A revision of the Dryinidae (Hymenoptera) Part 1

MEMOIRS of the AMERICAN ENTOMOLOGICAL INSTITUTE Number 37, Part 1 Pages I-XII, 1-946

A REVISION OF THE DRYINIDAE (HYMENOPTERA) by Massimo Olmi Universit::i degli Studi della Tuscia Viterbo, Italy

The American Entomological Institute 5950 Warren Road, Ann Arbor, Michigan 48105, U. S. A. 1984

TABLE OF CONTENTS

Introduction . . . . . .

............ 1

'.Iaterial

.. . . . . . . . . . 1 .4

examined.

~cknowledgments

3 i

0

.•

.5 .7 .. ... 7 .. .. 10 · .31 . . . . 31

logy . . . . . . . . . .

~osts

attacked ••..•

::'conomic

importance.

',',orphology

and

terminology.

::ollection . • . . . . . • . . • • . • • . • ::; reserva t ion . . . . . . . . . . . . . . . "ami Iy

size and distribution . . . . . . . . .

<ey

the

to

subfami lies

of

Sdbfami Iy Aphelopinae.. :;enus Aphelopus . . . . . . . . Palaearctic species. 1.

me I a I eu cu s .•..

2.

atratus ..

3.

serra tus.

4.

camu s ••..

5. 6.

nigriceps. n epa I en sis.

7. querceus ... Ethiopian species . . . . 1.

mediocarinatus.

2.

wittei . . . . . . .

3.

incisus . . . .

Oriental

species.

. .. 43 , •. 44

5. 6.

leucopus ..

.78

di ffusus ..

. . . 46

7.

t r in ita tis .

· .47 · .50

8.

alvarenganus . . . . . .

.79 .80 .80

9.

jamaicanus . . . .

.52 .54 .56 .57

Australian

1.

townesi . . . .

2. 3.

guineensis. australis ..

4.

ca I ed on i cu s .

.60 . ... 60

5.

a I bop i c t us ..

1. 2.

nearctica.

.. .. 62 .64 . ... 64

borneanus .. maculiceps.

.81 .. .81 · .. 82

species.

.59

.. .. 65 .66 .66 3. malayanus •.. 4.ochreus . . . . . .67 . 67 5. 0 r i en t a lis • • • . . . . . . .68 6. penanganus .. 1. 2.

.32 .34

Dry i n i dae •.

· .82 · .. 83 . . . . . 83

· .• 84 Genus Crovettia . . . . . . . . . . . . . . . 84 Nearctic species . . . . . . . . . . . 84

.... 85 · .87

the I i ae ..••.•

Neotropic species • . . . . 1. barbara . . . . • . . 2.

· .89 .89

4.

neotropica •••••••.••• 92 plaumanniana. · .92 brasiliana . . . . . . . . . . . 93

5.

3.

colombiana.....

..94

7.

birmanus . . . . . • . . . .

.69

Subfami Iy Biaphelopinae.

..94

8.

phi I ippinus ..

.69

Genus Biaphelopus.......

. .95

\Iearctic

species ...

.69

1.

ru f i v en t r is.

2.

bicolor . . . .

.70 .71

3. 4.

varicornis.

.72

a I bop i c t us. species.

.73 .76 .76

~eotropic

1.

tropicalis • . . . • • • .

2.

diaphanus . . . .

· .77 . .. 77

3.

su r i namen 5 is.

4.

fen t on i . . . . • . • . . . • • . . 7 B

Palaearctic 1.

species...

.96

masneri.........

.96

Subfami Iy Conganteoninae. Genus Conganteon . . . . . . .

Palaearctic species. 1.

nepalense . . . . .

Ethiopian

species.

1.

townesi .•..

2.

vulcanicum.

Genus Chelanteon . . . .

.96

.98 .. . . . 98 .. . . . 98 · .99

.100 .103 . . . . . . 105

II

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

species . • • • . • • . . . 105

22.

insignis . . . . . . . . . . . . 193

richardsi . . . . . . . . . . . 105

23.

pseudobilobus ..••••• 193

Genus Fiorianteon ••.••••••••. 108 speci es . . . . . . . 1 09

24. 25.

rubrolobatus . • • • . . . • 197 nigrolobatus • . • • . . . . 199

junonium . . . . . . . . . . . . . 109

26.

vagans . . . . . . . . . . . . . . 201

Subfami Iy Anteoninae . . . . . . . . . 110

27.

pilosifrons . . . . . . . . . 204

Oriental 1.

Pa I aearct i c 1.

28.

politus . • . . . . • • • . • • . 206

species •••••..•• 114

29.

claripes . . . . . . . . . . . . 206

1.

aerias . . . . . . . . . . . . . . 115

30.

chiapasi • • . . . . . • • . . . 209

2.

fuscum . . . . . . . . . . . . . . . 116

31.

n i 9 r i can s ••••••••••. 21 1

Genus Deinodryinus . . . . . . . . . . . 118

32.

actuosus • • • . . . • • . . . • 211

Palaearctic species . . . . . . . 121 1. areolatus . . . • • • • • . . . . 121

33.

incaicus . . . . . . . . . . . . 214

34.

bol ivianus ••...••••• 217

Ethiopian

eminens . • . • . • . . . . . . • 217

Genus Metanteon . . . . . . . . . . . . . . 113 Neotropic

species . . . . . . . . . 123

35.

1. 2.

insulanus . . . . . . . . . . . 124 madecassus ••..•••.•• 125

36.

trinidadi •••...•••.• 220

37.

rapax . . . • • • • • . . . . . • . 223

3.

madagascariensis .••• 128

38.

croceus . • • • • . . . . . • . • 223

4.

ben 0 i t i . . • • • • • • • • . . . 1 29

39.

5.

mon t i co Ius . • • • • . • . . • 130

40.

asper ••••.•••••••... 226 schl ingeri .•.••••.•• 230

sp e c i e s •••••••••. 1 3 1

41.

alexandrae . . . • • • . . . . 230

as i at i cu s •.•.•••••••. 131

42.

carpens .••••••.•.••• 232

species . . . . . . . . . . 133

43.

per I u c en s . . • • . • . • . . . 233

1. 2.

at r i v en t r is ••••••••• 1 33 quercicolus . . . . . . . . . 143

44. 45.

i nermi s .••••••••••.• 233 paranus . • . . . • • • • • • . . 236

3.

paradoxus . . . . . . . . . . . 145

46.

hi r tic 0 rn is •••..•••. 237

Or i en t a I 1.

Nearctic

species . . . . . . . . . 147

47.

rivularis • • • . . . . . . . . 239

e I egan s • • • • . . . . . . • • • 154

48.

aequa lis . • • . . . . . . . • • 240

Neotropic 1.

2•

sp e c i 0 su s ••••••••••• 1 57

49.

minor ••.••••••..•••• 241

3.

diaphanus . . . . . . . . . . . 158

50.

ben i anus • • . . . . . • • . • • 243

4.

n i grorufus ••••...••. 159

51.

i t en e z i . . . • • • • • • . . . . 245

5.

iphias ••....•••••••• 160 bicolor • • • • • • . . • . • . . 163

52.

b i I ob 0 ide s .••••••••• 245

53. 54.

f I uv i at i i i s . . • • . • . . . 248 cuzcanus ••••••.••.•• 249

6. 7.

maximus ••••••••••••• 165

8. 9.

melanocephalus . . . . • . 168

10.

11.

a t I an t i cu s .•.••••••• 172 achterbergi .•.•••••• 174

12.

peruvianus . . • • . • • . . • 174

13. 14.

hymenaeus . . . . . . . . . . . 178 colombianus . . . . . . . . . 180

15. 16. 17.

cascus . . . . . . . . . . . . . . 182 albopictus . . . . . . . . . . 183 amoenus •••••••••••.. 185

18.

pegnai .•.••••••••••• 186

19.

pseudoamoenus •••••.• 188

noyesi •••••••••••••• 170

55. panamensi s . . . • • . . . . . 251 Genus Lonchodry i nus . . . . . . • . . . 253 Palaearctic

1. 2. 3.

ruficornis • • . . • • . • • . 258 nepalensis ..•••••••• 267

Ethiopian

1. 2.

species . . . . . . . 255

subapterus . . • . • • • • . . 256

species • . . • • • . . . 269

seyrigi •••••••••••.• 269 paul iani . . • • • • . • . . . . 271

Oriental

species •.••.••••. 271

1.

sin en sis ••••••••.... 273 Nearctic species . . . . . . . . . . 273

20.

townesi . . . . . . . . . . . . . 190

1.

f I a vu s • • • • . . • . . • . • • • 274

21.

b i I obu s . . . . . . . . . . . . . 191

2.

masneri . . . . . . . . . . . . . 277

Table of contents

bakeri . . . . . . . . . . . . . . 278 3. Neotropic species • . . . . . . . . 282 1. tricolor . . . . . . . . . . . . 282 Australian species . . . . . . . . 284 1. guineensis . . . . . . . . . . 284 2. notogeicus . . . . . . • . . . 286 :;enus Anteon . . . . . . . . . . . . . . . . . 286 Palaearctic species . . . . . . . 290 1. jurineanum . . . . . . . . . . 294 2. brachycerum . . . . . . . . . 300 3. reticulatum . . . . . . . . . 304 4. nepalense . . • . . . . . . . . 304 5. arcuatum . . . . . . . . . . . . 305 6. flavicorne . . . . . . . . . . 309 7. ephippiger . . . . . . . . . . 313 8. pubicorne . . . . . . . . . . . 316 9. scapulare . . . . . . . . . . . 322 10. infectum . . . . . . . . . . . . 325 11. tripartitum . . . . . . . . . 329 12. crassifrons . . . . . . . . . 332 13. gaul I ei . . . . . . . . . . . . . 334 14. fulviventre . . . . . . . . . 337 15. hilare . . . . • . . . . . . . . . 342 16. ingenuum . . . . . . . . . . . . 342 17. corax . . . . . . . . . . . . . . . 345 18. muni tum . . . . . . . . . . . . . 345 19. japonicum . . . . . . . . . . . 347 Ethiopian species . . . . . . . . . 348 1. brachypterum . . . . . . . . 353 2. agile . . . . . . . . . . . . . . . 354 3. maritimum . . . . . • . . . . . 357 4. kasaicum . . . . . . . . . . . . 362 5. fiorii • . . . . . . . . . . . . . 362 6. kivuanum . . . . . . . . . . . . 363 7. taylori . . . . . . . . . . . . . 366 8. dayi . . . . . . . . . . • . . . . • 367 9. canabense . . . . . . . . . . . 370 10. ugandanum . . . . . . . . . . . 372 11. kawandanum • . . . . . . . . . 374 12. afrum . . . . . . . . . . . . . . . 375 13. turneri . . . . . . . . . . . . . 378 14. townesi . . . . . . . . . . . . . 379 15. decellei . . . . . . . . . . . . 380 16. anthereon . . . . . . . . . . . 381 17. namibiense . • . . . . . . . . 383 18. gutturnium . . . . . . . . . . 384 19. arab i cum . . . . . . . . . . . . 387

20. 21 . 22. 23. 24. 25. 26. 27. 28. 29. 30. 31. 32. 33. 34 . 35. 36 . 37. Or i en 1. 2. 3. 4. 5. 6. 7. 8.

III

natalense . . . . . . . . . . . 388 hoyoi . . . . . . . . . . . . . . . 390 mingoae . . . . . . . . . . . . . 391 long i scapum . . . . . . . . . 392 madagascolum . . . . . . . . 393 medleri . . . . . . . . . . . . . 395 hova . . . . . . . . . . . , . . . . 397 compressicoxale . . . . . 398 inf latrix . . . . . . . . . . . 399 rufonigrum . . . . . . . . . . 402 oriphi lum . . . . . . . . . . . 403 fabulosum • . . . . . . . . • . 404 abruptum . . . . . . . . . . . . 404 zairense . . . . . . . . . . . . 406 mo sse Ii . . . . . . . . . . . . . 408 capei . . . . . . . . . . . . . . . 409 eme r i tum . . . . . • . . . . . . 41 a garambanum . . . . . . . . . . 411 t a I sp e c i e s . . • . . . . . . . 41 2 borneanum . . . . . . . . • . . 416 thai . . . . . . . . . . . . . . . . 416 s i I v i co I um . . . . . . . . . . 419 indicum . . . . . . . . . . • . . 419 mysorense . . . . . . . . • • . 422 yasumatsui . . . . . . . . . . 424 sarawaki . . . . . . . . . . . . 425 bengalense . • . . . . . . . . 428 9. k r omb e i n i . . . . . . . . . . . 429 10. mui ri . . . . . . . . . . . . . . . 432 11. laotianum . • . . . . . . . . . 435 12. fyanense . . . . . . . . . . . . 435 13. javanum . . . . . . . . . . . . . 437 14. mun roe i . . . . . . . . . . . . . 439 lankanum . . . . . . . . . . . . 441 15. nemora Ie . . . . . . . . . . . . 442 16. si Ivestre . . . . . . . . . . . 444 17. expol itum . . . . • . . . . . . 444 18. pet ersen i . . . . . . . . . . . 445 19. bauense . . . . . . . . . . . . . 446 20. debi Ie . . . . . . . . . . • . . . 447 21. 22. phi I ippinum • . . . . . . . . 448 Nearct ic species •.•... '" •. 450 1. popenoe i . . . . . . . • . . . . 452 2. arizonense . . . . . . . . . . 455 3. canadense . . . . . . • • . . . 457 4. nebulosum . . . . . . . • . . . 461 5. funestum • . . . . . . . • • • . 462

IV

Mem. Amer.

En!.

6. wasbaueri . . . . . . . . . . . 464 7. minimum • . . . . . . . . . . . . 467 8. masoni . . . . . . . . . . . . . . 469 9. rugosiceps . . . . . . . . . . 471 10. puncticeps . . . . . . '" .474 11. xanthothorax . . . . . . . . 479 12. osborni . . . . . . . . • . . . . 481 13. durangoi . . . . . . . . . . . . 484 14. ciudadi . . . . . . . . . . . . . 486 15. menkei . . . . . . . . . . . . . . 486 16. hirtifrons . . . . . . . . . . 487 Neotropic species . . . . . . . . . . 488 1. clavatum . . . . . . . . . . . . 491 2. mo I Ie . . . . . . . . . . . . . . . 492 3. nigrorubrum . . . . . . . . . 494 4. pan amen se . . . . . . . . . . . 497 5. jamaicanum . . . . . . . . . . 497 6. mexicanum . . . . . . . . . . . 499 7. oliveirai . . . . . . . . . . . 502 8. noyesi . . . . . . . . . . . . . . 503 9 . I ob a tum . . . . . . . . . . . . . 506 10. propodeale . . . . . . . . . . 506 11. chi riquense . . . . . . . . . 508 12. nycteum . . . . . . . . . . . . . 510 13. annul icorne . . . . . . . . . 511 14. albitarse . . . . . . . . . . . 512 15. v i vax . . . . . . . . . . . . . . . 514 16. p iii corn e . . . . . . . . . . . 51 5 17. conterminum . . . . . . . . . 516 pi aumann i . . . . . . . . . . . 517 18. 19. bol ivianum . . . . . . . . . . 518 micros . . . . . . . . . . . . . . 519 20. 21. surinamense . . . . . . . . . 521 Austral ian species . . . . . . . . . 522 1. gi luwense . . . . . . . . . . . 526 2. chelogynoides . . . . . . . 527 myrmecophi lum . . . . . . . 529 3. 4. reticulaticeps . . . . . . 531 au I i cum . . . . . . . . . . . . . 533 5. sedlaceki . . . . . . . . . . . 534 6. 7. anxium . . . . . . . . . . . • . . 536 8. australe . . . . . . . . . . . . 538 9. parvu I um . . . . . . . . . . . . 539 10. orientale . . . . . . . . . . . 541 11. aculeatum . . . . . . . . . . . 545 12. corruptum . . . . . . . . . . . 545 13. miles . . . . . . . . . . . . . . . 547

Ins!.,

no. 37,

1984

14. destructor . . . . . . . . . . 547 15. caledonianum . . . . . . . . 551 16. typicum . . . . . . . . . . . . . 553 17. bougainvi I lei . . . . . . . 555 18. curradoi . . . . . . . . . . . . 556 19. firmum . . . . . . . . . . . . . . 558 20. leiosomum . . . . . . . . . . . 560 21. ni t idum . . . . . . . . . . . . . 561 22. bouceki . . . . . . . . . . . . . 564 23. mundum . . . . . . . . . . . . . . 565 24. bel lator . . . . . . . . . . . . 568 n i gri corne . . . . . . . , .. 568 25. fij ianum . • . . . . . . . . . . 569 26. dub i um . . . . . . . . . . . . . . 570 27. cor i aceum • . . . . . . . . . . 571 28. su ck lin 9 i . . . . . . . . . . . 573 29. bi smarckense . . . . . . . . 575 30. gloriosum . . . . . . . . . . . 575 31 . vulsum . . . . . . . . . . . . . . 576 32. 33. I uctuosum . . . . . . . . . . . 577 34. hagen i . . . . . . . . . . . . . . 578 haustum . . . . . . . . . . . . . 580 35. coral I inum . . . . . . . . . . 581 36. 37. tasmanianum . . . . . . . . . 582 permirum . . . . . . . . . . . . 583 38. involutum . . . . . . . . . . . 584 39. 40. superbum . . . . . . . . . . . . 585 41. rufiscapum . . . . . . . . . . 586 42. gi raul t i . . . . . . . . . . . . 587 Genus Prioranteon . . . . . . . . . . . . 589 Palaearctic species . . . . . . . . 589 1. biroi . . . . . . . . . . . . . . . 591 Ethiopian species . . . . . . . . . . 593 1. richardsi . . . . . . . . . . . 594 2. casalei . . . . . . . . . . . . . 596 Subfami Iy Bocchinae . . . . . . . . . . 599 Genus Bocchoides . . . . . . . . . . . . . 601 Ethiopian species . . . . . . . . . . 602 1. beki Iyensis . . . . . . . . . 602 Genus Chelothel ius . . . . . . . . . . . 604 Palaearctic species . . . . . . . . 605 1. gryps . . . . . . . . . . . . . . . 607 2. berlandi . . . . . . . . . . . . 609 Genus Bocchus . . . . . . . . . . . . . . . . 611 Palaearctic species . . . . . . . . 613 1. ita lieu s . . . . . . . . . . . . 6 1 4 2. umber . . . . . . . . . . . . . . . 614

v

Table of contents

3.

szelenyi . . . . . . . . . . . . 617

1.

4.

slovacus . • • . • • . . . . • • 618

2.

mi rabi lis . . • • . . • . . • . 687 infuscatus • . . . . . . . . . 690

5. 6.

pagl ianoi • • . . . . . . • . . 620 scaramozzinoi . . . . . . . 621

3.

dentatus . . . . . . . . . . . . 690

4.

townesi . . . • . . . . . . . . . 692

7. 8.

scobiolae • . • • . . . • . • • 622 europaeus . . . . . . . . . . . 624

5.

migratorius . . . . . . • . . 694

Oriental

9.

parthicus . . . . . . . . . . . 624

1.

species . . . . . . . . . . . 695

phi I ippinus . . . . . . . . . 697

species . . . . . . . . . . 627

2.

asiaticus . . . . . . . . . . . 697

1.

richardsi . . . . . . . . . . . 628

3.

malayanus . . . . . . . . . . . 699

2.

bini . . . . . . . . . . . . . . . . 629

Nearctic

3.

s ey rig i . • . . . . . . . . . . . 629

1.

:::thiopian

Jriental

species . . . . . . . . . . . 632

species . . . . . . . . . . . 700

perkinsi . . . . . . . . . . . . 700

Neotropic 1.

species . . . . . . . . . . 702

variegatus . . • . . • • . . . 703

1. 2.

laotianus . . . . . . . . . . 635 mu I u en sis . . . . . . . . . . . 635

2.

minimus . . . . . . • . . • . . . 704

3.

pedunculatus •.•••••• 636

3.

macilentus . . . . . . . . . . 706

4.

rubricus . . . . . . . . • . . • 639

'<earctic

species . . . . . . . . . . . 641

1.

mirabilis . • . • . . . . . . . 643

2. 3. 4. 5.

testaceus . . . . . • . . . . . 643 wasbaueri . . . • . . . • . . . 644

a r i z on i cu s . . . . . . . . . . 645

4.

rufus . . . . . . • . . • . . • . . 710

5.

bruchi . . . . • . . . . . . . . . 714

6.

clarus . • . . . • . • • . . • • . 715

Australian

species . . . . . . . . . 717

1.

flavus . . . . . . . . . . . . . . 718

2.

koebelei ••.••••••••• 721

f lavipes . . • . . . . • . . . . 650

Subfami Iy Dryininae . . . . . . . . . . 723

6. 7.

flavicoll is ••••.•••. 650

Genus Dryinus . . . . . . . . . . . . . . . . 728

I at i c ep s . . . . . . . . . . . . 652

8.

dubius • • . . . • • . . . . . . . 653

Palaearctic species . . . . . . . . 729 1. sanderi . . . . . . . . . . . . . 731

species . . . . . . . . . 655

2.

canariensis . . . . . . . . . 734

austral iae . . . . . . . . . . 656

3.

collaris . . . . . . . . . . . . 735

guineensi s ••••..••.. 658

4.

nepalensis . . . . . . . . . . 740

minimus . . . . . . . . . . . • . 658

5.

tarraconensis . . . . . . . 742

6.

koreanus . . . . . . . . . . . . 747

7.

an t i quu s . . • . . . • • . . • • 751

>\ustral ian

1. 2. 3. 4.

robustus . . . . . . . . . . . . 660

5.

bouceki . . • • . . . . • . . . • 661

:=""us Mirodryinus . . . . . . . . . . . . 663 "alaearctic species . . . . . . . . 664 1. gobiensis . . . . . . . . . . . 666

Ethiopian 1.

species . . . . . . . . . . 751

bisulcatus . . . • . . . . . . 754

2.

seyrigi . . . . . . . . . . . . . 756 alticolus . . . . . . . . . . . 758 spangleri . . . . . . . . . . • 762

3.

ungulatus . . . . . . . . . . . 671

3. 4.

4.

atlanticus . . . . . . . . . . 672

5.

deceptor . . . • • . . . . • • . 762

5.

ponomarenkoae . . • . . . . 674

6.

mahensis . . . . . . . . . . . . 764

:=enus Radi imancus . . . • • • • . . . . . 675

7.

sierranus . . . . . . . . . . . 766

species . . . . . . . . 676

8.

pretorianus . . . . . . . . . 767

1. olmii . . . . . . . . . . . . . . . 676 :::enus Mystrophorus . . . . . . . . . . . 678

9.

saussurei . . . . . . . . . . . 770

10.

hova • • • . . . . . . . . • . . . • 772

species . . . . . . . 678

11.

basilewskyi . . . . . . . . . 774

formicaeformis . . . • • . 678 Iy Thaumatodryininae .. 682

12.

chamaeleo . . • • . . . • • • . 775

13. 14.

schoutedeni . . . . . . . . • 778 mayogo . . . . • . . . . • . . . . 780

15.

orophi Ius • • . . . . • . . . • 781

2.

xerophi Ius • • . . . . • . . . 668

Palaearctic

Palaearctic 1. 5'~bfami

:=enus Thaumatodryinus • • • . . . . • 683 Ethiopian

species . . . . . . . . . 686

VI

Mem. Amer. Ent.

16 . mod est us . . . . . . • • . . . . 785 17. i ncertus . . . . . . . . . . . . 785 18. burgeoni . . . . . . . . . . . . 786 19. afer . . . . . . . . . . . . . . . . 787 Oriental species . . . . . . . . . . 791 1. irregularis . . . . . . . . . 794 2. kiefferi . . . . . . . . . . . . 795 3. longipes . . . . . . . . . . . . 799 4. pulcher . . . . . . . . . . . . . 799 5. ex iii s . . . . . . . . . . . . . . 802 6. k r omb e i n i . . . . . . . . . . . 802 7. stantoni . . . . . . . . . . . . 806 8. trifasciatus . . . . . . . . 807 9. latus . . . . . . . . . . . . . . . 808 10. indi cus . . . . . . . . . . . . . 810 11. praeclarus . . . . . . . . . . 814 12. fulvus . . . . . . . . . . . . . . 815 13. lucens . . . . . . . . . . . . . . 817 14. browni . . . . . . . . . . . . . . 817 15. lankanus . . . . . . . . . . . . 821 Nearctic species . . . . . . . . . . 822 1. mexicanus . . . . . . . . . . . 823 2. crawfordi . . . . . . . . . . . 825 americanus . . . . . . . . . . 826 3. alatus . . . . . . . . . . . • . . 828 4. Neotropic species . . . . . . , .. 831 1. constans . . . . . . . • . . . . 833 citricolus . . . . . . . . . . 835 2. f I a von i 9 e r . . . . . . . . . . 838 3. well ingensi s . . . . . . . . 839 4. surinamensi 5 • • • • • • • • 839 5. striatus . . . . . . . . . . . . 845 6. argentinus . . . . . . . . . . 846 7. 8. caraibicus • . . . . . . . . . 849 9. napensi s . . . . . . . . . . . . 850 opaci frons . . . . . . . . . . 851 10. 11. grand is . . . • • . . . . . , .. 852 a I va renganu s . . . . . . . . 857 12. 13. sin op en sis . . . . . . . . . . 858 14. bel i zen sis . . . . . . . . . • 858 anti I leanus . . . . . . . . . 860 15. 16. t eu t on i a e . . • . . . . . . . . 862 17. catarinae . . . • • • . . . . . 864 18. kimseyae . . . . . . . . . . . . 864 19. palaeodominicanus ... 866 Austral ian species . . . . . . . . 868 1. pallidus . . . . . . . . . . . . 871

Inst.,

no. 37,1984

2. speciosus . . . . . . . . . . . 874 3. subtilis . . . . . . . . . . . . 875 4. pacificus . . . . . . . . . . . 876 5. montanus . . . . . . . . . . . . 878 6. p seudophanes . . . . . . . . 878 7. insularis . . . . . . . . . . . 880 8. bouceki . . . . . . . . . . . . . 880 9. k 0 eb e lei . . . . . . . . . . . . 882 10. guineensis . . . . . . . . . . 884 11. gigas . . . . . . . . . . . . . . . 885 12. bi smarcki . . . . . . . . . . . 887 13. aterrimus . . . . . . . . . . . 888 14. biaki . . . . . . . . . . . . . . . 890 15. punctatus . . . . . . . . . . . 893 16. areolatus . . . . . . . . . . . 894 17.leptias . . . . . . . . . . . . . 895 18. quatei . . . . . . . . • . . . . . 897 19. dahmsi . . . . . . . . . . . . . . 897 20. glaber . . • . . . . . . . . . . . 900 21. austral ianus . . . . . . . . 900 22. venator . . . . . . . . . . . . . 903 23. papuanus . . . . . . . . . . . . 904 Genus Richardsidryinus . . . . . . . 907 Palaearctic species . . . . . . . 908 1. albrecht i . • . . . . . . . . . 909 maroccanus . . . . . • • . . . 910 2. corsicus . . . . . . . . . . . . 913 3. Ethiopian species . . . . . . . . . 914 1. e r ra t i cu s . . . . . . . . . . . 91 6 ca r i n i c ep s . . . . . . . . . . 91 8 2. 3. undu I a t us . . . . . . . . . . . 919 Ori ental speci es . . . . . . . . . . 922 1. indianus . . . . . . . . . . . . 922 2. pyri Ilae . . . . . . . . . . . . 925 Nearctic species . . . . . . . . . . 927 1. nearcticus . . . . . . . . . . 927 2. wetmorei . . . . . . . . . . . . 929 3. canadensis . . . . . . . . . . 930 Austral ian species . . . . . . . . 931 1. t r iii n ea t us • . . . . . . . . 932 Genus Tridryinus . . . . . . . . . . . . . 932 Ethiopian species . . . . . . . . . 934 1. ugandanus . . . . . . . . . . . 935 2. ampuliciformis . . . . . . 937 Nearctic species . . . . . . . . . . 940 1. serratus . . . . . . . . . . . . 941 2. testaceus . . • . . . . . . . . 941

VII

Table of contents

oculatus • . . • . . . • . • . . 945 3. Neotropic species •••.•••.• 947 1. maxiexophthalmicus .. 948 2.

ruficauda ..•••••.•.• 951

3.

poecilopterae ••••••• 952

4.

maximus . . • • • • . . . . . • • 954

5 •

s t ria tic ep s ••••••••• 956

6.

ru f i c ep s •••.•.•.•.•. 958

7. 8.

4.

barbarus ••.••••••.• 1018

Nearctic

species .•••.•.•. 1018

1.

inconsul tus . . . . . • . • 1019

2. 3.

do r sa lis ••••.•••••• 1 021 solaris . . . . . . • . • . . • ;f)24

4.

favreauae •••••••••. 1024

Neotropic

species ••.•.•.. 1027

1.

forestalis . • . • . . . • . 1027

nigroflavus ••.••••.. 961

2.

caraibicus .•.•.•.•• 1029

exophthalmicus •••••• 962

3.

ruber • • . . . . . . . . . . • . 1031

9.

quechuanus • . . . . • . • • • 965

4.

aterrimus • . • . • . • . • . 1031

10.

pi cescen s . • . • . • . . • . . 965

11.

andinus ..•.•.•••••.• 967

gibbosus • . • • . • . . • . . • 969 12. Genus Megadryinus .•••••.•.•.• 971

1.

species .•.•.•.•• 1033

amoenus .•.•••••.•.• 1 034

Neotropic

species • . . . . . • . 1037

1.

autumnalis •.•.•.•.• 1041

magnificus ••••.••... 971

2. 3.

bol ivianus . . • . . • • . • 1041 neotropicus •••....• 1044

4.

bicolor •••••••••.•. 1045

Genus Alphadryinus .•••••.•••• 973 Palaearctic

species . . • . . . • 973

1.

b ru e s i ••••••••••..•• 975

2.

balticus • . • . • • . • . . . • 978

Ethiopian

species ••••.•... 980

1. aberrans .•.••••••••• 980 Oriental species . . . • • • . • • . 982 1. asiaticus •••.•.•.••• 982 2.

Nearctic

species ..•.•.••• 971

Neotropic 1.

Genus Perodry i nuS . . • . • . • . . . • 1033

gibbosus . . . • . • . • • . . . 985

Neotropic

Palaearctic 1.

species . • . . . . 1046

zherichini •..•.•••. 1046

Subfami ly Transdryininae . . . . 1048 Genus Transdryinus •••••.•.•• 1048 Australian species . . • . • . . 1048 1.

notogeicus • . . . . • . . . 1048

species • • . • . • . . . 985

Subfami Iy Gonatopodinae ..•.• 1050 Genus Neodryinus • . . . • • . . • . • . 1065

1.

lamellatus •.•.•.•.•• 986

2.

panamensis . . . • . • . • . • 989

3.

parvus • • . • . • . . . . . . • • 989

4. 5.

pegnai . • . . • • • . • . • • . • 991 brasi I ianus . . • • • . . . . 992

6. 7.

rufus •••.••••••.•••• 995 ferrugineus •..•.•••• 995

8.

piceus • . • . • . • • • • . . . . 996

Australian

Genus Cretodryinus • . . . . • . . . • 1046

species ••.••••• 998 1. planus . . • . • . • • . • . • • . 998 Genus Mesodryinus ••.••.•••.• 1000 Palaearctic species .••••. 1001 1. dayi • • . • . • . . • . . . • . . 1003 2. niger •••••••••••••• 1005 Ethiopian species . • . . . . • . 1007 1. aethiopicus ••..•.•• 1008 Or i en t a I sp e c i e s . • . • . • • . • 1 01 1 1. 0 r i en t a lis • . • • . . . • . 1 0 1 1 2. peterseni •••••••••. 1015 3. viet. • . . • • . • . . • . . . • 1016

Ethiopian

species ••••.•.• 1067

1.

an t i quu s . . • . • • • • . • . 1069

2.

gigas • • . • . • . • • . . . . . 1071

Oriental species •.••.•.•. 1073 1.leptopus . . . . . . • . • • . 1076 2.

reticulatus . . • . . • . • 1076

3.

ch e I at us . • . • • . • • . . . 1 079

4.

rubescens . • • • . . • . . . 1081

5.

dolosus . • . • • • • . . • . • 1081

6.

robustus . • . • • . . . . • . 1082

7.

javanus . . • • • . . • . . . . 1085

8.

pseudodiffusus .•••• 1086

9.

diffusus . . . . • • . • • . . 1089

Nearctic species ••.•..•.• 1091 1.

typhlocybae • . . . . . . . 1091

2.

ball i • . • . . • . • • . . . . • 1095

Neotropic

species .•.•..•• 1097

1.

acuticol I is . • . • . . . . 1098

2.

dominicanus •.•..•.• 1101

Mem.

VIII

Amer.

Ent.

Inst.,

no.

37,

1984

albosignatus . . . . . . . 1 101 radialis . . . . . . . . . . . 1104

E t h i op i an

5.

peruvianus •••••.•.• 1107

Oriental

6.

maculicornis . . . . . . . 1107

3. 4.

7.

incaicus . . . . . . . . . . . 1109

8.

brachycerus ••••••.. 11 11

9.

trinitatis . . . . . . . . . 1111

Australian 1.

species •.•.•.. 1115

cheesmani . . . . . . . . . . 1116

2.

nelsoni . . . . . . . . . . . . 1118

3.

koebelei . . . . . . . . . . . 1120

4.

amboni . . . . . . . . . . . . . 1123

Genus Adryinus . . . . . . . . . . . . . . 1126 Ethiopian species . . . . . . . . 1126 1. cerrutii . . . . . . . . • . . 1128

1. 1.

sp e c i e s . . . . . . • . 1 1 77

ugandanus . . . . . . . . . . 1177 species . . . . . . . . . 1180

sinensis . . . . . . . . . . . 1180

Nearctic

species . . . . . . . . . 1180

1.

americanus . . . . . . . . . 1182

2.

mimus . . . . . . . . . . . . . . 1183

Neotropic 1.

species . . . . . . . . 1186

cubensis . . . . . . . . . . . 1188

Austral ian 1.

species . . . . . . . 1189

vitiensis . . . . . . . . . . 1191

Genus Pseudogonatopus . . . . . . . 1191 Palaearctic species . . . . . . 1193

1. 2.

fulgori . . . . . . . . . . . . 1194 camelinus . . . . . . . . . . 1197 rosellae •••...••.•• 1199

2.

nigerianus • . . • • . . . . 1130

3.

3.

bellicosus . . . . . . . . . 1132 owen i . . . • • . . . . . . . . . 1 135

4.

flavifemur . . . . . . . . 1202

5.

ortholabi s . . . . . . . . . 1203

4. 5.

6.

f

species . . . . . . . . . 1138

7.

dromedarius . . . . . . . . 1209

1.

gracilis . . . . . . . . . . . 1138

8.

albosignatus . . . . . • • 1209

2.

meridionalis . . . . . . . 1139

9. 10.

augustae . • • • • . . . . . . 1212 distinctus . . . . . . . . . 1212

1 1. 12.

P r i e sn e r i •••..•.••• 1 2 1 6 septemdentatus . . . . . 1217

rugidorsal is •••••.. 1135

Nearctic

Genus Echthrodelphax • . . . . . . . 1140 Palaearctic species . . . . . . 1140

0

ca r i lei . . . . • • • . . . 1 206

1. 2.

fa i r ch i I d i i . . . . . . . . 1 1 41 ba e t i cu s • . . . . . . . . . . 1 142

3.

longicollis ....•••• 1144

4.

hortusensis . . . . . . . . 1147

1.

pilosus . . . . . . • • . . . . 1222

5.

italicus . . . . . . . . . . . 1150

2.

perkinsiellavorus •• 1224

Ethiopian species . . . . . . . . 1152 1. migratorius . . . . . . . . 1152

3.

seyrigi . . . . . . . . . . . . 1225

4.

polli . . . • • . . . • • . . . . 1226

5.

ruens . . . . . . . . . . . . . . 1228 acutus . . • . . . . . . . • . . 1230

2. afer . . . . . . . . . . . . . . . 1155 3. obscurus . . . • . . . . . . . 1155 Ori enta I speci es • • • . . . . . • 1157 1. ru f us . . . . • . • . . . . . . . 1 1 57 2. fairchildii . . • . . . . . 1158 3. laotianus •...•••••• 1158 Nearctic species . . . . . • . . . 1160 1. krombeini . . . . . . . . . . 1160 Australian species . . . • . • • 1161 1. fa i r ch i I d i i . • • • . . . . 1 1 61 2. nigricoll is . . . • • . . • 1166 bi fasciatus . . . . . . • . 1169 4. pel op s . • . . . . . • • • . . • 1 1 69 Genus Acrodontochelys . . . . . . . 1172 Palaearctic species . . . . . . 1174 1. b ou c ek i . . . . . . . . . . . . 1 1 74

13.ligusticus •••..••.. 1219 Ethiopian

6.

species . . . . . . . . 1221

7 •

sp i ra cu I a r is .•••••. 1 233

8.

similis • • . . . . • . • • . . 1234

Oriental 1.

species . . . . . . . . . 1235

sarawaki . • . • . . • • . • • 1238

2.

sarawakensis . . . . . . . 1241

3.

nudus •.•••.•.••..•• 1245

4. 5.

malesiae . . . . . . . . . . . 1248 validus . . . . • • . . • . • • 1248

6. 7.

lankae • • . . . • • . . . . . . 1249 muiri . . . . . . . . . . . . . . 1251

8.

hospes . . . . • • . . . . . . . 1254

9.

sogatae . . . . . . . . . . . . 1254

10.

pusanus . . . . . . . . . . . . 1255

Nearctic

species .•.•••••• 1257

IX

Table of contents

1. 2. 3. 4. 5.

1258 freytagi canadensis . . . • • • . . . 1260 s joe s t ed t i . . . . . . . . . 1 263 ari zon i cus . . . • . . . . . 1266 ontarianus . . • • . . . . . 1269

6.

stenocrani . . . . . . . . . 1271

7. 8.

au t ox en ob ius .••.••. 1273 magnus . • . . . . . . . . • . • 1275

Neotropic

species . . . . . • . . 1276

1.

variistriatus . • . . . . 1276

2.

flavus . . . . . . . . • . . . . 1277

3.

delphacidi s . . . • • • . . 1279

4.

muesebeck i . . . . . . . . • 1280

5.

morenoi . . . • . . . . . . . . 1284

Australian species •••••.. 1284 1. nigricans . . . . . . . . . . 1287 2.

hospes . . . . . . . . . . • . • 1 290

3.

dichromus . . . . . . . . . . 1293

Neotropic 1.

species . . . . . • • . 1342

longichelatus

1342

Genus Apterodryinus . . . . . . . . • 1342 Palaearctic 1.

species . . . . • • 1344

gobiensis . . . • . • . . . . 1344

Ethiopian

species . . . . . . . . 1344

1.

somerset i . • . . . • • • . . 1345

2.

richardsi . . . . . . . . . • 1346

3.

rufulus • . • . . . . . . . • • 1347

Oriental 1. 2.

species . . . • • • • . . 1348

as i at i cu s . . . . . . . . • • 1 350 insularis . . . • . • • • • . 1350

Nearctic species . . . . • . . . . 1350 1. torvus . . . . . . . . . . . . . 1353 2.

secundus . . . . . . . • . . . 1355

3.

cal ifornicus . . . . . . . 1357

Neotropic

species ••••.••. 1358

1.

citrinus . . . . . . • • . . • 1359 arnaudi • . . . . . . . . • . . 1362

4.

saccharetorum . . . . . . 1294

2.

5.

kurandae •.••••••••. 1296

3 .

t est a c eu s ••••••..•• 1 362

6.

juncetorum . . . . . . . . . 1298

4.

menkei . . . . . • . • . . . • . 1364

7.

plurisulcatus . . . . . . 1299

5.

tijucanus . . . . . . • . . . 1365

8. opacus . . . . . . . . . . . . . 1302 '::;enus Donisthorpina . . . . . . . . . 1304

6. 7.

longicornis . . . . . . • . 1367 bellottii . . . . . . . . . . 1369

Palaearctic 1.

Oriental 1.

species •••..• 1305

pa I I ida . . . . . . • . • . • • 1306 species . . . . . . . . . 1308

asiatica • . . • • • • . . • . 1309

::;enus Agonatopoides . . . . . . . . . 1310 Palaearctic 1.

2. 3.

species •••.•• 1312

felix . . . . . . . . . . • . . 1312 sol idus . . . . . . . . . . . . 1316

canariensis . . . . . . • . 1318 Ethiopian species . . . . . . . . 1321

Austral ian species •••.••• 1371 1. not ogei cus . . . . . . . . . 1372 2.

insularis ••.••••••• 1374

Genus Dicondylus . . . . . • . . . . . . 1374 Palaearctic

dichromus . • • . • • . . . . 1376

2.

bicolor . . . . . . . . . . . . 1378

Ethiopian 1.

Oriental

fuscus • . . . . . . . . . . 1321

2. 3.

acuminatus . . . . . . . . . 1324 cafer . . . . . . . . . . . . . . 1325

Nearctic

4.

ruber . • . • • • . . . . . • • • 1327

Neot rop i c

5.

ceres . . . . . . . . . . . . . . 1328 species . . . . . . . . . 1330

1. 1.

1.

species . • . . . . . . 1383

pallidior . . . . . . . . . . 1383

1.

Oriental

species . . . . . . 1375

1.

species . . . . . . . . . 1384

indianus . . . • . • • • . • . 1384 species . . . . . . . . • 1387

americanus • . . . . . . . . 1388 spec i es •.••••.. 1390

nigrithorax . . . . . . . . 1390

Austral ian

species . . . . . . . 1392

1.

bengalensis . . . . . . . . 1333

2.

borneanus . . . . . . . . • . 1333

2.

rufus . . . . . . . . . . • . . . 1398

3.

insularis . . . . . . . . . . 1334

3. 4.

dubius . . . . . • . • . . • • . 1399 kiefferi . . . . . . . . . . . 1399

5.

primitivus . . . • . . • . . 1401

4. pyri Ilae • . • • . . . . . . . 1336 Nearctic species . . . . . . . . • 1338

1.

perkinsi . . . . . . . . . . • 1394

1.

synchromus . . . • . . • • . 1339

6.

oceanicus . . . . . • . . . . 1403

2.

calidus . . . . . . . . . . . . 1339

7.

alpinus . . . . . . . . . . . . 1404

x

Mem.

Amer.

En!.

Genus Haplogonatopus . • . • . . • . 1408 Palaearct i c

speci es ...••• 1408

1. 2.

apicalis ••.•••••.•• 1409 oratorius . . . . . . . . . . 1409

3•

at ra t us ••..••••.. • '. 1 41 3

Ethiopian

species . . . . . . . . 1415

Ins!.,

no. 37,

1984

Austral ian species . . . . . . . 1473 1. anomalus . . . . . . . . . . . 1473 Genus Trichogonatopus . . . . . . . 1473 Nearctic species . . . . . . . . . 1475 1. simulator . . . . . . . . . . 1475 Neotropic species . . . . . . . . 1475 1.

rich a rd s i • . . . • • . . . . 1 476

species . . . . . . . . . 1416

2.

hi sp i du s •..••••.••• 1478

ap i ca lis .•••••••••. 1 41 7

3.

pall idi tarsi s . . . . . . 1481

species . . . . . . . . 1421

4.

raptor . . . . • • • . . . • • • 1485

1.

insularis . . . . . . . . . . 1423

5.

albomarginatus . . . . . 1485

2.

hernandezae . . . . . . . . 1423

6.

neotropicus . . . • . . • . 1487

Australian species . . . . . . . 1424 1. v i t i en sis . . . . . . . . . . 1 425

7.

rubriceps . . . . . . . ... 1488

1.

katangae . . . . . . . . . . . 1415

Oriental 1.

Neotropic

2.

apicalis . . . . . . . . . . . 1427

Genus Pentagonatopus . . • • . • . • 1427 Nearctic 1.

species . . . . . . . . . 1428

borealis •••...••••. 1428

Genus Epigonatopus • . . . . • . . • • 1490 Austral ian species . . . . . . . 1490 1.

fallax . . . . . . . . . . . . . 1491

2.

sol i tarius . . . . . . . . . 1493 Genus Esagonatopus . . . • • • • . . • 14 93

montanus . . . . . . . . . . . 1430

Nearctic species . . . . . . . . . 1494 1. niger . . . . . . . . . . . . . . 1495

Genus Plectrogonatopoides ... 1431 Palaearctic species . . . . . . 1431

Genus Gonatopus . . • • • . . . • • • . . 1498

1. ch 0 r tin is •...•••••• 1431 Nearctic species . . . . . . . . . 1433 1. nearcticus . . . . . . . . . 1433

Palaearctic species . . . . . . 1502 1. vistosus . . . . . . . . . . . 1508 2. horvathi . . . . . . . . . . . 1509

Austral ian 1.

species . . . . . . . 1429

Genus Tetrodontochelys . . . . . . 1434 Palaearctic

species . . . . . . 1436

2.

perdebilis . . . . . . . . . 1497

3•

dod e r

4.

bi I ineatus .••...••• 1513 distinguendus . . . . . . 1513

0

i .••..••••••• 1 51 1

1.

p ed est r is ..••••••.. 1 437

5.

2.

sakai i . . . . . • • • • • . • . 1441

6.

striatus . . . . . . . . . . . 1518

3.

pulicarius . . . . . . . . . 1442

7.

vollenhoveni

4. 5. 6.

pseudoaudax •••••••. 1446 aegyptiacus . . . . . . . . 1446 audax ..•.••••••••.• 1 447

8. 9 •

iranicus . . . . . . . . . . . 1523 pop ov i • • . . . . • • • • . • . 1 523

10.

a Ib

Ethiopian species . . . . . ... 1449 1. ochreus . . . . . . . . . . . . 1450

11. 12.

suarezi . . . . . . . . . . . . 1528 dimidiatus . . . . . . . . . 1528

2.obscurus . . . . . . . . . . . 1452

13.

atlanticus . . . . . . . . . 1530

3.

14.

uni I ineatus . . . . . . . . 1532

rubripes . . . . . . . . . . . 1454

0

1521

lin ea t us . . . . • • . 1 527

4.

zai rensi s ••.•••...• 1454

15.

aegyptiacus . . . . . . . . 1533

5.

nigrior •.••.••.•••• 1458

16.

plumbeus . . . . . . . . . . . 1535

6.

afer . . . . . • • • • . . . . . . 1458

17.

barbatel Ius .•..•••. 1538

7.

varipes . . . . . . . . . . . . 1459

18.

ater •...•••..•••••• 1538

Oriental species . . . . . . . . . 1461 1. lucens . . . . . . . . . . . . . 1461

19.

I indbergi •••.•..••• 1541 lunatus • . . . • . . . • . . . 1541

20.

species . . . . . . . . . 1463

21 •

t en e r i f e i •.•••••..• 1 548

1.

plesius . . . . . . . . . . . . 1465

canari ensi s .•••••.. 1550

2.

unicus ..••••••••••• 1468

22. 23.

wit t me r i •...••••••• 1 551

3.

pecul iaris . . . . . . . . . 1470

24.

nigrosignatus . . . . . . 1554

Nearctic

Table of contents

25. planiceps •••••••••. 26. campestris •.•....•• 27. subtilis •••..•.•.•. 28. spectrum .••.• •.•• •• 29. bernard i . • . . . • . . . • . 30. formicarius ••.•.••. 31. mongolicus • . . • . . . . . 32. longus . • . . • . • . • . . . . 33. nivosus • . . • . . . . . . • . 34. brunneicollis .•.•.. 35. chersonesius •...••. 36. sepsoides . • . . . . . • • . 37. rhaensis • . . . . . . . . • . 38. barbieri •••.••....• 39. schenklingi . . . • . . . . 40. graecus • . . . • . . . . • . . Ethiopian species . . . . . . . . 1. meridionalis .•.••.. 2. longiusculus ••••... 3. austral is •• .•• ..•.. 4. pilosipes . . . . . . . . . . 5. guigliae ••.••.••••. 6. seyrigi . • . . . . . . . . • . 7. ankaratrus . . . . . . . . . 8. capensis •.••.••.... 9. mo sse Ii . . . • . . . . . . . • 10. rubrithorax . . . . . . . . 11. hyalinus ••.•.•••••. 12. ridens . . . . . -. . . . . . . . 13 • c ommu n is ••..••.•••. 14. luteipes . . . . . . . . . . . 15. bekilyanus . . . . . . . . . 16. incogni tus . . • . . . . . . 17. I eptothorax . . . . . . . . 18. a e t hi op i cu s ..•.••.. 19. tananarivei . . . . . . . . 20 . a rma t us . . . • . • • • . . . . 21. okahandjae . . . . . . . . . 22 . j oh n s i . . . . . . . . . . . . . 23. taylori . . . . . . . . . . . . 24. upembanus . . . . . . . . . . 25. benoi t i • • • . . . . . . . • . 26. trochantericus ...•. 27. opacus . . . . . . . . . . . . . 28. patrizii .••••••••.. 29. turneri . . . . . . . . . . . . 30. ericeti . . . . . . . . . . . .

1554 1556 1558 1560 1562 1564 1567 1567 1570 1570 1572 1573 1582 1585 1587 1587 1587 1590 1590 1593 1595 1596 1598 1601 1601 1 603 1605 1607 1607 16 10 1611 1612 1613 1615 1619 1620 1 623 1623 1 626 1628 1628 1631 1631 1634 1634 1637 1637

XI

Oriental species . . . . . . . . . 1. yasumatsui •..••.•.• 2. attenuatus . . . . . . . . . 3. schenkl ingi . . . . . • . • 4. mysorensis . . . . . . . . . 5. plebeius . . . . . . . . . . . 6. perpolitus . . . . . . . . . 7. javanus . . . . . . . . . . . . 8. cristatus . . . . . . . . . . 9. abjectus . . . . . . . . . . . 10. daunus . . . . . . . . . . . . . 11. phi I ippinus •.••..•• 12. maurus . . . . . . . . . . . . . 13. medius . . . • • . . • • . . . • 14. lucidus . . . . . . . . . . . . 15. magnus . . . • . • . . . • . . . 16. sitae . . . . . . . . . . . . . . 17. indianus . . . . . . . . . . . 18. thai . • • . . • . • . . . . . . . Nearctic species . . . . . . . . . 1. clavicornis . . . . . • . . 2. cyphonotus . . . . . . . . . 3. mayori . . . . . . . . . . . . . 4. portalensis . . . . . . . . 5. translucidus . . . . . . . 6. echo . . . . . . . . . . . . . . . 7. elongatus •••.••••.. 8. freytagi . . . . • . • • • . . 9. contortulus . . . . . . . . 10. planipes . . . . . . . . . . . 1 1. m i mo ide s . . . . . • . . • . . 12. agropyrus . . . . . . . . . . 13. frequens . . . . . . . . . . . 14. areolatus . . . . . . . . . . 15. harpax ••.•••..••... 16. solitarius . . . . . . . . . 17. herbarum . . . . . . . . . . . 18. paraleptias . . . . . . . . 19. pseudochromus . . . . . . 20. flavifrons . . . . . . . . . 21. rap ax .•..••.....••• 22. ashmeadi . • . . . . • • . . . 23.0mb rodes . . . . . . . . . . . 24. moni lis ••..••..•••. 25. argyrias . . . . . . . . . . . 26. curriei . . . . . . . . . . . . 27. innitidus . . . . . . . . . .

1639 1641 1641 1645 1647 1647 1650 1652 1652 1655 1656 1656 1660 1662 1662 1665 1665 1667 1668 1670 1673 1676 1679 1679 1682 1683 1684 1685 1688 1689 1 693 1695 1697 1700 1 703 1703 1704 1709 1712 1712 1715 1715 1720 1723 1724 1724 1 727

XII

Mem.

Amer.

En!.

28.

brooksi . . . . . . . • . • • . 1728

29.

pal I idiceps •••••••. 1729

Neotropic

species .•••.•.. 1730

1.

fernandinae • . . . . . . . 1732

2.

breviforceps .•••••• 1735

Ins!.,

no. 37,

1984

cont ortus . . . • • • . . . . 1768

22. 23.

bol ivianus . • • . . . . . . 1768

24.

autumnalis .•••••••• 1770

Australian

species . . . • . . . 1771

1.

melanias . . • • . . . . . . . 1772 williamsi . . . . . • • • . • 1774

3.

a9ilis • • . . . . • • . . . . • 1737

2.

4.

peruvianus . • . . • . . . . 1738

3.

zealandicus . . . . • • • . 1776

5. 6.

sp i no I a i . . . . . • • • • • • 1 739 wi II inki . . . . . . • • . . . 1783

4. 5.

9i9as •...•••••••••• 1779 p I i ca t us . • . . . . . . . . . 1 783

7.

pseudorbitalis . . . . • 1742

6.

rossi • . • . • . . . . . . . . . 1783

8.

orbitalis .••••••... 1745

7. 8.

naumann i • . . . . . . • • • • 1785 papuanus . . . . . • . . . . . 1786

9.

opacithorax . . . . • • • • 1745

10.

purpurascens •..•••• 1746

1 1. 12.

town e 5 i . . . . . . . . • . . . 1 748 apicalis . . . . . • • • • • • 1750

13. 14.

bartletti ..•.•••••• 1751 flavipes . . . . . • . • • . . 1752

9.optabilis . • . • • • . . . . 1786 10. 11.

decoratus . . . . . • . . . • 1788 austral iae . . . • • . . . • 1790

12.

pulcherrimus ..••••• 1791

Genus Gynochelys . . . . . • . . . • . . 1792 Ethiopian

species . . . . • • . . 1793

15.

campbelli .•••.••••• 1755

16.

tristis . • • • • . . . . • . . 1757

17.

doellojuradoi ••...• 1757

Subfami Iy Apodryininae . . • . . . 1796

18.

si Ivestri i •.••••..• 1759 lacualis . • . . • • . . . . • 1762

Genus Apodryinus .•••••••••.. 1796

1.

braunsi . . . . • . . • • • • • 1793

Neotropic species . . • . • . . • 1796 19. 1. masneri •..••••••••. 1796 20. arlei ••••.•..•••••• 1764 cilipes . • . . • . . • • . . • 1764 21 • Nomina dubia and incertae sedis species • • . . • . . . • . . . . . . . . . . • • . . 1798 Nomina nuda • • • • • • • • • • • • • . • • • • • • • • • • • • . . . . . . . . . • • • • . . . . . • • . • • • • 1829 F0 ss i I

dry i n ids . . . . . . . . . . • • • • . . . . . • . • . . . • • . . . . . . • . . . • • . . • . • . . . 1 830

Parasites of Host

hopper

dryinids . . . . . . • . • • . . . . . . . . . . . • . • . • • • . . . • . • . . . . • . • 1831 records • . . • . . • . • . . • • • . . . . . . . . . . . . • • . . • • . • . . • . . • • • • 1834

Ref e r en c e s . . . • . . . • . . . . . . . . . . . . . . . . • • . . . . . . . . . . . . . . . . . . . . . . . . . . 1 847 Abb rev i at i on s . . . . • • • • • • • • • . . . . • . . • • • • • • • • • • . . . . . . . . • • • • • • • • . . . 1869 Index to hosts . . . • . . . . . . • . • • • . . • . . . . . . . . . . • . . . . . . . . • • . . . . . . . . • 1869 Index to the parasites and predators of dryinids . . . . . . . . . • • • • . 1875 Index

to dryinids . . • • • . • • . • . . . . . . • . . . . . • . . . • • • • . • . . . . . • . . . . . • • l877

A REVISION OF THE

DRYINIDAE

(HYMENOPTERA)

by f.iassimo Olmi

Institute of Plant Protection, Via 5.

Camillo de Lellis,

University of Tuscia, 01100 Viterbo,

Italy

I iHRODUCT I ON

The Dryinidae, one of the smaller fami I ies of the acu leate ,:jymenoptera, is included in the superfamily Chrysidoidea (sensu Day 1977). The

literature on dryinids is wide.

or unrel iable because of

inc'lr rect

Jnrel iable taxonomic descriptions. :roublesome. ~evision

of

::>y Kieffer .J,

c),

a

The more useful

key

.c,ustralian,

however,

(1939),

the family,

is a

to the European

is useless

identifications or

incorrect and

Desides Richards'

monograph of the world species

treatment of the I~earctic dryinids by

a

Oriental

it,

The nomenclature is often

literature on

the British species (1914b),

Much of

or questionable

Fenton

(1918a,

Aphelopus by Jervis (1977), some studies on

and Nearctic dryinids by

R.C.L.

Perkins

(1903,

1906a,

1906b, 1907, 1912), another revision of the British species by J.F. Perkins (1976), a study on the classification of the dryinids by Richards (1953), a -<ey

to

the Russian species by

c!ryinids by rreytag useful

Freytag

(1982).

(1977,

Kieffer's work

because of

Ponomarenko

1978,

1980),

is unreliable for

its completeness.

><earctic species by Ashmead

(1978)

and studies on Nearctic

Freytag and Back

The rudimentary

(1893)

(1977),

identifications,

Giri

but

it

and is

revision of the

is not recommended.

The object of this paper is to revise the species already t.iescribed, to assemble the biological information about them and to describe the new species represented

in

the collections.

world species reasonably Little

The user wi II

find

treatment of the

complete.

is known of the males and the keys are therefore

unsatisfactory.

The classification

association of the sexes this problem.

Only

is based mostly on

the females.

is difficult and much further work

breeding experiments can

finally

The

is needed on

answer these

questions.

MATERIAL EXAMINED

The keys and the descriptions are based on numerous collections.

Those

individuals and

material

examined

institutions from which

in

Mem.

2

Amer.

Ent.

Inst.,

material was borrowed are listed below abbreviation used throughout the text: AD

South Austral ian Museum,

AM

American Museum of Natural

AS

Academia Sinica,

37,

1984

together with

Adelaide

Zoological

no.

the appropriate

(South Austral ia)

History,

New York

Institute,

Peking

(N.Y.)

(China)

B

Bernice P. Bishop Museum,

BA

Museo Argentino de Ciencias Naturales "Bernardino Buenos Aires

Honolulu

(Hawaii) Rivadavia",

(Argentina)

BE

Department of Entomology,

University of Wageningen

BI

Ferdinando Bin collection,

Perugia

BM

British Museum

BN

Zoologisches Forschungsinstitut und Museum Alexander Koenig, (German Federal

(Natural

(Holland)

(Italy)

History),

London

(England)

BO BR

Istituto di Entomologia dell'Universita, Bologna (Italy) Department of Primary Industries, Indooroopilly (Queensland)

BU

Nat iona I Museum of Natura I History,

BY

Department of Bio!ogy,

CA

Cal ifornia Academy of SCiences,

CB

Australian National Entomology,

Bonn

r,epublic)

Internacional

(Hungary)

Waco

San Francisco

Collections of

Canberra

Budapest

Baylor University,

(Texas) (Cal ifornia)

Insects (ANIC),

CSIRO,

Division of

(Austral ia)

CI

Centro

CM

Museum of Comparative Zoology, (Massachusetts)

de Agricultura Tropical,

CO

Zoologisk Museum,

DA

Department of Entomology,

KObenhavn

Cali

(Colombia)

Harvard UniverSity,

Cambridge

(Denmark)

University of Cal ifornia,

Davis

(California) DL

Jiri

DU

National

Dlabola collection,

EB

Institut fur Pflanzenschutzforschung

Museum of

Praha

Ireland,

(Czechoslovakia) Dubl in

Landwirtschaftswissenschaften,

(I reland) der Akademie der

Eberswalde (O.O.R.)

ED

South African Sugar Association (South Africa)

Experiment Station,

EN

Istituto di Entomologia agraria ed Apicoltura dell'Universita, (I taly)

FO

Alessandro Focarile collection,

GE

Museo Civico di Storia i~aturale "Giacomo Doria",

Aosta

GV

Museum d'Histoire Naturelle,

Mount Edgecombe Torino

(Italy) Genova

(Italy)

Geneve (Switzerland)

HA

Martin-Luther-Universitat,

HE

Zoological Museum of the University,

Halle-Wittenberg

(D.D.R.)

HU IF~

Zoologisches Museum an der Humboldt-Universitat, Berlin (O.D.R.) Institut Royal des Sciences Naturelles de Belgique, Bruxelles

IT I(E

Cornell University, Ithaca (I"ew York) College of Agriculture, University of Kentucky,

KY

Entomological

LA

~Jatural

Helsinki

(Finland)

(Belgium)

Laboratory,

Kyushu University

History Museum of Los Angeles County,

(Cal ifornia)

Lexington (Kentucky)

(Japan) Los Angeles

3

Introduction

LE

Rijksmuseum

LI

Linnean Society,

van Natuurlijke Historie,

LN

Zoological

LO

(U.S.S.R. ) Museum d'Histoire naturelle,

LU

Zoological

MA

Department of Entomology,

London

Institute,

U.S.S.R.

Institute,

Leiden

(Holland)

(England)

Lund

Academy of Sciences, Lausanne

Leningrad

(Switzerland)

(Sweden) University of Wisconsin,

Madison

(Wisconsin) MD

Instituto Espanol

ME

National

de Entomologia,

MI

Museo Civico di Storia Naturale,

ML

Milwaukee Public Museum,

Museum of Victoria,

Madrid

Melbourne

(Spain)

(Victoria)

Milano (Italy)

Milwaukee (Wisconsin)

MO

Zoological

MP I: MR

Moscow Palaentological Institute, Moscow (U.S.S.R.) Claud Morley collection (part kept in Ipswich h'luseum,

Museum,

Moscow

(U.S.S.R.) Ipswich,

England) MT

Ecole Nationale Superieure Agronomique,

Montpell ier

MU

M.

(Japan)

Miura collection,

Shimane University

NA

Museo di

ND

Division of Entomology, Delhi

Zoologia dell'Universita,

Napoli

(France)

(Italy)

Indian Agricultural

Research

Institute,

New

(I ndia)

NI

Entomological

NZ

Department of Scientific and

Laboratory,

Kochi

University,

I ndustrial

Nangoku-City (Japan)

Research,

Auckland

(New

Zealand) OL OT

Massimo Olmi collection, Viterbo (Italy) Biosystematics Research Institute, Ottawa

OX

Hope Museum,

Oxford

(Canada)

(england)

P

Museum National

d 'Histoire ,',aturelle,

PG

National

\I'"atural

PH

Academy of r,atural

PK

:,adezdha Ponomarenko collection,

PL PO

Guido Pagl iano collection, Torino (I taly) Istituto di entomologia agraria dell 'Universita,

PT

Facultad de Ciencias Naturales y '''1useo,

QU RY

Queensland r'l,useum, Fortitude Valley (Queensland) Biological and Chemical Research Institute, Rydalmere

Museum

History),

Sciences,

Paris

Praha

Philadelphia r"oscow

(France)

(Czechoslovakia) (Pennsylvania) (U.S.S.R.) Portici

La Plata

(I taly)

(Argentina) (,',ew South

Wales) SA

Friedrich Sander collection,

SC

Department of Food and Agriculture,

SP

Spinola collection,

c/o

Jena

r..1useo

(D.

D.

R.)

Sacramento

f,egionale di

(California)

Scienze Natural i,

Torino

(I taly) ST SU

Naturhistoriska Riksmuseet, Stockholm (Sweden) Instituto de Aclimatacion, Almeria (Spain)

SW

Museo Nacional

SZ

Pier Luigi

TE

Department of Entomology, Station

de Historia Natural,

Scaramozzino collection,

(Texas)

Texas A.

Santiago Torino

(Chi Ie)

(I taly)

and M.

University,

College

4

Mem.

Amer.

Ent.

Inst.,

TN

Museo de Ciencias Naturales,

TO

Istituto e Museo di (I taly)

TR

Museo Regionale di Scienze Naturali,

TU

Instituto de Zoologia,

no.

S.ta Cruz

37,

1984

(Canary

Islands)

Zoologia sistematica dell'Universita,

Torino

Torino (Italy)

Universidad Nacional

de Tucuman,

Tucuman

(Argentina) TV

Musee Royal

TW

American

de I 'Afrique centrale,

UQ

Department of Entomology,

Entomological

Institute,

Tervuren

(Belgium)

Ann Arbor

(Michigan)

University of Queensland,

SI.

Lucia

(Queensland) VE

Museo Civico di Storia Naturale,

VI

Naturhistorischen Museum,

Wien

VR

"Iuseo Civico di Storia Naturale,

WA

National

WS

~jatuurhistorisch

Museum of Natural Museum,

Venezia

(Italy)

(Austria) Verona

History,

(Italy)

Washington

Maastricht

(D.C.)

(Holland)

ZO

Zoolog ische Staatssamm lung,

Munchen

YA

Keizo Yasumatsu collection,

Oonojo-shi

(German Federa I Repub I i c) (Japan)

ACKNOWLEDGMENTS

I have been much indebted to the following persons who helped with the collections, either by sending dryinids on loan, or acting as host when thei r museums were vis i ted: Kees van Achterberg (Le i den), Anders Albrecht (Helsinki), Paul H. Arnaud Jr. (San Francisco, California), Donald Azuma

(Philadelphia,

S. Baldridge (Waco, Texas), (Tervuren), Claude Besuchet Ferdinando Bin (Montpellier), A.J.M.

(Perugia),

G.R.

Brown

Pennsylvania),

Marcos Baez

(Tenerife),

Robert

Tony Bellotti (Cali), P. L. G. Benoit (Geneve), C. F. M. den Bieman (Wageningen),

Zdenek Boucek (Rydalmere),

Carnegie (Mount Edgecombe),

(London),

Ernst-G.

A.

Bournier

Burmeister (Munchen),

Achille Casale

(Torino),

Lennart

Cederholm (Lund), Raul Cortes (Santiago), I talo Currado (Torino), E.C. Dahms (Fortitude Valley), Roy Danielsson (Lund), Michael Day (London), Jean Decelle (Tervuren), (Bruxelles), Dorn

(Halle),

N.

F.

(New Delhi),

(Wien),

Ian Galloway

Delhi),

I
Mark A.

(Davis),

V.

Marjorie Favreau (Indooroopilly),

Antonio Giordani Soika

Karl-Johan Hedqvist (Praha),

(La Plata),

(Maastricht),

Christophe Dufour (Lausanne),

Farooqi (I~ew

Luis de Santis

Dingemans-Bakels

I
(Stockholm),

Jervis

Frank Koch

(Kobenhavn),

Matthews

(Adelaide),

(College Station,

N.

Arnold Menke

Y.),

(Gent),

M.K.

Giri

Steven Krauth F.

Leclant

(Praha),

(Praha),

(Santiago), Max Fischer

(Katmandu), J. Jelinek

(Paris),

(Madison,

(Montpellier),

Lynn S.

Wisconsin), Ole

Lubomir Masner (Ottawa),

(Washington,

Elvira Mingo Perez

D.C.),

(Madrid),

M. S.I.

Swaraj Ghai

(Fukuoka),

Simone Kelner-Pillault

D.C.),

Jan Macek

Texas),

(Venezia),

(Berl in),

(Washington,

Lomholdt

Geeraert

Yoshihiro Hirashima

(Cardiff),

Dessart

Dlabola

Mario Elgueta

(New York, E.

Paul

Jiri

Sammie J.

E.G.

Merritt

Laszlo Moczar

I,ltroduction

(Budapest),

Gordon

(Nangoku-City), O'Connor

(Dubl in),

Ibaraki), cheol

M.

Nishida

Gerald R.

Giuseppe

(Suweon,

Korea),

d'Entreves

(Torino),

L.

(Santiago), Ponomarenko (Pretoria),

(Moscow),

(Verona),

(Oxford),

Pechuman

Scaramozzino

(Ithaca,

(Oxford),

California),

C.N.

Francisco J.

(Ann Arbor,

(Auckland),

(Sydney),

Suarez

Michigan),

Carlo Vidano

Roy R.

Ulrich

(Torino),

Luis E.

(Bonn),

G.L.

Sander

(Gottingen),

(Jena),

Lucia,

Pier Luigi

Brisbane),

Snell ing

Gennaro Viggiani

Henry

Valentine

(Portici),

C.

Vogt

(Cambridge), Annette K. Walker (Auckland), Marius S. V,asbauer (Sacramento, California), Ingrid Wegener (Berlin), V. B. Whitehead Town),

S.C.

(Oonojo-shi, Prof.

Willemstein Japan), Paul

H.

(Leiden),

K.IV.R.

A.

Zwart

Freytag

Willink helpful

in sending

The scann i ng electron photographs were I wish especially disposal

to thank Prof.

the facilities of the

The

(Tucuman),

(Cape

Keizo Yasumatsu

(Wageningen).

has been

problems about Gonatopodinae and

M.J.

(Los Angeles,

(Sapporo),

E..IV.

Pena Nadezdha

Prinsloo

Ritzkowsky

S. Takagi

(Almeria), H.

S.

(St.

Jong-

(Genova),

(Bologna),

(London),

Schneider

(Torino),

Pietro Passerin

New York),

Friedrich VI.

rvrargaret A. Smithers

(Yatabe,

Roberto Poggi

Maria Matilde Principi

(Torino),

James P.

Akio Otake

(Budapest),

(Kobenhavn),

(Buenos Aires),

Toshihiko I,ishioka

Wisconsin),

Guido Pagliano

Jeni:i Papp

Owain Westmacott Richards

Arturo Roig Aisina

Townes

L.

Borge Petersen

Hawaii),

(Milwaukee,

Osella

Christopher O'Toole

Paik

Scobie

(Honolulu,

Noonan

5

in discussing

taxonomic

material. taken

at Pi acenza Un i vers i ty.

Giorgio Domenichini

for putting at my

Institute of Agricultural

Entomology.

illustrations of whole insects were drawn mostly

by Dr.

Falchi, partly by Mr. Luciano Couvert, the remainder by myself. indebted to Dr. Falchi and Mr. Couvert for their figures.

Nicolo I am

BIOLOGY

Most of what

is known of the biology of the Dryinidae

the research of R.C.L. Perkins (1903, and North America, Fenton (1918a, b,

is based on

1905, 1906a, 1906b, 1907) in Hawaii c,) in the U.S.A., Abdul-~
in France, Chandra (1978b, 1980a, 1980b, 1980c) in the Phi I ippines, Barrett, Westdal and Richardson (1965) in Canada, Behrisch (1952) and Strubing

(1956)

in

the German Federal

(1930,

1931,

1932,

1940),

Esaki

and Mochizuki

1975)

and Jervis

1933,

1934,

(1979a,

1935,

(1941)

1979b,

Republic,

1936),

Esaki

in Japan,

1980b,

Esaki

Richards

1980c)

and Hashimoto

and Sameshima (1939),

(1939,

Waloff

(1974,

in Great Britain,

Ponomarenko (1971, 1975b) in the u.s.s.n., Olmi (1976) and Currado and Olmi (1972) in Italy, Lindberg (1950) in Finland. Fragmentary notes have been

added by other workers. The dryinids are all parasitoids of iiolllopter<1 Auchenorrhyncha.

her chelate forelegs, She stings

it,

the female catches an

causing

insensibi I ity,

and

With

adult or nymph of the host.

lays an egg between

its two

Mem.

6

Amer.

Ent.

Inst.,

no.

37,

1984

overlapping thoracic or abdominal sclerites •• She may also bite the hopper with her mandibles (Richards 1939). Only the female of the Aphelopinae does not have chelate foretarsi, her front

and middle pairs of

and must catch and hold the hopper with

legs (Jervis 1980c).

The hopper soon

recovers and runs off. The egg

is usually

invisible.

I t may be regarded as morphologically

external. The larva brown,

is at first

green or yellow

composed of cast off added

to the sac.

invisible,

but within a

sac begins to appear.

larval

skins,

few

days a black,

This spheroidal

dark

sac is

and with each moult an extra piece is

There are normally five moults.

The host remains active,

but does not moult while its parasitoid

is

develop i ng. The more

Aphelopinae is rather different. The egg is much

life history of

internal

and at

least

the first-stage larva

pecul iar hypertrophied mass of host persists until consumed by

the parasite larva is in the

larva must pass

is surrounded by a

(trophamnion).

tissue

its final

through

it.

instar,

The

The

trophamnion

and all

food

larva breaks out of the

trophamnion in its late second instar to project from the host in a sac composed of

larval

skins.

The sac

is cyl indrical

its distal

in other dry in ids.

end

and of a

host,

trophamnion continues to isolate the larva from the host's

the

much softer tex ture than

with

rounded,

I nsi de the

haemocoel.

Just before the larva is fully grown, it eats out all the contents of the host, which usually dies firmly fixed to the food-plant. On completion of feeding, ground. soil.

the

larva splits open

I t moves quite actively

The white cocoon

the sac and crawls out or drops on and pupates on

is of dense silk,

lining. The manner of development of different from all other dryinids in is passed within the external

the host body,

larval

polyembryonic,

sac on

one egg

often with

an

inner closely-fitting

that

the entire

larval

feeding period

the consequent complete absence of

parasitized hosts.

In addition

to produce up

the species

to seventy

young

(Kornhauser 1919). The mature larvae bore their way through abdominal sternites of the host and pupate in the ground. There seems to be ei ther one or two generations a species wich are found the cocoon as mature

in

the the

Crovettia theliae (Gahan) is strikingly

with

dividing

the food-plant or in

temperate regions,

larvae or prepupae.

is

larvae

the

year of the

the winter being passed

in

Some dryinids overwinter as

first instar larvae (Jervis 1980b). Crovettia be in the pupal stage during the winter. I n

theliae (Gahan) is believed to tropical

regions

to be several

overlapping generations without diapause.

the dryinids,

however,

is considerably minimized during

there appear

The activity of the dry season

(Chandra 1980c). Many Authors have recorded

Gonatopodinae) this

in association with

is an example of mimicry,

the occurrence of several ants.

Donisthorpe

(1927)

species

(mostly

suggests that

because dryinids obtain protection from

Introduction

:heir likeness instance,

to the ants.

Anteon myrmecophilum (R.C.L. Perkins), for

appears to be definitely myrmecophi lous.

noppers which

frequently

resemblance

in

its actions:

"if a

vial,

are attended by

appearance to the ants.

four hind

legs,

face

to face,

soliciting food"

dryinids,

ants,

It parasitises

This resemblance

according

in pairs,

(R.C.L.

where ants and Gonatopodini

in a

glass

standing sub-erect on

Perkins 1905). (1939),

is enhanced by

together

stroking one another,

to Richards

leaf-

and bears some sl ight

number of the parasites be bred

they may be seen gathered

mouth,

7

their

licking each other's

In other species of

"it appears more probable that

have been found

together

it

is due merely

to

some overlapping of their habitats". The fema les of the dry i n i ds are parthenogenet i c as well In

(1919)

that mated females produce female progeny

progeny.

According

to Waloff

(1974)

female or male. Many fema I es are predatory feeding hosts

as sexua I.

Crovettia theliae (Gahan) it is assumed by Kornhauser

the case of

purposes only.

Adult

and virgin

females male

parthenogenetic offspring can be

as adu I ts and kill

leaf-hoppers for

Aphelopus, however, are not predacious upon

(Jervis 1980c). The hosts frequently

exhibit changes,

The reproductive organs are often pigmentation, reproduction

often

toward

both

internal

greatly modified,

that of the opposite sex,

usually ceases shortly

and external.

and changes in cuticle occur.

Host

after attack.

HOSTS ATTACKED

The dryinids are all

parasitoids of

fJomoptera Auchenorrhyncha

(Tables 1a,lb). Many published host records the host or the parasitoid was probably

ECONOMIC

IMPORTANCE

From the economic point of view reducing

the hopper popUlation by that a

parasitization.

greater mortality Only

adult

the value of the dryinids in

parasitization

the predaceous habits of the females. opinion

are unreliable because either

misidentified.

Several

is effected

in

is greatly enhanced by

Authors have expressed this way

than

Aphelopus are not predaceous upon hosts.

A number of species of dryinids exert an appreciable effect natural

control

the

by

of their hosts,

which

in

the

is often of considerable economic

importance.

fJaplogonatopus vitiensis R. C. L. Perk ins an d Pseudogona topus hos{Jes R.C.L.

Perkins were unsuccessfully

China,

respectively,

in connection

introduced with

into Hawaii

the biological

from Fiji

control

of the

and

8

Memo

Amero

Ent. Ins!., no.

37,

1984

(])

(])

(]) CO CO CO U U

QJ

u

._

0-

C

ro .- :0

0-

U

(])

