1 906c ) ; Deltocep h a l u s sp . ? ( R . C . L . P e r k i n s 1 906c ) ; Deltocep h a l u s aifinis G . -B . ( F e n ton 1 9 1 8b , 1 92 1 ) ;

Deltocep h a l u s i n imicus ( S a y ) ( F en t o n 1 9 1 8b ) ; Graminella n igrifron s ( F orbes ) , Endria i n imica ( S ay ) , Ps ammotettix stri a t u s ( L . ) , Macrosteles fasc ifrons ( S t a I ) , L a ta l u s s ayi ( F i t c h ) , Deltocep h a l u s fl a vicosta ( S t a I ) ( F rey t a g a n d B a c k 1 9 77 ) . D I S T R I B U T I O N : N e a r c t i c reg i on : U . S . A . : Ames ( I o w a ) , WA ! L a f a y e t t e ( I n d . ) , WA ! C o l u m b u s ( O h i o ) , B ! B a y V i ew ( O h i o ) , WA ! Lex i n g ton ( K y . ) , O L ! Se l m a ( A l a . ) ( As h me a d 1 893 ) ; Sp r i n g C reek ( G a . ) , I T ! R a l e i g h ( N . C . ) , TW !

23 . Gon a top u s ombrodes ( R . C . L .

Perk i n s )

Neogon a top u s ombrodes R . C . L . P e r k i n s 1 905 : 43 . Gon a topus ombrodes ( R . C . L . Per k i n s ) : K i effer 1 9 1 4 b : 1 24 . Gon a top u s p u n c t a tu s Fen ton 1 9 1 8b : 268 ( sy n . p roposed b y F rey t a g 1 97 7 ) ; locus typ icus C o l u m b u s ( O h i o , U . S . A ) . nee Neogon a top u s ombrodes R . C . L . Perk i n s : S t r u b i n g 1 9 56 : 1 48 .

F E M AL E : a p terou s ; l e n g t h 2 , 45- 3 , 29 m m ; h e a d t es t aceou s , w i t h v e r t e x b l ack ; a n t e n n a e b ro w n , w i t h segmen t s 1 - 2 ye l l ow ; p r o n o t u m b l ac k , w i t h re d d i s h s p o t s on a n t e r i or m a r g i n , pos t e r i or m a r g i n , s i des a n d t r a n s v e r s a l i m p ress i o n ; scu t u m , sc u te l l u m a n d m e t a t h o r a x + p ropodeum b l ack ; a b domen b r o w n - b l ac k ; l e g s t e s t aceo u s ; head ( F i g . 1 1 86 B) e x c a v a ted , s h i n y , w e a k l y g r a n u l a t ed ; p ro n o t u m crossed b y a t r a n s v e r s a l i m p ress i o n , sh i n y , g r a n u l a te d ; scu t u m g r a n u l a t e d , w i t h some l o n g i t u d i n a l k ee l s a n d l a ter a l l y w i t h 2 p o i n t s ; met a n o t u m d u l l , t r a n sverse l y s t r i a t e , g r a n u l a ted ; m e t a t h o r a x + p ropodeum du I I , scu I p t u red as m e t a n o t u m ; p l e u r a a n d pos ter i o r s u r f a c e of p ropodeum n o t t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re o b so l e t e ; segmen t 1 o f f ro n t t a r s u s a s l o n g a s segm e n t 4 ; en l a r ged c l a w ( F i g . 1 1 96 ) w i t h o u t s u b a p i c a l too t h , w i t h a sma l l too t h at the end of a l on g i t u d i n a l f u rrow and 6 p e g - l i k e h a i r s ; segmen t 5 of fron t t a r s u s ( F i g . 1 1 96 ) w i t h 1 r o w of 1 4 l a me l l a e ( l a me l l ae l on g e s t n e a r p rox i m a l e n d of segmen t ) ; a p e x w i t h a g r o u p of 8 l a me l l a e ( 2 of t h ese v e r y l o n g ) ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l spurs 1 , 0 , 1 . MA L E : f u l l y w i n ged ; l e n g t h 2 , 0 -2 , 5 m m ; b l a ck o r b l ac k i s h - b r o w n o r b r ow n , w i t h a n t e n n a e b r ow n , m a n d i b l es a n d t e g u l ae y e l l o w ; l e g s n o t b a n d e d , y e l l ow , w i t h coxae b r o w n i s h ; a n t e n n a I segme n t s i n fo l l ow i n g p ropor t i o n s : 5 : 6 : 1 5 : 1 5 : 1 3 : 1 3 : 1 2 : 1 1 : 1 1 : 1 4 . a n ten n a I segmen t 3 more t h a n s i x t i mes a s l on g a s b r o a d ( 7 , 5 ) ; h e a d sh i n y , smoo t h , w i t h o u t scu l p t u re , w e a k l y p u n c t a t e , h a i r y ; fron t a l l i n e a b se n t ; P O L = 9 ; OL = 3 ; O O L = 3 ;

1 72 1

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M a l e ge n i t a l i a o f Gon a top us ombrodes ( R . C . L . Co l u m b u s ( O h i o ) ; N : n o t c h .

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b re a d t h of t h e oce l l i : 3 ; t e m p l e s d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e ; scu t u m h a i ry , g r a n u l a t e d , d u l l ; n o t a u l i ces com p l e t e , poster i o r l y sep a r a ted ; m i n i m u m d i s t a nce b e tween t h e n o t a u l i ces a p p rox i m a t e l y as l o n g as bre a d t h o f oce l l i ; sc u t e l l u m a n d m e t a n o t u m sh i n y , smoo t h , w i t h o u t sc u l p t u re , w e a k l y p u nc t a t e ;

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Mem . Amer . E n t .

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1 1 98 - C h e l a of Gon a top u s monilis n . sp .

( type)

h y a l i n e ; dorsa l process of gonoforceps ( F i g . 1 1 97 ) n ea r l y a s l o n g a s gonoforcep s , a p i c a l l y b ro a de n ed , p a d d l e- s h a p e d ; b a s i vo l se l l a w i t h n o t c h ; m a x i l l a r y p a l p i w i t h 4 s e g m e n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . LOCUS TYP I CU S : Col u m b u s ( O h i o , U . S . A . ) TYP I C A L M A T E R I AL : l ec t o t y p e F ! a n d 1 p a r a l ec t o t y p e F ! ( d es i g n a t ed b y O l m i 1 98 2 ) of N . om 'Jrodes i n B ; ho l o t y p e F ! , a l l o t y p e M ! a n d 1 p a r a t y ;o e F ! o f G . p u n c t a t u s i n IVA . HOST S : i n U . S . A . Cic a dula sexn o t a t a F a l l . ( e rror ) ( T ho m p son 1 9 54 ; A i n s l i e 1 920 ) ; L a t a l u s sayi ( F i t c h ) ( F rey t a g 1 97 7 ) = Deltocep h a l u s sayi ( R . C . L . Perk i n s 1 906c ) ; Ps ammotettix stri a t u s ( L . ) ( F re y t a g 1 977 ) . B I OLOG Y : i n form a t i on s i n A i n s l i e ( 1 9 20 ) . D I ST R I B U T I ON : N e a rc t i c reg i on : U . S . A . : W i scon s i n ( F rey t a g 1 977 ) ; I o w a ( F rey t a g 1 977 ) ; Co l u m b u s ( O h i o ) , B ! IVA ! K e n t u c k y ( F rey t a g 1 97 7 ) . P a l a e a rc t i c re g i on : t h e record from Germ a n y b y S t r u b i n g ( 1 9 56 ) i s erroneo u s .

1 72 3

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P e rk i n s ) f r o m T a n b a r k

24 . Gon a top u s monilis n . sp .

F E M.A l E : a p tero us ; l e n g t h 3 , 37 m m ; h e a d b l a c k , w i t h m a n d i b l es , c l y p e u s a n d m a I a r s p a ce t e s t aceou s ; a n te n n ae b row n ; t h o r a x a n d p ropodeum b l ac k , excep t for s i des a n d po s t e r i o r s u r face of p r o n o t u m w h i c h are red d i s h ; a b domen b l ac k ; l e g s t e s t aceou s , w i t h coxae p a r t I y b row n ; a n te n n ae d i s t a l l y t h i c k e n e d ; a n t en n a I seg men t s i n fo l l o w i n g p ropor t i o n s : 9 : 6 : 1 6 : 9 : 8 : 7 : 6 : 5 : 5 : 6 ; h e a d e x c a v a ted , d u l l , g r a n u l a t ed ; fro n s w i t h a med i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 2 ; Ol = 3 , 5 ; O O l = 9 ; p ro no t u m crossed b y a s t ro n g t r a n s vers a l i m p ress i on , s h i n y , w i t h co l l a r g r a n u l a t ed a n d d i sc w i t ho u t sc u l p t u re ; sc u t u m s h i n y , w i t h some l on g i t u d i n a l k ee l s ; m e t a no t u m h o l l o w b e h i n d sc u t e l l u m ; m e t a t h o r a x + p ropodeum d u l l , g r a n u l a ted , w i t h pos t e r i o r s u rface t r a n s verse l y s t r i a t e ; p l eu r a n o t t r a n s v erse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re d i s t i n c t a n d c o m p l e t e ; fore t a rs a l seg m e n t s i n fo l l o w i n g p roport i o n s : 1 3 : 3 : 5 : 1 9 : 26 ; en l a rged c l a w ( F i g . 1 1 98 ) w i t ho u t s u b a p i c a l too t h , w i t h a s m a l l t o o t h a t t h e e n d o f a l o n g i t u d i n a l fu rrow a n d w i t h 4 b r i s t l es ; s e g m e n t 5 o f fron t t a rs u s ( F i g . 1 1 98 ) w i t h 2 rows of 4 + 29 l a me l l a e ( s h o r t e s t row n o t

1 724

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Ent.

I ns t . ,

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1 984

e x t e n d i n g b e y o n d 0 , 5 l en g t h of segme n t ) ; a p e x w i t h a g ro u p of 1 4 l a m e l l a e ; m a x i I l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k nown L O C U S T Y P I C U S : E I Sa i t o ( Me x i c o ) T Y P I CAL M A T E R I AL : ho l ot y pe F ' i n O T . HOST S : u n k no w n D I S T R I BU T I O N : Nearc t i c reg i on : M E X I C O :

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F E MAL E : a p t erou s ; l e n g t h 3 , 00-3 , 66 m m ; red d i sh- y e l l ow , w i t h p e t i o l e a n d a b domen b l a c k ; a n t e n n a I s e g me n t s 8- 1 0 b ro w n i s h ; h e a d e x c a v a t e d , s h i n y , g r a n u I a t e d ; p r o n o t u m crossed b y a t r a n sversa I i m pres s i on , g r a n u I a t e d ; s c u t u m l o n g e r t h a n b roa d ; m e t a t h o r a x + p ropodeum g r a n u l a ted ; pos t e r i o r s u rface of p ropodeum t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re d i s t i n c t a n d comp l e t e ; s e g me n t 4 o f fron t t a r s u s s l i g h t l y shorter t h a n s e g me n t 1 ( 30 : 32 ) ; e n l arged c l a w ( F i g . 1 1 99 ) w i t ho u t s u b a p i c a l too t h , w i t h a sma l l t oo t h a t t h e e n d of a l o n g i t u d i n a l f u r row a n d 0-5 peg - l i k e h a i rs ; segmen t 5 of f ro n t t a rs u s ( F i g . 1 1 99 ) w i t h 1 row of 1 1 or 3 ( p rox i m a l ) + 1 9 l a me l l a e ; a p e x w i t h a g ro u p of 1 3- 1 4 l a me l l ae ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u r s 1 , 0 , 1 . M A L E : u n k no w n L O C U S T Y P I C U S : Nog a l es ( A r i z . , U . S . A . ) T Y P I CAL M A T E R I A L : l ec t o t y p e F ! ( d es i g n a t ed b y O l m i 1 982 ) i n B . H O ST S : u n k no w n D I ST R I BU T I O N : N e a rc t i c reg i o n : U . S . A . : T a n b a r k F l a t ( Los A n g e l es Co . , C a l i f . ) , O L ! Nog a l es ( A r i z . ) , B !

2 6 . Gon a top u s c u rriei K rombe i n

Gon a top u s curriei K ro m b e i n 1 962 : 4 .

F E MAL E : a p tero u s ; l en g t h 3 , 00-3 , 43 m m ; red d i s h - y e l l o w ; i n some spec i me n s w i t h a s m a l l b ro w n oce l l a r spot , p e t i o l e b l ac k a n d ab domen b ro w n -red d i s h ; a n t en n ae redd i s h - y e l l ow ; i n some spec i me n s w i t h segmen t 1 0 b row n ; a n t e n n a e d i s t a l l y t h i ck e n e d ; a n t e n n a I s e g m e n t s i n fo l l ow i n g propor t i on s : 9 : 5 : 1 7 : 1 0 : 7 : 7 : 5 : 5 : 5 : 8 ; h e a d e x c a v a t e d , s h i n y , w ea k l y g r a n u l a t e d , w i t h frons u n s c u l p t u red ; occ i p i t a l c a r i n a i n v i s i b l e ; fro n t a l l i n e comp l e t e ;

1 725

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w i t ho u t scu l p t u re ; seg m e n t 4 of f ro n t t a r sus l o n g e r t h a n segmen t 1 ( 27 : 22 ) ; e n l arged c l a w ( F i g . 1 200 ) w i t ho u t s u b a p i c a l toot h , w i t h a sma l l too t h a t t h e e n d o f a l on g i t u d i n a l f u rrow a n d w i t h 8- 1 1 peg - l i k e h a i rs ; segmen t 5 of fro n t t a rs u s ( F i g . 1 200 ) w i t h 1 row of 1 7 l a m e l l ae or 2 rows of 3 ( p ro x i m a l ) + 2 1 l a me l l a e ; a p e x w i t h a g ro u p of a t l e a s t 1 4 - 1 5 l a m e l l a e ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : P l u m m ers I s l a n d ( M d . , U . S . A . ) TYP I C A L MAT E R I A L : h o l o t y p e F ! a n d 1 p a r a t y pe F ! i n WA . H O ST S : u n k n o w n D I ST R I B U T I O N : Nearc t i c reg i o n : U . S . A . : Co l u m b i a ( Mo . ) , O L ! Lo r i n g ( V a . ) , WA ! P l u m mers I s l a n d ( M d . ) , IVA !

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1 72 7

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F E M AL E : a p t e ro u s ; l en g t h 2 , 56-3 , 4 3 m m ; red d i s h - t e s t aceo u s , w i t h p e t i o l e b l ac k a n d a b d omen b row n , h e a d ( F i g . 1 1 86 A ) a l most f l a t , d u l l , g r a n u l a t ed ; p ro n o t u m crossed b y a t r a n s v e r s a l i m p ress i o n , g r a n u l a t ed , d u l l ; sc u t u m s l e n de r , g r a n u l a ted ; m e t a n o t u m g r a n U l a t ed , n o t t r a n sverse l y s t r i a t e ; m e t a t h o r a x + p ropode u m d u l l , g r a n u l a t ed ; p l e u ra a n d pos t e r i o r s u rface of p ropod e u m n o t t r a n s v e r se l y s t r i a t e ; meso-m e t a p l e u r a l s u t u re d i s t i n c t a n d comp l e t e ; fore t a rsa l segmen t s i n fo l l ow i n g p ropor t i o n s : 1 5 : 2 : 5 : 1 5 : 23 ; en l a r g ed c l a w ( F i g . 1 20 1 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d 4-8 peg - l i k e h a i rs ; segmen t 5 of f ro n t t a r s u s ( F i g . 1 20 1 ) w i t h 1 row of 1 0 l a m e l l ae ; a p e x w i t h a g r o u p of 7-1 3 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MA L E : u n k n o w n L O C U S T Y P I C U S : No g a l es ( A r i z . , U . S . A . ) TYP I C A L M A T E R I A L : l ec t o t y p e F ' ( des i g n a t ed b y O l m i 1 98 2 ) of A . i n n i tidu s i n B ; h o l o t y p e F ! of A . s u t uralis i n 'NA . HOS T S : i n U . S . A . n o t i d e n t i f i e d C i c a d e l l i d s ( R . C . L . Perk i n s 1 907 ) . D I ST R I BU T l m� : Nearc t i c reg i o n : U . S . A . : No g a l es ( A r i z . ) , B ' T u cson ( A r i z . ) , WA !

1 7 28

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F E MAL E : a p t e rou s ; l e n g t h 3 m m ; h e a d t e s t aceo u s , w i t h a b ro a d b ro w n t ra n s v e r s a I b a n d o n v e r t e x ; a n t e n n a e b ro w n , w i t h segme n t s 1 -2 t e s t aceou s ; t h o ra x , p ropode u m , pet i o l e a n d a b domen b l ac k ; l eg s t e s t a c eo u s , w i t h c l u b s o f femora d a r k ; a n t e n n a e d i s t a l l y t h i c k e n e d ; a n t en n a l segmen t s i n fo l l ow i n g proport i o n s : 6 : 5 : 1 0 : 7 : 6 : 6 : 5 : 5 : 5 : 6 ; h e a d s l i g h t l y exca v a t ed , d u l l , g ra n u l a t e d ; frons w i t h a med i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 2 ; O L = 2 ; O O L = 7 p ro n o t u m crossed b y a t r a n s v e rsa l i m p ress i on , d u l l , g ra n u l a t e d ; sc u t u m d u l l , g r a n u l a t e d ; m e t a t ho r a x + p ropode u m d u l l , g ra n u l a t ed ; pos t e r i o r s u rface of p ro p o d e u m t r a n s v erse l y s t r i a t e o n l y on pos t e r i o r h a l f ; p l e u r a n o t t r a n sverse l y s t r i a te ; meso me t a p l e u r a l s u t u re d i s t i nc t a n d comp l e t e ; fore t a r s a l seg m e n t s i n fo l l o w i n g p ropor t i o n s : 1 1 : 2 , 5 : 4 : 1 2 : 1 9 ; e n l arged c l a w ( F i g . 1 202 ) w i t h o u t s u b a p i c a l too t h , w i t h a sm a l l too t h a t t h e e n d o f a l o n g i t u d i n a l fu rrow and w i t h 2 b r i s t l es ; seg m e n t 5 of f ron t t a r s u s ( F i g . 1 20 2 ) w i t h i n ner s i de p ro x i m a l l y n o t serra t e , w i t h 1 row of 20 l a me l l ae ; a p e x w i t h a g ro u p of 1 3 l am e l l ae ; m a x i l l a ry p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 2

1 7 29

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1 204 - Segme n t 5 of f ro n t t a rs u s of fem a l e o f Gon a top us p a l l idiceps ( R . C . L . P e r k i n s ) from F a rgo ( No r t h D a k o t a )

segme n t s ; t i b i a l s p u r s 1 , 0 , 1 . M A L E : u n k no w n L O C U S T Y P I C U S : S a s k a toon ( Sa s k a t c h ew a n , TYP I C AL M A T E R I A L : h o l o t y p e F ! i n OT .

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b row n , w i t h occ i p u t and f ro n t p a r t of v e r t e x t e s t aceou s ; a n t e n n ae y e l l o w , w i t h s e g m e n t s 4 - 1 0 b ro w n i s h ; p rono t u m b l ac k or y e l l o w or ferr u g i n o u s , u s u a l l y w i t h d i sc a n d m o s t p a r t of s i des b row n ; scu t u m , sc u te l l u m , m e t a t h o r a x + p ropod e u m , p e t i o l e a n d a b domen u s u a l l y b l a ck ; a p e x of p ropodeum r e d d i s h ; somet i mes scu t u m , s c u t e l l u m and m e t a to r a x + p ropodeum t e s t ac eo u s , w i t h b ro w n spo t s ; l eg s y e l l ow o r f e r ru g i n o u s ; h e a d exca v a ted , s h i n y , g r a n u l a t ed ; p r o n o t u m crossed b y a t r a n s ver s a l i m p ress i o n , s h i n y , g r a n u l a t e d ; me t a t ho r a x + p ropodeum d u l l , w e a k l y o r s t ro n g l y g r a n u l a ted ; p os t e r i or s u r f a ce of p ropodeum a n d p l eu ra t ra n sverse I y s t r i a t e ; meso m e t a p l e u ra l s u t u re obso l e t e o r w e a k l y d i s t i n c t ; fore t a rsa l s e g m e n t s i n fo l l ow i n g p ro p or t i o n s : 1 0 : 2 : 4 : 1 1 : 1 7 ; en l a r g e d c l a w ( F i g . 1 20 3 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d of a l o n g i t u d i n a l fu rrow a n d w i t h 2-7 p e g- l i k e h a i rs ; seg m e n t 5 of f ro n t t a rs u s ( F i g s 1 20 3 , 1 204 ) w i t h o n e row of 8-2 1 l a m e l l ae ; some t i mes t h i s row of l a me l l ae i s on d i s t a l h a l f o f segmen t ; a p e x w i t h a g r o u p o f 7 - 1 3 l a me l l a e ; m a x i l l a ry p a l p i w i t h 4-5 s e g me n t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 .

1 730

Mem . A m e r .

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1 984

MALE : u n k n o w n L O C U S TYP I C U S : A l a m e d a ( C a l i f . , U . S . A . ) TYP I C AL MAT E R I AL : l ec t o t y pe F ! ( de s i g n a t e d b y O l m i 1 982 ) i n B . HOST S : i n U . S . A . D e l t o c ep h a l u s fuscinervosus V . D . ( F rey t a g 1 97 7 )

A thys a n u s s p . ( R . C . L . Perk i n s 1 906c ) ; Endria in im ic a . D I S T R I B U T I O N : N e a rc t i c reg i o n : CANADA : C h e r r y v i l l e ( B . C . ) , T E ' She l b u r n e ( N . S . ) , OT ! u . S . A . : A l a m e d a ( C a l i f . ) , 3 ! D a n v i l l e ( C a l i f . ) , CA l F a rgo ( N . D . ) , OL! IV A ! L a w rence ( D o u g l a s C o . , K a n s . ) , OL ' J o h n son C i t y ( T e n n . ) , CA l Pa l k C a . ( F l a . ) , IVA !

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. 1 2 Me s o -me t a p l eu r a l s u t u r e ob s o l e t e . . . . . . . . . . . . . . . . . . . . . . . . . . S c u t u m l a t e r a l l y w i t h t wo p a i n t s ( F i g . 1 2 3 4 B ) . . . . . . . . 5 S c u t u m l a t e r a l l y w i t h o u t t wo p o i n t s ( F i g . 1 2 3 4 A ) . . . . . . . . . . . . . 9 S t a l k b e t w e e n p r on a t u m a n d m e t a t h a r a x + p r o p ad e u m v e r y s l en d e r , mo r e t h a n t w i c e a s l o n g a s p r an a t u m y e l l ow . . . . . . . . . . . S t a l k b e t w e en p r on o t u m a n d l e s s t h a n t w i c e a s l on g a s

b r oad ( F i g . 1 2 3 4 B ) ; s t a l k a n d . 3 . a gilis n . s p . m e t a t h o r a x + p r o p o d e u m l e s s s l en d e r , b r o a d ; s t a l k a n d p r o n o t u m mo s t l y

6

b l a ck An t e r i o r s u r f a c e o f m e t a t h o ra x + p r op o d e u m w i t h

7

f u r r ow a n d w i t h n u me r o u s t r a n s v e r sa l k e e l s ; m e t a n o t u m h o l l ow b eh i n d s c u t e l l u rn . . . . . . 5 . spinol a i O l m i A n t e r i o r s u r f a c e o f m e t a t h o ra x + p r op o d e u rn srno o t h , w i t h o u t s c u l p t u r e o r t r a ck s o f med i a n f u r r ow ; m e t a n o t u rn n o t h o l l ow b eh i n d s cu t e l l u m 7 Me s op l e u r a a n d me t a p l eu ra On d i f f e r e n t p l a n e s , b e c a u s e t h e k e e l

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6 t ra c k o f med i a n •

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1 73 1

Gon a topod i n ae

a l o n g t h e me s o -me t a p l e u ra l shaped •

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su t u r e i s v e ry b r o a d a n d 4 . p eruvianus

s t ep

8

. . . . . . . . . .8 Me s op l e u r a a n d me t a p l eu ra on t h e s a m e p l a n e . S c u t u m f u l l y b l a c k ; v e r t e x o f h e a d d u l l , g ra n u l a t ed . . . . . . . . . .

9

2 3 . b oli via n u s n . sp . S c u t u m a l mo s t f u l l y y e l l ow ; v e r t e x o f h ea d s h i n y , w i t h on l y o c e l l a r t r i a n g l e g r a n u l a t e d ; s i d e s o f v e r t e x s mo o t h a n d w i t h o u t s c u l p t u r e . . . . . . . . . . . . . . . . . 2 0 . arlei n . sp . Me t a t h o ra x + p r op od e u m w i t h a s t r o n g med i a n f u r r ow . •

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2 2 . contortus n . sp . p r op od e u m w i t h ou t med i a n f u r r o w . 1 0 T h o ra x a n d p r op od e u m r ed d i s h - t e s t a c e o u s . . . . . . . . . . . . . . . 6 . willinki n . sp . T h o r a x a n d p r op od eu m f u l l y o r mo s t I y b l a ck . 1 1 Me s op l e u r a a n d m e t a p l e u ra on t h e s a m e p l a n e ·

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. . . . . . . . 7 . p seudorb ita lis n . sp . Me s op l e u ra a n d m e t a p l eu ra on d l f f e r e n t p l a n e s , b e c a u s e t h e k e e l a l o n g t h e me s o - me t a p l eu ra l s u t u r e i s v e r y b r o a d a n d s t ep s h a p ed 8 . orb italis C a m e r o n 1 2 E n l a rg e d c l a w w i t h a p e x n o t p o i n t ed , r o u n d ed , w i t h ou t s ma l l t oo t h ( F i g s 1 2 1 5 , 1 2 1 6 , 1 2 1 7 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 3 E n l a r g e d c l a w w i t h a p e x p o i n t ed a n d w i t h s ma l l t o o t h ( F i g . ·

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1 218) 15 1 3 E n l a rg ed c l a w w i t h l a m e l l a e ( F i g . 1 2 1 5 ) ; t h o ra x a n d p r op od eu m du I I , g ranu I a t ed 9 . op a cithorax n . sp . E n l a rg ed c l aw w i t h o u t l a m e l l a e , w i t h b r i s t l e s ( F i g s 1 2 1 6 , 1 2 1 7 ) ; t h o r a x a n d p r op od e u m s h i n y , s m o o t h , w i t h ou t s c u l p t u r e . 1 4 1 4 H e a d w i t h OL n e a r l y a s l o n g a s P O L ; t h o r a x a n d p r op o d e u m m o s t l y r e d d i sh - t e s t a c e ou s d a rk . . 1 0 . p urp urascens n . sp . H ea d w i t h OL n ea r l y t w i c e a s l o n g a s POL ; t h o ra x a n d p r op od e u m mo s t l y b l a c k 1 1 . townesi n . sp . 1 5 S t a l k b e t we e n p r o n o t u m a n d m e t a t h o r a x + p r op od e u m v e r y l on g , l on g e r t h a n d i s c o f p r on o t u m ( F i g . 1 2 2 1 B ) . . . . . . . . . . . . . . . . . . . . •

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. 1 2 . apica lis C a m e r on S t a l k b e t w e en p r o n o t u m a n d me t a t h o ra x + p r op od e u m s h o r t , l on g a s , o r sh o r t e r t h a n d i s c o f p r o n o t u m ( F i g . 1 2 2 1 A ) . 1 6 T h o r a x a n d p r op od e u m mo s t l y y e l l ow - t e s t a c e o u s o r r e d d i s h T h o ra x a n d p r op od e u m f u l l y o r a l mo s t f u l l y b l a c k . . . . . 1 7 Me t a t h o r a x + p r op od e u m w i t h a med i a n f u r r ow . . . . . 2 1 . cilipes K i e f f e r Me t a t h o ra x + p r o p o d e u m w i t h o u t med i a n f u r r ow . •

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1 8 H ea d f I a t 1 3. H ea d e x c a v a t e d . 1 9 H e a d , t h o r a x a n d p r op od e u m f u I w i t h ou t s c u l p t u r e ( e x c e p t f o r H ea d , t h o r a x a n d p r o p od e u m d u I 2 0 S i d e s o f m e t a n o t u m r o u n d ed ( F i .

as . . . . . 16 . . . . 17 . . . . . 22 . . . . . .

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b artletti n . .

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I y o r a I mo s t f u I I y striae) . . I , g r a n u l a t ed . g . 1 22 1 A ) . . . . . . . . .

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smo o t h , . . . . . . .

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19 20

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1 7 32

Mem . Amer .

En!.

I n s! . ,

1 984

1 4 . fl a vipes n . s p . S i d e s o f me t a n o t u m p r o t r u d i n g 1 5 . camp b elli n . s p . P o s t e r i o r s u r f a c e o f p r op o d eu m f u l l y t ra n s v e r s e l y •

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st r i a t e. . . . . .

1 6 . tristis n . s p . P o s t e r i o r s u r f a c e o f p r op o d eu m w i t h o n l y p o s t e r i o r h a l f t ra n s v e r s e l y s t r i a t e . . . . . . 1 7 . doellojura doi ( O g l ob l i n ) 2 2 P o s t e r i o r su r f a c e o f p r op o d eu m s t r o n g l y t ra n s v e r s e l y s t r i a t e ; me t a t h o ra x + p r op od e u m w i t h o u t s c u l p t u r e a m o n g s t r i a e 1 9 . l a c u a li s n . s p . P o s t e r i o r su r f a c e o f p r op o d eu m w i t h o u t t ra n sv e r s a l s t r i a e , •

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w i t h on l y a f e w s t r i a e n ea r a p ex ; me t a t h o r a x + p r op o d e u m g ra n u l a t ed , d u l l 23 2 3 Me t a t h o ra x + p r op od e u m s l e n d e r , w i t h a n t e r i o r s u r f a c e s l i g h t l y i n c l i n ed ( F i g . 1 2 2 6 A ) 1 8 . s ilvestrii K i e f f e r Me t a t h o ra x + p r op o d e u m sq u a t , w i t h a n t e r i o r s u r f a c e m o r e i n c l i n ed ( F i g . 1 2 2 6 8 ) . . . . 2 4 . a u tumna lis n . s p . •

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MALES

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N o t au N o t au D o r sa 1 206 ) D o r sa 1 208 )

l i c e s c o mp l e t e . . 13. l i c e s i n c omp l e t e l p r o c e s s of g o n o f o r c e p s 1 . l p r o c e s s of g o n o f o r c ep s 2. .

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2 mu c h s h o r t e r t h a n p en i s ( F i g . fern a n di n a e n . s p . s l i g h t l y sh o r t e r t h a n p en i s ( F i g . breviforceps K i e f f e r •

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T h e m a l e s o f t h e o t h e r s p ec i es a r e u n k n ow n .

1 . Gon a top u s fern a n di n a e n . s p .

F E MAL E : a p terou s ; l e n g t h 1 , 87-2 , 00 m m ; t e s t a c eo u s , w i t h p e t i o l e b l ac k a n d a b domen p a r t l y b row n i s h ; h e a d f l a t , s h i n y , w i t ho u t scu l p t u re ; p ronot u m s h i n y , crossed b y a t r a n s ve rs a l f u rrow , w i t h o u t scu l p t u re , h a i r l es s ; s c u t u m d u l l , g r a n u l a ted ; m e t a n o t u m t r a n s v e rse l y s t r i a t e , met a t h o r o x + p ropodeum s h i n y , w e a k l y g r a n u l a ted ; p o s te r i o r s u rface of p ro p o d e u m a n d p l eu r a t r a n s v erse l y s t r i a t e ; meso-me t a p l eu r a l s u t u re obso l e t e ; segmen t 1 o f f ro n t t a rs u s n e a r l y t w i ce a s l o n g a s segmen t 4 ( 8 : 4 ) ; en l a rged c l a w ( F i g . 1 205 ) w i t ho u t s u b a p i c a l too t h , w i t h a s m a l l tooth a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d w i t h 5 p e g- l i ke h a i r s ; segmen t 5 of fron t t a rs u s ( F i g . 1 205 ) w i t h i n n er s i de p ro x i m a l l y serra t e , w i t h 2 r o w s of 8 + 9 l a m e l l ae ; a p e x w i t h a g r o u p of 7 l a me l l ae ; m a x i l l a r y p a l p i w i t h 4

G o n a topo d i n ae

Fig.

1 205 - C he l a of Gon a top u s fern a n din a e n . s p .

1 73 3

( t y pe )

segme n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : fu l l y w i n ged ; l en g t h 1 , 68 m m ; b ro w n - l i g h t ; a n ten n a l seg m e n t s i n fo l l o w i n g p roport i o n s : 4 : 3 5 , 5 ; 5 , 5 : 4 , 5 : 4 : 4 : 5 : 4 , 5 : 6 ; an t e n n a l segmen t 3 more t h a n t h ree t i mes a s l o n g a s b ro a d ( 3 , 5 ) ; head s h i n y , h a i r y , a l u t aceou s ; fro n t a l l i ne abse n t ; temp l es d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 7 ; OL = 3 ; OOL = 2 ; brea d t h of t h e oce l l i : 4 scu t u m

1 734

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Mem . Amer .

En\.

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n o . 37 ,

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1 206 - M a l e g en i t a l i a o f Gon a top u 5 fern a n din a e n .

sp .

from G a l a p agos

I s l ands

s h i n y , s h i n y g ra n u l a ted , h a i r y ; n o t a u l i ce s i n v i s i b l e ; scu te l l u m a n d m e t a n o t u m s h i n y a n d smoot h ; p ro p o d e u m smoo t h a n d sh i n y , w i t h o u t sc u l p t u re , w i t h a m ed i a n f u r row o n dors a l s u rface ; dors a l p rocess o f gonoforceps ( F i g . 1 20 6 ) short a n d s l e n d e r , w i t h a p e x rounded ; m a x i l l ary p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . LOCUS TYP I C U S : Fer n a n d i n a I s l a n d ( G a l a p a g os I s l a n d s , E c u a dor ) TYP I CAL MA T E R I A L : ho l o t y p e F ! a n d 1 p a r a t y p e F ! i n CA ; 2 p a r a t y p e s ( 1 F , 1 M ) ! i n OL. HOST S : u n k n o w n D I S T R I B U T I O N : GALAPAGOS I SLA N D S : F e r n a n d i n a I s l a n d , C A ! OL ! S a n t a C r u z I s l a n d , CA ! OL !

Gon a topod i n ae

Fig.

1 73 5

1 207 - C he l a o f Gon a top u s bre viforcep s K i e ffer ( t y p e )

2 . Gon a topus bre viforceps K i effer

Gon a top u s bre viforceps K i effer 1 904 a : Pa chygon a top u s brev i forceps K i effer ) :

356 . K i e ffer i n K i effer e t

M a rs a h a l l 1 906 : 50 . Cyrtogon a top u s breviforceps

( I< i effer ) : K i e ffer 1 907b : 1 9 . ( K i e ffer ) : K i effer 1 9 1 4 b : 70 . Cyrtogon a to p u s breviforceps ( K i e ffe r ) : B r u c h 1 9 1 5 : 443 . Netagon a top u s brasiliensis O g l o b l i n 1 9 32 : 264 ( n . s y n . ) ; l o c u s typ i c u s G u a y r a ( B r a z i I ) . Cyrtogo n a top u s breviforceps ( K i e ffe r ) : R i c h a r d s 1 969 : 80 . Cyrtogon a top us bre viforceps ( K i e ffe r ) : O l m i a n d C u rrado 1 976a : 1 65 . Cyrtogo n a top u s breviforceps

F E MA L E : a p terou s ; l e n g t h 2 , 4 3-3 , 00 m m ; b l ac k , w i t h fron t p a r t of v er t e x , c l y p e u s a n d m a n d i b l e s y e l l o w ; a n t e n n a e b ro w n , w i t h segmen t s 1 -3 y e l l ow ; l e g s y e l l o w , w i t h co xae , c l u b s of femora a n d b o t h a p exes of m i d a n d h i n d t i b i ae b row n ; h e a d f l a t , s h i n y , h a i r l es s , g r a n u l a te d ; p ro n o t u m crossed b y a t r a n svers a l i m p res s i o n , s h i n y , g r a n u l a ted ; scu t u m g r a n u l a ted , s h i n y ; m e t a n o t u m n o t t r a n s v erse I y s t r i a te ; m e t a t h o r a x + p ropodeum sh i n y , g r a n u l a te d ; p l e u r a a n d p o s te r i or s u rface of p ro p o d e u m n o t t r a n s v erse l y s t r i a t e ; meso-met a p l e u r a l s u t ure obso l e t e ; seg men t 1 o f front t a r s u s n e a r l y t w i ce a s l o n g a s segmen t 4 ( 1 24 : 7 ) ; en l a r ged c l a w ( F i g . 1 20 7 ) w i t ho u t s u b a p i c a l too t h a n d l a me l l ae , w i t h a s m a l l t o o t h a t t h e e n d of a l o n g i t u d i n a l fu rrow a n d 3-6 p e g - I i ke h a i r s ; s e g m e n t 5 of fron t t a r s u s ( F i g . 1 207 ) w i t h i n n e r s i de p ro x i m a l l y not serra t e , w i t h 2 r o w s of 8 - 1 2

1 736

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1 984

1 20 8 - M a l e gen i t a l i a of Gon a topus breviforceps K i e f f e r f ro m R i o P i e d r a s ( Pe r u } { r i g h t ha l f remo v e d )

v e r y s m a l l l a m e l l a e ; a p e x b roadened , w i t h a g r o u p of 6-8 l am e l l ae ; m a x i l l a ry p a l p i w i t h 3-4 segmen t s ; l a b i a l p a l p i w i t h 2 seg men t s ; t i b i a l s p u rs 1 , 0 , L MALE : fu l l y w i n ged ; l e n g t h 1 , 3 1 - 2 , 00 m m ; b l ac k ; m a n d i b l e s t e s t aceo u s ; a n te n n a e a n d l e g s n o t b a n de d , b row n ; a b domen b row n ; a n te n n a e n o t d i s t a l l y t h i c j k e ned ; a n te n n a l se g m e n t s i n fo l l o w i n g p ro p or t i o n s : 5 : 4 : 8 : 8 : 7 : 7 : 6 : 6 : 6 : 7 ; a n t e n n a l seg men t 3 more t h a n t h ree t i m e s a s l o n g a s

Gon a topod i n a e

1 737

b ro a d ( 3 , 25 ) ; h e a d s h i n y , w e a k l y g r a n u l a t e d ; fron t a l I i ne a b se n t ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 6 ; O L = 2 ; OOL = 2 ; b r e a d t h of t h e oce l l i : 4 ; sc u t u m w e a k l y g r a n u l a t e d ; not a u l i ces i n comp l e t e , reac h i n g a p p ro x i m a te l y 0 , 5 l e n g t h of sc u t u m ; sc u t e l l u m , m e t a n o t u m a n d p ropode u m smoo t h , s h i n y , w i t ho u t scu l p t u re ; dors a l s u r f ace o f p ropodeum w i t h a m e d i a n f u rrow ; fore w i n g s h y a l i n e , w i t ho u t d a rk t r a n sv e r s a l b a n d s ; d i s t a l p a r t o f r a d i a l v e i n l on g e r t h a n p ro x i m a l p a r t ; gon oforceps ( F i g . 1 208 ) w i t h a dors a l p rocess l o n g a n d n a r row ; b a s i v o l se l l a w i t ho u t notc h ; m a x i I l a r y p a l p i w i t h 3-4 segmen t s ; l ab i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 1 , 2 . L O C U S T Y P I C U S : S . t a C r u z ( A r g en t i n a ) T Y P I CAL M A T E R I A L : h o l o t y p e F ! of G. breviforceps i n G E ; h o l o t y pe F ! a n d 4 p a ra t y pes F F ! o f M. bra siliens is i n P T . HOST S : u n k n o w n D I ST R I BU T I ON : Neo t rop i c reg i on : BAHAMAS : New P rov i d ence , BM ! HA I T I : P t . P r i n ce , WA ! D O M I N I CA N R E P U B L I C : R a n c ho A rr i b a , O T ! E I R i o ( L a V e g a ) , O L ! O T ! P E R U : R i o P i e d r a s , O L ! WA ! BRAZ I L : E s c a d a ( P e r n a m b u co ) , B ! G u a y ra ( P a r a n a ) , P T ! A R G E N T I NA : S . t a C r u z ( P a t agon i a ) , GE !

3 . Gon a top u s agilis n .

sp .

F E MALE : a p terou s ; l e n g t h 4 , 0 m m ; h e a d t e s t aceou s ; a n te n n ae y e l l o w t e s t aceou s ; p ro n o t u m b l a c k or fu l l y tes t aceou s , w i t h red d i s h n u a nces on s i des ; sc u t u m and sc u te l l u m ye l l ow ; m e t a t ho r a x + p ropodeum b l ac k , w i t h a p e x r e d d i s h ; a bdomen b l ac k or re d d i s h - b ro w n ; l e g s test aceou s n , w i t h coxae a n d c l u b s of femora p a r t l y b rown or fu l l y tes t aceou s ; h e a d exca v a te d , s h i n y , w i t hou t scu l p t u r e ; p rono t u m crossed b y a s t r o n g t ra n s v e rs a l i m p ress i on , smoo t h , s h i n y , w i t ho u t scu l p t u re ; sc u t u m sh i n y , w i t h some l o n g i t u d i n a l k ee l s , l a t e ra l l y w i t h two po i n t s ( F i g . 1 234 a ) ; s t a l k b e tween p rono t u m a n d me t a t ho r a x + p ropodeum more t h a n t w i ce a s l o n g a s b roa d ; m e t a n o t u m s h i n y , n o t t r a n sverse l y s t r i a t e ; m e t a t ho r a x + p ropodeum s h i n y , smoo t h , w i t ho u t scu l p t u r e ; p l e u ra a n d pos ter i o r s u rface of p ropodeum t r a n s v e r se l y s t r i a t e ; meso-me t a p l eu r a l s u t u re d i s t i nc t a n d comp l e t e ; fo re t a rsa l seg m e n t s i n fol l ow i n g p ropor t i o n s : 1 1 : 2 : 3 : 1 3 : 1 9 ; en l a r g e d c l a w ( F i g . 1 209 ) w i t h o u t s u b a p i c a l toot h , w i t h a sma l l too t h a t t h e e n d of a l o n g i t u d i n a l f u r row a n d 8 peg- l i k e h a i r s ; seg m e n t 5 o f f ron t t a r s u s ( F i g . 1 209 ) w i t h 2 rows of 4 + 20 l a me l l ae ; apex w i t h a g roup of 1 3 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : T i b u ( C o l om b i a ) T Y P I CA L MAT E R I A L : h o l o t y p e F ! i n BM ; 1 p a ra t y p e F ! i n LE . HOST S : u n k n o w n D I ST R I BU T I O N : Neo t rop i c reg i o n : PANAMA : G a m b o a ( C a n a l Zone ) , L E ! COLOMB I A : T i b u ( M a r ac a i bo B a s i n ) , B M !

1 738

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1 209 - C h e l a o f Gon a top u s a gilis n . s p .

4.

( type )

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F E MALE : a p terou s ; l en g t h 3 , 2 5 mrn ; h e a d b row n , w i t h occ i p u t , c l y p e u s , frons a l o n g orb i t s a n d m a n d i b l es t e s t a c eo u s ; a n t e n n a e t e s t a ceo u s , w i t h segmen t s 6- 1 0 b row n ; t h o r a x a n d p ropodeum b l ac k ; scu t u m l i g l> ' , p e t i o l e b l ac k ; a b domen b l a c k - b ro w n ; l e g s b row n , w i t h t a r s i l i g h t ; h e a d exca v a ted , s h i n y , smoot h , w i t ho u t scu l p t u r e ; p ron o t u m crossed b y a t r a n s v e rs a l i m p ress i on , h a i r y , smoo t h , w i t ho u t SCU l p t u re ; scu t u m s h i n y , w i t h some l o n g i t u d i n a l k ee l s ; s t a l k b e t ween p ron o t u m a n d m e t a t h o r a x + p ropode u m l es s t h a n t w i ce as l o n g as b roa d ; me t a t h o r a x + p ro p o d e u m s h i n y , smoo t h , w i t h o u t scu l p t u re , h a i r y , w i t h n u merous t r a n s v e r s a l k ee l s o n pos t e r i o r s u rface ; p l e u r a w i t ho u t t r a n s v e r s a l k ee l s ; meso-me t a p l eu r a l s u t u r e d i s t i n c t a n d comp l e t e ; mesop l e u ra a n d met a p l e u r a o n d i f f e re n t p l a nes ; se g m e n t 1 o f fro n t t a r s u s s h o r t e r t h a n seg m e n t 4 ( 9 : 1 1 ) ; e n l a r ged c l a w ( F i g . 1 2 1 0 ) w i t ho u t s u b a p i c a l too t h , w i t h sm a l l too t h at the e n d of a

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( type)

s e g rren t 5 o f front t a r s u s ( F i g .

1 2 1 0 ) w i t h i n n e r s i de pro x i m a l l y serr a t e , w i t h 2 rows of 5 + 1 7 l a m e l l ae ; a p e x w i t h a g r o u p of 1 5 l a me l l a e ; m a x i l l a ry p a l p i w i t h 5 segmen t s ; l a b i a l p a l p ; w i t h 2 segmen t s ; l a b i a l s p u r s 1 , 0 , 1 . MAL E : u n k n o w n LOCUS TYP I C U S : T i ngo M a r i a ( Peru ) TYP I CAL MAT E R I A L : h o l o t y p e F ! i n C A . HOS T S : u n k n o w n D I ST R I B U T I O N : Neo t rop i c re g i o n : P E R U : T i n go M a r i a ( Monzon V a l l e y ) , CA '

5.

Con a top u s spin o l a i O l m i

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F EJ,lA L E :

a p t e ro u s ;

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l e n g t h 3 , 00 m m ;

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c l y p e u s , m a n d i b l es a n d fron t p a r t o f v e r te x ; a n t e n n a e t e s t aceo u s ; t h o r a x , p ropod e u m , p e t i o l e a n d a b domen b l ac k ; p ro n o t u m w i t h b ro w n s p o t s on s i des ; l e g s tes t aceo u s , w i t h cox a e , t i b i a e a n d c l u b s of femora b row n ; h e a d

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exca v a te d , s h i n y , g r a n u l a t ed ; p rono t u m crossed b y a t r a n s v e rs a l i mp ress i on , s h i n y , smoo t h , w i t hou t sc u l p t u r e ; sc u t u m s h i n y , w i t h some l o n g i t u d i n a l k ee l s ; sc u te l l u m s h i n y and smoo t h ; me t a t hor'ax + p ropodeum

1 742

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s h i n y , w i t h n u m erous t r a n s v e r s a ! k ee l s on p o s t e r i o r s u r face , p l e u r a a n d a n te r i o r s u r f ace ; a n t e r i o r s u r f a c e w i t h t ra c k of med i a n f u rrow ; m e t a n o t u m h o l l o w b eh i n d sc u t e l l u m ; meso-me t a p l e u r a l s u t u re d i s t i n c t a n d co m p l e t e ; fore t a rs a l segm e n t s i n fo l l o w i n g p ropo r t i o n s : 8 : 2 : 3 : 8 : 1 4 ; en l a r g e d c l a w ( F i g . 1 2 1 1 ) w i t ho u t s u b a p i c a l t oo t h , w i t h a s m a l l too t h a t t h e e n d o f a l o n g i t u d i n a l fu rrow a n d 8 p e g - l i k e h a i rs ; segmen t 5 of f r o n t t a rs u s ( F i g . 1 2 1 1 ) w i t h 2 r o w s o f 5-1 6 l a m e l l a e ; a p e x w i t h a g r o u p of 1 1 l a me l l ae ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l spurs 1 , 0 , 1 . MALE : u n k n o w n L O C U S T Y P I C U S : L a Seren a ( C h i l e ) TYP I C A L M A T E R I A L : ho l o t y p e F ! i n CA ; 1 p a r a t y p e F ! i n SP . HOST S : u n k n o w n D I ST R I B U T I O N :

N e o t r o p i c reg i o n :

CH I L E :

L a Seren a ,

6 . Con a to p u s w i l l i n k i n .

CA !

Ch i l e ,

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sp .

F E MA L E : a p t e ro u s ; l e n g t h 3 , 7 5 m m ; red d i s h , w i t h a n t e n n a I seg men t s 7 - 1 0 b ro w n i s h , p e t i o l e b l ac k , a b do m e n b l ac k ( e x c e p t for red d i s h n u a nces o n t e r g i t e 1 ) ; l e g s re d d i s h , w i t h c l u b s o f femora a n d c o x a e b ro w n i s h ; h e a d e xc a v a te d , d u l l , g r a n u l a te d ; p r o n o t u m crossed b y a t r a n s v e rs a l i m p ress i o n , s h i n y , w i t h o u t scu l p t u re ; sc u t u m s h i n y , w i t h some l o n g i t u d i n a l k ee l s , l a t e r a l l y w i t h o u t p o i n t s ; m e t a n o t u m f l a t , s h i n y , w i t h o u t scu l p t u re a n d t r a n s v e rs a l s t r i a e , not or s l i g h t l y ho l l o w b e h i n d sc u t e l l u m ; m e t a t h o r a x + p r o p o d e u m s h i n y , w i t h o u t s c u l p t u re ; me t a p l e u r a a n d p os t e r i o r s u r face of p ro p o d e u m t r a n sverse l y s t r i a t e ; mesop l e u r a n o t t r a n s v e rse l y s t r i a t e ; meso m e t a p l e u r a i s u t u re d i s t i n c t and comp l e t e ; fore t a rs a l seg men t s i n fo l l o w i n g p ro p or t i o n s : 1 2 : 2 : 4 : 1 6 : 24 ; en l a r ge d c l a w ( F i g . 1 2 1 2 ) I-V i t hou t s u b a p i c a l too t h a n d s m a l l too t h a t t h e e n d of a l o n g i t u d i n a l f u rrow , on l y w i t h 9 peg - l i k e h a i rs ; a p e x of en l a r g e d c l a w p o i n t e d a n d n o t b e n t a t r i g h t a n g l e ; se g m e n t 5 o f f ro n t t a rs u s ( F i g . 1 2 1 2 ) w i t h 2 rows of 7 + 20 l a m e l l a e ; a p e x w i t h a g r o u p of 1 0 l a m e l l ae ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 seg men t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k nown LOCUS TYP I C U S : T ucuman ( Argen t i n a ) TYP I CAL MAT E R I A L : ho l o t y p e F ! i n T U . HOST S : u n k n o w n D I ST R I B U T I O N : Neot rop i c reg i o n : I" RGE N T I NA : T u c u m a n , TIJ ! 1-1 0 T E S : t he spec i es i s n a med i n h o n o r of M r . A . W i l l i n k .

7 . Gon a top u s p s e u dorb i t a l i s n .

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v e r t e x , c l y p e u s a n d ma n d i b l e s y e l l o w ; a n t e n n ae y e l l ow , w i t h segme n t s 7- 1 0 b row n ; pron o t u m b l a c k , w i t h re d d i s h s i d es ; scu t u m , scu t e l l u m a n d m e t a t h o r a x + p ropodeum b l a c k ; abdomen b r ow n , w i t h redd i s h n u a nces ; l e g s red d i s h ; h e a d exc a v a t e d , s h i n y , w i t h o u t s c u l p t u r e , excep t f o r occ i p u t a n d f r o n t p a r t of v e r t e x g r a n u I a ted ; p r on o t u m c rossed b y a t r a n s v e r s a I i mp ress i o n , smoo t h , h a i r y , sh i n y , w i t h o u t s c u l p t u re ; s c u t u m s h i n y , w i t h

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some l o n g i t u d i n a l kee l s ; m e t a n o t u m smoo t h , n o t t r a n s v e r se l y s t r i a t e ; m e t a t h o r a x + p ropodeum sh i n y , smoo t h , w i t h o u t scu l p t u re ; p l e u ra a n d p o s t e r i o r s u r face o f p ropodeum t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re d i s t i n c t a n d comp l e t e ; mesop l e u r a a n d met a p l e u r a o n t h e s a m e p l a n e ; segm e n t 1 of f ro n t t a rs u s s h o r t e r t h a n segmen t 4 ( 1 3 : 1 9 ) ; e n l a r g e d c l a w ( F i g . 1 2 1 3 ) w i t ho u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d o f a l o n g i t u d i n a l fu rrow a n d 1 2 p e g -- l i k e h a i rs ; seg m e n t 5 of f ro n t t a rs u s ( F i g . 1 2 1 3 ) w i t h 2 rows of 4 + 2 3 l a me l l ae ; a p e x w i t h a g ro u p of a t l e a s t 1 3 l a me l l ae ; m a x i l l a ry p a l p i w i t h 6 s e g m e n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : S . J u a n ( Co s t a R i c a ) TYP I CAL MA T E R I A L : ho l o t y p e F ! i n OL ; 1 p a r a t y p e F ! i n A M . HOST S : u n k n o w n

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1 745

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Gona top us orb it alis C a m e ron 1 888 : 442 .

F E MALE :

a p tero u s ;

l e n g t h 2 , 56-4 , 06 m m ;

h e a d b l ac k or b row n ,

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a n d m a n d i b l es red d i s h - y e l l o w ; i n s o m e spec i mens a l so f rons p a r t l y testaceo u s ; a n te n n a e redd i s h , w i t h seg men t s 3 - 1 0 o r 7- 1 0 b row n ; t h or a x , p ropod e u m ,

pe t i o l e a n d a b domen b l ac k ;

l eg s b r o w n o r b l ac k o r t e s t a c eo u s ;

t a r s i l i g t h ; h e a d e x c a v a ted , d u l l , g r a n u l a ted ; i n some spec i me n s s h i n y a n d w i t ho u t sc u l p t u re ; p rono t u m crossed b y a t r a n s v ers a l i mp res s i on , s h i n y , smoo t h , w i t ho u t s c u l p t u re ; sc u t u m ( F i g . 1 2 34 A ) d u l l , w i t h SOme l o n g i t u d i n a l k e e l s ; m e t a n o t u m very short , smoo t h , not t r a n s verse l y s t r i a t e ; m e t a t h o r a x + p ro p o d e u m s h i n y , smoo t h , w i t h 0 L1 t scu l p t u re ; p l e u r a a n d poste r i o r s u r f a c e o f propod e u m t r a n s v e rse I y s t r i a t e ; meso- m e t a p l e u r a l s u t u re d i s t i n c t a n d comp l e t e ; mesop l e u r a a n d m e t a p l eu r a o n d i f f e ren t p l a nes , b ec a u se t h e kee l a l o n g t h e meso-me t a p l e u r a l s u t u re is very b road and s t e p - s h a p e d ; segmen t 1 of f ron t t a r s u s shorter than segmen t 4 ( 1 4 : 1 8 ) ; e n l a r g e d c l a w ( F i g . 1 2 1 4 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d of a l o n g i t u d i n a l f u r ro w a n d 8 - 1 1 l a m e l l ae , p l u s 1 b r i s t l e ; seg m e n t 5 of fron t t a r s u s ( F i g . 1 2 1 4 ) w i t h 2 rows o f 28-32 l a me l l ae ; a p e x w i t h a g r o u p of a t l e a s t 1 5-30 l a m e l l a e ; m a x i l l a ry p a l p i w i t h 6 segme n t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n l O C U S TYP I C U S : C h i r i q u i vo l c a n o ( P a n a m a ) TYP I CAL MA T E R I A L : h o l o t y p e F ! i n BM . H O ST S : u n k no w n D I ST R I B U T I O N : N eo t rop i c re g i on : PANA M A : C h i r i q u i v o l c a n o , BM ! C h i r i q u i H a r t m a n n s ( 5 . t a C l a r a ) , OT ! V E N E Z U E LA : S a n E s te b a n , WA I

9 . Gon a topus op a cithorax n . sp .

F E MA L E : a p terou s ; l e n g t h 3 , 62 m m ; h e a d t e s t aceous-d a r k , w i t h m a n d i b l e s t e s t a c eo u s ; a n te n n ae t e s t a c eo u s ; t h o r a x a n d p ro p o d e u m b l ack , w i t h s i de s o f p rono t u m test aceo u s - d a r k a n d w i t h pos t e r i o r h a l f of sc u t u m y e l l o w ; a p e x of p ropode u m test aceo u s ; a b domen b l ac k ; l e g s t e s t aceo u s , w i t h coxae p a r t l y b row n , fore femora a n d fore t i b i a e a l most f u l l y b ro w n , m i d a n d h i n d femora p a r t l y b row n ; a n t e n n a e d i s t a l l y t h i c k e n ed ; a n t e n n a I segmen ts i n fo l l o w i n g p ropor t i o n s : 1 0 : 7 : 28 : 1 5 : 1 2 : 1 1 : 9 : 7 , 5 : 6 : 1 0 ; h e a d s h i n y , esc a v a t e d , smoo t h , w i t h o u t sc u l p t u re ; f rons w i t h an i n com p l e t e med i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 2 ; Ol = 2 , 5 ; O O l = 8 , 5 ; p ron o t u m

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d u l l , crossed b y a s t ro n g t r a n svers a l i m p ress i on , g r a n u l a ted ; sc u t u m s h i n y ; met a n o t u m f l a t , g r a n u l a t e d ; m e t a t h o r a x + p ropodeum d u l l , g ra n u l a te d , w i t h p o s te r i o r su rface t r a n s v erse l y s t r i a te ; meso-me t a p l e u r a l s u t u re obso l e te ; sc u t e l l u m w i t h t w o l a tera l p o i n t s ; fore t a rs a l s e g me n t s i n fo l l o w i n g p ropor t i o n s : 1 6 : 4 : 8 : 1 7 : 30 ; en l a r ged c l a w ( F i g . 1 2 1 5 ) w i t ho u t s u b a p i c a l too t h , w i t h 3 b r i s t l e s a n d 6 l a me l l ae ; segmen t 5 of fro n t t a rs u s ( F i g . 1 2 1 5 ) w i t h i n n er s i de p ro x i m a l l y not serra t e , w i t h 1 row o f 6 l a m e l l a e ; a p e x w i t h a g r o u p of n e a r l y 23 l a me l l a e ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MAL E : u n k n o w n L O C U S T Y P I C U S : M e t a t e ( Me x i co ) TYP I C AL MAT E R I A L : ho l o t y p e F ! i n T W . HOST S : u n k n o n w D I ST R I B U T I O N : N e o t rop i c reg i o n : M E X I C O : M e t a t e ( T u x tepec , O a x a c a ) , TW !

1 0 . Gon a top u s p u rp urascens n .

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F E M AL E : a p t e rou s ; l e n g t h 2 , 7 5 m m ; re d d i s h - t e s t aceo u s , w i t h p e t i o l e b l ac k a n d a b domen d a r k ; a n t e n n a e d i s t a l l y t h i c k e n e d ; a n te n n a l seg m e n t s i n fo l l o w i n g p ropor t i o n s : 7 : 6 : 1 9 : 1 0 : 8 : 7 : 6 : 6 : 5 : 7 ; head e xc a v a ted , s h i n y , smoo t h , w i t h o u t sc u l p t u re ; fro n s w i t h w e a k med i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 2 ; O L = 2 ; OOL = 9 ; p ron o t u m crossed b y a s t ro n g t r a n s v e rs a l i m p res s i o n , s h i n y , smoo t h , w i t h o u t scu l p t u re ; sc u t u m w i t h o u t l a te r a l p o i n t s ; meso t ho r a x a n d m e t a t ho r a x + p ropodeum s h i n y , smoo t h , w i t ho u t sc u l p t u re , excep t for pos t e r i o r s u r face o f p ropodeum a n d p a r t of me t a p l e u r a t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re obso l e t e ; fore t a rsa l segmen t s i n fo l l o w i n g proport i o n s : 1 3 : 3 : 6 : 1 2 : 23 ; en l a rged c l a w ( F i g . 1 2 1 6 ) w i t ho u t s u b a p i c a l too t h , w i t h o u t sm a l l tooth a t t he e n d of a l o n g i t u d i n a l f u r row , w i t h a p e x n o t po i n ted , rou n ded , w i t h 1 row of 5

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F E MA L E : a p terou s : l e n g t h 2 m m ; h e a d b l ac k , w i t h m a n d i b l es t e s t aceou s ; a n t e n n ae t e s t a ceou s ; t h o r a x a n d p ro p o d e u m b l ac k , w i t h s i d e s of pron o t u m t e s t a c eo u s ; a b domen b row n - t e s t aceou s ; l e g s t e s t a ceou s ; a n t e n n a e d i s t a l l y t h i c k e n e d ; a n t e n n a l se g m e n t s i n p ropor t i o n s : 6 : 5 : 1 2 : 7 : 7 : 6 : 5 : 4 : 4 : 5 , 5 ; h e a d sh i n y , smoo t h , exc a v a t e d , w i t h o u t sc u l p t u re ; f r o n s w i t h o u t med i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; t e m p l e s p rom i n e n t ; P O L = 1 , 5 ; O L 3 ; O O L = 6 ; p rono t u m crossed b y a s t ro n g t r a n sv e rs a l i m p ress i o n , s h i n y , smoo t h , w i t h ou t scu I p t u re ; scu t u rn , rne t a n o t u rn a n d m e t a t h orax + p ro p o d e u m sh i n y , smoot h , w i t h o u t sc u l p t u re , exce p t for s o m e t r a n sversa l s t r i a e o n poster i o r s u r f ace of p ropod eu m ; rneso-met ap l e u r a l s u t ure obso l e t e ; sc u t e l l u m a n d m e t a n o t u m w i t h o u t l a t e r a l po i n t s ; fore t a rs a l seg men t s i n f o l l ow i n g propor t i o n s : 1 0 : 2 : 5 : 9 : 1 6 ; en l a r g e d c l a w ( F i g . 1 2 1 7 ) w i t ho u t s u b a p i c a l t oo t h , w i t h a p ex rou n d e d , w i t h 4 b r i s t l es ; s e g m e n t 5 o f f r o n t t a r s u s ( F i g . 1 2 1 7 ) w i t h i n n er s i d e p rox i m a l l y not serra t e , w i t h t w o rows o f 2 =

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F E M A L E : a p t erou s ; l e n g t h 4 , 37 m m ; h e a d b l a ck , w i t h f ro n t p a r t of ver tex , c l y p e u s an d m a n d i b l es y e l l ow ; a n t e n n ae y e l l ow , w i t h segme n t s 8- 1 0 b l a ck ; t h o r a x a n d p ro p o d e u m b l a ck ; a b domen b row n-b l ac k ; l eg s y e l l ow , w i t h coxae a n d c l u b s of femora p a r t l y b l ac k or b r ow n ; h e a d exca v a t ed , sh i n y , smoo t h , w i t h o u t scu l p t u re ; p ro no t u m crossed b y a t r a n sversa l i m p res s i o n , sh i n y , smoo t h , w i t h o u t sc u l p t u re j scu t u m sh i n y , smoo t h , w i t h o u t scu l p t u r e , l a ter a l l y w i t h 2 p o i n t s ; s t a l k b e t ween p rono t u m a n d met a t ho r a x + p ropod e u m ( F i :1 . 1 22 1 B ) l on g er t h a n d i sc of p rono t u m ; met a n o t u m smoo t h , n o t t r a n s v e rse l y t r a n sverse l y ; rn e t a t h o r a x + p ropodeum smoo t h , sh i n y , w i t h o u t sc u l p t u re ; p l e u r a n o t t r a n sverse l y s t r i a t e ; pos t e r i o r su rface of p ropod e u m t r a n sverse l y s t r i a t e ; meso-met a p l e u ra l s u t u re obso l e t e ; seg m e n t of f ro n t t a r s u s s h o r t e r t h a n segmen t 4 ( 1 8 : 2 2 ) ; en l a rg e d c l a w ( F i g . 1 2 1 8 ) w i t h o u t s u b a p i c a l toot h , w i t h a sma l l too t h a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d 9 p e g - I i k e h a i rs ; segmen t 5 o f f ron t t a r s u s ( F i g . 1 2 1 8 ) w i t h 2 rows of 7 + 1 8 l a me l l ae ; apex w i t h a g ro u p of 9 l a m e l l a e ; m a x i I l ary p a l p i

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F E MAL E : a p t e ro u s ; l e n g t h 2 , 62 m m ; y e l l ow - t e s t a c eo u s , w i t h p e t i o l e b l ack and a b domen d a r k ; a n t e n n a e d i s t a l i y t h i c k e n e d ; a n t e n n a I seg m e n t s i n fo l l ow i n g p ropo r t i o n s : 7 : 4 : 1 2 : 8 : 6 : 6 : 5 : 5 : 5 : 7 ; h e a d f l a t , d u l l , g r a n u l a t ed ; f r o n s w i t h a med i a n k ee l ; POL � 1 ; O L � 2 ; OOL � 6 ; occ i p i t a l c a r i n a i n v i s i b l e ; p r o no t u m crossed b y a t r a n s v e r s a l i m p ress i on , d u " , g ra n u l a t e d ; scu t u m , sc u t e l l u m , m e t a n o t u m a n d m e t a t h o r a x + p ropodeu m d u l l , very w e a k l y g r a n u l a t ed ; m e t a n o t u m n o t t r a n s v erse l y s t r i a t e ; meso m e t a p l e u r a l s u t u re o b so l e t e ; pos t e r i o r s u rf a c e of p ropode u m t r a n sverse l y s t r i a t e ; p l e u r a n o t t r a n s v e r se l y s t r i a t e ; sc u t u m w i t h o u t l a t e r a l po i n t s : fore t a rsa l s e g me n t s i n fo l l ow i n g p ropor t i on s : 1 4 : 2 : 3: 1 1 : 1 6 ; en l a r g e d c l a w ( F i g . 1 2 1 9 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l t o o t h a t t h e e n d of a l o n g i t u d i n a l f u r row a n d w i t h 3 b r i s t l e s j s e g m e n t 5 of f ro n t t A r s u s ( F i g . 1 2 1 9 ) w i t h i n n e r s i d e prox i m a l l y w e a k l y serra t e , w i t h 2 rows o f 8 + 1 4 I a me l i a e ex t e n d i n g b e y o n d 0 , 5 l e n g t h of segmen t a n d p a r t I y l oc a t ed o n a p ro m i n e n ce ; a p ex w i t h a g r o u p of 1 3 l a me l l a e ; m a x i l l a ry p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . M AL E : f u " Y w i n ged ; l e n g t h 2 , 1 2 - 2 , 7 5 m m ; b l a c k or b ro w n , w i t h l eg s n o t b a n d e d , t e s t aceou s ; c o x a e b r ow n ; a n te n n ae n o t d i s t a l l y t h i c k e n e d ; a n te n n a l se g m e n t s i n fo l l ow i n g p ropo r t i o n s : 5 : 4 : 9 : 9 : 9 : 9 : 7 : 7 : 6 : 8 , 5 ; a n t en n a l se g m e n t 3 more t h a n t h ree t i mes a s l o n g a s b r o a d ( 6 ) ; h e a d h a i r y , s h i n y , w e a k l y g r a n u l a t e d ; f ro n t a l l i n e comp l e t e ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L � 6 , 5 ; OL � 2 , 5 ; OOL � 1 , 5 ; b r e a d t h of t h e oce l l i : 5 ; scu t u m s h i n y , g r a n u l a ted ; n o t a u l i ces comp l e t e , p o s t e r i o r l y se p a r a t e d ; m i n i m u m d i s t a n ce b e t ween t h e n o t a u l i ces s h o r t e r t h a n a n t e n n a l se g men t 2 ( 2 : 4 ) ; sc u t e l l u m a n d rne t a n o t u rn s h i n y , smoo t h , w e a k l y p u n c t a t e , w i t h o u t s c u l p t u re a m o n g p u n c t u re s ; p ropodeurn s h i n y , a l mo s t smoo t h , w e a k l y s t r i a te , w i t h a med i a n f u rrow o n dor s a l s u rf a c e ; fo re-w i n g h y a l i n e , w i t h o u t d a r k t r a n s v e r s a l b a n d s ; g o n oforceps ( F i g . 1 220 ) w i t h d o r s a l p rocess l o n g a n d s l e n d e r ; b a s i vo l se l l a w i t h o u t n o t c h ; m a x i l l a ry p a l p i 4 s e g m e n t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u rs 1 , 1 , 2 . L O C U S TYP I C U S : M a y a g uez ( P u e r t o R i c o ) TYP I C A L M A T E R I A L : h o l o t y p e F ! a n d 5 p a r a t y p es ( 1 p a r a t y p es ( 1 F , 1 M ) ! i n O L .