co

U ._

~

6

~~~E-~

~ u 0 QJ :J -O:ZOLL

APHELOPINAE

x

x x

Aphelopus Crovettia

x

B I APHELOP I NAE Biaphelopus CONGANTEONINAE Conganteon Chel anteon Fiorianteon

ANTEONINAE

x x x x

Metanteon Deinodryinus

x x

Lonchodryinus Anteon

x x

x

Prioranteon BOCCHINAE

x

x

Bocchoides

Chelothel ius Bacchus

x

Mirodryinus Rad i i mancus Mystrophorus

x x

THAUMATODRYININAE Thaumatodryinus DRYININAE Dryinus Richardsidryinus Tri dry i nus Tabl e lao Hosts af Dryinidae

x xxxxxxxx

xx

xxxx x x x x

x x

x x x

9

Introduction

I

I

I

I

GJ

GJ

ro ro c ,-

GJ

C ,-

'GJ

U

0

ro C

,-

.D

'-

I

»

if)

0. 0

I

U

-0 L

GJ

ro

u 0. v ro » v - ::;;: f- -- roGJ GJ V GJ GJ GJ ro ro ro v v,- v,- v,- uro '-ro ,-

,-

GJ

I~

GJ

-

-

'- L U 0. 0

-

L GJ U ro :J

ro

GJ

V

'ro"

GJ ro ro v ,~I~ V

,,-

v

GJ

c

GJ 0. ro GJ GJ v v o ~ ro ro c v v v '- » ,v ro C ,,o.L ro ro GJI E u u ro '" u x 0 0. ro ro <Jl u ,,u if) '- 0 ,-

0

,-

,-

.D

'"

::;;:

U

U

G5

,-

0

U

~IGJ

GJ

ro

v,-

,-

f- --.J l1..

~egadryi nus

«

-

cr:

c u ,_ ,-

ro

V

L 0.

0

-

01 0

'-

0 01

-

GJ :J 0 l1..

z

x

Alphadryinus x

Mesodry i nus Perodryinus

Ix

Cretodry i nus TRANSDRYININAE Transdryinus

x

GONATOPODINAE Neodryinus

x x x x x

x x x x x

Adryinus Echthrodel phax

I

x

x

Acrodon toche I y s

x x

Pseudogonatopus

x x x

Don i sthorp i na

I

Agon a topo ides

x

Apterodry i nus

x x x

D i condyl us Haplogonatopus Pentagonatopus P lectrogona topoi des

x

Tetrodon tochel ys

I

Tr i chogona top us Epigonatopus Esagona topus

x

Gonatopus

x

Gynochelys APODRYININAE Apodryinus Table lb.

I

Hosts of

Dryinidae

I

I

10

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

Perkinsiella saccharicida K i rka I dy. rlphelopus albopictus Ashmead (= typhlocybae Muesebeck) was

sugprcane leaf-hopper, unsuccessfully with

introduced into New Zealand from

the biological

control

of the yellow apple

the U.S.A. leaf-hopper,

in connection

Typhlocyba

australis Frcgg.

MORPHOLOGY AND TERMINOLOGY

The terminology Reid

(1941)

except

in this work follows

that

that of Richards

(1977)

and

the naming of the wing cells and veins follows

system proposed by Townes (1969). SCULPTURE. The surface of the body

in

Dryinidae

is often sculptured

the in a

variety of ways. The principal terms used in describing the sculpture are listed as follows: without sculpture (smooth); punctate; granulate (Fig. 1); rugose; Eady

(1968)

reticulate rugose is useful

(Fig.

2);

striate

(Fig.

3).

The paper by

in explaining the descriptive terms.

HEAD AND I TS APPENDAGES. The head of Dryinidae is characteri st i ca II y orthognathous with its long axis vertical and the mouthparts directed downwards. head.

The romnoU:-Jrl e'J,c, occupy

the greater part of the side of the

Sometimes numerous hairs arise from between

the head between

the eyes

lower face and upper face.

is termed

face;

it

the facets.

The front of

is made up of the c I ypeus,

The antennae are articulated

in sockets

(antennal sockets) situated very low down, near the clypeus. The antenna I socket is surrounded by a sulcus defining a narrow antennal sclerite. The clypeus includes the greater part of the face below the antenna I sockets. Laterally and dorsally the clypeus is defined by the curved epistomal suture. On each side, near the top of the epistomal suture, is a small pit, the anterior tentorial r'it. The pit marks the point where an arm of the tentorium, the internal skeleton of the head, is invaginated. The ventral margin of the clypeus appears rounded, bidentate or shallowly emarginated as follows: 1) bidentate clypeus: only in females of Dryininae (except for Alphadryinus and jljesodryinus) , Prioranteon (Anteoninae), part of Gonatopus, Trichogonatopus, Apterodryinus, Adryinus, iVeodryinus (Gonatopodinae) ; 2) shallowly emarginated clypeus: in females of Alphadryinus and jljesodryinus (Dryininae), Mirodryinus and l?adiimancus (Bocchinae), Thaumatodryininae, TransdryininiJe,. in males of Dryininae (except for part of Dryinus and Mesodryinus) , Thaumatodryininae, Anteoninae (except for part of Anteon and Lonchodryinus), part of iVeodryinus (Gonatopodinae),. 3) rounded c I ypeus: in fema les of Apodryininae, Aphelopinae, Biaphelopinae, Conganteoninae, Gonatopodinae (except for part of Gonatopus, Trichogonatopus, Apterodryinus, Adryinus, Neodryinus), Anteoninae (except for Prioranteon), Bocchinae (excep t for J1lirodryinus and Radiimancus),. in ma I es of Aphelopinae, Conganteoninae, Bocchinae, part of

"'~,

Fig,

1- Variety

of sculpture:

............

'-"'-~,

granulate

, ,

...... "

(male of Gonatopus lunatus

Klug).

I/i ern.

12

.

Fig.

'

Ent.

I nst.,

no.

37,

1984

, ...

2 -

Variety of sculpture:

oratorius Westwood).

,

Amer.

reticulate rugose

(male of

IJaplogonatopus

13

Introduct i on

Fig.

3 -

Variety of sculpture:

Westwood) .

s triate (female of

Gonatopus s epsoides

14

Mem.

Amer.

En!.

Inst.,

no.

37,

1984

Dryinus and 1\1esodryinus (Dryininae), Gonatopodinae (except for part of Neodryinus) , part of Anteon and Lonchodryinus (Anteoninae). The ocelli I ie near the top of the face; they are three in number. They vary

much

in position;

the distance from the outer edge of a

lateral

ocular-ocellar line, OOL; the distance between the inner edges of the two I atera I ocell i is the postocellar line, POL; the distance from the inner edge of the anterior ocellus to the inner edge of a lateral ocellus is the ocellar line, OL; the distance of the lateral ocell i from the occipital margin is the ocellar-occipital line, OPL. In Apodryininae the ocell i are absent. The labrum ish i nged to the ventra I marg i n of the cl ypeus and is a transverse fl ap. In Dryinidae there is a ocellus to the compound eye

tendency inner)

for

the

is the

I ine of attachment to move upwards on the under (or

side of the clypeus so that much or all

The part of the face under the antennal

lower face.

The epistomal

sclerites but antennal

it

in

labrum

is hidden. as the

suture sometimes runs close to the antenna I

is often well

sclerites

of the

sockets may be known

separated from

the following

them.

It

is close to

the

dryinids:

females of Thaumatodryininae, Lonchodryinus (Anteoninae), Aphelopus (Aphelopinae) ; b) ma I es of Anteoninae and Aphelopus (Aphelopinae); a)

The epistomal

suture

is separated from

the antennal

sclerites in

the

following dryinids: a) fema I es of Dryininae,

Gona topodinae, Anteoninae (except for Lonchodryinus), Conganteoninae, Transdryininae, Biaphelopinae, Crovettia (Aphelopinae), Bocchinae, Apodryininae; b) males of Dryininae, Thaumatodryininae, Gonatopodinae, Bocchinae, Conganteoninae, Crovettia (Aphelopinae); The part of

the face above the antennal

upper face.

It

area between

is possible to define the the antennal

sockets and

sockets may be known as the

frons as the central part of the the median ocellus.

In

various

Dryinidae keel s or impressed lines may be developed on parts of the frons. Sometimes frontal

keels and

I ines are found

to be taxonomically

important.

The main central area of the upper face is often divided by a central keel (the frontal line). In many Dryinidae (males in particular) the frons is more or

less excavated

excavations are the which bears

the

to receive the basal

part of

the antennae;

these

antennal scrobes. The top of the head is the vertex

lateral

ocelli

and

is usually

limited posteriorly

by

the

occipital carina. The occipital carina is absent in the following dryinids: a) females of Transdryininae, part of Gonatopodinae, part of Dryinus (Dryininae), part of Tridryinus (Dryininae), Apodryininae; b) males of Gonatopodinae (except for Echthrodelphax and part of Neodryinus) ; The occipital

carina

is incomplete in

the following

dryinids:

Chelothelius (Bocchinae) , part of Thaumatodryininae, part of Gonatopodinae, part of Tridryinus, Alphadryinus, Mesodryinus, Dryinus (Dryininae) ; b) males of Mesodryinus, i2ichardsidryinus, part of Tridryinus and Dryinus a)

females of

Introduction

(Dryininae),

part of

The occipital

carina

15

Neodryinus (Gonatopodinae); is complete

in

the following

dryinids:

Anteoninae, Conganteoninae, Aphelopinae, Biaphelopinae, Bocchinae (except for Chelothelius), part of Thaumatodryininae, )jegadryinus, Richardsidryinus, Perodryinus, part of Tridryinus, Alphadryinus, fllesodryinus, Dryinus (Dryininae); b) ma I es of Aphelopinae, Anteoninae, Conganteoninae, Bocchinae, Thaumatodryininae, Perodryinus, part of Tridryinus and Dryinus (Dryininae) ; The foramen is an approximately circular gap in the skeleton of the back a)

fema les of

of the head

through

which

thorax.

If the occipital

off from

the

the

carina

internal

organs pass from

is effaced

the vertex

will

the head

to

the

not be marked

occiput which I ies between the occipital carina and the

foramen. The area between the compound eyes and the occipital carina the gena; the upper part of the gena is known as the temple. Viewed

is

dorsally the distance from the posterior edge of a compound eye to the occipital carina is the ocular-occipital line, TL. Sometimes the occipital carina

1s absent

region

between

and

vertex,

occiput

and

gena

are

the bottom of the compound eye and

then

continuous.

The

the articulation of the

mandibles is the malar space. In many Dryinidae a well-marked subocular sulcus (Fig. 4) connects the lower corner of the eye to the mandibular articulation.

The subocular sulcus

is always present

in males.

Only

in

Gonatopodinae, Apodryininae, Conganteon (Conganteoninae) it absent. The mouthparts of Dryinidae are found to be taxonomically important. The mandibles (Fig. 5) stand apart from the labio-maxi Ilary

females of

complex. cocoon, insects.

Perhaps because of their' use for such purposes as opening killing or handling prey, Within

the

they

Dryinidae, however,

"nay be greatly reduced

have not been reduced as

IS

the

in other

the number of mandibular teeth

as follows:

:;onatopodinae: female: 4; male: 3; Jryininae: female: 4; male: 3; Anteoninae: female and male: 4; Conganteoninae: female and male: 3 + 1 smaller; 30cchinae: female and male: 1, 2, 3, 3 + 1 smaller; Transdryininae: female:

1;

male unknown;

.-'iphelopinae: female and male: 3 irregular; 3iaphelopinae: female: 3 + 1 smaller; male unknown; Thaumatodryininae: female and male: 4; .~podryininae:

female:

The max ill ae and

4

irregular;

lab i um are fi rm I y

male unknown; un i ted

to one another by

membranes

labio-maxillary c0:!,,:lex. Each maxilla consists of a ::oasal piece or cardo which articulates with a process of the hypostoma. At and form a

single

tne distal (ventral) end of the cardo a stipes its distal end, the stipes bears the maxillary six segments but frequently

have fewer

is articulated.

Externally

at

palpi which primitively have

in special ized genera or species.

(" e rn . A me r.

16

:: n t.

Inst.,

!l0 .

3 7,

1984

•

•

•

•

f

1P' F

-4 .. -

",,_

Fig.

4 -

. _.

Subocular su lcus

the mandi bu lar art i culation

(S)

connecting

(male of

the

l ower corner of the eye to

Anteon fulviventre

Haliday).

Introduction

- l c,

'of

Fig.

5 -

posterior

Mandible with (female of

four

>

41

,

teeth progressing

Anteon gaullei

17

Kieffer).

larger from anterior on e to

18

Mem.

I nternally and a

Amer.

Ent.

Inst.,

no.

37,

1984

two lobes, a more proximal one, the lacinia, galea. The labium consists of a submentum and The distal end of the prernenturn bears externally the labial

the stipes bears

more distal

one,

the

a prementum. palpi, which characteristically have three segments though the number may be reduced

in particular genera and species.

palp-segments may

be concisely

maxi Ilary palp-segments;

The data as to reduction of

summarised as follows

second

number =

labial

(first number

palp-segments):

Dryininae: female and male: 6/3; ThaUluatodryininae: female and male: 6/3; Anteoninae: female and male: 6/3; Bocchinae: female and male: 6/3; Conganteoninae: female and male: 6/3, 5/3; Transdryininae: female: 6/3; male unknown; Biaphelopinae: female: 6/3, 5/3; male unknown; Aphelopus (Aphelopinae): fema I e and ma Ie: 5/2; Crovettia (Aphelopinae): fema I e and ma Ie: 5/3; Gonatopodinae: Gynochelys: female: 6/3; male unknown; Epigonatopus: female: 2/2; male unknown; Esagonatopus: female and male: 6/2; Gonatopus: female and male: 3/2, 4/2, 4/3, 5/2, 5/3, 6/3; Tetrodontochelys: female and male: 2/2, 3/2, 4/2, 5/2; Trichogonatopus: female: 6/3; male unknown; Haplogonatopus: female and male: 2/1; Dicondylus: female and male: 2/2, 3/2, 4/2; Pentagonatopus: female: 5/2; male unknown; Plectrogonatopoides: female: 5/3; male unknown; Pseudogonatopus: female and male: 2/2, 3/2, 4/2; Donisthorpina: fema Ie and ma Ie: 5/2; Agonatopoides: female and male: 5/3; Apterodryinus: female and male: 6/3; Acrodontochelys: female and male: 2/2, 3/2, 4/2; Adryinus: female: 6/3; male unknown; Echthrodelphax: fema Ie: 3/2, 4/2, 5/2, 5/3, 6/3; rna Ie: 4/2; Neodryinus: female and male: 6/3; Apodryininae: female: 4/3; male unknown. The maxillary

and

labial

palpi

are usually borne on special

stipes and prementum respectively. like a

basal

segment of the palp.

segment of the palpi

really

there can be no doubt that watching a

narcosis)". palpi

In

The question

is a palpiger or a

in the present paper. According seen by

These lobes,

to R.C.L.

it is a

the

as to whether the basal true joint

Perkins

the present paper,

(1905)

true moveable joint

species under the microscope,

is always considered

the first small

a true joint.

lobes of the

palpigers, may look

as

is not discussed "in many forms

(as can easi Iy be

it recovers from

basal

The maxillary

segment of the and

labial

palpi

Introduction

have been

largely used in

generic differentiation. palpal

segments

distinguishing pal pal

variable

segments

in

however,

Moreover,

the

last

labium shows

it now

lobes,

in

the genus.

longer of prime appears that palpi

The

importance in

the number of

except

two segments of the maxillary three central

the number of

the species and

is therefore no

the genera.

that

segments agrees between males and females,

particular, The

Gonatopodinae for

the classification of the

I t now appears,

is usually

number of palpal

19

that

in males

in

may be fused.

the outer ones being

the

paraglossae and the inner one the glossa. The Dryinidae have fi I iform or distally

thickened antennae,

segment or is small

and

with

segments. when

10 segments

the scape forms a

part of the antenna, of various

with

in both sexes.

The first

scape is articulated in a socket. The second segment or pedicel flagellum.

the

The surface of types.

ball

and socket joint for

the antennae

is closely covered

These are sometimes taxonomically

large enough

Thaumatodryininae,

the distal

The flagellum consists of eight with

important,

to be visible without great magnification. the females have tufts of

long hairs on

sense-organs especially In

the antennal

Gonatopodinae the females have longitudinal groove-I ike structures, the rhinaria (Fig. 6), on the distal

segments 5-10. segments. in

In some genera of

The number of distal

the genus

(2,

3,

4 or 5),

segments showing

but constant

in

the rhinaria

the species.

is variable

Rhinaria are

also present in the females of Apodryininae (on antenna I segments 3-10), Transdryininae (on the I as t three antenna I segmen ts) and Dryininae (on the last six antenna I segments: except for fiiesodryinus, which seem to be always without rhinaria). distal

The data as to distribution of rhinaria on

antenna I segments may

be concisely summarised as follows

(within

brackets number of rhinaria per segment):

Gonatopodinae Agonatopoides: 9th (1), 10th (2); Haplogonatopus: 9th (1), 10th (2); Acrodontochelys: no rhinaria; Tetrodontochelys: no rhinaria; Gonatopus: no rhinaria; Dicondylus: 9th (1), 10th (2); Pseudogonatopus (A): 9th (1), 10th (2); (8):

8th

(1),

9th

(1),

10th

(C):

7th

(1),

8th

(1),

9th

(2); (1),

10th

(2);

Trichogonatopus: no rh i nari a; Apterodryinus: 7th (1), 8th (1), 9th (1), 10th (2); Donisthorpina: 9th (1), 10th (2); Esagonatopus: no rhinaria; Plectrogonatopoides: no rhinaria; Echthrodelphax: 9th (1), 10th (2); Adryinus: 6th (1), 7th (1), 8th (1), 9th (1), 10th (2); Neodryinus: 6th (1), 7th (1), 8th (1), 9th (1), 10th (2); Pentagonatopus: 7th (1), 8th (1), 9th (1), 10th (2);

""

•

20

Mem . Amer. En!.

I ns!.,

no. 37,

1984

Gynochel ys : 6th ( 1 ), 7th ( 1 ), 8th (1), 9 th ( 1 ) , 10th (2); Tr an sdryininae Transdryinus : 8th ( 1) , 9th ( 1 ), 10th (2); Dry ininae Dryinus : 5th ( 1 ), 6th (1), 7th ( 1 ), 8 t h ( 1 ), 9th (1), 10 th (2); Richardsidryinus: 5th ( 1 ), 6 th ( 1) , 7th ( 1 ), 8 t h (1) , 9th ( 1 ), 10th (2); Al phadryinus : 5th (1), 6th (1), 7th (1), 8th (1), 9th (1), 10 th (2); Tr i d r y i nus: 5 t h ( 1), 6 t h ( 1), 7 t h ( 1 ), 8 t h ( 1 ), 9 t h ( 1 ), l Ot h ( 2 ) ;

Fig. 6 -

Last antennal segme nt s showing rh in ar i a Haplogo natopus oratorius Westwood).

(fema le of

Perodryinus : 5th ( 1 ) , 6 th ( 1 ) , 7th ( 1 ) , 8 th ( 1 ) , 9th ( 1 ) , 10 t h ( 2) ; Me sod ryinus: n o rh i nar i a j Megadryinu s : 5th ( 1 ) , 6th ( 1 ) , 7th (1 ) , 8th ( 1 ) , 9th ( 1 ) , 10th ( 2) ; Apodryininae Apodryinus : 3th ( 1 ) , 4th ( 1 ) , 5th ( 1 ) , 6 t h ( 1 ) , 7 th ( 1 ), 8 t h ( 1 ) , 9th ( 1 ) , lath THORAX.

( 2) ;

T h e thor ax consists of t h ree segm ent s (pr',ihorax,

mesothorax ,

I ntroduct ion

21

meta thorax} ; the two hinder ones normally bear wings. Some females of Gonatopodinae are apterous. In Gonatopodinae, Anteoninae and Bocchinae, females may be brachypterous or micropterous.

The first

abdominal

segment

(propodeurn) has become firmly attached to the thorax and is connected to the rest of the abdomen by a narrow petiole. Some authors propose the term mesosoma for the thorax + propodeum. Usually the segments of the mesosoma are easily visible. fused

(only

propectus

PROTHORAX.

less

is distinct).

The dorsal

the mesonotum, the

recognizable because the sutures are more or

Apodryininae, however, the mesosoma sclerites are completely

In

sclerite

whi Ie the

lateral

(pronotum) is rather closely attached to sclerites (propleura, composed entirely of

proepisterna) are loosely attached to their notum. The propleura form the prothorax and are termed propectus. They meet

the ven tra I part of along

the median

I ies the small

ventral

I ine.

Quite often hidden between

the first coxae,

prosternum. Anteriorly the propectus supports the head,

sharing this duty with the pronotum; posteriorly it is attached to the first coxae. The pronotum is commonly divided by an anterior transverse furrow into an a

anterior part

rather higher level

(anterior collar) and a more posterior part (disc) at (in

the plane of the mesonotum).

Sometimes a

posterior collar. Gonatopodinae no transverse furrow is visible. I n the ma les of Dryinidae as well as the fema I es of Aphelopinae, Biaphelopinae and Conganteoninae, the p ronotum is very reduced dorsa II y posterior transverse furrow

is also visible,

definining a

In some females of apterous

and

is almost

invisible from above.

This condition

is primitive.

In

the

more specialized females of Dryinidae the pronotum is distinctly visible dorsa II y. It is produced backwards and swoll en into pronotal tubercles. They may

reach back as far as the tegulae or be widely

separated from

Gonatopodinae are the pronotal tubercles absent. In females of the most specialized forms (Dryininae, Gonatopodinae, Transdryininae) the pronotum is remarkably elongated, them.

Only

in

the females of

further forward extension

is caused

of other prothoracic elements is unusually mobile. the prothorax adaptive,

in

by

the dorsal

(propleura

According

to Reid

in particular). (1941)

and

position and development The pronotum

itself

the peculiar modifications of

Gonatopodinae and Dryininae are in a large measure

being concerned

in

the raptorial

habits of the females.

In

Apodryininae the pronotum is fused with the posterior sclerites of the mesosoma. MESOTHORAX.

contains

The second

thoracic segment

the principle flight-muscles.

Only

is usually in

the

largest,

since

it

the apterous females of

Gonatopodinae is it more slender than the other thoracic segments. Dorsa II y the segment is covered by the mesonotum which is divided into two principa I sc I eri tes, the scutum and the scutellum; taxonomi sts often reserve the term mesonotum for the scutum alone. scutum

two sulCi,

the

From the front margin of the

notaulices, often converge backwards, defining a V-

shaped prescutal area. The sulci may be deep and strong and meet at about the centre of the posterior margin of the scutum. In other dryinids they extend back

to the hind margin of the scutum without meeting;

in

22

Mem.

many

groups,

they

are

weak

Amer.

and

very

Gonatopodinae median scutal line

apterous fema I es of

Aphclopinae bisects less

scutum. in

each

and

are the

37,

1984

i ncomp I ete or even

absen t.

the notau I ices are absen t. is often

each

present,

si de of

about

sulcus may

half way

connected

aspect of

anterior,

which

forwards

sternal that

It

very

narrow

are

the

it

scutum.

areas

and

and

They

I a tera I

pleural

to

are absent

regions of

is convenient

to

treat

mesopleuron

is

the

pleural sulcus, lateral

reduced,

the pleural

narrow and wh i ch

articulation Since

sulcus runs

near and

the mesopleuron

the mesopleura.

are partly

visible.

marked off by

a

Just

the

the mesepisternum

an

the

is

tegula

cutting

function

of fl ight,

forms

long

entirely of

anterior

very

iJryininae. marked.

much

while

dorsum of

the

scutellum

to Reid

(1941

be seen

l,

Gonatopodinae,

among there

prothorax,

In

meta thorax

METATHORAX.

Apodryininae and

runs from of

the

subserv ience and

the

It

the bulky,

is formed

reduced. by

In

to

and special ised.

female

the

The

almost

lateral

the prepectus,

stages of thoraces

this

among

various modifications of

is one

this

non-adapta t i ve one.

is

It

region

prepectus. The Gonatopodinae.

from

that

the

be obsol ete or strong I y

This condition

scutellum.

area and

antero-dorsal

visible.

The early

the reduction

the

the

are formed

some phylogenetic significance; of

At

or

is much

suture may

median,

transverse

the mesonotum

in

a

trochantin

the pronotum

importance.

the

the

apterous

the mesothoracic waist

The meso-metap I eura I apterous

is a

modified

The

the prepectus Can

According in

in

from

is

to

and continuous

antero-ventral

rei eased

parallel

parts by

is

epicnemium

developed

hO's assumed considerable

special isation of

area

waist connecting

thorax.

walls of

the

the mesothorax,

slender the

the scutum,

ventral

off an

has become

and

posterior part of

this

the mid

sterno-pleural sutures

trochantinal sulcus. epicnemial sulcus is towards

well

lateral

spec ies the

that

down

particularly

Gonatopodinae

developed

of each mid coxa

is

of

into two

many

in front

it off

apterous

thorax

In

seems

mesepisternum,

which

divided

It

the region prepectus

is

sulcus.

the sulcus cutting corner of

It

sternal sulcus.

vis ib I e

runs

of

the mesepimeron

well

suture, which divides mesosternum is relatively

The

are

The

into an

very

the

the forewings.

together.

divided

posterior,

d i v i ded by beneath

in

the mesothorax

them

the nieso-I~etapleural

and

two more or

the paraps i da I furrows

metapleuron.

a

the

Gonatopodinae. Arising on parapsidal furrows which

upwards from

is

and

l"ing-process

to

with

In

the

scutellar sulcus.

pair of

across the

The scuta I

the

apterous

be a

the mesothorax or

mesepimeron.

coxa

part I y

In

longitudinally

the scu tell um

with

in

large mesepisternum and a

very

obliquely

coincident

sulcus are absent

Gonato;Jodinae. parapsides. The

intimately

reduced

no.

the

the scutal

forward

apterous

lateral

On

is centrally

scutal

side of

extend

so

areas,

axi Ilae and

Axillae

area.

Ins!.,

axillae, which belong morphologically to the The scutellar sulcus runs behind the axi Ilae and, on its central front of the scutellum. The scutal sulcus divides the scutum from

triangular

part, the

a

the prescuta I

En!.

that

may

possibly

the enlargement of may

the scuturn

perhaps

the mesothorax

the

have and

be regarded

is fused

as

with

propodeum.

The metathorax,

which bears

the relatively

small

hind wings,

Introduction

is smaller and segment area,

less differentiated

is covered

usually

Gonatopodinae between

the

a

suture from In

region

metasternum fused

Vv

i th

The

tergite.

the propodeum

to

The

the a

the dorsal

is a

small

muscle of the

Dryinidae

together on

have

each

front edge of

micropterous. is

the

of

the

In

wings When

"hich

articulate

the subfamilies.

'.'eins

and

Irig.

7)

I

text

I

term

in front

of

as

the

inclined

No suture between

Apodryininae

In

the meta thorax

is

to

the first

abdominal

spiracles

lie near the

segment

is articulated

and

above

The

the point

lateral by

margins

its petiole

i nterna I organs where

they

leave

the mesosoma.

a

row

of

wings,

the fore

hooks or

hamuli,

and engaging

Dryinidae, 3-5 wing

the

follow

hamuli

margin

thorax

the

by

is

the

a

system

the

hind margin

some females of

are brachypterous or margin

dorsal margin, the

by

a

of

the

and

series of

axillary sclerites. the tegula.

(1969)

for for

wing

the

large plate,

single characteristics Townes

In

held

arise from

downfolded

is supported by

hind ones,

which

the front

the

means of

hidden

and

are found.

Some females

The membrane

imoortant

with

is outstretched

hind

are normally

.\ ing-veins provide

Two

are absent.

with

sclerites

the

and

and

is

orifice of the propodeum, through which passes petiole. In Apodryininae the propodeum is fused

costal margin, the wing is the apex.

axillary

In

wh ich

separated

metepisternum

into

the metap I euron

the name given

two pairs of

the

is not

the meSQsoma.

ventrally,

the hind wing

the

divided

groove

the metanotum,

doub Ie ba II-and-socket joi n t.

side by

the fore-wing.

:Jryinidae

is visible. is

the other segments of

NII~GS.

not

is what

+ propodeum. The metasternum is a very

abdominal

the petiole somewhat

the propodeum

transverse

wetapleuron

The

is

A

area

apterous

is probab I y

the scutellum

thorax.

second

with

the

two posterior pairs of coxae.

propodeum

attached

to

of

the

and

In

the

single central

This central

defined.

the propodeum.

behind

the other segments of

the

with

with

area

between

well

Gonatopodinae

the meta thorax

of

enter

not

the propodeum.

and metapleura

PROPODEUM. segment

is

Dorsally

consists of a

depressions.

the propodeum

ap terous

fused

the mesothorax. which

postscutellum.

the

the propodeum

the flat

surface of

narrow

with

the

indistinguishably fore

lateral

the metanotum fused

metepimeron. metanotum

by

termed

scutell um and

is more often by

has

than

metanotum.

the

defined

(1939)

Richards

by

23

tip

veins The The

the classification labelling

of

the wing-

cells which they enclose. The fore-wing of the Dryinidae pterostigma, 1-3 basal cells enclosed by pigmented veins, 2 costal vein, the metacarpus joining the pterostigma to the apex, and in SOme species an enclosed radial cell. The radial cell is open when the -adial

the

shows a

vein

is

incornplete.

In I~adiimancus

Moczar

the radial

vein

is

Gonatopodinae, Dryininae, Anteoninae, Bocchinae, -::lauIllatodryininae, Transdryininae have three basal cells (costal, median 3"ld submedian) enclosed by pigmented veins (costa, subcosta, medius, s:1!Jmea'ius, basal, nervulus). COI1ganteoninae and iliaphelopinae have two :casal cells {costal and median} enclosed by pigmented veins (costa, .o:..;:'costa, medius, basal). A;;helopinae have basal cell (costal) enclosed by ::igmented veins (costa and subcosta). The first brachial cell rarely 38sent.

Vvinged

, '"

24

Mem.

appear s

En!.

Ins!.,

no.

37,

1984

Th e male of My strophorus formicaeformis Ruthe has

to be enclosed.

fore-wings reduced,

Amer.

spoon-shaped;

the hind-wings a re absent.

"

Fig.

7 -

Fore-wing of Gonatopodinae : C

submedian cell;

=

ra di al

ve in,"

basal vein;

LEGS.

There

F cs

=

costal

cell;

M

=

median cell;

5 -

= first brachial cell; R = radial cell; t = metacarpus; r = costa," b = subcosta", e = medius", h = submedius", k =

= nervulus.

0

is a pair of

legs for each

thoracic segment.

The

legs consist

of a coxa,

trochanter, femur, tibia and a segmented tarsus. The females of the Dryinidae (except for Aphelopinae and Biaphelopinae) have relatively

elongated fore-coxae.

segments.

The

l ast tarsal

the foot-pad or arolium

T he tarsus always consists of five

segment bears the claws.

(Fig.

8).

Its dorsal

Between the claws

surface contains a

plate,

li es the

orbicula. The f ron t tarsi of most fema l e dryinids (excep t for Aphelopinae and Biaphelop inae) are formed into a chela (Fig. 9) for gripping the homopterous prey.

The chela

is composed of the 5th segment,

and an

enlarged claw. A second claw, the rudimentary claw (Fig. 8), i s reduced (in Thaumatodryininae, Bocchinae, Dryininae, Gonatopodinae) or absent (in Anteoninae, Conganteoninae and Transdryinillae) The trochanter is usua II y a srnall segment. I n the females of Dryininae, Transdryininae, Thaumatodryininae and Gonatopodinae it is very elongated. The femur usua l ly consists of a prox i ma l swol len part {club of femur} and a distal slender part {stalk of femur}. The tibiae are provided with articulated spurs o n

the ventral

apex of the tibia

in

s ide.

Typically,

the following

the

three legs have spurs at the

numbers:

1,

1,

2

(males,

females of

25

Introduction

Aphelopinae, Biaphel opin ae, Conganteoninae, Anteoninae, Thaumatodryininae, some Bocchinae, most of Dryininae ) ; 1, 1, 1 (females of Apodryininae, some Dryininae, most of Bocchinae); 1, 0, 1 (females of Transdryininae, most of Gonatopodinae) or 1, 0, 2 (some females o f Gonatopodinae). The fore-tibia always has only one apical spur. This spur i s mod ifi ed, tarsus,

Fig.

together with

into an

8 -

Arolium

the ve ntral

s i de of the first segment of the

antenna-cleaner ( Fig. 10). The first segment of the fore-

(A)

and rudimentary

claw

(C)

of

Haplogonatopus oratorius

(Westwood) .

(bas itarsus ) may be short or very elo ngated. The modified fifth segmen t of the fore-tarsus con s ist s of a proximal part (basal part, tarsus

d irec tl y distal m ay

art icul ated

part

form a

with

the fourth

proximal

anterior

segmen t of the fore-tarsus ABDOMEN.

and a more or

less

long,

free

(ap ical part) . Segments 2 or 3 (rarely 1) of the fore-tarsus (inner)

fore-tarsus may be short or very bris tles,

segment)

and

hook.

e longa ted.

The fourth The

seg men t o f

the

inner side of the fifth

the enlarged claw may bear

lamellae,

ha i rs or peg-I ike ha i rs. The abdomen consists of

ten

segments,

but this number may

not

~iem.

26

Fig. N ;

9 5th

Arner.

En!.

I ns!.,

Chela o f Anteon g aullei Kieffer t a r s al

segment).

(A ;

no.

37,

1984

aro liu m;

E ;

enlarged claw;

Introduction

be visible without careful of a dorsal

27

study or dissection.

Each

is normally

made up

tergite and a ventral plate or sternite. T he

plate or

....

;'1' ,-

""

j" "

\ ,

j

•

, •

•

,

L F i g.

10 -

Antenna-cleaner of

Jiaplogonatopus or atorius (Westwood) (female):

T = tib i a l spur.

•

spiracular openings

lie at the sides of the

segmen t

into the propodeum wh ic h

is modi fied

tergites.

The first abdominal

is fu nctiona ll y

part of t he

gaster is used f or all the abdominal segments beh in d the propodeum. Some authors propose the term metasoma for t he post-propodeal part of the abdomen. Th e petiole is the narrowed, anterior, mor e or less long stalk. Accord i ng to R ich ards (197 7 ) t he Dryinidae have 8 pairs of thorax.

The

abdom in al

ter m

spiracles.

are normall y

T erg it e 10 is

to which

ov ipos itor, segment

which

is reduced the

only 8 abdom inal Th e ninth

t he seventh

tergites

(7 gastral)

tergite is entire ly

indistinguishably fused

side of the abdomen

The eighth sternite

val vifer,

the female,

developed and v i s i ble.

and depigmented. the vent ral

In

is the

with

retracted

tergite 9.

On

last vis ible sternite.

and membranous and on each s i de is a

first

first valvula is attached. They are parts of the

is adapted for sti nging.

T he sternite of the ninth

is also redu ced and on each s ide has a

second valvifer with the

,

28

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

second valvula attached to its anterior end. The valvifers described as appendage bases. the ovipositor,

forming

immovably joined first The

valvulae.

the

The first

can be

valvulae are the active part of

lancets of the sting.

to one another and form

a

The second valvulae are

sheath

(sting-sheat)

for

the

terebra.

The two pairs of valvulae together constitute the

third valvulae each arise fror,; the posterior end of the second

valvifer.

The genital

orifice

is anterior to the first

longer act as an ovipositor.

The sting and

plates have not so far proved as useful corresponding parts of the male. gastral)

In

are exposed and normally

are available.

The first

next six sternites The external

(up

sternite

in separating

the male developed.

No data on

is not visible as a

to sternite 7 = sixth

last abdominal

species as have the

the first eight

tergites

(7

tergites 9 and

definite sclerite.

gastral)

genital ia of the male are attached

ninth segment.

valvifers which no

the modified

are fully

10

The

developed.

to the hind margin of the

The shape of the ninth sternite may

have taxonomic

importance in Dryininae, Anteoninae and Aphelopinae in separa t i ng species. The male genital ia (Fig. 11) consist of a basal ring, two lateral a median penis (consisting of two penis valves) and two volsellae on the sides of the penis. Each volsella consists of a proximal basivolsella and a d i sta I distivolsella. I n the Gonatopodinae the dorsa I end of each paramere forms an appendage termed the dorsal process of gonoforceps (Fig. 12). I ts shape has taxonomic importance in separating species. In the males of Anteoninae the parameres may bear proximally more or less large dorsal membranous lobes. LENGTH OF THE BODY. The adu I ts of the Dryinidae range in size from 1,00 mm (Gonatopus ashmeadi Kieffer) to 10,93 mm (i'legadryinus magnificus

parameres,

Richards).

Measurements of total

the front of the head (excluding

the an tennae)

to

length from

the apex of the abdomen

the sting).

COLOUR.

Most males are black.

gasters,

but head,

black,

length of adults refer to the

(exc Iud i ng

thorax

Most females

have black

to reddish-black

and propodeum are highly variable

in colour -

reddish or yellowish.

EGG AND POSTEMBRYON I C STAGES. The morpho logy of the egg has been described by Fenton (1918a). The egg appears oval or kidney-shaped and varies .in color from sculpturing on

light yellow

the chorion.

of the egg were added by Westdal,

Richardson

has been described by (1965).

The

larva

third and fourth and serves each molt, usually the oral process

Fenton

(1918a)

is apterous. instars.

It

is

but soon darkens in

the second,

(1961),

process

Westdal,

is present

located near the anterior end the host.

light brown.

pink or green.

third and fourth

The

The first-instar

the

larva

Richardson in

the second,

the

larva

larva is white after

The mature larvae are A pair of fleshy oral

instars.

In

the fifth

lobes are replaced with biting-chewing mouthparts and has disappeared.

is no

Barrett,

The structure of

and Barrett,

in

There

notes on the morphology

Raatikainen

(1980c).

A ventral

to a

but occasionally

grey or dark brown.

fragmentary

(1940),

and Jervis

to anchor the parasitoid

white,

are present

Clausen

(1965)

to dark

Additional

lobes

instar the ventral

larva has a sacciform or

-

Introduct i on

29

•

•

, .

1

"

•

Fig. ring;

,

-

11

- Ma Ie gen i ta I i a of Deinodryinus atriventris (C resson):

N -

penis;

P ;

parameres;

S ;

basivolsella;

V ;

B ;

distivolsella.

basa I

30

Mem.

embryo n ic shape. or U-shaped.

12 -

En!.

third -

T he f i fth -i nstar larva

w i der posteriorly

F i g.

The second - ,

Amer.

I ns!.,

no.

is straigh t ,

and more pointed anteriorly.

N = penis;

1984

and fourth - ins t ar l arvae are curved d i stinctly

segmen t ed,

Other notes on

Male genitalia of Gonatopus lunatus Klug :

gonoforceps;

37,

0 -

the structure

dorsa l

process of

V = dist i vo l sella. •

of the ( 1905) ,

larva were published by Mik Jeanne l

( 19 13),

(1882),

Keilin and Thompson

G i ard

(1889),

( 1915),

Haupt

R.C.L. ( 19 16,

Perk i ns 1932),

Fenton ( 1918b), Maneval (1935), Esaki and Sameshima (1939, 1940), Clausen (1940), Pruthi and Mani ( 1942), L i ndberg (1950), Behrisch (1952), Heik in heimo (1957), Raa t ika i nen ( 1961), Guagliumi (1962), Chaudhary and Ramzan

( 1968),

Ponomarenko

(1975b)

and Jerv i s

(1980c).

Introduction

31

COLLECTION

Best collecting Townes

1972;

of flying

will

be from

Malaise traps

(classical

or

for

Dryinids.

near the ground, collecting

results

instance Townes ' and rVl asner l s models: 1\1alaise 1937; Masner and Goulet 1981). They are used mostly for sampling

modified models,

Pan

i.

traps may

e.

the

methods such

be used

normally

as sweeping or Malaise trapping

communication).

Apterous

Sweeping

recommended for collecting

is not

to sample Dryinids

spectrum of species not

Dryinids are best obtained

oy

active on or

obtained

by

(Masner personal pan

adult Dryinids,

traps. as

the results

are unsatisfactory. Collecting satisfactory Dryinids, obtained rearing

nor by

their hosts.

sweeping.

parasitized

parasitization Chandra's rearing

as

pantrapping

Dryinids

hoppers collected 1980c).

hoppers

Dryinids may

will by

be studied

Chandra

parasitized

or Malaise trapping

the behavior and

For these purposes,

Adult

abilities may

models:

Adult from

methods such

techniques for studying

in

resul ts

be obtained

sweeping. in

Good

glass

best

cages

may

tubes plugged

also be obtained by

wi II

from

be

the field

Biology,

rearing

results

are not

development of by

behavior and

(for

instance

also be obtained with

holding

by

cotton.

cocoons collected

plants. 'ilinged

Dryinids or parasitized

hosts may

be attracted

oy

light

traps. P,c<ESERVATION

Small

Dryinids are best mounted on

are pushed using

through

stainless steel

pinned

pins.

away

to obscure are relaxed genitalia

from

the body

the sides and it

with

is as a

rectangular cards.

and

to open needle.

identify

many males,

prepare genitalia relaxed capsule should be removed

examination

in

Faure

glycerol

20 cc,

and clearing

GOlla topodillae, identify

many

Arabic

and

the

the mouth

a

hooked

w hi ch have very

and

c I ear

dorsal

pin,

genitalia

be

wings

so as not

the

male

be glued it

to

has the

and

in

is essential.

To

The entire male genital

50 cc,

for

dorsa I the

the

specimens

pUllout

Dryinids Can

the

Maceration

Gonato:.JoJinae,

should

have

Vihi Ie the

which

if dropped.

water

recommended

the adul ts should

is recommended;

the specimen

gum 30 g).

the

parts

examination of

(distilled

pins

pinned

antennae positioned

specimen.

Alternatively,

using

are not

males of

easy examination of

legs

they

specimens are required.

liquid

solution

is used,

thorax,

This method of mounting

advantage of protecting To

the

the

underside of

well

hooked

minute stainless steel

Larger Dryinids are best

Whichever method

to one side of the centre of

pulled

added

Plastazote strips.

10

mounted

Chloral

for

microscope

hydrate 50 g,

% potassium

hydroxide

male genitalia of processes of gonoforceps.

volsellae should

process of gonoforceps.

In

be removed many

To for

species of

Dr/ininae, Anteoninae and Aphelopinae the n i n th stern i te shou I d a I so be

32

Mem.

removed from To

Amer.

the male genital

identify females,

En!.

removed

using

Maceration

a

pin

and

in

37,

1984

capsule for microscope examination. are required.

mounted

is essential.

The fore-tarsi

cleared

This method of mounting

the examination of which

To

should be

for microscope examination

10% potassium hydroxide solution

The chela should be dehydrated, examination.

no.

examination of the chela

prepare chelae relaxed specimens balsam.

Ins!.,

in

Canada

is not necessary.

and mounted for microscope

is recommended

is important for

also for the palpi,

identifying females and males of

Gonatopodinae. The palpi should be removed using a hooked pin or two fine needles. 1939)

Richards'

is too complicated

boi ling of the speci men amount of boiling, which

method for removing and in

the mouth-parts

is not recommended.

water or very

the mouth-parts can

weak

potash.

Dryinid

larvae should be preserved

acetic acid

(absolute alcohol

After

be pulled out with

must be made to pierce the gula area and

in Carnoy's fluid

then

is based on the proper

a

fine needle,

tear forwards.

in 90% alcohol

60 ml,

(Richards

This method

after being

chloroform 30 ml

fixed

and glacial

10 ml).

FAMILY SIZE AND DISTRIBUTION

This revision records 844 species grouped

probably comprises approximately distribution of subfami I ies and Krombein,

Hurd,

North of Mexico)

(1979:

Nearctic;

Neo.

=

tables 2a and 2b. names

Catalog of Hymenoptera

in America

Eth.

Ori.

Nea.

Neo.

46 37 5 2

24 22 1 1

22 16 3 3

79 55 1

-

-

2 3 3

-

2

1 7 7

-

-

8 8

-

-

-

6 4 2

-

-

1 1

-

-

-

-

-

-

I

Geographic distribution of Pal.

in

geographical

25 20 1 3

Pal.

2a -

world fauna

The geographical

have been follows.

Anteoninae Anteon Deinodryinus Lonchodryinus Metanteon Prioranteon Aphelopinae Aphelopus Crovettia Apodryininae Apodryinus Biaphelopinae Biaphelopus

per region:

is shown

and Canadian

Smith and Burks

ncmina dubia and nUda)

The actual

1,000 species.

genera

For abbreviations of U.S.

Tab.

(except for

into 46 genera and 10 subfamilies.

=

Palaearctic;

Neotropic;

Aus.

Eth. =

I

Dryinids and = Ethiopian;

Australian.

21

Aus.

44 42 2

14 9 5 1 1

-

-

-

5 5

-

number of known Ori.

=

Oriental;

species

Nea.

=

33

Introduction

Bocchinae Bocchoides Bocchus Chelothelius Mirodryinus Mystrophorus Radiimancus Conganteoninae Chelanteon Conganteon Fiorianteon Dryininae Alphadryinus Cretodryinus Dryinus Megadryinus Mesodryinus Perodryinus Richardsidryinus Tridryinus Gonatopodinae Acrodontochelys Adryinus Agonatopoides Apterodryinus Dicondylus Donisthorpina Echthrodelphax Epigonatopus Esagonatopus Gonatopus Gynochelys Haplogonatopus Neodryinus Pentagonatopus Plectrogona topoides Pseudogonatopus Tetrodontochelys Trichogonatopus Thaumatodryininae Thaumatodryinus Transdryininae Transdryinus Tab.

2b -

Eth.

Ori.

Nea.

Neo.

18

4 1 3

4

8

-

-

-

9 2

-

5

-

1 1 2

-

-

-

1 1 15 2 1 7

2

-

2

Pal.

Neo.

~

~

8

-

1 1

-

-

26 1

23 2

-

-

-

-

15

48 8

-

4

-

-

-

-

2

1

4

-

-

-

3

3 2 67 1 5 5 3 1

4 1 3 3 59 2 2 2 3 1

3 1 2 1 5

3

-

-

-

40

3 -

1 13 6

-

Aus.

Eth. ~

51 1

4 2 1 1 3

-

30 1 1 2

18

-

-

8 7

10 1

-

-

5 5

-

1 9

-

-

-

-

-

-

5 -

-

15

76 1

5

-

-

-

2

Aus.

-

19

Palaearctic;

Neotropic;

4

-

-

Geographic distribution of

per region: Nearctic;

Pal.

1

2 29

-

-

25 1

-

-

19 1 4 4

23

-

1

12 57 1

43 1

-

-

-

-

1 7 1

-

-

-

24

12

2 7 4 2

-

-

2 9

1 4 1

3 1 1

-

8

5 -

3 3

3 1 1 1

-

-

7 6 6

-

-

-

8 1

2 2 1 1

Dryinids and number of known species ~

Ethiopian;

Australian.

Ori.

~

Oriental;

Nea.

~

34

;"em.

Amer.

t<EY TO THE

En!.

Ins!.,

no.

37,

SUBFAi\', I LIE 5 Or=-

1984

DNYldI JA~'

F Er'IALE 5

Front

tarsus not

Front

tarsus

Fig.

13 -

Female of

2 Fore-wing

chelate

chelate

wi th

(Fig.

(Fig.

13).

.2 .3

16)

Aphelopus atratus (Dalman) from Caluso (Italy). only

costal

cell

enclosed by

pigmented

veins

(Fig.

13)

1. AphelopinaeR.C.L. Perkins -

Fore-wing vei ns

with

(F i g.

3 Mesosoma

costal

composed by

completely apparent

and median

cells

14)..................... fused

obliterated,

pronotum and

segments

except

a

for

lateral

pronotum and rnesopleura; -

Mesosoma not pronoturn,

because 4 Chela

or

scutum,

the

partly

composed

metanotum and

rudimentary

rudimentary

claw

claw

mesosoma

of

• 1 O. by

ocelli

15) . . . . . . . . . . . . . . . . .

Apodryininae n. fused

segments

visible

easily

(Fig.

subf. (Fig.

16);

recognizable,

16).

.4

9)

••...•.••••• 5

8)...

• .•..••..••. 6

(Fig.

(Fig.

sutures

suture between apparent

(Fig.

propodeum always

sutures are visible;

wi thout

Chela wi th

only

15);

pigmented

subf.

suture between propectus and

absent

. .•. • . . . • • ••• •. • ••• . • •• . • • . • • , ••• '"

enclosed by

13iaphelo,;;inae n.

(Fig. the

track

ocelli

clearly 2.

35

Introduction

f'i9.

g.

14 -

Female of BiaphelopliS Inasneri

n.

15 ~ Female of ArJoc--{r/inus n,d~neri n.

sp.

sp.

(type)

from

from Godavari

t,lalalcahuello

(I'epal).

(Chile).

36

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

Fig.

16 -

Female of

Conganteon vuic2nicum Benoit (type) from Ruanda.

Fig.

17 -

Fema I e of

Transdryinus notogeicus n. sp.

(type)

from

Queens I and.

Introduction

Fig.

18 -

Female of

37

j'letanteon aerias (Walker) from EI Coigo (Chile).

5 Median leg without tibial spur (tibial spurs 1,O,1)(Fig. 17); occipi tal carina absent • . . . . . • • . . • . • . 8. Tra.rJsdryininae n. subf. -

Median

leg with

tibial

spur

(tibi",1

spurs

1,1,2)(Fig.

18);

occipital

carina complete • . • . . • . . • • • . . . . . . • . . . . 4. Anteoninae ."I.C.L. Perkins 6 Antennae with tufts of long hairs on segments 5-10 (Fig. 19) .•.• ..• • . . .• . . . .••. . • .. . . . . . '" -

Antennae wi thout

7 Median

tufts

leg without

of

•....•.... long hai rs

tibial

spur

Thaumatodryininae n.

6.

(Fig.

(tibial

spurs

-

Median

leg with

tibial

four

posterior

(Fig.

teeth,

at

most

tarsus

(Fig.

5);

(tibial

spurs

progressing

enlarged

with

21)

spur

teeth,

claw

one preapical

1,0,1

1,1,1

larger

(Fig.

21)

tooth;

or

1,0,2)(Fig.

20)

00natopodinae K i ef fer

. . • • • . . . . . . . • • • • . • . • • . . . . • . • • . . . . • • . • . 9. 8 Mandible with

subf.

16) . • • • . . . . • • . . . . . • . . . . . 7

or

1,1,2)(Fig.

from anterior never with a

fifth

segment

16).8

one

to

row of

of

front

always wi th more

than one preapical lamella . . . . . . . . • . • • • . . . • • • • • • . . . . . . • . . . . . • . . . . . . • . . . . 7. Dryininae K i ef f er

-

Mandible with

1-4

large

teetn and a

(Fig.

22);

teeth

(Fi;j.

23)

front

tarsus

(Fig.

wi th veins

quadridentate mandible with

tooth between

winged; costal, (Fig.

23)

has

(Fig.

rarely

to posterior,

row of

fore-wing

pigmented veins

Fully

one

always has a

9 Fully winged; -

22);

the

two posterior

occasionally quadridentate mandible with

larger from anterior

by

(Fig.

rudimentary

strong

teeth

one preapical

with

costal

submedian

in

in

and

24) . . . . . . . . . . . . . . . . . . . . . . . 5.

the

lamella

progressing

enlarged

fifth

segment

cells

winged

cl early

claw of

only . . . . . . . . . . . . . . 9 clearly

Conganteoninae n.

fully

cell s

teeth

this case

and median

16) . . . . . . . . . . 3.

micropterous;

median and

but

three teeth

forms

enclosed

subf. fore-wing

enclosed by oigrnented

lJoccninae ,
38

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

Fig.

19 -

Female of

Thaumatodryinus flavus n. sp. from New Guinea.

Fig.

20 -

Female of

Trichogonatopus neotropicus n. sp. from Brazil.

Introduction

Fig.

Fig.

22 -

21

-

Typical

Varieties of mandibles

39

chela of

in

jjryininae.

Bocchinae and Conganteoninae.

MALES

Fully winged; veins -

(Fig.

fore-wing with

25);

enclosed, 2 Fully by

but

winged;

pigmented

Usually

(Figs

in

this

costal

cell

enclosed

by

pigmented

complete . . . . . . . . . . . . . . . . . . . . . . . . .

costal,

pi grnen t ed ve ins

(F i g.

four

occasionally

case without with

(Fig.

winged,

with

3 Mandible with

26,27);

fore-wing veins

fully

fore-wing

-

only

carina

........... .. . . . . . . . . . . . . . . • 1. ilphelopinae R.C.L. Perkins Usually fully winged, rarely micropterous; fully winged forms wi th fore-wing with costal and 1-2 basal cells clearly enclosed by pigmented veins

-

occipital

costal

26) . . . . . . . . . 3.

only

costal

cel

carina.........

and median

cells

clearly

..2 enclosed

Conganteoninae n. subf.

rarely micropterous; median and

with

occipital

submedian

fully

winged

cells

forms with

clearly

enclosed by

27) . . • . . . . . . . . . • . • • • . . . • • • • • • . . • . • . . • . . • • • • • • 3

teeth progressing

larger

from anterior one

to

posterior (Fig. 5) • . . . . • . • • • . . . . • . • . • • • . • . . • . • . • . . • . . . . . . . . . • • • • • . • 4 Mandible with 1-4 teeth (Fig. 22); quadridentate mandibles with three large

teeth and a

teeth

(Fig.

rudimentary

tooth between

the

two posterior

22) • • • . . . . . . . . . . . • . . • . . . . . . . . . . . . • . . . . • • . • • . • • . • . . . . . • • 5

40

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

/~

/

~ ~~~

/// /

Fig.

Fig.

24 -

23 -

Typical

chela of Bocchinae.

Female of Bacchus uwber n.

sp.

(type).

Introduction

Fig.

25 -

Fig.

Male of

26 -

Crovettia nearctica n. sp. from Portal (Arizona).

Male of

Fiorianteon junonium n. sp. (type).

41

42

Mem.

Fig.

Fig.

27 -

28 -

Male of

iviale of

Amer.

Ent.

Inst.,

no.

37,

1984

Esagonatopus niger (Fenton) from Lexington (Kentucky).

Thaumatodryinus flavus n. sp. from f
4 Fore-wi ng wi th metacarpus as long as or longer than the pt erost i gma (F i g. 28) . . . . . . . . . . . . • . . . . . • . . . . . . • . . . 6. Thaumatodryininae n. subf. -

Fore-wing

with

metacarpus

shorter

than

.................•...•.•............. 5 Mandible with not

1-4

progressing

teeth

larger

(Fig.

22);

the pterostigrna

4.

tridentate mandibles with

from anterior

one

. . • . . . . . • • . . . . • . . . . • . • • . • . . • . . • . • . . • . • 5. -

Mandible with p

0

s t e rio r

three

( rig.

teeth

(Fig.

progressing

29) . . .

Anteoninae R.C.L. Perkins to posterior

(Fig.

teetn 220) •.

ilocchinae R i cha rd s

larger

from anterior

one

to

2 2A ) • . • . . . . . . . • . . • . • . . • • . . . . . . . . . . . • . . • • • . • • • • . . . . • . 6

Introduction

6 Occipital -

carina

complete or

43

incomplete;

dorsal

7.

Occipital

complete,

dorsal

carina absent;

process

of

occasionally

gonoforceps present

(Fig.

•..•.•••. . . • • . • • . • • • • • • • • . . . . . . • . • • • •• 9. The males of Transdryininae,

process

of

Dryininae Kieffer

gonoforceps absent • • • • • • • • • • . . . . . • • • •

but

in

this

case

12) . . . . . . . . . . . . . . . . . . . . .

Gonatopodinae Kieffer

Apodryininae and Biaphelopinae are

unknown.

Fig.

29 -

Male of Anteon

tripartitum Kieffer from Italy.

1. SUBFAr/,ILY APHELOPIN!l8 R.C.L.

PeRKINS

Dryininae partim Kieffer in Kieffer et ~'larshall 1905: 215. Anteoninae partim Kieffer in Kieffel' et Marshall 1906: 510. Aphelopinae R.C.L. Perkins 1912: 7. Aphelopini Kieffer 1914b: 214. Aphelopinae Richards 1939: 278. Aphelopinae Muesebeck and Walkley 1951: 1042. Aphelopinae J. F. Perk i ns 1976: 11. Aphelopinae Krombe in 1979: 1250.

TYPE GC:NUS: Aphelopus Da I man 1823. FEMALE: fully winged; fore-wing with only costal veins;

front

tarsus not chelate;

occipital

cell

enclosed by

carina complete;

pigmented

maxi Ilary palpi

,~',em,

44

5-segmented;

labial

MALE:

fully

veins;

occipital

witn

DISH
with

Ent.

tibial

I nst.,

2-3 segments;

fore-wing

no,

spurs

maxillary

37,

tiClial

with only costal

carina complete;

2-3 segments;

HOSTS:

palpi

winged;

Amer,

cell

palpi

1984

spurs

1,1,2.

enclosecJ

oy

5-segmented;

pigmented

labial

palpi

1,1,2.

worldwide

i',eworacidae, Cicadellidae T/phloc/Jinae.

Gc"E"A: 2 SI"ECIC:S: 42

KEY TO TH:=' GEf\lEi=<:A OF APHELOPLVA2

FEt.~ALE5

Labial large

palpi teeth

radial

vein

with and

2

segments;

1-2

regularly

-

clypeus

I ess broad,

antennal

sci eri tes

curved

fully

wi th (Fig.

MA~~S

mandibles quadridentate,

rudimentary

and C1etanotuf1l usually

AND

teeth

(Fig.

(Fig.

30);

granulatecJ,

epi stomal

22Cl);

head,

not

with

fore-wing

2-3 with

scutum,

scutellum

ret iculate

rugose;

suture cJel1

separated

31A) . • • • . . . . . . • • • • . . . .

1.

from

the

ilphelopus Dalman

Labial

palpi with 3 segments; mandibles tricJentate, witn teeth progressing larger from anterior one to !-,osterior (Fig. 22A); fore-wing with radial vein less regularly curved (Fig. 43); usually distal and proximal part of radial vein forming an angle; at least the

anterior

broader, sclerites

with

surface

of

epistalnal

(Fig.

the

scutum

suture

reticulate

running

close

rugose;

to

313).,.,.,., . . . . . . . . . . . . . . . . . . .

clypeus

the antennal 2.

Crovettia n. gen.

1. Aphelopus Ualman

Aphelopus Oa I man 1823: 8.

Ceraf)hron parti", Katzeburg 1848: 141 (syn. proposed by r
Aphelopinae

45

AtJhelojJus Dalman; r(rombein 1979; 1250.

TYPt:: SPt::C I t::S; FEr!,ALt::

(Fig.

segments; teeth; and

Drylnus a tra tus Da I man 1823, des i 9. by Viestwood 1840. 30);

maxillary palpi

mandibles quadridentate,

fore-wing

metanotuDi

with radial usually

fully

vein

with 5 segments; with 2-3

regularly

granulated,

labial

large teeth cCJrved;

not

head,

reticulate

palpi

and

with 2

1-2 rudimentary

scutum,

rugose;

scutellum

epistornal

suture well separated from the antennal sclerites; antennae distally thickened. w:ALE; structurally resembles the female; antennae not distally thickened. DISTkIBUTIO:,; HOSTS;

worldwide

Clcadelllche Typ/llocyblnae

SPC:C I t::S;

Fig.

35

30 -

Female of Aphelo,)us atratus

(Dalman)

from

Caluso

(Italy).

46

Mem.

NOTES:

Amer.

Ent.

Inst.,

no.

in some species of Aphelopus a medial

prescutal

area,

is often present.

37,

1984

scutal

I ine,

which

bisects the

This characteristic has no specific value.

PALAEARCTIC SPECIES OF APEfELOPU5

Seven species are known.

KEY TO THE SPECIES

FEMALES AND MALES

Head with mandibles, surface white -

Head with

at

2 Notaulices

or

clypeus and a more or

most

mandibles and

reaching at

basivolsella with

one

most

Notaul ices

length

two

0,?5

strong bristles

only mandibles white or

uniformly dark; with -

1-2

frontal

males with

scutum;

inner part

of.the.

32A) . . . . . . . . . . . . . . . . . . . . .

of

scutum;

(Fig.

inner part

of

the

39A) . . . . . . . . . . . . . . . . . . ..

querceus n.

7.

testaceous;

inner part

testaceous . . . . . . . . 3

melaleucus (Da I man)

1.

length

· .. , . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. 3 Head wi th

of

(Fig.

•••••••••.•

reaching beyond

basivolsella with

clypeus white or

0,6

strong bristle

•• ••• ••••• .•••••• • ••• • . '" -

less conspicuous

testaceous • • • • • • . . • • • • • . • • . • • • • . • . . . • • . . . . • • • • • . . • 2

of

sp. clypeus black

or

the basivolsella without

or

strong bristles . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4

Head wi th

clypeus and mandibles white

somet i mes part I y

dark

the basivolsella without 4 Notaulices

reaching

or

testaceous;

(never un i f orml y);

at

or with

most

0,5

ma I es wi th

clypeus inner part

of

1 strong bristle . . . . . . . . . . . . . . . . . . ? length

of

scutum . . . . . . . . . . . . . . . . . . . 5

- Notaulices reaching beyond 0,65 length of scutum . . . . . . . . . . . . . . . . . . . 6 5 Males with penis distally tridentate! (Fig. 33A) •..••••••••••••••• -

• ••••••••.•••••••.•.•••••••••.•••••.. 2. atratus (Da I man) Males with penis distally trumpet-shaped! (Fig. 38A) . . . . . . . . . . . . ..

· . . . . . . . . . . . . . . . • • . . . . . . . . . . . . . . . . . .. 6 Notaulices

reaching beyond

the posterior margin long

0,75

of

the

scutum;

straight

rod

(Fig.

form of

a

narrow,

distally pointed

(Fig.

of

scutum;

sp. sometimes

reaching

males with distivolsella

34A);

basivolsella

in

the

long and

34A) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

• • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • ••

! The females of A. distinguish.

nepalensis n.

6.

length

3.

serratus R i cha rd s

atratus and A. nepaiensis are difficult to

Aphelopinae

-

Notaulices

reaching about

distivolsella differently (Fig.

reaching beyond

the posterior margin

of

straight

width

equal

to,

(Fig.

35A);

or greater

distivolsella

basivolsella -

length

formed

in

0,75

the

of

the

form of

long and narrow,

medially narrowest Fig.

curved point

(Fig.

35C);

distinctly

of

scutum;

epistomal

height a

long

width less

of

than

space at of

sometimes

reaching

its narrowest

clypeus

straight

(Fig. rod

(Fig.

(Fig.

males

34A);

34A) ••••••••••.

serratus Richards scutum;

epistomal

the malar space at medial 4.

point

35A);

height

of

suture

its

clypeus

males with distivolsella and basivolsella differently 36A) . . . . . . . . . . . . . . . . . . . . . . . . . . . .

broad

suture medially

distally pointed 3. of

males with

basivolsella very

nigriceps Kieffer

5.

the malar

than medial

scutum;

37);

length

scutum;

........... ........ .................. Notaulices reaching about 0,65 length

35C);

of

(Fig.

37) . . • . . . • • • . . . • . . . . . . . . • • . • • . .

7 Notaulices

with

0,65

47

(Fig.

formed

camus Richards

1. Aphelopus melaleucus (Dalman)

Gonatopus melaleucus Dalman 1818: 82. Dryinus (Aphelopus) melaleucus (Dalman): Dalman 1823: 14. Aphelopus melaleucus (Dalman): Hal iday 1833: 273. Ceraphron albipes Ratzeburg 1848: 141 (n. syn.); locus typicus Rugen

Island

(Germany).

Aphelopus melaleucus (Dalman): Giard 1889b: 708. Aphelopus albipes Kieffer in Kieffer et Marshall 1905: 217 (n. syn.); locus typicus forest near B i tche and Lemberg (Germany). Aphelopus melaleucus (Dalman): Kieffer in Kieffer et Marshall 1905: 218.

Aphelopus trisulcatus Kieffer 1914b: 216 (n. syn.) ; nomen novum for A. albipes Kieffer. Aphelopus melaleucus (Dalman) : Kieffer 1914b: 217. Aphelopus albipes (Ratzeburg) : Kieffer 1914b: 217. Aphelopus melaleucus (Dalman) : Berland 1928 : 169. Aphelopus melaleucus (Dalman) : Richards 1939: 280. Aphelopus melaleucus (Dalman) : Cavro 1950: 6. Aphelopus melaleucus (Dalman) : J.F. Perkins 1976: 11. Aphelopus melaleucus (Dalman) : Jerv i s 1977: 301. Aphelopus melaleucus (Dalman) : Ponomarenko 1978: 18. Aphelopus melaleucus (Dalman) : Jervis 1979b: 191.

FEMALE: fully winged; length 1,50-2,00 mm; black; mandibles, clypeus, malar space, lower face and a U-shaped mark (embracing bases of antennae with arms brown,

with

lying along

segments 8-10

I ight,

inner margins of eyes) or fully

testaceous;

white;

legs

antennae

testaceous-

48

Mem.

whitish,

Amer.

with hind coxae,

Ent.

Inst.,

no. 37,

1984

hind clubs of femora and hind tibiae partly

brown;

sometimes also mid coxae and mid and fore clubs of femora partly

brown;

antennae distally

proportions:

4:

3;5:

4,5:

thickened; 5:

5:

5:

antennal

4:

4,5:

4:

segments in following 7;

head dull,

smooth,

fully

31 - Head of females of Aphelopus melaleucus (Dalman)(A), Crovettia theliae (Gahan)( B), Biaphelopus masneri n. sp. (C) and Conganteon vulcanicum Benoit (D).

Fig.

granulate; 4;

=

OL

dull,

frontal

2;

line present or absent;

OOL = 2,6; OPL

smooth,

granulate;

=

2;

TL

=

2;

occipital scutum,

notaulices incomplete,

carina complete;

=

POL

scutellum and metanotum

reaching about 0,5 length

of scutum; propodeum reticulate rugose, with 2 longitudinal keels on the posterior surface; median area shiny, less rugose than the part outside the area,

almost smooth,

fore-wings hyaline,

with few

irregular weak keels,

without dark transversal

bands;

without areolae;

tibial

spurs 1,1,2.

MALE: fully winged; length 1,75-2,37 mm; black; mandibles, clypeus and some small spots between the antennal sockets and on the malar space yellowish or whitish; 9-10 light; distally 4,5:

5:

thickened; 5,5:

present; = 2.

antennae brown or black,

legs brown, 6:

sometimes with segments

and fore tibiae testaceous;

antenna I segments in following proportions:

7:

occipital

scutum,

with tarsi

7:

7:

10;

head dull,

carina complete;

smooth,

POL = 7;

scutellum and metanotum dull,

granulate;

OL = 3; smooth,

antennae not 3,5:

frontal

OOL = 4; granulate;

4: line

OPL = 2;

TL

notaulices

incomplete, reaching about 0,5 length of scutum; sometimes median scutal line present; propodeum reticulate rugose with 2 longitudinal keels on the posterior surface; the area, hyaline, (Fig.

median area shiny,

almost smooth,

with few

without dark transversal

32B)

less rugose than the parts outside

weak

irregular keels;

bands;

posteriorly strongly emarginated;

with 1 strong bristle (Fig.

32A);

32A); tibial spurs 1,1,2. LOCUS TYPICUS: Vastergotland

fore-wings

ninth sternite of the abdomen inner part of the basivolsella

apex of the penis not

tridentate (Fig.

(Sweden)

of G. melaleucus in STj types of C. albipes Ratzeburg and A. albipes Kieffer (= A. trisulcatus) probably lost. HOSTS: in England Edwardsiana crataegi (Edw.){Richards 1939; J.F. TYPICAL MATERIAL:

holotype F!

Aphelopinae

49

Perkins 1976); according to Jervis (1980c): Fagocyba carri and cruenta, Ossiannilssonola callosa, Edwardsiana avellanae, flavescens, geometrica, hippocastani, plabeja and rosae, Ribautiana ulmi, Typhlocyba quercus, Alnetoidia alneti; in France Edwardsiana hippocastani (Edw.) (Giard

B

Fig. 32 - Male of Aphelopus melaleucus (Dalman) from Vastergotland (Sweden): A: genitalia (viewed ventrally); B: ninth abdominal sternite.

1889b), Typhlocyba douglasi Edw. (Giard 1889b), Ribautiana ulmi (l.) (G i ard 1889b), Edwardsiana rosae; in Germany Fagocyba cruenta var.

douglasi (Edw.); in U.S.S.R. Typhlocyba douglasi Edw., Edwardsiana hippocastani (Edw.), Ribautiana ulmi (l.) (Ponomarenko 1978). BIOLOGY:

informations in Jervis

(1979b,

1980b,

species; overwintering stage: prepupa. DISTRIBUTION: Palaearctic region: U.S.S.R.

c);

in England bivoltine

(Ponomarcnko 1978); FINLAND:

Aminne (Ab, Pojo), HE! Dyrehaven (Sjall.), HE! SWEDEN: Vastergotland, ST! Skane, TW! Messaure, TW! Ol!; HUNGARY: Ugod, BU! Nagyvisnyo (BUkk), BU! Szigetsztmiklos, BU! SzSztMiklos, BU! Nagymezo, BU! Visegrad, BU! Oserdo (BUkk), BU! Szar-Fanniv., BU! Mecsek hgys., BU! Nagykovacsi,

50

BU!

Mem.

Fenyofo,

BU!

Amer.

Matra,

BU!

Ent.

Csakvar,

Inst.,

BU!

no.

37,

Budapest,

1984

BU!

Vertesh.

(Fanier

vgy.), BU! ROMANIA: Ba Victoria (Nagy 1967b; 1972); CZECHOSLOVAKIA (Ogloblin 1924b); YUGOSLAVIA: Kastel Novi (Dalmacija), BU! Prespa Geul (Otesevo,

Makedonija),

BM!

GERMANY:

Bot.

Garten

(Berl in),

HU!

Nymphenburg (MUnchen), EB! Hamburg, HE! Rhondorfer Tai (Siebengebirge), OT! Lohrberg-Fuss (Siebengebirge), OT! RUgen Island (Ratzeburg 1848); AUSTRIA: Huben (0 Tirol), HU! Wien, BU! Laaerb, P! DENMARK: Almindingen (Bornholm), CO! Espe, CO! Korsor, CO! Vicmou, CO! Bcrumaa, CO! Basnaes, CO! Grevind Scov, CO! Langeland I., CO! HOLLAND: Putten (Gld.), LE! Wijster (Dr.), BM! BELGIUM (Crevecoeur et Marechal 1933); SWITZERLAND: Schuls (Vulpera), GV! Epalinges (Vaud), LO! Martigny (Valais), GV! Engadine Nat. Park (Grigioni), BM! FRANCE: Montpellier (Herault), P! Foret de Clenay (Cote-d'Or), P! Maisons-Laffitte, P! Wimereux (Pas-de-Calais) (Berland 1928); Le Puy, P! Foret de Fontainebleau (Seine-et-Marne), P! Lille (Nord), P! Huelgoat (Fl.), BM! Meudon (Seine)(Berland 1928); Foret de Dabo (Moselle), VI! ENGLAND: Minsmere Reserve (Suffolk), BM! Spratton (Northants), BM! Oxford, BM! TW! Wight Island, GE! Dawlish Warren (Devon), OT! Leicester, TW! Esher Common (Surrey), BM! Bix (Henley, Oxfordshire), BM! Hounslow (Mddx.), BM! Burnham Beeches (Bucks.), BM! Dorking (Surrey), BM! OL! Hanworth (Mddx.), BM! Bald Hill (Lewknor, Oxfordshire), BM! Bedford Park (London), BM! Crave I Pits (Theale, Berks.) BM! Horsley (Surrey), BM! Radcliffe-on-Trent (Notts.), BM! Beaconsfield (Bucks.), BM! Epping Forest, B! numerous other localities in Richards 1939; SCOTLAND: Beinn Eighe Forest (R.W.), BM! Tentsmuir Nat. Res. (Fife), BM! Aviemore (EI.), OL! other localities in Richards 1939; IRELAND: Donaghadee (Co. Down), BM! Agher (Co. Meath), BM! other localities in Richards 1939; SPAIN: Valladolid (Ceballos 1959); ITALY: Mt. Bracco (Barge, Cuneo), OL! Oropa (Vercelli), GE! Pizzighettone (Cremona), TW! Fori) (Zangheri P., 1969); CYPRUS: Kilani-Krios R., BM! Pera Pedi-Krios R., BM! LEBANON: Becharre, BM!

Nearctic region

(error):

CANADA:

Port Hope (Ottawa)(Kieffer 1914b).

2. Aphelopus atratus (Dalman)

Dryinus (Aphelopus) atratus Dalman 1823: 15. Aphelopus atratus (Dalman): Hal iday 1833: 273. Aphelopus melalecus (Dalman) var. atratus (Dalman): Kieffer in Kieffer et Marshall

1905: 219.

Aphelopus atratus (Dalman): Kieffer 1914b: 218. Aphelopus holomelas Richards 1939: 289 (n. syn.); locus typicus Lustleigh

(England).

Aphelopus holomelas Richards: J. F. Perkins 1976: 11, 12. Aphelopus holomelas Richards: Jervis 1977: 301.

Aphelopinae

FEMALE:

fully

antennae fully legs fully

winged; black,

black;

segment

mandibles testaceous;

1 or segments 1-2 testaceous;

sometimes with hind coxae and hind clubs of femora

partly brown or only thickened;

length 1,5-2,0 mm; sometimes with

testaceous,

51

with clubs of hind femora brown;

antenna I segments

in following proportions:

antennae distally 5:

4,5:

4,5:

6:

6:

6: 5: 5: 5: 8; head dull, fully granulate; frontal line present; occipital carina complete; POL = 7; OL = 3; OOL = 4; OPL = 4; TL = 3; scutum, scutellum and metanotum dull, about 0,5 lengt of scutum; reticulate rugose, median area rugose, MALE:

fully

proportions:

weak

bands;

notaulices scutal

keels on

length

5:

6:

propodeum

not reticulate without dark

spurs 1,1,2. 1,50-2,25 mm;

black or brown; 4:

reaching

the posterior surface;

fore-wings hyaline, black;

legs brown,

with

antennae not distally thickened; 4:

incomplete, line present;

the parts outside the area,

irregular keels;

tibial

winged;

antennae fully testaceous;

with 2 longitudinal

less rugose than

with few

transversal

granulate;

sometimes median

6,5:

7:

7:

8:

8:

mandibles testaceous; tarsi

and fore tibiae

antenna I segments in following

12;

head dull,

granulate;

frontal

line present or absent; occipital carina complete; POL = 8; OL = 5; OOL 5; OPL = 3; TL = 3; scutum, scutellum and metanotum dull, granulate; notaul ices incomplete,

reaching

about 0,5 length of scutum;

sometimes

B

Fig.

33 -

Male of

Aphelopus atratus abdominal

median scutal longitudinal

line present;

(Dalman):

sternite

genitalia

(A)

and ninth

(B).

propodeum reticulate rugose with 2

keels on the posterior surface;

median area smooth,

less

rugose than the parts outs i de the area, wi th few weak i rregu I ar keel s; fore-wings hyaline, without dark transversal bands; ninth sternite of the abdomen not posteriorly emarginated

(Fig.

33B);

apex of the penis

52

Mem.

tridentate (Fig.

(Fig.

33A);

33A);

tibial

lOCUS TYPICUS:

Amer.

Ent.

Inst.,

no.

37,

1984

inner part of the basivolsella with 2 strong bristles

spurs 1,1,2.

Vastergotland

TYPICAL MATERIAL:

(Sweden)

lectotype M!

and 1 paralectotype M!

(hereby

of D. atratus in ST; holotype M, allotype F and 66 paratypes (29 FF, 37 MM) of A.holomelas in BM; 6 para types (4 MM, 2 FF)! of A. holomelas in OX; 1 para type F! of A. holomelas in DU. HOSTS: in England Typhlocyba opaca (Richards 1939; J.F. Perkins 1976); accord i ng to Jerv i s (1980c) Alebra albostriella and wahlbergi, Fagocyba cruenta, Edwardsiana hippocastani, Ribautiana tenerrima and ulmi, Typhlocyba bifasciata and quercus, Eupteryx aurata, cyclops, melissae and urticae, Zygina sp.; in I ta I y Eupteryx stachydearum, Typhlocyba rosae and Typhlocyba froggatti; in France Typhlocyba rosae. designated:

BIOLOGY:

to melaleucus)

the para lectotype actually belonging

informations in Jervis

overwintering

stage:

DISTRIBUTION:

(1980b,

c):

in England bivoltine species;

prepupa.

Palaearctic region:

FINLAND

(Hellen

1953);

SWEDEN:

Degaberga (Sk.), BM! Skane, TW! Ol! Vastergotland, ST! HUNGARY: Szod, BU! ReyfUlop, BU! Vacz (Tudosdomb), BU! Kelebia, BU! SzSzT Miklos, BU! Gezahaza

(Bakony),

ROMANIA

(Nagy

BU!

1972);

Fej ervarcsurgo,

YUGOSLAVIA:

Bosna),

BU!

Postojne (Slovenija),

a. Rh.,

OT!

AUSTRIA:

(Wien),

P!

BM!

Semmeringgebeit

Eisenzicken,

VI!

DENMARK:

LEI SWITZERLAND: Auvernier, lO! (Vaud), BM! FRANCE: Mt. Mezenc, BM!

Morlaix

(Fl.),

Maisons-laffitte, Rhinefield (Bx.),

BM!

P!

(Hants),

BM!

lustleigh

(Dartmoor),

OT!

leicester,

TW!

Korsor,

(Suffolk),

BM!

BM!

l ill afUred, Krupa

Heiligensee,

(Reichenau

distr.),

CO!

BU!

(Pazaric,

HU!

BM!

HOllAND:

BM!

ENGLAND: Spratton

Kew

Silwood Park

(D.T.),

GERMANY:

BU! BU!

Ingelheim

Prater

Putten

(Gld.),

lutry, lO! Nyon, GV! les Pleiades P! Arles-sur-Tech (Pyrenees Orientales),

Thames Ditton

Ol!

BM! 8M!

P!

Minsmere Reserve

Studland Heath

Budapest, (Bosna),

Mouleydier (Oordogne),

le Puy, BM!

BU!

Trebevic

(Surrey), BM!

Huelgoat

8M!

Oxford,

Esher Common

(Sy.),

BM!

(Berks.),

BM!

Watbarrow Point

Wimbledon Castle,

BM;

Com.

BM!

BM! Brickhill

(Surrey),

Bedford Park

(Wray

(Fl.),

(Northants),

BM!

(london),

(london),

BM!

lancashire),

Dartmoor, BM! Monks Wood (Cambs.), BM! SCOTLAND: Aviemore (EI.), Tentsmuir Nat. Res. (Fife), BM! Rannoch Forest (Perths.), BM!

IRELAND: Drumchory (Co. Wd.), BM! Ol! ITALY: 5. Benedetto Belbo (Cuneo), Pl! SZ! Peveragno (Cuneo), SZ! Caluso (Torino), EN! Pizzighettone (Cremona),

TW!

CYPRUS:

Pera Pedi-Moniatis R.,

3. Aphelopus serratus Richards

Aphelopus serratus Richards 1939: 284. Aphelopus serratus Richards: J.F. Perkins 1976: 11, 12. Aphelopus serratus Richards: Jervis 1977: 302.

BM!

Aphelopinae

FEMALE:

fully

winged;

testaceous or whitish; legs testaceous,

length

=

OOL

frontal

3,5;

OPL

antennae distally thickened; 6: 4: 6:

6:

6:

6:

I ine present;

occipital

height of clypeus;

=

epistomal

3;

=

TL

scutum,

notaulices reaching

3;

5:

6:

reticulate rugose,

beyond 0,75

fully

tibial

winged;

black;

legs brown,

Fig.

34 -

distally

thickened; 6:

6:

6:

6:

9;

median scutal

epistomal

fully

7;

OL

or greater than medial granulate;

line present;

keels on

=

width

sometimes reaching

the

propodeum

the posterior surface;

fore-wings hyaline,

1,75-2,12 mm;

black;

without dark

mandibles and clypeus

and fore tibiae testaceous;

and ninth abdominal

granulate;

England:

carina complete;

POL

space at

its narrowest point equal

clypeus;

scutum,

to,

genitalia

sternite (8).

frontal

suture medially straight

antennae

antennae not

antenna I segments in following proportions: head dull,

incomplete; occipital TL = 3;

to,

Male of Aphelopus serratus Richards from

6:

=

POL

sometimes clypeus uniformly dark or black;

with tarsi

(A)

head dull,

spurs 1,1,2.

length

testaceous or whitish;

8;

scnure medially straight;

length of scutum;

with 2 longitudinal

median area reticulate rugose; bands;

5:

sometimes

antenna I segments

scutellum and metanotum dull,

posterior margin of the scutum;

MALE:

antennae black;

carina complete;

of the malar space at its narrowest point equal

transversal

mandibles and clypeus

with hind coxae and hind femora partly brown;

in following proportions: 4;

black;

sometimes uniformly black or dark;

mid femora partly brown; granulate;

1,5-2,5 mm;

53

=

6;

(Fig.

OL

=

4;

35A);

4:

5:

6:

OOL

=

4;

OPL

=

3;

width of the malar

or greater than medial

scutellum and metanotum dull,

5:

line complete or

granulate;

height of

notaulices

54

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

complete and posteriorly separated or incomplete and reaching length of scutum;

median scutal

with 2 longitudinal

Fig.

35 -

line present;

keels on the posterior surface;

median area

Heads of males of Aphelopus serratus Richards and camus Richards

(Dalman)(B)

beyond 0,75

propodeum reticulate rugose, shining,

atratus

(A),

(C).

smooth, little rugose; fore-wings hyaline, without dark transversal bands; ninth sternite of the abdomen posteriorly strongly emarginated (Fig. 34B); distivolsella in the form of a long straight rod (Fig. 34A); basivolsella (Fig. 34A) long and narrow and distally pointed; basivolsella with two bristles distally; tibial spurs 1,1,2. LOCUS TYPICUS: Norwood (England) TYPICAL MATERIAL:

holotype M,

in BM;

(2 FF,

5 para types

allotype F and 8 paratypes (5 FF,

3 MM)!

in DU;

6 para types MM!

3 MM)

in OX.

according to Jervis (1980c), Alebra albostriella Kybos smaragdula (Fallen), Fagocyba cruenta (H.-S.), Edwardsiana geometrica (Schrank), Lindbergina aurovittata (Doug I as), Ribautiana tenerrima (H .-5.), Eupterycyba jucunda (H .-5.), Typhlocyba quercus (F.), Alnetoidia alneti (Dah I born), Zygina sp. HOSTS:

in England,

(Fallen),

BIOLOGY: informations in Jervis (1980b, c); in Egland univoltine species; overw i nteri ng stage: prepupa and first instar larva. DISTRIBUTION: HE!

HUNGARY:

Bellinchen

Palaearctic region: TiszasUly,

(Oder),

HA!

FINLAND:

BU! Szabohegy

Lohrbery-Fuss.

Aminne

(Koszeg),

(Ab.,

BU!

Pojo),

GERMANY:

(Siebengebirge),

OT!

HE!

Lojo,

Kiel,

DENMARK,

HU! CO!

HOLLAND: Putten (GLD.), LE! SWITZERLAND: Besazio (Tessin), GV! Follaterres (Valais), LO! ENGLAND: Norwood (London) (Richards 1939); Minsmere Reserve (Suffolk),

BM!

Rhinefield

(Hants),

BM!

Esher Common

(Surrey), BM! Leicester, TW! OL! Watey End (Boxmoor, HT.), BM! Bedford Park (London), BM! Slough (Bucks), BM! Staunton (GW.), BM! Wye (Kent), BM!

Kingsteingnton

SCOTLAND Agher

(SD.),

(Richards 1939);

(Co. Meath),

BM!

BM!

other localities in Richards

IRELAND:

D'dee (DO.),

Donaghadee (Co.

(1939);

Down),

BM!

OL!

BM!

4. Aphelopus camus Richards

Aphelopus heidelbergensis Richards 1939: 286 (syn. proposed by J.F.

Aphelopinae

55

Perkins 1976); locus typicus Heidelberg (Germany). Aphelopus camus Richards 1939: 287 (heidelbergensis has page priority over camus, but J.F. Perkins 1976, the first reviser, placed the former as a synonym of the latter); locus typicus Storeys Way

(Cambridge,

England).

Aphelopus camus Richards: J.F. Perkins 1976: 11, 12. Aphelopus camus Richards: Jervis 1977: 302.

FEMALE:

fully

winged;

clypeus whitish; legs testaceous, tibiae

length

light brown;

granulate;

mm;

black;

mandibles testaceous;

with segment 1 partly

with mid and hind femora dark brown and with antennae distally thickened;

following proportions: 4: = 3;

1,68-1,81

antennae black or brown,

frontal

3,5: 5:

6: 7:

I ine incomplete;

occipital carina complete;

6:

6:

POL = 6;

scutum,

7:

antennal 6:

7;

OL = 3;

yellow hind

segments in

head dull, OOL = 3;

OPL = 3;

TL

scutellum and metanotum dull,

granulate; notaulices incomplete, reaching 0,65 length of scutum; propodeum reticulate rugose, with 2 longitudinal keels on the posterior surface;

fore-wings hyaline,

1,1,2. MALE: fully

winged;

length 1,75-2,25 mm;

testaceous or whitish;

Fig.

36 -

"9nt;

=

bands;

tibial

spurs

mandibles and clypeus legs brown,

with tarsi

Male of Aphelopus camus Richards from Carlsbad genital ia

antennae not distally 3:

black;

antennae black or brown;

(Czechoslovakia):

::-oportions: 4:

without dark transversal

5:

6:

6:

(A)

and ninth abdominal

thickened; 6:

6:

6:

6:

antennal 8;

segments

head dull,

sternite (6).

in following

granulate;

frontal

-e incomplete; occipital carina complete; POL = 6; OL = 4; OOL = 3; OPL 3; TL = 3; scutum, scutellum and metanotum dull, granulate; notaulices

-~omplete,

reaching about 0,65

length of scutum;

propodeum reticulate

56

Mem.

rugose,

with 2 longitudinal

slightly

less rugose than

without

(Fig.

36A);

spurs 1,1,2.

tibial

LOCUS TYPICUS:

keels on

bands;

36B);

hoi otype M!

Inst.,

no.

37,

1984

the posterior surface;

median area

fore-wings hyaline,

ninth sternite of the abdomen posteriorly

inner part of the basivolsella with

Storeys Way

TYPICAL MATERIAL:

Ent.

the parts outside the area;

dark transversal

emarginated

BM;

Amer.

(Cambridge,

holotype M!

England) of A.

and allotype F!

and 4 para types MM

(1

1 bristle (Fig.

heidelbergensis in camus in BM.

!) of A.

seen

in Germany Chlorita sp.

HOSTS:

DISTRIBUTION: Miklos,

BU!

Palaearctic region:

Vacz

CZt=:CHOSLOVAKIA: Heidelberg,

BM!

HUNGARY:

(Tudosdomb),

BU!

Carlsbad,

Bohemia,

Bellinchen

P!

FenyOfb,

Rbjtbkmuzsaj,

(Oder)

TW!

BU!

Bartfa,

BU!

Szod,

BU!

Hoi ics., OL!

SzSzt

BU!

GERMANY:

(holotype and two para types of

Aphelopus chloritae Haupt, spec i es never descr ibed), HA! SW I TZERLAND: Saas-Fee (Valais), Casale Roletto

BM!

ENGLAND:

(Mongrando,

Storeys Way

Vercelli),

OL!

(Cambridge),

BM!

TURKEY: Ankara,

BU!

ITALY:

5. Aphelopus nigriceps Kieffer

A,0,':elopus melaleucus (Da I man) var. nigriceps Kieffer in Kieffer et Marshall

Aphelopus Aphelopus Aphelopus Aphelopus

FEMALE:

fully

1905:

nigriceps nigriceps nigriceps nigriceps

winged;

antennae black,

219.

Kieffer: Kieffer:

Kieffer 1914b: 216. Richards 1939: 291.

Kieffer: Kieffer:

J.F. Perkins 1976: Jervis 1977: 301.

length 1,75-2,25 mm;

black;

with segments 1-2 testaceous;

legs

12.

11,

mandibles

testaceous;

testaceous,

with hind

coxae and hind femora partly brown;

antennae distally thickened;

antennal

segments in following proportions:

5,5:

head

fully POL

granulate,

=

8;

OL

=

dull;

4;

OOL

frontal

=

4;

6:

5:

6:

I ine incomplete;

OPL

=

3,5;

=

TL

6:

6:

occipital

3;

scutum,

6:

5:

5:

7;

carina complete; scutellum and

metanotum smooth, granulate; notaul ices incomplete, reaching about 0,65 length of scutum; median scutal I ine sometimes present; propodeum reticulate rugose,

with 2

median area shiny, wings hyaline, MALE:

fully

longitudinal

less rugose than

without dark transversal

winged;

legs brown,

distally thickened;

antennal

7:

head dull,

7:

8:

8:

11;

segments

carina complete; POL = 9; OL scutellum and metanotum dull, about 0,65 length of scutum;

the posterior surface;

bands;

length 1,75-2,37 mm;

antennae brown or black; 7:

keels on

the parts outside the area; black;

with

tarsi

tibial

mandibles testaceous; light;

antennae not

in following proportions:

granulate;

frontal

fore-

spurs 1,1,2.

line present;

4: 4: 6: 6: occipital

=

4; OOL = 4,5; OPL = 3; TL = 4; scutum, granulate; notaulices incomplete, reaching

median scutal

line sometimes present;

57

Aphelopinae

propodeum reticulate rugose,

with 2 longitudinal

surface;

almost smooth,

median area shiny,

keels on

the posterior

slightly rugose;

fore-wings

8

Fig.

Male of Aphelopus nigriceps Kieffer:

37 -

abdominal

hyal ine,

without dark transversal

genitalia

(A)

and ninth

sternite (B).

bands;

ninth sternite of the abdomen

posteriorly emarginated (Fig. 37B); inner part of the basivolsella with 2 strong bristles (Fig. 37A); tibial spurs 1,1,2. LOCUS TYP ICUS:

Dumfries

(Scotland)

TYPICAL MATERIAL: holotype F! in BM. HOSTS: in England Eurhadina concinna (Germar)(Jervis 1980b, BIOLOGY:

informations in Jervis

JISTRIBUTION: Pulchauki

Palaearctic region:

(Katmandu),

Lemland

(AI.),

Tihany,

BU!

(1980b,

HE!

Vacz.

OT!

OL!

HUNGARY: Cscrcg.,

c);

NEPAL:

FINLAND:

Fenycfc,

BU!

Godavari Lojo,

BU!

c).

in England bivoltine species. (Katmandu),

HE!

Kangasala,

OT!

Istvinkut (Hegyalja),

SzSzt Miklos,

BU!

ROMANIA

OL!

HE! BU!

(Nagy

1972);

CZECHOSLOVAKIA: Donovali (Zvolan), OL! YUGOSLAVIA: Prespa Geul (Otesevo, Makedon ij a), BM! GERMANY: Aachen, P! AUSTR I A: Huben (0 Tirol),

HU!

Semmeringgebeit

(Gld.),

LE!

FRANCE:

=oret de Dabo (Moselle), JL!

Wimbledon Com.

'.:--iorthants),

BM!

'.Surrey),

BM!

iRELAND:

Co.

VI!

(London),

Norfolk,

SCOTLAND: Kerry,

(Reichenau

Le Puy,

TW!

BM!

P!

ENGLAND: OT!

(Fl.),

Rhinefield

Wicklow,

BM!

HOLLAND:

BM!

BM!

Putten

Maisons-Laffitte,

Minsmere Reserve

Bedford Park

Bonar Bridge, Co.

distr.),

Huelgoat

(Suffolk),

(Hants),

BM!

Spratton

(London),

BM!

Horsley

Claddich,

BM!

Dumfries,

TW!

6. Aphelopus nepalensis n. sp.

P!

BM!

BM!

58

Mem.

FEMALE:

fully

winged;

Amer.

Ent.

Ins!.,

no.

length 1,81- 1,87 mm;

37,

black;

1984

mandibles

antennae fully brown or with segments 1-2 testaceous; antennae distally 5:

6,5:

7:

POL; 6;

6:

6:

thickened; 5:

6:

Ol = 3,5;

5:

antennal

9;

segments in following proportions:

head dull,

OOl = 5;

OPl ;

3;

testaceous;

legs testaceous;

granulate; Tl ;

4;

frontal

scutum,

5:

I ine incomplete;

scutellum and

B \ ) (

/ /

/

A Fig.

Male of Aphelopus nepalensis n.

38 -

genitalia

(A:

metanotum dull,

penis and

granulate;

length of scutum; on

MALE:

median area shiny

without dark transversal

fully

winged;

from Bokaihunde (Nepal):

notaulices incomplete,

propodeum reticulate rugose,

the posterior surface;

hyaline,

sp.

left paramere), ninth abdominal left volsella (e)

length

bands;

1,87-1,93 mm;

sternite (B)

reaching

about 0,5

with 2 longitudinal

and smooth;

tibial black;

and

keels

fore-wings

spurs 1,1,2. mandibles testaceous;

antennae brown, sometimes with segment 1 testaceous; legs testaceous, with base of hind coxae black; sometimes also hind femora and hind tibiae

59

Aphelopinae

brown;

antennae not distally

proportions: 4: frontal ;

3,5;

4:

6: 6: 7:

line incomplete; OPL ;

granulate;

2,5;

TL ;

thickened;

6,5:

occipital 3,5;

7:

9;

head dull,

carina complete;

scutum,

notaulices incomplete,

propodeum reticulate rugose,

antenna I segments in following

6,5: 6,5:

POL; 6;

granulate; OL ;

3,5;

OOL

scutellum and metanotum dull,

reaching

about 0,5 length of scutum;

with 2 longitudinal

keels on

the posterior

surface; median area shiny and smooth; fore-wings hyaline, without dark transversal bands; apex of the penis not tridentate (Fig. 38A); inner part of the basivolsella with 2 bristles LOCUS TYPICUS:

TYPICAL MATERIAL: paratype F! HOSTS:

(Fig.

38C);

tibial

spurs 1,1,2.

Bokaihunde (Nepal) holotype M!

and 2 paratypes

(1

F,

1 M)!

in B;

~T.

in

unknown

DISTRIBUTION: Trisuli),

Palaearctic region:

NEPAL:

Bokaihunde (20 Km N of

B! Syabrubens (35 Km N of Trisuli),

B!

27° 58'

N 85° 00'

E,

OT!

7. Aphelopus querceus n. sp.

FEMALE: MALE:

unknown

fully

winged;

length 2,18 mm;

black;

mandibles and clypeus

B

Fig.

39 -

Male of Aphelopus querceus n. ninth abdominal

:estaceous;

head black,

sp.

(type):

genitalia

(A)

and

sternite (B).

with some whitish

spots between

the antennal

sockets; malar space white; antennae black; legs testaceous, with hind "ernora darkened; antennae not distally thickened; antennal segments in

60

Mem.

following proportions: granulate; 3,5; OOL

frontal

Amer.

5:

5:

7:

Ent.

8:

line complete;

9:

Inst.,

9:

9:

no.

9,5:

37,

1984

9,5:

14;

head dull,

occipital carina complete;

POL = 6;

OL

= 5; OPL = 2,5; TL = 2,5; scutum dull, granulate; notaulices

incomplete,

reaching about 0,75

length of scutum;

scutellum granulate;

metanotum shiny, without sculpture; propodeum reticulate rugose, without transversal or longitudinal keels, with a median shiny and smooth area on the posterior surface;

fore-wings hyaline,

without dark transversal

bands;

ninth sternite of the abdomen posteriorly weakly emarginated (Fig. 39B); inner part of the basivolsella with 2 bristles (Fig. 39A); apex of the penis not

tridentate

LOCUS TYPICUS:

TYPICAL MATERIAL: HOSTS:

(Fig.

39A);

Bhurumche

tibial

spurs 1,1,2.

(Nepal)

holotype M!

in OT.

unknown

DISTRIBUTION:

Palaearctic region:

NEPAL:

Bhurumche (Katmandu),

ETHIOPIAN SPECIES OF

OT!

APHELOPUS

Three species are known.

KEY TO THE SPECIES

FEMALES AND MALES

Notaulices margin

of

invisible or the

slightly

visible

scutum •••••••••••••••••

only near

the anterior

l.mediocarinatus (Benoi t)

Notaulices distinct, reaching at least 0,5 2 Notaulices reaching approximately 0,5-0,65

length length

of of

scutum ••••••• 2 scutum •••..•••

2.wittei Benoi t -

Notaulices

reaching

approximately

the posterior margin

scu tum. • • • • • • • • • • • • • • • • • • • • • • • • . • • • ••

3. incisus n.

of

the

sp.

1. Aphelopus mediocarinatus (Benoit) n. comb.

Antaphelopus Antaphelopus locus typicus Antaphelopus

FEMALE:

fully

mediocarinatus Beno it 1951 d: 23. madagascariensis Benoi t 1953b: 429 (n. syn.); Esira

(Madagascar).

mediocarinatus Benoit: Benoit 1953b: 430.

winged;

length 1,81-1,87 mm;

black;

mandibles testaceous,

Aphelopinae

with

teeth reddish;

white;

antennae distally 4:

5:

clypeus black or white;

antennae brown, 6:

5,5:

incomplete;

5:

5:

occipital

face fully

with segments 1-3 testaceous;

thickened;

5:

61

5:

antenna I segments

8;

head dull,

carina complete;

fully

black or partly legs fully

testaceous;

in following proportions: granulate;

POL = 6,5;

frontal

OL = 4;

4:

line

OOL = 3,5;

OPL

=

3; TL = 3; scutum dull, fully granulate; notaulices invisible; scutellum and metanotum shiny, without sculpture or very sl ightly granulate; sometimes dull and distinctly granulate; propodeum reticulate rugose; dorsal surface with wide areolae; 2 longitudinal keels on the posterior

surface;

median

area usually

separated from

transverse keel,

occasionally

reticulate rugose,

rugose along

the margins with central

propodeum reticulate rugose; granulate;

the dorsal

surface by

usually shiny,

area smooth;

lateral

areas of the

mesopleura and metapleura smooth,

fore-wings hyaline,

without dark

transversal

a

reticulate

bands;

fully tibial

spurs 1,1,2. MALE: fully winged; length 1,25-1,62 mm; black; with teeth reddish; clypeus black or light; fully antennae brown,

with segments 1-2 testaceous;

antennae not distally

thickened;

proportions:

5:

frontal

3:

3:

4:

line absent;

OPL = 0,5;

6:

7:

occipital

TL = 2;

scutum,

legs fully

antenna I segments

6,5:

7:

6:

10;

carina complete;

POL = 6;

granulate or reticulate rugose;

usually not reticulate rugose, tibial

transversal

bands;

fully

lateral

granulate;

ninth abdominal

granulate;

fully

OOL = 4; granulate;

2 longitudinal keels on keel; median area dull areas of the propodeum

fore-wings hyaline,

sternite variable

without

(Fig. 40);

spurs 1,1,2.

rig.

40 -

sternite

Male of

Aphelopus mediocarinatus (Benoit): ninth abdominal

(specimens from Garamba National Esira on

_=:US TYPICUS:

Paulis

Park on

the

left and from

the right).

(Zaire)

of A. mediocarinatus in TV; holotype M! of madagascariensis in P; 1 para type M! of A. madagascariensis in TV.

-v"ICAL MATERIAL: ~.

fully

OL = 3;

scutellum and metanotum dull,

ices invisible; propodeum reticulate rugose; the posterior surface and an anterior transversal

dark

testaceous;

in following

head dull,

~otaul

or shiny,

mandibles testaceous, black or partly white;

holotype F!

62

Mem.

HOSTS:

Entebbe, Butahu,

OT! Mt.

Ethiopian region: ZAIRE:

Paulis

Ruwenzori,

Hoyo (Albert Nat.

Ile-ife

(W.

State),

SOUTH AFRICA: Natal), BM!

Ent.

Inst.,

no.

37,

1984

unknown

DISTRIBUTION:

Mt.

Amer.

BM!

Albert Nat.

Park),

BM!

OL!

TV!

Mossel

Bay

Afgoi

TV!

Park), OT!

(Benadir),

Kalonge

TV!

Rwankwi

Ibadan,

Pietermaritzburg

MADAGASCAR:

SOMALIA:

(Haut-Uele),

(N of Kivu Lake),

Umuahia CRIN

Drakensberg

(Cape Province),

BM!

OL!

Fianarantsoa,

Park, TV!

Aff.

TV!

NIGERIA:

(EC State),

TW!

P!

UGANDA:

Katauleko,

Garamba Nat.

(Natal),

Esira,

OL!

(Riv.

BM!

(Van Reenen,

Ceres (Cape Province),

TV!

2. Aphelopus wittei Benoit

Aphelopus wittei Benoit 1951c: 16. Gymnaphelopus wittei (Benoit): Benoit 1951d: 23.

FEMALE: fully winged; length 1,62-2,50 mm; black; head usually with a more or less wide whitish frontal spot; sometimes face black; clypeus white;

antennae fully

legs fully

testaceous;

testaceous or brown,

with segments 1-2 testaceous; is usually black:

in

specimens with wide frontal spots the ventral side of head and body fully whitish; antennae not distally thickened; antennal segments in

is

following proportions:

the ventral

5:

5:

6:

side of the body

8:

8'

7:

5:

5:

granulate; frontal line incomplete; occipital OL = 4; OOL = 5; OPL = 4; TL = 5; scutum, fully

granulate;

notaulices

incomplete,

scutum.

sometimes reaching only 0,5

rugose,

usually with 2 longitudinal

area usually smooth and shiny, dorsal

surface and on

metapleura smooth,

5:

7;

head dull,

usually reaching 0,65 length of

length of scutum; keels on

propodeum reticulate

the posterior surface;

not reticulate rugose;

the sides of the posterior surface;

fore-wings with only

with segments 1-2 testaceous and

legs fully testaceous (at most with stalks of hind femora darkened); some specimens head with a more or less wide frontal whitish spot; sometimes this spot

is very

ocellar region brown; some specimens only

wide:

occasionally head fully

the antennae in the clypeus

is whitish;

antenna I segments

in following proportions:

7:

fully granulate;

occipital scutum,

head dull,

carina complete;

POL

=

7,5;

scutellum and metanotum dull,

incomplete,

OL

=

4;

fully

frontal

in

OOL

approximately 0,5

=

5,5:

5:

5,5:

6:

7:

line incomplete; 5;

granulate;

usually reaching approximately 0,65

occasionally reaching only

testaceous;

antennae not distally

thickened;

10;

in

whitish with

this case are fully

7,5: 7:

7:

the

mesopleura and

without sculpture or sl ightly granulate;

antennae brown

median

areolae only on

hyaline, without dark transversal bands; tibial spurs 1,1,2. MALE: fully winged; length 1,68-2,25 mm; black; head usually mandibles testaceous,

fully

carina complete; POL = 5,5; scutellum and metanotum dull,

OPL

=

4;

TL

=

5;

notaulices

length of scutum;

length of scutum;

propodeum

Aphelopinae

reticulate rugose, shiny

usually

and smooth,

with 2 longitudinal

not reticulate rugose,

occasionally posterior surface of and fully

reticulate rugose;

reticulate rugose, hyaline,

tibial

keels;

median area usually

granulate or without sculpture;

the propodeum without

longitudinal

mesopleura and metapleura smooth,

sl ightly granulate or without sculpture;

without dark transversal

(Fig. 41);

63

bands;

ninth abdominal

keels

not

fore-wings sternite variable

spurs 1,1,2.

B

"'-9.

41 - Male of Aphelopus wittei Benoit: specimens from Ngesho (A), Kirungu

ninth abdominal sternite in (B) and Burunga (C).

_~=~S TYPICUS:

-.=

Kashwa (Zaire) :::AL MATERIAL: holotype F!,

:~-=.:ype

-':::-5:

M!

and 90 paratypes!

in TV;

unknown

::: 5-:;IBUTION: :::- ,,-awasha

=_"',:A.:

allotype M!

in BM. Ethiopian region:

(Salisbury),

Ruhengeri

(Kirii

BM!

ANGOLA:

NIGERIA:

sources),

',',agera Lake (Albert Nat.

TV!

Park),

Gabela,

BM!;

ZIMBABWE:

Ile-ife (W State), ZAIRE:

TV!

BM!

Garamba Nat.

Yolohafiri

Cave

(Mt.

OL! Park, Hoyo,

TV!

1

64

Mem.

Albert Nat. Nord,

Park),

TV!

Albert Nat.

Kivu), (Riv.

TV!

Mt.

Amer.

Hoyo

Inst.,

TV!

(Albert Nat.

Albert Nat.

Park),

Park),

Park),

TV!

OL!

Kalonge

(Riv.

LE!

TV!

TV!

37,

(Riv.

Lumumbashi,

Albert Nat. Park), TV! Mt. Musimba Park), TV! Kalonge (Riv. Kamahoro, Nat.

no.

Secteur Tshiaberimu

Park),

Talia Nord,

Ent.

1984

Kalivina,

OL!

(Secteur Tshiaberimu,

(Secteur Tshiaberimu, Albert Nat. Aff. Butahu, Mt. Ruwenzori, Albert Kamusonge,

aff.

Butahu,

Albert Nat.

Park),

TV!

Kiondo ya Kwanza

Ruwenzori,

Albert Nat.

Park),

TV!

Riv.

Ruwenzori,

Ruwenzori, Nat.

Albert Nat.

Park),

Park),

Albert Nat.

TV!

TV!

Riv.

Park),

Park),

Burunga

(Mokoto,

Mt.

Ruwenzori,

Tshiaberimu,

TV!

Albert Nat.

OL!;

Riv.

Kyanyamu,

(Aff. Mt.

Mt.

Park),

Park),

Park),

TV!

TV!

TV!

Musabaki

Kalonge,

Bombi,

Ruwenzori,

Albert Nat. Park),

(Kyandolire,

TV!

(Aff.

Mt.

Albert

Albert Nat.

Bomboka

Elisabethville,

Riv.

(Aff.

Ruwenzori,

BM!

Mt.

Butahu,

Kakalari

(Kalonge,

Mt.

(Kalonge,

Mt. Sesero (Bitashimva,

Albert Nat.

Albert Nat.

Nyamwamba

Kikyo

Lusilube (Mt.

Ngesho (Kivu),

TV!

TV!

TV!

Talia

(Munagana,

Secteur Tshiaberimu

Kirungu

Ruwenzori, Mt.

aff.

Tshengalero

Hintumo (Secteur dr.

Talia Nord,

Secteur Tshiaberimu, Albert Nat. Park), TV! OL! Rutshuru (Kivu), TV! Mt. Ibale (Kyandolire, Mt. Ruwenzori, Albert Nat. Park), TV! N'Dalage (Kivu), TV!

Rweru

(1951c);

(Volc.

Mikeno,

MADAGASCAR:

Matjesfontein

Albert Nat.

Mandraka,

(Cape Province),

Park),

TV!

BM!

OL!

BM!

Bekily,

other localities in Benoit P!

SOUTH AFRICA:

OL!

3. Aphelopus incisus n. sp.

FEMALE: MALE:

unknown

fully

winged;

teeth brown; testaceous,

with hind coxae,

distally

thickened;

6:

6:

6:

6:

occipital scutum,

length 1,68 mm;

antennae brown,

6:

9;

POL = 6;

granulate; OL = 3;

almost reaching

OOL = 3;

tibial spurs 1,1,2. LOCUS TYPICUS: Umuahia CRIN

6:

antennae 4:

4:

line almost complete; OPL = 2,5;

granulate;

TL = 2,5;

notaulices almost

the posterior margin of the scutum;

fore-wings hyal ine,

TYPICAL MATERIAL:

frontal

with

legs

hind femora and hind tibiae brown;

head dull,

complete;

reticulate rugose;

mandibles testaceous,

antenna I segments in following proportions:

scutellum and metanotum dull,

complete,

HOSTS:

black;

with segments 1-2 testaceous;

without dark transversal

propodeum bands;

(Nigeria)

holotype M!

in BM.

unknown

DISTRIBUTION:

Ethiopian region:

NIGERIA:

OR I ENTAL SPEC I E5 GF

Eight species are known.

Umuahia CRIN

APHELOPUS

(EC State),

BM!

5:

65

Aphelopinae

KEY TO THE SPECIES

FEMALES AND MALES

Species -

fully

testaceous,

with part

of

the antennae brown . . . . . . . . . . .

• • . • • • • • • • . • • • • • • • • • • • • • • • • • • • • • • • • •• 1. borneanus n. sp. Species partly or fully black or brown •••••••••••••.•••••••••..•••• 2

2 Head testaceous, at most with ocellar region or vertex dark •••••••• 3 Head mostly or fully black or brown • • . • • • • • . • • • • • . • • • . • • . . • • . • • • • • • 5

-

3 Notaulices

invisible •••••••••••••••••

2. maculiceps Bergman

Notaulices distinct ••.••••••••.••••••..•.•••••••••••••••••••••••••• 4

4 Notaulices short, reaching at most 0,5 length of scutum . . . . . . . . . . . .. · . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. 3. malayanus n. sp. - Notaul ices reaching approximately 0,80 length of scutum . . . . . . . . . . . .. · . . . . . . . . • . . • . . . • . . . • . . . . . . . . . • . . . . .. 4. ochreus n. sp. 5 Notaulices short, reaching at most 0,5 length of scutum . . . . . . . . . . . . 6 - Notaulices long, reaching the posterior margin of scutum . . . . . . . . . . . 7 6 Notaulices reaching approximately 0,5 length of scutum . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5. orientalis n.sp. - Notaulices reaching approximately 0,3 length of scutum . . . . . . . . . . . . ..

6. penanganus n. sp. 7 Metapleura separated from the propodeum by a strong distinct longitudinal keel (Fig. 42 A) . . . . . . . . 7. birmanus n. sp. -

Metapleura distinctly areolae

Fig.

(Fig.

separated

from

the propodeum by

42 B) . . . . . . . . . . . . . . . . . . 8.

a

series

philippinus n. sp.

42 - Metathorax + propodeum and abdomen of Aphelopus birmanus n.

sp.

(A)

and

philippinus n. sp. (B).

1. Aphelopus borneanus n. sp.

::::::MAlE:

fully

winged;

length 2,37 mm;

:-10 brown;

antennae distally

::;roportions:

5:

'ne incomplete;

4,5:

6:

7:

POL = 7;

5:

testaceous,

thickened; 6:

6:

Ol = 3;

6:

5:

antennal 8;

OOl = 4;

with antennal segments

head dull, OPl = 5;

segments

in following

granulate;

Tl = 5;

frontal

occipital

of

66

Mem.

carina complete; notaulices

scutum,

Amer.

En!.

Ins!.,

no.

37,

scutellum and metanotum dull,

reaching approximately 0,5

incomplete,

1984

propodeum reticulate rugose,

with 2 longitudinal

surface;

without dark transversal

1,1,2. MALE:

fore-wings hyaline,

granulate;

length of scutum; keels on

the posterior

bands;

tibial

spurs

unknown

LOCUS TYPICUS:

Tenompok

TYPICAL MATERIAL: HOSTS: unknown DISTRIBUTION:

(Sabah,

holotype F!

Oriental

Malaysia)

in B.

region:

MALAYSIA:

Tenompok

(Sabah),

B!

2. Aphelopus maculiceps Bergman

Aphelopus maculiceps Bergman 1957: 9.

FEMALE

and MALE:

fully

winged;

with ocellar region brown; testaceous;

4:

4:

= 3;

not

4:

absent;

1,25-1,37 mm;

thorax and propodeum black,

abdomen brown-black; female,

length

antennae brown,

thickened

4:

3,5:

3:

occipital

scutum,

3:

legs testaceous; in male; 3:

2,5:

sometimes with scutum brown;

antennae distally thickened

antenna I segments 5;

carina complete;

head granulate, POL

head testaceous,

with segments 1-3 or 1-4

=

5; OL

scutellum and metanotum dull,

=

in following dull;

frontal

3; OOL

granulate;

=

in

proportions: line

4; OPL

=

3; TL

notaulices

invisible; propodeum reticulate rugose, with 2 longitudinal keels on the posterior surface; median area partly smooth; fore-wings hyaline, without dark transversa I bands; LOCUS TYPICUS:

t i b i a I spurs 1,1,2.

Bogor (Java)

holotype M and allotype F in LE; 1 paratype F! in 1 paratype F! in BM; paratypes in Bogor (Division of Entomology, Faculty of Agriculture). HOSTS: in Java Empoasca sundaica Bergman (Bergman 1957). DISTRIBUTION: Oriental region: INDONESIA: Bogor (W Java), BM! WA! TYPICAL MATERIAL:

WA;

3. Aphelopus malayanus n. sp.

FEMALE: occiput in

fully dark;

length

1,80 mm;

the only female specimen known);

with dorsal granulate; 4;

winged;

OOL

=

regions brown; frontal

5;

incomplete,

head

testaceous,

antennae with segment 1 testaceous

OPL

=

5;

TL

=

5;

vertex and

thorax and propodeum testaceous,

abdomen brown;

line complete;

with

(other segments missing

occipital

scutum dull,

reaching approximately 0,5

legs testaceous;

carina complete; granulate;

head dull, POL = 6;

OL

notaulices

length of scutum;

scutellum dull,

Aphelopinae

granulate;

metanotum shiny,

reticulate rugose,

with 2

fore-wings hyal ine, MALE:

fully

lengt

keels on

transversal

abdomen

propodeum

the posterior surface;

bands;

tibial

1,43-1,56 mm; head testaceous,

and part of vertex dark;

three segments dark; or brown; propodeum or black;

without sculpture;

longitudinal

without dark

winged;

ocellar region

smooth,

67

spurs 1,1,2. sometimes with

antennae testaceous,

with

last

thorax testaceous, usua II y with dorsa I reg ions dark testaceous, with dorsal and posterior surface brown

brown-testaceous;

legs testaceous;

antennae not d ista II y

4: 4: 5: 5: 5: 5: 5: 5: 5: 5; head dull, granulate; frontal line complete; ocipital carina complete; POL = 6; Ol = 3; OOl = 3; OPl = 4; Tl = 3; scutum, scutellum and metanotum dull, granulate; notaulices incomplete, reaching at most 0,5 thickened;

antennal

length of scutum;

segments

without dark

_OCUS TYPICUS:

TYPICAL MATERIAL:

with 2 longitudinal

median area smooth and shiny;

transversal

Klang Gates

:0; 1 paratype F! ..,OSTS:

proportions:

propodeum reticulate rugose,

on the posterior surface; -,yal ine,

in following

bands;

(Malaya,

holotype M! in OL.

tibial

keels

fore-wings

spurs 1,1,2.

Malaysia)

and 1 paratype F!

in B;

1 paratype M!

unknow n

JISTRIBUTION: Oriental region: MALAYSIA: Klang Gates (Malaya, Kuala _,-,mpur), B! Gomak For. Res. (Malaya, Kuala lumpur), Ol! Kuala Tahan "',alaya,

Pahang),

B!

PHiliPPINES:

Pinigisan

(Mantalingajan,

Palawan),

:O!

=:::':AlE:

fully

winged;

-=;ion and vertex dark; :==taceous and

dorsal

=:;:;omen brown; ==;'71ents

=_ :, c.::

n,

length 1,81

testaceous ,

mm;

head

antennae testaceous;

regions brown;

legs testaceous;

sp.

thorax

with

with ocellar ventral

side

propodeum brown or blackened;

antennae distally

thickened;

antenna I

3,5: 3,5: 4: 5: 5: 6: 6: 6: 6: 8; head line complete; POL = 5; OL = 3; OOl = 4; OPL =

in following proportions:

granulate; Tl = 3;

__

4. Aphelopus ochreus

frontal

occipital

granulate;

carina complete;

notaulices reaching

scutum,

:-::)odeum reticulate rugose,

with 2 longitudinal

o_-;ace;

without dark

fore-wings hyaline,

scutellum and metanotum

approximately 0,8

length of scutum;

keels on

transversal

the posterior

bands;

tibial

spurs

.. 2. ~_:::

unknown

_::.,;S TYPICUS: Fraser's Hill (Malaya, -.:; CAL MATERIAL: holotype F! in B.

-:::':::-5: :

Malaysia)

unknown

:-=
Oriental

region:

MALAYSIA:

Fraser's Hill

5. Aphelopus orientalis n. sp.

(Malaya),

B!

in

68

Mem.

FEMALE:

fully

winged;

antennae brown, distally 4:

4:

4:

4:

9;

antennal

no.

black;

~

POL

6;

approximatelyO,5-0,65

granulate;

Ol

1984

mandibles testaceous; legs testaceous;

~

3,5;

frontal ~

OOl

granulate;

antennae

length of scutum;

6:

line incomplete;

3,5;

~

OPl

2;

~

Tl

4:

5:

4:

occipital 2;

notaulices incomplete,

scutum,

reaching

propodeum reticulate rugose,

keels on the posterior surface;

fore-wings hyaline,

37,

segments in following proportions:

scutellum and metanotum dull,

shiny;

Inst.,

length 1,56 mmj

head dull,

carina complete;

2 longitudinal

Ent.

with segments 1-2 testaceous;

thickened;

4:

Amer.

with

median area smooth and

without dark transversal

bands;

tibial

spurs

1,1,2. MALE:

fully

winged;

antennae brown,

length 2,68 mm;

black;

mandibles testaceous;

with segment 1 testaceous;

hind femora brownj

legs testaceous,

antennae not distally thickened;

following proportions:

5:

6:

6:

7:

7:

8:

8:

8:

8:

9;

with clubs of

antennal

segments in

head dull,

granulate;

line complete; occipital carina complete; POL ~ 10; Ol ~ 5; OOl ~ 5; Tl ~ 4; scutum, scutellum and metanotum dull, granulate;

frontal 4; OPl

notaul ices incomplete, reaching approximately 0,5 length of scutum; propodeum ret icu I ate rugose, with a strong transversa I keel between and posterior surface; median area shiny, transversal

bands;

lOCUS TYPICUS:

not rugose;

two

longitudinal

fore-wings hyaline,

dorsa I

keels;

without dark

spurs 1,1,2. (Taiwan)

holotype F!

in B;

paratype M!

in WA.

unknow n

DISTRIBUTION: SRI

smooth, tibial

Fenkihu

TYPICAL MATERIAL: HOSTS:

posterior surface with

lANKA:

Oriental

region:

Adams Peak Trail

TAIWAN: (4,5 mi.

Fenkihu

(Chiayi Hsien),

W Maskeliya,

Kan.

B!

Dist.),

WA!

6. Aphelopus penanganus n. sp.

FEMALE:

fully

winged;

length

1,68 mm;

black;

mandibles testaceous;

antennae testaceous, with segments 7-10 dark; legs testaceous; antennae distally thickened; antenna I segments in following proportions: 4: 4: 5: 5: 4,5:

4,5:

occipital scutum,

4:

4:

4:

5;

head dull,

carina complete;

POL

~

granulatej 6;

Ol

scutellum and metanotum dull,

~

smooth; 1,1,2. MALE:

line incomplete; ~

3;

OPl

~

Penang

(Malaya,

holotype F!

bands;

tibial

Malaysia) in B.

unknown

DISTRIBUTION:

Oriental

region:

MALAYSIA:

Tl

median area shiny

without dark transversal

unknown

TYPICAL MATERIAL:

2,5;

~

3;

notaul ices incomplete,

propodeum reticulate rugose,

keels on the posterior surface;

fore-wings hyaline,

lOCUS TYP ICUS: HOSTS:

OOl

granulate;

reaching approximately 0,3 length of scutumj with 2 longitudinal

frontae

3,5;

Penang

(Malaya),

B!

and

spurs

Aphelopinae

69

7. Aphelopus birmanus n. sp.

FEMALE:

fully

winged;

1,87 mm;

length

black-brown;

mandibles testaceous;

antennae brown, with segments 1-2 testaceous; legs testaceous, with hind femora and hind tibiae dark; antennae distally thickened; antenna I segments complete; TL = 2;

in

occipital scutum,

reaching

the holotype);

= 5;

POL

= 3;

OL

the posterior margin of the scutum;

fore-wings hyaline,

without dark

two

= 4;

OOL

granulate;

line OPL

= 2;

notaulices

propodeum reticulate rugose,

keels on the posterior surface; strong distinct

(last frontal

metapleura separated

longitudinal

transversal

keel

bands;

42 A);

(Fig.

tibial

spurs 1,1,2.

unknow n

LOCUS TYP ICUS:

Kambatti

TYPICAL MATERIAL: HOSTS:

granulate;

scutellum and metanotum dull,

the propodeum by a

MALE:

head dull,

carina complete;

with 2 longitudinal from

5: 4: 5: 5: 5: 5: 4: 4,

in following proportions:

segments missing

(Burma)

holotype F!

in BM.

unknown

DISTRIBUTION:

Oriental

region:

BURMA:

Kambatti

(NE Burma),

BM!

8. Aphelopus philippinus n. sp.

FEMALE: unknown MALE: fully winged;

length 1,60 mm;

and ma 1ar space testaceous; ::lropodeum brown; "airy, 4: 4:

abdomen brown-testaceous;

not distally 5:

6:

:occipital

6:

7:

7:

thickened; 7:

7:

carina complete;

scutum dull,

head brown,

granulate;

with mandibles,

antennae testaceous-brown;

7;

antennal

head dull,

POL

=

6;

4; OOL

notaulices complete,

scutellum and metanotum shiny,

smooth,

antennae

in following proportions:

granulate;

=

OL

legs testaceous;

segments

clypeus

thorax and

=

frontal

3;

OPL

=

I ine complete;

4; TL

=

very

sl ightly granulate;

propodeum

-eticulate rugose, with 2 longitudinal keels on the posterior surface; -etapleura separated from the propodeum by a series of areolae (Fig. 3); fore-wings hyal ine, . , 1,2. _JCUS TYPICUS:

Pinigisan

-vPICAL MATERIAL: -JSTS:

without dark

transversal

(Philippine

holotype M!

bands;

tibial

spurs

Islands)

in CO.

unknown

= STRIBUTION: ::alawan),

Oriental

region:PHILIPPINES:

4;

posteriorly separated;

Pinigisan

CO!

NEARCT I C SPEC I ES OF APHELOPU5

(Mantalingajan,

42

70

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

Four species are known.

KEY TO THE SPECIES

FEMALES AND

Notau Ii ces -

MALES

i nv is i b Ie ••••.•••••.••.•••

2 Fore-wing with a dark

transversal

band;

testaceous ••••••••••••••••••••••••••• -

rufiventris Ashmead

1.

Notaulices distinct •••••••••••••••••••.••••••••.••.••.••••••••.••.• 2

Fore-wing hyaline,

without

dark

at

least

transversal

thorax and propodeum mostly black

or

the

scutum mostly

bicolor Fenton

2.

band;

dorsal

reddish-brown;

side of

occasionally

scutum partly testaceous • • • • • • . . • . • • • • . . • . . • • • • . • • • . • . • . • • • • • • . • . • • 3 3 Notaulices reaching beyond 0,65 length of scutum •••••••••••••••••••. ••••••••••••••••.••.••.••••••••.••••. -

Notaulices

4 Notaulices

reaching at

most

0,5

reaching approximately

0,5

. • • • • • • • • . • • • • • • • • • • • • • • • • • • . • • • • . • •. -

Notaulices

reaching

at

most

0,25

of

scutum •••.••••••••.•••••. 4

length

of

scutum .••••••••.•.••

albopictus Ashmead

4.

length

• • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • ••

varicornis Brues

3.

length

of

1.

scutum •••••••••••.••••.••

rufiventris Ashmead

1. Aphelopus rufiventris Ashmead

Aphelopus rufiventris Ashmead 1893: 100. Aphelopus affinis Ashmead 1893: 102 (no syn.); locus typicus Canada.

Aphelopus rufiventris Ashmead : Fenton 1918b: 276. Aphelopus rufiventris Ashmead: Muesebeck and Walkley 1951: 1042. Aphelopus rufiventris Ashmead: Krombei n 1979: 1250.

FEMALE AND MALE:

fully

winged;

length

with antennae brown or testaceous; segments 1-2 or 1 testaceous; reddish or black, testaceous;

hind

1,31-1,81

mm;

dark-red or black,

sometimes antennae brown,

clypeus black,

reddish or white;

with face

sometimes with a more or less wide white spot; legs with coxae,

antennae distally thickened

legs

clubs of femora and tibiae brown;

in female,

not thickened

in male;

antennal

segments in following proportions: (F) 3: 3,5: 4,5: 4,5: 5: 5: 5: 6: 6: 10; (M) 3: 4: 4: 4: 4,5: 5: 6: 6: 6: 8; head dull, fully granulate; frontal I ine absent; scutum,

occipital

carina complete;

POL=8;

OL=5;

scutellum and metanotum sl ightly granulate;

reaching only approximately 0,3

length of scutum;

OOL=S;

OPL=3;

TL=4;

notaul ices invisible or

propodeum reticulate

Aphelopinae

71

rugose, with or without longitudinal keels and median area on the posterior surface; fore-wings hyaline, without dark transversal bands; tibial

spurs 1,1,2.

lOCUS TYPICUS: Jacksonville (Florida,

U.S.A.)

TYPICAL MATERIAL: holotype F! of A. rufiventris in WA; lectotype F! (hereby designated) of A. affinis in WA (in WA a female labelled Aphelopus tibialis Ashmead Type agrees with the original description of A. affinis and is hereby designated as lectotype of A. affinis: A. tibialis was never described). HOSTS: unknown DISTRIBUTION: Nearctic region: CANADA: Gagnon Rd. (Terrace, B. C.), Mc Murray (Alta.), OT! Ottawa (Ont.), OT! Ol! St. lawrence Is. Nat.

OT!

Park (Ont.), OT! Ol! Kouchibouguac N. P. (N. B.), OT! Ol! U.S.A.: Portal (Chiricahua Mts., Cochise Co., Ariz.), SCI Speir Rch. (Uvalde, Uvalde Co., Tex.), SC! Jacksonville (Fla.), WA! Wattacoo (Pickens Co., C.),

TW!

OLl

Pink Beds (N. C.),

TW!

Washington

(D.

C.),

WA!

S.

IT!

2. Aphelopus bicolor Fenton

Aphelopus bicolor Fenton 1924: 192. Aphelopus bicolor Fenton: Muesebek and Walkley 1951: 1042. Aphelopus bicolor Fenton: Krombein 1979: 1250.

FEMALE AND MALE: fully winged; length 1,37-1,59 mm; testaceous or reddish dark, with propodeum, pet iole and abdomen reddi sh-brown; antennae brown, with segments 1-2 or 1-3 testaceous; sometimes propodeum fully black or with only the dorsal surface reddish-brown; antennae distally thickened in female, not thickened in male; antennal segments in following proportions: (F) 3,5: 3: 4: 4: 4: 4: 4: 4: 4: 6; (M) 5: 5: 7: 7: 7: 7: 6,5: 6,5: 6: 10; head dull, granulate; frontal line absent; occipital carina complete; POL = 6; Ol = 3,5; OOl = 3; OPl = 3,5; Tl = 3; scutum, scutellum and metanotum dull, granulate; notaulices incomplete,reaching approximately 0,5 length of scutum; propodeum reticulate rugose, with two longitudinal keels on the posterior surface; median area partly smooth; fore-wings with a dark transversal band beneath the pterostigma; tibial s p u rs 1, 1 , 2 • lOCUS TYPICUS: Spring Hill (Iowa, U.S.A.) TYPICAL MATERIAL: holotype F! in WA. HOSTS: in U.S.A. Hymetta trifasciata (Say) (Fenton

1924;

Krombein

1979).

ClISTRIBUTION: Nearctic region: CANADA: Kouchibouguac N. P. (N. B.), OT! Jl! U.S.A.: Spring Hill (Iowa), WA! Arkansas (Muesebeck and Walkley 1951; Krombein 1979); Cleveland (S. C.), TW! Yorklin (Del.), Ol!

72

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

3. Aphelopus varicornis Brues

Aphelopus varicornis Brues 1906b: 143. Aphelopus varicornis Brues: Fenton 1918b: 277. Aphelopus microtomus Wh i ttaker 1928: 389 (n. sy n. ); locus typicus Chilliwack (Canada). Aphelopus pilicornis Whittaker 1928: 389 (n. syn.); locus typicus Ch i II iwack (Canada). Aphelopus microtomus Whittaker: Muesebeck and Walkley 1951: 1042.

Aphelopus varicornis Brues: Muesebeck and Walkley 1951: 1043. Aphelopus microtomus Wh i ttaker: Krombei n 1979: 1250. Aphelopus varicornis Brues: Krombei n 1979: 1250.

FEMALE AND MALE:

fully

winged;

length

1,25-2,20 mm;

black;

mandibles

testaceous; usually clypeus also testaceous or whitish; face sometimes fully black, usually with a more or less wide whitish or testaceous spot; antennae fully occasionally

brown or with segments 1-2 or only

scutum partly

testaceous;

and stalk of hind femora brown; antennae distally

thickened

absent or present;

in female,

occipital

TL = 5;

scutum,

notaulices incomplete,

not thickened

testaceous;

in male;

antennal

(F) 5: 4: 5: 5: 5,5: 5: 5,5: 4: 4: head dull, granulate; frontal line

carina complete;

POL = 5;

OL = 3;

scutellum and metanotum dull,

5;

OOL = 4;

granulate;

reaching approximately 0,65-0,75 length of scutum;

propodeum reticulate rugose,

with 2 longitudinal

surface;

smooth;

median area partly

LOCUS TYPICUS:

with hind tibiae

sometimes hind tibiae fully

segments in following proportions: (M) 5: 4: 5: 6: 6: 6: 6: 5: 5: 8; OPL = 4;

1 testaceous;

legs testaceous,

tibial

keels on

the posterior

spurs 1,1,2.

Woods Hole (Massachusetts,

U.S.A.).

varicornis in ML; holotype M! and 1 paratype F! of A. microtomus in BM; 1 para type M! of A. microtomus in LA; holotype M! and 8 para types (6 FF, 2 MM)! of A. pilicornis in BM; 3 paratypes MM! of A. pilicornis (two very damaged) in LA. TYPICAL MATERIAL:

HOSTS:

of A.

holotype F!

unknown

DISTRIBUTION:

Nearctic region:

ALASKA:

OT!

Mile 140 Alaska Steese Hwy.,

OT!

Hixon

(B.C.),

(B.

OT!

C.),

OT!

Oliver (B.

OT!

Kleanza Cr. C.),

OT!

M.

289 Richard Hwy.

CANADA: (Terrace,

Spring Creek

Norman Wells B.

C.),

OT!

(Terrace,

(Shaw Crk.), (N.

T.),

W.

Burnaby

B.

C.),

OT!

Terrace (B. C.), OT! Cultus Lake (B. C.), OT! Robson (B. C.), OT! Chilliwack (B.C.), BM! LA! Galiano (B. C.), BM! Quadra Is. (B. C.), Chowichan L. C.),

OT!

OT!

Hatzic Lake (B.

Diamond Head Trail

(Banff Nat. OT!

(B. C.),

(Squamish,

B.

C.),

OT!

C.),

OT!

Pitt Meadows

Johnston Canyon

Park,

Alta.),

OT!

Medicine Hat.

(Alta.),

Elkwater Lk.

(Alta.),

OT!

Oldman River

(Lethbridge,

Jumping Pd.

Cr.

(Calgary,

Alta.),

OT!

SC!

(B.

OT!

Kenosee (Sask.),

Waterton Alta.), OT!

(Alta.), OT!

La Ronge

Aphelopinae

73

(Sask.), AM! Int. Peace Gardens (Turtle Mtn. For. Res., Man.), OT! Innisville (Ont.), OT! Ottawa (Ont.), OT! Ol! One Sided lake (Ont.), Marmora

(Ont.),

OT!

Ol!

Finland

(Ont.),

OT!

Bell's Corners

(Ont.),

OT!

OT!

Oxford Mills (Ont.), OT! North Branch (Ont.), OT! Ol! Maynooth (Ont.), OT! Tillsonburg (Ont.), OT! Dundas (Ont.), OT! Hull (Que.), OT! Ol! Cap Rouge (Que.), OT! Summit of King Mt. (Old Chelsea, Que.), OT! Kouchibouguac N. P. (N. B.), OT! U.S.A.: Corvallis (Oreg.), TW! Ochoco Creek (Oreg.), TW! Selma (Oreg.), TW! Mt. Hood (Oreg.), TW! Pinehurst (Oreg.), TW! H)'att Reservoir (Oreg.), TW! lowman (Idaho), TW! Stanley (Idaho), TW! OLl Salmon Creek Campground (Sierra City, Sierra Co., Calif.), CAl Bilby (Alta, Calif.), CAl Arroyo Seco Campgr. (Monterey Co., Calif.), SCI Oynard (Ventura Co., Calif.),SC! Crescent City (Calif.), TW! Somerset (EI Dorado, Calif.), TE! Clark Fork River (26 mi. NE Strawberry, Alpine Co.,

Calif.),

AM!

S.l.O. Reservoir (S.l.O.

Co.,

Calif.),

TE!

Atescadero (S.l.O. Co., Calif.), TE! Portal (Cochise Co., Chiricahua Mts., Ariz.), TW! SCI lee Creek (1,5 mi. SW lolo Hot Springs, Missoula Co., Mont.), AM! Doolittle Ranch (Mt. Evans, Colo.), OT! Speir Rch. (Uvalde, Uvalde Co., Tex.), SCI Heep Farm (Austin, Travis Co., Tex.), TE! lawrence (Kans.), TW! Williamsville (Mo.), OT! Ann Arbor (Mich.), TW! Ol! Indian Gap (Tenn.), OT! Dade Co. (Fla.), Ol! Pine Mountain (Rabun Co., Ga.), OT! Wattacoo (Pickens Co., S.C.), Ol! TW! Greenville (S.C.), TW! Cleveland (S.C.), TW! Highlands (N.C.), TW! OT! Clingman's Dome (Grt. Sm. Mt. Nat. Park, N.C.), OT! Pink Beds (N.C.), TW! Pisgah Mt. (N. C.), TW! Reed's Gap (Augusta Co., Va.), OT! Mt. Jackson (Shenandoah Co., Va.), OT! Takoma Park (Md.), TW! High Point St. Pk. (N.J.), TW! Moorestown (N.J.), TW! Farmingdale (N.Y.), TW! Ithaca (N.Y.), TW! Pughkeepsie (N.Y.), TW! Shokan (N. Y.), TW! Milford Center (N.Y.), TW! Hancock (N.Y.), TW! Kalbfleisch Field Station (Huntington, long Island, N.Y.), AM! lake Placid (N.Y.), OT! Cornell University (Ithaca, N.Y.), IT! Woods Hole (Barnstable Co., Mass.), Ml! Mt. Ktaadn (Maine), WA! Lakes of the Clouds (Mt. Washington, N. H.), OT! MEXICO: EI Saito (Durango), OT! la Ciudad (Durango), OT! EI Palmito (Sinaloa), OT! Concordia (Sinaloa), OT! Toluca (Mexico), OT! Ixtapan de la Sal (Mexico), OT!

4. Aphelopus albopictus Ashmead

Aphelopus albopictus Ashmead 1893: 101. nec Aphelopus albopictus Ashmead: Ashmead 1895: 787 Aphelopus arizonicus R.C.l. Perkins 1907: 53 (n. syn.); locus typicus Nogales (U.S.A.). Aphelopus viduus R. C. l . Perk i ns 1907: 54 (n. syn.); locus typicus Nogales (U.S.A.). Aphelopus pulcherrimus R.C.l. Perkins 1907: 54 (n. syn.); locus typicus Nogales (U .S.A.). Aphelopus microleucus R.C.l. Perkins 1907: 55 (n. syn.); locus

74

Mem. Amer.

Ent.

Inst.,

no.

37,

1984

typicus Nogales (U.S.A.). Aphelopus dikraneuri Fenton 1918b: 276 (n. syn.); locus typicus Columbus (U.S.A.).

Aphelopus Aphelopus Aphelopus Aphelopus

microleucus R.C.L. Perkins: Fenton 1918b: 276. arizonicus R.C.L. Perkins: Fenton 1918b: 277. viduus R.C.L. Perkins: Fenton 1918b: 277. comesi Fenton 1918b: 277 (n. syn.); locus typicus Columbus

(U.S.A.) •

Aphelopus albopictus Ashmead: Fenton 1918b: 278. Aphelopus pulcherrimus R.C.L. Perkins: Fenton 1918b: 278. Aphelopus typhlocybae Muesebeck 1935: 167 (n. syn.); locus typicus Poughkeepsie (U.S.A.).

Aphelopus typhlocybae Muesebeck: Dumb I eton 1937: 869. Aphelopus albopictus Ashmead: Muesebeck and Walkley 1951: 1042. Aphelopus arizonicus R.C.L. Perkins: Muesebeck and Walkley 1951: 1042.

Aphelopus comesi Fenton: Muesebeck and Walkley 1951: 1042. Aphelopus dikraneuri Fenton: Muesebeck and Walkley 1951: 1042. Aphelopus microleucus R.C.L. Perkins: Muesebeck and Walkley 1951: 1042.

Aphelopus pulcherrimus R.C.L. Perkins: Muesebeck and Walkley 1951: 1042.

Aphelopus typhlocybae Muesebeck and Walkley 1951: 1042. Aphelopus viduus R.C.L. Perkins: Muesebeck and Walkley 1951: 1043. Aphelopus albopictus Ashmead: Krombein 1958: 100. Aphelopus albopictus Ashmead: Krombei n 1979: 1250. Aphelopus arizonicus R.C.L. Perkins: Krombein 1979: 1250. Aphelopus comesi Fenton: Krombei n 1979: 1250. Aphelopus dikraneuri Fenton: Krombein 1979: 1250. Aphelopus microleucus R.C.L. Perkins: Krombein 1979: 1250. Aphelopus pulcherrimu sR. C. L. Perk i ns: Krombe i n 1979: 1250. Aphlopus typhlocybae Muesebeck: Krombei n 1979: 1250. Aphelopus viduus R.C.L. Perkins: Krombein 1979: 1250.

FEMALE AND MALE: fully winged; length 1,31-2,25 mm; black or reddishbrown; mandibles testaceous or whitish; clypeus black, brown, testaceous, or whitish; face fully black or with a more or less wide whitish spot; antennae fully brown or with segments 1-2 or only 1 testaceous; legs fully testaceous or with some parts more or less brown; sometimes legs almost fully brown, with only tarsi testaceous; sometimes only hind tibiae and stalk of hind femora brown; antennae distally thickened in female, not thickened in male: antenna I segments in following proportions: (F) 4: 4: 4: 5: 4,5: 5: 4: 4: 4: 6; (M) 5: 4,5: 5,5: 6: 6: 6: 6,5: 6: 6: 8; head dull, granulate; frontal I ine absent or present; occipital carina complete; POL = 6; OL = 4; DOL = 4; OPL = 3; TL = 3; scutum, scutellum and metanotum dull, granulate; notaulices incomplete, reaching approximately

Aphelopinae

0,5 length of scutum;

propodeum reticulate rugose,

75

with 2 longitudinal

keels on the posterior surface and with median area rugose or partly smooth; fore-wings hyaline, without dark transversal bands; tibial spurs 1,1,2. LOCUS TYPICUS: Washington

(D.C.,

U.S.A.).

TYPICAL MATERIAL: holotype F! of A. albopictus in WA; lectotype F! and 2 para lectotype FF! (designated by Olmi 1982) of A. arizonicus in B; lectotype M! and 1 para lectotype M! (designated by Olmi 1982) of A. viduus in B; holotype F! of A. pulcherrimus in B; lectotype M! (designated by Olmi 1982) of A. microleucus in B; holotype M! and 2 para type FF! of A. dikraneuri in WA; holotype M! of A. comesi in WA; holotype F! and 49 paratypes (3 seen !) of A. typhlocybae in WA; 2 paratypes (IF, 1M)! of A. typhloc~Tbae in BM •• HOSTS: in U. S.A. Typhlocyba pomaria Mc Atee (Muesebeck 1935), Dikraneura sp. (R.C.L. Perkins 1907), Typhlocyba sp. (R.C.L. Perkins 1907), Erythroneura sp. (R.C.L. Perkins 1907), Forcipata sp. (probably loca De L.

and C.)

(Krombein 1979); according to Fenton 1918b the host of A.

dikraneuri is Dikraneura fieberi; Erythroneura spp. (Krombein 1979); according to Fenton (1918b) the host of A. comesi is Erythroneura comes; according to Fenton (1924) also Erythroneura vulnerata Fitch; in New Zealand (imported: see Dumbleton 1937) Typhlocyba froggatti Baker. DISTRIBUTION: Nearctic region: CANADA: Rampart House (Yukon), aT! Chilliwack (B.C.), BM! Ketchum L. (B. C. ), aT! Gagnon Rd. (Terrace,

B.

C.), aT! Hixon (B. C.), aT! Terrace (B. C.), aT! Robson (B. C.), aT! Morley (Alta.), aT! Ninette (Man.), aT! Ottawa (ant. ), aT! Merivale (ant.), aT! Vineland Sta. (Ont.), aT! Marmora (Ont.), aT! Rondeau Provo Pk. (Ont.), aT! Finland (Ont.), aT! Brighton (ant.), aT! Blackburn (Ont.), aT! Cumberland (ant.), TW! Stittsville (Ont.), TW! Old Chelsea (Que.), aT! Summit King Mt. (Old Chelsea, Que.), aT! Mistassini (Que.), aT! Hull (Que.), aT! aLI Abbotsford (Que.), aT! Kouchibouguac N. P. (N. B.), aT! Saulok (N. 5.), aT! U.S.A.: Selma (Oreg.), TW! OL! Corvallis (Oreg.), TW! Seaside (Oreg.), TW! Needles (Calif.), IT! Alameda (Calif.), B! OL! Ogilby Rd. (Chocolate Mts., Imperial Co., Calif.), SCI Somerset (EI Dorado, Calif.), TE! Nogales (Ariz.), B! Portal (Ariz.), TW! Speir Rch. (Uvalde, Uvalde Co., Tex.), SCI Ames (Iowa), WA! Columbia (Boone Co., Mo.), WA! Bentonville (Ark.), WA! Lake Oachita S. P. (Ark.), aT! Ann Arbor (Mich.), BM! Columbus (Ohio), WA! Elm-Maple For. (Nashville, Tenn.), WA! Greenville (S. C.), TW! Wattacoo (Pickens Co., S. C.), TW! Cleveland (S.C.), OL! TW! Highlands (N. C.), TW! OL! Virginia (Krombein 1958); Mt. Lk. Bioi. Sta. (Va.), TW! Spring Br. (Pa.), TW! Maryland (Muesebeck and Walkley 1951); Poughkeepsie (N. Y.), TW! WA! BM! New York Agr. Exp. Stn. (N. Y.), NZ! Ithaca (N. Y.), aT! Otsego L. (N. Y.), TW! Canajoharie (N. Y.), TW! Shokan (N. Y.), TW! Milford Center (N. Y.), TW! Lewisboro (Westchester Co., N. Y.), AM! South Haven (Long Island, N. Y.), AM! Washington (D. C.), WA! Mt. Washington (N. H.), aT! MEXICO: 6 Km S of Maolinco (Veracruz, 1700 m), AM! Neotropic region: the record from Grenada I. (Ashmead 1895) is referable

76

Mem.

En t.

Amer.

I nst.,

to the new species Aphelopus diffusus Austra I i an reg ion: probably

NEW ZEALAND

unsuccessfully

no.

37,

1984

(see Neotropic species).

(imported as A.

typhlocybae: Dumb I eton 1937;

introduced).

APHELOPUS

NEOTROPIC SPECIES OF

Nine species are known.

KEY TO THE SPECIES

FEMALES AND MALES

-

Frontal

line complete ••••••••••••••••

Frontal

I ine absent

2 Scutum testaceous, -

Scutum black,

or

whi te

brown

or

3 Species fully whitish, antennae almost

1.

tropicalis n. sp.

incompl ete •••••••••••••••••••••••••••.•••••• 2 or yellow •••••••.•.••••••••••••••••••••••• 3

reddish-black ••••••••••••••••.•.•.••••••.••. except

completely

dorsal

dark ••••••

surface of

2.

7

the propodeum brown;

diaphanus n.

sp.

Species at least partly dark, differently colored •••••••••••••••••• 4 4 Prop odeum redd i sh • • . • • • • . • • • • • . . • • • •• 3. surinamensis n. sp. -

-

Propodeum black •••.••••••••••••••••••••••••••••••••.••••••••••••..• 5

5 Scutellum testaceous ••••.••••.•..•••• 4. -

6 Notaulices reaching approximately 0,5

I ength

8. -

fentoni n. sp.

Scu tel I um brown •••••••••.•••••••••••••••••••••••••••••••••••••.•••• 6

Notaulices

reaching

approximately

0,3

of

scutum •••••••••••.•.

alvarenganus n.

length

of

9. jamaican us n.

7 Notaulices almost

invisible,

slightly

sp.

scutum .•••••••••••••

visible near

sp.

the anterior

margin of the scutum •••••••••••••••••• 5. leucopus Kieffer Notaulices reaching at least 0,5 length of scutum •.•••••••••••••••• 8

8 Notaulices reaching about

0,5

length

.. . . •. . .. . . .. . . . . . . . . . . . .. . . .. . . . . . " -

of

6.

scutum •••••••••••••••••.••••

diffusus n.

sp.

0,65-0,75 length of scutum •••••••• . . . . . . . . . . . . . . . . . . . • . . . . . . • . . • . . • • • .• 7. trinitatis n. sp.

Notaulices

reaching approximately

1. Aphelopus tropicalis n. sp.

FEMALE AND MALE:

fully

winged;

clypeus and mandibles whitish; whitish;

antennae brown;

length 1,81-2,25 mm;

black;

in male malar space and

sometimes antennal

in female

lower face also

segment 1 testaceous;

legs

Aphelopinae

77

testaceous; sometimes coxae, clubs of femora and tibiae of mid and hind legs brown; clubs of fore femora dark; antennae distally thickened in female, (F)

not

thickened

in male;

antennal

5: 5: 7: 8: 9: 9: 9: 7: 7:

dull,

granulate,

frontal

2,5; TL = 3; scutum,

11;

segments in following proportions:

5: 5: 6: 9: 8: 8: 8: 7: 7: 9; head

(M)

line complete;

scutellum

POL = 7;

OL = 4;

and metanotum dull,

OOL = 4;

granulate;

incomplete,_ reaching approximately 0,65

length of scutum;

reticulate rugose,

keels on

with

median area smooth transversal

bands;

LOCUS TYPICUS: in AM;

HOSTS:

propodeum

the posterior surface;

fore-wings hyal ine,

without dark

spurs 1,1,2.

Chimborazo (Ecuador) holotype F!

1 paratype M!

2 paratypes MM!

in CA;

1 paratype

in TW;

in OL ..

unknown

DISTRIBUTION:

Neotropic region:

Chimborazo (30 mi. Puerto

longitudinal

and shiny; tibial

TYPICAL MATERIAL: M!

two

OPL =

notaulices

Ibanez

TRINIDAD:

SW of Alausi),

(Aysen),

CAl

Arima Valley,

Ascazubi-Pito,

TW!

AM!

ECUADOR:

OL!

CHILE:

TW!

2. Aphelopus diaphanus n. sp.

FEMALE:

fully

winged;

length

1,68 mm;

whitish,

with dorsal

surface of the

propodeum and antennal segments 5-8 brown; antennae distally thickened; antennal segments in following proportions: 4: 4: 4,5: 5: 5: 5: 5: 4,5 (last 2 segments missing line absent;

occipital

3; TL = 2,5; scutum, incomplete,

in

the holotype);

with 2 longitudinal

median area smooth;

granulate;

Ol = 4;

scutellum and metanotum dull,

reaching approximately 0,5

reticulate rugose,

head dull,

POL = 5;

carina complete;

granulate;

length of scutum; keels on

fore-wings hyaline,

frontal

OOl = 3;

OPl =

notaulices

propodeum

the posterior surface;

without dark transversal

bands;

tibial spurs 1,1,2. MALE: unknown lOCUS TYPICUS:

Gamboa

TYPICAL MATERIAL:

(Panama)

holotype F!

HOSTS: unknown DISTRIBUTION: Neotropic region: Zone),

in LEo PANAMA:

Gamboa

(Pipeline Road,

Canal

LE!

3. Aphelopus surinamensis n. sp.

FEMALE: unknown MALE: fully winged; and vertex and

lateral

dark;

length 1,56 mm;

antennae brown,

head whitish,

with

side of the thorax whitish;

with ocellar region

segments 1-3 testaceou5; scutum,

ventra I

scutellum and propodeum

78

Mem.

testaceous-dark;

abdomen

distally thickened; 4,5: 5:

5:

occipital scutum,

5:

Amer.

4,5:

En!.

Inst.,

testaceous-brown;

antenna I segments 4:

4:

6,5;

carina complete;

head dull,

POL; 5;

granulate;

OL ;

3,3;

DISTRIBUTION:

Neotropic region:

3;

4:

3,5:

line absent;

OPL ;

2,5;

TL ;

propodeum reticulate rugose, median area smooth;

bands;

tibial

spurs 1,1,2.

in LE. SURINAME:

Billiton,

LE!

4. Aphelopus fen toni n. sp.

FEMALE:

fully

region brown;

winged;

length 2,06 mm;

antennae brown,

and clypeus testaceous; metap leura,

scutum and scutellum

legs testaceous;

5;

Ol ;

granulate;

3,5;

OOl ;

metanotum dull,

3;

frontal OPl ;

granulate;

0,5 length of scutum;

6:

testaceous;

4,5: 5,5:

line absent; 2,5;

Tl ;

5:

mandibles

mesopleura,

propectus testaceous; 4,5: 4,5:

occipital

2,5;

hyaline,

without dark transversal

tibial

4:

6; POL

scutellum and

with 2

longitudinal

median area smooth and shiny; bands;

4:

reaching approximately

propodeum reticulate rugose,

the posterior surface;

antenna I 4:

carina complete;

scutum,

notaulices incomplete,

keels on MALE:

with ocellar

antennae distally thickened;

segments in following proportions: head dull,

head testaceous,

with segments 1-2 testaceous;

metanotum and propodeum brown-b lack;

abdomen brown;

fore-wings

spurs 1,1,2.

unknown

lOCUS TYPICUS:

Nova Teutonia (Brazil)

TYPICAL MATERIAL: HOSTS:

holotype F!

in BM.

unknown

DISTRIBUTION:

Neotropic region:

BRAZil:

Nova Teutonia (S.ta Catarina),

BM! NOTE:

the species is named

in honor of the

late special ist Mr.

F.

A.

Fenton.

5. Aphelopus leucopus Kieffer

Aphelopus leucopus Kieffer 1906a: 239.

FEMALE:

fully

winged;

antennae brown,

length

1,25-1,62 mm;

black;

with segments 1-2 testaceous;

2;

notaulices incomplete,

keels on the posterior surface;

holotype M!

antennae not

frontal

OOL ;

granulate;

fore-wings hyaline, without dark transversal LOCUS TYP I CUS: Bill i ton (Suri name) TYPICAL MATERIAL: HOSTS: unknown

1984

legs testaceous;

approximately 0,65 length of scutum;

with 2 longitudinal

37,

in following proportions:

scutellum and metanotum dull,

reaching

no.

mandibles testaceous;

legs testaceous,

with hind

Aphelopinae

coxae,

hind femora and hind tibiae brown;

antennal 3,5:

79

6;

segments

in following proportions:

head dull,

complete;

=

POL

granulate;

=

6; OL

3;

frontal

invisible,

slightly

=

OOL

scutellum and metanotum dull,

4:

4,5: 4:

2,5;

OPL

granulate;

=

4: 4:

3,5:

occipital

carina

=

scutum,

3,5;

TL

notaulices

3;

incomplete,

3,5:

almost

visible near the anterior margin of the scutum;

propodeum reticulate rugose,

Managua

TYPICAL MATERIAL: HOSTS:

4:

line absent;

with 2 longitudinal

surface; median area smooth and shiny; transversal bands; tibial spurs 1,1,2. MALE: unknown LOCUS TYPICUS:

antennae distally thickened;

keels on

the posterior

fore-wings hyaline,

without dark

(Nicaragua)

holotype F!

in CA.

unknown

DISTRIBUTION: (Tabasco),

Neotropic region:

NICARAGUA:

Managua,

CAl

MEXICO:

Teapa

BM!

6. Aphelopus diffusus n. sp.

;"ec Aphelopus albopictus Ashmead 1893: 101. Aphelopus albopictus Ashmead: Ashmead 1895: 787. Aphelopus albopictus Ashmead: Ashmead 1900: 235.

FEMALE AND MALE: reddish-dark;

fully

winged;

face usually

sometimes head almost fully testaceous or whitish; testaceous;legs fully tibiae brown; male; 8:

antennal

7,5:

frontal OPL =

10;

1,56-2,12 mm;

testaceous;

testaceous,

TL = 2;

4:

5:

6:

occipital

scutum,

notaulices incomplete,

reddish or

clypeus and mandibles black or

sometimes with hind femora and hind thickened

in female,

segments in following proportions:

(M) 4,5:

black,

antennae fully brown or with segments 1-2

antennae distally

line absent; 1;

length

with a more or less wide whitish spot;

6:

7:

6:

6:

6:

carina complete;

7;

not

(F) 4:

4:

5:

head dull,

POL

=

7;

scutellum and metanotum dull,

reaching

thickened

approximately 0,5

6:

in 7:

8:

8:

granulate;

=

OL

3;

OOL

=

4;

granulate;

length of scutum;

propodeum reticulate rugose, with 2 longitudinal keels on the posterior surface; median area smooth and shiny; fore-wings hyaline, without dark transversal

bands;

tibial

spurs 1,1,2.

LOCUS TYPICUS: Nova Teutonia (Brazil) TYPICAL MATERIAL: holotype M! and 1 para type F!

in BM;

2 paratypes

MM! in WA; 5 para types (4 MM, 1 F)! in OT; 1 paratype M! paratypes MM! in AM; 5 paratypes (2 FF, 3 MM)! in OL. HOSTS: unknown D I STR I BUT I ON: Neotrop ic reg ion:

in TW;

2

GRENADA I.: SI. John's River (Aphelopus albopictus Ashmead misident.: Ashmead 1895), WA! Balthazar (Aphelopus albopictus Ashmead misident.: Ashmead 1895), WA! ANTIGUA I.: 51. John's,

80

Mem.

AM!

COLOMBIA:

ECUADOR: OT!

Amer.

Ol!

Insl.,

Municipio la Cumbre

Portete de Tarqui,

BM!

Enl.

TW!

Estacao Forestal

no.

(Hda.

BRAZil:

37,

1984

la Maria,

Valle),

Nova Teutonia

Cabeca do Veado

Ol!

(S.ta Catarina),

(Distrito Federal),

OT!

Ol!

7. Aphelopus trinitatis n. sp.

FEMALE AND MALE:

fully

winged;

more or less wide whitish

spot;

length 1,62-1,87 mm;

black;

face with a

clypeus and mandibles whitish or

testaceous; antennae brown, with segmen ts 1-2 or 1-3 testaceous, somet i mes with mid and hind coxae and femora brown; sometimes also hind tarsi brown;

antennae distally thickened

antennal 5:

7;

(M)

absent; Tl

=

in female;

segments in following proportions: 6: 8: 8:

6:

occipital

8:

9:

9:

not

(F)

thickened

4:

3,5:

8: 9; head dull,

8:

carina complete;

POL; 6;

Ol ;

4:

in male;

5:

6:

granulate;

3;

6:

7:

6:

frontal

OOL = 3,5;

line

OPL = 3;

3; scutum, scutellum and metanotum dull, granulate; notaulices

incomplete, reaching approximately 0,65-0,75 length of scutum; propodeum reticulate rugose, with 2 longitudinal keels; median area smooth and shiny;

fore-wings hyaline,

1,1,2. lOCUS TYP ICUS:

without dark transversal

Chaguaramas Bay

bands;

tibial

spurs

(Trinidad)

TYPICAL MATERIAL: holotype M! and 8 paratypes (1 F, 7 MM)! in BM; paratypes (8 MM, 7 FF)! in aT; 5 paratypes (4 MM, 1 F)! in TW; 1 paratype M! in AM; HOSTS: unknown DISTRIBUTION:

4 paratypes (1

Neotropic region:

3 MM)!

F,

TRINIDAD:

15

in Ol.

(51.

Chaguaramas Bay

George),

BM! Arima Valley (51. George), BM! Ol! Curepe (S.ta Margarita), BM! GRAND CAYMAN: Georgetown, OT! MEXICO: Teapa (Tabasco), BM! S. Cristobal OT!

de las Casas (Chiapas),

COLOMBIA:

ECUADOR: Quito-S.

TW!

Ol!

Yerba Buena

(Hda.

la Maria,

(Bochil,

Valle),

Chiapas),

AM!

Tulcan,

TW!

loja,

Tarqui,

Ol!

Vn.

Chiles,

TW!

Troya,

Domingo,

OL!

Portete de Tarqui,

Ol!

Papallacta,

TW!

BRAZil:

Nova Teutonia Veado,

OT!

Municipio La Cumbre

(S.ta Catarina),

Distrito Forestal),

OT!

OT!

BM!

Ol!

ARGENTINA:

Estacao Forestal

TW!

(Cabeca do

Horco Molle (Tucuman),

OT!

8. Aphelopus alvarenganus n. sp.

FEMALE:

fully

of the face,

winged;

1-2 testaceous;

scutum testaceous;

propodeum black;

abdomen brown;

distally thickened; 6:

6:

5:

5:

length 2 mm;

head testaceous,

clypeus and mandibles whitish;

5:

9;

carina complete;

antennal head dull,

POL = 5,5;

with anterior region

antennae brown,

with segments

scutell um and metanotum brown; legs fully

testaceous-whitish;

segments in following proportions: granulate; Ol = 4;

frontal

OOl = 3;

line absent;

OPL = 2;

antennae 5:

4:

6:

occipital

Tl = 2;

scutum,

6:

Aphelopinae

scutellum and metanotum dull, approximately 0,5 central

granulate area on

without MALE:

dark transversal fully

winged;

OPL = 1,5;

4,5: 4: TL = 2;

LOCUS TYP I CUS:

tibial

with

a

spurs 1,1,2.

structurally resembles the female;

antenna I segments in following

6: 6: 6: 6: 6: 9;

POL

=

5; OL

=

4;

OOL

=

4;

spurs 1,1,2.

holotype M!

in TW;

Braz i I)

1 paratype F!

in OL.

unknown

DISTRIBUTION: TW!

thickened;

reaching

fore-wings hyal ine,

Represa Rio Grande (Guanabara,

TYPICAL MATERIAL: HOSTS:

bands;

5: 6: tibial

notaul ices incomplete,

propodeum reticulate rugose,

the posterior surface;

length 2 mm;

antennae not distally proportions:

granulate;

length of scutum;

81

Neotropic region:

BRAZIL:

Represa Rio Grande (Guanabara),

OL!

9. Aphelopus jamaican us n. sp.

FEMALE: MALE:

unknown

fully

winged;

region of the face, scutum

length

testaceous;

thickened;

antennal 8,5;

legs testaceous;

head dull,

granulate;

with 2 longitudinal

median area rugose;

frontal

propodeum black; 3:

4: 4,5: 5:

line absent;

slightly granulate;

keels on

fore-wings hyaline,

Hardwar Gap

TYPICAL MATER IAL: HOSTS:

6:

occipital

without dark

transversal

(Jamaica)

holotype M!

Neotropic region:

in OT. JAMAICA:

Hardwar Gap,

AUSTRAL I AN SPEC I ES OF APllELOPUS

Five species are known

(one introduced).

KEY TO THE SPECIES

FEMALES AND MALES

propodeum

the posterior surface;

unknown

DISTRIBUTION:

anterior

antennae not di sta II y

spurs 1,1,2.

LOCUS TYPICUS:

with

antennae testaceous;

segments in following proportions:

scutellum and metanotum dull,

reticulate rugose, tibial

testaceous-dark,

POL = 5,5; OL = 4; OOL = 3,5; OPL = 1; TL = 2; scutum, notaulices incomplete, reaching approximately 0,30 length

carina complete; dull, granulate; of scutum;

head

scutellum and metanotum brown;

abdomen brown-testaceous; 6,5: 6,5: 7: 7:

1,56 mm;

clypeus and mandibles whitish;

OT!

bands;

82

Mem.

Head and

thorax

Amer.

Ent.

no.

37,

1984

testaceous . . . . . . . . . . . . . . . . .

townesi n.

• • . • • • • • • • • • • • • • • • • • • • • • • 1. -

Inst.,

sp.

thorax mostly black . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2

Head and

2 Notaulices reaching approximately 0,65-0,75 length of -

Notaulices

3 Frontal

reaching

line absent

0,5

or

(invisible near

incomplete

oce I Ius) .••.•••••.••••••• 3.

length

australis n.

of

scutum . . . . . . . 3

scutum . . . . . . . . . . . . . 4

approximately

the anterior

sp.

- F ron t a I lin e comp let e ••.• 4. caledonicus n. sp. 4 Posterior surface of the propodeum with median area shiny and smooth, not rugose . . . . . . . 2. guineensis n. sp. -

Posterior

surface of

the propodeum with median area dull

and

albopictus Ashmead (see Nearct ic species)

rugose . . . . . . . . . . . . . . . . . . . 5.

1. Aphelopus townesi n. sp.

FEMALE: MALE:

unknown

fully

winged;

length

segments 4-10 dark, antennae not

distally

proportions: 5: I ine complete;

5:

7:

thickened; 8:

= 8;

POL

carina complete;

1,75-2,43 mm;

metanotum brown,

scutum

8:

9:

antennal

8:

= 6;

OL dull,

testaceous,

7:

7:

OOl

segments

9; head dull, granulate; frontal OPl = 6; Tl = 5; occipital

granulate;

notaulices incomplete,

length of scutum;

sl ightly

propodeum reticulate rugose,

granulate;

between

shiny dark

dorsal

and smooth,

lOCUS TYP I CUS:

Atherton

with a strong

transversal

posterior surface centrally

laterally reticulate rugose;

TYPICAL MATERIAL:

reaching

scutellum and metanotum shiny,

and posterior surface;

transversa I bands;

abdomen dark;

in following

= 4;

approximately 0,65 keel

with antennal

propodeum black,

fore-wings hyal ine,

without

t i b i a I spurs 1,1,2. (Austra I i a)

holotype M!

and 1 paratype M!

in TW;

1 paratype M!

in Ol. HOSTS:

unknown

DISTRIBUTION: NEW GUINEA: NOTE:

Australian region: Baiyer R.,

the species

TW!

is named

AUSTRALIA:

Atherton

(Queensland),

TW!

OL! Henry K.

in honor of Mr.

Townes.

2. Aphelopus guineensis n. sp.

FEMALE AND MALE: testaceous;

fully

winged;

length 1,56-1,87 mm;

clypeus brown or testaceous-dark;

black;

antennae brown,

mandibles with

segments 1-2 testaceous and segment 3 partly or fully testaceous; legs testaceous; antennae distally thickened in female, not thickened in male; antennal 5;

(M)

3:

segments 4:

4,5:

in following proportions: 6:

6:

7:

7:

6,5:

6,5:

9;

(F)

5:

4:

head dull,

5:

5:

5:

4:

granulate;

4:

4:

frontal

4:

Aphelopinae

I ine absent; 2,5;

occipital

carina complete;

83

=

POL

5;

=

Ol

3;

=

OOl

3;

OPl

= 2,5; scutum, scutellum and metanotum dull, granulate;

Tl

notaul ices

incomplete,

reaching

propodeum reticulate rugose, suerfacej

approximately 0,5 length of scutum;

with 2 longitudinal

median area smooth and shiny;

transversal

bands;

lOCUS TYPICUS:

tibial

keels on

without

dark

spurs 1,1,2.

Kundiawa

(New Guinea)

TYPICAL MATERIAL: holotype F! and 1 paratype F! in TW; 4 paratypes (1F, 3MM)! in Ol. HOSTS:

the posterior

fore-wings hyaline,

in B;

6 paratypes MM!

unknown

DISTRIBUTION:

Australian

NE New Guinea),

B!

(NE New Guinea),

region:

Hospital

B!

Tari

NEW GUINEA:

Sk.

Gap

(Wau,

(Mt.

Kundiawa

NE New Guinea),

Hagen),

TW!

(East Highlands, 8!

Kassam Pass

Ol!

3. Aphelopus australis n. sp.

FEMALE AND MALE:

fully

testaceousj

whitish

clypeus

1-2 testaceous;

winged; or

legs testaceous;

femora brown;

length 1,43-1,75 mm;

testaceousj

antennae distally thickened

antenna I segments

in following

4:

4:

6:

frontal

7;

(F)

5:

line absent;

4:

5:

mandibles

or with

segments

somet i mes with mid and hind coxae and

male; 4:

black;

antennae brown

5:

5:

in female,

proportions:

4,5:

4,5:

4,5:

occipital carina complete;

not

thickened

(M) 4,5: 7;

POL

4:

4:

head dull,

=

5,5;

5:

5:

granulate;

=

Ol

in

5:

3;

=

OOl

4; OPl = 2,5; Tl = 2; scutum, scutellum and metanotum dull, granulate; notaulices incomplete, reoaching approximately 0,65-0,75 length of scutum; propodeum reticulate rugose, surface;

transversal

bands;

tibial

lOCUS TYP ICUS: Casula TYPICAL MATERIAL: in B;

spurs

keels on

the posterior

fore-wings hyaline,

without dark

1,1,2.

(Austral ia)

holotype M!

9 paratypes MM!

HOSTS:

with 2 longitudinal

median area smooth and shiny;

in TW;

and 1 para type F! 4 paratypes

(1F,

in BM;

3MM)!

1 paratype F!

in Ol.

unk now n

DISTRIBUTION:

Australian

BM!

(Victoria),

Warburton

Yangaborra

region: TW!

(NE Queensland),

Nebo (Queensland),

TW!

AUSTRALIA:

Casula

Foot of Black Mtn. B!

Mt.

Port Arthur

Glorious

(New South Wales),

(Canberra),

(Queensland),

(Tasmania),

TW!

BM!

TW!

Ol!

Mt.

Ol!

4. Aphelopus caledonicus n. sp.

FEMALE:

fully

winged;

length

1,43-1,68 mm;

black;

mandibles testaceous;

clypeus and part of the face or part of the malar space whitish; br·own, distally

with segments 1-2 or 1-3 testaceousj thickened;

antennal

legs testaceous;

segments in following

antennae

antennae

proportions:

3,5:

3:

4:

84

5:

Mem.

5:

5:

5:

5:

5:

carina complete;

7;

Amer.

head dull,

POL; 5;

OL ;

scutellum and metanotum dull, approximately 0,65-0,75 with 2

longitudinal

Ent.

Inst.,

granulate; 3;

OOL ;

granulate;

length of scutum;

keels on

smooth or slightly rugose;

no.

37,

1984

frontal

3;

line complete;

OPL ;

2;

TL ;

4;

occipital

scutum,

notaulices incomplete,

reaching

propodeum reticulate rugose,

the posterior surface;

fore-wings hyaline,

median area shiny,

without dark

transversal

bands; tibial spurs 1,1,2. MALE: unknown LOCUS TYPICUS:

Col

TYPICAL MATERIAL: HOSTS:

d'Amieu

(New Caledonia)

holotype F!

in B;

1 para type F!

in OL.

unknown

DISTRIBUTION: NEW GUINEA:

Australian region: Umboi

I.

(1

NEW CALEDONIA:

Km N of Awelkom,

Col

d'Amieu,

NE New Guinea),

B!

DL!

5. Aphelopus albopictus Ashmead (see Nearctic species)

2. Crovettia n. gen.

TYPE SPEC I ES:

Crovettia theliae (Gahan 1918)

FEMALE: maxillary pal,:" with 5 segments; labial palpi with 3 segments; mandibles tridentate, with teeth progressing larger from anterior one to posterior; fore-wing with radial vein forming a very pronounced curve; usually distal and proximal part of radial vein forming an angle; at least the anterior surface of the scutum reticulate rugose; running close to the antennal MALE:

sclerites;

structurally resembles the female;

thickened

(in Cr.

antennae slightly DISTRIBUTION:

epistomal

suture

antennae distally thickened. antennae not usually distally

barbara from the Neotropic region, the male has distally

Nearctic,

thickened).

Neotropic

HOSTS: Membra cidae SPECIES: 7

NEARCTIC SPECIES OF CROVETTIA

Two species are known.

KEY TO THE

SPECIES

85

Aphelopinae

FEMALES AND MALES

Head,

scutum,

scutellum and metanotum fully

maxillary palpi

with

segment

1 much

longer

reticulate than

rugose; 2 ................. .

1. nearctica n. -

Head,

scutellum and metanotum not

granulate;

scutum granulate,

with

reticulate

sp. fu I I y

the anterior surface

rugose; maxillary palpi with segment •• • • • • •• • • • • •• • • • • • • • • •• • • • •• . . • • •• •

Fig. 43 -

rugose,

1 nearly as 2.

Male of Crovettia nearctica n.

theliae

sp.

reticulate

long as (Gahan)

from Forsyth

2

....... .

(Georgia).

1. Crovettia nearctica n. sp.

=EMALE:

fully

:eeth reddish;

winged;

length 2,25-3,12 mm;

antennae brown,

black;

mandibles testaceous,

with segment 1 testaceous;

coxae and clubs of femora brown;

legs testaceous,

antennae distally thickened;

antennal

with with

86

Mem.

segments palpi

Ent.

in following proportions:

with segment

1 much

and propodeum dull, carina comlete; invisible;

fully

=

POL

10;

Inst.,

8:

6:

no.

6:

longer than 2;

6:

=

5;

=

OOL

5:

9;

1984

4:

head,

reticulate rugose; OL

37,

4:

without dark transversal

shorter than proximal part

(2:8),

4:

scutum,

frontal

=

OPL

bands;

4:

7;

maxi Ilary

scutellum,

metanotum

line incomplete;

8;

TL

posterior surface of the propodeum without

wings hyaline, much

Amer.

=

10;

longitudinal

distal

occipital

notaul ices keels;

part of radial

fore-

vein

the two parts forming an angle;

tibial

spurs 1,1,2. MALE

(Figs.

testaceous;

25,

43):

fully

winged;

antennae testaceous;

Fig.

44 -

length 2,50-2,81

legs testaceous,

mm;

black;

mandibles

with coxae black and femora

Male genitalia of Crovettia nearctica n. sp.

(type).

brown; antennae not distally thickened; antennal segments in following proportions: 7: 6: 5: 7: 6: 6: 7: 6: 6: 9; maxi Ilary palpi with segment longer than 2; occipital scutum,

carina complete;

fully POL

=

reticulate rugose; 9;

OL

scutellum and metanotum dull,

invisible;

propodeum reticulate rugose,

longitudinal

keels;

part of radial forming

head dull,

fore-wings hyaline,

=

4,5 OOL

fully dull,

=

8;

OPL

without

=

8;

TL

transversal (2:7);

=

6;

notaulices

without dark transversal part

much

line incomplete;

reticulate rugose;

vein much shorter than proximal

an angle;

frontal

or bands:

distal

the two parts

gonoforceps with apex not externally emarginated

(Fig.

44);

87

Aphelopinae

spurs 1,1,2.

tibial

LOCUS TYPICUS:

Forsyth

TYPICAL MATERIAL:

(Georgia,

holotype M!

U.S.A.)

and 3 para types

paratype F!

in lO;

1 paratype F!

(1

in AM;

4 para types MM!

F,

1 M)!

HOSTS:

in SC;

(2 MM,

F)!

3 paratypes MM!

in OT;

in TW;

1

2 paratypes

in OL.

unknown

DISTRIBUTION: Ariz.),

SCi

Cleveland

Nearctic region:

Forsyth (S.

(Ga.),

C.),

TW!

Conn.), AM! South (Sinaloa), OT!

OL!

Haven

U.S.A.:

OT!

OLl

Portal

Wattacoo

Wilmington

(Long

(Chiricahua Mts., (Pickens Co.,

lO!

(Del.),

Island,

N.

Y.),

S.

Westport

AM!

OL!

Cochise Co.,

C.),

TW!

OLl

(Fairfield Co.,

MEXICO:

EI

Palmito

2. Crovettia theliae (Gahan) n. comb.

Aphelopus theliae Gahan 1918: 151. Aphelopus theliae Gahan: Muesebeck and Walkley 1951; 1042. Aphelopus theliae Gahan: Krombei n 1979: 1250.

FEMALE: teeth with

fully

winged;

reddish-brown; tarsi

light;

length 2,06-2,20 mm;

antennae distally

thickened;

7: 6: 5: 4: 4: 4: 4: 4: 4: 6;

proportions:

maxi Ilary palpi

with segment

black;

mandibles testaceous,

with segments 1-;2

antennae brown,

1 nearly

as

testaceous;

antenna I segments

head dull, long

fully

as 2;

with

legs brown;

in following

granulate;

frontal

I ine

incomplete;

occipital carina complete; POL = 9; OL = 5; OOL = 5; OPL = 3; TL = 4; scutum dull, granulate; with anterior third reticulate rugose; notaulices invisible; scutellum fully

granulate;

metanotum reticulate rugose;

propodeum fully

reticulate rugose, without longitudinal keels on the posterior surface; forewings hyaline, without dark transversal bands; distal part of radial vein

(2:4),

much shorter than proximal

part

sp u rs 1, 1 , 2. MALE: fully winged;

1,93-2,18 mm; black; mandibles testaceous,

teeth brown; light;

antennae not distally

proportions: segment

occipital

scutum dull,

invisible;

legs brown,

thickened;

two parts forming

with fore

an

angle;

tibiae and fore

antenna I segments

tibial with

tarsi

in following

8: 5: 4,5: 4: 4: 3,5: 3,5: 3,5: 3,5: 6; maxillary palpi with

1 nearly as

incomplete; = 4;

length

antennae brown;

the

long

granulate,

scutellum dull,

reticulate rugose,

as 2;

head dull,

carina complete;

without

with

POL

fully

=

9;

granulate;

OL

=

5;

OOL

frontal

=

4;

anterior third reticulate rugose;

granulate;

metanotum shiny,

longitudinal

keels on

punctate;

line

OPL

=

3;

propodeum

the posterior surface;

fore-

wings hyal ine, without dark transversal bands; distal part of radial vein much shorter than prox i ma I part (2:4), the two parts form i ng an ang Ie; gonoforceps with apex externally emarginated LOCUS TYPICUS:

Cold Spring Harbor

TYPICAL MATERIAL:

holotype F!

(New

(Fig.

York,

45);

tibial

spurs 1,1,2.

U.S.A.)

and 4 paratypes

(1

F,

3 MM)!

TL

notaulices

in WA.

88

Mem.

Fig. 45 -

Amer.

Ent.

Ins!.,

no. 37,

1984

Male genitalia of Crovettia theliae (Gahan) (New York)

from Cold Spring Harbor

HOSTS: in U.S.A. Thelia bimaculata Fabr. (Membracidae) (Gahan 1918; Kornhauser 1919). BIOLOGY: informations in Kornhauser (1916, 1917, 1919); the species is polyembryonic; the external larval sac is absent. DISTRIBUTION: Nearctic region: CANADA: Richter Pass (OsoyOOS, B. C.), OT! OL! Rockport (Ont.), OT! U.S.A.: Glenville (Kern Co., Calif.), TE! lone (Amador Co.,

Calif.),

SCI

5 mi.

W Portal

(Cochise Co.,

Ariz.),

AM!

Alexandria

(La.),

89

Aphelopinae

aT!

Ol!

Harbor Haven

Roscommon (Long

(long

(Mich.),

Island, Island,

N.

OL!

Y.),

N.

Y.),

TW!

WA!

Washington

lewisboro

C.),

Ol!

Cold Spring N.

Y.),

Ol!

South

AM!

NEOTROPIC SPECIES OF

Five species

(D.

(Westchester Co.,

CROVETTIA

are known.

KEY TO THE SPECIES

FEMALES AND MALES

Fore-wing with distal

part

of

radial

vein much

shorter

than proximal

••••• 2 part • • . . • . . .• .•. . . . •. . • . . .••••••. ••• 4 2 Head, scutum, scutellum and metanotum fully ret iculate rugose. 3. plaumanniana n. sp. - At most head, anterior surface of the scutum and metanotum reticulate rugose; scutellum never reticulate rugose .•••..••.•••••• 3 3 Head only granulate, without irregular keels; pal much longer than Ol. • • • • • • • • • • • • • • • • • . • • • • . • . • • • • • • . • 4. brasiliana n. sp. - Head granulate with irregular keels; pal shorter than Ol . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . " 5. colombian a n. sp. 4 Head ret iculate rugose . . . . . . . . . . . . . . . 1. barbara n. sp. - Head not reticulate rugose, fully granulate •••••••••. 2. neotropica n. sp. part .• '" •••...•••••••••••••.••••••• Fore-wi ng wi th d i sta I part of rad i a I

vein

longer

than

proximal

1. Crovettia barbara n. sp.

FEMALE:

fully

winged;

antennae brown,

length

and fore tibiae testaceous; in following

1,50-1,87 mm;

black;

with segments 1-2 testaceous;

proportions:

antennae distally

8:

4:

3:

2,5:

3:

3:

mandibles testaceous;

legs testaceous with thickened;

:J: 3:

3:

antennal

4;

tarsi

segments

head dull,

::wanulate and fully reticulate rugose, less strongly reticulate rugose than 'C1 male; frontal line incomplete; occipital carina complete; POL = 6; Ol 3;

OOl = 4,5;

-..Jgose only ,;-anulate, -.Jgose;

OPl = 1,5;

Tl = 2;

scutum dull,

near the anterior margin; not reticulate rugose;

granulate,

notaulices

reticulate

invisible;

scutum dull,

metanotum and propodeum reticulate

propodeum without transversal

or longitudinal

keels;

fore-wings

90

Mem.

hyaline,

fully

part

antennal

Fig.

head dull,

length

invisible;

genitalia:

hyaline,

46;

tibial

part of radial

black;

vein

longer

mandibles testaceous; 6:

4:

with

Ol = 3;

3:

3:

3:

3:

frontal

3:

3:

4,5;

line complete;

OOl = 4; OPl = 2;

Tl = 2,5;

numerous areolae on the anterior half; granulate;

metanotum and propodeum

longitudinal

keels on

without dark transversal

twice as long as proximal spurs 1,1,2.

Nova Teutonia

3:

barbara n. sp. type.

reticulate rugose;

scutellum dull,

vein more than fig.

mm;

propodeum without

fore-wings

lOCUS TYPICUS:

distal

1984

spurs 1,1,2.

POL = 6;

granulate and

37,

antennae slightly distally thickened;

granulate and fully

reticulate rugose;

bands;

no.

Male genitalia of Crovettia

46 -

scutum dull,

of radial

1,81

Ins!.,

in following proportions:

carina complete;

notaulices

tibial

legs brown;

segments

occipital

(6:4);

winged;

antennae brown;

surface;

En!.

without dark transversal

than proximal MALE:

Amer.

(Brazil)

the posterior

bands;

part

distal

(7,5:3);

part

91

Aphelopinae

TYPICAL MATERIAL: F!

HOSTS:

in BM;

1 paratype F!

in OT;

1 paratype

unknown

DISTRIBUTION: BM!

holotype M!

in OL.

Caraca

Sao Paulo),

Fig.

Neotropic region:

(S. OT!

47 -

Barbara,

BRAZil:

Minas Gerais),

Nova Teutonia OT!

Bocaina

(S.ta Catarina),

(S.

Jose Barreiro,

Ol!

Male genital ia of Crovettia neotropica n.

sp.

type

92

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

2. Crovettia neotropica n. sp.

FEMALE AND MALE: testaceous-brown; testaceous; antennal

fully

length 1,43-1,56 mm;

winged;

antennae brown;

legs brown,

antennae distally thickened

segments

with

in female,

in following proportions:

black;

tarsi

not

mandibles

and fore tibiae

thickened

in male;

5,5: 4: 4: 4: 4: 4: 4: 3: 3:

5,5; head dull, granulate; frontal line absent; occipital carina complete; POL = 5; Ol = 2,5; OOl = 3; OPl = 2,5; TL = 3; scutum dull, granulate and with nUmerous areolae near the anterior margin; scutellum and metanotum dull, without

longitudinal

without

dark

than proximal

LOCUS TYPICUS:

distal

holotype M!

Neotropic region:

invisible;

fore-wings hyaline,

part of radial

(4,5:4); male genital ia: fig. 47;

Nova Teutonia

TYPICAL MATERIAL: HOSTS: unknown DISTRIBUTION:

bands;

notaulices

propodeum reticulate rugose,

keels on the posterior surface;

transversal part

granulate;

vein sl ightly tibial

longer

spurs 1,1,2.

(Brazil) and 1 paratype F! BRAZil:

in BM.

Nova Teutonia

(S.ta Catarina),

BM!

Fig.

48 - Male genitalia of Crovettia plaumanniana n. sp. paratype

3. Crovettia plaumanniana n. sp.

93

Aphelopinae

FEMALE:

fully

winged;

length 2,18 mm;

black;

antennae brown,

with segments 1-5 testaceous;

tarsi

antennae distally

testaceous;

following

proportions:

granulate,

fully

8:

6:

5:

5:

thickened; 5:

reticulate rugose;

3,5:

antennal

4:

frontal

mandibles testaceous; legs brown,

3,5:

I ine

4:

with

tibiae and

segments in

6,5;

head dull,

incomplete; occipital

complete; POL = 9; DL = 5; DOL = 8; OPL = 8; TL metanotum dull, fully reticulate rugose; notaulices

carina

=

5; scutum, scutellum, invisible; propodeum

reticulate rugose, without transversal or longitudinal keels; fore-wings hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part MALE:

fully

winged;

antennae testaceous-dark; partly brown;

(2:5);

length 2,31

tibial mm;

legs testaceous,

antennae not distally

following proportions: reticulate rugose;

8:

spurs 1,1,2.

black;

5:

frontal

5:

6:

mandibles testaceous;

with coxae and cl ubs of femora

thickened;

5,5:

5:

5:

I ine incomplete;

antenna I segments

5:

5:

in

8; head dull, fully

face without

lateral

keels;

occipital carina complete; POL = 9; OL = 6; OOL = 9; OPL = 6; TL = 6; scutum, scutellum and metanotum dull, fully reticulate rugose; notaulices invisible;

propodeum reticulate rugose,

between dorsal longitudinal

and posterior surface;

keels,

reticulate rugose;

transversal

bands;

part

genitalia:

(2:6);

LOCUS TYPICUS:

distal

HOSTS:

48;

tibial

transversal

keel

without dark

vein much shorter than

proximal

spurs 1,1,2.

(Brazil)

holotype F!

in OT;

paratype M!

in BM.

BRAZIL:

Nova Teutonia

unknown

DISTRIBUTION: OT!

fig.

a strong

fore-wings hyaline,

part of radial

Nova Teutonia

TYPICAL MATERIAL:

with

posterior surface without

Neotropic region:

(S.ta Catarina),

BM!

Cr. plaumanniana structura II y resemb les Cr. nearctica (Nearct ic

NOTES: region).

It

is named in honor of Mr.

Fritz Plaumann.

4. Crovettia brasiliana n. sp.

FEMALE:

fully

winged;

antennae brown, light;

length

antennae distally

proportions:

8:

1,75-1,81

mm;

black;

with segments 1-2 testaceous; 5:

3:

3:

thickened;

antennal

4:

3,5:

4:

3,5:

mandibles testaceous;

legs brown,

segments

3,5:

5;

with

tarsi

in following

head dull,

granulate;

frontal line incomplete; occipital carina complete; POL = 7; OL = 4; OOL 4,5; OPL = 2; TL = 2; scutum dull, granulate, reticulate rugose only near the anterior margin;

notaul ices

incomplete,

visible only

near the anterior

Clargin of the scutum; scutellum dull, granulate; metanotum shiny, smooth, .\ ithout sculpture; propodeum reticulate rugose, without transversal and :ongitudinal distal

keels;

fore-wings hyal ine,

part of radial

S::lurs 1,1,2. '!ALE: unknown

without dark transversal

vein much shorter than

proximal

part

bands;

(2:4);

tibial

94

Mem.

LOCUS TYP ICUS:

Nova Teutonia

TYPICAL MATERIAL: HOSTS:

En!.

Ins!.,

no. 37,

1984

(Brazi I)

holotype F!

in OT;

paratype F!

in OL.

MEXICO:

Yerba Buena

(Bochil,

unknown

DISTRIBUTION: OL!

Amer.

BRAZIL:

Neotropic region:

Nova Teutonia

(S.ta Catarina),

Chiapas),

OT!

5. Crovettia colombiana n. sp.

FEMALE: brown,

fully

thickened; 4,5:

4,5:

frontal ;

8;

winged;

antennal 4,5:

6;

3;

TL ;

5;

reticulate rugose; longitudinal

with

very

carina complete;

scutum dull,

granulate,

scutellum dull,

fore-wings hyal ine,

part of radial

9:

weak

6:

8:

7:

6:

5:

irregular keels;

POL; 7,5;

OL ;

4;

OOL near

reaching approximately metanotum dull,

fully

without transversal

without dark

vein much shorter than

legs

antennae distally

reticulate rugose only

granulate;

propodeum reticulate rugose,

keels;

mandibles testaceous;

notaulices almost invisible,

length of scutum;

distal

granulate,

line incomplete; occipital

OPL ;

black;

antennae black;

segments in following proportions:

head dull,

the anterior margin; 0,5

length 2,50 mm;

with fore tibiae testaceous;

transversal

proximal

or

bands;

part (4:8);

tibial

spurs 1,1,2. MALE:

unknown

LOCUS TYPICUS:

1°10'

TYPICAL MATERIAL: HOSTS: unknown DISTRIBUTION:

N,

7°5'

1°10'N,

7°5'W

SUBFAM I L Y BIAPHELOPINAE N.

SUBF.

TYPE GENUS: Biaphelopus n. fully

winged;

tibial

maxi Ilary palpi

COLOMBIA:

(Putumayo),

OT!

gen.

fore-wing

enclosed by pigmented veins; complete;

Colombia)

in OT.

Neotropic region:

2.

FEMALE:

W (Putumayo,

holotype F!

with costal

front

tarsi

and median cells clearly

not chelate;

with 5-6 segments;

labial

occipital palpi

with

carina 3 segments;

spurs 1,1,2.

MALE: unknown DIST.RIBUTION: Palaeartic HOSTS:

unknown

GENERA:

1

SPECIES: NOTES:

1

the new subfami I y Biaphelopinae structura II y

resemb les

Conganteoninae (ma in I y due to the scu I pture of head and thorax and venation of the fore-wing). In females of Biaphelopinae, however, the fore tarsi are not chelate. Differences regarding the males of Biaphelopinae (as yet unknown)

probably

wi II

concern

the radial

vein

(in females and males

95

Biaphelopinae

of

Conganteoninae the distal part is as long as or longer than the in females of Biaphelopinae the distal part is shorter than

proximal part; the proximal

part).

Biaphelopus n. gen.

TYPE SPEC I ES:

Biaphelopus masneri n. sp.

FEMALE

14,

(Figs.

with 3 segments; wing

with

head,

proximal

scutum,

49):

maxi Ilary palpi

scutellum and metanotum fully sclerites

vein forming

an

reticulate rugose; (Fig.

31

epistomal

C).

Palaearctic

1

Fig.

49 -

Female of

fore-

angle;

unknown

SPEC I ES:

palpi

teeth;

unknown

DISTRIBUTION: HOSTS:

labial

with 2 rudimentary

and distal parts of radial

suture running close to the antennal MALE:

with 5-6 segments;

mandibles quadridentate,

Biaphelopus masneri n. sp. (type).

96

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

1. Biaphelopus masneri n. sp.

FEMALE:

fully

winged;

antennae brown,

coxae basally black;

antennae distally

following proportions: reticulate rugose, complete;

=

POL

invisible;

8:

4:

10: 8:

granulate;

5;

OL

and metanotum dull, keels;

length 2,43 mm;

black;

with segments 1-2 testaceous;

=

3;

fully

8:

frontal

=

OOL

thickened; 6,5:

5:

5:

7;

OPL

=

propodeum reticulate rugose,

6;

5:

=

TL

vein shorter the right, MALE:

than

proximal

maxillary palpi 6 on

the

(4:6);

fore tarsi

left);

labial

palpi

(in

hind

8;

fully

carina

scutum,

scutellum

notaul ices

bands, distal

or transversal with costal

and

part of radial

not chelate;

tibial

spurs

the holotype 5 segments on

with 3 segments.

unknown

LOCUS TYPICUS:

Godavari

TYPICAL MATERIAL: HOSTS:

(Nepal)

holotype F!

in OT.

unknown

DISTRIBUTION: NOTE:

part

with 5 or 6 segments

head dull,

longitudinal

without dark transversal

median cells clearly enclosed by pigmented veins; 1,1,2;

7;

occipital

granulate;

without

with

antenna I segments in

I ine absent;

reticulate rugose,

fore-wing hyal ine,

mandibles testaceous; legs testaceous,

Palaearctic region:

the species

is named

3.

NEPAL:

Godavari

in honor of Mr.

SUBFAM I L Y

OT!

CONGANTEONINAE N. SUBF.

TYPE GENUS: Conganteon Benoit 1951. FEMALE: fully winged; fore-wing with costal enclosed by pigmented veins

(Katmandu),

Lubomir Masner.

(Fig.

16);

and median cells clearly

maxi Ilary palpi

with 5-6 segments;

labial palpi with 3 segments; front tarsus chelate (Fig. 16); mandibles quadridentate, with an intermediate rudimentary tooth (Fig. 22 B); chela with rudimentary claw carina complete; MALE:

fully

tibial

winged;

(Fig.

fore-wing

enclosed by pigmented veins labial

palpi

pronotal

with costal

(Fig.

with 3 segments;

intermediate rudimentary tibial

53);

tubercles present;

tooth

26);

and median cells clearly

maxillary

palpi

mandibles quadridentate, (Fig.

22 B); occipital

with 5-6 segments; with an

carina complete;

spurs 1,1,2.

DISTRIBUTION: HOSTS:

Palaearctic,

Ethiopian,

Oriental

unknown

GENERA: SPECIES:

occipital

spurs 1,1,2.

3 5

KEY TO THE GENERA OF

CONGANTEONINAE

97

Conganteoninae

FEMALES

Enlarged

claw wi th

••• • • • •• • • • . • • • • . " -

subapi cal

tooth

•••.••.••• 2.

Enlarged

claw without subapical . • . . . . . . . . . . . . . . • . . • • . . • . • • • . 1. The females of Fiorianteon

n.

(Fig.

59) . . . . . . . . . . . . . . . . .

Chelanteon n. gen. tooth

(Fig.

56) . . . . . . . . . . . . ..

Conganteon Benoit

gen.

are unknown.

MALES

Distal

part

of

radial

vein

longer

• • • • • • • • • • • • • • • • • • • • . • • . • • •• -

Distal

part

of

I'adial

1.

vein much

60) ••••••.•••••.•••••.••.••. The males of Chelanteon

Fig. 50 -

n.

3.

gen.

than

proximal

part

(Fig.

50).

Conganteon Ben 0 i t shorter

than proximal

Fiorianteon n. gen. are unknown.

Male of Conganteon nepalense n.

sp.

(type)

part

(Fig.

98

Mem.

Amer.

Ent.

Inst.,

no. 37,

1984

1. Conganteon Benoit

Conganteon Benoit 1951c: 11.

TYPE SPECIES:

Conganteon vulcanicum Benoit 1951, orig. desig.

FEMALE:

winged;

fully

segmented;

maxi Ilary palpi

mandibles quadridentate,

with 5-6 segments;

with

an

labial

palpi

3-

intermediate rudimentary

22 B); fore-wing with short and oval pterostigma (Fig. 55); vein longer than proximal part (Figs. 16, 55); enlarged claw without subapical tooth, lamellae, bristles or hairs (Fig. 53); pronotal tubercles reaching tegu lae. MALE: fully winged; maxillary palpi with 5-6 segments; labial palpi 3segmented; mandibles quadridentate, with an intermediate rudimentary tooth (Fig. 22 B); fore-wing withshort and oval pterostigma (Fig. 50); distal part of radial vein longer than proximal part (Fig. 50); occipital carina complete. DISTRIBUTION: Palaearctic, Ethiopian HOSTS: unknown SPECIES: 3 tooth

(Fig.

distal

part of radial

PALAEARCTIC SPECIES OF

One species

CONGANTEON

is known.

Conganteon nepalense n. sp.

FEMALE: MALE

unknown

(Fig.

50):

fully

tegulae brown; OOL

=

7;

OPL

winged:

legs brown;

=

4;

reticulate rugose;

TL

=

7;

reticulate rugose; longitudinal distal

proximal

carina complete;

reticulate rugose;

with central

a curve;

genitalia:

figs.

51,

holotype M!

52 A;

(Nepal)

in B.

OL

2;

invisible;

and smooth;

metanotum

without transversal

without dark transversal

without distal

35 Km N of Trisuli

TYPICAL MATERIAL:

POL = 4;

temples fully

notaul ices

area shiny

longer than proximal

membranous process,

mandibles black;

keels

posterior surface without

fore-wings hyal ine, vein

black;

reticulate rugose;

and posterior surface;

keels;

spurs 1,1,2. LOCUS TYPICUS:

1,87 mm;

propodeum reticulate rugose,

part of radial

forming

occipital

scutum fully

scutellum reticulate rugose, between dorsal

length

head fully

part;

bands;

the two parts

gonoforceps without dorsal inner pointed process;

tibial

99

Conganteoninae

HOSTS:

unknown

DISTRIBUTION: Syabrubens),

Fig.

51

Palaearctic region:

NEPAL:

35 Km N of Trisuli

(Nawakot,

B!

-

Conganteon nepalense n. sp. (type)

Male genitalia of

ETH lOP I AN SPEC I ES OF

COiv'GANTEON

Two species are known.

KEY TO THE SPECIES

FEMALES

Segment

of

fore

tarsi

I ess

than

twi ce as

. •. . . •. . . . •. •. . . . . ••. . . •. •. . . •. ••. . . •. " -

Segment

1 of

fore

tarsi

more

than

1.

twi ce as

........................................

2.

long

as

segment

townesi n. long as

4 ...

sp.

segment

4 ...

vulcanicum Benoit

100

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

MALES

Vertex

of

the head,

reticulate

rugose,

near with

the

occipi tal

longitudinal

carina,

. . . . . . . . . . . . . . . . . . . . . . . . • . . . . . . . . • . . . . •• Vertex

of

rugose,

the head,

without

near

the occipital

longitudinal

shiny,

not

keels •.••.••••..•.••

townesi n.

1.

carina,

sp. reticulate

vulcanicum Benoit

2.

keels ••••••

dull,

B

/

52 - Gonoforceps (laterally viewed) of Conganteon nepalense (A: type) and vulcanicum Benoit (B: specimen from Mt. Hoyo)

Fig. n.

sp.

1. Conganteon townesi n. sp.

FEMALE:

fully

winged;

length 2,81

anterior part of face testaceous; coxae basally black; following proportions: reticulate rugose; shiny

and with

complete;

POL

very short;

=

black;

mandibles, testaceous,

antennae distally thickened; 10:

5:

12: 9:

9:

8:

6:

6:

5:

antennal 5:;

clypeus and with

hind

segments

head dull,

posterior region of the vertex near the occipital

longitudinal 7;

OL

=

4;

scutum shiny,

scutellum shiny,

mm;

antennae fully

smooth,

keels; OOL

fully finely

=

frontal 8;

OPL

line present;

=

8;

TL

reticulate rugose; punctate,

=

9.

occipital

in

fully carina carina

pronotum hairy,

notaul ices invisible;

without sculpture among

the

Conganteon i nae

Fig. 53 -

Chela of

Conganteon townesi n. sp.

101

(type)

102

Mem.

punctures;

metanotum sh i ny,

the punctures; longitudinal distal

(Fig.

53)

Inst.,

vein

a curve;

no.

puncta te,

longer than

tooth,

without

without dark proximal

in following proportions:

15:

bristles,

1984

wi thout scu I pture among

segment 3 of fore tarsi

without subapical

37,

reticulate rugose,

fore-wings hyal ine,

part of radial segments

Ent.

strong I y

propodeum fully

keels;

parts forming tarsal

Amer.

part

(13:10);

produced

3,5:

3:

transversal

transversal

10:

the

into a 15;

and

bands; two

hook;

fore

enlarged claw

hairs or lamellae;

segment 5

of fore tarsi (Fig. 53) with 1 row of nearly 52 small lamellae extending continuously to the apex; apex with 3 long lamellae; rudimentary claw present; MALE:

tibial

fully

spurs 1,1,2.

winged;

length 2,75-2,87 mm;

anterior part of face testaceous; hind coxae basally segments dull,

antennae not distally

in following proportions:

fully

reticulate rugose;

carina shiny,

with

carina complete;

Fig.

black;

54 -

=

9;

Ol

mandibles,

10:

5:

10:

8:

thickened; 8:

8:

clypeus and

legs testaceous, 6:

6:

with

antenna I 5:

7;

head

posterior region of the vertex near occipital

longitudinal

POL

black;

antennae testaceous;

=

Male genitalia of

keels; 5;

OOl

frontal

=

8;

line present;

OPl

=

9;

occipital

TL = 10;

pronotum

Conganteon tOlvnesi n. sp. (paratype)

Conganteon i nae

103

hairy, very short; scutum shiny, fully reticulate rugose; notaulices invisible; scutellum and metanotum strongly punctate, without sculpture among

the punctures;

longitudinal

keels;

part of radial

vein

forming a curve;

propodeum reticulate rugose,

fore-wings hyal ine,

longer than proximal

genitalia:

fig.

54;

LOCUS TYP ICUS:

as in vulcanicum;

Pietermaritzburg

TYPICAL MATERIAL: M!

holotype F!

part

without

dark

transversal

transversal

(13:11);

or

bands;

the two parts

gonoforceps without proximal

membranous process and without distal (laterally viewed)

without

inner pointed process;

tibial

spurs

gonoforceps

1,1,2.

(South Africa) and 2 para types MM!

in TW;

paratype

in OL.

HOSTS:

unknown

DISTRIBUTION: NOTE:

Ethiopian region:

the species is named

SOUTH AFRICA:

in honor of Mr.

Pietermaritzburg,

Henry K.

TW!

OL!

Townes.

2. Conganteon vulcanicum Benoi t

Conganteon vulcanicus Benoit 1951c: 12.

FEMALE

(Figs.

16,

55):

fully

winged;

mandibles testaceous;

antennae brown,

and propodeum black;

legs testaceous,

black;

head hairy,

complete; = 8;

length 2,37 mm; with

head black,

segments 1-2 testaceous;

shiny,

fully

reticulate rugose;

occipital

fully

reticulate rugose;

POL = 2,5;

antenna I segments in following

Fig.

55 -

thorax

with coxae and hind femora partly

temples distinct,

OPL = 6;

with

Female of Conganteon

proportions:

9:

carina OL = 5; 5:

vulcanicum Benoit

10:

7:

OOL 7:

104

6:

Mem.

6:

6:

5,5:

8;

pronotal

Amer.

En!.

Ins!.,

tubercles reaching

no.

37,

tegulae;

1984

scutum shiny,

fully

reticulate rugose, hairy; notaulices invisible; scutellum shiny, with anterior half smooth and without sculpture; posterior half reticulate rugose;

metanotum reticulate rugose;

without keels;

fore wings hyal ine,

pterostigma short and oval,

brown;

enclosed by pigmented veins; than proximal fore tarsi

part

produced

propodeum fully

without dark

radial

costal vein

reticulate rugose,

transversal

bands;

and median cells clearly with

distal

(9:7);

the two parts forming

into a

hook;

fore tarsal

part slightly

an 2ngle;

segments

longer

segment 3 of

in following

proportions: 11: 2: 2: 3: 7; enlarged claw (Fig. 56) as long as segment 5; segment 5 of fore tarsi (Fig. 56) with a long row of 27 lamellae extending continuously very hairs;

long;

rudimentary

Fig.

MALE:

fully

antennae not

56 -

distally 3:

8:

apex with 5

length

tibial

thickened; 7:

one of these

lamellae,

bristles or

spurs 1,1,2.

1,81 mm; black; mandibles black,

antennae brown; 7:

lamellae,

tooth,

Conganteon vulcanicum Benoit (type)

Chela of

winged;

6:

the apex;

without subapical

claw present;

half testaceous-brown; proportions:

to

enlarged claw

7:

6:

legs black-brown,

antenna I segments 5:

6:

7;

with distal

with

tarsi

in following

head dull,

fully

reticulate

light;

Conganteon inae

=

4; Ol = 2; OOl = 5; OPl = 4; Tl scutellum and metanotum dull, fully

rugose; frontal I ine present: POL occipital carina complete; scutum, reticulate rugose; without

notaulices invisible;

transversal

surface without transversal

bands,

the

keels;

with costal

distal

an

vein

angle;

Volcano Muhavura

TYPICAL MATERIAL: HOSTS:

hyaline,

posterior

without dark

holotype F!

longer than proximal

part

gonoforceps without proximal inner pointed process;

tibial

spurs

(Ruanda)

in

TV.

unknown

DISTRIBUTION: ZAIRE: Hoyo,

7;

and median cells clearly enclosed by

membranous process and without distal 1,1,2. lOCUS TYP ICUS:

and posterior surface;

fore-wings

part of radial

two parts forming

=

propodeum reticulate rugose,

between dorsal

longitudinal

pigmented veins;

(6:5);

keel

105

Ethiopian region:

Bomboka TV!

Ol!

(Kyandolire,

MADAGASCAR:

RUANDA:

Mt.

Volcano Muhavura

Ruwenzori,

Ankaratra,

Albert Nat.

(Burambi),

Park),

TV!

TV!

Mt.

P!

2. Chelanteon n. gen.

TYPE SPEC I ES:

Chelanteon richardsi n. sp.

FEMALE

57,58):

(Figs.

fully

winged;

maxillary palpi

with 5 segments;

labial palpi with 3 segments; mandibles quadridentate, with an intermediate rudimentary tooth (Fig. 22 B); fore-wing with short and oval pterostigma

(Figs.

part

57,

(Figs.

57,

58);

bristles or hairs (Fig. MALE:

58);

distal

part of radial

enlarged claw

vein

with subapical

longer than proximal

tooth,

without

lamellae,

59); pronotal tubercles reaching tegulae.

unknown

DISTRIBUTION: HOSTS:

Oriental

unknown

1

SPECIES:

Chelanteon richardsi n. sp.

FEMALE

(Figs.

57,

58):

fully

winged;

anterior part of face (particularly

length

2,81

along orbits),

mm;

head black,

with

clypeus and mandibles

yellow; antennae brown, with segments 1-2 yellow; thorax, propodeum, petiole and abdomen black; legs yellow, with hind coxae, hind clubs of femora

and hind

and shiny,

two areas on where

brown;

the sides of the frontal

the surface

temples distinct, sculpture;

tibiae partly

tegulae yellow;

anterior part of face hairy,

POL

=

I ine

is smooth and finely reticulate rugose, 5;

Ol

=

2;

OOl

=

fully

head slightly

reticulate rugose,

swollen

except for

in front of the anterior ocellus,

punctate;

occipital

carina complete;

except for a wide smooth area without OPl;

antenna I segments

in following

106

Mem.

Fig.

proportions: with

57 -

8:

transversal

dull,

fully

finely nearly

brown; radial

14:

10:

keels;

10:

Ins!.,

no.

37,

1984

10:

7:

6:

6:

8;

pronotum rugose,

tubercles reaching

notaulices invisible;

the tegulae;

metanotum shiny,

reticulate rugose, transversal

scutum

fully

smooth,

scutellum

reticulate rugose;

hairy,

without keels;

bands;

pterostigma short and oval,

and median cells clearly enclosed by

vein curvilinear,

hairy,

scutellum shiny,

with some areolae near the posterior margin;

without dark costal

En!.

Chelanteon richardsi n. sp. (type)

pronotal

long as metanotum;

propodeum fully hyaline,

Female of

reticulate rugose;

punctate, as

4:

Amer.

with distal

part

fore-wings

pigmented veins;

longer than proximal

part;

Conganteon i nae

segmen t 3 of fore

tars i produced

following proportions:

Fig.

shorter than

lamellae,

the apex;

8:

hook;

59 -

(12:13);

segment 5 of fore tarsi

enlarged claw

Dalat

with a

rudimentary

Chela of

fore tarsa I segments

13; enlarged claw

(Fig.

Chelanteon richardsi n. sp.

present;

(Viet Nam)

in slightly

(type)

(Fig.

to the apex;

subapical

claw

59)

59)

with

a

apex with 5

tooth and without tibial

spurs 1,1,2.

Chelanteon richardsi n. sp. (type)

unknown

_OCUS TYPICUS:

into a

4:

lamellae extending continuously

bristles or hairs;

Fig.

\1ALE:

3,5:

Female of

segment 5

long row of 44 lamellae at

58 -

17:

107

108

Mem.

TYPICAL MATERIAL: HOSTS:

holotype F!

En!.

Inst.,

no.

37,

1984

in B

unknown

DISTRIBUTION: NOTE:

Amer.

Oriental

the species

region:

is named

VIET NAM:

Dalat,

B!

in honor of Mr. Owain W.

Richards.

3. Fiorianteon n. gen.

Fiorianteon junonium n. sp.

TYPE SPEC I ES: FEMALE:

unknown

Fig.

MALE labial

(Figs. palpi

26,

60 -

60):

Male of

fully

pterostigma proximal

winged;

with 3 segments;

intermediate rudimentary (Figs.

part

26,

(Figs.

tooth

60); 26,

Fiorianteon junonium n. sp. (type)

(Fig.

distal

60);

maxillary palpi

with 6 segments;

mandibles quadridentate, 22 B);

fore-wing

part of radial

occipital

with an with short and oval

vein much shorter than

carina complete.

Conganteon i nae

109

DISTRIBUTION: Palaearctic HOSTS: unknown SPECIES: NOTES:

1

the genus

is named

in honor of the

late Mr. Giorgio Fiori.

Fiorianteon junonium n.

FEMALE: MALE black;

unknown

(Figs.

c I ypeus

sp.

26,

60):

testaceous;

fully

winged;

antennae and

antennae not distally

proportions: punctate,

Fig.

11:

5:

18:

14:

thickened;

13:

11:

without sculpture among

61

-

length 3,81

Male genitalia of

mm;

legs testaceous, 9:

black;

antenna I segments 9:

8:

10;

the punctures;

mandibles and

with hind coxae basa II y in following

head shiny, frontal

Fiorianteon junonium

n.

strongly

I ine complete;

sp.

(type)

110

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

occipital carina complete; POL = 6; OL = 4; OOL = 11; OPL = 6; TL = 12; scutum shiny, very strongly punctate, without sculpture among the punctures; scutum;

notaulices incomplete,

scutellum punctate,

metanotum rugose, keel

between

longitudinal distal

du II;

dorsal keels;

genital ia:

fig.

wi thou t

61;

without dark

much shorter than

gonoforceps without

transversal

proximal

dorsal

inner pointed process;

Yamada

transversa I

posterior surface without part

and median cells clearly enclosed by

TYPICAL MATERIAL: HOSTS:

vein

length of

the punctures;

propodeum ret i cu I ate rugose,

fore-wings hyal ine,

and without distal LOCUS TYPICUS:

approximately 0,5

and posterior surface;

part of radial

wings with costal

reaching

without sculpture among

fore-

pigmented veins;

proximal

tibial

bands;

(4: 10);

membranous process

spurs 1,1,2.

(Japan)

holotype M!

in OT.

unknow n

DISTRIBUTION:

Palaearctic region:

4.

JAPAN:

Yamada

(Nagano),

OT!

ANTHONINAE R. C. L. PERKINS

SUBFAMILY

Dryininae partim Kieffer in Kieffer et Marshall 1905: 130. Anteoninae partim Kieffer in Kieffer et Marshall 1906: 510. Anteonini R. C. L. Perkins 1912: 7. Anteonini Kieffer 1914b: 137. Anteonini Richards 1939: 190. Anteonini Muesebeck and Walkley 1951: 1040. Anteoninae J. F. Perkins 1976: 13. Anteoninae partim Krombei n 1979: 1240.

TYPE GENUS: Anteon Jurine 1807. FEMALE: fully winged, brachypterous or micropterous; winged species)

with costal,

by pigmented veins palpi

(Fig.

with 3 segments;

rudimentary (Fig.

62);

claw

(Fig.

62);

anterior one to posterior MALE:

fully

winged;

labial

teeth progressing carina complete; 29);

tibial

enlarged claw

(Fig.

5);

fore-wing

with

palpi

with

pronotal

teeth

(Fig.

median 29);

labial

without

and

lamellae

larger from

tibial

occipital spurs 1,1,2.

and submedian cells

maxi Ilary palpi

with 6

mandibles quadridentate,

larger from anterior one to posterior fore-wing

(in fully

chela

tubercles present; long hairs;

with costal,

3 segments;

without

18);

teeth progressing

tufts of

pigmented veins

with 6 segments;

(Fig.

(Fig.

5);

with

occipital

with metacarpus shorter than pterostigma

spurs 1,1,2.

DISTRIBUTION: HOSTS:

maxi Ilary palpi tarsus chelate

antennae without

enclosed by

segments;

18); front

fore-wing

and submedian cells clearly enclosed

mandibles quadridentate,

carina complete; clearly

median

worldwide

Cicadellidae (i nc Iud i ng I diocerinae, Macropsinae and

(Fig.

Anteoninae

occasiona II Y Typhlocybinae) GENERA: SPEC I ES:

5 239

Fig.

62 -

Chela of Metanteon

fuscum n. sp. (type)

111

112

Mem.

Amer.

Ent.

Ins!.,

no.

37,

1984

ANTEONINAE

KEY TO THE GENERA OF

FEMALES

62) •..•..•.........••..•••.••......••

of at least 4 bristles 1. Metanteon n. gen.

Inner

1-2 bristles

Inner

2

side of

enlarged

side of

claw with

enlarged

a

claw with

row

only

(Fig.

or peg-like hairs

located proximally (Fig. 63) •.•••.•••••••••.•••••••••.••••••..••••• 2 Inner side o-f enlarged claw wi th a proximal prominence bearing 1

b r i s tie (F i g. 64) •••••••••.•••••••••••••••••••••.••••.••••.••.••••• 3 Inner side of enlarged claw wi th a proximal prominence not bearing bristles (Fig. 63); 1-2 short bristles or peg-like hairs located further distally than the proximal prominence (Fig. 63) •••••••.••.• 6 3

Species brachypterous

-

Species

fully

winged

4 Propodeum wi th posterior -

a

strong

5 Fore-wi ng proximal

transversal

keel

a

st rong

4.

transversa I

surface •••••••••••••.•.•••• wi th d i sta I part

(Fig.

part

158);

case propodeum without -

162) .••••••••••••••••••••••••••••••••.• 4 158) ••••••••••.•••..••••.••••.•••••••.•• 5

surface . • • • • • • • • . • • . • . • . • . •

Propodeum wi thou t posterior

(Fig. (Fig.

a

of

rad i a I

vei n

strong

and

Anteon Jurine

kee I 3.

occasi onally

between dorsal

between dorsa I

and

Lonchodryinus Kieffer as

long as

sl ight Iy

transversal

or

longer

shorter,

keel

but

between

in

dorsal

than that and

posterior surface ••••.••••••••••••••• 3. Lonchodryinus Kieffer Fore-wing wi th di stal part of radial vein much shorter than proximal or

longer

strong

part

(Fig.

178),

than proximal

transversal

keel

occasionally

part,

but

between

in

sl ightly

that

dorsal

shorter,

as

long as,

case propodeum with a

and posterior

surface ••.•••••

.••••.•••.••.•..•.•••...••••.•....••• 4. Anteon Jurine 6 Micropterous

species with

thoracic

structure

Gonatopodinae (pronotum remarkably elongated; between pro- and metanotum; propodec:m humped)(Figs. -

Fully

winged

species with

Anteoninae (pronotum not the pronotum;

metanotum

385,386) •••• thoracic

metanotum visible;

invisible; 5.

of stalk

+

to

that

scutum broader

propodeum not 2.

that

metathorax

similar

elongated;

.•••••••••••••••••••••••••.••••••••••

to

Prioranteon n. gen.

structure

remarkably

similar

scutum forming

humped)(Fig.

of than 67) •••••

Deinodryinus R.C.L. Perkins

MALES

Pterostigma

four

or more

times as

long as broad

• • • • • • • • • • • • • • • • • . • • . • •• • . • • • • • • • • • •• -

Pterostigma

less

2 Fore-wing with

than

four

times as

distal

part

of

radial

2.

(see fig.

176) ••..•

Deinodryinus R.C.L. Perk ins

long as broad vein much

(see

shorter

fig.

176) ••• 2

than proximal

Anteoninae

113

.,/

Fig.

part

(Figs.

longer

Chela of Deinodryinus atriventris

63 -

29,

180);

than proximal

occasionally part,

but

in

slightly that

(Cresson)

shorter,

case with a

as

long as,

or

strong

transversal

-

keel between dorsal and posterior surface . . . . . . . . . . . . .. . . . . . • . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4. Anteon Jurine Fore-wing wi th distal part of radial vein as long as or longer than proximal part (Fig. 160); occasi onally sl ight Iy shorter, but in that case propodeum without a strong transversal keel between dorsal and posterior surface . . . . . . . . . . . . . . . . . . . . 3. Lonchodryinus Kieffer

The males of Metanteon

n.

gen.

and Prioranteon

n.

gen.

are unknown.

1. Metanteon n. gen.

TYPE SPEC I ES: Metanteon FEMALE palpi

(Figs.

18,

65):

with 3 segments;

proximal

part

(Fig.

aerias (Wa I ker 1839). fully

winged;

fore-wing

65);

maxi Ilary palpi

with distal

with 6 segments;

part of radial

vein

labial

longer than

inner side of enlarged claw with a row of at

least 4

114

Mem.

Fig.

bristles (Fig. 62); MALE: unknown D I STR I BUT I ON: HOSTS:

64 -

Amer.

Ent.

Inst.,

no.

37,

1984

Chela of Anteon jurineanum Latreille

occipital

carina complete.

Neotrop ic

unknown

SPECIES:

2

NEOTROP I C SPEC I ES OF NETANTEON

Two species are known.

KEY TO THE SPECIES

FEMALES

Head and pronotum fully -

testaceous-reddish . . . . . . . . . . . . . . . . . . . . . . ..

•••••••••••••••••••••••••••••••••••••

1.

Head and pronotum partly black . . . . . . .

2.

MALES

Unknown

aerias (Walker) fuscum n. sp.

Anteoninae

115

1. Metanteon aerias (Walker) n. comb.

Dryinus aerias Walker 1839: 92 Anteon aerias (Walker): Dalla Torre 1898: 537.

FEMALE

(Figs.

18, 65): fully winged;

length 2,81-3,00 mm;

pronotum and scutum

testaceous-reddish

propectus,

metanotum,

Fig.

scutellum,

(last

mesopleura,

head,

antenna I segment metapleura,

antennae,

slightly dark);

propodeum,

petiole,

65 - Female of Metanteon aerias (Walker) from EI Coigo (Chile)

abdomen black;

legs testaceou5,

with hind coxae proximally

brown;

tegulae

116

Mem.

testaceous; 7: = 7;

8;

antenna I segments

head dull,

OPl = 5;

with a

weak

reaching

smooth,

Tl = 5;

En!.

Ins!.,

frontal

1984

9: 5: 11: 9: 9: 8: 7: 7:

line absent;

carina complete;

impression,

scutum,

37,

no.

in following proportions:

granulate;

occipital

transversal

tegulae;

punctate, reaching

Amer.

stronger on

POl= 3;

pronotum dull, the sides;

scutellum and metanotum shiny,

without sculpture among punctures; approximately 0,65

without transversa I keel

pronotal smooth,

notaulices distinct,

length of scutum;

propodeum fully

OOl

tubercles

weakly incomplete,

reticulate

rugose,

du II,

without rugose;

longitudinal keels on the posterior surface; metapleura reticulate mesopleura smooth, weakly granulated; fore-wings hyaline, with distal

part of radial parts formi ng

vein slightly a curve;

between dorsa I and

Ol = 2;

granulate,

longer than proximal

pterost i gma

10: 2: 3:

part;

long and narrow;

posterior surface and

distal

fore

and proximal

tarsa I segments

with

lamellae in addition

row

of 6 short bristles MALE: unknown LOCUS TYPICUS:

11:

in

17; segment 5 of fore tarsi (Fig. 66) some bristles and a row of nearly 26 lamellae-like bristles; apex with 3

following proportions:

to some bristles;

(Fig.

66);

inner side of enlarged claw

Chile

TYPICAL MATERIAL:

holotype F!

in BM.

HOSTS: unknown DISTRIBUTION: Neotropic region:

CHILE:

(Cord.

Nahuelbuta,

Arauco),

OTt

Chile,

EI

Coigo

fully

winged;

length 3,00-3,87 mm;

half of face yellow-testaceous; side somet imes testaceous; wide black spot on

(Curico),

OT!

OL!

n. sp.

head fully

black or with

anterior

malar space and anterior half of the ventral

dorsa l s i de of head somet i mes testaceous,

vertex;

Pichinahuel

BM!

2. Metanteon fuscum

FEMALE:

with a

spurs 1,1,2.

tibial

mandibles and clypeus

with

antennae brown, with segments 1-2 or only 1 testaceous; prothorax black, the posterior margin of the pronotum I ight or testaceous; mesothorax, metathorax, basally distally

propodeum and abdomen black;

black

occipital

thickened;

reaching

antennal

punctate and granulc:te;

POL = 4;

OL = 3;

granulate and rugose,

tegulae;

scutum,

with

black;

with

frontal

OOl = 8;

OPL = 7;

transversal

keels;

scutellum and metanotum shiny,

with

hind coxae antennae

10: 7:

segments in following proportions:

10; head dull,

carina complete;

pronotum dull,

legs testaceous,

and sometimes with clubs of femora partly

12: 11: 9: 7: 7:

a

testaceous or black;

15:

15:

line absent; TL = 7;

pronotal

tubercles

punctate and

granulate; notaulices incomplete, reaching approximately 0,65 length of scutum; propodeum dull, reticulate rugose, without transversal and longitudinal keels; fore-wings hyal ine, without dark transversal bands; distal part of radial longer than proximal part (17:10); fore tarsal segments in following

vein

15: 3: 5: 16: 24; inner part of enlarged claw (Fig. 62) with a segment 5 of fore tarsi (Fig. 62) with 2 rows of nearly 55 lamellae and a group of 5 lamellae at apex; tibial spurs 1,1,2.

proportions:

row of 6 bristles;

Anteoninae

Fig.

MALE:

66 -

Chela of Metanteon

117

aerias (Walker)(type)

unknown

LOCUS TYPICUS:

Cobquecura

TYPICAL MATERIAL: TW; 4 paratypes FF! HOSTS: unknown JISTRIBUTION:

(Chile)

holotype F!

and 3 paratypes FF!

in OT;

1 paratype F!

in OL.

Neotropic region:

CHILE:

Cobquecura

(Nuble),

OT!

Butamalal

in

118

Mem.

(Araueo),

Ol!

Piehinahuel

las Cabras

(Cord.

Amer.

Ent.

Ins!.,

(Chillan Vuleain,

Nahuelbuta,

Araueo),

aT!

no.

37,

1984

Nuble), EI

aT!

TW!

Ol!

Coigo (Curieo),

Ol!

2. Deinodryinus R.C.l. Perkins

Fig.

67 -

Female of

Deinodryinus paradoxus R. C. l . Perkins from Nogales

(Arizona)

Anteoninae

119

Deinodryinus R.C.L.Perkins 1907: 45. Trisanteon Kieffer 1913b: 300 (n. syn.); type species Trisanteon hirticornis (Kieffer 1911b), monotypic and orig. desig. Deinodryinus R.C.L. Perkins: Kieffer 1914b: 138. Trisanteon Kieffer: Kieffer 1914b: 196. Electrodryinus Ponomarenko 19 75c: 126 (n. syn.); type speci es Electrodryinus areolatus Ponomarenko 1975c, monotypic.

Fig.

68 -

Female of Deinodryinus paradoxus R.

C.

L.

Perkins from Nogales

(Arizona)

TYPE SPECIES: Deinodryinus paradoxus R.C.L. Muesebeck and Walkley 1951. FEMALE labial

(Figs. palpi

67,

68):

fully

with 3 segments;

winged;

Perkins 1907,

maxi Ilary palpi

fore-wing

longer or slightly shorter than proximal

with part

distal (Fig.

desig.

by

with 6 segments;

part of radial

vein

67)(the length of the

distal and proximal parts of the radial vein in females is not a specific characteristic: in laboratory bred females the length is variable in the same species);

inner side of enlarged claw with a proximal

bearing bristles

(Fig. 63);

further distally than complete;

the proximal

prominence

(Fig.

63);

elongated;

scutum broader than pronotum;

propodeum not humped);

to occipital

carina (pronotum

metanotum

vertex of head frequently

oblique keels connecting posterior ocelli

located

occipital

thoracic structure not simi lar to that of Gonatopodinae

not remarkab I y viSible;

prominence not

1-2 short bristles or peg-like hairs

carina.

with

two strong

120

MALE

Mem.

(Figs.

69,

labial

palpi

broad

(see fig.

slightly

70):

Amer.

fully

winged;

with 3 segments; 176);

En!.

wrapping

37,

maxillary palpi

fore-wing with distal

shorter than proximal branch

no.

HOSTS:

part

penis

(Fig.

(Fig. (Fig.

69);

79); 83);

Ethiopian,

vein

long as longer or

antenna I hairs usually much

gonoforceps usually

with

vertex of head frequently

Oriental,

to occipital

Nearctic,

an with

carina.

Neotropic.

Cicadellidae

SPEC I ES:

Fig.

Palaearctic,

with 6 segments;

part of radial

two strong oblique keels connecting posterior ocelli DISTRIBUTION:

1984

pterostigma four or more times as

longer than breadth of segments internal

Inst.,

65

69 -

Male of

Deinodryinus paradoxus R. C. L. Perkins from Nogales (Arizona)

Anteoninae

Fig.

70 -

121

Deinodryinus paradoxus R. C. L. Perkins from Nogales

Male of

(Arizona)

PALAEARCT I C SPEC I ES OF

One species

(fossil)

DEINODRYINUS

is known.

Deinodryinus areolatus (Ponomarenko) n. comb.

Electrodryinus areolatus Ponomarenko 1975c: 128

FEMALE

(Fig.

71):

fully

winged;

anterior part of face brown; spot;

mandibles testaceous,

antennae testaceous; testaceous-brown;

with

thorax, 17:

(length of head

bidentate;

30:

distally 22:

18:

less than

15:

three

from occipital

157:63;

antennal

clypeus

with

11:

brown

surface brown; black;

thickened;

antenna I hairs very

view

the apex of the mandibles): emarginated,

10:

antennae

in dorsal

head black, with a central

propodeum and abdomen

densely hairy;

breadth of segments;

testaceous,

teeth and proximal

antennae slightly

in following proportions: 10-segmented,

length 4,48 mm;

clypeus

11:

short,

times as

legs

antennal 10:

13;

segments antennae

much shorter than long

carina behind

as head the ocelli

to

with anterior margin weakly

sclerites well

separated from epistomal

122

Mem.

Fig.

suture;

Ent.

occipital

Ins!.,

no.

37,1984

Deinodryinus areolatus (Ponomarenko)(type)

Female of

teeth progressing

eyes apparently bare;

protruding; carina;

-

mandibles with 4

posterior; sulcus;

71

Amer.

carina complete;

posterior ocelli

larger from anterior one to

malar space with a temples prominent;

not easily

visible,

well-marked subocular eyes normally

not reaching

sculpture of vertex and frons not easily

visible;

the occipital

pronotum not

crossed by transversal impressions; thoracic structure simi lar to that of living Deinodryinus; pronotal tubercles reaching tegulae; posterior margin of pronotum visible;

longer than

scutum shiny,

posteriorly separated; than pronotum very

broad;

(20: 15);

dorsal

anterior margin;

finely punctate,

sculpture of pronotum not easily

with

notaul ices complete and

scutellum and metanotum

invisible;

propodeum reticulate rugose;

surface of propodeum nearly

as

scutum

longer

areolae of propodeum long

as posterior

Anteoninae

surface;

123

posterior surface not easi Iy visible;

posterior surface of

propodeum very steep; fore-wings hyaline, without dark transversal bands, with the usual venation of Anteoninae; pterostigma narrow, more than four times as

long as broad

(40:8);

radial

nearly as long as proximal part wings with cells);

the usual

three cells enclosed

hind wings hyal ine,

typical of Deinodryinus; slender stalk,

slightly

cell

(18:17);

open;

distal

radial (costal,

without dark bands; (10:6);

vein

fore-

median and sUbmedian shape of wings and

fore trochanters not slender, longer than broad

part of radial

vein not s-shaped;

legs

without proximal

segment 3 of fore

tarsi

produced into a hook; rudimentary claw absent; enlarged claw much longer than arolium (32:7), without subapical tooth and other teeth; segment 5 of fore tarsi visible;

nearly as long as enlarged claw,

segment 1 of fore

tarsi

nearly as

abdomen without a distinct petiole; labial

palpi

MALE:

unknown

with 3 segments;

LOCUS TYPICUS:

tibial

with

lamellae not easily

long as segment 4

maxi Ilary palpi

(15:17);

with 6 segments;

spurs 1,1,2.

Baltic amber

TYPICAL MATERIAL:

holotype F!

in MPI.

HOSTS: unknown DISTRIBUTION: only known from Baltic amber. NOTES:

this species seems

to have the structure of the

living Deinodryinus,

and therefore belongs to the genus Deinodryinus. There is no known species of living Deinodryinus from the Palaearctic region.

ETH lOP I AN SPEC I ES OF DEINODRYINUS

Five species are known.

KEY TO THE SPECIES

FEMALES

Posterior segment

surface

(Fig.

fore

of

pronotum with

tarsi

bearing

sharp

lateral

lamellae of

margins;

same 1. insulanus (Benoit) - Posterior surface of pronotum with rounded lateral margins .••.• 2 2 Antennal segment 3 nearly twice as long as segment 2; segment 5 of fore tarsi bearing small lamellae, some longer than others (Fig. 73) •••••••••••••••••••••••••••• 2. madecassus (Benoit) - Antennal segment 3 more than three times as long as 2 •••••••••• 3 3 Segment 5 of fore tarsi bearing lamellae of approximately same length

5 of

72) ••••••••...•••.•••..•

approximately

124

Mem.

Amer.

Ent.

Ins!.,

no.

37,

1984

length (Fig. 76); species mostly black . . . . . . . . . . . . . . . . . . . . . . ... . • • • • • • • • • . • • • • • • • • . • • • • • • • • • • • • • • •• 4. benoiti n. sp. -

Segment

5 of

others

(Fig.

fore

tarsi

75)

species mostly

bearing

smal

lamellae,

some

longer

than

reddish-testaceous • • . . . • . . • • • • •

3. madagascariensis (Ben 0 i t )

The female of

D. monticolus n. sp. is unknown.

MALES

Distal

part

of

reticulate

radial

rugose,

vein

punctures.. . . . . . . . -

Distal

part

of

ret i cu I ate

longer

strongly

than

punctate,

. ...............

radial

vein much

proximal without 2.

shorter

rugose ••••••••••••••••••••

part;

head not

sculpture among

madecassus (Benoi t) than proximal

5.

part

monticolus n.

head

sp.

The males of D. insulanus (Benoit), D. benoiti n. sp. and D. madagascariensis (Benoi t) are unknown.

1. Deinodryinus insulanus (Benoit) n. comb.

Prenanteon insulanum Benoi t 1953b: 419. FEMALE:

fully

winged;

length 5,00 mm;

triangle and petiole black,

reddish-testaceous,

abdomen partly

brown,

with ocellar

last 3 antenna I

segments dark; antennal segments in following proportions: 18: 5: 8: 8: 9: 9: 9: 11; head shiny, strongly reticulate rugose; frontal

18: 10: line

present;

=

=

3;

ocellar triangle convex,

=

OOL

shiny,

10;

OPL

strongly

=

10;

TL

=

with keels among

8;

occipital

punctate and with

impression separating an

transversal

anterior collar and a

margins of posterior surface sharp; scutum shiny, punctures;

scutum;

smooth,

notaul ices

incomplete,

scutellum shiny,

the punctures; without a

smooth,

metanotum sh i ny,

margin very convex and transversal

surface with

a central

raised

keel

distal part of radial

approximately 0,65

OL

lateral tegulae; the

length of

without sculpture among

without sculpture,

with

anterior

propodeum reticulate rugose,

and posterior surface;

reticulate rugose, with

5;

transversal

tubercles reaching

into a carina;

vein

with a

posterior disc;

weakly punctate, smooth,

between dorsal

area r,ot

keels,

POL

pronotum

without sculpture among

reaching

mesopleura and metapleura rugose, darkened;

pronotal

strongly punctate,

the ocelli;

carina complete;

smooth,

weak keels;

posterior

without keels;

fore-wings fully

longer than proximal

part

(11:7);

Anteoninae

fore tarsal

segments

claw

72)

(Fig.

prominence;

with

in following

segment 5 of fore tarsi same

segment 2 produced

Fig.

MALE:

proportions:

72 -

10:

4:

6:

1 bristle located further distally

lamellae of approximately lamellae:

125

Chela of

(Fig.

length;

into a

72)

26;

enlarged

the proximal

with 2-3 rows of nearly 27

apex with

hook;

11:

than

tibial

a

group of nearly 6

spurs 1,1,2.

Deinodryinus insulanus (Benoit)(type)

unknown

LOCUS TYP I CUS:

I vondro

TYPICAL MATERIAL: HOSTS: unknown

(Madagascar)

holotype F!

DISTRIBUTION: Ethiopian region: (Benoit 1953b).

in P;

1 para type F

MADAGASCAR:

in TV.

Ivondro,

P!

La Mandraka

2. Deinodryinus madecassus (Benoit) n. comb.

Hirtanteon madecassum Benoi t 1953b: 422. FEMALE: fully winged; length 3-5 mm; reddish-dark; mandibles testaceous, with teeth brown; antennae testaceous, with segments 7-10 brown; propodeum black; abdomen brown; an tenna I segments in follow i ng proportions: 11: 5: 11: 6: 6: 6: 6: 6: 5: 7; head shiny, strongly punctate, = 7;

especially anterior part,

OL = 5;

OOL = 9;

pronotum shiny, punctures,

smooth,

with a

OPL = 7; strongly

transversa I

without sculpture among

TL = 7;

occipital

punctate,

impress i on

punctures;

POL

carina complete;

without sculpture among

between

the anteri or coil ar and

126

Mem.

Amer.

En!.

Ins!.,

no.

37,1984

the posterior disc; lateral margins of posterior surface rounded; pronotal tubercles reaching tegulae; scutum shiny, smooth, finely punctate, without sculture among punctures; 0,65

length of scutum;

puncta te, convex

incomplete,

without scu I pture among punctures;

and raised

transversal with

notaul ices

keel

a central

reaching

scutellum and metanotum shiny,

into a carina;

between

dorsal

approximately

smooth,

finely

anterior marg i n of metanotum

propodeum reticulate rugose,

and posterior surface;

smooth area not reticulate rugose;

without a

posterior surface

mesopleura shiny,

strongly punctate, with a tuft of hairs on the anterior third; metapleura partly smooth, without sculpture; fore-wings with 2 dark transversal bands, one on the basal cells and another beneath the pterostigma; distal part of radial vein longer than proximal part (14:8); fore tarsal segments in following proportions: bristle

8:

3: 4:

located further distally

fore tarsi

(Fig.

73)

8:

than

with 2 rows of

19;

enlarged claw

the proximal lamellae

(about 8

shorter);

apex with a group of nearly 7 lamellae;

produced

into a

Fig.

MALE

hook;

73 -

(supposed;

Chela of

it

mandibles testaceous,

D. madecassus

to as fully

winged;

teeth brown;

tegulae testaceous;

and hind tibiae testaceous;

with

1

segment 5 of

longer and 21

no segment of fore

Deinodryinus madecassus

breeding): with

73)

tarsi

spurs 1,1,2.

was referred

was not obtained by testaceous;

tibial

(Fig.

prominence;

(Benoit) (type)

by

antennae brown,

abdomen brown;

Benoi t

length 2,37 mm;

1953b;

it

black;

with segments 1-2

legs brown,

antennae with hairs at most as

with

tarsi,

long as

mid

Anteoninae

127

general breadth of segments; head shiny, strongly punctate, without sculpture among punctures, hairy; POL; 8; OL ; 4; OOL ; 6; OPL ; ;

4;

occipital

carina complete;

scutum shiny,

without sculpture among punctures; approximate:y 0,50 smooth, fully

finely

punctate,

reticulate rugose,

Fig.

74 -

notaul ices

length of scutum;

smooth,

incomplete,

Male genital ia of

reaching

scutellum and metanotum shiny,

without sculpture among dull;

5;

finely punctate,

punctures,

propodeum

mesopleura and metapleura smooth,

without

Deinodryinus madecassus (Benoit) (allotype)

TL

128

Mem.

scu Ipture;

propodeum without a

posterior surface; slightly without

Amer.

Ins!.,

proximal

wrapping penis;

tibial

LOCUS TYPICUS:

Bekily

TYPICAL MATERIAL:

no.

37,

transversa I keel

fore-wings hyaline,

longer than proximal dorsal

En!.

part

with

between dorsa I and

distal

(10:9);

1984

part of radial

genitalia:

membranous process,

fig.

74;

with an apical

vein

gonoforceps branch

spurs 1,1,2. (Madagascar)

holotype F!,

allotype M!

and 2 para types FF!

in P;

2

paratypes FF! in TV. HOSTS: unknown DISTRIBUTION:

Ethiopian

region:

MADAGASCAR:

Bekily,

P!

TV!

3. Deinodryinus madagascariensis

(Benoi t)

n.

comb.

Hirtanteon madagascariensis

FEMALE:

fully

winged;

length 5,0 mm;

with segments 6-10 brown; segments shiny,

1953b: 423.

testaceous;

petiole black;

in following proportions:

with face fully

Benoi t

12:

antennae testaceous,

abdomen partly

7:

reticulate rugose;

25:

10:

8:

8:

vertex and

brown;

8:

8:

8:

antennal 11;

head

temples not reticulate

rugose, but strongly punctate; frontal I ine present; occipital carina complete; ocellar triangle convex; POL = 5; Ol = 4; OOl = 13; OPL = 12; Tl = 8;

pronotum shiny,

punctures;

smooth strongly punctate,

pronotum with

a strong

transversa I

anterior collar and the posterior disc; rounded;

pronotal

strong I y

punctate,

incomplete,

tubercles reaching

approximately 0,5

smooth,

scutum shiny,

smooth,

notau I ices

length of scutum;

without sculpture;

the

margins of posterior surface

without scu I pture among punctures;

reaching

metanotum shiny,

lateral

tegulae;

without sculpture among

impress ion between

scutellum and

anterior margin of the

metanotum very convex and raised into a carina; propodeum shiny, reticulate rugose, without a transversal keel between dorsal and posterior surface; posterior surface with a central smooth area not reticulate rugose; mesopleura not reticulate rugose, strongly punctate, with the anterior third bearing a rugose, on

strong I y

dense tuft of hairs;

punctate;

the basa I cell s and another beneath

segments

in following proportions:

75)

1 bristle

with

metapleura not reticulate

fore-w i ngs with 2 dark

transversa I bands,

the pterost i gma;

11: 4:

located further distally

7:

16:

than

30;

one

fore tarsa I

enlarged claw

the proximal

(Fig.

prominence;

segment 5 of fore tarsi (Fig. 75) with 2 rows of lamellae, many small (nearly 30), some much longer (nearly 9); apex with a group of nearly 9 lamellae

(one of these very

long);

no segment of fore tarsi

a hook; tibial spurs 1,1,2. MALE: unknown lOCUS TYPICUS:

Tananarive

TYPICAL MATERIAL:

(Madagascar)

holotype F!

in TV.

produced

into

129

Anteoninae

Fig.

HOSTS:

Chel a

75 -

Deinodryinus madagascariensis (Benoi t) (type)

of

unknown

DISTRIBUTION:

Ethiopian region:

MADAGASCAR:

Tananarive,

TV!

4. Deinodryinus benoiti n. sp.

FEMALE:

fully

the face

(mainly

testaceous,

winged;

length 6,62 mm;

along orbits),

head black,

with anterior part of

clypeus and mandibles testaceous;

with segments 9-10 brown;

margins of pronotum and apex of propodeum testaceous; abdomen segment

with segment 1 mostly

reddish-testaceous;

1 and other segments black;

testaceous,

with distal

tegulae

posterior part of

testaceous;

legs reddish-

region of fore and hind clubs of femora brown;

anterior region

among punctures; frontal

pronotum shiny,

smooth,

punctures,

lateral

with

of face strongly

occipital

I ine incomplete;

with

petiole black;

antennae not distally thickened; antenna I segments in following proportions: 18: 7: 30: 19: 16: 14: 13: 12: 12: 17; head smooth, punctate;

antennae

thorax and propodeum black,

POL

punctate,

carina complete;

=

finely

6;

OL

=

5;

punctate,

frons without

OOL

=

12;

shiny,

without sculpture OPL

lateral

=

13;

TL

keels;

=

9;

without sculpture among

margins of posterior surface rounded;

pronotal

130

Mem.

Fig.

76 -

tubercles reaching smooth,

finely

incomplete,

Amer.

Chela of

tegulae;

punctate,

reaching

reticulate rugose, region,

one on

Inst.,

no.

37,

1984

Deinodryinus benoiti n. sp. (type)

scutum,

scutellum and metanotum shiny,

without sculpture among punctures;

approximately 0,65

without

transversal

fore-wings with 3 dark

surface;

Ent.

the basal

length of scutum;

keel

between

transversal

dorsal

notaul ices

propodeum fully and posterior

bands,

on the proximal

cells and another beneath

the pterostigma;

part of radial vein longer than proximal part (18:13); fore tarsal segments in proportions: 15: 5: 10: 32: 48; enlarged claw (Fig. 76) bristle

located further distally

fore tarsi same

(Fig.

length;

MALE:

76)

Perinet

TYPICAL MATERIAL:

2 rows of about 90

a

prominence;

lamellae;

tibial

(Madagascar)

holotype F!

Ethiopian

the species

region:

is named

in BM. MADAGASCAR:

in honor of Mr.

Perinet,

P.L.G.

unknown

BM!

Benoit.

5. Deinodryinus monticolus n. sp.

FEMALE:

with

1

segment 5 of

lamellae of approximately

group of nearly 8

unknown

DISTRIBUTION: NOTE:

the proximal

unknown

LOCUS TYPICUS: HOSTS:

with

apex with

than

distal

spurs 1,1,2.

Anteoninae

MALE:

fully

winged;

antennae brown; brown;

length 2,43 mm;

legs testaceous,

antennae not distally

general

breadth of

9: 5: 6: 6: 6: 6: rugose;

frontal

OOl = 5; hairy,

segments;

line present;

without

TL = 3;

finely

with

head dull,

occipital scutum

without

posterior surface;

posterior surface with

transversal part

bands;

distal

proportions:

smooth,

POL = 7;

finely

notaulices

transversal

areas;

Ol = 3;

punctate and

incomplete,

sculpture among punctures;

strong

lateral

following

reaching

scutellum and metanotum shiny,

with

as rugose as

in

keel

between

2 longitudinal

fore-wings

part of radial

hyal ine,

vein

propodeum

dorsal

keels; without

much shorter

and

median

branch

lOCUS TYPICUS:

wrapping

Kayove

TYPICAL MATERIAL: HOSTS:

penis;

tibial

spurs

area

dark

than proximal

(3,5: 8,5); gonoforceps without proximal membranous process,

wide apical

partly

long as

granulate and reticulate

carina complete;

shiny,

reticulate rugose, nearly

a

hairs at most as

antenna I segments

length of scutum;

punctate,

mandibles testaceous;

hind coxae and hind femora

sculpture among punctures;

approximately 0,5 smooth,

black;

with

thickened,

5,5: 5: 6: 8;

OPl = 2;

131

with a

1,1,2.

(Ruanda)

holotype M!

in TV.

unknown

DISTRIBUTION:

Ethiopian

region:

RUANDA:

Kayove

OR I ENTAL SPEC I ES OF

One spec ies

(Kisenyi),

TV!

DEINODRYINUS

is known.

Deinodryinus asiaticus n. sp.

FEMALE:

fully

winged;

antennae testace0us, testaceous; with

lateral

keels;

the sides of

and orbits;

the

occipital

temples

surrounded by invisible;

around orbits; transversal punctate, shorter

keels;

without

than

temples

furrow

triangle convex;

wide furrow

POL = 4;

sculpture among

the scutum

(15:22);

this keel

OOl

= 11;

pronotal

with

sculpture;

OPl

the

short the

behind posterior ocelli, carina; with

occipital short keels;

a

strong

crossed by

hairy on

punctures;

without

crossed by

side of head

smooth,

tarsi

keels and between

to occipital

transversely

Ol = 3;

shiny,

frons

longitudinal

and ventral

with

in following

starts at anterior ocellus;

carina complete;

between eyes and

pronotum

legs brown,

head shiny,

triangle;

2 oblique keels connect posterior ocelli carina

mandibles testaceous;

antenna I segments

median keel

at sides of ocellar

keels on

keels

thickened;

6: 4: 4: 6: 6: 6: 5: 9;

7:

longitudinal

keels start

transversal lateral

13:

3

black;

segments 7-10 brown;

antennae distally

proportions: frons

length 3,75 mm;

with

=

6;

Tl

the sides,

keel

short

=

4;

ocellar

very

finely

posterior surface of pronotum

tubercles

reaching

the

tegulae;

132

Mem.

scutum

shiny,

notaulices

smooth,

incomplete,

scutellum very punctures;

Amer.

dark

humped,

fore

bands,

distal

humped, with

one on

segments tarsi

length of scutum;

without sculpture among

without sculpture, longitudinal

keels;

shiny;

smooth,

without

fore-wings with 2

cells and another beneath

vein much

shorter than proximal

in following proportions:

produced

smooth,

posterior surface shiny,

the basal

part of radial

tarsal

segment 3 of fore

37,1984

without sculpture among punctures;

punctate,

posterior surface without

transversal

no.

approximately 0,25

strongly

metanotum very

pterostigma; (3:8);

Inst.,

finely punctate, reaching

propodeum reticulate rugose, sculpture;

Ent.

into a hook;

7:

1:

6:

enlarged claw

12: (Fig.

the

part 25; 77)

with

1 bristle located further distally than the proximal prominence; segment 5 of fore tarsi with a few rows of nearly 39 lamellae; apex with a group of 7

lamellae;

tibial

Fig.

MALE: unknown LOCUS TYP I CUS:

77 -

spurs 1,1,2.

Chela of

Deinodryinus asiaticus n. sp. (type)

Phou Kou Khouei

TYPICAL MATERIAL:

holotype F!

HOSTS: unknown DISTRIBUTION: Oriental

region:

(Laos) in B.

LAOS:

Phou

Kou Khouei

(Vientiane),

B!

Anteoninae

133

NEARCT I C SPEC I ES OF DEINODRYINUS

Three species are known.

KEY TO THE SPECIES

FEMALES

Propodeum strongly

reticulate rugose, mainly on dorsal surface .. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1. atriventris (Cresson) - Propodeum not reticulate rugose; dorsal surface mainly smooth wit h on I y weak irregular keels ....•••••••••••••••••••••.••••••• 2 2 Face with a few irregular keels near the clypeus . . . . . . . . . . . . . . .. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2. quercicolus R.C.L. Perkins - Face fully sculptured by irregular keels •...••••••••••••..••.••• . . . . .. . . . . . . . . . . . . .. . . . . . . . . . . . . . . . 3. paradoxus R.C.L. Perk i ns

MALES

Gonoforceps without head

strongly

long apical

reticulate

branch

rugose ..••

wrapping

penis

(Fig.

86);

3. paradoxus R.C.L. Perkins

long apical branch wrapping penis (Figs. 81, 84) . . • • • . • • . . • • . . • • . . . • . . . . . . . • . • . . . • • . . . . • • . . • • • . . . . • . • 2 2 Head strongly reticulate rugose (Fig. 78) . . • . . . . . . . • . . • • . . . . • . • . • • . . . . . • • • • • . . • . . • • • . • . . . • . . • • • . . . • 1. atriventris (Cresson) -

Gonoforceps wi th a

82,

-

83,

Head punctate,

without

sculpture among punctures,

rugose . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

2.

not

reticulate

quercicolus R.C.L. Perkins

1. Deinodryinus atriventris (Cresson)

Dryinus atriventris Cresson 1872: 193. Chelogynus henshawi Ashmead 1893: 93 (n. syn.); locus typicus Mil ton (Massachusetts,

U.S.A).

Anteon rugosus Ashmead 1893: 96 (n. syn.); locus typicus III i noi s (U.S.A) .

Bocchus atriceps Brues 1904: 118 (n. syn.); locus typicus Moshol u (New York,

U.S.A).

Chelogynus ferrugineus Brues 1905: 183 (n. syn.); locus typicus Texas (U.S.A) •

-

134

Mem. Amer.

Fig.

78 -

Male of

En!.

Ins!. ,

37,

1984

Deinodryinus atriventris (Cresson): sculpture of the head

•

no .

Anteon i nae

Fig. 79 - Male of

•

,

135

Deinodryinus atriventris (Cresson): antennal hairs

136

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

Chelogynus atriceps (Brues): Brues 1905: 183. Chelogynus grandis Brues 1905: 184 (n. syn.); locus typicus Riverside (Massachusetts,

U.S.A).

Anteon atriventris (Cresson): Kieffer 1907b: 25. Anteon ferrugineus (Brues): Kieffer 1907b: 26. Anteon grandis (Brues): Kieffer 1907b: 26. Anteon henshawi (Ashmead): Kieffer 1907b: 26. Chelogynus schaefferi Brues 1907a: 101 (n. syn.); locus typicus Huachuca Mts

(Arizona,

U.S.A).

Deinodryinus variabilis Fenton 1927: 10 (n. syn.); locus typicus Falls Church

(Virginia,

U.S.A).

Deinodryinus variabilis Fenton var. carinatus Fenton 1927: 11 syn.); locus typicus Flatbush (New York, U.S.A). Deinodryinus pilosus Fenton 1927: 12 (n. syn.); locus typicus Chiricahua Mts

(Arizona,

(n.

U.S.A).

Deinodryinus atriceps: (Brues): Muesebeck and Walkley 1951: 1041. Deinodryinus atriventris (Cresson): Muesebeck and Walkley 1951: 1041. Deinodryinus ferrugineus (Brues): Muesebeck and Walkley 1951: 1041. Deinodryinus grandis (Brues): Muesebeck and Walkley 1951: 1041. Deinodryinus henshawi (Ashmead): Muesebeck and Walkley 1951: 1041. Deinodryinus pilosus Fenton: Muesebeck and Walkley 1951: 1041. Deinodryinus schaefferi (Brues): Muesebeck and Walkley 1951: 1041. Deinodryinus variabilis Fenton: Muesebeck and Walkley 1951: 1041. Trisanteon rugosus (Ashmead): Muesebeck and Walkley 1951: 1042. Deinodryinus atriceps (Brues): Krombein 1979: 1242. Deinodryinus atriventris (Cresson): Krombein 1979: 1242. Deinodryinus ferrugineus (Brues): Krombei n 1979: 1242. Deinodryinus grandis (Brues): Krombein 1979: 1242. Deinodryinus henshawi (Ashmead): Krombein 1979: 1242. Deinodryinus pilosus Fenton: Krombein 1979 : 1242. Deinodryinus schaefferi (Brues): Krombei n 1979: 1242. Deinodryinus variabilis Fenton: Krombein 1979: 1242. Trisanteon rugosus (Ashmead): Krombei n 1979: 1242. r

FEMALE:

fully

the varieties); fully 15: 8:

winged;

length 5-7 mm;

body fully

iestaceous or blackened; 17:

13: 9:

10: 9:

colour very

variable

testaceous or almost fully

9:

7:

antennal 9;

black;

(see key

to

legs almost

segments in following proportions:

head shiny,

with sculpture very

variable; head fully reticulate rugose or with numerous irregular keels, sometimes with areolae and keels; frons with 3 median and parallel longitudinal keels or without visible keels; the old specific taxonomy based on the presence or absence of these keels is not recommended, because of the wide variability of sculpture in

the same species;

always with a keel

carina complete;

shiny,

hairy,

with

strongly punctate;

around a

strong

pronotal

the orbits;

occipital

transversal

anterior

tubercles reaching

impression;

tegulae;

frons pronotum

pronotum

scutum shiny,

Anteon i nae

Fig.

80 -

Male of Deinodryinus atriventris

137

(C re sson):

notaulices

Mem. Amer . En!.

138

Ins!. ,

no. 37,

1984

:. _>C."

":-, . ',¥ :

Fig .

81

-

Male of

Deinodryinus atriventris (Cresson): genital ia (A: apical branch)

Anteon i nae

139

-••

A

!

•

I ,

Fig.

82 -

Male of

Deinodryinus atriventris {Cresson}: genitalia {A: apical branch}

140

Mem.

smooth,

finely

usually

almost complete,

punctate,

only 0,65-0,75

Amer.

En!.

Inst.,

no.

37,

1984

without sculpture among punctures; reaching posterior margin;

length of scutum;

scutellum and metanotum shiny,

without

sculpture or finely

rugose;

areolae always present on dorsal

surface;

usually

with

without areolae;

2 dark

an excavated median

pterostigma; (10:14);

distal

one on

sometimes slightly

hook;

distally

3:

enlarged claw

than

7:

tibial

spurs 1,

usually

(13:14); 28;

63)

1,

posterior surface fore-wings

with

shorter than proximal

fore tarsal

segments

segment 3 of fore tarsi

with

prominence;

with 2-3 rows of nearly 30-51 lamellae;

vein

12:

(Fig.

the proximal

smooth,

less reticulate

the basa I cell s and another beneath

shorter

11:

propodeum more or

region

part of radial

following proportions: into a

punctate;

transversa I ban ds,

notaul ices

sometimes reaching

1 bristle

apex

with

in produced

located further

segment 5 of fore tarsi

lamellae;

the part

(Fig.

63)

a group of nearly 9-16

2.

MALE: fully winged; length 3,0-4,5 mm; black; antennae fully testaceous or brown, with segment 1 testaceous; mandibles testaceous, with teeth brown; fully

clypeus black or testaceous;

testaceous,

and clubs of femora black, shiny,

hairy,

strongly

antennae not distally proportions:

15:

7:

in addition

to occipital connecting invisible; smooth, (Fig.

thickened;

15:

12:

carina

carina;

with frons

the ocell i

12:

occipital

scutum)

11:

(Fig.

79);

11:

areolae

tibiae;

in following

14;

antennal

hairs much

region of head between ocell i

with 2 strong

longitudinal

sometimes

temples prominent;

and smooth,

keels

scutum shiny,

punctures;

notaul ices

the posterior margin of

sometimes reaching only 0,65

scutellum and metanotum shiny

parallel

these keels are almost

without sculpture among

incomplete (reaching approximately

length of scutum;

without sculpture or finely

punctate; propodeum strongly reticulate rugose, without a transversal between dorsal and posterior surface; posterior surface without longitudinal

keels;

fore-wings hyaline,

without dark

transversal

(Fig.

LOCUS TYPICUS:

Texas

holotype F! holotype holotype para type

wrapping penis;

tibial

(U.S.A)

holotype F of

of

WA. HOSTS:

of

spurs 1,1,2.

D. atriventris not found (not in PH); Ch. hensha"ri in WA; holotype M! of A. rugosus in WA; F! of B. atriceps in CM; holotype F! of Ch. ferrugineus in PH; F! of Ch. grandis in PH; holotype F! of Ch. schaefferi in WA; F! and 2 para types FF of D. variabilis in WA; hoi otype F! and F! of D. variabilis carinatus in WA; hoi otype F! of D. pilosus in

TYPICAL MATERIAL:

holotype

83),

keel

bands;

distal part of radial vein sl ightly or much shorter than proximal part (9: 10 or 10: 14); gonoforceps with a long apical branch (Figs. 81, 82), variable shape

head

very deep;

2 strong obi ique keels connecting posterior ocell i

carina complete;

or complete;

legs with coxae

mid and hind

(Fig.78); 11:

legs sometimes

sometimes

antenna I segments 12:

to the clypeus;

strongly punctate,

80)

testaceous;

to black

reticulate rugose

longer than breadth of segments and occipital

tegulae

with hind cOXae basally black;

unknown

DISTRIBUTION:

Nearctic region:

CANADA:

Bothwell

(Ont.),

OT!

Dresden

Anteoninae

141

\

'

,, , '\ ,

~j!) Fig.

83 -

Male genitalia of

shape of the apical Arbor,

branch

Michigan

(B),

Deinodryinus atriventris (Cresson): variable (specimens from Cochise Co.,

Uvalde Co.,

Texas

(C),

Arizona

Salmon,

Texas

(A), (D).

Ann

142

Mem.

Amer.

(Ont.),

aT!

(Ont.),

aT!

Hamilton

(Ont.),

aT!

Innisville (Ont.),

Cumberland

(Ont.),

Stittsville (Ont.), Que.), OL!

aT!

aT!

Hull

Lytle Creek

Co.,

Calif.),

TW!

IT!

SCi

(S.

Salmon

(Anderson Co.,

Bentsen

Bdo.

Huachuka Mts

Chiricahua Mts

(Ont.),

OL!

(Uvalde Co.,

Tex.),

N.P.

(Tex.),

Co.,

Tex.),

Whitewood

aT!

(5.

TE!

Alfalfa Co. Wisconsin BM!

PH!

WA!

Genesee Co. aT!

Mo.),

(Ont.),

CAl

S.

Portal

(Old Chelsea, (N.

(Tex.),

WA!

Tex.),

Tex.),

TE!

TW!

WA!

Tex.),

TE! WA!

TE!

Speir Rch. WA!

Big Bend

Elkhart

(Brazos Co.,

(Kans.),

Ariz.), TW!

(Tex.),

(Tex.),

aT!

Diego

(Ariz.),

Presidio

Hidalgo Co.

B.),

Marcos (5.

(Cochise Co.,

Hidalgo Co.,

aT!

aT!

(Anderson

Tex.),

TE!

Marysville (Kans.),

Big Fork

(Minn.),

TW!

Milwaukee Co.

(Mich.),

TW!

BM!

OL!

Lake Odessa

County 4 (Iowa),

WAr

Illinois,

(Mich.),

WA!

(Boone Co.,

Golden Pond P.

TW!

(Lexington,

WAr

Pine Heath

Tenn.),

aT!

(Gatlinburg,

Rapides Parish Ol! TW!

Mathews Co.

Bethesda

(Rabun Co.,

C.),

TW!

lost River State Park

(Va.),

WA!

Glencarlun

Falls Church Hummelstown

Pine Mountain

WA!

Montevallo

aT!

(N.

aT!

(Ky.),

Mo.),

(Wis.),

Cleveland (5. C.), TW! Ol! Wattacoo (Pickens Co., S. C.), TW! Mc Clellanville (S. C.), TW! Greenville (S. C.), TW! Wake Co. (N. C.), Highlands

(Ga.),

Ancaster

Ann Arbor

Natchez Trace S. Forsyth

WAr

(Erath Co.,

Columbia

TE!

One Sided Lake

Kouchibouguac N.P.

College Station

WA!

aT!

Foxboro (Ont.),

Stephenville (Erath Co.,

WAr

Lawrence

(Okla.),

(Mo.),

(Vernon Co., TW!

Bluffdale

(Brues 1905);

Williamsville WA!

Tex.),

aT!

Parker Canyon Lk.

Brownsville

Texas,

Oak.),

TW!

Tenn.),

TW!

TE!

ML!

(Mich.),

SCI

(Brownsville,

(Ont.),

Calif.),

(Rio Grande Valley State Park,

Esprza Rch.

1984

Summit of King Mt.

(Ariz.), WA!

37,

(On!.), CAl

OL!

Co.,

(Ariz.), Tex.),

aT!

aT!

no.

Park

Ottawa

(Que.),

(Que.),

Inst.,

Rondeau

aT!

Wainfleet

Hemmingford

U.S.A.:

(la.),

TW!

Ent.

(Va.), (Pa.),

WAr WAr

Pk.

Md.),

(N.

J.),

(Va.),

(Pa.),

laurel

(Montg'y Co.,

High Point 51.

Dupont

TW!

CAl

WAr

(Md.), WAr

(W.

Va.)(Krombein

Ol!

(Pa.),

Plummers

Takoma Pk.

Moorestown

1979);

Martinsville (Va.),

Spring Br.

aT!

Ga.),

(Md.),

(N.

J.),

lsi.

TW! TW!

aT!

TW! (Md.),

WAr

New Jersey, WAr

Princeton

WA! (N.

J.), 'IVA! (N. Y.),

Brown's Mills Jc (N. J.), WAr Westerly (R. I.), TW! Poughkeepsie TW! Farmingdale (N. Y.), TW! Millwood (N. Y.), TW! Armonk (N.

Y.),

Milford Center

AM!

TW!

Flatbush

(N.

Y.),

(N.

AM!

Y.),

TW!

Ithaca

South Haven

(N.

Y.),

AM!

Station (Huntington, N. Y.), AM! Huntington Kerhonkson (Cherry town, Ulster Co., N. Y.), N.

Y.),

AM!

Lewisboro

(Westchester Co.,

N.

(long

Island,

Y.),

AM!

Newburgh

N.

Y.),

lO!

Six-Mile (Ithaca,

N.

(Tompk Co.,

N.

Y.),

ZO!

lake Ridge Road

(Tompkins Co.,

(N.

West Point Riverside

Y.),

(N.

WA!

Y.),

(Mass.),

Mosholu

WAr PH!

(N.

Y.),

CM!

Wash'n

(N.

H.),

Y.),

lO!

Me.

Mt Whiteface

Pigeon Cove (Mass.), Mt.

Y.),

(N. Y.), AM! 4 mi. NNW AM! Brainard (Rensselaer Co.,

(Fostertown, Flatbush

N.

Kalbfleisch Field Res.

CM!

AM!

Milton

MEXICO:

lean Bogs

N. (N.

Y.),

aT!

Y.),

WAr

(Mass.),

'IVA!

Iturbide (Nuevo

Leon), TE! Colima (Manzanillo), SC! EI Palmito (Sinaloa), aT! Valle Nacional (Oaxaca), TW! Miltepec (Oaxaca), TE! Tehuacan (Puebla), OL! NOTES:

the colour of the female of D.

variable. Though

atriventris (Cresson) is very

Some varieties were described

there are intermediate colours

in

the past as good species.

I am proposing

the following key

to

Anteoninae

the main colour varieties

Body

fully

(without

testaceous with

143

taxonomic

value):

petiole black . . . . . . . . . . . . . . . . . . ..

· . . • . . . . . • . . . . . . . . . . . . . . . . ferrugineus (B ru es) - Body at least partly dark or black . . • . . . . . • . . . . . . . . . . . . . . . . ... 2 2 Thorax and propodeum fully or almost fully black . . . . . . . . . . . . . . 3 -

Thorax and propodeum at

3 Legs

fully

testaceous,

least at

partly

most

wi th

testaceous hind

or

reddish . . . . . 5

coxae basally

black . . . .

· . . . . • . . . . . . . . . . . . . . . . . . . . henshawi (Ashmead) (= variabilis Fen t on) - Legs most Iy black • • • • . • • • . • • • • . • • • • • • • • • • • • • • • . • • . • • • . • • • . . • • . . 4 4 Frons with three distinct longitudinal and parallel median keels

. grandis (Brues) -

Frons without

three distinct

longitudinal

and parallel

median

keel s •••..••••••••••..••• . pilosus Fenton 5 Head,

thorax and propodeum testaceous;

on propodeum;

abdomen

occasionally brown brown . . . . . . . . . . . . . . . . . .

spots

· . . • . . . . . . . . . . . • • . . . . . . . . . atriventris (C res son) -

Thorax and propodeum part Iy

6 Prothorax

testaceous;

testaceous . . . . . . . . . . . . . . . . . . . . . . . . . 6

mesothorax,

metathorax and propodeum black

. . • . . . . . . . . . . . . • . . • . . . . . . . schaefferi (Brues) -

Thorax and propodeum mostly

testaceous,

spots . . . . . . . . . . . . . . . . . . . . • atriceps

with

(Brues)

irregular dark

(= carinatus Fenton)

The presence or absence of keel s on the frons (see grandis and pilosus) has no taxonomic value: this characteristic is very variable,

with

intermediate cases.

2. Deinodryinus quercicolus R.C.L. Perkins

Deinodryinus quercicola R.C.L. Perkins 1907: 47. Deinodryinus quercicola R.C.L. Perk i ns: Muesebeck and Walkley 1951 : 1041.

Deinodryinus quercicola R.C.L. Perkins: Krombein 1979: 1242.

FEMALE:

fully

winged;

length 4,57 mm;

near the tegulae and with segments 4-10 dark; face with

a few

complete;

OPL

segment 3 internal keels;

body

petiole black; shiny,

hairy;

longer than OL;

head shiny,

antennae thin,

organs visible;

(10:6);

body

impress ion,

tegulae;

weakly

occipital

distally

granulate; carina

thickened,

the head,

with

as

long

with

but without

pronotum crossed by

approx ima te I y

spot

with

almost transparent,

pronotum as sculptured as

tubercles reaching

anterior transversa I

with a black

antennae testaceous,

irregular keels near the clypeus;

longer than segment 4

pronotal

yellow-reddish,

as broad;

an scutum

144

Mem.

as sculptured as

the pronotum;

approximately 0,65 propodeum with with 2 dark beneath distally

weak

than

1,1,2. MALE:

fully

8:

black;

distally

7:

7:

7:

occipital

shorter than 4 1 bristle

mandibles,

with coxae, thickened;

7:

6:

without sculpture among punctures;

vertex;

carina complete;

fore-wings

cells and another

6:

8;

(22:36);

located further

segment 5 of fore tarsi

legs testaceous, 5:

reaching

group of about 15 lamellae;

antennae not 8:

19B4

not reticulate rugose; the basal

1 of fore tarsi

length 2,37 mm;

following proportions: than

a

37,

incomplete,

tooth and with

prominence;

apex with

testaceous;

tibiae brown;

punctate,

keels,

segment

no.

scutellum and metanotum smooth;

one on

without sUbapical

winged;

antennae fully hind

bands,

the proximal

Ins!.,

notaul ices

transversal

the pterostigma;

of 13+26 lamellae;

En!.

length of scutum;

transversal

enlarged claw

Amer.

with 2 rows

tibial

spurs

clypeus and clubs of femora

antenna I segments head shiny,

and in

smooth,

frons more strongly punctate

POL = 5;

OL = 3;

OOL =

7;

OPL

5; TL = 6; scutum, scutellum and metanotum shiny, smooth, very finely puncta te, without scu I p ture among the punctures; notau Ii ces strong,

/

Fig.

BL, - Male genitalia of Deinodryinus quercicolus R. C. L. Perkins from Nogales,

Arizona

(right half removed).

Anteoninae

incomplete,

reaching

reticulate rugose,

approximately 0,65

with

a

keel

between

posterior surface without

longitudinal

without

bands;

dark

as proximal ( Fig.

84);

transversal part

(7:7);

length of scutum; dorsal

keels,

distal

rugose;

fore-wings hyal ine,

part of radial

gonoforceps with

propodeum

and posterior surface;

an apical

vein nearly

branch

as

long

wrapping penis

t i b i a I s p u rs 1, 1 , 2 •

LOCUS TYPICUS:

Nogales

TYPICAL MATERIAL: HOSTS:

145

(Arizona,

holotype F!

U.S.A)

in B.

unknown

DISTRIBUTION:

Nearctic region:

U.S.A.:

Nogales

(Ariz.),

B!

3. Deinodryinus paradoxus R.C.L. Perkins

Deinodryinus paradoxus R.C.L. Perkins 1907: 46.

FEMALE: small

fully

on segment 10 dark; 6:

6:

5:

keels; 12;

winged;

black spot

finely

1 of the abdomen;

antenna I segments 7;

head shiny,

face fully ~

OPL

length 4,02-5,30 mm;

near tegulae and a

8;

TL

punctate,

antennae yellow-testaceous

weakly

alutaceous,

10;

hairy,

tubercles reaching

the

with a

weak

without sculpture among punctures; reaching 0,5- 0,65 finely

punctate,

shiny,

almost

unsculptured,

13:

3;

8:

7:

7:

irregular weak ~

OL

impression;

smooth,

2;

~

OOL

smo')th,

pronotal

finely

punctate,

incomplete,

scutellum and metanotum smooth,

without sculpture among with

6:

pronotum shiny,

notaulices distinct,

length of scutum;

shiny,

transversal

scutum shiny,

~

POL

carina complete;

tegulae;

with segments 9-

11:

'with numerous

irregular keels;

occipital

with a

the petiole and partly

in following proportions:

sculptured by ~

yellow-testaceous,

black spot on

weak

the punctures;

transversal

keels,

propodeum

without a strong

transversa I keel between dorsa I and posterior surface; posterior surface without longitudinal keels; fore·-wings with 2 dark transversal bands, one on

the basal

radial

vein

proximal narrow;

cells and another beneath longer or sl ightly

parts forming fore tarsal

(Fig.

85)

radial

proximal

cell

open;

into a hook;

lamellae;

apex with

with a medial

distal

part;

part of

distal

pterostigma

in following proportions:

produced

with 2 rows of 30-39

enlarged claw

curve;

segments

segment 3 of fore tarsi 85)

a

the pterostigma;

shorter than

9:

3:

and

long and

7:

14:

30;

segment 5 of fore tarsi

(Fig.

a group of 10-15 lamellae;

peg-like bristle;

tibial

spurs 1,1,2.

MALE: fully winged; length 2,56-2,81 mm; black; antennae yellow; mandibles testaceous with teeth reddish; clypeus and tegulae testaceous; legs testaceous with coxae, antennae very segment 3; 10: keel

10:

10:

hairy;

clubs of femora

hairs very

antenna I segments 9:

11;

head shiny,

around the orbits;

long,

and

nearly

tibiae partly

twice as

in following proportions: strongly

long 14:

reticulate rugose,

frons with 2 parallel

longitudinal

6:

brown; as breadth of 14:

with

9:

10:

a strong

keels and

146

Mem.

Fig.

85 -

Chela of

Amer.

En!.

Inst.,

no.

37,

1984

Deinodryinus paradoxus R. C. l . Perkins from Nogales (Arizona)

frontal

I ine;

connecting

vertex behind posterior ocell i with 2 short obi ique keels

the posterior ocelli

to the occipital

there are 2 areolae and the occipital

carina

carina;

between

is excavated;

these keels

POL

=

7;

=

Ol

4; OOl = 11; OPl = 7; Tl = 11; scu tum, scu tell um and metanotum smooth, shiny, finely punctate, without sculpture among punctures; notaul ices distinct,

incomplete,

reaching

propodeum reticulate rugose, posterior surface, surface; of radial and

with 2

incomplete

fore-wings hyaline, vein slightly

narrow;

approximately 0,75 without a

longitudinal

without dark

(Fig.

86)

without

keel

part

long apical

and 2 paralectotypes

(designated by Olmi

1 paralectotype F!

in B;

dorsal

and

posterior

bands;

(8:10);

penis; tibial spurs 1,1,2. lOCUS TYPICUS: Nogales (U.S.A) TYPICAL MATERIAL: lectotype F! 1982)

between

keels on

transversal

shorter than proximal

gonoforceps

length of scutum;

transversal

distal

part

pterostigma

long

branch wrapping

(IF,

in BM.

1M)!

Anteoninae

147

B Fig.

86 -

Nogales,

Male genitalia of Deinodryinus paradoxus R. Arizona

(A);

left gonoforceps

(B)

C.

L.

Perkins from

of a specimen from EI

Naranjo

(Mexico)

HOSTS:

in U.S.A.

Cicadell ids

(R.C.L.

Perkins 1907).

DISTRIBUTION: Nearctic region: U.S.A.: Nogales (Ariz.), B! BM! OL! Portal (Ariz.), TW! OL! Fripp Island (Beaufort Co., S.C.), WA! MEXICO: EI I~aranjo

(San Luis Potosi),

TE!

San Pedro

Iturbide

(Linares,

TW!

NEOTROP I C SPEC I ES OF DEINODRYINUS

Fifty five species are known.

KEY TO THE SPECIES

Nuevo Leon),

148

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

FEMALES

Head

fully

rugose,

smooth,

wi thout

without

areolae

sculpture,

or keel s,

not

except

granulated, for

the

not

frontal

line

• ••••••••••••••••••••••••••••••••••••••••••••••••••••••••••••• 2

Head more

2 3

or

less

rugose,

0,5

reaching

approximately posterior margin of scutum .. 4 transversal keel between dorsal and

Species

a

st rong

1. elegans n.

transversa I

testaceous,

at

most

Head,

least

pronotum and

scutum fully

pronotum and

scutum fully

or almost

dark

fully

..5

nigrorufus n.

4.

or almost

fully

bands,

sp.

black . . .

iphias n.

5.

transversal

reddi sh-

sp.

fully

hyal ine

or

1-2 distinct

dark

transversal

bands . . . . . . . . . . 9

two

strong

obi ique keels

connecting posterior

ocell i

to occipi tal carina • • • . • . • • . . . • • • • • • • • • • • . . • • • • • • • • • . • • . • • • . . • 8 Frons with two median longitudinal keels; notaulices short, reaching

less

0,5

than

length

of

scutum . . . . . . . . . . . . . ..

.................................. Frons with almost

three median

reaching posterior margin

of

Antennal

segment

3

sl ight Iy

shorter

bicolor (Olmi

6.

longitudinal

· . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. 9

sp.

Head without two strong oblique keels connecting posterior ocell i to occipi tal carina . . . . . . . . 49. minor n. sp. Head with

8

and

darkened . • • • • • • • • • • • • • • • • • . . • • • • • • • • • • • • . • . • • • • • • • • • • • • 7

Fore-wings with

7

diaphanus n.

3.

· . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. Fore-wings wi thout fully

dorsa I

sp.

with propodeum b l a c k . . . . . . . . . . . . . . . . .

testaceous . . . . . . . . . . . . . . . . . . . . . . . . Head,

sp.

between

with petiole black.

· . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. Species at

keel

speciosus n.

2.

surface . . . . . . . . . . . . . . . . .

fully

of

strong

surface ••.••••••••••••••

posterior

6

smooth.6

Notaulices

Propodeum wi thout

5

never

approximately

Propodeum with a

length

scutum . . . . . . . . 3

reaching

posterior

4

with areolae or keels,

Notaulices

keels;

surface of

smooth median

7.

finely

maximus n.

than

propodeum almost

region,

longer,

scutum . . . . . .

••• • • ••• . •••••. . •. . •••••• •••• ••• •. 8. Antennal segment 3 longer than segment 10 Posterior

and Currado)

notaulices

punctate,

segment

sp. 1 •••••••••••••

melanocephalus (Cameron) 1 • . • . . • • • • • • . • • • . . • • • • 10 fully

smooth

without

or wi th

keels

a

or

areolae ••••••••••••••••••••••••••...••••••••••••.•••••••..••• 11 Posterior keels 11

surface

of

Vertex of head wi th po s t e rio roc e I l i t 0 Vertex

propodeum rugose,

dul I,

sculptured by

or areolae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ..

of

.. 27

two strong obi ique keel s connect ing ') C c i pit a I ca r ina •..•••••••••••••••..••••. 1 2

head without

two obi ique keels

connecting posterior

ocelli to occipital carina . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 20 12 Posterior surface of pronotum rugose, wi th numerous irregular transversal

keels ••••••••••••••••••••••••••••••••••••..•••••• 13

Anteoninae

Posterior

surface

sculpture,

or

of

finely

149

pronotum smooth,

without

keels,

without

punctate . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 14

13 Thorax and propodeum black . . • • • . . • Thorax and propodeum fully

9.

or almost

noyesi n. fully

sp.

reddish-testaceous ••

• ........••....•......••.••••... " 10. atlanticus n. sp. 14 Posterior surface of propodeum wi th two strong longitudinal keels and a shiny

strong

transversal

keel

median area • • • . • • . • • • • . . • • • .

Posterior smooth

surface

area

of

surrounding

propodeum without

surrounded by

a

smooth and

11. achterbergi n.

sp. keels;

longitudinal

irregular keels •••••••• 15

areolae and

15 Head mainly b l a c k . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Head

fully

or mainly

16 Thorax and propodeum fully black .• Thorax and propodeum mainly

17 Pronotum with

two

strong

18 Vertex of head reticulate Vertex

of

punctate,

two

transversal

strong

(behind and

not

(behind and reticulate

dorsal

• •• 17

lobes ••••••••

fluviatilis n.

53.

on

the

sides

on

the

of

the ocell i)

fully

atlanticus n. sp.

10. sides

rugose .••

sp.

lobes . • • . . • • . . • 18

transverse dorsal

rugose •.••••.•••••••••• head

peruvianus n. sp.

12.

testaceous...... .••••..•

· ............................... " Pronotum without

. ••• 16 • ••• 19

reddish-testaceous. ••••••..••••

of

the

ocell i)

13. hymenaeus n.

sp.

19 Pronotum, scutum, scutellum and metanotum fully black ••. • •••.••••••••..•••..•••.•••.•••• " 14. colombian us n. sp. Pronotum,

scutum,

scutellum and metanotum mainly

testaceous ••••••••••••••••••••••••

20 Region

of

head behind

ocell i

15.

smooth,

punctate,

of

head behind

longitudinal

ocell i

rugose,

wi thout

sculptured by

numerous

..25 ..22 • .24

keels..............

21 Prothorax fully or mainly black. Prothorax

reddishsp.

keels ••••••••••.••••••. 21

sculpture among punctures and wi thout Region

cascus n.

fully

testaceous

22 Frons wi thout

areolae, pronotum with two very

pronotum black,

with

•••••••

.••••••••.•••.••••••

wi th only a few I ongi tudinal prominent dorsal transversal

dorsal

lobes and pronotal

wh i t i sh ••••••••••••••••.•••••••• "

16.

keel s; lobes;

tubercles

albopictus n.

sp.

Frons reticulate rugose, sometimes with longitudinal keels; pronotum without dorsal lobes or with only slightly prominent dorsal

23 Frons

transverse reticulate

claw with

lobes;

pronotum fully

rugose and with

one peg-like hair

black •••••••.•••••••. 23

longi tudinal

(Fig.

keel s;

enlarged

110) . . . • • . • • . . . • . . • • • • . • . • . . .

17. amoenus n. sp. Frons without very broad,

longitudinal

keels,

reserrbl ing areolae;

strongly

enlarged

punctate;

claw wi th

punctures

two peg-I ike

hairs (Fig. 111) ....•••••..•••.•.• 18. pegnai n. sp. 24 Mesothorax, metathorax and propodeum black ••••••.••••••.••• . ••••• . ••••••••••• . • . . ••••••. . . . " 19. pseudoamoenus n. sp. Mesothorax,

metathorax and propodeum

testaceous ••••••••••••••••

150

Mem.

Amer.

Ent.

Inst.,

· ....•••........••...•.........• " 25 Notaulices scutum;

thin,

reaching

pronotal

Notaulices deep, pronota I

dorsa I

of

reaching

posterior margin

reddish;

22.

notaulices

transversal

21.

of

scutum;

52.

furrow

about

approximately

0,50

biloboides n. sp. edges

pseudobilobus n.

of

sp.

Head without

a

of

ocell i • • • • • . • • . • • • • • • . • • • • • • • • • . • • • • • • • . . • • . • • • . 28

posteri or

furrow

0,75

bilobus (Fenton)

joining posterior 23.

transversal

sp.

reaching

reaching

ocell i . . . . . . . . . . . . . . . deep

insignis n.

notaul ices

scutum . . . . . . . . . . . . . . . . . .

posterior

of

less promi nen t . . . . . . . . . . . . . . . . . . . . . . . . ..

fully

27 Head with a deep

length

almost

scutum . . . . . . . . . . . . . . . . . .

of

sp.

0,50-0,75

lobes more prominent . . . . . . . . . . . . . . . . . 26

Pronotum mostly black; length

1984

townesi n.

20.

approximately

· . . . . . •• . . . . • •• . . •• . . ••. •• . . . . length

37,

dorsal lobes

26 Pronotum almost

no.

joining posterior