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H OST S : i n P u e r t o R i co Baldulus m a idis ( De l . a n d W . ) ( B a r t l e t t 1 939 ) . D I ST R I BU T I O N : Neot rop i c reg i o n : P U E R TO R I C O : M a y a g ue z , WA ! DL ! N OT E S : t h e spec i es i s n a m e d i n honor of M r . K . A . B a r t l e t t . Accord i n g to B a r t l e t t ( 1 939 ) t h e p a ras i t i z a t i on of Baldulus ma idis by G. b artletti a r r i ved to 1 5 percen t . The effec t i v e n e s s of G . b a rtletti was somew h a t reduced b y a h y p e r p a r a s i t e , Ooencyrtus sp . , w h i ch a t t ac k e d t h e cocoo n s .

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F E MAL E : a p t e ro u s ; l e n g t h 2 , 75-2 , 93 mm ; y e l l ow - t e s t aceou s , w i t h p e t i o l e b l ac k a n d a b domen brow n i s h , h e a d exc a v a ted , sh i n y , g r a n u l a t e d , excep t for f ro n s w i t h o u t sc u l p t u re ; p ron o t u m crossed by a t r a n sver s a l i m p ress i o n , smoo t h , sh i n y , w i t h o u t scu l p t u re ; sc u t u m sh i n y , w i t h o u t scu l p t u r e ; s t a l k b e t w ee n p rono t u m a n d m e t a t h o r a x + p ro p o d e u m ( F i g . 1 22 1 A ) shorter t h a n d i sc of prono t u m ; m e t a n o t u m smoo t h , n o t t r a n sv erse I y s t r i a te ; m e t a t h o r a x + p ropodeum sh i n y , smoo t h , w i t ho u t sc u l p t u re ; p l e u r a a n d p o s te r i or su rf ace of p ro p o d e u m t r a n s v erse l y s t r i a t e ; meso-me t a p l eu r a l s u t u re obso l e t e ; segme n t 1 of f ro n t t a r s u s shorter t h a n seg m e n t 4 ( 1 0 : 1 2 ) ; en l arged c l a w ( F i g . 1 2 2 2 ) w i t hou t s u b a p i c a l too t h , w i t h a sm a l l tooth at t h e e n d of a l o n g i t u d i n a l f u rrow a n d 5-7 p e g - I i k e h a i r s ; segmen t 5 of f ro n t t a r s u s ( F i g . 1 22 2 ) w i t h 2 r o w s o f 1 3-23 l a m e l l ae ; a p e x w i t h a g rou p o f 9 - 1 2 l a me l l a e ; m ax i l l a ry p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l spurs 1 , 0, 1 . M AL E : u n k n o w n L O C U S TYP I C U S : L a P a l m a ( Co s t a R i c a )

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TYP I C A L M A T E R I AL : h o l o t y p e F ! i n OL ; 1 p a r a t y p e F ! i n B ; 1 p a r a t y p e F ! i n WA . HOST S : u n k no w n D I ST R I B U T I O N : Neot rop i c reg i o n : COSTA R I CA : L a P a l m a , OL ! J A M A I C A : Mon y m u s k E s t a t e , WA ! E C UADOR : B a n o s , B !

1 5 . Gon a top us c a mp b elli n . sp .

F E MALE : a p t e ro u s ; l e n g t h 3 m m ; t e s t aceou s , w i t h a n te n n a I s e g rnen t s 5-8 b row n , v e r t e x of h e a d d a r k , pet i o l e b l ack , poste r i o r t h i r d of each a b dom i n a l se g m e n t d a r k , b o t h e x t re m i t i es of m i d a n d h i n d femora b row n ; a n t e n n a e d i s t a l l y t h i c k e n e d ; a n t e n n a I seg m e n t s i n fo l l o w i n g p ropor t i o n s : 6 : 5 : 1 5 : 9 : 6 : 6 : 5 : 4 : 4 : 6 , 5 ; head sh i n y , exca v a t ed , w i t h o u t s c u l p t u re ; f ro n s w i t h a med i a n k e e l ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 1 , 5 ; OL = 1 , 5 ; OOL = 6 ; p ron o t u m c ross sed b y a s t ro n g t r a n s v e rs a l i m press i o n , smoo t h , s h i n y , w i t ho u t sc u l p t u re ; sc u t u m d u l l , w e a k l y g r a n u l a te d , w i t h 2 l a t e r a l

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p o i n t s ; m e t a n o t u m sh i n y , t r a n s v e rse l y s t r i a t e , w i t h 2 l a t e r a l po i n t s ; m e t a t h o r a x + p ropodeum sh i n y , w i t h o u t scu l p t u r e , except for p o s te r i o r s u rface a n d p l e u r a w h i c h are t r a n s v e r se l y s t r i a t e ; meso-me t a p l e u r a l s u t u re o b so l e t e ; fore t a r s a l se g m e n t s i n fo l l ow i n g p roport i o n s : 1 3 : 3 : 4 : 1 4 : 20 ; e n l a r g e d c l a w ( F i g . 1 22 3 ) w i t ho u t su bap i c a l too t h , w i t h a s m a l l too t h a t t h e e n d of a l o n g i t u d i n a l fu rrow a n d w i t h 8 peg- l i k e h a i r s ; s e g m e n t 5 o f f r o n t t a r s u s ( F i g . 1 22 3 ) w i t h i n n er s i de prox i m a l l y n o t serra t e , w i t h 2 rows of 1 + 7 l a me l l a e e x t e n d i n g b e y o n d 0 , 5 l en g t h of segmen t ; apex w i t h a g r o u p of 1 2 l a m e l l ae ; m a x i l l a r y p a l p i w i t h 4 se�' r r , e n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MAL E : u n k n own L O C U S TYP I C U S : R e s . Mon teverde ( Cos t a R i c a ) T Y P I CAL MAT E R I AL : h o l o t y p e F ! i n � T . H O ST S : u n k n o w n D I ST R I B U T I ON : N e a rc t i c reg i o n : C OSTA R I CA : Res . M o n t e v e r d e , " OT E S : t h e spec i es i s n a med i n honor o f M r . J . M . C a m p b e l l .

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1 6 . Gon a top u s tristis n . sp .

F E MAL E :

a p tero u s ;

l e n g t h 3 , 25 m m ;

y e l l ow - t e s t aceo u s w i t h pe t i o l e b l ac k ;

head exc a v a ted , d u l l , h a i r l es s , g r a n u l a t ed ; p ro no t u m crossed by a t r a n sv e rs a l i m p ress i on , d u l l , g r a n u l a ted ; sc u t u m d u l l , g r a n u l a t ed ; s t a l k b e t w ee n p r o n o t u m a n d m e t a t h o r a x + p ropod e u m s h o r t e r t h a n d i sc of p ronot u m j m e t a no t u m d u I I , t r a n s verse I y s t r i a t e ; meta t h o r a x + p ropod e u m d u l l , g r a n u l a ted ; p l e u r a a n d poste r i o r su rface o f p ropode u m t r a n sverse l y s t r i a te ; meso-met a p l e u r a l su t u r e obso l e t e ; s e g m e n t 1 of f ro n t t a r s u s shorter than se g m e n t 4 ( 1 3 : 1 5 ) ; en l a r g e d c l a w ( F i g . 1 22 4 ) w i t h o u t s u b a p i c a l toot h , w i t h a sma l l too t h a t t h e e n d of a l o n g i t u d i n a l f u r row a n d 6 p e g - l i k e h a i rs ; se g m e n t 5 of f ro n t t a rs u s ( F i g . 1 2 24 ) w i t h 2 r o w s of 3 + 2 0 l a m e l l ae ; apex w i t h a g r o u p of 7 l a me l l ae ; m a x i l l a ry p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 seg men ts ; t i b i a l sp u r s 1 , 0 , 1 . M AL E : u n k no w n l O C U S TYP I C U S : L a P a l m a ( C o s t a R i c a ) TYP I C A L M AT E R I AL : h o l o t y p e F ! i n Ol . HOSTS: u n k nown D I ST R I B U T I O N : Neot rop i c reg i o n : C O S T A R I C A : L a P a l m a , O l !

1 7 . Gon a top u 5 doellojura doi ( O g l ob l i n ) n . com b .

Dicon dyl u s doello-jura doi O g l ob l i n 1 9 38 : 38 .

F E M AL E :

a p t e ro u s ;

l e n g t h 4 mm ;

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C he l a of Gon a top u 5 doellojura doi ( Og l o b l i n ) ( t y p e )

abdomen brow n , a n t e n n a I segme n t s 4- 1 0 b ro w n , v e r t e x of h e a d w e a k l y d a r k ; a n t e n n a e d i s t a l l y t h i ck e n e d ; a n t e n n a I segmen t s i n fo l l ow i n g p ropor t i on s : 6 : 5 : 1 5 : 8 : 8 : 6 : 5 : 5 : 5 : 7 ; h e a d exc a v a t e d , d u l l , w e a k l y o r s t r o n g l y g r a n u l a ted ; f r o n s w i t h a med i a n k ee l ; occ i p i t a l c a r i n a i n v i s i b l e ; POL = 2 ; OL = 3 ; OOL = 8 ; p ron o t u m crossed by a s t ro n g t r a n s v e r s a l i m p ress i o n , d u l l , w e a k l y g r a n u l a t ed ; scu t u m d u l l , g r a n u l a ted ; s c u t u m a n d m e t a n o t u m w i t h o u t I a t e r a I poi n t s ; m e t a n o t u m du I I , n o t t r a n s v e r se l y s t r i a te , g r a n u l a t ed ; m e t a t h o r a x + p ropodeum d u l l , g r a n u l a ted ; pos ter i o r s u rf ace w i t h t r a n s v er s a l s t r i ae on l y on post e r i o r h a l f ; p l e u r a n o t t r a n s v e r se l y s t r i a t e ; meso-me t a p l e u r a l s u t u re o b so l e t e ; f o r e t a r s a l segmen t s i n fo l l ow i n g p ropor t i o n s : 1 3 : 3 : 3 , 5 : 1 1 : 20 ; en l a r g e d c l a w ( F i g . 1 225 ) w i t h o u t s u b a p i c a l too t h , w i t h a sm a l l too t h a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d w i t h 7 peg - l i k e h a i r s ; segmen t 5 o f f ro n t t a rs u s ( F i g . 1 2 2 5 ) w i t h o n e row of 25 l a me l l a e ; a p ex w i t h a g r o u p of 1 0 l a me l l ae ; m a x i l l a ry p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 .

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Gon a topus sil vestrii K i effer f rom Rezende ( B r a z i I ) ( A )

MALt=. : u n k n o w n L O C U S TYP I C U S : T u c u m a n ( A rge n t i n a ) TYP I C AL MAT E R I AL : ho l o t y p e F ! i n B A . HOST S : u n k n o w n D I ST R I B U T I O N : N eo t ro p i c reg i o n : ARG E N T I N A :

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1 8 . Gona top u s silvestrii K i e ffer

Gon a topus s i l v e s trii K i effer 1 9 1 2b :

1 74 . K i effer 1 9 1 4 b : 1 28 . Gon a topus sil vestrii K i effer : B r u c h 1 9 1 5 : 443 . Gon a top us c a rettei B r u c h 1 9 1 5 : 443 ( n . s y n . ) ; l o c u s typ i c u s S. Ju l i an ( S . ta Cruz, Argent i na ) .

Gon a topus silvestrii K i eff e r :

F E MA L E : ap terou s ; l e n g t h 2 , 87-3 , 6 2 mIT! ; h e a d b row n - b l ac k , w i t h f r o n t p a r t o f v e r t e x , c l y p e u s a n d m a n d i b l es y e l l ow ; a n t e n n a e b row n , w i t h segmen t s 1 - 2 ye l l ow ; p ron o t u m b l ac k , i n s o m e spec i me n s w i t h b rown n u a n ces o n d i sc ; sc u t u m , sc u t e l l u m a n d m e t a t h o r a x + p ropode u m b l ac k , w i t h apex o f p ropode u m l i g t h ; a b domen b ro w n -redd i sh ; l e g s redd i sh , w i t h coxae a n d c l u b s of f e m o r a p a r t l y b row n ; h e a d exc a v a t e d , g r a n u l a te d , h a i r l e ss ; p ro no t u m crossed b y a s t r o n g t r a n s v e r s a I i m p res s i on , g r a n u I a ted ; sc u t u m d u l l , g r a n u l a ted ; m e t a t h o r a x + p ropod e u m ( F i g . 1 226 A ) d u l l , - � a n u l a t ed ;

1 760

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p l e u r a a n d pos ter i o r s u rf ace of propodeum not t r a n s v e rse l y s t r i a t e or w i t h some s t r i a e o n l y near a p e x ; rneso-m e t a p l e u r a l s u t u re obso l e t e ; s e g me n t 1 of f ro n t t a r s u s a p p rox i m a te l y a s l o n g a s seg m e n t 4 ; e n l arged c l a w ( F i g .

1 2 27 ) w i t h o u t s u b a p i c a l toot h , w i t h a s m a l l too t h a t t h e e n d of a l on g i t u d i n a l f u rrow a n d w i t h 6-7 pe g - I i k e h a i r s ; s e g m e n t 5 of fro n t t a r s u s ( F i g . 1 2 2 7 ) w i t h t w o rows o f 1 9 -25 l a m e l l a e e x t e n d i n g b e y o n d 0 , 5 l e n g t h of segmen t ; apex w i t h a g rou p of n e a r l y 9 l a m e l l a e ; m a x i l l a ry p a l p i w i t h 4-5 segmen t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k n o w n L O C U S TYP I C U S : S . t a C r u z ( Ar g en t i n a ) TYP I CAL MAT E R I AL : l ec to t y p e F ! ( h ereby des i g n a ted ) of G . sil vestrii i n P O ; l e c t o t y p e F ! a n d 1 p a r a l ec t o t y p e F ! ( h ereby d e s i g n a ted ) o f G . carettei in P T . HOST S : u n k n o w n D I S T R I B U T I O N : Neotrop i c reg i o n : A R G E N T I N A : S . t a C r u z ( P a t a g o n i a ) , P O ! S . J u l i a n ( S . t a C r u z , P a t a gon i a ) , P T ! BRAZ I L : R e z e n d e ( R i o d e J a n e i ro ) , B ! NOT E S : t h e l ec to t y p e of G . s il vestrii i s i n b a d cond i t i o n s ( i t w a s k e p t i n a l co h o l t h ro u g h o u t 70 y e a r s ! t h e c o l o u r i s now tes t a ceou s ) .

1 9 . Gon a top u s l a c u a l i s n . sp .

F E M A L E : a p t e ro u s ; l e n g t h 3 , 43 m m ; h e a d redd i s h - te s t aceo u s , w i t h b row n spot on oce l l a r reg i on ; a n t e n n ae brow n , w i t h se g m e n t s 1 - 2 y e l l ow ; p ro no t u m redd i s h , w i t h med i a n brow n s p o t s on c o l l a r a n d a n t e r i or m a r g i n o f d i sc ; scu t u m , s c u t e l l u m , m e t a t h o r a x + propod e u m a n d a b d omen b l a c k ; l e g s b l a c k , w i t h t roc h a n te r s , fore t i b i ae a n d t a r s i red d i s h ; h e a d exc a v a te d , s h i n y , g r a n u l a t e d , excep t f o r frons w i t h ou t s c u l p t u re ; prono t u m crossed b y a t r a n s v e rs a l i m press i o n , s h i n y , g r a n u l a ted ; scu t u m g r an u l a t e d , w i t h some l o n g i t u d i n a l k ee l s ; s t a l k b e t w een pro n o t u m a n d m e t a t h o r a x + p ropodeu m shorter t h a n d i sc of pron o t u m ; m e t a n o t u m s t ro n g I y t r a n s v e rse l y s t r i a te ; m e t a t horax + propod e u rn s h i n y , w i t h o u t s c u l p t u r e , w i t h t r a c k of med i a n f u rrow ; p l e u r a a n d p o s t e r i o r s u rface o f p ropod e u rn t r a n s v e rse l y s t r i a t e ; meso-rne t a p l eu r a l s u t u re o b so l e t e ; seg men t 1 of fro n t t a r s u s as l o n g as s e g m e n t 4 ; en l arged c l a w ( F i g . 1 228 ) w i t hou t s u b a p i c a l toot h , w i t h s m a l l too t h at t h e e n d of a l o n g i t u d i n a l f u rrow a n d 7 peg- l i k e h a i r s ; se g m e n t 5 of fro n t t a r s u s ( F i g . 1 2 28 ) w i t h 2 row s of 3 + 2 0 l am e l l a e ; apex w i t h a group of a t l e a s t 1 1 l a m e l l a e ; m a x i l l ar y p a l p i 4 s e g me n t s ; l a b i a l p a l p i w i t h 2 segmen ts ; t i b i a l s p u r s 1 , 0 , 1 . MAL E : u n k n o w n L O C U S TYP I C U S : V i l l a r i c a L a k e ( C h i l e ) TYP I C A L MAT E R I A L : h o l o t y p e F ! i n OL . HOST S : u n k n o w n D I S TR I B U T I O N : I'Jeo t rop i c reg i o n : C H I L E : V i l l a r i c a L a k e ( P u o i n ) , OL !

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F E MAL E :

a p t e ro u s ;

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l e n g t h 2 , 87 m m ;

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p r o t h o r a x b l ac k ;

s c u t u m y e l l o w ; m e t a t h o r a x + p ro p o d e u m b l ac k ; pe t i o l e b l ac k ; a b domen b row n - t e s t aceo u s ; legs t e s t a c eo u s ; a n t e n n a e d i s t a l l y t h i c k e n e d ; a n ten n a l segmen t s i n fo l l ow i n g p ropo r t i o n s : 7 : 5 : 1 5 : 5 : 4 , 5 : 5 : 4 , 5 : 4 : 4 ; h e a d exca v a t e d , sh i n y , w i t h o u t scu l p t u r e ; f r o n s w i t h a med i a n ke e l ; o c c i p i t a l c a r i n a i n v i s i b l e ; P O L � 2 , 5 ; OL � 2 ; OOl � 8 ; p ro n o t u m c rossed b y a t r a n s v e r s a l i m p ress i o n , sh i n y , smoo t h , w i t h o u t scu l p t u re ; sc u t u m d u l l , w i t h some l o n g i t u d i n a l k ee l s ; m e t a n o t u m smoo t h , w i t h o u t t r a n sv e r s a l k ee l s ; meso-me t a p l e u r a l s u t u re d i s t i n c t a n d comp l e t e ; mesop l e u r a a n d m e t a p l e u r a on t h e s a me p l a ne ; me t a t h o r a x + p ropodeum s h i n y , smoo t h , w i t h o u t sc u l p t u re , except for p l e u r a a n d pos t e r i o r s u rface w h i c h a re t r a n s v erse l y s t r i a te ; fore t a rsa l segmen t s i n fo l l ow i n g p ropo r t i o n s : 1 2 : 2 : 4 : 1 3 : 1 9 ; en l a r g e d c l a w ( F i g . 1 22 9 ) w i t h o u t s u b a p i c a l t oo t h , w i t h a sm a l l too t h a t t h e e n d of a l o n g i t u d i n a l f u rrow ; segmen t 5 of f r o n t t a r s u s ( F i g . 1 229 ) w i t h i n n er s i d e p rox i m a l l y se r r a t e , w i t h 2 rows of 1 4 l a me l l ae ; apex w i t h a gro u p of 1 0 l a m e l l a e ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segm e n t s ; t i b i a l s p u r s 1 , 0 , 1 . MAL E : u n k n o w n L O C U S TYP I C US : N o v a T e u t o n i a ( B r az i l ) TYP I CAL MAT E R I AL : h o l o t y p e F ! i n O T . H O ST S : u n k n own D I ST R i b U T I O N : Neot rop i c reg i o n : B R A Z I L : N o v a Teu t o n i a , N O T E S : t h e spec i e s i s n a m ed ; n honor of M r . R . Ar I e .

OT !

2 1 . Gon a top us cilipes K i effer

Gon a topus cilipes K i effer 1 904a : 355 . Neogon a topus cilipes ( K i e ffer ) : O l m i a n d C u r r a d o 1 976a :

1 75 .

F E MALE : a p te ro u s ; l e n g t h 3 , 8 mm ; h e a d b r ow n , w i t h occ i p u t , c l y p e u s , m a n d i b l e s a n d f ro n t p a r t o f v e r t e x red d i s h - y e l l ow ; a n t e n n ae y e l l ow re d d i s h , w i t h seg m e n t s 4-6 bro w n i s h ( l a s t segme n t s m i s s i n g in t h e on l y k n own spec i men ) ; p ro n o t u m red d i s h , w i t h a sma l l b ro w n s p o t o n co l l a r ; d i sc of p ro n o t u m w e a k l y b r o w n i s h ; sc u t u m a n d s c u t e l l u m r e d d i s h ; m e t a t h o r a x + p ro p o d e u m re d d i s h , weak l y bro w n i s h o n d i sc , met a p l e u r a a n d a n te r i o r s u r f a c e o f p ropo d e u m ; p e t i o l e b row n ; a b domen b r ow n , w i t h re d d i s h n u a nces ; l e g s y e l l ow - red d i s h ; a n t en n a l seg men t s i n fo l l ow i n g p ropor t i on s : 1 0 : 6 : 20 : 8 : 8 : 7 ( o t h er s e g me n t s m i s s i n g i n t h e o n l y k n o w n spec i me n ) ; h e a d exc a v a t e d , h a i r l es s , w i t h p a r t of f r o n s w i t h o u t scu l p t u re ; occ i p u t a n d v e r t e x g r a n u I a t e d ; POL � 1 , 5 ; O L � 3 ; OOL � 1 0 ; p ro n o t u m c rossed b y

G o n a topod i n a e

Fig.

1 230

-

C h e l a of Gon a top u s cilipes K i effer ( t y p e )

1 765

1 766

Mem . Amer .

Fig.

1 231

a s t ro n g t r a n s ver s a l

En! .

I ns! . ,

no . 37,

1 984

- C h e l a of Gon a top u s contortu s n .

i m p ress i o n ,

h a i ry ,

smoo t h ,

sh i n y ,

sp .

( t y pe )

w i t hou t scu l p t u r e ;

sc u t u m h a i r y , g r a n u l a ted ; s c u t e l l u m sh i n y , smoo t h , h a i ry ; m e t a n o t u m rugo s e , h a i r l e ss ; m e t a t h o r a x + p ropodeum w e a k l y g r a n u l a t ed , w i t h med i a n fu rrow ; pos t e r i o r s u rface of p ropodeum t r a n sverse l y s t r i a t e ; meso m e t a p l e u r a l s u t u re obso l e t e ; en l a r ged c l a w ( F i g . 1 230 ) w i t hou t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d 7 - 8 pe g - l i k e h a i rs ; se g m e n t 5 of fro n t t a r s u s ( F i g . 1 2 30 ) w i t h 2 row s o f 4 + 22 l a me l l ae be g i n n i n g at prox i m a l t h i r d of segmen t ; apex w i t h a grou p of 1 4 l a me l l a e ; m a x i l l a ry p a l p i w i t h 4 seg men t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . M AL E : u n k n o w n L O C U S TYP I C U S : P . t o 1 4 d e M a y o ( P a r a g u a y ) TYP I C A L M A T E R I AL : h o l o t y p e F ! i n G E . H O ST S : u n k n o w n D I ST R I B U T I O N : Neotrop i c reg i o n : PARAGUAY :

P . to 1 4 de M a y o ,

GE !

Gon a to pod i n ae

Fig.

1 232

-

C h e l a of Gon a topus bolivia n u s n . s p .

1 767

( t y pe )

1 768

Mem . A mer .

En t .

I n st . ,

no.

37,

22 . Gon a t op u s contortus n .

F E MAL E :

ap terou s ;

l e n g t h 2 , 20 m m ;

redd i sh-d a r k ,

1 984

sp .

w i th a n tennae,

m a n d i b l e s , t a r s i a n d s t a l k s o f femora l i g h t ; a n te n n a e d i s t a l l y t h i c k ened ; a n t e n n a I segmen t s i n fo l l o w i n g p ropor t i on s : 8 : 5 : 1 0 : 5 : 4 , 5 : 4 , 5 : 4 , 5 : 4 : 3 , 5 : 6 ; h e a d w e a k l y ex c a v a t e d , s h i n y ; frons w i t h a com p l e t e m e d i a n k ee l , w i t h o u t scu l p t u re ; v e r t e x d u l l , w e a k l y g r a n u l a t ed ; occ i p i t a l c a r i n a a b sen t ; P O L � 3 ; O l � 2 ; OOl � 9 ; p ron o t u m crossed b y a s t r o n g t r a n s v e r s a I i m p ress i o n , sh i n y , smoo t h , w i t h o u t scu l p t u r e ; scu t u m a n d sc u t e l l u m smoo t h , sh i n y , w i t hou t scu l p t u re ; m e t a n o t u m f l a t , h o l l ow b e h i n d s c u t e l l u m , s h i n y , r u gose ; meso-me t a p l e u r a l s u t u re d i s t i n c t ; mesop l e u r a a n d m e t a p l e u r a o n d i ffere n t p l a n e s ; mesop l e u r a n o t t r a n svers e l y s t r i a t e ; m e t a p l eu r a t r a n s v erse l y s t r i a t e ; m e t a t h o r a x + p ropodeum sh i n y , w i t h a s t r o n g med i a n f u rro w , w i t h pos t e r i o r s u r f a ce t r a n s v e r se l y s t r i a t e ; a n t e r i or s u r f ace of p ropod e u m w i t h ou t sc u l p t u r e , smoo t h ; fore t a r s a l seg men t s i n fo l l ow i n g p ropor t i o n s : 1 2 : 3 : 5 : 1 5 : 2 ; en l a r ged c l a w ( F i g . 1 23 1 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t the e n d o f a l o n g i t u d i n a l f u r row a n d w i t h a row of 4 l a m e l l ae ; segmen t 5 of f ron t t a rs u s ( F i g . 1 23 1 ) w i t h t w o rows of 3 ( not p rox i m a l ) + 1 5 l a me l l a e ; apex w i t h a g r o u p of n e a r l y 1 0 l a m e l l ae ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 seg men t s ; t i b i a l spurs 1 , 0, 1 . M AL E : u n k no w n L O C U S TYP I C U S : Cl i o I te n e z ( Bo l i v i a ) TYP I C AL MAT E R I AL : h o l o t y p e F ! i n A M . H O ST S : u n k n o w n D I S TR I B U T I O N : Neo t r o p i c reg i on : BO L I V I A :

Rio

I t enez ( D e p t . Ben i ) ,

AM !

23 . Gon a topu5 bolivi a n u 5 n . sp .

F E MAL E : ap terou s ; l e n g t h 3 , 40 m m ; b l ac k ; m a n d i b l e s , c l y p e u s , m a l a r s p ace and a n a rrow reg ion a ro u n d orb i t s y e l l ow ; a n t e n n a e t e s t aceou s , w i t h segme n t s 8- 1 0 d a rk e n e d ; l e g s t e s t aceo u s , w i t h m i d a n d h i n d coxae d a rk e n e d ; a n t e n n ae d i s t a l l y t h i k e ned ; a n te n n a l seg men t s in fo l l ow i n g p ropor t i o n s : 1 2 : 7 : 2 4 : 1 3 : 1 1 : 9 : 7 : 6 : 6 : 8 ; h e a d exc a v a t e d , sh i n y ; f r o n s w i t h a co mp l e t e m e d i a n kee l ; occ i p i t a l c a r i n a a b sen t ; v e r t e x g r a n u l a ted ; frons w i t h o u t sc u l p t u re , exce p t for a n a rrow reg i o n on t h e s i des of t h e m ed i a n kee l w h i c h i s g r a n u l a te d ; POL � 2 , 5 ; OL � 2 , 5 ; D O L 9 ; p ro n o t u m c rossed by a t r a n sv e r s a l r m p r es s i on , s h i n y , smoo t h , a l mo s t h a i r l es s , w i t ho u t sc u l p tu r e ; sc u t u m d u l l , r u g ose, w i t h t w o l a t e r a l p o i n t s ; s c u te l l u m i n v i s i b l e ; meso-me t a p l e u r a l su t u re w ea k , b u t d i s t i n c t ; m e t a n o t u m f l a t , smoo t h , w i t hou t sc u l p t u r e , n o t ho l l ow beh i n d s c u t e l l u m ; m e t a t h o r a x + p ropodeum sh i n y , smoo t h , w i t h o u t sc u l p t u r e , excep t for n u me ro u s t r a n s v e r s a l s t r i ae o n pos t e r i o r s u rface o f p ropod e u m ; meso p l e u r a a n d m e t a p l e u r a o n t h e s a me p l a n e ;

m e t a p l e u r a n o t t r a n s v erse l y s t r i a t e ,

smoo t h ;

G o n a topod i n a e

Fig.

1 233 - C h e l a of Gon a topus a u t umna 1is n . sp .

1 769

( t y pe )

1 770

Mem .

Amer .

Ent .

no.

37,

1 984

M JJ A

Fig.

I nst . ,

B

1 234 - Sc u t u m a n d sc u t e l l u m of f em a l e s of Gon a top u s orb ita lis C a meron ( t y p e ) ( A ) and agilis n . sp . ( t y p e ) ( B )

mesop l e u r a w i t h on l y a few t r a n svers a l s t r i ae ; fore t a r s a l se g me n t s i n fo l l o w i n g p ropor t i o n s : 20 : 4 : 6 : 25 : 35 ; en l arged c l a w ( F i g . 1 232 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d o f a l o n g i t u d i n a l f u r row , w i t h a row of 1 0 l a me l l a e ; segmen t 5 of f ron t t a r s u s ( F i g . 1 232 w i t h t w o r o w s of 4 ( p rox i m a l ) + 2 1 l a m e l l a e ; a p e x w i t h a g ro u p of near l y 1 3 l a m e l l a e ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M AL E : u n k n o w n L O C U S TYP I C U S : 1 0 Km E S a n A n ton i o ( R i o M a mo r e , Bo l i v i a ) TYP I C A L M A T E R I AL : h o l o t y p e F ! i n A M . HOST S : u n k n o w n D I ST R I B U T I O N : Neot rop i c re g i o n : BO L I V I A 1 0 K m E S a n A n ton i o ( f, i o M a mo r e , D e p t . Be n i ) , A M !

24 . Gon a t op u s a u tumn a lis n .

sp .

F E MAL E : a p terou s ; l e n g t h 3 , 20 m m ; h e a d b l ac k , w i t h m a n d i b l e s , c l y p e u s a n d m a l a r s p ace t e s t aceou s j a n t e n n ae t e s t aceO U 5 , w i t h segme n t s 7- 1 0 b row n ; t ho r a x a n d p ropodeum b l ack ; a b domen b l ack-brow n ; l e g s b row n , w i t h coxae b l ac k a n d t a r s i l i g t h ; a n te n n ae d i s t a l l y t h i c k e n e d ; a n t e n n a l se gmen t s i n fo l l ow i n g p ropor t i o n s : 1 5 : 6 : 2 1 : 1 1 : 9 : 8 : 7 : 7 : 6 , 5 : 8 ; h e a d d u l l , f u l l y g r a n u l a ted ; f r o n s w i t h a comp l e t e m ed i a n k ee l ; oc i p i t a l car i n a i n v i s i b l e ; POL � 2 ; OL � 2 ; OOL � 1 1 ; p ro n o t u m crossed by a s t ro n g t r a n s vers a l i m p ress i o n , h a i r y , sh i n y , smoo t h , w i t h o u t sc u l p t u re ; sc u t u m d u l l , g r a n u l a t e d a n d w i t h some l o n g i t u d i n a l k ee l s ; s c u te l l u m a n d m e t a t h o r a x + p ropodeum d u l l , g r a n u l a ted ; meso-me t a p l eu r a l s u t u re obso l e t e ; p l e u r a n o t t r a n s v e rse l y s t r i a te ; p o s t e r i o r s u r f a c e of p ro p o d e u m a l mo s t f u l l y n o t t r a n sverse l y s t r i a t e , e x c e p t for re g i o n n e a r bot tom o f p ro p o d e u m a n d s i des o f p o s te r i o r s u r f a c e , w h ere some s t r i a e a r e v i s i b l e ; a n t e r i or s u r f ace of p ro p o d e u m very i n c l i n ed ( F i g . 1 226 B ) ; fore t a rsa l segme n t s i n fo l l ow i n g propor t i o n s : 1 6 : 3 : 6 : 2 1 : 32 ; en l arged c l a w ( F i g . 1 233 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l too t h at t h e e n d of a l on g i t u d i n a l f u r row

G o n a topod i n ae

1 77 1

a n d w i t h a row o f 1 1 peg - I i k e h a i r s ; seg m e n t 5 o f fron t t a r s u s ( F i g . 1 23 3 ) w i t h t w o rows o f n e a r l y 2 1 l a me l l a e ; apex w i t h a g roup of n e a r l y 1 8 l a m e l l a e ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 s e g me n t s ; t i b i a l spurs 1 , 0 , 1 . MALE :

unknown

L O C U S TYP I C U S : Mou t h o f R i o B a u re s ( Bo l i v i a ) H OSTS : u n k no w n D I ST R I BU T I O N : Neotrop i c re g i o n : BO L I V I A : Mou t h o f R i o B a u re s ( R i o D ep t . Be n i ) , A M !

I te n e z ,

A U S T R A L I AN S P E C I E S OF GONA TOPUS

T w e l v e spec i e s are k n ow n .

K E Y TO T H E S P E C I E S

F EMALES

S e g men t s egmen t S e g men t

2

3

o f f r on t t a r s u s a p p r ox i ma t e l y t w i c e a s l o n g a s 4 1 . m el a n ias ( R . C . L . P e r k i n s ) 1 o f f r an t t a r s u s s h o r t e r t h a n , a s l on g a s , o r 2 s I i gh t I y l o n g e r t h a n s e g m e n t 4 S eg m e n t 5 o f f r on t t a r s u s w i t h t wo r ow s o f l a m e l l a e ex t e n d i n g b ey o n d 0 , 5 l en g t h o f s eg m e n t ( F i g s 1 2 3 7 , 1 2 3 8 , 1 2 4 0 , 1 2 4 1 , 1 242 ) 3 S e g m e n t 5 o f f r on t t a r su s w i t h o u t l a m e l l a e o r w i t h 1 - 2 r o w s o f l a m e l l a e ; i f t w o r o w s , t h e s h o r t e s t r o w d o e s n o t ex t e n d b ey on d 0 , 5 l e n g t h o f s eg m e n t ( F i g s 1 2 4 3 , 1 2 4 4 , 1 2 4 5 , 1 2 4 7 , 1 2 4 8 , 1 249 ) 7 •

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. . 2 . williamsi n . sp . V e r t ex o f h e a d a l mo s t f l a t ; s c u t u m l a t e r a l l y w i t h ou t p o i n t s ; p o s t e r i o r su r f a c e o f p r op o d e u m a l mo s t f u l l y s mo o t h , t r a n s v e r s e l y s t r i a t e on l y n e a r a p e x . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 . zeala n dicus n . sp . H e a d w i t h f r on s a n d v e r t ex d u l l , g r a n u l a t e d , w i t h ou t t r a n s v e r s a l f o l d s o r l on g i t u d i n a l k e e l s 4 . gigas ( R . C . L . P e rk i n s ) H e a d w i t h f r o n s a n d v e r t ex sh i n y , n o t g r a n u l a t ed , w i t h s t r o n g .

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Me s o -m e t a p l eu r a l su t u r e d i s t i n c t a n d c omp l e t e 4 Me s o -me t a p l eu r a l su t u r e ob s o l e t e 5 V e r t ex o f h ea d v e r y e x c a v a t ed ; s c u t u m l a t e r a l l y w i t h 2 p o i n t s ; p o s t e r i o r su r f a c e o f p r op od e u m f u l l y t r a n s v e r s e l y s t r i a t e •

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1 772

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En!.

I ns! . ,

n o . 37 ,

t r a n sv e r s a l f o l d s o r l o n g i t u d i n a l k e e l s 6 H ea d w i t h f r on s s c u l p t u r ed by n u m e r ou s t r a n sv e r s a l f o l d s . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 . plic a tus n . sp . H e a d w i t h f r o n s s c u l p t u r e d by n u me r o u s I o n g i t u d i n a l k e e l s , w i t h ou t t r a n sv e r s a l f o l d s 6 . rossi n . s p . S e g m en t 5 o f f r on t t a r s u s w i t h o u t l a m e l l a e , w i t h on l y a r o w o f b r i s t l e s ( F i g . 1 2 4 3 ) ; h e a d f l a t . 7 . n a um a n n i n . s p . S e g men t 5 o f f r on t t a r s u s w i t h 1 - 2 r o w s o f l a m e l l a e ; ( F i g s 8 1 2 4 4 , 1 2 4 5 , 1 2 4 7 , 1 2 4 8 , 1 2 4 9 ) ; h e a d e x c a v a t ed T h o r a x a n d p r o p od e u m b l a ck o r b r own - b l a ck ( ex c ep t f o r ap ex o f p r o p od e u m l i g h t ) 9 Th o r a x a n d p r op od e u m p a r t I y o r f u l l y r ed d i sh o r t e s t a c e o u s o r y e I l ow . 10 Me s o -m e t a p l e u r a l s u t u r e ob s o l e t e ; h e a d g r a n u l a t ed •

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. . . . . . . . . . . . . . . . 8 . papua nus n . sp . Me s o - me t a p l eu r a l s u t u r e d i s t i n c t a n d c o mp l e t e ; h e a d w i t h o u t scu l p t u r e 9 . op t a bilis ( R . C . L . P e r k i n s ) 1 0 Me s o - m e t a p l eu r a l s u t u r e d i s t i n c t a n d c o mp l e t e . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1 0 . decor a t u s ( R . C . L . P e rk i n s ) M e s o - me t a p I eu r a I s u t u r e ob s o l e t e 1 1 1 1 Me t a t h o r a x + p r op od e u m r e d d i sh -b r own ; a n t e n n a l s eg me n t 1 0 b r own ; b od y s l e n d e r l 1 . a u s tr a l i a e R . C . L . P e rk i n s M e t a t h o r a x + p r o p o d e u m r e d d i s h - t e s t a c e ou s ; a n t e n n a l s eg m e n t 1 0 wh i t e ; b od y r ob u s t 1 2 . pulcherrimus ( R . C . L . P e rk i n s ) •

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MALES

P r o p o d e u m s mo o t h a n d sh i n y ; p r ox i ma l p a r t o f ra d i a l v e i n mu ch sh o r t e r t h a n d i s t a l p a r t 1 . mela n i a s ( R . C . L . P e rk i n s ) P r op od e u m r e t i c u l a t e r u g o s e , d u l l ; d i s t a l p a r t o f r a d i a l v e i n n e a r l y a s l o n g a s p r ox i ma l p a r t 2 D o r s a l p r o c e s s of g o n o f o r c e p s n e a r l y a s l o n g a s g on o f o r c e p s (F i g . 1 239 ) 3 . zea l a n dicus n . s p . D o r s a l p r o c e s s o f g on o f o r c e p s mu ch sh o r t e r t h a n g on o f o r c e p s ( F i g . 1 2 4 6 ) . . . . . . . . . . . . . . . . . . . . . . 9 . op ta bilis ( R . C . L . P e rk i n s ) ! 1 0 . decora tus ( R . C . L . P e rk i n s ) ! .

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T h e m a l es of t h e o t h e r spec i e s are u n k n ow n .

1 . Gona topus mela n i a s ( R . C . L . Perk i n s ) n . com b .

!

T h e on l y k n o w n · m a l e s p ec i m en c a n b e l o n g to G.

op t a b ilis or decora tus .

.

Gon a topod i n ae

Fig.

1 235

-

C h e l a of Gon a topus mela n i a s ( R . C . L . I s l ands )

1 7 73

Perk i n s )

Pa chygon a top u s mela n i a s R . C . L . Perk i n s 1 90 5 : 4 6 .

f rom S u v a ( F i j i

1 774

Mem . A mer . E n t .

I nst . ,

no .

37,

1 984

F E MAL E : ap t e r o u s ; l e n g t h 2 , 25-2 , 38 mrn ; b l a ck , w i t h c l y p e u s , m a n d i b l e s a n d f ro n t p a r t of v e r t e x y e l l o w ; a n t e n n a e b row n , w i t h seg men t s 1 - 3 l i g h t ; l eg s b r o w n , w i t h t r oc h a n t e rs a n d t a r s i l i g h t ; h e a d f l a t , sh i n y , w i t h o u t scu l p t u re , exce p t f o r occ i p u t a n d f r o n t p a r t o f v e r t e x w h i c h a re g r a n u I a t e d ; prono t u m crossed b y a t r a n s v e r s a I i m p ress i o n , w i t h o u t scu l p t u re o r w e a k l y a l u t a ceou s ; sc u t u m sh i n y , w i t h o u t scu l p t u re , w i t h some l o n g i t u d i n a l k ee l s ; m e t a n o t u m f l a t , not t r a n s v e r se l y s t r i a te ; m e t a t h o rax + p ropodeu m sh i n y , w i t h o u t scu l p t u re or weak l y a l u t a ceou s ; p l e u r a a n d p o s t e r i o r su rface of p r o p o d e u m n o t t r a nsver se l y s t r i a te o r w e a k l y s t r i a te ; seg m e n t 1 of f ro n t t a r s u s a p p rox i m a t e l y t w i ce as l o n g as segmen t 4 ( 1 4 : 6 ) ; en l arged c l a w ( F i g . 1 2 35 ) w i t h o u t s u b a p i c a l too t h a n d l a m e l l a e , w i t h a sm a l l too t h a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d 4-5 p e g - l i k e h a i r s ; segme n t 5 o f f r o n t t a r s u s ( F i g . 1 2 35 ) w i t h 2 rows o f 1 2- 1 4 s m a l l l a m e l l ae ; apex b ro a den e d , w i t h a g r o u p o f 5 - 1 0 l a me l l a e ; m a x i I l a r y p a l p i w i t h 3-4 segmen ts ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MAL E : fu l l y w i n g ed ; l e n g t h 2 , 2 -2 , 5 m m ; bro w n ; h e a d , t h o rax a n d p ropodeu m d a r k ; a n t e n n a e , a b d o m e n a n d l e g s i n some spec i m e n s I i gil t ; l e g s never b a n d e d ; a n t e n n a l se gmen t s i n fo l l o w i n g p ropo r t i o n s : 4 : 4 : 8 : 7 : 6 : 7 : 6 , 5 : 6 , 5 : 6 , 5 : 9 ; a n te n n a l seg men t 3 more t h a n t h ree t i mes as l o n g as b r o a d ( 3 , 6 ) ; h e a d s h i n y , g r a n u l a t e d , h a i r y ; fron t a l l i n e absen t ; POL � 7 ; O L = 3 ; OOL = 3 ; bread t h of t h e oce l l i : 3 ; t e m p l e s d i s t i n c t ; occ i p i t a l ca r i n a i n v i s i b l e ; scu t u m g r a n u l a t e d ; n o t a u l i c es i n comp l e t e , rea ch i n g a p p rox i m a te l y 0 , 3-0 , 5 l e n g t h of sc u t u m ; sc u t e l l u rn a n d m e t a n o t u m h a i r y , smoo t h , sh i n y , w i t h o u t scu l p t u re ; p ropodeu m sh i n y , smoo t h , w i t h o u t scu l p t u re ; d or s a l s u r face w i t h o u t med i a n f u r ro w ; dorsa l p rocess of gonoforceps ( F i g . 1 2 36 ) sho r t , s l e n d e r , w i t h apex not po i n t e d ; b a s i vo l se l l a w i t ho u t n o t c h ; ma x i l l a ry p a l p i w i t h 3-4 seg men t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . LOCUS TYP I C US : t y p i c a l m a t e r i a l w i t h o u t l oc a l i t y l a b e l ; B u n d a b e r g a n d Rock h a m p ton ( Qu e e n s l a n d , A u s t r a l i a ) i n o r i g i n a l d e sc r i p t i o n . TYP I C AL MAT E R I AL : l ec t o t y p e F ! a n d 2 p a r a l ec t o t y pes MM ! ( de s i g n a t e d b y O l m i 1 982 ) i n B . HOSTS : i n A u s t r a l i a Euleimonios s p . , A thys a n u s s p . ( R . C . L . Perk i n s 1 906c ) • D I ST R I B U T I O N : A u s t r a l i a n reg i o n : AUSTRAL I A : B r i s b a n e ( Q ueens l a n d ) , QU ! B u n d aberg ( Qu e e n s l a n d ) , B ! R oc k h a m p t o n ( Queens l a n d ) , B ! O x l e y ( Qu ee n s l a n d ) , O L ! BR ! F I J I I SLANDS : S u v a ( V i t i L e v u ) , B ! L a u t o k a ( V i t i Lev u ) , BM ! I N D O N E S I A : Ambon ( Ce r a m ) , OL !

2 . Gon a topus williamsi n . sp .

F E MAL E : ap t e r ou s ; l e n g t h 4 , 6 m m ; h e a d brow n , w i t h c l y p eus , m a n d i b l e s a n d f ro n t p a r t o f v e r t e x y e l l o w ; a n t e n n a e b r ow n , w i t h se g m e n t s 1 - 2 y e l l o w ; t h o r a x , p ropodeu m a n d a b domen b l ack ; l e g s b r o w n - b l ack , w i t h t a r s i l i g h t ; h ea d ex c a v a t e d , sh i n y , w i t h o u t scu l p t u re , e x c e p t for occ i p u t

Gon a topod i n a e

Fig.

1 77 5

1 236 - M a l e ge n i t a l i a of Gon a top u s mel a n i a s ( R . C . L . Perk i n s ) f ro m Ox l e y ( Qu ee n s l a n d } ( l e f t h a l f removed )

a n d f r o n t p a r t of vertex wh i c h a re g r a n u I a t e d ; occ i p u t a I so w i t h some short k ee l s ; occ i p i t a l ca r i n a i n v i s i b l e ; p ro n o t u rn crossed b y a t r a n s v e r s a l i m p ress i o n , sh i n y , smoo t h , w i t h o u t sc u l p t u r e ; sc u t u m sh i n y , w i t h some l o n g i t u d i n a l k ee l s , l a te r a l l y w i t h 2 po i n t s ; m e t a n o t u m h o l l ow b e h i n d sc u te l l u m , t r a n sverse l y s t r i a te ; m e t a t h o r a x + p ro p o d e u m sh i n y , w i t h o u t sc u l p t u re , w i t h w e a k t r a c k of med i a n f u rrow ; p l e u r a a n d p o s te r i o r su rface of propodeum t r a n s v e r se l y s t r i a t e ; meso-rnet a p l e u r a l su t u r e d i s t i n c t a n d co m p l e t e ; se g m e n t 1 of f ro n t t a r s u s s h o r ter segmen t 4 ( 1 1 : 1 5 ) ; en l a r g e d c l a w ( F i g . 1 237 ) w i t h o u t s u b a p i c a l t oo t h , w i t h a s rn a l l too t h a t t h e e n d o f a l o n g i t u d i n a l f u r r o w a n d 8 peg- l i k e h a i rs ; segmen t 5 o f fron t t a r s u s ( F i g . 1 237 ) w i t h 2 r o w s of 8 + 2 1 l a m e l l ae e x t e n d i n g b e y o n d 0 , 5 l e n g t h o f segmen t ; apex w i t h a g r o u p o f a t l e a s t 1 5 l a rn e l l a e ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 .

Mem . A mer .

1 776

Fig.

1 237

-

Ent.

I nst . ,

no.

37,

1 984

C h e l a of Gon a top us will i a m s i n . s p .

( type )

MALE : u n k n o w n LOC U S TYP I C U S : H a l i f a x ( A u s t r a l i a ) TYP I C AL MA T E R I AL : ho l o t y p e F ! i n OL . HOST S : u n k n o w n D I ST R I B U T I O N : A u s t r a l i a n reg i o n : A U S T R A L I A : H a l i f a x ( Queen s l a n d ) , N O T E S : t h e s p ec i e s is n a med i n honor of M r . F . X . W i l l i a m s .

3 . Gon a top u s ze a l a n dicus n .

OL !

sp .

F E MA L E : a p t e r o u s ; l e n g t h 2 , 1 8 - 3 , 25 m m ; h e a d b l a c k , w i t h c l y p e u s , m a n d i b l e s a n d f ro n t p a r t o f v e r tex ( more a l o n g orb i t s ) y e l l ow ; a n te n n ae b l ac k , w i t h seg men t s 1 - 2 a n d a n a rrow p rox i m a l p a r t of s e g me n t 3 y e l l ow ; t h o r a x a n d p ropodeum b l ac k ; a b domen b l ac k or b row n ; l e g s t e s t aceOU 5 , w i t h coxae p a r t l y b r o w n o r b l ac k a n d c l u b s o f femora b r o w n i s h ; h e a d w e a k l y e xc a v a te d , a l m o s t f l a t , sh i n y , h a i r l e s s , w i t h o u t sc u l p t u re , smoo t h ; occ i p u t h a i r y ; occ i p i t a l c a r i n a v i s i b l e on l y b eh i n d pos t e r i o r oce l l i ; p ro n o t u m crossed b y a t r a n s v e r s a l i m p ress i o n , h a i r l e s s , smoo t h , w i t h o u t scu l p t u re ; sc u t u m shor t , a s l o n g as b roa d ; sc u t e l l u m a s l o n g a s scu t u m , r u gose ; m e t a n o t u m ru gose, w i t h i r reg u l a r a n d s h o r t k ee l s ; m e t a t h o r a x +

G o n a t opod i n ae

1 7 77

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C h e l a of Gon a top u s zea l a n dicus ( N ew Z e a l a n d )

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1 239 - M a l e g en i t a l i a of Gona top u s zealandicus n . sp . f rom Hooker V a l l ey ( New Z ea l a n d )

p ropodeum s m oo t h , w i t h o u t scu l p t u re , excep t for a p i c a l reg i on of pos ter i or s u r face w h i ch i s t r a n sverse l y s t r i a te ; p l e u r a n o t t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l su t u r e d i s t i n c t a n d comp l e t e ; se g m e n t 1 o f f ro n t t a r s u s a s l o n g a s se g m e n t 4 ; en l a r g e d c l a w ( F i g . 1 23 8 ) w i t ho u t s u b a p i c a l t oo t h , w i t h a sm a l l too t h a t t h e e n d of a l o n g i t u d i n a l f u rrow , w i t h o u t l a me l l ae or b r i s t l es ; se g m e n t 5 of f ro n t t a r s u s ( F i g . 1 2 38 ) w i t h 2 rows of 1 6 l a me l l ae ; apex w i t h a g r o u p o f 3 l a me l l ae ; m ax i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : f u l l y w i n g e d ; l e n g t h 1 , 75-2, 2 5 m m ; h e a d b l a c k , u s u a l l y w i t h m a n d i b l es redd i s h ; a n te n n ae , t h o r a x , propodeum a n d a b domen b l ac k ; l e g s b l ac k , w i t h t a r s i l i g t h ; a n t e n n a l segmen t s i n f o l l ow i n g propo r t i o n s : 5 : 5 : 9 : 9 : 9 : 8 : 8 : 8 : 7 : 9 ; a n te n n a l seg m e n t 3 n e a r l y t h ree t i mes a s l on g a s b ro a d ( 3 ) ; h e a d h a i r y , s h i n y , g r a n u l a t e d ; temp l e s d i s t i nc t ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L � 7 ; O L � 3 ; OOL � 5 ; b re a d t h o f t h e oce l l i : 3 ; scu t u m s h i n y , g r a n u l a ted ; n o t a u l i ces d i s t i n c t , i ncom p l e t e , rea c h i n g a p p ro x i m a te I y 0 , 5 l e n g t h o f scu t u m ; sc u te l l u m a n d met a n o t u m s h i n y , smoo t h , w e a k l y p u n c t a t e ; p ropode u m ret i c u l a te ru gose ; fore- w i n g h y a l i n e , w i t ho u t d a r k t r a n s vers a l

bands ;

d i s t a l p a r t o f ra d i a l v e i n n e a r l y a s l o n g

G o n a to pod i n ae

Fig.

1 240 - C h e l a of Gon a top us gigas

1 779

(R.C.L.

Perk i n s ) ( l ec t o t y p e )

as p rox i m a l p a r t ; dors a l p rocess of gonoforce p s ( F i g . 1 239 ) l o n g , w i t h a p ex rou n d e d ; b a s i v o l se l l a w i t h o u t n o t c h ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 1 , 2 . lOCUS TYP I C U S : K a w a r a u Gorge ( An n i ec k , Ce n t r a l O t a g o , New Ze a l a n d ) TYP I CAL MA T E R I AL : ho l o t y p e F ! a n d 9 p a r a t y p es ( 1 F , 8 M M ) ! i n N Z ; p a r a t y p e M ! i n BM ; 4 p a r a t y pes ( 2 F F , 2 M M ) ! i n Ol . HOST S : u n k n o w n D I ST R I BU T I ON : A u s t r a l i a n reg i o n : N E W Z E A LA N D : K a w a r a u G o r g e ( An n i ec k , C e n t r a l O t a go ) , N Z ! D u n s t a n ( R a n g e , C e n t r a l O t a go ) , N Z ! M t . D a l ge t y ( M a c k e n z i e ) , O l ! T h a r lodge ( He r m i t a g e , M t . Cook , M a c k e n z i e ) , N Z ! Hooker V a l l e y ( H erm i t a g e , M t . Cook N a t . P a rk , M a c k e n z i e ) , BM ! N e r v o u s K n o b ( C ra i g i e b u r n Fores t , M i d C a n t e r b u r y ) , Ol ! N Z ! B i r d l i n g s F l a t ( C h r i s t c h u rc h , M i d C a n t e r b u r y ) , O l ! A k a roa ( B a n k s Pen i n su l a , M i d C a n t e r b u r y ) , N Z ! Tower P e a k ( T a k i t i m u R a n g e , Sou t h l a n d ) , N Z ! Hedge hope s u mm i t ( Hokon u i H i l l s , So u t h l a n d ) , N Z ! Stephens I s l a n d ( M a r l borou g h Sou n d s ) , N Z ! Bee b y ' s K n o b , O l !

4 . Gon a top u s gigas ( R . C . l .

Perk i n s )

n . comb .

Cha lcogon a top u s gigas R . C . l . P e r k i n s 1 90 5 : 47 . C h a lcogona topus a lb iscapus Dodd 1 9 1 4c : 3 ( n . sy n . ) ; locus typicus N e l son ( Quee"" l a n d , A u s t r a l i a )

F E MAL E : a p terou s ; l e n g t h 4 , 00-5 , 85 m m ; b l a c k or b r o w n - b l ac k , w i t h m a n d i b l e s y e l l ow ; a n te n n a e brow n , w i t h s e g me n t 1 p a r t l y y e l l o w ; l e g s

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- C h e l a of Gon a top u s p l i c a t u s n . s p .

( type )

brow n , w i t h c h e l ae y e l l o w ; h e a d exc a v a te d , g r a n u l a te d , d u l l , w i t ho u t t r a n s v e rs a l k ee l s ; p rono t u m crossed by a t r a n sver s a l i mp re ss i on , g r a n u l a ted ;

meta thorax

+

p ropodeum g r a n u l a te d ; meso-me t a p l eu r a l su t u r e

Gon a topod i n ae

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1 242 - C h e l a o f Gonatopus rossi n .

1 78 1

sp .

( type)

obso l e t e o r w i t h on l y pos t e r i or h a l f v i s i b l e ; segm e n t 4 o f fron t t a r s u s n e a r l y as l o n g a s segmen t 1 ( 50 : 49 ) ; en l a rged c l aw ( F i g . 1 240 ) w i t hou t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d 4-6 p e g- l i ke h a i rs ; segmen t 5 of f ron t t a rs u s ( F i g . 1 240 ) w i t h 2 rows of at l e a s t 24-30 I a me l i ae ex t en d i ng beyon d 0 , 5 l e n g t h of segmen t ; apex w i t h a g ro u p of a t least 1 0-20 l a me l l a e ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MAL E : u n k no w n

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L O C U S TYP I C U S : B u n d a b e r g ( A u s t r a l i a ) TYP I C A L MAT E R I AL : l e c t o t y pe F ! a n d 1 p a r a t y pe F ! ( d e s i g n a t e d b y O l m i 1 982 ) of Ch . gigas i n B ; 1 p ar a l ec t o t y p e F ! of Ch . gigas i n B ; h o l o t y pe F ! of Ch . a l b iscap u s i n AD . HOSTS : i n A u s t r a l i a Tartessus syrtidis ( R . C . L . Per k i n s 1 906c ) . D I ST R I B U T I O N : A u s t r a l i a n reg i on : AUSTRAL I A : B u n d aberg ( Q uee n s l a n d ) , B ! BM ! B r i s b a n e ( Quee n s l a n d ) , UQ ! Ne l so n ( C a i r n s , N Quee n s l a n d ) , AD !

5 . Gon a top u s plic a t u s n . sp .

F E MALE : a p t e ro u s ; l e n g t h 4 , 68 m m ; b a l c k , w i t h m a n d i b l es , che l ae a n d s i d es o f c l y p e u s y e l l ow ; a n te n n a e w i t h se g m e n t 1 y e l l o w , seg men t s 2-3 b l ac k , other se g m e n t s m i s s i n g i n the o n l y k n o w n fema l e s p ec i men ; h e a d exca v a t e d , s h i n y , smoo t h , w i t h o u t scu l p t u r e , w i t h s o m e t r a n s v e rs a l k ee l s o n frons a n d v e r t e x ; pro n o t u m crossed b y a t r a n s v e r s a I i m press i o n , s h i n y , smoo t h , w i t h o u t scu l p t u re ; scu t u m sh i n y , w i t h some l on g i t u d i n a l k ee l s , l a tera l l y w i t h 2 po i n t s ; m e t a no t u m sh i n y , w i t h o u t t r a n sv e rs a l k ee l s ; m e t a t h o r a x + propodeum s h i n y , w i t ho u t scu l p t u re ; mesop l e u r a g r a n u l a t ed and w i t h a few t r a n sversa l s t r i ae ; m e t a p l e u r a g r a n u l a t e d and w i t h n u me r o u s s t r i ae ; d i sc of m e t a t h o r a x + propodeum w i t h t w o l a t e r a l s t ro n g c u r v i I i n ear f u rrow s ; pos t e r i o r s u r f ace o f p ropodeum t r a n sverse l y s t r i a t e ; meso-me t a p l e u r a l s u t ure o b so l e t e ; s e g m e n t 1 o f fron t t a r s u s a s l o n g a s s e g me n t 4 ; en l arged c l a w ( F i g . 1 24 1 ) w i t h o u t s u b a p i c a l t oo t h , w i t h a s m a l l tooth a t t h e e n d of a l on g i t u d i n a l fu rrow a n d 7 p e g - l i k e h a i rs ; s e g me n t 5 of fro n t t a r s u s ( F i g . 1 24 1 ) w i t h 2 rows of 1 8 + 22 l a me l l ae e x t e n d i n g b e y o n d 0 , 5 l e n g t h o f segmen t ; apex w i t h a g roup o f a t l e a s t 2 0 l a me l l ae ; m a x i I l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l spurs 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : W a u ( Ne w G u i n ea ) TYP I C A L M A T E R I AL : h o l o t y p e F ! i n B . HOSTS : u n k n o w n D I ST R I B U T I O N : A u s t r a l i a n re g i o n : N E W G U I N E A : W a u ,

B!

6 . Gon a top u s rossi n . s p .

F EMAL E : a p t e r ou s ; l e n g t h 5 , 00 m m ; head b l ac k , w i t h frons ( more a l o n g or b i t s ) , c l y p e u s , m a n d i b l e s t e s t aceou s ; t h o r a x , propodeum a n d a b domen b l ac k ; l e g s t e s t aceo u s , w i t h fore cox a e , c l u b s of femo r a , fore t i b i ae p a rt l y b ro w n i s h ; h e a d exc a v a t e d , sh i n y , w i t h n u merous l on g i t u d i n a l k ee l s , w i t h o u t t r a n s v e rs a l fo l d s , n o t g r a n u l a ted ; p rono t u m sh i n y , crossed b y a t r a n sv e rs a l i m p ress i o n , smoo t h , w i t h o u t scu l p t u re , h a i r l ess ; sc u t u m sh i n y , w i t h n u merou s l o n g i t u d i n a l k ee l s ; sc u te l l u m very s h o r t ; met a n o t u m w i t h

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n u mero u s t r a n sversa I kee l s ; m e t a t horax + p ropodeum d u I I , w i t h n u mero u s l on g i t u d i n a l k ee l s on a n t e r i o r su rface a n d w i t h n u m erou s t r a n s v ersa l k ee l s o n p l e u r a a n d pos t e r i o r su rface ; meso-me t a p l e u r a l s u t u re obso l e t e ; segmen t 1 of fron t t a r s u s s h orter t h a n segmen t 4 ( 1 1 : 1 4 ) ; e n l arged c l a w ( F i g . 1 242 ) w i t h o u t su b a p i c a l too t h , w i t h a sm a l l too t h a t the e n d of a l on g i t u d i n a l fu rrow a n d w i t h 6 peg - l i k e h a i r s ; segmen t 5 of fro n t t a r s u s ( F i g . 1 242 ) w i t h 2 r o w s of 3 2 l a me l l ae exte n d i n g beyond 0 , 5 l e n g t h o f segmen t ; apex w i t h a t l e a s t 1 5 l a me l l ae ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l spurs 1 , 0 , 1 . MAL E : u n k n o w n LOCUS TYP I C U S : G l encoe ( A u s t r a l i a ) TYP I CAL MAT E R I AL : h o l o t y p e F ! i n C A . HOSTS : u n k n o w n D I ST R I BU T I ON : A u s t r a l i a n reg i o n : AUSTRAL I A : G l encoe ( New Sou t h W a l es ) , CA l NOT E S : t h e spec i es i s named i n honor of M r . E . S . Ros s .

7 . Gona topu5 n a um a n n i n .

sp .