28 Posterior surface of pronotum approximately long l and with two strong transverse dorsal Posterior

surface

less

than

twice as broad as

wi th

weak

29 Region

of

paral lei

of

pronotum approximately

(not

longl,

keels;

ocell i

pronotum fully

of

rugose,

head behind posterior

without

longitudinal

as

as broad as

without

dorsal

long

lobes

or or

with

numerous

reddish-testaceous.

· .•.•....••..•.•...............•.• Region

as broad

lobes . . . . . . . . . . . . 29

lobes • • . . . • • • . . • . . • • • . . • 31

transversal)

head behind posterior

longitudinal

and

twi~e

edges

24.

ocel I i

rubrolobatus n. punctate

or

sp.

reticulate

keels...................

.. .. 30

nigrolobatus n. sp. Prothorax fully reddish . . . . . . . . . . . 26. vagans n. sp.

30 Prothorax

fully

31

surface

Posterior rugose,

or with

or mainly of

Posterior

surface of wi thout

32 OOL approximately 33 Head

than

or

pronotum fully

twice as long

rugose,

posterior

long

rugose, to

reticulate wi thout

occipital

34 Head almost keels

on

Head more

or

longitudinal 35 Posterior

two

smooth, not

le~s

with

ocell

two to

or almost

strong

fully

smooth,

rugose,

wi th

a

.38 .33

27.

granulate,

median

reticulate

ret iculate

carina . . . . . . . . .

pilosifrons Fen t on almost

smooth,

weakly

connect ing posteri or 28. keel

.34

oblique keels

occipital

obi ique keel s

ocell i

politus n. sp. and

two weak

lateral

rugose . . . . . . . . . . . . . . . . . . . . . . . . 35

rugose

or

sculptured by

numerous

keels • • • . • • . . • • . . • • • • • • • . • • . . • • • • . • • • • • . • . . • . . . . 36

surface

of

pronotum slightly

Breadth measurements refer surface.

reticulate

keels . . . . . . . . . 32

as POL......... ••••••. •••.

carina . . . . . . . . . . . . . . .

frons,

or

as POL.............

· . . . . . • . . . . . . . • . . . . . . . . • • • . • • • . . .. Head not

rugose

longitudinal

sculpture among punctures........

twice as

reticulate

connecting

25.

pronotum fully

transversal

punctate, OOL more

black ..

to

broader

the anterior margin of

than

long,

the posterior

Anteoninae

smooth,

with

margin,

without

Posterior long,

of

numerous

and with

short

longitudinal

transversal

surface

with

margin

numerous

keels.

short

some

less

than

three

times as

than

three

least

long

as POL... • • . . • . . . . •

reddish

or

31.

keels

5 of

chiapasi n.

tarsi

wi th

a very

long

• •• 37

nigricans (Cameron) actuosus n.

of head behind posterior ocel I i without connecting posterior ocelli to occipital

fore

sp.

testaceous •••.•.•••••••

32. 38 Region

sp.

as POL ••••••. 30.

times as

partly

keels . . . . . . . . .

rivularis n.

long

37 Species fully black •••••.••••.•••• Species at

47.

sp.

times as broad as

keels near posterior

transversal

. . • . • . • • • • • . • . • • . • . • • • . . • OOL more

1,5

longitudinal

strong

keels near posterior

claripes n.

29.

pronotum nearly

.............................. 3600L

151

I ame I I a

on

sp.

two oblique carina;

segmen t

(F i g •

129) •..

ap ex

33. incaicus n. Region

of

head behind posterior ocelli

connect ing posterior

ocell

fore

very

39 Head

tarsi

without

sculptured by

a

to long

with

occipi tal lamella

longitudinal

keels,

• •• . . • • • • • • • • . • • • . . . . . . . • . . . . • .

34.

on

sp. oblique keels

two

carina; apex

segment

(Figs.

5 of

131,

132)

.•. 39 not reticulate rugose ••• bolivianus n. sp.

Head fully or mainly reticulate rugose ••••••••••.•••••••••••• 40 40 Vertex of head behind ocell fully reticulate rugose . . . . . . . . . 41 Vert ex 41

of

Prothorax

head beh i nd

oce I

not

ret i cu I ate

reddish . . . . . . . . . . . . . . . . .

35.

rugose...

• ...•••• 42

Prothorax b l a c k . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

42 Head fully

or almost

fully

testaceous

• • . • • . • • • • • • • • • . • • • • • • • • . . • • • . . •• , Head ma i n I y 43 Vertex among

of

b I a ck • • • • • • • • • • • • • • . ••

head behind

punctate,

of

head behind

long i t ud i na I ful Iy

ocel I i

strongly

kee Is • . . . • . • . . . . • . . • • • . . .

reddish-testaceous.

Head most I y black •••.•••..•••.••• 45 Prothorax black . . . . • • • • • • • . . . . • Prothorax

reddi sh-testaceous •••

sp.,

sp.,

reddish ..

trinidadi n. sp. rapax n. sp. and wi thout

sculpture •••. 44

sculptured by . . . . . . . . . 45

croceus n. sp. asper n. sp. 40. schlingeri n. sp. 41. alexandrae n. sp. 38. 39.

D. hirticornis (K i ef f er), inermis n. sp., cuzcanus aequalis n. sp., paranus n. sp., perlucens n. sp., carpens n. benianus n. sp., itenezi n. sp., panamensis n. sp. are unknown. The females of

n.

or

punctures . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

Vertex 44 Head

ocell i

36. ;J 7.

.. . . . 43

eminens n. sp.

152

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

MALES

Antennal

hairs

Antennal

hairs as

shorter

segments

(see

than breadth

long as

or much

of

segments ••••••••.•••••• 2

longer

than breadth

2

Head

3

Head at least partly rugose, reticulate rugose Gonoforceps wi th reduced internal branch (Fig.

smooth,

not

rugose,

wi thout

· . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. 1 17, 4

5

large

internal

26.

122) . . . . . . . . . . ..

vagans n.

sp. (Figs.

1 4 2 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. 4 Internal branch of gonoforceps subapical (Fig. 141.) . . . . . . · ............................... " 42. carpens n. sp. Internal branch of gonoforceps apical (Figs. 117,142) . . . . . 5

Posterior

surface of

propodeum smooth

surface

of

43.

Gonoforceps much Gonoforceps

shorter

slightly

than penis

23.

than penis

shorter,

94,

(Figs.

and

shiny ••••••••

perlucens n.

sp.

propodeum rugose and dul I ••••••••••..

· ............................... " as

pseudobilobus n. 96,

(Figs.

long as,

or

137,

sp.

156) . . . . . . 7

slightly

longer

143) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9

Head not reticulate rugose, with longitudinal keels, punctate and slightly rough; very strong and distinct oblique keels connecting posterior

ocell i

to

occipi tal

....... Head

fully

reticulate

connecting posterior

8

sp.

or with keels .. 3

141,

Posterior

7

.. . . . . ..

branch wrapping penis

· ............................... "

6

.. . . . . . 6

areolae or keel s.

4. nigrorufus n.

Gonoforceps wi th

of

79)............................

fig.

Gonoforceps without

rugose; ocelli

internal

indistinct branch

internal

branch

carina . . . . . . . . . . . . . . .. and Currado)

bicolor (Olmi

to occipital

· ............................... " Gonoforceps with an

6.

oblique keels carina . . . . . . . . . . . . . . . 8 (Fig.

137) ..

sp. wrapping penis (Fig.

156) ..

wrapping penis

38.

croceus n.

· . . . . . • . . . . . . . . . . . . . . . . . . . . . . . . . . . 55. panamensis n. 9

Gonoforceps wi thout (Figs.

94,

branch

keels,

large

internal

branch

except

fully for

wrapping penis

granulate and punctate, few

short

rugose,

wi th

without

keels near antennal

. . . . . . . . •. . . . . . . . . . . . . . . . •"

11

very

(Figs.

144) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .' . . . . . . . . 14

10 Head smooth,

Head

or wi th

143) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 10

Gonoforceps wi th

130,

internal

sp. reduced branch

5.

iphias n.

areolae or

sockets . . . . . . . . sp.

areolae and keel s . . . . . . . . . . . . . . . . . . . . . . . . . . 11

reaching approximately 0,80 length of scutum . . . . . . .. · ............................... " 44. inermis n. sp. Notaulices reaching at most 0,50 length of scutum . . . . . . . . . . . . 12 Notaulices

12 Posterior surface of propodeum without longitudinal keels . . . . .. · ............................... " 1. elegans n. sp. Posterior

surface

of

propodeum with

strong

longitudinal

keels

153

Anteoninae

· . . . • . . . • . . . • • . • • . . . • • • • . • • . . . . . . . . . • • • • . • • . . . . . . . . • • . . • • • • . . 13 13 Gonoforceps with distal inner pointed process (Fig. 155) . . . . . . . · •.....••.••...••••••••••..•.•..•. 54. cuzcanus n. sp. Gonoforceps without distal inner pointed process (Fig. 152) ••.. • • • • . • . • . . . . • • . . • . . • • . . . . • • • . . . . •• 51. itenezi n. sp . 14 Head without two oblique keels connecting posterior ocelli to occipital

carina;

vertex

of

head behind

ocelli

never

reticulate

rugose •••••••••••••••••••••.••••••••••••••••••••••••••••••••• 15 Head with

two

to occipital

15 Head fully

strong

smooth,

punctures,

not

region

keels

sometimes

finely

reticulate

Head with anterior posterior

oblique

carina,

region

connecting posterior

indistinct

punctate,

among

without

45.

of

strongly

frons

sculpture among

paranus n. sp.

rugose ..

and vertex

ocelli

areolae . . . . . . 16

reticulate

rugose;

smooth . . . . . . . . . . . . . . . . . . . . . . . . . . ..

• ••.•••••.•••...••••.•••.•••...••. 33. incaicus n. sp. 16 Gonoforceps with internal branch on proximal region (Fig. · . . . . . . . . . . . . . . . . • • . • • • • . . . . . . . . •. 25. nigrolobatus n. sp. Gonoforceps with (Figs.

104,

17 Frons with

internal

branch

on

apical

or

subapical

two median Some

Frons wi th

or wi thout

longitudinal

transversal

folds,

keels; not

region between

reticUlate

region

of

segment

two di st inct

frons always

reticulate

3 approximately as

....................••.....•...... Antennal

segment

19 Posterior

surface

3

longer of

excavated

than

Posterior

surface of

than

lateral

20 Distal

part

than

segment

propodeum wi th

lateral

areas . . . . . .

propodeum wi th

1 .... .

segment

hirticornis (Kieffer) 1 •••..•••••••••.•••••• 19

median

14.

keel s;

rugose . . . . . . . . . . . . . . 18

long as

46.

sp.

I ongi tudinal

median

these

rugose . . . . . .

12. peruvianus n.

18 Antennal

region

145) • • • • • . . . . • • . . . . . • . • • • . . . . • . . • . • • • . . . . • . . . . . . . 17

keels with

median

120)

area more deeply

colombianus n.

median

area

sp.

less

excavated

areas . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 20 of

radial

vein

shorter

or at

most

as

long

as

proximal p a r t . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . • .21 Distal part of radial vein longer than proximal part . . . . . . . . . 22

21

Notaulices

reaching

approximately posterior margin

(F i g.

157 B ) . . . . . . . . . . . . . . . . . . . Notaulices shorter (Fig. 157 A) . . . 22 Posteri or surface of

propodeum wi thout

· . . . . . . . . . • • . . . . . • • . . . . . • • . . . . . . •. Posterior

surface

of

propodeum with

· . . • . . • . . . . . . . • . . . . . . . . . . . . . . . . . ..

of

scutum

9. noyesi n. sp. 36. trinidadi n. sp. longitudinal

50. benianus n. two

48.

keels . . . . . .

sp.

longitudinal

aequalis n.

keels . . . . .

sp.

The males of D. achterbergi n. sp., schlingeri n. sp., eminens n. amoenus n. sp., asper n. sp., diaphanus n. sp., bolivianus n. sp., melanocephalus (Cameron), nigricans (Cameron), pegnai n. sp., alexandrae n. sp., bilobus (Fenton), pilosifrons Fenton, maximus n. sp., insignis n. s::>., townesi n. sp., albop'-ctus n. sp., hymenaeus n. sp., fluviatilis n. sp.,

154

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

sp., biloboides n. sp., rivularis n. sp., minor n. sp., politus n. sp., c1aripes n. sp., rubrolobatus n. sp., rapax n. sp., speciosus n. sp., atlanticus n. sp., pseudoamoenus n. sp., chiapasi n. sp., cascus n. sp.,

actuosus n. sp. are unknown.

1. Deinodryinus elegans n. sp.

FEMALE: region

fully

winged;

blackish,

length 3,0 mm;

sides of pronotum

reddish-testaceous,

with

a

black

scutum

and metanotum blackish,

black,

abdomen and petiole brown-testaceous;

antenna I segments head

dull,

complete;

fully

=

POL

3;

=

OL

behind

posterior ocelli

strong

anterior

granulated and

with

tegulae;

scutum

shiny,

reaching

approximately 0,5

shiny,

smooth,

with

2 dark

beneath part

nearly

strong

without

fore

(Fig.

produced

into a

winged;

partly

legs

brown

in following fully OL

=

3;

=

7;

punctate,

reaching

5:

=

6;

tibial

scutum,

without

10:

6,5: some

vein

nearly

branch 88);

hyal ine, as

6;

penis,

spurs

LOCUS TYPICUS:

(Fig.

dark

part a

keel

fore-wings

longer than

87)

6:

with

4:

7:

7:

7:

7:

segment

and

19; the

3 of

hind

6,5:

hairs much

tibiae

antenna I segments 8,5;

head dull,

region

behind

carina complete;

punctures;

(7:7);

12:

than

mandibles and antennae

thickened;

POL = 5;

longer than

notaul ices

smooth, incomplete,

propodeum reticulate rugose, without bands;

longitudinal distal

keels;

part of radial

gonoforceps without

short apical

(Trinidad)

proximal

2:

2 rows of

lamellae;

irregular keels;

transversal

with only

a

1,1,2.

black;

length of scutum;

Augustine

transversal

keels;

clubs of femora

antennal

1,1,2.

St.

strong

scutellum and metanotum shiny,

long as proximal

wrapping

tibial

without

with

incomplete,

located further distally

with posterior surface reticulate rugose and fore-wings

a

vein

occipital

sculpture among

approximately 0,5

8;

tubercles reaching

proportions:

spurs

coxae,

=

6:

of head

hairy,

notaulices

group of nearly 6

with

TL

region

longitudinal

following

2 obi ique keels;

OPL

breadth of segments; finely

10: frons

7;

6:

carina

and posterior surface fully

antennae not distally

proportions:

OOL

in

with

6:

the basa I Ce II s and another

length 2,31-2,50 mm;

reticulate rugose; with

with

dorsal

part of radial

a

thickened;

6:

scutellum and metanotum

propodeum

hook;

testaceous,

or black;

posterior ocell i

with

=

TL

6:

occipital

pronotal

segment 5 of fore tarsi

fore tarsi

7:

pronotum dull,

keels;

with one bristle

apex

11:

granulated;

one on

distal

lamellae;

fully

7;

length of scutum;

segments

87)

prominence;

testaceousj

=

posterior surface without

tarsal

6:

posterior surface of pronotum

weakly

sculpture;

54

MALE:

OPL

transversal

smooth,

the pterostigma;

enlarged claw

7;

furrow;

transversa I bands,

(13:9);

proximal

=

OOL

and posterior surface;

reticulate rugose;

10:

without oblique keels;

transversal

between dorsal

proportions:

propodeum

antennae distally

without areolae or keels;

3;

with ocellar

anterior margins of

mesopleura distally blackened,

in following

granulated,

spot,

an apical

membranous band

(Fig.

Anteoninae

Fig. 87 -

Chela of

Deinodryinus eJegans n. sp.

155

(type)

156

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

v

Fig.

88 -

Male genitalia of Deinodryinus elegans n.

(Trinidad);

A:

left half (dorsally viewed); (ventrally

B:

and 1 paratype M!

DISTRIBUTION:

TRINIDAD:

BOLIVIA:

Neotropic region:

Mouth of Rio Baures

(Rio

from St.

Augustine

viewed).

TYPICAL MATERIAL: holotype F! in AM; 1 para type M! in OL. HOSTS: unknown OL!

sp.

right gonoforceps

st.

Itenez,

in BM;

Augustine Dept.

1 paratype F!

(St.

Beni),

George),

AM!

BM!

157

Anteoninae

2. Deinodryinus speciosus n. sp.

FEMALE:

fully

not black;

winged;

length 3,50 mm;

antennae distally

proportions:

9:

6:

14:

7:

frons without median or Ol

~

3;

~

OOl

7;

OPL

numerous transversal scutum

shiny,

incomplete,

6,5:

lateral 8;

keels,

TL

6:

6:

keels; ~

7;

hairy;

reaching

5,5:

transversa I keel

posterior surface small;

petiole testaceous,

pronotal

in following

head dull,

granulate;

carina complete;

pronotum dull,

granulated,

tubercles reaching

weakly granulated; length of scutum;

without sculpture; between

6,5;

occipital

approximately 0,5

smooth,

testaceous;

antenna I segments

without sculpture or very

metanotum shiny, without

~

6:

fully

thickened;

POL

~

3;

with tegulae;

notaulices

scutellum and

propodeum reticulate rugose,

dorsa I and posteri or surface;

posterior surface without

longitudinal

areol ae on keels;

fore-

wings with 2 dark transversal bands, one on the basal cells and another beneath the pterostigma; distal part of radial vein longer than proximal part 21;

(14:8);

fore

tarsal

enlarged claw

(Fig.

segments 89)

in following

proportions:

with one bristle

the proximal

prominence;

segment 5 of fore tarsi

41

apex with a

group of 7

lamellae;

Fig.

MALE:

89 -

2,5:

5:

lamellae;

(Fig.

tibial

89)

15: than

with 2 rows of

spurs 1,1,2.

Deinodryinus speciosus n. sp. (type)

unknown

lOCUS TYPICUS:

Teodoro Sampaio

TYPICAL MATERIAL: HOSTS:

Chel a of

7:

located further distally

holotype F!

(Sao Paulo,

Brazil)

in OT.

unknown

DISTRIBUTION:

Neotropic region:

BRAZIL:

Teodoro Sampaio

(Sao Paulo),

OT!

158

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

3. Deinodryinus diaphanus n. sp.

FEMALE:

fully

petiole black;

Fig.

winged;

length 3-4 mm;

antennae slightly

90 -

fully

testaceous;

distally thickened;

sometimes with

antennal

Chel a of Deinodryinus diaphanus n.

sp.

segments

(type)

in

159

Anteoninae

following proportions: finely

punctate,

granulated, POL

~

3;

without

OL

~

12:

2;

tegulaej

transversal

proximal 2:

4:

part

15:

than

10:

9:

and

lateral

OPL

~

8;

furrow,

8;

weakly

scutellum

granulated;

7:

7:

10;

~

bands; 11);

distal fore

enlarged claw

the proximal

and

lamellae;

carina complete;

pronotum shiny,

granulated; metanotum

pronotal shiny,

tarsal

radial with

vein

in following

with

without

shorter than

proportions:

9:

located furter distally

segment 5 of fore tarsi

apex

without

weakly

sl ightly

1 bristle

with

almost

fore-wings hyaline,

segments 90)

smooth, tubercles

smooth,

propodeum shiny,

weak;

part of

(Fig.

prominence;

rows of nearly 40

occipital

8;

head shiny,

frons weakly

notaulices almost complete,

with sculpture very

(10:

24;

7:

keels;

TL

the posterior margin of scutum;

reticulate rugose, dark

11:

~

scutum,

sculpture or weakly reaching

15:

median

OOL

an anterior transversal reaching

6:

without sculpture among punctures;

(Fig.

a group of nearly 5

90)

with 2

lamellae;

tibial

spurs 1,1,2. MALE: unknown LOCUS TYPICUS:

Nova Teutonia

TYPICAL MATERIAL: HOSTS:

(Brazil)

holotype F!

and

1 paratype F!

in 8M.

unknown

DISTRIBUTION:

Neotropic region:

BRAZIL:

Nova Teutonia

(Sta Catarina),

BM!

4. Deinodryinus nigrorufus n.

FEMALE:

fully

winged;

length 3,12-3,25 mm;

testaceous;

antennae fully

4-10 dark;

scutum reddish,

anterior margin;

testaceous,

head and prothorax

occasionally

without or with 2

scutellum fully

black,

=p.

reddish-

with segments 3-10 or

lateral

brown spots near the

occasionally fully

or partly

reddish; metanotum and propodeum black; mesosternum, metasternum, mesopleura reddish; metapleura black, with reddish central spot; abdomen brown; femora

legs testaceous,

with

hind coxae brown;

and extremities of hind tibiae brown;

occasionally

hind clubs of

antenna I segments

in

following proportions: 10: 5: 13: 8: 7: 7: 7: 5: 6: 9; head dull, smooth, granulated, without areolae or keels; occipital carina complete; frons without median or ~

8;

lateral

pronotum dull,

impression; smooth,

pronotal

very

weakly

with

a

tubercles reaching granulated;

posterior margin of scutum; without sculpture;

POL ~ 4;

keels;

granulated,

OL ~ 2;

weak

tegulae;

notaul ices

scutellum

propodeum dull,

OOL ~

10;

OPL ~ 7;

TL

anterior transversal scutum hairless,

incomplete,

and metanotum shiny,

reticulate rugose,

shiny,

almost reaching

hairy;

the

smooth and dorsal

surface

with wide areolae; posterior surface with small areolae; no longitudinal keel on posterior surface; fore-wings hyal ine, without dark transversal '::lands; tarsal (Fig.

distal

part of radial

segments 91)

with

vein

longer than

in following proportions: 1 bristle

8:

proximal

3:

located further distally

5:

13:

than

part 21;

(11 :9);

fore

enlarged claw

the proximal

160

Mem.

Fig.

prominence; of nearly 31 spurs

1,

MALE:

1,

fully

91

-

Chela of

Amer.

Ent.

no.

37,

1984

Deinodryinus nigrorufus n. sp.

segment 5 of fore tarsi lamellae;

Inst.,

apex with

a

(Fig.

91)

(type)

double-S shaped,

with 2 rows

group of nearly 4 lamellae;

tibial

2. winged;

length 2,81

mm;

black;

mandibles and antennae

black; legs brown; antennae not distally thickened; antenna I hairs shorter than breadth of segments; antenna I segments in following proportions: 8: 6:

9,5:

10:

9,5:

9:

9:

9:

without sculpture among occipital scutum,

8:

9;

head shiny,

punctures;

carina complete;

POL

=

7;

=

Ol

scutellum and metanotum shiny,

sculpture among punctures;

smooth,

finely

punctate,

frons without median or 2;

=

OOl

smooth,

6,5;

finely

notaulices complete,

lateral

OPl

=

5;

punctate,

posteriorly

Tl

keels;

=

5;

without

separated;

propodeum dull, reticulate rugose, without transversal or longitudinal keel s; fore-w i ngs hya line, without dark transversa I bands; d i sta I part of radial

vein slightly

shorter than proximal

92) with long apical branch wrapping lOCUS TYPICUS: Puyehue (Chile) TYPICAL MATERIAL:

3 para types FF!

HOSTS:

unknown

DISTRIBUTION:

in OT;

3paratypes

Neotropic region:

Santiago Province), OT!

Pichinahuel

OT!

Ol!

TU!

(Cord.

Cobquecura

Puyehue

CHilE:

(2 FF,

Maipu

(Ososno),

Nahuelbuta,

(Nuble),

(9:10);

tibial

and 1 paratype F!

holotype F!

in TU;

part

penis;

in CA;

1 M)!

Ol!

OT!

(Fig.

5. Deinodryinus iphias n. sp.

Que.

Tregualemu

Ol!

OT!

1 paratype F!

in Ol.

(Rinconada,

CAl

Arauco),

gonoforceps

spurs 1,1,2.

la Reina

la Plata, (Maule), (Santiago),

Anteoninae

Fig.

92 -

Male genitalia of Pichinahuel

FEMALE:

fully

testaceous;

winged;

Deinodryinus nigrorufus n. sp. from

(Chile):

right half removed

length 3,75 mm;

antennae brown,

161

black;

with hind coxae and clubs of femora partly margin of pronotum testaceous; segments

clypeus and mandibles

with segments 1-3 testaceous; antennae distally

in following proportions:

12:

7:

legs

brown or black;

17:

12:

testaceous,

posterior

thickened;

antenna I

10:

10:

10:

9:

9:

10;

head dull, punctate and granulated, without areolae or keels, smooth; frons without median and lateral keels; occipital carina complete; region behind ocelli

without

occipital carina; shiny,

with

reaching

a

strong

tegulae;

transversal nearly

as

weakly

granulated;

transversal

smooth

as broad;

margin of scutum; sculpture;

OL = 4;

OOL = 8;

impression,

posterior ocelli

OPL = 8;

TL = 7;

with pronotal

posterior surface of pronotum rugose,

striae, long

two oblique keels connecting

POL = 4;

near. the posterior margin; scutum shiny,

notaulices

smooth,

incomplete,

punctate,

almost reaching

without

tubercles

with some

posterior surface

finely

scutellum and metanotum shiny,

propodeum reticulate rugose,

to

pronotum

smooth,

transversal

very

the posterior without or

longitudinal

keels; posterior surface fully reticulate rugose; fore-wings with two weak dark transversal bands, one on the basal cells and another beneath the distal

part of radial

fore tarsal

pterostigma;

segments

in following

claw

93)

(Fig.

prominence;

with

vein shorter than proportions:

11:

3:

1 bristle located further distally

segment 5 of fore tarsi

(Fig.

93)

with

proximal 7:

20:

than

part

27;

(12:15);

enlarged

the proximal

two rows of 39

162

Mem.

lamellae;

apex with a

fully

winged;

antennae brown,

group of 5

no.

37,

11:

thickened;

antenna I segments

11:

not reticulate rugose,

tibial

black;

1984

11:

13;

spurs 1,1,2.

legs testaceous,

without keels,

brown;

antennal dull,

with

fore and

hind tibiae

hairs

in following

head smooth,

(type)

mandibles testaceous;

clubs of femora partly

antennae not distally

10: 11:

lamellae;

segment 1 testaceous;

brown;

breadth of segments;

17: 9:

Ins!.,

length 2,68-3 mm;

with

hind coxae partly darkened;

En!.

93 - Chela of Ueinodryinus iphias n. sp.

Fig.

MALE:

Amer.

longer than

proportions:

punctate,

13:

7:

granulated,

except for a short keel

between eyes

and antenna I sockets; frons without median and lateral keels; region behind ocell i without two obi ique keels connecting posterior ocell i to occipital = 8;

carina;

TL = 7;

incomplete,

occipital

without

smooth,

distal (Fig.

94)

with

or

sculpture;

longitudinal

fore-wings hyal ine,

part of radial very

vein

keels;

reduced

apical

branch;

unknown

OPL

scutellum and reticulate

posterior surface fully

without dark

shorter than proximal

OOL = 8;

notaul ices

propodeum dull,

transversal

part

tibial

LOCUS TYPICUS: Campina Grande (Brazil) TYPICAL MATERIAL: holotype F! and 1 para type M! in OL. HOSTS:

OL = 3;

punctate;

the posterior margin of scutum;

without

transversal

reticulate rugose;

POL = 6;

weakly granulated,

almost reaching

metanotum shiny, rugose,

carina complete;

scutum shiny,

(9:14);

bands;

gonoforceps

spurs 1,1,2. in TW;

1 paratype M!

Anteoninae

DISTRIBUTION:

Neotropic region:

Serra do Caraca

Fig.

94 -

(5.

Barbara,

BRAZIL:

163

Campina Grande

Minas Gerais),

(Curitiba),

TW!

OL!

Male genitalia of Deinodryinus iphias n. sp. Grande (Brazi I): right half remov",C;.

from

Campina

6. Deinodryinus bicolor (Olmi and CurradoJ n. comb.

Lonchodryinus bicolor 0 I m i and Currado 1979a: 340.

FEMALE: fully winged; length 3,75-5,81 mm; head black, with mandibles black or yellow; antennae yellow, with segments 6-10 or 7-10 brown-black; segment 3 brownish; black;

pronotum,

segment

scutum,

1 with a brown

scutellum,

sometimes anterior collar of pronotum black; propodeum black; brown or black;

dorsal

metanotum

mesopleura,

somet i mes mesop I eura part I y legs testaceous,

with a

streak;

dorsal

propleura

reddish-testaceous; red;

metapleura and

pet io I e black;

basal

abdomen

black spot on

hind

164

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

I

Fig.

coxae;

95 -

Chela of Deinodryinus bicolor (Olmi

tegulae testaceous;

antennal

and Currado)(type)

segments in following proportions:

12:

7: 15: 7: 7: 7: 7: 6: 5: 8; head shiny, smooth, hairy on frons and clypeus, without sculpture; frons with 2 longitudinal parallel keels connecting ocellar region region

to clypeus and with keel

behind ocellar triangle with

around the orbits;

two obi ique keels connecting posterior

Anteoninae

ocelli

to occipital

very 15;

strong

carina;

and keel

pronotum shiny,

scutum,

smooth,

scutum;

=

3;

small

longer than proximal segments

claw

95)

(Fig.

with

approximately 34

testaceous; distally

legs

segments

part;

with

distal

distal

apex

with

length 3,5 mm; with

with

hairs

frons with

longitudinal

reticulate rugose;

two

keels;

vertex

a

length of

2,5:

with

5:

8:

11:

lateral

an

19;

than

the proximal

lamellae;

tibial

mandibles and antennae brown;

antennae not

11:

10:

11:

11:

10:

antenna I

10:

keels around the orbits and

frons weakly

angle;

enlarged

than breadth of segments; 7:

without

rectilinear,

three rows of

hind coxae basally

longer

smooth,

vein

group of nearly 40

black;

TL

(areolae very

parts forming

9:

95)

15;

the sides;

without sculpture;

located further distally

in following proportions:

head shiny;

hairy on

part of radial

(Fig.

carina

=

OPL

approximately 0,30

and proximal

two bristles

testaceous,

thickened,

strongly

in following proportions:

lamellae;

spurs 1,1,2. MALE: fully winged;

15;

surface reticulate rugose

segment 5 of fore tarsi

prominence;

occipital

=

OOL

mesopleura and metapleura shiny,

fore-wings hyaline,

fore tarsal

hollow; 3;

smooth,

reaching

dorsal

posterior surface,

=

OL

finely punctate,

incomplete,

propodeum hairy;

sculpture;

median

with a

POL

scutellum and metanotum shiny,

notaulices distinct, wide);

vertex

shaped;

165

rugose among

not reticulate rugose,

12;

8: with

the keels,

two

not

punctate and without

sculpture among punctures; region of head behind ocell i with two obi ique keels connecting posterior ocelli to occipital carina; vertex with some deep furrows

= 9;

around

OPL

finely

the ocell i;

= 9;

TL

punctate,

complete,

surface smooth distal

(12:11); broad

10;

without

and shiny;

transversal vein

longer than

holotype F!

Quincemil

(Marcapata),

sl ightly

fully

part of frons,

winged;

with

96);

posterior

transversal

longer than proximal (Fig.

OOL

smooth,

propodeum

keels,

without dark

part

gonoforceps with

spurs 1,1,2.

in GE. COLOMB I A:

TW!

BRAZIL:

Meta

length 5,31-7,50 mm;

(Cord.

Itajtuba

7. Deinodryinus maximus n.

FEMALE:

OL = 3;

notaul ices almost

longitudinal

gonoforceps

(Fig. 96); tibial I tajtuba (Braz i I)

HOSTS: unknown 01 STR I BUT I ON: Neotrop ic reg ion: PERU:

or

fore-wings hyal ine,

branch

TYPICAL MATERIAL:

punctures;

the posterior margin of scutum;

part of radial

LOCUS TYP I CUS:

POL = 5;

carina complete;

scutellum and metanotum shiny,

without sculpture among

penis much

apical

occipital

scutum,

almost reaching

reticulate rugose, bands;

=

8M!

GE!

sp.

head black,

clypeus and mandibles testaceous;

Macarena),

(Para),

with

anterior

antennae brown,

with

segmen ts 1-3 or 1-5 testaceous; thorax and propodeum black; tegu I ae testaceous; abdomen black; legs with coxae black (distally testaceous), trochanters

testaceous,

clubs of femora black,

stalks of femora,

tibiae and

166

Mem.

Fig.

96 -

Male genitalia of Quincemil

tarsi

testaceous

segments

in

head shiny,

finely

punctate,

clypeus;

clypeus with the orbits;

a

=

smooth,

right

5;

OL

=

finely

5;

=

OOL

25:

10:

27:

punctate,

impression;

tubercles reaching

without

14:

19;

keel;

scutum

TL

=

shiny,

head

with

17;

strong

with

to keels

carina

to occipital carina; lateral

20;

longitudinal

pronotum

smooth,

incomplete,

punctures;

13: frons

anterior ocellus

hairy,

punctures, finely

with

furrow;

shiny, anterior

pronotal

punctate,

reaching

scutellum and metanotum shiny,

without sculpture among

13:

and occipital

sculpture among

notaulices

antennal 14:

posterior disc centrally excavated;

tegulae;

length of scutum;

=

17:

keels connecting

triangle with

OPL

brownish);

20:

posterior ocell i

ocellar

sculpture among punctures; punctate,

1984

sculpture among punctures;

longitudinal

between

18;

37,

half removed.

tibiae distally

without

median

carina complete;

transversal

0,8

no.

2 oblique keels connecting posterior ocelli

occipital POL

(Peru):

and parallel

region

Ins!.,

Deinodryinus bicolor (Olmi and Currado) from

(except for hind

longitudinal

with

En!.

following proportions:

3 median around

Amer.

without

approximately

smooth,

finely

propodeum with posterior

Anteoninae

surface smooth,

shiny,

strongly

with 2-3 rows of areolae anterior surface dull finely dark

punctate;

dorsal

surface reticulate rugose,

in front of posterior surface;

and

strongly punctate;

metapleura dull

transversa I bands,

punctate;

167

fu II y

part of radial

vein

nearly

Fig.

97 -

Chela of

and strongly

darkened;

as

mesopleura with

posterior surface shiny and punctate;

hind

long as proximal

fore-wings without

wings a I so darkened; part;

fore tarsal

Deinodryinus maximus n. sp. (type)

d i sta I

segments

168

Mem.

in following proportions:

Amer.

12: 4:

6:

En!.

Ins!.,

15:

26;

one bristle located further distally than 5 of fore tarsi lamellae;

(Fig.

97)

LOCUS TYPICUS:

Gourdonville

DISTRIBUTION: Tingo Maria

1984

enlarged claw

(Fig.

97)

prominence;

tibial

with segment

with one row of nearly 23

with a group of nearly 22 lamellae;

with a group of nearly 9 lamellae; MALE: unknown TYPICAL MATERIAL: HOSTS: unknown

37,

the proximal

double-S shaped,

intermediate region

no.

apex

spurs 1,1,2.

(French Guyana)

holotype F!

Neotropic region:

(Monzon Valley),

in P;

1 paratype F!

FRENCH GUYANA:

in CA.

Gourdonville,

P!

PERU:

CA!

8. Deinodryinus melanocephalus (Cameron) n. comb.

Dryinus melanocephalus Cameron 1888: 446. Bocchus melanocephalus (Cameron): Kieffer 1914b: 46.

FEMALE:

fully

winged;

mandibles testaceous;

length 3,75 mm; antennae brown,

head black, with

with clypeus and

segments 1-4 testaceous;

thorax

and propodeum testaceous, with brown spot near tegulae; petiole and abdomen bl ack-brown; fore legs testaceous, with coxae and basa I part of clubs of femora brown; mid legs brown; hind legs brown, with coxae and proximal

half of femora

testaceous;

in following proportions: fully

reticulate rugose,

longitudinal keels

there

parallel is a

13: 6: with

tegulae testaceous;

11: 7:

7:

7:

a strong keel

6:

around

keels connecting ocell i

6:

segments

head dull,

the orbits;

frons

between

carina;

hairy, with 2

these

vertex behind ocelli

to occipital

carina complete and strongly carinate; POL 9; TL = 11; pronotum shiny, smooth, hairy,

antennal 8;

to clypeus;

track of another median keel;

2 obi ique keels connecting posterior ocell i

6:

without

occipital

=

5; OL = 3; OOL = 12; OPL finely punctate, with a

=

transversal impression; pronotal tubercles reaching tegulae; scutum, scutellum and metanotum shiny, smooth, finely punctate, without sculpture among punctures;

notaulices distinct,

0,75 length of scutum; transversal without on

between

longitudinal

the basal

radial

keel

incomplete,

propodeum shiny, dorsal

keels;

reaching approximately

with weak keels,

and posterior surface;

without a

posterior surface

fore-wings with 2 dark transversal

cells and another beneath

vein nearly as long as proximal

the pterostigma; part

(8,5:9);

distal

bands,

one

part of

pterostigma

long and

narrow; fore tarsal segments in following proportions: 12: 5: 9: 14: 30; segment 5 of fore tarsi (Fig. 98) with two rows of 34 lamellae; apex with a group of 8 lamellae;

enlarged claw

bristle; segment 3 of fore MALES: unknown LOCUS TYPICUS:

tarsi

(Fig.

98)

with a hook;

Volcan de Chiriqui

(Panama)

with one medial

tibial

spurs 1,1,2.

peg-like

169

Anteoninae

Fig.

98 -

Chela of Deinodryinus melanocephalus

TYPICAL MATERIAL: FF! in BM. HOSTS:

lectotype F!

(hereby

designated)

(Cameron) (lectotype)

and 2 paralectotypes

unknown

DISTRIBUTION: 3M!

Neotropic region:

PANAMA:

Volcan de Chiriqui,

BM!

Bugaba,

170

Mem.

Fig.

99 -

Amer.

Chela of

En!.

Ins!.,

no.

37,

1984

Deinodryinus noyesi n. sp. (type)

9. Deinodryinus noyesi n. sp. FEMALE:

fully

wingedj

and clypeus

testaceousj

6-10 brownj

legs brown,

length 4,12-5,40 mmj antennae testaceous, with

blackj

mandibles,

malar space

with segments 8-10,

fore tibiae and fore tarsi

7-10 or

testaceous;

An teon i nae

sometimes also fore coxae segments

ha irs much

longer

numerous areolae, behind ocell i particularly carina;

testaceous;

in following proportions:

171

antennae distally

13:

7:

20:

8:

than breadth of segmen ts; in addition

to 3 median

not reticulate rugose,

with

8:

thickened;

8:

7:

7:

head sh i ny;

and 2 lateral

numerous

with 2 oblique keels connecting

7:

frons keels;

longitudinal

posterior ocelli

antenna I

9;

antennal

with region keels,

to occipital

temples finely punctate and without sculpture among punctures;

occipital

carina complete;

pronotum hairy,

shiny,

=

POL

with an

3;

posterior surface of pronotum weak I y transversal tegulae;

=

OL

3;

scutellum

pronotal

and metanotum shiny,

without sculpture among punctures;

12;

OPL

=

transversal

10;

TL

=

11;

furrow;

granu I a ted and with numerous

keels mainly on anterior half;

scutum,

=

OOL

anterior strong

notaul ices

tubercles reaching

smooth,

finely

incomplete,

punctate,

reaching

approximately 0,65 length of scutum; propodeum reticulate rugose; posterior surface with 2 incomplete longitudinal keels; median area smooth; forewings with 2 dark beneath proximal 15:

34;

than

transversal

the pterostigma; (10,5:8);

fore tarsal

enlarged claw

the proximal

rows of nearly 30

(Fig.

fully

segments

prominence;

winged;

one on

the basal

part of radial

99)

lamellae;

segment 3 of fore tarsi MALE:

bands,

distal

with

vein

in following proportions:

1 bristle a

into a

99)

(Fig.

group of nearly 26 hook;

length 3,20-3,37 mm;

9: 2: 9:

located further distally

segment 5 of fore tarsi

apex with

produced

cells and another

I ittle longer than

tibial

black;

with 2

lamellae;

spurs 1,1,2.

mandibles and

antennae

fully testaceous; legs testaceous, with hind coxae proximally black; antennae not distally thickened; antenna I segments in following proportions:

14:

6,5:

14:

10:

10:

10:

10:

10:

10:

12;

antenna I hairs much

longer than breadth of segments; head ret icu I ate rugose; frons with 3 median and 2 lateral keels around the orbits; occipital carina complete; frons

(in front of anterior ocellus)

than breadth of ocellus;

with

a

transversal

furrow

region behind posterior ocelli

connecting posterior ocelli

to occipital

carina;

POL

=

less broad

with 2 oblique keels

5;

OL

=

3;

OOL

=

10;

OPL = 9; TL = 7; scutum, scutellum and metanotum shiny, smooth, strongly punctate, without sculpture among punctures; notaul ices almost complete, almost

reaching

the posterior margin of scutum

reticulate rugose, surface;

few

median

area

irregular keels;

distal

part of radial

proximal tibial

part;

(Fig.

with 2 incomplete longitudinal almost

fully

smooth,

fore-wings hyaline, vein

gonoforceps

nearly

as

(Fig.

100)

not

157 B);

reticulate

without dark

rugose,

transversal

long as or sl ightly with apical

propodeum

keels on posterior

branch

shorter

with

a

bands; than

wrapping penis;

spurs 1,1,2.

LOCUS TYP ICUS:

SI. Augustine (Trinidad) paratyoe M! in TYPICAL MATERIAL:holotype F! and 1 paratype M! in BM; LE; 1 paratype F! in WA; 6 paratypes (5 FF, 1 M)! in AM; 3paratypes (2 FF, 1 M)! in OL. HOSTS: unknown DISTRIBUTION: Curepe

Neotropic region:

(Sta Margarita),

BM!

TRINIDAD:

SURINAME:

SI.

Krepi

Augustine

(SI.

(Charlesburg,

George),

BM!

Paramaribo),

172

Mem.

Fig.

100 -

Amer.

Male genitalia of

Ent.

VENEZUELA:

no.

37,

1984

Deinodryinus noyesi n. sp. from Curepe

(Trinidad):

LE!

Inst.,

left half removed.

Ocumare de La Costa

(Araqua),

WA!

BOLIVIA:

Baures (Rio Itenez, Beni), AM! Ol! laranjeiras (Dept. Beni), NOTE: the species is named in honor of Mr. J.S. Noyes.

Mouth of Rio AM!

10. Deinodryinus atlanticus n. sp.

FEMALE:

fully

winged;

testaceous with ctypeus

and

a

anterior region

segments 8-10 dark; with black spots on pronotum

and

length 4,37-5,00 mm;

head fully

large black spot on frons and vertex; of frons

testaceous;

mandibles,

antennae

testaceOU5,

with

thorax and propodeum testaceous-reddish, sometimes the sides of pronotum, on pleura, between scutum and

between metanotum and propodel1m;

black-testaceous;

black or reddish-

legs testaceous;

petiole black;

antennae distally

thickened;

abdomen antennal

Anteoninae

Fig.

segments

101

-

173

Deinodryinus atlanticus

Chela of

in following proportions:

16:

8:

20:

12:

10:

n.

sp.

9:

9:

(type)

9:

8:

11;

head

dull, fully reticulate rugose; frons with a complete median keel; region behind ocelli with 2 slightly visible oblique keels connecting posterior ocelli

to occipital

OOL = 13; pronotal

occipital

TL = 10;

tubercles reaching

weak dorsal shi ny,

carina;

OPL = 9;

lobes parallel

smooth,

notaulices

fi nel y

incomplete,

carina complete;

pronotum shiny,

tegulae;

OL = 3;

not rugose,

with

posterior surface of pronotum with

to the sides;

punctate,

POL = 3,5;

smooth,

scutum,

two

scutellum and metanotum

wi thout scu I pture among punctures;

almost reaching

the posterior margin of scutum;

propodeum reticulate rugose, without transversal or longitudinal keels; posterior surface with median area smooth, not rugose; fore-wings with dark

transversal

pterostigma;

bands,

distal

one on

the basal

part of radial

vein

cells and another beneath

longer than proximal

part

two

the

(13:9);

fore tarsal segments in following proportions: 12: 3: 9: 13: 30; segment 3 of fore tarsi produced into a hook; enlarged claw (Fig. 101) with one bristle

located further distally

fore tarsi

(Fig.

lamellae; tibial MALE: unknown

101)

than

the proximal

with 2 rows of 28

prominence;

segment 5 of

apex with a

group of 14

spurs 1,1,2.

LOCUS TYPICUS: Nova Teutonia (Brazil) TYPICAL MATERIAL: holotype F! in OT;

."!

lamellae;

1 para type F!

in TW;

1 paratype

in OL.

'-IOSTS:

unknown

JISTRIBUTION:

Neotropic region:

BRAZIL:

Nova Teutonia

(Sta Catarina),

174

OLl

Mem.

OT!

S.

Amer.

En!.

Ins!.,

no.

J. Barreiro (Serra da Bocaina),

TW!

37,

1984

11. Deinodryinus achterbergi n. sp.

fully

mandibles and almost fully frons

black;

winged;

length 4,18-4,31

FEMALE: clypeus,

antennae testaceous,

propodeum black; black,

abdomen

brown;

thickened;

head black,

testaceous;

legs

testaceous,

and distal

antennal

with malar space,

ventral

segments 8-10 brown;

with

stalk of hind femora brown

antennae distally

mm;

with coxae partly

apex of hind

segments

face of head

thorax and

in following

tibiae brown; proportions:

14: 7: 19: 13: 9: 9: 8: 8: 7: 11; head fully reticulate rugose shiny; with 3 median longitudinal keels; 2 lateral keels around the orbits;

frons

occipital carina complete; region behind posterior ocelli reticulate rugose and with 2 oblique keels connecting posterior ocelli to occipital carina; POL = 3;

OL = 3;

DOL = 11;

OPL = 10;

strong anterior transversal punctate, pronotal shiny,

furrow;

without sccdpture among punctures, tubercles reaching

smooth,

tegulae;

finely punctate,

notaul ices almost complete, median

area

with

pronotum hairy,

scutum,

reaching

anterior

finely lobes;

scutellum and metanotum the punctures;

the posterior margin of scutum;

with 2 longitudinal

an

with a

smooth,

2 weak parallel

with

without sculpture among

almost

propodeum reticulate rugose, surface;

TL = 9;

posterior surface shiny,

keels on posterior

transversal

keel,

smooth;

fore-wings

with 2 dark transversal bands, one On the basal cells and another beneath the pterostigma; distal part of radial vein longer than proximal part (16:11); fore tarsal segments in following proportions: 11: 3: 7: 15: 30; enlarged claw (Fig. 102) with 1 bristle located further distally than the proximal prominence; segment 5 of front tarsus (Fig. 102) with 2 rows of nearly 29 lamellae; apex with a group of 12 lamellae; tibial spurs 1,1,2. MALE:

unknown

LOCUS TYPICUS:

Gamboa

TYPICAL MATERIAL: HOSTS:

NOTE:

and 1 para type F!

in LE.

unknown

DISTRIBUTION: Zone),

(Panama)

holotype F!

Neotropic region:

PANAMA:

Gamboa

(Pipeline Rd.,

Canal

LE! the species

is named

in

honor of Mr.

Kees van Achterberg.

12. Deinodryinus peruvianus n. sp.

FEMALE: and

fully

somet i mes

winged;

length

4,68-5,12 mm; black;

anterior reg ion of frons

with segments 7-10,

6-10 or 3-5;

with coxae fully black,

testaceous j

mandibles,

antennae

segments 9-10 brown;

clypeus

testaceous,

legs testaceous,

clubs of femora and mid and hind tibiae partly

175

Anteoninae

Fig.

102

-

Chela of Deinodryinus achterbergi n.

sp.

(type)

176

Mem.

black; distally 13:

11:

Amer.

Ent.

Inst.,

sometimes fore clubs of femora thickened; 10:

8:

with 3 median

Fig.

8:

antennal 7:

10;

head shiny,

longitudinal

103 -

segments keels;

2

fully

no.

37,

1984

testaceous;

antennae slightly

in following proportions:

14:

8:

reticulate rugose and punctate; lateral

keels around

Chela of Deinodryinus peruvian us n.

the orbits;

sp.

(type)

18: frons

Anteoninae

occipital

carina complete;

posterior ocelli Tl = 16;

vertex with 2 obi ique strong keels connecting

to occipital

pronotum hairy,

punctures;

pronotal

transversal

furrow;

carina;

dull,

punctures;

margin of scutum;

tubercles reaching smooth,

dorsal finely

one on

part of radial

front

distal

tarsus

proximal

in following proportions:

(Fig.

103)

(Fig.

prominence;

with basal

103)

apex with a

group of 12

tibial

spurs 1,1,2.

MALE:

fully

Fig.

winged;

lamellae;

with

blaCk;

7:

13:

(Peru):

strong scutum,

the posterior between dorsal

posterior surface

another benea th 30;

the

part; part

(13:17); than

with 2 rows of 28

mandibles,

the

lamellae;

produced

antennae and

antennae not distally

right half removed.

fore

segment 5 of

apical

104 - ·Male genitalia of Deinodryinus peruvianus n. Maria

13;

1-2 dark

segment 2 of fore tarsi

length 3,0 mm;

with hind coxae brown;

keel

located further distally tarsus

=

without sculpture

long as proximal 5:

OPl

lobes;

areolaej

part shorter than

1 bristle

segment 5 of front

hook;

testaceous,

with

dorsal

punctate,

wide

11:

12;

pronotum with a

fore-wings as

=

OOl

almost reaching

the basa I cell s and vein

4;

without sculpture among

transversal

surface with punctate;

transversa I bands,

enlarged claw

finely

propodeum with a strong

almost smooth,

segments

=

Ol

tegulae;

notaul ices almost complete,

pterostigma; tarsal

3;

posterior surface with 2 small

and posterior surface; hairy,

=

POL

finely punctate,

scutellum and metanotum shiny, among

177

into a legs

thickened;

sp.

from Tingo

178

Mem.

antennal 11;

segments

head shiny,

median

Ent.

Ins!.,

no.

37,

in following proportions:

10:

6:

reticulate rugose or with

longitudinal

carina complete; ocelli

Amer.

keels;

vertex

to occipital

2

keels around

= 5,5;

POL

longer than

and metanotum shiny,

smooth,

propodeum with a

punctate,

longitudinal with

transversal

(8:

10);

bands;

spurs 1,1,2. lOCUS TYPICUS:

104)

Tingo Maria

8:

8:

with 2

occipital posterior

scutum,

=

Tl

11;

scutell um

the posterior margin

between dorsal

and

posterior surface with shiny,

fore-wings hyaline,

finely

without dark

vein shorter than proximal

with apical

branch wrapping penis;

part tibial

(Peru)

TYPICAL MATERIAL:

holotype F!

paratype F!

1 para type F!

HOSTS:

9:

= 9;

OPl

area almost smooth,

hairy;

part of radial

(Fig.

8;

keel

wide areolae;

median

keels,

distal

in OT;

9:

frons

without sculpture among

strong transversal

keels;

=

almost reaching

surface with

transversal

gonoforceps

9:

the orbits;

OOl

finely punctate,

of scutum; 2 complete

9:

keels;

breadth of segments;

notaul ices almost complete, dorsal

= 3;

Ol

punctures;

posterior surface;

10:

transversal

with 2 obi ique strong keels connecting

carina;

antenna I ha irs much

lateral

1984

and 2 paratypes in AM;

(1M,

IF)!

1 para type F!

in

BM;

in OL.

unknown

DISTRIBUTION:

Neotropic region:

Circular Rd.),

OT!

Tingo Maria,

BM!

(Dept.

AM!

Beni),

COLOMBIA: BOLIVIA:

TRINIDAD:

la Hormiga

Rurenabaque

Curepe

(Sta.

(Mocoa,

(Beni),

Margarita,

Putumayo),

BM!

Ol!

BM!

PERU:

laranjeiras

13. Deinodryinus hymenaeus n. sp.

FEMALE: frons,

fully

vinged;

length 3,87 mm;

head black,

cl ypeus and mand i b les testaceous;

1-2 testaceous;

with anterior region of

antennae brown,

thorax and propodeum reddish-testaceous;

wi th segments petiole black;

abdomen brown, with segment 1 testaceous; legs testaceous; antennae distally thickened; antenna I segments in following proportions: 14: 7: 10:

9:

9:

9:

9:

7:

10;

head shiny,

19:

with frons sculptured by numerous

areolae and some longitudinal keels; frontal line present; regions behind and on the sides of ocell i not reticulate rugose or keeled, punctate and without sculpture among punctures, except for two short obi ique incomplete keels connecting posterior ocelli to occipital carina; occipital carina complete; smooth, nearly

POL

=

3;

Ol

with strong as

long

=

3;

as broad,

tubercles reaching transverse dorsal

lobes, and

=

11;

=

10;

Tl

=

10;

pronotum shiny,

posterior surface of pronotum

without sculpture,

with

pronotal

posterior surface of pronotum without

only

with

metanotum

notaulices almost complete,

OPl

impression;

not rugose,

tegulae;

scutum,

scutellum

OOl

transversal

two dorsal

shiny,

reaching

propodeum reticulate rugose,

without

posterior surface with a central

weak

smooth,

almost

lobes;

the posterior margin of scutum;

longitudinal

smooth

longitudinal

without sculpture;

region;

or transversal

fore-wings with

keels; two dark

179

Anteoninae

Fig.

105 -

Chela of Deinodryinus hymenaeus n.

sp.

(type)

180

Mem.

Amer.

En!.

transversa I bands,

one on

pterostigma;

part of radial

fore

tarsal

distal segments

enlarged claw proximal 26

lamellae;

(Fig.

HOSTS:

105)

37,

1984

the basa I ce II s and another benea th vein nearly as

with

1 bristle

group of 8

Quatro Barros

the

long as proximal

11:

2,5:

5,5:

11:

part;

26;

located further distally

than

tarsus

two rows of

segment 5 of front

apex with a

TYPICAL MATERIAL:

no.

in following proportions:

prominence;

MALE: unknown lOCUS TYPICUS:

Ins!.,

lamellae;

(Fig. tibial

105)

with

the

spurs 1,1,2.

(Brazil)

holotype F!

nv.

in

unknown

DISTRIBUTION:

Neotropic region:

BRAZil:

Quatro Barros

(Curitiba),

TW!

14. Deinodryinus colombian us n. sp.

FEMALE:

fully

testaceous;

winged;

propodeum black; partly

black;

abdomen

with 3 median

testaceous-dark;

15:

and

7:

17:

keels;

temples not

10:

8:

keels,

occipital rugose,

2

antennal

8:

8:

8:

lateral

mandibles

thorax and

legs testaceous,

thickened;

13:

longitudinal

and numerous transversal posterior ocell i

head reddish-testaceous;

with segments 8-10 dark;

antennae not dis'tallY

following proportions: rugose,

length 4 mm;

antennae testaceous,

with coxae

segments

10;

in

head shiny;

keels around

carina complete;

region behind

punctate and without sculpture

among punctures; behind posterior ocelli two oblique keels connect posterior ocell i to occipital carina; POL ~ 3; Ol ~ 3; OOl ~ 12; OPl TL

~

10;

furrow;

pronotum hairy,

shiny,

posterior surface finely

punctures, pronotal

with 2 weak dorsal

tubercles reaching

smooth,

shiny,

with a

strong

punctate,

tegulae;

transversal

smooth,

lobes parallel

finely punctate,

frons

the orbits

scutum,

~

10;

anterior

without sculpture among

to the sides of pronotum; scutellum and metanotum

without sculpture among punctures;

notaul ices almost complete, almost reaching the posterior margin of scutum; propodeum with dorsal surface reticulate rugose; posterior surface almost fully

smooth and finely punctate;

bands;

distal

part of radial

vein

fore-wings

with 2 dark

transversal

little longer than proximal

part

(15:14);

fore tarsal segments in following proportions: 12: 3: 8: 13: 33; enlarged claw (Fig. 106) with 1 bristle located further distally than the proximal prominence;

segment 5 of front

lamellae;

apex with

produced

into a

tarsus

(Fig.

a group of nearly

hook;

tibial

18

106)

with 2 rows of nearly 26

lamellae;

fore tarsal

segment 3

spurs 1,1,2.

MALE: fully winged; length 2,81-3,12 mm; black; mandibles and part of c I ypeus testaceous; antennae fu II y testaceous; legs testaceous, wi th coxae partly

black;

antennae not distally

following proportions:

11:

7:

15:

9:

thickened; 9:

much

longer than breadth of segments;

frons

with 3 median

behind ocelli

longitudinal

keels;

9:

9:

antennal

9:

9:

head shiny, 2 lateral

11;

segments antennal

fully

reticulate rugose;

keels around

two oblique keels connect posterior ocelli

in hairs

the orbits;

to occipital

carina;

Anteoninae

Fig.

106 -

Chela of Deinodryinus colombianus n.

181

sp.

(type)

182

Mem.

occipital scutum,

Amer.

carina complete;

En!.

POL = 5;

Ins!.,

OL = 3;

scutellum and metanotum shiny,

no.

37,

1984

OOL = 11;

smooth,

OPL = 9;

sculpture among punctures;

notaul ices

posterior margin of scutum;

propodeum reticulate rugose,

longitudinal

keels on posterior surface;

an anterior transversal hyaline,

without dark

shorter than

proximal

keel,

transversal part

branch wrapping penis; LOCUS TYPICUS:

Fig.

107 -

Tame

strongly

(Roraima),

TW!

(Fig.

107)

vein

little

with an apical

left half removed.

and 1 paratype M!

in BM;

1 para type M!

in OL.

HOSTS: unknown DISTRIBUTION: Neotropic region: Surumu

fore-wings

part of radial

Deinodryinus colombianus n. sp. from Tame

holotype F!

1 paratype M!

area clearly del imited by

spurs 1,1,2.

(Colombia):

in TW;

gonoforceps

the

with 2 complete

(Colombia)

Male genitalia of

TYPICAL MATERIAL:

distal

without

almost reaching

transversely striate;

bands;

(10:11);

tibial

incomplete, median

TL = 10;

finely punctate,

COLOMBIA:

Tame

(Arauca),

OL!

15. Deinodryinus cascus n. sp.

BM!

BRAZIL:

Anteoninae

FEMALE:

fully

ocellar spot,

winged;

antennae testaceous, testaceous,

length 5,0 mm;

occipital

carina black,

with segment

183

head reddish-testaceous,

ventral

10 darkened;

black;

propodeum black,

posterior surface reddish-testaceous;

black;

pronotum reddish-

with anterior surface and sides partly

and metanotum reddi sh-testaceous;

with brown

face almost fully

scutum,

with med i an

abdomen black;

scutellum area of

legs testaceous,

with

mid and hind coxae and mid and hind clubs of femora partly black; antennae sl ightly proportions:

16:

distally 8:

reticulate rugose; and with

20:

two

antenna I segments

10:

9:

12;

lateral

occipital

in following

head shiny,

weakly

keels around

carina complete;

the orbits region

with 2 obi ique keels connecting posterior ocell i

=

POL

10:

tracks of

tracks of keels;

behind posterior ocell i carina;

12:

11:

frons with

two median

occipital

thickened;

15:

5;

=

OL

4;

=

OOL

13;

=

OPL

12;

TL

=

14;

to

pronotum

hairy, shiny, smooth, finely punctate, without sculpture among punctures; posterior surface of pronotum nearly as broad as long; pronotal tubercles reaching

tegulae;

punctate,

scutum,

scutellum and metanotum shiny,

without sculpture among punctures;

reaching approximately 0,65

length of scutum;

with posterior surface smooth, longitudinal basal vein

keels;

part

the pterostigma; (14:

10);

distal

fore tarsal

9: 25: 44; enlarged claw

4:

11:

posterior surface without

fore-wings with 2 dark transversal

longer than proximal

proportions:

finely

propodeum reticulate rugose,

without areolae;

cells and another beneath

smooth,

notaulices incomplete,

(Fig.

bands,

segments

108)

one on

the

part of radial in following

with one bristle

located further distally than the proximal prominence; segment 5 of front tarsus (Fig. 108) with 2 rows of 43 lamellae; apex with a group of 8 lamellae; tibial MALE: unknown

spurs 1,1,2.

LOCUS TYPICUS:

San Cristobal

TYPICAL MATERIAL: HOSTS:

de

holotype F!

las Casas

(Mexico)

in OT.

unknown

DISTRIBUTION: (Chiapas),

Neotropic region:

MEXICO:

San Cristobal

de

las Casas

OT!

16. Deinodryinus albopictus n. sp.

FEMALE:

fully

winged;

mandibles testaceous; Clrothorax black,

length 6 mm;

:7:

15:

thickened; 13:

12:

,ongitudinal

antennal

10:

keels;

10:

11;

JOL

=

to occipital 12;

OPL

=

head shiny,

carina; 13;

TL

=

without occipital 13;

mesothorax,

legs testaceous;

in fai lowing proportions:

antennae 16:

8:

29:

with frons sculptured by some

region of head behind ocell i

sculpture among punctures, Gcell i

lobes whitish;

abdomen and

segments

with clypeus and

with segments 1-3 testaceous;

with posterior and dorsal

Cletathorax and propodeum black; distally

head black,

antennae brown,

punctate,

without

two obi ique keels connecting carina complete;

pronotum shiny,

POL = 4,5;

smooth,

posterior OL = 2,5;

finely punctate,

184

Mem.

Fig.

108 -

Amer.

Chela of

En!.

no.

twice as broad

prominent dorsal

and transverse

lobes;

tegulae;

scutellum and metanotum shiny,

without sculpture among punctures; approximately 0,80

1984

pronotum with a strong

posterior surface nearly

scutum,

37,

Deillodryillus cascus n. sp. (type)

without scu I pture among punctures; impression;

Ins!.,

pronotal

notaul ices

length of scutum;

as

transversa I

long,

with

two

tubercles reaching smooth,

finely punctate,

incomplete,

reaching

propodeum reticulate rugose,

without

transversal or longitudinal keels, with posterior surface smooth and shiny; fore-wings with two weak dark transversal bands, one on the basal cells and another beneath the pterostigma;

distal

part of radial

vein shorter

185

Anteoninae

part (14:20); fore tarsal segments in following proportions: 14: 3,5: 10: 17: 37; enlarged claw (Fig. 109) with 1 bristle located

than proximal

further dis ta II y

than

the prox i ma I prom i nence;

segment 5 of front

109) with two rows of approximately 45 lamellae; of approximately 10 lamellae; tibial spurs 1,1,2. (Fig.

Fig.

'.1ALE:

109 -

sp.

(type)

unknown

_OCUS TYP I CUS:

Encruzi I hada

-YPICAL MATERIAL: --1JSTS:

Chela of Deinodryinus albopictus n.

tarsus

apex with a group

(Brazi I )

holotype F!

in TW.

unknown

JISTRIBUTION:

Neotropic region:

BRAZIL:

Encruzilhada

17. Deinodryinus amoenus n. sp.

(Bahia),

TW!

186

Mem.

FEMALE:

fully

winged;

Amer.

mid and

legs black;

17:

occipital

not reticulate rugose,

pronotum shiny,

9:

black;

distally

27:

20:

carina;

hairy,

15:

longitudinal

antenna I

15:

13:

16;

keels and by

region behind ocell i

OOl = 11;

Ol = 3;

OPl = 10;

with a strong anterior transversal punctate, lobes;

with 2 dorsal

transverse

without sculpture among punctures;

tubercles reaching

smooth,

finely

notaulices incomplete,

length of scutum;

Tl = 12;

furrow;

without sculpture among

pronotal

scutellum and metanotum shiny,

approximately 0,5

16:

without oblique keels connecting posterior

POL = 4;

punctures,

scutum,

antennae occasionally

thickened;

18:

temples and

posterior surface of pronotum finely tegulae;

occasionally anterior testaceous;

legs testaceous;

numerous

carina complete;

1984

punctate and without sculpture among punctures;

behind posterior ocelli to occipital

37,

mand i b les

antennae slightly

with frons sculptured by

some areolae; region

testaceous;

in following proportions:

head shiny,

ocell i

no.

with segments 3-10 or 6-10 dark;

hind

segments

Ins!.,

length 5,00-5,62 mm;

reg ion of frons and cl ypeus testaceous,

En!.

punctate,

reaching

propodeum reticulate rugose,

except

for the posterior surface almost fully smooth, finely punctate, without scu I pture among punctures; fore-w i ngs darkened benea th the pterost i gma; distal

part of radial

segments 110)

vein

shorter than

in following proportions:

with

1 bristle

segment 5 of front apex with a

12:

proximal 3:

9:

located further distally tarsus

(Fig.

group of nearly

110)

16: than

part 35;

(15:19);

the proximal

with 3 rows of nearly

16 lamellae;

fore tarsal

enlarged claw 52

(Fig.

prominence; lamellae;

segment 3 of fore tarsi

produced

into a hook; tibial spurs 1,1,2. MALE: unknown LOCUS TYP ICUS: Nova Teutonia (Brazi I) TYPICAL MATERIAL: F! in Ol. HOSTS:

in BM;

1 paratype F!

in TlV;

1 paratype

unknown

DISTRIBUTION: BM!

holotype F!

Azul

Neotropic region:

(Minas Gerais),

TW!

BRAZil:

Nova Teutonia

Alto da Serra

(Sta Catarina),

(Morretes),

Ol!

18. Deinodryinus pegnai n. sp.

FE:MAlE:

fully

winged;

length 4,5 mm;

and part of malar space yellow; thickened; 9:

9:

9:

9:

antenna I segments 11;

head du II,

black;

mandibles,

chelae testaceous;

in following proportions:

strong I y

punctate;

sides of clypeus

antennae distally 14:

8:

punctures very

20:

12:

11:

dense and

deep, resembl ing areolae; frons without median and lateral keels; occipital carina complete; region of head behind ocelli without two oblique keels connecting posterior ocell i to occipital carina POL = 7; Ol = 5; OOl = 10; OPl = 10; Tl = 14; region of head behind ocelli punctate, not reticulate rugose or keeled;

pronotum hairy,

shiny,

with a

weak

transversal

impression and with pronotal tubercles reaching tegulae; posterior surface of pronotum smooth, finely punctate, without sculpture among punctures,

Anteoninae

187

I

/

Fig.

110 -

Chela of Deinodryinus amoenus n.

sp.

(type)

Mem.

188

nearly

as

long as broad,

metanotum shiny,

smooth,

Amer.

Ent.

Inst.,

without dorsal finely

lobes;

punctate,

punctures;

notaul ices almost complete,

of scutum;

propodeum reticulate rugose,

no.

37,

1984

scutum,

scutellum and

without sculpture among

almost

reaching

without

the posterior margin

transversal

or

longitudinal

keels, with posterior surface almost fully smooth and shiny; fore-wings with one dark transversal band beneath the pterostigma; distal part of radial vein longer than proximal part (16:11); fore tarsal segments in following proportions: 11: 3: 8: 14: 29; enlarged claw (Fig. 111) with 2 bristles

located further distally

of front

tarsus

a

(Fig.

group of nearly

MALE:

11

111)

with

than

the proximal

lamellae;

tibial

segment 5

lamellae;

apex with

spurs 1,1,2.

unknown

LOCUS TYPICUS:

EI

Canelo

TYPICAL MATERIAL:

(Chile)

holotype F!

in TW.

HOSTS: unknown DISTRIBUTION: Neotropic region:

CHILE:

NOTE:

prominence;

two rows of nearly 38

the species

is named

EI

in honor of Mr.

Canelo

(Santiago),

Luis E.

TW!

Pegna.

19. Deinodryinus pseudoamoenus n. sp.

Fig.

111

-

Chela of

Deinodryinus pegnai n. sp.

(type)

189

Anteoninae

FEMALE:

fully

winged;

length 3,87-5 mm;

testaceous;

clypeus black,

testaceous,

with segments 7-10 dark;

meta thorax and testaceous;

propodeum black;

antennae distally

proportions:

17:

10:

25:

head black;

with anterior margin

12:

prothorax

testaceous;

antennae mesothorax,

abdomen brown-testaceous;

thickened;

antenna I segments

12:

11:

9:

reticulate rugose and with some

(at

least one median)

vertex not reticulate rugose,

mandibles

testaceous;

strongly

9:

9:

11;

punctate;

legs

in

head dull,

following strongly

longitudinal

frontal

keels;

I ine complete;

occipital carina complete; POL = 7; Ol = 3,5; OOl = 13; OPl = 11,5; Tl = 11; pronotum hairy, shiny, weakly rugose, with two dorsal lobes; pronotal tubercles reaching half,

tegulae;

scutum shiny,

without sculpture among

approximately 0,75 sculpture,

punctures;

length of scutum;

punctate mainly on anterior notaulices

incomplete,

scutellum shiny,

smooth,

reaching without

only with SOme punctures near the anterior margin;

metanotum

shiny, smooth, without sculpture; propodeum reticulate rugose, with posterior surface rugose and centrally smooth; propodeum without transversal beneath part 37;

or

longitudinal

the pterostigma;

(15:23);

fore

keels;

distal

tarsal

fore-wings

with a

part of radial

segments

112)

with one bristle

dark

transversal

into hooks;

located further distally

than

11:

3:

9::15:

en I arged claw

the proximal

prominence; segment 5 of front tarsus (Fig. 112) with 2 rows of 33 lamellae; apex with a group of 14 lamellae; tibia! spurs 1,1,2. MALE: unknown LOCUS TYPICUS:

Fig.

112 -

Nova Teutonia

Chela of

band

shorter than proximal

in following proportions:

segments 2 and 3 of fron t tarsus produced

(Fig.

vein

(Brazil)

Deinodryinus pseudoamoenus n. sp.

(type)

190

Mem.

TYPICAL MATERIAL: HOSTS: unknown

holotype F!

DISTRIBUTION: OT!

Amer.

Neotropic region:

Campina Grande

(Curitiba),

Ent.

Ins!.,

no.

37,

1984

in OT;

paratype F!

BRAZil:

Nova Teutonia

in TW. (Sta Catarina),

TW!

20. Deinodryinus townesi n. sp.

FEMALE:

fully

testaceous; yellow;

winged;

mesothorax,

testaceous-brown; coxae;

15:

legs testaceous, 7:

22:

frons without

with small

thickened; 13:

10:

9:

longitudinal

without sculpture among posterior ocelli

head black,

with mandibles

with segments 1-2 testaceous;

prothorax

meta thorax and propodeum redd i sh-testaceous;

antennae distally

proportions: rugose;

length 5 mm;

antennae brown,

antennal 9:

8:

keels;

punctures,

to occipital

brown spots on

8:

in following

head dull,

reticulate

region behind ocell i

without

carina;

segments

9;

occipital

abdomen

mid and hind

punctate,

two obi ique keels connecting carina complete;

POL = 5;

= 3,5;

OOl = 13; OPl= 11; Tl= 10; pronotum shiny, finely punctate, without scu I pture among punctures; pronotum wi th a strong transversa I impression;

posterior surface nearly

prominent dorsal

Fig.

transverse

113 -

lobes;

Chela of

twice as broad as

pronotal

long,

tubercles reaching

with

two

tegulae;

Deinodryinus tOI.rnesi n. sp. (type)

OL

Anteoninae

scutum,

scutellum and metanotum shiny,

sculpture among punctures; 0,75

length of scutum;

longitudinal wi th

two dark

beneath part 31;

keels,

113)

prominence;

tibial

spurs

MALE:

unknown

one on

punctate,

reaching without

transversal

1 bristle

apex with a

proportions:

proximal

12:

3:

9:

located further distally

segment 5 of front

lamellae;

or

fore-wings

vein shorter than

in following

with

without

approximately

the basa I cell s and another

part of radial

segments

(Fig.

rows of nearly 31

tarsus

(Fig.

group of nearly

113) 12

15:

than

with

two

lamellae;

1,1,2.

LOCUS TYP I CUS:

Encruzi I hada

TYPICAL MATERIAL: HOSTS:

distal

fore tarsal

enlarged claw

finely

posterior surface smooth and shiny;

transversa I bands,

(15:23);

smooth,

incomplete,

propodeum reticulate rugose,

with

the pterostigma;

the proximal

notaul ices

191

(Brazi I )

holotype F!

in TW.

unknown

DISTRIBUTION: NOTE:

Neotropic region:

the species

is named

in

BRAZIL:

Encruzilhada

honor of Mr.

(Bahia),

Henry K.

TW!

Townes.

21. Deinodryinus bilobus Fenton

Deinodryinus bilobus Fenton 1927: 11.

FEMALE:

fully

winged;

length 6,87 mm;

clypeus and mandibles testaceous;

black,

with anterior part of frons,

antennae testaceous,

brown;

propectus and pronotum redd i sh-testaceous,

black;

mesothorax,

tegulae testaceous; coxae;

propodeum black;

legs testaceous,

11;

segments head dull,

mid and

in following proportions: hairy,

rugose,

with 3 incomplete longitudinal

tubercles

abdomen brown-redd i sh;

with black spots on mid and

stalks of hind femora black;

antennal 10:

meta thorax,

with segments 7-10

with pronota I

20:

10:

with numerous

median keels;

hind

hind tibiae and tarsi 30:

14:

14:

longitudinal

occipital

brown;

14:

11:

keels;

11:

frons

carina complete;

head without keels around the orbits; POL = 5; OL = 3; OOL = 15; OPL 13; TL= 10; pronotum shiny, smooth, hairy, without sculpture, with an anterior transversal impression; posterior disc with 2 lateral distinct dorsal

lobes;

pronotal

metanotum shiny,

tubercles reaching

smooth,

finely

punctures;

notaulices incomplete,

of scutum;

propodeum reticulate rugose,

smooth,

finely

punctate,

without sculpture; basal

hairy;

vein shorter than proximal 7:

3:

weak,

scutum,

7:

12:

part

26;

located further distally than

scutellum and

without sculpture among

reaching with

approximately 0,65

mesopleura and metapleura shiny, the pterostigma; (15:23);

the proximal

(Fig.

bands,

distal

fore tarsal

enlarged claw

length

posterior surface shiny,

fore-wings with 2 dark transversal

cells and another beneath

proportions:

tegulae;

punctate,

prominence;

the

part of radial

segments

114)

smooth,

one on

with

in following

1 bristle

segment 5 of front

192

Mem.

Fig.

tarsus

(Fig.

114 -

Amer.

En!.

Ins!.,

114) double-S shaped,

Santarem

TYPICAL MATERIAL: HOSTS:

unknown

37,1984

Chel a of lJeinodryinus bilobus Fenton

with 2 rows of nearly

apex with a group of nearly 10 lamellae; MALE: unknown LOCUS TYPICUS:

no.

(Brazil)

holotype F!

in VIA.

tibial

(type)

33 lamellae;

spurs 1,1,2.

Anteoninae

DISTRIBUTION:

Neotropic region:

BRAZIL:

193

Santarem

(Para),

IVA!

22. Deinodryinus insignis n. sp.

FEMALE:

fully

of frons,

winged;

clypeus,

testaceous,

petiole black;

antennae distally

proportions:

15:

8:

rugose and with keels;

ventral

with segments 7-10 brown;

testaceous-reddish; black;

length 4,03-4,62 mm;

mandibles and

two

24:

9:

9:

lateral

keels around

7:

7:

10;

frons with

=

3;

=

OOL

transversal

12;

OPL

=

12;

striae on median

surface; pronotal metanotum shiny, almost reaching

=

TL

10;

region,

in following

head dull,

three median

the orbits;

occipital

fully

reticulate

longitudinal

carina complete;

longitudinal

pronotum shiny,

with

antennae

with hind coxae partly

segments

region behind posterior ocell i with numerous OL

with anterior half

propodeum and abdomen

antennal

7:

irregular keels;

thorax,

legs testaceous,

thickened;

13:

head black,

side testaceous;

two dorsal

keels;

smooth,

lobes on

POL = 3; with

weak

posterior

tubercles reaching tegulae; scutum, scutellum and smooth and without sculpture; notaulices almost complete, the posterior margin of scutum;

propodeum with dorsal

surface reticulate rugose and with posterior surface smooth and without fore-wings with 2 dark

sculpture;

transversal

cells and another beneath

the pterostigma;

longer than

proximal

(24:17);

proportions:

12:

4:

8:

part 12:

27;

bands,

distal

fore tarsal

enlarged claw

one on

segments

(Fig.

the basal

part of radial

vein

in following

115)

with

1 bristle

located further distally than the proximal prominence; segment 5 of front tarsus (Fig. 115) with 2 rows of 26 lamellae; apex with a group of 12 lamellae; MALE:

tibia!

spurs 1,1,2.

unknown

LOCUS TYPICUS:

Sinop

TYPICAL MATERIAL: HOSTS:

(Mato Grosso,

holotype F!

Brazil)

in OT;

1 paratype F!

in AM.

unknown

DISTRIBUTION: BOLIVIA:

Neotropic region:

BRAZIL:

Sinop

(Mato Grosso),

OT!

5 km NIV mouth of Rio Grande (Rio Mamore, Dept. Beni), AM!

23. Deinodr:linus pseudobilobus n. sp.

FEMALE:

fully

winged;

length 4,50-5,62 mm;

head black,

with clypeus and

mandibles testaceous; antennae brown, with segment 1 and partly 2 testaceous; prothorax reddish-testaceous, occasionally black with reddish margins

(in

a

paratype from Horco Molle);

mesothorax,

metathorax and

propodeum black; abdomen black and partly testaceous; legs fully testaceous, occasionally with hind coxae and stalks of hind femora partly black; antennae distally thickened; antennal segments in following :lroportions: 12: 5: 15: 9: 9: 9: 7: 6: 6: 7; head shiny, rugose; frons

194

Mem.

Fig.

115 -

sculptured by strong

Amer.

Chela of

Ent.

no.

37,

1984

Deinodryinus insignis n. sp. (type)

longitudinal

sculpture among keels;

Inst.,

keels;

posterior third of frons without

anterior and median

vertex and region between ocelli

third granulated among keels;

and occipital

carina punctate,

without

keels and without sculpture among punctures; posterior edges of posterior ocelli joint by a transversal furrow; occipital carina complete; POL ~ 5; OL ~ 3,5;

OOL ~ 15;

without sculpture;

OPL ~

TL ~ 10;

13;

pronotum shiny,

sides transversely striate,

on posterior surface;

pronotal

tubercles reaching

scutellum and metanotum shiny, sculpture among punctures;

smooth,

notaul ices,

very

hairy,

dorsally

with 2 prominent dorsal tegulae;

finely punctate,

reaching

lobes

scutum, without

approximately 0,75

length

Anteoninae

of scutum;

propodeum reticulate rugose,

between dorsal longitudinal

Fig.

and posterior surface;

keels,

116 -

with

strong

Chela of

195

without strong

transversal

keels

posterior surface without

transversal

keels,

dull;

fore-wings with 2

Deinodryinus pseudobilobus n. sp. (type)

196

Mem.

dark

transversal

pterostigma; fore

tarsal

claw

(Fig.

prominence; 60

lamellae;

bands,

distal

one on

Ent.

with

Inst.,

the basal

part of radial

segments 116)

Amer.

no.

37,1984

cells and

another beneath the

vein shorter than proximal

in following proportions:

14:

3:

9:

13:

1 bristle arising further distally than

segment 5 of front apex with a

tarsus

(Fig.

group of nearly

24

116)

part

34;

(6:15);

enlarged

the proximal

with some rows of nearly

lamellae;

tibial

spurs 1,1,2.

\

\ \ !

\

II

\ \

d

\ I

I

\1

Fig.

117 -

I J

I

Male genitalia of Deinodryil1us pseudobilobus n. Molle (Argentina):

left half removed.

sp.

from Horco

197

Anteoninae

MALE:

fully

testaceous, brown,

with

th ickened; segments

tarsi

and

trochanters

testaceous;

14: 7:

19:

13:

with frons strongly sculptured by punctate

and

without

carina complete;

scutum shiny, notaulices,

~6;

POL

13:

12:

strongly punctate,

reaching

metanotum shiny,

~

4;

OOL

~

antenna I

10: 9:

punctures;

vertex

to occipital

12; OPL

12;

keels and with

~

12;

carina;

TL

~

11;

without sculpture among punctures;

approximately 0,80

punctate,

10:

longitudinal

sculpture among

OL

legs

antennae not di sta II y

strong oblique keels connecting posterior ocelli

occipital

antennae

1 brown;

an tenna I ha i rs much shorter than breadth of segmen ts;

strongly

without

mandibles testaceous;

sides of segments 2-3-4 and partly

in following proportions:

head shiny, vertex

length 3,87-5,00 mm;

winged;

with dorsal

length of scutum;

scutellum and

without sculpture among punctures;

propodeum

reticulate rugose, without a transversal keel between dorsal and posterior surface; posterior surface reticulate rugose; fore-wings weakly crossed by a transversal dark band beneath the pterostigma; distal part of radial vein shorter than proximal part (9:18); gonoforceps (Fig. 117) with apical branch

wrapping penis;

LOCUS TYPICUS:

TYPICAL MATERIAL: in B;

spurs

1,1,2.

(3 FF,

and 1 paratype F!

holotype F!

2 paratypes FF!

paratypes

tibial

Horco Molle (Argentina)

4 para types

in TU;

4 ,'I,M)!

(3 FF,

in BM;

1 '''1)!

1 paratype F! 7

in TW;

in OL.

HOSTS: unknown DISTRIBUTION: Neotropic region:

BOLIVIA:

Rurenabaque

(Beni),

B~'.!

BRAZIL: Pitfall (Cerrado), OL! Rezende (Estado de Rio), B! Campina Grande (Curitiba), OL! Teodoro Sampaio (Sao Paulo), TW! Serra do Caraca (S. BM!

Barbara, TW!

Minas Gerais),

Las Cejas

OL!

(Tucuman),

ARGaHINA:

OL!

TU!

Horco ,'.,101Ie (Tucuman),

Los Puestos

(Tucuman),

OL!

OL!

24. Deinodryinus rubrolobatus n. sp.

FEMALE:

fully

winged;

length 5,25 mrn;

head

black,

with mandibles and

c I ypeus testaceous; antennae brown, with segment 1 testaceous; prothorax reddi sh; mesothorax, meta thorax and propodeum black; abdomen black, with some regions testaceous; femora, hind tibiae and hind distally

13: 13: keels;

thickened; antenna I segments in following proportions: 17: 8: 24: 12: 9: 8: 7: 9; head shiny, sculptured by numerous longitudinal

region

behind posterior ocell i

temples without keels,

4; OOL

legs testaceous, with hind coxae, hind tarsi partly brown or black; antennae

~12;

OPL ~

punctate;

10; TL

~

8;

with numerous

occipital

pronotum shiny,

without sculpture among punctures;

longitudinal

carina complete; smoooth,

keels;

POL ~ 4;

finely

OL ~

punctate,

posterior surface of pronotum with

two

dorsal transverse lobes; pronotal tubercles reaching tegulae; scutum, scutellum and metanotum shiny, smooth, finely puntate, without sculpture among punctures; of scutum;

notaul ices

incomplete,

propodeum reticulate rugose,

reaching dull,

approximately 0,65

without transversal

or

length

198

Mem.

Fig.

longitudinal rugose;

118 -

keels;

Amer.

Chela of

En!.

Ins!.,

no.

37,

1984

Deinodryinus rubrolobatus n. sp.

posterior surface of propodeum dull,

fore-wings with

two dark

cells and another beneath

transversal

the pterostigma;

bands,

distal

(type)

fully reticulate

one on

the basal

part of radial

vein

Anteoninae

shorter than

proximal

part

proportions:

13: 4:

14:

8:

located further distally tarsus

(Fig.

118)

(12:21); 31;

than

spurs 1,1,2.

LOCUS TYPICUS:

Pompeya

TYPICAL MATERIAL: HOSTS: unknown

tarsal

enlarged claw the proximal

with 2-3 rows of 53

lamellae; tibial MALE: unknown

DISTRIBUTION:

fore

199

segments in following

(Fig.

118)

prominence;

lamellae;

with

1 bristle

segment 5 of front

apex with a

group of

18

(Ecuador)

holotype F!

Neotropic region:

in OT. ECUADOR:

Pompeya

(Napo R.,

Pastaza),

OT!

25. Deinodryinus nigrolobatus n. sp.

FEMALE:

fully

winged;

pronotum fully testaceous,

length 3,75-4,75 mm;

black or with margin

darkened;

legs testaceous,

hind clubs of femora partly antennal 14;

segments

head with

keels;

brown;

black;

testaceous;

occasionally 14:

reticulate rugose,

vertex behind ocell i

with hind coxae and

antennae slightly distally

in following proportions:

frons fully

mandibles testaceous;

antennae brown or

7:

18:

10:

9:

without median

not reticulate rugose,

thickened; 10:

and

9:

9:

strongly punctate,

without sculpture among punctures, without obi ique keels connecting posterior ocell i to occipital carina; occipital carina complete; POL = 4: =3; OOL = 10; OPL = 9; TL = 9 pronotum shiny, with an anterior transversal

furrow;

posterior surface with 2 strong

anterior half strongly

transversely striate;

surface finely punctate, scutum,

not striate;

punctures; propodeum

reaching

reticulate rugose,

lobes;

tubercles reaching

punctate,

posterior surface of pronotum nearly

notaulices incomplete,

transversal

OL

posterior half of the posterior

pronotal

scute II um and metanotum fi nel y

9:

lateral

tegulae;

without scu I pture among

twice as broad as

approximately 0,5

long;

length of scutum;

with posterior surface partly

reticulate rugose

and transversely striate; fore-wings darkened beneath the pterostigma; distal part of radial vein little shorter than proximal part (14:16); fore tarsal (Fig.

segments 119)

with

prominence; lamellae;

in following proportions: 1 bristle

segment 5 of front

apex with a

10:

tarsus

group of 12

(Fig.

or brown;

legs testaceous,

rugose, around ocelli

15:

shiny;

8:

15:

14:

13:

thickened; 13:

frons with 3 median

the orbits;

occipital

10:

than

25;

enlarged claw

the proximal

with 2 rows of nearly 29

segment

black;

3 of fore

tarsi

mandibles and

antennae

with coxae and clubs of femora partly black

antennae not distally

proportions:

6:

119)

lamellae;

produced into a hook; tibial spurs 1,1,2. MALE: fully winged; length 3,75-4,31 mm; testaceous;

3:

located further distally

11:

11:

antennal 9:

12;

longitudinal

carina complete;

segments

head fully

keels;

region

with 2 oblique keels connecting posterior ocelli

2

in following reticulate

lateral

behind

keels

the posterior

to occipital

carina;

200

Mem.

Fig.

POL =7;

119 -

OL = 3,5;

metanotum shiny, antennal complete,

Amer.

Chela of

hairs much

OPL = 6;

punctate,

longer than

almost reaching

Ins!.,

no.

37,

1984

Deinodryinus nigrolobatus n. sp. (type)

OOL = 11; smooth,

En!.

TL = 10;

scutum,

scutellum and

without sculpture among punctures;

breadth of segments;

notaul ices almost

the posterior margin of scutum;

propodeum fully

Anteoninae

201

/

Fig.

120 -

Male genitalia of Teutonia

reticulate rugose, wings hyaline,

without

subapical

proximal

holotype F!

in TU;

2 paratypes MM!

HOSTS:

unknown

in

TW;

DISTRIBUTION:

Neotropic region:

:'Jova Teutonia

(S.ta Catarina),

(Tucuman),

TU!

TW!

las Cejas

bands;

(13:14);

part

Nova Teutonia

TYPICAL MATERIAL:

keels on

transversal

branch wrapping penis;

lOCUS TYP ICUS:

right half removed.

longitudinal

without dark

little shorter than

Deinodryinus nigroiobatus n. sp. from Nova

(Brazi I):

tibial

posterior surface; distal

gonoforceps

fore-

part of radial (Fig.

120)

with

vein a

spurs 1,1,2.

(Brazi I) and 1 para type M! 2 paratypes

PERU: BM!

Ol!

(1

in BM;

F,

M)!

Machu Picchu, ARGENTINA:

(Tucuman),

TW!

1 paratype F!

in Ol. Ol'

BRAZil:

Horco Molle

Ol!

26. Deinodryinus vagans n. sp.

202

Mem.

FEMALE:

fully

winged;

mandibles and clypeus partly 3 testaceous; mesothorax,

Amer.

Ent.

Inst.,

length 4,18-6,00 mm; testaceous;

no.

prothorax reddish-testaceous,

antennae distally

following proportions:

12:

with frons sculptured by region behind ocelli

8:

19:

thickened; 16:

13:

occipital OL

=

=

OOL

puncta te, broad as reaching punctate,

OPL

impression;

long,

with

tegulae;

9;

TL

=

9;

du II;

occipital

two strong

or

transverse dorsal

length of scutum;

longitudinal

vein shorter than proximal

-

carina complete; hairy,

lobes;

region

keels;

two dark

notaulices

part

Chela of

to POL

=

8;

finely strong

twice as

pronotal

scutellum and metanotum shiny,

fore-w i ngs with

121

areolae;

pronotum with a

tubercles

smooth,

finely

incomplete,

propodeum reticulate rugose,

posterior surface fully

transversa I bands,

cells and another beneath the pterostigma;

Fig.

in

head shiny,

keels and with

posterior surface of pronotum nearly

approximately 0,65

rugose,

10;

resembling

pronotum shiny,

without sculpture among punctures;

transversal

basal

line complete;

=

scutum,

without

10:

two obi ique keels connecting posterior ocell i

frontal

13;

10:

longitudinal

wi thout scu I pture among punctures;

transversal

reaching

without

carina;

4;

testaceous-brown; segments

punctate and without sculpture among punctures;

occasionally punctures very strong and broad, behind ocell i

2 and

occasionally darkened;

abdomen 10:

with malar space,

with segments 1,

antennal

13:

nUmerous parallel

1984

head black,

antennae black,

meta thorax and propodeum bl ack;

legs testaceous;

37,

(18:22 or 18:24);

distal

one on

reticulate the

part of radial

fore tarsal

segments

Deinodryinus vagans n. sp. (type)

in

Anteoninae

following proportions: bristle

fore tarsi

(Fig.

of nearly MALE:

12:

located further 15

fully

121)

winged; ventral

9:

20:

37;

than

enlarged claw

the proximal

with 2 rows of nearly

lamellae;

antennae with

3:

distally tibial

203

31

(Fig.

121)

prominence;

lamellae;

with

1

segment 5 of

apex with

a

group

spurs 1,1,2.

length 3,43 mm;

black;

side testaceous,

mandibles partly

dorsal

side brown;

testaceous;

legs testaceous,

with femora, hind coxae, mid and hind tibiae partly brown; antennae not distally thickened, with hairs shorter than breadth of segments; antennal segments

in following proportions:

head shiny,

and with region behind ocelli posterior ocelli

to occipital

sculpture among punctures; complete;

POL =7;

Ol = 4;

and metanotum shiny, punctures;

Fig.

122 -

13:

7:

16:

10:

10:

10:

with frons sculptured by numerous parallel

notaul ices

without

carina; frontal

region behind ocelli OPl = 8:

punctate,

incomplete,

9:

12; keels

Tl = 6;

without

carina

scutum,

scutellum

without sculpture among

almost reaching

gonoforceps

punctate,

occipital

the posterior margin of

Male genitalia of Deinodryinus vagans n. (Brazil)(A);

10:

two oblique keels connecting

I ine complete;

OOl = 10:

smooth,

10:

longitudinal

sp.

from

(laterally viewed)(B).

Teresopolis

204

Mem.

scutum;

Ent.

propodeum reticulate rugose,

posteriorsurface; distal

Amer.

vein

no.

with small

fore-wings hyal ine,

part of radial

Inst.,

37,

1984

smooth area on

without dark transversal

shorter than

proximal

part

(12:

(Fig. 122) with small apical branch \Vrapping penis; LOCUS TYPICUS: Teresopolis (Brazil)

tibial

TYPICAL MATERIAL: holotype F! and 4 paratypes FF! in AM; 4 paratypes (3 FF, 1 M)! in OL. HOSTS: unknown

in TW;

DISTRIBUTION:

Neotropic region:

(Romansos,

Dept.

Teresopolis,

OL!

Jatai

Serra do Caraca ARGENTINA:

Beni), (S.

AM!

(Goias),

Barbara,

Horco Molle

BOLIVIA:

Rio

Ichilo

OU

S.

Junction Rio

(Santa Cruz), J.

Barreiro

Minas Gerais),

(Tucuman),

TW!

bands;

14);

gonoforceps

spurs 1,1,2. paratype F!

Itenez-Rio Paraguay OL!

(Serra

BRAZIL: da Bocaina),

Caruaru,

OL!

TW!

TW!

27. Deinodryinus pilosifrons Fen ton

Deinodryinus pilosifrons Fenton 1927: 11.

FEMALE: frons,

fully

winged;

length 5,0 mm;

head black,

clypeus and mandibles testaceous;

segment weak I y

dark;

with anterior part of

antennae testaceous,

propectus and pronotum redd i sh;

with

last

mesothorax,

meta thorax and propodeum black; abdomen reddish; legs testaceous, with mid and hind clubs of femora brown; tegulae testaceous; antennal segments in following proportions: reticulate rugose; connecting 10;

~

TL

18:

anterior ocellus

10;

8:

23:

17:

frons with 3 median

occipital

punctate and rugose,

to clypeus;

14:

with

13:

POL

~

4;

OL

~

12:

14;

OOL

hairy,

impression;

tubercles reaching

head fully

irregular keels

3;

pronotum dull,

an anterior transversal pronotal

13:

longitudinal

carina complete;

disc centrally excavated;

14:

and

tegulae;

~

10;

OPL

strongly posterior scutum

shiny, smooth, finely punctate, without sculpture among punctures; notaulices not fully visible because hidden by ,he pin (in the only known specimen); scutellum and metanotum shiny, smooth, finely punctate, without sculpture among punctures; propodeum reticulate rugose; posterior surface not smooth, reticulate rugose, without longitudinal keels; fore-wings with two dark transversa I bands, one on the basa I cell s and another beneath the pterostigma; radial cell: fig. 147 B; distal part of radial vein longer than proximal part (21:15); fore tarsal segments in following proportions: 12: 4:

7:

distally

14: than

26;

enlarged claw

the proximal

double-S shaped,

123)

with one bristle

tibial

Semacoch

TYPICAL MATERIAL:

spurs 1,1,2.

(Gua tema I a)

holotype F!

in WA.

located further

segment 5 of fore tarsi

with some rows of nearly 40

of nearly 4 lamellae; MALE: unknown LOCUS TYP I CUS:

(Fig.

prominence;

lamellae;

(Fig.

apex with a

123) group

Anteoninae

Fig.

HOSTS:

123 -

205

Chela of Deinodryinus pilosifrons Fenton

(type)

unknown

DISTRIBUTION:

Neotropic region:

GUATEMALA:

Semacoch

(Alta Vera,

Paz),

206

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

WA!

28. Deinodryinus politus n. sp.

FEMALE: brown

fully

pronotum margin a

winged;

and ventral reddish,

brown;

reddish

10: 9:

with hind coxae and

9: 9: 8:

9:

weak short complete; TL = 6;

with sides partly brown; abdomen

head dull,

irregular striae,

pronotum ha i ry,

mainly

du II,

very

antennae

strong

tegulae;

14:

weakly rugose,

near the antennal POL = 4;

OL = 3;

sockets; OOL = 8;

transversa I

with

legs

in following proportions:

with a

tubercles reaching

testaceous-brown;

granulated,

occipital carina complete;

with pronotal

with anterior

mesopleura black,

hind clubs of femora brown;

antenna I segments 11;

with ocellar region

propectus black;

scutum reddish,

propodeum black;

metapleura black;

thickened;

head reddish,

antennae testaceous;

scutellum reddish;

spot;

testaceous, distally

length 4,87 mm;

side black;

8:

18:

with

frontal

line

OPL = 9;

impression and

posterior surface of pronotum

nearly as long as broad, flat, strongly transversely striate, rugose; scutum dull, granulated; notaul ices incomplete, almost reaching the posterior margin of scutum; granulated;

propodeum dull,

longitudinal

keels;

scutellum and metanotum shiny, reticulate rugose,

very

weakly

without transversal

posterior surface of propodeum fully

or

reticulate rugose,

dull; fore-wings with two dark transversal bands, one on the basal cells and another beneath the pterostigma; distal part of radial vein little shorter than proximal part (12: 14); fore tarsal segments in following proportions: 12: 3: 6: 14: 25; enlarged claw (Fig. 124) with 1 bristle located further distally tarsi

(Fig.

nearly 6 MALE:

124)

lamellae;

the proximal

tibial

prominence; lamellae;

segment 5 of fore

apex with a

group of

spurs 1,1,2.

unknown

LOCUS TYPICUS:

Serra do Caraca

TYPICAL MATERIAL: HOSTS:

than

with 2 rows of nearly 50

holotype F!

(Brazil) in TW.

unknown

DISTRIBUTION: Neotropic region: Minas Gerais), TW!

BRAZIL:

Serra do Caraca

(S.

Barbara,

29. Deinodryinus c1aripes n. sp.

FEMALE: pronotum,

fully

winged;

length 4,38 mm;

sides of pronotum,

scutum,

testaceous,

propodeum,

with collar of

partly

metapleura,

dorsal

side of abdomen black; metanotum brown; antennae not or slightly distally thickened; antennal segments in following proportions: 9: 5: 15: 13: 11: 10: 9: 10: 9: and with two

11; head shiny, granulated; frons with a weak frontal line lateral longitudinal keels; occipital carina complete; POL =

207

Anteoninae

Fig.

124 -

Chela of Deinodryinus politus n.

sp.

(type)

208

3,5;

Mem.

OL = 4,5;

OOL= 11;

anterior surface rugose; numerous short

Amer.

Ent.

OPL = 10;

TL = 8;

lobes;

scutum;

37,

1984

pronotum partly

hairy,

pronotal notaul ices

as

tubercles reaching long

incomplete,

long

(26:22);

almost

scutellum and metanotum shiny,

as broad,

tegulae;

flat,

scutum shiny,

reaching smooth,

without

posterior surface of weakly

the posterior margin of

finely punctate,

sculpture among punctures; propodeum dull, reticulate rugose, longitudinal keels on anterior surface and without transversal

Fig.

with

granulated and with

long i tud i na I keel s near the posterior marg i n of pronotum;

pronotum slightly broader than granulated;

no.

posterior surface smooth,

posterior surface of pronotum nearly dorsal

Inst.,

125 -

Chela of

Deinodryinus c1aripes n. sp.

without

with some or

(type)

Anteoninae

longitudinal fully

beneath

posterior surface; dull;

the pterostigma;

proximal 85:

keels on

reticulate rugose, part;

117;

fore

tarsal

enlarged claw

posterior surface of propodeum

fore-wings

distal

with one dark

part of radial

segments

(Fig.

209

125)

vein

transversal

about as

in following proportions:

with

1 bristle

band

long as 45:

9:

21:

located further distally

than the proximal prominence; segment 5 of fore tarsi (Fig. 125) with two rows of nearly 29 + 21 lamellae; apex with a group of nearly 8 lamellae (one of these very MALE: unkonwn LOCUS TYPICUS:

long);

Essequibo

TYPICAL Mf\TERIAI: HOSTS:

tibial

spurs

1,1,2.

(Guyana)

holotype F!

in CO.

unknown

DISTRIBUTION:

Neotropic region:

GUYANA:

Essequibo,

CO!

30. Deinodryinus chiapasi n. sp.

FEMALE:

fully

of frons,

head black,

with anterior region

clypeus and mandibles reddish-testaceous;

winged;

length 5,62 mm;

antennae testaceous,

with segments 7-10 brown; pronotum and posterior testaceous, black;

thorax and propodeum black,

lobes of pronotum testaceous;

with hind coxae,

hind clubs of femora and hind

antennae slightly distally

proportions:

15:

rugose;

frons

median

keels;

9:

21:

with

two

occipital

14:

12:

lateral

~

with

4;

OOL~

14;

OPL~

irregular keels,

with pronotal shiny,

12;

with

punctate,

reaching

reticulate rugose,

area rugose;

10:

10:

keels around

TL

~

15;

tegulae;

located further distally

fore tarsi

(Fig. to

MALE:

unknown

LOCUS

TYP ICUS:

126)

tibial

Yerba Buena

TYPICAL MATERIAL:

OT!

POL

as broad as

long and

scutellum and metanotum notaulices

propodeum dull, median

bands, one on the basal

fore tarsal

segments (Fig.

prominence;

lamellae

in

126)

with one

segment 5 of

(extending

spurs 1,1,2.

in OT. MEXICO:

5;

reticulate rugose and

enlarged claw

the proximal

with ~

(Mexico)

holotype F!

HOSTS: unknown ::JISTRIBUTION: Neotropic region: Chiapas),

than

32;

with 2 rows of nearly 38

the apex);

carina;

keels on posterior surface;

the pterostigma;

continuously

to occipital

length of scutum;

4:

15:

two

behind posterior ocelli

scutum,

following proportions:

7:

in following reticulate

the orbits and with

fore-wings with 2 dark transversal 12:

segments

without sculpture among punctures; longitudinal

legs

tibiae partly

head shiny,

posterior surface nearly

cells and another beneath bristle

10;

pronotum dull,

approximately 0,75

with 2

antennal 8:

region

posterior ocell i

tubercles reaching

smooth,

incomplete,

thickened;

11:

carina complete;

two obi ique keels connecting OL

with sides of

abdomen bl ack;

Yerba Buena

(20 mi.

N Bochil,

210

Mem. Amer.

Fig.

126 -

En!.

Ins!.,

no. 37,

1984

Chela of Deinodryinus chiapasi n. sp.

(type)

Anteoninae

211

31. Deinodryinus nigricans (Cameron) n. sp.

Dryinus nigricans Cameron 1888: 446. Plastodryinus nigricans Cameron): Kieffer 1914b: 43.

FEMALE: fully winged; length 4,5 mm; head black, with clypeus and mand i b les testaceous (wi th teeth brown); antennae testaceous-brown, wi th segments 7-10 brown;

thorax,

propodeum,

of abdomen

light;

testaceous;

antenna I segments

8:

7:

8:

7:

longitudinal

10;

head dull,

keels,

with 3 median POL = 3; a

weak

legs brown,

hairy;

pronotal

OPL = 11;

tubercles reaching

scutum;

with

tegulae;

incomplete,

numerous

reticulate rugose, surface;

without a

around

fore-wings

smooth,

keel

transversal

12:

9:

to clypeus; hairy,

with

keels;

granulated;

approximately 0,75

granulated;

transversal

with 2 dark

22:

the orbits and

irregular short

length of

propodeum shiny,

between dorsal

posterior surface with 2 irregular longitudinal

smooth;

9:

pronotum dull,

scutum dull,

reaching

scutellum and metanotum dull,

14:

apex

tegulae

keels connecting ocell i

TL = 12;

with

black;

light;

numerous irregular

strong keel

and parallel

impression,

notaulices distinct,

following proportions:

frons with a

OOL= 12;

transversal

in

pet iol e and abdomen

last segments of tarsi

granulated,

longitudinal

OL = 3;

with

bands,

and

keels;

one on

posterior

median

the basal

area

cells

and another beneath the pterost i gma; d i sta I part of rad i a I vei n much longer than proximal part (20:12); radial cell very wide (Fig. 147); pterostigma (9:

15);

long and

narrow;

segment 5 of fore

lamellae;

apex with

127)

1 peg-I ike bristle

with

prominence; MALE:

fore tarsal

tarsi

(Fig.

a group of at

segment

127)

least

16

lamellae;

located further

segment 3 of fore

tarsi

1 shorter than

segment 4

with some rows of nearly 44 distally

produced

enlarged claw than

into hook;

(Fig.

the proximal tibial

spurs 1,1,2.

unknown

LOCUS TYP ICUS: Volcan de Chiriqui (Panama) TYPICAL MATERIAL: holotype F! in BI.i. HOSTS:

unknown

DISTRIBUTIOI'j:

Neotropic region:

PANAMA:

Volcan

de Chiriqui

BM!

32. Deinodryinus actuosus n. sp.

FEMALE:

fully

winged;

length 4,37-4,68 mm;

less wide testaceous frontal occasionally

two reddish

area;

spots on

head black,

with a

more or

clypeus and mandibles testaceous; temples;

antennae

testaceous,

with

segments 7-10 brown; prothorax reddish-testaceous; scutum reddishtestaceous, usually with region near the anterior margin darkened; sometimes almost fully

darkened;

propodeum redd i sh-testaceous,

scutellum,

metanotum,

with furrows between

pleura and

scutum and scutell um,

212

Mem.

Fig.

127 -

Chela of

scutell um and metanotum, brown-reddish; brown, distally 13:

10:

hind

Ent.

Inst.,

propodeum and metanotum

9:

antennal 8:

10;

37,

1984

darkened;

abdomen

with clubs of hind and mid femora partly

tibiae and sometimes also mid 9:

no.

Deinodryinus nigricans (Cameron)(type)

legs testaceous,

thickened; 10:

Amer.

segments

head dull,

tibiae brown;

antennae slightly

in following proportions:

fully

reticulate rugose;

13:

frons

6: with

18: two

Anteoninae

lateral region ocelli

keels around

the orbits and with

behind posterior ocell i to occipital

OOl =12;

carina;

OPl = 10;

tracks of three median keels;

with 2 obi ique keels connecting

occipital

Tl = 13;

213

carina complete;

pronotum dull,

posterior

POL = 3,5;

rugose,

Ol =3;

with numerous

transversal keels on posterior surface; posterior surface nearly as broad as long; pronotal tubercles reaching tegulae; scutum, shiny, smooth, finely

punctate,

without sculpture among punctures;

reaching approximately 0,65 length of scutum; shiny,

smooth,

without sculpture;

posterior surface rugose, dark transversal pterostigma;

bands,

distal

without one on

the basal

part of radial

vein

keels;

with

two the

fore-wings

longer than proximal

claw

128)

13: 3:

8:

segment 5 of fore tarsi

apex with a

group of 12 lamellae;

(Fig. tibial

128)

(20: 16);

enlarged

the proximal

with 2 rows of 26 lamellae;

spurs 1,1,2.

MALE: unknown lOCUS TYPICUS: San Cristobal de las Casas (Mexico) TYPICAL MATERIAL: holotype F! and 1 paratype F! in OT;

128 -

part

13: 30;

with one bristle located further distally than

prominence;

with

cells and another beneath

segments in following proportions:

Fig.

incomplete,

propodeum reticulate rugose, longitudinal

fore tarsal (Fig.

notaul ices

scutellum and metanotum

Chela of Deinodryinus actuosus n.

sp.

2 paratypes FF!

(type)

214

Mem.

Amer.

En!.

Inst.,

no.

1984

37,

in OL. HOSTS:

unknown

DISTRIBUTION: (Chiapas),

Neotropic region:

OL!

MEXICO:

San Cristobal

de

las Casas

OT!

33. Deinodr:linu5 incaicU5 n. sp.

FE,'-'lALE:

fully

winged;

length 4,37-4,81

sometimes with anterior half of frons testaceous;

antennae fully

7-10 brown;

legs fully

mm;

head

testaceous;

testaceous;

testaceous or black, clypeus and mandibles

occasionally

testaceous or with coxae,

with segments 6-10 or

mid and hind

and hind clubs of femora black;

occasionally only mid and

partly

and abdomen

black;

thorax,

propodeum

of pronotum testaceous;

antennae distally

black;

thickened;

tibiae,

occasionally margins antenna I segments

following proportions: 14: 7: 21: 11: 9: 10: 8: 8: 7: 10; head shiny; in front of anterior ocellus with a central region sculptured by longitudinal regions on finely

keels;

frons

without

lateral

the sides of the ocell i

punctate,

temples smooth;

keels;

and behind

surface smooth, dorsal

flat, lobes,

with

finely

tegulae;

areolae smaller, bands,

pterostigma; fore tarsal claw

with

without sculpture among punctures,

scutellum

rugose;

surface with longitudinal

the basal

areolae wide;

keels;

propodeum

posterior surface

fore-wings with 2 dark

longer than proximal

in following proportions: 1 bristle

notaulices

cells and another beneath

vein

pronotal

and metanotum shiny,

the posterior margin of scutum;

part of radial

segments

(Fig - ,29)

without

one on

distal

11; OPl = 10; TL = 9; anterior furrow; posterior

without sculpture among punctures; dorsal

11:

2:

6:

located further distally

20:

the

part 33;

than

(21:17);

enlarged

the proximal

prominence; segment 5 of fore tarsi (Fig. 129) with 2 rows of nearly lamellae; apex with a group of nearly 5 lamellae (one of these very long);

tibial

MALE:

fully

52

spurs 1,1,2. winged;

legs testaceous;

length 3,37 mm;

black

antennae not distally

following proportions: much

to

=

anterior surface weakly

scutum,

almost reaching

reticulate rugose;

transversal

punctate,

occasionally

finely punctate,

almost complete, dull,

carina complete;

posterior ocell i smooth,

without sculpture among punctures or weakly granulated;

tubercles reaching smooth,

occipital

in frons

vertex xithout oblique keels connecting posterior ocelli

occipital carina; POL = 2,5; Ol = 3; OOl pronotum shiny, with a strong transversal without

mid

hind coxae

14:

6:

15:

9:

9:

longer than breadth of segments;

reticulate rugose;

frontal

sides of the ocelli

I ine present;

without sculpture among punctures; carina complete;

10:

POL

=

Ol

=

3;

11:

10:

antennae,

segments in

11;

antennal

without keels,

and on

to occipital

=

7;

OPl

carina;

=

7;

the

punctate and

region behind ocell i without OOl

hairs

wi th frons fu II y

regions of head behind

posterior ocelli 5,5;

10:

clypeus,

antennal

head sh i ny,

not reticulate rugose,

oblique keels connecting

mandibles,

thickened;

Tl

two

occipital

=

5;

scutum,

Anteoninae

/

\

Fig.

129 -

215

r.! .1

~/

Chela of Deinodryinus incaicus n.

sp.

(type)

216

Mem.

Amer.

scutellum and metanotum shiny, among punctures;

internal

proximal

branch;

3 paratypes

para type M! HOSTS:

in AM;

without sculpture

almost reaching

the posterior

without transversal

part

tibial

(2 FF,

bands;

(12:14);

distal

gonoforceps

or

fore-

part of radial (Fig.

130)

vein

with

spurs 1,1,2.

1 M)!

1 para type F!

in TW;

2 paratypes FF!

in BM;

2 paratypes FF!

1 paratype F! in OT;

1

in OL.

unknown

DISTRIBUTION:

Fig.

1984

with posterior surface fully reticulate rugose;

LOCUS TYP ICUS: Mitu (Colombia) TYPICAL MATERIAL: holotype F! and in LE;

37,

finely punctate,

without dark transversal

little shorter than broad

smooth,

no.

propodeum reticulate rugose,

keels,

wings hyaline,

Ins!.,

notaul ices almost complete,

margin of scutum; longitudinal

Ent.

130 -

Neotropic region:

PANAMA:

Gamboa

(Pipeline Road,

Male genitalia of Deinodryinus incaicus n. (Brazil):

right

half removed

sp.

from

Canal

Vila Vera

217

Anteoninae

Zone),

LE!

Catarina), IV

(Sinop,

COLOMBIA:

Mitu

IV 50° 30'

BM!

Mato Grosso),

Tucuman,

OT!

Jujuy,

(Vaupes),

S 12° 30' TIV!

TIV!

BM!

BRAZIL:

Rio Guapore

Horco Molle

Nova Teutonia

(Sta

OL!

TIV!

12° 31' S 55° 37'

(Rondonia),

AM!

ARGENT INA:

(Vi la Vera), (Tucuman),

OT!

OL!

34. Deinodryinus bolivianus n. sp.

FEMALE:

fully

length 5,0 mm;

winged;

anterior region of frons

testaceous;

black;

10 black; abdomen and legs fully testaceous; thickened;

antennal

7: 7: 7: 7: 9; keels;

segments

head dull,

in

mandibles,

antennae slightly

sculptured by

occipital carina complete;

numerous

region behind ocell i

distally

15: 8: 20:

following proportions:

hairy,

clypeus and wi th segments 9-

antennae testaceous,

13:

10:

longitudinal

with 2 obi ique keels

connecting posterior ocelli to occipital 13: OPl ; 9; Tl ; 9; pronotum sh i ny,

carina; POL; 3; Ol ; 2,5; OOl ; crossed by a strong transversa I

furrow;

finely

posterior surface smooth,

among punctures;

pronotal

and metanotum shiny, punctures; scutum;

flat,

punctate,

tubercles reaching

smooth,

tegulae;

finely punctate,

notaulices incomplete,

reaching

propodeum reticulate rugose;

approximately 0,75

dorsal

transversa I bands,

scutellum

without sculpture among

surface with

posterior surface with areolae smaller and without fore-w i ngs with 2 dark

without sculpture scutum,

one on

length of

areolae wide;

longitudinal

keels;

the basa I cell sand

another beneath the pterostigma; distal part of radial vein longer than proximal part (20: 14); fore tarsal segments in fOllowing proportions: 12:

3: 8:

13: 32; enlarged claw (Fig.

131) with 1 bristle located further segment 5 of fore tarsi (Fig. 131) with 2 rows of nearly 32 lamellae; apex with a group of nearly 27 lamellae; segment 3 of fore tarsi produced into a hook; tibial spurs 1,1,2.

distally

MALE:

than

the proximal

unknown

LOCUS TYPICUS:

Rurenabaque

TYPICAL MATERIAL: HOSTS:

prominence;

(Bolivia)

holotype F!

in

BM.

unknown

DISTRIBUTION:

Neotropic region:

BOLIVIA:

Rurenabaque

(Beni),

8M!

35. Deinodryinus eminens n. sp.

FEMALE:

fully

winged;

surface of frons,

prothorax redd i sh; brown-black;

length 4,56-4,81

mm;

head black,

clypeus and mandibles testaceous; mesothorax,

legs testaceous;

metathorax and

with anterior

antennae testaceous,

propodeum black;

antennae slightly distally

abdomen

thickened;

segments in following proportions: 15: 8: 20: 18: 14: 13: 10: 13; head dull, fully reticulate rugose; frons with 3 weak median

antennal

10:

keels;

2

lateral

keels around the orbits;

occipital

carina complete;

10:

region

218

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

r

/

Fig.

131

-

Chela of Deinodryinus bolivianus n.

sp.

(type)

Anteoninae

219

/

/

/

II / 'Y~

,

If~. l'

/

Fig.

132 -

Chela of Deinodryinus eminens n.

of head behind posterior ocelli ocelli

to occipital

carina;

sp.

(type)

with 2 oblique keels connecting posterior

POL = 5; Ol = 4; OOl = 13; OPl = 12; Tl = 12;

220

Mem.

pronotum shiny,

hairy,

Amer.

En!.

Ins!.,

no.

37,

1984

with a strong anterior transversal

furrow;

posterior surface of pronotum nearly as long as broad,

finely

without

lobes parallel

sculpture among punctures,

the sides of pronotum;

pronotal

with 2 weak dorsal

tubercles reaching

scutellum and metanotum shiny,

smooth,

tegulae;

very finely

punctate, to

scutum,

punctate,

without

sculpture among punctures; notaul ices almost complete, almost reaching the posterior margin of scutum; propodeum reticulate rugose, with posterior surface dull,

reticulate rugose;

one on

the basal

radial

vein

little shorter than proximal

in following proportions: bristle

12:

3:

7:

fore tarsi

(Fig.

132)

the pterostigma;

part 30;

(13:14);

the proximal

tibial

(Fig.

part of

segments

132)

with 1

segment 5 of

prominence;

lamellae;

bands,

distal

fore tarsal

enlarged claw

with 2 rows of nearly 26

15 lamellae;

apex with a group

spurs 1,1,2.

unknown

LOCUS TYPICUS:

Nova Teutonia

TYPICAL MATERIAL:

(Brazil)

holotype F!

HOSTS: unknown DISTRIBUTION: Neotropic region: BM!

12:

located further distally than

of nearly MALE:

fore-wings with 2 dark transversal

cells and another beneath

Sta Barbara

in BM; BRAZIL:

(Minas Gerais),

paratype F!

in TW.

Nova Teutonia

(Sta Catarina),

TW!

36. Deinodryinus trinidadi n. sp.

FEMALE: fully winged; length 3,75-4,75 mm; head testaceous or reddishtestaceous, with ocellar region brown or dark; antennae testaceOU5, with segments 8-10 or 9-10 dark; legs testaceous;

thorax and propodeum black;

occasionally extremities of hind tibiae brown; mid tibiae and hind coxae partly brown; thickened; 8:

8:

8:

lateral

antennal

11;

abdomen brown;

stalk of hind femora and extremities of hind tibiae brown;

segments

head shiny,

keels around

reticulate rugose;

in following proportions:

reticulate rugose;

the orbits;

region

occasionally extremities of

antennae slightly distally 15: 7:

21:

13:

9:

frons with 3 median keels;

region behind ocelli

behind posterior ocell i

8: 2

weakly or strongly

with 2 obi ique keels

connecting posterior ocelli to occipital carina; occipital carina complete; POL = 3; OL = 3; OOL = 12; OPL = 12; TL = 12; pronotum shiny, hairy, with a strong finely dorsal

anterior transversal

punctate,

lobes parallel

tegulae;

scutum,

punctate, reaching

furrow;

to the sides of pronotum;

the posterior margin of scutum

one on

part of radial

the basa I cells and vein slightly

with 2 weak

tubercles reaching

smooth,

very finely

notaul ices incomplete,

(Fig.

157 A);

almost

propodeum with

fore-wings with 2 dark transversal

another beneath

longer than proximal

segments in following proportions:

smooth,

pronotal

scutellum and metanotum shiny,

without sculpture among punctures;

posterior surface almost fully rugose; bands,

posterior surface of pronotum

without sculpture among punctures,

12: 4:

9:

15:

the pterost i gma;

part 35;

(17:15);

fore

enlarged claw

di sta I tarsal (Fig.

Anteoninae

221

133) with 1 bristle located further distally than the proximal prominence; segment 5 of fore tarsi (Fig. 133) with 2 rows of nearly 34 lamellae; apex with a MALE:

group of nearly 15 lamellae; fully winged;

Fig.

133 -

tibial

length 2,60-3,56 mm;

Chela of

spurs 1,1,2. black;

mandibles and antennae

Deinodryinus trinidadi n. sp. (type)

222

Mem.

Amer.

with

Ent.

Inst.,

testaceous;

legs testaceous,

hairs much

longer than breadth of segments;

37, 1984

no.

hind coxae proximally black; antennae not

antennal

distally

antennal segments in following proportions: 15: 7: 15: 10: 12: 12: 12: 12: 12: 15; head shiny, fully reticulate rugose; frons with 3

thickened; median

keels;

region behind ocelli

2

lateral

to occipital

scutum,

keels around

posterior ocelli carina;

the orbits;

occipital

POL

=

4;

=

Ol

scutellum and metanotum shiny,

3;

OOl

smooth,

=

5;

finely

OPl

notaul ices almost complete,

posterior margin of scutum;

propodeum reticulate rugose,

surface with 2

complete keels;

longitudinal dull,

rugose;

bands or weakly darkened;

distal

part of radial

8;

Tl

=

7;

without

almost reaching

the

with posterior

area with a

without dark

transversal

transversal

vein nearly as

long as

(14:14); gonoforceps (Fig. 134) with apical branch wrapping tibial spurs 1,1,2.

proximal penis;

median

fore-wings hyal ine,

=

punctate,

sculpture among punctures;

anterior keel,

carina complete;

with 2 oblique keels connecting posterior

part

lOCUS TYPICU5:

5t.Augustine

TYPICAL MATERIAL:

Fig.

(Trinidad)

holotype F!

and 4 paratypes MM!

in 8M;

1 paratype

134 - Male genital ia of Deinodryinus trinidadi n. sp. from 51. Augustine

(Trinidad):

right

half removed.

Anteoninae

F!

in OT;

HOSTS:

2 paratypes MM!

in OL.

unknown

DISTRIBUTION: BM!

223

Simla

Neotropic region:

(Arima),

TRINIDAD:

St.

Augustine

(SI.

George),

OL!

OT!

37. Deinodryinus rapax n. sp.

FEMALE:

fully

length 6,56 mm;

winged;

anterior region of frons

testaceous;

black;

mandibles,

antennae testaceous,

clypeus and with segments 7-

10 brown; fore legs testaceous; mid and hind legs brown; antennae thickened; antenna I segments in following proportions: 19: 9: 29: 22: 20: 19: 17: 17: 16: 18; head shiny, fully reticulate rugose; frons distally

with 3 median keels;

2 lateral

keels around

the orbits;

region of head

behind posterior ocelli reticulate rugose and with 2 oblique keels connecting posterior ocell i to occipital carina; occipital carina complete; POL; 5; OL ; 2,5; OOL ; 12; OPL ; 13; TL ; 15; pronotum hairy, dull, crossed by

an

anterior strong transversal

pronotum smooth,

strong I y

punctate,

posterior surface of pronotum dorsal

weak parallel

lobes;

less than

pronotal

scutellum and metanotum shiny, among punctures; of scutum;

posterior surface of

twice as broad as

long,

tubercles reaching tegulae;

smooth,

finely

punctate,

with 2 scutum,

without sculpture

reaching approximately 0,80

notaul ices incomplete,

propodeum dull,

furrow;

wi thout scu I pture among punctures;

reticulate rugose;

length

posterior surface dull,

reticulate rugose, without longitudinal keels; fore-wings with 2 dark transversal bands, one on the basal cells and another beneath the pterostigma; fore tarsal

distal

part of radial

vein

longer than

segments in following proportions:

part (30:25); 19: 39; enlarged

proximal

14: 5:

10:

135) with one bristle located further distally than the proximal segment 5 of fore tarsi (Fig. 135) strongly S-curved, with some rows of at least 60 lamellae (extending continuously to the apex); tibial spurs 1,1,2.

claw

(Fig.

prominence;

MALE: unknown LOCUS TYP ICUS:

Chiriqui

TYPICAL MATERIAL: HOSTS:

(Panama)

holotype F!

in LE.

unknown

DISTRIBUTION: Panama),

Neotropic region:

PANAMA:

Chiriqui

(Fortuna,

North

LE!

38. Deinodryinus croceus n. sp.

FEMALE:

fully

winged;

length 5 mm;

reddish-testaceous,

with abdomen

black and antennal segments 9-10 brown; antennae distally thickened; antennal segments in following proportions: 14: 8: 17: 13: 9: 9: 8: 8:

8:

224

Mem. Amer. En!.

Ins!., no. 37,

1984

/

Fig.

135 -

Chel a of Deinodl'yinus l'apax n. sp.

(type)

Anteoninae

Fig.

136 -

Chela of Deinodryinus croceus n.

225

sp.

(type)

226

10;

Mem.

head shiny,

longitudinal

Amer.

with frons

keels;

and

occipital

En!.

Inst.,

37,

1984

vertex reticulate rugose and with some

carina complete;

oblique keels connecting posterior ocelli without

no.

sculpture among punctures,

region behind ocell i with

to occipital

carina,

two

punctate and

not reticulate rugose or keeled;

POL

4,5; OL = 3,5; OOL = 13; OPL = 12; TL = 13; pronotum shiny, with a strong transversal impression and with pronotal tubercles reaching tegulae; smooth,

posterior surface of pronotum nearly as with

metanotum

almost

shiny,

reaching

rugose, dull

two weak dorsal

dull,

smooth,

without

vein

sculpture;

without transversal

the basal

radial

lobes;

long as broad,

scutum,

or

with

shorter than

keels;

two dark

cells and another beneath

slightly

propodeum reticulate

longitudinal

fore-wings

proximal

shiny,

scutellum and

notaulices almost complete,

the posterior margin of scutum;

and reticulate rugose;

one on

parallel

posterior surface

transversal

the pterostigma; part

(14:15);

bands,

distal

part of

fore tarsal

segments in following proportions: 12: 3: 8: 15: 32; enlarged claw (Fig. 136) with 1 bristle located further distally than the proximal prominence; segment 5 of fore a

group of 17

MALE:

fully

tarsi

(Fig.

lamellae;

winged;

136)

tibial

length 3,31

testaceous;

antennae testaceous;

femora and

hind

hairs

tibiae brown;

proportions:

15:

8:

the orbits;

15:

12:

mm;

black;

lamellae;

with coxae,

11:

11:

11:

carina complete;

antenna I segments

POL

=

6;

=

OL

3;

=

OOL

clubs of

11:

9:

12;

keel,

region

head dull,

with

two

10;

=

OPL

antennal

in following fully

lateral

keels around

of head behind ocelli

tracks of two oblique keels connecting posterior ocelli

carina;

apex with

mandibles and clypeus

antennae not distally thickened;

frons without median

occipital

two rows of 30

legs testaceous,

longer than breadth of segments;

reticulate rugose; with

with

spurs 1,1,2.

9;

=

TL

13;

only

to occipital

scutum,

scutellum

and metanotum shiny, smooth, finely punctate, without sculpture among punctures; notaulices almost complete, almost reaching the posterior margin of scutum; longitudinal

propodeum reticulate rugose, keels,

for-ewings hyaline,

without dark

vein shorter than proximal apical

part

branch wrapping penis;

LOCUS TYPICUS:

Amaicha del

TYPICAL MATERIAL:

without transversal

with posterior surface dull transversal (11 :13); tibial

Valle

holotype F!

HOSTS: unknown DISTRIBUTION: Neotropic region:

and

and

reticulate rugose;

bands;

distal

part of radial

gonoforceps

(Fig.

137)

without

spurs 1,1,2.

(Argentina)

and

1 paratype M!

ARGENTINA:

in TW.

Amaicha del

Valle,

TW!

39. Deinodryinus asper n. sp.

FEMALE:

fully

winged;

length 3,56 mm;

and anterior half of frons mandibles testaceous;

head black,

(mainly along orbits)

antennae testaceous,

prothorax reddish-testaceous,

with

temples reddish

testaceous;

clypeus and

with segments 8-10 brown;

with sides partly black;

mesothorax and

Anteoninae

Fig.

137 -

Male genitalia of Valle

Deinodryinus croceus n. sp. from Amaicha del

(Argentina):

meta thorax reddish-testaceous; testaceous-brown;

227

right half removed.

propodeum black;

legs testaceous,

pet iol e black;

with hind coxae partly

abdomen

brown;

antennae

sl ightly distally thickened; antenna I segments in following proportions: 13: 7: 19: 13: 9: 9: 7,5: 7,5: 7: 10; head with frons reticulate rugose and with 3 median vertex,

longitudinal

keels;

temples and region

2

lateral

behind ocelli

and without sculpture among punctures; 2 ob I i que keel s connect i ng carina complete; shiny,

hairy,

POL

with

=

3;

OL

of pronotum without sculpture, ::Jronotum, scutum,

nearly

as

long

region

posteri or ocell i

=

3;

=

OOL

an anterior strong

keels around

10;

scutellum and metanotum shiny,

,-.ithout sculpture among punctures;

OPL

transversal

pronotal

punctate

behind posterior ocell i

to occi pita I cari na;

with 2 dorsal

as broad;

the orbits;

not reticulate rugose,

=

8;

TL

furrow;

=

11;

pronotum

posterior surface

lobes parallel

to the sides of

tubercles reaching

smooth,

with

occi pita I

very finely

tegulae;

punctate,

notaulices almost complete,

almost

228

Mem.

Amer.

Ent.

Inst.,

reaching the posterior margin of scutum; posterior surface reticulate rugose, transversal

bands,

pterostigma; (15:13);

distal

segments

(Fig.

138)

prominence;

nearly 36 lamellae;

37,

1984

propodeum reticulate rugose,

with

dull; fore-wings with 2 dark

the basal

part of radial

fore tarsal

enlarged claw proximal

one on

no.

cells and another beneath the

vein slightly

longer than proximal

in following proportions:

12: 3:

7:

with 1 bristle located further distally

segment 5 of fore tarsi

(Fig.

apex with a group of nearly

138)

9:

part

29;

than

the

with 3 rows of

10 lamellae;

tibial

spurs

~ \

Fig.

138 -

Chela of Deinodryinus asper n.

sp.

(type)

\

229

Anteoninae

Fig.

139 -

Chela of

Deinodryinus schlingeri n. sp.

(type)

230

1,1,2. MALE:

Mem.

Amer.

En!.

Inst.,

no.

37,

1984

unknown

LOCUS TYPICUS:

Nova Teutonia

TYPICAL MATERIAL: HOSTS:

(Brazil)

holotype F!

in BM.

unknown

DISTRIBUTION: BM!

Neotropic region:

BRAZIL:

Nova Teutonia

(Sta Catarina),

40. Deinodryinus schlingeri n. sp.

FEMALE:

fully

winged;

length 3,62 mm;

malar space testaceous; legs brown, testaceous;

with fore tarsi, 13:

6:

17:

8:

mandibles,

clypeus and

with segments 7-10 brown;

fore tibiae and stalks of fore femora

antennae distally

proportions:

black;

antennae testaceous, thickened;

8:

8:

6:

6:

antennal

6: 9;

segments in following

head shiny;

frons

with

numerous longitudinal keels and areolae; occipital carina complete; region of head behind ocell i with numerous longitudinal keels, among which two oblique keels connecting posterior ocelli

=

2,5; OOL transversal

to occipital

carina;

POL = 3,5;

OL

=

15; OPL = 11; TL = 11; pronotum shiny, with an anterior furrow; posterior surface nearly as long as broad, with 2

weak parallel

lobes,

tubercles reach i ng

hairy,

tegu I ae;

weakly scutum,

rugose,

almost smooth;

pronotal

scute II um and metanotum smooth,

shiny,

finely punctate,

without sculpture among punctures;

almost

complete,

reaching

almost

the

posterior

margin

of

notaulices

scutum;

propodeum

reticulate rugose; posterior surface reticulate rugose, without longitudinal keels; fore-wings with 2 dark transversal bands, one on the basal cells and another beneath the pterostigmaj distal part of radial vein as long as proximal part (15:15); enlarged claw (Fig. 139) with 1 bristle located further distally than the proximal prominence; segment 3 of fore tarsi produced nearly

into a

22

hook;

lamellae;

segment 5 of fore tarsi

apex with

a

(Fig.

group of nearly

139)

with 2 rows of

18 lamellae;

tibial

spurs

1,1,2. MALE:

unknown

LOCUS TYPICUS:

Tingo Maria

TYPICAL MATERIAL: HOSTS:

in CA.

unknown

DISTRIBUTION: NOTE:

(Peru)

holotype F!

Neotropic region:

the species

is named

in

PERU:

Tingo Maria

honor of Mr.

E.I.

(Monson Valley),

CAl

Schlinger.

41. Deinodryinus alexandrae n. sp.