F E MAL E : a p terou s ; l e n g t h 2 , 25 m m ; head brow n , w i t h c l y p e u s , m a n d i b l es a n d a n te r i o r reg i o n of frons test aceou s j a n t en n ae test aceou s ; p ropec t u 5 b row n ; prono t u m , meso t ho r a x , m e t a t horax a n d propodeum test aceou s ; p e t i o l e a n d a b domen b l ack ; l e g s t e s t aceou s , w i t h coxae a n d c l u b s of femora b row n ; a n t e n n ae d i s t a l l y t h i ckened ; a n ten n a l segmen t s in fo l l ow i n g porpor t i on s : 6 : 4 : 5 : 3 , 5 : 3 : 3 : 3 , 5 : 3 , 5 : 3 , 5 : 5 , 5 ; head f l a t , s h i n y , smoo t h , w i t ho u t scu l p t ure ; frons w i t h a med i a n i n comp l e te short kee l i n fro n t of t h e a n te r i o r oce l l u s ; occ i p i t a l car i n a v i s i b l e on l y p a r t l y o n the s i des of t h e poster i o oce l l i ; POL = 2; OL = 2; OOL = 5 , 5 ; p ro n o t u m crossed by a t r a n s v e rs a l i mp ress i o n , sh i n y , smoo t h , w i t h o u t scu l p t u r e ; sc u t u m a n d m e t a t horax + propodeum sh i n y , smoo t h , w i t h o u t scu l p t u r e , exce p t for poste r i o r s u rface of propodeum t r a n sv erse l y s t r i a t e ; meso-me t a p l eu r a l s u t u re obso l e t e ; met a n o t u m w i t h s i des not rou n d e d , w i t h 2 l a tera l po i n t s ; fore t a r s a l segme n t s i n fo l l ow i n g propor t i on s : 7 : 2 : 3 : 6 : 1 1 ; e n l arged c l aw ( F i g . 1 243 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d of a l on g i t u d i n a l fu rrow and w i t h 5 b r i s t l es ; segme n t 5 of fron t tarsus ( F i g . 1 243 ) w i t h i n n er s i d e prox i m a l l y serr a t e , w i t h o u t l a me l l ae on p rox i m a l a n d m ed i a l reg i o n , on l y w i t h a row o f 5 b r i s t l es ; l a me l l a e are v i s i b l e on l y on d i s t a l reg i o n , w here there is a g roup of 5 l a me l l ae , in a d d i t i on to 1 separa ted l a me l l a l oc a ted on curve b e t ween d i s t a l a n d med i a l reg i o n s ; m a x i l l a ry p a l p i w i t h 4 segme n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l spurs 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I CU S : Forest S t a t i on ( B u l b u r i n S t a t e Fores t , Quee n s l a n d , Austra l i a ) TYP I CAL MATER I AL : h o l o t y pe F ! i n UQ .

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1 245 - Se g men t 5 of fron t t a r s u s of fema l e of Gon a top u s op t a bilis ( R . C . L . Perk i n s ) ( l ec t o t y p e )

H OS T S : u n k n o w n D I ST R I B U T I O N : A u s t r a l i a n reg i on : A U S T R A L I A : F o r e s t S t a t i o n ( B u l b u r i n S t a t e Fore s t , v i a M a n y Pea k s , Q u e e n s l a n d ) , U Q ! N OT E S :

t h e s p ec i e s i s n a med

i n honor o f M r .

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8 . Gon a top us p a p u a n u s n .

Nauma n n .

sp .

F E M AL E : a p tero u s ; l e n g t h 3 , 0 m m ; h e a d b row n , w i t h c l y e p u s , m a n d i b l es a n d f ro n t p a r t of v e r t e x y e l l ow ; a n t en n ae y e l l ow ( o n l y se g men t s 1 -4 p resen t , o t h e r seg m e n t s m i ss i n g i n t h e on l y k n own fem a l e ) ; t h o r a x a n d p ro p o d e u m b l ac k ; a b domen b row n ; l e g s y e l l o w , w i t h cox a e a n d c l u b s o f femora b row n ; h e a d exca v a te d , sh i n y , g r a n u l a t ed ; p rono t u m crossed b y a t r a n s v e rs a l i m p ress i o n , s h i n y , smoo t h , w i t h o u t scu l p t u re or w e a k l y s c u l p t u red on d i sc ; scu t u m sh i n y , w i t h some l on g i t u d i n a l k ee l s ; met a n o t u m s h i n y , smoo t h , n o t t r a n sv erse l y s t r i a t e ; m e t a t h o r a x + p ropodeum sh i n y , w i t hou t scu l p t u re ; p l eu r a a n d p o s te r i o r s u rf a ce of p ropod e u m t r a n s v erse l y s t r i a t e ; meso-me t a p l e u r a l s u t ure o b so l e t e ; s e g m e n t 1 of f ro n t t a r s u s a s l o n g a s seg m e n t 4 ; en l a r g e d c l a w ( F i g . 1 24 4 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d of a l o n g i t u d i n a l f u rrow a n d 6 peg - l i k e h a i r s ; segme n t 5 o f fron t t a r s u s ( F i g . 1 244 ) w i t h 1 row o f 1 7 l a me l l ae ; apex w i t h a g ro u p of 1 2 l a me l l ae ; m a x i l l a r y p a l p i w i t h 5 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : Aroa E s t a t e ( N ew G u i n e a ) TYP I C A L MAT E R I AL : h o l o t y p e F ! i n B . HOST S : u n k n o w n D I ST R I B U T I ON : A u s t r a l i a n reg i o n : N E W G U I N E A : Aroa E s t a te ( Re d sc a r B a y , Papu a ) , B !

9 . Gon a top u s op ta bilis ( R . C . L .

Perk i n s ) n . com b .

G o n a topod i n a e

Fi g .

1 246 - M a l e ge n i t a l i a o f Gon a top u s op tabilis ( R . C . L . decor a t u s ( R . C . l . Perk i n s ) ( p a r a l ec t o t y p e )

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Ch alcogon a top u s op t a b ilis R . C . l . Perk i n s 1 905 : 47 .

F E M AL E : a p terou s ; l e n g t h 3 , 47-4 , 2 1 m m ; b row n -b l ac k , w i t h m a n d i b l es , c l y p e u s , a n t e n n a I segmen t s 1 -2 a n d fro n t p a r t o f vertex ye l l ow ; l e g s b ro w n -y e l l ow , w i t h f o r e t i b i a e a n d t a r s i y e l l ow ; h e a d smoo t h , w i t h o u t scu l p t u r e , exca v a te d j p ron o t u m crossed by a t r a n s v e rsa l i m p ress i o n , w i t ho u t sc u l p t u r e or w e a k l y scu l p t u red ; m e t a t h o r a x + p ropodeum w i t h o u t scu l p t u re ; p o s te r i o r surface of p ropodeum t r a n sverse l y s t r i a t e ; meso m e t a p l e u r a l s u t ure d i s t i n c t and comp l e t e ; se g men t 4 of fron t t a rs u s s l i g h t l y l o n ger t h a n se g m e n t 1 ( 35/31 ) ; en l arged c l a w w i t h o u t s u b a p i c a l too t h , w i t h a sm a l l toot h a t t h e e n d o f a l o n g i t u d i n a l f u r row a n d some b r i s t l e s ; segmen t 5 of fro n t t a r s u s ( F i g . 1 245 ) w i t h 1 row of 1 8 l a me l l a l e ; apex w i t h a group of 4 l a m e l l a e ; m a x i l l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . M A L E ( see n o t e s ) : f u l l y w i n g e d ; l e n g t h 2 , 27 m m ; b l a ck , w i t h h e a d a n d a n t e n n a e b row n ; a n t e n n ae n o t d i s t a l l y t h i c k e n e d ; a n t e n n a l segmen t s i n fo l l o w i n g proport i o n s : 5 : 6 : 1 0 : 9 : 9 : 9 : 9 : 8 : 7 : 1 1 ; a n te n n a I segmen t 3 more t h a n t h ree t i mes as l o n g as broad ( 3 , 33 ) ; head d u l l , g r a n u l a ted ; fro n t a l l i n e a b sen t ; t e m p l e s d i s t i n c t ; occ i p i t a l ca r i n a i n v i s i b l e ; POL = 8 ; O l = 4 ; OOl = 5 ; sc u t u m d u l l , g r a n u l a t e d ; n o t a u l i ces i n comp l e te , reach i n g a p p rox i m a te l y 0 , 3 l e n g t h of scu t u m ; sc u t e l l u m g r a n u l a ted , d u l l , w i t h p o s te r i or s u rf ace smoo t h a n d w i t ho u t sc u l p t u re ; m e t a n o t u m smoo t h , sh i n y , w i t h o u t scu l p t u re ; p ropod e u m ret i c u l a t e r u g ose , g r a n u l a t ed , w i t h o u t k ee l s ; fore-w i n g h y a l i n e , w i t h o u t d a r k t r a n sver s a l b a n d s ; d i s t a l p a r t of rad i a l v e i n near l y a s l o n g a s p rox i m a l p a r t ( 1 0 : 9 ) ; dorsa l p rocess of gonoforceps ( F i g . 1 246 ) p o i n te d , m u c h shorter t h a n gonoforce p s ; m a x i I l ary p a l p i w i t h 6 segme n t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . lOCUS T Y P I C US : Bu n d aberg ( A u s t r a l i a ) TYP I CAL M A T E R I AL : l ecto t y p e F ! a n d 2 p a r a l ec t o t y p e s ( 1 F , 1 M ) ! ( de s i g n a ted b y O l m i 1 98 2 ) i n B .

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HOST S : i n A u s t r a l i a Deltocephalus sp . , Nephotettix sp . ( R . C . L . Perk i n s 1 906c ) D I ST R I B U T I O N : A u s t r a l i a n re g i o n : AUSTRAL I A : B u n d a b erg ( Queen s l a n d ) , B ! N O T E S : t h e on l y k n o w n m a l e c a n b e l o n g to G . op t a b il i s or decoratus ( R . C . L . Perk i n s 1 9 1 2 ) . •

1 0 . Gonatopus decoratus ( R . C . L . P erk i n s ) n . comb .

Ch alcogon a topus decora tus R . C . L .

Perk i n s 1 905 : 47 .

F E MAL E : ap terou s ; l e n g t h 3 , 29 m m ; head brow n , w i t h m a n d i b l e s , c l y p e u s , fron t p a r t o f ver tex a n d a n ten n a l seg men ts 1 - 2 y e l l ow ; p ronot u m brow n , w i t h a n t e r i o r m a r g i n of co l l a r , s i des a n d poster i or marg i n of d i sc y e l l o w ; scut u m , sc u te l l u m a n d m e t a t horax + p ropodeum redd i s h ; head exc a v a t e d , smoo t h , w i t h ou t scu l p t u re ; prono t u m crossed b y a t r asvers a l i m pres s i on ;

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met a t horax + p ropodeum g r a n u l a te d ; meso-me t a p l e u r a l s u t u re d i s t i nc t a n d comp l e t e ; poster i o r 3 u rf ace o f propode u m t r a n s verse l y s t r i a te ; seg men t 4 of fro n t t a r s u s near l y as l on g as segme n t 1 ( 30 : 29 ) ; en l arged c l a w ( F i g . 1 247 ) w i t h o u t s u b a p i c a l too t h , w i t h a sm a l l too t h a t the end of a l on g i t u d i n a l fu rrow a n d 1 peg - l i k e h a i r ; segment 5 of fron t t a r s u s ( F i g . 1 247 ) w i t h 1 row of 1 8 l a me l l ae ; apex w i t h a g roup of a t l ea s t 5 l a me l l ae ;

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m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segme n t s ; t i b i a l spurs 1 , 0 , l . MALE : u n k no w n ( see note a t descr i p t i o n of m a l e of G . op ta bilis ) LOCUS TYP I C U S : B u n d a b e r g ( A u s t r a l i a ) TYP I C A L M A T E R I AL : l ec t o t y p e F ! ( d es i g n a t e d b y O l m i 1 982 ) i n B . HOST S : i n A u s t r a l i a Phryn ophyes s p . ( R . C . L . Perk i n s 1 906c ) . D I ST R I B U T I O N : A u s t r a l i a n reg i o n : AUSTRAL I A : B u n d a berg ( Qu e en s l a n d ) ,

1 1 . Gon a top u s a u stra l i a e R . C . L Perk i n s

Gon a top u s a u s traliae R . C . L . Perk i n s 1 90 5 : 4 1 . Neogona top u s dubiosus R . C . L . Perk i n s 1 90 5 : 44 ( n . s y n . ) ; locus typicus B u n d a b erg ( Q ueen s l a n d , Austra I i a ) . Gon a top u s koebelei R . C . L . Perk i n s 1 906a : 489 ( n . sy n . ) ; locus typicus P a r a m a t t a ( N ew Sou t h Wa l es , A u s t r a l i a ) . Dicondylus a u s traliae ( R . C . L . Perk i n s ) : K i effer 1 907b : 2 2 .

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1 79 1

F E MAL E : a p terou s ; l e n g t h 2 , 63-3 , 36 m m ; h e a d , p ro n o t u m a n d scu t u m t e s t aceous-red d i s h ; a n t e n n a e t e s t aceo u s , w i t h seg m e n t s 4- 1 0 o r 6- 1 0 d a r k ; sc u te l l u m , m e t a t h o r a x + p ropodeum a n d a b domen re d d i s h -b row n , exce p t for apex of propodeum w h i c h is y e l l o w ; l e g s t e s t aceou s ; head exc a v a te d , sh i n y , w i t h o u t scu l p t u re ; p ron o t u m crossed b y a t r a n sver s a l i m p ress i o n , sh i n y , smoo t h , w i t h o u t scu l p t u re ; sc u t u m s h i n y , w i t h some l on g i t u d i n a l k ee l s ; met a n o t u m sh i n y , smoo t h , not t r a n sverse l y s t r i a t e ; m e t a t h orax + p ropode u m w i t hou t scu l p t u r e , s h i n y ; p l e u r a not t r a n s v erse l y s t r i a t e ; poste r i o r su rface o f propodeum t r a n sve rse l y s t r i a t e ; meso-me t a p l e u r a l s u t u re obso l e t e ; segmen t 1 of fron t t a r s u s as l on g as s e g m e n t 4 ; en l a r ged c l a w ( F i g . 1 248 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l tooth at the end of a l o n g i t u d i n a l f u rrow and 4-6 p e g- l i k e h a i r s ; se g m e n t 5 of fro n t t a r s u s ( F i g . 1 248 ) w i t h 1 r o w of 1 4- 1 6 l a m e l l ae ; a p e x w i t h a g r o u p of 7 - 1 4 l a me l l ae ; m a x i l l a ry p a l p i w i t h 4-5 segmen ts ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : Bu n d aberg ( A u s t r a l i a ) TYP I C A L M A T E R I AL : l ec t o t y p e F ! a n d 2 p a r a l ec t o t y pes F F ! ( d es i g n a ted b y O l m i 1 98 2 ) of G . a u stra l i a e i n B ; l e c t o t y pe F ! a n d 2 p a r a l ec t o t y pes F F ! ( des i g n a t ed by O l m i 1 982 ) of N. dubiosus i n B ; l ec t o t y p e F ! ( d es i g n a t e d b y O l m i 1 982 ) of G . koebelei i n B . HOSTS : i n A u s t r a I i a Deltocep h a l u s sp . , A thysa n u s sp . , Nephotettix sp . ( R . C . L . P e r k i n s 1 906c ) . D I ST R I BU T I ON : A u s t r a l i a n reg i o n : AUSTRAL I A : B u n d a berg ( Q ueens l a n d ) , OL ! B ! M y o r a Sp r i n g s ( N . S t r a d b ro k e I s . , Queen s l a n d ) , UQ ! P a r a m a t t a ( Ne w Sou t h W a l e s ) , B !

1 2 . Gon a top us pulcherrimus

(R.C.L.

Perk i n s ) n . com b .

Neogon a top u s pu lcherrimus R . C . L . Perk i n s 1 90 5 : 4 4 .

F E M AL E : a p terou s ; l e n g t h 3 , 29 m m ; red d i s h - t e s t aceou s , w i t h p e t i o l e b l ac k a n d a b domen b row n - b l ack ; a n ten n a e b row n , w i t h seg men t s 1 -3 a n d 1 0 y e l l o w ; h e a d e x c a v a ted ; p ro n o t u m crossed b y a t r a n sversa I i m p res s i o n ; poste r i o r s u rf ace of p ro p o d e u m t r a n sverse l y s t r i a t e ; meso - m e t a p l e ura l s u t u re obso l e t e ; se g m e n t 4 of fro n t t a r s u s n e a r l y as l o n g as s e g m e n t 1 ( 20 : 22 ) ; en l a r g e d c l a w ( F i g . 1 249 ) w i t h o u t s u b a p i c a l too t h , w i t h a s m a l l too t h a t t h e e n d of a l o n g i t u d i n a l fu rrow a n d 4 p e g - l i k e h a i r s ; se g m e n t 5 of fro n t t a r s u s ( F i g . 1 249 ) w i t h 1 row of 1 6 l a m e l l ae ; a p e x w i t h a g r o u p of 1 3 l a m e l l a e ; m a x i l l a r y p a l p i w i t h 4 segme n t s ; l a b i a l p a l p i w i t h 2 s e g me n t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k no w n L O C U S TYP I C US : B u n d a b e r g ( Au s t r a l i a ) TYP I C A L MAT E R I AL :

l ec to t y p e F !

( des i g n a ted b y O l m i 1 982 )

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1 792

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HOST S : in A u s t r a l i a Nep h o tettix plebeius ( R . C . L . Perk i n s 1 906c ) . D I ST R I B U T I ON : A u s t r a l i a n reg i on : AUSTRAL I A : B u n d a be r g ( Qu e e n s l a n d ) ,

Fi g .

1 250 - F em a l e of Gynochelys braunsi Brues ( t y p e )

1 8 . Gynochelys B r u e s

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T Y P E S P E C I E S : Gyn ochel u s b r a u n s i B r u e s 1 90 6 , monot y p i c . F E MALE ( F i g s 1 250, 1 25 1 ) : m i crop terou s ; m a x i I l a ry p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; p ron o t u m ( F i g . 1 25 1 ) crossed b y a s t ro n g t r a n s v e rs a l i m p ress i o n ; e n l a rged c l aw ( F i g . 1 252 ) w i t h s u b a p i c a l too t h a n d l a m e l l ae ; p ro n o t u m not p rod uced i n to l obes d i rec ted tow a r d s t h e t e g u l a e ; fore troc h a n ters a p p rox i m a te l y four t i mes a s l o n g a s b road ; 1 , 0, 1 . M A L E : u n k no w n D I ST R I BU T I O N : E t h i o p i a n H O ST S : u n k n o w n SPEC I E S : 1

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1 25 2 - C h e l a of Gynochelys b r a u n s i Brues ( t y p e )

F EM A L E ( F i g s 1 25 0 , 1 25 1 ) : m i crop t erou s ; l en g t h 4 , 00 m m ; redd i s h , w i t h b ro w n s p o t s on ver tex , prono t u m a n d p l e u r a ; p e t i o l e b l a ck ; h e a d d u l l , g r a n u l a ted ,

w i t h i r reg u l a r r u g os i t i e s a n d s t r i ae ;

v er tex sw o l l e n ; occ i p i t a l

Gon a t opod i n a e

1 79 5

c a r i n a i n com p l e te, o n l y v i s i b l e beh i n d post e r i or oce l l i a n d on t h e s i d es of t h e oce l l a r t r i a n g l e ; a n t e n n a e very s l e n de r a n d l o n g ; a n te n n a l seg m e n t s i n fo l l ow i n g p ropor t i on s : 1 3 : 7 : 30 : 28 : 27 : 1 2 : 1 1 : 9 : 8 : 1 2 . seg m e n t 3 a p p ro x i m a te l y e l e v e n t i me s as l o n g as b ro a d ; POL = 6 ; OL = 5 ; OOL = 1 0 ; O P L = 1 ; t e m p l e s d i s t i n c t ; pron o t u m crossed b y a t r a n sver s a l i m p ress i o n , d u l l , g r a n u l a t e d , poste r i o r l y n o t prod u ced i n to l ob e s ; sc u t u m d u l l , g r a n u l a te d , a p prox i m a t e l y c i rc u l a r , a s l o n g as co l l a r ; n o t a u l i ces d i s t i n c t a n d co m p l e t e , poste r i o r l y se p a r a t ed ; fore w i n g s red uced , a s l o n g a s sc u t u m ; h i n d w i n g s a b s e n t ; sc u t e l l u m sh i n y a n d smoot h , n e a r l y 0 , 5 a s l o n g a s sc u t u m ; met a n o t u m sh i n y a n d smoo t h , n e a r l y 0 , 5 a s l o n g a s sc u te l l u m ; p ro p o d e u m h u m p e d ( a s i n a p terous spec i e s o f G o n a topod i n es ) , g r a n u l a te d , d u l l ; p l e u r a a n d pos te r i or s u r f ace t r a n sverse l y s t r i a t e ; a n te r i o r s u rf ace g r a n u l a te d a n d r u g ose ; meso-me t a p l e u r a l s u t u re com p l e t e , d i s t i n c t ; fore t roc h a n ters a p p rox i m a t e l y f o u r t i me s a s l o n g a s b ro a d ; fore t a rsa l s e g me n t s i n fo l l o w i n g propor t i o n s : 1 6 : 3: 5 : 1 8 : 28 ; en l a r ged c l a w ( F i g . 1 25 2 ) w i t h s u b a p i c a l tooth a n d 7 l a m e l l a e ; seg m e n t 5 of fron t t a r s u s ( F i g . 1 25 2 ) w i t h 2 r o w s o f 2 0 l a me l l ae ; a p ex w i t h a g ro u p o f 1 2 l a m e l l a e ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 0 , 1 . MALE : u n k n o w n L O C U S TYP I C U S : A l goa B a y ( So u t h Af r i c a ) TYP I CAL M A T E R I AL : h o l o t y p e F ! i n M L . H OSTS : u n k n o w n D I ST R I B U T I O N : E t h i o p i a n reg i o n : SOU T H A F R I C A : A l go a Bay Prov i n ce ) , M L !

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1 0 . S U B FA M I L Y A PODRYININAE N . S U B F .

T Y P E G E N U S : Apodryi n u s n . gen . F E MA L E ( F i g s 1 5 , 1 253 ) : ap terou s ;

m a x i l l a r y p a l p i w i t h 4 segme n t s ;

l a b i a l p a l p i w i t h 3 segmen t s ; m a n d i b l es w i t h four i r reg u l a r tee t h ; occ i p i t a l c a r i n a a b sen t ; oce l l i a b sen t ; suboc u l a r s u l c u s a b sen t ; meSOSoma composed b y f u sed segmen t s ; meso soma su t u re s comp l e t e l y ob l i t e r a t e d , excep t f o r a s u t u re b e t ween prope c t u s a n d a p p aren t p ron o t u m a n d a l a tera l t r ack of s u t u re b e t w een a p p a re n t pro n o t u m a n d mesop l eu r a w h i c h are v i s i b l e ; fore troc h a n t e rs a p p ro x i m a t e l y t w i ce as l o n g as b ro a d ; segmen t 3 of fron t t a r s u s p roduced i n to a hook ; fron t t a r s u s ( F i g . 1 25 4 ) w i t h o n e on l y prea p i c a l l a m e l l a ; a n t en n ae w i t h o u t t u f t s of l on g h a i rs ; t i b i a l spurs 1 , 1 , 1 MALE : u n k no w n D I ST R I B U T I O N : Neot rop i c H O ST S : u n k n o w n GENERA : 1 SPEC I E S :

Apodryin u s n .

gen .

T Y P E S P E C I E S : Apodryinus masneri n . sp . F E M A L E ( F i g s 1 5 , 1 2 53 ) : a p t e ro u s ; m a x i l l a ry p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 3 segme n t s ; occ i p i t a l c a r i n a a b sen t ; oc e l l i a b sen t ; mesosoma composed b y f u sed segme n t s ; mesosoma su t u res comp l e t e l y ob l i t e r a t e d , excep t f o r a s u t u re b e t ween propec t u s a n d a p p a ren t pro n o t u m a n d a l a t e r a l t r ack of su t u re b e t ween a p p a r e n t pro n o t u m a n d mesop l eu r a w h i c h a re v i s i b l e ; a n t e n n a e w i t h rh i n a r i a on seg men t s 3- 1 0 ; en l a r g e d c l a w ( F i g . 1 254 ) w i t h s u b a p i c a l too t h a n d l a m e l l ae ; segmen t 5 of fron t t a rs u s ( F i g . 1 254 ) w i t h row s of l a me l l ae ; a p p aren t p ron o t u m n o t crossed b y t r a n s v e r s a l i m p ress i o n ; t i b i a l s p u r s 1 , 1 , 1 . MAL E : u n k no w n D I ST R I BU T I O N : Neot rop i c HOST S : u n k n o w n SPE C I E S :

Apodryin u s m a sneri n . sp .

F E MA L E ( F i g s 1 5 , 1 253 ) : a p t e ro u s ; l e n g t h 3 , 20-3 , 40 m m ; h e a d b l ac k , w i t h m a n d i b l e s , c l y p e u s a n d a n a rrow fron t a l reg i o n n e a r c l y p e u s t e s t aceo u s d a r k ; a n te n n ae t e s t aceo u s ; t h o r a x a n d p ropodeum b l ac k , excep t f o r p ropec t u s a n d a n t e r i o r re g i o n of a p p a re n t p r o n o t u m w h i c h are t e s t aceou s ;

1 797

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a b domen b l ack , w i t h a p e x t e s t aceo u s ; l eg s b row n , w i t h t a r s i l i g h t ; h e a d swo l l e n , d u l l , f u l l y ret i c u l a t e ru gose ; temp l es a n d face h a i r y ; temp l e s very l o n g , a p prox i m a t e l y t w i ce as l o n g as e y e s ; occ i p i t a l c a r i n a a b sen t ; oce l l i a b sen t ; fron t a l l i n e d i s t i nc t from v e r t ex to c l y p eu s ; c l y p e u s w i t h ven t r a l m a r g i n rou n d e d ; su boco l a r s u l c u s a b s e n t ; a n t e n n a l sock e t s touch i n g c l y p e u s ; a n te n n a e n o t d i s t a l l y t h i c kened ; a n t en n a I segmen t s i n f o l l ow i n g p ropo r t i o n s : 1 6 : 4 : 8 : 6 : 5 : 5 : 5 : 5 : 5 : 8 ; r h i n a r i a on segmen t s 3 - 1 0 ( t w o r h i n a r i a o n l a s t segmen t ) ; p ropec t u s n o t dorsa l l y v i s i b l e , v i s i b l e u n d er a n te r i or m a r g i n o f a p p a ren t p ron o t u m ; mesosoma comp l e t e l y composed of f u sed s e g me n t s ( excep t for p ropec t u s ) , so t h a t p ro no t u m , scu t u m , s c u te l l u m , met a n o t u m a n d p ropodeum a re n o t recog n i z a b l e ; on l y a t r ack of s u t u re i s l a ter a l l y v i s i b l e b e t w een a p p aren t p ron o t u m a n d a p p aren t mesop l eu r a ; mesosoma composed of a n a n te r i o r sh i n y , smoo t h

1 798

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I nst . ,

n o . 37,

1 984

reg i o n , prob a b l y t h e a p p a re n t pronot u m ; a p p aren t pron o t u m w i t h o u t scu l p t u r e , l a te r a l l y w i t h some l o n g i t u d i n a l fo l d s ; pos t rer i o r l y t h e mesosoma is f u l l y re t i c u l a te ru gose , sh i n y , w i t h o u t sc u l p t ure a mo n g areo l ae ; t h e sp i rac l es a r e v e r y prom i n e n t ; a w i d e dor s a l a rea beh i n d a p p aren t pro n o t u m is l e s s re t i c u l a t e ru gose t h a n o t h e r reg i on s : i t is per h a p s t h e a p p a re n t scu t u m ; t h e mesosoma i s cy l i n d r i c , w i t h a pos t e r i o r re g i on very i n c l i n e d ; pos t e r i o r re g i o n of a p p a re n t propode u m ret i c u l a te rugose, l a ter a l l y w i t h two dors a l prom i n e n t p o i n t s and two o t her l a tera l l e ss p rom i n e n t poi n t s ; meso-me t a p l e u r a l su t u re i n v i s i b l e ; fore t a r s a l segmen t s i n fo l l o w i n g proport i o n s : 1 8 : 3 : 4 : 1 5 : 1 8 ; segmen t 3 of fron t t a rs u s produced i n to a hook ; en l a r ged c l a w ( F i g . 1 254 ) w i t h a s u b a p i c a l too t h , a row of n e a r l y 1 8 l a me l l a e on i n ner s i d e a n d n u merous h a i rs on ex ter n a l s i d e ; segmen t 5 of f r o n t t a r s u s ( F i g . 1 254 ) w i t h two rows of 4 ( l o n g ) + 1 3 ( sh or t ) l a m e l l ae ; apex w i t h a g r o u p of n e a r l y 5 l a me l l ae ; m ax i l l a ry p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 1 , 1 . M AL E : u n k n o w n L O C U S TYP I C U S : N e a r Ma l a l c a h u e l l o ( M a l l ec o , C h i l e ) TYP I C A L M A T E R I AL : ho l o t y p e F ! a n d 1 p a r a t y p e F ! i n O T ; 1 p a r a t y p e F ! i n OL . HOST S : u n k n o w n ( t h e t y p i c a l m a t e r i a l w a s co l l ec t e d i n Nothofagus fores t ) D I ST R I B U T I O N : Neot rop i c reg i o n : C H I L E : N e a r M a l a l c a h u e l l o ( M a l l eco ) , OT ! OL ! 22 Km ESE Rec i n to ( M u b l e ) , O T !

NOMINr� lJUBIA A N D INCERTAE SEDIS S P E C I E S

T h e fo l l o w i n g n ames h a v e n o t been a s soc i a ted w i t h spec i e s i n t h e b o d y o f t h e tex t , b ec a u se e i t he r t h e t y pe spec i me n s seem to b e l o s t a n d t h e or i g i n a l desc r i p t i o n s a r e u n re l i a b l e , or b ec a u se t h e y a r e ma l es o r fem a l es n o t a s soc i a ted to a d i s t i n c t ge n u s .

a b dom ina lis ( Chelogynus a l b i dicollis v a r . ) O g l ob l i n 1 924b : 1 26 ; locus typicus K r a l ove H r a d ec ( Czechos l ov ak i a ) . T h e t y p i c a l m a te r i a l seems to be l o s t ( i t ' s n o t i n P G , P T , B A ) . T h e or i g i n a l descr i p t i o n i s u n re l i a b l e . a ffi n is (Prodryin u s ) A r l e 1 935 : 44 ; locus typ i c u s P e t ropo l i s ( A l to d a Serr a , Brazi I ) . Accord i n g to M r . A r l e ( v er b a l comm u n i c a t i on ) t h e t y pe spec i men F ( on l y k n o w n spec i men ) i s l o s t . T h e or i g i n a l descr i p t i on i s u n re l i a b l e . For t h i s spec i e s A r l e ( 1 9 35 : 46 ) proposed t h e n e w g e n u s Ctenodryin u s . a frica n u s (Aphelop u s ) Ben o i t 1 9 5 1 c : 1 5 ; locus typ icus M a y -y a-Moto ( Z a i re ) = A n taphelop u s a fri c a n u s ( Be n o i t ) : Beno i t 1 95 1 d : 2 3 . T h e h o l o t y p e M ! ( o n l y k n o w n spec i me n ) i s i n T V . T h e e m a r g i n a t ed o cc i p u t and u n sc u l p t u red p ropodeum p l ace t h i s spec i e s w i t h

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Gon a topodin a e ( t h e same group o f Aphelop u s inexpec t a t u s Ben o i t , w h i ch h a s on l y t h e cos t a l c e l l enc l osed ) . Desc r i p t i o n : fema l e u n k n o w n ; m a l e f u l l y w i n ged ; l e n g t h 1 , 8 m m ; h o l o t y p e very d a m a ged : h e a d , a b domen , l eg s a n d r i g h t w i n g s m i ss i n g ( or i g i n a l l y t h e y w e re p r e se n t ) ; accord i n g to t h e or i g i n a l desc r i p t i o n , co l o u r b l ac k , w i t h t i b i ae a n d t a r s i ye l l ow ( a c t u a l l y t h o r a x a n d p ropodeum a re b r o w n ) ; accord i n g t o t h e o r i g i n a l descr i p t i o n , h e a d w i t h f r o n s a n d vertex smoo t h , s h i n y ; fron t a l l i ne i ncom p l e t e , on l y a n t e r i o r l y v i s i b l e ; m a n d i b l es t r i d en t a t e ; c l y p e u s regu l a r l y r o u n d e d a n d convex ; a n t e n n a l segmen t s of a p p ro x i m a t e l y e q u a l l e n g t h s ; eyes h a i r y ; P O L l o n g e r t h a n O O l ; occ i p u t e m a r g i n a t e , c o n c a v e ; accord i n g t o my observ a t i o n s , scu t u m sh i n y , w e a k l y g r a n u l a t ed ; n o t a u l i ces i n comp l e t e , reach i n g a p p rox i m a te l y 0 , 5 l e n g t h of scu t u m ; scu te l l u m a n d m e t a n o t u m sh i n y , smoo t h , w i t h o u t scu l p t u r e ; p ro p o d e u m sh i n y , smoo t h , w i t h o u t scu l p t u re , n o t re t i c u l a t e r u gose ; meso p l e u ra a n d me t a p l e u r a s h i n y , smoo t h , w i t h o u t scu l p t u r e ; fore-w i n g h y a l i n e , w i t h o u t d a rk t r a n s versa l b a n d s , w i t h o n I y cos t a l ce l l c l e a r l y v i s i b l e ; acco r d i n g t o t h e o r i g i n a l descr i p t i o n , a b domen l a t e r a l l y f l at . a l b icoxis (A n teon) K i effer i n K i effer et M a r s h a l l 1 905 : 1 95 ; locus typicus Foret de 5 t -Germ a i n ( F r a nce ) . = A n teon a l b icoxis K i effer : K i effer 1 9 1 4 b : 1 58 . T h e t y p e ( on l y k n o w n spec i men ) w a s a m a l e or i g i n a l l y i n t h e D e G a u l l e co l l ec t i on . I t i s n o t i n P . T h e o r i g i n a l desc r i p t i on i s unrel i a b l e . a l b ipennis ( '., a beo) K i effer i n K i effer e t M a r s h a l l 1 90 5 : 222 ; locus typicus S i ofok ( H u n g a r y ) . = '., a berius a l b ipennis ( K i effer ) : K i effer 1 9 1 4 b : 6 1 . T h e type ( o n l y known spec i m en ) was a m a l e o r i g i n a l l y in the Szep l i g e t i co l l ec t i o n . I t is n o t in B U . T h e or i g i n a l desc r i p t i o n i s u n re l i a b l e . a l b i tarsis ( L a b eo ) K i effer 1 906a : 239 ; loc u s t yp icus O r m s b y Co . ( N ev . , U. S. A. ) . = L a berius a l b i t arsis ( K i effer ) : K i effer 1 9 1 4b : 64 . T h e h o l o t y p e M ! ( on l y k n o w n spec i me n ) i s i n C A . I t i s a G o n a topod i n e . Descr i p t i o n : f u l l y w i n g e d ; l e n g t h 1 , 6 m m ; b l a c k ; m a n d i b l es t e s t aceou s ; a n t e n n ae b r ow n ; l e g s b ro w n , w i t h t a r s i l i g h t ; a n t en n a e n o t d i s t a l l y t h i c k e n ed ; a n t e n n a l segme n t s i n fo l l ow i n g p ro p o r t i o n s : 5 : 3 : 6 : 6 : 6 : 6 : 6 : 6 : 5 ( l a s t segmen t m i s s i n g i n h o l o t y p e ) ; a n t e n n a l segm e n t 3 l ess t h a n t h ree t i m es a s l o n g a s b ro a d ; h e a d sh i n y , g r a n u l a t e d ; fron t a l l i n e a b sen t j f r o n s w i t h a med i an i m p ress i on i n f r o n t of a n t e r i o r oce l l u s ; occ i p i t a l c a r i n a i n v i s i b l e ; P O L = 7 ; O l = 3 , 5 ; O O l = 3 ; bre a d t h o f t h e oce l l i : 3 ; scu t u m sh i n y , g r a n u l a t ed ; n o t a u l i ces i n comp l e t e , rea c h i n g a p p ro x i m a t e l y 0 , 3 l e n g t h o f sc u t u m ; sc u t e l l u m a n d met a n o t u m sh i n y , very we a k l y g r a n u l a te d ; p ropodeum smoo t h , sh i n y , w i t h o u t scu l p t u re ; d o r s a l s u rf ace w i t h a med i a n f u rrow ; fore-w i n g h y a l i n e , w i t h o u t

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1 25 5 - M a l e ge n i t a l i a of C a "Yeo a l bosign a t u s K i effer ( n eo t y p e ) ; notch .

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d a rk t r a n svers a l b a n d s ; d i s t a l p a r t of ra d i a l v e i n l on g e r t h a n p rox i m a l p a r t ; a b domen , i n c l u d i n g g en i t a l i a , m i ss i n g i n ho l o t y p e ; ma x i l l a r y p a l p i w i t h 3 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 1 , 2 . T h e u n scu l p t u red propodeu m s u g ge s t s t h a t t h e spec i es m a y b e a n k n ow n : A crodontochelys . I n t h e Nearc t i c reg i o n on l y t w o spec i e s a r e mimus a n d americ a n u s . T h e m a l e of m im u s i s k now n : i t i s d i ffere n t from a l b i tarsis . T h e m a l e o f american u s i s u n k n ow n . I t i s possi b l e t h a t a l b itarsis i s t h e m a l e of american u s . a lbosign a t u s {La beo} K i effer i n K i effer e t M a r sh a l l 1 905 : 224 ; locus typicus S v a b h e g y ( H u n g a r y ) . = L a b erius a l b o s ign a tu s ( K i effer ) : K i effer 1 9 1 4 b : 6 2 . T h e t y p e ( on l y k n o w n spec i men ) w a s a m a l e i n t h e Szep l i g e t i co l l e t i o n . I t i s n o t i n B U . T h i s spec i es seems ea s i l y recog n i z a b l e b y i t s p a r t l y y e l l ow - t e s t aceous face . Some m a l es i n BU a n d P ag ree w i t h t h e or i g i n a l desc r i p t i o n . I h a v e des i g n a t e d a neo t y p e f ro m V acz

Nom i n a d u b i a

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( H u n g a r y ) i n BU . D i s t r i b u t i o n : P a l aearc t i c reg i o n : H U NGARY : V a c z , B U ! SzSz t M i k l o s , BU ! S v a b h e g y ( Bu d a p e s t ) ( K i effer i n K i effer e t M a r sh a l l 1 905 ) ; SW I T Z E R L A N D : L u try , LO ! FRAN C E : Hort u s ( He r a u l t ) , P ! H os t s : i n F r a n c e Rib a u todelp h a x imitans R i b a u t ( Delp h acidae ) . T h i s spec i es i s p r o b a b l y a Pseu dogon a top u s . Descri p t i on : fema l e u n k n ow n ; m a l e f u l l y w i n ged ; l en g t h 1 , 68 - 1 , 84 m m ; b row n , w i t h v e n t r a l s i d e of h e a d , c l y pe u s , m a n d i b l e s , a n t e r i or s u rf ace a n d l a t e r a l reg i o n s of f r o n s ( espec i a l l y a l o n g orb i t s ) y e l l ow - t e s t aceo u s ; a n t en n a l segmen t s i n fo l l ow i n g p roport i o n s : 5 : 5 : 7 : 7 : 6 , 5 : 6 , 5 : 6 : 6 : 6 : 9 ; a n ten n a I segmen t 3 l es s t h a n t h ree t i mes a s l o n g a s b ro a d ( 2 , 8 ) ; h e a d d u l l , g r a n u l a ted ; fron t a l l i n e a b sen t ; occ i p i t a l c a r i n a i n d i s t i n c t b e h i n d oce l l i ; POL = 7 ; O L = 2 ; OOL = 3 ; smoo t h ov a l a re a v r s i b l e b e t w een e y e s a n d poster i or oce l l i ; scu t u m , scu t e l l u m a n d m e t a n o t u m g ra n u l a ted ; n o t a u l i ce s comp l e t e , poster i or l y sep a r a ted ; m i n i m u m d i s t a n ce b e t w een n o t a u l i ces shorter t h a n b rea d t h of oce l l i ( 1 : 2 ) ; p ropode u m ret i c u l a te ry gose, exce p t for a sma l l s u rface n e a r met a n o t u m w h i ch i s smoo t h a n d w i t h o u t scu l p t u re ; dor s a l s u rf ace w i t h a deep med i a n f u rrow ; fore- w i n g h y a l i n e , w i t h o u t d a r k t r a n svers a l b a n d s ; r a d i a l v e i n re g u l a r l y c u rved ; r a d i a l ce l l open ; dor s a l p rocess o f gon oforceps ( F i g . 1 25 5 ) s l i g h t l y shorter t h a n gonoforce p s , b ro a d e n e d ; b a s i v o l se l l a w i t h notch ; m a x i l l a ry p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . americ a n u m (Prenan teon ) K i effer 1 9 1 4 a : 9 1 ; locus typicus W i sco n s i n (U. S. A. ) . = Pren a n teon american u s K i effe r : K i effer 1 9 1 4b : 2 1 2 . T h e t y p e ( on l y k n o w n spec i m en ) w a s a m a l e i n t h e B a k e r co l l ec t i on . I t seems to be l o s t ( i t i s n o t i n W A , A M , B M , C A , e M , D A , L A , PH ) . T h e or i g i n a l desc r i p t i o n i s u n re l i a b l e . a n doi (Pachygon a top u s ) E s a k i a n d H a sh i m o t o 1 93 5 : 2 2 ; locus typ icus O i t a ( Kyushu, Japan ) . T h e t y pe F ( on l y k n o w n spec i men ) seems to b e l o s t . T h e or i g i n a l descr i p t i o n i s u n re l i a b l e . T h e h o s t i s Deltocep h a lu s dors alis Mo t s . ( E s a k i a n d H a sh i moto 1 935 ) . angus ticollis (Chelogyn u s ) Og l ob l i n 1 9 24b : 1 25 ; locus typ ic u s C e s k a L i p a ( C zecho s l o v a k i a ) . T h e t y p e spec i me n s of t h e O g l o b l i n spec i es from Czechos l o v a k i a seem t o be l os t ( t hey a re not in P G , P T , BA ) . The or i g i n a l desc r i p t i o n s a r e u n re l i a b l e . areola tu s ( A p h elop u s mela leucus v a r . ) K i effer i n K i effer e t M a r s h a l l 1 90 5 : 2 1 8 ; locus typ i c u s M a i so n s - L a f f i t t e ( F r a n ce ) . = A p h elop u s areola tu s K i effer : K i effer 1 9 1 4b : 2 1 6 . T h e t y p e ( o n l y k n o w n spec i men ) w a s a m a l e i n t h e D e G a u l l e co l l e c t i o n . I t i s n o t i n P . T h e or i g i n a l descr i p t i o n i s u n re l i a b l e . a terrimus (A n teon ) H e l l e n 1 9 1 9 : 284 ; locus typ icus u n k n ow n . T h e t y p e M shou l d b e i n H E . I d on ' t k n o w n t h i s form ( d e sc r i bed a s a b e r r a t i o n of An teon longicornis Da l m a n ) . T h e ori g i n a l desc r i p t i o n i s u n re l i a b l e .

1 802

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berl a n di (Dryin u s ) Be n o i t 1 953b : 4 1 1 ; locus typ icus Pe r i n e t ( M a d a g a scar ) . T h e h o l o t y p e ( on l y k n own spec i men ) i s a fema l e i n P . I t i s v e r y d a m a g e d : t horax a n d propodeum are m i ss i n g . T h e or i g i n a l d esc r i p t i o n i s u n r e l i a b l e . T h e s y s tem a t i c po s i t i o n i s n o t recog n i z ab l e . b ifa s c i a t u m (A n teon ) K i effer 1 908 : 7 ; locus typicus Be l i ze . A n teon b ifa scia t u s ( K i ef f e r ) : K i effer 1 907b : 2 5 . = Callian teon b ifa s c i a t u s ( K i ef f er ) : K i effer 1 9 1 3 b : 300 . = Chelogyn u s i:J ifa s c i a t u s ( K i effe r ) : K i e ffer 1 9 1 4b : 1 88 . T h e t y pe ( o n l y k n o w n spec i men ) w a s a fema l e i n t h e B a k e r c o l l ec t i o n . I t see ms to b e l o s t ( i t i s n o t i n WA, A M , BM , C A , C M , D A , LA, P H ) . T h e or i g i n a l desc r i p t i on i s u n r e l i a b l e . K i effer ( 1 9 1 3 b ) p roposed for t h i s spec i e s t h e n e w g en u s Ca llian teon . b ifa s c i a t u s (Dryin u s ) Say 1 828 : 8 1 ; locus typicus I n d i a n a . ( U . S . A . ) . = Prodryin u s ( ?) bifa s c i a t u s ( S ay ) : K i effer i n K i e ffer e t M a r s h a l l 1 906 : 498 . = Prodryin u s ( ? ) bifa s c i a t u s ( S ay ) : K i effer 1 9 1 4b : 5 3 . T h e t y p e ( o n l y k n o w n spec i men ) w a s a fem a l e i n t h e S a y co l l ec t i o n . I t seems to be l os t . T h e or i g i n a l descr i p t i on i s u n re l i a b l e . brasiliensis (Ch elogyn u s ) A r t e 1 9 3 5 : 48 ; locus t ypicus R i o d e J a n e i ro- G a v e a ( Brazi I ) . Accord i n g to Mr . R . A r l e ( v erb a l commu n i c a t i on ) t h e t y p e F ( on l y k n o w n spec i men ) i s l o s t . T h e or i g i n a l desc r i p t i on i s u n re l i a b l e . brevifilis (A n teon ) K i effer i n K i effer e t Mars h a l l 1 906 : 5 1 7 ; locus typicus Eng l and . = Chelogyn u s brevifilis ( K i effe r ) : K i effer 1 9 1 4 b : 1 83 . T h e t y p i c a l m a t e r i a l ( a t l e a s t t w o fem a l es ) w a s i n t h e M a r s h a l l co l l ec t i o n . I t seems to be l os t ( i t i s n o t i n BU ) . T h e or i g i n a l descr i p t i on i s u n re l i a b l e . capicola (Gon a top u s ) K i effer 1 9 1 3 : 324 ; locus typicus C a p e T o w n ( Sou t h Afr i c a ) . = Gon a top u s cap icola K i effer : K i effer 1 9 1 4b : 1 20 . T h e t y p e ( on l y k n o w n spec i men ) w a s a fem a l e i n t h e Sou t h Af r i c: C' t : M u se u m ( C a pe Tow n , Sou t h A f r i c a ) . Accor d i n g to t h e c u r a t o r , t h e.. t y pe seems to be l o s t . T h e or i g i n a l desc r i p t i o n i s u n r e l i a b l e . carin a tu s ( Ap helop u s melaleucus v a r . ) K i effer i n K i e ffer e t M a r sh a l l 1 90 5 : 2 1 9 ; locus typicus C a d der ( Scot l a n d , U n i t e d K i n g do m ) . = Aphelop u s carin a t u s K i e f fe r : K i effer 1 9 1 4 b : 2 1 7 . T h e t y p e ( o n l y k n o w n spec i men ) w a s a m a l e i n t h e C a meron c o l l ec t i on . It see ms to be l os t ( i t i s n o t i n BM ) . T h e or i g i n a l descr i p t i o n i s u n re l i a b l e . citrinicollis (An teon / K i effer i n K i effer e t M a r s h a l l 1 905 : 1 59 ; locus typicus B u d a p e s t ( H u n g ary ) . = A n teon citrinicollis K i effe r : K i e ffer 1 9 1 4 b : 1 50 . T h e t y p e ( o n l y k n o w n spec i men ) w a s a fem a l e i n t h e Sze p l i g e t i co l l ec t i o n . I t seems t o b e l os t ( i t i s not i n BU ) . accord i n g t o t h e or i g i n a l desc r i p t i o n , t h e spec i es shou l d b e a s y n o n y m of

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1 256 - M a l e gen i t a l i a of Pseu dogon a top u s corn u t u s Beno i t

1 803

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A n teon eph ippiger ( D a I m a n ) o r A . ful viventre ( H a I i d a y ) . c1aricollis (A n teon ) K i effer i n K i effer e t M a r s h a l l 1 90 6 : 5 1 4 ; locus typicus E n g I a n d . = Chelogynus c1aricollis K i effer : K i effer 1 9 1 4b : 1 8 1 . T h e t y p i c a l m a t er i a l ( a t l e a s t f o u r spec i me n s ) w a s composed of fem a l e s in t h e M a r s h a l l c o l l ec t i on . I t seems to b e l os t ( i t is n o t i n BU ) . Acco r d i n g t o t h e o r i g i n a l descr i p t i o n , p rob a b l y the spec i es i s a s y n o n y m of A n teon gaullei K i effer . conjunctus ( L a beo) K i effer i n K i effer et M a r s h a l l 1 905 : 223 ; locus typicus M a t y a sfe l d ( H u n g a r y ) . = L a b erius conjunctus ( K i effe r ) : K i ef f e r 1 9 1 4b : 6 2 . T h e t y p e ( on l y k n o w n spec i men ) w a s a m a l e i n t h e Szep l i g e t i co l l ec t i on . I t seem s to be l os t ( i t i s n o t i n BU ) . T h e o r i g i n a l desc r i p t i o n i s u n re l i a b l e . corn u t u s (Pseudogo n a top u s ) Beno i t 1 95 1 b : 1 59 ; loc u s typicus S w e l l e n d a m ( C a pe P rov i n c e , Sou t h A f r i c a ) . T h e t y p e ( on l y k n o w n spec i men ) i s a m a l e i n B M . I t b e l o n g s to the

1 804

Mem . Ame r . E n ! .

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1 984

G o n a topod i n e s . The gen u s is not i d en t i f i a b l e at the p rese n t t i me . Desc r i p t i on : fema l e u n k n ow n ; m a l e f u l l y w i n g ed ; l e n g t h 2 , 0 m m ; h e a d b l a ck , w i t h m a n d i b l e s y e l l ow ; a n t en n a e b r ow n ; t h o r a x a n d p ropodeum b l a ck ; a b domen red d i sh-brow n ; l e g s b row n - y e l l o w ; h e a d s h i n y , h a i ry , s t ro n g I y p u n c t a t e , w i t h o u t scu I p t u re a m o n g p u n c t u res ; f ro n t a I I i n e i n comp l e t e , shor t , v i s i b l e o n l y i n f ro n t of a n te r i o r oce l l u s ; smoo t h , sh i n y a r e a v i s i b l e b e t ween eyes a n d pos ter i or oce l l i ; i n te rn a l edges of th i s a rea n e a r i n te r n a l m a r g i n s of e y e s , p rom i n e n t a n d p o i n t ed ; en t i re a r e a s u rrounded b y a k ee l ; temp l e s d i s t i n c t ; occ i p i t a l c a r i n a i n v i s i b l e ; s c u t u m s h i n y , s t ron g l y p u n c t a t e ; n o t a u l i ce s com p l e t e , poste r i o r l y sep a r a ted ; m i n i m u m d i s t a n c e bet ween n o t a u l ices a s l o n g a s b r e a d t h of oce l l i ; scu t e l l u m and m e t a n o t u m sh i n y , smoo t h , w e a k l y p u n c t a t e ; p ropodeum v e r y w e a k l y re t i c u l a t e r u gose, w i t h o u t l o n g i t u d i n a l k ee l s o n post e r i or s u rf ace ; forew i n g s h y a l i n e , w i t h o u t d a rk t r a n s v e r s a l b a n d s ; dorsa l s u rface w i t h a med i a n f u rrow ; dor s a l p rocess of gonoforcep t s ( F i g . 1 256 ) s l en d e r , s h o r t a n d po i n t ed ; b a s i v o l se l l a w i t h o u t n o tch ; m a x i l l a ry p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 1 , 2 . decipiens (Gon a top u s ) P rov a n cher 1 887 : 1 79 ; loc u s typ icus C a p Rouge ( Q u e b ec , C a n a d a ) . = Gon a top u s decipiens P ro v a n ch e r : K i effer 1 9 1 4b : 1 23 . T h i s spec i e s , w h i c h former l y w a s a s soc i a ted w i t h d r y i n i d s , act u a l l y b e l o n g s to t h e Bethylidae (Bethylus de cipien s ) ( K rom b e i n 1 979 ) . deletus ( Th a u m a todryin u s ) B r u e s 1 93 3 : 1 47 ; locus typicus Ba I t i c a m b e r . T h e t y p e spec i me n s of t h e B r u e s foss i I spec i es were i n t h e a m b e r co l l ec t i on a t A l b e r t u s U n i v e rs i t a t , Kon i g s b e r g ( n ow K a l i n i n g r a d ) . T h e y seem to b e l os t . The o r i g i n a l desc r i p t i o n s a re u n r e l i a b l e . dors alis (A n teon ) K i effer i n K i effer e t M a r s h a l l 1 90 5 : 20 2 ; locus typicus M a i so n s - L a ff i t t e ( F r a nce ) . See neglectus (A n teon ) K i effer . dors a li s (Dryin u s ) Nees 1 83 4 : 372 ; loc u s typ ic u s Si ckersh a usen ( German y ) • nec A nteon dors alis K i effer i n K i effer et M a r s h a l l 1 905 : 202 . = A n teon dors a lis ( N ee s ) : K i effer 1 9 1 4 b : 1 5 1 . T h e t y p e spec i me n s of the Nees spec i e s seem to be l ost . The o r i g i n a l desc r i p t i on s are u n re l i a b l e .

dorsettensis (ffomopterophagus) D a l e 1 878 : 304 ; locus typ ic u s G l a n v i l l e ' s Wootton ( E n g l a n d ) . T h e t y p e spec i men seems l ost ( i t i s not i n BM ) . Acco r d i n g t o K i r k a l d y ( 1 906 ) , t h e descr i p t i o n of t h i s spec i es w a s b a sed o n e a r l y s t a g e . Accor d i n g t o R i c h a r d s ( 1 939 ) , i n t h e D a l e co l l ec t i on ( i n OX ) t h ere are t w o fem a l e s of Gona topus sepsoides West wood f rom G l a n v i l l e ' s Woo t to n . P rob a b l y ff. dorsetten s is is a j u n i o r s y n o n y m of G . sepsoides a n d t h erefore ffomop terophagu s i s a j u n i o r s y n o n y m of Gon a top u s . draco (Dryin u s ) M a rs h a l l i n K i effer et M a r sh a l l 1 90 5 : 83 ; loc u s typicus Se v i l l a ( Sp a i n ) .

1 805

Nom i n a d u b i a

�rodryin us ( ?) draco ( M a r s h a l l ) : K i effer 1 9 1 4b : 50 . T h e t y p e ( on l y k no w n spec i men ) w a s a fema l e i n t h e Me d i n a co l l ec t i o n . I t seems t o be l o s t ( i t i s n o t i n M O ) . T h e or i g i n a l descr i p t i o n i s u n re l i a b l e . =

dra c u n c u l u s (Dryin u s ) M a r s h a I I

i n K i effer e t M a rs h a I I

locus typicus C i u d a d Rea l ( Sp a i n ) . = Prodryin u s ( ?) dra c u n c u l u s ( M a r sh a l l ) :

1 905 : 82 ;

K i effer 1 9 1 4b :

50 .

T h e t y p e ( on l y k no w n spec i men ) w a s a fema l e i n t h e Med i n a co l l ec t i on . I t seems to be l os t ( i t i s n o t i n M O ) . T h e or i g i n a l descr i p t i on i s u n r e l i a b l e . dryiniformis (Gon a topu s ) M a r sh a l l i n K i ef fer e t M a r s h a l l 1 90 5 : 1 00 ; locus typicus C i u d a d Rea l ( Sp a i n ) . = Gonatopus dryiniformis M a r s h a l l : K i e ffer 1 9 1 4 b : 1 1 0 . T h e t y p e ( on l y k no w n spec i men ) w a s a fem a l e i n t h e Med i n a co l l ec t i on . I t seems t o b e l os t ( i t i s n o t i n MD ) . T h e or i g i n a l descr i p t i o n i s u n re l i a b l e . empoascae (Aphelop u s ) Ponomare n k o 1 970 : 428 ; locus typicus K a m e n k a ( Moscow P rov i nc e , U . S . S . R . ) . The t y p i c a l m a t e r i a l was n o t s t u d i ed , b ec a u se m i s s i n g in PK, M O , L N . T h e or i g i n a l desc r i p t i o n i s u n re l i a b l e . erythrocep h a l u s (Gonatop u s ) Nees 1 834 : 386 ; locus typ ic u s F r a n c e . = Gonatopus erythrocep h a l u s Nees : K i effer et M a r s h a l l 1 90 5 : 1 06 . = Gonatop u s erythrocep h a l u s Nees : K i effer 1 '; 1 4b : 1 1 1 . T h e t y p e s p ec i me n s of t h e Nees spec i es seem t o b e l os t . T h e o r i g i n a l descr i p t i on s a r e u n re l i a b l e . esakii (Anteon) Y a s u m a t s u 1 960 : 2 1 ; locus typicus H i k o s a n ( F u k u o k a P ref . , Kyushu, Japan ) . T h e t y p i c a l m a t e r i a l ( co m posed of m a l e s a n d fe m a l e s ) seems to be l os t . The or i g i n a l desc r i p t i on i s u n re l i a b l e . fennicus (Anteon) H e l l e n 1 9 1 9 : 285 ; locus typicus L ok a l a k s ( Ab , F i n l a n d ) . T h e t y p e ( on l y k no w n spec i me n ) w a s a fem a l e i n H E . I don ' t k n o w n t h i s spec i es . T h e or i g i n a l descr i p t i o n i s u n re l i a b l e . fen toni (Neogona top u s ) F re y t a g 1 977 : 574 ; locus typicus Lex i n g to n ( Ky . , U. S . A. ) . T h e h o l o t y p e M ! a n d 1 p a r a t y p e M ! are i n W A ; 3 p a r a t y p es MM are in K E . F re y t a g ( 1 977 ) desc r i bed fen toni a s Neogon a top us on the b a s i s o f some morpho l o g i c c h a r a c te r i s t i c s ( see d i sc u ss i on o n t a xonomy of m a l e Gon a topod i ne s o n p a g e 1 054 ) . Accord i n g to m y s t u d i e s , these c h a racte r i s t i cs a re not s a t i sfac tory . The rea l g e n u s is t h e refore n o t i de n t i f i a b l e . Desc r i p t i on : f em a l e u n k n o w n ; m a l e f u l l y w i n ged ; l en g t h 2 , 0- 2 , 5 m m ; b l ac k , w i t h m a n d i b l e s y e l l ow ; 3 n t e n n ae a n d a b domen b row n ; l eg s y e l l o w , w i t h cox a e p a r t l y b row n ; a n t e n n a I segmen t s i n fo l l o w i n g p ropor t i o n s : 8 : 6 : 1 0 : 1 1 : 1 1 : 1 1 : 1 0 : 1 0 : 1 0 : 1 3 ; a n t e n n a l seg m e n t 3 more t h a n t h ree t i mes a s l o n g a s b ro a d ( 4 , 4 ) ; h e a d sh i n y , h a i ry , g r a n u l a t ed ; f ro n t a l l i n e comp l e t e ; P O L = 8 ; O L = 3 ; OOL = 2 ; b r e a d t h o f oce l l i : 4 ; occ i p i t a l c a r i n a a b sen t ; t e m p l es d i s t i n c t ; scu t u m s h i n y , h a i ry , g r a n u l a ted ; n o t a u l i ces com p l e t e , poster i or l y

Mem . Ame r .

1 806

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no .

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1 984

sep a r a t ed ; m i n i m u m d i s t a nce b e t w een not a u l i ces shorter t h a n b r e a d t h of oce l l i ( 1 , 5 : 4 ) ; scu te l l u m a n d m e t a n o t u m sh i n y , h a i r y , w e a k l y p u n c t a t e ; p ropod e u m re t i c u l a te r u g o se , w i t h o u t k ee l s ; forew i n g s h y a l i ne ,

w i t h o u t d a r k t r a n svers a l b a n d s ;

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1 25 7 - M a l e ge n i t a l i a of Neogon a top u s fen toni F re y t a g ( p a r a t y p e )

gono forceps ( F i g . 1 25 7 ) n e a r l y a s l o n g a s gonoforce p s , a p i c a l l y b roaden ed ; b a s i v o l se l l a w i t h notc h ; m a x i l l a ry p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 1 , 2 . H o s t s u n k n ow n . D i s t r i b u t i on : N e a rc t i c reg i on : U . S . A . : Lex i n g t on ( K y . ) , WA ! K E ; Mercer Co . , ( K y . ) , K E !

filicorn is (A n teon ) K i effer i n K i effer et M a r s h a l l 1 905 : 1 97 ; locus typicus u n cer t a i n ( Scot l a n d or I t a l y ) . = A n teon filicornis K i e ffe r : K i effer 1 9 1 4 b : 1 58 . T h e t y p i c a l m a t er i a l ( m a l es ) w a s composed of spec i m e n s from Scot l a n d i n t h e C a meron co l l ec t i o n , a n d from T r i es t e ( I t a l y ) i n t h e G r aeffe co l l ec t i on . These spec i me n s seem to b e lost ( t h e y are n o t in 8 M , M I a n d V I ) . T h e or i g i n a l desc r i p t i o n i s u n re l i a b l e .

Nom i n a d u b i a

1 807

filicornis (Dryin u s ) Brues 1 92 3 : 343 ; locus typ icus B a l t i c amber . = Th a u m a todryin us filicornis ( B r u es ) : Brues 1 9 33 : 1 47 . = Harp a c tosphecion filicornis ( B rue s ) , H a u p t 1 944 : 92 . T h e t y p e spec i me n s of t h e Brues fos s i l spec i es were i n t h e amber c o l l ec t i o n a t A l b e r t u s U n i vers i t a t , Kon i g s b e r g ( no w K a l i n i n g r a d ) . T h e y seem to be l o s t . T h e or i g i n a l desc r i p t i o n s a re u n re l i a b l e . flavipes (A n teon) K i ef f e r i n K i e ffer e t M a r s h a l l 1 905 : 1 49 ; locus typicus M a i so n s - L a f f i t t e ( F r a n ce ) . T h e t y p e ( on l y k n o w n spec i men ) w a s a m a l e i n t h e De G a u l l e c o l l ec t i o n . I t seems to be l os t ( i t i s n o t i n P I . T h e o r i g i n a l descr i p t i o n i s u n re l i a b l e . Accor d i n g to K i e ffer ( 1 9 1 4 b ) A . flavipes i s t h e m a l e o f A . fla vicorne ( D a l m a n ) . formicaria (Mu til l a ) J u r i n e 1 807 : 266 ; locus typ icus Sw i t z er I a n d . T h e t y p e spec i men see ms to be l o s t . I t i s n o t i n GV . T h e or i g i n a l descr i p t i o n i s u n re l i a b l e : see a l so ichneumonides (Me thoc a ) L a t r e i l l e . fulvicollis (Chelogyn u s ) Hed i c k e 1 926 : 244 ; locus typicus Be l l i n c h e n ( Germany ) • T h e t y p e ( on l y k n o w n spec i men ) w a s a fem a l e . I t shou l d be i n H U . I d on ' t k n ow t h i s spec i es . T h e o r i g i n a l descr i p t i o n i s u n re l i a b l e . fuscicollis (A n teon ) K i effer i n K i effer e t M a r s h a l l 1 90 6 : 5 1 3 ; locus typ icus Botu s f l e m i n g ( Co r n w a l l , E n g l a n d ) . = A n teon fu scicollis K i ef f e r : K i e ffer 1 9 1 4b : 1 50 . T h e t y p e ( on l y k n o w n spec i m en ) w a s a fem a l e i n t h e M a r s h a l l co l l ec t i o n . I t seems to be l os t ( i t i s n o t i n B U ) . T h e o r i g i n a l desc r i p t i on i s u n re l i a b l e . gir a u di (A n teon) K i effer i n K i e ffer e t M a r s h a l l 1 90 5 : 204 ; locus typicus u n cer t a i n ( H u n g a r y or A u s t r i a ) . T h e t y p i c a l m a t e r i a l ( m a l e s ) w a s composed of spec i me n s f rom H u n g a r y i n t h e Szep l i ge t i co l l ec t i o n , a n d from A u s t r i a i n t h e G i r a u d co l l ec t i o n . T h ese spec i m ens seem t o b e l o s t ( t hey a r e n o t i n B U , V I , M T , P I . T h e o r i g i n a l descr i p t i o n i s u n r e l i a b l e . gracilis (Agon a top u s ) E s a k i a n d H a s h i moto 1 93 2 : 30 ; locus typicus C h i k uz e n - W a k a s u g i y a m a ( K y u s h u , J a p a n ) . T h e t y pe F ( on l y k n o w n s p ec i men ) seems to b e l o s t . T h e o r i g i n a l descr i p t i o n i s u n r e l i a b l e . gracilis ( Th a u m a to dryin u s ) B r u e s 1 9 33 : 1 48 ; locus typ icus B a I t i c amber . T h e t y p e spec i mens of t h e Brues foss i I s p ec i es were i n t h e a m b e r c o l l ec t i o n a t A l b er t u s U n i v e r s i t a t , Kon i g s b e r g ( n ow K a l i n i n g r a d ) . T h ey seem to be l o s t . T h e o r i g i n a l desc r i p t i o n s are u n r e l i a b l e . graeffi ( Gon a topus form icarius v a r . ) K i ef f e r i n K i ef f e r e t M a r sh a l l 1 90 5 : 92 ; locus typicus T r i e s t e ( I t a l y ) . = Gon a top u s graeffi K i effer : K i ef f e r 1 9 1 4b : 1 09 . T h e t y p e ( on l y k n o w n spec i men ) w a s a fem a l e i n t h e G r aeffe co l l ec t i o n . I t seems to be lost ( i t is n o t in V I and M I ) . T h e or i g i n a l descr i p t i o n i s u n re l i a b l e . gramineus (Neogo n a top u s ) Ponomarenko 1 975 a : 3 1 8 ; locus typicus U sc i u g h u n -O b o ( A n d a - X u d u k , U v er-X a n g a i a i m a k , M o n g o l i a ) .