FEMALE: frons,

fully

winged;

length 6 mm;

head black,

clypeus and mandibles testaceous;

1-4 testaceous; propodeum,

prothorax reddish-testaceous;

abdomen

black;

fore

with anterior region of

antennae black, mesothorax,

legs testaceous,

with

with segments metathorax,

tarsi

brown;

mid

Anteoninae

and hind

legs black;

antennae distally

following proportions;

18;

8:

30:

23:

231

thickened antenna I segments

16:

15:

13:

with frons reticulate rugose and sculptured by behind ocelli carina POL

=

and 5;

with

=

12:

15;

longitudinal

4;

=

OOl

longitudinal

16;

=

OPl

keels;

13;

=

Tl

occipital 15;

in

head shiny, keels;

two oblique keels connecting posterior ocelli

with numerous

Ol

13:

region

to occipital

carina complete;

pronotum shiny,

smooth,

with a strong transversal impression and with pronotal tubercles reaching tegulae; posterior surface of pronotum nearly as long as broad, with two dorsal

parallel

longitudinal

lobes;

scutum,

scutellum and metanotum shiny,

smooth, without sculpture; notaul ices almost complete, almost reaching posterior margin of scutum; propodeum reticulate rugose, without transversal

or

longitudinal

reticulate rugose; basal vein

keels;

fore-wings with

cells and another beneath longer than proximal

proportions;

11:

Fig.

4:

140 -

11:

(Fig.

140)

42;

Chela of

located further distally tarsi

part

17:

with

than

posterior surface of propodeum dull, two dark

(22: 19);

HOSTS;

bands,

distal

tarsal (Fig.

segments 140)

one on

in following

with

1 bristle

Deinodryinus alexandrae n. sp. (type)

the proximal

two rows of 29

holotype F!

Neotropic region:

the species

is named

the

part of radial

prominence;

lamellae;

segment 5 of fore

apex with a

is the male of

group of 13

D. alexandrae)

(Ecuador)

in TW.

unknown

DISTRIBUTION: NOTE;

fore

enlarged claw

Napo and Coca Ri vers

TYPICAL MATERIAL:

transversal

the pterostigma;

lameillae; tibial spurs 1,1,2. MALE: unknown (perhaps D. hirticornis LOCUS TYP I CUS;

the

ECUADOR:

Napo and Coca Rivers,

in honor of Mrs.

Alessandra Mucci.

TW!

232

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

42. Deinodryinus carpen,s n. sp.

FEMALE: MALE:

unknown

fully

winged;

length 3,68 mm;

black;

antennae not distally thickened antennal breadth of segments; 17:

11:

11:

11:

10:

frons with numerous OL = 3,5;

OOL = 9;

Fig.

141

9:

9:

11;

head shiny,

longitudinal OPL = 8;

keels;

TL = 7;

to occipital

mainly on anterior half,

-

much shorter than

antenna I segments in following proportions:

connecting posterior ocelli punctate,

fore tibiae testaceous;

hairs short,

Male genitalia of

punctate,

12: 8:

without sculpture;

occipital carina complete;

POL

6;

vertex without two oblique keels carina;

scutum shiny,

strongly

without sculpture among punctures;

Deinodryinus carpens n. sp. (type): left half removed.

Anteon i nae

notaulices

incomplete,

almost reaching

scutellum and metanotum shiny, among punctures;

233

the posterior margin of scutum;

smooth,

finely

propodeum reticulate rugose,

punctate, without

without sculpture

longitudinal

posterior surface of propodeum reticulate rugose and with small shiny central distal

area;

fore-wings hyaline,

part of radial

vein

without dark transversal

about as long as proximal

part;

keels;

smooth bands;

gonoforceps

(Fig. 141) with subapical branch wrapping penis; tibial spurs 1,1,2. LOCUS TYPICUS: S.J. Barreiro (Brazil) TYPICAL MATERIAL: holotype M! and 2 paratypes MM! in Ol; 2 paratypes MM!

in TW ••

HOSTS:

unknown

DISTRIBUTION: OL!

Neotropic region:

BRAZil:

S.J.

Barreiro (Serra da Bocaina),

TW!

43. Deinodryinus perlucens n. sp.

FEMALE: MALE:

unknown

fully

winged;

antennae fully testaceous, thickened,

length 3,75-4,12 mm;

testaceous,

black;

mandibles testaceous;

sometimes with segments 3-10 darkened;

with stalks of hind femora brown;

with hairs much shorter than breadth of segments;

segments in following proportions: head sh i ny,

15: 7:

20:

legs

antennae not distally 12:

12:

12:

11:

antennal 11:

with frons scu I ptured by numerous strong para Ilel

11:

14;

and

longitudinal keels; region behind ocelli strongly punctate, without sculpture among punctures, without two strong oblique keels connecting posterior ocelli complete; finely

to occipital

POL = 4;

punctate,

incomplete,

smooth,

occipital

carina complete;

OOl = 11; OPL = 11;

TL = 6;

without sculpture among punctures;

smooth,

finely

punctate,

propodeum with dorsal

surface smooth,

shiny,

fore-wings hyaline,

finely

lOCUS TYPICUS: HOSTS:

notaul ices

Avispas

scutellum and

without sculpture among

without sculpture among punctures;

transversal

bands;

distal

vein shorter than proximal part (12:19); gonoforceps (Fig. wide apical branch wrapping penis; tibial spurs 1,1,2. TYPICAL MATERIAL:

part of radial 142)

with a

(Peru)

holotype M!

in TW;

1 para type M!

in OL.

unknown

DISTRIBUTION:

Neotropic region:

PERU:

Avispas

(Marcapata),

44. Deinodryinus inermis n. sp.

FEMALE:

unknown

line

surface reticulate rugose and posterior

punctate,

without dark

frontal

scutum shiny,

almost reaching the posterior margin of scutum;

metanotum shiny, punctures;

carina;

OL = 2,5;

Ol!

TW!

234

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

"

/

I

Fig.

Male genitalia of Deinodryinus perlucens n.

142 -

ha I f

MALE:

fully

winged;

length 3,37 mm;

black;

antennae testaceous;

legs

with coxae,

tibiae partly black;

antennae not distally thickened;

following proportions: rugose; POL = 7;

frons

17:

testaceous, 8:

with 3 median

OL = 4;

OOL = 12;

15:

13:

sp.

(type):

left

removed.

14:

longitudinal OPL = 11;

oblique keels connecting posterior ocelli

14:

mandibles testaceous; clubs of femora and hind

12: 9:

keels;

TL = 11;

antenna I segments in 11:

occipital

11;

head reticulate

carina complete;

vertex with 2 strong

to occipital

carina;

antennae with

Anteoninae

ha i rs much

longer than breadth of segments;

metanotum sh i ny,

smooth,

punctate,

notaul ices almost complete, strong

transversal

keel

almost smooth;

between

fore-wings

scutell um and

approximately 0,80

with some transversal dorsal

posterior surface with 2 slightly visible shiny,

scutum,

without scu I pture among punctures;

reaching

propodeum reticulate rugose, and a

235

surface

and posterior surface

longitudinal

hyaline,

length of scutum;

keels on dorsal keels;

without dark

median area

transversal

bands;

distal part of radial vein nearly as long as proximal part (13:13); gonoforceps (Fig. 143) without apical branch wrapping penis, only with a small

Fig.

apical

143 -

lobe;

tibial

spurs 1,1,2.

Male genitalia of half removed);

lOCUS TYP I CUS:

las Cej as

TYPICAL MATERIAL: M! in TW. HOSTS:

Deinodryinus inermis n. sp. (type)(A: right

B:

gonoforceps

(laterally

viewed).

(Argen tin a)

holotype M!

in TU;

1 para type M!

in Ol;

paratype

unknown

DISTRIBUTION:

Neotropic region:

ARGENTINA:

las Cejas

(Tucuman),

Ol!

TU!

236

Mem.

Jujuy,

Amer.

En!.

Inst.,

no.

37,1984

TW!

45. Deinodryinus paranus n. sp.

FEMALE: unknown MALE: fully winged; antennae testaceous;

length 1,62-2,68 mm; black; mandibles, legs and stalk of hind femora and distal half of hind tibiae

dark; antennae not distally thickened; antenna I segments in following proportions: 9: 5: 7: 4: 5: 5: 5: 5: 5: 6,5; head shiny, smooth, punctate, without sculpture among punctures; occipital

carina complete;

keels connecting posterior ocelli = 4;

OPl = 4;

scutum,

Tl = 3;

frons without median and

region behind posterior ocelli to occipital

antenna I hairs almost reaching

keels;

POL = 4;

Ol = 2;

OOl

longer than breadth of segments;

scutellum and metanotum shiny,

notaul ices incomplete,

carina;

lateral

without oblique

smooth,

without sculpture;

the posterior margin of scutum;

propodeum reticulate rugose, without transversal or longitudinal keels; fore-wings hyal ine, without dark transversal bands; distal part of radial vein slightly shorter than proximal an apical

Fig.

branch wrapping penis;

144 -

part tibial

(7:8)

gonoforceps

Male genital ia of Deinodryinus paranus n. ha I f

lOCUS TYPICUS:

Faz.

Taperinha

removed.

(Para,

(Fig.

144)

with

spurs 1,1,2.

Brazil)

sp.

(type):

right

237

Anteoninae

TYPICAL MATERIAL: M!

holotype M!

in OT;

2 paratypes MM!

in TW;

1 paratype

in Ol.

HOSTS:

unknown

DISTRIBUTION: ARGENTINA:

Neotropic region:

Horco Molle

BRAZil:

(Tucuman),

Ol!

Faz.

Taperinha

(Para),

OT!

TW!

46. Deinodryinus hirticornis (Kieffer) n. sp.

/

I

/

I

\/

\

I

/

/

/

\

II \

\

I

,I '-

\ '-

/

Fig.

145 -

Male genital ia of

Deinodryinus hirticornis (Kieffer) (type):

gonoforceps removed.

238

Mem.

Fig.

146 -

Amer.

Male genital ia of de las Cenyas

Ent.

Inst.,

no.

37,

1984

Deinodryinus hirticornis {Kieffer} from Rio {Mexico}:

left half removed.

Anteon hirticornis Kieffer 1911b: 231. Trisanteon hirticornis {Kieffer}: Kieffer 1913b: 300. Trisanteon hirticornis {Kieffer}: Kieffer 1914b: 197.

FEMALE: MALE:

unknown

fully

winged;

antennae testaceous;

length 3,25-4,18 mm;

black;

occasionally clypeus black;

abdomen brown-black;

legs testaceous,with coxae,

mandibles,

clypeus and

tegulae testaceous; clubs of femora,

mid and

hind tibiae partly black or brown; antennal segments in following proportions: 13: 7: 13: 12: 12: 11: 10: 10: 8: 12; antennal hairs longer

Anteoninae

than breadth of segments; with

two median

head shiny,

longitudinal

keels;

keels connecting posterior ocell i complete;

=

POL

7;

=

OL

5;

notaul ices

margin of scutum;

strongly reticulate rugose;

region behind ocell i

to occipital

=

OOL

scutellum and metanotum se;iny, among punctures;

239

12;

smooth,

carina;

=

OPL

11;

finely

incomplete,

keels on

12;

punctate,

frons

two obi ique

occipital

=

TL

almost reaching

propodeum reticulate rugose,

incomplete longitudinal

with

carina

scutum,

without sculpture the posterior

without or with 2

posterior surface;

area between

these

two

keels rugose, dull, with strong transversal keels; fore-wings hyaline, without dark transversal bands; distal part of radial vein nearly as long as proximal proximal

part

(10:9);

gonoforceps

membranous process,

(Figs.

with apical

145,

146)

without dorsal

branch wrapping penis;

tibial

spurs 1,1,2. LOCUS TYPICUS:

Atoyac

TYPICAL MATERIAL: HOSTS:

(Mexico)

holotype M!

in

BM.

unknown

DISTRIBUTION:

Neotropic region:

MEXICO:

Atoyac

(Vera Cruz),

BM!

Rio de

las Cenyas (8 mi. NW of Acaponeta, Nayarit), CAl ECUADOR: Coca, TW! Napo and Coca Rivers, OL! TW! BRAZIL: Teodoro Sampaio (Sao Paulo), OL! TW!

Nova Conquista

NOTES:

(Bahia),

this species

is very

TW!

Surumu

(Roraima),

near the Nearctic

chief difference is concerning

the presence of two

posterior surface of propodeum

OL!

longitudinal

D. atriventris no keel

(in

TW!

D. atriventris (Cresson). The keels on

is visible).

47. Deinodryinus rivularis n. sp.

FEMALE: fully winged; length 3,80 mm; testaceous-reddish, with antennal segments 9-10 darkened; posteri or edges of eyes and part I y temp les blackened;

a

small

anterior marg in;

darkened median

sutu re between

mesosternum black; thickened;

antennal

10:

10:

9:

10:

frontal

=

clubs of femora

3,5;

scutum

brown;

antennae weakly distally

in following

proportions:

head dull,

granulate;

frons rugose,

two

lateral

weak

longitudinal

keels connecting ocellar triangle

OOL

=

is visible near the

segments

line and with

longitudinal OL

13;

area on

metanotum and propodeum blackened;

10;

OPL

=

10;

TL

=

6;

15:

keels;

to occipital

occipital

8:

with

19:

14:

11:

incomplete

vertex without carina;

POL = 3;

carina complete;

pronoturn

shiny, weakly granulated, with pronotal tubercles reaching tegulae; posterior surface of pronotum nearly 1,5 times as broad as long, with numerous short

long i tud i na I keel s near the posterior marg in;

surface sculptured by some strong metanotum shiny, reaching without

without sculpture;

approximately 0,75

radial

bands, cell

one beneath open;

distal

keels;

keels;

scutum,

posterior scutellum and

notaulices incomplete,

length of scutum;

longitudinal or transversal

transversal cells;

smooth,

transversal

propodeum reticulate rugose,

fore-wings with

two dark

the pterostigma and another on part of radial

vein

about as

the basal

long as

240

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

".::: :.:-

A

Fig.

147 -

cells of females of Deinodryinus nigricans

Radial

and pilosifrons Fenton

proximal 3:

7:

part

15:

27;

distally than with

(20:20);

fore tarsal

enlarged claw the proximal

lamellae

Rio

HOSTS:

Itenez

148)

lamellae;

(one of these very

TYPICAL MATERIAL:

segments in following proportions:

prominence;

two rows of nearly 40

MALE: unknown LOCUS TYPICUS:

(Fig.

long);

(Cameron) (A)

(B).

with

13:

1 bristle located further

segment 5 of fore

tarsi

(Fig.

148)

apex with a group of nearly 7

tibial

spurs 1,1,2.

(Bolivia)

holotype F!

in AM.

unknown

DISTRIBUTION:

Neotropic region:

BOLIVIA:

Rio

Itenez (Dept.

Beni),

AM!

48. Deinodryinus aequalis n. sp.

FEMALE: unknown MALE: fully winged; testaceous;

distally thikened, segments

length 3,25-4,25 mm;

antennae testaceous; with

black;

legs brown,

with

clypeus and mandibles tarsi

ligth;

antennae not

hairs longer than breadth of segments;

in following proportions:

15: 9:

20:

10:

11:

13:

12:

antenna I

12:

12:

14;

head dull, reticulate rugose; frons fully reticulate rugose, with frontal line; region behind ocelli punctate or reticulate rugose and with two obi ique keels connecting carina complete;

posterior ocell i

POL = 5;

OL = 3,5;

scutellum and metanotum shiny, among punctures; margin of scutum;

to occipital

carina;

OOL = 11; OPL = 6;

smooth,

finely punctate,

notaulices almost complete,

occipital

TL = 8;

scutum,

without sculpture

reaching almost the posterior

propodeum reticulate rugose;

posterior surface of

Anteoninae

241

propodeum with median area weakly rugose, two

lateral

longitudinal

keels and by

wings with one weak dark

Fig.

distal (Fig.

almost smooth,

vein

longer than

with broad apical

band beneath

proximal

branch wrapping

part penis;

LOCUS TYPICUS; S.J. Barreiro (Brazil) TYPICAL MATERIAL: holotype M! and 3 paratypes MM! MM! in OL. HOSTS:

surrounded by

anterior transversal

keel;

fore-

the pterostigma;

Chela of Deinodryinus rivularis n.

148 -

part of radial 149)

transversal

an

sp.

(type)

(17:15); tibial

gonoforceps

spurs 1,1,2.

2 paratypes

in TW;

unknown

DISTRIBUTION:

Neotropic region:

Bocaina), OL!

TW!

BRAZIL:

S.

J.

Barreiro (Serra da

49. Deinodryinus minor n. sp.

FEMALE: fully winged; length 3,60 mm; head black, with clypeus and mand i b les testaceous; antennae and legs testaceous; thorax, propodeum and abdomen

testaceous;

anterior half of pronotum brown;

antennae long,

weakly distally thickened;

antenna I segments in following proportions:

7:

11:

17:

14:

12:

head rugose; longitudinal

11:

11:

frons

11:

with

keels;

14;

head dull,

incomplete frontal

occipital

granulated;

I ine and with

carina complete;

two

vertex without

lateral

POL ~ 3;

OL ~ 4;

keels connecting ocellar triangle to occipital

carina;

an anterior transversal

OPL

furrow;

~

8;

TL

pronotal

~

5;

pronotum shiny,

lobes reaching

tegulae;

with

weak

longitudinal

~

7;

13:

anterior half of

pronotum weakly granulated,

OOL fully

242

Mem.

Fig.

149 -

Amer.

Male genitalia of

En!.

sculptured by strong than

long

(22:20);

sculpture;

transversal

scutum; keels; open;

(15:17); proximal

prominence;

nearly 41

lamellae;

very MALE:

long);

tibial

150)

almost reaching without

1 bristle

apex with a

Itenez

(Bolivia)

holotype F!

(Fig.

group of nearly 6

spurs 1,1,2.

Rio

TYP:CAL MATERIAL:

transversal or bands;

in AM.

without

longitudinal radial

14:

3:

6:

located further distally

segment 5 of fore tarsi

broader

smooth,

the posterior margin of

in following proportions:

with

unknown

LOCUS TYPICUS:

area near the

vein slightly shorter than proximal

segments

(Fig.

except for a small

without dark transversal

part of radial

fore tarsal

removed.

keels,

propodeum reticulate rugose,

enlarged claw

1984

scutellum and metanotum shiny,

incomplete,

fore-wings hyaline, distal

37,

posterior surface of pronotum slightly

scutum,

notaulices

no.

Deinodryinus aequalis n. sp. (type): right ha I f

posterior margin smooth;

Ins!.,

150)

cell

part 19: than

30; the

with 2 rows of

lamellae

(one of these

Anteoninae

150 -

Fig.

HOSTS:

243

Chela of Deinodryinus minor n.

sp.

(type)

unknown

DISTRIBUTION:

Neotropic region:

BOLIVIA:

Rio

Itenez

(Dept.

Beni),

AM!

50. Deinodryinus benianus n. sp.

FEMALE:

unknown

(probably

other species known from

this species

the same

is the male of one of the two

locality:

D. rivularis and minor; the

males of these two species are unknown; only breeding experiments can finally answer this question). MALE:

fully

winged;

length 3,20 mm;

and mandibles testaceous; hind femora partly brown;

black;

anterior margin of clypeus

antennae testaceous;

legs yellow,

hind coxae partly brown;

hairs nearly as long as breadth of segments;

with mid and

antennae hairy,

antennae not distally

with

Mem.

244

thickened; 11:

12:

antenna I segments

10:

12:

the occipital longitudinal

16;

En!.

frontal

parallel

posterior ocelli scutum,

keels;

scutellum

than distal

bands;

sl ightly

shorter than penis;

lOCUS TYPICUS:

Rio

TYPICAL MATERIAL:

Fig.

=

Itenez

areas;

151)

with

tibial

8:

21:

punctate;

track of two

4,5;

Ol

=

3;

OOl

smooth,

notaulices incomplete, median

12:

11:

occipital

incomplete

vein

8;

OPl

=

almost reaching

rugose,

without

longer than

without dark proximal

part

branch wrapping penis,

1,1 ;2.

(Bolivia)

holotype M!

in AM.

unknown

151

-

Male genitalia of

Deinodryinus benianus ha I f

removed.

8;

area of posterior surface

large internal

spurs

=

strongly punctate,

fore-wings hyaline,

part of radial

gonoforceps

16:

temples and vertex near

propodeum reticulate

lateral

(Fig.

with

and metanotum shiny,

(14:12);

1984

two oblique keels connecting

POL

keels on posterior surface;

not more excavated

HOSTS:

frons

with

cari'1a;

without sculpture among punctures;

transversal

37,

not reticulate rugose,

vertex

to occipital

no.

reticulate rugose;

line complete;

the posterior margin of scutum;

longitudinal

Ins!.,

in following proportions:

head shiny,

carina almost fully

carina complete;

Tl = 7;

Amer.

n.

sp.

(type):

left

Anteoninae

DISTRIBUTION:

Neotropic region:

BOLIVIA:

245

Rio

Itenez

(Beni),

AM!

51. Deinodryinus itenezi n. sp.

FEMALE:

unknown

(probably

this species

is the male of one of the two

other species known from the same locality: D. rivularis and minor; the males of these two species are unknown; only breeding experiments can finally MALE:

answer this question). fully

testaceous,

winged;

length 2,0 mm;

black;

with coxae little darkened;

distally;

antenna I hairs

segments

in following proportions:

segments 4-9 nearly

mandibles,

antennae and

antennae not distally

longer than breadth of segments;

as

antennal

7: 7: 5: 5: 5: 5: 5: 6:

12:

long as broad;

head shiny,

legs

thickened; 11;

antennal

reticulate rugose;

frons without longitudinal keels; occipital carina complete; vertex with little visible longitudinal keels connecting posterior ocelli to occipital carina; POL = 10; OL and metanotum shiny,

= 8;

punctures;

incomplete,

scutum; without

notaul ices

dark

median

transversal

than proximal penis,

OOL = 7; OPL = 6; TL = 5; scutum, scutellum smooth, finely punctate, without sculpture among reaching

propodeum reticulate rugose,

posterior surface; part

without

two

two

longitudinal

area shiny and smooth; bands;

(3:10);

internal

approximately 0,30

with

distal

branch

fore-wings hyal ine,

part of radial

gonoforceps

(Fig.

wrapping

152)

penis;

length of

keels on

vein much shorter

slightly

tibial

shorter than

spurs

1,1,2.

LOCUS TYPICUS: Rio Itenez (Bolivia) TYPICAL MATERIAL: holotype M! in AM. HOSTS:

unknown

DISTRIBUTION:

Neotropic region:

BOLIVIA:

Rio

Itenez

(Beni),

AM!

52. Deinodryinus biloboides n. sp.

FEMALE:

fully

winged;

length 5,80 mm;

cl ypeus and ma I ar space testaceous; 6-10 brown;

propectus black,

with anterior, whitish;

posterior and

pronotal

abdomen

mid and hind

12:

12:

11:

lateral

reddi sh;

13;

margins

testaceous;

legs testaceous, partly

dull;

without

to occipital

vertex behind posterior ocell i

but with numerous complete frontal

longitudinal

line;

POL

=

6;

keels; OL

=

lobes

meta thorax and hind coxae part I y

antennae distally 23:

occipital

7:

45:

20:

20:

carina complete;

keels connecting posterior ocell i not reticulate rugose,

frons reticulate rugose, 3;

segmen ts

pronotum black,

dorsal

with

black;

in following proportions:

head reticulate rugose, longitudinal

with

mesothorax,

region behind ocell i carina;

with mandibles,

side testaceous;

testaceous;

tibiae and femora

thiCkened antenna I segments 16:

with dorsal

tubercles partly

propodeum black; black,

head black,

antennae testaceous,

OOL

=

18;

OPL

=

13;

with TL

=

12;

246

Mem.

Fig.

152 -

Amer.

En!.

Ins!.,

no.

37,

1984

Deinodryinus itenezi n. sp. (type): right

Male genital ia of

half removed.

pronotum with pronotal

tubercles reaching

transverse impression and pronotum dull, punctate, reaching

alutaceous;

tegulae,

two dorsal

scutum,

longitudinal

the basal

radial

vein shorter than proximal

cells and another beneath

following proportions:

18:

8:

18:

part 24:

notaulices thin,

lobes;

smooth,

incomplete,

propodeum reticulate rugose;

keels and with median area shiny,

fore-w i ngs wi th

one on

anterior strong prominent

scutellum and metanotum shiny,

length of scutum;

not ret icu I ate rugose;

with an

transverse very

without sculpture among punctures; approximately 0,5

posterior surface without smooth,

with

two dark

(22:28);

54;

transversa I bands,

the pterostigma; fore tarsal

enlarged claw

distal

part of

segments

(Fig.

153)

in

with one

bristle located further distally than the proximal prominence; segment 5 of fore tarsi (Fig. 153) with two rows of nearly 40 lamellae; apex with a group of nearly 25 MALE:

lamellae;

tibial

spurs 1,1,2.

unknown

lOCUS TYPICUS:

Mouth of Rio Baures

TYP ICAl MATER IAl:

holotype F!

(Bolivia)

in AM.

Anteoninae

247

I

Fig.

HOSTS:

153 -

sp.

(type)

unknown

DISTRIBUTION: Dep t.

Chela of Deinodryinus biloboides n.

Ben i ),

Neotropic region:

AM!

BOLIVIA:

Mouth of Rio Baures (Rio

Itenez,

248

Mem.

Amer.

Ent.

Ins!.,

no.

37,

1984

53. Deinodryinus fluviatilis n. sp.

FEMALE:

fully

clypeus,

antennal

testaceous,

winged;

antennae distally

without

occipital

keels,

smooth,

OPl = 16;

Tl = 14;

tuberc I es reach i ng broad as

long,

with some punctures, to occipital

tegu I ae;

beneath

except

carina;

shiny,

smooth,

without transversal

transversal

the pterostigma;

frontal

bands, distal

line

two oblique keels

POL = 4;

Ol = 4;

without sculpture; lobes;

without sculpture;

OOl = 14;

pronotal tw i ce as

scutum, notaulices

the posterior margin of scutum; or

longitudinal

surface with median area shiny and smooth, with two dark

and

in

posterior surface of pronotum near I y

almost reaching

reticulate rugose,

antenna I segments

two prominent transverse dorsal

scutellum and metanotum shiny, almost complete,

antennae

abdomen

region of head behind ocell i partly

pronotum :1airy,

with

propodeum,

keels on frons and vertex;

carina complete;

connecting posterior ocelli

with mandibles,

13: 10: 10: 9, 9: 9: 11; head shiny,

with numerous longitudinal

complete;

thorax,

thickened;

17: 10: 28:

following proportions:

head black,

malar space testaceous;

wi th segments 7-10 brown;

legs testaceous; rugose,

length 4,40 mm;

sockets and partly

keels;

without areolae;

one on the basal

part of radial

propodeum

posterior fore-wings

cells and another

vein nearly

as

long as

(21 :21); fore tarsal segments in following proportions: 15: 4: 11: 17: 41; enlarged claw (Fig. 154) with 1 bristle located further distally than the proximal prominence; segment 5 of fore tarsi (Fig. 154) proximal

part

Fig.

with

154 - Chela of Deinodryinus fluviatilis n. sp.

two rows of nearly 23

lamellae; MALE:

tibial

lamellae;

spurs 1,1,2.

unknown

lOCUS TYPICUS:

Rio Mamore

(Bolivia)

apex with a

(type)

group of nearly

17

Anteoninae

TYPICAL MATERIAL: HOSTS:

holotype F!

249

in AM.

unknown

DISTRIBUTION: Rio Grande,

Neotropic region:

Dept.

Beni),

BOLIVIA:

Rio Mamore (5 km NW mouth of

AM!

54. Deinodryinus cuzcanus n. sp.

FEMALE: MALE:

unknown

fully

winged;

testaceous;

length 2,25 mm;

legs brown,

with

tarsi

black;

ligth;

antennae and mandibles

antennae not distally

with segments 3-5 triangular and with distal

Fig.

Male genitalia of Deinodryinus cuzcanus n.

155 -

ha I f

apex;

antennal

6:

6:

7:

10;

segments with

sp.

proximal

(type):

right

removed.

segments in following proportions:

antennal

thickened,

apex broader than

hairs

10: 6,5:

5,5:

5:

4:

5:

longer than breadth of

segments; antennal segments 3-5 not uniformly hairy, but with groups of hairs on distal apex of each segment; head dull, fully reticulate rugose; frontal

I ine absent;

occipital

frons with two

carina complete;

lateral

obi ique keels connecting posterior ocell i 4,5; OOL punctate,

=

5;

OPL

=

5;

TL

keels around

the orbits;

region of head beind posterior ocell i

=

5;

to occipital

scutum shiny,

without sculpture among punctures;

carina;

smooth,

without

POL = 7;

hairy,

finely

notaulices incomplete,

OL

250

Mem.

Fig.

156 -

Amer.

En!.

reaching

approximately 0,30

smooth,

fi nel y

median

area shiny,

transversal part

(4:11);

without

bands;

smooth,

gonoforceps

internal

1984

with a

strong

not rugose;

part of radial (Fig.

scutellum and metanotum

without scu I pture among punctures; transversal

posterior surface with

distal

37,

removed.

length of scutum;

punctate,

propodeum reticulate rugose, and posterior surface;

no.

Deinodryinus panamensis n. sp. (type): right

Male genitalia of

ha I f

sh i ny,

Inst.,

155)

two

fore-wings hyaline,

between dorsal without dark

vein much shorter than

with distal

branch wrapping penis;

keel

long i tud i na I keel s;

tibial

proximal

inner pointed process and spurs 1,1,2.

251

Anteoninae

LOCUS TYPICUS:

Quincemil

TYPICAL MATERIAL: HOSTS:

(Peru)

holotype M!

in OT;

1 paratype M!

in AM.

unknown

DISTRIBUTION: Turrialba,

Fig.

Neotropic region:

PERU:

Quincemil

(Cuzco),

OT!

COSTA RICA:

AM!

Males of Deinodryinus

157 -

(B):

trinidadi n. sp.

and noyesi n.

(A)

sp.

length of notaul ices.

55. Deinodryinus panamensis n. sp.

FEMALE: MALE:

unknown

fully

testaceous,

winged;

legs black,

18:

16:

without two

head shiny,

OPL = 16;

keels,

TL = 16;

occipital

hairy,

almost reaching punctate,

propodeum reticulate rugose,

part of radial

gonoforceps (Fig.

vein with distal

spurs 1,1,2.

~OCUS

TYPICUS:

Barro Colorado

TYPICAL MATERIAL:

holotype M!

-IOSTS: unknown JISTRIBUTION: Neotropic region: JA!

4;

carina;

or

longitudinal

part nearly as it's,

long as proximal

however, open,

because

is partly broader and partly effaced;

156) shorter than penis,

tibial

=

two

band beneath the pterostigma;

apparently closed: vein

OL

with

to occipital

without transversal

cell

the distal

frons 7;

the posterior margin of scutum;

radial

radial

=

without sculpture among

fore-wings with one dark transversal radial cell

vertex not

without sculpture among punctures;

keels;

open;

POL

region behind posterior ocelli

finely punctate,

notaul ices almost complete,

(10:10);

antennae not

carina complete;

fully reticulate rugose;

scutellum and metanotum shiny,

part

mandibles

last segment dark;

ligth;

oblique keels connecting posterior ocelli

scutum shiny,

punctures;

partly

with

strongly reticulate rugose;

strongly punctate;

longitudinal

longitudinal

clypeus testaceous;

strongly hairy, with hairs longer than breadth of segments in following proportions: 21: 10: 29: 17: 16:

15: 20;

reticulate rugose, OOL = 15;

black;

antennae testaceous,

with fore tibiae and fore tarsi

distally thickened, segments; antennal 18:

length 4,80 mm;

with teeth dark;

I.

with branch wrapping penis;

(Panama)

in DA. PANAMA:

Barro Colorado

I.

(Canal

Zone),

252

Fig.

Mem.

158 -

Amer.

En!.

Ins!.,

no.

Female of Lonchodryinus ruficornis (England)

37,

1984

(Dalman)

from

Point of Ayr

Anteoninae

253

3. Lonchodryinus Kieffer

Lonchodryinus Kieffer 1905a: 95. Prenanteon Kieffer 1913b: 301 (syn. proposed by Olmi 1983); type species Prenanteon melanocerum (Kieffer 1905), design. by Richards 1935.

Psilanteon Kieffer 1913b: 301 (n. syn.); type species Psilanteon aequalis (Kieffer 1905), orig. desig. Lonchodryinus Kieffer: Kieffer 1914b: 14. Prenanteon Kieffer: Kieffer 1914b: 199. Psilanteon Kieffer: Kieffer 1914b: 212. Prenanteon Kieffer: Berland 1928: 153. Prenanteon Kieffer: Richards 1937: 119. Psilanteon Kieffer: Richards 1937: 119. Prenanteon Kieffer: Richards 1939: 235. Prenanteon Kieffer: Muesebeck and Walkley 1951: 1041. Prenanteon Kieffer: J.F. Perkins 1976: 23. Prenanteon Kieffer: Krombein 1979: 1242. Lonchodryinus Kieffer: Olmi and Currado 1979a: 342. Prenanteon Kieffer: Moczar 1983a: 186. Lonchodryinus Kieffer: Olmi 1983: 81.

Fig.

159 -

Female of Lonchodryinus ruficornis (Dalman) from Point of Ayr (England)

254

Fig.

Mem. Amer.

160 -

Male of

Ent.

Inst.,

no. 37,

1984

Lonchodryinus ruficornis (Dalman) from Hunstanton (Norfolk, England)

Type species: FEMALE (Figs.

Lonchodryinus tricolor Kieffer 1905, monotypic. 158,

159):

fully winged

(Fig.

158) or occasionally

brachypterous (Fig. 162); maxillary palpi with 6 segments; labial palpi with 3 segments; fore-wing with distal part of radial vein as long as or longer than proximal part (Figs. 158, 159); occasionally slightly shorter; inner side of enlarged claw with a proximal

prominence bearing one

255

Anteoninae

Fig.

161

Male of Lonchodryinus ruficornis

-

(Norfolk,

bristle

(Fig.

strong

transversal

MALE (Figs. labial palpi

164); occipital keel

from Hunstanton

propodeum usually without a

and posterior surface.

160, 161): fully winged; maxj Ilary palpi with 6 segments; with 3 segments; fore-wing with distal part of radial vein

shorter;

proximal

pterostigma less

propodeum usually posterior surface; 167);

carina complete;

between dorsal

as long as or longer than slightly

(Dalman)

England)

part

(Figs.

160,

than four times as

without a strong

transversal

gonoforceps without apical

keel

161); occasionally long as broad; between dorsal

branch wrapping penis

and (Fig.

vertex of head without two obi ique keels connecting posterior ocell i

to occipital antennal

carina;

DISTRIBUTION: HOSTS:

antennal

hairs at most as

long as breadth of

segments. worldwide

Cicadellidae (i nc I ud i ng Typhlocybinae)

SPECIES:

12

PALAEARCT I C SPEC I ES OF LONCHODRYINUS

Three species are known.

KEY TO THE SPECIES

256

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

FEMALES

Fore-wing

-

reduced, short, without pterostigma (Figs. 162, 163) .. .••••.•••••••••••....•...••.. 1. subapterus (Kieffer) Fore-wing not reduced, with pterostigma (Figs. 158, 159) . . . . . . . . . • • • • • • • • • • • • • • . • • • • • • • • • • • •• 2. ruficornis (Da I man) The female of L.

nepalensis n. sp. is unknown

MALES

Temples nearly emarginated -

twice as

(Fig.

long as POL;

167) . . . . . . .

Temples nearly as

long as,

3.

gonoforceps apically

nepalensis n. sp.

slightly

longer or

shorter

than POL;

gonoforceps apically not emarginated (Fig. 166) . . . . . . . . . . . . . . . . . • • • • • . . • • • • • • • • . • • • . • • • • • • • •• 2. ruficornis (Da I man) The male of L.

subapterus (Kieffer) is unknown.

1. Lonchodryinus subapterus (Kieffer)

n.

comb.

Anteon subapterus Kieffer in Kieffer et Marshall 1905: 138. Mystrophorus subapterus (Kieffer): Kieffer 1914b: 195. Prenanteon ruficornis (Dalman) var. subapterus (Kieffer): Richards 1939:

243.

Prenanteon subapterus (Kieffer): J.F. Perkins 1976: 23.

FEMALE

(Figs.

reaching 0,5

162,

163):

micropterous,

length of propodeum;

and tegulae yellow,

antennae fully

with fore-wing reduced and not

length 3,81

mm;

yellow-reddish,

black,

with mandibles

legs yellow-reddish,

with proximal region of hind coxae and distal part of hind femora brown; head shiny, smooth, finely punctate; pronotum hairy, rugose, strongly punctate; scutum shiny, smooth, finely punctate; notaulices incomplete, extending distinctly beyond 0,5 length of scutum; scutellum shiny, smooth, without sculpture;

metanotum rugose,

dull;

propodeum reticulate rugose;

posterior surface with 2 longitudinal keels; another median keel is visible between these two keels; fore tarsal segment 1 twice as long as 4 (10: 5); enlarged claw

(Fig.

164)

segment 5 of fore tarsi lamellae; MALE:

with a

(Fig.

proximal

164)

prominence bearing one bristle;

with 2 rows of 3 (proximal)

apex with a group of 7 lamellae;

unknown

tibial

spurs 1,1,2.

+ 17

257

Anteoninae

Fig.

162 -

Female of Lonchodryinus subapterus ( Ireland)

LOCUS TYPICUS:

Aviemore

TYPICAL MATERIAL:

(Scotland)

holotype F!

in BM.

(Kieffer)

from Mullanx

258

Mem. Amer.

HOSTS:

Ent.

Inst.,

no.

37,

1984

unknown

DISTRIBUTION:

Palaearctic region:

U.

S. S.

R.:

Russian Lapland

(Hellen

1919); ENGLAND: Church Stretton (Caer Caradoc, SP.), BM! Salop (J. F. Perkins 1976); Hopton Fen (Suffolk), BM! SCOTLAND: Aviemore (Inverness East),

BM!

Wichen sedge Fen

Hawksmoor (Richards 1939); IRELAND: Mullanx, BM! NOTES:

(Cambs),

BM!

Corrie (Gretna,

Lichfield Dumfries)

probably this species is a variety of L. answer th i 5 quest ion.

(Staffs),

OX!

(Richards 1939);

ruficornis. Only breeding

experi ments can

Fig.

163 -

Female of Lonchodryinus subapterus (Kieffer) from Mullanx (Ireland)

2. Lonchodryinus ruficornis (Da I man)

Gonatopus ruficornis Dalman 1818: 83. Gonatopus frontalis Dalman 1818: 84 (n. syn.); locus typicus Vastergotland

(Sweden) (discussion on synonymy:

see page 267).

Gonatopus basalis Dalman 1818: 84 (syn. proposed by Richards 1939); locus typicus Smaland (Sweden). Gonatop us fuscicornis Da I man 1818: 87 (syn. proposed by Richards 1939); locus typicus Vastergotland (Sweden). Dryinus iongicornis Dalman 1823: 10 (syn. proposed by Richards 1939); locus typicus Smaland (Sweden).

Anteon i nae

259

Dryinus crassimanus Haliday in Curtis 1828: 206 (syn. proposed by Richards 1939); locus typicus Holywood (Ireland). Dryinus daos Walker 1837: 423 (syn. proposed by Dlmi 1983); locus typicus London (England) (the M! selected as type by Richards 1935 is very damaged and not identifiable: a second male of daos in Haliday collection is a ruficorne). Dryinus ilus Walker 1837: 424 (syn. proposed by Richards 1939); locus typicus Eng I and. Dryinus misor Walker 1837: 424 (syn. proposed by Richards 1939); locus typicus Eng I and. Dryinus lapponicus Thomson 1860: 176 (n. syn.); locus typicus South part of Lap I and

(Sweden).

Dryinus retusus Thomson 1860: 176 (n. syn.); locus typicus Norway. Chelogynus lepidus Forster 1861: 39 (n. syn.); locus typicus Roseggthal

(Switzerland).

Chelogynus frontalis (Dalman): Dalla Torre 1898: 540. Anteon aequalis Kieffer in Kieffer et Marshall 1905: 177 (n. syn.); locus typicus Glen Lyon (Scotland). Anteon frontalis (Dalman): Kieffer in Kieffer et Marshall 1905: 179. Anteon melanocera Kieffer in Kieffer et Marshall 1905: 179 (syn. proposed by Richards 1939); locus typicus Mugdock (Scotland). Anteon crassiscapus Kieffer in Kieffer et Marshall 1905: 180 (syn. proposed by Richards 1939); locus typicus Dieppe (France). Anteon vitellinipes Kieffer in Kieffer et Marshall 1905: 181 (n. syn.)j locus typicus Clober (Scotland). Anteon proximus Kieffer in Kieffer et Marshall 1905: 182 (n. syn.) jlocus typicus Svabhegy (Hungary). Anteon foersteri Kieffer in Kieffer et Marshall 1905: 183 (n. syn.)j locus typicus unknown. Anteon procericornis Kieffer in Kieffer et Marshall 1905: 184 (syn. proposed by Richards 1939); locus typicus Sutherland (Scotland). Anteon longicornis (Dalman) var. ruficornis (Dalman): Kieffer in Kieffer et Marshall

1905:

186.

Anteon luteipes Kieffer in Kieffer et Marshall 1905: 187 (syn. proposed by Richards 1939); locus typicus Meudon (France). Anteon declivis Kieffer in Kieffer et Marshall 1905: 188 (syn. proposed by Richards 1939); locus typicus Clydesdale (Scotland). Anteon pallidinervis Kieffer in Kieffer et Marshal 1905: 189 (syn. proposed by Richards 1939); locus typicus Kill i ngdock (Scot I and). Anteon dolichocerus Kieffer in Kieffer et Marshall 1905: 190 (syn. proposed by Richards 1939); locus typicus Sevres (france). Anteon integer Kieffer in Kieffer et Marshall 1905: 206 (n. syn.); locus typicus Mugdock (Scotland). Anteon parvicollis Kieffer in Kieffer et Marshall 1905: 206 (syn. proposed by Richards 1939); locus typicus Mesnil-Ie-Roi (France). Anteon pyrenaicus Kieffer in Kieffer et Marshall 1905: 207 (syn. proposed by Richards 1939); locus typicus Ax-Ies-Thermes (France).

260

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

Anteon curvinervis Kieffer in Kieffer et Marshall 1905: 208 (n. syn.); locus typicus Bonar Bridge (Scotland). Anteon fractinervis Kieffer in Kieffer et Marshall 1905: 208 (syn. proposed by Richards 1939); locus typicus Clober (Scotland). Anteon hyalinipennis Kieffer in Kieffer et Marshall 1905: 209 (syn. proposed by Richards 1939); locus typicus Dairy (Scotland). Anteon fuscipennis Kieffer in Kieffer et Marshall 1905: 209 (n. syn.); locus typicus Maroth (Hungary). Anteon longifilis I
inval id;

type in BU

according

to

Moczar 1967).

Anteon daos (Walker): Kieffer in Kieffer et Marshall 1906: 521. Anteon vitellineipes Kieffer: Cameron 1907: 32. Anteon pallidicornis Kieffer: Cameron 1907: 33. Anteon crassimanus (Haliday): Kieffer 1907b: 25. Anteon ilus (Walker): Kieffer 1907b: 26. Anteon lapponicus (Thomson): Kieffer 1907b: 26. Anteon lepidus (Foerster): Kieffer 1907b: 26. Anteon retusus (Thomson): Kieffer 1907b: 27. Antaeon morleyi Chitty 1908: 144 (syn. proposed by Richards 1939); locus typicus Foxha II (Eng I and). Antaeon luffnessensis Chitty 1908: 145 (syn. proposed by Richards 1939); locus typicus Luffness Links (Scotland). Antaeon beaumonti Chitty 1908: 145 (n. syn.); locus typicus Chobham (England) •

Prenanteon frontalis (Dalman): Kieffer 1913b: 301. Anteon daos (Walker): Kieffer 1914b: 164. Chelogynus walkeri Kieffer 1914b: 185 (syn. proposed by Richards 1939); locus typicus Holywood (Ireland). Prenanteon vitellinipes longifilis (Kieffer): Kieffer 1914b: 203. Prenanteon longicornis (Dalman): Kieffer 1914b: 205. Prenanteon pyrenaicus (K i effer) var parcepunctatus Kieffer 1914b: 209 (syn. proposed by Richards 1939); locus typicus Clober (Scotland). PrelJanteon palustris Oglobl in 1924a: 102 (n. syn.); locus typicus Strbske pleso

(Czechoslovakia).

Prenanteon filicornis Oglobl in 1924b: 127 (n. syn.); locus typicus Pisek

(Czechoslovakia). frontalis (Dalman):

Prenanteon PrelJanteon PrelJanteolJ PrelJanteon Prenanteon Prenanteon Prenanteon

Berland

1928:

154.

luteipes (Kieffer): Berland 1928: 155. crassiscapus (Kieffer): Berland 1928: 156. pyrenaicus (Kieffer): Berland 1928: 156. parvicollis (Kieffer): Berland 1928: 156. longicornis (Kieffer): Berland 1928: 157. luteipes (Kieffer): Maneval 1935: 6.

Anteoninae

Prenanteon Prenanteon Prenanteon Prenanteon

261

daos (Walker): Richards 1939: 236. ruficornis (Dalman): Richards 1939: 240. frontalis (Dalman): Richards 1939: 245. euscelisi Haupt 1941: 61 (n. syn.); locus typicus Ha II e

(Germany) •

Prenanteon luteiceps (lapsus}(Kieffer): Menozzi 1942: 37. Prenanteon longicornis (Dalman): Cavro 1950: 5. Prenanteon semenovi Ponomarenko 1970: 428 (n. syn.); locus typicus Gremyachka

(U.

S.

S.

R.).

Prenanteon foveatus Richards 1971: 48 (syn. proposed by Olmi 1983); locus typicus Si I wood Park (Eng I and) • Prenanteon frontalis (Dalman): J.F. Perkins 1976: 23. Prenanteon ruficornis (Dalman): J.F. Perkins 1976: 23. Prenanteon longicornis (Dalman): J.F. Perkins 1976: 24. Prenanteon foveatus Richards: J.F. Perkins 1976: 24. Prenanteon basalis (Dalman): J.F. Perkins 1976: 25. Prenanteon ruficorne (Dalman): Ponomarenko 1978: 21. Prenanteon semenovi Ponomarenko: Ponomarenko 1978: 21. Prenanteon ruficornis (Dalman): Moczar 1983a: 186. Prenanteon pektusanense Moczar 1983a: 187 (n. syn.); locus typicus Mt.

Pektusan

(Korea).

Prenanteon basale (Dalman): Moczar 1983a: 189. Prenanteon c1avatum Moczar 1983a: 189 (n. syn.); locus typicus Mt. Pektusan

(Korea).

Prenanteon Prenanteon Prenanteon Prenanteon

FEMALE

(Figs.

occasionally dark;

melanocera (Kieffer): Moczar 1983b: 200. basale (Dalman): Moczar 1983b: 201. longicornis (Da I man): Moczar 1983b: 202. c1avatum Moczar: Moczar 1983b: 203.

158,

159):

fully

winged;

length 2,25-4,68 mm;

head and thorax brown-reddish;

occasionally frons mainly reddish;

black;

mandibles yellow,

with

teeth

sometimes malar space and frons

frontalis Dalman); clypeus yellow, black or (var. longicornis. daos and ret usus ) or brown, with segments 1-2 testaceous (var. ruficornis and basalis); mainly

whitish or yellow

redd ish-dark;

(var.

antennae testaceous

occasionally

antennae brown

occasionally

also segments 6-10 or 7-10 testaceous;

fu II y

testaceous or with

brown-testaceous; 14:

12:

shiny,

=

7;

12:

10:

strongly

Ol

=

4;

deep,

very

=

10;

black;

tegulae

testaceous;

abdomen

black or

in following proportions:

antennae slightly distally

smooth,

OPl

1-2 or 1-3 testaceous;

=

impression;

8;

12:

thickened;

6:

13:

head

without sculpture among punctures; Tl

=

8;

frons without median

sometimes a

deep groove

legs

pal

groove on

is visible

(in

foveatus and retusus); frontal impression more or less

variable,

pronotum shiny,

15;

punctate,

OOl of

10:

1,

hind coxae basa II y

antenna I segments

10:

disc or with discal holotypes FF!

with segment

with no specific value;

strongly

punctate;

pronotal

occipital

carina complete;

tubercles reaching

tegulae;

the

262

Mem.

Fig.

164 -

Amer.

En!.

Inst.,

no.

37,

1984

Lonchodryinus subapterus (Kieffer) from Mullanx

Chela of

(Ireland)

scutum shiny, strong I y

smooth,

punctate,

without punctures;

occasionally scutum finely or

wi thout scu I pture among punctures;

incomplete, occasionally

long,

the holotype F!

as in

Perkins

1976 notaulices at most reaching mid

as in

the holotype F!

of the var.

basalis (according to J. F.

scutum,

occasionally notaul ices short,

length of scutum);

reaching approximately 0,5

of the var.

typ ica I finely

as in

the

lectotype F!

reaching and

length of scutum,

longicornis (according to J. F. Perkins

1976 notaulices extending distinctly beyond mid occasionally notaul ices shorter, scutum,

notau I ices

reaching approximately 0,65 length of

in

length of scutum);

approximately 0,3

length of

the paralectotypes FF!

of the

ruficornis (i n types the notau I ices are not sha Ilower, narrow and aciculate,

metanotum shiny,

as written by J. smooth,

F.

Perkins 1976);

scutellum and

without sculpture or punctate;

propodeum

Anteoninae

reticulate rugose,

posterior surface with 2 longitudinal

usually as rugose as lateral and

longicornis)

rugose

(as

hyaline;

in

and dull;

nearly

(9:

7);

(as in

part;

of

retusus

vein as

and

long as,

fore tarsal

segment 5 of front

15-20 lamellae;

keels;

the holotypes FF!

occasionally median area shiny,

part of radial

longer than proximal segment 4

areas

the holotypes FF!

distal

263

fore-wing

slightly shorter or slightly

(Fig.

165)

longer than

with 1 row of

apex with a group of nearly 3-5 lamellae;

2 of front tarsus produced into a hook; enlarged claw (Fig. bristle located on proximal prominence; tibial spurs 1,1,2.

Fig.

165 -

basalis

smooth and not

euscelisi);

segment 1 slightly

tarsus

median area of

Chela of

Lonchodryinus ruficornis

(Dalman)

segment

165)

with one

from Granarolo

( Italy)

MALE

(Figs.

160,

161):

mandibles testaceous, testaceous;

5;

winged;

length 2,12-3,75 mm;

teeth brown;

tegulae testac'eous;

occasionally punctate,

fully

with

legs testaceous,

POL = 5;

OL = 2,5;

head shiny, OOl = 8;

frons with or without a more or less deep central

carina complete; among punctures; of scutum,

scutum shiny,

smooth,

notaulices incomplete,

finely

smooth,

finely

finely

OPl

=

4;

punctate,

without sculpture

reaching approximately 0,5

punctate,

TL =

impression; occipital

broad and deep or narrow and finely crenulate;

metanotum shiny,

occasionally

with coxae partly brown;

also clubs of femora partly brown;

without sculpture;

black;

antennae fully brown;

length

scutellum and

without sculpture among

264

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

, I

!

/

Fig.

166 -

(Dalman)

punctures;

Gonoforceps

areas,

long as,

(Switzerland) (B).

keels on

without dark transversal

slightly

almost without sculpture;

bands;

shorter or slightly

lateral

distal

longer than

view variable,

part of radial proximal

narrow or broad

part;

(Fig.

166);

spurs 1,1,2.

LOCUS TYPICUS:

Vastergotland

TYPICAL MATERIAL: lectotype F!

basalis,

(Sweden)

the numerous types are in following collections:

holotype M!

of G.

paralectotypes FF! lectotYpe M! of

lectotype F!

frontalis; in D. daos, lectotype

of G.

of G.

ruficornis,

in ST:

C. fuscicornis, lectotype F! (hereby desi gnated) D. longicornis, holotype F! of D. lapponicus,

and 3 paralectotypes FF!

and 1 paralectotype F! of holotype F! of D. retusus, DU:

Lonchodryinus ruficornis

with 2 longitudinal

occasionally smooth and shiny,

shape of gonoforceps in tibial

ru~ose,

of

and Valetta

usually median area on posterior surface as rugose as

fore-wing hyal ine, vein as

(Ireland) (A)

propodeum reticulate

posterior surface; lateral

(laterally viewed)

from Avimore

/

(hereby PK: F!

designated)

and 2

Pro semenovi; D. crassimanus, lectotype

holotype F! of

holotype F! of

of

in F!

Anteoninae

of Ch.

walkeri; in VI:

lectotype M!

265

(hereby

designated)

and 3

of Ch.

lepidus; in MR: holotype M! of A. luffnessensis; in BM: holotype F! of A. aequalis, holotype F! of A. vitellinipes, holotype M! of A. curvinervis, holotype M! of A. integer, holotype M! of A. beaumonti, holotype F! allotype M! and 2 paratypes FF! of Pl'. foveatus, holotype M! of A. fractznervis, holotype F! of A. pallidinervis, holotype F! of A. dechvis, holotype F! of A. melanocera, holotype F! of A. procericornis, holotype M! of D. ilus, holotype M! of D. misor, holotype M! of A. morleyi, holotype M! of A. hyalinipennis, holotype M! of Pl'. parcepunctatus; in P: holotype F! of A. crassiscapus, holotype F! of A. luteipes, holotype F! of A. dolichocerus, holotype M! of A. pyrenaicus, lectotype M! (hereby designated) of A. parvicollis; in HA: holotype F! of Pl'. euscelisi; in BU: holotype F! of A. halidayi (type in BU, not in DU); types of A. proximus. A. foersteri. A. fuscipennis. Pl'. palustris. Pl'. filicornis. A. longifihs probably lost. HOSTS: in Italy Empoasca flavescens (Flor), Empoasca decipiens Paoli, Asymmetrasca decedens Paol i (= Empoasca decedens) (Menozzi 1942; specimen from Granarolo (BO!), leg. Menozzi, determined by Menozzi as Pl'. luteiceps Kieffer), Jassargus bisubulatus Theny; in England Psammotettix confinis (Dahlb.) (Waloff 1974, 1975; J.F. Perkins 1976), Psamn;otettix nodosus (Rib.) (Richards 1971; Waloff 1975); Psammotettix cephalotes (H.-S.) (Richards 1971), t-'uscelis obsoletus (Waloff 1974, 1975), Euscelis plebejus (Waloff 1974, 1975), Macrosteles laevis Rib. (Richards 1971), £lymana sulph urella and Arthaldeus pascuellus (Wa loff 1974, 1975); in Germany Streptanus sordid us (Zett.) (= lo'uscelis sordidus: Haupt 1941) and, for var. frontalis. Jassargus obtusivalvis (= lJeltocephalus picturatus Fieb.) and t-'rrastun us ocellaris F. (= lJeltocephalus ocellaris: Haup t 1941); in U.S.S.R. Nacrosteles laevis Rib. (Ponomarenko 1978). paralectotypes

(2FF,

DISTRIBUTION:

Palaearctic region:

Pay plateau

1M)!

(Sam-zi-yan,

Provo

JAPAN:

Kamikochi,

Ryang-gang),

BU!

TW!

KOREA:

Sam-zi-yan

Chann-

(Mt.

Pektusan), BU! Mu-do-bong (Mt. Pektusan, Chann-Pay plateau, Provo Ryang-gang), BU! MONGOLIA: Semon Bajanzogt (Central aimak), BU! Bogdo ul.

(Ulan-Baator,

Central

(Changaj Gebirge, Central

aimak),

Namnan

ul

(Chovsgol

S.

HE!

Metsapirtti,

HE!

Ekenas, (AI.),

(Tudosdomb),

HE!

SWEDEN:

(Lpl.),

Eksharad

OL!

Fokstua

Lojo,

(Lpl.),

Opheimsbakken

OT!

Gremyachka HE!

HE! HE!

Mausgnsbyn BM!

R.:

ovoo

OT!

(Dovre),

HE!

Vastergotland, Lapland,

ST!

Messaure,

(Vmld.),

OT!

NORWAY:

Vacz-Szod,

ST!

27°

BU!

(Trondhj.),

Abisko

(Lpl.),

BU!

ST!

OT!

Csevharaszt,

00'

HE!

OT!

~~orway,

Munkacs,

OL!

Sordavala, HE!

Foglo

OT!

Dalaro

BU!

E,

Kantalaks,

(Sjall.),

(Lpl.),

Smaland,

BU!

NEPAL:

N 85°

FINLAND:

Dyrehaven

Hatfjelddal,

Visegrad,

aimak),

Somon Burenchaan

58'

PK!

Riksgransen TW!

HUNGARY: BU!

Hell

HE!

BU!

aimak),

TW!

Somon Culuut

(Ulan-Baator,

Central

(Ryazan Province),

OT!

BU!

(Bulgan

Phakding,

Tammerfors,

CO!

Ichtamir -

Pass Zosijn

Bulgan aimak),

OL!

(Aurdal),

Somon BU!

(Ulan-Baater,

Chutag,

Somon Abzaga

(Katmandu),

U.

Jebrenjokk

BU!

BU!

aimak),

Tosgoni (Somon

aimak),

OT!

Bano

S.

BU!

Gebirge

Bhurumche

aimak),

Archangaj

Skane, ST!

Godollo, BU!

TW!

(Malmen), BU!

Tasnad,

Vacz BU!

266

Mem.

Felsohagi,

BU!

Amer.

Bukkszentielek,

Tihany,

BU!

Mecsek hgs.,

(Bukk),

BU!

Irsa pta.,

Budapest,

BU!

Nagykovacsi,

Ent.

BU!

BU!

BU!

BU!

Bukk h.

BU!

BU!

BU!

Budai

BU!

Unoko,

BU!

Mecsekh,

Ocsa,

Fulophaza,

BU!

Inarcs,

37,

BU!

BU!

H.,

BU!

O-Sebeshely,

BU!

BU!

BU!

BU!

Kunfeherto,

Kurtaberc

BU!

Valko,

Velem,

Hoi ics,

BU!

BU!

Nagyvisnyo,

Budakeszi,

BU!

1984

Csakvar park,

Barszentmihaly,

Borzsony, BU!

no.

Alp Kudsir.,

Dobogoko,

Balvany,

Tompa,

Inst.,

BU!

Szeged,

BU!

Kormochanya, BU!

BU!

SzSztMiklos,

BU!

Rev., BU! Bi harfured, BU! Koszeg, BU! Szod, BU! Tata tovaros, BU! Cs i k Szepvis, BU! Svabhegy (Budapest) (Kieffer in Kieffer et Marshall 1905); Szaohegy (Koszeg), BU! Boroszno, BU! ROMANIA (Nagy 1972); CZECHOSLOVAKIA: Bohemia, TW! M. Tatra, BU! Kobyli (Moravia), BULGARIA:

Mts Rila,

(Slovenija),

CO!

BU!

Gruz

YUGOSLAVIA:

(Dalmacija),

Mosbrucher Hochmoor (Eifel), (Bohmerwald), Berlin,

HU!

HU!

HU!

Weinberg

Beuerburg

Westpreussen),

CO!

HU!

Postojne (Slovenija),

VI!

GERMANY! Halle,

Bellinchen

(Gross-Machnow),

(Oberbayern),

CO!

Westf.),

HU!

HU!

HU!

Veldes

HA!

HA! Schwarzer See

Frechland

(Elbe),

Sachsen),

HU!

Aachen,

Berkesnvercher (Berlin),

HU!

Huben

HU! (0.

Niemegk, Tirol),

HU!

HU!

Kuhhornhatte

Igls

(Tirol),

(N.

OT!

Siebenburgen),

St.

Anton,

BM!

P!

LE!

Finkenkrug

(Berlin), HU! Mecklenburg, HU! Feldwege I. (Unna, Westf.), HU! (Hoheneck, Vogesen), HU! Misdrog, HU! Gipfel (Gr. Wasserkuppe, Gbg.),

HU!

Salesche (Tucheler Haide,

Mehlsheuer (Vagsland,

Saline (Unna,

(Oder) ,

BM!

BM!

HU!

Wien,

Fischbodle Rhon-

AUSTRIA: WA!

P!

Prater (Wien), P! Austria, P! Obergurgl (Tirol), OT! Semmeringgebeit (Reichenau), BM! Reichenau, P! DENMARK: Gentafte, CO! Tipperne (Jylland),

CO!

Ordrup Naes

(Odsherred,

Sjaell.),

CO!

Korsor,

CO!

Hoisteinborg, CO! Aarhus, CO! Vesterhede (Romo, Sdr. Jylland), CO! Espe, CO! Ulfshale, CO! Danmark, CO! HOLLAND: Wijster (Dr.), LE! Asperen, LE! SWITZERLAND: (Neuchatel), Valley

Auvernier, LO!

Duillier

(Engadin),

BU!

LO!

Les Ponts

(Vaud),

LO!

Suchet-Issom.

(Neuchatel),

Aclens (Vaud),

LO!

(Vaud), LO!

Montmollin

LO!

Nyon,

Morteratsch

GV!

Vercorin

(Valais), LO! Arolla (Valais), BM! Aarau, BM! Derborence (Valais), GV! Foret d'Aletsch (Valais), GV! Epalinges (Vaud), LO! Boussens (Vaud), OL! Pleiades

(Vaud),

Roseggthal Cluozza), P!

GV!

Fopps

Dieppe (Seine

(Seine-et-Oise), (Gard),

GV!

Mayens de Syon

(Grigioni),

GV!

(Val

Inf.), P!

Sevres

VI!

(Valais),

Roseg Valley Cluozza),

P!

Meudon

LO!

(Engadin),

GV!

FRANCE:

P!

Ax-Ies-Thermes

P! P!

(Vaud),

Valetta

Le Puy,

(Seine-et-Oise),

Maisons-Laffitte (Seine-et-Oise), (Seine),

Gryon BU!

P!

GV!

(Val

BO!

Bolquere,

Mesnil-Ie Roi Pont du Gard

(Ariege),

P!

Lille

(Nord),

P! Vincennes, P! La Frette, P! Foret de Marassan (Hautes-Alpes) , P! ENGLAND: Boxmoor (HT.), BM! Nunton (Moczar 1967); Eastbourne (Sussex), BM!

Wi mb ledon Com.

(London),

OT!

Cambri dge,

BM!

Sherford Bri dge (DT.),

BM! Chobham (Surrey), BM! Oxford, OT! Rhinefield (Hants.), BM! Ben Rhydding (Yorkshire), HA! Spratton (Northants), BM! Si Iwood Park (Berks.), Fowhall

8M!

Claygate

(Suffolk East),

(Dartmoor,

Devon),

Richmond Park,

OX!

OT!

(SR.), BM!

8M!

Wotton-under-Edge

White Downs

Aldbury

Hunstanton

(HT.),

(Dorking,

BM!

(Norfolk),

(Gloster West),

Surrey),

Point of Ayr

8M!

SCOTLAND:

BM!

(Flint),

BM!

Hay tor BM!

Wood Walton

Fen

Anteoninae

(Cambs.),

BM!

Glen lyon,

Bonar Bridge,

BM!

BM!

(Perths.),

Enochdhu

Killingdock, BM!

BM!

BM!

DU!

Avimore

DU!

Glenasmole

SZ!

SZ!

(Senales, BO!

Res.

(Fife),

BM!

(WX.),

BM!

BM!

(Torino),

Ol!

Co.

TW!

SZ!

(Genova),

MI!

Ol!

(Dublin),

Cavehill (in

OT!

trout's stomach),

1903);

(Aosta), (Cuneo),

OLl Pl!

(Bussoleno

Unserfrau

Granarolo

Monte Fascie (Genova), (Perugia),

BM!

(Belfast),

Pietrabianca

(Cobelli

BM!

(Kirkcudbrigth),

Benedetto Belbo

(Torino),

Monte Peglia

TW!

BM!

(Perths.),

ITALY: Jovencan

S.

Trentino

BO!

Clober,

(Perths.),

Dairy

(Wicklow)

TW!

BM!

BM!

Gollierstown

Naturno (Bolzano),

Rezzoaglio d'Aveto (Genova),

lavagna

BM!

Wicklow,

Kerry,

Noasca

OT!

Ballinluig

IRELAND: Co.

Aosta),

Condove (Torino),

Bolzano),

BM!

Bally Smutton

BM!

(Perths.),

Birks of Aberfeldy

Sutherland,

Ballyteige (Dublin),

BM!

Aviemore, BM!

(Valsavaranche,

la Cassa

Torino),

BM!

(Harwood),

Eaux Rousses

Rannoch Forest

(Stirling),

Clydesdale,

Tentsmuir Nat.

Athdown,

BM!

Mugdock

267

(Bologna), Ol!

Cavi

di

Ol!

NOTES: specimens from Japan are bigger than specimens from other countries. Synonymy questions about L. frontalis were defined in consequence of the discovery of males. Previously I observed in females intermediate forms with frontal spot very wide and very small. Doubts were removed in consequence of the discovery of males. I observed no difference between males of L. ruficornis and frontalis. Frontalis is only a variety with part of frons whitish or yellowish in females. This variety is not frequent. I know the var. frontalis from the following local ities: SWEDEN (Vastergotland), GERMANY (Aachen, Bellinchen, Schwarzer See), SWITZERLAND (Aarau, Mayens de Syon), FRANCE (Pont du Gard), ENGLAND (Oxford, Matley Bog), ITALY (La Cassa, Monte Fascie, Monte Peglia).

3. Lonchodryinus nepalensis n. sp.

FEMALE: MALE:

unknown

fully

winged;

mandibles whitish; testaceous;

legs testaceous,

distally thickened; 12:

11:

length 3 mm;

10: 9:

9:

8:

11;

~

5,5;

OL

~

2,3;

metanotum shiny,

notaul ices incomplete,

malar space, ventral

frontal ~

8;

smooth, ~

5;

TL

LOCUS TYPICUS:

28°

00'

N.

part;

scutum,

8:

6:

13:

without

carina complete; scutellum and

bands;

with

spurs 1,1,2.

85° 00'

E.

length of scutum; or longitudinal distal

keels;

part of radial

the two parts forming a curve;

inner process,

tibial

punctate,

occipital 10;

without transversal

vein nearly as long as proximal 167);

antennae not

without sculpture among punctures;

fore-wing hyaline without dark transversal

(Fig.

~

reaching approximately 0,5

propodeum reticulate rugose,

emarginated

finely

line absent;

OPl

punctate,

gonoforceps without distal

clypeus and

side of segment 1

segments in following proportions:

head shiny,

OOL

finely

with

with hind coxae basally black;

antennal

sculpture among punctures; POL

black;

antennae brown,

(Nepal)

internal

apical

margin

268

Mem. Amer.

En!.

Ins!.,

\ I I [\

Fig.

167 -

no.

37,

1984

/ ./

Male genitalia of Lonchodryinus nepalensis n. I eft ha I f removed.

TYPICAL MATERIAL: holotype M! in aT. HOSTS: unknown DISTRIBUTION: Palaearctic region: NEPAL:

28° 00'

N 85° 00'

sp.

E,

(type):

aT!

Anteoninae

ETHIOPIAN SPECIES OF

269

LONCHUDRYINUS

Two species are known.

KEY TO THE SPECIES

FEMALES

Fore-wing hyal ine, -

wi thout dark transversal bands •.•••••••••• . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1. seyrigi (Benoi t) Fore-wing wi th two dark transversal bands . . . . . . . . . . . . . . . . . . .. ••••••••..•••.•••••••.••••.•• 2. pauliani (Benoi t)

MALES Unknown

1.

Lonchodryinus seyrigi (Benoit) n. sp.

ifirtanteon seyrigi Benoi t 1953b: 421.

FEMALE:

fully

damaged,

winged;

without

length 3 mm;

head and fore

holotype (only

legs;

margins of pronotum reddish;

abdomen black;

testaceous;

punctate,

reaching

pronotum strong I y

tegulae;

scutum,

approximately 0,5 distal

notaulices

length of scutellum;

without keels; part of radial

fore-wing hyaline, vein

mid and hind

sh i ny,

incomplete,

propodcum fully

mandibles and clypeus yellow;

Bekily

TYPICAL MATERIAL:

part

description the head antennae yellow,

(10:7);

HOSTS:

Ethiopian region:

in P. MADAGASCAR:

spurs

with palpi,

with segments 7-10 black;

unknown

DISTRIBUTION:

bands;

tibial

is black,

(Madagascar)

holotype F!

punctate, reaching

reticulate rugose,

fore legs yellow. MALE: unknown LOCUS TYPICUS:

legs fully

without dark transversal

longer than proximal

1,1,2. According to the original

very with

with pronota I tuberc I es

scutellum and metanotum shiny,

without sculpture among punctures; dull,

known specimen)

thorax and propodeum black,

Bekily,

P!

270

Mem. Amer. En!.

Fig.

168 -

Ins!.,

no. 37,1984

Chela of Lonchodryinus pauliani (Benoit)(type)

271

Anteoninae

2. Lonchodryinus pauliani (Benoit) n. comb.

Prenanteon pauliani Benoit 1953b: 420.

FEMALE:

fully

testaceous,

winged;

length 5,50 mm;

with segments 8-10 brown;

head

yellow-testaceous;

propectus,

pronotum,

antennae

scutum,

scutellum and metanotum reddish-dark; tegulae testaceous; propodeum and abdomen black; mesopleura black, with reddish spot on anterior half; metapleura black;

fore

black,

ligth;

27:

with

12:

8:

tarsi 8:

8:

8:

8:

legs fully

reddish-testaceous;

11;

head shiny,

without sculpture among punctures; around

the orbits;

OPL = 13;

strongly

and hind

punctate,

pronotum shiny,

=

POL

smooth,

5;

finely

3;

punctate,

scu I pture among punctures,

with a strong transversa I an terior tegulae;

tubercles reaching

without sculpture among punctures; smooth,

keel

with central

smooth area,

metapleura shiny,

without sculpture;

pterostigma;

part of radial

distal

almost reaching long 7:

as broad

16:

29;

the basa I ce II s and

(19:4);

fore tarsal (Fig.

segment 5 of front

45 lamellae

(some of these very

hook;

segment 2 not produced

MALE: unknown LOCUS TYPICUS:

Mt.

TYPICAL MATERIAL: HOSTS:

with a (Fig.

long);

Tsaratanana

reticulate rugose;

fore-wing

with 2 dark

another beneath

longer than

proximal 168)

the

proximal

hook;

tibial

Ethiopian region:

group of nearly 7 tarsus produced

spurs 1,1,2.

(Madagascar) in P. MADAGASCAR:

OR lENT AL SPEC I ES OF

One species is known.

Mt.

4:

prominence bearing

unknown

DISTRIBUTION:

part,

with 2-3 rows of nearly

apex with a

segment 3 of front

into a

holotype F!

without

pterostigma more than four times as

tarsus

apex very prominent;

reaching

segments in following proportions:13:

168)

one bristle; lamellae;

vein much

the margin of wing;

enlarged claw

impress ion;

posterior surface

mesopleura

smooth and without sculpture; one on

14;

finely punctate,

reticulate rugose,

and posterior surface;

transversa I bands,

=

OOL

without

scutellum and metanotum shiny,

propodeum shiny,

between dorsal

smooth,

notaulices incomplete,

length of scutum;

without scu Ipture;

transversal

scutum shiny,

14: 7:

without keels

=

OL

pronotal

approximately 0,75

legs

mainly on frons,

frons with median keel,

occipital carina complete;

TL = 11;

mid

antenna I segments in following proportions:

Tsaratanana,

LONCHOD12Y INUS

P!

into a

272

Mem. Amer. En!.

Ins!.,

no. 37, 1984

Fig. 169 - Chela of Lonchodryinus sinensis n. sp.

(type)

Anteoninae

273

Lonchodryinus sinensis n. sp.

FEMALE:

fully

winged;

length 2,12 mm, except for the abdomen,

holotype (only known specimen); reg ion of frons testaceous;

black;

antennae brown;

femora and hind coxae partly brown; antennal

mandibles,

clypeus and

legs testaceous,

missing

with cl ubs of

antennae not distally thickened;

segments in following proportions:

10:

7:

13:

13:

10:

11: 9:

9:

12; head shiny, smooth, finely punctate, without sculpture among punctures; frons without median and lateral keels; POL = 4; OL = 2,5;

=

7;

OPL

=

4;

=

TL

strongly punctate, smooth,

occipital

carina complete;

with a strong

shorter than scutum shiny,

5;

(6:16);

finely punctate,

notaulices incomplete,

reaching

approximately 0,5 smooth,

with a strong

part

(10:9);

bands;

pterostigam

following proportions: proximal 169)

distal 3:

keel

lateral

areas;

part of radial

2,5:

6:

propodeum

between

dorsal

fore-wing vein

(25:6);

hyal ine,

longer than fore tarsal (Fig.

segment 5 of front

part shorter than proximal

tarsus with 2 rows of 44 lamellae;

length of scutum;

14; enlarged claw

prominence bearing one bristle;

with basal

scutum

surface

posterior surface with 2 longitudinal

longer than broad

11:

OOL

hairy,

posterior surface

without sculpture;

transversal

median area as rugose as

dark transversal

impression;

tubercles reaching tegulae;

and median area of posterior surface; keels;

pronotum shiny,

9:

without sculpture among punctures;

scutellum and metanotum shiny, reticulate rugose,

transversal

pronotal

in

anterior

part

(6:8);

without

proximal segments in

169) tarsus

with a (Fig.

segment 5 of front

apex with a group of 5 lamellae;

tibial

spurs 1,1,2. MALE: unknown LOCUS TYPICUS:

Tainan Hsien

TYPICAL MATERIAL: HOSTS:

(Taiwan)

holotype F!

in B.

unknown

DISTRIBUTION:

Oriental

region:

TAIWAN:

Tainan Hsien

NEARCT I C SPEC I ES OF

(Kwantzeling),

B!

LONCHODRY INUS

Three species are known.

KEY TO THE SPECIES

FEMALES

At -

least

the head

testaceous . . . • . .

1.

flavus n. sp.

Head most Iy black . . . . • . . . . . . • . . • • . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2

274

Mem.

2 Head dull, -

Amer.

granulated;

Ent.

Inst.,

antennae very

+ pronotum •••.•••.••••••••••• 2.

Head

shiny,

punctate,

without

antennae very

37,1984

short,

head

granulated;

no.

sl ight Iy

[{;asneri n.

longer

sculpture among punctures,

long,

much

longer

+ scutum . . . . . . . . . . . . . . . . . . . . . . . . . . 3.

than

sp.

than

not

+ pronotum

head

bakeri (Kieffer)

MALES

Posterior -

OPL at

ocell i

nearly

as

,5 times as

I east

long as

or

slightly

shorter

• . • • • ••

1.

flavus n.

sp.

long

as posterior

........

than

POL.

ocell i . . . . . . . . . . . . ..

bakeri (Kieffer)

3.

L. masneri is unknown.

The male of

1. Lonchodryinus flavus n. sp.

FEMALE:

fully

scutum brown

winged;

length 2,68-3,25 mm;

and pronotum dark;

abdomen black or darkened; not distally

thickened;

occasionally

antennal

fully

testaceous;

occasionally

in some specimens also propodeum and also metanotum dark;

segments

antennae

in following proportions:

8:

5:

7: 8: 8: 7: 6: 5: 5: 7; head shiny, smooth, finely punctate, without sculpture among punctures; frons without median and lateral keels; occipital

carina complete;

=

POL

5;

=

OL

3;

OOL = 7;

=

OPL

4;

=

TL

4;

pronotum very short, almost invisible, shiny, smooth; scutum, scutellum and metanotum shiny, smooth, finely punctate, without sculpture among punctures;

notaul ices incomplete,

scutum;

propodeum dull,

between

dorsal

and posterior surface,

poste,-ior surface; distal

fore-wing

part of radial

fore

tarsal

claw

(Fig.

5 of front

vein

hyaline,

more than

with 2

longitudinal

without dark three

lamellae;

winged;

170)

located on

a

as proximal

2,5:

4:

with 2 rows of nearly 25

tibial

bands;

long

proximal

(Fig.

2:

keel

keels on

transversal

times as

170)

with one bristle

length of

without transversal

6:

tarsus

fully

approximately 0,5

segments in following proportions:

a group of 11 MALE:

reaching

reticulate rugose,

11;

prominence; lamellae;

part;

enlarged segment

apex with

spurs 1,1,2.

length 3,00-3,75 mm;

black;

mandibles testaceous,

with teeth reddish; antennae brown, with segment 1 testaceous; legs testaceous; somet i mes anterior ha I f of frons testaceous; antennae not distally 13:

14:

thickened; 14:

punctures; 3,5;

OOL

=

14:

15:

antenna I segments 13:

15;

frontal

I ine absent;

5;

=

OPL

4;

TL

slightly shorter than POL;

in following proportions:

head shiny,

=

4;

occipital

punctate,

carina complete;

posterior ocell i

scutum shiny,

smooth,

10:

7:

14:

without sculpture among POL = 7;

nearly as finely

OL =

long as or

punctate,

without

An teon i nae

Fig.

170 -

Chela of

sculpture among punctures; 0,5

length of scutum;

sculpture;

275

Lonchodryinus flavus n. sp. (type)

notaulices

incomplete,

reaching

scutellum and metanotum shiny,

propodeum fully

reticulate rugose;

approximately

smooth,

without

posterior surface with

two

276

Mem.

longitudinal distal 171;

171

Ent.

fore-wing hyal ine,

part of radial

tibial

Fig.

keels;

Amer.

vein

Inst.,

no.

37,

1984

without dark transversal

nearly as long as proximal

part;

bands;

genital ia:

fig.

spurs 1,1,2.

Male genital ia of Lonchodryinus flavus n. (Canada): right half removed.

-

lOCUS TYPICUs:

Forsyth

TYPICAL MATERIAL: paratypes (2 FF, 3 MM)!

in Ol.

HOSTS:

unknown

DISTRIBUTION:

(Georgia,

holotype F!

4 MM)!

Nearctic region:

from Alberta

U.S. A.)

and 6 paratypes

in TW;

sp.

1 paratype F!

CANADA:

(1

F,

in AM;

5 MM)!

in OT;

6

6 paratypes (3 FF,

Rampart House (Yukon),

OT!

Jumping Pd. Cr. (Calgary, Alta.), Ol! OT! Bothwell (Ont.), Ol! OT! Hull (Que.), OT! Kirks Ferry (Que.), OT! OL! lac Mondor (Ste. Flore, Que.), OT!

U.S.A.:

(Md.),

TW!

Kerhonkson TW!

Ann Arbor

(Mich.),

High Point St. (Cherry town,

Bemus Pt.

(N.Y.),

Pk.

TW!

Ulster Co.,

Ol!

TW!

Forsyth

(N.J.),

TW!

N.Y.),

(Ga.),

Shokan AM!

Pinkham Notch

OT!

Takoma Pk.

(N.Y.),

TW!

Poughkeepsie (N.H.),

TW!

4 mi.

(N.Y.),

NNW Ol!

Anteoninae

277

2. Lonchodryinus masneri n. sp.

FEMALE:

fully

winged;

length 2,31-3,00 mm;

black;

mandibles testaceous;

occasionally also clypeus and malar space testaceous; with segments 1 and 7-10 brown; testaceous,

with coxae and clubs of femora partly brown;

hind coxae,

hind femora

distally

thickened;

7:

5:

7:

5:

frontal

4:

4:

half granulate; punctures; scutum; keels; dark 12:

occipital

pronotal smooth,

posterior surface with transversal part

19;

(7:8);

punctate,

172 -

(Fig.

reaching

Chela of

OL = 4;

scutum,

approximately 0,5

without

segments

segment 5 of front

transversal or

OOL = 8;

scutellum and length of longitudinal

fore-wing hyal ine,

without

vein sl ightly shorter than

in following proportions:

with one bristle

tarsus

(Fig.

lamellae;

13:

posterior

located on

172) tibial

with 2 rows of 40 spurs 1,1,2.

6:

2:

a proximal

Lonchodryinus masneri n. sp. (type)

apex with a group of 5

unknown

OPL = 4;

tegulae;

part of radial 172)

5:

without sculpture among

areolae smaller;

distal

fore tarsal

enlarged claw

prominence; lamellae;

bands;

9:

with anterior half rugose;

tubercles reaching finely

antennae slightly

with some punctures on frons;

carina complete;

pronotum dull,

legs

occasionally only

in following proportions:

granulate,

notaul ices incomplete,

Fig.

MALE:

head dull,

propodeum reticulate rugose,

proximal 3:

5;

TL = 6,5;

metanotum shiny,

and hind tibiae partly brown;

antenna I segments

line absent;

POL = 4,5;

antennae testaceous,

occasionally only segment 10 dark;

278

Mem.

LOCUS TYPICUS:

Cave Creek Cyn.

TYPICAL MATERIAL: FF!

in SC;

HOSTS:

holotype F!

1 paratype F!

En!.

Inst.,

(Arizona, in OT;

no.

37,1984

U.S.A.)

1 paratype F!

in TW;

2 paratypes

in OL.

unknown

DISTRIBUTION: OT!

Amer.

Nearctic region:

Chiricahua Mts.

Canyon

(Culberson

NOTES:

the species

(Portal,

Co.,

Tex.),

is named

U.S.A.:

Cave Creek Cyn.

Cochise Co.,

Ariz.),

OL!

(Portal,

SCI

Mc.

Ariz.),

Kittrick

TW!

in

honor of Mr.

Lubomir Masner.

3. Lonchodryinus bakeri (Kieffer) n. comb.

Anteon bakeri Kieffer 1906a: 239. f'renanteon bakeri (Kieffer): Kieffer 1914b: 212. f'renanteon micropunctatus Fenton 1927: 15 (n. syn.); locus typicus Nerepis

(N.

Prenanteon Prenanteon Prenanteon f'renanteon

FEMALE:

fully

black-brown;

B.,

U.

S.

A.).

bakeri (Kieffer): Muesebeck and Walkley 1951: 1041. micropunctatus Fenton: Muesebeck and Walkley 1951: 1042. bakeri (Kieffer): Krombein 1979: 1242. micropunctatus Fenton: Krombein 1979: 1242.

winged;

length 2,50-3,00 mm;

occasionally

antennae dark or testaceous, pleura,

scutum,

testaceous;

head and pronotum black or

pronotum partly reddish;

scutellum,

mandibles testaceous;

sometimes with segments 3-10 darkened;

metanotum and propodeum blaCk;

abdomen black or brown;

tegulae

legs brown-testaceous or testaceous,

with coxae basally black; occasionally only hind coxae basally black; antennal segments in following proportions: 11: 6: 15: 17: 14: 13: 10: 9: 9: 11; head shiny, smooth, finely punctate on vertex, temples, behind ocelli, on the sides of ocelli, on posterior half of frons; anterior half of frons more strongly or very keels; 7;

occipital

pronotum shiny,

scutum shiny,

strongly

finely

anterior margin; scutum;

finely punctate;

carina complete;

=

POL

punctate;

punctate,

more strongly

notaulices incomplete,

irregular keel

posterior surface with 2

between

longitudinal

=

3;

OOL

=

longitudinal 7;

OPL

=

tubercles reaching

7;

smooth,

0,50-0,65

length of

without SCUlpture;

propodeum reticulate rugose,

dorsal

TL

tegulae;

punctate only near the

reaching

scutellum and metanotum shiny,

transversal

frons without

OL

pronotal

occasionally metanotum weakly rugose; a

6;

without

and posterior surface;

keels sometimes weakly

visible;

median area nearly as rugose as lateral areas; fore-wing usually very long, hyaline, without dark transversal bands; distal part of radial vein longer than proximal proportions: hOOk;

12:

3:

enlarged claw

prominence;

part

3:

6:

(Fig.

(11:7); 15;

fore

tarsal

segments in following

segment 2 of front

173)

segment 5 of front

with one bristle tarsus

(Fig.

173)

tarsus produced located on a with basal

into a

proximal part shorter

Anteoninae

Fig.

173 -

Chela of Lonchodryinus bakeri (Kieffer) (N.

than distal

part,

of 4-7

lamellae;

MALE:

fully

279

with 2 rows of nearly 23-27 lamellae; tibial

winged;

from Good Hope

W. T., Canada)

apex with a group

spurs 1,1,2.

length

1,75-3,25 mm;

black;

mandibles testaceous;

280

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

antennae brown, with segments 1-2 or only 1 testaceous; legs testaceous, occasionally with hind coxae proximally black; antennae not distally thickened; 14:

13:

antennal

13:

12:

13;

seglcents

sculpture among punctures; deep median 3;

OOl

=

longitudinal

6;

=

OPl

posterior ocelli;

in following proportions:

head shiny,

3;

Tl

scutum,

smooth,

frons

furrow;

=

4;

without median and occipital

OPl at

transversal

keel

with 2 weakly hyal ine,

Fig.

visible

longitudinal

Male genital ia of Odessa

incomplete,

14:

keels, POL = 6;

long

keels;

fore-wing distal

14:

with a Ol

as

length of

smooth,

punctate,

reaching

propodeum reticulate rugose,

bands;

15:

without

lateral

1,5 times as

and posterior surface;

without dark transversal

174 -

least

notaulices

length of scutum;

between dorsal

6:

carina complete;

scutellum and metanotum shiny,

'without sculpture among punctures; approximately 0,5

10:

strongly punctate,

without a

posterior surface

usually very

part of radial

long,

vein

nearly

Lonchodryinus bakeri (Kieffer) from lake

(Michigan):

right

half removed.

Anteoninae

as long as proximal proximal

dorsal

part

(9:9);

281

gonoforceps

174) without membranous tibial

(Fig.

process and without distal

inner pointed process;

spurs 1,1,2. lOCUS TYPICUS:

Ormsby Co.

TYPICAL MATERIAL:

(Nev.,

holotype Fr

U.

S.

A.)

A. bakeri in CA; holotype Fr of Pro

of

micropunctatu5 in WA. HOSTS:

unknown

DISTRIBUTION:

Nearctic region:

Salmon (Naknek), Rich. Hwy.), aT! Big Delta, House

aT!

Hope

(N.

T.),

Pk.

Sandspit

aT!

(B.C.

W.

Stone Mt. aT!

Paxson,

(Yukon),

Chilkat Pass

(B.C.),

(Mi.

aT!

TW!

Unalakleet,

Tombstone C.G.

aT!

Atlin

(Hedley,

(Yukon),

Pro

Watson lake (Yukon),

aT!

(Dawson,

Good

(N.W.T.), (B.C.),

aT! TE!

Johnston Canyon

(Banff,

(Mi.

Yukon),

Kitimat

aT!

(B.C.),

Stewart

aT!

Alta.),

aT!

Gagnon Road

aT!

Battle River

(B.C.),

B.C.), (B.C.),

aT!

Terrace B.C.),

(Terrace,

(Alta.),

aT! aT!

B.C.),

aT!

(Terrace,

Spring Creek

TE!

Hixon

(Terrace,

(B.C.),

lakelse l .

aT!

aT!

(B.C.),

Ketchum l .

Cultus lake

(B.C.),

aT!

392 Alaska Hwy.,

B.C.),

aT!

B.C.),

Rupert

aT!

Rampart

TW!

(B.C.),

(Terrace,

aT!

TE!

lk.

Summit l .

B.C.),

King

aT!

Kitwanga

aT!

OTr

Gravel

496 Alaska Hwy.,

Shames

(B.C.),

aT!

Norman Wells

(B.C.),

(B.C.),

aT!

(Yukon),

Yukon),

OTr

151., Queen Charlotte Islands, B.C.), AM!

(Moresby

Nickle Plate City Robson

Dawson

CAl

Cowichan l .

Pete lake

CANADA:

-

Umiat,

Anchorage, OTr Shaw Crk. (M. 289 206 Richardson Hwy.), aT! Haynes,

TW!

(B.C.),

Liard Hot Spg.

ALASKA:

OTr Naknek, OTr Isabel Pass (Mi.

aT!

aT!

B.C.), Morley

(Alta.), aT! Frank (Alta.), aT! Mc Murray (Alta.), aT! Cameron l . Rd. (Waterton Nat. Park, Alta.), aT! White Fox (Sask.), aT! Churchill (Man.), aT!

Brandon

(Ont.), (Kam.

(Man.),

aT!

TW!

Co.,

Que.),

Whale River Chelsea, Co.,

Que.),

Cartwright

aT!

Mistassini

aT!

aT!

aT!

Warkworth Cr. (Ont.),

aT!

(Que.),

Que.),

aT!

Ottawa

Hull

(labrador),

(N.B.), aT!

(Que.),

(Que.),

Old Chelsea

Nerepis

(Churchill,

Merivale

(Que.), WA!

Hebron

aT!

aT!

(Oreg.),

TW!

Stanley

(Mt.

Shasta,

Ormsby Co. aT!

(Nev.),

Beaver Ck.

(Idaho),

TW!

CAl

aT!

aT!

Idaho City Cal if.),

(23 mi.

NNW West Yellowstone,

aT!

(labrador),

TW! U. S. A.: Westport (Oreg.), TW! Ochoco Creek

Utah),

TW! aT!

Gallatin Co.,

Chicago Cr.

(Mt.

Evans,

Colo.),

aT!

Dool ittle Ranch

aT!

(Mt.

Evans,

Colo.),

aT!

West Chicago Cr.

Echo l .

(Papineau

(N.B.),

Mc Bride

Crescent City

(Vernal,

Great

(Old

Goose Bay

(Idaho),

OTr

Stittsville

aT!

Buckingham Twp.

Kouchibouguac N.P.

Big Brush Cr.

aT!

lac Crescence

Chima (Que.),

(labrador),

Siskiyou Co.,

aT!

Summit of King Mt.

aT! Raleigh (labrador), TW! Doyles (labrador), (Wash.), TW! Mt. Ranier (Wash.), TW' Mt. Hood Springs

Man.),

(Ont.),

(Mt.

(Cal if.),

Vernal Mont.),

Evans,

(9800',

TW!

(Utah), AM!

Colo.),

Clear Cr.

Co., Colo.), aT! Estes Park (Colo.), aT! Phantom Vy (Colo.), TW! Gould (Colo.), TW! loveland Pass (Colo.), aT! Idaho Springs (Colo.), aT! Horosier Pass

(Summit Co.,

Colo.),

aT!

(Nebr.),

aT!

Polk Co.

(Mich.),

TW!

Washtenaw Co.

(Mich.),

Columbus

Crystal

Falls

(Mich.),

Ol!

Iron Co.

(Mich.),

BM!

Ol!

Ol!

aT!

(Black Hi 115,

Sylvan l .

(Wis.),

CAl

Iron River

(Mich.),

lake Odessa

Ol!

Oak.), TW!

Benzie Co.

(Mich.),

Yellow Dog Plains (Marquette Co.,

S.

(Mich.),

BM!

Mich.),

Ann Arbor

TW!

Rapides

282

Mem.

Parish

(La.),

Hi II

(Pr.

Pk.

(N.J.),

Placid Mt.

TW!

Pisgah

Georges Co., TW!

(N.Y.), OT!

NOTES: L. (Dalman).

Mt.

Md.),

Ent.

Inst.,

(N.C.), WA!

TW!

Troy

(N.Y.),

(6100'-6288',

EI

Saito

TW!

N.H.),

(Durango),

37,

1984

(N.C.),

(Md.),

TW!

(Huntington,

Dryden OT!

no.

Highlands

Cabin John

Kalbfleisch Field Station OT!

Washington

Durango,

Amer.

N.Y.),

(Maine),

Durham

TW!

TW!

(N.H.),

OT!

Oxon

High Point St. AM!

8M!

WA!

Lake

Summit of

MEXICO:

OT!

bakeri is very near the Palaearctic species L. ruficornis

LONCHOf)J(Y1NUS

NEOTRJP I C SPEC I ES OF

One species

is known.

Lonchodry in us tricolor Kieffer

Lonchodryinus Lonchodryinus Lonchodryinus Lonchodryinus

FEMALE:

fully

tricolor tricolor tricolor tricolor

winged;

Kieffer 1905a: 96. Kieffer:

Kieffer

Kieffer:

Olmi

and Currado 1979a:

Kieffer:

Olmi

1983:

length 6,0-7,5 mm;

1914b:

15.

head black,

anterior region of face and malar space testaceous; with

teeth reddish;

antennae fully

testaceous;

scutum usually reddish-testaceous; margins reddish,

with clypeus,

mandibles testaceous,

propectus,

pronotum and

occasionally pronotum black,

and scutum black;

metapleura black;

342.

81.

scutellum,

metanotum,

occasionally scutellum reddish;

with

propodeum and

mesopleura black,

distal region reddish, or fully reddish; petiole black; abdomen, and legs testaceous; antenna I segments in following proportions: 19:

18:

15:

numerous around

14:

13:

the orbits;

pronotum hairy, reaching

12:

longitudinal

14;

POL

shiny,

tegulae;

among punctures;

head dull,

keels

=

6;

granulated,

(obi ique on OL

smooth,

=

2,5;

finely

scutum shiny,

OOL

smooth,

notaulices distinct,

with frons hairy,

the sides of ocell i);

=

13;

punctate;

with

tegulae 20: 8: 25:

OPL

=

17;

pronotal

keel

TL

=

with is

12;

tubercles

finely punctate,

almost complete,

a

without sculpture

reaching

almost the

posterior margin of scutum; scutellum and metanotum shiny, smooth, without sculpture; mesopleura and metapleura dull, rugose; propodeum fully

reticulate rugose,

surface; with a vein

dull,

weak dark spot beneath

longer than proximal

forming

without keels between

posterior surface without

an

proportions:

angle; 12: 4:

6:

13:

22;

curvi I inear; fig.

176;

dorsal

keels;

the pterostigma,

part,

pterostigma:

longitudinal

with distal

distal

fore tarsal

segment 5 of front

and posterior

fore-wing hyal ine, part of radial

and proximal segments

tarsus

(Fig.

parts

in following 175)

with

Anteoninae

Fig.

175 -

Chela of

Lonchodryinus tricolor

283

Kieffer

(type)

284

Mem.

Amer.

Ent.

Ins!.,

no.

37,

1984

some rows of nearly 57 enlarged claw prominence;

Fig.

176 -

lamellae; apex with a group of 9 lamellae; 175) with one bristle located on a proximal spurs 1,1,2.

(Fig.

tibial

Pterostigma of

Lonchodryinus tricolor Kieffer (type): length of

pterostigma measured from A to B.

MALE:

unknown

lOCUS TYPICUS:

Goias

TYPICAL MATERIAL:

(Brazil)

holotype F!

HOSTS: unknown DISTRIBUTION: Neotropic region: TW!

Caruaru,

Ol!

TW!

in P. BRAZil:

Encruzilhada

Goias,

(Bahia),

AUSTRAL IAN SPEC I ES OF

P!

Ol!

Vilhena

(Rondonia),

TW!

LONCHODl2YINUS

Two species are known.

KEY TO THE SPECIES

FEMALES Unknown

MALES

Head black,

wi th mandibles,

clypeus and anterior

region of face 1. guineensis n. sp. Head fully black; only mandibles testaceous . . . . . . . . . . . . . . . . . . .. . . . . . . . . . . . . . . . . . . . . • . . • • . . • . . . . . . . . . . . 2. notogeicus n. sp. testaceous . . . . . . . . . . . . . . . . . . . . . . . . . . . .

-

1. Lonchodryinus guineensis n. sp.

Anteoninae

FEMALE: MALE:

285

unknown

fully

winged;

length 2,37 mm;

and anterior reg ion of frons testaceous;

head black,

testaceous;

thorax and propodeum black;

antennae not distally

thickened;

proportions:

9:

7:

5:

6:

9:

9:

sculpture among punctures,

abdomen brown;

antenna I segments

8:

8:

8:

with mandibles,

ma I ar space testaceous;

12;

I ittle rugose on

clypeus

antennae

legs testaceous;

in following

head shiny,

punctate,

without

anterior half of frons;

frons

without median and lateral keels; occipital carina complete; POL; 5,5; OL ; 2,5; OOL ; 5; OPL ; 5; TL ; 3; scutum, scutellum and metanotum shiny, smooth, finely punctate, without sculpture among punctures; notaul ices almost complete, almost reaching the posterior margin of scutum; propodeum dull,

reticulate rugose,

wing

hyaline,

longer than proximal genitalia missing 1,1,2. LOCUS TYP I CUS:

(14:10);

Ba i yer R.

the (only

or

bands;

longitudinal distal

keels;

fore-

part of radial

two parts forming

an angle;

known

tibial

specimen);

vein

spurs

(New Gu i nea)

holotype M!

in TW.

unknown

DISTRIBUTION:

Fig.

part

transversal

transversal

in the holotype

TYPICAL MATERIAL: HOSTS:

without

without dark

177 -

Australian region:

Male genitalia of

NEW GUINEA:

Baiyer R.,

Lonchodryinus notogeicus half removed.

n.

TW!

sp.

(type):

right

286

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

2. Lonchodryinus notogeicus n. sp.

FEMALE: MALE:

unknown

fully

winged;

length 2,37 mm;

black;

antennae testaceous;

legs testaceous,

with hind coxae proximally brown;

antennae not 8:

5:

7:

8:

distally 8:

8:

8:

thikened; 8:

7:

9;

sculpture among punctures; occipital

carina complete;

scutum shiny, notaul ices

strongly

incomplete,

antennal

between dorsal

frons without median POL = 6;

punctate,

and

OOL = 5;

smooth,

finely

OPL = 5;

length of scutum;

punctate,

without sculpture

without a

transversal

without dark

vein

much

dorsal

longer than

gonoforceps

process and without distal Kogh i

transversal

proximal

(Fig.

TL = 3;

punctures;

fore-wing hyal ine,

proximal

Mt.

without keels;

posterior surface without

the margin of wing;

TYPICAL MATERIAL:

lateral

without sculpture among

reaching approximately 0,75

reaching

HOSTS:

OL = 3;

punctate,

and

posterior surface;

part of radial

spurs 1,1,2. LOCUS TYP I CUS:

in following proportions:

strongly

propodeum reticulate rugose,

longitudinal 'keels; distal

segments

head shiny,

scutellum and metanotum shiny, among punctures;

mandibles testaceous;

177)

part

keel

bands;

(21 :8)

and

without membranous

inner pointed process;

tibial

(New Ca I edon i a)

holotype M!

in B.

unknown

DISTRIBUTION:

Australian

region:

fjEW CALEDONIA:

Mt.

Koghi,

B!

4. Anteon Jurine

Anteon Jurine 1807: 302. Antaeon Haliday 1833: 275, invalid emend. Chelogynus Hal iday 1838: 518 (syn. proposed by Kieffer in Kieffer et Marshall 1905); type species Chelogynus infectus (Haliday 1837), desig. by Muesebeck and Walkley 1951 (desig. of Ch. gaullei by Kieffer

1914b invalid because the

type was not an originally

included species)(discussion on synonymy:

see page 288).

Anteon Jurine partilli: Kieffer in Kieffer et Marshall 1905: 130. Neochelogynus R.C.L. Perkins 1905: 60 (syn. proposed by R.C.L. Perkins 1907); type species Neochelogynus typicus R.C.L. Perkins 1905, orig.

desig.

Prosanteon R.C.L. Perkins 1905: 66 (syn. proposed by R.C.L. Perkins 1907); type species Prosanteon chelogynoides R.C.L. Perkins 1905,

monotypic and orig.

desig.

Paranteon R.C.L. Perkins 1905: 67 (n. syn.); type species Paranteon rnyrrnecophilus R.C.L. Perkins 1905, orig. desig. Chelogynus Haliday: R.C.L. Perkins 1907: 48. Chelogynus Haliday: R.C.L. Perkins 1907: 48.

Anteoninae

287

Anteon Jurine: R.C.L. Perkins 1907: 50. Lasianteon Kieffer 1913b: 300 (syn. proposed by Muesebeck and Walkley 1951); type species Lasianteon rubrifrons (Kieffer 1905), desig.

by Muesebeck and Walkley

1951.

Xenanteon Kieffer 1913b: 300 (syn. proposed by Richards 1939); type species Xenanteon reticulatun, (Kieffer 1905), desig. by Richards 1939.

Liodryinus Kieffer 1913b: 300 (syn. proposed by Richards 1953); type species Liodryinus doddi (Kieffer 1911b), orig. desig. Anteon Jurine: Kieffer 1914b: 140. Chelogynus Haliday: Kieffer 1914b: 172. Xenanteon Kieffer: Kieffer 1914b: 193. Lasianteon Kieffer: Kieffer 1914b: 197. Allanteon Kieffer 1914b: 198 (syn. proposed by Berland 1928); type species Allanteon punctatus (Kieffer 1905), monotypic. Anteon Jurine: Berland 1928: 157. Chelogynus Haliday: Berland 1928: 162. Anteon Jurine: Richards 1937: 118. Chelogynus Haliday: Richards 1937: 118. Anteon Jurine: Richards 1939: 245. Anteon subg. Anteon Jurine: Richards 1939: 246. Anteon subg. Chelogynus Haliday: Richards 1939: 246. Hirtanteon Benoit 1951b: 163 (n. syn.); type species Hirtanteon canabensis Benoit 1951b, orig. desig. Anteon Jurine: J.F. Perkins 1976: 13. Chelogynus Haliday: J.F. Perkins 1976: 18. Chelogynus subg. Chelogynus Haliday: J.F. Perkins 1976: 18. Chelogynus subg. Neochelogynus R.C.L. Perkins 1976: 19. Anteon Jurine: Krombein 1979: 1240. Anteon subg.Anteon Jurine: Krombein 1979: 1240. Anteon subg. Chelogynus Haliday: Krombein 1979: 1241.

Anteon jurineanum Latreille 1809, first included species.

Type species: FEMALE palpi

(Figs.

178,

179):

with 6 segments;

distal

part of radial

fully

labial

winged; palpi

rarely brachypterous;

with 3 segments;

vein much shorter than

occasionally sl ightly shorter,

as

long as,

or

proximal

strong MALE labial

(Fig.

64);

occipital

transversal (Figs. palpi

29,

times as

carina complete;

between

180): fully proximal

dorsal

winged;

with 3 segments;

much shorter than as long as,

keel

part

(Fig.

or longer than proximal

long as broad

transversa I keel

between

(see fig.

178); part;

propodeum usually

with a

and posterior surface. with distal

with 6 segments; part of radial

vein

180); occasionally sl ightly shorter,

part;

176);

with

(Fig.

prominence bearing one

maxi Ilary palpi

fore-wing

part

maxillary

longer than proximal

inner side of the enlarged claw with a proximal bristle

fore-wing

pterostigma less than four

propodeum usually with

dorsa I and posterior surface;

a strong

gonoforceps without

288

Mem.

Fig.

apical

Amer.

178 -

branch wrapping

En!.

Female of

the penis;

Ins!.,

hairs at most as HOSTS:

1984

vertex of the head usually to occipital

long as breadth of antennal

carina;

without two

antenna I

segments.

worldwide

Cicadellidae (including Idiocerinae and Macropsinae)

SPECIES: NOTES:

37,

Anteon gaullei Kieffer

oblique keels connecting posterior ocelli DISTRIBUTION:

no.

156

whereas the females of

Chelogynus Hal iday and Anteon Jurine are

easily separable, the males show less distinct characteristics. The problem of the separation or synonymy of the two genera is resolved only after careful

study of the males.

main difference concerned

the

According

to J.F.

Perkins

length of the temples and

the scape ("head viewed dorsally

with

(1976)

the

the breadth of

temples shorter than breadth of

Anteon; "head viewed dorsally with temples at least as long as breadth of scape" in Chelogynus). I have seen that this difference is not

scape"

in

289

Anteoninae

Fig.

179 -

Female of

Anteon gaullei Kieffer

always visible: in some males of A. infectum (Hal iday) and A. scapulare (Haliday) for instance, the temples are shorter than the breadth of the scape.

Other differences concern

the petiolar area

and

the ninth sternite.

flavicorne, arcuatum), the petiolar area is usually broader and more shiny than in species of Chelogynus. The lucidity, however, seems unimportant since I have seen males of pubicorne and ephippiger with petiolar areas fully smooth and shiny, although these species are Chelogynus. The breadth In European species of

and

Anteon (for instance brachycerum,

the shape of the petiolar area usually

However, I have seen that males, for instance, of A.

are helpful

characteristics.

these characteristics also are variable. The inflatrix Benoit from the Ethiopian region, A.

canadense Ashmead from the Nearctic region, A. thai n. sp. from the Oriental

region have narrow petiolar areas that are rugose,

shape characteristics similar to

Chelogynus,

three species have segment 4 of the front the basitarsus,

proving they are true

dull,

and

whereas the females of the

tarsus at most 0,5 as

Anteon.

long as

In European species of

Anteon, the distal lobe of the ninth sternite is truncated, whereas in Chelogynus it is rounded. I have seen however, that the males of A. canadense (Anteon) have the distal lobe of the ninth sternite rounded. In conclusion I think that Anteon and Chelogynus are synonyms, even if in a single zoogeographic region the two groups can be separated easi Iy (for instance in Europe). The differences, however, are not val id for the world fauna.

290

Mem.

Fig.

180 -

Amer.

Ent.

Inst.,

no.

37,

1984

Male of Anteon tripartitum Kieffer

PALAEARCTIC SPECIES OF

ANTEON

Twenty species are known.

KEY TO THE SPECIES

FEMALES

(type)

Anteoninae

Segment

4

segment

5 of

front

tarsus at

front

most

tarsus

291

0,5 as

with basal

long part

as

segment

longer

1;

than

distal

(see fig. 183); front tarsal segments 3 or 4 produced into a hook • . • • • • • • • • . . • • • . . • • . • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • . • • • • • 2

part

Segment

4

of

front

tarsus

at

segment

5 of

front

tarsus

wi th

than

distal

produced

2

surface

tarsal

Posterior front

3

(see of

segment

surface

tarsal

Scutum shiny,

of

keel

on

into a

centrally

least

mid-length fully

of

keels;

longitudinal for

keels;

hook . . . . . . . . . . . . . 4

into a

except

arabicum n. or

some weak

tracks

of

sp.

reticulate

(see Ethiopian rugose;

never

species)

wi thout

1. jurineanum Latrei lie

2.

occasionally ocellus;

with

fore

a

short

tarsal

median

3

segment .. . . . . . .

brachycerum (Dalman)

carinate;

median keel

frons;

strongly

into a

Head not

fore

complete

tarsal

or

reaching

at

3 or 4 produced

segments

reticulate

reticulate

3.

hook

rugose;

fore

tarsal

segment

4

reticu12tum Kieffer

rugose;

fore

tarsal

3 produced

segment

into

.6

hook • • • • • • • • . • • . • • • . • • . • • . • . . • . . . • . . . • • . . . • . •

Notaulices

reaching

Antennae at

approximately

most

least

4.

fully

smooth

and

5. 6.

median median

rugose,

as

pronotum, and

thorax

scutum.

sp. of

scu tum.... .

...7

or brown . . .

flavicorne (Dalman)

area

of

area

posterior

scutum and 7.

fully

part Iy

of

rugose as

· • • • • • • . . . . . . • • • • .. Head

of

arcuatum Kieffer

SCUlpture,

Propodeum with Head,

length

yellow . . . . . . . . . . . . . . . . . . . ..

wi thout

nearly

nepalense n.

partly black

•••••••••••••••.•••• Propodeum wi th

0,65

reaching mid-length

•....••••.•.....•••• Antennae

9

2

sides . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ..

carinate;

· . . . . . . . . . . . . . . . . . ..

8

sculpture,

the anterior

produced

Notaulices at

7

1;

shorter

hook • • • • • • • • • • • • • • • • • . . • • . • • • • • • . . • • . . • • . • • . . • . • . . • • • . . 5

into a

6

segment

hook...........................

Frons

a

two

3 or 4 produced

20.

not

visible near

Head

or

longitudinal

the

......... ...... .....

5

segment as

hook . . . . . . . . . . . . . . . . . . . 3

granulated

centrally

produced

as

long

tarsal

into a

propodeum with

sculpture . . . . . . . . . . . Frons

front

long as

4 produced

without

................... Scutum dull,

191);

part

propodeum without

segments

irregular keel s

4

fig.

0,66 as

basal

hook . . • • . • • • . • • • . • • . . • • • • . . • • . • . . • • . . . • . . . • . . • 8

into a

Posterior front

part

least

posterior

lateral

surface

shiny,

rugose........ surface

most Iy

.. . . . . . . . 9

dull,

fully

areas • • . • . . . • • . • • • • • . . . • • 12

scutellum yellow or

reddish . . . . . . . .

ephippiger (Da I man)

black;

at

most

clypeus and

frons part Iy

testaceous . . • . . • • • . • • • • • • • • • • . . • • • . . • . • • • • • • . . • • . • • • • • • • • • • • • 10 10

Species very and

fully

segment

5

small,

smooth; of

front

• • • • . • • • • • • • • • • • • •

long

frons

less

tarsus 8.

than

smooth, with

2,5 mm;

without one

head

keels

row of

pubicorne (Da I man)

finely

punctate

or areolae;

lamellae

(Fig.

193) ..

292

Mem.

En!.

Species bigger,

long more

punctate;

with keels

frons

tarsus wi th 11

Amer.

Antennal

two

rows

segments

of

segments

least

(Figs

1,5

head and

13 Frons almost

1984

with

head

segment 195,

strongly

5 of

front

197) •••••••••

times as

long

as

• .11

1. .•.

as

long as

1 or

little

longer •••..

infectum (Ha I i day) joining

slightly visible;

the

ocelli;

posterior keel

lateral

always

thorax black ••••..•••••.•••••••••••••••.••••.

tripartitum K i ef f er

11.

triangle not

or differently

37,

scapulare (Ha I i day)

9. 10.

.. .. .. .. .. .. .. . . . . . Ocellar

3,0 mm,

triangle delimited by keels

keels occasionally visible;

no.

or areolae;

2+3 nearly

. . • • • • • • • • • • • • . •• .• 12 Ocellar

than

lamellae

2+3 at

• . • • • • • • • • • • • • • • • • .. Antennal

Ins!.,

delimited by

keels;

head and

thorax black

colored..............................

fully

reticulate

•••••••••• ••••• •••• Frons never almost

..13

rugose . . . . . . . . . . . . . . .

crassifrons (Moczar)

12. fully

reticulate

rugose;

at

most

anterior

half rugose • . . . . • • • • • • • . . . • • • • • • • . . . . . . . . • . . . • . . • • . • • • • • • • • • • 14 14 Head, thorax and propodeum fully black, occasionally with cupreous tints; head .. .. .. .. .. .. .. .. . . . . Head,

without sculpture among 8. pubicorne (Da I man)

thorax and propodeum partly

or

fully

the punctures . . . . .

yellow,testaceous,

reddish or reddish-dark; occasionally fully brown-dark, almost black or fully black, but in this case head granulated among punctures ••••••••••••••••••••••••••••••.••••..•..•••••••••••• 15 15 Head

fully

black;

scutum fully

reddish,

yellow or

two

of

rows

or mainly black;

testaceous;

lamellae

(Fig.

segment

5 of

pronotum

front

tarsus

with

202) . . . . . . . . . . . . . . . . . . . . . . . . . . .

13. gaullei Kieffer At

least

the head partly

occasionally

head and

yel low,

case pronotum brown-dark, testaceous of

lamellae

16 Head dul I, segment

or yel low; (Figs. always

1 nearly

shiny,

weakly 17 Head, frons

segment

clearly and twice as 14.

punctate,

granulated;

slightly

longer

black

or blackish,

5 of

front

long

testaceous; not

tarsus

but

in

that

reddish, wi th

1-2

rows

4

SCUlpture among punctures

segment

1 nearly as

long as

or very or

........................ .

Pronotum never black; head and body partly or fully reddish, testaceous 7.

antennal

as 4 . . . . . . . . . . . . . . . . . . . . . . . . . . . .

fulviventre (Ha I i day)

without

antennal

than

strongly granulated;

thorax and propodeum fully black, testaceous ... 15. hilare n. sp.

• • • • • • •• • . •

or

or black,

191,204,207) . . . . . . . . . . . . . . . . . . . . . . . . . . . . 16

• • • • • • • • • • • • • • • • • •• Head

reddish

scutum brown-dark

with anterior half

•• 17 of

variably colored, usually or yel low . . . . . . . . . . . . ..

ephippiger (Da I man)

The fema I es of A. ingenuum n. sp., corax n. sp., japonicum n. sp., munitum n. sp. are unknown. Probab I y A. ingenuum, corax or munitum are

Anteoninae

the males of

293

A.hilare and A. nepalense. Only breeding esperiments can

answer the question.

MALES

Posterior

surface

of

propodeum without

longitudinal

keels ••••••

1. jurineanum La t re i I Ie 2

Posterior surface of propodeum with two longitudinal keels ••.• 2 Gonoforceps with no distal inner process more or less pointed

(F i g.

181) • • • • • • • • • • • • • • • • • • • . . • • • • . • • • • • • • . . • • • . . • • • • • • • • • . . • 3

Gonoforceps wi th distal (Fig.

3

Posterior almost

inner process more

or

less pointed

192) •••••••.••••••••••••••••••••••••••.••..•••.•••.••••• 9 surface of

fully

Posterior

smooth,

surface of

propodeum wi th median

area

shiny

and

rugose . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4

not

propodeum with median area dul I

and

rugose

• ••••••••••••••••••••••••••••••••••••••••••••••••••••••••••••• 7

4

Head more

or

punctures;

less

finely

frons without

areolae . . . . . . . . . . . . . . . Head more not

or

less

reticulate

6

18. but

rugose;

with

sculpture among

punctures

resembling

sp. occasionally apparently

strong punctures

resembling

frons . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 Hind femora fully black or brown..... . . . . . . . . . . ......... . . . . . . . . . . . . . . . . . . . . . . . 10. infectum (Hal iday) Hind femora fully testaceous . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6 Head partly

or

region

fully

Head partly

or fully

of

reticulate

9.

granulated •••••••••.•••

16.

rugose,

partly

or

fully

scapulare (Hal iday)

reticulate

· • . . • • . . • . • . . . • • • • • • .•

7

without

strong

munitum n.

reticulate

rugose,

areolae on anterior

5

punctate and areolae or

rugose,

ingenuum n.

not

granulated

sp.

Scutum finely punctate, wi thout sculpture among punctures, wi thout areolae, not granulated •••••••• .•••••••••••••••••.•

• • • • • . • • • • • • • • • • • • . . .. Scutum differently strongly

reticulate

1 7.

corax n.

sculptured, rugose

sp. granulated,

with anterior half

or with posterior half

strongly

punctate ••••••••••••••••••••••••••••••••••••••••••••.••••••••• 8

8

Scutum granulated;

antennae almost

fully

brown;

legs with

tibiae partly brown . . . . . . . . . . . . . . . . . . . . • . . . • • • • . . • . • . . • • . . . • •. 1. jurineanum La t re i I I e

coxae,

femora and

Scutum not granulated, but strongly anterior half and strongly punctate anrJ

legs

fully

testaceous,

. • . • • • • . . . • • • • • • •• 9

Gonoforceps wi thout head not

brown

or black . . . . . . . . .

for hind

1 9. japonicum n.

dorsal

184);

except

reticulate rugose on on posterior half; antennae

punctate,

proximal

granulated; 2.

coxae basally brown.

sp.

membranous process antennae and

brachycerum (Dalman)

(Fig.

legs mainly

294

Mem.

En!.

Gonoforceps wi th a

more or

membranous process

(Fig.

10 Distal

inner process

apical

11

Amer.

margin

of

excavated

Ins!.,

less

extended medially

(Figs.

190) •••••••••••••••••.••• 11

188,

inner process

of

gonoforceps

or

straight

testaceous,

at

Legs

testaceous,

most

wi th

proximal

gonoforceps

margin

tibiae partly brown . . . .

1984

large dorsal

Distal Legs

37,

192) ••.••••••..••••••..••••••••••••• 10

apical

convex

no.

hind 6.

extended apically

(Figs.

192,

legs with

hind

Coxae brown;

coxae,

femora

clubs

occasionally

of

femora

stalks

of

brown

coxae and

femora brown

13 Head not dorsal (Fig. Head

less brown

punctate,

femora brown

strongly granulated

206) . . . . . . . . . . . . strongly or

or

finely

14.

finely of

punctate,

strongly

or weakly

granulated,

15 Head

strongly

weakly

smooth,

granUlated,

205);

wi thout

without

long

proximal (Fig.

short

dorsal

203) . . . . . . . 14

sculpture among

without wi th

sculpture among punctures

short

or

long median keel

on

gaullei Ki effer occasionally with areolae and

tripartitum Kieffer

Head punctate,

sculpture among punctures 8.

proximal

very

sculpture among

granulated;

11.

granulated . . . . . . . . . . . .

hind

med ian kee I on frons. ephippiger (Da I man)

usually 13.

(Fig.

irregular keels . . . . . . . without

mid and

fulviventre (Hal iday)

punctate,

frons . . . . . . . . . . . . . . . . .

. . . . . . • • • • • 13 least

gonoforceps usually

gonoforceps very

punctures, usua I I y wi thou t • • • • • • • • . • • • • • • • • • . . • •. 7. Head more

of

punctate,

occasionally

membranous process

with at

or blackish . . . . . . . . . . . . . . . . . . . . . . . . . . . 15

membranous process

punctures 14 Head

or blackish,

or dark .

extremeties

hind

. • • • • • • . • • . . . • • •• . • . . . . . • . • • . • . • . • • • . • . . . . • • • or

and

flavicorne (Dalman)

coxae and

Legs more

and with

194) . . . . . . . . . . . . 12

. . . . . . . . . . . . . . . . . . . . . . . 5. arcuatum Kieffer 12 Legs fully yellow or testaceous, with proximal of

and with

or weakly

pubicorne (Dalman)

The males of A. reticulatum Kieffer, hilare n. sp., arabicum n. sp., nepalense n. sp. are unknown. Probably A. ingenuum n. sp., corax n. sp. and munitum n. sp. are the ma I es of A. hilare and A. nepalense. Only breeding experiments can answer the question.

1. Anteon jurineanum Latrei lie

Anteon jurineanum Latrei lie 1809: 35 Gonatopus brevicornis Dalman 1818: 85 (syn. proposed by Olmi 1983); locus typicus V astergot I and (Sweden). Dryinus cursor Haliday, in Curtis 1828: 206 (syn. proposed by Olmi 1983); locus typicus England. Dryinus otiartes Walker 1837: 425 (n. syn.); locus typicus England.

Anteoninae

295

Dryinus sisithrus Wa I ker 1837: 426 (n. syn.); locus typicus Eng I and. Dryinus nanus Haliday, in Walker 1837: 427 (n. syn.); locus typicus Wicklow

(Ireland).

Anteon crenulatus Kieffer in Kieffer et Marshall 1905: 141 {n. syn.)i locus typicus Bonar Bridge (Scotland). Anteon thomsoni Kieffer in Kieffer et Marshall 1905: 142 (syn. proposed by Richards 1939); locus typicus Sweden (new name for Dryinus brevicornis Thomson 1860 nec Gonatopus brevicornis Dalman 1818) •

Anteon vicinus Kieffer in Kieffer et Marshall 1905: 145 (n. syn.); locus typicus Dumfries (Scotland). Anteon marginatus Kieffer in Kieffer et Marshall 1905: 147 (syn. proposed by Olmi 1983); locus typicus Maisons-Laffitte (France). Anteon rectus Kieffer in Kieffer et Marshall 1905: 171 (n. syn.); locus typicus Dumfri es (Scot I and) • Anteon scoticus Kieffer in Kieffer et Marshall 1905: 174 (n. syn.); locus typicus Sutherland (Scotland). Anteon jurineanus Latreille: Kieffer 1907b: 26. Anteon nanus (Haliday): Kieffer 1907b: 27. Anteon otiartes (Walker): Kieffer 1907b: 27. Anteon sisithrus (Walker): Kieffer 1907b: 27. Antaeon barbatus Chitty 1908: 142 (n. syn.); locus typicus Bentley Wood

(England).

Xenanteon brunneipes Berl and 1928: 168 (n. syn.); locus typicus Maisons-Laffitte (France).

Dryinus sisithrus Walker: Richards 1935: 147. Anteon crenulatum Kieffer: Richards 1935: 147. Anteon scoticum I
296

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

FEMALE: fully winged; length 2,00-3,12 mm; black; mandibles testaceous, with teeth brown; antennae fully brown or with segments 1-2 or 1-5 testaceous;

legs brown,

testaceous,

with hind coxae partly brown,

with fore tibiae and

and tibiae partly brown or blackish; distally

thickened;

5:

4:

5:

5:

5:

5:

antennal 8;

lateral

tegulae testaceous;

granulate,

OL,

these

OOL,

lateral

OPL,

face with a frontal keels are incomplete;

variable;

posterior surface short,

temples visible;

shorter than scutum

reaching tegulae;

scutum dull,

occasionally only

granulate;

propodeum reticulate rugose,

181

-

longitudinal distal

and Tentsmuir Nat.

keels;

5:

carina complete;

pronotum dull,

metanotum shiny,

and posterior surface;

Res.

fore-wing hyaline,

part of radial

4:

sockets;

pronotal

rugose,

with

tubercles

reaching mid-length of smooth or rugose,

with a strong

transversal

posterior surface without

Male genitalia of Anteon jurineanum Latreille from

(England) (A)

12:

granulate and reticulate rugose;

without sculpture; between dorsal

clubs of femora antennae short,

with or without

occipital

notaul ices incomplete,

scutellum and

Fig.

line,

(4:18);

scutum or shorter;

keel

occasionally

with more or less numerous

keels near orbits directed towards the antennal

occasionally POL,

ligth;

segments in following proportions:

head dull,

areolae or irregular keels;

tarsi

or with coxae,

(Scotland) (8):

without dark transversal

vein much shorter than proximal

New Forest

right half removed.

part

bands;

(3:8);

fore

tarsal segments in proportions: 9: 2: 2: 2: 8; segment 4 of front tarsus produced into a hook; enlarged claw (Fig. 64) with one bristle located on a proximal

prominence;

segment 5 of front

tarsus

(Fig.

64)

with 1 row of

Anteoninae

1-4 lamellae; lamellae; MALE:

in addition

tibial

fully

winged;

Fig.

light;

182 -

apex with a

length 1,56-3,00 mm; legs more or

tegulae testaceous;

black;

dull,

in following proportions:

face with frontal directed

(laterally

viewed)

complete;

line,

6:

with or without

POL = 8;

OL = 4;

sockets;

7:

and fore

of

antennal

Anteon jurineanum

(Sweden).

8:

8:

weakly or strongly

lateral

8:

8:

8:

7:

12;

head

granulated;

keels near the orbits

occasionally

OOL = 7;

scutum dull or shiny,

sculptured;

12:

tarsi

thickened;

with more or less numerous areolae or irregular keels;

towards the antennal

incomplete;

with

antennae not distally

Shapes of gonoforceps

granulate,

mandibles testaceous;

less brown,

Latrei lie from Vastergotland

segments

group of 1-3

spurs 1,1,2.

antennae brown or black; tibiae

numerous bristles;

297

smooth,

OPL = 3;

these

TL = 4;

without sculpture;

lateral

keels are

occipital

carina

occasionally

anterior surface of scutum more strongly

notaul ices incomplete,

reaching mid-length of scutum or

298

Mem.

shorter;

Amer.

Inst.,

scutellum and metanotum shiny,

propodeum reticulate rugose, and posterior surface;

bands;

areas;

(Fig.

181)

without membranous proximal apex very rounded

long

(Fig.

smooth,

longitudinal

1984

without sculpture;

transversal

and narrow

keels;

fore-wing

distal of radial

gonoforceps

37,

keel

posterior surface usually without

area as rugose as lateral transversal

no.

with a strong

with 2 distinct

occasionally

(3:9);

Ent.

process;

keels;

this case with median

hyaline,

without dark

with

part

inner pointed process and

gonoforceps,

182);

182); occasionally

dorsal

vein much shorter than proximal

without distal

(Fig.

in

between

longitudinal

in

lateral

occasionally broad,

view,

with

pointed or

intermediate characteristics;

tibial

spurs 1,1,2. lOCUS TYPICUS:

Paris (France)

lectotype M! (designated by Olmi 1983) of A. jurineanum in SP; lectotype F! and 2 paralectotypes MM! (designated by Olmi 1983) of G. brevicornis in ST; holotype F of D. cursor in ME; holotype M! of D. otiartes in BM; holotype F! in of D. sisithrus in BM; lectotype M! of A. nanus in DU; holotype F! in of A. crenulatus in BM; holotype F! of A. vicinus in BM; lectotype M! (designated by Olmi 1983) of A. marginatus in P; holotype F! of A. rectus in BM; holotype M! of A. scoticus in BM; holotype M! of A. barbatus in BM; holotype F! of X. brunneipes in P; type of A. thomsoni not seen (probably in lU; not found TYPICAL MATERIAL:

in ST). HOSTS: in U. S. S. R. name A. brevicorne). DISTRIBUTION:

Oncopsis flavicollis l . (Ponomarenko 1978: under

Palaearctic region:

JAPAN:

Sapporo (male determined as

Anteon japonicum by Ashmead: undescribed species), WA! MONGOLIA: 3 km N form Somon Uenc aimak),

BU!

Ol!

lothar

OT!

Ignalina

(Chovd

NEPAL:

aimak),

Pulchauki

(Birganj),

(lietuva),

EB!

BU!

SO from Somon Bajanzogt

(Katmandu),

OT!

27° 58'

Ol!

OT!

N 85° 00'

other localities

Godavari

E,

Ol!

(Central

(Katmandu),

OT!

U.

S.

in Ponomarenko (1978);

S.

Ekenas, HE! lOjo, HE! SWEDEN: Vastergotland, ST! Hoor distr. (SK.), Ring Sjo, (SK.), BM! Degaberga (SK.), BM! Dagstorp sjo. (SK.), BM! Rostanga Duka BU!

(SK.),

BM!

(Csoroghegy), Rakos.,

BU!

Skaralid BU!

Nagykovacsi,

Duczfalva,

BU!

Nagyvisnyo

(Fenyofo),

BU!

ROMANIA:

Chodau Gora

(Bohemia),

(Karawanken,

AUSTRIA:

BM!

(SK.),

BM!

Mogyoroskert BU!

Rev.,

(BUkk),

Tasnad

Sornmeringgebeit

BU!

HU!

Engadine Nat.

Park

BU!

Kolasin

Tata,

BM!

BU! BU!

BU!

BM!

OT!

BM!

HUNGARY:

Bukk h. Omassa,

Asvany,

BU!

Kisszepalma

CZECHOSLOVAKIA:

Halle,

(Reichenau distr.),

(Grigioni),

BU!

(Crna Gora),

GERMANY:

Prater (Wien), P! Wien, ZO! DENMARK: CO! Ostiylland, CO! HOllAND: Utrecht, lO!

Narvik,

Budapest,

(Maramures),

YUGOSLAVIA:

Siovenija),

NORWAY:

(Bakony hgs.),

R.:

FINLAND:

HU!

Huben

BM!

Kranjiska

Mecklenburg, (0 Tirol),

HU!

HU!

Korsor, CO! Blldso, CO! Danmark, Ol! SWITZERLAND: Boussens (Vaud), FRANCE:

lille (Nord),

P!

Fontainebleau, P! Maisons-laffitte (Seine-et-Oise), P! Paris, SP! Morlaix (Fl.), BM! ENGLAND: Hengistbury Head (SH), Ol! Epping Forest (Essex), CA!

Coombe Wood

(Surrey),

BM!

(Surrey),

Southampton,

P!

Chobham Cornm.

BM!

New Forest,

(Surrey),

BM!

Ol!

Esher Cornmon

Farnham Cn.

(Bucks.),

Anteoninae

BM!

Bentley Wood

(Surrey),

BM!

Risborough Aldbury (SH),

(Suffolk East),

Rammamex Heath BM!

Whaddon

(Herts.),

BM!

Romsey

BM!

Gt.

Brickhi II

(BX.),

BM!

Lustleigh

(SO.,

Oartmoor,

BM!

Slough

(EK.),

BM!

(Romsey,

I<ew Gans.

BM!

(SH),

BM!

BM!

Ivinghoe

BM!

BM!

BM!

Boxmoor

Chase,

BM!

(HT.),

SH.),

Horsley

BM!

P. BM!

Lyndhurst

Beaconsfield (Exmoor), BM!

BM! Wye

Kent's Oak

Wendover

BM!

(HT),

(Surrey),

BM!

BM!

(New Forest,

(New Forest,

BM!

Porlock Oistr.

(Nairn),

BM!

Bricket Wood

(Bucks.),

BM!

Holland's Wood

Lyndhurst-Brockenhurst Rd.

BM!

Newton,

Wimbledon Common

Darnaway Forest

(London),

(Berks.),

BM!

Cannock

Oartmoor),

(HU.),

BX.),

Chase (BX.),

(Bucks.),

Woodwalton

SH.),

Silwood Park

(Brickhill,

(8X.),

(Bucks.),

BM!

299

(BX.),

SH.),

BM!

BM!

SCOTLAND:

Grantown

(Moray), BM! Tentsmuir Nat. Res. (Fife), BM! Inchnadamph N.R. (N.S.), Bf,~! Sutherland, BM! Aviemore (Inverness), BM! Loch Marech (Ross), BM! Kinlochewe

(R.W.),

BM!

Bonar Bridge,

(Inverness),

BM!

Lochinver.

N.R.

BM!

Gairloch

(RVV.),

Glen.,

8M!

Kincraig

Dowry

(Wicklow),

Glen

(Co. I'll),

DO),

BM!

S.

BM!

BM!

SPAIN:

(RW.),

(EI.),

Wicklow,

Manzarron

Limonetto (Cuneo), NOTES:

according

species without propodeum)

~~r.

Gretna,

~~ethy

DU!

(Co.

(Murcia), SZ!

BM!

Killin

TW!

BM!

Loch Morlich

Dumfries,

Bridge

Glenasmole

WI),

B/.l!

BM!

BM!

(EI.),

(Perthshire),

BM!

(Co.

Beinn Eighe

BM! DU),

Leixoes,

(Bussoleno),

Cavruber

IRELANO:

Tipperary Wa.

PORTUGAL:

Pietrabianca

BM!

Devilis

(~~ewcastle,

OLl

Torino),

ITALY: SZ!

GE! to Richards

longitudinal

(1939)

keels on

was composed of only

cursor Haliday) however,

BM!

BM!

Kippure Ho.

Benedetto Belbo (Cuneo),

BM!

(Sutherland),

the brevicorne group

(the complex of

the posterior surface of the

two species,

and marginatum Kieffer.

brevicorne (Dalman) (=

According

to J.F.

Perkins (1976),

the brevicorne group was composed of three spec i es,

cursor was considered a separate species.

Differences in

because

these species,

however,

were hardly recognizable with the keys of Richards and J.F.

Perk ins.

Nei ther author actua II y

saw

the type of marginatum,

and both

based their keys on unrel iable original descriptions. I found and studied the type of marginatum in the National Paris Museum and also studied numerous other specimens of the brevicorne group kept collections.

I

in European

have always found specimens exhibiting characteristics

intermediate to those I isted

in the keys for separating

the three species.

Variations are present in the proportions of the sides of the ocellar triangle, the frons, exist

I

breadth of the ocell i, frontal

think

keels,

sculpture of the scutum,

OPL,

sculpture of

colour and gonoforceps. Since intermediate forms

that the supposed brevicorne group

only one species (brevicorne Dalman)

which

is really composed of

is very variable.

As proposed

(1983), however, brevicorne (Dalman) is a junior synonym of jurineanum Latrei lie (nec jurineanum Latrei lie in Richards 1939 and J.F. by Olmi

Perkins 1976). This synonymy

is based on

the discovery of the type of

Anteon jurineanum Latrei lie in the Spinola collection. The classic jurineanum of Richards and J.F. Perkins has become Anteon arcuatum Kieffer. Another interesting variation

in the morphology of the brevicorne

300

Mem. Amer.

Ent.

Ins!.,

no.

37,

1984

group concerns the posterior surface of the propodeum. This is the main area used in identifying brevicorne - absence of longitudinal keels in this group, presence in the other species. I, however, have seen some males of the brevicorne group with tracks of two longitudinal keels, and modified my keys accordingly.

2. Anteon brachycerum (Dalman)

Dryinus brachycerus Dalman 1823: 12. Dryinus lyde Walker 1837: 427 (syn. proposed by Richards 1939); locus typicus Eng I and. Anteon nigricornis Kieffer in Kieffer et Marshall 1905: 172 (syn. proposed by Richards 1939); locus typicus Eccles (Scotland). Anteon triareolatus Kieffer in Kieffer et Marshall 1905: 175 (syn. proposed by Richards 1939); locus typicus Bonar Bridge (Scotland). Anteon brevicollis Kieffer in Kieffer et Marshall 1905: 176 (syn. proposed by Richards 1939); locus typicus Bonar Bri dge (Scot I and) • Anteon flavitarsis Kieffer in Kieffer et Marshall 1905: 176 (syn. proposed by Richards 1939); locus typicus Clober (Scotland). Anteon indivisus Kieffer in Kieffer et Marshall 1905: 199 (syn. proposed by Richards 1939); locus typicus Bishopton (Scotland). Anteon nigroclavatus Kieffer in Kieffer et Marshall 1905: 205 (syn. proposed by Richards 1939); locus typicus Kilsyth (Scotland). Anteon curvatus Kieffer in Kieffer et Marshall 1906: 517 (syn. proposed by Richards 1939); locus typicus Sutherland (Scotland). Anteon obscuricornis Kieffer in Kieffer et Marshall 1906: 523 (syn. proposed by Richards 1939); locus typicus Eccles (Scotland). Anteon brachycerus (Dalman): Kieffer 1907b: 25. Anteon nigroclavatus Kieffer: Cameron 1907: 33. Antaeon suffolciensis Chitty 1908: 144 (syn. proposed by Richards 1939); locus typicus Felden (England). Anteon curvus Kieffer 1914b: 155 (syn. proposed by Richards 1939); locus typicus Sutherland (Scotland). Anteon neglectus Kieffer var. suffolciensis (Chitty): Kieffer 1914b: 162.

Anteon Anteon Anteon Anteon Anteon Anteon

lyde (Walker): Richards 1935: 151. indivisum Kieffer: Richards 1935: 151. obscuricorne Kieffer: Richards 1935: 151. brachycerum (Dalman): Richards 1939: 250. brachycerum (Dalman): J.F. Perkins 1976: 16. brachycerum (Dalman): Ponomarenko 1978: 18.

FEMALE: fully winged; length 2,3-3,3 mm; black, with mandibles yellow (except for teeth brown) and legs brown (with tarsi and junctions light);

Anteoninae

Fig.

301

8

Chela of Anteon brachycerum

183 -

A:

tegulae testaceous;

apical

part;

antennae short,

(Dalman)

B:

basal

from Dowry

(Ireland);

part.

with segment 1 nearly twice as long

as 4; head dull, smooth, granulated; frontal line absent; scutum shiny, punctate, hairy, smooth; notaulices distinct, incomplete, reaching midlength of scutum; sculpture;

scutellum and metanotum smooth,

propodeum reticulate rugose,

between dorsal longitudinal

and posterior surface;

keels;

a

shiny,

transversal

without irregular keel

posterior surface with 2

median area very shiny

almost smooth or weakly rugose; radial

with

(more than

fore-wing hyal ine,

vein much shorter than proximal

part;

lateral

areas),

with distal

segment 1 of front

part of tarsus

much longer than segment 4; fore tarsal segment 3 produced into a hook; segment 5 of front tarsus (Fig. 183) with nearly 10-13 bristles; apex with a group of 4

lamellae; enlarged claw

(Fig.

183)

on a proximal prominence; tibial spurs 1,1,2. MALE: fully winged; length 2,3-3,3 mm; black, mandibles yellow and junctions

(except for teeth brown);

light;

tegulae testaceous;

with one bristle located with antennae brown

and

legs fully brown or with tarsi

head smooth,

dull,

granulated;

302

Fig.

Mem.

184 -

Amer.

Ent.

Ins!.,

no.

37,

Male genital ia of Anteon brachycerum

1984

(Dalman)

from Burnham

Beeches (Eng I and) •

frontal

line absent;

scutum shiny,

hairy,

punctate,

smooth;

notaulices

distinct, incomplete, reaching mid-length of scutum; scutellum and metanotum shiny, without sculpture; propodeum reticulate rugose, with transversa I i rregu I ar keel surface with 2

longitudinal

lateral

smooth;

areas),

keels;

fore-wing

much shorter than proximal inner pointed process,

between dorsa I and posterior surface; median area very shiny hyaline,

part;

with distal

gonoforceps

without proximal

dorsal

(Fig.

(more than

part of radial

184)

vein

with a distal

membranous process;

spurs 1,1,2. LOCUS TYPICUS: Smaland (Sweden) TYPICAL MATERIAL: lectotype M! and 1 para lectotype F!

a

posterior

tibial

(designated by

MO':Lar 1967) of D.

brachycerus in ST; holotype M! of A. suffolciensis in BM; holotype F! of A. nigricornis (= A. obscuricornis) in BM; lectotype M! of A. lyde in BM; holotype F! of A. triareolatus in BM; holotype F! of A. brevicollis in BM; holotype F! of A. flavitarsis in BM; holotype M! of A. indivisus in BM; lectotype M! of A. nigroc1avatus in BM; holotype M! of A. curvatus (= A. curvus) in BM. HOSTS: in England Oncopsis spp. (J.F. Perkins 1976); in U. S. S. R. Oncopsis flavicollis L. (Ponomarenko 1978). DISTRIBUTION:

Palaearctic region:

U.S.S.R.

(Ponomarenko 1978);

Smal and,

ST!

V I!

Umea.

NEPAL:

FINLAND:

meadow

L.

Godavari

(Katmandu),

Rago (Estl.),

(Vasterb.),

LE!

HE!

Stock hoi m,

OT!

SWEDEN: TW!

Ri ngsjon,

303

Anteoninae

Fig.

185 -

Chela of

Anteon reticula tum Kieffer from

M.

Peglia

(Italy).

304

TW!

Mem.

NORWAY:

ROMANIA

Laugesund,

HU!

En!.

Ins!.,

HUNGARY:

37,

no.

1984

Nagyszenas

(Budai

H.),

1972); GERMANY: Mecklenburg, HU! Annatal

(Nagy

Rheinland),

Amer.

OTt

DENMARK:

Biskelund,

CO!

Bildso,

CO!

BU!

(Siebengebirge,

FRANCE

(Richards

1939); ENGLAND: Felden (Herts), BM! Burnham Beeches (Bucks.), BM! Wimbledon Common Bishopton BM!

(Sussex),

(Renfrew),

BM!

BM!

VI!

Epping Forest

Kilsyth

Bonar Bridge (Sutherland South),

Sutherl and,

BM!

IRELAND:

Portmanock

(Dublin),

BM!

Dowry

BM!

BM!

CAl

SCOTLAND:

Clober

(Stirling),

Eccles (Dumfries),

(Wick low),

Woodbrook,

(Essex),

(Stirling), BM!

Athdown

BM!

(Wick low),

BM!

BM!

3. Anteon reticulatum Kieffer

Anteon reticulatus Kieffer in Kieffer et Marshall 1905: 140. Xenanteon reticulatus (Kieffer): Berland 1928: 168. Anteon reticulatum Kieffer: Richards 1939: 256.