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grandis (Neogona top u s ) F rey t a g 1 977 : 576 ; locus typicus Gordon N u r sery ( Dou g l a s Co . , W i s . , U . S. A . ) . T h e h o l o t y p e M ' i s i n MA . F re y t a g ( 1 977 ) desc r i bed grandis as Neogon a topus on t h e b a s i s of some morpho l og i c c h a r a c t e r i s t i c s � see d i sc u ss i o n on t a x o n o m y of m a l e Gon a topod i n es o n p a ge 1 054 ) . A c c o r d i n g to my s t u d i es , t h ese c h a r acter i s t i c s a re not s a t i sfacto r y . T h e rea l g e n u s i s t h e refor e , n o t i d en t i f i a b l e . Desc r i p t i o n : fem a l e u n k now n ; m a l e f u l l y w i n ged ; l e n g t h 3 m m ; b l ac k ; ma n d i b l es t e s t a ceo u s ; a n te n n a e , l e g s a n d a b domen b row n ; a n t e n n a e n o t d i s t a l l y t h i c k e n e d ; a n t en n a l segmen t s i n f o l l o w i n g p ropo r t i o n s : 1 0 : 7 : 1 7 : 1 6 : 1 8 : 1 6 : 1 5 : 1 4 : 1 3 : 1 6 ; a n t e n n a l seg m e n t 3 more t h a n f o u r t i mes a s l o n g as b road ( 5 , 66 ) ; h e a d d u l l , g r a n u l a ted ; fron t a l l i n e i n com p l e t e ; occ i p i t a l ca r i n a a b sen t ; t e m p l es d i s t i n c t ; POL = 9 , 5 ; Ol = 3 ; OOl = 4 ; occ i p u t con c a v e ; scu t u m d u l l , g r a n u l a ted ; n o t a u l i ces com p l e t e , p o s t e r i or l y s e p a r a ted ; m i n i m u m d i s t a nce b e t w een n o t a u l i ces shorter than b rea d t h of oce l l i ( 2 : 4 ) ; s c u t e l l u m a n d m e t a n o t u m d u l l , g r an u l a t ed ; p ropodeum d u l l , ret i c u l a t e r u g o s e ; forew i n g s h y a l i n e , w i t h o u t d a r k t r a n s v ersa l b a n d s ; ra d i a l ce l l open ; d i s t a l p a r t of ra d i a l v e i n l on g e r t h a n p rox i m a l p a r t ( 26 : 1 1 ) ; b a s i vo l se l l a w i t h

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1 2 59

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M a l e g e n i t a l i a of L a beo gre n a densis A s h m e a d

1 809

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notch ; dorsa l process o f gonoforceps ( F i g . 1 258 ) b ro a d ened ; d i s t a l p a r t w i t h n u me ro u s p a p i l l a e ; m a x i l l a r y p a l p i w i t h 4 segm e n t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 1 , 2 . H o s t s u n k n ow n . F o u n d o n l y i n t h e t y p i c a l loc a l i t y . gren a densis (Labeo) A s h m e a d 1 900 : 235 ; locus typicus M i r a b e a u E s t . ( G re n a d a , A n t i l l es ) . = L a berius gre n a densis ( A s h m e a d ) : K i effer 1 9 1 4b : 66 . T h e ho l o t y p e ( o n l y k n o w n spec i m s n ) i s a m a l e i n B M ! t h e t a xonom i c d i ff i c u l t i e s e n c o u n t e red w i t h r e g a r d to m a l e G o n a t opod i n e s , pre v e n t t h e a s s oc i a t i o n o f t h i s spec i e s w i t h a g en u s . Desc r i p t i o n : fema l e u n k now n ; m a l e f u l l y w i n ge d ; l e n g t h 1 , 87 mm ; h e a d , t h o r a x a n d p ropod e u m b ro w n -b l ack ; a n ten n ae b row n ; a b domen a n d l e g s b ro w n tes t aceou s ; a n t e n n a l seg men t s i n fo l l o w i n g propor t i o n s : 5 : 6 : 1 3 : 1 2 : 1 1 : 1 0 : 1 0 : 9 : 8 : 1 0 ; a n t e n n a l seg m e n t 3 more t h a n t h ree t i mes a s l o n g a s b ro a d ( 4 , 3 3 ) ; h e a d sh i n y , g r a n u l a ted ; temp l e s d i s t i n c t ; occ i p i t a l c a r i n a a b s e n t ; POL = 8 ; OL = 2 ; OOL = 4 ; brea d t h of oce l l i : 4 ; fron t a l l i n e a b s e n t ; v e r t e x w i t h o u t l a te r a l smoo t h a re a s

1 81 0

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1 260 - M a l e of Aph elop u s inexpectatus Ben o i t

( type)

b e t ween eyes a n d pos t e r i o r oce l l i ; scu t u m sh i n y , g r a n u l a ted ; n o t a u l i ces co m p l e t e , poster i or l y sep a r a ted ; m i n i m u m d i s t a n ce b e t ween n o t a u l i ces shorter t h a n a n te n n a I segme n t 2 ( 4 : 6 ) ; sc u t e l l u m a n d m e t a n o t u m sh i n y , smoo t h , w e a k l y p u nc t a te ; p ropod e u m ret i c u l a t e ru gose ; areo l a e sm a l l ; fore w i n g s h y a l i n e , w i t h o u t d a r k t r a n sver s a l b a n d s ; dors a l process o f g o n oforceps ( F i g . 1 259 ) l o n g , w i t h i n ner s i de ro u n ded ; b a s i v o l se l l a w i t h o u t n o t c h ; m a x i l l a r y p a l p i w i t h 6 segmen t s ; l a b i a l p a l p i w i t h 3 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . H o s t s u n k n ow n . ichneumon ides (Methoc a ) L a t r e i l i e 1 804 : 269 ; locus typicus u n k n ow n . ? Gon a topus m u tillarius Nees 1 834 : K i effer 1 9 1 4b : 1 1 8 . = = ? Mutilla formicaria J u r i n e 1 807 : K i effer 1 9 1 4b : 1 1 8 . T h i s spec i e s , w h i c h former l y w a s a s soc i a ted w i t h dry i n i d s , a c t u a l l y b e l o n g s to Tiphiidae s u b f . Methocidae . ignotus ( Gon a topus soc i a b ilis v a r . ) O g l ob l i n 1 924b : 1 2 1 ; locus typ icus Ji l o ve ( C zech o s l o v a k i a ) . The t y pe spec i me n s of t h e O g l ob l i n spec i e s from Czechos l o v a k i a seem to b e l os t ( t h e y are not i n PG, PT a n d BA ) . The or i g i n a l descr i p t i o n s a re u n re l i a b l e . impre s s u s ( A n teon trivi a lis v a r . ) K i effer i n K i effer e t M a r sh a l l 1 90 5 : 20 1 ;

N om i n a d u b i a

Fig.

1 26 1

181 1

- M a l e ge n i t a l i a of Ap helopus inexpecta tus Beno i t ( t y pe ) ( r i g h t h a l f removed )

l oc u s t y p i c u s L o i t sch ( A u s t r i a ) . A n teon trivia lis impressus K i effe r : K i effer 1 9 1 4b : 1 6 1 . T h e t y p i c a l m a t e r i a l ( a t l e a s t t w o m a l e s ) w a s i n t h e Gr aeffe c o l l e c t i o n . I t seems t o b e l os t ( i t is not in V I and M I ) . The or i g i n a l descr i p t i o n i s u n r e l i a b l e . incert u s ( Chelogyn u s a l b idicollis v a r . ) O g l o b l i n 1 924b : 1 26 ; locus typ i c u s K r a l ove H r a d ec ( C zechos l o v a k i a ) . T h e t y p e spec i me n s of t h e O g l ob l i n spec i es from Czechos l o v a k i a seem to b e l os t ( t h e y a re n o t i n PG, PT, BA ) . The or i g i n a l descr i p t i o n s =

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Ent .

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n o . 37,

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are u n re l i a b l e . incert u s ( L a beo) A s h m e a d 1 887a : 74 ; locus typicus F l or i d a ( U . S . A . ) . = L a berius incertus ( As h m e a d ) : K i effer 1 9 1 4b : 6 5 . Accord i n g to M r . Arno l d M e n k e ( p erson a l com m u n i c a t i on ) t h i s spec i e s b e l o n g s t o Ichneumonidae (Euphoriella incerta ) . indi visus (Ap helop u s mel a l e u c u s v a r . ) K i effer i n K i effer e t M a r s h a l l 1 90 5 : 220 ; locus typicus Sco t l a n d ( U n i ted K i n g dom ) . = Aphelop u s indivis u s K i effer : K i effer 1 9 1 4b : 2 1 8 . T h e t y p e ( o n l y k n o w n spec i men ) w a s a m a l e i n t h e C a meron c o l l ec t i o n . I t seems to be l o s t ( i t is not i n BM ) . The o r i g i n a l descr i p t i o n i s u n r e l i a b l e . inexpec t a tus (A ph elop u s ) Ben o i t 1 95 1 a : 302 ; locus typ icus G i a r a b u b o a s i s ( Libya) . T h e t y p e M ! ( o n l y k n o w n s p e c i men ) i s i n G E . T h e e m a r g i n a ted occ i p u t , dors a l p rocess of g o n oforceps and u n scu l p t u red p ropodeum p l ace t h i s spec i es w i t h G o n a topod i n e s ( t h e s a m e g ro u p of Aphelop u s a fricanus Beno i t , w h i ch h a s on l y t h e c o s t a l ce l l c l e a r l y v i s i b l e ) . Desc r i p t i o n : fema l e u n k n o w n ; m a l e ( F i g . 1 260 ) f u l l y w i n g ed ; l e n g t h 2 , 25 m m ; b l ac k ; m a n d i b l es a n d c l y p e u s y e l l o w ; some l i g h t spots around a n t e n n a l soc k e t s ; a n t e n n ae t e s t aceo u s ; l eg s b ro w n , w i t h t i b i ae a n d t a r s i l i g h t ; an t e n n ae n o t d i s t a l l y t h i c k e ned ; a n t en n a l seg men t s i n f o l l ow i n g p ropo r t i o n s : 4 : 3 : 5 : 6 : 5 : 5 : 5 ( a n t e n n a l segmen t s 8- 1 0 m i s s i n g i n h o l o t y p e ) ; a n t e n n a l segmen t 3 l ess t h a n t h ree t i mes a s l o n g a s broad ( 2 , 1 6 ) ; h e a d d u l l , g r a n u l a te d , w i t h n u merous i r re g u l a r k ee l s ; occ i p u t e m a r g i n a t e d , con c a v e ; occ i p i t a l c a r i n a a b sen t ; POL = 7 ; D L = 3 ; DOL = 3 , 5 ; b r e a d t h o f oce l l i : 3 ; t e m p l es d i s t i n c t ; fron t a l l i n e a b sen t ; scu t u m d u l l , g r a n u l a ted ; n o t a u l i ce s i n comp l e t e , rea ch i n g a p p rox i m a t e l y 0 , 75 l e n g t h of scu t u m ; sc u te l l u m a n d m e t a n o t u m sh i n y , smoo t h , w i t h o u t sc u l p t u re ; forew i n g s h y a l i n e , w i t h o u t d a r k t r a n s v e r s a I b a n d s ; p t erost i g m a l o n g a n d n a rrow ; o n l y cos t a l ce l l c l e a r l y v i s i b l e ; gonoforce p s ( F i g . 1 26 1 ) w i t h a l o n g a n d n a rrow dors a l p rocess ; b a s i v o l se l l a w i t h o u t n o tc h ; m a x i I l a r y p a l p i w i t h a t l e a s t 3 segmen t s ( l a s t segmen t s m i s s i n g i n h o l o t y p e ) ; l a b i a l p a l p i w i t h 2 segme n t s ; t i b i a l s p u rs 1 , 1 , 2 . Hos t s u n k n o w n . Fou n d on l y i n t h e t y p i c a l l oc a l i t y . japon ic u s (Neodryin u s ) Uc h i d a 1 927 : 3 2 ; locus typicus O i t a ( K y u sh u , Japan ) • T h e t y p e F ( o n l y k n o w n spec i men ) seems to be l os t . T h e o r i g i n a l descr i p t i o n i s u n re l i a b l e . jucundus (A n teon ) O g l ob l i n 1 924b : 1 2 2 ; locus typicus R a a b ( C zechos l o v a k i a ) . T h e t y pe s p ec i me n s of t h e O g l o b l i n s p ec i es from Czechos l ov a k i a seem to be l os t ( t hey a re not in PG , P T , BA ) . The or i g i n a l desc r i p t i o n s a re u n re l i a b l e . kevirae (Para dryin u s ) L a l 1 939 : 50 ; locus typicus M u s h e r i ( B i h a r , I n d i a ) . T h e t y p e ( o n l y k no w n spec i men ) w a s a m a l e i n t h e P u s a I m p er i a l C o l l eg e c o l l ec t i o n . I t seems t o b e l o s t ( i t i s n o t i n N O ) . T h e or i g i n a l

No m i n a d u b i a

Fig.

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1 26 2 - R a d i a l ve i n of m a l e of Phor 'J a s longicornis Brues ( t y p e )

descr i p t i on i s u n re l i a b l e . leuconeurus ( A n teon trivia l i s v a r . ) K i effer 1 9 1 4 b : 9 1 ; l o cu s typicus T r i e s t e ( I taly ) . = A n teon trivia lis leu coneuru s K i effer : K i effer 1 9 1 4b : 1 6 1 . T h e t y p e ( on l y k n o w n spec i men ) w a s a m a l e i n t h e G r aeffe co l l ec t i o n . I t seems to be l os t ( i t i s n o t i n V I a n d M I ) . The o r i g i n a l desc r i p t i on i s u n re l i a b l e . leviga t u s (A n teon ) K i effer i n K i effer e t M a r s h a l l 1 905 : 1 69 ; loc u s typ icus C h armey ( Sw i tzer l a n d ) . Xen a n teon leviga t u s ( K i effe r ) : K i effer 1 9 1 4b : 1 94 . = The t y pe ( on l y k no w n spec i men ) w a s a fema l e i n t h e De G a u l l e co l l ec t i on . I t seems to be l os t ( i t i s n o t i n P l . T h e o r i g i n a l desc r i p t i on i s u n r e l i a b l e . longicornis (Phorb a s ) B r u e s 1 907b : 1 5 2 ; locus typicus Saegerstown ( P a . , U. S. A. ) . = Phorb a s i a l ongic ornis ( Bru <e s ) : K i effer 1 9 1 4b : 59 . T h e h o l o t y p e M ! ( on l y k n o w n spec i men ) i s i n WA . T h e emarg i n a ted occ i p u t and t he n u m b e r of p a l p a l segmen t s p l ace t h i s spec i es w i t h Gon a topo d i n e s . Descr i p t i on : fema l e u n k n ow n ; m a l e f u l l y w i n ged ; l e n g t h 2 , 2 5 m m ; b l a c k , w i t h h e a d a n d a n t e n n a e b row n ; l eg s b row n , w i t h t a rs i , t i b i ae , s t a l k s o f femora a n d t roc h a n ters l i g h t ; a n te n n a e n o t d i s t a l l y t h i ck e n e d ; a n t e n n a l seg men t s i n fo l l o w i n g p roport i o n s : 8 : 7 : 1 8 : 1 7 : 1 6 : 1 6 : 1 4 : 1 2 : 1 0 : 8 ; a n t en n a l segmen t 3 more t h a n t h ree t i me s as l o n g as b ro a d ( 4 , 33 ) ; head sh i n y , g r a n u l a t e d ; fron t a l l i n e a b s e n t ; occ i p i t a l c a r i n a a b sen t ; POL = 1 1 ; OL = 4 ; OOL = 5 ; occ i p u t con c a v e , e m a r g i n a t e d ; b re a d t h o f oce l l i : 3 ; sc u t u m s h i n y , g r a n u l a t e d ; n o t a u l i ces co m p l e t e , p o s t e r i o r l y s e p a r a t e d ; m i n i m u m d i s t a n ce b e t w een n o t a u l i ces a p p rox i ma t e l y a s l o n g as b r ea d t h of oce l l i ; scu te l l u m a n d m e t a n o t u m sh i n y , smoo t h , w e a k l y p u n c t a t e ; p ropodeum d u l l , f u l l y ret i c u l a t e r u gose ; dors a l s u rface w i t h a med i a n f u rrow ; a reo l ae on t h e s i des of t h i s f u r row sm a l l e r t h a n a reo l ae of poster i or s u rface ; fore w i n g s h y a l i n e , w i t h o u t d a rk t r a n sver s a l b a n d s ; d i s t a l p a r t of ra d i a l v e i n ( F i g . 1 26 2 ) m u ch l on g er t h a n p ro x i m a l p a r t ; a b domen m i s s i n g i n h o l o t y p e ; m a x i l l a r y p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 1 , 2 . Hos t s u n k n ow n . F o u n d on l y i n t h e t y p i c a l l oc a l i t y . T h i s spec i es w a s n o t i nc l u ded i n g e n e r a l k e y s o f m a l e Gon a t opod i n es b e c a u s e the a b domen ( i n c l u d i n g gen i t a l a rm a t ure ) is m i ss i n g i n h o l o t y p e . T h e t a xonom i c d i ff i c u l t i es of m a l e G o n a topod i n es p reven t t h e

1 81 4

Mem . Amer . E n t .

Fig.

I nst . ,

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1 26 3 - M a l e of L a beo longitarsis A s h m e a d

( type)

a s soc i a t i o n o f t h i s spec i es w i t h a g e n u s . lo ng itar s i s (A n teon) K i effer i n K i effer e t M a r s h a l l 1 906 : 5 1 6 ; N e w Forest ( E n g l a n d ) . � Chelogyn u s longi tarsis ( K i e ffer ) : K i effer 1 9 1 4b : 1 8 2 .

locus typicus

T h e t y p e ( on l y k n o w n spec i men ) w a s a fem a l e i n t h e M a r sh a l l co l l ec t i o n . I t seems to be l os t ( i t i s n o t i n BU ) . T h e or i g i n a l desc r i p t i on i s u n re l i a b l e .

1815

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1 264 - M a l e g e n i t a l i a of L abeo longitarsis Ashmead

longitarsis ( L a b eo ) A s h m e a d 1 893 :

( type)

8 8 ; l o c u s typ i c u s J ac k son v i l l e ( F l a . ,

U . S. A . ) . = L a b er i u s longitarsis ( A s h m e a d ) : K i effer 1 9 1 4 b : 65 . T h e h o l o t y p e M ! ( on l y k n o w n spec i me n ) i s i n ViA . The e m a r g i n a t ed occ i p u t , dors a l p rocess of gonoforceps a n d n u m ber of p a l p a l segmen t s p l ace t h i s s p ec i e s w i t h Gona topod i n es . Descr i p t i on : f e m a l e u n k n ow n ; m a l e ( F i g . 1 263 ) f u l l y w i n ged ; l en g t h 3 m m ; b l a c k ; m a n d i b l es a n d l e g s y e l l ow - b row n ; a n t e n n a e b row n - b l a c k ; a n t en n a l segme n t s i n fo l l ow i n g p roport i o n s : 6 : 4 : 9 : 8 : 8 : 8 : 8 : 7 : 6 : 1 0 ; a n t en n a l seg men t 3 more t h a n t h ree t i mes a s l on g a s b ro a d ( 4 ) ; a n t e n n a l segmen t 1 0 a p p rox i m a t e l y s i x t i mes a s l o n g a s b road ; h e a d sh i n y , h a i r l e s s , w i t hou t sc u l p t u re ; f ron t a l l i ne a b sen t ; occ i p i t a l c a r i n a a b sen t ; t em p l es d i s t i n c t ; P O L = 6 ; O L = 3 ; OOL = 4 ; scu t u m sh i n y , w i t h o u t scu l p t u re or w e a k l y g r a n u l a t ed ; not a u l i ce s com p l e t e , p o s t er i o r l y sep a r a ted ; m i n i m u m d i s t a n c e b e t w een n o t a u l i ces a s l o n g a s b re a d t h of oce l l i ( 4 ) ; sc u t e l l u m a n d met a n o t u m smoo t h , sh i n y , w ea k l y p u n c t a t e , w i t hou t scu l p t u re among p u n c t u res ; p ropodeum

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g r a n u l a t ed and ret i c u l a te r u gose ; dor s a l s u r face of p ropodeum w i t h t r a c k of med i a n f u r ro w ; forew i n g s h y a l i n e , w i t h o u t d a rk t r a n sver s a l b a n d s ; dor s a l p rocess o f go noforce p s ( F i g . 1 264 ) s ho r t , d i s t a l l y e m a r g i n a t ed ; b a s i v o l se l l a w i t h o u t n o tc h ; m a x i l l a ry p a l p i w i t h 5 s e g me n t s ; l a b i a l p a l p i w i t h 2 s e g men t s ; t i b i a l s p u rs 1 , 1 , 2 . Hos t s u n k n ow n . K n o w n o n l y i n t h e t y p i c a l l oc a l i t y . T h e t a x o n o m i c d i f f i c u l t i e s of m a l e Gon a t opod i n es p re v e n t t h e a ssoc i a t i on o f t h i s spec i es w i t h a g e n u s . l u teicornis ( A n teon carin a t u s v a r . ) K i effer i n K i effer e t M a r sh a l l 1 905 : 1 5 2 ; locus typicus N e u - V a r a d ( H u n g a r y ) . = Chelogyn u s lu teicornis ( K i effer ) : K i effer 1 9 1 4b : 1 78 . T h e t y p e ( on l y k n o w n spec i men ) w a s a fem a l e i n t h e Szep l i g e t i co l l ec t i o n . I t seem s to b e l o s t ( i t i s n o t i n BU ) . T h e o r i g i n a l descr i p t i on i s u n r e l i a b l e . A n teon l u teicornis K i effer w a s i n c l u d e d i n t h e k e y s o f R i ch a rds ( 1 939 ) a n d J . F . Perk i n s ( 1 976 ) t o B r i t i s h spec i e s . R i ch a r d s a n d J . F . Perk i n s n ever s a w t h e t y p e spec i me n a n d b a sed t h e i r k e y s o n t h e o r i g i n a l desc r i p t i on o f K i effer . I t h i n k t h a t t h e n a m e l u teicornis i s b e s t i g n ored , b e c a u s e t h e t y p e spec i men i s l os t a n d t h e descr i p t i o n i s u n re l i a b l e . O n t h e o t her h a n d , i f t h e op i n i o n o f R i ch a r d s a n d J . F . Per k i n s i s correc t , l u teicornis i s a sen i o r s y n o n y m of A n teon p u n c t a tus K i effer . I s t u d i ed t h e t y p e spec i men of puncta t u s i n t h e N a t i on a l M u se u m of P a r i s ( F ra nce ) a n d d i d n o t see a n y d i fference b e t w een i t a n d A n teon infectum ( H a l i d a y ) . T h e ch a r a c te r i s t i c s p roposed by J . F . Perk i n s ( 1 97 6 ) for s e p a r a t i n g t h e t w o spec i e s ( h e a d d u l l o r sh i n y , scu l p t u re o f t h e f ro n s , co l o u r , n u m ber o f l a m e l l a e ) a re v e r y v a r i a b l e i n A n teon . I n conc l u s i o n , e v e n i f t h e op i n i on of R i c h a r d s a n d J . F . Perk i n s w a s correc t , l u teicornis c a n at most b e con s i dered a j u n i or s y n on y m of infec tum . l U l<'eli (Lestodryin u s ) Ben o i t 1 95 1 c : 6 ; locus typ i c u s Kom i ( S a n k u r u , Z a i re ) . T h e t y p e ( on l y k n o w n spec i men ) i s a m a l e i n T V . T h e t a xonom i c d i f f i c u l t i e s o f m a l e D r y i n i n es p re ve n t t h e assoc i a t i o n o f t h i s spec i es w i t h a g e n u s . Desc r i p t i on : fema l e u n k n ow n ; m a l e fu l l y w i n ged ; l en g t h 4 , 2 m m ; b l ack ; m a n d i b l es t e s t aceo u s ; a n t e n n a e w i t h seg men t s 1 - 2 t e s t aceou s , 3-6 d a rk e n e d , 7 - 1 0 m i s s i n g i n h o l o t y p e ; a b domen b row n ; l eg s y e l l o w ; a n t en n ae n o t d i s t a l l y t h i c k e n e d ; a n t e n n a I s e g me n t s i n fo l l ow i n g p ropo r t i o n s : 1 1 : 8 : 26 : 1 5 : 1 6 : 1 5 ( o t h er seg men t s m i s s i n g i n h o l o t y p e ) ; h e a d ret i c u l a t e ru gose, d u l l , g r a n u l a t ed , w i t h i rr e g u l a r k ee l s o n t h e v e r t e x ; t em p l e s i n v i s i b l e ; occ i p i t a l c a r i n a comp l e t e , v i ewed dorsa l l y reac h i n g t h e e y e s ; fron t a l l i n e a b sen t ; POL = 5 ; O L = 2 ; OOL = 3 ; O P L = 1 ; sc u t u m , sc u te l l u m a n d m e t a n o t u m d u l l , g r a n u l a t ed , ret i c u l a t e ru gose ; n o t a u l i ce s i n comp l e t e , reach i n g a p p rox i m a te l y 0 , 5 l en g t h of scu t u m ; p ropodeum r e t i c u l a t e r u g o s e , w i t h o u t k ee l s ; forew i n g s h y a l i n e , w i t h o u t dark t r a n sv e rs a l b a n d s ; d i s t a l part of r a d i a l v e i n l on g e r t h a n p rox i m a l p a r t ( 2 5 : 1 6 ) ; t i b i a l s p u rs 1 , 1 , 2 . H o s t s u n k n ow n . Fou n d o n l y i n t h e t y p i c a l l oc a l i t y . major (A n teon) K i effer 1 907b : 2 5 ; locus typ i c u s S m a l a n d ( Sweden ) .

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Dryin u s collaris D a I m a n v a r . beta D a I m a n 1 82 3 : 9 . A n teon major ( D a l m a n ) : K i effer 1 907 b : 2 5 . Dryin u s collaris v a r . major D a l m a n : K i effer 1 9 1 4b : 1 5 1 . T h e n a me major w a s a ssoc i a ted b y K i effer ( 1 907b , 1 9 1 4 b ) w i t h a s u p p osed v a r i e t y of Dryin u s collaris p ro p osed b y Da I m a n ( 1 823 ) . I n re a l i t y , h o w e v e r , D a l m a n w a s descr i b i n g o n l y a v a r i e t y , beta . Major i s o n l y t h e f i rs t word of t h e desc r i p t i o n . T h e A u t h o r , t h erefore , i s K i effer , a n d n o t D a l m a n . T h e t y p e ( on l y k n o w n s p ec i men ) i s p rob a b l y i n S T . I h a v e n o t seen t h i s m a t e r i a l . mayeti (A n teon ) K i effer i n K i effer et M a r sh a l l 1 90 5 : 1 70 ; locus typicus Austr i a . = A n teon mayeti K i effer : K i effer 1 9 1 4b : 1 52 . T h e' t y p e ( o n I y k n o w n spec i men ) w a s a fema I e fou n d b y G i r a u d a n d k e p t i n t h e M a y e t c o l l ec t i on . I t seems t o b e l os t ( i t i s n o t i n MT ) . T h e o r i g i n a l descr i p t i o n i s u n r e l i a b l e . medialis (An teon ) H e l l e n 1 9 1 9 : 284 ; locus typicus u n k no w n . T h e t y p e F shou l d be i n H E . I don 1 t k n o w t h i s form ( d escr i b ed a s a b e r ra t i o n of A n teon longicornis D a l m a n ) . The or i g i n a l descr i p t i o n i s u n re l i a b l e . minor ( A n teon vulgaris v a r . ) K i effer i n K i effer e t M a r sh a l l 1 90 5 : 203 ; locus typ ic u s Vo l osca ( I t a l y ) . = A n teon vulgaris minor K i effer : K i effer 1 9 1 4b : 1 62 . T h e t y p e ( on l y k n o w n spec i men ) w a s a m a l e i n t h e G r a effe co l l ec t i on . I t seems to be l os t ( i t i s n o t i n V I a n d M I ) . T h e or i g i n a l descr i p t i o n i s u n re l i a b l e . m in u tellus (A n teon ) K i e ffer i n K i effer e t M a r sh a l l 1 906 : 523 ; locus typ i c u s u n cert a i n ( Ke n m u i r or D u m f r i e s , Sco t l a n d , U n i ted K i n g dom ) . New name. n e c A n teon minutus A s h m e a d 1 893 : K i effer i n K i effer e t M a r s h a l l 1 906 : 5 23 . n = A n teon m in u tu s K i effer i n K i effer e t M a r s h a l l 1 905 : 204 ( sy n . p roposed b y K i effer i n K i effer e t M a rs h a l l 1 906 ) . T h e t y p e ( on l y k n o w n s p ec i men ) w a s a m a l e i n t h e C a meron co l l ec t i on . I t seems t o b e l o s t ( i t is not in BM ) . T h e or i g i n a l descr i p t i on i s u n re l i a b l e . m in u tu s (A n teon ) K i effer i n K i effer e t M a rs h a l l 1 905 : 204 . See min u tellus (An teon ) K i effer . miripes (A llodryin u s ) K i effer 1 92 2 : 66 ; locus typ ic u s B u t u a n ( P h i I i p p i n e s ) . = A llodryin u s miripes K i effer : B a l t a z a r 1 966 : 205 . T h e t y p e ( on l y k n o w n spec i men ) w a s a fema l e i n t h e B a k e r co l l ec t i on . I t seems t o b e l o s t ( i t i s n o t i n W A , L A , C A , A M , B M , C M , D A , P H ) . T h e or i g i n a l desc r i p t i o n i s u n re l i a b l e . F o r t h i s spec i e s K i effer ( 1 92 2 ) p roposed t h e new gen u s A llodryin u s . mortuorum (Les todryin u s ) Brues 1 93 3 : 1 49 ; locus typ i c u s B a I t i c a m b e r . T h e t y pe s p ec i me n s o f t h e B r u e s fos s i I s p ec i e s were i n t h e a m b e r co l l ec t i o n a t A l b e r t u s U n i ve r s i t a t , Kon i g s b e r g ( n ow K a l i n i n g r a d ) . T h e y seem t o b e l o s t . t h e or i g i n a l descr i p t i o n s a re u n re l i a b l e .

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mul ticolor (Gon a top u s ) K i e ffer i n K i effer et M a r s h a l l 1 90 5 : 1 02 ; loc u s typ icus P a l e r mo ( I t a l y ) . � Gon a top u s m u l ticolor K i effe r : K i effer 1 9 1 4 b : 1 1 1 . T h e t y p e ( o n l y k n o w n spec i men ) w a s a fe m a l e i n t h e De Stefa n i co l l ect i o n . I t seems t o b e l o s t ( i t i s n o t i n t h e en tomo l o g i c a l co l l e c t i o n s i n P a l e r mo ) . T h e D e S t ef a n i co l l ec t i o n seems t o b e l o s t . T h e o r i g i n a l desc r i p t i o n i s u n re l i a b l e . m u ltidens (Dicondyl u s ) O g l ob l i n 1 924b : 1 1 8 ; locus typicus Bo l e s l a v ( C zechos l o v a k i a ) . The t y p e spec i me n s of t h e O g l ob l i n spec i es from Czech o s l o v a k i a seem to b e l os t ( t hey a re not i n PG, P T , BA ) . The or i g i n a l desc r i p t i o n s are u n rel i ab l e . m u tillaria (Methoca ) L a t re i l l e 1 806 : t a b . 1 3 , f i g . 7 ; locus typicus F r a n ce . Gon a topus thoracic us Nees 1 834 : 387 ( s y n . proposed by Nees 1 834 ) . � Gon a top u s thora cicus Nees : K i effer i n K i effer e t M a r s h a l l 1 905 : 1 25 . � Gon a topus thoracicus Nees : K i effer 1 9 1 4b : 1 1 9 . The t y p i c a l m a te r i a l of m u tillaria L a t r e i l i e seems to b e l o s t . The or i g i n a l desc r i p t i o n is unrel i a b l e . Nees ( 1 834 ) bel i e ved t h a t Me thoca m u tillaria L a t re i l i e 1 806 was a spec i e s d i ffere n t from Methoca m u tillaria L a t re i l l e 1 809, a n d p roposed a new n a m e for i t ( G . thoracic u s ) . Accor d i n g t o K i effer i n K i effer e t M a r s h a l l ( 1 905 ) t h i s spec i e s b e l o n g s t o Mu tillida e . T h e t y p e spec i m e n s of t h e spec i es desc r i bed by Nees seem to be l os t . T h e or i g i n a l descr i p t i o n s a re u n r e l i ab l e . m u tillarius (Gon a top u s ) Nees 1 834 : 384 ; locus typ ic u s S w i t z e r" l a n d . T h e t y p e spec i m e n s of t h e spec i es descr i be d by Nees seem to be l o s t . T h e or i g i n a l desc r i p t i on s a re u n r e l i a b l e . n a s u t u s (Embolem u s ) A s h m e a d 1 887a : 7 5 ; locus typicus J a c k s o n v i l l e ( F l a . , U. S. A. ) . � L a b eo n a s u t u s ( A� h m e a d ) : Ash m e a d 1 893 : 89 . � L a b erius n a s u t u s ( As h m ea d ) : K i effer 1 9 1 4b : 64 . T h e h o l o t y p e M ! ( on l y k no w n spec i m en ) i s i n WA . T h e e m a r g i n a t e occ i p u t , dors a l p rocess o f gonoforceps a n d n u m b e r o f p a l p a l seg men t s p l ace t h i s spec i es w i t h Gon a topod i n es . Desc r i p t i on : fem a l e u n k n ow n ; m a l e fu l l y w i n ged ; l e n g t h 2 , 52 m m ; b l ac k ; a n t e n n a e , m a n d i b l es , a b domen a n d l eg s b row n ; a n t e n n a I s e g me n t s i n fo l l ow i n g p ropo r t i on s : 7 : 6 : 9 : 9 : 8 : 8 : 8 : 8 : 8 : 1 1 ; a n t e n n a I seg men t 3 l ess t h a n t h ree t i mes a s l o n g a s b road ( 2 , 8 ) ; head d u l l , g r a n u l a t e d ; fron t a l l i n e a b sen t ; occ i p i t a l c a r i n a a b sen t ; t e m p l e s d i s t i n c t ; occ i p u t conc a v e ; P O L � 8 ; OL � 3 ; OOL � 5 ; scu t u m sh i n y , g r a n u l a t ed ; no t a u l i ce s i n com p l e t e , reach i n g a p p rox i m a te l y 0 , 8 l e n g t h of scu t u m ; scu t e l l u m a n d m e t a n o t u m sh i n y , smoo t h , w e a k l y g r a n u l a ted ; p ropode u m sh i n y , e n t i re l y s l i g h t l y ru gose ; dor s a l s u rf a ce w i t h a t rack of med i a n fu rrow ; forew i n g s h y a l i n e , w i t h o u t d a r k t r a n s v e r s a l b a n d s ; d i s t a l p a r t o f r a d i a l v e i n ( F i g . 1 265 B ) m u c h l on ge r t h a n prox i m a l p a r t ; dor s a l p rocess of gon oforceps ( F i g . 1 26 5 A ) l o n g , s l e n d e r , p o i n t ed ; b a s i v o l se l l a w i t hou t notch ; m a x i l l a r y p a l p i w i t h 5 seg men t s ; l a b i a l

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-

M a l e of Embolemus n a s u tu s A s h m e a d r a d i a l v e i n ; N : notch .

( t y pe ) ; A :

ge n i t a l i a ;

B:

p a l p i w i t h 3 segmen t s ; t i b i a l s p u r s 1 , 1 , 2 . T h e t a xonom i c d i ff i c u l t i e s o f m a l e Gon a t opod i n es prev e n t t h e a ssoc i a t i on of t h i s spec i es w i t h a gen u s . Hos t s u n k n o w n . K n o w n on l y i n t h e t y p i c a l l oc a l i t y . neesi ( Gona topus m a rsh a l l i v a r . ) K i effer i n K i effer e t M a r sh a l l 1 905 : 1 2 1 ; locus typicus Ber l i n ( Ge r m a n y ) . = Gon a top u s l u n a t u s Nees ( n ec K I u g ) 1 834 : 383 . = Gon a top u s neesi K i effer : K i effer 1 9 1 4 b : 1 1 8 . K i effer ( i n K i effer e t M a r s h a l l 1 905 ) b e l i e ved t h a t t h e d ry i n i d i de n t i f i ed as Gon a top u s l u n a tus by Nees ( 1 834 ) w a s n o t the re a l G . l u n a t u s K I u g . He th erefore p roposed a n e w n a me f o r t h e N e e s spec i e s ,

Gon a topus marsh a l l i v a r . nees i . T h e t y p e ( o n l y k n o w n spec i me n ) w a s a fem a l e i n the Nees co l l ec t i on . T h i s co l l e c t i o n , however, seems to b e l o st . T h e o r i g i n a l descr i p t i o n i s u n re l i a b l e . neglectus (A n teon) K i effer i n K i effer e t M a rs h a l l 1 906 : 523 ; locus typ i c u s M a i so n s - L aff i t te ( F r a n ce ) . New n a me . nec Dryin u s dors alis Nees 1 83 4 : 372 . = A n teon dorsa l i s K i effer i n K i effer et M a rs h a l l 1 905 : 202 ( s y n . p roposed b y K i effer i n K i effer e t M a r s h a l l 1 906 ) . = A n teon neglec t u s neglectus K i effe r : K i effer 1 9 1 4b : 1 6 1 . T he t y p e ( o n I y k n o w n spec i men ) w a s a m a I e i n t he De G a u l i e c o l l ec t i o n . I t seems to be l os t ( i t i s n o t i n P l . T h e o r i g i n a l desc r i p t i on i s u n re l i a b l e . n igra (Heterolepis) Nees 1 834 : 272 ; locus typicus S i c k e rs h a u se n ( Ge r m a n y ) . A n teon n iger ( Nees ) : D a I I a Torre 1 898 : 539 . Heterolepis n igra Nees : K i effer 1 9 1 4 b : 2 2 1 .

Mem . Amer . E n t .

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1 266 - M a l e g e n i t a l i a of L a beri n u s p aranensis O g l ob l i n

( type)

T h e t y p e s p ec i me n s o f t h e spec i e s desc r i bed b y Nees seem t o b e l o s t . T h e or i g i n a l desc r i p t i o n s a re u n re l i a b l e . n igriventris (Gon a top u s ) Nees 1 834 : 3S6 ; loc u s typicus S i ckers h a u sen ( G erm a n y ) • ; Gon a top u s bicolor H a l i d a y : K i effer i n K i effer et M a r sh a l l 1 905 : l OS . ; Gon a top u s n igriventris Nees : K i effer 1 9 1 4b : 1 1 S . T h e t y p e spec i me n s o f t h e spec i es descr i b ed b y Nees seem t o b e l o s t . T h e o r i g i n a l descr i p t i on s a re u n re l i a b l e . n igrofemora tu s (A n teon) H e l l e n 1 9 1 9 : 284 ; locus typ i c u s u n kno w n . T h e t y p e M shou l d be i n H E .

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don ' t k n ow t h i s form ( descr i b ed a s

Nom i n a d u b i a

1 82 1

a berra t i on of A n teon longicornis D a l m a n ) . T h e or i g i n a l desc r i p t i o n i s u n re l i a b l e . n i tidell u s (A n teon) K i effer i n K i effer e t M a rs h a l l 1 906 : 523 ; l o c u s typ i c u s u n c e r t a i n ( F r a n ce or Sco t l a n d ) . New n a me . n ec Neochelogyn u s n i tidus R . C . L . Perk i n s 1 905 : 6 2 . A n teon n itidus K i effer i n K i effer e t M a r s h a l l 1 90 5 : 1 95 . � A n teon nitidus K i effe r : C a meron 1 907 : 3 3 , 34 .

A n teon n itidus K i effer : K i effer 1 9 1 4 b : 1 57 . T h e t y p i c a l m a te r i a l ( m a l es ) w a s composed of a m a l e f rom Sevres ( F r a n ce ) in the De G a u l l e co l l ec t i o n , and a m a l e from Scot l a n d i n

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t h e C a meron co l l ect i o n . T h e t y p e spec i me n s seem to b e l o s t ( t hey a re n o t i n BM a n d P l . T h e or i g i n a l desc r i p t i on i s u n re l i a b l e . n itidus (A n teon ) K i effer i n K i effer e t M a rs h a l l 1 905 : 1 95 ; l o c u s typi c u s u n cer t a i n ( F ra nce or Sco t l a n d ) . See nitidellus (A n teon ) K i effe r . paranensis (La berin u s ) O g l o b l i n 1 93 2 : 266 ; locus typicus G u a y r a ( P a r a n a , Br a z i I ) . T h e h o l o t y pe M ! ( o n l y k n o w n spec i me n ) i s i n P T . The e m a r g i n a t e occ i p u t , dors a l p rocess of g onoforceps a n d n u mb e r o f p a l p a l se g m e n t s p l ace t h i s spec i e s w i t h G o n a topod i n es . Desc r i p t i on : fema l e u n k n ow n ; m a l e f u l l y w i n g e d ; l en g t h 2 , 45 m m ( a ccord i n g t o t h e or i g i n a l desc r i p t i on , a b domen a n d most of t h e t h o r a x are m i s s i n g i n h o l o t y p e ) ; accord i n g t o t h e or i g i n a l desc r i p t i o n , co l o u r b l ack w i t h a n ten n a I segme n t s 1 - 2 , s t a l k s of femor a , e x t rem i t i es of t i b i ae , t a r s i , t e g u l ae a n d a b domen b ro w n ( t h e co l o u r i s n o t v i s i b l e i n t h e ho l o t y p e , b ec a u se i t i s comp l e te l y m o u n ted o n a s l i d e i n C a n a d a b a l sa m ) ; scu l p t u re o f h e a d n o t v i s i b l e i n t h e h o l o t y p e ; m a n d i b l es w i t h 3 tee t h ; temp l es i n v i s i b l e ; occ i p i t a l m a r g i n a b sen t ; scu t u m , sc u t e l l u m a n d m e t a n o t u m n o t re t i c u l a te r u gose ; n o t a u l i ce s comp l e te , poster i or l y sep a r a t e d ; p ropodeum ret i c u l a t e r u gose ; dors a l s u rface w i t h a med i a n f u rrow ; forew i n g s h y a l i n e , w i t h o u t dark t r a n sv e rs a l b a n d s ; d i s t a l p a r t of ra d i a l v e i n m u c h l o n g e r t h a n prox i m a l p a r t ; dors a l p rocess of g o n oforceps ( F i g . 1 266 ) l o n g , d i s t a l l y b ro a d e n e d ; m a x i I l a r y p a l p i w i t h a t l e a s t 3 segmen t s ( o t h er segmen t s m i s s i n g i n ho l o t y p e ) ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u r s 1 , 1 , 2 . H o s t s u n k now n . K n o w n on l y i n t h e t y p i c a l l oc a l i t y . T h e t a xonom i c d i ff i c u l t i es o f m a l e G o n a topod i nes p rev e n t t h e a s soc i a t i on o f t h i s spec i es w i t h a g e n u s . p a storalis (Go n a top u s ) O g l ob l i n 1 924b : 1 2 1 ; locus typ icus Nera tov i ce ( C zechos l ov a k i a ) •

T h e t y p e spec i m e n s of t h e O g l ob l i n spec i es from Czechos l o v a k i a seem t o be l os t ( t h e y a re not in PG, P T , BA ) . The or i g i n a l desc r i p t i o n i s u n r e l i ab l e . perkinsi (Ech throdelp h a x ) Dodd 1 9 1 4c : 2 ; locus typ ic u s Ne I son ( Qu ee n s I a n d , Austra l i a ) . T h e t y pe ( o n l y k n o w n spec i men ) w a s a fem a l e i n t h e D o d d co l l ect i on . I t seems to be l os t ( i t i s n o t i n AD a n d BM ) . T h e or i g i n a l

1 82 2

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no.

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desc r i p t i on i s u n r e l i a b l e . perk insi (" es todryin u s ) K i effer 1 9 1 4 a : 90 ; locus typ i c u s Los B a n o s ( L u zo n , P h i l i p p i n es ) . = "es todryin u s perkinsi K i effer : K i effer 1 9 1 4b : 2 3 . = [esto dryi n u s perkinsi K i e ffer : B a l t a z a r 1 966 : 203 . T h e t y p e ( on l y k no w n spec i men ) w a s a fema l e . I t seems to be l o s t . The or i g i n a l desc r i p t i o n i s u n re l i a b l e . p iceipes ( Aphelop u s melaleucus v a r . ) K i effer i n K i effer e t M a r s h a l l 1 90 5 : 2 1 9 ; locus typ ic u s L i l l e ( F r a n ce ) . = Aphelop u s piceipes K i effe r : K i effer 1 � 1 4b : 2 1 7 . = A p helop u s p iceipes K i effe r : C a v ro 1 950 : 6 . T h e t y p e ( on l y k n o w n spec i men ) w a s a m a l e i n t h e D e G a u l l e co l l ec t i o n . I t seems to be l o s t ( i t i s n o t i n P ) . T h e or i g i n a l descr i p t i o n i s u n re l i a b l e . p u n c t u l a tus (A n teon ) K i effer i n K i effer e t M a r s h a l l 1 90 5 : 1 94 ; loc u s typ ic u s u n cer t a i n ( F r a n c e or Y u g os l a v i a ) . = A n teon p u n c t u l a tus K i effe r : K i effer 1 9 1 4 b : 1 57 . T h e t y p i c a l m a t e r i a l ( a t l e ast t h ree m a l es ) w a s composed o f t w o m a l es i n t h e De G a u l l e co l l ec t i o n ( f rom V i n c e n n e s a n d Foret de S t Germa i n , F r a n c e ) a n d a m a l e i n t h e G r a effe c o l l ec t i o n ( f rom Pu l a , Y u g o s l a v i a ) . The t y pes seem to be l os t ( t h e y a re n o t i n V I , M I a n d P ) . T h e or i g i n a l descr i p t i o n i s u n re l i a b l e . radialis (A n teon ) K i effer i n K i effer e t M a r sh a l l 1 905 : 205 ; locus typicus Z u g l i ge t ( H u n g a ry ) . = A n teon ra dialis K i effe r : K i effer 1 9 1 4b : 1 64 . T h e t y pe ( on l y k n o w n spec i men ) w a s a m a l e i n t h e Szep l i ge t i c o l l ec t i o n . I t seems t o b e l os t ( i t i s not i n BU ) . T h e o r i g i n a l descr i p t i on i s u n re l i a b l e . ruficollis (A n teon ) K i effer i n K i effer e t M a rs h a l l 1 905 : 1 60 ; locus typicus Boroszno ( H u n g a r y ) . = Chelogynus ruficollis ( K i effe r ) : K i effer 1 9 1 4 b : 1 82 . T h e t y p e ( o n l y k no w n spec i men ) w a s a fema l e i n t h e Szep l i ge t i co l l ect i on . I t seems to b e l o s t ( i t i s n o t i n BU ) . t h e or i g i n a l desc r i p t i o n i s u n re l i a b l e . rufoniger (Chelogyn u s ) O g l o b l i n 1 9 24b : 1 26 ; locus typ icus K r a l u p y ( C zechos l o v a k i a ) . T h e t y p e spec i me n s of t h e O g l ob l i n spec i es descr i b ed from Czechos l o v a k i a seem to b e l o s t ( t h e y a re n o t i n P G , P T , BA ) . The o r i g i n a l desc r i p t i o n s a re u n r e l i a b l e . s a n ctivin cen ti ( [ a beo) Ashmead 1 894 : 1 97 ; locus typ icus S t . V i n c e n t (Anti l ies) . = L a b erius s a n c ti-vin centi ( As h m e a d ) : K i effer 1 9 1 4 b : 66 . T h e ho l o t y pe M ' i s i n WA ; a n o t her ma l e i s i n BM . T h e emarg i n a t e occ i p u t , dors a l process o f g o n oforeeps a n d n u m ber of p a l p a l segmen t s p l a ce t h i s spec i e s w i t h t h e Gona top o d i nes . Desc r i p t i on : fema l e u n k n o w n ; m a l e f u l l y w i nged ; l e n g t h 1 , 68 ; h e a d b l a c k , w i t h m a n d i b l e s y e l l ow ; a n t e n n a e b row n ; t h o r a x a n d propode u m b l a c k ;

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1 267 - M a l e g e n i t a l i a of i, a beo s a n c t iv i n c e n t i A s h m e a d ( t y p e )

a b domen redd i s h - b row n ; l e g s b row n - y e l l o w ; a n t en n a l seg men t s i n fo l l o w i n g p ropo r t i o n s : 5 : 4 : 8 : 8 : 7 : 7 : 6 : 6 : 6 : 8 ; a n te n n a l seg men t 3 l ess t h a n t h ree t i mes as l on g as b ro a d ( 2 , 66 ) ; h e a d sh i n y , g r a n u l a t ed ; frons w i t h a med i a n f u r row i n fro n t of a n te r i o r oce l l u s ; fron t a l l i n e a b s e n t ; POL = 7 ; O l = 3 ; OOl = 4 , 5 ; b re a d t h o f oce l l i : 2 ; v e r t e x w i t h a s m a l l smoo t h a rea b e t w een eyes a n d pos t e r i o r oce l l i ; t e m p l es d i s t i n c t ; occ i p i t a l c a r i n a a b s e n t ; occ i p u t con c a v e ; sc u t u m s h i n y , g r a n u l a ted ; n o t a u l i ce s com p l e t e , poste r i o r l y sep a r a ted ; m i n i m u m d i s t a n c e b e t ween n o t a u l i ces shorter t h a n a n t e n n a I segmen t 2 ( 2 : 4 ) ; scu te l l u m a n d m e t a n o t u m sh i n y , smoo t h , w e a k l y p u n c t a t e ;

1 8 24

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En \ .

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no. 37,

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p ro p o d e u m ret i c u l a te r u g ose ; dors a l s u rf a c e w i t h a med i a n f u rrow ; p o s t e r i o r s u rf a ce w i t h o u t k ee l s ; forew i n g s h y a l i n e , w i t hou t d a rk t r a n s v e r s a l b a n d s ; dor s a l p rocess o f gonoforce p s ( F i g . 1 267 ) l o n g , d i s t a l l y b e n t , w i t h p a p i I l ae o n t h e e x t e r n a l s i d e ; b a s i v o l se l l a w i t h n o t c h ; m a x i l l a ry p a l p i w i t h 4 s e g me n t s ; l a b i a l p a l p i w i t h 2 s e g m e n t s ; t i b i a l s p u rs 1 , 1 , 2 . H o s t s u n k n o w n . K n o w n on l y i n t h e t y p i c a l l oc a l i t y . T h e t a xon o m i c d i ff i cu l t i e s of m a l e G o n a topod i n es prevent t h e a s soc i a t i o n of t h i s spec i es w i t h a g e n u s .

sciop h a n e s (Gon a top u s ) M a r s h a l l i n K i ef f e r e t M a r sh a l l 1 905 : 1 07 ; loc u s typicus C i u d a d R e a l ( Sp a i n ) . = Gona top u s sciop hanes M a r s h a l l : K i e f fe r 1 9 1 4b : 1 1 2 . T h e t y p e ( o n l y k n o w n spec i m en ) w a s a fe m a l e i n t h e Med i n a c o l l ec t i o n . I t see m s to be l o s t ( i t i s n o t i n MD ) . T h e o r i g i n a l d e sc r i p t i o n i s u n re l i a b l e . sekerae (An teon ) O g l ob l i n 1 924b : 1 23 ; locus typ i c u s K r a l u py ( C zechos l o v a k i a ) . T h e t y p e spec i me n s of t h e Og l ob l i n spec i e s descr i b e d f ro m Czechos l ov a k i a seem to be l os t ( t hey a re n o t i n PG , P T , BA ) . T h e or i g i n a l d e sc r i p t i on s a re u n re l i a b l e . septen trion a l i s (Gon a top u s ) W h i t t a k e r 1 9 2 8 : 388 ; locus typ i c u s C h i l l i w a ck ( B . C . , Canada ) . T h e t y p e ( o n l y k n o w n s p ec i men ) w a s a f em a l e i n t h e W h i t t a ker co l l ec t i o n . I d o n ' t k n o w the p rese n t l oc a t i on of t h i s t y p e s p ec i m e n ( i t i s n o t i n B M , W A , LA ) . The o r i g i n a l desc r i p t i on i s u n re l i a b l e . simpl icipes (Dryinop sis) Brues 1 9 1 0 : 1 7 ; locus typicus S a m u t A p i ( Sarawak ) . T h i s spec i e s , accor d i n g to R e i d ( 1 94 1 ) , s h o u l d b e a M e t h oc i d . T h e a n t e n n a e a re 1 2-seg men ted a n d t h e t a r s i of t h e fem a l e ( on l y k n o w n s pec i men ) are n o t c h e l a t e . Brues ( 1 9 1 0 ) p roposed for t h i s spec i es t h e n e w gen u s Dryinop s i s . s im u l a n s ( � a beo) A s h m e a d 1 894 : 1 98 ; l o c u s typicus S t . V i nce n t ( A n t i l l es ) . = L a beri u s simulans ( A s h m e a d ) : K i e f f e r 1 9 1 4b : 66 . T h e ho l o t y p e M ! i s i n WA ; a n o t h e r m a l e i s i n BM . The e m a r g i n a t e occ i p u t , dors a l p rocess of gonoforce p s a n d n u m b e r of p a l p a l seg men t s p l a ce t h i s spec i e s w i t h t h e G o n a topod i n es . Desc r i p t i o n : f em a l e u n k n ow n ; m a l e f u l l y w i n g e d ; l e n g t h 1 , 87 m m ; h e a d b row n , w i t h m a n d i b l e s y e l l ow ; a n t e n n a e , t h o r a x a n d p ro p o d e u m b row n ; a b do m e n red d i h s- t e s t aceou s ; l eg s ye l l o w ; a n t en n a l seg men t s i n f o l l ow i n g p ropo r t i o n s : 5 : 5 : 9 : 1 0 : 9 : 9 : 9 : 8 : 8 : 1 1 ; a n t en n a l se g m e n t 3 more t h a n t h ree t i mes a s l on g a s broad ( 4 , 5 ) ; head s h i n y , smoo t h , w e a k l y p u n c t a te ; fron t a l l i n e a b sen t ; v e r t e x w i t h o u t smooth a re a s b e t ween ey es a n d p o s t e r i o r oce l l i ; POL = 7 ; O L = 3 ; OOL = 3; b r e a d t h of oce l l i : 3 ; t e m p l e s d i s t i n c t ; occ i p i t a l c a r i n a a b se n t ; sc u t u m sh i n y , g r a n u l a te d ; n o t a u l i ce s comp l e t e , p o s t e r i or l y sep a r� a ted ; m i n i m u m d i s t a n c e b e t w e e n n o t a u l i ces shorter t h a n a n t e n n a I seg men t 2 ( 2 : 5 ) ; scu t e l l u m a n d m e t a n o t u m smoo t h , sh i n y , w e a k I y p u n c t a t e ; p ro p o d e u m ret i c u l a t e ru g o se o n p o s t e r i or s u r f ace ; dor sa l s u r f a ce smoo t h a n d

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1\ \

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1 268

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M a l e g e n i t a l i a of '- a heo sim u l a n s A s h m e a d

( type )

w i t h ou t scu l p t u r e , w i t h a med i a n f u r row ; forew i n g s h y a l i n e , w i t h o u t d a r k t r a n s v e r s a l b a n d s ; dors a l p rocess o f gon oforce p s ( F i g . 1 268 ) l o n g , s l e n d e r , d i s t a l l y b ro a de n e d ; b a s i vo l se l l a w i t h o u t n o t c h ; m a x i l l a ry p a l p i w i t h 4 segmen t s ; l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . H o s t s u n k now n . K n o w n on l y i n t h e t y p i c a l l oc a l i t y . T h e t a xonom i c d i ff i c u l t i e s of m a l e G o n a topod i n es p r e v e n t t h e assoc i a t i o n o f t h i s spec i es w i t h a g e n u s . som n i a t u s (Neodryin u s ) B r u e s 1 933 : 1 52 ; locus typicus B a l t i c a m b e r . T h e t y pes of t h e Brues foss i I spec i es were i n t h e a m b e r c o i l ect i o n a t A l b e r t u s U n i v e rs i t i H , Kon i g sberg ( now K a l i n i n g r a d ) . T h e y seem to b e l o s t . T h e o r i g i n a l descr i p t i o n s a r e u n re l i a b l e . s u ccineipes (An teon ) K i effer i n K i effer e t M a rs h a l l 1 905 : 203 ; locus typ ic u s u n ce r t a i n ( A u s t r i a or Sco t l a n d ) . = A n teon succineipes K i effer : C a meron 1 907 : 34 . = A n teon succineipes K i effe r : K i effer 1 9 1 4b : 1 63 . T h e t y p i c a l m a t e r i a l ( a t l e a s t t w o m a l e s ) w a s composed of a ma l e from Au s t r i a i n t h e M a y e t c o l l ec t i o n , a n d a m a l e from Sco t l a n d i n

Mem . Ame r .

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v e i n of m a l e of L a beo texa n u s A s h m e a d

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t h e C a meron co l l ec t i on . T h e y seem to b e l o s t ( t h e y are n o t i n BM a n d M T ) . T h e or i g i n a l desc r i p t i o n i s u n re l i a b l e . suma tran u s (Neodryin u s ) E n d e r l e i n 1 907 : 325 ; locus typ i c u s Soek a r a n d a ( Su m a t e r a , I n d ones i a ) . Psilodryin u s s u m a tranus ( E n d e r l e i n ) : K i e ffer 1 9 1 4b : 54 . = Neodryin u s s u m a tranus E n de r l e i n : R i ch a rds 1 9 53 : 66 . T h e t y pe ( on l y k n o w n spec i men ) w a s a fe m a l e i n t h e E n d e r l e i n co l l ec t i on . I h a v e n o t fou n d t h e t y p e s p ec i men i n E B , H U , L N . The or i g i n a l descr i p t i o n is u n r e l i a b l e . t a m b i n i a e (Ap terodryin u s ) E s a k i a n d H a s h i moto 1 93 5 : 20 ; locus typ i c u s =

N a se ( Am a m i Os h i m a I s . , J a p a n ) . T h e fem a l e t y p e ( o n l y k n o w n spec i men ) seems to b e l os t . T h e or i g i n a l desc r i p t i o n i s u n re l i a b l e . T h e h o s t i s Tambinia debilis S t a l ( Tropiduchida e ) ( E s a k i a n d H a s h i moto 1 93 5 ) . tars a l is ( A n teon trivia lis v a r . ) K i effer i n K i effer et M a rsh a l l 1 906 : 520 ; l o c u s typ i c u s E n g l a n d . = A n teon trivia lis tars a l is K i ef fe r : K i effer 1 9 1 4 b : 1 60 . T h e t y p e ( on l y k n o w n spec i men ) w a s a m a l e i n t h e C a meron co l l ec t i o n . I t seems to b e l o s t ( i t i s not in BM ) . T h e or i g i n a l desc r i p t i on i s u n re l i a b l e . tauricus (Echthrodelp h a x ) P o n o m a r e n k o 1 9 70 : 432 ; locus typ i c u s Sevas topo I ( Crym . , U . S. S . R . ) . T h e t y p e ( o n l y k n o w n spec i men ) w a s a fem a l e i n t h e Ponom a ren ko co l l ect i o n . I t , however , was m i ss i n g i n PK, MO and L N . T h e or i g i n a l descr i p t i o n i s u n re l i a b l e . terrestris (Dicondyl u s ) O g l ob l i n 1 9 24b : 1 20 ; locus typicus C e k a l ov ice ( C zechos l o v a k i a ) . T h e t y p e spec i m e n s of t h e O g l ob l i n spec i e s from Czechos l o v a k i a seem t o b e l os t ( t hey are not i n P G , PT, BA ) . The or i g i n a l desc r i p t i on s a re u n r e l i a b l e .

testa ceipes ( L a beo) K i effer i n K i effer e t M a r s h a l l 1 905 : 224 ; locus typicus M a t y a s fo l d ( H u n g a r y ) . = l, a berius tes taceip e s ( K i effe r ) : K i effer 1 9 1 4b : 63 . T h e t y p e ( o n l y k n o w n spec i me n ) w a s a m a l e i n t h e Szep l i g e t i co l l ect i o n . I t seems to be l o s t ( i t i s n o t i n BU ) . T h e or i g i n a l descr i p t i o n i s u n re l i a b l e . texa n u s { L a b eo } A s h m e a d 1 893 : 89 ; locus typ i c u s T e x a s ( U . S . A . ) . = L a b erius tex a n u s ( As h m e a d ) : K i effer 1 9 1 4b : 65 .

Nom i n a d u b i a

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1 8 27

M a l e g e n i t a l i a of � a beG tpx a n u s A s h m e a d ( t y p e ) ( r i g h t h a l f remov ed )

T h e h o l o t y p e M ' ( on l y k n o w n s p ec i men ) i s i n WA . T h e e m a r g i n a t e occ i p u t , dor s a l p rocess o f gonoforceps a n d n u m ber o f p a l p a l segmen t s p l ace t h i s s p ec i e s w i t h t h e G o n a topod i n e s . Desc r i p t i on : fema l e u n k n ow n ; m a l e fu l l y w i n g ed ; l e n g t h 3 mm ; b r o w n - b l ac k , w i t h a b domen b row n ; l eg s b row n , w i t h fore t i b i ae a n d t a r s i test aceo u s ; a n te n n a I segmen t s 1 - 3 b row n ( o t h er se g m e n t s m i ss i n g i n h o l o t y pe ) ; a n t e n n a I segmen t 3 more t h a n t h ree t i mes as l o n g as b ro a d ( 4 , 4 ) ;

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h e a d d u l l , g r a n u l a ted ; fron t a l l i ne a b sen t ; occ i p i t a l c a r i n a a b s e n t ; P O L � 1 1 ; O L � 6 ; OOL � 4 ; b read t h of oce l l i : 3 ; temp l es d i s t i n c t ; occ i p u t conca v e ; scu t u m d u l l , g r a n u l a ted ; n o t a u l i ces comp l e t e , p o s t e r i o r l y sep a r a t e d ; m i n i m u m d i s t ance b e t ween n o t a u l i ces shorter t h a n bre a d t h of oce l l i ( 1 : 3 ) ; sc u t e l l u m and met a n o t u m sh i n y , smoo t h , w e a k l y p u n c t a t e ; propod e u m ret i c u l a t e r u g ose ; dor s a l s u rf ace w i t h a t r a c k of med i a n f u rrow ; forew i n g s h y a l i n e , w i t h o u t d a rk t r a n s vers a l b a n d s ; d i s t a l p a r t o f ra d i a l v e i n ( F i g . 1 269 ) m u c h l o n g e r t h a n p rox i m a l p a r t ; dor s a l p rocess o f g o n oforceps ( F i g . 1 2 70 ) l on g , po i n ted a n d s l e n d e r ; b a s i vo l se l l a w i t h o u t n o t c h ; m a x i l l a ry a n d l a b i a l p a l p i w i t h 2 segmen t s ; t i b i a l s p u rs 1 , 1 , 2 . Hos t s u n k n ow n . F o u n d o n l y i n t h e t y p i c a l l oc a l i t y . T h e t a xonom i c d i ff i c u l t i e s of m a l e G o n a t opod i n es prev en t t h e assoc i a t i o n o f t h i s spec i e s w i t h a ge n u s . thoracicus (Go n a top u s ) Nees 1 834 : 387 ; locus typ icus u n k now n . See m u tillaria (Meth o c a ) L a t r e i l i e . tibia lis (A n teon ) Say 1 836 : 284 ; locus typ i c u s u n ce r t a i n ( I n d i a n a or D . C . , U . S. A. ) . � A n teon tibia lis S a y : A s h m e a d 1 893 : 96 . � A n teon ? tib ialis Say : K i effer 1 9 1 4b : 1 69 . T h e t y p e spec i me n s o f t h i s Say spec i e s seem t o b e l o s t . T h e or i g i n a l descr i p t i o n i s u n re l i a b l e . tolhursti (Dryin u s ) Fe r n a n d o 1 9 59 : 242 ; locus typ i c u s S t . Coom b s ( Ta l awakele, Sri Lanka ) . T h e t y p e ( o n l y k no w n spec i men ) w a s a fem a l e i n t h e co l l ec t i o n s of t h e Tea Research I n s t i t u te of S r i L a n k a . I do n ' t k n ow t h i s spec i e s . trichotomma (A n teon) K i effer i n K i effer e t M a rsh a l l 1 90 5 : 200 ; loc u s typ i c u s H u n g a ry . � L a s i a n teon trichotomma ( K i effer ) : K i effer 1 9 1 4 b : 1 98 . T h e t y pe ( o n l y k n o w n spec i m en ) w a s a m a l e i n t h e Szep l i g e t i co l l ec t i on . I t seems t o b e l o s t ( i t i s n o t i n BU ) . T h e or i g i n a l desc r i p t i on i s u n r e l i a b l e . tridenta t u s (A n teon) K i e ffer 1 9 1 4 b : 1 56 ; locus typ icus u n c e r t a i n ( C l am a r t or Me s n i I - I e -Roy , F r a n c e ) . � A n teon l ongiforc e p s K i effer 1 905 partim : 1 5 1 . T h e t y p i c a l m a t e r i a l ( a t l e a s t t w o m a l es ) w a s composed of spec i me n s i n t h e De G a u l l e co l l ec t i on . T h e t y p e spec i me n s seem to be l o s t ( t h e y a re n o t i n P l . T h e or i g i n a l descr i p t i on i s u n r e l i a b l e . trinodis (Gon a top u s ) K i effer i n K i effer e t M a r s h a l l 1 90 5 : 1 22 ; loc u s typ i c u s S i c k e rs h a u sen ( G erm a n y ) � Gon a top u s form icarius v a r . beta Nees 1 83 4 : 382 . � Gon a top u s trinodis K i effe r : K i effer 1 9 1 4 b : 1 1 7 . T h e t y p e ( o n l y k n o w n spec i men ) w a s a fema l e i n t h e Nees co l l ect i o n . I t seems to be l os t . T h e or i g i n a l descr i p t.i on i s u n re l i a b l e . unicarin a tu s (A n teon ) K i effer i n K i e ffer e t M a rs h a l l 1 906 : 5 1 2 ; loc u s typ icus u n c e r t a i n ( W i g h t I s l a n d a n d o t h e r l oc a l i t i e s , E n g l a n d ) . � A n teon u n icarin a tu s K i effe r : K i effer 1 9 1 4 b : 1 48 . T h e t y p i c a l m a t e r i a l w a s composed of some fema l e spec i me n s i n t h e

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M a r s h a l l c o l l ec t i on ( f rom W i g h t I s l a n d , C o r n w o r t h y and N i t or ) . t h e t y p e spec i me n s seem to be l o s t ( t hey a r e n o t i n BU ) . T h e o r i g i n a l descr i p t i on i s u n re l i a b l e . ve tus (Lestodryin u s ) Brues 1 93 3 : 1 50 ; locus typ icus B a I t i c amber . T h e t y pes of t h e B r u e s fos s i I spec i e s w e re i n t h e a m b er c o l l ec t i o n a t A l b e r t u s U n i ver s i t a t , Ko n i g s b e r g ( n ow K a l i n i n g r a d ) . T h e y seem t o b e l os t . T h e o r i g i n a l desc r i p t i o n s a r e u n re l i a b l e .

A

B

F i g . 1 27 1 - f,\e t a t ho r a x + p ropodeum ( l a t er a l l y v i e wed ) of fem a l es of Gon a topus paralep t i a s ( R . C . L . Perk i n s ) from I d y l l w i l d ( C a l i for n i a ) ( A ) a n d pseu dochromus ( R . C . L . Perk i n s ) from W a rren ( Te x a s ) ( B ) .

NOM I NA N U DA

borealis (Pren a n teon ) He l l e n 1 93 5 : 7 . chlorotettixi (Chelogyn u s ) Fen ton 1 9 1 8 a : 202 ; see A n teon os horn i Fen ton . fl a viventre (A n teon ) He l l e n 1 93 5 : 7 . formicin u s (Gon a top u s ) K ro g e r u s 1 93 2 : 1 24 ; see Gon a top u s formicarius Lj u n g h . krogeri (Gon a top u s ) H e l l e n 1 93 5 : 8 ; see Gon a top u s b a r h a tellus R i c h a r ds . longicornis (A n teon) He l l e n 1 93 5 : 7 . n itidulus (A n teon ) H e l l e n 1 93 5 : 7 . opa cifrons (A n teon) H e l l e n 1 93 5 : 7 . rap ax (Dryin u s ) H a l i d a y i n C u r t i s 1 8 29-30 : 1 1 0 . rufescens (Gon a top u s ) He l l e n 1 93 5 : 8 ; see Dicondylus dichromus ( K i effer ) . rufom a c u l a tum (A n teon ) H e l l e n 1 93 5 : 7 .

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F OSS I L D R Y I N I D S

T h e fol l ow i n g spec i e s a re k no w n :

P A L A E AR C T I C R E G I O N B a l t i c amber Th a u m a todryinus filicornis ( B rues 1 92 3 ) Th a u m a to dryin us deletus Brues 1 93 3 Th a uma todryinus gracilis Brues 1 9 33 Les todryin u s mortuorum Brues 1 933 Les todryin u s vetus Brues 1 93 3 Neodryin u s somnia t u s Brues 1 933 A lp h a dryin u s bruesi n . sp . ( Dryin i n a e ) A l p h a dryin u s b a l ticus n . sp . ( Dryinin a e ) Deino dryinus areola tu s ( Po n o m a r e n k o 1 9 7 5 ) ( A n teonin a e ) T a i my r a m b e r Cretodryin u s zherichini Ponomare n k o 1 97 5 ( Dryinin ae ) Dryinus a n ti q u u s ( Po n o m a re n k o 1 98 1 ) ( Dryin in a e ) N EA R C T I C R E G I O N Med i c i n e H a t a m b e r 12ichardsidryin u s c a n a densis ( Po n o m a r e n k o 1 98 1 ) ( Dryinin a e ) N EO T R O P I C R E G I ON Do m i n i c a a m ber Dryin u s p a l aeodomin i c a n u s C u r rado a n d O l m i

( Dryin in a e )

T h e B r u e s spec i e s a re not recog n i z a b l e , because t h e t y pe spec i m e n s ( on l y k n ow n spec i mens ) s e e m to b e l os t . T h e o r i g i n a l desc r i p t i o n s are u n r e l i a b l e . B i b l i o g r a p h i c references a b o u t t h e Brues spec i es a re in the c h a p ter "Nom ina dubia a n d incert a e sedis spec i e s" . About t h e o t h e r spec i e s refere n ces i n t h e b o d y o f t h e t ex t . Acco rd i n g t o M r . A . G . Ponom a r e n k o ( v er b a l comm u n i c a t i on ) t h e a bove spec i es w e re l i v i n g i n t h e fo l l ow i n g per i od s : D om i n i c a a mb e r : B a l t i c a mb e r : M ed i c i n e H a t a mb e r : T a i my r a mb e r : C r . zh e r i c h i n i D . an t i q uu s :

2 7 - 3 0 , 0 0 0 , 0 0 0 y ea r s a g o 4 0 - 4 5 , 0 0 0 , 0 0 0 y ea r s a g o 7 0 - 7 5 , 0 0 0 , 0 0 0 y ea r s a g o 8 0 , 0 0 0 , 0 0 0 y ea r s a g o 1 0 0 , 0 0 0 , 0 0 0 y ea r s a g o

P a r a s i t es

1 83 1

PARAS I T E S OF D R Y I N I D S

T h e p u p a e of d ry i n i d s c a n be p a r a s i t i z ed by h y pe rp a r a s i t o i d s of t h e f a m i l i es Ceraphron ida e, Diapriidae a n d Encyrtida e .

CERA PHF!ONIDAE

Ceraphron a b normis R . C . L . Perk i n s i n H a w a i i a n I s l a n d s w a s b re d ou t of cocoo n s of Haplogon a topus vitien sis R . C . L . P e r k i n s a n d Ech throdelp h a x fairchildii R . C . L . Perk i n s . I t a t t ac k s i t s h o s t o n l y a f t e r t h e d r y i n i d l a r v a h a s m a d e i t s cocoon . I n a b o u t t h ree w e e k s Cer a p h ron i d s e m e r g e from cocoon s . Ceraphron idae a re a l so p a r a s i tes o f fJaplogon a top u s a n d Pseudogona top u s i n t h e P h i l i p p i n e s , acco r d i n g to C h a n d r a ( 1 980b ) .

DIA PRIIDAE

D i a p r i i d p a r a s i tes of d r y i n i d s b e l o n g to the ge n u s lsm a r u s H a l i d a y ( su b f a m i l y Belytin a e ) . I n E n g l a n d , accord i n g to Jerv i s ( 1 9 79 a , 1 980c ) a n d W a l o ff ( 1 97 5 ) , Ismarus fl a vicornis T homson a n d I . h a lidayi F o r s t e r a re p a r a s i tes of A n teon s p p . ; Ismarus rugulosus Forster i s p a r a s i t e of A n teon (Chelogyn u s g ro u p ) s p p . a n d Lonchodryin u s (= I'ren a n teon ) sp p . ; 1 smar' u s dorsiger C u r t i s i s p a r a s i t e o f A p helop u s s p p . A l l D i a p r i i d s w e re b r e d o u t of cocoo n s . Acco r d i n g to C h a m b e rs ( 1 95 5 ) Ismarus fla vicornis i s p a r a s i te of A n teon fl a vicorne ( D a l m a n ) . See a l so N i x on ( 1 9 57 ) . I n S w i t z e r l a n d ( W a l l 1 967 ) t h e s a me f o u r spec i e s of lsmarus a re k no w n . T h e h o s t s shou l d be A n teon sp p . lsmarus fla vicornis i n Sw i t zer l a n d s h o u l d b e p a r a s i t e o f A n teon flavicorne ( D a l m a n ) .

ENCYRTIDAE

T h e m o s t p a r t of p a r a s i tes of d r y i n i d s b e l on g s to t h i s f a rn i I y . A l l k no w n spec i e s were b red o u t of cocoo n s . T a c h i k a w a ( 1 9 8 1 ) p roposed t h e f o l l o w i n g l i s t o f wor l d E n c y r t i d g e n e r a p a r a s i t o i d s o f D ry i n i d s : 1 ) Cha lcerinys R . C . L . P e r k i n s 1 906b ; 2 ) Cheiloneurus West wood 1 833 ( = Bla tticida A s h m e a d 1 904 ; Chrysopop h a g u s A sh m e a d 1 894 ; Crista tithorax G i r a u I t 1 9 1 1 ; Netach eiloneurus Hoffer 1 95 7 ; Saron o tum R . C . L . Perk i n s 1 906b ) ;

1 8 32

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I n s! . ,

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3) Ech throdryln u s R . C . L . Perk i n s 1 906b ; 4 ) Ech throgon a top u s R . C . L . Perk i n s 1 906b ; 5 ) Helegon a top u s R . C . L . Perk i n s 1 906b ( = Hazmb urkla Hof fer 1 954 ; Masencyrtus Hoffer 1 960 ; Paludencyrtus Hof fer 1 965 ; Schedioides Mercet 1 91 9 ) ; 6 ) Hypergon a top u s T i m b er l a k e 1 92 2 . I n t h e P a I a e a r c t i c reg i on p a p ers on t h e a b o v e g e n e r a were p u b l i shed by H o f f e r ( 1 954, 1 95 7 , 1 96 0 , 1 96 5 , 1 969 ) , Merc e t ( 1 9 1 9 ) , T r j a p i tz y n ( 1 964 ) a n d Sze l e n y i ( 1 9 72 ) . I n t h e g e n u s flelegon a top u s Sze l e n y i ( 1 97 2 ) l i s ted t h e f o l l o w i n g P a l ae a rc t i c spec i es : ponomaren koi T rj a p i t z y n 1 96 4 , formo s u s Mercet 1 9 1 9 , r a s n i tzyn i Trj a p i t z y n 1 96 4 , n ikolskayae ( Hoffer 1 965 ) , citripes ( E rdos 1 957 ) , dimorp h a Hof fer 1 9 57, fa cia lis Sze l e n y i 1 9 72 . No i n for m a t i o n a bo u t t h e h o s t s w a s g i ven . I n t h e A u s t r a l i a n reg i o n t h e f o l l ow i n g spec i es a re k n o w n : Ech throdryin u s R . C . L . Perk i n s destructor R . C . L . Perk i n s 1 906b : ob t a i n e d f rom Neodryin u s R . C . L . Perk i n s a n d Dryi n u s L a t re i l l e ( = Para dryin u s R . C . L . Perk i n s ) ; Queen s l a n d ( Austra l i a ) . Ech throgona top u s R . C . L . Perk i n s exitio s u s R . C . L . Perk i n s 1 906b : o b t a i n e d f rom Gon a top u s L j u n g h ; B r i s b a n e , C a i r n s , B u n d a b e r g ( Qu e e n s l a n d , A u s t r a l i a ) ( R . C . L . Perk i n s 1 906b ) ; So l omon I s l a n d s ( M ac Q u i l l a n 1 974 ; C h i u 1 979 ) . Ch a lcerinys R . C . L . Perk i n s eximia R . C . L . Perk i n s 1 906b : ob t a i ned from Gona top u s Lj u n g h ; S y d ney ( New So u t h Wa l es , A u s t r a l i a ) , C h i l ders and B u n d a b erg ( Quee n s l a n d , Austra l i a ) . Cheiloneurus West wood americ a n u s ( R . C . L . Perk i n s 1 906b ) ( = Saron otum america n um } : ob t a i ned f rom many d r y i n i d s amon g w h i c h Haplogon a top u s vitien sis R . C . L . Perk i n s a n d Pseu dogon a top u s hospes R . C . L . Perk i n s ; H a w a i i a n I s l a n d s . Th i s spec i e s w a s b red i n Hon o l u l u f rom N e a rc t i c d ry i n i d s . Acco rd i n g t o T i m b e r l a k e ( 1 9 1 9 ) i t w a s " p rob a b l y e i t h e r a cc i d en t a l l y or perh a p s i n t e n t i on a l l y I i be r a ted , t ho u g h i g n ora nce of t h e i r n a t u re , from m a t er i a l se n t t o Hon o l u l u f rom O h i o by Koe b e l e i n 1 903" . T h e reprod u c t i on of eh . americ a n u s see ms re g u l a r l y p a r t h e n o g e n e t i c ( t h e l y o t o k y ) . See a l so F l a n ders ( 1 943 ) . a u stra l i a e ( R . C . L . Perk i n s 1 906b ) ( = Saron otum a u str a l i a e } : ob t a i n e d from Pseu dogon a top u s R . C . L . Perk i n s ; B u n d a berg ( Quee n s l a n d , A u s t ra l i a ) . C o l l ec t e d a l so at O l ow a l u ( M a u i , H a w a i i a n I s l a n d s ) ( Swezey 1 9 1 9 ) a n d i n So l omon I s l a n d s ( M ac Q u i l l a n 1 974 ; C h i u 1 979 ) . chlorodryini R . C . L . Perk i n s 1 906b : ob t a i n e d from Dryin u s L a t r e i l l e ( = Chlorodryinus R . C . L . Perk i n s ) ; K u r a n d a ( Q uee n s l a n d , A u s t r a l i a ) . gon a topodis R . C . L . Perk i n s 1 906b : ob t a i ned f rom Pseudogona top u s R . C . L . Perk i n s a n d Echthrodelp hax R . C . L . Perk i n s ; C a i rn s , K u r a n d a , B u n d a be r g , C h i I d ers ( Q ueens l a n d , A u s t r a l i a ) . q u a dricolor ( G i r a u l t 1 9 1 5 ) ( see Or i e n t a l spec i es ) .

P a r a s i tes

1 83 3

Helegona topus R . C . L . P e r k i n s pseudoph a n es R . C . L . P e r k i n s 1 906b : o b t a i n e d f rom Pseu dogona topus hospes R . C . L . P e rk i n s : P u u n e n e ( M a u i , H a w a i i a n ! s l a n d s ) ( Sw ezey 1 9 1 9 ) . Th i s spec i es w a s b red from N e a rc t i c Gon a top u s or a l l i ed form s . Accord i n g to T i m b e r l a k e ( 1 9 1 9 ) i t w a s " p rob a b l y e i t h e r a cc i d en t a l l y or perh a p s i n t en t i o n a l l y l i b e r a t e d , t h o u g h i g n ora nce of t h e i r n a t u r e , f rom m a t e r i a l se n t t o Hon o l u l u from O h i o b y Koebe l e i n 1 903" . Hypergona top u s T i m ber l a k e brunneipes T i m b e r l ak e 1 9 22 : ob t a i n e d f rom Dicondylus perk insi ( As h m e a d ) (= Pseudogon a top u s perkin s i l ; Ki l a u e a ( H a w a i i ) ( T i mber l a k e 1 9 22 ) . fl a vipes T i m b e r l a k e 1 92 2 : ob t a i n e d from Dicondylus perk insi ( As h m e a d ) ( = Pseu dogona top u s perkin si l ; H a l a w a ( O a h u , H a w a i i a n I s l a nds ) ( T i mber l ake 1 922 ) . h a vlaiiensis ( R . C . L . Perk i n s 1 9 1 2 ) ( = Eh throgona top u s h a ,,'aiiensis ) : o b t a i n e d from Dicondyl u s perk i n s i ( A s h m e a d ) ( = Pseu dogona topus perkin s i ) ( R . C . L . Perk i n s 1 9 1 2 ) and Ech throdelp h a x fairch ildii R . C . L . P e r k i n s ( T i m b e r l a k e 1 92 2 ) ; H a w a i i a n I s l a n d s ( Mo u n t a i n V i e w , H a w a i i ; P a p a i k ou , H a w a i i ; H i l o , H a w a i i ; Wa i a k e a , H a w a i i ; Hon o l u l u , Oahu ) . hemipterus T i m b er l a k e 1 9 2 2 : K a u m uohon a ( O a h u , H a w a i i a n I s l a n d s ) ; t h e host is u n k n ow n . moloka ien s is ( A s h m e a d 1 90 1 ) ( = Microterys moloka ien s is ) : Mo l ok a i ( H a w a i i a n I s l a n d s ) ( T i m b e r l a k e 1 9 22 ) ; t h e host i s u n k n o w n . o a h uensis T i m b e r l a k e 1 9 22 : M t . K a a l a ( O a h u , H a w a i i a n I s l a n d s ) ; t h e h o s t is u n k n ow n . vulcan u s T i m ber l ak e 1 9 22 : o b t a i n e d from Dicon dylus perki n s i ( As h m e a d ) ( = Pseu dogon a top u s perkin s i ) ; K i l a uea ( H a w a i i ) ( T i m b er l a k e 1 92 2 ) . I n t h e N e a rc t i c re g i o n t h e fo l l o w i n g spec i e s a re k n ow n : Ech throgon a top u s R . C . L . Perk i n s p a chycep h a l u s R . C . L . Perk i n s 1 906b : ob t a i ned f rom Gon a top u s or a l l i ed for m s ; O h i o ( U . S . A . ) . Helegon a top u s R . C . L . Perk i n s p s e u dop h a n es R . C . L . Perk i n s 1 906b ( see A u s t r a l i a n spec i e s ) . Cheilone urus Wes t w oo d americ a n u s ( R . C . L . Perk i n s 1 906b ) ( see A u s t r a l i a n spec i e s ) . swezeyi A s h m e a d 1 90 3 ; ob t a i n e d f rom Neodryin u s typh locyb a e ( A s h m e a d ) ( = Dryin u s ormenidis A s h m e a d ) ; O h i o ( U . S . A . ) ( Swezey 1 90 3 ; R . C . L . P e r k i n s 1 906b ) . I n t h e O r i e n t a l reg i o n t h e fo l l o w i n g spec i es a re k n o w n : Cheiloneurus Wes t woo d q u a dricolor ( G i r a u l t 1 9 1 5 ) ( = Crista tithorax q u a dricolor ) : ob t a i ned f rom a D r y i n i d p a r a s i t e of Pyril l a ( M a n i 1 9 39 ) ; L a k s h m a n a n ( De l h i , I n d i a ) ; t h i s spec i e s , h o w e v e r , w a s desc r i bed from Queen s l a n d ( A u s t r a l i a ) ( i n A u s t r a l i a t h e h o s t i s u n k n ow n ) . j a v a n u s R . C . L . Perk i n s 1 9 1 2 : ob t a i n e d from Dryin u s ( p ro b a b l y s t a n toni

1 834

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Ent.

I nst . ,

no.

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Ashmead ; = Paradryinus j a v a n u s R . C . L . Perk i n s 1 9 1 2 ) ; Pek a l o n g a n ( J a v a , I n dones i a ) . s p ec i es ? ( R a h m a n a n d N a t h 1 940 ) : a spec i es of Ch eiloneurus w a s ob t a i ned f rom cocoo n s o f Richards idryin u s pyrillae ( K i effe r ) ( = Les todryin u s pyrillae ) i n P u n j a b ( I n d i a ) . Helegon a top u s R . C . L . Perk i n s p u lchricornis H a y a t a n d Verma 1 978 : A l i g a rh ( U t t a r P r ade s h , I n d i a ) ; t h e h o s t i s u n k n ow n . I n the E t h i o p i a n reg i o n , accord i n g to C a r n e g i e 1 97 5 , a spec i e s o f Lestodryin u s p a ra s i t i z i n g Numici a viridis M u i r i s p a r a s i t i zed b y Cheilon eurus s p . ; Sou t h Af r i c a . I n the Neotrop i c reg i o n , accord i n g t o G u a g l i u m i ( 1 962 ) , a spec i es of Pseu dogon a top u s p a r a s i t i z i n g Saccharosydne s a cch arivora is p a r a s i t i z ed i n V e n ez u e l a b y Ech throdryin u s s p . I n P u e r t o R i co , accord i n g to B a r t l e t t ( 1 939 ) , Gon a top u s s p . ( hereby descr i b ed a s Gon a top u s b a rtletti n . sp . ) p a r a s i t i z i n g Baldulus ma idis i s p a r a s i t i zed b y a new spec i es of Ooencyrtus . De Sa n t i s ( 1 96 3 ) l i s t e d !!elegon a topus pseudop h a nes R . C . L . Perk i n s f rom Argen t i n a . I n V e n e z u e l a , accord i n g t o T e r a n ( 1 980 ) , a spec i es of Echthrodryin u s p a ra s i t i zes a n u n k n o w n spec i e s of Pseu dogon a top u s .

HOST H O P P E R R E C O R D S

PALAEAR C T I C R E G I O N

Cicadellidae Idioceri n a e

I d i o c e r u s s t i g ma t i c a l i s L ew i s . . . . . P op u l i c e r u s a l b i c a n s ( K i r s c h b a u m ) . P . c o n f u su s ( F l o r ) . . . . . . . . . . . . . . . . P . l am i n a t u s ( F l o r ) . . . . . . . . . . . . . . . P . p op u l i ( L i n n a e u s ) . • • . . . . • • . . . . . R h y t i d o d u s d e c i mu sq u a r t u s ( S ch r a n k T r e mu l i c e r u s d i s t i n g u e n d u s

. . . .

. . . .

. . . .

. . . .

. . . .

. An t e on f l a v i c o r n e ( D a l m . . An t eon f l a v i c o r n e ( D a l m . . An t e o n f l a v i c o r n e ( D a l m . . An t e on f l a v i c o r n e ( D a l m . . . . • . . An t e o n f l av i corne ( Da l m . ) . . . . . An t eon a r cua t u m K i e f f e r A . f l a v i c o r n e ( D a l ma n )

( K i r s chbau m ) . An t e on

f l a v i c o rn e

( Da l m . )

Cicadellidae /Ilacropsinae

On c o p s i s

f l av i co l l i s

( L i n n a e u s ) . . . . . . . . . A n t e o n b r a ch y c e ru m ( D a l . A.

) ) ) )

j u r i n eanum La t r e i I I e

1 83 5

Hosts

Cica dellidae Aphrodinae

A p h r od e s b i c i n c t a

( Sch rank )

.

•

•

•

G o n a t op u s b i l i n e a t u s K i e f f e r G . st r i a tus K i ef f er

Cicadellidae Deltocep h a l i n a e

Ada r ru s

t a u ru s R i b au t

•

.

.

.

.

•

.

.

.

.

G on a t op u s d i s t i n gu endu s K i e f f e r G. G. G. Tet

A r a l d u s p r op i n q u u s

l una t u s K l ug s e p s o i d e s We s t w o o d sp e c t r u m ( S n e l l en v a n V o l l e n h . ) r o d o n t o ch e l y s p u l i c a r i u s ( K l u g )

( F i eb e r ) . . . . G on a t o p u s d i s t i n g u e n d u s K i e f f e r G . s ep s o i d e s We s t w o o d G . sp e c t r u m ( Sn e l l e n v a n V o l l en h .

A r o c ep h a l u s l a n g u i d u s ( F l o r ) . . . G on a t op u s s e p s o i d e s We s t w o od A . sag i t t a r i u s R i b a u t . . . . . . . . . . G on a t o p u s s e p s o i d e s We s t w o od A r t h a l d e u s p a s c u e l l u s ( F a l l e n ) . G on a t o p u s l u n a t u s K l u g G . s ep s o i d e s We s t w o o d L o n c h od ry i n u s ru f i c o r n i s ( D a l ma n ) C i r c u l i f e r d u b i o s u s ( Ma t s u m . ) . . G on a t o p u s d i s t i n g u e n d u s K i e f f e r ( = D e l t o c e p h a l u s ru s c i n o n e n s i s R i b a u t ) D i p l o c o l en u s a b d om i n a l i s ( F . ) . . G o n a t o p u s f o r m i ca r i u s L j u n g h G . s e p s o i d e s We s t w o o d My s t r o p h o r u s f o rm i ca e f o r m i s R u t h e D . n i g r i f r o n s ( K i r s ch b a u m ) . . . . . G o n a t op u s f o r m i ca r i u s L j u n g h D o r a t u r a s p . . . . . . . . . . . . . . . . . . . . G o n a t op u s g r a e c u s O l m i E l y ma n a s u l p h u r e l l a ( Z e t t . ) . . . . L on c h od ry i n u s ru f i c o r n i s ( D a l man ) E n an t i o c e p h a l u s c o rnu t u s

(H . S . ) . . . . . . . . . . . . . . . . . . . . . . . . . • G on a t opu s l una t u s K l ug E r ra s t u n u s o c e l l a r i s ( F a l l e n ) . . G on a t o p u s s e p s o i d e s We s t w o o d L o n c h od r y i n u s r u f i c o r n i s ( D a l ma n ) ( = D e l t o c ep h a l u s o c e l l a r i s ) E . sp . . . . . . . . . . . . . . . . . . . . . . . . . G on a t op u s l u n a t u s K l u g E u s c e l i s i n c i su s ( K i r s c h b a u m ) . G on a t op u s l u n a t u s 1( l u g G . s e p s o i d e s We s t w o o d ( = E . p l eb e j u s F a l l e n ) L o n c h od r y i n u s ru f i c o r n i s ( D a l ma n ) E . ob s o l e t u s ( K i r s c h b a u m ) G o l d e u s h a rp a g o ( R i b au t ) . J a s s a rg u s b i s u b u l a t u s ( T h e n ) . f l o r i ( F i eb e r ) . . • . . . . . • . . . ob t u s i v a l v i s ( K i r s c h b a u m ) . ( = D e l t o c epha l u s p i c t u ra t u s ) p s eudoc e l l a r i s ( F l o r } . . . r ep l e t u s ( F i eb e r } . . . . . . . . .

. L o n c h od ry i . G o n a t op u s . L o n c h od ry i • G o n a t op u s . G o n a t op u s L on ch o d ry i . G on a t op u s . G on a t o p u s

n u s ru f i c o rn i s ( D a l ma n ) s e p s o i d e s We s t w o od n u s ru f i c o rn i s ( D a l ma n ) s e p s o i d e s We s t w o od s ep s o i d e s We s t w o o d n u s ru f i c o r n i s ( D a l ma n ) s ep s o i d e s We s t w o od s e p s o i d e s We s t w o o d

1 836

Mem . A me r .

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la b u r r u s g e n i c u l a t u s ( R i b a u t ) ( � Ada r ru s g en i cu l a t u s ) Ma c r o s t e l e s l a e v i s ( R i b a u t ) . •

•

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I nst . ,

no. 37,

1 984

G on a t o p u s s e p s o i d e s We s t w o o d

l o n c h o d r y i n u s r u f i c o r n i s ( D a l ma n ) T e t rodon t o c h e l y s p ed e s t r i s ( Da l m . ) M o cu e l l u s c o i l i n u s ( B oh e ma n ) . • • G on a t o p u s d i s t i n g u e n d u s K i e f f e r (Es. H . ) N e p h o t e t t i x c i n c t i c e p s ( U h l e r ) . T e t r o d on t o c h e l y s s a k a i N . n i g r o p i c t u s ( S t a l ) . . • . . . . . . . T e t r o d o n t o ch e l y s s a k a i i ( E s . H . )

( � N . a p i c a l i s Mo t s c h u I s k y ) Op s i u s sp . . . . . . . . . . . . . . • . . . . P a l u d a e l o n g a t a ( Wa g n e r ) .

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. G o n a t opu s b e r n a rd i ( P i c a rd ) . . G o n a t op u s d i s t i n g u e n d u s K i e f f e r

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( � R h op a l o p y x e l o n g a t u s ) P s ammo t e t t i x a l i e n u s

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cepha l o t e s ( H . - S . ) . . . . . . . . c o n f i n i s ( D a h l b om ) . . . . . . . .

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S o rh o a n u s a s s i m i l i s ( F a I I e n ) ( � D e l t o c ep h a l u s a s s i m i l i s ) S . x a n t h on eu r u s ( F i eb e r ) . . . . S t r e p t a n u s s o rd i d u s ( Z e t t . ) . ( � E u s c e l i s s o rd i d u s ) T h a mn o t e t t i x d i l u t i o r ( K i r s . T h a mn o t e t t i x sp . . . . . . • . . . . . . T u r ru t u s s o c i a l i s ( F l o r ) . . . .

. .

s e p s o i d e s We s t w o o d

G . sp e c t r u m ( Sn e l l e n v a n V o l l e n h . G on a t op u s f o rm i c a r i u s l j u n g h G . l u n a t u s K l ug G . s ep s o i d e s We s t w o od . l o n c h o d r y i n u s r u f i c o r n i s ( D a l ma n . G on a t op u s f o rm i c a r i u s l j u n g h G . s e p s o i d e s We s t w o o d l o n ch od r y i n u s r u f i c o r n i s ( D a l ma n • G o n a t opu s s ep s o i d e s We s t w o o d l o n c h od r y i n u s r u f i c o rn i s ( D a l ma n . G o n a t opu s s e p s o i d e s We s t w o o d . G o n a t o p u s s e p s o i d e s We s t w o o d . G o n a t op u s c a mp e s t r i s P o n oma r e n k o G . d i s t i ngu endu s K i e f f e r G . h 0 rv a t h i K i e f f e r G . l una t u s K l ug G . r h a e n s i s P o n o ma r e n k o G . s ep s o i d e s W e s t wo o d . G o n a t opu s s ep s o i d e s We s t w o od

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. . . G o n a t opu s s ep s o i d e s We s t w o o d . . . l on c h o d r y i n u s r u f i c o r n i s ( D a l ma n ) ) . . G o n a t op u s b i l i n e a t u s K i e f f e r • . G o n a t op u s p o p ov i P on oma r e n k o . . . G o n a t op u s s e p s o i d e s We s t w o o d

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( F a l l e n ) . . . Ap h e l op u s a t r a t u s ( D a l ma n ) A . s e r r a t u s R i cha rd s A . wah l b e rg i ( B oh ema n ) . . . . . . . . . A p h e l op u s a t r a t u s ( D a l ma n ) A l n e t o i d i a a l n e t i ( D a h l b o m ) . . . . A p h e l op u s me l a l eu cu s ( D a l ma n ) A . s e r ra t u s R i cha rd s

1 837

Hosts

A s y mme t r a s c a d e c ed e n s ( P a o l i ) . C h l o r i t a sp • . . . • • . • • • • • . . . . . . . E d wa r d s i a n a a v e l l a n a e ( E d w . ) . . E . c r a t a e g i ( D ou g l a s ) • • . . • • . • • ( = T y p h l o c y b a f r o g g a t t i B ak e r ) E . f l a v e s c en s ( F a b r i c i u s ) . . . • • E . g e o me t r i c a ( S c h r a n k ) . . • . . . . E.

h i pp o ca s t a n i

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. . . •

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L o n c h od ry i n u s r u f i c o r n i s ( D a l ma n ) Ap h e l op u s camu s R i c h a r d s . Ap h e l o p u s me l a l eu cu s ( D a l ma n ) • Ap h e l o p u s a t ra t u s ( D a l ma n ) A . me l a l eu cu s ( D a l ma n ) • Ap h e l op u s me l a l eu cu s ( D a l ma n ) . Ap h e l op u s me l a l eu cu s ( D a l ma n ) A . s e r r a t u s R i ch a rd s . Ap h c l op u s a t r a t u s ( D a l ma n ) A . me l a l eu cu s ( D a l ma n ) • Ap h e l op u s me l a l e u c u s ( D a l ma n ) . Ap h e l o p u s a t ra t u s ( D a l ma n ) A . me l a l e u c u s ( D a l ma n ) . L o n c h od r y i n u s r u f i c o rn i s ( D a l ma n ) . L o n c h o d r y i n u s r u f i c o r n i s ( D a l ma n ) . Ap h e l op u s s e r r a t u s R i ch a r d s . A p h e l op u s a t ra t u s ( D a l ma n ) . A p h e l o p u s a t ra t u s ( D a l ma n ) . Ap h e l o p u s a t ra t u s ( D a l ma n ) . Ap h e l op u s a t r a t u s ( D a l ma n ) . Ap h e l op u s a t ra t u s ( D a l ma n ) . Ap h e l op u s n i g r i c ep s K i e f f e r . A p h e l op u s me l a l e u c u s ( D a l ma n ) . Ap h e l o p u s a t r a t u s ( D a l ma n ) A . me l a l eu cu s ( D a l ma n ) A . s e r r a t u s R i ch a r d s . Ap h e l op u s s e r r a t u s R i ch a r d s . Ap h e l op u s s e r r a t u s R i ch a rd s . Ap h e l o p u s me l a l e u c u s ( D a l ma n ) . Ap h e l o p u s a t r a t u s ( D a l ma n ) •

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A . s e r r a t u s R i ch a rd s Ap h e l o p u s a t ra t u s ( D a A . me l a l e u c u s ( D a l ma n T y p h l o c y b a b i f a s c i a t a ( B oh . ) • . . Ap h e l op u s a t ra t u s ( D a T . q u e r c u s ( F a b r i c i u s ) . . . . . . . . . A p h e l op u s a t ra t u s ( D a A . me l a l eu c u s ( D a l ma n R.

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C i x i u s c on t a m i n a t u s F l o r . . . . . . . D ry i n u s c o l l a r i s C . n e r v o s u s ( L i n n a e u s ) . . . . . . . . . D ry i n u s c o l l a r i s T a c h y c i x i u s p i l o su s ( O l i v i e r ) . . D ry i n u s c o l l a r i s

( L i n n a eu s ) ( L i n n a eu s ) ( L i n n a eu s )

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C r i orno r p h u s a l b oma r g i n a t u s C . . . D i c o n d y l u s b i c o l o r D e l p h a c i n u s me s ome l a s ( B o h . ) . . . D i c o n d y l u s b i c o l o r D i c r a n o t r op i s d i v e r g en s K i r s D . h a ma t a ( B o h e ma n ) . . . . . . . • .

( H a l i d ay ) ( H a l i d ay )

P s e u d og on a t op u s r o s e l l a e C u r r o O . D i c o n d y l u s b i c o l o r ( H a l i d ay ) P s e u d og on a t op u s d i s t i n c t u s ( K . ) P s . r o s e l l a e Cu r rado and O l m i D i t r op i s p t e r i d i s ( Sp i n o l a ) . . . . D i c o n d y l u s b i c o l o r ( H a l i d a y ) P s e u d o g on a t opu s d i s t i n c t u s ( K . ) G r av e s t e i n i e l l a b o l d i ( S c o t t ) . . D i c o n d y l u s b i c o l o r ( H a l i d a y ) .

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P s e u d o g on a t opu s d i s t i n c t u s ( K . ) ( B oh . ) . D i c o n d y l u s b i c o l o r ( H a l i d a y ) P s e u d og on a t op u s d i s t i n c t u s ( K . ) ( = L a od e l p h a x e l e g a n t u l u s ) J a v e s e l l a d i s c o l o r ( B o h eman ) . . . P s e u d o g o n a t o p u s d i s t i n c t u s ( K . ) J ob s c u r e l l a ( B oh eman ) . . . . . . . . D i c o n d y l u s b i c o l o r ( H a l i d a y ) J . p e l l u c i d a ( F a b r i c i u s ) . . . . . . . D i condy l u s b i c o l o r ( H a l i d ay ) P s e u d o g on a t op u s d i s t i n c t u s ( K . ) K o s s w i g i a n e l l a ex i g u a ( B oh . ) . . . P s e u d o g on a t op u s d i s t i n c t u s ( K . ) L a od e l p h a x s t r i a t e l l u s ( F a l l . ) . A g o n a t op o i d e s s o l i d u s ( H a u p t ) E ch t h r od e l p h a x f a i r c h i l d i i P e rk i n s E . h o r t u s e n s i s ( Ab d u l - N o u r ) H a p l og on a t o p u s a t r a t u s E s . H a s h . H . o ra t o r i u s ( We s t wood ) H . a p i c a l i s R . C . L . P e rk i n s P s e u d o g o n a t op u s a l b o s i g n a t u s ( K . ) P s . f u l g o r i ( N a k a g a wa ) M e g a d e l p h a x s o rd i d u l u s ( S t a l ) . . D i c o n d y l u s b i c o l o r ( H a l i d a y ) D . d i ch r omu s ( K i e f f e r ) E c h t h r od e l p h a x h o r t u s e n s i s ( A . - N . ) H y l ed e l phax

e l eg a n t u l u s

H a p l og on a t op u s o r a t o r i u s ( We s t w . ) P s e u d o g on a t op u s r o s e l l a e C u r . 0 1 M e g a d e l p h a x s p . . . . . . . . . . . . . . . . . A g o n a t op o i d e s s o l i d u s ( H a u p t ) P s e u d o g on a t op u s a u g u s t a e C u r . 0 1 Me t a d e l p h a x p r op i n q u a ( F i eb e r ) . P s e u d o g on a t op u s s e p t ernd en t a t u s S . Mu e l l e r i a n e l l a f a i r ma i r e i ( P . ) . D on i s t h o r p i n a p a l l i d a ( C e b a l l o s ) ( = M . ex t r u s a S c o t t ) Mu i r o d e l p h a x a u b e i ( P e r r i 5 ) • • • . A g o n a t o p o i d e s s o l i d u s ( H a u p t ) N i l a p a r v a t a l u g e n s ( S t a l ) . . . • • . E c h t h r od e l p h a x f a i r c h i l d i i P e r k i n s

1 839

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( = N . o ry z a e Ma t s u mu r a ) P a ra l i b u rn i a d a l e i ( 5 c o t t R i b au t od e l ph a x a n g u l o s u s R . c o l l i n u s ( B oh eman ) . . . . R . i m i t an s ( R i bau t ) . . . . . .

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P s e u d o g o n a t op u s f l a v i f emu r E s . H . . P s e u d o g o n a t op u s d i s t i n c t u s ( K . ) . P s e u d o g o n a t op u s d i s t i n c t u s ( K . ) . D i condy l u s b i c o l o r ( H a l i day ) . E c h t h r o d e l p h a x h o r t u s en s i s ( A . - N . ) P s eu d og on a t op u s a l b o s i g n a t u s ( K . ) P s . l i g u s t i cu s C u r r a d o and O l m i . D i c o n d y l u s b i c o l o r ( H a l i d ay ) D o n i s t h o rp i n a p a l l i d a ( C eb a l l o s ) . E ch t h r od e l phax f a i r c h i I d i i P e rk i n s . D i c o n d y l u s b i c o l o r ( H a l i d ay ) . D i condy l u s b i c o l o r ( Ha l i day ) D . d i c h r o mu s ( K i e f f e r )

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C a l i s c e l i s sp . . . . . . . . . . . . . . . . . . B o c ch u s eu r o p a e u s ( B e r n a rd ) H y s t e r op t e ru m f l a v e s c en s ( 0 1 . ) . R i c h a r d s i d r y i n u s c o r s i c u s ( Ma r . ( = H . 9 ry I I 0 i d e s F . ) H . l a t i f r on s . . . . . . . . . . . . . . . . . . . D ry i n u s sand e r i O l m i I s s u s c o l e op t r a t u s ( F . ) . . . . . . . . D r y i n u s c o l l a r i s ( L i n n a e u s )

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. N e od r y i n u s j a v a n u s . N e o d ry i n u s j a v a n u s

( R o ep k e ) ( R oepk e )

Lophopidae

. D ry i n u s i n d i c u s ( K i e f f e r ) . R i c h a r d s i d ry i n u s p y r i l a e ( K . R i c h a rd s i d r y i n u s p y r i l l a e ( K . R i c h a r d s i d ry i n u s p y r i I l a e ( K . A g o n a t op o i d e s p y r i I l a e ( Ma n i D ry i n u s i n d i cu s ( K i e f f e r ) R i c h a rd s i d r y i n u s p y r i l l a e ( K

P h r omn i a ma rg i n e l l a . . . . . P y r i I I a a b e r ra n s . . • • • • • P . p e rp u s i I i a ( Wa l k e r ) P . pu sana . . . P y r i I I a sp • . • • • • • • . . . .

. ) . ) . ) )

. )

Delp h a c idae

D i c r a n o t r op i s sp . . . . . . . . . . N i l a pa rva t a l u g en s ( S t a l ) .

Soga t a

sp . . . . . . . . . .

Sog a t e l l a f u r c i f e ra

Soga t e l I a s p . . . . . . . S t e n o c r a n u s sp . . . . . T oy a a t t e n u a t a D i s t

.

( H o rv a t h )

. P s e u d og on a t op u s n u d u s P e r k i n s . D i c ondy l u s i n d i a n u s D l m i E c h t h r o d e l p h a x f a i r c h i l d i i P e rk i n s H a p l og on a t op u s ap i c a l i s R . P e r k i n s P s e u d o g o n a t op u s n u d u s R . P e rk i n s P s . s a r a wa k i Mo c z a r . H a p l og on a t op u s a p i c a l i s R . P e r k i n s P s e u d o g o n a t op u s s o g a t a e R oh w e r . E c h t h r od e l p h a x f a i r c h i l d i i P e rk i n s H a p l o g o n a t o p u s ap i c a l i s R . P e r k i n s P s e u d o g o n a t op u s n u d u s R . P e rk i n s . P s e u d o g o n a t o p u 5 sa r a wak i Mo c z a r . P s e u d o g o n a t o p u s n u d u s R . P e rk i n s . P s e u d o g o n a t op u s s a r a wa k i Mo c z a r

1 842

Mem .

Am er .

En t .

I nst . ,

n o . 37,

1 984

NEAR CT I C R EG I ON

Cica dellidile Deltocep h a l in a e

( F i t ch ) . Amb l y s e l l u s c u r t i s i ( = A t h y sa n u s cu r t i s i i ) B a l c l u t h a i mp i c t a ( V a n D u z e e ) C h l o r o t e t t i x u n i c o l o r ( F i t ch ) C i r cu l i f e r t en e l l u s ( B ak e r ) .

. G on a t o p u s a sh mead i

K i ef f er

. A c r o d on t o c h e l y s a me r i c a n u s ( F . ) . An t e o n o s b o r n i ( F en t o n ) . G o n a t op u s c o n t o r t u l u s P a t t on T e t r o d o n t o c h e l y s u n i c u s ( P e rk i n s ) ( = E u t e t t i x t en e l l a ) . G o n a t op u s a s h m e a d i K i e f f e r D e l t o c ep h a l u s f l a v i c o s t a ( S t . G on a t op u s p a l l i d i c ep s ( R . P e rk i n s ) D . f u s c i n e rv o su s Van D u z e e . . G o n a t op u s a g r op y r u s F e n t on E n d r i a i n i m i c a ( Sa y ) . . . . • • • G . a s h me a d i K i e f f e r ( = D e l t ocepha l u s i n i m i cu s ) G . c on t o r t u l u s P a t t on G . p a I l i d i c ep s ( R . C . L . P e rk i n s ) G r a m i n e l l a n i g r i f r on s ( F o rb e s ) . G o n a t opu s a g r op y r u s F e n t o n G . a shmead i K i e f f e r L a t a l u s s a y i i ( F i t c h ) . . . . . . . . . . G on a t opu s a g r op y r u s F e n t on G . a shmead i K i e f f e r

Ma c r o s t e l e s d i v i s u s ( U h l e r ) M . f a s c i f r on s ( S t a I ) . . . . . . .

G . c o n t o r t u l u s P a t t on G . o mb r od e s ( R . C . L . P e r k i n s ) . T e t r o d on t o c h e l y s p l e s i u s ( F e n t on ) . G o n a t op u s a s h me a d i K i e f f e r G . c y p h on o t u s B rad l ey T e t r o d o n t o c h e l y s p l e s i u s ( F e n t on ) T . u n i c u s ( R . C . L . P e rk i n s )

. G o n a t op u s p s e u d o c h r omu s . G o n a t op u s a g r o p y ru s F en t G . a shmead i K i e f f e r . G o n a t op u s a g r o p y ru s F e n t G . a sh m e a d i K i e f f e r G . omb r o d e s F e n t o n S c a p h o i d eu s p a l u d o su s B a l l . . . . . E s a g o n a t o p u s n i g e r ( F e n t

Ph I ep s i u s sp • • • • • . • • • . • • . . • P s a mmo t e t t i x a f f i n i s ( G . - B . ) ( = D e l t ocepha l u s a f f i n i s ) P s . s t r i a t u s ( L i n n a eu s ) . . . • .

( P e rk i n s ) on on

on )

Cicadellidae Typ hlocyb i n a e

E r y t h r o n e u r a c ome s ( Sa y ) . . . . E . vu l n e r a t a ( F i t ch ) . . . . . . . Forc i pa t a c i t r i n e l l a ( Z e t t . ) ( = D i k r a n eu r a f i eb e r i L o w ) H y me t t a t r i f a s c i a t a ( Sa y ) . . T y p h l o c y b a p o ma r i a Mc A t e e .

. Ap h e l o p u s a l b op i c t u s A s h m e a d . Ap h e l opu s a l b op i c t u s A s h m e a d . A p h e l opu s a l b op i c t u s A s h m e a d . A� h e l opu s b i c o l o r F e n t o n . A p h e l o p u s a l b op i c t u s A sh mead

1 843

Hosts

Fla tidae

. . N e o d r y i n u s t y p h l o c y b a e ( A sh mead ) Me t c a l f a p r u i n o sa ( Sa y ) . . . ( = O r men i s p r u i n o s a ) M i s t h a r n op h a n t i a s o n o r a n a . . . . . . Ap t e r od ry i n u s t o rv u s R . P e r k i n s D ry i n u s a l a t u s ( C r e s s on ) O r men i s s e p t e n t r i o n a l i s

( Sp . ) . . N e od ry i n u s

t y ph l o c y b a e

( A sh mead )

A c a n a Ion i ida e

. D ry i n u s a l a t u s ( C r e s s on P e r o d ry i n u s a m o e n u s R . C . A p t e r o d ry i n u s s e c u n d u s . Ap t e r od ry i n u s t o r v u s R .

A ca n a l on i a b i v i t t a t a ( Sa y ) . ( = A mp h i s c e p a b i v i t t a t a ) A . c on i c a ( Say ) . . . . . . . Amph i s c ep a i mma c u l a t a • . . . •

) . L . P e rk i n s ( F en t on ) P e rk i n s

I ssidae

. B o c c h u s rn i r a b i l i s ( R . C . L . P e r k i n s ) . o o c c h u s a r' i z o n i c u 5 ( i� . P e rk i n s )

B r u c h omo r p h a o c u l a t a N ewm . B r u ch orno r p h a sp . . . . . . . . . . .

Del[l h a ciclae

B o s t a e ra a p a c h e a n u s

( K i rk . ( = C o ch i s e a p a ch eanu s ) B . n a s u t a Ba l l . . . . . . . . . . . . D e I p h a c od e s c a rnp e s t r i 5 V a n D . l u t u l en t a ( V a n D u z e e ) . .

L i b u rn i a

) . .

. A g o n a t op o i d e s

. . .

. A g o n a t op o i d e s s y n c h r o mu s P e r k i n s

D. . . .

. D i c on d y l u s a me r i c a n u s ( R . P e rk i n s ) . D i c on d y l u s a m e r i c a n u s ( R . P e rk i n s ) P s e u d og on a t op u s f r ey t a g i O l m i P s . s t e n o c r a n i R . C . L . P e rk i n s . D i c o n d y l u s a me r i c a n u s ( R . P e rk i n s ) P s e u d o g o n a t o p u s s j o e s t ed t i ( K . ) . P s e u d o g o n a t op u s a r i z o n i c u s P e rk i n s

sp . . . . . . . . . . . . . . . .

Sa c c h a r o s y d n e sa c c h a r i v o r a . S t en o c r a n u s d o r s a l i s . S t o b a e r a sp . . . . . . • . . .

sy n c h r o mu s P e rk i n s

. P s e u d o g on a t o p u s s t e n o c r a n i P e rk i n s . P s e u d o g o n a t o p u s a r i z o n i c u s P e rk i n s P s . a u t o x e n o b i u s R . C . L . P e rk i n s

Mem brac iclae

Mem . Amer . E n ! .

1 844

Th e l i a b i ma c u l a t a F a b r

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C r ov e t t i a

1 984

the l i ae

( Gahan )

NEOTROP I C REG I ON

Cica dellidae Deltocep h a l in a e

Ba l d u l u s ma i d i s

( D e L on g } . . . . . . G on a t op u s b a r t l e t t i

O l mi

Fl a tidae

O rrnen i s sp

Me g a d ry i n u s ma g n i f i c u s R i c h a rd s N e od r y i n u s t r i n i t a t i s R i c h a r d s T h a u ma t od r y i n u s r u f u s R i ch a r d s T r i d r y i n u s ru f i c a u d a ( R i c h a rd s ) P o ek i l l op t e r a c omp l e t a J a c . . . . . T r i d r y i n u s p o e c i l op t e r a e ( R i ch . ) P . p h a l a e n o i d e s ( L . } . . . . . . . . . . . T r i d ry i n u s p o e c i l op t e r a e ( R i ch . ) ( = P o e c i l op t e r a p h a l a e n o i d e s ) .

.

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•

Delph a cidae

P e rk i n s i e l l a s a c c h a r i c i d a . . . . . . P s eu d o g on a t op u s mo r e n o i O l m i Sa c c h a r o s y d n e s a c c h a r i v o r a ( We s t w . } . . . . • • . . . . . . . . . . . . . . . . . . . . . . . . . • . . . . . . • • . . . . • • • • . . . . . . . P s e u d o g on a t op u s va r i i s t r i a t u s Fen . S og a t od e s o ry z i c o l a . . . . . . . . . . . . H a p l og on a t op u s h e r n a n d e z a e O l m i

AUSTRAL I A N R E G I O N

Cicadellidae I diocerin ae

I d i ocerus

sp . . . . . . . . . . . . . . . . . . . A n t e o n d e s t r u c t o r

(R.C.L.

P e rk i n s )

Cicadellidae Macrop s i n a e

Ma c r op s i s sp . . . . . . . . . . . . . . . . . . . A n t e o n d e s t ru c t o r

( R . C . L . P e rk i n s )

1 845

Hosts

Cic a dellidae Deltocep h a linae

A t h y s a n u s sp . . . . . . . . . . . . . . . . . . . G on a t op u s a u s t r a l i a e R . P e r k i n s G . me l a n i a s ( R . C . L . P e rk i n s ) . . . . . . A c r o d on t o c h e l y s v i t i e n s i s ( P e rk . ) D e l t o c ep h a l u s sp . . . . . . . An t e o n p a r v u l u m ( R . C . L . P e r k i n s ) G o n a t op u s a u s t r a l i a e R . P e r k i n s G . op t a b i i i s ( R . C . L . P e r k i n s ) E cJ r i n o s c op u s sp . . . . . . . . . . . . . . . . An t e on d e s t ru c t o r ( R . C . L . P e rk i n s ) A . l e i o s omum ( R . C . L . P e rk i n s ) A. E u t e t t i x sp . . . . . . . . . . . . . . E x i t i a n u s p l eb e i u s ( K i rk a ( = N ep h o t e t t i x p l e b e i u s ) I p o c o n f e r t a K i rk a l d y . . . . Ma c r o s t e l e s s p . . . . . . . . . . . ( = E u l e i mo n i o s sp . )

n i t i dum ( R . C . L .

P e rk i n s )

. . . . . . A n t e o n n i g r i c o rn e ( R . C . L . P e rk i n s ) l d y ) . . G o n a t op u s p u l c h e r r i mu s ( P e r k i n s ) . . . . . . A n t e o n my rme c o ph i l u m ( R . P e r k i n s ) . . . . . . A c r od on t o ch e l y s v i t i e n s i s ( P e r k . ) A n t e o n c o r i a c e u m ( R . C . L . P e rk i n s )

A . pa rvu l u m ( R . C . L . P e rk i n s ) G on a t opu s me l a n i a s ( R . C . P e rk i n s ) N ep h o t e t t i x sp . . . . . . . . . . . . . . . . . G o n a t op u s a u s t r a l i a e R . P e rk i n s G . op t a b i l i s ( R . C . L . P e rk i n s ) N e s o s t e l e s s a n g u i n e s c en s K i rk . . An t eon f i j i an u m O l m i P h r y n op h y e s sp . . . . . . . . . . T a r t e s su s f u l v u s ( Wa l k e r ( = T . s y r t i d i s K i rk a l d y T a r t e s s u s sp . . . . . . . . . . . .

. . . . . . . G on a t o p u s d e c o r a t u s ( R . P e r k i n s ) ) . . . . . . G on a t o p u s g i g a s ( R . C . L . P e rk i n s ) ) . . . . . . . A n t e o n c h e l og y n o i d e s ( R . P e r k i n s )

Dictyoph aridae

H a s t a h a s t a t a . . . . . . . . . . . . . . . . . . D ry i n u s

l ep t i a s

(R .C .L .

P e rk i n s )

Nogodin idae

Ga e t u l i a

sp

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.

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•

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N e o d r y i n u s n e l s on i

R .C .L .

P e rk i n s

Ric a n iidae

P r i v e s a a p h r op h o r o i d e s Wa l k e r S c o l y p opa

sp

.

•

•

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•

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.

. D r y i n u s v en a t o r ( R . C . L . P e r k i n s ) N e od r y i n u s n e l s on i R . C . L . P e rk i n s N e od ry i n u s n e l s on i R . C . L . P e rk i n s

Mem . A mer .

1 846

En t .

I nst . ,

no .

37 ,

1 984

Fl a tidae

. N e od r y i n u s k o eb e l e i R . C . L . P e rk i n s N . n e l s on i R . C . L . P e rk i n s sp . . . . . . . . . . . . . . . . . . . . . . D ry i n u s k 0 eb e l e i ( R . C . L . P e rk i n s ) D . l e p t i a s ( R . C . L . P e rk i n s ) D . v en a t o r ( R . C . L . P e rk i n s )

C o l g a r p e ra cu t u s . Co l gar

Ma s s i l a

sp . . . . . . . . . . . . . . . . . . . . . D r y i n u s k o e b e l e i ( R . C . L . P e r k i n s ) D . v e n a t o r ( R . C . L . P e rk i n s ) N e od r y i n u s k o eb e l e i R . C . L . P e rk i n s N . n e l s o n i R . C . L . P e rk i n s P h a n t i op s i s sp . . . . . . . . . . . . . . . . . D r y i n u s k o e b e l e i ( R . C . L . P e r k i n s ) N e od ry i n u s n e l s o n i R . C . L . P e r k i n s ( � Aphanophan t i a ) . N e od r y i n u s n e l s o n i R . C . L . P e rk i n s S e p h e n a sp . . . D r y i n u s k o e b e l e i ( R . C . L . P e rk i n s ) 5 i p h a n t a sp . . . . . . . D . pa l l i d u s ( R . C . L . P e rk i n s ) D . v e n a t o r ( R . C . L . P e rk i n s ) N e od ry i n u s n e l s o n i R . C . L . P e r k i n s T h a u ma t o d r y i n u s k o e b e l e i

P e rk i n s

Delph acidae

A I oh a i p o me a e . H a d e o d e l p h a x sp . I l b u rn i a c op r o s m i c o l a Mu i r . . l . k o a e K i rk . . . K e l i s i a swe z e y i L i b u r n i a sp . . . .

N i l a p a r v a t a my e r s i . . . . . . . . . . . . P e r eg r i n u s ma i d i s ( A s h rn e a d ) . . . P e rk i n s i e l l a s a c c h a r i c i d a K i rk

Vanua v i

i en s i s K i rk . . . . . . . . . .

. E ch t h r od e l phax f a i r ch i l d i i P e rk i n s . E c h t h r od e l p h a x n i g r i c o l l i s P e r k i n s H a p l o g o n a t o p u s ap i c a l i s I� . P e r k i n s . D i c o n d y l u s p e r k i n s i ( A s h me a d ) . D i c o n d y l u s p e rk i n s i ( A shmead ) . D i c o n d y l u s p e r k i n s i ( A s h me a d ) . E c h t h r o d e l p h ax b i f a s c i a t u s P e rk i n s H a p l o g o n a t o p u s ap i c a l i s R . P e rk i n s P s e u d og o n a t o p u s d i c h r o rnu s P e rk i n s P s . k u r a n d a e R . C . L . P e rk i n s P s . j u n c e t o r u m R . C . L . P e rk i n s P s . o p a c u s R . C . L . P e rk i n s . D i c o n d y l u s a l p i n u s ( G ou r l ay ) . H a p l o g o n a t o p u s v i t i e n s i s P e rk i n s . E c h t h r od e l p h a x f a i r c h i l d i i P e rk i n s H a p l o g o n a t o p u s v i t i en s i s P e r k i n s P s e u d o g o n a t op u s d i c h r omu s P e rk i n s P s . h o s p e s R . C . L . P e rk i n s P s . s a c ch a r e t o ru m R . C . L . P e rk i n s . D i c o n d y l u s k i e f f e r i ( R . P e rk i n s )

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E .A.

pes t s b y E . A .

Popenoe . A p p en d i x to O b serv a t i o n s o n t w o i n sect

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Bul l .

Exp .

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Swezey O . H . , 1 9 28 . P rese n t s t a t u s of ce r t a i n i n sec t p e s t s u n d e r b i o l og i c a l con t r o l i n H a w a i i . J o u r n . Econ . E n t . 2 1 : 669-67 6 . Sze l e n y i G . , 1 97 2 . N e u E n c y r t i d e n a u s U n g a r n . A n n i s h i s t . - n a t . M u s . n a t n . H u n g . 64 : 347-353 . T a c h i k a w a T . , 1 98 1 . Hos t s of E n c y r t i d g e n e r a i n t h e Wor l d . Mem . Co l l eg e A g r i c . , E h i me U n i v . , 2 5 ( 2 ) : 8 5 - 1 1 0 . Teodorescu I . , 1 967 . G o n a topod i n e d i n Rom a n i a ( H y m e n o p t e r a D r y i n i d ae ) . An a l . Un i v . T e r a n J . B . , 1 980 .

B u c u res t i , Se r . B i o i . , 1 6 : 77-80 . L i s t a p re l i m i n a r de H y m e n o p t e r a p a r a s i tos de o t ros

i n sectos e n V e n e z u e l a . R e v . F a c . A g ro n . M a r a c a y X I : 283-389 . T h o m p s o n IV . R . , 1 954 . A c a t a l og u e of t h e p a r a s i t e s a n d p re d a t o r s of i n se c t p e s t s , Sec t i o n 2 , P a r t 3 . C o m mo n . I n s t . B i o i . C o n t r . , O t t a w a , 332 pp. T h o m s o n C . G . , 1 860 . S v e r i g e s P r oc t o t r u pe r . O f v . k . s v . V e t e n s k . Ak a d . F ii r h . 1 7 : 1 69- 1 8 1 . T i m b e r l a k e P . H . , 1 9 1 9 . O b s e r v a t i o n s on t h e Sou rces of H a w a i i a n E n c y r t i d a e ( H y m e n o p t e r a ) . P roc . H a w . E n t . Soc . 4 : 1 83 - 1 96 . T i m b e r l a k e P . H . , 1 9 2 2 . D e s c r i p t i o n s o f N e w G e n e r a a n d Spec i es o f H a w a i i a n E n c y r t i d a e ( H y m e n o p t e r a ) , I I I . P roc . H a w . E n t . Soc . 5 : 1 35 - 1 6 7 . T o w n es H . , 1 969 . T h e g e n e r a o f I c h n e u mo n i d a e , p a r t 1 . Mem . Ame r . E n t . I n s t . 1 1 : 300 p p . T o w n e s H . , 1 9 7 2 . A l i g h t - w e i g h t � \ a l a i se t r a p . E n t . N e w s 83 : 239 ) 247 . T rj a p i t z y n V . A . , 1 964 . E n c y r t i d a e p a r a s i t e s of D r y i n i d a e i n U . S . S . R . Z oo l . Z . 43 : 1 4 2- 1 45 . T u r n e r R . E . , 1 928 . New H y m e n o p t e r a o f t h e F a m i l y B e t h y l i d a e . A n n . M a g . N a t . H i s ! . , Se r . 1 0 , V o l . 1 : 1 29- 1 5 2 . U c h i d a T . , 1 927 . E i n e n e u e a r t v o n D r y i n i d e n ( H y m . ) . I n sec t a lYI a t s u m . 1 9 27 : 32-33 . W a l k e r F . , 1 83 7 . On t h e D r y i n i d a e . E n t . M a g . 4 : 4 1 1 -435 . W a l k e r F . , 1 839 . M o n o g r a p h i a C h a l c i d i t u m , 2 . Spec i e s c o l l ec t e d by C . D a r w i n , E s q . H . B a i l l i e r e , L o n d o n , 1 00 p p . W a l l I . , 1 967 . D i e I sm a r i n a e u n d Be l y t i n a e d e r Sc h w e i z . E n t o ma l . A b h a n d l . 35 ( 2 ) : 1 23 - 265 . YV a l of f N . , 1 974 . B i o l og y a n d b e h a v i o u r of some spec i e s of D r y i n i d a e ( H y me n o p t e r a ) . J . E n t . ( A ) 4 9 ( 1 ) : 97- 1 09 . W a l of f N . , 1 97 5 . T h e p a r a s i t o i d s o f t h e n y m p h a l a n d a d u l t s t a g e s of l ea f h o p p e r s ( A u c h e n o r r h y n c h a : Homop t e r a ) of ac i d ic g r a s s l a n d . T r 8. I , s . R . en t . Soc . L o n d . 1 26 ( 4 ) : 637-686 . Wes t w ood J . O . , 1 83 3 a . F u r t h e r N o t i ce s of t h e B r i t i s h p a r a s i t i c H y me n o p t e r o u s I n sec t s ; t o g e t h e r w i t h t h e ' t T r a n s a c t i o n s o f a F l y w i t h a l o n g T a i l " , o b s e r v e d by M r . E . W . Le w i s ; O b s e r v a t i o n s . M a g a z . N a t . H i s t . 6 : 4 1 4-42 1 .

a n d a d d i t i on a l

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A B B R E V I AT I ONS

! = s p e c i men seen by t h e Au t ho r o r l oc a l i t y c h ec k e cl b y F = fem a l e spec i me n F F = f e m a l e spec i m e n s M = m a l e s p ec i me n M M = m a l e s p ec i me n s

F o r a b b re v i a t i o n s of U .

S.

the Au t h or .

a n d C a n a d i a n g eo g r a p h i c a l

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K r o m b e i n , H u r d , Sm i t h and B u r k s ( 1 97 9 : C a t a l o g o f H y m e n o p t e r a i n A m e r i c a N o r t h of M e x i co ) h a v e b e e n f o l l o w s .

I N D E X T O HOSTS

T h i s i n cl e x i n c l u cl e s f a m i l i e s , h o s t s of d r y i n i d s . a b d om i n a l i s ( D i p l o c o l e n u s ) F . 6 8 1 , 1 56 5 , 1 58 0 , 1 835 ab e r ra n s ( P y r i I I a ) 9 2 6 , 1 8 4 1 A c a n a l on i i d a e 8 , 9 , 7 2 4 , 7 2 9 , 8 3 1 , 1 03 3 , 1 05 1 , 1 34 4 ,

s u b f a m i l i es ,

1 aff i n i 1 aff ini 1

g e n e r a a n d spec i e s of

35 5 , 1 35 7 , 1 843 s ( D e l t o c ep h a l u s ) G . B . 720 s ( P s a mmo t e t t i x ) G . B . 697

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a u ra t a ( E u p t e ry x ) L . 5 2 , 1 8 3 7 a u r ov i t t a t a ( L i n d b e rg i n a ) D ou g l a s 5 4 , 1 8 3 7 au s t r a l i s ( T y p h I o c y b a ) F r ogg . 1 0

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Say

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573,

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Mem . Amer . E n t .