FEMALE:

fully

testaceous

winged;

length 4,25-4,56 mm;

black,

(except for segments 6-10 brown);

coxae proximally black, femora brown;

with antennae

legs testaceous,

hind coxae almost fully black,

tegulae testaceous;

twice as long as segment 4;

antennae short,

head shiny,

with fore

clubs of hind

with segment

1 almost

fully strongly reticulate rugose;

frontal line present; pronotum rugose; scutum shiny, smooth, punctate, without sculpture, among punctures; notaulices distinct, incomplete, reaching approximately 0,3 length of scutum; punctate or without punctures; sculpture;

propodeum reticulate rugose,

between dorsal longitudinal

and posterior surface;

keels;

scutellum shiny,

metanotum shiny,

smooth,

smooth,

without

with a transversal

irregular keel

posterior surface with 2

median area of posterior surface strongly rugose,

with

numerous transversal keels; fore-wing hyaline, with distal part of radial vein much shorter than proximal; segment 1 of front tarsus nearly three times as long as segment 4;

segment 4 produced into a hook; 185) with 4 bristles; apex with a group of 6 lamellae; enlarged claw (Fig. 185) with one bristle located on a proximal prominence; tibial spurs 1,1,2. segment 5 of front

MALE:

tarsus

fore tarsal

(Fig.

unknown

LOCUS TYPICUS:

Maisons-Laffitte (France)

TYPICAL MATERIAL:

holotype F!

in P.

HOSTS: unknown DISTRIBUTION: Palaearctic region: Oise), NOTES:

P!

ITALY:

only

Monte Pegl ia

FRANCE:

(Perugia),

two known specimens

Maisons-Laffitte (Seine-etOLl

!

4. Anteon nepalense n. sp.

Anteoninae

FEMALE:

fully

testaceous, thickened; 6:

6:

5:

frontal

length 3,0 mm;

antennal

antennae brown;

punctate,

pronotum shiny,

shiny,

smooth and

scutum

(6:13);

POL

~

5;

two OL

~

without sculpture;

the orbits;

~

3,5;

OOL

fore-wing hyaline,

radial

vein much shorter than proximal

in following proportions:

tegulae;

MALE:

8:

3:

2:

unknown Godavari

TYPICAL MATERIAL: HOSTS:

4:

~

TL

scutum shiny,

part

with 2

area as rugose as bands;

(3: 11);

fore

enlarged claw

prominence;

finely

scutellum and metanotum

median

13;

with 1 row of 15 lamellae;

LOCUS TYPICUS:

OPL

notau I ices i ncomp lete,

without dark transversal

one bristle located on a proximal spurs 1,1,2.

4,5;

propodeum reticulate rugose,

areas;

186)

6:

posterior surface shorter than

keels on posterior surface;

(Fig.

5:

keels around

tubercles reaching

without sculpture;

tibial

6:

lateral

without scu I pture among punctures;

longitudinal

9:

with anterior surface rugose and posterior surface

pronotal

smooth,

6:

4;

reaching approximately 0,65 length of scutum; shiny,

10:

without sculpture among punctures;

frons also with

carina complete;

legs

antennae distally

segments in following proportions:

head shiny,

I ine present;

puncta te,

black;

with coxae and clubs of femora darkened;

6;

occipital 3;

winged;

305

lateral

distal tarsal

(Fig.

part of segments

186)

segment 5 of front

apex with a groc'p of 7

with

tarsus

lamellae;

(Nepal)

holotype F!

in OT;

1 paratype F!

in OL.

unknown

DISTRIBUTION:

Palaearctic region:

NEPAL:

Godavari

(Katmandu),

OT!

Ol!

5. Anteon arcuatum Kieffer

Anteon arcuatum Kieffer in Kieffer et Marshall 1905: 144. Anteon imberbis Kieffer in Kieffer et Marshall 1905: 149 (n. syn.); locus typicus Rannoch (Scotland). Anteon arcuatus Kieffer: Kieffer 1907b: 25. Anteon flavicorne (Dalman) var. bensoni Richards 1939: 255 (n. syn.); locus typicus Brickell Wood (England). Anteon jurineanum Latreille partim (only F): Richards 1939: 256 (discussion on synonymy:

see page 299).

Anteon jurineanum latreille: J.F. Perkins 1976: 16. Anteon arcuatum Kieffer: Olmi 1983: 80. Anteon arcuatum Kieffer: Moczar 1983b: 198.

FEMALE:

fully

winged;

length 2,18-3,00 mm;

black,

with antennae brown

(sometimes partly brown and testaceous); legs testaceous, with coxae partly brown and clubs of hind femora brown; tegulae testaceous; mandibles yellow, nearly

with teeth brown;

twice as long as segment 4;

antennae short, head shiny,

with segment

smooth,

1

without sculpture

306

Mem.

Fig.

186 -

or weakly alutaceous,

Amer.

Chela of

with

En!.

Ins!.,

no.

37,

1984

Anteon nepalense n. sp. (type)

irregular keels at

least near the occipital

triangle; frons with frontal I ine and 2 lateral longitudinal keels near orbits; pronotum rugose; scutum smoooth, shiny, punctate on anterior half,

almost without punctures on posterior half;

incomplete,

approximately

metanotum shiny,

smooth,

notaulices distinct,

reaching mid-length of scutum; without sculpture;

scutellum and

propodeum reticulate rugose,

with a transversal irregular keel between dorsal and posterior surface; posterior surface with 2 lateral longitudinal keels; median area dull, rugose, nearly as rugose as lateral areas; fore-wing hyal ine, with distal part of radial vein much shorter than proximal part; segment 1 of front tarsus much hook;

longer than segment 4;

segment 5 of front

apex with a group of 4

tarsus

fore tarsal

(Fig.

lamellae;

187)

tibial

segment 3 produced

into a

with 1 row of 8-18 bristles;

spurs 1,1,2.

Anteoninae

Fig.

187 -

MALE:

fully

with dorsal

Chela of

winged;

Anteon arcuatum Kieffer from Portmanock {Ireland}

length 2,31-2,50 mm;

part of the segmen ts brown;

brown;

mandibles yellOW,

brown;

femora brown;

testaceous;

307

with

teeth brown;

sometimes only

head dull or shiny,

black;

antennae testaceous,

somet i mes on I y

segments 1-3

legs testaceous,

hind femora brown;

granulated,

with coxae

tegulae

with numerous areolae or

irregular keels; frontal line present; frons without lateral longitudinal keels; scutum shiny, smooth, without sculpture; hairy, punctate on anterior half, distinct,

almost without punctures on

incomplete,

metanotum shiny, with a

transversal

posterior half;

reaching mid-length of scutum;

smooth,

without sculpture;

irregular keel

posterior surface with 2

propodeum reticulate rugose,

between dorsal

longitudinal

keels;

notaulices

scutellum and

and posterior surface;

median

area shiny,

less

308

Fig.

Mem. Amer.

188 -

Ent.

Inst., no.

37,

1984

Male genitalia of Anteon al'cuatum Kieffer from Esher Common (England)

rugose than lateral areas; fore-wing hyal ine, with distal part of radial vein much shorter than proximal part; gonoforceps (Fig. 188) with distal inner process pointed and extended medially; its apical margin excavated; dorsal proximal membranous process present; tibial spurs 1,1,2. LOCUS TYPICUS: Eccles (Scotland) TYPICAL MATERIAL: holotype F! of A.

al'cuatum in BM; holotype F! of A. flavicol'ne var. bensoni in BM; holotype F! of A. imbel'bis in BM. HOSTS: in I ta I y Rhytidodus decimusqual'tus (Schrk.). DISTRIBUTION: Palaearctic region:

MONGOLIA:

15 Km S from Dalanzadgad

(Gurban Sajchan ul., SLid gobi aimak), BU! U.S.S.R. (Ponomarenko 1978: under name jUl'ineanum); FINLAND (Hellen 1953); HUNGARY: Batorliget, BU! Tiszang, BU! Asvany, BU! Osborokas (Bugac), BU! GERMANY: Ferchland (Elbe), HU! Eisteraue, HA! AUSTRIA: Eisenzicken, VI! DENMARK: Legind, CO! Gejlbjerz, CO! HOLLAND: Den Haag, LE! SWITZERLAND: Brig (Valais), HE! ENGLAND: Esher Common (Surrey), BM! Brickett Wood (Herts), BM! Ham Common, BM! Bix (Henley, Oxfordshire), BM! Siapton (Linslade, BX.), BM! Starks Pond (Kingsteignton, SO.), OL! SCOTLAND: Eccles, BM! IRELAND: Portmanock

(Dublin),

BM!

ITALY: Alfiano Natta (Alessandria),

Anteoninae

309

EN!

6. Anteon flavicorne (Dalman)

Gonatopus flavicornis Dalman 1818: 83. Anteon sericeus Kieffer in Kieffer et Marshall 1905: 198 (n. syn.); locus typicus Maisons-Laffitte (France). Anteon flavicornis (Dalman): Berland 1928; 159. Anteon sericeus Kieffer: Berland 1928: 161. Anteon flavicorne (Dalman): Richards 1939: 253. Anteon levigatus Kieffer: Haupt 1941: 40. Anteon flavicornis (Dalman): Haupt 1941: 42. Anteon subflavicornis Haupt 1941: 43 (n. syn.); locus typicus Be II i nchen 312) •

Anteon Anteon Anteon Anteon Anteon

FEMALE:

(Germany)( discussion on synonymy:

flavicorne (Dalman) : subfl a vicorne Haupt: flavicorne (Dalman) : subflavicorne Haupt: flavicorne (Dalman) :

fully

winged;

see notes on page

J. F. Perkins 1976: 17. J. F. Perkins 1976: 18. Ponomaren ko 1978: Olmi

18.

and Currado 1979a:

Currado 1983:

length 2,18-4,00 mm;

345.

127.

black;

antennae yellow,

sometimes with fuscous dorsal markings; legs yellow, with hind coxae partly black and stalk of hind femora brown; tegulae testaceous; mandibles yellow,

with teeth brown;

nearly

twice as long as segment 4;

weakly

granulated,

frontal

line present;

with few

antennae short, head shiny,

with segment

punctate,

irregular keels near occipital

1

strongly or carina;

frons with 2 lateral complete or incomplete keels

near orbits directed towards the antenna I sockets; strongly punctate on anterior third,

without

scutum shiny,

smooth,

punctures on posterior

surface; notaulices distinct and incomplete, reaching approximately 0,30 length of scutum; scutellum and metanotum shiny, smooth, without sculpture;

Fig.

propodeum reticulate rugose,

189a -

Propodeum of Anteon

with a

transversal

keel

between

flavicorne (Dalman) (A) and subflavicorne

Haupt

(B).

310

Fig.

Mem. Amer.

189b -

Chela of

Ent.

Ins!.,

no. 37,

1984

Anteon flavicorne (Dalman) from Courmayeur (Italy)

Anteon inae

dorsal

and posterior surface;

311

posterior surface with 2 long i tudi na I keels;

median area smooth, rugose near the margins, projecting or not forward beyond anterior margin of lateral areas (Fig. 189a: this difference, used in

identifying A.

flavicorne and subflavicorne, is only individual

in Anteon;

variation

it

is present

also in other species,

for

instance A.

ephippiger); fore-wing hyaline, with distal part of radial vein much shorter than segment 4; tarsus

(Fig.

lamellae; proximal MALE:

proximal

part;

fore tarsal 189b)

prominence;

punctate;

apex with a group of 8

with one bristle located on

length 2,00-2,62 mm;

black;

legs testaceous,

tegulae testaceous;

granulated,

scutum shiny,

incomplete,

189b)

smooth,

finely

posterior surface with 2 longitudinal fore-wing

much shorter than proximal

mandibles yellow,

with teeth

keels;

median area shiny,

with distal

gonoforceps

(Fig.

its apical

dorsal proximal membranous process present; LOCUS TYP I CUS: Vastergot I and (Sweden) lectotype F!

(hereby

scutellum and

propodeum reticulate rugose, dorsal and posterior surface;

hyal ine,

part;

notaulices distinct,

length of scutum;

process pointed and extended medially;

TYPICAL MATERIAL: (4 MM, 1 F)! of G.

antennae testaceous,

with coxae and clubs of

punctate;

reaching approximately 0,3

partly rugose;

a

fully or partly reticulate rugose and

metanoturn shiny, smooth, finely punctate; with a transversal irregular keel between smooth,

longer than

segment 5 of front

spurs 1,1,2.

markings;

femora partly brown; head dull,

(Fig.

tibial

winged;

with fuscous dorsal brown;

into a hook;

with 1 row of bristles;

enlarged claw

fully

segment 1 of front tarsus much

segment 3 produced

almost

part of radial

190)

with distal

vein inner

margin excavated;

tibial

designated)

spurs 1,1,2. and 5 paralectotypes

flavicornis in ST; lectotype M! (hereby designated) of A. sericeus in P; holotype F! and 2 para types FF! of A. subflavicornis in HA. in England Idiocerus sp. (J.F. Perkins 1976), Populicerus laminatus (Flor) (~ Idiocerus laminatus) and P. populi (L.) (~ Idiocerus populi) (Chambers 1955); in Germany Populicerus confusus (Flor) (~ I diocerus confusus) (Haupt 1941); in I ta I y Rhytidodus decimusquartus (Schrk.), Populicerus albicans (Kbm.), Tremulicerus distinguendus (Kbm.) (~ Idiocerus distinguendus) , Idiocerus stigmaticalis Lewis (Currado 1983); in U. S. S. R. Populicorus laminatus (Flor) (~ Idiocerus laminatus) (Ponomarenko 1978). PARASITES: in England and Switzerland Ismarus flavicornis Thomson HOSTS:

(Nixon

1957;

Chambers 1955).

DISTRIBUTION:

Palaearctic region:

U.

S.

S.

R.

(Ponomarenko 1978);

FINLAND (Hellen 1953); SWEDEN: Vastergotland, ST! Sweden, VI! HUNGARY: Ujpest, BUI Szolnok, BU! SzSztMiklos, BU! CZECHOSLOVAKIA: Chodau (Bohemia),

BM!

GERMANY:

(Kurland),

HU!

Holzhausen

(Halle), AUSTRIA:

HA!

Thuringen,

Voslau,

Kobenhavn,

CO!

P!

Niederlehme (Kyswusterh), (Leipzig),

GE!

EB!

Mecklenburg,

Prater (Wien),

SWITZERLAND:

P!

Chancy

Halle, EB!

Graz,

HA!

HU!

Jelowka-Lepel

Dolauer Heide

Bellinchen P!

(Geneve),

(Oder) ,

DENMARK: GV!

Ecogia

Korsor,

HA! CO!

(Geneve),

GV!

312

Mem.

Amer.

\"\

Ent.

Inst.,

no.

37,

1984

: .

"\

Fig.

190 -

Male genitalia of Anteon (England):

Mies M.

(Vaud),

(Vaud),

Crespian

La!

La!

(Gard),

Moremont

FRANCE: LE!

flavicorne (Dalman) from Oxford

right half removed.

(Vaud),

La!

La Ferte-Milon

La Sauge (Vaud), (Aisne),

P!

Maisons-Laffitte (Seine-et-Oise),

in Kieffer et Marshall

1905);

ENGLAND:

Kings Wood

OL!

Romand s.

Lille (Nord), P!

(BD.),

Cretai I BM!

P! (Kieffer

Wood

Walton Fen (Cambs.), BM! Northwood, OX! Oxford, aT! BM! Monks Wood NNR (Cambs.), BM! SCOTLAND: Sutherland, BM! IRELAND (Richards 1939); ITALY:

S.

Benedetto Belbo

S. Giorio (Torino), in Olmi

EN!

(Cuneo),

Ol!

and Currado 1979a);

Pl!

Courmayeur ALGERIA,

Alfiano Natta (Aosta)

(= A.

(Alessandria),

EN!

subflavicorne Haupt

LN!

the flavicorne group was flavicorne (Dalman) and subflavicorne Haupt. About jurineanum see notes on page 299 (jurineanum sensu Richards 1939 and J. F. Perk i ns 1976 is rea II y Anteon arcuatum Kieffer). About subflavicorne, accord i ng to J. F. Perk i ns (1976), the mai n characteristic used in identifying flavicorne and subflavicorne was the NOTES:

according to J.F.

Perkins

(1976)

composed of three species: jurineanum,

shape of the petiolar area ("petiolar area projecting forward beyond anterior margin of lateral areas" in flavicorne; "petiolar area not projecting"

in subflavicorne). According to me the different shape of the

petiolar area is only an in other species;

variation in Anteon; it's present also ephippiger (Dalman) can show petiolar

individual

for instance A.

Anteoninae

313

area projecting or not forward beyond anterior margin of lateral An other interesting characteristic used in

concerns antenna I segment 3 ("segment 3 about three times as broad"

in flavicorne;

areas.

identifying the two species

"segment 3 about 2,5 times as

long as

long as broad"

in

subflavicorne). The proportions of the antennal segments, however, are very variable in Anteon: in bred series I have seen antennal segments variously long and broad (see also notes on page 322 about Anteon pubicorne). Other characteristics proposed by J. F. Perkins (1976) (sculpture of petiolar area and frons;

Anteon.

I

think that subflavicorne

colour)

are very variable

in

is a junior synonym of flavicorne.

7. Anteon ephippiger (Da I man)

Gonatopus ephippiger Dalman 1818: Gonatopus collaris Dalman 1818: 82 1939); locus typicus Vastergot I and Dryinus facialis Thomson 1860: 177

81. (syn.

proposed by Richards

(Sweden). (n. syn.);

locus typicus Got I and

(Sweden) •

Anteon albidicollis Kieffer in Ki effer et Marsha II 1905: 157 (n. syn. ); locus typicus Volosca (I ta I y) • Anteon rubrifrons Kieffer in Kieffer et Marshall 1905: 158 (n. syn.); locus typicus K is-Pose (Hungary). Anteon collan's (Dalman): Kieffer 1907b: 25. Anteon facialis (Thomson): Kieffer 1907b: 26. Chelogynus rufovariegatus Berl and 1928: 164 (syn. proposed by Richards 1939); locus typicus France. Anteon fulviventre (Haliday) partim (only M): Richards 1939: 265. Anteon ephippiger (Dalman) partim (only F): Richards 1939: 267. Anteon albidocolle Kieffer: Richards 1939: 270. Chelogynus ephippiger (Dalman) partim Haupt 1941: 47. Anteon flaviscapus Jansson 1950 partim (only M): 221 (n. syn.); locus typicus unknown. Anteon rubrifrons Kieffer: Moczar 1967: 297. Anteon albidicolle Kieffer: Olmi and Currado 1976b: 73. Chelogynus ephippiger (Dalman): J.F. Perkins 1976: 22, 23. Chelogynus ephippiger (Dalman): Ponomarenko 1978: 21. Chelogynus albidocollis (Kieffer): Ponomarenko 1978: 21. nec Chelogynus ephippiger (Dalman): Moczar 1979: 77. Anteon pyonganensis Moczar 1983a: 184 (n. syn.); locus typicus Pyongyan

(Korea).

Anteon ephippiger (Da I man): Moczar 1983b: 198.

FEMALE: fully winged; length 1,81-2,87 mm; colour variable; head yellow or reddish (light or dark); occasionally black or brown; antennae yellow

Mem.

314

Fig.

191

-

Chela of

Amer.

En!.

Ins!.,

Anteon ephippiger

no.

37,

(Dalman)

1984

from Benicasim

(Spain)

or testaceous; occasionally with segments 1-2 testaceous and 3-10 brownish; pronotum whitish or yellow; occasionally reddish or reddishdark with brown spots;

propectus yellow or reddish;

occasionally

blackish; scutum yellow or reddish; occasionally brown or black; scutell um and metanotum as colored as scutum; mesop leura and metap leura

Anteoninae

bl ack or redd i sh; reddish; slightly

abdomen black or brown;

tegulae testaceous;

longer than segment 4

punctures, shiny,

propodeum black;

legs yellow; hairless,

315

(8:6);

antennae long,

head shiny,

smooth,

without sculpture;

scutum,

smooth,

shiny,

without sculpture;

notaulices distinct,

0,3-0,5

length of scutum;

irregular keel smooth;

keels;

proximal

part;

and posterior surface;

median area rugose,

fore-wing hyaline,

with distal

segment 1 of front

fore tarsa I segment 2 produced 191)

with

prominence; MALE:

fully

testaceous,

(Fig.

tibial

winged;

keels;

Fig.

black;

head shiny,

without frontal

smooth,

antennae brown;

(F i g.

line;

notaulices distinct,

smooth,

without

frons without incomplete,

scutellum and metanotum shiny, with a

lateral

right half removed.

without

reaching

smooth,

transversal

Male genitalia of Anteon ephippiger (Dalman) (Spain):

legs

tegulae testaceous;

strongly punctate on anterior half,

propodeum reticulate rugose,

192 -

tarsus

apex with a group of 3 lamellae;

length 1,75-2,50 mm;

punctures on posterior half; sculpture;

shorter than segment 4;

segment 5 of front

with one bristle located on a proximal

with teeth brown;

mid-length of scutum;

shiny and

vein shorter than

1,1,2.

finely punctate,

scutum shiny,

reaching

transversal

posterior surface

at most with hind cOXae proximally black;

mandibles yellow, sculpture,

191)

spurs

with a

occasionally

tarsus slightly

1 row of 16-19 lamellae;

enlarged claw

incomplete,

part of radial

into hook;

pronotum

scutellum and metanotum

propodeum reticulate rugose,

between dorsal

with 2 longitudinal

1

without

without sculpture or weakly alutaceous;

smooth,

occasiona II y

with segment

vithout

irregular keel

from Nerja

316

Mem.

between

Amer.

Ent.

Inst.,

dorsa I and posterior surface;

no.

37,

1984

posterior surface with 2

longitudinal keels; median area rugose; fore-wing hyaline, with distal part of radial vein much shorter than proximal part; gonoforceps (Fig. 192)

with distal

inner pointed process and with proximal

membranous process; LOCUS TYPICUS:

tibial

dorsal

spurs 1,1,2.

Vastergotland

(Sweden)

of G. ephippiger in ST; holotype F! of G. collaris in ST; holotype F! of A. albidicollis in VI; holotype F! of A. rubrifrons in BU; holotype F! and 2 paratypes FF (one specimen from Prechaq is an A. fulviventre according to Richards 1939) of Ch. rufovariegatus in P; holotype F! of D. facialis in ST; holotype of A. flaviscapus not found. HOSTS: in Germany Macropsis sp. (Haupt 1941); in U. S. S. R. Psammotettix striatus L. (Ponomarenko 1978, under name Ch. albidocollis). TYPICAL MATERIAL:

DISTRIBUTION: Prov.),

BU!

holotype F!

Palaearctic region:

Ryesan

Somon Uenc

(Chovd aimak),

BU!

23 Km NW from Somon Chutag

BU!

Mongol ia

Nora BU!

Ekenas,

(Stockholm), Karansebes,

BM! BU!

HE!

U.S.S.R.

SWEDEN:

HUNGARY: Holies,

(South Pyongan

MONGOLIA:

(Namnan ul

(Ponomarenko 1975a),

HE!

Pyongyan BU!

SO from Somon Bajanzogt

BU!

Tvarminne,

KOREA:

(Ryang-gang Prov.),

BUkk h,

(Central

BU! BU!

ST!

Tabdi, N.

aimak),

Bulgan aimak),

(Ponomarenko 1978);

Vastergotland,

Budapest,

BU!

Gebirge,

3 Km N from

BU!

Enyed,

FINLAND:

Gotland, Mt.

BU!

ST!

Meszes,

Kis-Pose,

BU! Tatatovaros, BU! Bartia, BU! Orsz Miklos, BU! R. Szombat, BU! CZECHOSLOVAKIA: Carlsbad, P! ROMANIA (Nagy 1973); BULGARIA: KUstendil,

BU!

Dalmacija: VI!

GERMANY:

Neue MUhle, Langeland (Vaud),

YUGOSLAVIA:

I.,

OL!

Bellinchen

HU!

(Oder) ,

AUSTRIA:

CO!

St.

Kostanjika

(Istra),

BM!

of Anteon zernyanus Fahringer,

type F!

Wien,

Kobenhavn,

Giez (Vaud),

HA! P!

CO! GV!

Aachen, Prater

HU!

Mainz,

(Wien),

SWITZERLAND: FRANCE:

Gruz (= Gravosa; undescribed species), P!

OT!

Auvernier,

Le Puy,

P!

Halle,

HU!

DENMARK: LO!

Huelgoat,

La Praz BM!

Palavas (Herault), GV! ENGLAND: Spratton (Northants), BM! Wimbledon Comm. (London), OT! Esher Common (Surrey), BM! IRELAND (R i chards 1939);

SPAIN:

Giorio

(Torino),

Benicasim EN!

(Castellon),

Volosca

BM!

(Trieste),

Nerja

VI!

(Granada),

TURKEY:

BM!

Smyrna,

ITALY:

S.

WA!

8. Anteon pubicorne (Dalman)

Gonatopus pubicornis Dalman 1818: 87. Dryinus tenuicornis Dalman 1823: 13 (n. syn.); locus typicus Vastergotland

(Sweden).

Gonatopus cephalotes Ljungh 1824: 267 (n. syn.); locus typicus Smaland

(Sweden).

Dryinus lucidus Haliday, in Curtis 1828: 206 (n. syn.); locus typicus England. Dryinus penidas Walker 1837: 423 (n. syn.); locus typicus Britain.

Anteoninae

317

Dryinus alorus Walker 1837: 425 (n. syn.); locus typicus Britain. Chelogynus alorus (Walker): Haliday 1838: 518. Antaeon alorus (Walker): Marshall 1873: 7. Anteon azorus (Walker): Cameron 1907: 33. Anteon fuscoclavatus Kieffer in Kieffer et Marshall 1905: 196 (n. syn.); locus typicus Cadder (Scot I and) • Anteon triangularis Kieffer in Kieffer et Marshall 1905: 197 (n. syn.); locus typicus Bonar Bridge (Scotland). Anteon divisus Kieffer inK ieffer et Marsha II 1905: 200 (n. syn.); locus typicus Mugdock (Scotland). Anteon vulgaris Kieffer in Kieffer et Marsha II 1905: 202 (n. syn.); locus typicus Dumfries (Scotland). Antaeon breviventralis Ch i tty 1908: 144 (n. syn.); locus typicus Walmer (England).

Antaeon delicatulus Chitty 1908: 215 (n. syn.); locus typicus Ranworth Broad

(Eng I and).

Anteon subarcticus Hellen 1935: 7 (nomen nudum). Anteon alorus (Walker): Richards 1935: 151. Anteon fuscoclavatum Kieffer: Richards 1935: 151. Anteon penidas (Walker): Richards 1935: 151. Anteon vulgare Kieffer: Richards 1935: 151. Chelogynus serratus Maneval 1935: 6 (n. syn.); locus typicus Mont Mezenc

(France).

Anteon lucidus (Haliday) partim (only F): Richards 1939: 270. Chelogynus exiguus Haupt 1941: 52 (n. syn.); locus typicus Bellinchen

(Germany)(discussion on synonymy:

see notes on page

322).

Chelogynus ephippiger (Dalman) partim (only M): Haupt 1941: 47. Anteon flaviscapus Jansson 1950 partim (only F): 221 (n. syn.); locus typicus unknown (discussion on synonymy: see notes on page 322) •

Anteon subarcticus Hellen 1953: 96 (n. syn.); locus typicus Ivalo (Finland) •

Anteon pubicornis (Dalman); Moczar 1967: 299. Anteon cephalotes (Ljungh): Olmi and Currado 1975: 51. Chelogynus lucidus (Haliday): J.F. Perkins 1976: 22. Anteon cameroni Kieffer: Olmi and Currado 1979a: 347. Anteon lucidum (Haliday): Olmi and Currado 1979a: 347. Anteon pubicorne (Dalman): Moczar 1983a: 184. Anteon pubicorne (Dalman): Moczar 1983b: 198. Anteon mongolicum Moczar 1983b: 198 (n. syn.); locus typicus Bogdo ul.

(Mongolia).

FEMALE: fully winged; length 2,00-2,81 mm; black; antennae brown, with segments 1-2 or only 1 yellow; sometimes antennae testaceous, with segments 3-6 brown; legs fully testaceous; sometimes with clubs of hind

318

Mem. Amer.

Fig.

193 -

Chela of

Ent.

Inst.,

no.

37,

1984

Anteon pubicorne (Dalman) from Genova (Italy)

femora and clubs of femora brown; tegulae testaceous; mandibles yellow, with teeth brown; antennae long, with segment 1 slightly longer than segment 4; proportions of the antennal segments very variable (see notes on page 322); head shiny, smooth, finely or strongly punctate, without sculpture among punctures;

frons usually with a short frontal

line;

Anteoninae

sometimes

(in small

specimens)

pronotum strongly punctate,

without frontal

shiny;

reaching mid-length of scutum; keel

longitudinal shiny,

proximal;

with distal

fore tarsal

tarsus

(Fig.

5-6 lamellae;

193)

long as segment 4; (Fig.

193)

sometimes

segment 1 of front

segment 5 of front apex with a

with a proximal

bearing one bristle;

tibial

length 1,37-2,87 mm;

winged;

irregular

vein much shorter than

into a hook;

with one row of 20- 23 lamellae;

enlarged claw

smooth,

posterior surface with 2

part of radial

MALE:

fully

with a transversal

occasionally rugose near the margins;

segment 2 produced

tarsus approximately 0,66 as

punctate,

incomplete,

median area usually rugose and dull,

smooth and without keels;

fore-wing hyaline,

keels present;

finely

scutellum and metanotum shiny,

and posterior surface;

keels;

lateral

smooth,

notaul ices distinct,

propodeum reticulate rugose,

between dorsal

line;

scutum shiny,

without sculpture among punctures; finely punctate;

319

group of

prominence

spurs 1,1,2. black;

antennae brown or

black; sometimes with antennal segment 1 yellow; legs brown or black,with tarsi and junctions ligth; tegulae testaceous; mandibles yellow,

with teeth brown;

variable sometimes

Fig.

proportions of the antennal

(see notes on page 322); (in small

194 -

specimens)

Male genitalia of Ditton

head shiny,

smooth,

finely

segments very

with sculpture variable; punctate,

without sculpture

Anteon pubicorne (Dalman) from Thames

(England):

right half removed.

320

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

among punctures; sometimes (in large specimens) rugose or smooth, strongly or finely punctate, with areolae or weak keels, without sculpture or keels among punctures; notaulices distinct,

and metanotum shiny, rugose,

with a

surface; shiny,

scutum hairy,

incomplete, smooth,

transversal

shiny,

finely punctate;

irregular keel

fore-wing hyaline,

shorter than proximal

part;

with distal

gonoforceps

pointed process and with

proximal

spurs 1,1,2. LOCUS TYPICUS:

(Sweden).

Smaland

TYPICAL MATERIAL:

finely punctate;

holotype M!

(Fig.

dorsal

of G.

scutellum

propodeum reticulate

between dorsal

posterior surface with 2 longitudinal

rugose;

smooth,

reaching mid-length of scutum;

keels;

and posterior

median area dull

part of radial 194)

or

vein much

with distal

inner

membranous process;

tibial

pubicornis in ST; lectotype M! and

2 paralectotypes MM! (hereby designated) of D. tenuicornis in ST; holotype F! of G. cephalotes in LU; holotype M! of A. breviventralis in BM;

lectotype F!

of D.

BM;

holotype F!

of Ch.

lucidus in DU; holotype F! of A. delicatulus in serratus in P; holotype M! of D. penidas in BM; holotype M! of A. vulgaris in BM; holotype F! of Ch. exiguus in HA; holotype of A. flaviscapus not found; lectotype F! and 2 paralectotypes FF! (hereby designated) of A. subarcticus in HE; holotype M! of A. fuscoclavatus in BM; holotype M! of A. divisus in BM; lectotype M! of D. alorus in DU; holotype M! of A. triangularis in BM. HOSTS: in Germany Macropsis sp. (Haupt 1941: host of a male! determined by Haupt as Ch. ephippiger); in England Streptanus sordidus (Zett. ), Psammotettix con finis (Dah lb. ), Euscelis plebejus (F a II. ) , Macrosteles sp. (Waloff 1974, 1975), Psammotettix nodosus (Rib.) (Richards 1948: under name lucidum). D I STR I BUT I ON: BU!

Road

plateau, Central

Pa I aearctic region:

to Mt.

Pektusan

Ryang-gang Prov.), aimak),

BU!

KOREA:

Hyesan

(Ryang-gang Prov.),

(24 Km NW from Sam-zi-yan, BU!

MONGOLIA:

Bogdo ul.

SO from Somon Bajanzogt

(Central

Chann-Pay (Ulan-Baator, aimak),

BU!

8 Km

W from Somon Urdtamir (Changaj Gebirge, Archangaj aimak), BU! 7 Km NW from Somon Chanzargalant (Bulgan aimak), BU! U.S.S.R. (Ponomarenko 1978: under name lucidum); FINLAND: Landiskar (Ab Nagu), HE! Lovisa, HE! Paanajarvi, HE! Lulto (Lps.), HE! Ivalo (W.-H.), HE! SWEDEN: Ljungby

(Sm.),

OT!

Eksharad

(Vmld.),

OT!

Fjellfota sjo.

(SK.),

BM!

Messaure, TW! Mausgnsbyn, OT! Solna (Bergshamra, Up.), BM! Smaland, ST! LU! Genarp (SK.), BM! NORWAY: Opheimsbakken (Aurdal), CO! Hatfjelddal,

HU!

POLAND:

Koscieliska Valley

(Tatra Mts.),

BM!

HUNGARY:

Budapest, BU! Retyezat, BU! HUvosvolgy, BU! Lesencistvan, BU! Nagykovacsi, BU! Matrasztistvan, BU! Bakony, BU! Vacz (Tudosdomb),

BU!

Borzsony hgs., BU! SUmeg (Sarvaly), BU! Holies, BU! Budai H., BU! Kisret (Szabadszallas), BU! Galyateto (Matra), BU! Velem (Koszegy hgs.), BU!

Meszes vgy.

PUspokladany, Matrasztlaszlo, BU!

(Koszeg),

BU! BU!

Szolnok, Gal-ret

CZECHOSLOVAKIA:

BU! BU!

Parad,

BU!

Inares,

Tabd i,

BU!

(Borzsony hgs.),

Bohemia,

TW!

Carlsbad,

BU!

R-Szombat,

Koroscsatoma BU! P!

(Gyula),

Tatatovaros, Bartfa,

BU!

BU!

BU! BU! Szod,

BULGARIA:

Anteoninae

Madara,

BU!

GERMANY: Rostock

Kuleftse,

Bell inchen

(Mecklenburg),

(Rhon-Gbg.), (Munster i. BM!

BU!

HU!

HU!

HA!

HU!

Kurpark

W.),

Weinberg

YUGOSLAVIA:

(Oder),

HU!

(Unna,

Vaake a. OT!

Rh.,

HU!

d.

(Slovenija), Ol!

lohr (Romberg),

Westf.),

Ingelheim a.

Schliersee (Bayern),

Ratece

Mosbrucher Hochmoor (Eifel),

Sendelbach b.

(Gross-Machnow),

Kgswusterh.,

321

HU!

OT!

Weser,

HU!

1m hart bei

Korheide u.

Harz Mts.

Oderberg

i.

M.,

Irsingen,

Gersfeld

Kanal

(Brocken Dist.),

HU!

Paunsdorf

CO!

HU!

Niederlehme b.

(leipzig),

HU!

Moorlake,

HU! HU!

Finkankrg., HU! Eisenach, HU! Roshenberg, HU! Halle, HU! Tegel (Berlin), HU! Fangschleuse (Berlin), HU! Freienwalde (Berlin), HU! Berl in,

HU!

Bot.

Tirol),

HU!

Obergurgl

(Tirol),

(Wien),

DENMARK:

Schneeberg CO!

Ostiylland,

Garten P!

CO!

(Berl in),

HOllAND:

Bennekomsa Meent,

Ol!

lE!

Gryon

SWITZERLAND:

(Vaud), lO!

lO!

GV!

Derborence Mt.

Moremont

Peney,

P!

(Cote-d'Or),

P!

Sollested

(Vaud),

GV!

lO!

GV!

Lolland),

Biithoven

Ardou

(Valais),

BELGIUM:

Buzenol,

GV!

Ferreyres

lO!

(Valais),

GV!

BM!

Auvernier,

GV!

lutry,

),

Korsor,

lO!

FRANCE:

Gorges du Rhoin

(det. Anteon viduum Maneval,

(H.-l.)

Arles-sur-Tech

CO!

lE!

(Vaud),

(Pyrenees Or.

(0.

P!

(Utrecht),

BM!

Follaterres

Huben

Reichenau,

lausanne (Vaud),

GV!

Chambon-sur-lignon P!

(Dyreh,

lE!

AUSTR I A:

P!

S.te Catherine

(Valais),

Arles-Corsavy road

undescribed species),

ZO!

(Stekelhoekduin),

(Vaud),

Valais),

Munchen,

Prater (Wien),

Utrecht,

Rockaje

Bois Noir

(Ardou,

Meczenc,

HU!

OT!

(Pyrenees Or.),

BM!

Dijon

(Cote-

d'Or), P! Biscarosse, P! Maisons-laffitte (Seine-et-Oise), P! Touraine, IN! Brout-Vernet, P! ENGLAND: Byflaet (Surrey), BM! Brabourne (Kent), BM! Wye (Kent), BO! Whaddon (Bucks.), BM! Ranworth Broad (Norfolk East),

BM!

(Surrey),

Colne Point BM!

Walmer

Matfieldlings

(S.

Oxfordshire),

BM!

Spratton

(NE.),

BM!

Boxmoor

(Kent East),

Yorks.),

BM!

(HT.),

Craswall

Chobham Comm.

Thames Ditton

(Northants),

BM!

(Surr-2Y),

BM!

leicester,

Hothfield Bog

(Kent),

BM!

Puttenham,

BM!

Mapledurham

(Oxon),

BM!

Ham Street

(Kent),

(Cambs.),

BM!

Brickleigh Vale

TW!

BM!

BM!

(Surrey), Torquay

Rhinefield

(Devon),

Richmond Park

(Herefordshire) , BM!

(Devon),

(Hants.),

Wimbledon Com.

BM!

BM!

Bix

BM! (Henley, OT!

BM!

(london),

OT!

Monks Wood NNR

Walford's Gibbet

(Quantock

Forest, SS.), BM! Banstead (SR.), BM! Dancer's End (Aston Clinton, BM! Walland Marsh (EK.), BM! Dawlish Warren (Devon), OT! WALES: LI anddona

(A.),

(Perths.),

BM!

Aberfeldy

BM!

SCOTLAND:

Mugdock,

BM!

Cadder

(Perths.),

BM!

Dumfries,

(Dublin),

BM!

Co. Kerry,

Glenasmole

(lanark),

Tentsmuir Nat. BM!

Res.

BM!

TW!

Athdown

Rannoch Forest

(Fife),

Bonar Bridge,

BX.),

BM!

BM!

Birks of

IRELAND:

(Wicklow),

BM!

Royal

Canal (leixlip, KD.), BM! Portmarnock (Co. Dublin), DU! Dowry (Wick low) , BM! Ballyknockan (Wicklow), BM! Agherlow, BM! Merrion Square S.

(Dubl in),

BM!

Benedetto Belbo

Don Bosco (Torino), Genova,

GE!

1979a), GE!

M.

ITALY:

(Cuneo), Pl!

Antola

Aspromonte

Currado and Olmi

Eaux Rousses Pl!

SZ!

la Cassa (Genova)

(Valsavaranche,

Perevagno (Cuneo),

(Torino),

Ol!

Siusi

SZ!

Aosta),

BM!

Castelnuovo

(Bolzano),

BM!

(=A. cameroni K. in Olmi and Currado

(Reggio Calabria),

1972 under name A.

BU!

the following

localities

vulgaris K.) are doubtful:

(in

322

Mem.

Farazzano

(Fori))

Amer.

(Zangheri

Ent.

1969)

Inst.,

no.

37,

1984

and Trieste (Kieffer in Kieffer et

Marshall 1905); ALBANIA: Ipek, BU! GREECE: Kifisia, BM! NOTES: Chelogynus exiguus Haupt 1941 is a very small species with antennal segment 3 nearly as long as segment 2; it's the same species later described by Jansson (1950) as Anteon flaviscapus (nec Anteon

trivia lis Kieffer var. flaviscapus Kieffer 1905) and by Hellen (1953) as Anteon (Chelogynus) subarcticus. I n European collections I have seen many males and females belonging to these forms. specimens with antennal

They are small

segment 3 as long as segment 2.

In the typical

pubicorne the antennal segment 3 is much longer than segment 2. A. exiguus should be considered a valid species. The study of many collections of pubicorne, however, showed that the proportions of antennal segments 2 and 3 were variable. (segment 3 twice as

These proportions vary from 2,00

long as segment 2)

to 1,66;

1,60;

1,50;

1,33;

1,30;

1,25; 1,00 (segment 3 as long as segment 2). This variability is easily visible if a large population from the same local ity is studied. In collections of HU there is a population from Huben showing

separation of Hellen)

(0.

Tirol,

these different proportions of antenna I segments.

Austria)

I think that the

A. exiguus Haupt (= flaviscapus Jansson; = subarcticus

is not proposable,

the range of variability

although the specimens at opposite extremes of

seem very different.

specimens emerge from hosts parasitized by

I

two

think that these small larvae of dryinids.

Usually under these conditions the only emerging specimen (generally one only specimen completes its metamorphosis) or the two emerging specimens are very sma II

and with colour,

scu I pture and antenna I proportions very

different.

9. Anteon scapu lare (Ha I i day)

Dryinus scapularis Haliday, in Walker 1837: 419. Anteon longiforceps Kieffer in Kieffer et Marshall 1905: 151 (syn. proposed by Richards 1939); locus typicus C I amart (France). Anteon carinatus Kieffer in Kieffer et Marshall 1905: 152 (syn. proposed by Richards 1939); locus typicus Maisons-Laffitte (France). Anteon scapularis (Haliday): Kieffer 1907b: 27. Chelogynus carinatus (Kieffer): Berland 1928: 163. Anteon scapulare (Haliday): Richards 1939: 260. Chelogynus longiforceps (Kieffer): Haupt 1941: 53. Chelogynus lanionis Haupt 1941: 55 (n. syn.); locus typicus Bellinchen

(Germany).

Chelogynus scapularis (Haliday): J.F. Perkins 1976: 18.

FEMALES:

fully

winged;

length 3,5-5,0 mm;

part of the frons testaceous;

black;

antennae testaceous,

rarely clypeus and with fuscous dorsal

Anteoninae

Fig.

195,- Chela of Anteon scapulare (Haliday) from Clamart (France)

markings;

antenna I segment

and clubs of femora brown; testaceous;

antennae

long,

antenna I segments 2+3 at shiny,

323

punctate,

10 brown;

legs testaceous,

mandibles yellow, wi th segment

usually with coxae

teeth brown;

1 sl i ghtl Y longer than

least 1,5 times as

with numerous

with

long as segment 1;

irregular short keels,

tegulae segment 4; head

without sculpture

324

Mem.

among keels, and 2

Amer.

Ent.

with frons usually

lateral

Inst.,

no.

granulated;

37,

1984

frons with a

keels near orbits directed towards

median

the antennal

keel

sockets;

pronotum dull, strongly rugose and reticulate rugose, smooth only near the posterior margin; scutum shiny, strongly punctate on anterior half, finely

punctate on posterior half;

reaching mid-length of scutum; finely punctate; keel

notaul ices distinct,

pronotum reticulate rugose,

between dorsa I and posterior surface;

longitudinal

keels;

near the margins;

incomplete,

scutellum and metanotum shiny,

median area smooth, fore-wing hyaline,

with a

transversal

smooth, irregular

posterior surface wi th

shiny,

2

not rugose or rugose only

with distal

part of radial

vein

much shorter than proximal part; fore tarsal segment 2 produced into a hook; segment 5 of front tarsus (Fig. 195) with 2 rows of 30-70 lamellae extending continuously proximal

Fig.

bristle;

196 -

tibial

to

the apex;

enlarged claw

(Fig.

195)

with

1

spurs 1,1,2.

Male genitalia of

Anteon scapulare (Haliday) from Boussens

(Switzerland):

right half removed.

Anteoninae

325

MALE: fully winged; length 2,62-3,25 mm; black; antennae testaceous, with segments 1-6 or 1-5 dorsally fuscous; legs testaceous, with hind coxae almost fully balck; brown;

head shiny,

tegulae testaceous;

punctate,

mandibles yellow,

without sculpture among keels with frons granulated; vertex granulated; pronotum rugose, smooth, half;

frons

dull,

with a median keel

strongly punctate on anterior half,

scutellum shiny, with a

smooth,

transversal

incomplete,

finely punctate;

irregular keel

finely punctate on posterior

propodeum reticulate rugose,

between

dorsal

keels;

not rugose or rugose only near margins; part of radial

inner process,

and posterior surface;

median area shiny,

fore-w;ng

vein much shorter than proximal

with no distal

keels; scutum shiny,

reaching mid-length of scutum;

posterior surface with 2 longitudinal

196)

teeth

sometimes also

and 2 lateral

smooth only near posterior margin;

notaulices distinct,

with

with nUmerous short keels or areolae,

hyaline,

part;

with proximal

smooth,

with distal

gonoforceps

dorsal

(Fig.

membranous

process; tibial spurs 1,1,2. LOCUS TYPICUS: England TYPICAL MATERIAL: lectotype F! of D. scapularis in DU; holotype F! of A. longiforceps in P; holotype F! of A. carinatus in P; holotype F! of Ch. lanionis in HA. HOSTS: in Germany Oncopsis lanio L. (Haupt 1941). DISTRIBUTION: GERMANY:

Palaearctic region:

Halle,

HA!

Bellinchen

HUNGARY:

(Oder),

HA!

BU!

Koszeg,

BU!

Dolauer Heide

Budapest,

(Halle),

HA!

Wiesbaden, HU! Aachen, HU! AUSTRIA, P! HOLLAND: F. Willemsen bijstation Santpoort, LE! SWITZERLAND: Viay (Vaud), LO! Boussens (Vaud),

OL!

Auvernier, Laffitte

La Sarraz

LO!

(Vaud),

FRANCE:

(Seine-et-Oise),

Brickhi II,

BX.),

BM!

LO!

Aclens

Brout-Vernet, P!

Clamart,

Spratton

P!

P!

(Vaud),

Dijon

ENGLAND:

(Northants),

LO!

Peney,

(Cote-d'Or),

P!

GV! Maisons-

Rammamex Heath

(Lit.

BM!

10. Anteon infectum (Haliday)

Dryinus infectus Haliday in Walker 1837: 419. Dryinus inclytus Haliday in Walker 1837: 421 (syn. proposed by Richards 1935); locus typicus England. Dryinus lateralis Thomson 1860: 178 (syn. proposed by Richards 1939); locus typicus Bohuslan (Sweden). Anteon fusiformis Kieffer in Kieffer et Marshall 1905: 166 (syn. proposed by Richards 1939); locus typicus Dumfries (Scotland). Anteon punctatus Kieffer in Kieffer et Marshall 1905: 168 (n. syn.); locus typicus Ma isons-Laffi tte (France). Anteon inclytus (Haliday): I
326

Mem. Amer.

Fig.

197 -

En!.

Ins!.,

no. 37,

1984

Chela of Anteon infectum (Haliday) from Maisons-Laffilte (France)

Anteon ellimani Chitty 1908: 143 (syn. proposed by Richards 1939);

Anteoninae

327

locus typicus Chesham (Eng I and). Chelogynus punctatus (Kieffer): Berland 1928: 166. Anteon infectum (Haliday): Richards 1939: 258. Chelogynus infectus (Haliday): J.F. Perkins 1976: 18.

FEMALE:

fully

winged;

length 3,00-6,43 mm;

with segments 6-10 or 7-10 dorsally fuscous; partly brown,

hind clubs of femora brown,

femora brown;

tegulae testaceous;

antennae short,

black;

antennae testaceous,

legs testaceous,

with coxae

sometimes also clubs of fore

mandibles yellow,

with segment 1 nearly twice as

with teeth brown;

long as segment 4;

segments 2+3 nearly as long as, or sl ightly longer than segment 1; head shiny, smooth, punctate, with frons reticulate rugose; frons with a short median keel and with 2 lateral keels; pronotum rugose, dull, smooth only near the posterior margin; scutum shiny, smooth, punctate; notaulices distinct, incomplete, reaching mid-length of scutum; scutellum and metanotum shiny, smooth, punctate; propodeum reticulate rugose, with a transversal irregular keel between dorsal and posterior surface; posterior surface with 2 longitudinal keels; median area shiny, smooth, rugose only near the margins; fore-wing with a dark transversal band near the pterostigma, with distal part of radial vein much shorter than proximal part; tarsal segment 1 not quite as long as segment 4; fore tarsal segment 2 produced into a hook; segment 5 of front tarsus (Fig. 197) with 2 rows of 12-36 lamellae extending continuously to the apex; enlarged claw (Fig. 197) with one bristle located on a proximal prominence; tibial spurs 1,1,2. MALE: fully winged; length 2,30-3,43 mm; black; antennae testaceous, with segments 1-8 or 1-10 dorsally fuscous; legs testaceous, with coxae partly brown or black and clubs of femora brown; sometimes clubs of fore femora testaceous; hind ti b i ae brown; tegu I ae testaceous; mand ib les yellow, with teeth brown; head shiny, punctate, with numerous short keels; frons with a median short keel and with 2 lateral keels; pronotum dull, rugose; scutum shiny, smooth, strongly punctate on anterior half, finely punctate on posterior half; notaul ices distinct, incomplete, reaching mid-length of scutum; scutellum and metanotum finely punctate; propodeum reticulate rugose, with a transversal irregular keel between dorsal and posterior surface; posterior surface with 2 longitudinal keels; median area shiny, smooth, rugose only near the margins; fore-wing hyaline, with distal part of radial vein much shorter than proximal part; gonoforceps (Fig. 198) with no distal inner process and with proximal dorsal membranous process; tibial spurs 1,1,2. LOCUS TYPICUS: England or Ireland (?) TYPICAL MATERIAL: lectotype F! of D. infectus in DU; holotype F! of D. lateralis in ST; holotype M! of A. ellimani in BM; holotype F! of A. punctatus in P; holotype F! of A. fusiformis in BM; lectotype M! of D. inc1ytus in DU. HOSTS: unknown antennal

328

Fig.

Mem. Amer.

198 -

Ent.

Inst.,

no.

37,

1984

Male genitalia of Anteon infectum (Haliday) (Switzerland): right half removed.

from Boussens

DISTRIBUTION: Palaearctic region: JAPAN: Kamikochi, TW! SWEDEN: Bohusl an, ST! HUNGARY: Badacsony, BU! Harsbokorhegy, BU! Budapest, BU! Mogyoroskest (Bakony), BU! Sranazug (Gyula), BU! ROMANIA (Nagy 1972); GERMANY: Eschwege, TW! AUSTRIA: Graz, P! Konigswarth, GV! DENMARK: Roden Sko.v (Lolland), CO! Kobenhavn, CO! HOLLAND: Putten (Gld.), LE! Rys-Goosterland, LE! SWITZERLAND: Follaterres (Valais), LO! Vitznau, BM! Boussens (Vaud), OL! Aclens (Vaud), LO! La Sarraz (Vaud), LO! Auvernier, LO! Peney, GV! FRANCE: Maisons-Laffitte (Seine-et-Oise), P! Vincennes, P! Foret de Chatillon (Cote-d'Or), P! Morlaix (Fl.), BM! ENGLAND: Chesham, BM! Dorking (Sr.), OT! Rhinefield (Hants), BM! Minsmere Reserve (Suffolk), BM! Fedlen (Herts), BM! Monks Wood (Cambs.), BM! Capler Wood (Fownhope, HF.), BM! Bricket Wood (HT.), BM! Bagley, BM! SCOTLAND: Dumfries, BM! IRELAND: Athdown (Co. Wick low) , BM! ITALY: Mombarcaro (Cuneo), PL! S. Benedetto Belbo

Anteoninae

(Cuneo), NOTES:

Oll

Tempio Pausania

(Sassari),

Oll

the only known specimen from Japan

(length 6,43 mm) 3,00-4,50).

than

329

(a female)

is much bigger

the specimens from western Palaearctic region

A Japanese sUbspecies? probably only

(mm

the study of males can

answer the question.

11. Anteon tripartitum Kieffer

Anteon tripartitus Kieffer in Kieffer et Marshall 1905: 194. Anteon tricarinatus Kieffer in Kieffer et Marshall 1906: 513 (n. syn.); locus typicus Botusfleming (England). Antaeon kiefferi Chitty 1908: 143 (n. syn.); locus typicus Ipswich (England) •

Chelogynus angusticoliis Berl and 1928: 165 (syn. proposed by Richards 1939); locus typicus Maisons-laffitte (France). Chelogynus prehensor Maneva I 1935: 9 (n. syn.); locus typicus Chambon-sur-lignon

(France).

Chelogynus berlandi Richards 1936a: 169 (syn. proposed by Richards 1939); locus typicus Ma isons-laffi tte (France). Anteon kiefferi (Chitty): Richards 1939: 274. Chelogynus silvaticus Ponomarenko 1970: 429 (n. syn.); locus typicus Kamenka (U.S.S.R.). Chelogynus kiefferi (Chitty): J.F. Perkins 1976: 22, 23. Anteon flavicorne (Dalman): Olmi and Currado 1976b: 75. Chelogynus silvaticus Ponomarenko: Ponomarenko 1978: 21.

FEMALE:

fully

winged;

segments 1-2 or only

length 3,12-3,50 mm;

1 yellow;

black;

legs testaceous,

antennae brown,

with

with hind coxae and

clubs of hind femora partly brown; tegulae testaceous; mandibles yellow, with proximal half black; teeth brown; antennae short, with segment, 1 nearly

twice as

long as segment 4;

head shiny,

smooth,

granulated,

with

frons and occiput strongly punctate; vertex not punctate; frons and occiput with numerous irregular keels; ocellar triangle with keels joining ocelli;

frons

rugose,

with a median keel

dull,

scutum shiny,

with transversal smooth,

punctate;

notaulices distinct,

keels;

incomplete,

scutellum and metanotum shiny,

propodeum reticulate rugose,

between dorsal

lateral

pronotum

smooth near the posterior margin;

strongly punctate on posterior half,

sculpture among punctures; length of scutum;

and 2 weak

keels,

with a

surface and posterior surface;

smooth,

transversal

without reaching midfinely irregular keel

posterior surface with 2

longitudinal keels; median area rugose; fore-wing hyaline, sometimes distally darkened, with distal part of radial vein much shorter than proximal

part;

enlarged claw

bearing one bristle;

(Fig.

199)

with a proximal

segment 5 of front tarsus

(Fig.

199)

prominence with 2 rows of

330

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

/

Fig.

199 -

Chela of Anteon tripartitum Kieffer from S. ( Italy)

Giovanni

Ilarione

Anteoninae

27-30 lamellae; tarsus MALE

apex with a group of 7-12 lamellae;

little slightly than segment 1;

a hook;

tibial

(Figs.

331

fore tarsal

segment 4 of front

segment 2 produced

into

spurs 1,1,2.

29,

180):

fully

winged;

length 2,37-3,00 mm;

black;

antennae

brown or black; legs brown or black, with junctions, tarsi and fore tibiae light; tegulae testaceous; mandibles yellow, with proximal half black

and teeth brown;

head dull,

mainly punctate on frons, with keels joining ocelli;

weakly granulated or alutaceous,

with numerous frons

irregular keels;

directed towards the antenna I sockets; strongly punctate on anterior half,

scutum shiny,

notaul ices distinct,

length of scutum;

scutellum and metanotum shiny,

longitudinal

and posterior surface;

keels;

part of radial 200)

vein

with distal

200 -

smooth,

irregular keel

posterior surface with 2 fore-wing hyaline,

much shorter than proximal tibial

punctate

finely

with a transversal

median area rugose;

Trieste

keels

hairy,

reaching 0,30-0,50

part;

with distal

gonoforceps

inner pointed process and with proximal

membranous process; LOCUS TYPICUS:

Fig.

incomplete,

propodeum reticulate rugose,

between dorsal

smooth,

without punctures or finely

on posterior half; punctate;

ocellar triangle

with a median keels and 2 lateral

(Fig.

dorsal

spurs 1,1,2.

(Italy)

Male genitalia of

Anteon tripartitum Kieffer from Colnbrook

(England):

right half removed.

332

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

TYPICAL MATERIAL: holotype F! of A. kiefferi in BM; holotype F! of Ch. silvaticus in PK; holotype F! of Ch. angusticollis in P; holotype F! of Ch. prehensor in P; lectotype M! and 3 paralectotypes MM! (hereby designated) of A. tripartitus in VI; lectotype M! (hereby designated) of A. tricarinatus in BU. HOSTS:

unknown

DISTRIBUTION: PK!

Palaearctic region:

HUNGARY:

Or Sz.

Matrasztistvan, (Nagy

Fu i renda I,

CO!

Korsor,

CO!

CO!

Nyon,

Chambon-sur-Lignon

Pelsocs, BU!

Altbuchhrst,

(Oberfranken),

SWITZERLAND:

U.S.S.R.:

BU!

SzSztMiklos,

1971); GERMANY:

Muggendorf LE!

BU!

Miklos,

HU!

Prater

FRANCE:

(Haute-Loire),

CO!

BU!

(Csakvar),

Scheffau

AUSTRIA:

(Perm Province),

Budaors,

Hajduvagas

North Zea I and,

GV!

Kamenka

BU!

(Bayern),

(Wien),

P!

HOLLAND:

Retyezat, BU! OT! DENMARK:

Putten

(G I d. ),

Maisons-Laffitte (Seine-et-Oise),

P!

ENGLAND:

New Forest

BU!

ROMANIA

(SH.),

P!

BM!

Colnbrook (Bucks.), BM! Botusfleming (Cornwall), BU! Ashford (Kent), BM! Ipswich (Suffolk East), BM! SCOTLAND: Forres (Moray), BM! Gretna, BM! ITALY: S. Benedetto Belbo (Cuneo), PL! Pietrabianca (Bussoleno, Torino), SZ! S. Giovanni Ilarione (Verona), OL! Naturno (Bolzano), OT! Trieste, VI!

12. Anteon crassifrons (Moczar) n. comb.

Chelogynus crassifrons Moczar 1983a: 181.

FEMALE:

fully

winged;

length 3,60 mm;

black,

with mandibles testaceous,

antennae testaceous,with segments 4-10 brown; brown;

antennae weakly distally

following proportions:

17:

10:

thickened;

13:

12:

frons almost fully reticulate rugose,

11:

antenna I segments

10:

dull,

legs testaceous; 10: 9:

rugose;

9:

12;

abdomen in

head with

vertex and occiput

shiny, smooth, finely punctate, without sculpture among punctures; frontal line complete; occipital carina complete; POL = 7; OL = 5; OOL 9; OPL = 8; TL tegulae; scutum,

=

7; pronotum shiny, rugose; pronotal tubercles reaching scutellum and metanotum shiny, smooth, finely punctate,

without sculpture among punctures;

notaulices incomplete,

pronotum nearly as

reticulate rugose;

posterior surface of propodeum with 2 longitudinal

keels;

median area as rugose as

transversal

keel

between dorsal

without dark transversal proximal 4: 5: (Fig.

part

bands;

(6: 13); fore tarsal

long as scutum

reaching mid-

length of scutum;

lateral

areas; vein

propodeum

propodeum also with a

and posterior surface; radial

(20:21);

fore-wing

with distal

hyaline,

part shorter than

segments in following proportions:

12:

12: 32; segment 2 of fore tarsi produced into a hook; enlarged claw 201) with one bristle located on a proximal prominence; segment 5

of front

tarsus

(Fig.

201)

with 2 rows of nearly 23

with a group of nearly 6 lamellae;

tibial

long

spurs 1,1,2.

lamellae;

apex

Anteoninae

Fig. 201 - Chela of Anteon crassifrons (Moczar) (type)

MALE: unknown LOCUS TYPICUS: Za-mo san (Korea)

333

334

Mem.

TYPICAL MATERIAL: HOSTS:

Amer.

holotype F!

Ent.

Inst.,

no.

37,

1984

in BU.

unknown

DISTRIBUTION: Palaearctic region: KOREA: Za-mo san NE from Pyongyan, pyong-sung Prov.), BU!

(Bek-sung-li,

60 Km

13. Anteon gaullei Kieffer

Dryinus dorsalis Nees (the types of Nees are lost: the name Dryinus dorsalis Nees is best ignored; see also nomina dubia): Thomson 1860:

177 (syn.

proposed by Richards 1939).

Anteon gaullei Kieffer in Kieffer et Marshall 1905: 161. Anteon cameroni Kieffer in Kieffer et Marshall 1905: 162 (n. syn.); locus typicus Thornhi II (Scotland). Anteon maculipennis Kieffer in Kieffer et Marshall 1905: 164 (syn. proposed by Richards 1939); locus typicus Smaland (Sweden). Anteon trivialis Kieffer in Kieffer et Marshall 1905: 201 (n. syn.); locus typicus Claddich (Scotland). Antaeon rufulocollis Ch itty 1908: 143 (n. syn.); locus typicus Tubney

(England).

Anteon dorsalis (Nees): Berland 1928: 159. Chelogynus gaullei (Kieffer): Berland 1928: 163. Anteon rufulocolle Chitty: Richards 1935: 150. Anteon maculipenne Kieffer: Richards 1935: 151. Anteon rufulocolle (Chitty): Richards 1939: 262. Anteon cameroni Kieffer: Richards 1939: 275. Anteon gaullei Kieffer: Richards 1939: 277. Chelogynus ephippiger (Dalman) partim (3 FF): Haupt 1941: 47. Chelogynus gaullei (Kieffer): J.F. Perkins 1976: 22. Chelogynus rufulocollis (Chitty): J.F. Perkins 1976: 22. Chelogynus cameroni (Kieffer): J.F. Perkins 1976: 22. Chelogynus rufulocollis (Chitty): Ponomarenko 1978: 21. Chelogynus cameroni (Kieffer): Ponomarenko 1978: 21. Chelogynus gaullei (Kieffer): Ponomarenko 1978: 21. nec Anteon cameroni Kieffer: Olmi and Currado 1979a: 347.

FEMALE

(Figs.

178,

179):

fully

winged

(sometimes fore-wings slightly

shortened); length 2,37-4,00 mm; black; antennae testaceous with segments 7-10 brown; occasionally brown with segments 1-2 or 1-3 testaceous; mandibles yellow, with teeth brown; propectus yellow or black (in the type yellow); pronotum yellow or reddish; scutum black; occasionally partly reddish-dark; femora brown;

legs fully

testaceous,

abdomen brown or black;

antennae variable;

head shiny,

smooth,

strongly punctate on vertex and occiput,

sometimes with stalk of hind

tegulae testaceous;

length of the

strongly punctate on frons, without sculpture among

less

Anteoninae

335

Fig. 202 - 'Chela of Anteon gaullei Kieffer from Dieppe (France)

punctures; frons with a short median keel, without lateral keels; pronotum shiny, strongly punctate on anterior half, finely punctate or without punctures on posterior half; punctate,

incomplete,

reaching mid-length of scutum;

length of scutum; punctate; between

scutum shiny,

without sculpture among punctures;

and posterior surface;

finely

occasionally only

scutellum and metanotum shiny,

propodeum reticulate rugose,

dorsal

smooth,

notaul ices distinct,

with a

smooth,

transversal

reaching 0,3

finely irregular keel

posterior surface with 2

longitudinal keels; median area rugose; fore-wing sometimes shortened, hyal ine or apically infuscate, with distal part of radial vein much shorter than proximal or

part;

longer than segment 4

segment

1 of front

(this characteristic

tarsus as

is very

long as,

variable;

shorter

it cannot

be used as specific character in identifying gaullei, cameroni and rufulocolle); fore tarsal segment 2 produced into a hook; segment 5 of front

tarsus

(Figs.

group of 7-20 located on a

9,

lamellae; proximal

202)

with 2 rows of 18-30

enlarged claw prominence;

(Figs.

tibial

9,

lamellae; 202)

apex with a

with one bristle

spurs 1,1,2.

MALE: fully winged; length 2,12-2,50 mm; black; antennae brown or brown-light; mandibles yellow, with teeth brown; legs testaceous, with hind coxae proximally black and sometimes with stalk of hind femora brown;

tegulae testaceous;

scu I pture among punctures,

head shiny,

strongly punctate,

in some areas rugose;

without

scutum sh i ny,

smooth,

336

Mem. Amer.

Fig.

203 -

Ent.

Inst.,

(Germany):

dorsal

without punctures or finely punctate

notaulices distinct,

incomplete,

scutellum and metanotum smooth,

propodeum reticulate rugose,

with a

and posterior surface;

1984

right half removed.

strongly punctate on anterior half, scutum;

37,

Anteon gaullei Kieffer from Reichenau

Male genitalia of

on posterior half;

no.

shiny,

reaching mid-length of without sculpture;

transversal

irregular keel

posterior surface with 2

between

longitudinal

keels;

median area rugose; fore-wing hyaline, with distal part of radial vein much shorter than proximal part; gonoforceps (Fig. 203) with distal inner pointed process and with dorsal spurs 1,1,2. LOCUS TYPICUS:

S.

Germain

TYPICAL MATERIAL:

proximal

membranous process;

tibial

(France)

holotype F!

of

A. gaullei in P (the supposed type in holotype F! of A. cameroni

DU designated by Richards 1939 is not valid); in BM;

lectotype F!

(hereby designated)

and 3 paralectotypes FF!

of A.

maculipennis in ST; holotype F! of A. rufulocollis in BM; holotype M! of A. trivia lis in BM. HOSTS: in Germany Macropsis sp. (Haupt 1941, under name Ch. ephippiger) . DISTRIBUTION: FINLAND: Gotland,

Palaearctic region:

Nystad, ST!

HE!

HUNGARY:

SWEDEN:

U.

S.

S.

Degaberga

Igenpataka,

BU!

R.

(Ponomarenko 1978);

(SK.),

BM!

CSipkeskut

Smaland,

(Bukk

hgs),

ST! BU!

VI! Alp.

Anteoninae

Kudsir, BU!

BU!

Csevharaszt,

Horvatkimle,

Templomter

BU!

(Tompa),

BU!

Nadudvar (Hortobagy), (Kdezegi

hgs.),

BU!

BU!

Bartia,

Nagyvisnyo,

Emlekerdd

BU!

Kisret

Szabohegy

BU!

BU!

337

Vacz

(Tudosdomb),

Nagyszenas (Budai

(Szeged),

BU!

Oroszmezd,

(Szabadszallas),

(Kdszeg),

BU!

BU!

BU!

H.),

BU!

Pelsocs,

Meszes vgy.

Koszeg,

BU! BU!

Velem

(Koszeg),

BU!

Holics, BU! Budapest, BU! Retyezat, BU! CZECHOSLOVAKIA: Karlovy Vary (Bohemia), P! GERMANY: Bellinchen (Oder), HA! Aachen, HU! Mainz, OT! Naunhof

(Leipzig),

HU!

(Schleswig-Holstein), AUSTRIA: (Wien),

Turnau P!

GV!

B.,

Nyon,

Mesni I-Ie-Roi Gui Ihem

(Steiermark),

(Sealand),

Ulvenhout N.

(Valais),

Reichenau

Kaltenleutgeben

Ruderhegn LE!

Fangschleuse (Berl in),

CO!

(Wien), HOLLAND:

LE!

Wijster

GV!

P!

Huben VI!

WA!

P!

S.

Berl in,

(0.

Tirol),

Waidbruck.

Den Haag,

(Dr.),

Corcelettes

(Seine-et-Oise),

(Herault),

WA!

HU!

(Baden-Wurttemberg),

LE!

Pont du Gard

HU!

(Tirol),

LE!

LO!

HU!

Utrecht,

FRANCE:

P!

ENGLAND:

EB!

P!

Prater

DENMARK:

LE!

Asperen,

Vouvry

Dieppe,

(Seine-et-Oise),

(Gard),

Malente

Halle,

Wien,

SWITZERLAND:

(Vaud), Germain

HU! P!

P!

P!

51.

Bald Hi II

(Lewknor, Oxfordshire), BM! Boxmoor (Herts), BM! Hay tor (Dartmoor, Devon), OT! Finningley (5. Yorks), BM! Thames Ditton (Surrey), BM! Skotover,

BM!

Leicester,

(Berks.),

BM!

Ham Street

TW!

(Kent),

BM!

Wimbledon Com.

Yorks),

BM!

Norfolk,

(Argyll),

BM!

BM!

IRELAND:

Spratton

(Kent),

(Northants),

BM!

Tubney

(London),

SCOTLAND:

Murlough Ho.

OT!

Windsor Forest BM!

Faversham

Hatfield Lings (Dunsville,

Thornhill (S.

BM!

(Berks.),

DO.),

(Dumfries), BM!

BM!

5.

Claddich

Tollymore Park

(Co.

DO.), BM! Kerry, TW! Co. Wicklow, TW! ITALY: 5. Benedetto Belbo (Cuneo), PL! Pietrabianca (Bussoleno, Torino), SZ! S. Mauro (Torino), OL!

Oropa

(Bolzano),

(Vercelli), OT!

MI!

Sestola

Pizzighettone (Cremona),

(Modena),

GE!

BO!

TW!

Monte Peglia

Naturno (Perugia),

OL!

14. Anteon fulviventre (Hal iday)

Dryinus fulviventris Hal iday in Curtis 1828: 206. Dryinus fuscipes Thomson 1860: 177 (n. syn.); locus typicus Stockholm (Sweden).

Anteon ephippiger (Dalman) var. similis Kieffer in Kieffer et Marsha II 1905: 156 (syn. proposed by Richards 1935); locus typicus Ireland.

Anteon gracilicollis Kieffer in Kieffer et Marshall 1905: 165 (syn. proposed by Richards 1939); locus typicus Poss i I Marsh (Scot I and) • Anteon flavinervis Kieffer in Kieffer et Marshall 1905: 192 (n. syn.); locus typicus Dumfries (Scotland). Anteon parvulus Kieffer in Kieffer et Marshall 1905: 193 (n. syn.); locus typicus Dumfries (Scotland). Anteon xanthostigma Kieffer in Kieffer et Marshall 1905: 198 (n. syn. ); locus typicus Dumfries (Scot I and) • Anteon trivialis Kieffer var. flaviscapus Kieffer in Kieffer et

338

Mem.

Marshall

1905: 201

Amer.

(n.

En!.

no.

37,

1984

locus typicus Scotland (holotype with

syn.)

indecipherable local ity

Ins!.,

label).

Anteon parvus Kieffer in Kieffer et Marshall 1906: 523 (n. syn.); locus typicus Dumfries (Scotland). An teon fu sc ipes (T homson ): Kieffer 1907b: 26. Chelogynus alutaceus Richards 1935: 152 (syn. proposed by Richards 1939); locus typicus unknown ("British" in the locality label of the holotype) •

Anteon jurineanum Latreille partim (only M): Richards 1939: 257. Anteon fulviventre (Haliday) partim (only M): Richards 1939: 264. Anteon fulviventris (Haliday): J.F. Perkins 1976: 20, 22.

FEMALE:

fully

winged;

length 2,0-3,0 mm;

or reddish more or less dark; fully

colour variable;

head yellow

occasionally black or brown;

yellow or brown with segments 6-10 or 7-10 yellow;

antennae

pronotum and

scutum yellow or reddish light; occasionally dark or brown; scutellum and metanotum yellow (sometimes with anterior margin of metanotum black)

or reddish;

yellow or black; brown or reddish; with

teeth brown;

segment 4; short;

occasionally blackish;

propodeum black, legs testaceous; antennae short,

head dull,

pronotum

dull,

mesopleura and metapleura

sometimes with sides yellow; tegulae testaceous; with segment

not punctate, granulated;

1 nearly

granulated, scutum

twice as

smooth,

shiny,

abdomen

mandibles yellow frontal

smooth,

long as

line

without

sculpture or weakly alutaceous; notaulices distinct, incomplete, reaching mid-length of scutum; scutellum and metanotum shiny, smooth, without sculpture;

propodeum reticulate rugose,

between dorsal longitudinal distal front

and posterior surface;

keels;

with a

transversal

irregular keel

posterior surface with 2

median area rugose,

dull;

fore-wing

hyal ine,

vein much shorter than

proximal

tarsus s light I y

shorter than segment 4;

fore tarsa I segment 2

produced

into a

hook;

segment 5 of front

tarsus

(Fig.

part;

with

part of radial

204)

segment with

1 of

1 row of

20-21 lamellae; apex with a group of 3-6 lamellae; enlarged claw (Fig. 204) with one bristle located on a proximal prominence; tibial spurs 1,1,2. MALE: fully winged; length 1,87-2,50 mm; black; antennae brown-yellow; mandibles yellow, with teeth brown; legs fully yellow, sometimes with stalks of hind femora brown; granulated, keels;

tegulae testaceous;

without punctures;

scutum shiny,

smooth,

frontal

hairy,

head dull,

line present;

smooth,

frons without

lateral

wothout sculpture or weakly

granulated; notaulices distinct, incomplete, reaching mid-length of scutum; scutellum and metanotum shiny, smooth, without sculpture; propodeum reticulate rugose,

with a

transversal

irregular keel

between

dorsal surface and posterior surface; posterior surface with 2 long i tud i na I keel s; med i an area rugose; fore-wi ng hya line, with d i sta I part of radial 206)

vein much shorter than proximal

with distal

part;

gonoforceps (Fig.

inner pointed process and with proximal

very

short

339

Anteoninae

!

/

..

C~j

~ \_;;1 ..

\

Fig. 204 -

dorsal

Chela of

//

,

/ ~/

/

Anteon fulviventre (Haliday) from Spratton (England)

membranous process;

LOCUS TYPICUS:

/'

)

tibial

spurs 1,1,2.

England

TYPICAL MATERIAL:

lectotype F!

of D.

fulviventris in DU; lectotype F!

Mem. Amer.

340

En!.

Ins!. , no .

37,

1984

•

Fig.

205 -

Head of

Anteon fuiviventre (Haliday) from Bennekomsa Meent (Holland) •

Anteoninae

341

\\V':

I

I

'

\

\

j

I

II) \ \

Fig.

Male genitalia of Anteon fulviventre (Haliday) (England): right half removed.

206 -

from Spratton

(hereby designated) of D. fuscipes in ST; flavinervis in BM; holotype M! of A. xanthostigma in BM; holotype M! of A. parvulus (~ A. parvus) in BM; holotype F! of A. gracilicollis in BM; holotype M! of A. trivia lis var. flaviscapus in BM; holotype F! of Ch. alutaceus in DU; hoi otype F! of A. similis never and 3 paralectotypes FF! holotype M!

of A.

existed (see Richards 1935). HOSTS: in England Nocydia crocea

H.S.

(J.F.

Perkins 1976;

Chambers

1967) • DISTRIBUTION: Palaearctic region: U. S. S. R. ? (Ponomarenko 1978)(in Ponomarenko, f,:oscow and Leningrad collections I haven't seen specimens of fulviventre from U.

S.

s.

R.;

were gaullei or ephippiger); ST!

HUNGARY:

N.

Barszentmihaly,

Veracze, BU!

the specimens determined as fulviventre

FINLAND

BU!

Budapest,

(Hellen 1953); SWEDEN:

Barlangliget, BU!

Gyula

BU!

Nagyenyed,

(Karas Csat.),

BU!

Stockholm, BU! Tihany,

BU!

Mecsek hgs., BU! Tatatovaros, BU! Rev" BU! Visegrad, BU! Lasztonya, BU! CZECHOSLOVAKIA: Kytin (Bohemia), TW! GERMANY: Halle, HA! Mainz, OT! CO!

Niemegk, HU! Thuringen, GE! Finkenkrug (Berlin), HU! Glucksburg, AUSTRIA: Prater (Wien), P! DENMARK: Langeland I., CO! Kobenhavn,

CO!

HOLLAND:

(Oosteinde 34),

Bennekomsa Meent, OL! LE!

SWITZERLAND:

Pullen

Cudrefin

(G I d. ), (Vaud),

LE! CA!

Waarder FRANCE:

342

Mem.

Amer.

Ent.

Maisons-Laffitte (Seine-et-Oise),

Inst.,

P!

37,

no.

Le Puy,

P!

1984

Prechacq

(Landes)

(del.

Ch. l'uioval'iegatus by Berland), P! ENGLAND: Wytham (Berks), BM! Spratton

(Northants),

(Cambs.),

BM!

BM!

Ramsey

Rhinefield

(SH.),

BM!

(Hants),

BM!

Wood Walton

Castor Hanglands

Fen

(Cambs.),

BM!

St.

Catherine's Point (Watershoot Bay, IW.), BM! Torquay (Devon), OT! Burnham Beeches (Buckes.), BM! SCOTLAND: Rannoch Forest (Perths.), BM!

Possil

TW!

Ireland,

Marsh

(Vercelli), road,

(Lanark),

BM!

ITALY:

Benedetto Belbo

DU!

SZ!

S.

Dumfries,

Pizzighettone (Cremona),

High Atlas),

8M!

IRELAND:

(Cuneo),

TW!

MAROC:

Co.

PL! Asni

Wicklow,

SZ!

Mongrando

(Marrakesh

BM!

15. Anteon hilal'e n. sp.

FEMALE:

fully

winged;

antennae testaceousj

length 3,75 mm;

clypeus and

black;

mandibles

anterior half of frons and

testaceous;

legs

testaceOU5,

with stalks of hind femora and hind coxae partly black; antennae slightly distally thickened; antennal segments in following proportions 10:

7: 14:

10: 10: 11:

10: 9: 9: 14; head strongly punctate, without

sculpture among punctures; complete;

POL

=

4; OL

=

frontal

=

4; OOL

I ine present;

7,5; OPL

=

occipital

5,5;

TL

=

carina

8; pronotum

shiny, hairy, punctate, without sculpture among punctures; posterior surface of pronotum nearly as long as scutum (14:14); scutum shiny, punctate,

without sculpture among punctures;

notaul ices

reaching approximately 0,65 length of scutum; smooth,

shiny,

longitudinal areas;

without

sculpture;

propodeum

keels on posterior surface;

fore-wing hyaline,

incomplete,

scutellum and metanotum

reticulate rugose,

with

2

median are as rugose as lateral

without dark transversal

bands;

distal

part of

(9:14); fore tarsal segments in following proportions: 8: 2,5: 4: 16: 28; enlarged claw (Fig. 207) with one bristle located on a proximal prominence; segment 5 of front tarsus (Fig. 207) with 2 rows of 30 lamellae; apex with a group of 5 lamellae; tibial spurs 1,1,2. radial

vein shorter than proximal

MALE:

unknown

LOCUS TYPICUS:

Godavari

TYPICAL MATERIAL: HOSTS:

part

(Nepal)

holotype F!

in OT;

1 paratype F!

in OL.

unknown

DISTRIBUTION:

Palaearctic region:

NEPAL:

Godavari

(Katmandu),

OT!

OL!

16. Anteon ingenuum n. sp.

FEMALE: MALE:

unknown

fully

winged;

length 2,81-3,43 mm;

antennae brown or testaceous;

black;

legs testaceous,

mandibles testaceous;

with

hind coxae basally

Anteoninae

Fig. 207 - Chela of Anteon hilare n. sp.

343

(type)

344

Mem.

Fig.

208 -

Amer.

Male genitalia of

Ent.

Inst.,

no.

37,

1984

Anteon ingenuum n. sp. (type): right half removed.

black;

antennae not distally thickened;

proportions:

12: 7:

9:

7:

7:

8:

8:

without sculpture among punctures, mainly near the orbits; the orbits;

occipital

9:

antenna I segments in following

9:

10;

head shiny,

punctate,

with more or less numerous areolae

frons with frontal

carina complete;

POL

I ine and

=

7;

OL

two

=

3;

lateral OOL

=

keels near 7;

OPL

3; TL = 3,5; scutum, scutellum and metanotum shiny, smooth, finely punctate, without sculpture among punctures; notaulices incomplete, reaching mid-length of scutum; propodeum reticulate rugose, with 2 longitudinal keels on posterior surface; median area smooth, shiny; lateral distal (Fig.

areas rugose; part of radial

208)

without dorsal

inner process;

tibial

LOCUS TYPICUS:

hyaline,

without dark transversal

membranous proximal

part

(3:10);

bands;

gonoforceps

process and without distal

spurs 1,1,2.

Godavari

TYPICAL MATERIAL: MM! in OL. HOSTS:

fore~wing

vein shorter than proximal

(Nepal)

holotype M!

and 3 paratypes MM!

in OT;

2 paratypes

unknown

DISTRIBUTION:

Palaearctic region:

NEPAL:

Godavari

(Katmandu), OT!

OL!

Anteoninae

345

17. Anteon coI'ax n. sp.

FEMALE:

unknown

MALE: fully winged; length 2,5-3,0 mm; black; mandibles testaceous; antennae testaceous-dark; legs testaceous, with hind coxae basally black; antennae not distally

thickened;

antenna I segments

proportions: 12: 8: 11: 10: 10: 10: 10: 11: without sculpture among punctures; frontal complete;

POL = 5;

OL = 3;

OOL = 7,5;

OPL = 3,5;

scutellum and metanotum finely punctate, punctures; scutum;

notaul ices incomplete,

reaching

propodeum reticulate rugose,

posterior surface; hyaline,

shorter than proximal membranous proximal

part

(8:12);

scutum,

approximately 0,65

with 2

bands;

TL = 5;

without sculpture among

median area as rugose as

without dark transversal

in following

10: 12; head shiny, punctate, I ine present; occipital carina

longitudinal lateral

distal

gonoforceps

length of

keels on

areas;

fore-wing

part of radial (Fig.

process and without distal

209)

vein

with dorsal

inner process;

tibial

spurs 1,1,2. LOCUS TYPICUS:

Godavari

TYPICAL MATERIAL: MM!

(Nepal) and 5 paratypes MM!

holotype M!

in OT;

3 paratypes

in OL.

HOSTS:

unknown

DISTRIBUTION: Pulchauki

Palaearctic region:

(I(atmandu),

NEPAL:

Godavari

(Katmandu),

OT!

OL!

OT!

18. Anteon munitum n. sp.

FEMALE: MALE:

unknown

fully

winged;

length 2,12-3,43 mm;

antennae testaceous or brown; black;

black;

legs testaceous,

mandibles testaceous;

with

occasionally testaceous with all coxae and

partly brown,

antennae not distally

following proportions:

9:

6:

8:

8:

thickened;

7,5:

8:

8:

hind coxae basally

all

clubs of femora

antennal

8:

8:

10;

segments in head shiny,

punctate, without sculpture among punctures; frontal line present; occipital carina complete; POL = 7; OL = 4; OOL = 6; OPL = 3; TL = 4; scutum,

scutellum and metanotum finely punctate,

punctures;

notaulices incomplete,

propodeum reticulate rugose,

with 2

surface;

median area smooth,

hyaline,

without dark transversal

shorter than proximal membranous proximal

part

without sculpture among

reaching mid-length of scutum; longitudinal

shiny;

(3:11);

lateral

bands;

keels on posterior

areas rugose;

distal

gonoforceps (Fig.

process and without distal

fore-wing

part of radial 210)

vein

with dorsal

inner process;

tibial

spurs 1,1,2. LOCUS TYPICUS: Godavari (Nepal) TYPICAL MATERIAL: holotype M! and 31

paratypes MM!

in OT;

16

Mem.

346

Amer.

En!.

Ins!.,

~

no.

37,

1984

/

/

Fig.

209 -

Male genitalia of

Anteon corax n. sp. (type): right half removed.

paratypes MM! HOSTS:

in OL.

unknown

DISTRIBUTION: Pulchauki

Palaearctic region:

(Katmandu), OT!

OL!

NEPAL:

Godavari

(Katmandu),

OL!

OT!

347

Anteoninae

I

5'~~

~

,

\

"\

'\

\

\

!

Fig.

210 -

Male genitalia of

Anteon munitum n. sp. (type): right half removed.

19. Anteon japonicum n. sp.

FEMALE: MALE: and dull,

unknown

fully

winged;

legs fully strongly

complete;

POL

length 2,50 mm;

testaceous

reticulate rugose; ~

4;

Ol

~

black;

mandibles,

antennae,

(except for hind cOXae basally black); 3;

OOl

frontal ~

7;

OPl

line absent; ~

4;

occipital

Tl = 5;

scutum

tegulae head

carina dull,

strongly reticulate rugose on anterior half and strongly punctate on posterior half; notaulices incomplete, reaching only approximately 0,25 length of scutum;

scutellum and metanotum shiny,

smooth,

strongly

348

Mem. Amer.

En!.

Ins!.,

no.

37,

1984

B

Fig.

211

punctate; dorsal

-

Male genitalia of Anteon japonicum n. sp. (type): (A: dorsally viewed; B: laterally viewed).

propodeum reticulate rugose,

and posterior surface;

(4:12);

bands;

gonoforceps

and without distal

a transversal

areas;

distal

part of radial

(Fig.

211)

fore-wing

hyaline,

between

inner process;

holotype M!

gonoforceps

JAPAN:

proximal

(laterally viewed)

in B!

HOSTS: unknown DISTRIBUTION: Palaearctic region:

keels;

without dark

vein much shorter than proximal

with membranous dorsal

apex broad; tibial spurs 1,1,2. LOCUS TYPICUS: Tokyo (Japan) TYPICAL MATERIAL:

keel

posterior surface with 2 longitudinal

median area as rugose as lateral transversal

with

gonoforceps

Tokyo,

B.

ETH I OP IAN SPEC I ES OF ANTEON

Thirty seven species are known.

KEY TO THE SPECIES

process with

Anteoninae

349

FEMALES

Species brachypterous Species

2

(Fig.

212) .•

4 of front tarsus slightly shorter, nearly as long as, than basal part of segment 5 (Fig. 223) ••••••••••••• 3 Segment 4 of front tarsus at most 0,5 as long as basal part of segment 5 (Fig. 243) •••••.•••••••••.••.•.•••.••.••••••••••••• 16 Posterior surface of propodeum with two longitudinal keels ••••• longer

. . . . . • . . . . • . . . . • . . . • . . . . • . . • • . . . •• Posterior surface of

4

brachypterum n. sp.

1.

winged •••••••••••••••••..•••.••••.••.•••••••••.• 2

Segment

or

3

fully

Segment

5 of

without

a distinct

front

tarsus with basal

agile n. sp.

2.

propodeum without

tarsus with apical part

inner

region

longer

longitudinal

side almost

(Fig.

217);

than apical

segment

part

keels ••••• 4

straight,

(Fig.

5 of front 217) ••••••

...•........••..•.•...•..........• 3. maritimum (Turner) 5 of front tarsus wi th inner side curvi I inear, wi th a apical region (Fig. 223); segment 5 of front tarsus with basal part shorter than apical part (Fig. 223) ••••••••••• 5 5 Fore-wing wi th distal part of radial vein more than 0,5 as long as proximal part •••••••••••••••••••••••••••••••••••••••••.•••• 6 Fore-wing wi th distal part of radial vein 0,5 or less than 0,5 as long as proximal part . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 6 Pronotum wi th posterior surface longer than anterior surface, more than 0,5 as long as scutum . . . 4. kasaicum (Benoit) Pronotum wi th posterior surface shorter than anterior surface, less than 0,5 as long as scutum . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 Species mostly reddish-testaceous. 5. fiorii n. sp. 7 Species mostly black . . . . . . . . . . . . . . 6. kivuanum (Benoit) 8 Fore-wing with 1-2 dark transversal bands . . . . . . . . . . . . . . . . . . . . . 9 Fore-wing hyaline, without dark transversal bands . . . . . . . . . . . . 12 9 Posterior surface of propodeum sculptured by strong transversal keels, not reticulate rugose; pronotum humped, with posterior surface shorter than anterior surface; head fully granulated, with areolae or irregular keels on vertex and frons . . . . . . . . . . . . • • . • • • • • • • • • . • • • • • • . . • • • . . • • • • • • •• 7. taylori n. sp. Posterior surface of propodeum weakly reticulate rugose, wi th or without transversal keels; pronotum flat, with posterior surface longer than anterior surface; head fully or almost fully granulated, with only weak tracks of areolae . . . . . . . . . . . 10 10 Head and thorax almost fully black ••••••••••••••••••••••••••••• · . . . . . . . . . . . . . . . . . . . . • . . • • . . . . . . • , 8. dayi n. sp. Head and thorax fully or almost fully reddish-testaceous . . . . . 11 11 Propodeum fully reddish-testaceous ••••••••••••••••••••••••••••• • • . • • . . • • . • • • • • . . • • . . . • • . • • • . • • •. 14. townesi n. sp. Propodeum almost fUlly brown ••••• 15. decellei n. sp. 12 Head with at least the frons reticulate rugose ••••••••••••••• 13 Segment

distinct

350

Mem. Amer.

Head

fully

not

reticulate

En!.

not

with

only

reticulate

9.

frons

1984

wi thout

reticulate

rugose;

canabense (Benoit)

reticulate

rugose;

• . . . . . . . . . . . . • . . . . . . . . • . . . . . . . . .• 14 Scutum weakly punctate,

37,

scutum partly

reddish-testaceous.

Head granulated, granulated,

no.

rugose ••••.•••••••••••.•••••••••••• 14

13 Head fully reticulate rugose; species mostly

Ins!.,

rugose;

scutum

species mostly black •••

10.

ugandanum n.

sp.

sculpture among punctures,

not

granu I a t ed . . . . . . . . . . . . . . . . . . . . . . . 11. kawandanum n. sp. Scutum fully or almost fully granulated •••••••••••.••••••.••• 15

15 Species mostly black; nctaulices reaching approximately 0,5 length of

scutum . . . . . . . . . . . . . . . . .

Species mostly

12.

reddish-testaceous;

visible near anterior margin

of

airum n. sp.

notaulices only

faintly

scutum . . . . . . . . . . . . . . . . . . . . . . . . .

· . • . . . . . • . . . . . . . . . . . . . . . . . . • . . . .. 1 3. turneri n. sp. 16 Posterior surface of propodeum without longitudinal keels or with tracks of only two longitudinal keels . . . . . . . . . . . . . . . . . . . 17 Posterior surface of propodeum with two strong longitudinal keels •••••••••••••••••••••••••••••••••••••••••••••••••••••••• 26 0,5 as long as

17 Pronotum with posterior surface very short, scutum or

shorter •••••••••••••••••••••••••••••••••••••••••••• 18

Pronotum with posterior long as

18 Notaul ices weak, scutum;

surface very

long,

more

0,5 as

than

scutum ••••••••••••••••••••••.••••••••••••••••.••••••• 24 species

but fully

reaching approximately

0,5

length

of

testaceous,

with petiole black . . . . . . . . . .. .•....•••.•......•••••••.....•... 16. anthereon Benoit

Notaulices invisible or faintly visible near anterior margin of scutum; species almost fully black . . . . . . . . . . . . . . . . . . . . . . . . . . . 19 19 Fore-wing with

1-2 dark

Fore-wing hyal ine,

20 Head granulated, not

21

transversal

wi thout

dark

reticulate

bands •••••••••••••••••••• 20

transversal

bands . . . . . . . . . . . . 21

rugose •••••••••••••••••••••••••

• • • • • • • • • • . • . • • • • • • • • • . • • • • • • • • ••

17. namibiense n.

Head

18.

reticulate

Head granulated, reticulate Head fully

rugose . . . . . . . . . . . with weak

irregular

sp.

gutturnium (Benoit) short

keels,

not

rugose •••••••••••••••• 19. arabicum n. sp. reticulate rugose ••••••••••••••••••••••••••••••••• 22

22 Scutum fully

reticulate rugose . . . 20. Scutum punctate or weakly granulated,

23 Scutum punctate,

wi thout

fully

reticulate

• • • • • • . • • • • • • • • • • . • . • • • • • • • • • . • •• Scutum weakly

granulated, fully

reticulate

sculpture among punctures;

surface of pronotum almost

pronotum almost

natalense n. sp. not

not

smooth,

21.

rugose.

hoyoi n. sp.

punctate; without

rugose •• 23 posterior

posterior surface of

sculpture ••••.•••••••••••

. . . . . . . . • . . . . . . . . . . . . . . • . . . . . . . . . 22. mingoae n.

sp.

24 Fore-wing wi th a dark subapical transversal band; posterior surface of propodeum almost smooth, wi th weak irregular keels . . . . . . . . . . . . . . . . . . . . . . . . . . . . Fore-wing hyaline,

without

dark

23.

longiscapum (Ceballos)

transversal

bands;

posterior

Anteoninae

surface of

propodeum strongly

reticulate

25 Head fully reticulate rugose ••••• Head

fully

granulated,

weak areolae near 26 Posterior with keels

not

surface of

25.

rugose •••••••••••••• 25

madagascolum (Benoit) rugose

or

only

with

very

medleri n. sp.

propodeum with median area most Iy

only around

smooth,

the margins •••••••••••••••••••••••••••••

••••••••..••.•.••.••••••••.•••..• Posterior

24.

reticulate

the clypeus ••••

surface of

351

26.

hovaBenoit

propodeum with median

with numerous keels also

in

the central

area

fully

rugose,

region . . . . . . . . . . . . . . . 27

27 Head wi th OL longer than POL; fore-wing hyal ine, not darkened .. · • • . • . • • • • • • • . . • • . • • • • • . • • • • • • • .• 27. compressicoxale (B en 0 i t ) Head with OL shorter than POL; fore-wing with 1-2 dark transversal bands •••••••••••••••••••••••••••••••••••••••••••• 28 28 Scutum shiny, smooth, weakly punctate, without sculpture among punctures;

species mostly black ••••••••••.•••••••.•••••••••

. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ... Scutum dull,

granulated,

not

28.

inflatrix Benoi t

reticulate

redd ish. • • • • • • • • • • • • • • • • • • • • • • • ••

29.

rugose;

species mostly

rufonigrum n. sp.

The females of A. zair2nse Kieffer, oriphilum Kieffer, fabulosum n. abruptum n. sp. mosseli n. sp., capei n. sp., emeritum n. sp., garambanum n. sp. are unknown. sp.,

MALES

2

two

long i t ud i na I

Posterior

surface of

propodeum wi th

Posterior

surface of

propodeum wi thout

Posterior

surface of

propodeum wi th median area

long i t ud ina I

kee Is •••

2

keels ••••• 7

sh i ny,

at

I east

smooth, not rug ose •••••••••••••••••••••••••••••• . . . . 3 Posterior surface of propodeum wi th median area du I I , rugose, with numerous keels or areolae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 cen t ra I I y

3

Head punctate, without distal

wi thout sculpture among punctures; gonoforceps inner pointed process (Fig. 252) . . . . . . . . . . . . . . ..

.................................. Head punctate, distal

inner pointed process

(Fig.

• •..•••..•••.•••••••••••••.••••• ,

4

Head

strongly

distal

30.

oriphilum Kieffer

granulated among punctures;

punctate,

not

inner pointed process

Head reticulate pointed process

reticulate (Fig.

gonoforceps with

253) . . . . . . . . . . . . . . . . . . . . . . .. 31. fabulosum n. sp. rugose;

gonoforceps with

216) •••••••••••••••••••••••. 2. agile n. sp.

rugose; gonoforceps without distal inner (Fig. 254) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ..

• •... " . • . • . . . . . • . . . • . . . . . . . . • . . . . . . . . . • . . . . . . . . . . . . • . . . . . . . . . 5 5

Posterior

surface of

propodeum with median area

sculptured by

352

Mem.

areolae much

smaller

Amer.

than

Ent.

Inst.,

those of

no.

I at era I

surface of

areolae as

6

large as

Gonoforceps

1984

areas •••••••••••••••

abruptum n.

32. Posterior

37,

propodeum with median area those of

slightly

shorter

sp.

sculptured by

areas ••••••••••••••••••.• 6

lateral

than penis

(Fig.

255) •••••••••••••

••••••••••••••••••••••••••••••••• 33. zairense Benoi t Gonoforceps much shorter than penis (Fig. 250) •••••••••••••••••

• •••••••• " 7

28. inflatrix Beno it inner pointed process (Fig.

••••••••••••••••••••••

Gonoforceps without

distal

Gonoforceps wi th distal

inner pointed process

(Figs.

259) •••••••••••••••••••••••• , •••••••••••••••• '"

8

Propodeum without posterior

a

strong

transversal

keel

227) ••• 8 258,

•.••••••••••• 17

between dorsal

and

surface ••••••••••••••••••••••••••••••••••••••••••••• 9

Propodeum wi th a strong transversal keel between dorsal and posterior surface •••••••••••••••••••••••••••••••••••••••••••• 13

9

Head granulated •••••••••••••••••••••••••••••••••••••••••••••• 10 Head punctate or without

10 Scutum dull, Scutum not only

11

near

fully

sculpture,

fully granulated,

most

granulated ••••••••••. 12

ugandanum n. sp. very

weakly granulated

and posterior margins •••••••••••••••••••••• l l

lateral

Scutum punctate,

at

not

10.

granulated ••••

without

sculpture among punctures,

not

kawandanum n.

sp.

granulated •••••••••••••••••••••••

11.

Scutum very weakly punctate, without sculpture among punctures, but very weakly granulated near lateral and posterior

1 2.

ma rg ins. • • • • • • • • • • • • • • • • • • • • • • • ••

afrum n. sp.

12 Head strongly punctate ••••••••••• 34. mosseli n. sp. Head without sculpture or very weakly punctate ••••••••••••••••• • •••••••••••••••••••••••••••••••• 13 Head Head

fully

granulated,

reticulate

rugose

35.

capei n. sp.

without

areolae or

or with

irregular keels,

irregular keels •••• 14 granulated

among areolae and keels •••••••••••••••••••••••••••••••••••••• 16 14 Scutum fully Scutum not lateral

granulated,

granulated,

dull ••••

at

most

10.

weakly

ugandanum n. sp. granulated

only near

and posterior margins •••••••••••••••••••••••••••••••• 15

15 Scutum punctate, wi thout sculpture among punctures, not granulated ••••••••••••••••••••••• 11. kawandanum n. sp. Scutum very weakly punctate, wi thout sculpture among punctures, weakly granulated near lateral and posterior margins •••••••••••

1 2. afrum n. sp. 18. gutturnium (Benoit)

• • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • •• 16 Scutum dull, strongly granulated. Scutum shiny,

without

sculpture or very

. . . • . . • . . •.•••. . . . • . . . . • . • . • . . . . . • 17 Apex of

the distal

6.

inner process of

apex

of

the gonoforceps

Apex

of

the distal

from

the apex

(Fig.

weakly granulated ••••••

kivuanum (Benoi t )

the gonoforceps nearer

36. emeritum n. of

the

258) •••••••••.•••••••••••••••••••

inner process of

the gonoforceps

sp.

the gonoforceps

(Figs.

218,

246,

farther

259) •••••••••

Anteoninae

353

••••••••.•••••••••••••••••••••••••••••••••••••••••••••••••••• 18 18 Head weakly granulated,

with numerous

•••••••••••••••••••••••••••••••••• Head granulated,

without

19 Head with TL nearly approximately

0,5

length

of

long as OPL;

notaulices

slightly

reaching

scutum . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

. . . . . . . . . . . . . . . . . . . . . . . . . . . • . . . . • 25. medleri n. Head with TL

keels ••••••.•

keels . . . . . . . . . . . . . . . . . . . 19

transversal

twice as

transversal

maritimum (Turner)

3.

longer

than OPL;

sp.

notaulices

reaching

0,3 length of scutum . . . . . . . . . . . . . . . . . . . . . . . . . . . .. • . . • . • • . • • • • . . . • • . . • • • . . . . • . . . . .. 37. garambanum n. sp.

approximately

The males of A. mingoae n. sp., brachypterum n. sp., fiorii n. sp., rufonigrum n. sp., namibiense n. sp., turneri n. sp., natalense n. sp., taylori n. sp., dayi n. sp., canabense (Benoit), anthereon Benoit, hova Benoi t, madagascolum (Benoi t), longiscapum (Ceba Ilos), compressicoxale (Benoit), townesi n. sp., arabicum n. sp., decellei n. sp., kasaicum (Benoit), hoyoi n. sp. are unknown.

1. Anteon brachypterum n. sp.

FEMALE

(Figs.

longer than

212,

tota I

213):

brachypterous;

length of scutum,

fore-wing

very reduced,

scutell um and metanotum:

sl ightly

hi nd-w ing

invisible; length 3,43 mm; black; mandibles testaceous-dark; antennae brown, wi th segments 1-3 testaceous; tegu I ae testaceous; legs testaceous, with coxae and clubs of femora partly brown; following proportions: granulated; TL = 5;

frontal

10:

6:

7:

7:

line present;

7:

=

POL

occipital carina complete;

pronotum

tubercles reaching tegulae; (5: 13), very reduced,

6: 6;

6: OL

antennal 6:

=

8;

3;

segments in

head dull,

OOL

=

5;

OPL

=

4,5;

mandibles with 4 teeth progressing

larger from anterior one to posterior; pronotal

6:

pronotum dull, scutum dull,

hairy,

granulated;

nearly 0,5 as

long as

granulated and reticulate rugose;

scutellum

shiny, small, without sculpture, slightly shorter than scutum (4:5); metanotum short, nearly 0,5 as long as scutellum (2:4); propodeum dull, granulated, with a transversal keel between dorsal and posterior surface; dorsal surface reticulate rugose; posterior surface almost smooth, with some irregular keels;

pleura transversely striate;

following proportions: 7: proximal lamellae;

prominence;

2:

2:

4:

segment 5 of front

apex with a group of nearly 4

rudimentary claw;

LOCUS TYPICUS:

tarsus with lamellae;

no segment of fore tarsi

spurs 1,1,2. MALE: unknown Ghinda

TYPICAL MATERIAL:

(Eritrea)

holotype F!

fore tarsal

segments in

10; enlarged claw with one bristle on a

in HE.

1 row of nearly 20

chela

produced

(Fig.

214)

into a hook;

without

tibial

354

Mem. Amer. En!.

Fig. 212 -

Ins!., no.

37,

1984

Female of Anteon brachypterum n. sp.

HOSTS: unknown DISTRIBUTION: Ethiopian region:

(type)

ETHIOPIA: Ghinda (Eritrea),

2. Anteon agile n. sp.

HE!

Anteoninae

Fig.

213 -

Female of

355

Anteon brachypterum n. sp. (type)

Chelogynus kivuanus Benoit 1951c (only M): 15.

FEMALE: fully winged; length 2,56-3,18 mm; black; mandibles testaceous; antennae brown, with segments 1-2 testaceous; pronotum reddish or brown; legs testaceous, sometimes with hind coxae partly brown; antennae not distally thickened; antenna I segments in following proportions: 10: 6: 9:

10:

8:

8:

8:

8:

8:

11;

head shiny,

sculpture among puncutures, without median and = 5;

OOL = 6;

OPL = 5;

posterior surface tegulae;

length of scutum;

weakly punctate,

carina complete;

pronotum shiny, (14:11);

weakly punctate, smooth,

pronotal

frons

POL = 5;

almost flat,

OL

with

tubercles reaching

without sculpture among

finely punctate,

notaulices incomplete,

without

punctate behind ocelli;

occipital

longer than scutum

scutum shiny,

punctures;

keels;

TL = 7;

pronotum smooth,

punctures; among

lateral

smooth,

more strongly

reaching

without sculpture approximately 0,65

scutellum and metanotum shiny,

smooth,

weakly

punctate, without sculpture among punctures; propodeum reticulate rugose, with a strong transversal keel between dorsal and posterior surface; posterior surface with 2 lateral

areas;

part of radial segments

longitudinal

fore-wing hyaline,

in following proportions: longer than basal

front

(Fig.

enlarged claw front 4

tarsus

lamellae;

median

vein much shorter than proximal

tarsus much tarsus

keels;

215)

9:

(Fig.

215)

proximal

with 2 rows of 28

spurs 1,1,2.

3,5:

12:

bands;

(3:6);

21;

prominence;

lamellae;

fore

distal

tarsal

segment 4 of front

(12:8);

segment 5 of

part shorter than apical

with one bristle on a

tibial

3:

part

part of segment 5

with basal

area as rugose as

without dark transversal

part

(8:13);

segment 5 of

apex with a

group of

356

Mem. Amer.

Fig.

MALE:

fully

winged;

legs testaceous, 10:

10:

length

1,68-2,68 mm;

9:

antennal 10:

10;

sculpture among punctures; 5; OL ; 3; DOL; 7; OPL ;

black;

1984

sp.

(type)

mandibles testaceous; tegu I ae testaceous;

head shiny strongly punctate,

12:

6:

without

frons without median and lateral keels; POL; 3; TL ; 5; ocellar triangle with weak keels

joining ocelli;

occipital

strong I y punctate,

notaulices incomplete,

antennae

segments in following proportions:

shi ny,

carina complete;

scutum,

scutellum and metanotum

wi thout scu I pture among punctures;

reaching approximately 0,65

propodeum reticulate rugose, dorsal

37,

sometimes with hind coxae proximally darkened;

10: 8:

smooth,

no.

somet imes wi th segment 1 testaceous;

not distally thickened; 10:

Inst.,

Chela of Anteon brachypterum n.

214 -

antennae brown,

Ent.

and posterior surface;

length of scutum;

with a strong transversal

keel

between

posterior surface with 2 longitudinal

keels;

median area nearly as rugose as lateral areas; fore-wing hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal

part

(4: 12);

gonoforceps (Fig.

membranous process and with distal 1,1,2. LOCUS TYPICUS:

216)

with a proximal

inner pointed process;

tibial

spurs

Mann's Quelle (Cameroon)

TYPICAL MATERIAL:

holotype F!

and

1 paratype M!

in BM;

4 paratypes (1

357

Anteoninae

Fig.

F,

3 MM)!

HOSTS:

in TV;

Chela of Anteon agile n.

2 paratypes (1

F,

1 M)!

sp.

(type)

in OL.

unknown

DISTRIBUTION: Cameroon), Kalonge

215 -

Ethiopian region:

BM!

(Mt.

SOUTH AFRICA:

Ruwenzori,

(Talia N River,

CAMEROON: Katberg

Albert Nat.

Tshiaberimu,

(E.

Park),

Albert Nat.

Mann's Quelle (Mt. Cape Prov.), TV!

Park),

Oll

BM!

ZAIRE:

Mbulikerere River

OL!

Magera Lake (Albert

Nat. Park), TV! Rweru (Volc. Mik<er,o), TV! Kashwa (Benoit 1951c, under name Chelogynus kivuanus: male); Rutshuru (Benoit 1951c, under name

Chelogynus kivuanus: male); Garamba Nat. Park, TV! NOTES: the supposed males of Ch. kivuanus Benoit (Benoit 1951c) have two longitudinal therefore,

keels on

actually belong

the posterior surface of the propodeum. to another species

(agile).

3. Anteon maritimum (Turner) n. comb.

Chelogynus maritimus Turner 1928: 151. Chelogynus maritimus Turner: Benoit 1951b: 162.

They,

358

Fig.

Mem.

216 -

FEMALE:

winged;

no.

37,

1984

reddish or black,

with mandibles

with segments 8-10 or

with dorsal

surface

with anterior and

posterior lobes of pronotum testaceous;

metanotum,

abdomen black;

head black,

antennae testaceous,

propectus reddish or brown,

pronotum fully

margi ns testaceous; scutellum,

Inst.,

length 2,50-3,25 mm;

(with teeth brown);

9-10 darkened; testaceous;

Ent.

Male genitalia of Anteon agile n. sp. from Mbulikerere River (Zaire): left half removed.

fully

testaceous

Amer.

mesopleura,

legs brown,

metapleura,

with fore coxae,

propodeum,

lateral

scutum, petiole and

fore tibiae and tarsi

testaceous; tegulae testaceous; antennae distally thickened; antennal segments in following proportions: 15: 5: 6: 5: 5: 5: 5: 5: 4: 5; head shiny; keel;

frons with numerous transversal two keels around orbits;

keels and a

longitudinal

vertex without sculpture,

oblique keels connecting posterior ocell i

to occipital

median

with only two

carina;

POL = 7;

OL

= 4;

OOL = 6; OPL = 5; TL = 5; pronotum shiny, hairy, smooth, without sculpture, with a raised transversal carina between anterior and posterior surface;

anterior surface nearly as

surface shorter than scutum smooth,

hairy;

notaulices distinct,

length of scutum; sculpture;

(5:15);

long as OPL

scutum shiny, incomplete,

reticulate rugose,

posterior

reaching approximately 0,3

scutellum and metanotum smooth,

propodeum dull,

(6:5);

without sculpture, shiny,

with a

without

transversal

keel

Anteoninae

Fig.

217 -

Chela of Anteon maritimum Africa)

(Turner)

359

from Mi Inerton

(South

360

Mem.

between

Ent.

dorsal and posterior surface;

longitudinal basal

Amer.

keels;

no.

37,

vein much shorter than

the pterostigma;

proximal

segment 4 of front

part;

1984

posterior surface without

fore-wing with 2 dark transversal

cells and another beneath

segment 4;

Inst.,

bands,

distal

fore tarsal

one on

the

part of radial

segment

1 as

long as

tarsus slinghtly shorter than basal

part of

segment 5; segment 5 with inner side almost straight, without a distinct apical region; basal part of segment 5 longer than apical part; segment 5 (Fig. long,

217)

with nearly 23 lamellae;

longer than segment 5,

segment 3 of front MALE:

fully

testaceous; weakly, with a

tarsus produced into a hook;

winged;

with teeth brown;

enlarged claw

with one bristle on

length 2,00-2,18 mm;

antennae brown,

legs brown,

granulated, longitudinal

with tarsi

with

black;

(Fig.

tibial

very

prominence;

spurs 1,1,2.

mandibles testaceous,

with segment 1 yellow; and fore tibiae light;

and with 2 lateral

tegulae head shiny,

numerous irregular transversal

median keel

217)

a proximal

keels;

frons

keels around orbits;

POL = 6; OL = 3; OOL = 5; OPL = 2; TL = 2; scutum, scutellum and metanotum shiny, smooth, without sculpture; notaulices distinct, incomplete,

reaching approximately 0,3 length of scutum;

a transversal

keel

reticulate rugose; rugose;

between dorsal

propodeum with

and posterior surface;

posterior surface without

longitudinal

areolae of posterior surface smaller than

dorsal keels,

surface reticulate

those of dorsal

surface;

mesopleura with posterior surface partly smooth and shiny; anterior surface dull, with numerous irregular keels; metapleura dull, with numerous irregular keels; fore-wing hyaline, with distal part of radial vein much shorter than proximal part (2:5); gonoforceps (Fig. 218)

Fig.

218 -

Male genitalia of Anteon maritimum (South Africa):

(Turner)

right half removed.

from Milnerton

Anteoninae

361

without dorsal proximal membranous process, with a dorsal distal process; gonoforceps much shorter than penis; tibial spurs 1,1,2. LOCUS TYPICUS:

Milnerton

(South Africa)

\

/

Fig. 219 -

Chela of

Anteon kasaicum (Benoit)(type)

pOinted

362

Mem.

TYPICAL MATERIAL: HOSTS:

Amer.

holotype Fl

Ent.

Inst.,

37,

no.

and 4 paratypes

1984

(2 MM,

2 FF)!

in BM.

unknown

DISTRIBUTION: AFRICA:

Ethiopian region:

Milnerton

ZAIRE:

(Cape Town),

TV!

Garamba Nat.

Ol!

BM!

Park, OLl

SOUTH

Ceres (Cape Province),

BM!

4. Anteon kasaicum (Benoit) n. comb.

Chelogynus kasaicus Benoit 1950: 222.

FEMALE: dark;

fully

length 3,75-5,62 mm;

winged;

black;

mandibles testaceous

with segments 1-4 or 1-6 testaceous;

antennae brown,

tegu I ae

testaceous; pronotum with posterior lobes and lateral margins reddish; abdomen brown-testaceous; legs testaceous, with femora and hind tibiae partly or fully brown,

with junctions,

trochanters and tarsi

light;

antennal segments in following proportions: 24: 8: 12: 8: 7: 7: 8: 7: 7: 10; head dull, granulated, with nUmerous irregular keels, frons with three strong keels, OOl = 12;

one median and

OPl = 8;

with an anterior transversal transversal

two around orbits;

Tl = 11; occipital

keels on

carina complete;

impression,

anterior surface;

granulated,

POL

= 6; Ol = 3;

pronotum dull,

with numerous

posterior surface of pronotum

shorter than scutum (13:18); pronotal tubercles reaching tegulae; scutum shiny, smooth, without sculpture or weakly alutaceous; notaulices almost invisible, reaching approximately 0,2 length of scutum; scutellum convex, without sculpture, smooth, shiny; metanotum shiny and smooth; propodeum reticulate rugose,

wi:h a strong

posterior surface;

posterior surface without

with 2 dark transversal beneath

the pterostigma;

bands, distal

transversal one on

keel

between dorsal

longitudinal

the basal

part of radial

keels;

and

fore-wing

cells and another

vein shorter than

proximal

(8: 13); pterostigma four times as long as broad; fore tarsal segments in following proportions: 7: 5: 6: 14: 39; enlarged claw (Fig. 219) with one bristle on a proximal prominence; segment 5 of front tarsus (Fig. 219) with 2 rows of nearly 34 lamellae, extending continuously to part

the apex;

apex weakly prominent;

a prominent hook; MALE:

segment 2 of front

tarsus produced

into

spurs 1,1,2.

unknown

lOCUS TYPICUS:

Kasai

TYPICAL MATERIAL: HOSTS:

tibial

(Zaire)

holotype F!

in TV.

unknown

DISTRIBUTION:

Ethiopian region:

ZAIRE:

Kasai,

TV!

Ol!

5. Anteon fiorii n. sp.

CAMEROON:

Victoria,

363

Anteoninae

FEMALE:

fully

winged;

petiole black;

length 3,00-3,62 mm;

sometimes ventral

region of thorax black;

with segments 7-10 brown; 6:

6,5:

5:

reddish-testaceous,

with

sometimes brown spots on dorsal surface of propodeum;

5:

5:

line and two

4:

5:

antennal

5:

7;

antennae reddish-testaceous,

segments

head dull,

lateral keels near orbits;

in following proportions:

reticulate rugose, POL = 8;

OL = 5;

13:

with a frontal OOL = 7;

OPL =

7; TL = 7; occipital carina complete; pronotum shiny, weakly granulated, with numerous transversal keels, hairy; pronotal tubercles reaching tegulae;

pronotum humped,

posterior surface (14:7); shiny,

smooth,

granulated;

0,3 length of scutum; humped; with a

with

anterior surface nearly twice as

pronotum shorter than scutum notaul ices incomplete,

scutellum smooth,

metanotum very short, transversal

surface without

keel

du II,

between

longitudinal

with 2 dark transversal

bands,

smooth;

dorsal

keels,

shiny,

(3:12);

reaching approximately

without sculpture,

very

propodeum reticu I ate rugose,

and posterior surface;

smooth,

long as

scutum

without

one on the basal

areolae;

posterior

fore-wing

cells and another

beneath the pterostigma; distal part of radial vein shorter than proximal part (6,5:9); fore tarsal segments in following proportions: 6: 3: 5: 9: 25;

enlarged claw

(Fig.

segment 5 of front apical

region

220)

tarsus

with one bristle on a proximal

(Fig.

distinct from

Port St.

TYPICAL MATERIAL: FF!

in OL;

HOSTS:

Johns

prominence;

inner side curvilinear,

and proximal

region;

with an

basal

part of

segment 5 of front tarsus with 2 rows

tarsus produced into a hook;

tibial

(South Africa)

holotype F!

1 paratype F!

and 2 paratypes FF!

in TW;

1 paratype F!

in BM;

2 paratypes

in AM.

unknown

DISTRIBUTION: (Pondoland), NOTES:

part;

segment 2 of front

spurs 1,1,2. MALE: unknown LOCUS TYPICUS:

with

the medial

segment 5 shorter than apical of 28 lamellae;

220)

Ethiopian region:

BM!

OL!

SOUTH AFRICA:

Magoebaskloof (Tzaneen),

this species is named in honor of the

Port St. TW!

Johns

Durban

late Mr.

(Natal),

AM!

Giorgio Fiori.

6. Atlteon kivuanum (Benoit) n. comb.

Chelogynus kivuanus Benoit 1951c (only F): 13; the male described by Benoit (1951c: 15) is not kivuanum, because the posterior surface of propodeum has 2 in females);

FEMALE:

fully

this male is agile n.

winged;

longitudinal

keels

(absent

sp.

length 2,68-3,00 mm;

black;

antennae black,

with

segments 1-5 brown or testaceous; fore legs brown or testaceous; mid and hind legs brown-black, with tarsi light and coxae partly light; antennae distally thickened;

antenna I segments in following proportions:

18:

7:

9:

364

Mem.

Fig.

220 -

Amer.

Chela of

Ent.

Inst.,

no.

37,

1984

Anteon fiorii n. sp. (type)

Anteoninae

Fig.

221

-

Chela of

365

Anteon kivuanum (Benoit) from Upemba Nat. Park (Zaire)

6:

6:

7:

7:

7:

6 (segment 10 missing

granulated and reticulate rugose;

scutum

(8:20);

incomplete, longitudinal

keels;

very reduced,

smooth,

weakly

granulated;

notaulices

visible only near anterior margin of scutum;

smooth,

keels;

head

frons with two

posterior surface of pronotum shorter than

scutum shiny,

scutellum shiny,

line present;

POL ~ 6; OL ~ 5; OOL ~ 8; OPL ~ 7; TL ~ 7; pronotum dull, granulated, punctate, with

lateral keels near orbits; occipital carina complete; strong transversal

in the only studied female);

frontal

without sculpture;

metanotum shiny,

propodeum with a strong

transversal

keel

with between

dorsal and posterior surface; dorsal surface reticulate rugose; posterior surface without longitudinal keels, reticulate rugose; fore-wing with 2 dark transversal

bands,

one beneath

the pterostigma and another on

the

basal cells; distal part of radial vein shorter than proximal part (4:6); fore tarsal segments in following proportions: 6: 3: 4: 8: 25; segment 5 of front

tarsus

(11:14);

segment 5 of front

enlarged claw

(Fig. (Fig.

tibial

spurs 1,1,2.

MALE:

fully

winged;

antenne brown,

221) 221)

with basal

part shorter than apical

tarsus curvilinear,

part

with two rows of lamellae;

with one bristle on a proximal prominence;

length 2,18-2,37 mm;

with segment 1 testaceous;

black;

mandibles testaceous;

legs testaceous,

wi th part of

hind coxae and part of hind clubs of femora brown; antennal segments in following proportions: 10: 4: 4: 4: 4: 4: 4: 4: 4: 5,5; head shiny, granulated, ~

3;

with weak areolae;

frons with a median keel,

POL

~

without

7;

OL

~

lateral

4;

OOL

keels;

~

5;

OPL

~

2,5;

scutum shiny,

TL

366

Mem.

Fig.

Amer.

Ent.

Inst.,

no.

37,

1984

222 - Male genitalia of Anteon kivuanum (Benoit) from Ceres (South Africa):

smooth,

left half removed.

without sculpture or weakly granulated;

notaulices incomplete,

reaching approximately 0,25 length of scutum; scutellum and metanotum shiny, smooth, without sculpture; propodeum with a transversal keel between dorsal

and posterior surface;

dorsal

surface reticulate rugosc;

posterior surface without longitudinal keels, fully reticulate rugose; areolae of posterior surface smaller than those of dorsal surface; forewing hyaline,

without dark transversal

much shorter than proximal

bands;

distal

part of radial

vein

(3:7); gonoforceps (Fig. 222) without

part

proximal membranous process and without distal

inner process;

tibial

spurs 1,1,2. LOCUS TYPICUS:

Rutshuru

TYPICAL MATERIAL: HOSTS: unknown DISTRIBUTION: River,

Mt.

River,

Tshiaberimu,

OL!

Ceres

ZAIRE:

(Upemba Nat.

Ruwenzori,

Albert Nat.

Albert Nat.

Rutshuru

(Cape Province),

BM!

BM! OL!

in TV. (Kivu),

TV!

TV!

Park),

Mbulikerere River

Park),

TV!

TV!

Mt.

Kakalari

OL!

Park), OL!

Bomboka (Kyandolire, Albert Nat.

(Cape Province), Town),

and 1 paratype F!

Ethiopian region:

Kiamokoto-Kiwakishi

TV!

(Zaire)

holotype F!

OL!

(Bombi (Talia N

Hoyo (Albert Nat.

Park),

TV! TW!

TW!

7. Anteon taylori n. sp.

Park),

SOUTH AFRICA:

Port St. Johns (Pondoland),

Pietermaritzburg,

River

BM!

Somerset East

Kirstenbosch

(Cape

Anteoninae

FEMALE:

fully

winged;

antennae testaceous, testaceous,

length 3 mm;

with hind coxae,

occipital

antennal

legs

segments in following proportions:

head dull,

granulated;

and with 2 lateral

carina complete;

tegulae testaceous;

hind femora and hind tibiae partly brown;

6: 6: 6: 6: 5: 5: 5: 8;

frons with a median keel

reticulate rugose;

keels around orbits;

region between occipital

with 2 oblique keels connecting the

ocelli

mandibles testaceous;

with segments 7-10 brown;

antennae distally thickened; 15: 6:

black;

367

carina and posterior

internal

side of posterior ocelli

to occipital carina; POL = 7; OL = 4; OOL = 7; OPL = 5; TL = 4; pronotum humped, dull, granulated, with numerous transversal keels; posterior surface nearly 0,5 as long as scutum anterior surface (7: 11); shiny,

smooth,

pronotal

granulated;

0,25 length of scutum;

notaulices incomplete,

scutellum shiny,

metanotum with some longitudinal a strong transversal surface without transversal

keel

smooth,

keels;

keels;

shorter than

tegulae;

scutum

reaching approximately

without sculpture;

propodeum reticulate rugose,

between dorsal

longitudinal

keels,

(8:16),

tubercles reaching

and posterior surface;

with

posterior

posterior surface with strong fore-wings with 2 dark

not reticulate rugose;

transversa I bands, one on the basa I cell s and another beneath the pterostigma; distal part of radial vein much shorter than proximal part (3:8);

fore tarsal

segments in following proportions:

segment 5 of front tarsus part

(8:12);

223)

(Fig.

enlarged claw

(Fig.

with basal

223)

5:

Kawanda

TYPICAL MATERIAL: HOSTS:

2,5:

20;

6:

apical

with one bristle on a proximal

prominence; segment 5 of front tarsus with 1 row of 11 vith a group of 8 lamellae; tibial spurs 1,1,2. MALE: unknown LOCUS TYPICUS:

3:

part shorter than lamellae;

apex

(Uganda)

holotype F!

in BM.

unknown

DISTRIBUTION: NOTES:

Ethiopian region:

UGANDA:

Kawanda,

this species is named in honor of Mr.

BM!

T .H.C. Taylor.

8. Anteon dayi n. sp.

FEMALE: fully winged; length 2,37-3,00 mm; black; mandibles testaceous; antennae brown, with segments 1-5 testaceous; legs testaceous, with hind coxae brownish; brown;

sometimes also mid and hind coxae,

antennae distally thickened;

proportions:

6:

5:

5:

4:

4:

4:

weak areolae near clypeus; occipital

carina complete;

pronotum flat, long as scutum

dull,

POL

=

granulated,

(10:14),

5: 7;

5:

7;

head dull,

line present; OL

=

femora and tibiae

segments in following

5;

OOL

granulated,

with

frons without keels;

=

5;

OPL

=

5;

TL

=

4;

with posterior surface more than 0,5 as

longer than anterior surface (10:5);

tubercles reaching tegulae; si des granu I ated;

5:

frontal

antennal

scutum shiny,

notau I ices weak,

smooth,

pronotal

without sculpture,

i nvi s ib Ie or fa int I y

v is ib Ie and

with

368

Mem.

Fig.

incomplete,

with

surface; keels;

Chela of

Ent.

Inst.,

no.

smooth and without sculpture;

a strong transversal

1984

keel

scutellum and

propodeum reticulate

between dorsal

posterior surface weakly reticulate rugose,

fore-wing

37,

Anteon taylori n. sp. (type)

almost reaching mid-length of scutum;

metanotum shiny, rugose,

223 -

Amer.

with 1 dark transversal

and posterior

without

longitudinal

band beneath the pterostigma;

distal part of radial vein much shorter than proximal part (2:6); fore tarsal segments in following proportions: 6: 2: 3: 8: 16; segment 5 of front

tarsus with basal

claw

(Fig.

224)

part shorter than apical

with one bristle on a proximal

part

(6:10);

prominence;

enlarged

segment 5 of

Anteoninae

Fig.

front tarsus 7 lamellae; MALE:

(Fig. tibial

224 -

224)

Chela of

369

Anteon dayi n. sp. (type)

with 2 rows of 26 lamellae;

apex with a group of

spurs 1,1,2.

unknown

----------

370

Mem.

LOCUS TYPICUS:

Regenstein

TYPICAL MATERIAL: HOSTS:

Amer.

En!.

Ins!.,

no.

37,

1984

(South West Africa)

holotype F!

and

1 paratype F!

in BM.

unknown

DISTRIBUTION:

Ethiopian region:

SSW Windhoek), NOTES:

BM!

SOUTH WEST AFRICA:

Okahandja,

Regenstein

(15 mls.

BM!

this species is named in honor of Mr.

Michael Day.

9. Anteon canabense (Benoit) n. comb.

Hirtanteon canabensis Benoit 1951b: 164.

FEMALE:

fully

winged;

length 2,75 mm;

reddish-testaceous,

with petiole

black; antennal segments in following proportions: 12: 4: 7: 6: 5: 6: 5: 8; antennae distally thickened; head shiny, granulated and reticulate rugose,

hairy;

frons

with a median

two keels around the orbits;

POL = 7;

4;

pronotum dull,

occipital carina complete;

transversal

keels;

granulated,

(7:14);

pronotal

incomplete,

keel

5:

and with

OPL = 4;

TL =

with numerous

longer than OPL

tubercles reaching tegulae;

reticulate rugose near lateral

notaulices distinct,

OOL = 5:

granulated,

posterior surface of pronotum

shorter than scutum dull,

longitudinal

OL = 5;

6:

(7:4) scutum

and posterior margins;

reaching mid-length of scutum;

scutellum

shiny, smooth, without scul"'cure; metanotum dull, not smooth, with numerous longitudinal keels; propodeum dull, with a transversal keel between dorsal and posterior surface; dorsal surface reticulate rugose; mesopleura and metapleura granulated and with short transversal keels; posterior surface without longitudinal keels, keels only near margins; fore-wing hyal ine, bands, (3:7)

with distal

part of radial

vein much shorter than proximal

(according to Benoit 1953b distal part

1 slightly

longer than segment 4

fully granulated, with some without dark transversal

(7:6);

longer);

fore tarsal

segment 5 of front

tarsus

part

segment (Fig.

225) with basal part shorter than apical part (7:8), with inner side curvilinear and with apical region distinct; segment 4 of front tarsus nearly as long as basal part of segment 5; segment 5 with 1 row of 23 lamellae; apex with a group of 4 lamellae; enlarged claw (Fig. 225) with one bristle on a proximal

prominence;

segment 3 produced into a hook;

tibial spurs 1,1,2. MALE: unknown LOCUS TYP ICUS:

Mundjungani

TYPICAL MATERIAL: HOSTS:

in TV.

unknown

DISTRIBUTION: TV!

(Zaire)

holotype F!

Ethiopian region:

ZAIRE:

Mundjungani

(Likimi,

Ubangi),

Anteon i nae

371

(1\ Ii \

I ,

1

Fig.

225 -

Chela of

\ '

Anteon canabense n. sp. (type)

Mem.

372

Amer.

Ent.

Inst.,

no.

37,

1984

10. Anteon ugandanum n. sp.

FEMALE:

fully

winged;

length 1,87-2,18 mm;

black;

mandibles testaceous;

antennae brown (in specimens from Uganda) or yellow (in specimens from Zaire); legs brown, with tarsi light (in specimens from Uganda) or yellow

(in specimens from Zaire);

segments in following proportions: 4,5;

head dull,

median keel, OL

=

3;

scutum

without

=

OOL

transversal

granulated; 4;

keels;

(5:9);

granulated;

lateral

OPL

=

3;

antennae distally thickened;

antennal

6:

3,5:

3,5:

3,5:

3:

2,5:

frons reticulate rugose, keels; TL

=

occipital

4;

3:

3,5:

with a

carina complete;

pronotum dull,

POL = 5;

granulated,

with some

posterior surface of pronotum more than 0,5 as

pronotal

tubercles reaching tegulae;

notaulices incomplete,

3,5:

longitudinal

scutum dull,

reaching approximately 0,25

long as

smooth, length of

scutum; scutellum and metanotum shiny, smooth, without sculpture; propodeum with a strong transversal keel between dorsal and posterior surface; dorsal surface reticulate rugose; posterior surface without longitudinal keels, with areolae smaller than those of dorsal surface; fore-wing hyaline,

without dark transversal

vein much shorter than proximal following proportions:

5:

2:

2:

3:

part sl ightly shorter than apical with one bristle on (Fig.

226)

tibial

spurs 1,1,2.

a proximal

part 9; part

Fig.

226 -

distal

fore tarsal

segment 5 of front (4:5);

prominence;

with 1 row of 10 lamellae;

bands;

(2:6);

part of radial

segments in tarsus with basal

enlarged claw

(Fig.

segment 5 of front

apex with a group of 4

Chela of Anteon ugandanum n.

sp.

(type)

226)

tarsus lamellae;

373

Anteoninae

Fig.

227 -

Male genitalia of Anteon ugandanum n. sp. from Ameib Farm (South West Africa): right half removed.

MALE:

fully

winged;

antennae fully

length

1,68-1,93 mm;

black;

mandibles testaceous;

testaceous or with segments 3-10 weakly darkened;

testaceous or with coxae,

femora and

tibiae brown;

legs

antennae not distally

thickened; antennal segments in following proportions: 6: 5: 5: 8; head dull, fully granulated; frons with a

9: 5: 5: 6: 5: 6: longitudinal median

keel, without lateral keels; occipital carina complete; POL = 6; OL = 4; OOL = 4; OPL = 2,5; TL = 2,5; scutum dull, granulated, with some weak areolae near anterior margin;

notaulices

incomplete,

reaching

nearly

mid-length of scutum; scutellum and metanotum shiny, smooth, without sculpture; propodeum reticulate rugose, with or without a strong transversal without

keel

between dorsal

longitudinal

surface;

and posterior surface;

posterior surface

keels and with areolae smaller than

fore-wing hyal ine,

without dark transversal

those of dorsal

bands;

distal

part

of radial vein much shorter than proximal part (3:7); gonoforceps (Fig. 227) without proximal membranous process and without distal inner pointed process; tibial spurs 1,1,2. LOCUS TYPICUS: Kawanda (Uganda) TYPICAL MATERIAL: holotype F! and 9 paratypes (1 para types HOSTS:

(7 MM,

in TV;

M,

9 paratypes (5 FF,

8 FF)!

4 MM)!

in BM;

9

in OL.

unknown

DISTRIBUTION: ZAIRE:

2 FF)!

Ethiopian region:

Paulis (Haut-Uele),

(Albert Nat. Albert Nat.

Park),

TV!

TV!

UGANDA:

Kakalari

River

Park),

TV!

OL!

Kiribata

Albert Nat.

Park),

OL!

TV!

Talya River

Park), BM!

TV!

OU

Kawanda

Garamba Nat.

SOUTH WEST AFRICA:

(Bombi

,

Park,

River,

(Moyenne Lume, (May

OL! TV!

BM! OL!

Mt. Mt.

BM!

Hoyo

Ruwenzori, Ruwenzori,

ya Moto River,

Ameib Farm (19 mls.

Serere, Mt.

Albert Nat.

NW Karibib),

374

Mem.

Amer.

Ent.

Inst.,

no.

11. Anteon kawandanum n.

FEMALE:

fully

antennae and thickened;

winged;

length 1,50-2,00 mm;

legs brown,

antennal

sometimes fully

37,

1984

sp.

black;

mandibles testaceous;

testaceous;

antennae distally

segments in following proportions:

6:

3,5:

2:

2:

2:

2:

2: 2: 2,5: 5; head dull, fully granulated; frons with a longitudinal median keel, without lateral keels; occipital carina complete; POL = 4; Ol = 3; OOl = 5; OPl = 3; Tl = 5; pronotum shiny, with anterior surface granulated and posterior surface smooth and without sculpture; posterior surface of pronotum shorter than scutum

(3:8);

pronotal

reaching tegulae;

finely

punctate,

scutum shiny,

sculpture among punctures; I ength of scutum; sculpture;

notaul ices incomplete,

dorsal

longitudinal

without

smooth and wi thout

keel

between

surface reticulate rugose;

keels,

tubercles

nearly reaching mid-

scutell um and metanotum sh i ny,

propodeum with a strong transversal

posterior surface; without

smooth,

dorsal

and

posterior surface

with areolae smaller than

those of dorsal

surface; fore-wing hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part (2:4,5); fore tarsal segments in following proportions: 4,5: tarsus

(Fig.

228)

enlarged claw

with basal

(Fig.

228)

1:

1: 3:

7;

segment 5 of front

part shorter than apical

with one bristle on a

part

proximal

segment 5 of front tarsus with 1 row of 10 lamellae; of 7 lamellae;

tibial

spurs 1,1,2.

Fig.

228 -

Chela of Anteon kawandanum n.

(3:4);

prominence;

apex with a group

sp.

(type)

Anteoninae

Fig.

MALE

Male genitalia of Anteon kawandanum n. (Uganda): left half removed.

229 -

(bred):

legs brown; antennal

fully

winged;

length 1,06-1,43 mm;

mandibles testaceous; smooth,

without sculpture; carina complete;

granulated;

black;

from Kawanda

antennae and

6: 4:

4:

4:

3:

4:

4:

4:

with a more or less wide frontal

frons without median and

POL = 4;

sp.

antennae not distally thickened;

segments in following proportions:

head shiny,

375

Ol = 2;

OOl = 3;

lateral

5:

6;

region

keels; occipital

OPl = 3 Tl = 4;

scutum,

scutellum and metanotum shiny, smooth, weakly punctate, without sculpture among punctures; notaulices incomplete, reaching mid-length of scutum;

propodeum reticulate rugose,

between dorsal longitudinal

and posterior surface;

keels,

fore-wing hyal ine,

with areolae smaller than those of dorsal

vein much shorter than proximal tibial

transversal

part

process,

(2:4);

bands;

distal

gonoforceps

without distal

surface;

part of radial

(Fig.

229)

inner pointed

spurs 1,1,2.

lOCUS TYPICUS:

Kawanda

(Uganda)

TYPICAL MATERIAL: holotype F! and 16 paratypes (11 4 paratypes (3 FF, 1 M)! in OL. HOSTS:

keel

posterior surface without

without dark transversal

without membranous proximal process;

with a strong

FF,

5 MM)!

in BM;

unknown

DISTRIBUTION:

Ethiopian region:

Kawanda,

Ol!

BM!

Serere,

BM!

SOMALIA:

Afgoi

(Benadir),

Ol!

UGANDA:

Ol!

12. Anteon air-um n. sp.

FEMALE:

fully

tegulae and

winged;

legs fully

length 2,00-2,68 mm; yellow-testaceous;

black;

antennae,

mandibles,

antenna I segments in following

376

Mem. Amer. Ent.

Inst.,

no. 37, 1984

Fig. 230 - Chela of Anteon afrum n. sp.

(type)

proportions: 8: 4: 2: 2: 2,5: 2: 2: 3: 2,5: 4,5; head dull, granulated, smooth; frons without median and lateral keels; POL = 4; OL = 3; OOL 4; OPL = 2,5; TL = 4; occipital carina complete; pronotum dull, granulated; pronotal tubercles reaching tegulae; posterior surface of

Anteoninae

377

pronotum

longer than anterior surface and more than 0,5 as

scutellum

(5:7);

scutum granulated,

reaching mid-length of scutum;

propodeum with a

posterior surface;

dorsal

longitudinal

transversal

keel

surface reticulate rugose;

keels,

long as

notaulices incomplete,

scutellum and metanotum shiny,

without sculpture; without

smooth;

not reticulate rugose,

between

smooth,

dorsal

granulated;

fore-wing

hyal ine, without dark transversal bands; distal part of radial vein shorter than proximal part (2:4); fore tarsal segments in following proportions: on a

rows of MALE: and

1,5:

16 lamellae;

fully

2:

3:

prominence;

winged;

legs fully

segments dull,

4:

proximal

length 1,43 mm;

line;

Tl = 4;

frons

which are very

-

7:

5:

5:

5:

keels,

carina complete;

scutum shiny,

scu I pture among punctures,

mandibles,

smooth,

POL

5:

5:

=

5;

5:

tegulae

antenna I 5:

7;

head

incomplete and Ol

=

3;

OOl

=

4,5;

without

latera I and posterior margi ns

notaulices

Male genitalia of Anteon afrum n. (Zaire):

6:

with 2

spurs 1,1,2.

antennae,

weakly punctate,

wi th areas near

weakly granulated;

230)

tibial

thickened;

with an

much

with one bristle

(Fig.

lamellae;

black;

lateral

230)

tarsus

antennae not distally

without

occipital

(Fig.

group of 3

in following proportions:

weak frontal

231

enlarged claw

segment 5 of front

apex with a

testaceous;

granulated;

OPl = 3;

Fig.

9;

and

posterior surface

incomplete,

sp.

left half removed.

reaching nearly

from Garamba Nat.

Park

378

Mem. Amer.

Ent.

Inst.,

no.

37,

1984

mid-length of scutum; scutellum and metanotum shiny, smooth, without sculpture; propodeum reticulate rugose, with a strong transversal keel between dorsal

and posterior surface;

posterior surface without

longitudinal keels, finely reticulate rugose; fore-wing hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part (1,5:5); gonoforceps (Fig. 231) without proximal membranous process and without distal inner pointed process; tibial spurs 1,1,2. LOCUS TYP I CUS: Okahandja (South West Africa) TYPICAL MATERIAL: holotype F! in BM; 1 paratype M! in TV; 1 paratype F! in OL. HOSTS:

unknown

DISTRIBUTION: Ethiopian region:

ZAIRE: Garamba Nat.

SOUTH WEST AFRICA:

BM!

Okahandja,

Park,

TV!

OL!

13. Anteon turneri n. sp.

FEMALE: fully winged; length 3,43 mm; reddish-testaceous, with petiole black, metanotum and dorsal surface of propodeum partly black; first abdominal segment partly black;

antenna I segments

in following

proportions: 10: 5: 6: 5: 5: 5: 5: 5: 4,5: 6; head dull, smooth, granulated; frons with a median longitudinal keel and with 2 lateral keels near orbits; POL = 4; OL = 5; OOL = 5; OPL = 5; TL = 5; occipital carina complete; pronotum shiny, hairy, weakly granulated; with

Fig.

232 -

Chela of Anteon turneri n.

sp.

(type)

Anteoninae

posterior surface much long as scutum shiny,

smooth,

379

longer than anterior surface,

more than 0,5 as

(10:11); pronotal tubercles reaching tegulae; scutum granulated;

notaul ices faintly

margin;

scutellum smooth,

shiny,

smooth,

without sculpture;

propodeum with a

visible near anterior

without sculpture;

metanotum shiny,

transversal

keel

between

dorsal and posterior surface; dorsal surface reticulate rugose; posterior surface without longitudinal keels, not reticulate rugose, granulated; fore-wing hyaline,

without dark

transversal

bands;

distal

part of radial

(2,5:7); fore tarsal segments in following proportions: 6: 2: 3: 7: 16; enlarged claw (Fig. 232) with one bristle on a proximal prominence; segment 5 of front tarsus (Fig. 232) with 2 rows of 30 lamellae; apex with a group of 4 lamellae; tibial spurs 1,1,2. vein much shorter than proximal

MALE:

part

unknown

LOCUS TYPICUS:

Mossel

Bay

(South Africa)

TYPICAL MATERIAL: holotype F! in BM. HOSTS: unknown DISTRIBUTION: Ethiopian region: SOUTH AFRICA: Province), NOTES:

Mossel

Bay

(Cape

BM!

this species is named in honor of Mr.

Rowland E.

14. Anteon townesi n. sp.

Fig.

233 -

Chela of Anteon townesi n.

sp.

Turner.

380

Mem. Amer.

FEMALE:

fully

proportions:

Inst.,

length 3-3,56 mm;

winged;

petiole black;

Ent.

antennae distally

no.

fully

thickened;

1984

37,

reddish-testaceous,

antennal

segments

10: 6: 8: 7: 6: 6: 6: 6: 6: 7; head shiny,

weakly reticulate rugose;

frons with an

with

in following

granulated and

incomplete median keel

and with

2 incomplete lateral keels directed towards antennal sockets; occipital carina complete; POL = 5,5; OL = 5; OOL = 6,5; OPL = 6; TL = 6; pronotum dull,

granulated,

smooth,

with a

weak

transversal

anterior

impression; posterior surface of pronotum nearly as long as scutum; pronotal tubercles reaching tegulae; pronotum flat; scutum shiny, without sculpture,

with weak

tracks of irregular keels on sides;

visible near anterior margin of scutum; smooth and without sculpture; between dorsal

little

propodeum with a strong transversal

and posterior surface;

posterior surface with small

notaul ices

scutellum and metanotum shiny,

dorsal

areolae and

irregular transversal

fore-w i ng with 2 dark transversa I bands,

keel

surface reticulate rugose;

one on

keels;

the basa I cell sand

another beneath the pterostigma; distal part of radial vein shorter than proximal part (4:9) fore tarsal segments in following proportions: 8: 2:

3,5: 10: 20; enlarged claw (Fig. 233) with one bristle on a proximal prominence; segment 5 of front tarsus (Fig. 233) with 2 rows of 40 lamellae; apex with a group of 4 lamellae tibial spurs 1,1,2. MALE:

unknown

LOCUS TYPICUS:

Royal

TYPICAL MATERIAL: HOSTS:

National

Park

in TW;

(South Africa)

1 paratype F!

in 0'-.

unknown

DISTRIBUTION: TW!

Natal

holotype F!

Ethiopian region:

Ken ton-on-Sea,

NOTES:

this species

SOUTH AFRICA:

Royal

Natal

Nat.

Park,

OL! is named

in

honor of Mr.

Henry K.

Townes.

15. Anteon decellei n. sp.

FEMALE:

fully

winged;

length 2,62 mm;

reddish-testaceous,

except for

metanotum and part of propodeum brown; antennae brown, with segments 1-2 testaceous; antennae distally thickened; antennal segments in following proportions:

9:

5:

5:

4:

4:

4:

4:

4

(last 2 segments missing

in

the honly known specimen); head dull, granulated, reticulate rugose only near clypeus; frons with a longitudinal median keel and with 2 lateral keels around orbits;

occipital

4;

pronotum shiny,

OPL = 5;

TL = 5;

some transversal

tubercles rc,6ching

metanotum shiny,

POL = 6;

OL = 4;

weakly granulated,

posterior surface very

OOL = with

long,

(12:10) and longer than anterior surface (12:3);

notaulices incomplete, with a

smooth,

keels on anterior surface;

longer than scutum pronotal

carina complete;

tegulae;

scutum shiny,

smooth,

reaching mid-length of scutum;

smooth,

strong transversal

posterior surface without

without sculpture; keel

between

longitudinal

propodeum reticulate rugose,

dorsal

keels,

granulated;

scutellum and

and posterior surface;

weakly

reticulate rugose;

381

Anteoninae

Fig.

234 -

Chela of Anteon decellei n.

fore-wing with 2 dark transversal another beneath the pterostigma; than proximal

part

(3,5:7);

bands, distal

fore tarsal

one on

sp.

the basal

part of radial segments

(type)

cells and

vein much shorter

in following proportions:

6: 2: 2,5: 8: 18; segment 5 front tarsus with basal part shorter than apical part (8: 10); enlarged claw (Fig. 234) with one bristle on a proximal

prominence;

29 lamellae; MALE:

tarsus

(Fig. tibial

234)

with 2 rows of

spurs 1,1,2.

unknown

LOCUS TYPICUS:

Garamba National

TYPICAL MATERIAL: HOSTS:

segment 5 of front

apex with a group of 3 lamellae;

holotype F!

Park

(Zaire)

in TV.

unknown

DISTRIBUTION: Ethiopian region: ZAIRE: Garamba Nat. Park, NOTES: this species is named in honor of Mr. Jean Decelle.

16. Anteon anthereon Benoit

TV!

382

Mem. Amer.

Anteon anthereon

FEMALE: black;

fully

winged;

antennal

Ent.

Inst.,

Benoit 1951b:

no.

37,

1984

160.

length 2,68 mm;

fully

testaceous,

segments in following proportions:

10:

with petiole

5:

5:

5:

5:

5:

5:

5: 5: 7; antennae distally thickened; head shiny, hairy, fully weakly reticulate rugose; POL = 5; OL = 4; OOl = 4; OPL = 4; TL = 2; frons with a weak longitudinal median keel, without lateral keels; pronotum shi ny, weak I y rugose, with posterior surface very short, much shorter than scutum

(5:15);

weakly punctate;

pronotal

length of scutum; dorsal

keel

between dorsal

surface strongly reticulate rugose; granulated,

incomplete,

scutellum and metanotum shiny,

propodeum with a transversal dull,

tubercles reaching tegulae;

notaulices weakly visible,

scutum shiny, reaching mid-

without sculpture; and posterior surface;

mesopleura and metapleura

weakly reticulate rugose;

posterior surface of

propodeum without longitudinal keels, granulated; fore-wing with a faint preapical dark transversal band; distal part of radial vein much shorter

Fig.

235 -

Chela of

Anteon anthereon

Benoit

(type)

Anteoninae

than proximal segment 4

part

(9:2,5);

(3: 10); fore tarsal segment 5 of front

almost rectilinear and with 4 basal

part of segment 5;

segment 1 much tarsus

lamellae;

(Fig.

longer than

235)

with

inner side

segment 4 nearly 0,5 as

enlarged claw

long as

with 1 bristle on a proximal

segment 4 of front tarsus produced into a

prominence; 1,1,2. MALE:

383

hook;

tibial

spurs

unknown

LOCUS TYPICUS: Port St. Johns (South Africa) TYPICAL MATERIAL: holotype F! in BM. HOSTS:

unknown

DISTRIBUTION: (Pondoland),

Ethiopian region:

SOUTH AFRICA:

Port St.

Johns

BM!

17. Anteon namibiense n. sp.

FEMALE:

fully

winged;

length 2-2,12 mm;

testaceous;

an tennae brown,

testaceous;

mid and hind

proportions: granulated; lateral

11:

4:

3:

frons with a

legs brown; 3,5: 3,5:

antennal

3,5:

mandibles and

1 tes taceous;

3,5:

fore

granu I a ted,

longitudinal median keel

POL; 8;

OL ;

4;

OOL ;

5;

4,5;

head dull,

and with

OPL ;

with posterior surface very short,

pronotal

Fig.

tubercles reaching tegulae;

236 -

tegulae

legs

segments in following

3,5:

keels near orbits directed towards antenna I sockets;

carina complete; (2:15);

4:

black;

with segmen t

3;

TL ;

tracks of 2 occipital 2;

pronotum

much shorter than scutum

scutum dull,

Chela of Anteon namibiense n.

sp.

granulated;

(type)

384

Mem.

notaul ices very short,

Amer.

Ent.

keel

37,

1984

without sculpture;

and posterior surface;

longitudinal

keels, bands,

granulated, one on

distal

sl ightly rugose;

the basal

part of radial

propodeum with

dorsal

posterior surface not reticulate rugose,

transversal pterostigma;

shiny,

between dorsal

reticulate rugose;

no.

faintly visible near anterior margin of scutum;

scutellum and metanotum smooth, a transversal

I nst.,

surface

without

fore-wing

with 2 dark

cells and another beneath

the

vein much shorter than proximal

part

(2:6); fore tarsal segments in following proportions: 6,5: 1,5: 1: 1,5: 5,5; enlarged claw (Fig. 236) with one bristle on a proximal prominence; segment 5 of front tibial

tarsus

(Fig.

236)

with 4

lamellae and some bristles;

spurs 1,1,2.

MALE:

unknown

LOCUS TYPICUS:

Aus

(South West Africa)

TYPICAL MATERIAL:

holotype F!

and 1 paratype F!

in BM;

1 para type F!

in OL. HOSTS: unknown DISTRIBUTION: Ethiopian region: AFRICA:

Glen

(0.

F. S.),

SOUTH WEST AFRICA: Aus,

BM!

SOUTH

BM!

18. Anteon gutturnium (Benoi t) n. comb.

Xenanteon gutturnium Benoit 1951b: 162. Xenanteon gutturnium Benoit ssp. rutshuricum Benoit 1951c: 13 (n. syn.); locus typicus Rutshuru (Zaire).

FEMALE: yellow

fully

(with

winged;

occasionally 5:

5:

6:

granulated, keel,

head black,

6:

light;

6:

6:

9;

tegulae testaceous;

antenna I segments in following proportions: antennae distally thickened;

fully reticulate rugose;

without

with mandibles

antennae brown with segment 1 testaceous;

with segments 1-3 or 1-5 testaceous;

brown with tarsi 5:

length 2,37-3,12 mm;

teeth brown);

lateral keels;

frons with a

POL ~ 8;

OL ~ 5;

head hairy,

longitudinal

OOL ~ 7;

legs 17: 4:

shiny,

median

OPL ~ 5;

TL ~ 4;

pronotum rugose, granu I a ted, wi th posterior surface very short, much shorter than scutum (4: 14); pronotal tubercles reaching tegulae; scutum shiny,

granulated,

scutellum shiny,

with weak

irregular keels;

weakly punctate,

smooth;

propodeum wi th a transversa I keel

notaulices invisible;

metanotum strongly rugose;

between dorsal

dorsal

surface reticulate rugose;

keels,

granulated and with numerous transversal

metapleura granulated, dark bands,

and posterior surface;

posterior surface without

with numerous transversal

keels; keels;

one beneath the pterostigma and another on

longitudinal

mesopleura and fore-wing with 2 the basal

cells

(only the band beneath the pterostigma in specimens from Uganda); distal part of radial vein much shorter than proximal part (2:8); fore tarsal segment 1 much longer than segment 4

(8:3);

segment 4 of front

tarsus

Anteoninae

Fig.

237 -

Chela of Anteon gutturnium Africa)

(Benoit)

385

from Mossel

Bay

(South

386

Mem.

produced into a lamellae;

hook;

Amer.

Inst.,

segment 5 of front

(Fig.

tibial

spurs 1,1,2.

MALE

(not reared):

237) fully

winged;

with

testaceous;

tegulae testaceous;

hind femora,

teeth brown;

tarsus

legs yellow, head dull,

frons with a short

keels around orbits;

scutum dull,

granulated;

POL

(Fig.

237)

black;

with hind coxae,

longitudinal

mandibles clubs of

granulated and weakly median keel,

incomplete,

without

dorsal keels,

3·,

reaching

scutellum and metanotum shiny,

propodeum wi th a transversal

and posterior surface;

prominence;

wi th segments 1-2

longitudinal

notaulices distinct,

wi thout scu I pture;

with 10

part of segment 5;

= 6; OL = 3; OOL = 5; OPL = 4; TL

approximately 0,3 length of scutum;

surface without

1984

length 2,18-2,50 mm;

antennae brown,

hind tibiae brown;

reticulate rugose;

dorsal

37,

with one bristle on a proximal

testaceous,

smooth,

no.

segment 4 nearly 0,5 as long as basal

enlarged claw

lateral

En!.

keel

between

surface reticulate rugose; reticulate rugose,

posterior

with areolae smaller

than those of dorsal surface; mesopleura and metapleura granulated, with irregular keels and areolae; fore-wing hyal ine, with distal part of radial vein much shorter than proximal part; gonoforceps (Fig. 238) with dorsal proximal

membranous process,

gonoforceps nearly as LOCUS TYPICUS:

Mossel

TYPICAL MATERIAL:

Fig.

238 -

without distal

long as penis; Bay

tibial

inner pointed process; spurs 1,1,2.

(South Africa)

holotype F!,

allotype M!

and 1 paratype F!

Male genitalia of Anteon gutturnium (South Africa):

(Benoit)

left half removed.

of X.

from Katberg

387

Anteoninae

gutturnium in BM; 1 paratype F! of X. gutturnium in TV; holotype F! of X. gutturnium rutshuricum in TV. HOSTS: unknown DISTRIBUTION: Ethiopian region:

UGANDA:

BM!

Beach,

Bushenyi,

BM!

KENIA:

Diani

Kawanda, BM!

ZAIRE:

Ol!

BM!

Kampala,

Rutshuru

(Kivu),

TV!

Ikonongi Cave (Mt. Hoyo, Albert Nat. Park), Ol! TV! SOUTH AFRICA: Grahamstown, TW! Hluhluwe Game Res., Ol! TW! Kenton-on-Sea, TW! Ol! Mossel

Bay

Katberg

(Cape Province),

(E Cape Province),

BM!

TV!

Port St.

Johns

(Pondoland),

BM!

BM!

19. Anteon arabicum n. sp.

FEMALE:

fully

winged;

length 2,43 mm;

black;

mandibles testaceous;

antennae black, with segment 1 brown; legs black, with tarsi brown; antennae distally thickened; antennal segments in following proportions: 10: 4: 3,5: 4: 3: 4: 4: 4: 4: 5; head dull, granulated, with weak irregular short keels; frons with a longitudinal median keel and with 2 lateral

incomplete keels near orbits directed towards antenna I sockets;

occipital

carina complete;

Fig.

239 -

POL = 7;

Chela of

Ol = 4;

OOl = 4;

OPl = 3;

Anteon arabicum n. sp. (type)

Tl = 3,5;

388

Mem.

pronotum dull,

Amer.

granulated,

Ent.

longitudinal

37,

1984

keels,

shorter than anterior surface;

surface also shorter than scutum scutum shiny,

no.

with numerous transversal

posterior surface very short, tegulae;

Inst.,

(5:14);

pronotal

very weakly alutaceous,

weak keels or furrows;

notaulices

with posterior

tubercles reaching

with numerous invisible;

scutellum and

metanotum shiny, smooth, without sculpture; propodeum reticulate rugose, with a strong transversal keel between dorsal and posterior surface; posterior surface without dark transversal proximal 1,5:

part

1,5:

apical

bands;

(2:8);

1,5: 7;

and some bristles; MALE:

fore tarsal

tarsus with basal

(Fig.

239)

segment 5 of front

tibial

without

vein much shorter than

segments in following proportions:

enlarged claw

prominence;

fore-wing hyaline,

part of radial

segment 5 of front

part (5:2);

proximal

longitudinal keels; distal

6:

part longer than

with one bristle on a

tarsus

(Fig.

239)

with 4

lamellae

spurs 1,1,2.

unknown

LOCUS TYPICUS: Jebel Jihaf (Southern Yemen) TYPICAL MATERIAL: holotype F! in BM. HOSTS: unknown DISTRIBUTION: Ethiopian Jihaf (Aden),

Palaearctic region:

SOUTHERN YEMEN:

Jebel

BM!

20. Anteon natalense n.

sp.

FEMALE: fully winged; length 2,56-2,81 mm; black; mandibles and legs testaceous; antennae brown, with segment 1 testaceous; antenna I segments in following proportions:

13:

4,5:

4:

reticulate rugose;

occipital

OPL = 4;

pronotum rugose,

short,

TL = 3;

shorter than scutum

scutum shiny, longitudinal smooth,

(5:16);

reticulate rugose,

keels;

4:

shiny,

5:

pronotal

5:

5:

5:

POL = 8;

7;

head shiny,

OL = 5;

OOL = 5;

with posterior surface very tubercles reaching

tegulae;

with or without numerous strong

notaulices invisible;

without sculpture;

4:

carina complete;

scutum and metanotum shiny,

propodeum with a strong transversal

keel

between dorsal and posterior surface; posterior surface without longitudinal keels and with areolae smaller than those of dorsal

surface;

fore-wing hyal ine, without dark transversal bands or spots; distal of radial vein much shorter than proximal part (3:9); fore tarsal segments in following proportions: 7: tarsus with basal tarsus

less than 0,5 as long as basal

enlarged claw

(Fig.

segment 5 of front tibial MALE:

1,5: 2:

2,5: 8;

part much longer than apical 240)

tarsus

part;

segment 5 of front segment 4 of front

part of segment 5

with one bristle on a proximal (Fig.

240)

with 8

part

(2,5:7); prominence;

lamellae and some bristles;

spurs 1,1,2. unknown

LOCUS TYPICUS: Van Reenen (South Africa) TYPICAL MATERIAL: holotype F! in BM; 1 paratype F!

in TV;

1 paratype

Anteoninae

Fig. 240 -

Chela of

389

Anteon natalense n. sp. (type)

F! in OL. HOSTS: unknown DISTRIBUTION: Ethiopian region: ZAIRE: Garamba Nat. SOUTH AFRICA: Van Reenen (Drakensberg, Natal), BM!

Park,

OL!

TV!

390

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

21. 11nteon hoyoi n. sp.

FEMALE:

fully

winged;

antennae brown,

length 2,50 mm;

black;

mandibles testaceous;

with segments 1-2 testaceous;

legs fully

with hind coxae and hind femora partly brown; thichened;

antenna I segments

testaceous or

antennae distally

in following proportions:

12:

5:

7:

5:

5:

6'

6: 6: 6: 8; head shiny, reticulate rugose, 'Nith a longitudinal median keel and with 2 lateral keels around orbits; occipital carina complete; POL

~

5;

rugose,

Ol

~

3;

OOl

~

5;

~

OPl

4;

posterior surface shorter than scutum tegulae;

Tl

with posterior surface nearly as scutum shiny,

punctures;

smooth,

notaulices faintly

transversal

keel

2;

pronotum dull,

reticulate

long as anterior surface

(7:15);

pronotal

weakly punctate,

tubercles reaching

without sculpture among

visible near anterior margin of scutum;

scutell um and metanotum sh i ny, sculpture among punctures;

~

smooth,

weak I y

punctate,

propodeum reticulate rugose,

between dorsal

and posterior surface;

without with a strong

posterior surface

without longitudinal keels; fore-wing hyal ine, without dark transversal bands; distal part of radial vein much shorter than proximal part (2:9); fore tarsal

segments

Fig.

241

in following proportions:

-

Chela of

10:

1,5:

2:

3:

Anteon hoyoi n. sp. (type)

8;

segment 5

Anteoninae

of front

tarsus with basal

enlarged claw

(Fig.

segment 5 of front lamellae; MALE:

tibial

241)

tarsus

391

part much longer than apical with one bristle on a proximal (Fig.

241)

part

(7:1);

prominence;

with some bristles and 4 apical

spurs 1,1,2.

unknown

LOCUS TYPICUS: ,
(1< i vu),

1 paratype F!

ZAIRE:

Kabambi

Cave

in OL. (Mt.

Hoyo),

TV!

OL!

22. Anteon mingoae n. sp.

FE\\ALE:

fully

winged;

length 1,87 mm;

black;

mandibles testaceous;

antennae testaceous, with segments 6-10 brown; legs testaceous, with hind coxae partly brown; antennae distally thickened; antennal segments in following proportions: reticulate rugose;

Fig.

10: 4:

3:

3:

3:

4:

4:

frons with a median keel

242 -

3,5:

4:

5;

head shiny,

and 2 lateral

Chela of Anteon mingoae n.

sp.

keels near

(type)

392

Mem.

Amer.

Ent.

Inst.,

orbits directed towards antenna I sockets; 4;

OL = 3;

OOL = 6;

OPL = 3;

TL = 3;

no.

37,

occipital

smooth and without sculpture;

shorter than scutum

(3: 12); pronotal

weakly granulated;

shiny,

smooth,

keel

POL

with posterior

posterior surface

tubercles reaching tegulae;

notaulices invisible;

without sculpture;

strong transversal

carina complete;

pronotum rugose,

surface almost fully shiny,

1984

scutum

scutellum and metanotum

propodeum reticulate rugose,

between dorsal posterior surface;

with a

posterior surface

without longitudinal keels; fore-wing hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part (1 :7); fore tarsal

segments in following proportions:

segment 5 of front (4,5:1,5);

enlarged claw

prominence;

(Fig.

242)

segment 5 of front

some bristles; MALE:

tarsus with basal

tibial

6:

part much

1,5:

1,5:

1,5: 6;

longer than apical

part

with one bristle on a proximal

tarsus

(Fig.

242)

with 6 lamellae and

spurs 1,1,2.

unknown

LOCUS TYPICUS: Ankaratra (Madagascar) TYPICAL MATERIAL: holotype F! in MD. HOSTS: unknown DISTRIBUTION: NOTES:

Ethiopian region:

this species is named

MADAGASCAR:

Ankaratra,

in honor of Mrs.

MD!

Elvira Mingo Perez.

23. Anteon longiscapum (Ceba Ilos) n. sp.

Xenanteon longiscapus Ceballos 1936: 63. Xenanteon longiscapus Ceballos: Benoit 1953b: 417.

FEMALE:

fully

teeth brown; testaceous;

winged;

length 3,75 mm;

antennae testaceous,

legs brown,

proportions:

17: 6:

7:

with tarsi 5:

5:

6:

6:

black;

mandibles testaceous,

with segments 9-10 brown; light; 6:

6:

antennal 11;

with

tegulae

segments in following

antennae distally thickened;

head shiny,

granulated and reticulate rugose;

median keel

and with 2 lateral

frons with a

keels around orbits;

longitudinal

POL = 6;

OL = 5;

OOL = 4; OPL = 6; TL = 4; pronotum shiny, smooth, weakly granulated, without sculpture near posterior margin; posterior surface very long, more than 0,5 as long as scutum tegulae;

scutum shiny,

smooth,

scutellum and metanotum shiny, a transversal

keel

reticulate rugose; granulated, smooth,

smooth,

between dorsal

pronotal

tubercles reaching notaulices invisible;

without sculpture;

and posterior surface;

propodeum with

dorsal

surface

posterior surface with weak track of longitudinal

with numerous weak

without sculpture;

fore-wing with an apical radial

(9:16);

weakly granulated;

transversal

keels;

metapleura with numerous wide dark transversal

vein much shorter than proximal

longer than segment 4 (7:4);

part

irregular keels;

band;

(3:10);

keels,

mesopleura shiny, distal

part of

fore tarsal

segment

segment 4 nearly 0,5 as long as basal

part

Anteoninae

Fig.

of segment 5; (Fi g.

243)

a proximal

243 -

Chela of

I ameli ae;

prominence;

MALE: unknown lOCUS TYPICUS:

tibial

Fanovana

TYPICAL MATERIAL: HOSTS: unknown DISTRIBUTION:

Anteon longiscapum (Ceballos)(type)

segment 4 produced into a

with 22

393

hook;

en I arged cl aw

segment 5 of front

(F i g;

243)

tarsus

wi th one brist Ie on

spurs 1,1,2.

(Madagascar)

holotype F!

in P.

Ethiopian region:

MADAGASCAR:

Fanovana,

P!

24. Anteon madagascolum (Benoit) n. comb.

Xenanteon madagascola Benoit 1953b: 418.

FEMALE:

fully

winged;

wi th teeth brown; brown;

tegulae testaceous;

brown with tarsi 4:

4:

length 2,50-3,00 mm;

4:

4:

5:

5:

light; 5:

7;

long,

segments in following proportions:

antennae distally

thickened;

Tl = 4;

pronotum shiny,

granulated; reaching

notaul ices invisible;

Ol = 4;

scutellum shiny,

occasionally visible:

5:

keel;

OOl = 5;

posterior surface very

(10:13);

approximately 0,3 length of scutum;

longitudinal

POL = 7;

granulated;

more than 0,5 as long as scutum

13:

head shiny,

frons with a median

keels around orbits present or absent;

OPl = 6;

mandibles testaceous,

legs testaceous with hind coxae brown or

antennal

granulated and reticulate rugose; lateral

black;

antennae brown wi th segments 1-6 testaceous or fu II y

smooth,

in this case

scutellum and metanotum

394

shiny,

Mem.

smooth,

Amer.

weakly granulated;

between dorsal

Inst.,

numerous irregular keels;

longitudinal

37,

with

irregular keels;

bands;

distal

part of radial

1984

keel

dorsal

surface reticulate rugose;

keels,

granulated and with

mesopleura granulated;

rugose,

segment

no.

propodeum with a transversal

and posterior surface;

posterior surface without

fore tarsal

Ent.

fore-wing hyaline,

metapleura strongly without dark transversal

vein much shorter than proximal

1 longer than segment 4

(8:3,5);

part

(3:9);

segment 4 of front

tarsus produced into a hook, nearly 0,5 as long as basal part of segment 5; segment 5 (Fig. 244) with 10 lamellae; enlarged claw (Fig. 244) with one bristle on a proximal MALE: unknown LOCUS TYPICUS:

Bekily

TYPICAL MATERIAL:

prominence;

244 -

spurs 1,1,2.

(Madagascar)

holotype F!

HOSTS: unknown DISTRIBUTION: Ethiopian region:

Fig.

tibial

Chela of

in P;

2 paratypes FF in TV.

MADAGASCAR:

Bekily,

Anteon madagascolum

P!

TV!

Ivondro,

(Benoit)(type)

395

Anteoninae

OL!

Ranomafana

(Benoit 1953b);

DIEGO SUAREZ

(Benoit 1953b).

25. Anteon medleri n. sp.

FEMALE: fully winged; length 2,37 mm; black; mandibles testaceous; antennae brown, with segments 1-2 testaceousj tegulae testaceous; legs brown, with tibiae and tarsi testaceous; antennae distally thickened; antennal segments in following proportions: 9: 5: 6: 5: 5: 5: 5: 5: 5: head dull, a

granulated,

longitudinal

complete;

median keel,

POL = 5;

granulated,

with some weak areolae near clypeus;

except for a

without sculpture;

without

Ol = 4; OOl = 5;

lateral

long,

occipital carina

Tl = 6;

narrow posterior region

numerous transversal

surface of pronotum

keels;

OPl = 6;

7;

frons with

which

pronotum dull, is smooth and

keels on anterior half;

more than 0,5 as long as scutum;

posterior

pronota I

tubercles reaching tegulae; scutum dull, granulated; notaul ices invisible; scutellum and metanotum shiny, smooth, without sculpture; propodeum reticulate rugose,

with a strong

posterior surface;

posterior surface without

Fig.

245 -

Chela of

transversal

keel

between dorsal

longitudinal

keels;

Anteon medleri n. sp. (type)

and

fore-wing

396

Mem.

Amer.

Ent.

Ins!.,

no.

37,

1984

hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part (2:5); fore tarsal segments in following proportions: basal

6:

2:

2:

2,5:

8;

segment 5 of front

part longer than apical

one bristle on a row of 8

proximal

lamellae;

part (5:3);

prominence;

apex with a

tarsus

(Fig.

enlarged claw

segment 5 of front

group of 6 lamellae;

245)

(Fig.

with

245)

with

tarsus with

tibial

1

spurs 1,1,2.

MALE: fully winged; length 1,37 mm; black; mandibles and antennae testaceous; legs testaceous, with hind coxae, hind femora and distal region of hind tibiae brownish;

antennae not distally thickened;

segments in following proportions: 4,5:

6;

occipital

head dull,

granulated;

carina complete;

scutum shiny, punctures;

smooth,

246 -

4,5: 4,5:

4,5:

4,5:

4,5:

frons without median and

POL = 4;

OL = 2,5;

weakly punctate,

notaulices incomplete,

scutellum and metanotum shiny,

Fig.

6:

OOL = 4;

antennal

4,5:

lateral

4,5:

keels;

OPL = 2;

TL = 4;

without sculpture among

reaching nearly mid-length of scutum; smooth,

without sculpture;

Male genitalia of Anteon medleri n. left half removed.

sp.

from

propodeum

Ile-ife

(Nigeria):

397

Anteoninae

reticulate rugose,

with a strong median transversal

and posterior surface; wing hyaline,

posterior surface without

without dark transversal

much shorter than proximal proximal

part

membranous process,

(2:4);

bands;

distal

gonoforceps

with distal

NOTES:

th i s species

is named

NIGERIA:

between dorsal keels;

fore-

part of radial

(Fig.

inner process;

1,1,2. LOCUS TYPICUS: Ile-ife (Nigeria) TYPICAL MATERIAL: holotype F! and 1 paratype M! HOSTS: unknown DISTRIBUTION: Ethiopian region:

keel

longitudinal

vein

246)

without

tibial

spurs

in BM.

Ile-ife (W State),

in honor of Mr. J. T.

BM!

Med I er

26. Anteon hova Benoit

Anteon hova Benoi t 1953b: 415.

FEMALE: fully winged; length 3 mm; black; mandibles yellow; antennae yellow; legs yellow; tegulae testaceous; head missing in the only known specimen;

pronotum shiny,

anterior surface; distinct,

incomplete,

metanotum shiny, keel

without sculpture,

scutum shiny,

with

transversal

without sculpture;

reaching mid-length of scutum;

smooth,

between dorsal

smooth,

without sculpture;

and posterior surface;

keels on

notaulices

scutellum and

prnpodeum with a transversal dorsal surface reticulate

rugose; posterior surface with 2 long i tud i na I keel s shorter than scutum (6:12); r·ledian area mostly smooth, with keels only around margins;

Fig.

247 -

Chela of Anteon hova Benoit

(type)

398

Mem. Amer.

lateral

areas reticulate rugose;

posterior surface smooth, hyaline, (2:7); front

with distal

fore tarsal

segment 5;

no.

37,

1984

mesopleura with anterior surface rugose;

without sculpture;

metapleura rugose;

into a hook,

segment 5 (Fig.

247)

one bristle on a proximal

fore-wing

vein much shorter than proximal part

segment 1 longer than segment 4

with numerous bristles;

MALE:

Inst.,

part of radial

tarsus produced

part,

Ent.

nearl yO,S as

without

segment 4 of

lamellae on proximal

apex with 4

prominence;

(7:2);

long as bas a I part of and medial

lamellae; enlarged claw with

tibial

spurs 1,1,2.

unknown

LOCUS TYPICUS:

Bekily

TYPICAL MATERIAL: HOSTS:

(Madagascar)

holotype F!

in Pi

1 para type F

in TV.

unknown

DISTRIBUTION:

Ethiopian region:

MADAGASCAR:

Bekily,

P!

TV!

27. Anteon compressicoxale (Benoi t) n. comb.

Xenanteon compressicoxale Benoit 1953b: 416.

FEMALE:

fully

teeth brown; testaceous;

winged;

legs testaceous,

partly brown; 4:

4:

4:

length 2,68 mm;

antennae brown,

4:

antennal

7;

black;

with

tegulae

with hind coxae and hind clubs of femora

segments in following proportions:

antennae distally thickened;

numerous transversal

mandibles testaceous,

with segments 1-4 testaceous;

irregular keels;

POL

head dull,

=

=

4; Ol

5;

11: 4:

5:

granulated, OOl

=

4;

4:

4:

with

OPl

=

5; Tl = 2; pronotum damaged in holotype, with transversal keels, except for the area near posterior margin which is smooth; scutum shiny, smooth,

weakly punctate;

scutum;

scutellum and metanotum shiny,

notaul ices reaching approximately 0,25

propodeum with a transversal

Fig.

248 -

keel

smooth,

length of

without sculpture;

between dorsal

and posterior surface;

Chela of Anteon compressicoxale

(Benoit)(type)

Anteoninae

399

dorsal

surface reticulate rugose;

posterior surface with 2 longitudinal

keels;

median

lateral

area

metapleura dull,

as rugose as

rugose,

without dark transversal shorter than proximal segment 4 basal

(9:2);

areas;

with transversal bands,

part

with distal

(2:7);

into a

segment 5

(Fig.

and

fore-wing hyal ine,

part of radial

fore tarsal

segment 4 produced

part of segment 5;

mesopleura

keels;

segment

hook,

248)

vein

much

1 longer than

nearly 0,5 as

without

long as

lamellae on median

and proximal parts, with bristles; apex with 6 lamellae; enlarged claw (Fig. 248) with one bristle on a proximal prominence; tibial spurs 1,1,2. MALE:

unknown

LOCUS TYP I CUS:

Bek i I y

TYPICAL MATERIAL: HOSTS:

(Madagascar)

holotype F!

in P.

unknown

DISTRIBUTION:

Ethiopian region:

MADAGASCAR:

Bekily,

P!

28. Anteon inflatrix Benoi t

Anteon inflatrix Benoit 1951b: 161.

FEMALE:

fully

winged;

length 3,00-3,12 mm;

black;

mandibles yellow,

with teeth brown; antennae fully testaceous; occasionally with segments 6-10 fully brown or with dorsal surface brown; anterior margin of pronotum reddish-dark; occasionally

tegulae testaceous;

legs fully

with mid clubs of femora partly brown;

testaceous, hind coxae,

clubs of

hind femora and hind tibiae brown;

antenna I segments in following

proportions:

5:

head dull, median 5;

9:

5:

5:

4:

granulated,

longitudinal

OPL = 5;

4:

keel,

TL = 4;

5:

5:

without

posterior surface nearl y

(5:12);

scutum;

shiny,

6;

lateral

pronotum rugose,

smooth;

antennae distally thickened;

smooth,

keels;

POL = 7;

frons

OL = 4;

with a OOL =

wi th posterior surface sh i ny and

as long as OPL and shorter than scutum without sculpture,

notaul ices invisible occasionally length of scutum;

5:

weakly or strongly reticulate rugose;

incomplete,

weakly punctate;

reaching approximately 0,25

scutellum and metanotum shiny,

smooth,

without

sculpture; propodeum with a transversal keel between dorsal and posterior surface; dorsal surface reticulate rugose; posterior surface with 2 longitudinal

keels;

median area as rugose as lateral

and metapleura rugose; band beneath than proximal (7:2,5);

on medial MALE

the pterostigma; part (3:7);

distal part of radial

and proximal parts;

with one bristle on a (not reared):

fully

segment

tarsus produced

part of segment 5;

mesopleura

segment 5

vein much shorter

1 longer than segment 4

into a hook, (Fig.

249)

apex with 8 lamellae;

proximal winged;

areas;

with a more or less dark transversal

fore tarsal

segment 4 of front

long as basal 249)

fore-wing

prominence;

tibial

length 2,37 mm;

nearly 0,5 as

without

lamellae

enlarged claw spurs 1,1,2.

black;

mandibles

(Fig.

400

Mem. Amer. Ent.

I nst., no. 37,

1984

Fig. 249 - Chela of Anteon inflatrix Benoit from Port St. Johns (South Africa)

Anteoninae

testaceous;

antennae brown,

junctions and proportions:

tarsi

10: 6:

with a median 4; OOl = 6;

401

with segment 1 testaceous;

legs brown,

testaceous;

antennal

segments in following

7:

6:

7;

6:

6:

longitudinal

OPl = 5;

6: keel

Tl = 3;

6:

6:

head shiny,

and without

occipital

lateral

reticulate rugose,

keels;

carina complete;

wi th

POL = 7;

Ol

scutum shiny,

smooth, weakly punctate, without sculpture among punctures; notaul ices incomplete, reaching approximately 0,3 length of scutum; scutellum and metanotum shiny,

smooth,

without sculpture;

propodeum reticulate rugose,

with a transversal keel between dorsal and posterior surface; posterior surface with 2 longitudinal keels; median area as rugose as lateral areas;

fore-wing hyaline,

radial

vein

250)

without dorsal

process;

tibial

lOCUS TYP ICUS: in BM;

proximal

part

bands;

(3:10);

membranous process,

distal

part of

gonoforceps

(Fig.

without distal

inner

spurs 1,1,2. Port St.

TYPICAL MATERIAL: MM)!

without dark transversal

much shorter than proximal

Johns (South Africa)

holotype F!,

allotype M!

and 5 paratypes

(3 FF,

HOSTS: unknown DISTRIBUTION: Ethiopian region: SOUTH AFRICA: Port St. Johns (Pondoland), BM! Jonkershoek (Stellenbosch), TW! Eshowe (Zulu land) , Mi Inerton

Fig.

(Cape Town),

250 -

2

4 paratypes in TV.

OLl

BM!

Male genital ia of Anteon inflatrix Benoit from Port St. (South Africa):

left half removed.

Johns

402

Mem. Amer.

En!.

Inst.,

no.

37,

1984

29. Anteon rufonigrum n. sp.

FEMALE:

fully

winged;

length 4,37 mm;

brown-b I ack spot on vertex; reddish;

scutum,

head reddish,

antennae and

scutellum and metanotum reddish,

propodeum black;

with a

legs testaceous;

wide

pronotum

with brown nuances;

mesopleura and metapleura reddish;

abdomen brown-

testaceous; antennal segments in following proportions: 16: 7: 8: 6: 6: 8: 7: 7: 10; head dull, fully reticulate rugose, with frontal line; POL 10; Ol = 7;

OOl = 6; OPl = 8;

pronotum hairy,

shiny,

shorter than scutum

Tl = 3;

occipital

reticulate rugose;

(8:20);

pronotal

granulated,

fully reticulate rugose;

dull,

granulated,

with some punctures;

propodeum reticulate rugose, posterior surface;

251

keel

fore-wing

band beneath the pterostigma;

Fig.

-

scutum

scutellum

metanotum reticulate rugose; between dorsal

posterior surface with 2 longitudinal areas;

tegulae;

notaulices invisible;

with a transversal

area nearly as rugose as lateral transversal

carina complete;

posterior surface of pronotum

tubercles reaching

dull,

7:

distal

keels;

median

with a dark

part of radial

Chela of Anteon rufonigrum n.

sp.

(type)

vein

and

Anteoninae

much shorter than proximal

part

proportions:

14; enlarged claw

(Fig.

segment 5 of front

tarsus

12: 2,5:

on a proximal

3:

4:

prominence;

bristles and 10 lamellae, part;

(5:11);

403

with basal region much

segment 4 of front tarsus produced

as basal

part of segment 5;

MALE: unknown LOCUS TYPICUS:

Port St.

TYPICAL MATERIAL:

tibial

Johns

segments in following 251)

with one bristle

(Fig.

251)

with some

longer than apical

into a hook,

nearly 0,3 as

long

spurs 1,1,2.

(South Africa)

holotype F!

HOSTS: unknown DISTRIBUTION: Ethiopian region: (Pondoland),

fore tarsal

in BM. SOUTH AFRICA:

Port St.

Johns

BM!

30. Anteon oriphilum Kieffer

Anteon oriphilus Kieffer 1912a: 47. Anteon oriphilus Kieffer: Kieffer 1914b: 166.

FEMALE: MALE:

unknown

fully

teeth brown;

Fig.

252 -

winged;

length 2,25 mmj

antennae,

tegulae and

blackj

mandibles testaceous,

legs fully

testaceous;

Male genital ia of Anteon oriphilum Kieffer removed.

with

head shiny,

(type):

left half

404

Mem. Amer.

strongly punctate, longitudinal

En!.

Inst.,

no.

37,

1984

without sculpture among punctures;

median keel

and with 2 lateral

POL = 5;

antenna I sockets;

OL = 3; OOL = 4;

scutellum and metanotum shiny, sculpture among punctures; length of scutum;

smooth,

very

frons with a

keels directed towards OPL = 2;

TL = 2;

weakly punctate,

notaulices distinct,

propodeum with a transversal

incomplete, keel

scutum, without

reaching

between dorsal

midand

posterior surface; dorsal surface reticulate rugose; posterior surface with 2 longitudinal keels; median area shiny, smooth, with keels only near margins; lateral areas reticulate rugose; mesopleura and metapleura rugose, with posterior surface smooth and without sculpture; fore-wing hyaline, with distal part of radial vein much shorter than proximal part (3:6); gonoforceps (Fig. 252) without dorsal proximal membranous process and without distal inner pointed process; tibial spurs 1,1,2. LOCUS TYPICUS: Mount Sebert (Seychelle Islands) TYPICAL MATERIAL: holotype M! in BM. HOSTS: unknown DISTRIBUTION: Ethiopian region: SEYCHELLE ISLANDS: Mount Sebert (Mahe), BM!

31. Anteon fabulosum n. sp.

FEMALE: MALE:

unknown

fully

testaceous; black;

winged;

length 2,18 mm;

legs testaceous,

black;

mandibles and antennae

with hind coxae basally black;

antennae not distally thickened;

antenna I segments

femora partly in following

proportions: 9: 5: 5: 5,5: 5: 5: 5: 5: 5: 8; head dull, granulated and punctate; frons with a median keel; occipital carina complete; POL = 7; OL = 5;

OOL = 5;

OPL = 4;

TL = 4;

scutum shiny,

smooth,

weakly

punctate, without sculpture among punctures, granulated only near anterior margin: notaulices incomplete, reaching approximately 0,65 length of scutum; scutellum and metanotum shiny, smooth, without sculpture; propodeum reticulate rugose, with a strong transversal keel between dorsa I and posterior surface;

posterior surface wi th 2

longitudinal keels; median area shiny, with central region smooth, with some keels around margins; fore-wing hyaline, without dark transversal bands;

distal

gonoforceps

part of radial

(Fig.

LOCUS TYP I CUS:

253)

Jonkershoek

TYPICAL MATERIAL: HOSTS:

vein shorter than proximal

with a distal

inner process;

part

tibial

(South Africa)

holotype M!

in TW.

unknown

DISTRIBUTION:

Ethiopian region:

(Stellenbosch),

TW!

SOUTH AFRICA:

Jonkershoek

32. Anteon abruptum n. sp.

(3:6,5);

spurs 1,1,2.

405

Anteoninae

FEMALE: MALE:

unknown

fully

wingedj

antennae brown,

coxae partly blackj tibiae brown j

length 2,37-2,62 mmj

antennae not distally

granulated and fully

253 -

9:

5:

6:

6:

6:

thickened; 7:

reticulate rugose;

6:

6:

mandibles testaceous;

legs testaceous,

stalks of hind femora brown;

following proportions:

Fig.

blackj

with segments 1-2 testaceousj

distal

antennal 6:

10;

segments

removed.

n.

sp.

in

head dull,

frons with a short

Male genitalia of Anteon fabulosum

with hind

half of hind

incomplete

(type):

right half

406

Mem.

median keel

Amer.

Ent.

I nst.,

no.

in front of the anterior ocellus;

37,

1984

occipital

carina complete;

POL = 6; Ol = 4; OOl = 7; OPl = 4; Tl = 3; scutum shiny, granulated; posterior region punctate, without sculpture among punctures; notaulices incomplete,

reaching approximately 0,3

metanotum shiny, punctures;

smooth,

length of scutum;

weakly punctate,

propodeum ret i cu I ate rugose,

between dorsal

and posterior surface;

scutellum and

without sculpture among

wi th a strong

transversa I keel

posterior surface of propodeum with

2 longitudinal keels; median area reticulate rugose,

dull,

with areolae

smaller than those of lateral areas; gonoforceps (Fig; 254) without dorsal proximal membranous process and without distal inner pointed process; tibial

spurs 1,1,2.

lOCUS TYPICUS:

Pietermaritzburg

TYPICAL MATERIAL:

holotype M!

(South Africa) and 1 paratype M!

in TW;

1 paratype M!

in Ol. HOSTS:

unknown

DISTRIBUTION: TW!

Ethiopian region:

Hluhluwe Game Res.,

TW!

SOUTH AFRICA:

Pietermaritzburg

(Natal),

Ol!

33. Anteon zairense Benoit

Fig.

254 -

Male genitalia of Anteon abruptum n.

sp.

(type)

Anteoninae

407

Anteon zairense Benoit 1951d: 21.

FEMALE: MALE:

unknown

fully

winged;

teeth brown;

tegulae testaceous; of femora,

length 1,87-2,38 mm;

black;

mandibles yellow,

with

antennae fully brown or with segments 1-3 testaceous; legs brown or testaceous,

hind tibiae brown;

head shiny,

with

hind coxae,

hind clubs

reticulate rugose and

granulated; frons with a longitudinal median keel, without lateral keels; POL ~ 8; OL ~ 4; OOL ~ 6; OPL ~ 4; TL ~ 4; scutum shiny, strongly rugose near anterior marg in, half;

notaul ices distinct,

smooth and weak I y

incomplete,

scutell um and metanotum sh i ny, a transversal

keel

reticulate rugose;

smooth,

between dorsal

posterior surface with 2 longitudinal

area as rugose as lateral

255)

areas,

with numerous keels;

without dorsal

LOCUS TYPICUS:

255 -

part

(5:10);

propodeum wi th dorsal

keels;

surface

median

mesopleura and with distal

gonoforceps

part of (Fig.

proximal membranous process and without distal

with apex very

narrow;

tibial

inner

spurs 1,1,2.

Basoko (Zaire)

TYPICAL MATERIAL:

Fig.

fore-wing hyaline,

vein much shorter than proximal

pointed process,

weak I y punctate;

and posterior surface;

metapleura fully reticulate rugose; radial

punctate on posterior

reaching mid-length of scutum;

holotype M!

in TV.

Male genitalia of Anteon zairense Benoit removed.

(type):

left half

408

Mem. Amer.

HOSTS:

Ent.

Inst.,

no.

37,

1984

unknown

DISTRIBUTION: Jonkershoek

Ethiopian region:

(Stellenbosch),

OL!

ZAIRE: TW!

Basoko,

TV!

Grahamstown,

SOUTH AFRICA:

TW!

34. Anteon mosseli n. sp.

FEMALE:

unknown

MALE: fully winged; length 2,25-3,43 mm; black; mandibles and tegulae testaceous; antennal segments in following proportions: 8: 5: 10: 9: 9: 9: 9:

9:

9:

punctate, OPL

9;

antennae not distally thickened;

without sculpture among punctures;

head shiny, POL = 8;

strongly

OL = 4;

OOL = 7;

= 3; TL = 5; occipital carina complete; scutum shiny, strongly

punctate,

without distinct sculpture among punctures;

incomplete, reaching mid-length of scutum;

Fig.

256 -

Male genitalia of Anteon mosseli n. removed.

notaulices

scutellum and metanotum

sp.

(type):

right half

409

Anteoninae

smooth,

shiny,

without sculpture;

without transversal hyal ine,

keel

propodeum fully

between

dorsal

without dark transversal

shorter than proximal

part

(3:9);

membranous process and without

bands;

distal

gonoforceps

distal

reticulate rugose,

and posterior surface; part of radial

(Fig.

256)

fore-wing vein much

without proximal

inner pointed process;

tibial

spurs

1,1,2. lOCUS TYPICUS: Mossel Bay (South Africa) TYPICAL MATERIAL: holotype M! in BM; 10 paratypes MM!

in TW;

4

para types MM! in Ol. HOSTS: unknown DISTRIBUTION: Province),

Ethiopian region:

BM!

Jonkershoek

SOUTH AFRICA:

(Stellenbosch),

TW!

35. Anteon capei n.

FEMALE: MALE:

winged;

antennae brown;

Fig.

Bay

(Cape

sp.

unknown

fully

segments

Mossel Ol!

length 3,12 mm;

legs brown;

in following proportions:

257 -

black;

mandibles testaceous;

antennae not distally thickened;

Male genitalia of

8:

5:

10:

Anteon capei removed.

11,5:

n.

11,5:

sp.

11,5:

(type):

antennal 10:

10: 9:

right half

410

Mem.

9; head shiny, punctures; furrow;

En!.

Inst.,

weakly punctate,

no.

= 7;

= 4;

OL

scutum shiny,

punctures;

= 6;

OOL

smooth,

smooth,

= 3;

OPL

keels,

1984

with a weak median

= 5;

TL

weakly punctate,

notaulices incomplete,

and metanotum shiny,

37,

without sculpture among

frons without median and lateral

POL

complete;

smooth,

Amer.

occipital

carina

without sculpture among

reaching mid-length of scutum;

weakly

punctate,

scutellum

without sculpture among

punctures; propodeum reticulate rugose, without a transversal keel between dorsal and posterior surface; posterior surface without longitudinal keels; fore-wing hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part (3:9); gonoforceps distal

(Fig.

257)

inner process;

without proximal tibial

LOCUS TYPICUS: Matjesfontein TYPICAL MATERIAL:

(South Africa)

holotype M!

HOSTS: unknown DISTRIBUTION: Ethiopian region: Province),

membranous process and without

spurs 1,1,2. in BM. SOUTH AFRICA:

Matjesfontein

(Cape

BM!

36. Anteon emeritum n. sp.

FEMALE: MALE:

unknown

fully

winged;

length 1,87 mm;

antenna I segments 1-5 testaceous

black;

mandibles,

tegulae and

(other segments missing

in

the only

(

, \

/

j../

/1

\

I

\ I

Fig.

258 -

Male genitalia of Anteon emeritum n. removed.

sp.

(type):

left half

Anteoninae

known specimen); tibiae brown;

legs testaceous,

antennal keels;

with hind coxae,

hind femora and hind

segments in following proportions: 7:

(other segments missing irregular weak

411

in

the holotype);

occipital

head granulated,

carina complete;

4:

Tl ;

punctures;

without

lateral

keels,

frons with a

notaulices incomplete,

3; OOl ;

4:

4

longitudinal

2; 3; scutum shiny, smooth, finely punctate, without sculpture among

median keel,

POL; 6; Ol ;

4:

with numerous

5; OPl ;

reaching approximately 0,25 length of

scutum; scutellum and metanotum shiny, smooth, without sculpture; propodeum reticulate rugose, with a strong transversal keel between aorsal

and posterior surface;

with areolae smaller than without dark transversal than proximal

part

posterior surface without

those of dorsal bands;

distal

tibial

longitudinal

keels,

fore-wing hyaline,

part of radial

vein much shorter

(2:6); gonoforceps (Fig. 258) with dorsal proximal

membranous process and with a distal apex;

surface;

inner pointed process very

near

spurs 1,1,2.

lOCUS TYPICUS:

Okahandja

TYPICAL MATERIAL:

(South West Africa)

holotype M!

HOSTS: unknown DISTRIBUTION: Ethiopian region:

in BM. SOUTH WEST AFRICA:

Okahandja,

BM!

37. Anteon garambanum n. sp.

FEMALE: MALE:

Fig.

unknown

fully

259 -

winged;

length 1,62 mm;

black;

mandibles testaceous;

Male genitalia of Anteon garambanum n. removed.

sp.

(type):

left half

412

Mem.

Amer.

Ent.

Ins!.,

no.

antennae brown,

with segment 1 testaceous,

and hind tibiae,

hind coxae partly brown;

37,

1984

with clubs of femora,

mid

antennae not distally

thickened; antennal segments in following proportions: 7: 4: 3: 3,5: 3: 3,5: 3,5: 3,5: 3,5: 6; head dull, fully granulated; frons with median keel and without lateral keels; occipital carina complete; POL =5; OL = 3; OOL = 4; OPL = 1,5; TL = 2; scutum shiny, smooth, without sculpture; notaul ices incomplete, reaching approximately 0,3 length of scutum; scutellum and me\anotum shiny, smooth and without sculpture; propodeum reticulate rugose, with a strong transversal keel between dorsal and posterior surface;

posterior surface without

longitudinal

keels;

fore-wing

hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part (1 :4); gonoforceps (Fig. 259) without proximal membranous process and with a distal

inner pointed process;

tibial

spurs

1,1,2. LOCUS TYPICUS:

Garamba National

TYPICAL MATERIAL: HOSTS:

holotype M!

Park

(Zaire)

in TV.

unknown

DISTRIBUTION:

Ethiopian region:

ZAIRE:

Garamba Nat.

ORIENTAL SPECIES OF

Twenty

Park,

TV!

ANTEON

two species are known.

KEY TO THE SPECIES

FEMALES

Segment

4 of front

tarsus at

most

0,5 as

long as

segment

1

(Fig. 260) •••••.••••••••••••••••••••••••••••••••••••••••••••• 2 Segment 4 of front tarsus slightly shorter, as long as, or longer

than

1

segment

277) ••••••••.•••••••••••••••••••• 7 longitudinal keels ••••• 1. borneanum n. sp.

(Fig.

2

Posterior surface of propodeum without

3

Posterior surface of propodeum wi th two longi tudinal keels ••• 3 Head fully strongly reticulate rugose •••••••.•••••••••••••••• 4 Head at

most

reticulate

4

wi th

rugose;

irregular keels or wi th never fully

Scutum sculptured by

longitudinal

weakly punctate ••••••••• Scutum and

only vertex rugose ••••••••••••• 5

reticulate keels;

scutellum smooth,

22. philippinum n.

sp.

scutellum strongly punctate wi th very wide

impressions,

similar

to areolae,

without

. . . . . . . . . . . . . . . . . . . . . . . . . . 2. thai n.

sp.

longitudinal

keels •••

413

Anteoninae

5

Posterior lateral

surface of propodeum wi th median area as

3.

areas . . . . . . . . . . . . . . . . . . . . . . . . .

Posterior

rugose as

silvicolum n. sp.

surface of propodeum with median area

smooth,

not

rugose ••••••••••••••••••••••••••••••••••••.•••••••••••••••••••• 6

6

Head punctate,

without

sculpture among punctures ••••••••••••••••

· . . . . . . . . . . . . . . . . . . . . • . . . . . . . . . . . . . . •. 4. indicum n. Head punctate,

weakly granulated and

with

weak

sp.

irregular keels •.

· . . . . . . • . . . . . . . . . . . . . . . . . . • . . . . . . . . • .. 5. mysorense n. 7

Posterior surface of propodeum without Posterior surface of propodeum wi th

8

Head granulated, not for

the frontal

rugose,

line and

two

without

to

9

rugose,

the

not

two frontal

frontal

Segment

of

granulated,

line and

front

two

lateral

lateral

tarsus nearly as

Segment

of

front

scutum;

head not

keels .••• 11

6.

except

keels near orbits ..

yasumatsui n. sp.

wi th areolae or keels

(in addition

keels near orbits) . . . . . . . . . 9 long as

• • . • • . . • . . • . . . • . • . . • . . . . • . . . . • . . . . • . .. 10 Notaulices shorter,

longi tudinal

areolae and keels,

• . . . . . . . . . • . . . • . . • . . . • . • • . . . • . . • . . • . •• Head

sp. keels . . . . . . 8

longitudinal

7.

segment

4 . . . . . . . . . ..

sarawaki n. sp.

tarsus much shorter than segment 4 •••••••.• 10 faintly visible near anterior margin of

reticulate

rugose on

sides of

ocellar

triangle

bengalense n. sp. Notaulices longer, reaching approximately 0,5 length of scutum; head fully reticulate rugose . . . . . . . . . . 9. krombeini n. sp. · . . . . . . . . . . . • . . . . . . • . . • . . . • . . • • . . . . • ..

8.

11 Head smooth, more or less punctate, wi thout sculpture among punctures,

not

reticulate

rugose . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 12

Head rugose, fully reticulate 12 Head with OPL slightly longer

rugose •••••••••••••••••••••••••• 14 than breadth of posterior ocelli ..

13. Javanus n. Head wi th OPL nearly

twice as

long as breadth

of

sp.

posterior

oce I Ii ••••••••••••••••••••••••••••••••••••••••.••.••.••••••••• 12

13 Head with OL slightly longer than POL; species fully testaceous, with petiole black . . . . . . . . . . . . . . . . . . . 14. munroei n. sp. Head wi th OL nearly

twice as

long as POL;

species almost

fully

black •••••••••••••••••••••• '" ••••••• 15. lankanum n. sp. 14 Fore-wi ng hya line, wi thou t dark transversa I bands . . . . . . . . . . . . . . .

• • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • •• 10. muiri n. sp. 1-2 dark transversa I bands . . . . . . . . . . . . . . . . . . . . . 15

Fore-wi ng wi th

15 Pronotum fully smooth, without sculpture or weakly punctate . . . .. • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • . •• 11. laotianum n. sp. Dorsal

surface of pronotum strongly

• ••.•..••.•••••.••••••••••••••••.•. "

sp.,

reticulate

12.

rugose •••••••••••

fyanense n.

sp.

The females of A. nemorale n. sp., silvestre n. sp., expolitum n. peterseni n. sp., bauense n. sp. and debile n. sp. are unknown.

414

Mem.

Amer.

En!.

Inst.,

no.

37, 1984

MALES

2

Posterior

surface of

propodeum without

Posterior

surface of

propodeum with

Gonoforceps with a

longitudinal

two

keels . . . . . . 2 keels . . . . . 7

longitudinal

281) 16. nemorale n. sp. inner process (Figs. 270, 282) . . . . . . 3

distal

inner pointed process

(Fig.

• • • • • • • • • • • • . • . • • • • • • • • • • • • • • . • • • • • •• 3

Gonoforceps wi thout di stal Scutum fully reticulate rugose;

notaulices

• • • • • • • • • • • • • • • • • • • • • • • • • • • • . • • • • • • .• Scutum mostly

4

invisible . . . . . . . . . . ..

silvestre n. sp.

sculpture or punctate,

not

fully

ret iculate

rugose;

notaul ices visible . . . . . . . . . . . . . . • • • . . . . . . . . . 4

Head dull,

smooth,

granulated . . . . . . . . .

Head

5

or fully without

17.

shiny,

rugose,

6. yasumatsui n. sp.

irregular keels . . . . . . . . . . . 5

with areolae and

6

reaching approximately 0,65 length of scutum . . . . . . . . . · . . . . . . . . . . . . . . . . . . . . . . . . • . . . . . . . . • . .. 7. sarawaki n. sp. Notaulices reaching at most 0,5 length of scutum . . . . . . . . . . . . . . . 6 Notaul ices reaching approximately 0,5 length of scutum . . . . . . . . .. · . . . . . . . . . . . . . . • . . . . . . . . . . . . . . . . . . . . .. 9. krombeini n. sp. Notaulices reaching approximately 0,3 length of scutum . . . . . . . . . . · . . • • • . . . . . . . . . . . . . • • . . . . . . • . . . . . . . . .. 2. thai n. sp.

7

Posterior

surface of

propodeum wi th median

fully

not

reticulate

rugose . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8

Posterior

surface of

propodeum wi th median

Notaulices

I ate ra I

8

area

smooth,

area as

almost

rugose as

area s ••••••••••••••••••••••••••••••••••••••••••••••••• 1 2

(Fig. 266) ••••.• 5. mysorense n. sp. Gonoforceps without distal inner process (Figs. 283, 285) . . . . . . 9 Notaul ices reaching approximately 0,5 length of scutum . . . . . . . . 10 9 Notaulices reaching approximately 0,65 length of scutum . . . . . . . 11 10 Head fully or almost fully reticulate rugose . . . . . • • . . . . . . . . . . • • • • • . • • . . • • • • . • • . • • . . . . . . • • • • • • . • • . . . •• 18. expolitum n. sp. Gonoforceps wi th a

distal

inner

rounded process

· . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ..

Head

fully

or almost

fully

weakly punctate •••••••••• ••••••••••••

• • • • • • . • • • • • • • . • • • • • • • . . . • • • • • . • • • • •• 11 Gonoforceps nearly as strongly

punctate;

long as penis

notaulices

punctate;

12 Head fully

shorter

than penis

Head punctate,

not

rugose . . . . . . . . . .

reticulate

20.

(Fig.

notaul ices weak . . . . . . . . . . . . reticulate

peterseni n.

sp.

284); head more

strong . . . . . . . . . . . . . . . . . . . . . . . . . . ..

• ••••..••......•••••••••••••..•.••••• Gonoforceps much

1 9.

(Fig.

21. 2.

bauense n.

sp.

285); head weakly

debile n. sp. thai n. sp.

rugose . . . . . . . . . . . . . . . . . . . . . . . . . . ..

• • • • . • . . • • • • • • . • • • • • • • • • • • • • • • • • • • • ••

13. javanum n.

sp.

The males of A. philippinum n. sp., munroei n. sp., silvicolum n. lankanum n. sp.; bengalense n. sp., laotianum n. sp., borneanum n. sp., muiri n. sp., fyanense n. sp. and indicum n. sp. are unknown. sp.,

Anteoninae

Fig.

260 -

Chela of Anteon borneanum n.

415

sp.

(type)

416

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

1. Anteon borneanum n. sp.

FEMALE:

fully

testaceous;

winged;

length 1,87 mm;

legs testaceous,

black;

mandibles and antennae

with hind coxae partly brown;

clubs of femora partly brown;

mid and hind

antennae distally thickened;

antenna I

segments in following proportions: 12: 5: 5: 4: 4: 4: 4: 4: 4: 5; head shiny, granulated, with numerous irregular keels; frons with a median keel and 2 lateral keels around orbits; occipital carina complete; POL = 5; OL

3; OOL keel s;

= 5; OPL = 3; TL = 3; pronotum shiny, with numerous transversal posterior surface of pronotum smooth, short, shorter than scutum

(3:14);

pronotal

tubercles reaching tegulae;

scutum shiny,

except for the anterior region near pronotum which smooth;

notaulices incomplete,

reaching approximately 0,5

scutell um and metanotum sh i ny, reticulate rugose, posterior surface;

smooth,

granulated,

is not sculptured and

wi thout scu I pture;

length of scutum; propodeum

with a strong transversal keel between dorsal and posterior surface without longitudinal keels; fore-wing

hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part (3:7); fore tarsal segments in following proportions: a proximal than apical

11

1,5: 2:

5:

2: 9; enlarged claw (Fig. 260) with one bristle on

prominence;

segment 5 of front

lamellae extending continuously

MALE: unknown LOCUS TYPICUS:

to the apex;

Nanga Pelagus (Sarawak,

TYPICAL MATERIAL: HOSTS:

tarsus with basal

(7:2); segment 5 of front tarsus (Fig. 260)

part

holotype F!

tibial

part

longer

with 2 rows of

spurs 1,1,2.

Malaysia)

in B.

unknown

DISTRIBUTION: Sarawak),

Oriental

region:

MALAYSIA:

Nanga Pelagus (near Kapit,

B!

2. Anteon thai n. sp.

FEMALE:

fully

winged;

length 4,6 mm;

black,

with mandibles and

legs

testaceous; antennae testaceous, wi th segments 4-10 brown; antennae distally thickened; antennal segments in following proportions: 9: 5: 4,5: 5: keel; 4;

5:

5:

5:

7;

head shiny,

head fully reticulate rugose;

OOL

hairy,

5:

=

5;

OPL

=

5;

TL

=

3;

hairy;

occipital

carina complete;

pronotum dull,

nearly twice as broad as

long;

strongly punctate,

pronotal

with punctures very

without sculpture among punctures; approximately 0,3 length of scutum; metanotum,

shiny,

anterior margin;

smooth,

= 6; OL

tubercles reaching tegulae; (5:13);

broad,

scutum shiny,

similar to areolae,

notaul ices incomplete,

reaching

scutellum as sculptured as scutum;

without sculpture,

propodeum fully

POL

fully reticulate rugose,

posterior surface of pronotum shorter than scutum hairy,

6:

frons with a track of median

with some areolae only near

reticulate rugose,

dull,

with a strong

Anteoninae

417

Fig.

261

-

Chela of Anteon thai n.

sp.

(type)

418

Mem.

transversal with two

keel

keels;

proportions:

50:

on a proximal

Inst.,

10:

37,

lateral

with some transversal

11:

no.

distal

part of radial

fore-wing vein much

segments in following

enlarged claw

segment 5 of front

group of 2 lamellae;

posterior surface

areas of posterior surface

keels on median area;

bands;

(5: 18); fore tarsal

16: 45;

prominence;

1984

and posterior surface;

median and

without dark transversal

shorter than proximal part

with a

Ent.

between dorsal

longitudinal

reticulate rugose, hyaline,

Amer.

(Fig.

261)

with one bristle

tarsus with 2 lamellae;

segment 5 of front

tarsus

(Fig.

261)

apex

with basal

part longer than apical part (37:8); tibial spurs 1,1,2. MALE: fully winged; length 1,68-2,88 mm; black; mandibles and antennae testaceous;

legs testaceous,

with

hind coxae proximally black;

ditally thickened;

antenna I segments in following proportions:

5:

head shiny,

5:

keel

5:

4:

5:

9;

fully reticulate rugose;

antennae not 10: 7:

6:

5:

frons with a median

and with 2 lateral keels near orbits directed towards antenna I sockets;

occipital carina complete; scutum,

POL

= 7; OL = 4; OOL = 7; OPL = 4; TL = 4;

scutellum and metanotum shiny,

sculpture among punctures; strongly punctate;

smooth,

weakly punctate,

without

anterior surface of scutum reticulate rugose or

notaulices incomplete,

reaching approximately 0,3 length

of scutum; propodeum reticulate rugose, with a strong transversal keel between dorsal and posterior surface; posterior surface with 2 strong or weak

(almost invisible)

lateral

areas;

part of radial

Fig.

262 -

longitudinal

fore-wing hyaline,

keels;

median area as rugose as

without dark transversal

vein much shorter than proximal

Male genital ia of Anteon thai

n.

sp.

left half removed.

part

bands;

(3,5:9);

distal

gonoforceps

from Chawang

(Thai land):

419

Anteoninae

262) without proximal membranous process and without distal inner tibial spurs 1,1,2.

(Fig.

process;

LOCUS TYPICUS:

Mudigere (India)

TYPICAL MATERIAL:

holotype F!

2 paratypes MM!

in B; HOSTS:

in WA;

and 1 paratype M!

3 paratypes MM!

in CO;

2 para types MM!

in OL.

unknown

DISTRIBUTION:

Oriental

region:

THAilAND:

Chawang

(Nabon,

Banna),

Banna, B! HONG KONG: Taipokau (Kowloon), OL! INDIA: Mudigere (Karnataka), OL! CO! SRI LANKA: Udawattakele Sanct. (Kan. Dist.), Ol!

Kande Ela

(NE Dist.),

B! WA!

WA!

3. Anteon silvicolum n. sp.

FEMALE: fully winged; length 3,20 mm; black, with mandibles, antennae and legs testaceous; antennae distally thickened; antenna I segments in following proportions: punctate,

14: 7: 12: 10: 10: 9: 9: 8: 8:

wi thout scu I pture among punctures,

anterior half of frons; reticulate rugose; keel;

POL = 7;

rugose;

occipital

OL = 6;

pronotal

punctate,

vertex sculptured by

with a strong transversal areas;

part of radial

keel

two

smooth,

notaul ices incomplete,

scutellum and metanotum shiny,

without sculpture among punctures;

posterior surface with

pronotum reticulate

scutum shiny,

without sculpture among punctures;

punctate,

punctate on

frons with a complete median TL = 4,5;

tubercles reaching tegulae;

reaching 0,5 length of scutum;

lateral

OPL = 5;

head shiny,

irregular keels and partly

carina complete;

OOL = 8;

11;

more strong I y

between dorsal

longitudinal

fore-wing hyaline,

keels;

and posterior surface; median area as rugose as

without dark transversal

vein much shorter than proximal

segments in following proportions:

smooth,

propodeum reticulate rugose,

part

bands;

distal

(5:15); fore tarsal

12: 3: 3: 6: 15; enlarged claw (Fig. 263)

with one bristle on a proximal prominence; segment 5 of front tarsus (Fig. 263) with 1 row of 18 lamellae; apex with a group of nearly 6 lamellae; tibial

spurs 1,1,2.

MALE unknown lOCUS TYPICUS:

Kanda-ela Reservoir

TYPICAL MATERIAL:

holotype F!

HOSTS: unknown DISTRIBUTION: Oriental region: Nuwara Eliya,

N.E.

Dist.),

(Sri

lanka)

in WA. SRI

lANKA:

Kanda-ela Reservoir

(5.6 mi. SW

WA!

4. Anteon indicum n. sp.

FEMALE:

fully

testaceous;

winged;

legs fully

length 2,18-3,12 mm;

black;

mandibles and antennae

testaceous or with hind coxae and hind clubs of

420

Mem. Amer. En!.

Inst., no. 37, 1984

Fig. 263 - Chela of Anteon silvicolum n. sp.

(type)

Anteoninae

Fig. 264 - Chela of Anteon indicum n. sp.

421

(type)

422

Mem. Amer.

femora

brown;

En!.

Inst.,

antennae slightly distally

following proportions:

10:

4:

5:

6:

5:

no.

5:

5:

without scu I pture among punctures;

punctate;

frons without median

median keel; TL = 3; short,

occipital

and

carina complete;

pronotum shiny,

with

shorter than scutum

keels;

keels,

pronotal

segments

head shiny,

in

weakly

somet i mes more strong I y

= 7;

POL

antennal

9;

5:

lateral

transversal

(6:16);

1984

thickened;

5:

puncta te,

keel

37,

OL

sometimes with

= 5;

= 4;

OOL

5-,

OPL

with posterior surface

tubercles reaching

tegulae;

scutum shiny, smooth, weakly punctate, without sculpture among punctures; notaul ices incomplete, reaching approximately 0,3 length of scutum; scutellum and metanotum shiny,

without sculpture;

with a

posterior surface;

posterior surface with 2

shiny,

smooth,

also in central

area;

part of radial

tarsal

segments

fore-wing hyaline,

front

tarsus with basal (Fig.

front

tarsus

part

(Fig.

tibial

longitudinal

264)

and

median

area

occasionally with keels part

bands;

(3:7);

fore

9: 2: 2: 2,5: 12,5; segment 5 of

proximal

part

(10:2,5);

prominence;

1 row of 5 lamellae;

with

keels;

without dark transversal

longer than proximal

with one bristle on a

propodeum

between dorsal

near margins;

in following proportions:

264)

keel

vein much shorter than proximal

claw

MALE:

transversal

with a few keels only

distal

lamellae;

strong

smooth,

reticulate rugose,

enlarged

segment 5 of

apex with a group of 16

spurs 1,1,2.

unknown

LOCUS TYPICUS:

Bangalore

TYPICAL MATERIAL: HOSTS:

in

India

DISTRIBUTION:

(India)

holotype F!

and

paratype F!

INDIA:

Bangalore

in BM.

ldiocerus sp.

Oriental

region:

(Mysore),

BM!

5. Anteon mysorense n. sp.

FEMALE:

fully

testaceous; thickened; 3:

4:

keels;

6;

winged;

length 2 mm;

legs testaceous, antennal

with

mandibles and antennae fully

segments in following proportions:

head shiny,

punctate,

keels;

POL

9:

antennae distally 4:

3:

3:

weakly granulated and with weak

occipital carina complete;

incomplete lateral

black;

hind coxae partly black;

=

frons with a median keel 6;

OL

=

4;

OOL

=

4;

TL

=

3:

3:

3:

irregular

and with 2 4;

pronotum

shi ny, wi th SOme transversa I keel s, with posterior surface shorter than scutum (4:10); pronotal tubercles reaching tegulae; scutum shiny, smooth, weakly punctate, without sculpture among punctures; notaulices incomplete, reaching approximately 0,3 length of scutum; scutellum and metanotum shiny, smooth, between dorsal keels;

without sculpture; propodeum with a strong transversal keel and posterior surface; posterior surface with 2 longitudinal

median area shiny,

fore-wing hyal ine,

smooth with a few keels only near margins;

without dark transversal

vein much shorter than proximal following proportions: basal

part

6:

1,5:

longer than apical

part

1,5: part

bands;

(3:5);

1,5: 9; (6:3);

fore

distal

tarsal

part of radial

segments

segment 5 of front enlarged claw

(Fig.

in

tarsus with 265)

with

Anteoninae

423

265 - Chela of Anteon mysorense n. sp.

Fig.

(type)

one bristle on a proximal prominence; segment 5 of front tarsus with 4 lamellae; apex with a group of nearly 9 lamellae; tibial

(Fig. 265) spurs

1,1,2. MALE: fully

fully

length 2,18 mm;

winged;

testaceous;

following proportions: 9: punctate, rugose, without

black;

mandibles,

antennae not distally thickened;

antennae and

antennal

legs

segments in

5: 5: 5: 5: 5: 5: 5: 5: 8; head shiny, strongly

with vertex and region between ocelli

and occipital carina

without sculpture among punctures; frons with a median keel, lateral keels; occipital carina complete; POL = 7; Ol = 3,5; OOl

4; OPl = 4;

Tl = 4;

anterior half,

scutum shiny,

smooth,

weakly

without sculpture among punctures;

reaching approximately 0,3 length of scutum;

punctate,

mostly on

notaulices incomplete,

scutellum and metanotum

shiny, smooth, without sculpture; propodeum with a strong transversal keel between dorsa I and posterior surface; posterior surface wi th 2 longi tud i na I keels; lateral distal

median area shiny, areas rugose; part of radial

gonoforceps (Fig.

smooth,

with a few keels only near margins;

fore-wing hyaline,

without dark transversal

vein much shorter than proximal

part

bands;

(3:5);

266) without proximal membranous process and with distal tibial spurs 1,1,2.

inner rounded process; lOCUS TYPICUS:

Malleswaram

TYPICAL MATERIAL:

(India)

holotype F!

and 1 paratype M!

in BM;

1 paratype F!

INDIA:

(Mysore),

Ol. HOSTS: unknown DISTRIBUTION: Oriental

region:

Malleswaram

BM!

OL!

in

424

Mem.

Amer.

En!.

Inst.,

no.

37,

1984

266 - Ma Ie gen i ta I i a of Anteon mysorense n. sp. from Ma II eswaram

Fig.

(India):

right half removed.

6. Anteon yasumatsui n. sp.

FEMALE: fully 2,5:

fully

winged;

length 2,00 mm;

yellow-testaceous; 2,5:

2: 3:

longitudinal

3:

3:

antennal

3:

5;

median keel;

black;

antennae,

tegulae and

segments in fol/owing proportions:

head dull,

occipital

granulated;

frons

carina complete;

legs

8:

4:

with an incomplete

POL = 4;

OL = 3;

OOL

4; OPL = 3; TL = 5; pronotum granulated, with posterior surface slightly longer than scutum (8:6) and with transversal keels; pronotal tUbercles reaching tegulae; incomplete,

scutum shiny,

metanotum shiny, rugose,

smooth,

posterior surface without dark transversal proximal

part 9;

bands;

(2:4);

propodeum fully

between dorsal

longitudinal

keels;

fore-wing hyaline,

distal part of radial

fore tarsal

reticulate

and posterior surface; without

vein much shorter than

segments in following proportions:

segment 2 of front tarsus produced into a hook;

(Fig.

267)

nearly 4 lamellae; MALE:

with 2 rows of nearly 18 lamellae; tibial

fully winged;

tegulae and 5;

keel

notaulices

scutellum and

5:

1,5:

enlarged claw

267) with one bristle on a proximal prominence; segment 5 of front

(Fig. tarsus

=

without sculpture;

without sculpture;

with a transversal

2: 3,5:

smooth,

reaching approximately 0,5 length of scutum;

OL

=

3;

length 1,50-2,00 mm;

legs fully OOL

=

apex with a group of

spurs 1,1,2.

yellow-testaceous;

4; OPL

=

3;

TL

=

4;

black;

mandibles,

antennae,

head fully granulated, occipital

dull;

carina complete;

POL

pronotum

Anteoninae

Fig.

incomplete, smooth,

Chela of Anteon yasumatsui n.

267 -

short and rugose;

scutum shiny,

without

keel

distal

keels;

gonoforceps (Fig. without distal

268)

F)!

without dorsal

without dark transversal

proximal tibial

HOSTS:

2 paratypes

(1

F,

1 M)!

with a

posterior surface

vein much shorter than proximal

inner pointed process;

in BM;

notaulices

reticulate rugose,

and posterior surface;

part

(2:5);

membranous process and

spurs 1,1,2.

LOCUS TYPICUS: Phi bun Mangsahan (Thailand) TYPICAL MATERIAL: holotype F! and 1 paratype M! M,1

(type)

scutellum and metanotum shiny,

fore-wing hyal ine,

part of radial

sp.

without sculpture;

propodeum fully

between dorsal

longitudinal

bands;

smooth,

reaching 0,5 length of scutum;

without sculpture;

transversal

425

in VA;

2 paratypes (1

in OL.

unknown

DISTRIBUTION: Ratchathani),

Oriental VA!

region:

Non Sa-ard

THAILAND:

Phibun Mangsahan

(Maha Sarakham),

(Java), OL! INDIA: Delhi, BM! OL! NOTE: this species is named in honor of the

VA!

late Mr.

(Ubol

INDONESIA:

Pekalongan

Keizo Yasumatsu.

7. Anteon sarawaki n. sp.

"EMALE:

fully

2nd legs fully 3:

14: 9:

9:

9:

winged;

length 3,62 mm;

testaceousj 8:

8:

7:

blaCk;

antennae segments 9;

head shiny,

mandibles,

antennae,

tegulae

in following proportions:

rugose,

12:

with numerous areolae on

426

Mem.

Fig.

268 -

Amer.

Ent.

Inst.,

no.

37,

Anteon yasumatsui

Male genital ia of

1984

n.

sp.

from

Non Sa-ard

(Thailand)

frons and with

irregular keels on vertex and behind ocelli;

among areolae and keels; lateral

frons with a

keels around orbits;

POL = 6;

longitudinal OL = 4;

no sculpture

median keel

OOL = 6;

and 2

OPL = 5;

TL = 3;

occipital carina complete; pronotum hairy, rugose, shiny, strongly transversely striate and punctate, smooth only near posterior margin, posterior surface nearly as reaching

tegulae;

scutum shiny,

sculpture among punctures; margin of scutum;

long as scutum hairy,

(8:9);

smooth,

notaulices long,

pronotal

with

tubercles

weakly punctate,

without

almost reaching posterior

scutellum and metanotum shiny,

smooth,

weakly punctate,

without sculpture among punctures; propodeum reticulate rugose, with a transversal keel between dorsal and posterior surface; posterior surface without longitudinal keels; fore-wing hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part (3:12); fore tarsal

segments

in following proportions:

10:

3:

5:

10:

23;

enlarged

claw (Fig. 269) with one bristle on a proximal prominence; segment 5 of front tarsus (Fig. 269) with basal part much shorter than apical part, with 2 rows of nearly 35 1,1,2. MALE:

fully

tegulae and shiny, keel;

winged;

lamellae;

apex with a group of 4

length 2,12-2,50 mm;

legs fully

testaceous,

fully reticulate rugose, POL = 8;

OL = 4,5;

black;

lamellae;

mandibles,

tibial

spurs

antennae,

except for fore coxae basally black;

granulated;

OOL = 6;

head

frons with no visible median

OPL = 5;

TL = 3;

occipital

carina

complete; scutum hairy, shiny, smooth, strongly punctate, without sculpture among punctures; notaulices long, incomplete, almost reaching posterior margin of scutum;

scutellum and metanotum shiny,

propodeum reticulate rugose, posterior surface; hyal ine,

with a

transversal

posterior surface without

without dark transversal

bands;

smooth,

keel

longitudinal

distal

weakly punctate;

between dorsal keels;

part of radial

and

fore-wing vein much

Anteoninae

Fig.

269 -

shorter than proximal proximal

part

Gunong Mulu

TYPICAL MATERIAL: OL.

Anteon sarawaki n. sp. (type)

(3:11);

membranous process,

1,1,2. LOCUS TYPICUS:

HOSTS:

Chela of

427

gonoforceps

without distal (Sarawak,

holotype F!

(Fig.

270)

without dorsal

inner process;

tibial

spurs

Malaysia)

and 1 paratype M!

in BM;

1 paratype M!

in

unknown

DISTRIBUTION:

Oriental

region:

MALAYSIA:

Gunong Mulu

(Sarawak),

BM!

OL!

428

Mem.

Fig.

Amer.

Ent.

Inst.,

no.

37,

1984

270 - Male genitalia of Anteon sarawaki n. sp. from Gunong Mulu (Malaysia)

8. Anteon bengalense n. sp.

FEMALE:

fully

antennae

winged;

length

testaceous-dark;

5 mm;

legs

black;

mandibles

testaceous,

with

testaceous;

femora,

tibiae and

hind coxae partly brown; antennae distally thickened; antennal segments in following proportions: 20: 8: 13: 10: 8: 8: 8: 8: 8:

11; head shiny,

rugose,

frons wi th a median keel

wi thout and

also with numerous areolae; with

some areolae and

occipi tal

carina

(as

two

sculpture among areolae and keels; lateral

region

of

two oblique keels in

keels around

orbits;

from posterior ocelli

to

Deinodryinus); SOme areolae also near occipital

carina and on temples; POL = 5; OL = 3; OOL = 8; OPL = 8; TL = 7; pronotum shiny, with numerous transversal striae on anterior region, on posterior region,

frons

head behind posterior ocelli

without sculpture,

of pronotum shorter than scutum

with sides hairy;

(14: 16); pronotal

smooth

posterior surface

tubercles reachi ng

tegulae; scutum shiny, weakly punctate, without sculpture among punctures; notaulices incomplete, only visible near anterior margin of scutum; scutellum and metanotum shiny, among punctures; keel

between

smooth,

weakly

propodeum reticulate rugose,

dorsal and posterior surface;

punctate,

without sculpture

with a strong transversal

posterior surface rugose,

without

Anteoninae

Fig.

271

-

429

Chela of Anteon bengalense n.

longitudinal

keels;

fore-wing with 1 dark transversal

pterostigma;

distal

part of radial

(5:16);

fore tarsal

sp.

band beneath

vein much shorter than

segments in following proportions:

(type)

5,5:

proximal 4:

5:

the part

12: 26;

enlarged claw (Fig. 271) with one bristle on a proximal prominence; segment 5 of front tarsus (Fig. 271) with 2 rows of 25 lamellae; apex with a group of 8 I ameli ae; tibial

spurs 1,1,2.

MALE:

unknown

segment 2 of front

tarsus produced

into a hook;

LOCUS TYPICUS: Ammatti (India) TYPICAL MATERIAL: holotype F! in OT. HOSTS: unknown DISTRIBUTION: Oriental

region:

INDIA:

Ammatti

(S.

Coorg),

OT!

9. Anteon krombeini n. sp.

FEMALE: fully winged; length 4,52 mm; head black, with mandibles testaceous; antennae brown, with ventra I si de of segment 1 and segments 5-6 testaceous;

thorax,

propodeum and abdomen black;

mid tibiae and mid tarsi and fore tarsi specimen;

brown;

testaceous;

occasionally

28:

10:

18:

11:

10:

10:

antennal

10: 9:

9:

segments in following

14;

wi th

with trochanters

posterior legs missing in the only known

antennae distally thickened;

proportions:

legs testaceous,

legs black,

head shiny,

flat,

430

Mem. Amer.

Ent.

Inst.,

no.

37,1984

fully reticulate rugose; frons with a faintly visible median keel and two lateral keels around orbits; occipital carina complete; POL = 9; OL = 4; OOL

=

9;

OPL

=

by transversal smooth;

pronotal

irregular keels,

Fig.

13;

TL

=

10;

pronotum hairy,

shiny,

strongly sculptured

keels, except for a short area near the scutum which tubercles reaching tegulaej

scutum shiny,

except for anterior half of median area which

272 -

Chela of

is

sculptured by

Anteon krombeini n. sp. (type)

is smooth,

Anteoninae

431

punctate and without sculpture among punctures; reaching approximately 0,5 punctate, by

without sculpture among punctures;

longitudinal

transversal without

keel

keels;

(4:8);

dorsal

keels;

beneath the pterostigma; part

notaulices incomplete,

scutellum shiny, metanotum shiny,

propodeum reticulate rugose,

between

longitudinal

proximal

length of scutum;

fore

distal

part of radial

tarsal

sculptured

with a strong

and posterior surface;

fore-wing crossed by a

smooth,

posterior surface

dark

transversal

band

vein much shorter than

segments in following proportions:

10:

5:

8: 20: 48; enlarged claw (Fig. 272) with one bristle on a proximal prominence; segment 5 of fore tarsi (Fig. 272) with some rows of nearly 50 lamellae extending continuously MALE:

fully

winged;

antennae testaceous, partly

Fig.

black;

273 -

to the apex;

length 3,00-3,28 mm;

with segments 4-10 dark;

antennae hairy,

Male genitalia of

not distally

tibial

black;

spurs 1,1,2.

mandibles testaceous;

legs testaceous,

thickened;

antennal

with coxae segments

Anteon krombeini n. sp. from Induruwa

Jungle (Sri Lanka).

432

Mem.

Amer.

Ent.

15:

8:

in following proportions: fully

reticulate rugose;

carina complete;

POL

8:

punctures;

=

=

9; OL

without dark

8:

=

smooth,

8:

8:

1984

8:

transversal

band;

(4:12);

11;

head shiny,

punctate,

occipital

=

7; TL

4; scutum,

without sculpture among

reaching approximately 0,5 distal

part

without distal

inner pointed process;

hairy;

273)

(Fig.

tibial

length of

fore-wing hyaline,

part of radial

gonoforceps

LOCUS TYPICUS: Weddagala (Sri TYPICAL MATERIAL: holotype F!

=

8; OPL

scutellum and metanotum very

than proximal

F!

8:

8; OOL

notaulices incomplete,

scutum,

8:

37,

no.

frons with a median complete keel;

scutellum and metanotum shiny, scutum;

Inst.,

vein much shorter

long and narrow,

spurs 1,1,2.

Lanka) and 2 paratypes MM!

1 paratype

in WA;

in OL.

HOSTS:

unknown

DISTRIBUTION: Rat.

Dist.),

Jungle

Oriental

WA!

(Gi I imale,

NOTES:

region:

SRI

LANKA:

Weddagala

Kandy Reservoir Jungle (Kan. Rat.

this species

Dist.),

WA!

Hasalaka

Dist.),

(Kan.

is named in honor of Mr.

(Sinharaja Forest, OL!

Dist.),

Induruwa WA!

V. Krombein.

Karl

10. Anteon muiri n. sp.

FEMALE:

fully

winged;

length 4,43 mm;

black;

mandibles and antennae

testaceous; legs testaceous, with coxae and clubs of femora partly brown; antennae distally thickened; antennal segments in following proportions:

13: 6: 8: 6: 5: 6: 6: 6: 6: 9; head shiny, fully strongly reticulate rugose;

frons with a median keel

occipital

c:c;rina complete;

POL

=

and with 2 5;

OL

=

3;

lateral OOL

=

8;

keels around orbits; OPL

=

6;

TL

=

6;

pronotum shiny, strongly transversely striate, except for an area near posterior margin which is smooth and without sculpture; posterior surface

(12:19); pronotal tubercles reaching

of pronotum shorter than scutum tegulae;

scutum,

punctate,

scutellum and metanotum shiny,

without sculpture among punctures;

smooth,

weakly

notaul ices incomplete,

reaching approximately 0,65 length of scutum; propodeum reticulate rugose, with a strong transversa I keel between dorsa I and posterior surface;

posterior surface with 2 longitudinal

rugose as bands;

lateral

areas;

fore-wing hyal ine,

distal part of radial

keels;

median area as

without dark transversal

vein much shorter than proximal

part

(4:15); fore tarsal segments in following proportions: 6: 3: 4: 9: 25; enlarged claw (Fig. 274) with one bristle on a proximal prominence; segment 5 of front tarsus with basal part shorter than apical part (9:16); segment 5 of front tarsus (Fig. 274) with 2 rows of 22 lamellae; apex with a MALE:

group of 4

LOCUS TYP I CUS:

Mowong

TYPICAL MATERIAL: HOSTS:

lamellae;

tibial

spurs 1,1,2;

unknown

unknown

(Western Borneo,

holotype F!

in B.

I ndonesi a)

Anteoninae

DISTRIBUTION: NOTES:

Oriental

region:

INDONESIA:

433

Mowong

this species is named in honor of Mr.

Fig.

274 -

Chela of

Anteon muiri

F.

n.

(W Borneo),

Muir.

sp.

(type)

B!

434

Mem. Amer. Ent.

Fig. 275 -

Chela of

Ins!', no. 37, 1984

Anteon laotianum n. sp. (type)

Anteoninae

Anteon laotianum n. sp.

11.

FEMALE:

fully

brown;

length 4,37 mm;

winged;

and testaceous;

antennae testaceous,

antennae distally

proportions:

thickened;

black;

frons

with a

keels around orbits;

mandibles partly brown

with segments 6-10 brown; antennal

4,5: 5: 5: 5: 8: 8: 6: 7:

20:

reticulate rugose; lateral

435

median

occipital

segments

13;

head rugose,

longitudinal

keel

carina complete;

legs

in following fully

and with 2

furrow

around

occipital carina crossed by short keels; POL = 8; OL = 5; OOL = 7; OPL 10; temples invisible; pronotum shiny, smooth, weakly punctate, without sculpture among punctures,

and a

with

a

weak

weak posterior transversa I

tegulae;

scutum shiny,

punctures;

smooth,

anterior transversal

impress ion;

weakly punctate,

notaul ices incomplete,

without sculpture among

reaching approximately 0,25

scutum;

scutellum and metanotum dull,

rugose;

rugose,

with a

between dorsal

strong transversal

posterior surface with 2

surface;

by transversal

keels;

fore-wing

keel

longitudinal

with 2 dark

vein much shorter than proximal

pronotum shorter than scutum

keels;

proximal

rows of 27 MALE:

and posterior

median area crossed

transversal

bands,

distal

one on

part of

(4:9); posterior surface of segments in following

part

fore tarsal

9: 2,5: 3,5: 8: 22; enlarged claw (Fig. 275) with one bristle

proportions: on a

(17:21);

length of

propodeum reticulate

the basal cells and another beneath the pterostigma; radial

impression

pronota I tubercles reach i ng

prominence;

lamellae;

segment 5 of front

apex with a

group of 4

tarsus

(Fig.

lamellae;

275)

tibial

with 2

spurs 1,1,2.

unknown

LOCUS TYPICUS:

Ban Van Eue

TYPICAL MATERIAL: HOSTS:

(Laos)

holotype F!

in B.

unknown

DISTRIBUTION:

Oriental

region:

LAOS:

Ban Van Eue

(Vientiane),

B!

12. Anteon fyanense n, sp.

FEMALE:

fully

testaceous; distally 10:

9:

winged;

thickened;

9:

8:

median keel

length 3,68 mm;

antennae testaceous, 8:

8:

antennal 10;

mandibles and

segments in following

head shiny,

and with 2

black;

wi th segments 7-10 brown;

lateral

fully

proportions:

reticulate rugose;

keels around orbits;

legs

antennae 16:

6:

14:

frons with a

occipital

carina

complete; POL = 6; OL = 4; OOL = 9; OPL = 10; TL = 8; pronotum shiny, hairy, strongly reticulate rugose, with posterior surface shorter than scutum smooth,

(15:20);

pronotal

tubercles reaching

strongly punctate,

incomplete,

reaching

metanotum shiny, punctures;

approximately 0,65

smooth,

tegulae;

scutum shiny,

without sculpture among punctures; weakly

length of scutum;

punctate,

propodeum reticulate rugose,

notaulices

scutellum and

without sculpture among

with a strong transversal

keel

436

Mem. Amer. Ent.

Inst.,

no. 37, 1984

between dorsal and posterior surface; posterior surface with 2 longitudinal keels; median area rugose, but with a narrow, shiny,

Fig. 276 -

Chela of Anteon fyanense n. sp.

(type)

Anteoninae

437

central region which is smooth and without sculpture; fore-wing with dark transversal band beneath the pterostigma; distal part of radial much shorter than proximal proportions: on a

9:

4:

proximal

5:

13:

part

31;

(8:19);

fore

enlarged claw

segment 5 of front

prominence;

rows of 24 lamellae extending continuously 1,1,2. MALE:

tarsal (Fig.

segments

276)

tarsus

1 vein

in following

with one bristle

(Fig.

to the apex;

276)

with 2

tibial

spurs

unknown

LOCUS TYPiCUS:

Fyan

TYPICAL MATERIAL:

(Viet Nam)

holotype F!

HOSTS: unknown DISTRIBUTION: Oriental

region:

in B. VIET NAM:

Fyan,

B!

13. Anteon javanum n. sp.

FEMALE: brown,

fully

winged;

length 2,62 mm;

with segments 1-3 testaceous;

testaceous-dark;

scutell um,

and metasternum part I y antennae distally

thickened; 6:

8:

8:

7:

6:

6:

6:

abdomen brown;

antennae

scutum reddi shmesosternum

legs testaceous;

antenna I segments in following proportions:

9;

sculpture among punctures;

reddish-testaceous;

metanotum and propodeum black;

reddi sh-dark;

9:

5:

head

prothorax yellow;

head shiny,

strongly

punctate,

frons with a median keel

without

and with 2

lateral

weak keels near orbits; occipital carina complete; POL; 3; OL ; 3; OOL ; 4; OPL ; 3,5; TL ; 4,5; OPL slightly longer than breadth of posterior ocelli; pronotum shiny, rugose, with posterior surface flat, longer than scutum (13:11); pronotal tubercles reaching tegulae; scutum shiny, smooth,

weakly punctate,

almost complete,

without sculpture among punctures;

reaching

and metanotum shiny,

approximately 0,65

smooth,

without sculpture;

rugose,

with a strong transversal

surface;

posterior surface with 2

keel

notaul ices

length of scutum;

between dorsal

longitudinal

scutellum

propodeum reticulate

keels;

and posterior

median

area nearly

as rugose as lateral areas; fore-wing hyal ine, without dark transversal bands; distal part of radial vein much shorter than proximal part (3:11);

fore tarsal

segment 5 of front (5:14);

enlarged claw

prominence; lamellae; MALE:

segments in following proportions: tarsus with basal (Fig.

277)

fully

winged;

occipital 5;

OL ;

punctate,

carina complete; 3;

OOL ;

5;

(Fig.

lamellae;

length 1,87 mm;

fu II y testaceous; antennae brown, shiny, smooth, strongly punctate,

277) tibial

black;

2;

10:

19;

part

spurs 1,1,2. tegulae and

legs

with segments 1-2 testaceous; head without sculpture among punctures;

TL ;

3;

longitudinal

scutum shiny,

without scu I pture among punctures;

reaching approximately 0,5

2:

with 2 rows of 20

mandibles,

frons with a median

OPL ;

2:

witr, one bristle on a proximal

segment 5 of front tarsus

apex with a group of 4

6:

part shorter than apical

length of scutum;

keel;

smooth,

POL

weakly

notau I ices i ncomp I ete, scutellum and metanotum

438

Mem.

Fig.

shiny,

smooth,

transversal with 2

277 -

Ent.

between dorsal

longitudinal

Inst.,

no.

37,

Chela of Anteon javanum n.

without sculpture;

keel

wing hyal ine,

Amer.

keels;

sp.

(type)

propodeum reticulate rugose,

and posterior surface;

part

(5:10);

bands;

distal

gonoforceps

with a

posterior surface

median area as rugose as lateral

without dark transversal

much shorter than proximal

1984

areas;

part of radial (Fig.

278)

forevein

without

Anteoninae

Fig.

439

Male genitalia of Anteon javanum n.

278 -

sp.

from Tjibodas

(Java):

left half removed.

dorsal

proximal membranous process,

spurs 1,1,2. lOCUS TYPICUS:

Tjibodas

TYPICAL MATERIAL:

without distal

inner process;

tibial

(Java)

holotype F!

and 1 paratype M! in BM; 1 para type M! 1 paratype M! in OT. HOSTS: unknown DISTRIBUTION: Oriental region: INDONESIA: Tjibodas (Java), BM! lAOS: Ban Van Eue (Vientiane), B! INDIA: Kodaikanal (Pulney Hills, S. India), OT! in B;

14. Anteon munroei n. sp.

Chelogynus ephippiger (Da I man): Moczar 1979: 77.

FEMALE: black;

fully

winged;

length 2,20-2,40 mm;

antennae distally thickened;

proportions:

9:

5:

without sculpture, median keel;

5:

5:

5:

with small

=

POL

carina complete;

7:

4; Ol

=

5:

tegulae;

5;

OOl

testaceous,

8;

head flat,

notaulices incomplete,

with petiole

=

5;

smooth,

OPl

=

5;

shiny,

smooth,

smooth,

frons with a complete Tl

=

6;

without sculpture;

pronotum longer than scutum

scutellum and metanotum shiny, sculpture;

6:

punctures on vertex;

pronotum shiny,

tubercles reaching

6:

fully

antenna I segments in following

occipital pronotal

(17: 10);

weakly alutaceous,

scutum,

without

reaching approximately 0,80 length of

440

Mem.

Fig.

scutum;

Amer.

propodeum reticulate rugose,

37,

with a

1984

sp.

(type)

transversal

posterior surface wi th two

without dark transversal

shorter than proximal proportions:

5:

2:

3:

part 9:

a proximal prominence; segment 5 of front apex with

(3:9);

16;

Sq.

TYPICAL MATERIAL:

bands;

fore tarsal

enlarged claw

segment 2 of front

tarsus

(Fig.

279)

a group of 3 lamellae;

MALE: unknown LOCUS TYPICUS:

keel

Kabah

areas;

part of radial

forevein

segments in following

(Fig.

279)

with one bristle on

tarsus produced into a hook;

with 1 row of nearly 17 lamellae;

tibial

(Sarawak,

holotype F!

distal

between

long i tudi na I

spurs 1,1,2.

Malaysia)

and 1 paratype F!

in BU.

unknown

DISTRIBUTION: Oriental Sarawak), NOTES:

no.

median area of posterior surface as rugose as lateral

wing hyal ine,

HOSTS:

Inst.,

Chela of Anteon munroei n.

279 -

dorsa I and posterior surface; keels;

En!.

region:

MALAYSIA:

Sq.

Kabah

(Ng.

BU!

this species is named

in honor of Mr.

D.D.

Munroe.

Tada,

3d Div.,

Anteoninae

441

15. Anteon lankanum n. sp.

FEMALE: frons,

fully

winged;

clypeus,

length 2,40 mm;

head black,

mandibles and part of ventral

legs yellow;

antennae distally thickened;

proportions:

11: 7:

7:

7:

7:

7:

6:

7:

7:

antennal 9;

with anterior half of

side yellow;

antennae and

segments in following

head shiny,

smooth,

flat,

hairless, except for yellow anterior ha If of frons whic has dense long hairs; head punctate only along orbits and along occipital carina; frons and vertex without sculpture; occipital carina complete; POL = 3; Ol = 6; OOl

=

5;

=

OPl

posterior ocelli punctate and

4;

Tl

(4:2);

=

5;

OPl nearly twice as

pronotum shiny,

long as breadth of

rugose near anterior margin,

without sculpture among punctures on disc and near

Fig.

280 -

Chela of Anteon lankanum n.

sp.

(type)

442

Mem.

Amer.

En!.

Inst.,

no.

37,

1984

posterior margin; pronotal tubercles reaching tegulae; scutum shiny, smooth, wearkly punctate, without sculpture among punctures; notaulices incomplete,

reaching approximately 0,65 lenth of scutum;

metanotum smooth, punctures;

shiny,

weakly punctate,

pronotum s light I y

a strong transversal surface with two transversal

keel

longer than scutum

between dorsal

longitudinal

keels,

keels;

scutellum and

without sculpture among (14: 11);

propodeum wi th

and posterior surface;

posterior

median area sculptured by

as rugose as lateral

areas;

fore-wing hyaline,

without

dark transversal bands; distal part of radial vein much shorter than proximal part (4,5:10); fore tarsal segments in following proportions: 9:

3: 3: 9:

19;

prominence;

enlarged claw

(Fig.

280)

with one bristle on a proximal

segment 5 of front tarsus

lamellae extending continuously

(Fig.

280)

to the apex;

with 2 rows of nearly 14

tibial

spurs 1,1,2.

MALE unknown LOCUS TYPICUS:

Adams Peak Trail

TYPICAL MATERIAL:

holotype F!

HOSTS: unknown DISTRIBUTION: Oriental Maskeliya,

Kan.

Dis!.),

region:

(Sri

Lanka)

in WA. SRI

LANKA:

Adams Peak Trail

(4.5 mi.

W

WA!

16. Anteon nemorale n. sp.

FEMALE: MALE:

unknown

fully

testaceous;

winged;

distally thickened;

Fig.

281

-

length 1,68 mm;

antennae brown, antennal

black;

mandibles and

with segments 1-2 testaceous;

segments in following proportions:

Male genitalia of Anteon nemorale n. removed;

legs antennae not

B:

sp.

apex of gonoforceps).

(type)(A:

9:

4:

right

4:

half

Anteoninae

4:

5:

4:

4:

4:

5:

8;

head shiny,

443

granulated and fully reticulate rugose;

frons with a median keel and with 2 lateral keels near orbits directed towards antenna I sockets; occipital carina complete; POL; 6; OL ; 3; OOL ;

4;

OPL ;

2;

TL ;

3;

scutum shiny,

without sculpture among punctures; approximately 0,3 length of scutum; smooth,

without sculptur,e;

transversal without

keel

keels;

and posterior surface;

fore-wing hyal ine,

(1,5:5);

gonofoceps (Fig.

inner pointed process;

LOCUS TYPICUS:

281)

Forest Camp

TYPICAL MATERIAL:

Fig.

reaching

scutellum and metanotum shiny, with a strong posterior surface

without dark transversal

di sta I part of rad i a I vei n much shorter than prox i ma I part

a distal

HOSTS:

weakly punctate,

incomplete,

propodeum reticulate rugose,

between dorsal

longitudinal

bands;

smooth,

notaulices,

without proximal tibial

(Sabah,

holotype M!

membranous process,

with

spurs 1,1,2. Malaysia)

in B.

unknown

282 -

Male genitalia of

Anteon silvestre removed.

n.

sp.

(type):

right half

':'44

Mem.

Amer.

DISTRIBUTION: Oriental Kalabakan, Sabah), B!

En!.

region:

Inst.,

MALAYSIA:

no.

37,

1984

Forest Camp

(19 Km N of

17. Anteon silvestre n. sp.

FEMALE:

unknown

MALE: fully winged; length 1,62 mm; black; mandibles testaceous; antennae brown; legs testaceous, with coxae and fore and mid clubs of femora brown;

antennae not distally thickened;

granulated and fully with 2

lateral

occipital

reticulate rugose;

frons

keels near orbits directed POL = 6;

carina complete;

scutum dull,

with a strong transversal

keel

posterior surface without

OL = 3;

OOL = 4;

keels;

OPL = 1;

TL = 2;

notaulices invisible;

and posterior surface;

fore-wing hyaline,

distal part of radial

gonoforceps

and

sockets;

propodeum reticulate rugose,

between dorsal

longitudinal

bands;

(2:6);

part

in

with a median keel

granulated and fully reticulate rugose;

dark transversal

segments

towards antennal

scutellum and metanotum reticulate rugose;

proximal

antennal

6: 4: 3: 4: 4: 4: 4: 4: 4: 6; head dull,

following proportions:

without

vein much shorter than

282) without proximal membranous

(Fig.

process, without distal inner process; tibial spurs 1,1,2. LOCUS TYPICUS: Tenompok (Sabah, Malaysia) TYPICAL MATERIAL: HOSTS:

holotype M!

in B.

unknown

DISTRIBUTION:

Oriental

region:

MALAYSIA:

Tenompok

(Sabah),

B!

18. Anteon expolitum n. sp.

FEMALE:

unknown

MALE: fully winged; antennae testaceous,

length 2,28-2,81 mm; black; mandibles testaceous; with segments 7-10 brown; legs testaceous, with hind

coxae proximally brown; segments shiny,

reticulate rugose,

ocellus which keels, long,

antennae not distally

in following proportions:

10:

except for a

6:

8:

small

7:

complete;

incomplete median keel,

occipital carina complete;

7:

area

is smooth and without sculpture;

with a short,

thickened; 7:

7:

antennal 7:

7:

10;

head

in front of anterior

frons without

lateral

occasionally median

POL = 7;

OL = 4;

keel

OOL = 7;

OPL

= 5;

TL = 5; scutum shiny, smooth, weakly punctate, without sculpture among punctures; notaulices incomplete, reaching 0,5 length of scutum;

scutellum and metanotum shiny, sculpture among punctures; transversal with 2

keel

between

longitudinal

keels;

smooth,

weakly punctate,

propodeum reticulate rugose,

dorsal

and posterior surface;

median area shiny,

except for some keels near margins;

smooth,

fore-wing hyaline,

without with a strong

posterior surface without sculpture, without dark

Anteoninae

Male genitalia of Anteon expolitum n.

283 -

Fig.

445

sp.

(type):

right half

removed.

transversal

vein much shorter than proximal 283) without proximal membranous process and without distal inner process; tibial spurs 1,1,2. LOCUS TYPICUS: Ban Van Eue (Laos) TYPICAL MATERIAL: holotype M! in B; paratype M! in CO. HOSTS: unknown DISTRIBUTION: Oriental region: LAOS: Ban Van Eue (Vientiane), B! part

bands;

(5:12);

distal

part of radial

gonoforceps (Fig.

PHILIPPINES:

Pinigisan

(Mantalingajan,

Palawan),

CO!

19. An teon peterseni n. sp.

FEMALE: MALE:

unknown

fully

testaceous; hairy; 4,5:

winged;

antennal

4,5:

6;

length 1,84 mm;

antennae brown,

with mandibles and

segments in following proportions:

head smooth,

among punctures;

black,

with segmen ts 1-2 testaceous;

shiny,

8:

weakly punctate,

frons with a complete

longitudinal

4:

legs

antennae 5:

4:

4:

4,5:

without sculpture median keel;

5:

446

Mem.

Amer.

Ent.

Inst.,

no.

37, 1984

occipital carina complete; POL = 5; OL = 3; OOL = 7; OPL = 2,5; TL = 3; scutum shiny, smooth, very weakly punctate, without sculpture among punctures; shiny,

weakly punctate,

keels on posterior

median area of posterior surface partly smooth and shiny,

rugose; radial

scutellum

without sculpture among punctures;

with 2 longitudinal

propodeum reticulate rugose, surface;

reaching 0,5 length of scutum;

notaulices incomplete,

smooth,

fore-wing hyal ine,

without dark transversal

vein shorter than proximal

part (4:8);

damaged in the only known specimen;

bands;

distal

partly part of

gonoforceps and penis

gonoforceps without distal

inner

process; tibial spurs 1,1,2. LOCUS TYPICUS: Pinigisan (Philippines) TYPICAL MATERIAL: HOSTS:

holotype M!

in CO.

unknown

DISTRIBUTION: Oriental Palawan), NOTES:

region:

PHILIPPINES:

Pinigisan

(Mantalingajan,

CO!

this species is named

in honor of Mr.

Borge Petersen.

20. Anteon bauense n. sp.

FEMALE:

unknown

MALE: fully winged; length 1,87-2,37 mm; black; mandibles, antennae and legs fully testaceous; sometimes antennal segments 3-10 darkened; antennae not distally thickened; antennal segments in following proportions: punctate, occipital lateral

=

3; TL

10: 5:

7:

7:

7:

6:

6:

6:

6,5:

10;

head shiny,

without sculpture among punctures, carina;

keels;

=

4;

strongly

irregular keels near

frons with a short incomplete median keel,

occipital

carina complete;

scutum shiny,

among punctures;

with

smooth,

weakly

notaul ices incomplete,

= 6;

POL

OL

punctate,

= 3;

without

OOL

= 5;

OPL

without sculpture

reaching approximately 0,65

I ength of scutum; scutell um and metanotum sh i ny, smooth, weak I y punctate, without sculpture among punctures; propodeum reticulate rugose, with a strong

transversal

keel

between

posterior surface with 2 longitudinal

dorsal

keels;

and posterior surface;

median area shiny,

with some keels near margins; lateral areas shiny, fore-wing hyal ine, without dark transversal bands; vein much shorter than proximal as penis, (Fig.

with proximal

part

(3: 11); gonoforceps nearly as long

membranous process,

without distal

inner process

284); tibial spurs 1,1,2.

LOCUS TYPICUS: Bidi (Sarawak, Malaysia) TYPICAL MATERIAL: holotype M! in B; 1 paratype M! M!

smooth,

with a few keels; distal part of radial

in WA;

1 paratype

in OL.

HOSTS: unknown DISTRIBUTION: Oriental B! OL!

Forest Camp

Udawattakele Sanct.

region:

MALAYSIA:

(N of Kalabakan, (Kan.

Dis!.),

WA!

Bidi

Sabah),

(Bau District, B!

SRI

LANKA:

Sarawak),

Anteoninae

Fig.

447

Male genitalia of Anteon bauense n.

284 -

(Sabah):

sp.

from Forest Camp

left half removed.

21. Anteon debile n. sp.

FEMALE: MALE:

unknown

fully

legs fully

in following smooth,

winged;

weak,

weakly punctate,

9:

4:

6:

5:

5:

5:

mandibles, 5:

5:

5:

7;

antennae and antennal

lateral

segments

head shiny,

without sculpture among punct<.1res;

frons without median and

keels;

occipital

POL = 4;

OL

= 4;

OPL = 2; TL = 3; scutum, scutellum and metanotum shiny, weakly punctate, without sculpture among punctures; notaulices

incomplete,

reaching approximately 0,65 length of scutum;

propodeum reticulate rugose, dorsal

black;

antennae not distally thickened;

proportions:

carina complete; 2; OOL smooth,

length 1,56 mm;

testaceous;

with a strong

and posterior surface;

median area shiny,

smooth,

transversal

keel

between

posterior surface with 2 longitudinal

with a few keels near margins;

lateral

keels; areas

448

Mem.

En!.

Inst.,

no. 37,

Male genital ia of Anteon debile n.

285 -

Fig.

Amer.

1984

sp.

(type):

rigth half

removed.

shiny,

with a

few keels;

bands;

distal

part of radial

fore-wing hyaline,

without dark transversal

part (3:7); 285) shorter than penis, without proximal membranous process and without distal inner process; tibial spurs 1,1,2. LOCUS TYPICUS: Ban Van Eue (Laos) TYPICAL MATERIAL: holotype M! in B. HOSTS: unknown DISTRIBUTION: Oriental region: LAOS: Ban Van Eue (Vientiane), B! gonoforceps

vein much shorter than proximal

(Fig.

22. Anteon philippinum n. sp.

FEMALE:

fully

winged;

length 3,00 mm;

black;

mandibles testaceous;

antennae and legs fully testaceous; abdomen brown; antennae distally thickened; antennal segments in following proportions: 15: 7: 8: 7: 8: 7:

7:

7:

10;

median keel; complete;

head shiny,

two keels are present around orbits;

POL = 8;

OL = 4;

fully reticulate rugose, which

OOL = 6;

OPL = 5;

7:

frons with a complete occipital

TL = 5;

carina

pronotum shiny,

except for a narrow area near posterior margin

is smooth and without sculpture;

tegulae; shiny,

fully reticulate rugose;

posterior tubercles reaching

posterior surface of pronotum shorter than scutum

fully sculptured by weak

longitudinal

keels;

(7:20);

scutum

notaulices incomplete,

Anteoninae

Fig.

286 -

Chela of

449

Anteon philippinum

n.

sp.

(type)

reaching approximately 0,5 length of scutum; scutellum shiny, smooth, without sculpture; metanotum reticulate rugose; propodeum reticulate rugose, wi th a strong transversa I keel between dorsa I and posterior surface; posterior surface with two longitudinal keels; median area as rugose and dull transversal part

(4:11);

as lateral

bands;

distal

fore tarsal

areas;

fore-wing hyaline,

part of radial

vein

without dark

much shorter than proximal

segments in following proportions 10:

4:

4:

5:

450

Mem.

Amer.

Ent.

Inst.,

37,

no.

1984

17; enlarged claw (Fig. 286) with one bristle on a proximal prominence; segment 5 of front tarsus (Fig. 286) with two rows of nearly 25 lamellae; group of nearly 4

apex with a MALE:

lamellae;

tibial

spurs 1,1,2.

unknown

LOCUS TYPICUS:

Camp Lookout

TYPICAL MATERIAL: HOSTS:

(Philippines)

holotype F!

in AM.

unknown

DISTRIBUTION: Oriental Negros lsi.), AM!

region:

PHILIPPINES:

Camp Lookout

(Dumaguete,

ANTEON

NEARCTIC SPECIES OF

Sixteen species are known.

KEY TO THE SPECIES

FEMALES

5 of front tarsus with basal part longer than apical 291); segment 4 of front tarsus at most 0,5 as long front tarsal segments 3 or 4 produced into a hook ••••••• 2

Segment part as

1;

(Fig.

Segment

5 of

than apical 0,66 as

long as

produced 2

front part

tarsus wi th basal

303);

(Fig. segment

1;

usually

part

as

4 of

front

front

long as

or

shorter

tarsus at

tarsal

segment

least

2

into a hook ••••••••••••••.••••.•••••••••••••••••••••• 4

Posterior

surface of

propodeum without

• •.•••.••...••••••.•..••.•••• Posterior

3

segment

surface of

Notaulices

reaching

longitudinal

keels ••••••

1. popenoei (Ashmead)

propodeum with approximately

. . . . . . . . . . , . . • . . . . . . . . . . . . . . • 2.

two

longitudinal

keels •••• 3

0,5

length of scutum ••••••••• arizonense R.C.L. Perk i ns

reaching at most 0,3 length of scutum ••••••••••.•••• · . . • . • . . . . . . . . . • . • • • • . . . . . . . . 3. canadense (Ashmead)

Notaul ices

4 5

Posterior

surface of

propodeum without

Posterior

surface of

propodeum with

Prothorax black •••••••••••••• Prothorax

6

reddish •••..•••••••

Fore and hind wings

fully

darkened, 7

Antennal

hind wings hyal ine; but

longitudinal

keels ••••• 5 keel s •••• 6

iunestum (R.C.L. Perkins) wasbaueri n. sp.

dark •••••••••••••••••••••••••••••••••

. . . . . . . . . . . . . . . . . . . . . . . . . . . . .. Fore and

5. 6.

longitudinal

two

4. at

nebulosum n. sp. most

in

that

case hind-wing

segment

2 as

long as

• • • • • • • • • • • • • • • • • • • • • • • . • • • ••

segment 7.

fore-wing

51 ight Iy

always hyal ine ••••••••••• 7 3 ••••••••.••••••.••••••••

minimum (F en t on)

451

Anteoninae

Antennal

8

Segment

9

segment

4 of 4 of

Segment

2

shorter

than

segment

front

tarsus nearly

front

tarsus nearly as

3 ••••••••••••••••••••• 8

twice as

long as

long as or

segment

sl ight Iy

1. .. 9

longer

than

segment

1 •.•••.....••..•..••.•....••••..•••.•••.•••••••. 10

Head

smooth,

weakly

thorax mostly Head

rugose,

around

punctate,

testaceous ••••• punctate,

frons and

wi th

fully granulated,

Head

shiny,

punctate,

11 Pronotum fully punctures;

without

smooth,

head and

thorax black ••••••••••••••

rugosiceps K i ef f er

9.

punctate and wi th

or wi thout

10. puncticeps Ashmaead sculpture among punctures.aSW •. l 1

weakly punctate,

wi thout

sculpture among

thorax black wi th prothorax yellow •••••

• • • • • • • • • • • • • • • • • • • • • • • • • • •• Pronotum rugose,

head and

irregular keel s mostly

head and

irregular keels ••••••••••••.

irregular keels;

masoni n. sp.

8.

numerus

on vert ex;

• • • • . • • • • • • • • • • • • • • • • • • • • • • • ••

10 Head dull,

without

xanthothorax (B rad I ey )

1 1.

irregular keels on anterior half and on posterior half; sometimes more strongly rugose; thorax differently colored ••••••••••••••••••••••••••••

punctate head and

wi th

12. osborni (Fent on)

A. durangoi n. sp., hirtifrons Whittaker, ciudadi n. menkei n. sp. are unknown.

The females of sp.

and

MALES

2

Posterior

surface of

propodeum without

Posterior

surface of

propodeum wi th

Gonoforceps with distal

longitudinal

two

inner pointed

longi tudinal

process

(Fig.

keels ••••• 2 keels •••.

4

308 B) •••••

1 3. durangoi n. sp. inner pointed process

. . . • . . . . . . . . . . . . . . . . . . . . . . ..

(Figs. 288 297) •••••.•••••••••••••••••••.••••••••••••.••••••••••••• 3

Gonoforceps without

A, 3

B,

Propodeumwith straight

4

short

transversal

keel

line ••••••••••••••••

Propodeum with a

distal

transversal

Gonoforceps without

distal

forming

a

short

funestum (R.C.L. Perkins) 6. wasbaueri n. sp.

keel

line •••••••••••••••••

centrally

5.

centrally

forming

an

angle,

not

1. popenoei (Ashmead)

inner pointed process

(Figs.

302,

308 A) ••••••••••••••••••••••••.•••••.•••••••.••••••••••••••••• 5 Gonoforceps with distal inner pointed process (Fig. 293

A,

B, C) •••••••.••••••••••••••••.•••••••••.•••.••••••••••••••• 6

have been unable to find characteristics for separating males of the two species

452

5

Mem. Amer.

Head weakly punctate,

Ent.

smooth,

punctures ••••••••••••••••••• Head

6

7

8

9

10

11

12

13

smoot h ••••••••••••• "

The males of A.

dull,

no.

shiny, 14.

37,

1984

wi thout

ciudadi n.

sculpture among

sp.

or strongly granulated, 9. rugosiceps K i ef f er Gonoforceps much short er than pen is (F i g. 293 A, B) ••••••••••• 7 Gonoforceps as long as or longer than penis (Figs. 293 C, 309). • ., ••••••••••••••••••••••••••••••••••••••••••••• " •••••••••••• 8 Gonoforceps with small distal inner process (Fig. 293 A) ••••••• . • . . . • • . . . . . . . . . . . . . . . . . . . . . .. 3. canadense (Ashmead) Gonoforceps wi th large distal inner process (Fig. 293 B) ••••••• 16. hirtifrons Whi ttaker Posterior surface of propodeum with indistinct longitudinal keel s •••••••••••••••••••••••• 8. masoni n. sp. Posterior surface of propodeum with distinct longitudinal keels • •••••.••••••• " ••••••••••••••••••..••••.•• '" ••.••••.•••••••• 9 Head sparsely punctate, wi thout sculpture among punctures •••• 10 Head densely punctate, weakly or strongly granulated among punctures, wi th strong or weak areolae or irregular keels •••• 12 Face with anterior half testaceous; posterior half black ••••••• • • • • • • • • • • • • • • • • • • • • • • • • • • •• 15. menkei n. sp. Face fully black ••••••••••••••••••••••••••••••••••••••••••••• 11 Legs testaceous, at most with hind coxae and hind femora partly brown ••••••••••••••••••••••• 12. osborni (Fenton) Legs brown, with tarsi light and sometimes also tibiae testaceous •••••••••••••••••• 11. xanthothorax (Bradley) Scutum dull, rugose, fully strongly punctate, wi th some irregular keels near margins ••••••••••••••••••••••••••••••••••• • • • • • • • • • • • • • • • • • • • • • • • • • • •• 10. puncticeps Ashmead Scutum shiny, punctate, without sculpture among punctures, at most wi th anterior half rugose and dul I •••••••••••••••••••••• 13 Distal inner pointed process of gonoforceps extended medially and with apical margin excavated (Fig. 290) •••••••••••••••••••• . . • • . . . • • . • • . . . . . . . . . . . . . . . . . 2. arizonense R.C. L. Perkins Distal inner pointed process of gonoforceps extended apically and with apical margin convex or straight (Fig. 293C) •••••••••• • • • • • • • • • • • • • • • • • • • • • • • • • • •• 10. puncticeps Ashmead not

reticulate rugose,

Inst.,

weakly

••••

nebulosum n. sp. and minimum (Fenton) are

unknown.

1. Anteon popenoei (Ashmead)

Dryin us popenoei Ashmead 1888: i. Anteon unifasciatus Ashmead 1893: 95 (n. syn.); locus typicus

Anteoninae

Biscayne Bay

(Florida,

U.

S.

453

A.).

Anteon minutus Ashmead 1893: 97 (n. syn.); locus typicus Washington

Anteon Anteon Anteon Anteon

FEMALE:

fully

antennae fully legs fully

(D.

C.,

U.

winged;

3:

3:

testaceous;

4,5;

or areolae;

A.).

length 1,5-3,0 mm;

black;

mandibles testaceous;

testaceous or with segments 5-10 brown; antennae distally thickened;

following proportions: 9: 4: 3:

s.

popenoei (Ashmead): Ashmead 1893: 98. minutus Ashmead: Krombein 1979: 1241. popenoei (Ashmead): Krombein 1979: 1241. unifasciatus Ashmead: Krombein 1979: 1241.

head dull,

5:

2:

3:

granulated,

occipital carina complete;

keel

and with 2 strong or weak

Fig.

287 -

Chela of

3:

3:

3:

4:

tegulae testaceous;

antennal 6;

or 8:

with weak or strong frons with a median

lateral

keels;

POL = 5;

segments in

3:

3:

3:

3:

3:

irregular keels longitudinal

OL = 3;

OOL = 4;

Anteon popenoei (Ashmead) from Biscayne Bay (Florida)

454

OPL

Mem.

=

2;

rai sed

=

TL

into a

than scutum

no.

37,

1984

with posterior surface convex and

posterior surface of pronotum shorter

pronota I tubercles reach i ng tegu I ae;

notaul ices

incomplete,

reaching

scutellum and metanotum shiny,

posterior surface; hya line,

Inst.,

pronotum rugose,

propodeum reticulate rugose,

beneath

Ent.

transversa I cari na; (4: 11);

granulated; of scutum;

4;

Amer.

with a

smooth,

transversal

posterior surface without distal

part of radial

length

without sculpture;

keel

between dorsal

longitudinal

wi thout dark transversa I bands or with the pterostigma;

scutum du II,

approximately 0,25-0,30

keels;

and

fore-wing

1 dark transversal

band

vein shorter than proximal

part (2:4 or 2:5); fore tarsal segments in following proportions: 5: 2: 1,5: 2: 7; enlarged claw (Fig. 287) with one bristle on a proximal prominence; segment 5 of front

tarsus

longer than apical

part,

to the apex;

spurs 1,1,2.

MALE:

fully

testaceOU5 j

segment with

tibial winged;

287)

with basal

1 row of 7-8

length 1,31-2,00 mm; tegu I ae testaceous;

and fore tibiae

testaceous;

part sl ightly or much

lamellae extending continuously

black or brown;

antennae brown or brown-testaceous j

1 testaceous;

tarsi

(Fig. with

mandibles

somet i mes brown

legs fu II y

wi th

testaceous or brown

antennae not distally

thickened;

B

Fig.

288 -

Anteon funestum osborni (Fenton) (C):

Male genitalia of (Ashmead) (B),

(R.C.L.

Perkins)(A),

right half removed.

popenoei

Anteoninae

antennal 5,5;

segments in following proportions:

head dull,

areolae;

granulated,

8:

4,5:

complete;

kee!

and with or without

POL = 5;

Ol = 3;

4:

3:

3:

3:

3:

3,5:

4:

with weak or strong irregular keels or

sometimes strongly reticulate rugose;

longitudinal smooth,

455

frons with a median

lateral

OOl = 4;

keels;

OPl = 3;

occipital

Tl = 4;

carina

scutum shiny,

weakly punctate and without sculpture among punctures or weakly

granulated;

rarely scutum strongly granulated;

reachi ng approxima tel yO, 3 I ength of scutum; shiny,

smooth,

without sculpture or weakly

notaulices incomplete,

scute II um and metanotum

granulated;

propodeum

reticulate rugose, with a transversal keel between dorsal and posterior surface; posterior surface without longitudinal keels; transversal keel of propodeum centrally forming an angle; transversal

bands;

part

gonoforceps (Fig.

(2:5);

distal

tibial spurs 1,1,2. lOCUS TYPICUS: Marlatt

fore-wing hyaline,

part of radial 288 B)

(Kansas,

without distal

U.

S.

without dark

vein much shorter than proximal inner pointed process;

A.)

popenoei in WA; holotype F! of A. minutum in WA; holotype F! of A. unifasciatus in WA. TYPICAL MATERIAL:

HOSTS:

holotype M!

of D.

unknown

DISTRIBUTION:

Nearctic region:

CANADA:

Scandia

(Alta.),

OT!

Lethbridge

(Alta.), OT! Ottawa (Ont.), OT! Mer Bleue (Ottawa, Ont.), aT! Ol! Britannia (Ont.), aT! OL! Cayuga (Ont.), OT! Beech Grove (Que.), OT!

u.

Ol! Kouchibouguac N. P. (N.B.), OT! Plas~r Rock (N.B.), OT! S. A.: Mc Kittrick Canyon (Culberson Co., Tex.), TW! Nebraska, WA! Marlatt (Kans.), (Ga.),

WA!

OT!

Wattacoo OT!

Ol!

Rapides Parish

Rabun Bald

(Pickens Co., S.C.), Washington

Research Station

(D.

(la.),

TW!

(Rabun Co., C.),

(Huntington,

TW! WA!

Biscayne Bay

Ga.),

OT!

Greenvi lie Farmingdale

long

Island,

N.

(Fla.),

Cleveland (S.C.),

TW!

(N.Y.), Y,),

(S.

Ol!

Forsyth OL!

Di Ilwyn

TW!

AM!

WA! C.),

TW!

(Va.),

Kalbfleisch

Ol!

2. Anteon arizonense R.C.l. Perkins

Anteon arizonensis R.C.l. Perkins 1907: 51. Anteon arizonensis R.C.l. Perkins: Krombein 1979: 1240.

FEMALE: yellow,

fully

winged;

length 2,56 mm;

with teeth brown;

black;

abdomen brown;

antennae yellow;

legs yellow,

mandibles

with hind coxae

and hind clubs of femora brown; body shiny, hairy on sides of thorax and on front part of vertex; head rugose, strongly punctate; pronotum short, more than

twice as

long as broad

(5:1),

as sculptured as head;

scutum

smooth, without punctures, without sculpture; notaulices incomplete, reaching approximately 0,5 length of scutum; scutellum and metanotum smooth;

propodeum reticulate rugose,

between dorsal

and posterior surface;

with an

irregular transversal

keel

posterior surface with 2 straight

456

Mem. Amer.

Fig.

289 -

Ent.

Inst.,

no.

37,

Chela of Anteon arizonense R.C.L.

1984

Perkins from Vermilion Bay

(Ontario) •

longitudinal hyaline,

keels;

median area as rugose as lateral

without dark transversal

much shorter than proximal; segment 4

(35: 10);

enlarged claw

segment 5 of front

distal

part;

tibial

MALE:

fully

winged;

darkened; distal

(Fig.

289)

areas;

fore-wing

part of radial

tarsus much

vein

longer than

with one bristle on a proximal

tarsus with basal

part much

longer than

spurs 1,1,2. length 2,10-2,37 mm; black; mandibles testaceous; with segments 2-3 and apical region of segment 1

testaceous,

legs testaceous,

with hind coxae,

part of hind tibiae brown;

antennae not distally thickened; proportions:

with distal

segment 1 of front

prominence;

antennae

bands,

10: 5:

strongly punctate,

6:

6,5:

7:

antennal

6,5:

segments in following

6,5: 7:

mostly on frons,

to areolae; occipital

stalk of hind femora and

sometimes thorax and propodeum reddish; 6,5:

9;

head dull,

granulated,

where the wide punctures are similar

carina complete;

frons with a median keel,

hairy,

without lateral keels; POL = 7; Ol = 3,5; OOl = 5; OPl = 3; Tl = 3; scutum, scutellum and metanotum shiny, smooth, without sculpture or very weakly punctate, without sculpture among punctures; notaulices incomplete, reaching approximately 0,5 length of scutum; propodeum reticulate rugose, with a strong

transversal

keel

between dorsal

and posterior surface;

posterior surface with 2 longitudinal

keels;

as

without dark transversal

lateral

distal

areas;

fore-wing hyal ine,

part of radial

gonoforceps (Fig. with a distal

290)

median area nearly as rugose

vein much shorter than without dorsal

proximal

proximal

part

membranous process and

inner pointed process extended medially and with apical

margin excavated; tibial spurs 1,1,2. LOCUS TYPICUS: Nogales (Arizona, U. S.

A.)

TYPICAL MATERIAL: lectotype F! and 1 para lectotype M! Olmi 1982) of A. arizonense in B. HOSTS:

in U.

S.

A.

(designated by

Cicadellids.

DISTRIBUTION:

Nearctic region:

CANADA:

OT!

(B.C.), OT!

Cowichan Bay

Crowsnest

bands;

(4,5:7);

Ol!

Hixon

(B.C.), OT! (Vancouver

Terrace

Is.,

(B.C.),

B.C.),

OT!

457

Anteoninae

Sandspit (Moresby lsi., Queen Charlotte Islands, B.C.), AM! Twin Buttle (Alta.), OT! Waterton (Alta.), OT! Ol! Cooking lake (Alta.), OT! Elkwater (Alta.), OT! Jumping Pd. Cr. (Calgary, Alta.), OT! Maple Creek (Sask.), OT! Ol! Ninette (Man.), OT! Stittsville (On!.), OT! Vermilion Bay (On!.), OT! OLl Marmora (On!.), OT! Mistassini (Que.), OT! Wood R.-Wood l . (Baie Comeau, Que.), OT! U. S. A.: Brinnon (Jeff Co., Wash.), TE! Idaho City (Idaho), OL! TW! Crescent City (Calif.), TW! Tuscarora (Nev.), Ol! TW! OLl

Nogales (Ariz.),

B!

Cleveland

(S.

C.),

OLl

TW!

lockport

(N.Y.),

TW!

3. Anteon canadense (Ashmead)

/

1/

~

"-

I

I

"

,.I

I

1 / ! l-

Fig.

290 -

i

Male genitalia of Anteon arizonense R.C.l. lockport

(New York):

right half removed.

Perkins from

458

Mem. Amer. En!.

Inst.,

no. 37,

1984

basal part

Fig. 291 - Chela of Anteon canadense (Ashmead) (type)

Chelogynus canadensis Ashmead 1893: 93. Anteon politus Ashmead 1893: 96 (n. syn.); locus typicus Toronto (Ontario, Canada).

Anteoninae

459

Anteon pallidicornis Ashmead 1893: 97 (n. syn.); locus typicus Utah Lake

(Utah,

U.

S. A.).

Chelogynus canadensis Ashmead: Fenton 1918b: 272. Anteon canadensis (Ashmead): Muesebeck and Walkley 1951: 1040. Anteon (Anteon) canadensis (Fenton): Krombein 1979: 1240. Anteon (Anteon) pallidicornis Ashmead: Krombein 1979: 1241. Anteon (Anteon) politus Ashmead: Krombein 1979: 1241.

FEMALE:

fully

winged;

antennae fully

length 2,25-2,75 mm;

brown or testaceous,

black;

mandibles testaceous;

sometimes brown

with segment

1

testaceous; legs fully testaceous, with hind coxae basally black; antennal segments in following proportions: 10: 4,5: 5: 5: 5: 4: 4: 4: 4: 6; head shiny,