Eu r i n o s c o p u s 5 5 1 , 1 845

561 ,

564,

eu r o p a e a ( D i c t y op h a ra ) L . 738 , 747, 1 838 E u t e t t i x 569 , 1 84 5 1 378, ex c i s a ( U n k a n od e s ) M e I 1 383, 1 839 exc i su s ( D e l ph a x ) Me l . 1 3 7 8 , 1 383, 1 839 e x i g u a ( K o s sw i g i a n e l l a ) B oh . 1 2 1 5 , 1 (1 3 8 e x i g u u s ( Mu i r o d e l p h a x ) 1 21 5

Boh .

e x t r u s a U:'u e l l e r i a n e l l a ) S e t 1 30 8 , 1 838 f a i rma i r e i ( Mu e l l e r i a n e l l a ) P e r' r i s 1 3 0 8 , 1 8 3 8 f El s c i f r o n s ( ,\ia c r 0 s t e l e s ) S t a I 1 46 8 , 1 4 7 0 , 1 6 79 , 1 7 20 f i eb e r i ( D i k r a n e u r a ) L ow 7 5 F l a t i da e 8 , 9 , 6 8 3 , 6 8 6 , 7 2 4 , 7 2 9 , 8 3 1 , 884 , 9 0 3 , 904 ,

933, 954, 970, 973, 1 05 1 , 1 06 7 , 1 08 6 , 1 1 20 , 1 34 4 , 1 355, 1 84 1 , 1 843 , 1 84 4 , 1 846 f l a v e s c e n s ( E d wa r d s i a n a ) F . 49, 1 837 f I a v e � c e n s ( E mp o a s c a ) F l o r 2 6 5 , 1 83 7 f l av e s c en s ( H y s t e r op t e ru m ) 01 iv . 914 f l a v i c o l l i s ( On c op s i s ) L . 2 9 8 , 302, 1 83 4 f l a v i c o s t a ( D e l t o c ep h a l u s ) S t a l 1 720 f l o r i ( J a s sa rgu s ) F i eb . 1 5 8 0 , 1 835 Forc i pa t a 75 f r o g g a t t i ( T y p h l o cy b a ) B a k e r 52, 75, 1 837 Fu l g o r i da e 8 , 9 , 7 2 4 , 7 2 9 , 821 , 1 84 1 f u l v u s ( T a r t e s s u s ) Vl l k . 1 8 4 5 f u r c i f c ra ( S og a t e l l a ) H o rv . 1 1 65 , 1 1 6 6 , 1 1 9 7 , 1 24 6 , 1 29 2 , 1 4 2 1 , 1 83 9 , 1 84 1

I n st . ,

no .

37,

1 984

f u s e i n e r v o s u s ( D e l t o c ep h a I u s ) V . D . 1 730 Ga e t u l i a 1 1 20 , 1 84 5 g e n i c u l a t u s ( Ad a r r u s ) R i b . 1 58 0 , 1 83 6 9en i cu I a tu s ( L a b u r r u 5 ) ,R i b . 1 836 g e ome t r i c a ( E d wa r d s i a n a ) S c h r . 4 9 , 54 , 1 837 g r y l l o i d e s ( H y s t e r op t e r u m ) F . 914 H a d e od e l p h a x 1 1 6 9 , 1 4 2 1 , 1 8 4 6 h a ma t a ( D i c r a n o t r op i s ) B o h . 1 2 0 2 , 1 2 1 5 , 1 38 3 , 1 8 38 h a rp a g o ( G o l d e u s ) R i b . 1 5 8 0 , 1 835 h a s t a t a ( Ha s t a ) 8 9 7 , 1 84 5 h ey d e n i ( A t h y sa n u s ) 1 5 6 4 h i p p o c a s t a n i ( E dwa r d s i a n a ) E d w . 49 , 5 2 , 1 837 I d i o c e r i na e 8 , 600 , 678 ,

9, 1 1 0 , 288, 1 051 , 1 1 72 , 1 43 6 , 1 49 4 , 1 50 2 , 1 83 4 , 1 84 0 , 1 844 I d i o c e ru s 3 1 1 , 4 2 2 , 5 5 1 , 1 8 4 0 , 1 844 i m i t a n s ( R i b a u t od e l p h a x ) R i b . 1 1 50 , 1 2 1 1 , 1 22 1 , 1 839 i mma c u l a t a ( A mp h i s c e p a ) 1 3 5 5 , 1 843 i mp i c t a ( B a l c l u t h a ) V a n D . 1 1 8 2 i n c i su s ( E u s c e l i s ) Kbm . 1 5 4 7 , 1 580, 1 835 i n i m i ca ( End r i a ) Say 1 6 8 8 , 1 697, 1 720, 1 730 i n i m i cu s ( D e l t o c ep h a l u s ) Say 1 720 i n su l a r i s ( R i can i a ) L a l l . 7 9 1 , 1 840 i p om e a e ( A l oh a ) 1 1 6 5 , 1 8 4 6 I s s i da e 8 , 9 , 600 , 6 1 3 , 6 4 3 , 647 , 724 , 729, 907 , 1 83 9 J a s s i da e 4 6 4 j u cu n d a ( E u p t e ry c i d a ) H . - S . 5 4 , 1 83 7 k oa e ( I l b u r n i a ) K i r k . 1 3 9 5 , 1 846

I n dex

l a e v i s ( Ma c r o s t e l e s ) R i b . 265, 1 44 0 , 1 83 6 l am i n a t u s ( I d i o c e ru s ) F l o r 31 1 l a m i n a t u s ( P op u l i c e r u s ) F l o r 3 1 1 , 1 834 l a n g u i d u s ( A r o c ep h a l u s ) F l . 1 580, 1 835 j an i e ( On c op s i s ) L . 3 2 5 l a t i f r on s ( H y s t e r op t e r u m ) 7 3 3 L i b u rn i a 1 1 6 9 , 1 2 6 6 , 1 2 9 0 , 1 2 94 , 1 2 9 8 , 1 29 9 , 1 30 4 , 1 39 0 , 1 3 9 6 , 1 4 2 1 , 1 84 3 , 1 846 L o p h op i d a e 8 , 9 , 7 2 4 , 9 0 7 , 9 2 6 , 1 05 1 , 1 1 9 3 , 1 84 1 l u g en s ( N i l a p a r v a t a ) S t a l 1 1 6 6 , 1 20 2 , 1 240, 1 246 , 1 38 7 , 1 42 1 , 1 83 8 , 1 84 1 I u r i d a ( E u ry s a ) F i e b . 1 5 1 8 l u t eo t ac t e l l a ( N . ) 5 4 1 l u t u l en t a ( D e l p h a c o d e s ) V . D . 1 25 8 , 1 27 2 , 1 39 0 , 1 843 Ma c r op s i n a e 8 , 9 , 1 1 0 , 2 8 8 , 600 , 6 7 8 , 1 05 1 , 1 1 72 , 1 43 6 , 1 49 4 , 1 50 2 , 1 834, 1 844 Ma c r op s i s 3 1 6 , 3 2 0 , 3 3 6 , 5 5 1 , 1 844 Ma c r o s t e l e s 3 2 0 , 1 8 4 5 ma i d i s ( B a l d u l u s ) D e L . 1 7 5 2 , 1 83 4 , 1 844 ma i d i s ( P e r e g r i n u s ) A s h m . 1 42 7 , 1 846 ma r g i n e l l a ( P h r omn i a ) 6 1 4 , 1 841 Ma s s i I a 8 8 4 , 9 0 4 , 1 1 2 0 , 1 1 2 3 , 1 846 Megad e l p h a x 1 2 1 2 , 1 3 1 8 , 1 8 3 8 me l i s s a e ( E u p t e r y x ) C u r t i s 5 2 , 1 837 Memb r a c i d a e 1 843

8,

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84 ,

me s ome l a s ( D e l p h a c i n u s ) 1 383, 1 838 mu i r i

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( D i c r a n o t r op i s ) K i rk . 1 23 5 , 1 389

1 873

my e r s ; ( N i l a p a r v a t a ) 1 4 0 7 , 1 8 4 6 n a su t a ( B o s t a e ra ) Ba l l 1 3 3 9 , 1 843 N eph o t e t t i x 1 1 6 6 , 1 7 8 8 , 1 7 9 1 , 1 845 n e r v o s u s ( A p h r od e s ) 1 2 1 9 n e rv o su s ( C i x i u s ) L . 7 3 8 , 1 8 3 7 n i g r i f r on s ( D i p l o c o l en u s ) K b m . 1 56 5 , 1 835 n i g r i f r on s ( G r a m i n e I I a ) F o r b e s 1 69 7 , 1 720 n i g r op i c t u s

( N eph o t e t t i x )

Sta l

1 24 8 , 1 46 3 , 1 8 3 6 , 1 8 40 N i l a p a rv a t a 1 1 6 6 , 1 3 8 7 n o d o s u s ( P sammo t e t t i x ) R i b . 320, 1 580, 1 836 N og od i n i d a e 8 , 9 , 1 0 5 1 , 1 0 G 7 , 1 1 20 , 1 845 n o t a t u s ( P s a mmo t e t t i x ) M e l . 1 580, 1 836 ob s c u r e l l a 1 4 40

( C a l l i g y p ona ) B oh .

ob 5 cu r e i I a ( J a v e 5 e I I a ) B oh . 1 38 3 , 1 8 38 ob s o l e t u s ( E u s c e l i s ) K i r s ch . 265, 1 835 ob t u s i v a l v i s ( J a s s a r g u s ) K i r s c . 265 , 1 580, 1 83 5 oc e l l a r i s ( D e l t o cepha l u s ) Fa l l . 265, 1 835 oc e I I a r i 5 265,

( E rr a5 t unu s ) 1 580, 1 835

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o c u l a t a ( B r u c h omo rp h a ) N e wm . 6 4 3 , 1 843 On c op s i s 3 0 2 opaca ( Ty p h l o c y b a ) Edw . 5 2 , 1 837 Op s i u s 1 8 3 6 rmen i 5 7 1 3 , 9 5 1 , 9 7 3 , 1 1 1 5 , 1 844 o r y z a e Uj i l a p a r v a t a ) r!la t s . 1 1 6 6 , 1 2 02 , 1 8 3 9 o r y z i c o l a ( S o g a t od e s ) 1 4 2 4 , 1 844 p a l u d o s u s ( S c a p h o i d e u s ) Ba l 1 496 p a s c u e l l u s ( A r t h a l d eu s ) F a l o

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p a s c u o r u m ( D e l t o c ep h a l u s )

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1 844 P h a n t i op s i s 8 8 4 , 1 1 2 0 , 1 8 4 6 P h i ep s i u s 1 7 1 2 P h r y n op h y e s 1 7 9 0 , 1 8 4 5 p i c t u ra t u s ( D e l t o c ep h a l u s ) F i eb . 2 6 5 , 1 8 3 5 p i l o s u s ( T i'l c h y c i x i u s l 0 1 . 738 , 1 837 p l e b e i u s ( E x i t i a n u s ) !< i r k . 1 84 5

( N e p h 0 t e t t i x I i( i r k . 1 79 2 , 1 84 5 p l eb e ] a ( E cJ wa r cJ s i a n a ) E d w . 4 9 , 1 83 7 p i ebe] u s ( Eu sc e l i s l Fa l l . 265 , 320, 1 580, 1 835 p o ma r i a ( T y p h l o c y b a ) M c A . 7 5 p opu l ( I d i o c e ru s l L . 3 1 1 p o p u l i ( P op u l i c e r u s ) L . 3 1 1 , 1 834 P r i v e sa 904 , 1 1 2 0 p r op i n q u a ( M e t a d e l p h a x ) F i eb . 1 21 9 , 1 838 p r op i n q u u s ( Ad a r r u s ) F i e b . 1 580 p r op i n q u u s ( A r a l d u s ) F i eb . 1 5 1 8 , 1 56 2 , 1 835 p r u i n o s a ( �1e t c a l f a ) S a y 1 0 9 5 , 1 84 3 p r u i n o s a ( O rme n i s ) S a y 1 0 9 5 , 1 84 3 p s eud o c e l l a r i 5 ( J a s s a rg u s ) F l or 1 580, 1 835 p t e r i d i s ( D i t r op i s ) S p i n . 1 2 1 5 , 1 38 3 , 1 838 p i e be i u s

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no.

1 984

p u n g en s

( R i b a u t od e l p h a x ) R i b . 1 308 , 1 38 3 , 1 839 pu sana ( Py r i I l a ) 9 2 6 , 1 84 1 p u t o n i ( P s a mmo t e t t i x ) T h en 1 580, 1 836 Py r i I i a 9 2 6 , 1 3 3 8 , 1 8 3 3 , 1 8 4 1 q u e r cu s ( T y p h l o c y b a ) F . 4 9 , 5 2 , 54, 1 837 r ep l e t u s ( J a s s a rg u s ) F i eb . 1 58 0 , 1 835 R i b i'l u t o d e l p h a x 1 3 8 3 R i ca n i i d a e 8 , 9 , 7 2 4 , 7 2 9 , 9 0 4 , 1 0 5 1 , 1 067 , 1 1 20 , 1 840 , 1 84 5 L . 4 9 , 1 83 7 52 ru s c i n o n en s i s ( D e l t o c ep h a l u s ) F( i b . 1 5 1 8 , 1 8 3 5 s a c c h a r i c i d a ( P e rk i n s i e l l a ) K i 1 0 , 1 1 65, 1 225, 1 234, 1 23 5 , 1 28 4 , 1 29 2 , 1 29 4 , 1 2 9 6 , 1 4 2 7 , 1 8 3 9 , 1 84 4 , rosae

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1 846 ( 5a c ch a r o s y d n e I 1 269 , 1 27 7 , 1 83 4 , 1 84 3 , 1 544 s a g i t t a r i u s ( A r o c ep h a l u s ) R i b . 1 58 0 , 1 835 s a n g u i n e s c en s ( �� e s o s t e l e s ) K i r . 570 , 1 845 say i ( D e l t o c ep h a l u s ) F . 1 7 2 2 say i ( La t a l u s ) F . 1 6 8 8 , 1 6 9 7 , 1 720, 1 722 S c o I y p op a 1 1 2 0 , 1 8 4 5 Seph ena 1 1 20 , 1 846 5 lO P t e n t r i o n a l i s ( 0 rmen i s ) S p i n . 1 09 5 , 1 543 s e x n o t a t a ( C i c a cJ u l a l F a l l . 1 7 2 2 S i phan t a 7 2 3 , 8 7 3 , 8 8 4 , 9 0 4 , 1 1 20 , 1 846 s a c c h a r i v o r i'l

( Kyb o s ) Fa l l 54 , 1 837 s o c i a l i s ( T u r r u t u s ) F l o r' 1 5 8 0 , 1 836 5 09 a t a l 2 5 5 , 1 1,. 2 1 , 1 8 4 1 Soga t e l l a 1 1 8 9 , 1 2 4 0 , 1 8 4 1 s o n o r a n a ( ii i s t h a r n op h a n t i a ) K i . sma r a g cJ u l a

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83 1 ,

1 355,

s o rd i d u l u s 1 1 50 ,

1 843

( Megad e l p h ax ) S t . 1 20 2 , 1 37 8 , 1 383,

1 4 1 3 , 1 838 s o rd i d u s ( E u s e e l i s ) Z e t t . 2 6 5 , 1 836 s o r d i d u s ( S t r ep t a n u s ) Z e t t 265, 320, 1 836 S t en o c ra n u s 1 2 4 6 , 1 2 9 0 , 1 8 4 1 s t a ch y d e a r u m ( E u p t e ry x ) H a r d . 52, 1 837 s t a c t og a l u s ( O p s i u s ) Fb . 1 5 6 4 s t i g ma t i c a l i s ( I d i o c e r u s ) L w . 3 1 1 , 1 834 S t i r oma 1 3 8 3 S t oba e ra 1 2 6 9 , 1 2 7 5 , 1 8 4 3 s t r a m i n e u s ( Xa n t h od e l p h a x ) S t a l 1 30 8 , 1 839 s t r i a t e l l u s ( L a od e l p h a x ) F a l 1 1 50 , 1 1 66 , 1 1 97 , 1 2 1 1 , 1 3 1 8 , 1 41 3 , 1 4 1 5 , 1 42 1 , 1 8 3 2> s t r i a t u s ( P s a mrno t e t t i x ) L . 3 1 6 , 1 5 1 1 , 1 5 1 8 , 1 5" 7 , 1 55 8 , 1 58 0 , 1 58 5 , 1 69 7 , 1 72 0 , 1 72 2 , 1 836 s u l p h u r a l l a ( E l y ma n a ) Z e t t . 265, 1 83 5 s u n d a i c a ( E mp o a s c a ) B e r g . 6 6 , 1 840 s w e z e y i ( I< e l i s i a ) 1 3 9 5 , 1 8 4 6 s y r t i cl i s ( T a r t e s s u s ) 1 7 8 3 , 1 845 T a r t essus 529 , 1 845 t a u r u s ( Ad a r r u s ) R i b . 1 4 4 6 , 1 5 1 8 , 1 5 1, 7 , 1 5 6 2 , 1 5 8 0 , 1 C: 3 5

1 875

t en e l l a ( E u t e t t i x ) Bak e r 1 688

1 470,

t e n e r r i ma ( R i b a u t i a n a ) H . - 5 . 52, 54, 1 837 T h a mn o t e t t ', x 1 5 2 5 , 1 8 3 6 T h a n a t o cl i e t y a 8 0 6 , 1 8 4 1 t r i f a s c i a t a ( H y m e t t a ) Say 7 1 T r op i d u ch i d a e e , 9 , 7 2 4 , 9 0 7 , 9 1 7 , 1 0 5 1 , 1 1 9 3 , 1 8 1, 0 T y p h l o cy b a 7 5 , 1 0 9 5 Typh l o cyb i n a e 8 , 9 , 4 4 , 4 5 , 1 1 1 , 255, 600, 678, 1 05 1 , 1 1 72 , 1 4 3 6 , 1 494, 1 50 2 , 1 836, 1 ( R i bau t 1 837 un i co l o r ( Ch 484 u r t i ea e ( E u p 1 837 u l mi

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l o r o t e t t i x ) F i t ch t e ry x )

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( N ep h o t e t t i x )

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1 2 48 , 1 46 3 , 1 840 v i r i d i s ( N u m i c i a ) �/:u i r 9 1 7 , 1 83 4 , 1 840 v i t i e n s i s ( P e ,�k i n s i e I I a ) K i r'k . 1 427 v i t i e n s i s ( V a n u a ) K i rk a l d y 1 846 v u I n e ra t a ( E ry t h r o n eu r a ) 75 w a h l b e r g i ( A l e b r a ) G oh eman 5 2 , 1 836 x a n t h o n e u r u s ( S o r h o a n u s ) F i eb . 1 580 , 1 836 zeae ( C i cadu l i na ) Ch i na 1 6 1 5 , 1 839 Zyg i na 5 2 , 5 4 , 1 837

I N D E X T O THE PARAS I T ES AND PR EDATORS D F D RY I N I DS

T h i s i n d e x i n c l u d e s f a m i l i e s , s u b f a rn i I i es , k n o Vv n p a r a s i t e s a n d p re d a t or�s o f d r y i n i d s . a b n o rm i s ( C e r a p h r o n ) 1 427 , 1 83 1

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g e n e r a a n d spec i e s of t h e

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1 876

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g ra c i I i p e s

1 984

( A ch o r o l op h u s ) R .

P e rk i n s 1 3 7 8 h a l i d a y i ( I s ma r u s ) F o r s t e r 1 8 3 1 h a wa i i e n s i s ( E e h t h r o g o n a t op u s ) R . P e rk i n s 1 8 3 3 h a wa i i e n s i s ( H y p e r g o n a t op u s ) R . P e rk i n s 1 1 6 6 , 1 3 9 6 , 1 8 3 3 H a zmb u r k i a H o f f e r 1 8 3 2 H e l e g o n a t op u s R . P e rk i n s 1 8 3 2 , 1 83 3 , 1 834 h em i p t e ru s ( Hy p e rg on a t opu s ) T i mb e r l a k e 1 8 3 3 H y p e r g on a t op u s T i mb . 1 8 3 2 , 1 8 3 3 I s ma r u s H a l i d a y 1 8 3 1 j a v a n u s ( C h e i l on eu ru s ) R . P e rk . 80 6 , 1 8 3 3

1 83 1 ,

C h e i l o n e u r u s We s t w o o d 9 2 6 , 1 83 1 , 1 83 2 , 1 83 3 , 1 834 c h l o r od ry i n i ( C h e i l o n e u r u s ) R . P e rk i n s 1 8 3 2 C h ry s o p o p h a g u s A s h m e a d 1 8 3 1 c i t r i p e s ( H e I egona t opu s ) E r d ii s 1 8 3 2 C r i s t a t i t h o r a x G i ra u l t 1 8 3 1 d e s t ru e t o r ( E c h t h rod ry i n u s ) R . P e rk i n s 1 8 3 2 D i ap r i i da e 1 83 1 d i mo r p h a ( H e l e g o n a t op u s ) H o f 1 832 d o r s i g e r ( I sma r u s ) C u r t . 1 8 3 E ch t h r o d ry i n u s R . P e r k i n s 1 83 2 , 1 83 4 E e h t h r o g o n a t op u s R . P e r k i n s 1 83 2 , 1 833 E n ey r t i d a e 1 2 3 5 , 1 8 3 1 E ry t h r a e i d a e 1 3 7 8 e x i m i a ( C h a l c e r i n y s ) R . P e rk 1 832 ex i t i o s u s ( E ch t h r o g o n a t op u s ) R . P e rk i n s 1 8 3 2 f a c i a l i s ( H e I e g o n a t op u s ) S z e 1 832

no . 37,

g o n a t op od i s ( C h e i l o n e u r u s ) R . P e rk i n s 1 2 3 5 , 1 8 3 2

1 832 a m e r i c a n u s ( C h e i I on eu ru s ) R . P e rk . 1 4 2 7 , 1 8 3 2 , 1 8 3 3 a u s t r a l i a e ( C h e i I o n e u ru s ) R . P e rk . 1 8 3 2 a u s t ra l i a e ( Sa r on o t u m ) R . P e rk . 1 8 3 2 Be l y t i na e 1 83 1 B l a t t i e i d a A s h me a d 1 8 3 1 b r u n n e i p e s ( H y p e r g o n a t op u s ) T i mb e r l " k e 1 3 9 6 , 1 8 3 3 C e raph ron 1 8 3 1 C e raph ron i da e 1 8 3 1 Cha l c e r i ny s R . 1 832

I nst . ,

Ma s e n c y r t u s H o f f e r 1 8 3 2 M e t a ch e i I o n e u r u s H 0 f f e r 1 8 3 1 mo l o k a i e n s i s ( H y p e r g on a t op u s ) A shmead

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1 833

mo l o k a i e n s i s ( M i e r o t e r y s ) A s h m . 1 833 n i k o l sk a y a e ( H e l e g o n a t op u s ) Hof f e r 1 832 o a h u e n s i s ( H y p e rg on a t op u s ) T i mb e r l a k e 1 8 3 3 O o e n cy r t u s 1 7 5 2 , 1 8 3 4 p a e h y e ep h a l u s ( E c h t h r o g o n a t op u s R . P e rk i n s 1 8 3 3 P a l u d en ey r t u s H o f f e r 1 8 3 2 p o n oma r e n k o i ( H e I e g o n a t op u s ) T r j a p i t zy n 1 8 3 2 p s e u d o p h a n e s ( H e l e g on a t o p u s ) R . P e rk . 1 2 9 2 , 1 8 3 3 , 1 8 3 4 p u l c h r i e o r n i s ( C h e i l on eu r u s ) H a y a t a n d V e rma 1 8 3 4 q u a d r i c o l o r ( C h e i I on e u r u s ) G i r . 1 832, 1 833 q u ad r i c o l o r ( C r i s t a t i t h o r ax ) G i rau l t 1 8 3 3 r a S n i t z y n i ( H e I e g o n a t op u s ) T r j a p i t zy n 1 8 3 2 r u g u l o s u s ( I s ma ru s ) F o r . 1 8 3 1 Sa r on 0 t u m R . P e rk i n s 1 8 3 1 S e h ed i o i d e s Me r e e t 1 8 3 2

1 877

I n dex

sw e z e y i ( C h e i l o n e u r u s ) A s h m . 1 09 5 , 1 833

v u l c an u s ( H y p e rg o n a t opu s ) 1 396 , 1 833

T i mb .

I N D E X T D D RY I N I DS

T h i s i n d e x i n c l u de s s u b f a m i I i e s , g e n e r a a n d s p ec i e s of d r y i n i d s . An a s t e r i s k m a r k s a s y n o n y m , m i ss p e l l i n g , i n correc t i de n t i f i c a t i o n , o r nomen

n u du m . abdomi na l i s 1 798

( Ch e l ogy n u s ) O g l

a b e r r a n s ( A l p h a d r y i n u s ) B en . 980 " a b e r ra n s ( D ry i n u s ) B e n o i 980 a b j e c t u s ( G o n a t op u s ) D l m i 1 65 5 a b r u p t u m ( An t e o n ) O l m i 4 0 4 a c h t e rb e rg i ( D e i n o d ry i n u s ) 0 1 m i 1 74 A c r od on t o c h e l y s C u r r a d o 1 0 5 2 , 1 1 72 , 1 800 a c t u o su s ( D e i n od r y i n u s ) O l m i 211 a c u l e a t u m ( An t e o n ) O l m i 5 4 5 a c u m i n a t u s ( Ag on a t o p o i d e s ) O l m i 1 32 4 " a cu t i c o l l i s ( D ry i n u s ) K i e f f . 1 098 a c u t i co l l i s ( N eod ry i n u s ) K i e f f e r 1 09 8 " a c u t i c o l l i s ( P s i l od r y i n u s ) K i ef f e r 1 06 5 , 1 098 a c u t u s ( P s e u d og o n a t op u s ) D l m i 1 230 Ad ry i n u s D l m i 1 0 5 4 , 1 1 2 6 a eg y p t i a c u s ( G on a t op u s ) O l m i 1 533 a eg y p t i a c u s ( T e t r od on t o c h e I y s ) O l mi 1 446 " a eq u a l i s ( An t e o n ) K i e f f . 2 5 9 a eq u a l i s ( D e i n od ry i n u s ) D l m i 240 " a eq u a l i s ( P s i l an t e on ) 2 5 3 " a e r i a s ( A n t e o n ) Wa l k e r 1 1 5

':' a e r i a s ( D r y i n u s ) Wa I k e r . 1 1 5 a e r i a s ( M e t a n t e o n ) Wa l k e r 1 1 3 , 1 15 a e t h i op i c u s ( G on a t o p u s ) O l m i 1619 a e t h i op i cu s ( Me s od ry i n u s ) O l � i 1 008 , 1 83 9 a f e r ( D ry i n u s ) O l m i 7 8 7 , 1 8 4 0 a f e r ( E ch t h r o d e l p h a x ) O l m i 1 1 5 5 " a f e r ( P a rad ry i n u s ) O l m i 7 8 8 a f e r ( T e t r o d on t o ch e l y s ) O l m i 1 458 " a f f i n i s ( A p h e l op u s ) A s h me a d 7 0 " a f f i n i s ( G o n a t op u s ) F e n t . 1 7 1 7 a f f i n i s ( P rod ry i nu s ) A r l e 1 79 8 " a f r i c a n u s ( An t a p h e l op u s ) B e n . 1 79 8 a f r i c a n u s ( Ap h e l o p u s ) B e n o i t 1 79 8 , 1 8 1 2 a f rum ( An t e on ) O l m i 3 7 5 a g i l e ( An t e on ) O l m i 3 5 4 a g i l i s ( G o n a t op u s ) D l m i 1 7 3 7 A g o n a t op o i d e s R . P e r k i n s 1 3 1 0 * A g o n a t op u s R . P e r k i n s 1 0 5 4 , 1 05 5 , 1 50 0 a g r op y r u s ( G on a t o p u s ) F en t o n 1 0 5 5 , 1 69 5 , 1 8 4 2 " a g r opy r u s ( N e o g o n a t opu s ) F en t 1 695 a l a t u s ( D ry i n u s ) C r e s s o n 8 2 8 , 1 84 3 ':' a l a t u s ( G o n a t op u s ) C r e s s o n 8 2 8 ':' a l a t u s ( M e s o d ry i n u s ) C r e s s o n 828 " a l a t u s ( P r o d ry i n u s ) C r e s s o n 828

1 878

Ivlem . A m e r .

Ent.

a l b i c ox i s ( A n t e o n ) K i e f . 1 7 9 9 ':' a l b i d i c o l l e ( A n t e o n ) K i e f f e r 313 ':' a l b i d i c o l l i s ( A n t e o n ) K i e f f e r 313 ':' a l b i d o c o l l e ( An t e o n ) K i e f f e r 313 " a l b i d o c o l l i s ( C h e l ogy n u s ) 3 1 3 ':' a l b i d u s ( G on a t o f.l u s ) K i e f f e r 1 560 ':' a l b i d u s ( 01 e o g o n a t op u s ) K i e f . 1 5 60 a l b i p enn i s ( Lab e o ) K i e f . 1 79 9 ':' a l b i p e n n i s ( L a b e r i u s ) K i e f f e r 1 799 * a l b i p e s ( A p h e l op u s ) K i e f . 4 7 " a l b i p e s ( Ap h e l op u s ) R a t z eb u r g 47 ':' a l b i p e s ( C e r a p h r o n ) R a t z eb u r g 47 ':' a l b i s c a p u s ( C h a l c o g on a t op u s ) D od d 1 7 7 9 a l b i t a r s e ( An t e o n ) C a me r on 5 1 2 ':' a l b i t a r s i s ( D r y i n u s ) C a m e r o n 51 2 a l b i t a r s i s ( Lab eo ) K i e f f e r 1 79 9 , 1 8 00 * a l b i t a r s i s ( Lab e r i u s ) K i e f f e r 1 799 " a l b i t a r s i s ( Me s o d r y i n u s ) C a me r o n 5 1 2 " a I b 0 l i n ea t u s ( C h a l c og o n a t op u s ) K i e f f e r 1 527 a l b o l i n ea t u s ( G on a t op u s ) K i ef f e r 1 527 ':' a l b o l i n e a t u s ( Ma d e c a g o n a t o p u s ) K . 1 5 2 7 ':' a l b o l i n e a t u s ( P l a t y g o n a t op u s ) K i ef f e r 1 527 " a I b oma r g i n a t u s ( G o n a t o p u s ) C a me r o n 1 4 8 5 a l b oma r g i n a t u s ( T r i c h o g o n a t op u s ) C a me r on 1 4 8 5 a l b o p i c t u s ( A p h e l op u s ) A s h m e a d 7 3 , 1 9 , 84, 1 8 42

I nst . ,

no .

37,

1 984

a l b op i c t u s ( O e i n od ry i n u s ) O l m i 1 83 ':' a l b o s i g n a t u s ( G on a t o p u s ) K i e f 1 209 a l b o s i gn a t u s ( Lab e o ) K i e f f e r 1 800 * a l b o s i gna t u s ( La b e r i u s ) K i ef f e r 1 800 a l b o s i g n a t u s ( 01 e od r y i n u s ) D l m i 1 101 a l b o s i g n a t u s ( P s e u d o g o n a t op u s ) K i e f f e r 1 20 9 , 1 2 1 7 , 1 8 3 8 , 1 839 a l b r e c h t i ( R i ch a rd s i d ry i n u s ) D l mi 909 a l exan d ra e ( D e i n od r y i n u s ) D l m i 230 ':' A l l a n t e o n K i e f f e r 2 8 7 A I I odry i nus K i ef f e r 1 8 1 7 ':' A l l o g o n a t op u s H a u p t 1 5 0 0 ;' a l o r u s ( A n t a e o n ) V,a l k e r 3 1 7 ':' a l o r u s ( A n t e o n ) " ,a l k e r 3 1 7 ':' a l o r u s ( C h e l og y n u s ) lia l k e r 3 1 7 ':' a l o r u s ( D r y i n u s ) V'la l k e r 3 1 7 A l phad ry i nu s D l m i 9 7 3 a l p i n u s ( O i c on d y l u s ) G o u r l a y 1 40 4 , 1 8 4 6 * a l p i n u s ( N e o g o n a t op u s ) G o u r l a y 1 404 ':' a l t i c o l a ( D r y i n u s ) C a me r o n 1 1 07 ':' a l t i c o l a ( L e s t od r y i n u s ) B e n o i t 758 ':' a l t i c o l a ( N e o d r y i n u s ) C a me r o n 1 1 07 ':' a l t i c o l a ( P r od r y i n u s ) C a me r o n 1 1 07 a l t i c o l u s ( D ry i n u s ) B e n o i t 7 5 8 * a l u t a c eu s ( Ch e l o g y n u s ) R i ch a rd s 3 3 8 a l v a r en g an u s ( Ap h e l op u s ) D l m i 80 a l v a r e n g a n u s ( D ry i n u s ) D l m i 8 5 7 a mb o n i ( N e o d r y i n u s ) D l m i 1 1 2 3 ame r i c a n u m ( P r e n a n t e on ) K i e f f e r 1 80 1 a me r i c a n u s ( A c r od o n t o c h e l y s )

1 879

I ndex

F en t on

1 1 82,

1 80 0 ,

1 842

C eb a l l o s

1 50 0 ,

1 60 1

* a me r i c a n u s ( A p h e l o p u s ) A shmead 4 7 5

a n n u l i c o r n e ( An t e o n ) B r u e s 5 1 1 ':' a n n u l i c o rn i s ( An t e on ) B r u e s

a me r i c a n u s ( D i c o n d y l u s ) R . C . L . P e rk i n s 1 3 8 8 , 1 8 4 3 a me r i c a n u s ( D r y i n u s ) A s h m e a d 826 a me r i c a n u s ( E p i g o n a t o p u s ) F en t on 1 1 8 2 * a me r i c a n u s ( H a p l o g on a t op u s ) R . C . L . P e rk i n s 1 3 8 8

51 1 * a n oma l a ( G o n a t o p u s ) R . P e r k i n s 1 473 a n oma l u s ( T e t r o d o n t o c h e l y s ) R . P e rk i n s 1 4 7 3 ':' An t a e o n H a l i d a y 2 8 6 " An t ap h e l opu s B en o i t 44 An t eon J u r i n e 1 1 0 , 286 , 2 8 7 , 288 , 1 83 1 " A n t eon e l l a Dodd 6 1 2 An t e o n i n a e R . C . L . P e rk i n s 1 1 0 a n t h e r e on ( A n t e o n ) B e n o i t 3 8 1 a n t i l l ea n u s ( D r y i n u s ) E v a n s 8 6 0 ':' a n t i l l e a n u s ( Me s o d r y i n u s ) E v an s 8 6 1 a n t i q u u s ( D r y i n u s ) P o n o ma r e n k o 7 5 1 , 1 8 30

a me r l c a n u s ( Me s o d r y i n u s ) A shmead 8 2 6 " a me r i c a n u s ( My s t r o p h o r u s ) A s h me a d 8 2 6 a me r i c a n u s ( P r e n a n t e o n ) K i e f . 1 80 1 * a me r i c a n u s ( P r o d ry i n u s ) A shmead 8 2 6 * a me r i c a n u s ( P s eu d o g on a t op u s ) R . C . L . P e rk i n s 1 2 6 3 a m o en u s ( D e i n od ry i n u s ) O l m i 1 85 a m o en u s ( P e r o d ry i n u s ) R . C . L . P e rk i n s 1 0 3 3 , 1 0 3 4 , 1 8 4 3 * amp h i s c e p a e ( H e s p e r o d r y i n u s ) R . C . L . P e rk i n s 8 2 8 " a mp u l i c i f o r m i s ( C a mp y l o n y x ) We s t w o od 7 2 8 , 7 3 5 , 7 3 6 * amp u l i c i f o r m i s ( L e s t o d r y i n u s ) Turner 937 * a mp u l i c i f o r m i s ( M e s od r y i n u s ) Turner 937 a mp u l i c i f o rm i s ( T r i d r y i n u s ) Turner 937 and i nu s ( T r i d ry i nu s ) O l mi 967 a n d o i ( P a ch y g on a t op u s ) E s a k i a n d H a s h i mo t o 1 8 0 1 *ang u s t i co l l i s ( Ch e l ogynu s ) B e r l and 329 a n g u s t i c o i l i s ( Ch e l o g y n u s ) Og I ob l i n 1 8 0 1 * a n k a r a t r u s ( C h a l c o g on a t op u s ) C eb a l l o s 1 6 0 1 a n k a r a t r u s ( G on a t o p u s ) C eb a l l o s 1 6 0 1 '� a n k a r a t r u s ( Ma d e c a g on a t op u s )

a n t i q u u s ( L a b e r i u s ) P o n oma r e n k o 751 a n t i q u u s ( I� e od r y i n u s ) B en o i t 1 06 9 a n x i u m ( An t e o n ) O l m i 5 3 6 A p h e l op i n a e R . C . L . P e r k i n s 4 3 Ap h e l o p u s D a l ma n 4 4 , 1 8 3 1 a p i c a l i s ( G o n a t o p u s ) C a me r on 1 750 a p i c a l i s ( H a p l o g o n a t op u s ) R . C . L . P e rk i n s 1 4 0 9 , 1 4 1 7 , 1 4 1 8 , 1 425, 1 427, 1 83 8 , 1 8 4 1 , 1 8 46 Apod ry i n i n a e D l m i 1 7 9 6 Apod ry i n u s D l m i 1 7 9 6 Ap t e r o d ry i n u s R . C . L . P e rk i n s 1 34 2 ':' a p t e r u s ( An t e o n ) D od d 5 7 1 a r a b i c u m ( An t e o n ) D l m i 3 8 7 a r c u a t u m ( An t e o n ) K i e f f e r 2 8 9 , 299 , 3 0 5 , 3 1 2 , 1 8 34 a r c u a t u s ( An t eon ) K i e f f e r 3 0 5 a r e o l a t u s ( Ap h e l op u s ) K i e f f e r 1 80 1 " a r e o l a t u s ( C h a l c o g o n a t op u s ) F en t on 1 7 0 0 a r e o l a t u s ( D e i n od ry i n u s )

1 880

Mem .

Amer .

Ent.

P on oma r e n k o 1 2 1 , 1 8 3 0 a r e o l a t u s ( D ry i n u s ) O l m i 894 " a r e o l a t u s ( E l e c t r o d ry i n u s ) P on oma r e n k o 1 1 9 , 1 2 1 a r e o l a t u s ( G on a t op u s ) F en t o n 1 700 a rg en t i n u s ( D r y i n u s ) O l m i 8 4 6 * a rg y r i a s ( C h a l c og on a t op u s ) R . C . L . P e rk i n s 1 7 2 4 a r g y r i a s ( G o n a t op u s ) R . C . L . P e rk i n s 1 7 2 4 a r i z o n en s e ( A n t e on ) R . C . L . P e rk i n s 4 5 5 " a r i z o n e n s i s ( An t e on ) R . C . L P e rk i n s 4 5 5

•

* a r i z on i c a ( B o c c h u s ) R . C . L . P e rk i n s 6 4 5 " a r i z o n i c a ( E u k o eb e l e i a ) R . C . L . P e rk i n s 6 4 5 * a r i z o n i c a ( Ph o rb a s ) R . C . L . P e rk i n s 6 4 5 * a r i z on i c u s ( Ap h e l op u s ) R . C . L . P e rk i n s 7 3 a r i z on i cu s ( B o c c h u s ) R . C . L . P e rk i n s 6 4 5 , 1 8 4 3 " a r i z on i c u s ( H e s p e r od r y i n u s ) R . P e rk i n s 7 2 8 , 8 2 8 "' a r i z o n i c u s ( N e o d r y i n u s ) R . C . L . P e rk i n s 1 0 9 1 , 1 0 9 4 a r i z o n i c u s ( P s e u d o g o n a t op u s ) R . P e rk i n s 1 26 6 , 1 8 4 3 * a r i z on i cu s ( P s i I o d r y i n u s ) R . P e rk i n s 1 0 9 3 a r l e i ( G on a t op u s ) D l m i 1 7 6 4 a rma t u s ( G on a t o p u s ) O l m i 1 6 2 3 a r n a u d i ( A p t e r o d ry i n u s ) O l m i 1 36 2 * a r n o l d i i ( P a ch y g on a t o p u s ) P o n oma r e n k o 1 4 3 8 , 1 4 3 9 a s h me a d i ( G o n a t o p u s ) K i e f f e r 2 8 , 1 7 1 5 , 1 7 1 7 , 1 842 a s i a t i c a ( D o n i s t h o rp i n a ) D l m i 1 30 9 a s i a t i c u s ( A l p h a d ry i n u s ) D l m i 9 7 3 , 982 a s i a t i cu s ( Ap t e r o d r y i n u s ) D l m i 1 350

I nst . ,

n o . 37 ,

as i at i cu5 1 31

1 984

( D e i n o d ry i n u s ) D l m i

a s i a t i cu s ( T h a u ma t od r y i n u s ) D l mi 697 a s p e r ( D e i n o d ry i n u s ) O l m i 2 2 6 a t e r ( G on a t opu s ) D l m i 1 5 3 8 a t e r r i mu s ( A n t e o n ) H e l l e n 1 8 0 1 a t e r r i mu s ( D ry i n u s ) D od d 8 8 8 a t e r r i mu s ( Me s od ry i n u s ) O l m i 1 03 1 * a t e r r i mu s ( P a r a d ry i n u s ) D o d d 888 a t l a n t i c u s ( D e i n od ry i n u s ) D l m i 1 72 a t l a n t i cu s ( G o n a t op u s ) D l m i 1 530 a t l a n t i c u s ( M i r o d ry i n u s ) D l m i 672 a t r a t u s ( Ap h e l op u s ) D a l ma n 5 0 , 1 83 6 , 1 83 7 * a t r a t u s ( D r y i n u s ) D a l ma n 5 0 a t r a t u s ( H a p l o g o n a t op u s ) E s a k i a n d H a sh i mo t o 1 4 1 3 , 1 8 3 8 * a t r i c ep s ( B o c ch u s ) B r u e s 1 3 3 * a t r i c ep s ( C h e l og y n u s ) B r u e s 1 36 " a t r i c ep s ( D e i n od ry i n u s ) 1 3 6 " a t r i v e n t r i s ( An t e on ) C r e s s on 1 36 a t r i v e n t r i s ( D e i n od ry i n u s ) C r e s s on 1 3 3 , 1 3 6 a t r i v e n t r i s ( D r y i n u s ) C r e s s on 1 33 a t t en u a t u s ( G o n a t op u s ) D l m i 1 64 1 * a u d a x ( H e sp e r o d r y i nu s ) R . C . L . P e rk i n s 8 2 8 a u d a x ( T e t r od o n t o ch e l y s ) D l m i 1 44 7 a u g u s t a e ( P s e u d o g on a t op u s ) C u r ra d o a n d D l m i 1 2 1 2 , 1 8 38 au l i cu m ( An t eon ) O l m i 5 3 3 au s t r a l e ( An t e o n ) O l m i 5 3 8 a u s t ra l i a e ( B o c ch u s ) D l m i 6 5 6 * a u s t ra l i a e ( D i condy l u s ) R . C . L . P e rk i n s 1 7 9 0

I n dex

au s t ra l i a e

( G on a t op u s ) R . C . L .

P e rk i n s a u s t ra l i a n u s

1 79 0 , 1 84 5 ( D ry i n u s ) O l m i

900 * a u s t r a l i s ( An t e on ) D od d 5 8 5 a u s t r a l i s ( Ap h e l op u s ) O l m i 8 3 * au s t ra l i s ( E u c a mp t on y x ) C eb a l l o s 1 5 9 3 a u s t r a l i s ( G on a t op u s ) C eb a l l o s 1 59 3 a u t ox en ob i u s ( P s eu d og on a t o p u s ) R . P e rk i n s 1 2 7 3 , 1 8 4 3 a u t u mn a l i s ( G o n a t op u s ) O l m i 1 7 70 a u t u mn a l i s ( P e r od ry i n u s ) O l m i 1 04 1 * A v od ry i n u s P on oma r e n k o 9 0 7 * a z o r u s ( A n t e on ) Wa l k e r 3 1 7 * b a e t i c a ( Me r c e t i a ) C eb a l l o s 1 1 42 b a e t i c u s ( E ch t h r od e l p h a x ) C eb a l l o s 1 1 4 2 * b a k e r i ( An t e on ) K i e f f e r 2 7 8 b a k e r i ( L on c h od ry i n u s ) K i e f f e r 278 * b ak e r i ( P r e n a n t e on ) K i e f f e r 278 b a l l i ( N e od ry i n u s ) O l m i 1 0 9 5 b a l t i cu s ( A l p h a d ry i n u s ) O l m i 978, 1 830 b a rb a r a ( C r o v e t t i a ) O l m i 8 4 , 89 b a rb a ru s ( Me s od ry i n u s ) O l m i 1018 b a r b a t e l l u s ( G on a t op u s ) R i ch a r d s 1 5 3 8 * b a r b a t u m ( An t e o n ) C h i t t y 2 9 5 * b a rb a t u s ( A n t a e on ) C h i t t y 2 9 5 b a rb i e r i ( G on a t o p u s ) O l m i 1 5 8 5 b a r t l e t t i ( G on a t op u s ) O l m i 1 0 54 , 1 75 1 , 1 8 3 4 , 1 844 * b a s a l e ( P r e n a n t e on ) D a l . 2 6 1 * b a s a l i s ( G on a t op u s ) D a l . 2 5 8 * b a sa l i s ( P r e n a n t e on ) D a l ma n 26 1 b a s i I e w s k y i ( D r y i n u s ) B en o i t 774, 775

1 88 1

b a u en s e ( A n t eon ) O l m i 4 4 6 * b e a u m o n t i ( An t a e on ) C h i t t y 2 6 0 b ek i I y a n u s ( G on a t op u s ) B e n o i t 1612 * b ek i l y a n u s ( N e o g o n a t op u s ) B en . 1 61 2 b ek i l y e n s i s ( B o c c h o i d e s ) B e n . 60 2 , 6 0 4 b e l i z en s i s ( D ry i n u s ) O l m i 8 5 8 b e l l a t o r ( A n t e on ) O l m i 5 6 8 b e l l i c o s u s ( Ad ry i n u s ) B en . 1 1 3 2 * b e l l i c o su s ( N e od ry i n u s ) B e n o i t 1 1 32 b e l l o t t i i ( A p t e r od ry i n u s ) O l m i 1 369 b e n g a l e n s e ( An t e on ) O l m i 4 2 8 b en g a l en s i 5 ( Ag o n a t op o i d e s ) O l m i 1 3 33 b e n i a n u s ( D e i n od ry i n u s ) O l m i 243 b en o i t i ( D e i n od r y i n u s ) O l m i 1 2 9 b e n o i t i ( G o n a t op u s ) O l m i 1 6 3 1 * b e n s o n i ( An t e on ) R i c h a r d s 3 0 5 * b e r l a n d i ( C h e l ogy n u s ) R i c h a rd s 329 b e r l a n d i ( C h e l o t h e l i u s ) B e r n a rd 609 b e r l a n d i ( D ry i n u s ) B e n o i t 1 8 0 2 b e r n a r d i ( G on a t op u s ) P i ca rd 1 56 2 , 1 8 3 6 * b e r n a r d i ( P l a t y g o n a t op u s ) P i c . 1 562 b i ak i ( D r y i n u s ) O l m i 8 9 0 B i a p h e l op i n a e O l m i 9 4 B i a p h e l opu s O l m i 9 4 , 9 5 * b i c o l o r ( A n t e o n e l l a ) D od d 6 6 0 b i c o l o r ( A p h e l op u s ) F e n t on 7 1 , 1 842 b i c o l o r ( D e i n od r y i n u s ) O l m i a n d C u r ra d o 1 6 3 b i c o l o r ( D i c on d y l u s ) H a l i d ay 1 37 5 , 1 378, 1 37 9 , 1 380, 1 838, 1 839 * b i c o l o r ( D r y i n u s ) Ha l i day 1 3 7 8 1715 * b i c o l o r ( E c h t h r od e l p h a x ) E s ak i a n d H a sh i mo t o 1 1 6 1 , 1 1 6 3

1 882

Mem . Amer . E n t .

*b i c o l o r ( G o n a t o p u s ) A s h m e a d 1 71 5, 1 7 1 7 " b i c o l o r ( G on a t opu s ) H a l i day 1 379, 1 820 * b i c o l o r ( L on ch od r y i n u s ) O l m i a n d C u r ra d o 1 6 3 b i c o l o r ( P e r od ry i n u s ) O l m i 1 04 5 " b i d e n s ( L e s t od ry i n u s ) H a u p t 744 * b i f a r i u s ( G on a t op u s ) K i e f f e r 1 379 " b i f a r i u s ( H a p l o g on a t o p u s ) K i e f f e r 1 3 7 9 , 1 3 8 1) " b i f a r i u s ( P I a t y g on a t op u s ) K i ef f er 1 37 9 " b i f a r i u s ( T r i g o n a t opu s ) K i e f 1 3 7 5 , 1 37 9 b i f a s c i a t u m ( An t e o n ) K i e f f e r 1 802 * b i f a s c i a t u s ( An t eon ) K . 1 8 0 2 ':' b i f a s c i a t u s ( C a l l i a n t e on ) K i e f f e r 1 80 2 * b i f a s c i a t u s ( C h e l ogy n u s ) K i ef f er 1 802 b i f a s c i a t u s ( D ry i n u s ) Say 1 8 0 2 b i f a s c i a t u s ( E ch t h r od e l p h a x ) R . P e rk i n s 1 1 6 9 , 1 8 4 6 ;' b i f a s c i a t u s ( G o n a t op u s ) K i e f . 1 54 1 , 1 543 * b i f a s c i a t u s ( N eogona t op u s ) K i e f f e r 1 54 3 " b i f a s c i a t u s ( P rod ry i n u s ) S a y 1 80 2 b i I i n e a t u s ( G on a t op u s ) K i e f f e r 1 5 1 3 , 1 83 5 , 1 836 " b i I i n e a t u s ( N e o g on a t op u s ) K i ef f er 1 5 1 3 b i l ob o i d e s ( D e i n o d r y i n u s ) O l m i 245 b i l ob u s ( D e i n od ry i n u s ) F en t o n 1 91 b i n i ( B oc chu s ) O l m i 6 2 9 b i rma n u s ( A p h e l op u s ) O l m i 6 9 * b i r ma n u s ( Me g a g o n a t o p u s ) O l m i and C u r ra d o 1 5 0 1 , 1 6 5 2 b i r o i ( P r i o r a n t e on ) O l m i 5 9 1 ,

I nst . ,

no.

37,

1 984

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( A n t eon ) O l m i

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" b i s u l c a t a ( Me r c e t i a ) B e n . 7 5 4 b i su l c a t u s ( D ry i n u s ) B en . 7 5 4 B o c ch i n a e R i c h a rd s 5 9 9 * B o c ch i n i 5 9 9 B o c ch o i d e s B en o i t 60 1 , 6 0 2 B o c c h u s A sh m e a d 6 1 1 * b og o t en s i s ( T r i c h og o n a t op u s ) K i ef f er 1 485 " b o l i v a r i ( G o n a t op u s ) C eb . 1 4 4 2 0 0 l i v i a n u m ( An t e on ) O l m i 5 1 8 b o l i v i a n u s ( D e i n o d ry i n u s ) O l m i 217 b o l i v i a n u s ( G on a t o p u s ) O l m i 1 768 b o l i v i a n u s ( P e r od ry i n u s ) O l m i 1 04 1 " b o r e a l i s ( C y r t o g o n a t op o i d e s ) P on om . 1 5 0 0 , 1 5 0 9 ':' b o r e a l i s ( G o n a t op u s ) J . S a h l . 1 57 7 b o r e a l i s ( P en t a g on a t o p u s ) O l m i 1 428 " b o r e a l i s ( P r e n a n t e on ) H e l l e n 1 829 b o r n e a n u m ( An t e o n ) O l m i 4 1 6 b o r n e a n u s ( A g on a t op o i d e s ) O l m i 1 333 b o rn e a n u s ( A ph e l op u s ) O l m i 6 5 b o u c e k i ( A c r od o n t o ch e l y s ) C u r r o 1 1 7 2 , 1 1 74 b o u c ek i ( An t eon ) O l m i 5 6 4 b o u c ek i ( B o c c h u s ) O l m i 6 6 1 b o u c ek i ( D r y i n u s ) O l m i 8 8 0 b o u g a i n v i l l e i ( A n t e on ) O l m i 5 5 5 b r a c h y c e ru m ( A n t e on ) D a l ma n 2 8 9 , 300, 1 8 3 4 ';' b r a c h y c e r u s ( An t e on ) D a I . 3 0 0 " b ra c h y c e ru s ( D r y i n u s ) D a l ma n 300 ':' b ra c h y c e ru s ( D r y i n u s ) K i e f f e r 1 1 1 1 b r a c h y c e r u s ( N e od ry i n u s ) K i e f f . 1 1 1 1

1 883

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*b r a c h y c e r u s

( P rod ry i n u s )

K i ef f er 1 06 5 , 1 1 1 1 b r a c h y p t e r u m ( An t e on ) O l m i 353 b r a s i l i a n a ( C r ov e t t i a ) O l m i 93 b ra s i I i a n u s ( A l p h a d ry i n u s ) 0 1 m i 992 b ra s i l i en s i s ( C h e l ogynu s ) A r l e 1 80 2 b r a s i i i en s i s ( M e t a g on a t o p u s ) Og l . 1 5 0 0 , 1 7 3 5

b r u e s i ( A l p h a d ry i n u s ) O l m i 1 830

975,

b r u n n e i c o l l i s ( G on a t o p u s ) R i ch a r d s 1 5 7 0 * b r u n n e i c o l l i s ( N e o g o n a t op u s ) R i ch a rd s 1 5 7 0 * b r u n n e i p e s ( X e n a n t e on ) B e r l a n d 295 " b ru n n e s c e n s ( N e og o n a t o p u s ) R . P e rk i n s 1 7 1 7 b u rg e on i ( D ry i n u s ) B e n o i t 7 8 6 * b u y s s o n i ( P l a t y g o n a t op u s ) B e r l and 1 5 54 c a f e r ( A g o n a t op o i d e s ) O l m i 1 325 c a l ed on i a n u m ( A n t e on ) O l m i 5 5 1 c a l ed on i c u s ( Ap h e l o p u s ) � i m ; 83 ca l i d u s ( A g o n a t op o i d e s ) O l m i 1 339 c a l ; f o r n i cu s ( Ap t e r od r y i n u s )

b raun s i ( G y n o c h e l y s ) B ru e s 1 79 3 " b r ev i c o l l i s ( A n t e on ) K i e f f e r 300 " b r ev i c o rn e ( An t e on ) D a l ma n 2 9 5 , 299 " ' b r e v i c o rn i s ( A n t eon ) D a I ma n 295 * b r e v i c o rn i s ( D r y i n u s ) D a l ma n 295 " b r ev i c o r n i s ( G o n a t op u s ) D a l ma n 2 9 4 , 2 9 5 * b r ev i c o r n i s ( H a p l o g o n a t op u s ) R . P e rk ; n s 1 4 1 8 b r e v i f i i i s ( A n t e on ) K i e f f e r 1 802 " ' b r e v i f i l i s ( C h e l ogy n u s ) K i e f

A shmead 1 3 5 7 "' c a l ; f o rn i cu s ( G on a t op u s ) A sh m . 1 357 C a l l i a n t e on K i e f f e r 1 8 0 2 " came l i n u s ( 0 ; c ondy I u s ) K i e f f . 1 1 99 "' c a m e l i n u s ( G o n a t o p u s ) K i e f f e r 1 1 97

1 802 '� b r ev i f o r c e p s ( C y r t o g o n a t op u s ) K i ef f e r 1 500, 1 73 5 b r ev i f o r c e p s ( G o n a t op u s ) K i e f . 1 735 "' b r ev i f o r c e p s ( P a c h y g on a t o p u s ) K i ef f . 1 735 "'b r e v i v e n t r' a l i s ( A n t a e o n ) Ch i t t Y 3 1 7 "' b r i t t a n i cu s ( Me s o d ry i n u s ) R i ch a r d s 1 0 0 5 b r o ok s i ( G on a t opu s ) O l m i 1 7 2 8 b r own i ( D r y i n u s ) A s h m e a d 8 1 7 , 1 84 1 "' b r own i ( L e s t od r y i n u s ) A s h m e a d 818 b r u c h i ( T h a u ma t o d r y i n u s ) D e Sa n t i s a n d V . S a r m . 7 1 4

came I i n u s ( P s eu d o g o n a t op u s ) K i ef f . 1 1 97, 1 1 99 "' c a m e l u s ( G ona t o p u s ) D a l l a Torre 1 485 * came r on i ( An t e on ) K i e f f e r 3 1 7 , 334 "' c a m e r on i ( C h e l o 9 y n u s ) K i e f f . 334 c a mp b e l l i ( G o n a t o p u s ) O l m i 1 755 c a mp e s t r i s ( G on a t op u s ) P on oma r . 1 556 , 1 5 5 7 , 1 8 36 * C a mp y I o n y x We s t w o o d 7 2 8 , 7 2 9 c a mu s ( Ap h e l op u s ) R i ch a r d s 5 4 , 1 837 c a n a b en s e ( An t e o n ) B e n o i t 3 7 0 * c a n a b en s i s ( H i r t a n t e o n ) B en o i t 287, 370

1 884

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Ent.

ca n a d e n s e ( An t e o n ) A sh m e a d 2 8 9 , 457 * c a n a d e n s i s ( An t e o n ) A s h m e a d 459 * c a n a d e n s i s ( Av od ry i n u s ) P on oma r e n k o 9 3 0 * ca n a d en s i s ( C h e l o g y n u s ) A sh me a d 4 5 8 , 4 5 9 c a n a d en s i s ( P s eu d o g on a t o p u s ) a l m i 1 26 0 c a n a d en s i s ( R i ch a rd s i d ry i n u s ) P on oma r en k o 9 3 0 , 1 8 3 0 c a n a r i e n s i s ( A g on a t op o i d e s ) a I mi 1 3 1 8 c a n a r i e n s i s ( D ry i n u s ) C eb a l l o s 734 c a n a r i e n s i s ( G on a t o p u s ) a l m i 1 550 * c a n a r i en s i s ( P a r a d ry i n u s ) C eb a I l o s 7 3 4 c ap e i ( A n t eon ) a l m i 4 0 9 c a p en s i s ( G on a t op u s ) B r u e s 1 60 1 c a p i c o l a ( G on a t op u s ) K i e f f e r 1 80 2 c a ra i b i cu s ( D ry i n u s ) a l m i 8 4 9 c a r a i b i cu s ( M e s od r y i n u s ) a l m i 1 02 9 * ca r e t t e i ( G on a t op u s ) B r u c h 1 759 * ca r i n a t u s ( An t e o n ) K i e f . 3 2 2 c a r i n a t u s ( A p h e l op u s ) K i e f f e r 1 80 2 * ca r i n a t u s ( C h e l ogynu s ) K i e f . 322 * c a r i n a t u s ( D e i n o d ry i n u s ) F en . 1 36 * c a r i n a t u s ( N eod r y i n u s ) F ou t s 1 09 4 * c a r i n a t u s ( P s i l od ry i n u s ) F ou t s 1 0 9 3 * ca r i n i c e p s ( D ry i n u s ) Came ron 91 8 * c a r i n i c ep s ( L e s t od r y i n u s ) Came r on 9 1 8 c a r i n i c e p s ( R i c h a rd s i d ry i n u s ) C ame r on 9 1 8

I nst . ,

n o . 37 ,

1 984

c a rp e n s ( D e i n od r y i n u s ) a l m i 2 3 2 c a s a l e i ( P r i o ra n t e on ) a l m i 5 8 9 , 5 9 3 , 596 c a s c u s ( D e i n od ry i n u s ) a l m i 1 8 2 c a t a r i n a e ( D ry i n u s ) a l m i 8 6 4 * c e p h a l o t e s ( An t e on ) L j u n g h 3 1 7 * c e p h a l o t e s ( G on a t op u s ) L j u n g h 316 c e r e s ( A g on a t op o i d e s ) a l m i 1 3 2 8 c e r r u t i i ( Ad ry i n u s ) B e n o i t 1 1 26 , 1 1 28 * c e r r u t i i ( N e od ry i n u s ) B en o i t 1 1 28 * C h a l c og on a t op u s R . P e rk i n s 1 499 c h a ma e l e o ( D ry i n u s ) B e n . 7 7 5 c h e e sma n i ( N eod ry i n u s ) a l m i 1 1 16 C h e l a n t e on a l m i 1 0 5 ch e l a t u s ( N e o d ry i n u s ) a l m i 1 0 7 9 ch e l ogy n o i d e s ( A n t e on ) R . P e rk i n s 5 2 7 , 1 8 4 5 * ch e l ogy n o i d e s ( P r o s a n t e on ) R . P e rk i n s 2 8 6 , 5 2 7 * C h e l ogyn u s H a l i day 2 8 6 , 2 8 7 , 288, 1 83 1 C h e l o t h e l i u s R e i n h a rd 6 0 4 , 6 0 5 ch e r s on e s i u s ( G on a t o p u s ) P on oma r en k o 1 5 7 2 ch i a p a s i ( D e i n od r y i n u s ) a l m i 209 c h i r i q u en s e ( A n t e on ) C a m . 5 0 8 * ch i r i q u en s i s ( D r y i n u s ) C a m . 508 * ch i r i q u e n s i s ( Me s od ry i n u s ) C ame r o n 5 0 8 * C h l o r od ry i n u s R . P e rk i n s 728, 1 832 * ch l o r o t e t t i x i ( C h e l o g y n u s ) F en t on 4 8 1 , 1 8 2 9 c h o r t i n i s ( P l e c t r o g o n a t op o i d e s ) P on oma r en k o 1 4 3 1 , 1 4 3 2 c i I i p e s ( G o n a t op u s ) K i e f . 1 7 6 4 * c i I i p e s ( N e o g o n a t op u s ) K i e f f e r 1 764 c i t r i c o l u s ( D ry i n u s ) a l m i 835

I n dex

c i t r i n i c o l l i s ( An t e on ) K i e f f . 1 80 2 c i t r i n u s ( Ap t e r od ry i n u s ) D l m i 1 359 c i udad i (An t eon ) D I m i 486 c l a r i c o l l i s ( An t e o n ) K i e f f e r 1 80 3 * c l a r i c o l l i s ( Ch e l ogynu s ) K i ef f e r 1 80 3 c l a r i p e s ( D e i n od ry i n u s ) D l m i 206 c l a ru s ( T h a u ma t od ry i n u s ) D l m i 715 * c l a u d i a e ( N e og on a t op o i d e s ) Abdu l - N ou r 1 4 3 4 , 1 4 4 2 c l a v a t u m ( An t e o n ) D l m i a n d C u r ra d o 4 9 1 * c l a v a t u m ( P r en a n t e on ) Mo c z a r 261 * c l a v i c o rn i s ( C y r t o g o n a t op u s ) F en t on 1 6 7 3 c l a v i c o r n i s ( G on a t op u s ) F e n t on 1 673 * c o g n a t u s ( An t e o n ) R . P e rk i n s 549 * c og n a t u s ( C h e l o g y n u s ) R . P e rk i n s 5 2 7 , 5 4 9 * c o g n a t u s ( N e o ch e l ogy n u s ) R . P e rk i n s 5 4 8 * c o l l a r i s ( A n t e o n ) D a l ma n 3 1 3 * c o l l a r i s ( G on a t op u s ) D a l . 3 1 3 c o l l a r i s ( D ry i n u s ) L i n n . 7 3 5 , 736 , 1 83 7 , 1 83 8 , 1 83 9 * c o l l a r i s ( Sp h ex ) L i n n a e u s 7 3 5 c o l omb i a n a ( C r ov e t t i a ) D l m i 9 4 c o l omb i a n u s ( D e i n od ry i n u s ) D l m i 1 80 * c ome s i ( Ap h e l op u s ) F en t on 7 4 c orrmu n i s ( G on a i op u s ) D l m i 1 6 1 0 c o mp r e s s i c o x a l e ( An t e o n ) B en . 398 * c omp r e s s i c ox a l e ( X en a n t e on ) B en o i t 3 9 8 C on g a n t e o n B en o i t 9 8 C on g a n t e o n i n a e D l m i 9 6 * C on g od ry i n u s B en o i t 1 4 0 8 * c on j u n c t u s ( G on a t op u s ) K i e f .

1 885

1 3 7 9 , 1 38 0 c on j u n c t u s ( L ab e o ) K i e f . 1 8 0 3 * c on j u n c t u s ( L ab e r i u s ) K i e f f e r 1 803 c on s t a n s ( D r y i n u s ) D l m i 8 3 3 c o n t e r m i n u m ( An t e on ) D l m i 5 1 6 c o n o r t u l u s ( G o n a t op u s ) P a t t on 1 68 8 , 1 8 4 2 * c on t o r t u l u s ( N e o g on a t op u s ) P a t t on 1 6 8 8 c on t o r t u s ( G on a t op u s ) D l m i 1 7 6 8 c o ra l I i n u m ( An t e on ) D l m i 5 8 1 c o ra x ( An t e on ) D l m i 3 4 5 c o r i a c eu m ( An t e o n ) R . P e rk i n s 5 7 1 , 1 845 * c o r i a c eu s ( An t e on ) R . P e rk i n s 571 * c o r i a c eu s ( C h e l o g y n u s ) R . P e rk i n s 5 7 1 * c o r i a c eu s ( N e o ch e l o g y n u s ) R . P e rk i n s 5 7 1 c o r n u t u s ( P s eu d og on a t op u s ) B e n . 1 80 3 c o r r u p t u m ( An t e o n ) D l m i 5 4 5 * c o r s i c a e ( L e s t od r y i n u s ) K i e f f . 735 * c o r s i c u s ( D ry i n u s ) Ma r sh . 9 1 3 * c o r s i c u s ( Me s o d ry i n u s ) Ma r s h a l l 9 1 3 c o r s i cu s ( R i ch a rd s i d ry i n u s ) Ma r sh . 9 0 7 , 9 1 3 , 1 8 3 9 c r a s s i f r o n s ( A n t e on ) Moc z a r 3 3 2 * c ra s s i f r o n s ( C h e l o g y n u s ) Mo c z a r 3 3 2 * c ra s s i ma n u s ( An t e on ) H a l i d a y 260 * c r a s s i man u s ( D r y i n u s ) H a I i d a y 259 * c ra s s i s c a p u s ( An t e on ) K i e f f e r 259 * c ra s s i s c a p u s ( P r en a n t e on ) K i ef f er 260 c r a w f o rd i ( D ry i n u s ) K r om . 8 2 5 * c r a w f o rd i ( Me s od ry i n u s ) K r omb e i n 8 2 5 * c r e n u l a t u m ( An t e on ) K i e f . 2 9 5 * c r en u l a t u s ( An t e on ) K i e f . 2 9 5

1 886

M e m . Amer .

Ent .

C r e t o d r y i n u s P o n o ma r e n k o 1 0 4 6 * c r i s t a t u s ( C h a l c o g o n a t op u s ) K i ef f e r 1 65 2 c r i s t a t u s ( G o n a t op u s ) K i e f f e r 1 65 2 * c r i s t a t u s ( H a p l og o n a t op u s ) K i ef f . 1 652 c r o c e u s ( D e i n od r y i n u s ) D l m i 223 C r ov e t t i a D l m i 8 4 C t en od r y i n u s A r l e 1 7 9 8 c u b en s i s ( A c r o d on t o c h e l y s ) R i ch a r d s 1 1 8 8 " c u b en s i s ( C y r t og on a t o p u s ) R i ch a rd s 1 1 8 8 c u p r ep e s ( C h a l c o g o n a t op u s ) K i e f f e r 1 70 9 * c u p r ep o i d e s ( C h a l c o g on a t op u s ) K i e f f e r 1 70 9 c u r ra d o i ( An t e o n ) D l m i 5 5 6 c u r r i e i ( G on a t op u s ) K r o mb e i n 1 724 · cu r s o r

( An t e o n ) H a l i d a y

295 ,

299 * cu r s o r ( D ry i n u s ) H a l i d a y 2 9 4 * c u rv a t u s ( An t e on ) K i e f f . 3 0 0 * c u r v i n e rv i s ( A n t e on ) K i e f f e r 260 '� c u r v u s ( An t e on ) K i e f f e r 3 0 0 cu z c a n u s ( D e i n od r y i n u s ) D l m i 249 cy p h on o t u s ( G o n a t op u s ) B r ad l e y 1 67 6 , 1 8 4 2 " C y r t o g on a t op o i d e s P on om . 1 5 0 0 * C y r t o g o n a t op u s K i e f f e r 1 5 0 0 dahms i ( D ry i n u s ) D l m i 8 9 7 * d a o s ( An t e on ) Wa l k e r 2 6 0 * d a o s ( D r y i n u s ) Wa l k e r 2 5 9 * d a o s ( P r e n an t e on ) Wa l k e r 2 6 1 d a u n u s ( G on a t opu s ) D l m i 1 6 5 6 d a y i ( An t e o n ) D l m i 3 6 7 d a y i ( Me s od r'r i n u s ) D l m i 1 0 0 3 d e b i I e ( An t e on ) D l m i 4 4 7 d e c e l l e i ( An t e o n ) D l m i 3 8 0 d e c ep t o r ( D ry i n u s ) T u r n e r 7 6 2 * d e c e p t o r ( L e s t od r y i n u s ) T u rn e r 762

I ns ! . ,

no.

37 ,

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d e c i p i e n s ( G on a t o p u s ) P rovan c h e r 1 804 '� d e c l i v i s ( A n t e o n ) K i e f f e r 2 5 9 * d e c o r a t u s ( C h a l c o g o n a t op u s ) R . P e rk i n s 1 7 8 8 d e c o ra t u s ( G on a t opu s R . P e rk . 1 7 88 , 1 8 4 5 * d e c r e t o r i u s ( G on a t op u s ) H a u p t 1 379 D e i n od r y i n u s R . P e rk i n s 1 1 8 d e l e t u s ( T h a u ma t od ry i n u s ) B r u e s 1 804 , 1 8 3 0 " d e l i c a t u l u s ( An t e o n ) C h i t . 3 1 7 d e l p h a c i d i s ( P s eu d og o n a t op u s ) D l mi 1 279 * d en t a t i f o r c e p s ( G o n a t op u s ) K i ef f e r 1 1 97 " d en t a t i f o r c ep s ( P r i s t og o n a t opu s ) K i e f f e r 1 1 9 2 , 1 1 97 d e n t a t u s ( T h a u ma t o d ry i n u s ) B en o i t 6 9 0 * d e s e r t i c u s ( P l a t y g o n a t opu s ) B en o i t 1 5 2 7 d e s t r u c t o r ( A n t e o n ) R . P e rk i n s 547 , 549 , 1 844 �' d e s t ru c t o r ( C h e l og y n u s ) R . P e rk i n s 5 4 9 "' d e s t r u c t o r ( N e o c h e l o g y n u s ) R . P e rk i n s 5 4 7 d i a p h a n u s ( Ap h e l op u s ) D l m i 7 7 d i a p h a n u s ( D e i n od ry i n u s ) D l m i 1 58 d i ch r o mu s ( D i c o n d y l u s ) K i e f f e r 1 37 6 , 1 83 8 , 1 83 9 * d i c h r omu s ( G o n a t op u s ) K i e f . 1 376 d i ch r omu s ( P s e u d o g o n a t op u s ) R . P e rk i n s 1 2 9 3 , 1 8 4 6 " d i ch r ou s ( L e s t o d ry i n u s ) F e n t on 958 D i condy l u s H a l i day 1 0 5 2 , 1 3 7 4 , 1 375 d i f f u s u s ( A p h e l op u s ) D l m i 7 6 , 79 d i f f u s u s ( N eodry i nu s ) D l mi 1 089 * D i g on a t op u s K i e f f e r 1 5 0 0

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( A n t e on ) R . C . L .

* d o r s a l i s ( D ry i n u s ) N e e s 3 3 4 , 1 80 4 , 1 8 1 9 d o r sa l i s ( Me s od ry i n u s ) D l m i 1 021 d o r s e t t e n s i s ( H omop t e r op h a g u s ) Da l e I 804 d ra c o ( D ry i n u s ) Ma r sh a I I 1 8 0 4 * d r a c o ( P r od ry i n u s ) Ma r sh a l l 1 805

P e rk i n s 5 6 1 " d i m i d i a t u s ( Ch e l ogynu s ) R . P e rk i n s 5 6 1 d i m i d i a t u s ( G on a t op u s ) J . Sah I b e rg 1 5 2 8 * d i m i d i a t u s ( N e o ch e l o g y n u s ) R . P e rk i n s 5 6 1 * d i m i d i a t u s ( N e o g on a t op u s ) J . Sah l b e rg 1 5 2 8

Nees

334 ,

d i s t i n g u en d u s ( G on a t o p u s ) K i ef . 1 51 3 , 1 5 1 5 , 1 835 , 1 836

d r a c u n c u l u s ( D ry i n u s ) Ma r sh a l 1 805 * d ra c u n cu l u s ( P r od r y i n u s ) Ma r s h a I I 1 8 0 5 * d r o m ed a r i u s ( D i c on d y l u s ) C o s t a 1 209 ':' d r ome d a r i u s ( G on a t op u s ) C a me r on 1 4 8 5 * d r omed a r i u s ( G on a t o p u s ) C o s t a 1 20 9 , 1 560

* d i s t i n g u e� � u s ( N e o g on a t op u s ) K i ef . 1 5 1 5 * d i v i s u s ( An t e o n ) K i e f . 3 1 7 * d od d i ( A n t e on ) K i e f f e r 5 5 4 * d od d i ( L i od r y i n u s ) K i e f f e r 287 , 554 d od e r o i ( G on a t op u s ) D l m i a n d C u r ra d o 1 5 1 1 " d od e r o i ( N e o g on a t op u s ) D l m i and C u r ra d o 1 5 1 1 * d o e l l o j u r a d o i ( D i c on d y l u s ) D g l ob l i n 1 7 5 7 d o e l l o j u r a d o i ( G on a t o p u s ) D g l ob l i n 1 7 5 7 " d o l i c h o c e r u s ( A n t e on ) K i e f f e r 259 * D o l i c h o ch e l y s D l m i a n d C u r r o 1 34 2 d o l o s u s ( N e o d ry i n u s ) D l m i 1 08 1 d o m i n i c a n u s ( N e od r y i n u s ) E v a n s 1 101 * d om i n i c a n u s ( P r o d ry i n u s ) Ev a n s 1 1 0 1 D on i s t h o r p i n a R i c h a rd s 1 3 0 4 * d o r sa l i s ( A n t e on ) K i e f . 1 80 4

d r omeda r i u s ( P s e u d o g on a t op u s ) C o s t a 1 209 d ry i n i f o rm i s ( G on a t o p u s ) Ma r s h a l l 1 8 0 5 D ry i n i n a e K i e f f e r 7 2 3 D r y i n op s i s B r u e s 1 8 2 4 * D ry i n u s F ab r i c i u s 7 2 8 D ry i n u s L a t r e i I I e 7 2 8 , 7 2 9 , 1 832 d u b i o s u s ( A g on a t o p u s ) R . P e rk . 1 697 * d u b i o su s ( N e od ry i n u s ) R . P e rk i n s 1 0 9 1 , 1 0 9 4 * d u b i o su s ( N e og on a t op u s ) R . P e rk i n s 1 7 9 0 * d u b i o su s ( P s e u d o g o n a t op u s ) R . P e rk i n s 1 2 7 1 ':' d u b i o s u s ( P s i l od ry i n u s ) R . P e rk i n s 1 0 9 3 , 1 0 9 4 d u b i u m ( A n t e on ) F ou t s 5 7 0 * d u b i u s ( An t e on ) F ou t s 5 7 1 d u b i u s ( B o c ch u s ) D l m i 6 5 3 d u b i u s ( D i c on d y l u s ) D l m i 1 3 9 9 d u r a n g o i ( An t e o n ) D l m i 4 8 4 * e ch o ( C h a l c og on a t op u s ) R . P e rk i n s 1 6 8 3

* d i s t i n c t u s ( D i c ond . ) K . 1 2 1 3 " d i s t i n c t u s ( G on a t op u s ) K i e f 1213 d i s t i n c t u s ( P s e u d o g on a t op u s ) K i ef . 1 2 1 2, 1 2 1 3 , 1 2 1 7 , 1 83 8 , 1 839

1 888

Mem . A mer .

En!.

ec h o

( G on a t op u s ) R . P e r k i n s 1 68 3 E c h t h r od e l p h a x R . P e rk i n s 1 05 3 , 1 054, 1 1 40, 1 83 2 * E l e c t r od ry i n u s P o n oma r e n k o 1 19 e l eg a n s ( D e i n od ry i n u s ) D l m i 1 54 " e I I i man i ( A n t e on ) C h i t t y 3 2 6 e l o n g a t u s ( G on a t op u s ) D l m i 1 684 e me r i t u m ( An t e o n ) D l m i 4 1 0 e m i n en s ( D e i n od r y i n u s ) D l m i 217 e mp oa s c a e ( Ap h e l op u s ) P o n oma r . 1 80 5 e p h i p p i g e r ( An t e o n ) D a l ma n 2 8 9 , 3 1 2 , 3 1 3 , 1 80 3 * ep h i p p i g e r ( C h e l o g y n u s ) D a l m . 3 1 3 , 3 1 7 , 334 , 439 * ep h i p p i g e r ( G on a t op u s ) D a l m . 313 * E p i g o n a t op o i d e s R i ch a rd s 1 4 3 4 E p i g on a t op u s R . P e rk i n s 1 4 9 0 e r i c e t i ( G o n a t op u s ) D l m i 1 6 3 7 ':' e r r a t i cu s ( D ry i n u s ) T u rn e r 916 " e r r a t i c u s ( N e od r y i n u s ) T u r n e r 916 e r r a t i c u s ( R i ch a rd s i d r y i n u s ) T u rn e r 9 1 6 , 1 8 4 0 e ry t h r o c ep h a l u s ( G on a t o p u s ) N e e s 1 805 " e r y t h r od e s ( N e og on a t o p u s ) R . P e rk i n s 1 7 1 7 E s a g on a t op u s D l m i 1 0 5 4 , 1 4 9 3 e s a k i i ( An t e on ) Ya s u ma t s u 1 8 0 5 * e s c o r i a l en s i s ( Me s o d r y i n u s ) C eb a l l o s 9 1 3 * E u c a mp t on y x R . P e rk i n s 1 0 5 2 , 1 499 * E u g on a t op u s R . P e rk i n s 1 4 9 9 * E u k o eb e l e i a R . P e rk i n s 6 1 2 * e u p r e p e s ( C h a l c o g on a t op u s ) R . P e rk i n s 1 7 0 9 * eu p r e p o i d e s ( C h a l c og o n a t o p u s ) R . P e rk i n s 1 7 0 9

I ns t . ,

n o . 37 ,

1 984

eu r o p a e a ( B o c c h u s ) B e r n a rd 6 2 4 * e u r o p a e a ( E u k o eb e l e i a ) B e r n a r d 624 eu r o p a e u s ( B o c ch u s ) B e rn a rd 624, 1 839 " eu s c e l i s i ( P r en a n t e on ) H a u p t 261 * ex c a v a t u s ( G on a t op u s ) J . S a h l 1515 * e x c i s u s ( A n t a e o n ) We s t w o o d 1 378 * e x c i s u s ( L a b e o ) We s t w o od 1 3 7 9 * e x c i s u s ( L a b e r i u s ) We s t . 1 3 7 9 * ex i g u u s ( A n t e on ) 3 2 2 * ex i g u u s ( C h e l og y n u s ) H a u p t 3 1 7, 322 ex i l i s ( D r y i n u s ) D l m i 8 0 2 e x o p h t h a l m i c u s ( T r i d ry i n u s ) D I m i 962 exp o l i t u m ( A n t e o n ) D l m i 444 f ab u l o s u m ( A n t e o n ) D l m i 4 0 4 * f a c i a l i s ( A n t e on ) T h oms o n 3 1 3 * f a c i a l i s ( D r y i n u s ) T h om s on 3 1 3 f a i r ch i l d i ( E c h t h r od e l p h a x ) R . P e rk i n s 1 1 4 0 , 1 1 4 1 , 1 1 5 8 , 1 1 6 1 , 1 1 63 , 1 83 1 , 1 8 3 3 , 1 83 8 , 1 83 9 , 1 84 1 , 1 846 f a l l a x ( E p i g o n a t opu s ) R . P e rk i n s 1 4 9 1 f a v r ea u a e ( Me s o d r y i n u s ) D l m i 1 024 f e l i x ( Ag o n a t op o i d e s ) D l m i 1 3 1 2 , 1 344 f e n n i c u s ( An t e on ) H e l l e n 1 8 0 5 f e n t on i ( A p h e l op u s ) D l m i 7 8 f en t o n i ( N e og o n a t op u s ) F r ey t a g 1 805 f e rn a n d i n a e ( G on a t o p u s ) D l m i 1 73 2 f e r r u g i n eu s ( A g on a t op u s ) R . P e rk i n s 1 7 0 3 f e r r u g i n eu s ( A l p h a d r y i n u s ) D l m i 995 * f e r r u g i n eu s ( An t e o n ) B r u e s 1 3 6 * f e r r u g i n eu s ( C h e l o g y n u s ) B r u e s 1 33 * f e r r u g i n e u s ( D e i n od r . ) S r . 1 3 6

1 889

I ndex

f i j i a n u m ( An t e o n ) O l m i 5 6 9 1 845 f i l i c o rn i s ( An t e o n ) K i e f . 1 8 0 6 f i l i c o r n i s ( D ry i n u s ) B r u . 1 8 0 7 * f i l i c o rn i s ( G o n a t op u s ) K i e f . 1 54 1 , 1 543 * f i l i c o rn i s ( H a rp a c t o sp h e c i on ) B ru e s 1 8 0 7 * f i l i c o rn i s ( P r en a n t e o n ) O g l ob l i n 2 6 0 * f i i i c o rOn i s ( T h a u ma t od ry i n u s ) B ru e s 1 8 0 7 , 1 8 3 0 F i o r i an t e on O l m i 1 0 8 f i o r i i (An t eon ) O l mi 3 6 2 f i rmu m ( An t e o n ) O l m i 5 5 8 * f l a v i c e p s ( T e t rad r y i n u s ) K i ef . 650 * f l a v i c o l l i s ( B o c ch u s ) A s h m e a d 6 1 2 , 650, 6 5 1 f l av i c o rn e ( An t e o n ) D a l ma n 289 , 305 , 309, 3 1 2 , 3 1 3 , 3 2 9 , 1 83 1 , 1 834 * f l a v i c o rn i s ( A n t e on ) D a l ma n 309 * f l av i c o rn i s 309 , 1 5 1 * f l a v i c o rn i s T h om s on

I

( G on a t op u s ) D a l m . 3, 1 5 1 5 ( G on a t op u s ) 1515

f l a v i f emu r ( P s e u d o g o n a t op u s ) E s ak i a n d H a sh i mo t o 1 20 2 , 1 83 9 * f l a v i f r o n s ( C h a l c o g on a t op u s ) A sh m e a d 1 7 1 2 f l a v i f r o n s ( G on a t op u s ) A sh m e a d 1 71 2 * f l a v i n e r v i s ( An t e on ) K i e f f e r 337 f l a v i p e s ( A n t e on ) K i e f . 1 8 0 7 f l a v i p e s ( B o c ch u s ) K i e f . 6 5 0 f l av i p e s ( G on a t o p u s ) O l m i 1 7 5 2 * f l av i p e s ( T e t rad ry i n u s ) K i e f . 6 1 2 , 650 * f l a v i s c a p u s ( An t e o n ) K i e f f e r 322 , 337 * f l a v i s c a p u s ( An t e on ) J a n s s o n 3 1 3 , 3 1 7 , 322 * f l a v i s c a p u s ( An t e on )

Wh i t t a k e r 4 8 1 * f l a v i t a r s i s ( A n t e on ) K i e f . 3 0 0 * f l a v i v en t r e ( A n t e o n ) H e l l e n 1 829 f l a v on i g e r ( D ry i n u s ) O l m i 8 3 8 * f l a v u s ( G on a t o p u s ) H e d i ck e 1 56 0 f l av u s ( L on ch od ry i n u s ) O l m i 2 7 4 f l a v u s ( P s eu d og o n a t op u s ) O l m i 1 277 f l av u s ( T h a u ma t od r y i n u s ) O l m i 718 f l u v i a t i l i s ( D e i n od r y i n u s ) O l m i 248 f o ca r i l e i ( P s eu d og o n a t op u s ) D l m i 1 20 6 * f o e r s t e r i ( A n t e on ) K i e f . 2 5 9 f o r e s t a l i s ( Me s od ry i n u s ) O l m i 1 027 f o r m i c a e f o rm i s ( My s t r o p h o r u s ) Ru t h e 6 7 8 , 6 7 9 , 1 8 3 5 f o r m i ca r i a 1 80 7 ,

( Mu t i l l a ) 1810

Jurine

* f o rm i c a r i u s ( D ry i n u s ) L a t r . 729, 735, 736 f o r m i ca r i u s ( G o n a t op u s ) L j u n g h 1 50 2 , 1 56 4 , 1 57 7 , 1 82 8 , 1 835, 1 836 * f o rm i ca r i u s ( L e s t od r y i n u s ) Lat rei I I e 728, 735, 736 * f o rm i c i c o l a ( Do n i 5 t h o r p i n a ) R i cha r d s 1 30 4 , 1 3 0 6 * f o r m i c i n u s ( G on a t op u s ) K r og . 1 56 4 , 1 829 * f o u t s i ( G on a t op u s ) Wh i t t a k e r 1 676 * f ov e a t u s ( P r en a n t e on ) R i c h a r d s 261 * f ra c t i n e rv i s 260

(An t eon ) K i e f f e r

* f r eq u e n s ( C h a l c o g on a t op u s ) R . P e rk i n s 1 6 9 7 f r e q u en s ( G o n a t o p u s ) R . P e rk i n s 1 69 7 f r ey t a g i ( G o n a t op u s ) O l m i 1 6 8 5 f r e y t a g i ( P s eu d o g on a t o p u s ) D l m i 1 25 8 , 1 25 9 , 1 84 3

Mem . Amer . E n t .

1 890

* f r on t a l i s * f r on t a l i s

( An t e o n ) D a l m . 2 5 9 ( Ch e l og y n u s ) D a l m .

259 * f r on t a l i s ( G on a t op u s ) D a l ma n 258 * f r on t a l i s ( P r e n a n t e o n ) D a I m . 260 , 261 * f u l g o r i ( G on a t op u s ) N a k a g a wa 1 1 94 * f u l g o r i ( H a p l o g on a t op u s ) N a k a g a wa 1 1 9 4 * f u l g o r i ( P a r a g on a t op u s ) N a k a g awa 1 1 9 5 f u l g o r i ( P s e u d og o n a t op u s ) N a k a g a wa 1 1 9 4 , 1 8 3 8 f u l v i c o l l i s ( Ch e l ogynu s ) H ed i c k e 1 8 0 7 f u l v i v e n t r e ( An t e on ) H a l i d a y 3 1 3 , 337, 338, 1 803 * f u l v i v en t r i s ( An t e o n ) H a l i d a y 338 * f u l v i v en t r i s ( D ry i n u s ) H a l 337 f u l v u s ( D ry i n u s ) O l m i 8 1 5 f u n e s t u m ( A n t e o n ) R . P e rk i n s 462 * f u n e s t u s ( An t e on ) R . P e rk i n s 463 * f un e s t u s ( Ch e l ogynu s ) R . P e rk i n s 4 6 3 f u s c i c o l l i s ( An t e on ) K i e f f e r 1 80 7 * f u s c i c o rn i s ( G on a t op u s ) D a l m . 258 * f u s c i f o r m i s ( An t eon ) K i e f f e r 325 * f u s c i p en n i s ( An t e o n ) K i e f f e r 260 * f u s c i p e s ( An t e o n ) T h o m . 3 3 8 * f u s c i p e s ( D ry i n u s ) T h omson 337 * f u s c o c l a v a t u m ( An t e on ) K i e f . 31 7 * f u s c o c l a v a t u s ( An t e on ) K i e f . 31 7 f u s c u m ( Me t a n t e o n ) O l m i 1 1 6 f u s cu s

( Ag on a t op o i d e s ) O l m i

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no.

37,

1 984

1 32 1 * f u s i f o rm i s ( An t e on ) K i e f . 3 2 5 f y a n e n s e ( An t e on ) O l m i 4 3 5 g a r a mb a n u m ( An t e o n ) O l m i 4 1 1 g a u l l e i ( An t e o n ) K i e f . 3 3 4 , 1 80 3 " gau l l e i ( Ch e l ogynu s ) K i ef f e r 286 , 334 g i b b o s u s ( A l p h a d ry i n u s ) O l m i 985 g i b b o su s ( T r i d ry i n u s ) O l m i 9 6 9 * g i g a s ( C h a l c o g on a t op u s ) R . P e rk i n s 1 4 9 9 , 1 7 7 9 g i g a s ( D r y i n u s ) R . P e rk i n s 8 8 5 g i g a s ( G on a t op u s ) R . P e rk i n s 1 77 9 , 1 845 g i g a s ( N e o d ry i n u s ) C eb . 1 0 7 1 * g i g a s ( P a r a d ry i n u s ) R . P e rk i n s 885 g i l u w e n s e ( A n t e on ) O l m i 5 2 6 g i r a u d i ( An t e on ) K i e f . 1 8 0 7 g i ra u l t i ( An t e o n ) D od d 5 8 7 g l ab e r ( D ry i n u s ) O l m i 9 0 0 * g l a b e r ( G on a t op u s ) K i e f . 1 5 1 3 * g l a b e r ( N e og on a t op u s ) K i e f f e r 1 51 3 g l o r i o su m ( An t e o n ) O l m i 5 7 5 g ob i e n s i s ( Ao : e r o d r y i n u s ) P on oma r e n k o 1 3 4 4 g ob i e n s i s ( M i r o d ry i n u s ) P o n o m . 666 * g o b i en s i s ( N og a t op u s ) P o n o m . 1 34 2 , 1 344 G on a t o p od i n a e K i e f f e r 1 0 5 0 * G on a t op od i n i 1 0 5 0 G on a t op u s L j u n g h 1 0 5 0 , 1 0 5 2 , 1 0 5 3 , 1 0 54 , 1 0 5 5 , 1 49 8 1 500, 1 83 2 , 1 833 * g ra c i l i c o l l i s ( An t e o n ) K i e f . 337 * g ra c i I i c o r n i s ( G o n a t op u s ) K i e f f e r 1 54 1 " g ra c i I i p e s ( G on a t op u s ) K i ef f e r 1 543 * g ra c i l i s ( Ad ry i n u s ) F e n t on 1 1 3 8 g ra c i I i s ( Ag o n a t op u s )

I n dex

E sak i 1 80 7 * 9rac i i i s 1 543

a n d H a sh i mo t o ( G o n a t op u s ) K i e f f e r

* g ra c i l i s ( P s i l od r y i n u s ) F e n . 1 1 38 g r a c i l i s ( T h a u ma t od r y i n u s ) B ru e s 1 8 0 7 , 1 8 3 0 g ra e c u s ( G on a t o p u s ) O l m i 1 5 8 7 , 1 835 g r a e f f i ( G o n a t op u s ) K i e f f e r 1 80 7 g r a m i n e u s ( N e o g o n a t op u s ) P on .

1 80 7 * g r a n d i s ( A n t e o n ) B ru e s 1 3 6 * g rand i s ( Ch e l ogynu s ) B r . 1 3 6 * g r a n d i s ( D e i n od ry i n u s ) B r u e s 1 36 g ra n d i s ( D ry i n u s ) Og l o . 8 5 2 * g r a n d i s ( Me s od r y i n u s ) O g l ob . 852 g r a n d i s ( N e o g o n a t op u s ) F r ey t a g 1 80 8 ( L e s t od ry i n u s ) H 0 f f e r * g r eg 0 r i 744 g r e n a d en s i s ( L a b e o ) A s h m e a d 1 80 9 * g r en a d en s i s ( L a b e r i u s ) A s h m . 1 80 9 g ry p s ( C h e l o t h e l i u s ) R e i n h a r d 605 , 607 * g r y p s ( D r y i n u s ) R e i n h a rd 6 0 7 " g u i g l i a e ( C h a l c o g on a t op u s ) B en o i t 1 5 9 6 * g u i g l i a e ( D i g on a t op u s ) B en o i t 1 59 6 g u i g l i a e ( G on a t o p u s ) B e n . 1 5 9 6 g u i n e en s i s ( Ap h e l op u s ) O l m i 8 2 g u i n e en s i s ( B o c c h u s ) D l m i 6 5 8 g u i n e e n s i s ( D ry i n u s ) O l m i 8 8 4 g u i n e en s i 5 ( L on c h od ry i n u s ) O l mi 284 g u t t u rn i u m ( An t e on ) B en . 3 8 4 " g u t t u rn i u m ( X en a n t e o n ) B e n o i t 384 * G y mn a p h e l o p u s B en o i t 4 4 G y n o c h e l y s B ru e s 1 7 9 2 , 1 7 9 3

1 89 1

h a g en i ( An t e on ) O l m i 5 7 8 * h a l e a k a l a e ( G on a t op u s ) A sh mead 1 394 * h a l i d a y i ( A n t e on ) K i e f f e r 2 6 0 H a p l o g o n a t op u s R . P e rk i n s 1 0 5 3 1 05 4 , 1 40 8 , 1 83 1 * h a p l o t h o r a x ( G on a t opu s ) R . P e rk i n s 1 4 6 8 H a rp a c t o s p h e c i on H a u p t 1 8 0 7 * h a rp a x ( C h a l c o g o n a t op u s ) K r om . 1 703 h a r p a x ( G on a t op u s ) K r o m . 1 7 0 3 h au s t u m ( A n t e on ) D l m i 5 8 0 * h a wa i i en s i s ( L a b e o ) A sh m . 1 3 9 4 * h e i d e l b e r g e n s i s ( Ap h e l op u s ) R i c h a rd s 5 4 " h e I I e n i ( D i c on d y I u s ) R a a t i k a i n en 1 3 7 6 " h en s h a w i ( A n t e o n ) A s h m e a d 1 3 6 * h e n s h a w i ( C h e l o g y n u s ) A s h m ead 1 33 * h e n s h a w i ( D e i n od ry i n u s ) A shmead 1 3 6 * h e r b a r u m ( C h a l c o g o n a t op u s ) R . P e rk i n s 1 7 0 5 h e r b a r u m ( G o n a t op u s ) R . P e rk i n s 1 704 h e r n a n d e z a e ( H a p l o g o n a t op u s ) D l mi 1 42 3 , 1 844 * H e s p e r od r y i n u s R . P e rk i n s 7 2 8 , 729 H e t e r o l ep i s N e e s 1 8 1 9 " h e t e r o t h o r a x ( A g on a t op u s ) R . P e rk i n s 1 4 7 0 h i l a r e ( An t e on ) D l m i 342 " H i r t a n t e o n B en o i t 2 8 7 * h i r t i c o r n i s ( An t e o n ) K i ef f er 238 h i r t i c 0 r n i s ( D e i n od r y i n u s ) K i ef f er 237 " h i r t i c o rn i s ( T r i san t e on ) K i e f . 1 1 9 , 238 h i r t i f r o n s ( An t e on ) Wh i t t a k e r 487 , 4 8 8 * h i s p a n i c u s ( G o n a t op u s ) K i e f f e r 1 538, 1 573

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1 892

Ent.

h i sp i d u s ( T r i c h o g on a t op u s ) O l m i 1 47 8 * h o l ome l a s 50

( Ap h e l o p u s ) R i c h .

H omop t e r o p h a g u s D a l e 1 8 0 4 h o r t u s e n s i s ( E ch t h r od e l p h a x ) A b d u l -N o u r 1 1 4 7 , 1 8 3 8 , 1 839 * h o r t u s en s i s ( P t e r og on a t op u s ) A b d u l -N ou r 1 1 4 0 , 1 1 4 7 h o rv a t h i ( G o n a t op u s ) 1 50 9 , 1 8 3 6

K i ef f er

* h o rv a t h i ( P l a t y g on a t op u s ) K i ef f er 1 509 h o s p e s ( P s eu d og o n a t op u s ) R . P e rk . 7 , 1 2 5 4 , 1 29 0 , 1 29 1 , 1 83 2 , 1 83 3 , 1 84 6 , h ov a ( An t e on ) B e n o i t 3 9 7 h ov a ( D ry i n u s ) C e b a l l o s 7 7 2 * h ov a ( M e s od r y i n u s ) C eb . 7 7 2 h oy o i ( An t e on ) O l m i 3 9 0 * h y a l i n i p en n i s

( An t e o n ) K i e f .

260 loy a l i n u s ( G on a t op u s ) O l m i 1 6 0 7 h y me n a e u s ( D e i n od r y i n u s ) O l m i 1 78 * Hy me n od r y i n u s B e n o i t 6 1 2 i ch n e u mon i d e s ( Me t h o c a ) L a t r . 1 807, 1 8 1 0 * I d o l o g on a t op u s O g l ob l i n 1 3 7 5 * i g i I i i ( P s e u d og on a t op u s ) O l m i and Cu rrado 1 2 0 6 * i gn o t u s ( Ch e l ogynu s ) R . P e rk i n s 5 4 9 i g n o t u s ( G on a t o p u s ) D g l ob l i n 1 81 0 * i g n o t u s ( N e o ch e l o g y n u s ) R . P e rk i n s 5 4 9 * i l u s ( A n t e o n ) Wa l k e r 2 6 0 * i I u s ( D ry i n u s ) Wa l k e r 2 5 9 * i mb e rb i s ( A n t e on ) K i e f . 3 0 5 i mp r e s su s ( A n t e o n ) K i e f f e r 1 8 1 0, 1 8 1 1 i n ca i c u s ( D e i n od r y i n u s ) O l m i 214 i n ca i cu s ( N e od ry i n u s ) D l m i 1 1 09

I nst . ,

no . 37 ,

1 984

i n c e r t u s ( C h e l o g y n u s ) D g l ob l i n 1 81 1 i n c e r t u s ( D ry i n u s ) D l m i 7 8 5 i n c e r t u s ( L a b e o ) A sh m e a d 1 8 1 2 * i n c e r t u s ( L a b e r i u s ) A sh . 1 8 1 2 i n c i s u s ( Ap h e l op u s ) O l m i 6 4 * i n c l y t u s ( An t e on ) H a l i d a y 3 2 5 * i n c l y t u s ( D ry i n u s ) H a l i d ay 3 2 5 i n c og n i t u s ( G o n a t op u s ) O l m i 1 6 1 3 , 1 839 i n c on su l t u s ( Me s o d ry i n u s ) O l m i 1019 i n d i an u s ( D i condy l u s ) D l m i 1 384 , 1 84 1 i n d i a n u s ( G on a t op u s ) D l m i 1 6 6 7 i n d i a n u s ( R i c h a rd s i d ry i n u s ) D l mi 922 i n d i c u m ( An t eon ) O l m i 4 1 9 , 1 8 4 0 i n d i cu s ( D ry i n u s ) K i e f f e r 8 1 0 8 7 1 , 1 84 1 * i n d i c u s ( Me s od r y i n u s ) K i e f f e r 81 0 * i n d i v i s u m ( An t e o n ) K i e f f e r 300 * i n d i v i su s ( A n t e on ) K i e f f e r 3 0 0 i n d i v i s u s ( Aph e l op u s ) K i e f f e r 1 81 2 i n e r m i s ( D e i n od ry i n u s ) O l m i 2 3 3 i n exp e c t a t u s ( Ap h e l op u s ) B e n o i t 1 79 9 , 1 8 1 2 i n f e c t u m ( An t e on ) H a l i d a y 2 8 9 , 325, 327, 1 8 1 6 * i n f e c t u s ( An t eon ) H a l i d a y 3 2 5 * i n f e c t u s ( C h e l ogy n u s ) H a l i d a y 286, 327 * i n f e c t u s ( D ry i n u s ) H a l i d a y 3 2 5 i n f l a t r i x ( An t e o n ) B e n o i t 2 8 9 , 399 i n f u s c a t u s ( T h a u ma t od r y i n u s ) D l mi 690 i n g e n u u m ( An t e on ) D l m i 3 4 2 * i n i m i c i s ( G o n a t op u s ) F e n t on 1717 * i n n i t i d u s ( A g on a t op u s ) R . P e rk i n s 1 7 2 7 i n n i t i d u s ( G on a t op u s ) R . P e rk . 1 727

1 893

I n dex

i n s i g n i s ( D e i n od r y i n u s ) O l m i 1 93

* j a v a n u s ( P a r a d ry i n u s ) R . C . L . P e rk i n s 8 0 6 , 1 8 3 4

* i n su l a n u m ( P r en a n t e on ) B en o i t 1 24

* j a v a n u s ( P h a n e r od ry i n u s ) R o ep k e 1 0 6 5 , 1 0 8 5

i n su l an u s B en o i i n su l a r i s 1 334 i n su l a r i s 1 350, i n su l a r i s

* j oh a n n a e ( Ag on a t op o i d e s ) C u r rad o and O l m i 1 3 1 6 j oh n s i ( G on a t opu s ) O l m i 1 6 2 6 j u c u n d u s ( An t e on ) O g l ob l i n 1 8 1 2 * j u j u y en s i s ( A g on a t op o i d e s ) Og l ob l i n 1 3 6 2 j u n c e t o r u m ( P s e u d o g on a t op u s ) R . P e rk i n s 1 2 9 8 , 1 8 4 6 j u n o n i u m ( F i o r i a n t e on ) O l m ; ' 1 0 8 , 1 09

( D e i n od ry i n u s ) t 1 24 ( A g on a t op o i d e s ) O l m i ( A p t e r od r y i n u s ) O l m i 1 374 ( D ry i n u s ) D od d 8 8 0

i n su l a r i s ( H ap l ogona t op u s ) O g l ob l i n 1 4 2 3 * i n su l a r i s ( P a ra d r y i n u s ) D od d 880 " i n t eg e r ( A n t e on ) K i e f . 2 5 9 * i n t e r ru p t u s ( G on a t op u s ) K i ef f e r 1 527 * i n t e r r u p t u s ( P l a t y g on a t op u s ) K i ef f e r 1 527 i n v o l u t u m ( An t e on ) O l m i 5 8 4 * i ow en s i s ( P s eu d og on a t op u s ) F e n t on 1 2 7 1 i p h i a s ( D e i n od ry i n u s ) O l m i 1 6 0 i r a n i c u s ( G on a t o p u s ) O l m i 1 5 2 3 i r r e g u l a r i s ( D ry i n u s ) O l m i 7 9 4 i t a l i c u s ( B o c ch u s ) O l m i 6 1 4 i t a l i c u s ( E c h t h r od e l p h a x ) O l m i 1 1 50 i t en e z i ( D e i n o d r y i n u s ) O l m i 245 j a ma i c a n u m ( An t e on ) O l m i 4 9 7 j a ma i c a n u s ( Aph e l op u s ) O l m i 8 1 j a p on i c u m ( An t e o n ) O l m i 3 4 7 * j ap on i cu s ( H a p l o g o n a t op u s ) E sa k i a n d H a sh i mo t o 1 4 1 8 j a p on i c u s ( N e od ry i n u s ) U ch i d a 1 81 2 j a v a n u m ( An t e on ) O l m i 4 3 7 * j a v a n u s ( D i c on d y l u s ) R . P e rk i n s 1 6 5 2 * j av a n u s ( D i g on a t op u s ) R . P e rk i n s 1 5 0 0 , 1 6 5 2 j av a n u s ( G o n a t op u s ) R . P e rk i n s 1 65 2 , 1 840 j a v a n u s ( N e od r y i n u s ) R o ep k e 1 08 5 , 1 8 4 1

j u r i n e a n u m ( A n t e on ) L a t r e i l l e 2 8 7 , 294, 2 9 5 , 299 , 305 , 3 1 2 , 338, 1 834 * j u r i n e a n u s ( An t e on ) L a t r e i l i e 295 k a s a i c u m ( A n t e o n ) B en o i t 3 6 2 * k a sa i cu s 362

( C h e l o g y n u s.) B en o i t

* k a t a n g a e ( C on g o d r y i n u s ) B e n o i t 1 40 8 , 1 4 1 5 k a t a n g a e ( H a p l o g o n a t op u s B en o i t 1 4 1 5 k a wa n d a n u m ( An t e o n ) O l m i 3 7 4 k ev i r a e ( P a rad r y i n u s ) L a l 1 8 1 2 " k i e f f e r i ( An t a e o n ) C h i t t y 3 2 9 * k i e f f e r i ( An t e o n ) C h i t t y 3 2 9 * k i e f f e r i ( Ch e l ogynu s ) C h i t t y 329 k i e f f e r i ( D i condy l u s ) R . P e r . 1 399, 1 846 k i e f f e r i ( D r y i n u s ) t ou t s 7 9 5 * k i e f f e r i ( L e s t od ry i n u s ) F o u t s 796 *k i ef f e r i

( P s eu d og on a t op u s ) R . P e rk i n s 1 3 9 9 k i ms e y a e ( D ry i n u s ) O l m i 8 6 4 k i v u a n u m ( A n t e on ) B en o i t 3 6 3 * k i v u a n u s ( C h e l o g y n u s ) B en o i t 3 5 5 , 363 * k o e b e l e i ( C h a l c og o n a t opu s ) R . P e rk i n s 1 4 7 5 k o e b e l e i ( D ry i n u s ) R . P e rk i n s 8 8 2 , 1 846

1 894

Mem . Amer . E n t .

*k o eb e l e i ( G on a t o p u s ) R . P e r k . 1 7 90 k o eb e l e i ( N e o d r y i n u s ) R . P e rk . 1 0 6 5 , 1 0 8 6 , 1 1 20 , 1 8 4 6 * k o eb e l e i ( P a r a d r y i n u s ) R . P e rk i n s 7 2 8 , 8 8 2 k o eb e l e i ( T h a u ma t od r y i n u s ) R . P e rk i n s 6 8 4 , 7 2 1 , 1 8 4 6 " k o e b e I i ( N eod ry i n u s ) R . P e rk . 1 1 20 * k o r e a n u s ( C h l o r od r y i n u s ) Mo c z a r 7 4 7 k o r e a n u s ( D r y i n u s ) Mo c z a r 7 4 7 * k r og e r i ( G on a t o p u s ) H e l l e n 1 53 8 , 1 82 9 k r omb e i n i ( A n t e o n ) D l m i 4 2 9 k r omb e i n i ( D r y i n u s ) P o n o m . 802, 803 k r omb e i n i ( E ch t h r od e l p h a x ) D l m i 1 1 60 k u r a n d a e ( P s e u d o g on a t op u s ) R . P e rk i n s 1 1 9 2 , 1 2 9 6 , 1 8 4 6 * L a b e a B l a n c h a rd 1 3 7 5 * La b e o C uv i e r 1 3 7 5 * La b e o H a l i d a y 1 3 7 5 * L a b e o l a H a l d e ma n 1 3 7 5 * L a b e r i n u s O g l ob l i n 1 3 7 5 * L ab e r i u s K i e f f e r 1 3 7 5 l a cu a l i s ( G on a t op u s ) O l m i 1 7 6 2 * I a e t i c o r n i s ( An t e on ) K i e f f e r 539 l ame l l a t u s ( A l p h a d ry i n u s ) D I m i 986 * I a n i on i s ( C h e l o g y n u s ) H a u p t 322 l a n k a e ( P s e u d og on a t op u s ) P on oma r e n k o 1 2 4 9 l a n k a n u m ( A n t e on ) O l m i 4 4 1 l an k an u s ( D ry i n u s ) O l m i 8 2 1 l a o t i a n u m ( An t e on ) O l m i 4 3 5 l a ot i an u s ( Bocchu s ) D l mi 635 l a o t i a n u s ( E ch t h r od e l p h a x ) D l mi 1 1 58 * I ap p on i cu s ( A n t e on ) T h o m s on 260 * I a p p o n i c u s ( D ry i n u s ) T h omson 259

I nst . ,

no . 3 7 ,

1 984

* L a s i a n t e on K i e f f e r 2 8 7 " I a t e r a l i s ( A n t e on ) T h om s o n 3 2 5 * I a t e r a l i s ( D ry i n u s ) T h om . 3 2 5 l a t i c ep s ( B o c c h u s ) A s h m e a d 6 5 2 * I a t i c ep s ( P h o r b a s ) A s h m e a d 6 1 2 , 652 * I a t i c ep s ( P h o r b a s i a ) A sh m e a d 612 l a t u s ( D ry i n u s ) O l m i 8 0 8 * I a w a n a e ( N e o d ry i n u s ) R i ch a r d s 1 085 l e i o s omu m ( A n t e on ) R . P e rk i n s 560, 1 845 * l e i o s omu 5 ( An t e on ) R . P e rk i n s 560 * I e i o s omu s ( C h e l o g y n u s ) R . P e r . 560 * I e i o s omu s ( N e o c h e l o g y n u s ) R . P e rk i n s 5 6 0 * I ep i d u s ( A n t eon ) F o r s t e r 2 6 0 * I ep i d u s ( C h e l ogynu s ) F o r s t e r 259 " I ep t i a s ( C h a l c o g o n a t opu s ) R . P e rk i n s 1 7 1 2 l ep t i a s ( D r y i n u s ) R . P e rk i n s 895 , 1 84 5 , 1 846 * I ep t i a s ( P a ra d r y i n u s ) R . P e rk . 895 * L e s t od r y i n i n i 7 2 3 * L ep t od r y i n u s R i c h a r d s 1 0 6 5 * I ep t op u s ( L ep t od ry i n u s ) R i c h . 1 06 5 , 1 076 I ep t op u s ( N e o d ry i n u s ) R i c h a rd s 1 0 76 l ep t o t h o r a x ( G on a t op u s ) C eb a l . 1 05 2 , 1 6 1 5 * L e s t od ry i n u s K i e f f e r 7 2 8 , 7 2 9 , 1 834 l e u c on e u ru s ( A n t e o n ) K i e f . 1 8 1 3 l e u c op u s ( Ap h e l op u s ) K i e f f e r 78 " I eu c o s t omu s ( G on a t op u s ) J . S a h l b e rg 1 4 3 7 l e v i g a t u s ( A n t e on ) K i e f f e r 3 0 9 , 1 81 3 * I ev i g a t u s ( X en a n t e on ) K i e f f e r 1813

I n dex

* 1 i ch t en s t e i n i ( G o n a t o p u s ) P i c a rd 1 5 1 5 i g u s t i c u s ( P s e u d o g on a t op u s ) C u r r a d o and D l m i 1 2 1 7 , 1 2 1 9, 1 839 ':' 1 i n d b e r g i ( D i c o n d y l u s ) H e i k i n h e i mo 1 3 7 9 , 1 3 8 0 i n d b e r g i ( G on a t opu s ) H e l l e n 1 54 1 * 1 i n d b e r g i ( T r i c h og o n a t op u s ) H e I I en

1 50 0 ,

1 54 1

* L i od ry i n u s K i e f f e r 2 8 7 " I j u n g h i i ( D i c on d y l u s ) We s t w . 1 437 * I j u n g h i i ( G on a t op u s ) We s t w o o d 1 437 * I j u n g h i i ( T e t r a d o n t o ch e l y s ) We s t w o od 1 4 3 9 " I j ungh i i ( T e t rodon t oche l y s ) We s t w o od 1 4 3 4 , 1 4 3 7 , 1 4 3 8 l ob a t u m ( A n t e on ) D l m i 5 0 6 L on ch od ry i n u s K i e f f e r 2 5 3 , 1 8 3 1 l on g i c h e l a t u s ( A g o n a t o p o i d e s ) F en t o n 1 3 4 2 * I on g i c h e l a t u s ( D i c o n d y l u s ) F e n t on 1 3 4 2 " I on g i c o l l i s ( D r y i n u s ) K i e f f e r 1 1 44 l on g i c o l l i s ( E c h t h r od e l p h a x ) K i e f f e r 1 1 44 * I on g i c o l l i s ( Me r c e t i a ) K i e f f . 1 1 40 , 1 1 44 * I on g i c o r n i s ( An t e on ) D a l ma n 2 5 9 , 1 80 1 , 1 8 1 7 , 1 8 2 1 * I on g i c o rn i s ( A n t e on ) H e l l e n 1 829 I o n 9 i c o rn i s ( Ap t e r od r y i n u s ) K i e f . 1 36 7 * I on g i c o r n i s ( D i c o n d y l u s ) K i ef . 1 367 * I on g i c o rn i s ( D o l i ch o c h e l y s ) K i ef . 1 34 2 , 1 36 7 * I on g i c o rn i s ( D r y i n u s ) D a l ma n 258 * I on g i c o rn i s ( G o n a t op u s ) K i e f . 1 367 * I on g i c o rn i s ( P h o r b a s ) B r u e s

1 895

1813 * I on g i c o rn i s ( P h o rb a s i a ) B r u e s 1813 * I o n g i c o r n i s ( P r en a n t e on ) D a l m . 260, 261 ':' I on g i f i I i s ( A n t e on ) K i e f . 2 6 0 * l ong i f i i i s ( P renan t eon ) K i ef f 260 * I on g i f o r c ep s ( A n t e on ) K i e f f e r 322, 1 828 •

* I on g i f o r c e p s ( C h e l o g y n u s ) K i ef . 322 l on g i p e s ( D ry i n u s ) D l m i 7 9 9 l on g i s c a p u m ( An t e o n ) C e b a I . 3 9 2 " I on g i s c a p u s ( X en a n t e o n ) C eb a l . 392 l on g i t a r s i s ( An t e o n ) K i e f . 1 8 1 4 * I on g i t a r s i s ( C h e l ogy n u s ) K i e f . 1814 l on g i t a r s i s ( L a b e o ) A s h m . 1 8 1 5 * I on g i t a r s i s ( L a b e r i u s ) A s h m e a d 1 81 5 l on g i u s c u l u s ( G on a t o p u s ) O l m i 1 590 l on g u s ( G o n a t opu s ) P o n oma r e n k o 1 567 * I o n g u s ( N e o g on a t op u s ) P o n o m . 1 567 l u c en s ( D ry i n u s ) D l m i 8 1 7 l u c en s ( T e t r od on t o ch e l y s ) D l m i 1 46 1 , 1 84 0 * I u c i d u m ( A n t e on ) H a l i d a y 3 1 7 " I u c i d u s ( An t eon ) H a l i d ay 3 1 7 * I u c i d u s ( C h a l c og on a t op u s ) R oh w e r 1 6 6 2 * I u c i d u s ( C h e l og y n u s ) H a l . 3 1 7 * I u c i d u s ( D i g on a t op u s ) R oh w e r 1 662 * I u c i d u s ( D r y i n u s ) H a l i d ay 3 1 6 l u c i d u s ( G on a t opu s ) R oh w e r 1 6 6 2 1 840 l u c t u o s u m ( An t e o n ) D l m i 5 7 7 * I u f f n e s s en s i s ( An t a e on ) Ch i t t y 260 l u n a t u s ( G o n a t opu s ) K l u g 1 0 5 5 , 1 54 1 , 1 54 3 , 1 54 4 , 1 8 1 9 , 1 83 5 , 1 836

1 896

Mem . A me r . E n t .

I ns t . ,

n o . 37 ,

1 984

* I u n a t u s ( N e o g on a t op u s ) K l u g 1 46 1 , 1 54 3 , 1 544 * I u su s ( An t e on ) R . P e r k i n s 4 6 3 * I u su s ( C h e l o g y n u s ) R . P e rk i n s

* ma d a g a s c o l a ( X e n an t e on ) B e n o i t 393 mad a g a s c o l u m ( An t e on ) B e n . 3 9 3 * Ma d e c a g o n a t op u s B e n . 1 5 0 0

463 " I u t e i c ep s ( P r e n a n t e on ) K i e f . 261 I u t e i c o rn i 5 ( An t e on ) K i e f f e r 1 81 6 " I u t e i c o rn i s ( C h e l o g y n u s ) K i e f . 1816 ':' I u t e i c o r n i s ( G o n a t op u s ) K i e f . 1515 " I u t e i p e s (An t eon ) K i e f . 2 5 9

* ma d e c a s s u m ( H i r t a n t e o n ) B e n o i t 1 25 mad e c a s s u s ( D e i n o d r y i n u s ) B e n . 1 25 mag n i f i c u s ( Me g a d ry i n u s ) R i c h . 2 8 , 969, 971 , 1 844 ma g n u s ( G on a t op u s ) O l m i 1 6 6 5 ma g n u s ( P s e u d o g o n a t op u s ) B r own 1 275

l u t e i p e s ( G o n a t op u s ) B e n o i t 1 61 1 * I u t e i p e s ( P l a t y g on a l o p u s ) B en o i t 1 6 1 1 " I u t e i p e s ( P r e n a n t e on ) K i e f f e r 260 l u w e l i ( L e s t od r y i n u s ) B en o i t 1 81 6

764 * ma j o r ( A n t e on ) D a l ma n 1 8 1 6 ma j o r ( A n t e on ) K i e f f e r 1 8 1 6 * ma j o r ( D ry i n u s ) D a l ma n 1 8 1 6 ma l a y a n u s ( A p h e l op u s ) O l m i 6 6 ma l a y a n u s ( T h a u ma t od r y i n u s )

" I u z on i c u s

ma l e s i a e

( L e s t od ry i n u s )

K i e f f er 806 * I y d e ( An t e o n ) Wa l k e r 3 0 0 * I y d e ( D ry i n u s ) Wa l k e r 3 0 0 ma c i I en t u s ( T h a u ma t od ry i n u s ) D e San t i s a n d V i d a l Sa r m i en t o 7 0 6 ma cu l i c ep s ( Ap h e l op u s ) B e r g ma n 66, 1 840 " ma c u l i c o rn i s ( D r y i n u s ) C am e r o n 1 1 0 7 ma cu l i c o r n i s ( N e od r y i n u s ) Cameron 1 1 07 " ma cu l i c o rn i s ( P r o d r y i n u s ) C a me r o n 1 1 0 7 " ma cu l i p en n e ( A n t e on ) K i e f f e r 334 " ma c u l i p e n n i s ( An t e o n ) K i e f f e r 334 * mad a g a s c a r i e n s i s ( An t a ph e l op u s ) B e n o i t 6 0 m a d a g a s ca r i e n s i 5 ( D e i n o d ry i n u s ) B en . 1 2 8 * mad a g a s c a r i e n s i s ( H i r t a n t e o n ) B en o i t 1 2 8

ma h e n s i s ( D ry i n u s ) K i e f . 7 6 4 * ma h e n s i s ( Me s od r y i n u s ) K i e f .

O l mi

699 ( P s e u d o g on a t op u s )

0 1 m i 1 248 * ma r g i n a t u m ( An t e o n ) K i e f f e r 295, 299 * ma r g i n a t u s ( A n t e on ) K i e f f e r 295 ma r i t i mu m ( A n t e on ) T u r n e r 357 * ma r i t i mu s ( C h e l ogy n u s ) T u r n e r 357 ma r o c c a n u s ( R i c h a r d s i d r y i n u s ) 0 1 mi 9 1 0 " ma r sh a l l i ( G on a t op u s ) K i e f f e r 1 543, 1 8 1 9 ma s n e r i ( Ap o d ry i n u s ) O l m i 1 7 9 6 ma s n e r i ( B i a p h e l op u s ) O l m i 9 5 , 96 ma sn e r i ( L o n c h od r y i n u s ) O l m i 277 ma s on i ( A n t e on ) O l m ; 4 6 9 * ma u r i t i a n U 5 ( P s e u d o g o n a t op o i d e s ) W i I I . 1 1 9 2 , 1 234 * ma u r u s ( C h a l c o g o n a t op u s ) K i e f . 1 66 0

1 897

I n dex

mau r u s ( G on a t op u s ) K i e f . 1 6 6 0 * mau ru s ( P l a t y g o n a t op u s ) K i e f . 1 66 0

max i ex op h t h a l m i c u s ( T r i d ry i n u s ) O l mi 948 max i mu s ( D e i n od ry i n u s ) O l m i 1 65 ma x i mu s ( T r i d ry i n u s ) O l m i 9 5 4 ma y e t i ( An t e on ) K i e f f e r 1 8 1 7 * may e t i ( G on a t op u s ) K i e f . 1 4 0 9 may og o ( D r y i n u s ) B en o i t 7 8 0 * ma y o g o ( L e s t od ry i n u s ) B e n o i t 786 ma y o r i ( G on a t op u s ) 0 I m i l 6 7 9 med i a l i s ( A n t e o n ) H e l l e n 1 8 1 7 * med i o c a r i n a t u s ( A n t a p h e l op u s ) B en o i t 4 4 , 6 0 med i o ca r i n a t u s ( Ap h e l op u s ) Beno i t 60 med i u s ( G on a t op u s ) O l m i 1 6 6 2 med l e r i ( An t e on ) O l m i 3 9 5 Meg a d r y i n u s R i c h a r d s 9 6 9 * Me g a g on a t op u s O l m i a n d C u r r o 1 50 1 me l a l e u c u s ( Ap h e l op u s ) D a l ma n 47, 1 83 6 , 1 83 7 * me l a l eu c u s ( D ry i n u s ) D a l ma n

( P r en a n t e on ) K i e f .

261

*me l a n o c e r u m ( P r e n a n t . ) K . 2 5 3 * me l a n o s t i g mu s ( An t e on ) R . P e rk i n s 5 2 7 * me l a n o s t i g mu s ( P r o s a n t e o n ) R . P e rk i n s 5 2 7 menk e i ( A n t e o n ) O l m i 4 8 6 menk e i ( Ap t e r o d r y i n u s ) O l m i 1 364

* Me r c e t i a K i e f f e r 1 0 5 3 , me r i d i on a l i s ( Ad r y i n u s ) 1 1 39 me r i d i on a l i s ( G on a t op u s 1 5 90 * m e r i d i on a l i s ( R h y n ch og on a t op u s ) 1 500,

1 1 40

O l mi ) B en o i t

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1 590

Me s od ry i n u s K i e f f e r 7 2 9 , 1 0 0 0 * Me t a g on a t op u s O g l ob l i n 1 5 0 0 Me t a n t e on O l m i 1 1 3 mex i c a n u m ( An t e on ) O l m i 4 9 9 mex i c a n u s ( D r y i n u s ) R . P e rk i n s 823

* mex i c a n u s ( P a ra d ry i n u s ) R . P e rk i n s 8 2 3 * m i c r o l e u c u s ( An t e on ) R . P e rk . 73

47

* me l a l e u c u s

* me l a n o c e r a

( G on a t op u s ) D a l ma n

47

* me l a n a c r i a s ( An t e on ) R . P e rk i n s 4 6 3 * me l a n a c r i a s ( C h e l o g y n u s ) R . P e rk i n s 4 6 3 * me l a n a c r i a s ( P s e u d o g on a t op u s ) R . P e rk i n s 1 2 8 7 me l a n i a s ( G on a l op u s ) R . P e rk i n s 1 7 7 2 , 1 8 4 5 * me l a n i a s ( P a c h y g o n a t op u s ) R . P e rk i n s 1 4 9 9 , 1 7 7 3 * me l a n o c ep h a l u s ( B o c c h u s ) Came r o n 1 6 8 me l a n o c ep h a l u s ( D e i n od r y i n u s ) C ame r o n 1 6 8 * me l a n o c e p h a l u s ( D ry i n u s ) C a me r o n 1 6 8 * me l a n o c e ra ( A n t e o n ) K i e f . 2 5 9

* m i c r opu n c t a t u s ( P r enan t eon ) F e n t on 2 7 8 m i c r o s ( An t e on ) O l m i 5 1 9 * m i c r o t omu s ( Ap h e l op u s ) Wh i t t a k e r 7 2 m i g ra t o r i u s { E c h t h r od e l p h a x Beno i t 1 1 52 m i g ra t o r i u s ( T h a u ma t od r y i n u s ) B en o i t 6 9 4 m i l e s ( An t e on ) O l m i 5 4 7 m i mo i d e s ( G on a t op u s ) R . P e rk i n s 1 69 3

* m i mo i d e s ( N e o g o n a t op u s ) R . P e rk i n s 1 6 9 3 m i mu s ( A c r od on t o c h e l y s ) R . P e rk i n s 1 1 8 3 , 1 8 0 0 * m i mu s ( N e o g on a t op u s ) R . P e rk . 1 1 84

mi ngoae

( An t e on ) O l m i

391

1 898

Mem . Amer . E n t .

m i n i mu m ( An t e on ) F en t on 4 6 7 * m i n i mu s ( A n t e on ) F en t o n 4 6 7 m i n i mu s ( B o c c h u s ) O l m i 6 5 8 * m i n i mu s ( C h e l o g y n u s ) F e n t on 467 * m i n i mu s ( P a c h y g on a t op u s ) F en t on 1 4 6 8 m i n i mu s ( T h a u ma t od r y i n u s ) O l m i 704 m i n o r ( A n t e on ) K i e f f e r 1 8 1 7 m i n o r ( D e i n od ry i n u s ) O l m i 2 4 1 m i n u t e l l u s ( A n t e on ) K i e f f e r 1817 * m i n u t u s ( An t e on ) A sh me a d 4 5 3 , 1817 " m i n u t u s ( An t e o n ) K i e f . 1 8 1 7 m i ra b i l i s ( B o c c h u s ) R . P e rk i n s 6 4 3 , 1 84 3 * m i r a b i l i s ( E u k o eb e l e i a ) R . P e rk i n s 6 1 2 , 6 4 3 * m i r a b i l i s ( Ph o r b a s ) R . P e rk . 643 m i r a b i l i s ( T h a u ma t od ry i n u s ) O l mi 687 m i r i p e s ( A l l od r y i n u s ) K i e f f e r 1 81 7 M i r od r y i n u s P on oma r en k o 6 6 3 * m i s o r ( D r y i n u s ) Wa l k e r 259 mod e s t u s *moes t u s P e rk mo l l e ( An

( D ry ( Hap ins t e on

i nu s ) O l m i 785 l og o n a t op u s ) R . 1 41 8 ) O l mi 492

':' mon g o l i c u m ( An t e on ) Mo c z a r 317 m o n g o l i c u s ( G o n a t op u s ) P on o ma r e n k o 1 5 6 7 mon i I i s ( G on a t op u s ) O l m i 1 7 2 3 * Mon o g on a t op u s R i c h a r d s 1 4 0 8 mon t a n u s ( D ry i n u s ) O l m i 8 7 8 mon t a n u s ( P en t a g on a t op u s ) O l m i 1 4 2 7 , 1 43 0 mon t i c o l u s ( D e i n o d r y i n u s ) O l m i 1 30 m o r e n o i ( P s e u d og on a t op u s ) O l m i 1 284 , 1 8 4 4 " mo r l ey i ( An t a e on ) C h i t t y 2 6 0

I ns t . ,

no . 37,

1 984

m o r t u o ru m ( L e s t od ry i n u s ) B r u e s 1 8 1 7 , 1 830 mo s s e l i ( A n t e on ) O l m i 4 0 8 mo s s e l i ( G o n a t o p u s ) O l m i 1 6 0 3 mu e s eb e c k i mu i r i mu i r i mu l t i mu l t i

( P s eu d o g on a t op u s )

O l m i 1 2 80 ( A n t e on ) O l m i 4 3 2 ( P s e u d o g on a t op u s ) O l m i 1 25 1 c o l o r ( G on a t op u s ) K i e f f e r 1 818 d en s ( D i c o n d y l u s ) O g l ob l i n 1 81 8

mu l u e n s i s ( B o c c h u s ) O l m i 6 3 5 mu n d u m ( An t e on ) a I m l 5 6 5 mu n i t u m ( An t e on ) O l m i 3 4 5 mu n r o e i ( A n t e o n ) O l m i 4 3 9 mu t i l a r i a ( Me t h o c a ) L a t r e i l l e 1 8 1 8 , 1 828 mu t i l a r i u s ( G o n a t op u s ) N e e s 1 81 0, 1 81 8 my r m e c o p h i l u m ( A n t e on ) R . P e rk . 7 , 529, 1 845 * my rme c op h i I u s ( An t e o n ) R . P e rk i n s 5 2 9 " my rme c op h i I u s ( G o n a t o p u s ) K i ef f e r 1 54 1 , 1 543 "'my r m e c op h i l u s ( P a r a n t e on ) R . P e rk i n s 2 8 6 , 5 2 9 my s o r e n s e ( A n t e on ) O l m i 4 2 2 my s o r e n s i s ( G on a t op u s ) O l l11 i 1 64 7 , 1 8 4 0 My s t r op h o r u s F o r s t e r 6 7 8 n a m i b i e n s e ( An t e on ) O l m i 3 8 3 "' n a n u m ( A n t e o n ) H a I i d a y 2 9 5 " n a n u s ( A n t e on ) H a l i d a y 2 9 5 " n an u s ( D r y i n u s ) H a l i d a y 2 9 5 n a p en s i s ( D ry i n u s ) O l m i 8 5 0 n a s u t u s ( E mb o l emu s ) A s h me a d 1 81 8 * n a s u t u s ( L a b e o ) A s h me a d 1 8 1 8 * n a s u t u s ( L a b e r i u s ) A s h mead 1818 n a t a l e n s e ( A n t e on ) O l m i 3 8 8 n a u ma n n i ( G on a t op u s ) O l m i 1 7 8 5 n ea r c t i c a ( C r ov e t t i a ) O l m i 8 5 * n e a r c t i c u s ( P a c h y g on a t op u s )

1 899

I n dex

F e n t on 1 1 8 2 n e a r c t i cu s ( P l e c t r o g o n a t op o i d e s ) D l mi 1 43 3 n e a r c t i c u s ( R i ch a r d s i d ry i n u s ) D l mi 927 n eb u l o s u m ( A n t e o n ) D l m i 4 6 1 n e e s i ( G on a t op u s ) K i e f f e r 1 8 1 9 n eg l e c t u s ( A n t e o n ) K i e f . 3 0 0 , 1 80 4 , 1 81 9 n e l son i ( N eod ry i n u s ) R . P e rk . 1 1 1 8 , 1 84 5 , 1 846 n em o r a l e ( An t e o n ) D l m i 4 4 2 * N e o a n t e o n F ou t s 6 1 2 * N e o ch e l og y n u s R . P e r k i n s 286, 287 N e o d r y i n u s R . P e rk i n s 1 0 5 3 , 1 0 5 4 , 1 06 5 , 1 8 3 2 * N e o g on a t op o i d e s A b d u l -N ou r 1 4 34 * N e o g o n a t op u s R . P e rk i n s 1 0 5 4 , 1 05 5 , 1 49 8 n e o t r op i c a ( C r ov e t t i a ) D l m i 9 2 n e o t r o p i c u s ( P e r od ry i n u s ) D l m i 1 04 4 n e o t r op i c u s ( T r i c h og o n a t op u s ) D l mi 1 487 n ep a l en s e ( A n t eon ) D l m i 3 0 4 n ep a l e n s e ( C o n g a n t e on ) D l m i 98 n ep a l en s i s ( Ap h e l o p u s ) D l m i 5 7 n e p a l e n s i s ( D ry i n u s ) D l m i 7 4 0 n ep a l en s i s ( L on c h od ry i n u s ) D l mi 267 " n i 9 e r ( An t e on ) N e e s 1 8 1 9 * n i g e r ( C h a l c o g o n a t op u s ) F en t on 1 4 9 5 * n i g e r ( D ry i n u s ) K i e f f e r 1 0 0 5 n i g e r ( E sa g on a t o p u s ) F en t on 1 4 9 3 , 1 49 5 , 1 84 2 n i g e r ( Me s od ry i n u s ) K i e f f e r 1 00 0 , 1 005 * n i g e r ( N e o g on a t op u s ) F en t on 1 49 5 n i g e r i a n u s ( Ad r y i n u s ) D l m i 1 1 30 * n i g e r r i mu s ( L a b e o ) F o r s t e r

1 573 n i g r a ( H e t e r o l ep i s ) N e e s 1 8 1 9 * n i g r e l l u s ( D ry i n u s ) B r u e s 1 09 1 *n i

9

r e i I u s ( P s i I od ry i n u s ) B ru e s 1 09 1 n i g r i c a n s ( D e i n o d ry i n u s ) C a me r o n 2 1 1 * n i g r i c a n s ( D ry i n u s ) Cameron 21 1 * n i g r i c a n s ( P a ra g on a t o p u s ) R . P e rk i n s 1 1 9 1 , 1 2 8 7

* n i g r i c a n s ( P l a s t od ry i n u s ) C a me r on 2 1 1 n i g r i c a n s ( P s eu d o g on a t op u s ) R . P e rk i n s 1 2 8 7 n i g r i c o l l i s ( E c h t h r od e l p h a x ) R . P e rk i n s 1 1 6 6 , 1 8 4 6 n i g r i c e p s ( Ap h e l op u s ) K i e f f e r 56, 1 837 n i g r i c o r n e ( An t e o n ) R . P e r k i n s 568, 1 845 * n i g r i c o r n i s ( An t e on ) K i e f f e r 300 ':' n i g r i c o rn i s ( An t e on ) R . P e r k . 568 * n i g r i c o r n i s ( C h e l ogy n u s ) R . P e rk i n s 5 6 8 * n i g r i c o rn i s ( N e o c h e l o g y n u s ) R . P e rk i n s 5 6 8 * n i g r i c o r p u s ( P a r a d ry i n u s ) G i rau l t 8 8 5 n i g r i o r ( T e t r od o n t o ch e l y s ) D l m i 1 458 n i g r i t h o ra x ( D i c ondy l u s ) Dg l ob l i n 1 3 9 0 * n i g r i t h o r a x ( I d o l o g o n a t op u s ) D g l ob l i n 1 3 7 5 , 1 3 9 0 n i g r i v e n t r i s ( G o n a t op u s ) N e e s 1 8 20 * n i g r o c l a v a t u s ( An t e o n ) K i e f f e r 300 n i g r o f emo r a t u s ( A n t e on ) H e l l e n 1 8 20 n i g r o f l a v u s ( T r i d ry i n u s ) D l m i 961 n i g r o l oba t u s ( D e i n o d r y i n u s )

1 900

Mem . A m e r . E n t .

D l mi 1 99 n i g r o ru b rum ( A n t e on ) D l m i n i g r o ru f u s 1 59

494

( D e i n od ry i n u s ) D l m i

n i g r o s i gn a t u s ( G o n a t opu s ) J . Sah l b e rg 1 5 5 4 * n i g r u s ( C h a l c o g o n a t op u s ) F en t on 1 4 9 5 n i t i d e I I u s ( An t e on ) K i e f f e r 1 82 1 " n i t i d u l u s ( An t e o n ) H e l l e n 1 829 n i t i d u m ( A n t e on ) R . P e r k i n s 561 , 1 84 5 " n i t i d u s ( An t e on ) K i e f f e r 5 6 1 , 1 82 1 " n i t i d u s ( C h e l og y n u s ) R . P e rk i n s 5 6 1 ':' n i t i d u s ( N e o c h e I ogy n u s ) R . P e rk i n s 5 6 1 , 1 8 2 1 n i v o s u s ( G on a t op u s ) D l m i 1 5 7 0 * N og a t op u s P o n oma r e n k o 1 3 4 2 * n o t a t u s ( E p i g on a t o p o i d e s ) P on o ma r e n k o 1 4 4 2 n o t o g e i c u s ( Ap t e r o d ry i n u s ) D l mi 1 37 2 n o t og e i c u s ( L o n c h o d ry i n u s ) D I m i 286 n o t og e i c u s ( T r a n s d ry i n u s ) D l m i 1 04 8 n oy e s i ( An t e o n ) D l m i 5 0 3 n oy e s i ( D e i n od ry i n u s ) D l m i 1 7 0 nudu s

( P s e u d og on a t op u s ) R . P e rk i n s 1 24 5 , 1 8 4 1 n y c t e u m ( An t e o n ) D l m i 5 1 0 " ob s c u r a ( M e r c e t i a ) C e b a I l o s 1 1 55 * o b s c u r i c o r n e ( An t e on ) K i e f f e r 300 ':' ob s c u r i c o r n i s ( An t e o n ) K i e f f . 300 * ob s c u r i s s i mu s ( N e o g on a t op u s ) R . P e rk i n s 1 7 1 7 ob s c u r u s ( E c h t h r od e l p h a x ) C eb a l l o s 1 1 5 5 ob s c u r u s ( T e t r o d on t o ch e l y s ) D l mi 1 45 2

I ns t . ,

no. 37,

o c ea n i cu s 1 403 o c h r eu s

1 984

( D i c on d y l u s ) D l m i

( Ap h e l op u s ) D l m i

67

o ch r e u s ( T e t r o d o n t o c h e l y s ) D l m i 1 450 o c u l a t u s ( T r i d ry i n u s ) D l m i 9 4 5 o k a h a n d j a e ( G o n a t opu s ) D l m i 1 62 3 o l i v e i ra i ( An t eon ) D l m i 5 0 2 o l m i i ( R a d i i ma n c u s ) M o c z a r 6 7 6 omb r od e s ( G o n a t o p u s ) R . P e rk i n s 1 49 8 , 1 72 0 , 1 84 2 * omb r od e s ( N e o g o n a t op u s ) R . P e rk i n s 1 7 2 0 o n t a r i a n u s ( P s e u d og o n a t op u s ) D I m i 1 269 * op a c i f r o n s ( A n t e on ) H e l l e n 1 8 29 op a c i f r o n s ( D ry i n u s ) D l m i 8 5 1 o p a c i t h o r a x ( G o n a t op u s ) D l m i 1 745 o p a c u s ( G o n a t opu s ) D l m i 1 6 3 4 o p a c u s ( P s e u d o g on a t op u s ) R . P e rk i n s 1 3 0 2 , 1 8 4 6 * op t a b i l i s ( C h a l c og on a t op u s ) R . P e rk i n s 1 7 8 7 op t a b i l i s ( G o n a t op u s ) R . P e r k . 1 78 6 , 1 84 5 * o ra t o r i u s ( D i condy l u s ) We s t w o od 1 4 0 9 * o ra t o r i u s ( G o n a t op u s ) We s t wo od 1 409 o r a t o r i u s ( H a p l o g on a t op u s ) We s t w o od 1 4 0 9 , 1 8 3 8 * o ra t o r i u s ( Mo n o g o n a t op u s ) We s t w o o d 1 4 0 8 , 1 4 0 9 o rb i t a l i s ( G o n a t op u s ) C a me r on 1 745 o r i e n t a l e ( An t e o n ) D l m i 5 4 1 o r i en t a l i s ( Ap h e l op u s ) D I mi 67 " o r i en t a l i s ( H a p l og o n a t op u s ) R ohwe r 1 4 1 8 o r i e n t a l i s ( Me s od ry i n u s ) D l mi 1 0 1 1 " o r i en t a l i s ( Mo n o g o n a t op u s ) R oh w e r 1 4 1 8

I n dex

1 90 1

or i p h i l u m ( A n t e o n ) K i e f f e r 403 * o r i ph i I u s ( An t e o n ) K i e f f e r 403

* p a l l i d i c o rn i s

* o r m e n i d i s ( D ry i n u s ) A s h m e a d 1 09 1 , 1 833 * o r men i d i s ( P r o d r y i n u s ) A shmead 1 0 9 3 o r op h i I u s ( D ry i n u s ) B en o i t 781 * o r o ph i I u s ( L e s t o d r y i n u s ) B en o i t 7 8 1

* p a l l i d i n e rv i s ( A n t eon ) K i e f f e r 259 p a l l i d i o r ( D i c on d y l u s ) O l m i 1 383 * p a l l i d i t a r s i s ( G o n a t op u s ) Cameron 1 48 1 p a l l i d i t a r s i s ( T r i ch o g on a t op u s ) C a me r o n 1 4 8 1

* o r t h o l ab i s ( D i c on d y l u s ) K i ef f er 1 203 * o r t h o l a b i s ( G on a t op u s ) K i ef f e r 1 20 3 o r t h o l a b i s ( P s e u d o g on a t op u s ) K i e f f e r 1 20 3 , 1 2 0 6 o s b o rn i ( A n t e o n ) F e n t on 4 8 1 , 1 842 * o sb o r n i ( C h e l og y n u s ) F en t on

* p a l l i d u s ( C h l o r od r y i n u s ) R . C . P e rk i n s 7 2 8 , 8 1 0 , 8 7 1 · p a l l i d u s ( D i c on d y l u s ) C eb a l l o s 1 306 p a l l i du s ( D ry i n u s ) R . C . P e rk i n s 8 7 1 , 1 8 46 * p a l u s t r i s ( P r en a n t e on ) O g l ob l i n 2 6 0 * p a l u s t r i s ( P s e u d o g on a t op u s ) R .

481 * o t i a r t es 295

P e rk i n s 1 2 9 8 p a n a m e n s e ( An t e o n ) O l m i

( A n t e on ) Wa l k e r

* o t i a r t e s ( D ry i n u s ) Wa l k e r 294 owen i ( A d r y i n u s ) O l m i 1 1 3 5 * p a c e i ( H ap l og on a t op u s ) C u r r o and O l mi 1 376 * P a ch y g on a t op u s R . C . L . P e rk . 1 499 p a c i f i c u s ( D ry i n u s ) O l m i 8 7 6 p a g l i a n o i ( B o c ch u s ) O l m i 6 2 0 p a l a e od om i n i c a n u s ( D ry i n u s ) Cu r r o and O l mi 866, 1 830 p a l l i d a ( D on i s t h o rp i n a ) C eb . 1 30 6 , 1 8 3 8 , 1 8 3 9 p a l l i d i c e p s ( G o n a t op u s ) R . C . P e rk i n s 1 7 2 9 , 1 8 4 2 * p a l l i d i c e p s ( N e o g on a t op u s ) R . C . L . P e rk i n s 1 7 2 9 * p a l l i d i c o rn i R . P e rk i *pa I l i d i corn i A sh m e a d * p a l l i d i c o rn i K i ef f e r

s ( Ag on a t op u s ) n s 1 500, 1 697 s (An t eon ) 459 , 539 s ( A n t eon ) 260

( C h e l og y n u s )

R . P e rk i n s 5 4 0 * p a l l i d i c o rn i s ( N e o ch e l o g y n u s ) R . P e rk i n s 5 3 9

pan amen s i O l mi p a n a me n s i 251 papu a n u s pap ua n u s

497

s

( A l p h a d ry i n u s ) O l m i 989 s ( D e i n o d ry i n u s ) O l m i ( D ry i n u s ) O l m i 904 ( G o n a t op u s ) O l m i

1 786 p a r a d ox u s ( D e i n od ry i n u s ) R . C . P e rk i n s 1 1 9 , 1 4 5 * P a r a d ry i n u s R . C . L . P e rk i n s 728, 1 832 * P a ra g on a t op u s R . C . L . P e rk i n s 1 191 * p a ra l ep t i a s ( C h a l c o g o n a t op u s ) R . P e rk i n s 1 7 0 9 p a ra l e p t i a s ( G on a t op u s ) R . C . L . P e rk i n s 1 7 0 9 p a r a n e n s i s ( L a b e r i n u s ) O g l ob l i n 1 821 * P a r a n t e on R . C . L . P e r k i n s 2 8 6 p a r a n u s ( D e i n od ry i n u s ) O l m i 236 * p a r c e p u n c t a t u s ( P r e n a n t e on ) K i ef f er 260

1 902

Mem.

Amer . E n t .

I nst . ,

no . 37 ,

1 984

pa r t h i c u s ( B o c c h u s ) P on oma r en k o 6 2 4 , 6 2 5 " p a rv i c o l l i s ( An t e on ) K i e f f e r 259

1 86 * p ek t u s a n en s e ( P r e n a n t e on ) Mo c z a r 2 6 1 p e l o p s ( E ch t h r od e l p h a x ) O l m i

" p a rv i c o l l i s ( P r e n a n t e on ) K i ef f e r 260 p a rv u l u m ( An t e o n ) R . C . L . P e rk i n s 5 3 9 , 1 8 4 5 * p a rv u l u m ( C h e l o g y n u s ) R . C . L . P e rk i n s 5 4 0 * p a r v u I u s ( An t e on ) K i e f f e r 337, 539

1 1 69 p e n a n g a n u s ( Ap h e l op u s ) O l m i 6 8 " p e n i d a s ( A n t eon ) Wa l k e r 3 1 7 * p e n i d a s ( D r y i n u s ) Wa l k e r 3 1 6 P e n t a g on a t op u s O l m i 1 4 2 7 " p e rd eb i I i s ( C h a l c o g on a t op u s ) R . P e rk i n s 1 4 9 7 p e rd e b i l i s ( E s a g o n a t op u s ) R . C .

* p a rv u l u s ( A n t e on ) R . C . L . P e rk i n s 5 4 0 * p a r v u l u s ( N e o c h e l ogy n u s ) R .

P e rk i n s 1 4 9 7 p e rk i n s i ( D i c on d y l u s ) A s h mead 1 39 4 , 1 83 3 , 1 846 p e rk i n s i ( E c h t h r o d e l p h a x ) D od d 1 82 1 " p e rk i n s i ( G on a t op u s ) A s h m e a d 1 394 p e rk i n s i ( L e s t od ry i n u s ) K i e f f e r 1 82 2 * p e rk i n s i ( P s eu d og o n a t o p u s ) A shmead 1 3 9 4 , 1 8 3 3 p e rk i n s i ( T h a u ma t od r y i n u s ) K r o mb e i n 7 0 0 p e r k i n s i e l l a v o ru s ( P s e u d og on a t op u s ) B en o i t 1 224, 1 839 p e r l u c e n s ( D e i n od ry i n u s ) O l m i 233 p e rm i r u m ( An t e o n ) O l m i 5 8 3 P e r od ry i n u s R . C . L . P e r k i n s 1 03 3 * p e r p o l i t u s ( G i c on d y l u s ) R . C . P e rk i n s 1 6 5 0 p e rp o l i t u s ( G on a t op u s ) R . C . L . P e rk i n s 1 6 5 0 p e ru v i a n u s ( D e i n od r y i n u s ) O l m i

P e rk i n s 5 3 9 p a rv u s ( A l p h a d ry i n u s ) O l m i 989 " p a rv u s ( An t e on ) K i e f f e r 3 3 8 p a s t o r a l i s ( G o n a t op u s ) O g I ob l i n 1 8 2 1 p a t r i z i i ( G o n a t op u s ) B e n o i t 1 6 34 p a u l i a n i ( L on ch od r y i n u s ) B en o i t 2 7 1 * p a u l i a n i ( P r e n a n t e on ) B e n o i t 271 " p e c u l i a r i s ( G o n a t op u s ) B ru e s 1 470 p e cu I i a r i s ( T e t r o d on t o c h e I y s ) B ru e s 1 4 7 0 * p ed e s t r i s ( D i c o n d y l u s ) D a l ma n 1 4 3 7 , 1 5 1 8 * p ed e s t r i s ( G on a t op u s ) D a l ma n 1 437 " p ed e s t r i s ( P a ch y g on a t op u s ) D a l ma n 1 4 3 7 " p ed e s t r i s ( P l a t y g on a t op u s ) D a l ma n 1 4 3 7 , 1 4 3 8 p ed e s t r i s ( T e t r o d on t o c h e l y s ) D a l ma n 1 4 3 4 , 1 4 3 7 , 1 4 3 9 , 1 8 36 p ed u n c u l a t u s ( B o c c h u s ) N a g y 636, 639 p e g n a i ( A l p h a d ry i n u s ) O l m i 991 p e g n a i ( D e i n od ry i n u s ) O l m i

1 74 p e ruv i a n u s 1 738 p e ru v i an u s 1 1 07

( G on a t opu s ) O l m i ( N e od r y l n u s ) O l m l

p e t e r s en l ( An t eon ) O l m i 4 4 5 p e t e r s e n l ( Me s od r y i n u s ) O l m i 1015 * P h a n e r od ry i n u s R o epk e 1 0 6 5

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ph i I i p p i n u m ( An t e o n ) O l m i 4 4 8 p h i I i p p i n u s ( Ap h e l op u s ) O l m i 69 p h i I i p p i n u s ( G o n a t op u s ) O l m i 1 656 p h i I i p p i n u s ( T h a u ma t od ry i n u s ) O l mi 697 * P h o r b a s A sh mead 6 1 1 * P h o rb a s i a K i e f f e r 6 1 2 p i c e i p e s ( Ap h e l o p u s ) K i e f f e r 1 82 2 * p i c e s c en s ( A g o n a t op u s ) R . C . P e rk i n s 1 7 0 3 p i c e s c en s ( T r i d ry i n u s ) O l m i 965 p i c eu s ( A l p h a d ry i n u s ) O l m i 996 p i l i c o r n e ( A n t e on ) O g l ob l i n 51 5 *p i

i c o rn i s ( A n t e o n ) Og l ob l i n 515 '� p i l i c o r n i s ( Aph e l op u s ) Wh i t t a k e r 7 2 p i l o s i f r o n s ( D e i n od ry i n u s ) O l mi 204 p i l o s i p e s ( G o n a t op u s ) B r u e s 1 595 " p i l o su s ( D e i n od ry i n u s ) F en t on 1 3 6 ';' p i l o s u s ( G o n a t o p u s ) T h om s o n 1 573 p i l o s u s ( P s eu d og on a t op u s ) B en o i t 1 2 2 2 p l a n i c e p s ( G on a t op u s ) K i e f f e r 1 5 3 8 , 1 554 * p l a n i c e p s ( P l a t y g on a t op u s ) K i ef f e r 1 499 , 1 53 8 , 1 554, 1 556 p l a n i p e s ( G on a t o p u s ) O l m i 1 689 p l a n u s ( A l p h a d ry i n u s ) O l m i 998 * P l a s t od ry i n u s K i e f f e r 7 2 8 * P l a t y g on a t op u s K i e f f e r 1 4 9 9 p l a u ma n n i ( An t e o n ) O l m i 5 1 7 p l a u ma n n i a n a ( C r ov e t t i a ) O l m i 92

1 903

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( D i c on d y l u s ) R . C .

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p l eb e i u s ( G on a t o p u s ) R . C . L . P e rk i n s 1 6 4 7 P l e c t r o g on a t op o i d e s P o n oma r e n k o 1 43 1 * P l e c t r o g on a t op u s R i ch a rd s 1 5 0 0 * p l e s i u s ( E p i g o n a t op u s ) F en t o n 1 465 p i e s i u s ( T e t rod on t o c h e I y s ) F en t on 1 46 5 , 1 8 4 2 * p l e s u i s ( E p i g on a t op u s ) F e n t on 1 465 p l i c a t u s ( G on a t opu s ) O l m i 1 7 8 3 p l u mb e u s ( G on a t opu s ) O l m i 1 5 3 5 p l u r i s u l c a t u s ( P s e u d o g on a t op u s ) 0 1 m i 1 299 * p o e c i l op t e r a e ( Me s od ry i n u s ) R i c h a rd s 9 5 2 p o e c i l op t e r a e ( T r i d r y i n u s ) R i c h a rd s 9 5 2 , 1 8 4 4 " p o l i t u s ( An t e o n ) A s h m e a d 4 5 8 , 459 p o l i t u s ( D e i n od ry i n u s ) A s h m e a d pol l

206 ( P s e u d o g on a t op u s ) B e n o i t

1 226 * p o l y c h r omu s ( C h a l c og o n a t op u s ) Ma r sh a l l 1 5 2 7 * p o l y c h r omu s ( G o n a t opu s ) Ma r sh a I I 1 5 2 7 * p o l y c h r omu s ( P a c h y g on a t op u s ) Ma r sh a I I 1 5 2 7 * p o l y c h r omu s ( P l a t y g on a t op u s ) 1 527, 1 562 p on oma r en k o a e ( M i r o d ry i n u s ) O l mi 674 " p on oma r e n k o i ( P s e u d o g o n a t op u s ) Mo c z a r 1 2 3 8 p op e n o e i ( An t e on ) A sh m e a d 4 5 2 , 453 ':' p op e n o e i ( D r y i n u s ) A s h m e a d 4 5 2 p op ov i ( G o n a t opu s ) P on oma r e n k o 1 523, 1 836 p o r t a l en s i s ( G on a t op u s ) O l m i 1 679 p ra e c l a ru s ( D ry i n u s ) O l m i 8 1 4

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*p r eh en s o r ( C h e l og y n u s ) Ma n e v a l 3 2 9 *P renan t eon K i e f f e r 2 5 3 ,

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p r e t o r i an u s ( D ry i n u s ) O l m i 767 p r i e s n e r i ( P s e u d o g on a t o p u s ) O l mi 1 2 1 6 p r i mi t i vu s ( D i condy l u s ) O l mi 1 40 1 P r i o r a n t e on O l m i 5 8 9 * P r i s t o g on a t o p u s K i e f f e r 1 1 9 2 * p r o c e r i c o r n i s ( An t e o n ) K i ef f er 259 " p r o c e ru s ( A l l og on a t opu s ) Hau p t 1 50 0 , 1 5 1 5 * P rod ry i n u s K i e f f e r 1 0 6 5 p r op od e a l e ( A n t e o n ) F e n t on 506 " p r o p od ea l i s ( C h e l og y n u s ) F en t on 5 0 6 * P r o s a n t e on R . C . L . P e rk i n s 286 * p r o x i mu s ( A n t e on ) K i e f f e r 259 p s eu d o a mo e n u s ( D e i n od ry i n u s ) 0 1 mi 1 88 p s eu d o a u d a x ( T e t r o d on t o c h e l y 0 1 m i 1 44 6 p s e u d o b i l o b u s ( D e i n od r y i n u s ) 0 1 m i 1 93 " p s e u d o c h r a mu s ( C h a l c o g on a t o p u s ) R . C . L P e rk i n s 1 4 9 9 , 1 7 1 2 * p s eu d o c h r omu s ( E u g on a l op u s ) R . P e rk i n s 1 4 9 9 , 1 7 1 2 p s eu d o c h r o mu s ( G on a t o p u s ) R . P e rk i n s 1 7 1 2 , 1 8 4 2 p s eudod i f f u su s ( N eod ry i n u s ) O l m i 1 086 * P s e u d og on a t o p o i d e s W i I I i am s 1 1 92 P s eu d og on a t o p u s R . C . P e rk i n s 1 1 91 , 1 1 9 2 , 1 83 1 , 1 83 2 , 1 834 * p s e u d op h a n e s ( C h l o r od ry i n u s R . P e rk i n s 8 7 8 p s eu d o ph a n e s ( D ry i n u s ) R . C . L

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P e rk i n s 8 7 8 p s eu d o r b i t a l i s ( G o n a t op u s ) O l m i 1 74 2 * P s i l a n t e on K i e f f e r 2 5 3 ':' P s i l od ry i n u s K i e f f e r 1 0 6 5 * P t e r o g on a t op u s A b d u l - N ou r 1 1 40 p u b i c o r n e ( An t e o n ) D a l ma n 2 8 9 , 3 1 3 , 316, 3 1 7, 322 * p u b i c o r n i s ( An t e on ) D a l ma n 3 1 7 * p u b i c o rn i s ( G on a t op u s ) D a l ma n 316 pu l ch e r ( D ry i nu s ) O l m i 7 9 9 * p u l c h e r r i mu s ( A p h e l op u s ) R . C . P e rk i n s 7 3 p u l c h e r r i mu s ( G on a t op u s ) R . C . L . P e rk i n s 1 7 9 1 , 1 8 4 5 " p u l c h e r r i mu s ( N e o g o n a t op u s ) R . P e rk i n s 1 7 9 1 " p u l i c a r i u s ( E p i g on a t op o i d e s ) K l ug 1 44 2 p u l i c a r i u s ( T e t r od on t o ch e l y s ) K l ug 1 44 2 , 1 83 5 * p u n c t a t u s ( A l l a n t e on ) K i e f f e r 287 * p u n c t a t u s ( An t e o n ) K i e f f e r 325, 1 8 1 6 * p u n c t a t u s ( C h e l og y n u s ) K i e f f e r 327 p u n c t a t u s ( D ry i n u s ) O l m i 8 9 3 " p u n c t a t u s ( G on a t o p u s ) F e n t o n 1 720 p u n c t i c e p s ( An t e o n ) A s h m e a d 474 , 4 7 5 p u n c t u l a t u s ( An t e o n ) K i e f f e r 1 822 p u rp u r a s c en s ( G o n a t o p u s ) O l m i 1 746 p u sa n u s ( P s eu d o g on a t op u s ) O l m i 1 255 * p u s i l l u s ( D i c on dy l u s ) 5zep l i g e t i 1 5 77 * pu s i I l u s ( La b e o ) Szep l i g e t i 1 573 * pu s i I l u s ( La b e r i u s ) S z e p l i g e t i 1 577 * p y o n g a n e n s i s ( A n t e on ) Mo c z a r

1 905

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* p y r en a i cu s ( P r e n a n t e o n ) K i ef f e r 260 p y r i I l a e ( A g on a t o p o i d e s ) Ma n i 1 336, 1 84 1 * p y r i I l a e ( L e s t od ry i n u s ) K i ef f er 9 2 5 , 1 834 * p y r i I l a e ( P s e u d o g o n a t op u s ) Man i 1 3 3 7 p y r i I l a e ( R i c h a rd s i d ry i n u s ) K i e f f e r 925, 1 83 4 , 1 84 1 q u a t e i ( D ry i n u s ) O l m i 8 9 7 q u e ch u a n u s ( T r i d r y i n u s ) O l m i 965 q u e r c e u s ( A ph e l o p u s ) O l m i 5 9 * q u e r c i c o l a ( D e i n od ry i n u s ) R . P e rk i n s 1 4 3 q u e rc i co l u s

( D e i n od ry i n u s ) R . P e rk i n s 1 4 3 r a d i a l i s ( An t e on ) K i e f f e r 1 822 r ad i a l i s ( N e od r y i n u s ) O l m i

1 1 04 R a d i i ma n c u s Mo c z a r 6 7 5 * ra p a x ( C h a l c o g on a t op u s ) R . C . P e rk i n s 1 7 1 5 rap ax ( D e i n o d ry i n u s ) D l m i 2 2 3 * ra p a x ( D r y i n u s ) H a l i d a y 1 8 2 9 r a p a x ( G on a t o p u s ) R . C . L . P e rk i n s 1 7 1 5 * ra p t o r ( C h a l c o g on a t op u s ) R . P e rk i n s 1 4 8 5 * rap t o r ( N eod ry i n u s ) R . C . L . P e rk i n s 1 1 1 8 r a p t o r ( T r i c h o g o n a t op u s ) F en t on 1 0 5 2 , 1 4 8 5 * ra p t o r i p e s ( G on a t op u s ) 5 t rand 1 5 4 3 * r e c t u s ( A n t e on ) K i e f f e r 2 9 5 r e t i c u l a t i c ep s ( A n t e o n ) D od d 531 , 532 r e t i c u l a t u m ( An t e o n ) K i e f f e r 304 * r e t i c u l a t u m ( X en a n t e on ) K i ef f e r 2 8 7

* r e t i c u l a t u s ( An t e o n ) K i e f f e r 304 r e t i c u l a t u s ( N e o d ry i n u s ) F ou t s 1 076 * r e t i c u l a t u s ( P s i l od r y i n u s ) F ou t s 1 0 7 6 * r e t i cu l a t u s ( X e n a n t e on ) K i ef f er 304 * r e t u s u s ( An t e o n ) T h om s o n 2 6 0 * r e t u s u s ( D r y i n u s ) Th oms o n 2 5 9 rh a e n s i s ( G o n a t op u s ) P on o ma r enk o 1 5 8 2 , 1 8 3 6 * R h y n c h o g on a t op u s B en o i t 1 5 0 0 r i ch a r d s i ( Ap t e r o d ry i n u s ) D l m i 1 34 6 r i c h a r d s i ( B o c ch u s ) D l m i 6 2 8 r i c h a rd s i ( C h e l a n t e on ) D l m i 1 0 5 * r i c h a rd s i ( G on a t op u s ) Mo c z a r 1 539 * r i c h a rd s i ( P l a t y g on a t op u s ) D l m i a n d C u r ra d o 1 5 3 8 * r i c h a rd s i ( P I e c t r o g o n a t op u s ) Moc z a r 1 5 1 8 , 1 5 1 9 , 1 5 3 8 , 1 539 r i c h a rd s i ( P r i o r a n t e on ) D l m i 594 r i c h a rd s i ( T r i ch o g o n a t op u s ) D l mi 1 476 R i ch a rd s i d ry i n u s Mo c z a r 9 0 7 r i d e n s ( G on a t op u s ) D l m i 1 6 0 7 r i v u l a r i s ( D e i n od ry i n u s ) D l m i 239 * r ob u s t a ( An t e on e l l a ) D od d 6 1 2 , 660 robu s t u s ( B o c chu s ) D odd 6 6 0 " r o b u s t u s ( D i c o n d y l u s ) C eb a l l o s 1 21 3 r o b u s t u s ( N e od ry i n u s ) D l m i 1 08 2 r o s e l l a e ( P s e u d o g on a t op u s ) C u r r o and D l mi 1 1 99 , 1 8 38 r o s s i ( G o n a t op u s ) D l m i 1 7 8 3 * r o s s i cu s ( G on a t opu s ) P on oma r enk o 1 5 1 5 ru b e r ru b e r

( A g on a t op o i d e s ) 1 3 2 7 ( Me s od ry i n u s ) D l m i 1 031

1 906

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* ru b r i c a ( N e o a n t e o n ) F o u t s 6 1 2 , 639 ru b r i c ep s ( T r i c h og o n a t o p u s ) K i e f f e r 1 47 4 , 1 488 ru b r i c u s ( B o c c h u s ) F o u t s 6 3 6 , 639 " ru b r i f r o n s ( A n t e o n ) K i e f f e r 313 * ru b r i f r o n s ( L a s i a n t e o n ) K i ef f e r 287 r u b r i p e s ( T e t r o d o n t o ch e l y s ) O l m i 1 4 54 ru b r i t h o r a x ( G on a t opu s ) B en o i t 1 6 0 5 "' ru b r i t h o r a x ( N e og on a t o p u s ) B en o i t 1 6 0 5 ru b r o l o b a t u s ( D e i n od ry i n u s ) 0 1 m i 1 97 ru e n s ( P s eu d og on a t op u s ) O l m i 1 22 8 * r u f e s c en s ( G on a t o p u s ) H e l l e n 1 3 7 6 , 1 82 9 * r u f i c a u d a ( Me s od ry i n u s ) R i chards 951 ru f i c a u d a ( T r i d r y i n u s ) R i c h a rd s 9 5 1 , 1 8 4 4 " ru f i c e p s ( D ry i n u s ) C a me r o n 958 * ru f i c e p s ( P r o d ry i n u s ) C a me r o n 9 5 8 ru f i c e p s ( T r i d ry i n u s ) C a me r on 958 ru f i c o l l i s ( A n t e o n ) K i e f f e r 1 822 * ru f i c o l l i s ( C h e l o g y n u s ) K i ef f e r 1 822 " ru f i c o l l i s ( G on a t op u s ) H e l l en 1 55 4 "' ru f i c o r n e ( P r e n a n t e o n ) D a I ma n 2 6 1 " ru f i c o r n i s ( An t e o n ) D a l ma n 2 5 9 * ru f i c o r n i s ( G on a t opu s ) D a l ma n 2 5 8 ru f i c o r n i s ( L on ch od r y i n u s )

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D a l ma n 2 5 8 , 1 8 3 5 , 1 8 3 6 , 1 837 * r u f i c o r n i s ( P r e n a n t e on ) D a l ma n 261 ru f i s c a p u m ( An t e o n ) D od d 5 8 6 " r u f i s c a p u s ( An t e on ) D od d 5 8 6 ru f i v e n t r i s ( Ap h e l opu s ) A shmead 7 0 "' r u f oma c u l a t u m ( A n t e o n ) H e l l e n 1 829 ru f on i g e r ( C h e l o g y n u s ) O g l ob l i n 1 822 ru f on i g ru m ( An t e o n ) O l m i 4 0 2 ;' r u f ov a r i e g a t u s ( C h e l o g y n u s ) Ber l and 3 1 3 "' ru f u l o c o l l e ( An t e on ) C h i t t y 334 " r u f u l o c o l i s ( An t a e o n ) C h i t t y 334 " r u f u l o c o l i s ( C h e l og y n u s ) Ch i t ty 334 ru f u l u s ( Ap t e r od ry i n u s ) O l m i 1 34 7 ru f u s ( A l p h a d r y i n u s ) O l m i 9 9 5 ru f u s ( D i c on d y l u s ) F ou t s 1 3 9 8 ru f u s ( E ch t h r o d e l p h a x ) O l m i 1 1 57 * r u f u s ( P a r a g on a t opu s ) O l m i 1 228 " r u f u s ( P s e u d o g o n a t op u s ) F o u t s 1 228, 1 398 " r u f u s ( P s e u d og on a t op u s ) P on oma r en k o 1 3 7 6 ru f u s

( T h a u ma t od r y i n u s ) R i c h a rd s 7 1 0 , 1 8 4 4 r u g i d o r s a l i s ( Ad r y i n u s ) T u rn e r 1 1 3 5 "' r u g i d o r s a l i s ( N e od r y i n u s ) T u r" n e r 1 1 3 5 * ru g i d o r s a l i s ( P s i l od r y i n u s ) T u rn e r 1 1 3 5 ru g o s i c ep s ( An t e o n ) K i e f f e r 47 1 , 4 7 3 * ru g o s u s ( An t e o n ) A s h m e a d 1 3 3 "' r u g o s u s ( P s eu d og o n a t op u s ) F ou t s 1 3 9 8 * r u g o s u s ( T r i s a n t e on ) A sh mead

1 907

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( An t e o n ) F e n t on

* ru gu l o s u s ( C h e l o g y n u s ) F en t o n 4 8 1 * ru t sh u r i c u m ( X e n a n t e on ) Beno i t 384 s a c ch a r e t a r u m ( P s eu d o g on a t op u s ) R . C . L . P e rk i n s 1 29 4 , 1 8 4 6 ;'� 5 a c c h a r i v o r a e ( P s e u d og on a t op u s ) R i ch a rd s 1 2 7 6 * s a k a i i ( E p i g on a t o p u s ) E s a k i a n d H a s h i mo t o 1 4 4 2 s a k a i i ( T e t r o d o n t o ch e l y s ) E s . a n d H a sh . 1 4 4 1 , 1 8 3 6 san c t i v i n c e n t i ( La b e o ) A shmead 1 8 2 2 * sa n c t i v i n c e n t i ( L a b e r i u s ) A shmead 1 8 2 2 sand e r i ( D ry i n u s ) O l m i 7 3 1 , 1 8 39 s a rawak en s i s ( P s e u d og on a t op u s ) 0 1 m i 1 24 1 s a rawak i ( An t e o n ) O l m i 4 2 5 s a r a wa k i ( P s e u d o g on a t op u s ) Mo c z a r 1 2 3 8 , 1 8 4 1 s a u s s u r e i ( D ry i n u s ) C eb a l l o s 770 * sa u s s u r e i ( L e s t od ry i n u s ) C eb a I l o s 7 7 0 · sa u t e r i ( D i c on dy l u s ) S t rand 1 29 1 ':' s a u t e r i ( G on a t o p u s ) S t r a n d 1 29 1 * sax e t an u s ( L a b e o ) K i e f f e r 764 '� sa x e t a n u s ( L a b e r i u s ) K i e f f e r 764 s c a p u l a r e ( A n t e o n ) H a l i d ay 289 , 322 " s c a p u l a r i s ( An t e o n ) H a l i d ay 322 * s capu l a r i s ( Ch e l ogynu s ) H a l i d ay 3 2 2 " s c a p u l a r i s ( D ry i n u s ) H a l i d a y

322 s c a r a mo z z i n o i ( B o c c h u s ) O l m i 621 * s ch a e f f e r i ( Ch e l ogynu s ) B ru e s 1 36 * s c h a e f f e r i ( 0 e i n od r y i n u s ) B ru e s 1 3 6 s c h en k l i n g i ( G o n a t op u s ) S t r a n d 1 5 8 7 , 1 64 5 , 1 8 4 0 s c h l i n g e r i ( D e i n od ry i n u s ) O l m i 230 s c h ou t e d en i ( D ry i n u s ) B en o i t 778 * s c h o u t e d e n i ( L e s t od ry i n u s ) B en o i t 7 7 8 s c i op h a n e s ( G o n a t op u s ) Ma r sh a l l 1 8 2 4 s c ob i o l a e ( B o c c h u s ) N a g y 6 2 2 , 624 * s c o t i c u m ( An t e on ) K i e f f e r 295 * s c o t i cu s ( An t eon ) K i e f f e r 295 * s c u t e l l a r i s ( G o n a l op u s ) Beno i t 1 63 1 s 2 c u n d u s ( Ap t e r od ry i n u s ) F en t on 1 3 5 5 , 1 8 4 3 " s e cu n d u s ( E u c a mp t o n y x ) F en t on 1 3 5 5 s ed l a c ek i ( A n t e on ) O l m i 5 3 4 s ek e r a e ( A n t e on ) O g l o b l i n 1 8 2 4 " s erilen ov i ( P r e n a n t e on ) P o n oma r en k o 2 6 1 " s em i p o l i t a ( N e od ry i n u s ) G i rau l t 8 8 5 " s ep a r a t u s ( P s e u d og on a t op u s ) R i cha rd s 1 2 1 1 , 1 2 1 7 , 1 2 1 9 " s ep s o i d e s ( D i c o n d y l u s ) We s t w o od 1 5 7 7 s ep s o i d e s ( G o n a l o p u s ) We s t w o od 1 0 5 3 , 1 564 , 1 573, 1 57 7 , 1 8 0 4 , 1 8 3 5 , 1 8 36 " s ep t emd en t a t u s ( G o n a t op u s ) J . Sah I b e rg 1 2 1 7 s e p t emd en t a t u s ( P s e u d o g on a t op u s ) J . Sah l b e rg 1 2 1 7 , 1 8 3 8

1 908

Mem . Amer .

En t .

sep t e n t r i on a l i s ( G on a t op u s ) Wh i t t ak e r 1 8 2 4 * s e r i c eu s ( An t e o n ) K i e f f e r 309 s e r r a t u s ( Ap h e l op u s ) R i ch a rd s 52, 1 83 6 , 1 8 37 * s e r r a t u s ( C h e l og y n u s ) Ma n ev a l 3 1 7 s e r ra t u s ( T r i d r y i n u s ) D l m i 94 1 " s e y r i 9 i ( Ag on a I op o i d e s ) B en 0 i t 1 2 2 5 s ey r i g i ( B o c c h u s ) B en o i t 6 2 9 s e y r i g i ( D ry i n u s ) B e n o i t 7 5 6 * s ey r i g i ( E u c a mp t o n y x ) C eb a l l o s 1 5 9 8 s ey r i g i ( G on a t op u s ) C eb a l l o s 1 598 * s ey r i g i ( H i r t a n t e on ) B en o i t 269 " s e y r i g i ( H y men od r y i n u s ) B en o i t 6 1 2 , 6 2 9 s e y r i g i ( L on c h od r y i n u s ) B en o i t 2 6 9 s ey r i g i ( P s e u d o g o n a t op u s ) B en o i t 1 2 2 5 * s ey r i g i ( T r i c h o g on a t o p u s ) C eb a l l o s 1 5 9 8 * s i b i r i c u s ( D i c on d y I u s ) P on oma r en k o 1 3 7 6 s i e r r a n u s ( D ry i n u s ) D l m i 7 6 6 ':' s i l v a l i c u s ( C h e l og y n u s ) P on oma r e n k o 3 2 9 s i l v e s t r e ( An t e on ) K i e f f e r 444 s i l v e s t r i i ( G o n a t op u s ) K i e f . 1 759 s i I v i c o l u m ( An t e on ) D l m i 4 1 9 * s i m i l i s ( An t e o n ) K i e f f e r 3 3 7 " s i m i l i s ( G on a l op u s ) B r u e s 1 234 ':' s i m i l i s ( G on a t o p u s ) F en t on 171 7 s i m i l i s ( P s e u d o g on a t opu s ) B ru e s 1 2 3 4 , 1 2 5 8 , 1 8 3 9 ':' s i m i I i s ( P s e u d o g o n a t op u s ) F r ey t a g 1 2 3 4 , 1 2 5 8

I ns t . ,

no.

37,

1 984

s i mp l i c i p e s ( D r y i n op s i s ) B r u e s 1 824 s i mu l a n s ( L a b e o ) A s h m e a d 1 8 2 4 * s i mu l a n s ( L a b e r i u s ) A s h m e a d 1 824 * s i mu l a t o r ( C h a l c o g o n a t o p u s ) R . C . P e rk i n s 1 4 7 5 s i mu l a t o r ( T r i ch o g on a t op u s ) R . P e rk i n s 1 4 7 5 s i n e n s i s ( A c r o d on t o c h e l y s ) O l m i 1 1 80 s i n e n s i s ( L o n c h od ry i n u s ) O l m i 273 s i n op en s i s ( D r y i n u s ) D l m i 8 5 8 * s i s i t h ru s ( D r y i n u s ) Wa l k e r 2 9 5 * s i s y t h ru s ( An t e o n ) Wa l k e r 2 9 5 s i t a e ( G o n a t op u s ) P o n o ma r en k o 1 66 5 , 1 666 * s j o e s t ed t i ( G o n a t op u s ) K i e f f e r 1 263 s j o e s t ed t i ( P s e u d og o n a t op u s ) K i e f f e r 1 26 3 , 1 8 4 3 s l ov a c u s ( B o c ch u s ) S t r e j c ek 6 1 8 " s o c i a b i I i s ( G o n a t op u s ) K i e f f e r 1 577 s o g a t a e ( P 5 e u d o g on a t op u 5 ) R oh w e r 1 2 5 4 , 1 8 4 1 * s o g a t ea ( P s e u d o g o n a t op u s ) R oh w e r 1 2 5 4 s o l a r i s ( M e s od r y i n u s ) O l m i 1 0 24 5 0 1 i d u s ( Ag o n a t o p o i d e s ) H a u p t 1 3 1 6 , 1 838 " s o l i d u s ( A l l og o n a t op u s ) H a u p t 1 316 * s o l i t a r i u s ( C h a l c o g o n a t op u s ) R . C . L . P e rk i n s 1 7 0 3 s o l i t a r i u s ( E p i g on a t op u s ) R . C . P e rk i n s 1 4 9 0 , 1 4 9 3 s o l i t a r i u s ( G o n a l op u s ) R . C . L . P e rk i n s 1 7 0 3 s o m e r s e t i ( Ap t e r od ry i n u s ) 0 1 m i 1 345 s omn i a t u s ( N e od r y i n u s ) B ru e s 1 82 5 , 1 830 s p a n g l e r i ( D ry i n u s ) O l m i 7 6 2 sp e c i o s u s ( D e i n od ry i n u s ) O l m i

1 909

I n dex

1 57 sp e c i o s u s

( D ry i n u s ) D od d 8 7 4

* sp e c i o s u s ( P a ra d r y i n u s ) D od d 874 * sp e c t r u m ( D ry i n u s ) S n e l l en v a n V o l I e n h ov en 1 5 6 0 s p e c t r u m ( G on a t o p u s ) Sn e l l e n v a n V o l l e n h ov e n 1 5 6 0 , 1 83 5 , 1 836 * s p e c t r u m ( N e og on a t opu s ) Sn e l l en v a n V o l l e n h o v en 1 560 s p i n o l a i ( G on a t op u s ) D l m i 1 739 s p i r a cu l a r i s ( P s e u d o g on a t op u s ) Beno i t 1 233 s t a n t on i ( D r y i n u s ) A sh m e a d 806, 1 8 3 3 , 1 8 4 1 * s t a n t on i ( L e s t od r y i n u s ) A shmead 8 0 6 * s t a n t on i ( N e o d r y i n u s ) A sh m e a d 8 0 6 * s t a n t on i ( P r od ry i n u s ) A shmead 8 0 6 s t e n o c r a n i ( P s e u d o g on a t op u s ) R . C . P e rk i n s 1 2 7 1 , 1 8 4 3 * s t r i a t i c ep s ( P r o d r y i n u s ) K i ef f e r 956 s t r i a t i c ep s ( T r i d r y i n u s ) K i e f f e r 956 * s t r i a t u s ( Ag on a t o p o i d e s ) K i ef f er 1 5 1 8 , 1 5 1 9 s t r i a t u s ( D r y i n u s ) F en t on 8 4 5 s t r i a t u s ( G on a t o p u s ) K i e f f e r 1 052 , 1 5 1 8 , 1 835 * s t r i a t u s ( L e s t od ry i n u s ) F en t on 8 4 5 * s t r i a t u s ( P l e c t r o g on a t op u s ) K i e f f e r 1 50 0 , 1 5 1 8 s u a r e z i ( G on a t op u s ) D l m i 1 5 2 8 * su b a p t e r u s ( An t e o n ) K i e f f e r 256 s u b a p t e ru s ( L on c h od r y i n u s ) K i ef f e r 256 * su b a p t e r u s ( My s t r op h o r u s ) K i ef f er 256 * subap t e ru s ( P r e n a n t eon )

K i ef f er 256 " su b a r c t i c u s ( An t e o n ) H e l l e n 3 1 7 , 322 * s u b f l a v i c o r n e ( An t e o n ) H a u p t 309, 3 1 2 , 3 1 3 * s u b f l a v i c o r n i s ( An t e o n ) H a u p t 309 s u b t i l i s ( D ry i n u s ) D l m i 8 7 5 s u b t i l i s ( G on a t o p u s ) D l m i 1 558 s u c c i n e i p e s ( An t e o n ) K i e f f e r 1 825 * s u ch ov i ( H a p I og o n a t o p u s ) P on oma r en k o 1 4 1 3 s u ck l i n g i ( An t e o n ) D l m i 5 7 3 * s u f f o l c i en s i s ( A n t a e o n ) Ch i t t y 3 0 0 * s u f f 0 I c i e n s i s ( An t e o n ) C h i t t y 300 " su l c i s c u t e l l u m ( D i c o n d y l u s ) G i rau I t 6 6 0 s u ma t r a n u s ( N e o d r y i n u s ) End e r l e i n 1 8 26 * s u ma t r a n u s ( P s i l od ry i n u s ) Ender l e i n 1 826 s u p e rb u m ( An t e o n ) D od d 5 8 5 * su p e rb u s ( A n t e o n ) 5 8 5 s u r i n a me n s e ( An t e o n ) D l m i 521 s u r i n a me n s i s ( Ap h e l o p u s ) D l m i 77 s u r i n a men s i s ( D r y i n u s ) D l m i 839 * s u t u ra l i s ( Ag on a t op u s ) F e n t on 1 727 sy n c h r omu s ( Ag on a t op o i d e s ) R . P e rk i n s 1 3 1 0 , 1 3 3 9 , 1 8 4 3 s z e l e n y i i ( B o c ch u s ) Mo c z a r 617 * s z ep l i g e t i i ( D ry i n u s ) K i e f f e r 742, 744, 806, 8 1 8 * s z e p l i g e t i i ( P l a s t od ry i n u s ) K i ef f e r 7 2 8 , 744 t a mb i n i a e ( Ap t e r od r y i n u s ) E s ak i a n d H a sh i mo t o 1 8 2 6 t a n a n a r i v e i ( G o n a t op u s ) D l m i 1 6 20

1910

Mem . Amer . E n t .

ta r ra c on en s i s ( D r y i n u s ) Ma r s h a l l 7 4 2 , 7 4 4 , 1 8 3 8 ':' t a r r a c on e n s i s ( L e s t od ry i n u s ) Ma r sh a I I 7 4 4 t a r sa l i s ( An t eon ) K i e f f e r 1 826 t a sman i a n u m ( An t e on ) O l m i 5 8 2 * t a u r i cu s ( Ag on a t op o i d e s ) P on oma r en k o 1 5 1 8 t a u r i cu s ( E ch t h r o d e l p h a x ) P on o ma r e n k o 1 8 2 6 t a y l o r i ( An t e on ) O l m i 3 6 6 t a y l o r i ( G ona t opu s ) O l m i 1 6 2 8 t en e r i f e i ( G on a t o p u s ) O l m i 1 548 " t e n u i c o r n i s ( D r y i n u s ) D a l ma n 31 6 * t en u i c o r n i s ( Me r c e t i a ) K i ef f er 1 1 4 2 ':' t en u i p e s ( G o n a t op u s ) E s a k i a n d H a s h i mo t o 1 6 4 5 * t en u i s ( E p i g on a t op u s ) F en t on 1 71 7 t e r r e s t r i s ( D i c on d y l u s ) O g l ob l i n 1 8 2 6 * t e r ry i ( P a r a d ry i n u s ) R . C . L . P e rk i n s 8 1 7 , 8 1 8 t e s t a c e i p e s ( Lab e o ) K i e f f e r 1 826 * t e s t a c e i p e s ( Lab e r i u s ) K i e f f er 1 826 t e s t a c eu s ( A p t e r od ry i n u s ) Cameron 1 36 2 t e s t a c e u s ( Bocchu s ) 0 I m i 643 ;' t e s t a c eu s ( E u c a mp t o n y x ) R . C . P e rk i n s 1 4 9 9 , 1 6 8 2 * t e s t a c e u s ( G on a t opu s ) C a me r o n 1 3 6 2 , 1 6 8 2 * t e s t a c e u s ( G o n a t op u s ) R . C . L . P e rk i n s 1 6 8 2 t e s t a != eu s ( T r i d ry i n u s ) O l m i 941 * T e t r a d on t o c h e l y s J . F . P e rk i n s 1 4 3 4 * T e t r a d ry i n u s K i e f f e r 6 1 2 T e t r o d o n t o c h e l y s R i ch a r d s 1 434

I nst . ,

n o . 37,

1 984

t eu t on i a e ( D ry i n u s ) O l m i 8 6 2 t exan u s ( L ab e o ) A shmead 1 8 2 6 * t ex a n u s ( L a b e r i u s ) A s h m e a d 1 826 t h a i ( An t e o n ) O l m i 4 1 6 t h a i ( G o n a t op u s ) O l nc i 1 6 6 8 T h a u ma t od ry i n i n a e O l m i 6 8 2 T h a u ma t od r y i n u s R . C . L . P e rk i n s 682, 683 ':' t h e l i a e ( Ap h e l op u s ) G a h a n 8 7 t h e l i a e ( C r ov e t t i a ) Gahan 6 , 8 4 , 87 , 1 844 * t h o m s o n i ( An t e o n ) K i e f f e r 2 9 5 " t h o m s on i ( G o n a t op u s ) H e l l e n 1 51 5 t h o r a c i cu s ( G o n a t op u s ) N e e s 1 8 1 8 , 1 82 8 * t h o r a c i c u s ( N e od ry i n u s ) F ou t s 1 085 " t h o r a c i c u s ( P s i l od r y i n u s ) F ou t s 1 0 8 5 * t h r e n o d e s ( P a r a d ry i n u s ) R . C . P e rk i n s 9 0 3 t i b i a l i s ( A n t e o n ) Say 1 8 2 8 t i j u c a n u s ( Ap t e r od ry i n u s ) A r l e 1 36 5 * t i j u c a n u s ( P s e u d o g on a t op u s ) Ar l e 1 365 t o l h u r s t i ( D ry i n u s ) F e rn a n d o 1 828 t o rv u s ( Ap t e r o d ry i n u s ) R . C . L . P e rk i n s 1 3 4 2 , 1 3 5 3 , 1 8 4 3 t own e s i ( An t e on ) O l m i 3 7 9 t own e s i ( Ap h e l op u s ) O l m i 8 2 t own e s i ( C o n g a n t e on ) O l m i 1 0 0 t own e s i ( D e i n od ry i n u s ) O l m i 1 9 0 t own e s i ( G o n a t op u s ) O l m i 1 7 4 8 t own e s i ( T h a u ma t od ry i n u s ) 0 I m i 692 T r a n s d ry i n i n a e O l m i 1 0 4 8 T ra n s d r y i n u s O l m i 1 0 4 8 t ra n s l u c i d u s ( G o n a t op u s ) R . C . L . P e rk i n s 1 6 8 2 * t r i a n g u l a r i s ( A n t e on ) K i e f f e r 317 * t r i a r e o l a t u s ( An t e on ) K i e f f e r 300

I n dex

t u rn e r i ( An t e o n ) a l m i * t u rn e r i ( D i c o n d y l u s )

* t r i c a r i n a t u s ( An t e on ) K i ef f er 329 " T r i c h o g on a t op u s H e l l e n 1 50 0

191 1

1 47 3 ,

T r i c h o g on a t op u s K i e f f e r 1 4 7 3 t r i ch o t omma ( A n t e o n ) K i e f f e r 1 8 28 * t r i c h o t omma ( L a s i a n t e on ) K i ef f er 1 828 t r i c o l o r ( L on c h od r y i n u s ) K i e f f e r 2 5 4 , 282 t r i d en t a t u s ( An t e o n ) K i e f f e r 1 828 T r i d ry i n u s K i e f f e r 9 3 2 t r i f a s c i a t u s ( D ry i n u s ) K i e f f e r 807 , 8 2 1 * T r i g on a t o p u s K i e f f e r 1 3 7 5 * t r i l i n e a t u s ( N e o d ry i n u s ) D odd 9 3 2 t r i i i neat u s ( R i ch a r d s i d ry i n u s ) D od d 932 t r i n i d a d i ( D e i n od ry i n u s ) a l m i 220 t r i n i t a t i s ( Ap h e l o p u s ) a l m i 80 t r i n i t a t i s ( N e od ry i n u s ) R i ch a r d s 1 1 1 1 , 1 1 1 2 , 1 844 t r i n od i s ( G o n a t op u s ) K i e f f e r 1 828 t r i a r t i t u m ( An t eon ) K i e f f e r 329 ':' t r i p a r t i t u s ( An t e on ) K i e f f e r 329 " T r i san t e o n K i e f f e r 1 1 9 t r i s t i s ( G on a t o p u s ) a l m i 1 7 5 7 ':' t r i s u I c a t u s ( A p h e I o p u s ) K i ef f er 47 " t r i v i a l i s ( An t e on ) K i e f f e r 334 ':' t r o c h a n t e r i c u s ( E u c a mp t ony x ) B en o i t 1 6 3 1 t r o c h a n t e r i c u s ( G on a t o p u s ) B en o i t 1 6 3 1 t r op i c a l i s ( Ap h e l o p u s ) a l m i 76

1 637 t u r n e r i ( G o n a t op u s )

378 B en o i t

B en o i t

1 637

* t y p h l o c y b a e ( A ph e l op u s ) Mu e s e b e ck 7 4 , 7 6 * t y p h l o c y b a e ( G o n a t op u s ) A sh m e a d 1 0 9 3 * t y p h l o c y b a e ( L a b e o ) A sh mead 1 09 1 * t y p h l o cy b a e ( L a b e r i u s ) A shmead 1 0 9 3 t y p h l o cy b a e ( N e od ry i n u s ) A shmead 1 0 9 1 , 1 0 9 4 , 1 8 3 3 , 1 843 * t y p h l o cy b a e ( P s i l od r y i n u s ) A sh m e a d 1 0 9 4 t y p i c u m ( An t e o n ) R . C . L . P e rk i n s 5 5 3 * t y p i c u s ( An t e o n ) R . C . L . P e rk i n s 5 5 4 �' t y p i c u s ( C h e l o g y n u s ) R . C . L . P e rk i n s 5 5 4 * t y p i cu s ( N e o ch e l ogynu s ) R . C . P e rk i n s 2 8 6 , 5 5 4 u g a n d a n u m ( An t e on ) a l m i 3 7 2 u g a n d a n u s ( A c r o d on t o c h e l y s ) Beno i t 1 1 77 * u g a n d a n u s ( P l a t y g on a t o p u s ) Beno i t 1 1 7 8 * u g a n d a n u s ( R h y n c h og o n a t op u s ) Ben o i t 1 1 7 8 u g a n d a n u s ( T r i d ry i n u s ) a l m i 935 u mb e r ( B o c c h u s ) a I m i 6 1 4 " u mb r a t u s ( N e od ry i n u s ) R . C . L . P e rk i n s 1 1 1 8 * u n d u l a t u s ( L e s t od ry i n u s ) B en o i t 9 1 9 u n d u l a t u s ( R i c h a r d s i d ry i n u s ) B en o i t 9 1 9 u n g u l a t u s ( M i r od ry i n u s ) P on oma r en k o 6 6 4 , 6 7 1 , 6 7 4 u n i c a r i n a t u s ( An t e on ) K i e f f e r 1 828 * u n i cu s

( C h a l c o g o n a t op u s ) R . C .

P e rk i n s

1 468

Mem .

1912

Amer .

Ent.

un i c u s ( T e t r od on t o c h e l y s ) R . C . P e rk i n s 1 4 6 8 , 1 8 4 2 * u n i f a s c i a t u s ( A n t eon ) A sh m e a d 4 5 2 ,

453

u n i I i n e a t u s ( G o n a t op u s ) K i e f f e r 1 5 1 1 , 1 53 2 * u n i I i n e a t u s ( N e o g o n a t op u s ) K i ef f e r 1 532 u p emb a n u s ( G o n a t op u s ) O l m i 1 628 v a g a n s ( D e i n o d ry i n u s ) O l m i 201 v a l i d u s ( P s e u d o g on a t op u s ) 0 1 m i 1 248 * v a r i ab i l i s ( D e i n od ry i n u s ) F en t o n 1 3 6 v a r i c o r n i 5 ( Ap h e l op u s ) B r u e s 72 v a r i e g a t u s ( T h a u ma t od r y i n u s ) D e S a n t . a n d V i d a l S a rm . 703, 704 vari i s t r i a t u s ( P s e u d o g on a t op u s ) F en t o n 1 27 6 , 1 844 " v a r i p e s ( G o n a t op u s ) B r u e s 1 459 * v a r i p e s ( P a rad ry i n u s ) R . C . L . P e rk i n s 9 0 3 v a r i p e s ( T e t r od o n t o ch e l y s ) B ru e s 1 4 5 9 v en a t o r ( D r y i n u s ) R . C . L . P e rk i n s 9 0 3 , 1 8 4 5 , 1 8 4 6 * v en a t o r ( P a r a d r y i n u s ) R . C . L . P e rk i n s 9 0 3 v e t u s ( L e s t od ry i n u s ) B r u e s 1 829 , 1 830 * v i c i n u m ( An t e o n ) K i e f f e r 2 9 5 * v i c i n u s ( A n t e on ) K i e f f e r 2 9 5 * v i c t o r ov i ( D ry i n u s ) P on oma r e n k o 1 1 4 2 * v i d u u s ( Ap h e l op u s ) R . C . L . P e rk i n s 7 3 v i e t ( Me s od ry i n u s ) O l m i 1 0 1 6 * v i r g i n i en s i s ( An t e o n ) F en t on 475 * v i rg i n i e n s i s ( C h e l o g y n u s ) F en t on 4 7 5

I ns \ . ,

no . 37,

1 984

v i s t o s u s ( G o n a t op u s ) O l m i 1 5 0 8 * v i t e l l i n e i p e s ( An t eon ) K i e f f e r 260 * v i t e l l i n i p e s ( An t e on ) K i e f f e r 259 * v i t e l l i n i p e s ( P r e n a n t e on ) K i ef f er 260 v i t i en s i s ( A c r o d o n t o c h e l y s ) R C • L . P e rk i n s 1 1 9 1 , 1 8 4 5 * v i t i en s i s ( G o n a t op u s ) R . C . L . P e rk i n s 1 1 9 1 * v i t i en s i s ( H a p l o g o n a t op u s ) R . C . L . P e rk i n s 7 , 1 4 2 5 , 1 8 3 1 , 1 8 3 2 , 1 846 * v i t i en s i s ( N e o g o n a t op u s ) R • C L . P e rk i n s 1 1 9 1 " v i t r i p en n i s ( L a b e o ) H a l i d a y 1 37 5 , 1 378 * v i t r i p en n i s ( L a b e r i u s ) H a l i day 1 3 7 9 * v i v a r i e n s i s ( An t e o n ) B r a d l ey 475 * v i v a r i e n s i s ( C h e l og y n u s ) B r a d l ey 4 7 5 v i v a x ( A n t e on ) O l m i 5 1 4 v o l I en h ov en i ( G o n a t o p u s ) O l m i a n d Cu r ra d o 1 5 2 1 * v o l l en h ov e n i ( N e og on a t op u s ) O l m i a n d C u r ra d o 1 5 2 1 vu l can i cum ( C ongan t eon ) B en o i t 9 8 , 1 0 3 * v u l c a n i c u s ( C on g a n t e on ) Beno i t 1 03 *vu l g a r e ( An t eon ) K i e f f e r 3 1 7 * v u l g a r i s ( An t e o n ) K i e f f e r 3 1 7 v u l s u m ( An t e on ) O l m i 5 7 6 * wa g n e r i ( G on a t op u s ) S t r a n d 1 577 * wa l k e r i ( C h e l og y n u s ) K i e f f e r 260 wa s b a u e r i ( An t e o n ) O l m i 4 6 4 w a s b a u e r i ( B o c ch u s ) O l m i 6 4 4 w e l l i n g en s i s ( D ry i n u s ) O l m i •

•

839 w e t mo r e i ( R i c h a r d s i d ry i n u s ) O l mi 9 2 9 * wh i t t a k e r i ( An t e o n ) Mu e s e b e ck

I n dex

a n d Wa l k l ey 4 8 1 w i I I i a m s i ( G on a t op u s ) O l m i 1 774 w i I I i n k i ( G on a t o p u s ) O l m i 1 742 w i t t e i ( Ap h e l op u s ) B en o i t 6 2 ':' w i t t e i ( G y mn a p h e l op u s ) B en o i t 4 4 , 6 2 w i t t me r i ( G on a t op u s ) O l m i 1 55 1 "' x a n t h o s t i g ma ( A n t e on ) K i ef f er 337 x a n t h o t h o r a x ( An t e on ) B r a d l ey 479 '� x a n t h o t h o r a x ( C h e l o g y n u s ) B r ad l e y 4 7 9 * X en a n t e on K i e f f e r 2 8 7 ':' x e r o p h i l u m ( An t e o n ) B e n o i t 668

1913

* x e r o p h i I u s ( An t e on ) B en o i t 668 x e r op h i I u s ( M i r o d ry i n u s ) Ben o i t 668 * x e s t o c ep h a l u s ( C h a l c o g on a t op u s ) R . C . L . P e rk i n s 1 7 0 3 y a s u ma t s u i ( A n t e o n ) O l m i 424 y a s u ma t su i ( G o n a t op u s ) O l m i 1 64 1 z a i r e n s e ( A n t e on ) B e n o i t 4 0 6 , 407 z a i r en s i s ( T e t rodon t o ch e l y s ) O l mi 1 454 z e a l a n d i c u s ( G o n a t op u s ) O l m i 1 05 5 , 1 776 z h e r i ch i n i ( C r e t od ry i n u s ) P on oma r e n k o 1 0 4 6 , 1 8 3 0

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