granulated or not granulated,

centra I area of the frons smooth frons,

region behind ocelli,

and granulated; ocell i; =

4;

occipital

fronta I

POL = 7;

pronotum du II,

pronotal

weakly punctate,

scutum;

sculpture among punctures;

keel

Fig.

292 -

also among

OPL = 4;

smooth on I y

tegulae;

TL

near

scutum shiny,

scutum sometimes with areolae near

keels;

smooth,

propodeum reticulate

between dorsal

Chela of

rugose,

OOL = 5;

reaching approximately 0,3-0,5

scutellum and metanotum shiny,

with 2 longitudinal

OL = 4;

tubercles reaching

notaulices incomplete,

transversal

less wide

anterior ha If of

strongly punctate only near anterior margin,

without sculpture among punctures; margins;

more

with numerous irregular keels,

carina complete;

posterior margin; smooth

with a

punctate;

sometimes also sides of frons weakly rugose

sometimes head

line present;

usually

and weak I y

length of

weakly punctate, rugose,

and posterior surface;

median area as rugose as,

with

an

without irregular

posterior surface or

less rugose

than

Anteon canadense (Ashmead) from Toronto (Ontario):

variety

with

lamellae on tarsal

segment 5.

460

Mem.

Fig.

293 -

Amer.

Male genital ia of Whittaker

lateral

areas;

part of radial segments

fore-wing

En!.

Ins!.,

no.

37,

Anteon canadense (Ashmead) (A), hirtifrons puncticeps Ashmead (C).

(B),

hyal ine,

without dark transversal

vein much shorter than proximal

in following proportions:

tarsus produced

into a hook;

1984

6:

1,5:

1,5:

enlarged claw

part 2:

(Figs.

9;

bands;

(3:10);

distal

fore tarsal

segment 3 of front

291,

292)

with one

bristle on a proximal prominence; segment 5 of front tarsus (Figs. 291, 292) without medial lamellae and with bristles only, or with 3 lamellae; apex with a than distal MALE:

group of 2-5 part;

fully

tibial

winged;

lamellae;

segment 5 with basal

part much

longer

spurs 1,1,2.

length 2,62 mm;

black;

mandibles testaceous;

antennae brown; legs testaceous, with hind coxae basally black; antennae not distally thickened; antenna I segments in following proportions: 10: 5: 7: 8: 8: 8: 8: 8: 8: 11; head shiny, strongly punctate, without sculpture among punctures, with numerous irregular keels mostly on vertex and among ocell i;

frons

with

an

incomplete median

keel

visible only

in front of

anterior ocellus; occipital carina complete; POL = 5; Ol = 3; OOl = 7; = 4; Tl = 5; scutum shiny, weakly punctate, without sculpture among punctures,

strongly

OPl

punctate and almost rugose only near anterior margin;

notaul ices incomplete, reaching approximately 0,5 length of scutum; scutellum and metanotum shiny, smooth, weakly punctate, without sculpture among punctures; keel

between

longitudinal

propodeum ret icu I ate rugose,

dorsal keels;

and posterior surface;

with a

strong transversa I

posterior surface with 2

median area as rugose as

lateral

areas;

fore-wing

Anteoninae

461

slightly darkened; hind-wing hyaline; distal part of radial shorter than proximal part (5:11); gonoforceps (Fig. 293 A) pointed process; gonofoceps much shorter than penis; LOCUS TYPICUS: Ottawa (Canada)

vein much with a distal

tibial

spurs 1,1,2.

TYPICAL MATERIAL: holotype F! of Ch. canadensis in WA; holotype F! of A. pallidicornis in WA; holotype F! of A. politus in WA. HOSTS:

unknown

DISTRIBUTION:

Nearctic region:

Terrace (B.C.), OT! Crowsnest

(B.C.),

Burnaby

OT!

C.),

Mac Gillivray

Zymoetz

(Terrace,

B.C.),

(Alta.),

OT!

View

Mtn.

CANADA:

(B.

OT!

Gagnon Road

OT!

Shames

(Game Reserve,

Upper Hay River Post

(Alta.),

OT!

Ninette

(Terrace,

(Terrace,

B.C.),

B.C.),

Chilliwack, (Alta.),

(Man.),

OT!

B.C.),

CA!

OL!

OT!

OT! OT!

Morley

Ottawa

(Ont.), WA! Innisville (Ont.), OT! Toronto (Ont.), WA! Stittsville (Ont.), OT! Merivale (Ont.), OT! Nominingue (Que.), OT! OL! Laniel (Que.), OT! U. S.

A.:

Lowman

(Calif.), TE!

OL!

OT!

TW!

Idaho City

(Calif.),

OT!

Washoe Lake (Washoe Co.,

Estes Park

(Colo.),

Co.

(Mich.),

OL!

Maryland

Long

(Idaho),

Victorville

OL!

Island,

OT!

Golden

I ndian Gap

Nev.),

(Colo.),

TE!

OT!

OL!

OL!

TW!

Mc Cann

(Placer Co.,

Utah Lake

(Great Smoky Mt.

(Krombein 1979); N.Y.),

(Idaho),

Goose Meadow

Bena

(Minn.),

N.P.,

Calif.),

(Utah),

N.C.

-

Kalbfleisch Research Station

WA!

OT!

Benzie

Tenn.),

OT!

(Huntington,

AM!

4. Anteon nebulosum n. sp.

FEMALE:

fully

reddi sh-dark; testaceous;

winged;

length 3,31

mm;

antennae testaceous,

black;

mandibles and clypeus

with segments 1-4 brown;

antennae distally sl ightly thickened;

following proportions:

11: 7:

8,5:

8,5:

7:

6,5:

antennal

6:

6,5:

legs

segments in

6,5:

8;

head dull,

granulated and punctate,

mostly near clypeus;

keel

keels around orbits directed towards antennal

and with two

lateral

frons with a short median

sockets; occipital carina complete; POL = 5; OL = 4; OOL = 6; OPL = 5; = 6; pronotum dull, strongly sculptured by transversal keels; pronotal tubercles reaching tegulae; scutum

(10:18);

punctures; scutum;

posterior surface of pronotum shorter than

scutellum shiny,

notaul ices incomplete,

punctate,

without sculpture among

reaching approximately 0,65 length of

scutellum and metanotum shiny,

propodeum reticulate rugose, posterior surface;

TL

smooth,

with a transversal

without sculpture; keel

between dorsal

posterior surface with 2 longitudinal

keels;

and

median area

as rugose as lateral areas; wings fully darkened; fore-wing with distal part of radial vein much shorter than proximal part (7: 14); fore tarsal segments in following proportions: 294)

with one bristle on a

(Fig.

294)

tibial

spurs 1,1,2.

MALE:

10:

proximal

4:

with 2 rows of 2+23 lamellae;

unknown

4,5:

13:

prominence;

25;

enlarged claw

segment 5 of front

apex with a group of 4

(Fig.

tarsus

lamellae;

462

Mem. Amer.

Fig. 294 -

En!.

Ins!.,

no.

37,

1984

Chela of Anteon nebulosum n.

sp.

(type)

LOCUS TYPICUS: Lyme (Connecticut, U. S. A.) TYPICAL MATERIAL: holotype F! in WA. HOSTS: unknown DISTRIBUTION: Nearctic region: U. S. A.: Lyme (Connecticut),

5. Anteon funestum (R.C.L. Perkins)

WA!

Anteoninae

463

Chelogynus funestus R.C.L. Perkins 1907: 48. Chelogynus melanacrias R.C.L. Perkins 1907: 49 (n. syn.); locus typicus Nogales (Ariz. U. S. A.). Chelogynus lusus R.C.L. Perkins 1907: 50 (n. syn.); locus typicus Tucson (Ariz., U. S. A.). Chelogynus funestus R.C.L. Perkins: Muesebeck and Walkely 1951: 1040.

Anteon (Chelogynus) funestus (R.C.L. Perkins): Krombein 1979: 1241.

Anteon (Chelogynus) lusus (R.C.L. Perkins): Krombein 1979: 1241. Anteon (Chelogynus melanacrias (R.C.L. Perkins): Krombein 1979: 1241.

FEMALE:

fully

winged;

antennae yellow, segment 10 brown; femora brown;

length 1,83-2,59 mm;

black;

mandibles yellow;

with segments 6-10 or 7-10 brown; tegulae testaceous;

sometimes all

legs yellow,

occasionally only with stalk of hind

coxae, clubs of femora and tibiae brown;

occasionally only clubs of femora and hind tibiae brown; segments in following proportions: shiny,

granulated,

frons with one

11:

5:

8:

3:

4:

4:

4:

antennal 4:

4:

6;

head

with a few or numerous irregular keels or areolae;

longitudinal

towards antenna I sockets;

median keel

POL

=

5;

OL

and 2 keels near orbits directed

=

3;

OOL

=

6;

OPL

=

4;

TL

=

6;

occipital carina complete; pronotum granulated, except for an area near posterior margin which is smooth and without sculpture; pronotal tUbercles

Fig.

295 -

Chela of Anteon funestum

(R.C.L.

(Arizona)

Perkins)

from Nogales

464

Mem.

reaching

tegulae;

Amer.

scutum shiny,

notaulices incomplete and approximately 0,20 metanotum shiny, with an

part

(2:5);

37,

1984

keel

distal

scutellum and

segments

and posterior surface;

fore-wing hyaline,

part of radial

fore tarsal

smooth;

propodeum reticulate rugose,

between dorsal keels;

granulated,

reaching only

of scutum;

without sculpture; longitudinal

bands;

no.

very reduced,

(rarely 0,30)

length

smooth,

irregular transversal

proximal

Ins!.,

weakly or strongly

usually

posterior surface without dark transversal

En!.

without

vein much shorter than

in following proportions:

5:

2:

2: 5:

14; enlarged claw (Fig. 295) with one bristle on a proximal prominence; segment 5 of front tarsus (Fig. 295) with 2 rows of nearly 16 lamellae extending continuously

to the apex;

shorter than distal

spurs 1,1,2.

MALE:

fully

part;

winged;

teeth brown;

tibial

length

antennae fully

coxae,

clubs of femora and

shiny,

granulated,

longitudinal

TL = 3;

scutum weakly granulated;

keel

with

tibiae

tegulae testaceous;

legs yellow,

(except for the fore tibiae)

occipital

smooth,

not

notaulices incomplete,

head

frons with a

POL = 5;

OL = 3;

granulated; very

OOL

occasionally

reduced,

reaching

scutellum and metanotum shiny,

propodeum reticulate rugose,

between dorsal

brown;

keels near orbits directed

carina complete;

length of scutum;

without sculpture;

transversal

with

brown;

scutum shiny,

only approximately 0,25 smooth,

1,64-1,90 mm; black; mandibles yellow,

and with 2 lateral

median keel

OPL = 2;

part of segment 5

with some irregular keels or areolae;

towards antenna I sockets; = 4;

basal

and posterior surface;

with an

irregular

posterior surface

without longitudinal keels; propodeum with transversal keel centrally forming a short straight line; fore-wing hyaline, without dark transversal bands;

distal

part of radial

vein much shorter than proximal

288 A) without dorsal membranous process, not pointed distal inner process; tibial spurs 1,1,2. gonoforceps

(Fig.

LOCUS TYPICUS:

Tucson

TYPICAL MATERIAL:

(Arizona,

U.

(2:5);

S. A.)

and 5 paralectotypes

lectotype F!

part

with a wide,

(3 FF, 2 MM)!

iunestus in B; lectotype F! and 1 paralectotype F! (designated by Olmi 1982) of Ch. melanacrias in B. lectotype F! and 1 paralectotype M! (designated by Olmi 1982) of Ch. lusus in B. (designated by Olmi

1982) of Ch.

HOSTS: in U. S. A. Jassids. DISTRIBUTION: Nearctic region: OT!

U.

S.

(Ariz.),

A.:

OL!

Tucson

TW!

(Ariz.),

Ramsey Cyn.

CANADA: B!

CAl

(Sierra Vista,

Cochise Co.

(Portal,

Co.,

TE!

Gainesville (Fla.),

TW!

Monterey

Tex.),

(S.C.),

OL!

Kouchibouguac N.P.

Nogales (Ariz.),

Chiricahua Mts., (Va.),

OT!

Ariz.),

OT! OU

OL!

B!

Huachuca Mts., SCI

(N.B.),

WA!

Ariz.),

College Station

Forsyth

Takoma Pk.

(Ga.), (Md.),

OL!

Portal

OT! OL!

OT!

(Brazos Cleveland TW!

BERMUDA IS.: Bermuda, OT! MEXICO: La Ciudad (Durango), OT! OL! EI Saito (Durango), OT! OU Monterrey (Nuevo Leon), OL! TE! Teziutlan ·(Puebla),

OT!

6. Anteon wasbaueri n. sp.

Anteoninae

Fig.

296 -

winged;

465

Chela of Anteon wasbaueri n.

FEMALE:

fully

reddish;

clypeus,

mandibles,

length 2,06 mm;

distally

thickened;

antennal

black,

antennae and segments

sp.

(type)

with prothorax fully

legs fully

testaceous;

in following proportions:

antennae

6:

3,5:

4,5: 4: 4: 3,5: 3,5: 3,5: 3,5: 5; head dull, granulated, with frons weakly sculptured by longitudinal keels; frons also with an incomplete median keel; occipital carina complete; POL ~ 4,5; Ol ~ 3; OOl ~ 4,5; OPl ~ 3;

Tl ~ 4; pronotum dull, granulated, weakly transversal striate, with posterior surface shorter than scutum (7:10); pronotal tubercles reaching tegulae;

scutum shiny,

sculpture; scutum;

notaul ices

smooth,

very

incomplete,

weakly

faintly

scutellum and metanotum shiny,

propodeum reticulate rugose, and posterior surface;

granulated or almost without

visible near anterior margin of smooth,

without sculpture;

with a strong transversal

posterior surface without

keel

longitudinal

between keels;

dorsal fore-

wing hyaline, without dark transversal bands; distal part of radial vein shorter than proximal part (1,5:8); fore tarsal segments in following proportions:

5,5:

bristle on a

proximal

with

1 row of 25

1:

2,5:

5:

10,5; enlarged claw

prominence;

lamellae;

(Fig.

segment 5 of front

apex with a

group of 5

296)

tarsus

lamellae;

with one (Fig. tibial

296) spurs

466

Mem.

Amer.

En!.

Inst.,

no.

37,

1984

\ \

\

A

Fig.

Male genitalia of Anteon wasbaueri n.

297 -

(Texas}(B:

left half removed);

A:

gonoforceps

sp.

from Speir Rch.

(laterally viewed).

1,1,2. MALE:

fully

winged;

length 1,75-2,81

mm;

black;

mandibles testaceous;

antennae brown; legs brown, with tarsi light; antennae not distally thickened; antenna I segments in following proportions: 7: 3: 3,5: 3: 4: 4: 4: 4: 6; head dull, granulated, and irregularly reticulate rugose; frons

with a complete median keel

and with

two

directed towards antenna I sockets;

occipital

carina complete;

=

3;

OOl

=

granulated;

5;

OPl

=

3.

notaul ices

Tl

=

3;

scutum shiny,

incomplete,

visible only

lateral

4:

keels around orbits POL = 6;

weakly or strongly near anterior margin of

Ol

Anteoninae

scutum;

scutellum and metanotum shiny,

propodeum reticulate rugose, and posterior surface; wing hyaline,

not pointed distal

MM!

part

(Uvalde,

holotype F!

bands;

(1,5:5,5);

inner process;

Speir Rch.

TYPICAL MATERIAL:

weakly granulated;

posterior surface without

much shorter than proximal wide,

smooth,

with a strong transversal

without dark transversal

LOCUS TYPICUS:

467

keel

distal

keels;

fore-

part of radial

gonoforceps

tibial

between dorsal

longitudinal (Fig.

297)

vein

with a

spurs 1,1,2.

Uvalde Co.,

Texas,

and 3 paratypes MM!

U.

S.

A.)

in SC;

3 paratypes

(Uvalde,

Uvalde Co.,

in OL.

HOSTS:

unknown

OISTRIBUTION: Texas),

SC!

Nearctic region:

U.

S.

A.

Speir Rch.

OU

the females of A. wasbaueri is clearly different from the funestum for its reddish prothorax, the males are a real headache. I was not able to find differences between males of A. wasbaueri and A. funestum. I fA. wasbaueri proves to be a jun ior synonym of A. funestum, then its female will be considered as a rare colour variety of AAA. funestum. NOTES:

whereas

A.

female of

The species

is named

in

honor of Mr.

Marius S.

Wasbauer.

7. An teon minimum (Fen ton)

Chelogynus minimus Fenton 1927: 13. Anteon (Chelogynus) minimus (Fenton): Krombein 1979: 1241.

FEMALE:

fully

winged;

antennae fully

length 2,00-2,50 mm;

black;

mandibles testaceous;

brown or with segments 1-2 testaceous;

abdomen brown;

legs testaceous or brown,

with

tarsi

tegulae testaceous; light;

antennal

segments in following proportions: 9: 4: 4: 3: 3: 4: 3,5: 3: 3: 5,5; head shiny, smooth, weakly punctate and alutaceous; frons with a longitudinal POL ~ 4;

median keel;

carina complete;

OL ~ 3;

OOl ~ 5,5;

pronotum shiny,

without sculpture on posterior half; scutum shi ny, notaul ices

smooth,

incomplete,

weak I y

pronotal

punctate, smooth,

with an

posterior surface;

posterior surface with 2

proportions: nearly 20

4:

2,5:

2:

prominence; lamellae;

part 4:

(2:6); 12;

bands;

keel

distal

enlarged claw group of 6

keels;

area;

(Fig.

and

median area

fore-wing

part of radial

segments

tarsus

propodeum

between dorsal

longitudinal

fore tarsal

segment 5 of front

apex with a

tegulae;

length of scutum;

with a smooth and shiny central

shorter than proximal a proximal

tubercles reaching

irregular transversal

without dark transversal

occipital

smooth and

without sculpture;

reticulate rugose,

hyaline,

TL ~ 4;

wi thout scu I pture among punctures;

reaching approximately 0,5

scutellum and metanotum shiny,

only partly rugose,

OPl ~ 4,5;

rugose on anterior half,

vein much

in following

298 with one bristle on (Fig.

lamellae;

298)

basal

with part of

row of

468

Mem. Amer. Ent.

Inst.,

no. 37,

1984

\

Fig. 298 - Chela of Anteon minimum (Fenton)(type)

Anteoninae

segment 5 shorter than distal MALE:

part;

469

tibial

spurs 1,1,2.

unknown

LOCUS TYPICUS:

Barneveld

TYPICAL MATERIAL: HOSTS:

(N.

Y.,

holotype F!

s.

U.

A.)

in WA.

unknown

DISTRIBUTION: Nearctic region: U. S. Kouchibouguac N.P. (N.B.), OT! OL!

A.:

Barneveld

(N.Y.),

WA!

CANADA:

8. An teon masoni n. sp.

FEMALE:

fully

scutellum,

winged;

metanotum,

reddish-brown;

length 2,31-2,50 mm;

abdomen

testaceous or brown;

scutum with reddish or brown spots; segments shiny,

in following proportions:

smooth,

occipital 3;

very

9:

5:

OOL = 6;

TL = 5;

6:

rugose on I y

tuberc les reachi ng

scutum,

tegu I ae;

5:

6:

5:

and

5:

lateral

pronotum shiny,

scu I pture among punctures, smooth,

6,5:

thickened; 5:

head and antennal

7;

head

without sculpture among punctures;

frons without median

OPL = 4;

with

mesopleura

somet i mes pronotum,

antennae distally

weakly punctate,

carina complete;

OL = 3;

reddish-testaceous,

propodeum and metapleura black;

keels;

punctate,

near anterior marg in;

POL = without

pronota I

scute II um and metanotum sh i ny,

without sculpture or weakly punctate and without sculpture among

punctures; scutum;

notaulices incomplete,

reaching

propodeum reticulate rugose,

approximately 0,65

with 2

longitudinal

length of

keels on posterior

surface; median area as rugose as lateral areas; fore-wing hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal

part

proximal

(4:10);

3:

11:

part

(4: 10);

19; enlarged claw

prominence;

distal

fore tarsal (Fig.

segment 5 of front

299)

apex with a group of 4

produced

into a

tibial

vein much shorter than

in following proportions:

6:

2:

with one bristle on a proximal

tarsus

lamellae;

hook;

part of radial segments (Fig.

lamellae;

299)

with

1 row of 24

segment 2 of front

tarsus

spurs 1,1,2.

MALE (not reared): fully winged; length 3,25 mm; black; mandibles testaceous; antennae black; legs testaceous, with coxae partly black and femora partly

brown;

antennae not distally

in following proportions: shiny,

11:

strongly punctate,

weakly granulated;

frons

sculpture among punctures; sculpture; transversal with

10:

10:

antennal

10:

10:

segments

11;

incomplete median keel, lateral

head

carina complete;

scutum shiny, notaul ices

POL = 7,5;

weakly punctate,

incomplete,

reticulate rugose,

reaching

between dorsal

with a

fore-wing

OL = 3,5;

without approximately

smooth,

without

track of a

and posterior surface;

tracks of 2 longitudinal keels;

visible only

keels directed

scutellum and metanotum shiny,

propodeum dull, keel

thickened;

10:

frons also with 2

occipital

TL = 5;

0,5 length of scutum;

10:

with a short

towards antenna I sockets; OPL = 5;

10:

without sculpture among punctures or very

in front of anterior ocellus; OOL = 8;

7:

hyaline,

posterior surface without dark

470

Mem.

Fig.

transversal (5:12);

bands;

Amer.

299 -

Chela of

distal

gonoforceps (Fig.

membranous process and

En!.

Ins!.,

distal

Ottawa

(Ontario,

wide,

dorsal

Canada)

part

proximal

inner pointed process;

1,1,2. LOCUS TYPICUS:

1984

vein shorter than proximal

with a very

with a

37,

Anteon masoni n. sp. (type)

part of radial 300)

no.

tibial

spurs

Anteoninae

Fig.

300 -

Male genitalia of

TYPICAL MATERIAL:

holotype F!

in TW; 1 paratype F! HOSTS: unknown DISTRIBUTION: (Ont.),

OLl

OL!

A.:

Corvall is

(Ont.),

OT!

Parke Reserve (Kam. (Oreg.),

Poughkeepsie (N.Y.),

OL! TW!

Anteon masoni n. sp. from Hull (Quebec)

and 5 paratypes FF!

5 paratypes FF!

Nearctic region:

Finland

OT!

in AM;

471

C,\NADA: Ottawa

TW!

(Ont.),

Summit of King Mt.

Co.,

Que.),

Highlands

OT!

in OT;

1 paratype F!

in OL.

OL!

(N.C.),

OT!

OL!

Rockport

(Old Chelsea,

Hull OL!

(Que.), Laurel

Que.),

OT!

(Md.),

U.

S.

OT!

4 mi. NNW Kerhonkson (Cherry town, Ulster Co.,

N.Y.),

AM!

NOTES:

the species is named in honor of Mr.

W.

R.

9. Anteon rugosiceps Kieffer

Anteon rugosiceps Kieffer 1906a: 238.

M.

Mason.

472

Mem. Amer.

Fig.

301

-

Chela of

En!.

Ins!.,

no.

37,

1984

Anteon rugosieeps Kieffer from Me Bride Springs (California)

473

Anteon i nae

Anteon rugosiceps Kieffer: Kieffer 1914b: 168. Anteon rugosiceps Kieffer: Muesebeck and Walkley 1951: 1040. Anteon rugosiceps Kieffer: Krombein 1979: 1241.

FEMALE

(not reared):

testaceous;

fully

winged;

antennae testaceous,

3,75-4,00 mm;

with hind coxae almost fully black; distally thickened; 9:

9:

9:

9:

9:

9:

antennal 11;

mandibles legs testaceous,

abdomen partly testaceous;

segments in following proportions:

head shiny,

numerous irregular keels, frons,

black;

with segments 6-10 brown;

not granulated,

antennae 12: 7:

11:

punctate and with

very numerous particularly on anterior part of

where there are numerous areolae;

frons with a complete median

keel; occipital carina complete; POL = 9; Ol = 6; OOl = 9; OPl = 7; Tl = 6; pronotum shiny, with anterior half rugose and posterior surface of pronotum shorter than scutum scutum,

(6:22);

pronotal

scutellum and metanotum shiny,

sculpture among punctures; 0,3 length of scutum; transversal

keel

part

(6:10);

reaching

propodeum reticulate rugose,

keels;

median area smooth,

distal

fore tarsal

part of radial

without

approximately

with a strong

and posterior surface;

at most with some areolae around margins; or less darkened;

weakly punctate,

notaul ices incomplete,

between dorsal

with 2 longitudinal

tubercles reaching tegulae;

smooth,

posterior surface

shiny,

without sculpture,

fore-wing with distal

half more

vein much shorter than proximal

segments in following proportions:

6:

6:

5:

13: 30;

enlarged claw (Fig. 301) with one bristle on a proximal prominence; segment 5 of front tarsus (Fig. 301) with 2 rows of 29 lamellae extending continuously to the apex; MALE:

fully

winged;

antennae brown;

legs brown,

thickened;

antennal

6,5: 7:

9;

7:

Ol = 3;

shiny,

black;

with tarsi

mandibles testaceous;

light;

antennae not distally 9:

5:

6:

7:

granulated and fully reticulate rugose;

and 2 lateral

OOl = 5;

smooth,

spurs 1,1,2.

segments in following proportions:

head dull,

with a median keel 5;

tibial

length 2,18 mm;

OPl = 4;

keels;

Tl = 4;

occipital scutum,

very weakly punctate,

7:

7:

frons

carina complete;

POL =

scutellum and metanotum

without sculpture among punctures;

notaulices incomplete, reaching at most 0,5 length of scutum; propodeum reticulate rugose, with a transversal keel between dorsal and posterior surface;

posteior surface with 2 longitudinal

as lateral distal

areas;

fore-wing hyal ine,

part of radial

gonoforceps (Fig. without distal

lOCUS TYPICUS:

302)

DISTRIBUTION:

Shasta,

holotype M!

Nearctic region:

Santa Clara Co. (Mt.

without dorsal

Santa Clara Co.

Kouchibouguac N.

P.

median area as rugose

vein much shorter than proximal

inner pointed process;

TYPICAL MATERIAL: HOSTS: unknown

keels;

without dark transversal

(N.

Siskiyou Co.,

CAl

bands;

(6:4);

membranous process and

spurs 1,1,2.

(California,

U.

S.

A.)

in CA. CANADA:

B.), OT!

(Calif.),

proximal tibial

part

u.

Onyx

Calif.),

S.

Dirleton A.:

(Cal if.), OT!

Ol!

(Ont.),

Pinehurst Ol!

OT!

OT!

Ol!

(Oreg.),

TW!

OL!

Mc Bride Springs

Washoe lake (Washoe Co.,

474

Mem.

Fig.

302 -

Amer.

Ent.

Inst.,

no.

37,

1984

Ma Ie gen i ta I i a of Anteon rugosiceps Kieffer (type)

Nev.), TE! OLl Parker Canyon Lk. (Ariz.), TW! Mt. Vernon Cn. Colo.), OT! Golden (Colo.), OT! OL! MEXICO: Ixtapan de La Sal OT!

10. Anteon puncticeps Ashmead

(Golden, (Mexico),

475

Anteoninae

Anteon puncticeps Ashmead 1893: 97. Aphelopus americanus Ashmead 1893: 100 (n. syn.); locus typicus Jacksonvi lie

(Florida,

U.S.

A.).

Chelogynus vivariensis Brad ley 1926: 7 (n. syn.); locus typicus Mc Lean Bogs Bioi.

Res.

(N.

Y.,

U.S.

A.).

Chelogynus virginiensis Fen ton 1927: 14 (n. syn.); locus typicus I"osslyn

(Virginia,

U.

S.

A.).

Chelogynus vivariensis Bradley: Muesebeck Anteon puncticeps Ashmead: Krombein 1979: Anteon (Chelogynus) virginiensis (Fenton): Anteon (Chelogynus) vivariensis (Bradley):

and Walkley

1951:

1041.

1241. Krombein

1979:

1241.

Krombein

1979:

1241.

FEMALe: fully winged; length 2,5-3,0 mm; colour variable; usually black; sometimes head reddish-testaceous, with a brown spot on ocellar triangle and behind ocelli; testaceous; black;

antennae fully

mandibles testaceous;

testaceous or brown,

mesopleura black or testaceous;

abdomen black or brown; brown;

pro;:lOdeum

metapleura black or reddish-dark;

legs testaceous or with stalk of hind femora

occasionally body fully

propodeum brown;

with segments 1-2

thorax black or reddish-dark;

antennal

testaceous,

segments

with dorsal

surface of

in following ,oroportions:

10:

5:

9:

7:

6: 6: 6: 6: 5: 8; head dull, weakly granulated, with a few irregular short keels mainly near orbits; frons with a median keel, without lateral keels, occipital carina complete; POL = 5; OL = 4; OOL = 7; OPL = 6; 7; pronotum dull, strongly transversely striate; posterior surface of pronotum shorter than scutum tegulae;

scutum,

(11:16);

posterior

scutellum and metanotum shiny,

TL

lobes of pronotum reaching smooth,

finely punctate,

part Fig.

303 -

Chela of

Anteon puncticeps Ashmead from Hc Lean Bogs Bioi. Res.

(New York)

476

Mem.

Amer.

En!.

Ins!.,

no.

37,1984

\

\\

Fig.

304 -

Chela of Anteon puncticeps Ashmead from Rosslyn variety with one row of lamellae.

(Virginia):

Anteoninae

without sculpture among punctures;

477

notaul ices

incomplete,

reaching nearly

0,50-0,66 length of scutum; propodeum reticulate rugose, transversal

keel

between

with 2 longitudinal

dorsal

keels;

and posterior surface;

with a strong

posterior surface

median area nearly as rugose as lateral

areas;

fore-wing hyal ine, without dark transversal bands; distal part of radial vein much shorter than proximal part (6:12); fore tarsal segments in

9: 3: 3: 9: 20; enlarged claw (Figs. 303, 304) with 1 bristle on a proximal prominence; segment 5 of front tarsus (Figs. 303, 304) with 1-2 rows of nearly 22 lamellae; apex with a group of 4-8 following proportions:

I ameli ae;

segment 5 of front tarsus with basa I part shorter than apica I

(8:12); segment 2 of front tarsus produced into a hook;

part

tibial

spurs

1,1,2. MALE:

fully

length 1,56-2,50 mm;

winged;

teeth brown;

antennae brown,

black;

mandibles yellow,

with

1 testaceous or with ventral

with segment

side yellow; sometimes segments 2-5 brown; tegulae testaceous; legs yellow, sometimes with basal part of hind coxae brown, always with hind clubs of femora and hind tibiae more or less brown; rugose and punctate;

frons with a

head dull,

longitudinal

granulated,

weakly

POL = 6;

median keel;

OL

= 3; OOL = 7; OPL = 4; TL = 5; occipital carina complete; scutum shiny, smooth,

finely punctate,

rugose only near anterior margin;

notaulices

incomplete, reaching mid-length of scutum; scutellum and metanotum shiny, smooth, without sculpture; propodeum reticulate rugose, with an irregular transversal keel between dorsal and posterior surface; posterior surface with 2 longitudinal straight keels; median area as rugose as lateral areas;

fore-wing hyaline,

without dark transversal

bands;

distal

part of

radial vein much shorter than proximal part (4:10); gonoforceps (Fig. C) with a wide dorsal proximal membranous process and with a distal inner pointed process; LOCUS TYPICUS:

TYPICAL MATERIAL:

spurs 1,1,2.

tibial

Arlington

(Virginia,

U.

S.

A.) of A.

and 1 para type M!

holotype M!

handwritten by Ashmead Anteon punctaticeps:

label

293

puncticeps (type

in the original

description Anteon puncticeps) in WA; holotype F! of Ch. vivariensis in IT; holotype M! of A. american us in IVA; holotype F! of Ch. virginiensis in WA. HOSTS:

unknown

DISTRIBUTION: Quadra

Is.

Nearctic region:

(B.C.),

OT!

Robson

(B.

OT!

Morley

(Alta.),

TW!

C.),

Stittsvi lie (Ont.),

OT! OT!

OT!

Funkstown

Ancaster

OT!

One Sided Lake

(Ont.),

OT!

Tillsonburg

Beach

(Madoc,

Knowlton

BM!

of King Mt.

(Old Chelsea,

(Que.),

Cap Chat

OT!

Lawrencetown

N.

C.),

S.),

Golden L.

Brighton

Braeside (Ont.), aT! OT!

Old Chelsea

(ant.),

U.

S.

A.:

OT!

OT!

aT!

P.

Mt.

aT!

(N.

Park

Crystal (Ont.),

aT!

Summit

Quyon

B.),

Ranier

OL!

Innisvi lie

Rondeau

OT!

(Que.),

(Que.),

C.),

(Alta.),

Chaffeys Locks Hull

Kouchibouguac N. OT!

WAr (B.

(ant.),

(Ont.),

Bothwell

(Que.),

C.),

Kananaskis

OL!

OT!

aT!

(B.

Cultus Lake

OT!

WAr

Dundas (Ont.),

Que.),

Is.

OT!

OT!

OT!

Cap Rouge

(Que.),

(Hal ifax Co.,

(B.

(Man.),

(Ont.),

OL!

C.),

(Ont.), OT!

(Ont.),

Ont.), OT! (Que.),

Vancouver

(B.

Mission City

(ant.),

TW!

CANADA:

Kowichan L.

OT!

(Wash.),

TW!

478

Mem. Amer. Ent.

Inst.,

no. 37,

1984

Fig. 305 - Chela of Anteon xanthothorax (Bradley)(type)

Ochoco Creek (Oreg.), TW! Corva II is (Oreg.), TW! Mc Bri de Spri ngs (Mt. Shasta, Siskiyou Co., Calif.), OT! Texas, PH! Salmon (Anderson Co., Tex.), TE! College Station (Brazos Co., Tex.), TE! Washtenaw Co. (Mich.),

Anteoninae

Ol!

lake Odessa

lake Co.,

(Mich.), Ohio),

Ridge,

(Mich.),

BM!

AM!

Tenn.),

Ol!

Columbia OT!

(Boone Co.,

Ol!

TW!

TW!

Greenville (5.

(N.

C.),

(Va.),

WA!

Island

(Md.),

TW!

Wattacoo

OT!

C.),

WAr

(D.

laurel

(Md.),

TW!

(V",.), C.),

(Md.),

TW!

WAr

WA! S.

C.),

TW! C.),

FAils Church

TW!

(Del.),

Gun

Morgan (East

Cleveland

Mc Clellanville TW!

OT!

Wayah

WA!

TW!

(Md.),

ZO!

OT!

(Va.),

(Pa.),

Cabin John

Wi Imington

BM!

(Ga.),

Ol! (N.

Spring Br.

OT!

Ol!

Chapin Sanctuary

Forsyth

Highlands

WA!

WA!

(Mich.),

NNW Mc Connelsvi lie,

Mo.),

(Pickens Co.,

Arlington

Washington

Takoma Pk.

(8 mi.

Jacksonville (Fla.),

(5. Bald

Ann Arbor

Muskingum River

(5. C.), C.),

BM!

479

Rosslyn

Plummers

IT!

Bowie (Md.),

Moorestown

(N.

J.),

TW! Westerly (R. I.), TW! Mc lean Bogs Biological Reservation (N. Y.), IT! lake Placid (N. Y.), OT! Keene Valley (Essex Co., N. Y.), OT! Poughkeepsie TW!

Shokan

(N. (N.

Y.),

TW!

Y.),

TW!

South Haven

(long

(Huntington,

N.Y.), AM!

Y.),

AM!

Island,

lewisboro

Washington

(N.

Farmingdale (N. Syracuse N.

4 mi.

Y.),

OT!

TW!

TW!

Hancock

AM! N.

Bigelow lawn

Bemus PI. (N.

(N.

Y.),

Y.),

TW!

Kalbfleisch Field Station

NNW Kerhonkson

(Westchester Co.,

H.),

Y.),

(N. Y.),

Y.),

(Mt.

(Cherry town,

AM!

Washington,

11. Anteon xanthothorax

Ulster Co.,

N.

Alpine Garden of Mt. N.

H.),

OT!

(Brad ley)

Chelogynus xanthothorax Brad ley 1926: 8. Chelogynus xanthothorax Brad ley: Muesebeck and Wa I k I ey 1951: 1041.

Anteon (Chelogynus) xanthothorax (Bradley): Krombein 1979: 1241.

FEMALE:

fully

winged;

antennae brown,

length 2,50-3,37 mm;

black;

with segments 1-3 testaceous;

mandibles testaceous;

prothorax yellow;

abdomen

sometimes brown-testaceous; legs testaceous; antennae distally thickened; antennal segments in following proportions: 11: 5: 8: 7: 7: 6: 7: 7: 7: 9; head shiny,

smooth,

puncta te on frons, median and = 7;

lateral

OPl = 6;

punctate,

weakly punctate on vertex,

wi thout scu I pture among punctures; keels;

Tl = 6,5;

occipital carina complete; pronotum shiny,

wi thout scu I pture among punctures; than scutum smooth,

(12:13);

pronotal

weakly punctate,

incomplete and weakly

smooth,

more strongly

frons

very weakly

Ol = 4;

OOl

punctate,

posterior surface of pronotum shorter

tubercles reaching tegulae;

scutum shiny,

without sculpture among punctures;

impressed,

wi thout

POL = 5;

but only on

notaulices

the anterior third;

scutellum

and metanotum shiny, smooth, without sculpture; propodeum reticulate rugose, with a strong transversal keel between dorsal and posterior surface;

posterior surface with 2 longitudinal

rugose as

lateral

the pterostigma; part

(5:10);

areas; distal

fore tarsal

keels;

fore-wing almost hyal ine,

part of radial

median area as

with a pale band near

vein much shorter than proximal

segments in following proportions:

8:

4:

4:

7:

19;

480

Mem.

Fig.

306 -

Odessa

Amer.

En!.

Inst.,

no.

37,

1984

Male genitdlia of Anteon xanthothorax

(Michigan):

the

left half is ventrally

(Bradley)

viewed;

from Lake

the right half

is

dorsally viewed.

segment 5 of front (9:

10);

tarsus

enlarged claw

prominence; lamellae;

with basal

(Fig.

305)

segment 5 of front

part sl ightly

tarsus

(Fig.

apex with a group of nearly 7

tarsus produced

into a hook;

tibial

305)

and tibiae partly

antennal

segments

head shiny, without

testaceous;

longitudinal

punctate,

segment 3 of front mandibles testaceous; light; sometimes also

antennae not distally 8:

5:

8:

7:

thickened;

8:

7:

7:

7:

without sculpture among punctures;

median keel;

part

spurs 1,1,2.

in following proportions:

smooth,

apical

proximal

with 2 rows of nearly 33

lamellae;

MALE: fully winged; length 1,56-2,75 mm; black; antennae black or brown; IpSlS brown, with tarsi femora

shorter than

with one bristle on a

occipital carina complete;

POL

6:

8;

frons

=

6;

Ol

4; OOl = 6; OPl = 4; Tl = 5; scutum, scutellum and metanotum punctate, wi thout scu I pture among punctures; notau I ices i ncomp lete, reach i ng approximately 0,5 strong transversal surface with hyal ine,

two

length of scutum; keel

longitudinal

keels;

without dark transversal

shorter than proximal

propodeum reticulate rugose,

between dorsal

part

(4:8);

and posterior surface;

median area rugose;

bands;

distal

gonoforceps

fore-wing

part of radial

(Fig.

306)

with a

posterior

with a

vein much large

Anteoninae

dorsal

proximal

process;

membranous process and with a distal

tibial

Taughannock Falls (New York,

TYPICAL MATERIAL:

holotype F!

U.S.A.).

and 1 paratype F!

in

IT.

unknown

DISTRIBUTION: C.),

OL!

Ottawa (Ont.),

Brighton (Ont.),

OT!

Nearctic region:

(B.

OT!

inner pointed

spurs 1,1,2.

LOCUS TYPICUS: HOSTS:

481

Kleanza Creek

(On!.), OT!

Chelsea, (Que.),

OT!

Que.), OT!

(Ont.), OT!

OT!

OT!

(Que.),

Kouchibouguac N. St.

P.

John's

(Que.), OT!

(N.

OT!

OT!

OL!

Stittsville (Ont.),

Innisville

B.), OT!

(On!.), OT!

St.

(Ont.), OT!

Bonaventure OT!

Is.

(Kam.

Aylmer

Andrews

OT!

U. S.

(Alta.), OT!

OT! OT!

Wakefield

Effingham

(Ont.),

Summit of King Mt.

Parke Reserve

(Newfoundland),

Robson

Waterton

Hami Iton

Fabre (Que.),

OT!

OT!

OT!

OT!

(Yukon),

C.),

Belleville (Ont.),

Chimo (Que.),

Cap Rouge

Sheldon L.

B.

(Ont.),

Marmora (Ont.),

(Ont.),

New Richmond S.), OT!

OT! Golden L.

Frankford

Chatterton

CANADA:

(Terrace,

(Que.), Co.,

(Que.),

(N.

B.),

A.:

Lowman

OT!

Que.), OT!

OT!

(Old Hull OT!

OLl

Lockeport (Idaho),

(N.

TW!

Benzie Co. (Mich.), OL! Washtenaw Co. (Mich.), OL! Iron Co. (Mich.), Ann Arbor (Mich.), OL! BM! Lake Odessa (Mich.), OL! BM! Hartford (Conn.),

OT!

Keuka Lake

Falls (Ithaca, Y.),

N.

Y.),

(Penn Yann,

IT!

Y.)(Bradley

Oneonta

(N.

Canajoharie (N. Y.), TW! Rome (N.Y.), TW! Milford Center South Haven (Long Island, N. Y.), AM! 4 mi. NNW Dryden

Ulster Co.,

(Maine),

OL!

N.

Y.),

AM!

Alpine Garden

TW!

Taughannock

TIV! TW!

(Cherry town,

(N.Y.),

OT!

(N. Y.), (N. Y.),

AM!

Millwood

1926);

lO!

Shokan

Y.),

TW!

Y.),

TW!

Kerhonkson

Y.),

N.

OLl

N.

(N.

N.

Six-Mile (Ithaca,

OL!

Westchester Co.

Brainard (Mt.

(Rensselaer Co.,

Washington,

N.

H.),

OT!

12. Anteon osborni (Fenton)

Chelogynus chlorotettixi Fenton 1918a: 202 (nomen nudum). Chelogynus osborni Fenton 1918a: 272. Chelogynus rugulosus Fenton 1927: 14 (n. syn.); locus typicus Saint John

(New Brunswick,

Canada).

Anteon flaviscapus Whittaker 1930: 67 (n. syn.); locus typicus Hollyburn

(British Columbia,

Canada).

Anteon whittakeri Muesebeck and Walkley 1951: 1041; new name for A. flaviscapus Whittaker 1930 preoccupied by A. flaviscapus Kieffer 1905.

Anteon whittakeri Muesebeck and Walkley: Krombein 1979: 1241. Anteon (Chelogynus) osborni (Fenton): Krombein 1979: 1241. Anteon (Chelogynus) rugulosus (Fenton): Krombein 1979: 1241.

FEMALE:

fully

winged;

length 2,25-3,18 mm;

occasionally reddish-light or reddish-brown;

head black or brown; sometimes with anterior half

482

Mem. Amer.

Ent.

Inst.,

no.

37,

1984

\

I

/

Fig.

307 -

Chela of Anteon osborni

I

/

/

/

/

(Fenton) (type)

Anteoninae

of frons

testaceous and with posterior half black;

testaceous;

mandibles testaceous;

testaceous,

thorax black;

with anterior half of pronotum brown;

legs testaceous,

clypeus sometimes

antennae brown or

segments 2-10 or 3-10 or 4-10 darkened; yellow,

483

with

sometimes prothorax

abdomen black or brown;

someti mes with hind coxae basa II y

black or with cl ubs of

mid femora, hind coxae, clubs of hind femora, hind tibiae brown; antennal segments in following proportions: 11: 5: 7: 7: 6: 6: 5: 5: 5: 8; head shiny,

smooth,

frons;

no sculpture among punctures;

OL

=

4;

=

OOL

striate,

finely punctate, 5;

=

OPL

5;

=

TL

6;

occipital carina complete;

pronotum shiny,

smooth only near posterior margin;

tegulae;

scutum shiny,

punctures; scutum;

smooth,

notaulices incomplete,

reaching

with an

and posterior surface;

rugose,

pronotal

weakly punctate,

scute II Um and metanotum shi ny,

propodeum reticulate rugose, dorsal

strongly punctate only on anterior half of POL = 6;

transversely

tubercles reaching

without sculpture among

approximately 0,65

smooth,

length of

without scu I pture;

irregular transversal

keel

between

posterior surface with 2 longitudinal

keels;

median area partly rugose, with a shiny and smooth central area; forewing hyal ine, without dark transversal bands; distal part of radial vein much shorter than proximal proportions: a proximal nearly 29

7:

2:

3:

8:

prominence; lamellae;

18;

part

tibial

MALE:

10);

(Fig. tarsus

segments

307)

in following

with one bristle on

(Fig.

group of 4 lamellae;

307)

with 2 rows of

segment 5 with basal

segment 2 of front tarsus produced

into a

spurs 1,1,2.

fully winged;

length 1,50-2,62 mm;

antennae testaceous or testaceous-brown; fully

fore tarsal

segment 5 of front

apex with a

part shorter than d i sta I part; hook;

(3:

enlarged claw

black;

mandibles testaceous;

sometimes segment

1 I ight;

legs

testaceous or with hind COXae partly black and femora partly brown;

antennae not distally thickened; proportions: punctate,

8:

5:

7:

7:

7:

7:

antenna I segments in following

6,5:

6,5:

7:

10;

without sculpture amongpunctures;

median keel

head shiny,

smooth,

frons with a short

in front of anterior ocellus and with a

weakly

incomplete

weak median furrow,

without lateral keels; occipital carina complete; POL = 5,5; OL = 3; OOL 7; OPL = 4; TL = 4; scutum shiny, smooth, weakly punctate, without sculpture among punctures,

weakly rugose only near anterior margin;

notaulices incomplete, reaching approximately 0,5 length of scutum; scutellum and metanotum shiny, smooth, without sculpture; propodeum reticulate rugose,

with a

posterior surface;

posterior surface with 2

strong

as rugose as

lateral

bands;

part of radial

distal

gonoforceps

(F i g.

areas;

288 C)

transversal

keel

between dorsal

longitUdinal

fore-wing hyal ine,

keels;

without dark

vein much shorter than proximal

with a

and

median

area

transversal part

(4: 10);

I arge dorsa I prox i ma I membranous process

and with a distal inner pointed process; tibial LOCUS TYPICUS: Sandusky (Ohio, U. S. A.).

spurs 1,1,2.

TYPICAL MATERIAL: holoytpe F! of Ch. osborni in WA; holotype F! of Ch. rugulosus in WA; holotype M! and 3 paratypes MM! of A. flaviscapus Whittaker in BM; 1 paratype M! of A. flaviscapus Whittaker in WA; 1 paratype M! of A. flaviscapus in LA.

484

Mem.

HOSTS:

in U.

S.

DISTRIBUTION: CANA:JA: (B.

B.

C.),

(Yukon),

Hixon

OT!

(B.

B.

C.),

OT!

Atlin

(Osoyoos,

B.

OT!

aT!

Golden L.

(Ont.),

aT!

Maynooth

Stittsville

aT!

Hollyburn

(Ont.), OT! (Ont.), aT!

aT!

OT!

P.

(Oreg.),

TIV!

Stanley

Elk Creek (Cal if.),

(3 mi. TW!

(N.

Brandon

Is.,

OT!

(Man.),

(Ont.),

aT!

aT!

(Que.),

OT!

OL!

Hull

B.),

WA!

Saint John

TW!

Corvall is

(Idaho), (Ariz.),

TW!

Somerset

Siskiyou Co., TW!

Estes Park

Ottawa

Dresden (Ont.),

(Ont.), aT!

One

(Ont.),

aT!

Fort Chimo

OT!

Parke Reserve (Kam. OT!

Chimo

Old Chelsea

(Wash.),

aT!

TIV!

(Que.),

Que.), OT!

C.),

Marmora

Niagara Glenn

(Ont.),

B.

Richter Pass

OT!

E Mc Cloud,

Portal

C.),

Frances (Ont.),

(Old Chelsea,

Ranier

LA!

Bancroft

Lac Brule (Que.),

Mt.

Chilliwack

IVA!

aT!

aT!

aT!

OT!

BM!

Innisville (Ont.), OT!

Meach Lake (Que.),

Kouchibouguac N.

C.),

Ancaster (ant.),

Chaffeys Locks

Mistassini

B.

Brighton

TW!

B.),

Ft.

OLl

(Ont.), OT!

OT!

Spring Creek

(Vancouver

(Terrace,

Cumberland Que.),

B.

c.),

(B.

Cowichan Bay

Lakelse L.

aT!

Summit of King Mt.

OT!

Galiano (B. C.), BM! Robson (B. C.), OT! C.), aT! Burnaby (B. C.), OT! Shames (Terrace,

Ninette (Man.),

(Ont.),

(Fairbanks),

(Yukon),

Sided Lake (Ont.),

Co.,

37,1984

Birch L.

Sheldon L.

(B. C.), OT!

C.),

no.

Madden Lake (Oliver,

C.),

C.), OT!

(Ont.), OT!

ALASKA:

aT!

Cowichan L. (B. c.), OT! Gagnon Road (Terrace, B. Cultus Lake (B.

Inst.,

Chlorotettix unicolor (Fitch) (Fenton 1918b).

A.

C.), OL!

BM!

Ent.

Nearctic region:

Dawson

(Terrace,

Amer.

(N.

B.),

(Oreg.), (EI

AM!

(Colo.),

(Que.),

Fredericton

(~J.

Rhododendron

Dorado Co.,

Calif.),

aT!

(Que.), OT!

WA!

TW!

(Que.),

Calif.),

TE!

Crescent City

TW!

aT!

Lawrence

(Kans.), TW! (Mich.), BM!

Benzie Co. (Mich.), OL! Iron Co. (Mich.), OL! Ann Arbor TW! Washtenaw Co. (Mich.), Sandusky (Ohio), WA! Cleveland

(S. C.), OL!

TW!

C.), OT!

(State Forest, (Md.),

lVattacoo (Pickens Co.,

Indian Gap

OT!

IV.

(Gr.

Va.),

Smoky Mt.

aT!

High Point St.

Hawksbi II

Pk.

(N.

S. C.),

Nat.

J.),

Pk.,

aLl

N.

(Shenandoah, TIV!

TW!

C.),

Va.),

Lakehurst

Highlands (N.

aT!

Cooper's Rock

aT!

(N. J.),

Laurel TW!

(R. I.), TW! Syracuse (N. Y.), TIV! South Haven (Long Island, N. Y.), AM! Ithaca (N. Y.), ZO! Sloansville (N. Y.), OT! Farmigdale (N. Y.), TIV! Bemus PI. (N. Y.), TIV! Shokan (N. Y.), TIV! Millwood (N. Y.), TIV! Canajoharie (N. Y.), TW! So. Hadley (Mass.), TIV! Alpine Garden (Mt. Washington, N. H.), OT! MEXICO: EI Saito (Durango), OT! La Ciudad (Durango), aT!

lVesterly

13. Anteon durangoi n. sp.

FEMALE:

unknown

MALE: fully winged; length 2 mm; black; mandibles testaceous; antennae and legs brown; antennae not distally thickened; antennal segments in following proportions: 7: S: 6: 6: 7: 5: 6: 6: 6: 9; head dull, granulated, with track of areolae on vertex and with numerous longitudinal striae on frons; frons without median and lateral keels; occipital

carina complete;

POL

=

5;

OL

=

3;

OOL

=

4;

OPL

=

3;

TL

=

5;

Anteoninae

Fig.

308 -

Male genitalia of n.

scutum shiny, notaul ices

sp.

Anteon ciudadi n. sp.

(type)(B):

weakly punctate,

incomplete,

faintly

without

(type)(A)

and

durangoi

right half removed.

without sculpture among punctures;

visible near anterior margin of scutum;

scutellum and metanotum shiny, reticulate rugose,

485

smooth,

without sculpture;

longitudinal keels on

propodeum

posterior surface,

with a

transversal keel between dorsal and posterior surface; fore-wing hyal ine, without dark transversal bands; distal part of radial vein shorter than proximal

part

(2,5:9);

membranous process,

gonoforceps

with a distal

(Fig.

308 B)

without dorsal

iner pointed process;

1,1,2. LOCUS TYPICUS: La Ciudad (Mexico) TYPICAL MATERIAL: holotype M! in OT.

tibial

proximal spurs

486

Mem.

HOSTS:

Amer.

En!.

Ins!.,

no.

37,

1984

unknown

DISTRIBUTION:

Nearctic region:

MEXICO:

La Ciudad

(Durango),

OT!

14. Anteon ciudadi n. sp.

FEMALE: MALE:

unknown

fully

winged;

length 2,50-2,87 mm;

antennae testaceous-brown; brown;

antennae not distally

puncta te, lateral

black;

mandibles testaceous;

testaceous or wi th coxae and femora

thickened;

antenna I segments

10: 7: 8: 9: 9: 8: 8: 8: 8:

proportions:

=

legs fu II y

10;

head shiny,

without scu I pture among punctures;

=

occipital carina complete; POL = 6; OL = 3,5; OOL = 6; OPL 5; scutum, scutellum and metanotum weakly punctate, without

sculpture among punctures;

notaul ices

incomplete,

reaching

0,3 length of scutum; propodeum reticulate rugose, keels on posterior surface; fore-wing hyal ine,

median area dull,

without dark transversal

vein shorter than proximal tibial

weakly

frons wi thout med i an and

keels;

3,5; TL

dorsal

in following

smooth,

proximal

with 2

approximately longitudinal

as rugose as

bands;

distal

lateral

(5:9); gonoforceps (Fig. 308 A)

part

membranous process,

without distal

areas;

part of radial without

inner pointed process;

spurs 1,1,2.

LOCUS TYPICUS: La Ciudad (Mexico) TYPICAL MATERIAL: holotype M! in OT; HOSTS: unknown

paratype M!

DISTRIBUTION:

Pinehurst

La Ciudad

Nearctic region:

(Durango),

U.

S.

A.:

in TW.

(Oreg.),

TW!

MEXICO:

OT!

15. Anteon menkei n. sp.

FEMALE:

unknown

MALE: fully winged; length I,BI-2,12 mm; head black, with anterior half of frons, clypeus and mandibles testaceous; antennae and legs fully testaceous;

somet i mes antennae darkened;

abdomen black;

antennae not distally

following proportions: punctate, lateral

B:

6:

6,5:

7:

thorax and propodeum black;

thickened;

7:

7:

7:

7:

without scu I pture among punctures;

keels;

occipital

carina complete;

POL

antenna I segments 7:

9;

head shiny,

frons without med i an

=

5;

OL

=

3;

OOL

=

in weakly and

6;

OPL

=

3; TL = 4; scutum shiny, smooth, weakly punctate, without sculpture among punctures; notaul ices incomplete, reaching approximately 0,5 length of scutum;

scutellum

and

metanotum shiny,

smooth,

weakly

punctate,

without sculpture among punctures; propodeum reticulate rugose, with 2 longitudinal keels on posterior surface; median area as rugose as lateral areas;

fore-wing hyal ine,

radial

vein shorter than proximal

without dark part

transversal (4:10);

bands;

distal

gonoforceps (Fig.

part of 309)

Anteoninae

Fig.

without dorsal

309 -

Male genitalia of Anteon menkei n.

proximal

membranous process,

process; tibial spurs 1,1,2. LOCUS TYPICUS: Gatlinburg (Tennessee, TYPICAL MATERIAL: holotype M! paratypes MM! in OL. HOSTS: unknown DISTRIBUTION: Nearctic region: Cleveland NOTES:

(S.

C.),

487

TW!

Wattacoo

in WA;

sp.

with a distal

(type)

inner pointed

U.S.A.). 2 paratypes MM!

U. S. A.:

Gatlinburg

(Pickens Co.,

probably this species is a male variety

S.

in TW;

(Tenn.),

C.),

TW!

2

WA!

OLl

(with frons partly

testaceous) of A. osborni. The species is named in honor of Mr. Arnold S.

Menke.

16. Anteon hirtifrons Whittaker

Anteon hirtifrons Wh i ttaker 1930: 68. Anteon hirtifrons Whittaker: Muesebeck and Walkley 1951: 1040.

488

Mem. Amer.

Anteon hirtifrons

FEMALE: MALE:

Ent.

Ins!.,

no.

37,

Krombein 1979:

Whittaker:

1984

1241.

unknown

fully

winged;

antennae testaceous, with hind coxae,

length 2,52 mm;

black;

mandibles testaceous;

wi th segments 1-3 dorsa II y

darkened;

femora and tibiae partly darkened;

legs testaceous,

antennae not distally

thickened; antenna I segments in following proportions: 13: 7: 9: 9: 9: 8: 8: 8: 8: 12; head strongly punctate and reticulate rugose, with irregular keels;

frontal

line complete;

occipital carina complete;

POL; 8;

OL ;

4;

6; TL ;

3,5; scutum, scutellum and metanotum shiny, smooth, without sculpture; notaulices incomplete, reaching approximately 0,5 length

OOL ;

of scutum;

propodeum reticulate rugose;

longitudinal

keels;

posterior surface with

median area as rugose as lateral

areas;

two

fore-wing

hyaline, without dark transversal bands; distal part of radial vein much shorter than prox i ma I part ( 5: 10); gonoforceps (F i g. 293 B) with a very large distal

inner pointed process;

LOCUS TYPICUS:

Hollyburn

TYPICAL MATERIAL: very damaged, HOSTS:

(B.

C.,

holotype M!

without body,

spurs 1,1,2.

tibial

Canada)

in BM;

1 paratype M! in LA (specimen legs and wings).

only with antennae,

unknown

DISTRIBUTION: Nearctic region: Galiano (B. C.), BM!

CANADA:

Hollyburn

NEOTROPIC SPECIES OF

(B.

C.),

LA!

BM!

ANTEON

Twenty one species are known.

KEY TO THE SPECIES

FEMALES

4 of front tarsus at most 0,5 as long as segment 310, 317) ••••••••.•••••••••••••.•••••.•.••.•••••••..... 2 Segment 4 of front tarsus slightly shorter, as long as, or longer than segment 1 (Figs. 311, 313) . . . . . . . . . . . . . . . . . . . . . . . . 4 Segment (Figs.

2

3

Posterior surface of

propodeum wi th

two longi tudinal keels ••••• 1. c1avatum Olmi and Currado Posterior surface of propodeum without longitudinal keels . . . . . 3 Scutum fully granulated; frons reticulate rugose and granulated . . • . . . • • . • • • • . • . • . . . • • • .. •• 6. mexicanum n. sp • Scutum weakly punctate, wi thout sculpture among punctures;

Anteoninae

frons

rugose,

without

sculpture among keels,

5

Posterior

surface of

propodeum without

Posterior

surface of

propodeum with

Fore-wing hyaline, Fore-wi ng wi th

6

without

1-2 dark

Pronotum and head

dark

keels . . . . 5

longitudinal

keels •• 13

bands.....

..6

bands.............

.10

transversal at

granulated •.•

sp.

longitudinal

two

transversa I

fully black,

not

oliveirai n.

• •.•••.•••••••••••••.••.•••.•• 7.

4

489

most

with mandibles . •• 7

testaceous............................................. Pronotum and head at

7

Head by

granulated,

numerous

Head

8

fully

not

part I y

red

reticulate

t estaceous •••••••••• 8

or

rugose,

sculptured by numerous areolae and

molle n.

Head

red . . . . .

pronotum fully

sp.

· . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3.

nigrorubrum n.

Head with anterior half

testaceous-whitish;

pronotum fully Scutum fully

of

of

frons

frons black

punctate and wi thout

Scutum with anterior rugose,

or partly

sp.

punctate,

region

wi th

SCUlpture among

panamense n. sp.

smooth;

irregular

posterior half

lobes;

surface of

posterior without

Posterior

surface of

surface

of

pronotum wi th

surface of

punctate,

dull,

striae •••••••••••

· . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5. jamaicanum n. 10 Posterior

vertex

reddish-dark;

or most Iy black . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9

smooth,

punctures • •••.•••••••.•••••••• 4.

11

sculptured

irregular keels •••• •••

· . . . • . . • . • . • . • . . . . • . . . . . . . . . . . 2. fully black;

not

micros n. sp.

irregular keels .. 20.

and posterior half 9

I east

two dorsal

propodeum smooth,

sp. transversal shiny,

weakly

sculpture among punctures . . . . . . . . . . . . . . . . . 11 pronotum wi thout

propodeum dull,

fully

dorsal

reticulate

lobes; rugose

posterior or with

transversal

keels . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 12

Head

without

shiny,

sculpture,

or punctate and without

noyesi n. sp. surinamense n. sp.

sculpture among punctures •.•.. 8. Head du I I, 12 Posterior

g ranu I at ed •••••••• 21. surface of

pronotum shiny,

without

sculpture,

smooth; posterior surface of propodeum wi th strong keels, not reticulate rugose.13. annulicorne Brues Posterior

surface of

surface of

pronotum dull,

propodeum reticulate

Pronotum not 14 Head

fully

two dorsal

forming

albitarse (Came ron)

lobes near posterior margin . . . . . 14

two dorsal

lobes • •••••••••••••••••••

granulated . . . . . . . . 19.

Head punctate,

without

posterior

rugose ••• ••••••••.••••••••

• •••••••••.•••••••••••••••••• 14. 13 Pronotum forming

granulated;

transversal

•• 15

bolivianum n. sp.

sculpture among punctures,

not

granulated . . . . . . . . . . . . . . . . . . . . 9. lobatum n. sp. 15 Anterior surface of pronotum nearly as long as posterior surface (Fig. 322 B); head fully except for the frontal I ine .. 10. Anterior

surface of

pronotum much

smooth,

wi thout

sculpture,

propodeale (Fenton) shorter

than posterior

490

Mem.

surface 16 Head as,

(Fig.

fully or

Amer.

En!.

Ins!.,

rugose,

longer

frons

only

with OPL

longitudinal

median

vertex

striae

occipital

rugose,

smooth,

nearly

as

long

in

the

with OPL nearly

three

wi th numerous

region

between

anterior

carina . . . . . . . . . . . . . . . . . . . . . . ..

nycteum n.

• •••••••••••••••..•••..•.•••• 1 2.

The females of

shorter,

chiriquense (Cameron)

reticulate

long as POL;

ocellus and

1984

than POL . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

•• • . •• ••••••••• •• • •• ••• • ••••• 11. times as

37,

322 A) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 16

reticulate

sl ightly

Head with

no.

sp.

A. pilicorne, plaumanni, conterminum and vivax are

unknown.

MALES

2

Posterior

surface of

propodeum with

Posterior

surface of

propodeum without

Head

fully

reticulate

rugose.18.

Head punctate and wi thout reticulate

two

longitudinal longitudinal

plaumanni n. sp.

sculpture among punctures,

rugose . . . . . . . . . . . . 15.

keels ••• 2 keels . . . . 3 not

vivax n. sp.

3

Scutum granulated •••••••••••••••••••••••••••••••••••••••••••• 4 Scutum weakly punctate, without sculpture among punctures . . . . 5

4

Gonoforceps wi thout

di stal

inner rounded pro<:css

(Fig.

329)

• •••••••••••••••••••••••••••• 1 6. pilicorne (Og I ob lin) Gonoforceps with distal inner rounded process (Fig. 318) . . .

· . . . . . . . . . . . . • • . . • • . . . . . . . . . . . 6.

5

Head with numerous

longitudinal

inner pointed process

(Fig.

6

reticulate

rugose,

sp.

gonoforceps with distal

312) . . . . . . . . . . . . . . . . . . . . . . . . . . . . ..

· • . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2. Head

mexicanum n. keels;

without

molle n. sp. longitudinal keels;

gonoforceps wi th or wi thout di stal inner pointed process (Figs 316, 330) ••••••••••••••••••••••••••••••••••••••••••••••••••.• 6 Gonoforceps wi th di stal inner pointed process (Fig. 330) . . . . . . • •••••••••••••••••••••••••••• 17. conterminum n. sp. Gonoforceps without distal inner pointed process (Fig.

. . . . . . . . . . . 5. jamaicanum n.

316) . . .

sp.

The males of A. clavatum, albitarse, lobatum, propodeale, annulicorne, chiriquense, oliveirai, micros, nigrorubrum, panamense, noyesi, surinamense, bolivianum and nycteum are unknown.

Anteoninae

491

1. Anteon clavatum Dlmi and Currado

Anteon c1avatum Dlmi and Currado 1979a: 343.

Fig.

310 -

Chela of Anteon c1avatum Dlmi

and Currado

(type)

492

Mem.

FEMALE:

fully

Amer.

winged;

Ent.

length 4,37 mm;

antenna I segments 1-2 testaceous; redd i sh;

legs testaceous,

antennae distally 17:

10:

9:

frontal

OOl = 6; fully

7:

7:

7:

8:

OPl = 6;

scutellum shiny, margin;

7:

7:

9;

segments

head fully

with

teeth

pronotal

smooth,

reticulate rugose,

tubercles reaching

weakly

punctate,

smooth,

with a

shiny;

POL = 8;

Ol = 4;

pronotum shiny,

tegulae;

scutum dull,

notaulices invisible;

reticulate rugose near posterior

without sculpture;

transversal

brown;

in following proportions:

occipital carina complete;

posterior surface with 2

surface;

with antennae brown;

reticulate rugose only on sides;

metanotum shiny,

reticulate rugose,

black,

1984

mandibles testaceous,

antennal

Tl = 3;

granulated,

37,

frons with 2 keels around orbits;

reticulate rugose;

smooth,

no.

with fore coxae and fore femora part I y

thickened;

line complete;

Inst.,

keel

propodeum shiny,

between dorsal

longitudinal

keels;

and posterior

median area as

reticulate rugose as lateral areas; fore-wing hyal ine, without dark transversal bands; distal part of radial vein much shorter than proximal part

(4:13);

pterostigma

following proportions:

one bristle located on a (Fig.

310)

3:

2,5:

4:

proximal

with some bristles;

part of segment 5 much MALE:

long and narrow;

15:

12;

fore tarsal

segments

enlarged claw

(Fig.

segment 5 of front

prominence;

apex with a

longer than apical

in

310)

group of 13 lamellae; part;

tibial

with tarsus basal

spurs 1,1,2.

unknown

lOCUS TYPICUS:

Liangtelau

TYPICAL MATERIAL:

(Brazil)

holotype F!

HOSTS: unknown DISTRIBUTION: Neotropic region:

in GE. BRAZil:

liangtelau,

GE!

2. Anteon molle n. sp.

FEMALE:

fully

winged;

length 2,12-2,62 mm;

black;

mandibles testaceous;

antennae brown; legs testaceous, with hind coxae basally brown; antennae distally thickened; antennal segments in following proportions: 10: 6: 9: 9: 9: 8: 8: 8: areolae and longitudinal

8: 10; head dull, granulated, with numerous keels; frontal I ine complete; occipital carina

complete;

=

POL = 4,5;

Ol

3;

OOl

=

7;

OPl

=

4;

Tl

=

5,5;

pronotum dull,

granulated, rugose, with transversal keels, smooth only near posterior margin; posterior surface of pronotum shorter than scutum (5:14); pronotal

tubercles reaching

smooth,

shiny,

notaul ices

tegulae;

weakly punctate,

incomplete,

scutum,

scutellum and metanotum

without sculpture among punctures;

reaching approximately 0,3

length of scutum;

propodeum reticulate rugose, with a transversal keel between dorsal and posterior surface; posterior surface without longitudinal keels; fore-wing hyal ine,

without dark transversal

bands;

distal

part of radial

vein

shorter than proximal part (4:9); fore tarsal segments in following proportions: 9: 2,5: 3,5: 5,5: 15; enlarged claw (Fig. 311) with one bristle

located on a

proximal

prominence;

segment 5 of front

tarsus

(Fig.

Anteoninae

Fig.

311)

with

311

-

1 row of 37

493

Chela of Anteon molle n.

lamellae;

apex with a

sp.

(type)

group of 5

lamellae;

tibial

spurs 1,1,2. MALE:

fully

winged;

length 1,87-2,00 mm;

black;

mandibles testaceous;

antennae brown, with segments 1-2 testaceous; legs testaceous, with coxae and clubs of femora partly brown; antennae not distally thickened; antenna I segments in following proportions: 8: 4: 5: 6: 6: 6: 5: 5: 6: 8; head shiny,

with frons sculptured by numerous

behind posterior ocelli 5,5;

OL = 2,5;

OOL = 6;

metanotum shiny, punctures; scutum; dorsal

reticulate rugose; OPL = 3;

smooth,

reaching

propodeum reticulate rugose, and posterior surface;

fore-wing hyal ine, proximal

part

scutum,

de

transversal bands;

gonoforceps

with distal

las Casas

POL =

without sculpture among

with a

(3:7);

region

scutellum and

approximately 0,25

transversal

membranous process,

tibial spurs 1,1,2. LOCUS TYPICUS: San Cristobal

carina complete;

posterior surface wi thout

without dark

vein shorter than proximal dorsal

TL = 4;

weakly punctate,

notaulices incomplete,

longitudinal keels;

occipital

keel

length of between

long i tud i na I keel s;

distal (Fig.

part of radial 312)

without

inner pointed process;

(Mexico)

494

Mem.

Fig.

Amer.

Ent.

Inst.,

no.

37,

1984

312 - Male genitalia of Anteon molle from S. Cristobal de las Casas (Mexico):

TYPICAL MATERIAL: holotype F! (4 MM, 1 F)! in OL. HOSTS: unknown

left half removed

and 9 paratypes MM!

DISTRIBUTION:

Neotropic region:

MEXICO: San Cristobal

(Chiapas),

! OT!

(Sinaloa),

OT!

OL!

Ol

COLOMBIA:

EI Cali

Palmito

(Valle del

Cauca),

Ol!

OT!

5 paratypes

in OT;

de las Casas

Concordia

(Sinaloa),

Ol!

3. Anteon nigrorubrum n. sp.

FEMALE:

fully

testaceous;

winged;

length 1,25 mm;

antennae testaceous;

and propodeum black;

head black,

prothorax red;

abdomen and

with mandibles

mesothorax,

legs testaceous;

meta thorax

antennae distally

Anteoninae

Fig.

thickened;

313 -

Anteon nigrorubrum

Chela of

antenna I segments

495

n.

sp.

in following proportions:

7:

(type)

3,5:

4:

4:

3,5:

3,5: 3,5: 4: 4: 6; head dull, fully granulated and with numerous longitudinal weak striae; frons without median and lateral keels; occipital 4;

carina complete;

pronotum dull,

hairy,

POL = 5;

OL = 2,5;

granulated and

OOL = 4,5;

posterior surface of pronotum shorter than scutum tubercles reaching smooth,

very

tegulae;

scutum,

weakly punctate,

OPL = 2,5;

with some transversal (7:10);

TL

keels;

pronotal

scutellum and metanotum shiny,

without sculpture among punctures;

notaul ices incomplete, reaching approximately 0,65 length of scutum; propodeum reticulate rugose, with a strong transversal keel between dorsal and posterior surface; posterior surface without longitudinal keels, fully

reticulate rugose,

bands;

distal

fore tarsal

segments

enlarged claw prominence; lamellae, MALE:

(Fig.

fore-wing hyaline,

in following proportions: 313)

lamellae;

with one bristle tarsus

5:

1:

1,5: 3,5:

located on a

(Fig.

313)

part shorter than apical tibial

without dark transversal

vein much shorter than proximal

segment 5 of front

with basal

group of 4

dull;

part of radial

with

part

part

(1:6);

10;

proximal 1 row of 23

(4:6);

apex with a

spurs 1,1,2.

unknown

LOCUS TYPICUS:

St.

TYPICAL MATERIAL:

Augustine

(Trinidad)

holotype F!

HOSTS: unknown DISTRIBUTION: Neotropic region:

in BM. TRINIDAD:

St.

Augustine

(St.

George),

496

Mem. Amer. En!.

Ins!.,

no. 37,

1984

BM!

Fig. 314 - Chela of Anteon panamense n. sp.

(type)

Anteoninae

497

4. Anteon panamense n. sp.

FEMALE:

fully

half of frons testaceous;

winged;

clypeus

antennae brown, reddish-dark;

testaceous;

abdomen and

8:

4:

with numerous complete; rugose,

with anterior

ventral

mandibles testaceous,

face of head

with

teeth reddish;

6 testaceous;

thorax and

except for anterior margin of pronotum which

antennae distally

proportions:

head black,

with segments 1-5 and part I y

propodeum black, dark;

length 2,06-2,12 mm;

testaceous and temples reddish-dark;

6:

6:

antennal

segments

6:

4:

head dull,

4,5:

4:

4:

8;

in following granulated and

keels on frons and vertex;

OL = 3;

OPL = 2;

OOL = 4;

is

with stalk of hind femora

thickened;

longitudinal

POL = 4,5;

legs testaceous,

occipital

TL = 4;

carina

pronotum dull,

except for a narrow region near posterior margin smooth;

posterior surface of pronotum shorter than scutum tubercles reaching

tegulae;

scutum,

(5:10);

pronotal

scutellum and metanotum shiny,

smooth, weakly punctate, without sculpture among punctures; notaul ices incomplete, reaching approximately 0,3 length of scutum; propodeum reticulate rugose,

with a

posterior surface;

posterior surface without

rugose;

hyaline,

fore-wing

radial

strong

transversal

without dark

in following (Fig.

314)

proportions: located on a

with

apical

part

1,1,2. MALE:

unknown

part 11;

(3:8);

29

holotype F!

keels,

bands;

lamellae,

and

reticulate

distal

fore tarsal

enlarged claw

prominence;

with basal lamellae;

part of

segments

(Fig.

314)

segment 5 of front

apex with a group of 4

Barro Colorado

TYPICAL MATERIAL:

2: 5:

1,5:

1 row of nearly

(4,5:5,5);

LOCU5 TYPICUS: HOSTS:

6:

proximal

between dorsal

transversal

vein much shorter than proximal

one bristle

keel

longitudinal

with

tarsus

part shorter than tibial

spurs

Island (Panama) in LE;

1 paratype F!

in OL.

unknown

DISTRIBUTION:

Neotropic region:

PAt,AMA:

Barro Colorado

Island,

LE!

OL!

5. Anteon famaicanum n. sp.

FEMALE:

fully

winged;

length 2,62 mm;

head black,

clypeus and anterior half of frons testaceous; segment

1 testaceous;

antenna I segments 6:

8;

head dull,

dull,

OL = 4;

antennae distally

in following proportions:

10:

5:

8:

with

thickened;

7:

7:

6:

5,5:

reticulate rugose and with nUmerous longitudinal

frons without median 5,5;

legs testaceous;

with mandibles,

antennae brown,

and

OOL = 5,5;

lateral OPL = 3;

keels;

occipital

TL = 5;

carina complete;

5,5: keels; POL

=

pronotum with anterior region

rugose, striate; posterior surface shiny,

smooth,

not rugose,

shorter

than scutum (7:13); pronotal tubercles reaching tegulae; scutum with anterior region near anterior margin smooth and shiny, without sculpture;

498

Mem.

Fig.

315 -

Amer.

Ent.

no.

37,

Anteon jamaicanum

Chela of

posterior half of scutum dull,

Ins!.,

punctate,

with

1984

n.

sp.

(type)

irregular striae,

notaulices visible only near anterior margin of scutum;

rugose;

scutellum and

metanotum shiny,

smooth, without sculpture; propodeum reticulate rugose, strong transversa I keel between dorsa I and posterior surface; posterior surface without longitudinal keels; fore-wing hyaline, without wi th a

dark transversal part 18;

(4:8);

bands;

fore tarsal

enlarged claw

prominence;

(Fig.

distal

part of radial

segments

in following

315)

vein shorter than proportions:

with one bristle

segment 5 of front

lamellae extending continuously

tarsus

(Fig.

located on a

315)

to the apex;

7:

proximal

2,5:

3:

9:

proximal

with 2 rows of nearly 26

tibial

spurs 1,1,2.

MALE: fully winged; length 1,56 mm; black; mandibles testaceous; antennae testaceous-dark; legs testaceous; antennae distally thickened; antennal segments in following proportions: 8: 3: 6: 5: 5: 5,5: 5,5: 5: 8; head dull, reticulate rugose and granulated; frons with a median keel; TL

occipital

=

2;

carina complete;

scutum,

scutellum

POL = 4,5;

without sculpture among punctures; margin of scutum; keel

distal

and posterior surface; fore-wing hyaline,

part of radial

gonoforceps tibial

keels; (Fig.

316)

3;

=

OOL = 4;

smooth,

near anterior

with a strong transversal

posterior surface without

without dark

vein much shorter than without distal

OPL = 2;

weakly punctate,

notaulices visible only

propodeum reticulate rugose,

between dorsal

longitudinal

OL

and metanotum shiny,

5:

transversal

proximal

part

bands;

(2:7);

inner pointed or rounded process;

spurs 1,1,2.

LOCUS TYPICUS: Hardwar Gap (Jamaica) TYPICAL MATERIAL: holotype F! and 1 paratype M!

in OT.

Anteoninae

Fig.

316 -

499

Male genital ia of Anteon jamaicanum n.

HOSTS: unknown DISTRIBUTION: Neotropic region:

JAMAICA:

sp.

Hardwar Gap,

(paratype)

OT!

6. Anteon mexicanum n. sp.

FEMALE:

fully

winged;

antennae brown,

length 2,00-3,18 mm;

coxae and clubs of femora partly brown; antennal

black;

with segments 1-3 testaceous;

mandibles testaceous;

legs testaceous,

with hind

antennae distally thickened;

segments in following proportions:

10: 5:

5:

4:

4:

4:

5:

5:

5:

6,5; head dull, granulated and reticulate rugose; sometimes head with a few areolae, mostly smooth on vertex; frons with a median keel; occipital carina complete; POL = 5,5; Ol = 3,5; OOl = 6; OPL = 4; Tl = 4; pronotum du II, granu I ated and with transversa I keel s; pronota I tubercles reaching tegulae;

posterior surface of pronotum shorter than scutum

500

Mem.

Fig.

(6:12);

317 -

scutum dull,

Amer.

Chela of

En!.

granulated;

without sculpture;

transversal without

keel

keels;

37,

1984

notaulices

indistinct,

and posterior surface;

fore-wing

faintly

visible

scutellum and metanotum shiny,

propodeum reticulate rugose,

between dorsal

longitudinal

no.

Anteon mexicanum n. sp. (type)

only near anterior margin of scutum; smooth,

Ins!.,

hyaline,

without

with

a

posterior surface dark

transversal

Anteoninae

501

318 - Male genitalia of Anteon mexicanum n. sp. from S. Cristobal

Fig.

de las Casas

(Mexico):

right

half removed.

bands; distal part of radi21 vein shorter than tarsal segments in following proportions: 5: 2:

proximal

2:

part

2,5: 9,5;

(2:6); fore enlarged claw

317) with one bristle located on a proximal prominence; segment 5 tarsus (Fig. 317) with 2 rows of 6 lamel!ae; apex with a group of 8 lamellae; tibial spurs 1,1,2. MALE: fully winged; length 1,81-2,12 mm; black; mandibles testaceous; (Fig.

of front

antennae black; thickened; 5:

5:

5:

median

legs black,

antenna I segments

7; keel;

head dull,

with

tarsi

I ight;

antennae not distally

in following proportions:

9:

granulated and reticulate rugose;

occipital carina complete;

POL = 5;

OL = 3;

5:

5:

5:

5:

5:

frons with OOL = 5;

OPL

3; TL = 3,5; scutum dull, granulated; notaul ices indistinct, faintiy visible only near anterior margin of scutum; scutellum and metanotum

502

Mem.

shiny,

smooth

transversal without

Amer.

distal

(1,5:6);

between dorsal

(Fig.

with distal

LOCUS TYPICUS:

1984

propodeum reticulate rugose,

and posterior surface;

318)

de

TYPICAL MATERIAL:

holotype F!

3 paratypes

las Casas

tibial

2 MM)!

F,

part

membranous

spurs 1,1,2.

(Mexico)

and 5 paratypes

(1

transversal

proximal

proximal

inner rounded process;

with a

posterior surface

without dark

without dorsal

paratype F! HOSTS:

37,

vein much shorter than

San Cristobal

in TW;

no.

fore-wing hyaline,

part of radial

gonoforceps

process and

I nst.,

and without sculpture;

keel

longitudinal keels;

bands;

Ent.

(1

in

F,

4 MM)!

in OT;

OL.

unknown

DISTRIBUTION: (Chiapas),

Neotropic region:

OT!

OL!

JAMAICA:

MEXICO:

Blue Mt.,

San Cristobal

OT!

PERU:

de

las Casas

Machu Picchu,

TW!

7. Anteon oliveirai n. sp.

FEMALE:

fully

winged;

antennae brown,

distally 7:

7:

thickened;

8:

with a

with

7:

7:

length 4 mm; segments

antennal

7:

10;

median keel

segments

head shiny,

and

two

black;

mandibles testaceous;

1-2 testaceous;

legs

testaceous;

in following proportions:

antennae

19:

9:

irregularly sculptured by folds;

lateral

keels around orbits;

occipital

10:

frons carina

complete; POL = 6,5; OL = 4; OOL = 6; OPL = 6,5; TL = 3; region behind ocelli weakly granulated, with irregular folds and with tracks of two oblique keels connecting posterior ocelli to occipital carina; pronotum shiny, with a strong transversal impression between anterior and posterior surface; transversal reaching

anterior part of posterior surface forming

carina,

tegulae;

shiny,

with

pronotal

posterior surface shorter than scutum

scutellum and metanotum shiny, sculpture among punctures; 0,3

irregular folds;

length of scutum;

smooth,

weakly

notaul ices incomplete,

narrow dark band beneath proportions:

11:

located on a

proximal

with

2:

3:

the pterostigma;

part 3:

(6:10);

11,5; enlarged claw

prominence;

tibial

spurs 1,1,2. unknown

LOCUS TYP I CUS:

Serra do Caraca

TYPICAL MATERIAL: HOSTS:

holotype F!

approximately

with a strong posterior surface fore-wing with a

part of radial

vein

segments in following (Fig.

319)

segment 5 of front

two rows of nearly 10 lamellae;

MALE:

distal

fore tarsal

scutum,

without

reaching

transversa I keel between dorsa I and posterior surface; without longitudinal keels, dull and reticulate rugose; shorter than proximal

raised

(7,5:19);

punctate,

propodeum reticulate rugose,

a

tubercles

apex with a

with one bristle

tarsus

group of 2

(Fig.

319)

lamellae;

(Brazi I) in TW.

unknown

DISTRIBUTION:

Neotropic region:

Minas Gerais), TW! NOTES: the species is named

BRAZIL:

Serra do Caraca

in honor of Mr.

F.M.

(5.

Oliveira.

Barbara,

Anteon i nae

Fig.

319 -

503

Anteon oliveirai n. sp. (type)

Chela of

8. Anteon noyesi n. sp.

FEMALE: black

fully

winged;

segments 7-10 brown; distally 4:

length 2,62-3,00 mm;

3,5:

thickened; 5:

7:

6:

6:

9;

punctures,

hairy,

reddish;

antennae

segments in following proportions:

head shiny,

occipital carina complete;

pronotum shiny,

with propodeum

antennae sometimes with

propodeum sometimes partly

antennal

weakly punctate,

without sculpture among punctures; orbits; = 6;

testaceous,

(mesopleura and metapleura testaceous);

smooth,

frons with only 2 lateral

POL = 5;

punctate,

Ol = 3;

7:

6:

keels around

OOl = 6;

without sculpture,

OPl = 5;

transversal

Tl

among

with posterior surface shorter than scutum

posterior surface of pronotum with 2 dorsal

10:

mainly on frons,

(4:20);

lobes;

pronotal

tubercles reaching tegulae; scutum, scutellum and metanotum shiny, smooth, wihtout sculpture; notaulices invisible; propodeum shiny, with a strong

transversal

keel

between dorsal

and posterior surface;

dorsal

surface very short, with a few areolae; posterior surface shiny, smooth, without sculpture, without areolae or keels; fore-wing with 2 dark transversal pterostigma; (4: 10);

bands, one on the basa I cell s and another beneath the distal

fore tarsal

enlarged claw

(Fig.

part of radial segments 320)

with one bristle

prominence;

segment 5 of front

than apical

part

nearly 31

(6:11);

lamellae;

MALE: unknown lOCUS TYPICUS:

St.

TYPICAL MATERIAL:

vein much shorter than proximal

in following proportions: tarsus

located on a

(Fig.

segment 5 of front

apex with a Augustine

7:

group of 6

320)

2,5:

3:

7:

part 17;

proximal

with basal

part shorter

tarsus with some rows of lamellae;

tibial

spurs 1,1,2.

(Trinidad)

holotype F!

and

1 paratype F!

in BM;

1 paratype F!

504

Mem. Amer.

En!.

Inst.,

no.

37,

1984

, \

\ I

t Fig.

in aT;

320 -

1 paratype F!

Chela of Anteon noyesi n.

TRINIDAD:

BM!

Amazonas,

NOTES:

(Arima),

J/

sp.

(type)

in TW.

HOSTS: unknown DISTRIBUTION: Neotropic region: Simla

)

OT!

BRAZIL:

SI.

Augustine

TW!

the species is named in honor of Mr. J.S.

Noyes.

(St.

George),

505

Anteoninae

Fig.

321 -

Chela of Anteon lobatum n.

sp.

(type)

506

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

9. Anteon lobatum n. sp.

FEMALE:

fully

winged;

reddish-testaceous; brown;

length 2,5-3,0 mm;

antennae testaceous,

scutum reddish-testaceous,

scutellum reddish-dark; metapleura black, reddish-brown; segments

in

head and prothorax fully

with segments 7-10 or 9-10

with sides and anterior half black;

metanotum and propodeum black;

with more or

less wide reddish

legs testaceous-dark;

following proportions:

tegulae

11:

6:

spots;

testaceous;

6:

4:

4:

6:

7:

mesopleura and abdomen antennal 7:

7:

9;

head

smooth, hairy, shiny, weakly punctate, without sculpture among punctures; frons without longitudinal median keel, with two lateral around orbits on

the frons POL = 6;

carina complete; shiny,

smooth,

pronotal

and metanotum shiny, visible only

reticulate rugose,

forming

tUbercles reaching

smooth,

lateral

sockets;

two dorsal

tegulae;

without sculpture;

transversal

keel

areas reticulate rugose;

lobes near

scutum,

scutellum

notaulices indistinct,

near anterior margin of the scutum;

with a

keels

occipital

8; OPL = 5; TL = 6; pronotum

OOL =

hairy,

between dorsal

posterior surface with 2 longitudinal

sudace; shiny;

OL = 3;

weakly punctate,

posterior margin; faintly

directed towards antennal

keels;

propodeum and posterior

median area smooth,

mesopleura and metapleura smooth;

fore-wi ng with 2 dark transversa I bands, one on the basa I cell sand another beneath the pterostigma; distal part of radial vein much shorter than proximal part (4:9); fore tarsal segments in following proportions: 3: 2: 3: 5: 14; enlarged claw (Fig. 321) with one bristle located on a proximal

prominence;

nearly 26

lamellae;

segment 5 of front

tarsus

(Fig.

apex with a group of nearly 6

321)

with 2 rows of

lamellae;

tibial

spurs

1,1,2. MALE: unknown LOCUS TYPICUS:

Gamboa

TYPICAL MATERIAL: F!

(Panama)

holotype F!

and 2 paratypes FF!

in LE;

1 paratype

in OL.

HOSTS:

unknown

DISTRIBUTION:

Neotropic region:

PANAMA:

Gamboa

(Canal

Zone),

LE!

OL!

10. Anteon propodeale (Fenton) n. comb.

Chelogynus propodealis Fenton 1927: 13.

FEMALE: tegulae, brown; 4:

6:

with a

fully winged; length 2,25 mm; black; mandibles, antennae, legs testaceous, except hind stalks of femora and hind tibiae antenna I segments

8;

in following proportions:

head very shiny and smooth,

median

longitudinal

keel

11:

5:

without sculpture,

and with 2

lateral

5:

4:

hairy;

4:

4:

4:

frons

keels around orbits;

Anteoninae

507

o

A

Fig.

322 -

Anteon chiriquense (Cameron) propodeale (Fenton) (type) (B) (laterally viewed).

Head and prothorax of females of

(type) (A)

and

occipital carina complete; POL = 5; OL = 3; OOL = 5; OPL = 3,5; TL = 3,5; pronotum hairy, very shiny and smooth, without sculpture, convex and raised (Fig.

into a

322 B);

transversal

pronotal

carina between

tubercles reaching

metanotum very shiny and smooth, invisible;

longitudinal

dorsal

keels;

scutum scutellum and

without sculpture,

propodeum with a transversal

and posterior surface; with 2

anterior and posterior half

tegulae;

hairy;

irregular keel

surface reticulate rugose;

median area shiny,

with short keels only near margins;

lateral

smooth,

notaulices

between dorsal posterior surface without areolae,

areas with

wide areolae;

fore-wing with 2 dark transversal bands, one on the basal cells and another beneath the pterostigma; distal part of radial vein much shorter than proximal 4:

1,5:

2:

proximal nearly as

4:

(2:6);

segments in following proportions:

(Fig.

323)

segment 5 of front

long as distal to

fore tarsal

enlarged claw

prominence;

continuously MALE:

part 10;

part,

with nearly

the apex;

tibial

Santarem

(Brazil)

with one bristle

tarsus 14

(Fig.

323)

located on a

with basal

lamellae extending

spurs 1,1,2.

unknown

LOCUS TYPICUS:

TYPICAL MATERIAL: HOSTS: unknown DISTRIBUTION:

holotype F!

Neotropic region:

in WA. BRAZIL:

Santarem

(Para),

WA!

part

508

Mem.

Fig.

323 -

Amer.

En!.

Ins!.,

no.

37,

1984

Anteon propodeale (Fenton)(type)

Chela of

11. Anteon chiriquense (Cameron) n. comb.

Dryinus chiriquensis Cameron 1888: 447. f'4esodryinus chiriquensis (Cameron): Kieffer 1914b: 32.

FEMALE:

fully

winged;

legs testaceous, basally black;

length 3,62-6,00 mm;

with stalks of hind femora antennae testaceous,

testaceous or brown-testaceous;

black; dark;

mandibles testaceous;

sometimes hind coxae

with segments 6-10 dark;

head hairy,

dull,

fully

abdomen

reticulate rugose;

occipital carina complete; POL = 5; OL = 3; OOl = 7; OPl = 6 or 4; Tl = 9; pronotum hairy, punctate, forming a raised transversal carina between anterior and posterior surface sometimes with an

(Fig.

322 A);

incomplete median keel;

than posterior surface;

pronotal

posterior surface of pronotum

anterior surface much shorter

tubercles reaching

tegulae;

sides of

posterior surface not rounded, but raised into longitudinal carinas; scutum, scute II um and metanotum smooth, sh i ny, weak I y puncta te, without sculpture among punctures; 0,25

length of scutum;

notaul ices short,

propodeum with a

incomplete,

transversal

keel

reaching only

between dorsal and posterior surface; dorsal surface reticulate rugose; areolae very wide, with internal surface granulated; posterior surface with 2 longitudinal keels; median and lateral areas almost fully smooth, shiny; mesopleura and metapleura shiny and smooth; fore-wing with 2 light or dark transversa I bands, one on the basa I cell s and another beneath the

Anteoninae

Fig.

pterostigma; (4: 13);

distal

fore tarsal

enlarged claw prominence; lamellae; MALE:

324 -

(Fig;

Chela of

509

Anteon chiriquense

part of radial

(Cameron) (type)

vein much shorter than proximal

segments in following proportions: 324)

8:

4:

4:

part

10: 24;

with one bristle located on a proximal

segment 5 of front

tarsus

(Fig.

apex with a group of 5 lamellae;

324) tibial

with 2 rows of nearly spurs 1,1,2.

unknown

LOCUS TYPICUS:

Volcan de Chiriqui

TYPICAL MATERIAL: HOSTS:

holotype Fl

(Panama)

in BM.

unknown

DISTRIBUTION:

Neotropic region:

PANAMA:

Volcan de Chiriqui,

BM!

BOLIVIA: 10 Km E San Antonio (Rio Mamore, Dept. Beni), AM! BRAZIL: Nova Teutonia (S.ta Catarina), OT! 12° 31' S 55° 37' W (Sinop, Mato Grosso),

TWl

18

510

Mem.

Fig.

Amer.

En!.

Inst.,

no.

37,

1984

325 - Chela of Anteon nycteum n. sp.

(type)

12. Anteon nycteum n. sp.

FEMALE:

fully

winged;

of frons and a small reddish;

length 3,37 mm;

head black,

spot between posteior ocelli

clypeus and mandibles testaceous;

testaceous,

with hind coxae basally black;

and abdomen

black;

antennae distally

with anterior region

and occipital

carina

antennae testaceous; thorax,

thickened;

propodeum,

legs petiole

antenna I segments

in

following proportions: 11: 9: 8: 8: 8: 8: 8: 8: 8: 11; head shiny, with frons fully reticulate rugose; frons without median keel, but with a median furrow connecting anterior ocellus to clypeus;

vertex smooth,

punctate,

numerous parallel

median striae connecting anterior ocellus to occipital

carina; = 3,5;

head

with

OL = 5,5;

without sculpture among punctures,

not

reticulate rugose,

two keels around orbits; OOL = 8;

OPL = 12;

with

occipital carina complete;

TL = 9;

pronotum shiny,

POL

with a

strong transversa I impression;

anterior surface of pronotum much shorter

than posterior surface

pronotum form i ng a

(8: 17);

ra i sed transversa I

carina between anterior and posterior surface; posterior surface smooth, shiny, weakly punctate, without sculpture among punctures, except for SOme transversa I str; ae near the transversal reaching

tegulae;

scutum,

cari na;

pronota I tuberc I es

scutellum and metanotum shiny,

smooth,

weakly

Anteoninae

punctate,

approximately 0,5

transversal with

two

keel

length of scutum;

between dorsal

longitudinal

511

keels;

propodeum with a

and posterior surface;

dorsal

surface reticulate rugose;

areas of posterior surface rugose;

median area smooth,

hyal ine,

bands;

without dark

transversal

shorter than proximal proportions:

11:

located on a

3:

part

4:

10:

proximal

lamellae;

tibial

22;

fore tarsal

enlarged claw

prominence;

with 2 rows of nearly 28 MALE:

(6: 14);

distal

lamellae;

strong

posterior surface lateral

shiny;

fore-wing

part of radial

vein

segments in following (Fig.

325)

segment 5 of front apex with a

with one bristle

tarsus

(Fig.

325)

group of nearly 6

spurs 1,1,2.

unknown

LOCUS TYPICUS:

12° 31'

TYPICAL MATERIAL: HOSTS:

S 55°

37'

holotype F!

W (Sinop,

Mato Grosso,

Brazil)

in TW.

unknown

DISTRIBUTION: Grosso), TW!

Neotropic region:

BRAZIL:

12° 31'

S 55° 37'

W (Sinop,

Mato

13. Anteoll annulicorne Brues

Anteon annulicornis Brues 1905: 185.

FEMALE;

fully

winged;

length 2,75 mm;

head black,

with mandibles

testaceous; antennae testaceous, with segments 2-4 brown, dark; thorax, propodeum and abdomen black; legs brown, light;

antennae distally thickened;

proportions:

11;

numerous short

5:

8;

=

OPL

forming

a

surface; reaching

5;

5:

5:

irregular keels;

keels around orbits; 5,5;

4:

=

TL

median

occipital 5;

6:

antennal 5:

5:

segments median

carina complete;

POL

smooth,

carina between

scutum shiny,

granulated;

granulated,

keel

=

7;

hairy,

and OL

=

with

two lateral 4,5;

OOL

=

without sculpture,

anterior and posterior

posterior surface shorter than scutum tegulae;

in following

head dull,

frons with a

pronotum shiny,

transversal

8;

8-10 brownwith tarsi

(7,5: 17);

pronota I tubercles

notaulices very reduced,

faintly visible near anterior margin of scutum; scutellum and metanotum shiny, amooth, without sculpture, punctate; propodeum with a strong transversal

keel

between dorsal

reticulate rugose;

and posterior surface;

posterior surface without

numerous transversal

irregular keels;

dorsal

longitudinal

fore-wing

with

surface

keels,

with

two dark transversal

bands, one on the basal cell s and another benea th the pterost i gma; distal part of radial vein shorter than proximal part (2:10); fore tarsal segments tarsus with

in following proportions:

(Fig.

326)

with basal

two rows of nearly

enlarged claw prominence;

(Fig.

tibial

326)

part

20:

8:

8,5:

22:

46;

longer than apical

segment 5 of front part

17 lamellae extending continuously with one bristle

spurs 1,1,2.

(28: 18)

and

to the apex;

located on a proximal

512

Mem.

Fig.

MALE:

326 -

Amer.

Chela of

Ent.

Inst.,

no.

37,

1984

Anteon annulicorne Brues (type)

unknown

lOCUS TYPICUS:

Mexico

TYPICAL MATERIAL:

holotype F!

HOSTS: unknown DISTRIBUTION: according

in PH.

to the morphology of the holotype,

I

think

that

A. annulicorne is a Neotropic species. The Mexican locality is unknown. The morphology is very similar to A. albitarse (Cameron).

14. Anteon albitarse (Cameron) n. comb.

Dryinus albitarsis Cameron 1888: 447. Nesodryinus albitarsis (Cameron): Kieffer 1914b: 33.

FEMALE:

fully

winged;

or partly black;

length 1,81-3,00 mm;

antennae testaceous,

segments 6-10 black;

tegulae testaceous;

hind coxae partly black; brown;

with

mid,

black;

mandibles testaceous

last segment brown or with

legs testaceous,

with all or only

hind and sometimes fore clubs of femora

mid and hind tibiae brown;

antenna I segments in following

proportions: 12: 6: 7: 5: 4: 5: 5: 5: 5: 7; head dull, granulated, smooth; frons with a longitudinal median keel and with 2 lateral keels near orbits directed towards antennal POL = 6;

Ol = 4;

OOl = 6;

OPl = 5;

sockets; Tl = 6;

occipital

carina complete;

pronotum dull,

granulated,

Anteoninae

Fig.

327 -

transversal

keels on anterior half;

pronotum forming

med i an cari na between anterior and posterior ha If;

tubercles reaching not visible;

Anteon albitarse (Cameron) (type)

Chela of

with numerous transversal

513

tegulae;

scutum dull,

smooth,

scutellum and metanotum smooth,

granulated,

shiny,

a

pronota I

notaul ices

without sculpture or

weakly punctate; propodeum reticulate rugose, with a transversal between dorsal and posterior surface; posterior surface without

keel

longitudinal keels; mesopleura and anterior half of metapleura dull and rugose; posterior half of metapleura smooth, shiny, without sculpture; fore-w i ng

wi th 2 dark

another beneath than proximal 6:

2:

3:

Bol ivia);

5:

part

16

transversa I bands,

the pterostigma; (in

(2,5:8);

(Fig.

lamellae; apex with a MALE: unknown LOCUS TYPICUS:

DISTRIBUTION:

1,5:

the basa I cell sand

3:

8:

(Fig.

327)

vein much shorter

in following 20

with one bristle

tarsus

group of nearly 17

Panajachel

TYPICAL MATERIAL: HOSTS: unkown

segments

or 5:

327)

segment 5 of front

one on

part of radial

fore tarsal

the holotype)

enlarged claw

prominence;

distal

located on

with

lamellae;

proportions:

(in a specimen from a proximal

1 row of nearly

tibial

spurs 1,1,2.

(Guatemala)

holotype F!

Neotropic region:

in BM. MEXICO:

Vista Hermosa

(Tuxtepec,

12

514

Mem.

Fig.

328 -

Amer.

Ent.

Inst.,

no.

37,

Male genitalia of Anteon vivax n.

1984

sp.

(type):

right

half

removed.

Oaxaca),

OL!

TW!

GUATEMALA:

(5t. George),

OL!

BM!

Panajachel,

COLOMBIA:

Domingo, TW! BOLIVIA: Chulumani Notes; in the borderlands between

Cal i

BM!

(Valle),

TRINIDAD: CAl

51. Augustine

ECUADOR:

Quito /

5.

(La Paz), BM! the Neotropic and Nearctic regions,

A. albitarse can be confused with A. iunestum (Nearctic). The two species are distinguished by the fore wings (hyal ine in iunestum and very dark in albitarse).

15. Anteon vivax n. sp.

FEMALE: MALE:

unknown

fully

winged;

antennae brown,

length 1,93 mm;

black;

mandibles testaceous;

with segments 1-2 testaceous;

coxae basally brown;

legs testaceous,

antennae not distally thichened;

antennal

with hind segments

Anteoninae

Fig.

515

Male genitalia of Anteon pilicorne Ogloblin from Burzaco

329 -

(Argentina) (A:

left

half removed);

B:

gonoforceps.

in following proportions: 8: 6: 7: 6: 7: 7: 7: 7: 6,5: 9; head shiny, punctate, without sculpture among punctures; frons without median and lateral =

2,5;

keels;

occipital

Tl = 4;

scutum,

carina complete;

sculpture among punctures; 0,3

length of scutum;

proximal

notaulices incomplete,

part

(3:

7);

TYPICAL MATERIAL:

de

holotype M!

bands;

OOl

=

punctate,

with 2

gonoforceps

las Casas

3;

reaching

(Fig.

5,5;

approximately areas;

part of radial 328)

without

inner pointed

(Mexico)

in OT.

unknown

DISTRIBUTIOI',: (Chiapas),

Neotropic region:

MEXICO:

San

Cristobal

OT!

16. Anteon pilicorne Ogloblin

Anteon pilicornis Ogloblin 1938: 39.

de

OPl

without

longitudinal

lateral

distal

membranous process and without distal

process; tibial spurs 1,1,2. lOCUS TYPICUS: San Cristobal HOSTS:

=

Ol

median area as rugose as

without dark transversal

vein shorter than proximal dorsal

5;

propodeum reticulate rugose,

keels on posterior surface; fore-wing hyal ine,

=

POL

scutellum and metanotum weakly

las Casas

516

Mem.

FEMALE: MALE:

Amer.

Ent.

Inst.,

no.

37,

1984

unknown

fully

winged;

antennae brown,

mm; black; mandibles testaceous;

length 2,75-3,43

with segment

1 testaceous;

legs testaceous,

and sometimes clubs of femora partly brown;

with coxae

sometimes also hind

tibiae

brown; antennae not distally thichened; antenna I segments in following proportions: 14: 6: 6: 6: 7: 7: 5: 6: 5: 9; head dull, fully reticulate rugose;

frons with a median keel

and with 2

;

5; OOL ;

7; OPL ;

5,5; TL ;

some areolae on anterior third; approximately 0,3 scu I pture; transversal without

keel

between

longitudinal

dark transversal proximal

part

occipital

notaul ices

length of scutum;

metanotum rugose;

bands;

(2:9);

membranous process

incomplete,

reaching

scutellum shiny,

smooth,

and posterior surface;

reticulate rugose;

distal without

(Fig. distal

329)

without

with a strong

posterior surface

fore-wing hyaline,

part of radial

gonoforceps and

keels near orbits

carina complete;

propodeum ret icu I ate rugose,

dorsal

keels,

lateral

POL; 8; OL 5; scutum dull, fully granulated, with

directed towards antenna I sockets;

without

vein much shorter than

without dorsal

proximal

inner pointed process,

with

a

series of papi Ilae on inner face; tibial spurs 1,1,2. LOCUS TYPICUS: General Urquiza (Buenos Aires, Argentina) TYPICAL MATERIAL: HOSTS:

DISTRIBUTION: Veado, BM!

holotype M!

in BA.

unknown Neotropic region:

Distrito Federal),

General

Urquiza

OL!

BRAZIL:

aT!

Estacao Forestal

ARGENTINA:

(Buenos Aires),

(Cabeca do

Burzaco (Buenos Aires),

BA!

17. Anteon conterminum n. sp.

FEMALE: "'·,ALE:

unknown

fully

winged;

antennae brown,

testaceous,

length 2,06

with

segment

mm; black; mandibles testaceous;

1 testaceous;

legs brown,

fore tibiae and femora partly testaceous;

with

tarsi

antennae not

distally thickened; antennal segments in following proportions: 10: 4,5: 4: 4,5: 5: 5: 5: 5: 5; antennal segment 10 missing in the only known specimen;

head dull,

granulated and fully

median complete keel antennal

sockets;

OPL; 2,5; weak I y

TL ;

two

occipital 3;

puncta te,

incomplete,

and

scutum,

lateral

carina complete;

without scu I pture among punctures;

faintly

posterior surface;

posterior surface without

without dark trasversal

shorter than proximal LOCUS TYPICUS:

tibial

part

(1 :7);

trasversal bands;

keel

distal

(Brazil)

3;

OOL ;

4;

smooth, propodeum

between dorsal keels;

part of I'adial (Fig.

towards

notau I ices

longitudinal

gonoforceps

spurs 1,1,2.

Nova Teutonia

frons with a

OL ;

visible near anterior margin of scutum; with a strong

pointed process;

POL; 5,5;

scutellum and metanotum shiny,

reticulate rugose, hyal ine,

reticulate rugose;

keels near orbits directed

and fore-wing vein much

330) with distal

inner

Anteoninae

Fig.

330 -

517

Anteon conterminum n. sp. (type): left half

Male genitalia of

removed.

TYPICAL MATERIAL: HOSTS:

holotype M!

in OT.

unknown

DISTRIBUTION: OT!

Neotropic region:

BRAZil Nova Teutonia

(Sta Catarina),

18. Anteon plaumanni n. sp.

FEMALE: MALE:

unknown

fully

winged;

length 2,0 mm;

head black;

testaceous;

thorax black or brown-reddish;

testaceous;

antennae not distally

following proportions: rugose;

frons

with

9:

5:

7:

a median

thickened;

6:

sockets;

~

Tl

OOl

weakly

~

5,

OPl

punctate,

~

2,5;

without

6:

~

2;

scutum,

transversal with 2

keel

wing hyal ine,

6:

9;

and

keels;

with 2

membranous process and

reticulate keels

POL ~ 5;

Ol

scutellum and metanotum

notaulices

in addition scutum

incomplete, with

and posterior surface;

(3:7);

in

lateral

carina com,:Jiete;

median area as rugose as

without dark trasversal

legs

segments head fully

propodeum reticulate rugose,

much shorter than proximal proximal

6:

sculpture among punctures;

between dorsal

longitudinal

antennal

keel

occipital

reticulate rugose near anterior margin; mid-length of scutum;

6:

longitudinal

directed towards antennal 3;

6:

mandibles and antennae

abdomen black;

bands;

gonoforceps

distal

posterior surface

lateral

areas;

part of radial

(Fig.331)

without distal

reaching

a strong forevein

without dorsal

inner pointed process;

518

Mem.

Fig.

331

Amer.

Ent.

Inst.,

no.

37,

Male genitalia of Anteon plaumanni n.

-

1984

sp.

(type):

right

half

removed.

tibial

spurs 1,1,2.

LOCUS TYPICUS:

Nova Teutonia

TYPICAL MATERIAL: M! in OL. HOSTS:

(Brazil)

holotype M!

in BM;

1 paratype M!

in OT;

1 paratype

unknown

DISTRIBUTION:

Neotropic region:

Nova Teutonia

(S.ta Catarina)

NOTES:

the species is named

JAMAICA:

Hardwar Gap,

OT!

OL!

BRAZIL:

BM!

in honor of Mr.

Fritz Plaumann.

19. Anteon bolivianum n. sp.

FEMALE:

fully

winged;

distally thickened; 3:

2:

5:

7:

only keel POL = 5;

7:

6:

length 2,52 mm;

antennal 9;

head dull,

is around the orbits), OL =

3,5;

fully

reddish-testaceous;

segments in following proportions:

OOL = 9;

not reticulate rugose, granulated;

OPL = 5;

occipital

TL = 6;

with posterior tubercles reaching

with a

impression;

transverse

furrow;

long,

scutum,

sculpture,

very

scutell urn

approximately 0,30

tegulae;

smooth,

pronotum

lobes separated by

and metanotum sh i ny,

weakly punctate;

smooth,

notaul ices incomplete,

length of scutum;

5: (the

carina complete;

posterior surface of pronotum nearly

with two prominent dorsal

5:

without keels

pronotum shiny,

without sculpture, as broad as

antennae 11:

twice

a median

wi thout

reaching

propodeum reticulate rugose,

with a

Anteoninae

332 - Chela of Anteon bolivianum n. sp.

Fig.

strong

transversal

surface with weakly

keel

two weak

rugose,

the basal

519

shiny;

between dorsal

and posterior surface;

longitudinal keels; fore-wing

(type)

median and

lateral

with two dark transversal

cells and another beneath

the pterostigma;

distal

posterior areas

bands,

one on

part of

part (3:9); fore tarsal segmens 7: 3: 3: 8: 21; enlarged claw (Fig. 332) with one bristle located on a proximal prominence; segment 5 of front tarsus (Fig. 332) with two rows of nearly 40 lamellae; apex with a group of nearly 17 lamellae; tibial spurs 1,1,2. radial

vein much shorter than proximal

in following proportions:

MALE:

unknown

LOCUS TYPICUS:

Mouth of Rio Baures (Bolivia)

TYPICAL MATERIAL: HOSTS:

DISTRIBUTION: Itenez,

holotype F!

in AM.

unknown Dept.

Neotropic region: Beni),

BOLIVIA:

Mouth of Rio Baures

AM!

20. Anteon micros n. sp.

(Rio

520

Mem.

Fig.

333 -

Amer.

En!.

Ins!.,

no.

37,

1984

Anteon micros n. sp. (type)

Chela of

FEMALE: fully winged; length 1,64 mm; black; mandibles testaceous; legs brown, with fore tibiae and tarsi yellow; antennae distally thickened; antennal 5;

segments

head dull,

carina complete; dull,

granulated;

POL = 5;

granulated,

more than

in following proportions:

fully

Ol =

frons 3;

7:

3:

3:

3:

3:

2,5:

without median keel;

OOl = 3;

OPl = 4;

Tl 4;

with posterior surface shorter than scutum

twice as broad

as

long

(15:4);

3:

3:

4:

occipital pronotum (4: 10)

and

posterior tubercles reaching

tegulae; scutum dull, granulated; notaul ices incomplete, reaching approximately 0,30 length nf scutum; scutellum and metanotum shiny, smooth,

without

transversal

keel

sculpture;

propodeum

between dorsal

reticulate

rugose,

and posterior surface;

with

a

strong

posterior surface

Anteoninae

without

longitudinal

bands;

distal

keels;

521

fore-wing hyaline,

part of radial

without dark

fore tarsal

segments in following proportions:

5:

claw

333)

proximal

(Fig.

with one bristle

segment 5 of front

group of 4

apex with a MALE:

tarsus

transversal

vein much shorter than proximal

(Fig.

located on a 333)

lamellae;

2:

3,5:

part

9;

(2:5);

enlarged

prominence;

two rows of nearly 8 lamellae;

with

tibial

2:

spurs 1,1,2.

unknown

LOCUS TYPICUS:

Miranda

TYPICAL MATERIAL: HOSTS:

(Venezuela)

holotype F!

in AM.

unknown

DISTRIBUTION:

Neotropic region:

VENEZUELA:

Miranda

(Caracas),

AM!

21. Anteon surinamense n. sp.

FEMALE:

fully

winged;

length 3,20 mm;

testaceous,

with posterior margin

of metanotum and med i an area of propodeum brown; thickened; 8:

8:

7:

9;

head dull,

lower face; keel

antennae d i sta II y

antenna I segments in following proportions: occipital

POL

smooth,

of pronotum nearly

with a

carina complete;

around orbits;

pronotum shiny,

granulated,

=

8;

OL

with a

=

6:

5:

OOL

transversal long,

=

10;

=

OPL

impression;

4:

keels on

frons without median keel,

4;

twice as broad as

12:

track of weak

6;

=

TL

3:

7:

the with a

9;

posterior surface

shorter than scutum

(9:18),

wi th two dorsa I transverse lobes separated by a med I an furrow; posterior tubercles reaching tegulae; scutum shiny, smooth, without sculpture; notaul ices

indistinct,

faintly

visible near anterior margin of scutum;

scutellum and metanotum shiny, a strong transversal

keel

smooth,

without sculpture;

between dorsal

surface of propodeum reticulate rugose; without

longitudinal

sculpture,

keels,

and

propodeum with

posterior surface;

dorsal

posterior surface of propodeum

with median area shiny,

surrounded by areolae and,

smooth,

near the bottom,

by

without transversal

keels; fore-wing with two dark transversal bands, one on the basal cells and another beneath the pterostigma; radial vein with distal part shorter than proximal 9:

3,5:

4:

proximal nearly 37 1,1,2. MALE:

9:

part 23;

(4: 11);

fore tarsal

enlarged claw

prominence; lamellae;

(Fig.

segments 334)

segment 5 of front apex with a

in following proportions:

with one bristle

tarsus

(Fig.

group of nearly

12

334)

located on a

with 2 rows of

lamellae;

unknown

LOCUS TYPICUS:

holotype F!

TYPICAL MATERIAL:

in AM.

holotype F!

HOSTS: unknown. DISTRIBUTION: Neotropic region: Paramaribo,

AM!

in AM. SURINAME:

Botanic Gardens of

tibial

spurs

522

Mem. Amer.

Fig.

334 -

Chela of

Ent.

Inst.,

no.

1984

Anteon surinamense n. sp. (type)

ANTEON

AUSTRALIAN SPECIES OF

Fourty

37,

two species are known.

KEY TO THE SPECIES

FEMALES

Segment

4

of

front

tarsus at

Segment

4

of

front

tarsus

longer 2 3

than

most

0,5 as

slightly

surface of

propodeum with

Posterior

surface of

propodeum without

fully

Head at

reticulate

most

than,

segment as

•• 1..2

long as,

or

1 .•.••••....•••.••••••.••••••..•...•.••.. 10

segment

Posterior Head

long as

shorter

rugose.

reticulate

1.

rugose

two

longitudinal longitudinal

keels •••• 3 keels ••••• 4

giluwense n. sp.

only

on anterior half

of

frons

or behind ocell i, wi th a wide, punctate and smooth frontal area in front of the ocell i •.•••.• 2. chelogynoides (R.C.L. Perkins) 4

Head

smooth,

reticulate

punctate,

rugose;

without

posterior

sculpture among punctures,not

surface

of

propodeum almost

523

Anteoninae

smooth,

not

reticulate

rugose,

keel s •••••••••••••••••••••••• Head

reticulate rugose;

reticulate

5

posterior

or with

strong

Scutum reticulate

rugose,

or with

wi thout

surface

of

transversal strong

propodeum keels ••••••••••.• 5

or weak

sculpture or very

Propodeum wi th a

strong

surface;

longitudinal

transversal

scutum very

keels •••••••••••

Propodeum usually without dorsal

and posterior

transversal

keel

Scutum fully

weakly

faintly

irregular punctate

or

between dorsal sculptured by

transversal

occasionally

visible among

aulicum n. irregular

sides ..

keel

between

weak

the other keels . . . . . . . . 7

5.

the

with a

rugose ••••••••••.••.••••••••••

sculptured by weak

mainly visible on

and

irregular

reticulaticeps Dodd

4.

reticulate

• ..•...••..•......•..........

Antennal

keel

strongly

a distinct

surface;

strongly

Scutum shiny,

8

longitudinal

weakly granulated • • • • • • • • • • • • • • . . • • • . . • • • • . . • • . • • • • • • . • • • 8

posterior

7

short

keels •••••••••••••••••••••••••••.••.••••••..••.•• 6

Scutum smooth, very

weak and

myrmecophilum (R.C.L. Perkins)

rugose

longitudinal

6

with 3.

sp. longi tudinal

keels

sedlaceki n. sp.

6.

1 nearly as long as segments 2+3 . . . . . . . . . . . . . .

segment

. . . . . . . . . . . . . . . . . . . . . . . . • . . . . 8. australe n. sp. Antennal segment 1 nearly as long as segments 2+3+4 . . . . . . . . . . . 9

9

Fore-wing wi thout

dark

transversal

· . . . . . . . • . . . . . . . . . . . . . • . . . . .. Fore-wing with a dark

bands •••••••••.•••••••••••••

anxium n.

7.

transversal

• • • . . • • . • • . • • • . . • . . • • • . • • . •• 41. 10 Posterior surface of propodeum with Posterior

surface of

sp.

band •••••••••.•••••••••.. rufiscapum n. sp. two

propodeum without

longi tudinal longitudinal

keels ••. 11 keels . . . . 23

11 Fore-wing hyaline, without dark transversal bands . . . . . . . . . . . . 12 Fore-wi ng wi th 1-2 dark transversa I bands . . . . . . . . . . . . . . . . . . . . 20 12 Head granulated, not reticulate rugose or reticulate rugose only behind Head at 13 Head

ocell i . . . . . . . . . . .

least

partly

9.

reticulate

reticulate rugose

parvulum (R.C.L. Perkins) rugose

on anterior half

on of

frons and vertex . . . 13 frons,

punctate

on

vertex •••••••••••••••••••••••••••••••••••••.••••••••••••••••• 14 Head

fully

reticulate

rugose

or with

numerous

longitudinal

k eel s on f ron s • • • • . . • • . • • • • • • • • • • . • • • • • • • • • • • • • • . • • . • • • . • • • . • 1 5 14 Median area of posterior surface of propodeum smooth •••••••••••

• • • • • • • • • • • • • • • • • . • • • • • • • • •• Median

area of

posterior

· . . . . . . . . . . . . . . . . . . . . . . . . . .. 15 Scutum fully

reticulate

Scutum punctate,

not

16 Scutellum smooth; anterior margin Scutellum partly

orientale n. sp. propodeum rugose ••••••••.••

11. aculeatum n.

sp. rugose . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 16

reticulate

notaulices

of

10.

surface of

scutum . . .

reticulate

rugose ••••.•••••.•••••••••••• 17

invisible or faintly

12.

visible near

corruptum n. sp.

rugose;

notaulices

reaching

0,75 length of scutum •••••••••••••••••••••••••••• • • . • • . . • . . • . • • . . • . . • . . . . . . .. 1 3. miles n. sp.

approximately

524

Mem.

Amer.

17 Frons with numerous

Ent.

Inst.,

longitudinal

no.

37,

1984

keels . . . . . . . . . . . . . . . . . . . ..

• • • • • • • • • • • • • • • • • • • • . • • • • • •• 18. curraJoi n. sp. Frons fully reticulate r u g o s e . . . . . . . . . . . . . . . . . . . . . . . 18 Head with OPL much

shorter

· • • • • • • • • • • • • • . • • • • • • • . • . • •• Head with OPL 19 Segment

4

of

longer front

4

of

front

• • • • . . • • • • . . • • • . 20 Fore-wing wi th

tarsus

longer

wi th

sp.

tarsus

shorter

. ...•.•..• dark

than

1 ••••.••••••••••

than

segment

1 •••••••••••

superbum Dodd

40.

transversa I

band beneath

the

typicum (R.C.L. Perkins)

16.

2 dark

segment

destructor (R.C.L. Perk i ns)

14.

pt eros! i gma ••••• Fore-wing

caledonianum n.

15.

than TL . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 19

.. .. .. . . . . . . . . . . . . . . . . . . . . Segment

.18

than TL . . . . . . . . . . . . . . . . ..

transversal

bands,

one

on

the basal

cell s

21

and another beneath the pterost igma . . . . . . . . . . . . . . . . . . . . . . . . . . 21 Head with frons sculptured by strong longitudinal keels, not

22

Head fully reticulate rugose . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 22 Scutum reticulate rugose . . . . 19. firmum n. sp.

reticulate

rugose . . . . . . . . . . .

Scutum punctate,

not

reticulate

............................ 23 Fore-wing

wi th

Fore-wing 24 Segment

4

1-2 dark

of

tarsus nearly

front

tarsus

wi th

1 dark

leiosomum (R.C.L. Perkins)

transveral

wi thout

5 of

rugose •••••••••••••••••••••••••

20.

hyal ine, front

bougainvillei n. sp.

17.

dark

wi th basal

bands . . . . . . . . . . . . . . . . . . . . . 24

transversal as

part

bands . . . . . . . . . . . . 25

long as

long

segment

(Fig.

363);

1;

segment

fore-wing

Segment

4

of

front

band . . . . . . . . . . . . . . . . . . . . . . . . . . . .. 21. nitidum (R.C.L. Perkins) tarsus nearly twice as long as segment

segment

5

of

front

tarsus with basal

fore wing

transversa I

wi th

•...•.•.

2 dark

transversal

•.••.•.•••••••••••

25 Head

fully

reticulate

Head

fully

or almost

26 Head with numerous

rugose fully

22.

part

(Fig.

bands.......

1

365)

.. . . . . . . .

bouceki n. sp.

or with

longitudinal

granulated,

longitudinal

short

keels,

keels . . . . . . 26

not

reticulate

rugose.28

not

reticulate

rugose

. . . . . . . . . . . . . . . . . . . . . . . . . . . . 23. mundum n.

sp.

Head fully reticulate rugose . . . . . . . . . . . . . . . . . . . . . 27 Segment 1 of front tarsus shorter than segment 4 • •••••••••••••••••••••••••••• 28

42.

Segmen t

of

front

tarsus

Segmen t

5 of

front

tarsus almost

d i st i nct

longer

5 of front

29 Scutum wi thout

sculpture,

areas very

sp

segment

rect i I inear,

4 . . . . . . . . . . . . . 28 wi thout

a

25.

bellator n. sp. distinct

curved apex

(Fig.368)

nigricorne (R.C.L. Perk i ns)

shiny and

smooth,

except

for

the

weakly granulated •••••••••••••••••••••

• • • • . • • • • • • • • • • • • • • • • • • • • • •• Scutum fully

24.

tarsus with a

•••••••••••••••••••••••••••• lateral

than

curved apex. (Fig .367) •••••••••••••••••••••••• ••••••••••••••••••••

Segment

giraulti n.

• .27

26.

fiji anum n.

sp.

granulated ••••••••••••••••••••••••••

.. .. 29

Anteoninae

525

30 Species shiny; notaulices reaching approximately 0,5 length of scutum . . . . . . . . . . . . . . . . . . . . . . 27. dubium Fouts Species dull; notaul ices shorter, reaching approximately 0,25 length of scutum ..••••.•.••• 28. coriaceum (R.C.L. Perkins)

The females of A. sucklingi, bismarckense, gloriosum, vulsum, luctuosum, hageni, haustum, corallinum, tasmanianum, permirum and involutum are unknown.

MALES

2

3

4

5 6

7

8

Propodeum wi thout a strong transversal keel between dorsal and posteri or surface . . . . . . . . . . . . . . . . . . . . . .2 Propodeum wi th a strong transversal keel between dorsa I and posterior surface . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 Posterior surface of propodeum with two longitudina keels .•

· . . . . . . . . . . . . . . . . . . . . . . . . . . . 29. sucklingi n. sp. Posterior surface of propodeum wi thout longi tudinal keel s . . . . • 3 Head smooth, weakly punctate, wi thout sculpture among punctures • • • . . . . . . • • . . . • • • . . 3. myrmecophilum (R.C.L. Perkins) Head rugose, fully reticulate rugose • • . • . . • . • . . . . . • . . . . . • . . . · . . . . . . . . . . . . . . . . . . . . . . . . . .. 5. aulicum n. sp. Posterior surface of propodeum with two longitudinal keels . . . . . . . . . . . . ... 5 Post eri or surface of propodeum wi thout longitudinal keels . . . . 15 Gonoforceps wi th a long dorsal process (Figs 356, 357, 372) . . . 6 Gonoforceps wi thout long dorsal process (Figs 373, 374, 375) .. 7 Dorsa I process of gonoforceps very long and narrow, wi th apex curved (Figs 356, 357); scutum very weakly punctate . . . . . . . . . .. · . . . . . . • . •. . . ••••. . . . . . 15. caledonianum n. sp. Dorsal process of gonoforceps shorter and broad, wi th apex not curved (Fig.372) scutum very strongly punctate; punctures on scutum very broad . . . . . . . . . . 30. bismarckense n. sp. Head fully granulated . . . . . . 31. gloriosum n. sp. Head reticulate rugose, or punctate or with numerous longitudinal keels on frons . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 Head fully or mostly reticulate rugose or with longitudinal keels on frons . . . . . . . . . . • . . . . . . . . . . . . . . • . . . . . • • • . . . . • • . . . • • . . . 9 Head

9

punctate,

at

most

with

areolae near

occipital

carina,

on malar space and on anterior half of frons . . . • . • . . . . . . . • • . • 13 Scutum almost fully reticulate rugose or with weak irregular keels . • . . . . . • • • • • . • • . . . • • • . 32. VUlSUIll n. sp. Scutum punctate and wi thout sculpture among punctures, not . . . 10 reticulate rugose . . . . . . . . . .

526

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

10 Median area of

posterior

surface of

propodeum smooth ...•••••• 11

Median area

posterior

surface of

propodeum rugose • • • • • • • • 12

11

Head

fully

of

reticulate

rugose ••••••••..•••••••••••••••••••

• •••••••••••••••••••••••••• 33. luctuosum n. sp. Head with frons sculptured by numerous longitudinal

keel •••.••.

hageni n. sp. haustum n. sp. 14. destructor (R.C.L. Perkins)

· ••. . . . . . . . . . . . . . . •. . . . . . .

34. 35.

12 Species mostly reddish •.•. Species mostly black •••••••

13 Notaulices reaching approximately 0,5 length of scutum •••••• •. • . ••. . . . . . •. . . •. . • . . •. 36. corallinum n. sp. Notaulices very reduced, at most reaching 0,3 length of scutum • •• 14 14 Median area

of

posterior

surface of

• • • • • • • • • • • • • • • • • • • • • • • • • •• Median area of

posterior

37.

propodeum smooth ••••.

tasmanianum n. sp.

surface of

• . . . • . . . • . . . • • . • • . • • • • • . • •.

1 0.

propodeum rugose. sp.

orientale n.

15 Head punctate, without sculpture among punctures •••• • . . . . . . . . • . • • • • • • • • • . . • • . . . 21. nitidum (R.C.L. Perkins) Head ret iculate rugose or with longi tudinal keels, or granu I at ed ••••••••.•••••••••...•••••••••••..••••••••••••••••• 16 16 Head granulated, sometimes also wi th a few irregular keels •••

· . . . . . . . . • . . . • . . . . . . . . . . . . . • 8.

australe n.

Head with numerous

keels

longitudinal

on

sp.

frons

or

reticulate

rugose •••.•.•••••••••••••••.••••••••••••••••••••••••••••••••• 17

17 Head with numerous longitudinal keels on frons ••••••••••••••••• · . . . . . . . . . . . . . . . . . . . . . . . . . . 38. permirum n. sp. Head

fully

or partly

18 Scutum punctate,

reticulate

rugose ••••••••••••••••••••••• 18

sculpture among punctures •••••• • . . . . . • • • . . . . . . . • • • • . • . . . . . 21. nitidum (R.C.L. Perkins)

Scutum fully

wi thout

weakly granulated.

39. involutum n.

· . . . . . . . . . . . . . . . . . . . . . . . . ..

sp.

The males of A. dubium, coriaceum, superbum, hjianum, nigricorne, reticulaticeps, bouceki, typicum, leiosomum, parvulum, chelogynoides, bougainvillei, anxium, corruptum, giraulti, giluwense, curradoi, sedlaceki, aculeaturn, miles, firmum, mundum, bellator and rufiscapum are unknown.

1. Anteon giluwense sp.

FEMALE:

fully

light;

length 2,50 mm;

winged;

antennae black,

with segment 1 partly

antennae distally thickened;

proportions:

8:

5:

8:

6:

6:

5:

5:

5:

black;

antennal 5:

mandibles testaceous;

testaceous; 8;

legs brown,

segments

head dull,

with

tarsi

in following fully

reticulate

Anteoninae

Fig.

335 -

527

Chela of Anteon giluwense n.

sp.

(type)

rugose; frons with a complete median keel; occipital carina complete; OL = 3; OOL = 4,5; OPL = 4; TL = 3; pronotum dull, reticulate

POL

= 5;

rugose and with transversal

keels;

pronotal

tubercles reaching

posterior surface of pronotum shorter than scutum scutellum and metanotum shiny, sculpture among punctures; 0,65 length of scutum; transversal

keel

smooth,

scutum,

weakly punctate,

without

notaul ices incomplete,

reaching approximately

propodeum reticulate rugose,

between dorsal

tegulae;

(3:11);

with a strong

and posterior surface;

posterior surface

with two longitudinal keels; median aerea rugose; fore-wing hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part (3:7); fore tarsal segments in following proportions: 8:

2:

2:

2:

12; enlarged claw

proximal prominence; nearly 12 bristles; MALE:

(Fig.335)

with one bristle located on a

segment 5 of front tarsus

(Fig.

apex with a group of 10 lamellae;

335)

with 2 rows of

tibial

spurs 1,1,2.

unknown

LOCUS TYPICUS:

Mt.

TYPICAL MATERIAL:

Giluwe (New Guinea) holotype F!

HOSTS: unknown DISTRIBUTION: Australian region:

in TW. NEW GUINEA:

Mt.

Giluwe,

TW!

2. Anteon chelogynoides (R.C.L. Perkins)

Prosanteon chelogynoides R.C.L. Perkins 1905: 66. Prosanteon melanostigmus R.C.L. Perkins 1906a: 493 (n. syn.); locus typicus Mittagong (Austral ia). Anteon chelogynoides (R.C.L. Perkins): R.C.L.Perkins 1907: 50. Anteon melanostigmus (R.C.L. Perkins): Kieffer 1914b: 170. Chelogynus cognatus (R.C.L. Perkins) partim: Kieffer 1914b: 193.

528

Fig.

Mem. Amer.

336 -

Chela of

En!.

Ins!.,

no.

37,

1984

Anteon chelogynoides (R.C.L. Perkins) from Pimel ia (Western Austral ia)

FEMALE:

fully

winged;

with teeth brown;

length 2,25-3,43 mm;

antennae fully

part of segment 2 yellow;

black;

mandibles yellow,

testaceous or brown with segment 1 and

legs yellow,

with only hind coxae brown

(in

chelogynoides) or with all coxae, femora and mid and hind tibiae brown (in melanostigmus); antennae distally thickened; antennal segments in

Anteoninae

following proportions: punctate, keels;

9:

4,5:

4,5:

4:

4:

529

4:

4:

4:

4:

with frons strongly punctate and rugose; OL = 3;

POL = 5;

OOL = 4;

OPL = 4,5;

numerous areolae and transversal than scutum

(5:11)

scutum smooth,

keels,

notaulices incomplete,

reaching

head shiny,

TL = 4,5;

with

irregular

pronotum with

with posterior surface shorter

and slightly shorter than

weakly punctate,

6;

vertex

anterior surface

without sculpture among approximately 0,3-0,5

(5:6)

punctures;

length of scutum;

scutellum and metanotum smooth, weakly punctatae, without sculpture among punctures; propodeum reticulate rugose, with a transversal keel between

dorsal

longitudinal rugose; radial

and posterior surface;

keels;

fore-wing

median hyal ine,

posterior surface with or shiny,

with basal

part

numerous bristles; proximal MALE:

2:

2:

3:

part

10;

enlarged claw tibial

(Fig.

(3:6);

two

lateral

bands;

areas

distal

fore tarsal

part of

segments

segments 5 of front

tarsus

part

lamellae and

longer than apical

prominence;

smooth;

without dark transversal

vein much shorter than proximal

in following proportions:6: 336)

area dull

336)

(6:4),

with 9

with one bristle

(Fig.

located on a

spurs 1,1,2.

unknown

LOCUS TYPICUS:

Bundaberg

TYP I CAL IvlATER I AL:

(Queensland,

lectotype F!

Australia)

Pro chelogynoides in B; lectotype F!

of

Pro melanostigmus in B. in Australia Tartessus sp. (?)(R.C.L. Perkins 1906c) for the var. chelogynoldes. of

HOSTS:

DISTRIBUITION: B!

Mittagong

Australian

region:

(New South Wales),

AUSTRALIA:

B!

Wanneroo

Bundaberg (32 mls.

( Queensland),

NE of Perth,

Western Australia), 8M! Pimelia (Penberton, Western Australia), OL! Kununurra (North Western Austra I i a), OL! Mt. Bundey Road (v i a Darw in, Northern Territory),

UQ!

3. Anteon myrmecophilum (R.C.L. Perkins)

Paranteon myrmecophllus R. C. L. Perk i ns 1905: 67 Anteon myrmecophilus (R.C.L. Perkins): Kieffer 1907b: 27

FEMALE:

fully

winged;

with metanotum, not distally 7:

8:

8:

8:

length 2,74-2,93 mm;

propodeum,

thickened; 7:

7:

7:

antenna I segments

8;

yellow-reddish,

petiole and abdomen

head convex,

sometimes

reddish-brown;

antennae

in following proportions:

strongly punctate,

14:

6:

without sculpture

among punctures; frons without median and lateral keels; POL = 5; OL = 3; OOL = 7; OPL = 6; TL = 5; pronotum short, faintly visible in dorsal view;

scutum convex,

dull,

weakly punctate,

without sculpture among

punctures, with weak longitudinal keels on anterior half; notaul ices distinct, incomplete, reaching approximately 0,3 length of scutum; scutellum shiny,

smooth,

very convex,

without sculpture,with a

weak

530

Mem.

Fig.

337 -

Chela of

Amer.

Ent.

Inst.,

metanotum shiny,

propodeum with dorsal surface by a

weak

1984

(Queensland)

smooth,

surface dull,

transversal

without sculpture,

rugose,

keel

posterior margin of propodeum; lateral

37,

Anteon myrmecophilum (R.C.l. Perkins) from Bundaberg

median furrow;

no.

very convex;

separated from posterior

semicircle shaped,

not reaching

posterior surface smooth and shiny;

areas rugose and dull; dorsal surface shorter than breadth of fore-wing with a dark transversal band beneath the

metanotumj

pterostigma; part

distal

(5:7 or 4:5):

(5:2,2); tarsus basal

part of radial segment

fore tarsal (Fig.

337)

segment 3 produced

long as segment 4;

located on a MALE:

fully

tarsus much into hook;

proximal winged;

twice as

segment 5 of front

prominence;

tibial

length 2,56-2,74 mm;

lamellae at apex;

long as apical

enlarged claw

(

proximal

longer than segment 4

with numerous bristles and some

part of segment 5 nearly

nearly as

vein slightly shorter than

1 of front

Fig.

337)

part;

distal

part

with one bristle

spurs 1,1,2. brown-black;

antennae brown;

tibiae and tarsi light; tegulae testaceous; mandibles brown, with teeth light; antenna I segments in following proportions: 9: 4,5: 5: 6: 6: 6: 5: 5:

5:

6;

head shiny,

smooth,

convex,

without sculpture among punctures; Tl = 3;

scutum sh i ny,

anterior third;

smooth,

and sparse punctures,

Ol = 3;

OOl = 6;

OPl = 2,5;

with strong and sparse punctures on

posterior surface without punctures or weakly punctate,

without sculpture among punctures; reaching

with strong

POL = 6;

approximately 0,5

notaul ices distinct,

length of scutum;

shiny,

smooth,

without sculpture,

shiny,

smooth,

not reticulate rugose;

convex;

semicircle shaped separating dorsal the posterior marg i n of propodeum;

incomplete,

scutellum and metanotum

mesopleura and metapleura

propodeum with a

transversal

and posterior surface, dorsa I surface rugose,

keel

not reaching shorter than

531

Anteoninae

Fig.

338 -

Male genitalia of Anteon myrmecophilum Bundaberg

breadth of metanotum; near dorsal surface;

(Queensland):

right

(f'.C.L.

posterior surface almost smooth, lateral

without dark transversal

areas reticulate rugose;

bands,

with distal

Perkins)

from

half removed.

sl ightly rugose

fore-wing

part of radial

hyal ine,

vein shorter

than proximal part (4:7); gonoforceps (Fig. 338) without inner distal process and without dorsal proximal membranous process, narrow in dorsal

view;

tibial

LOCUS TYP I CUS:

TYP I CAL MATER I AL: HOSTS:

(Austra I i a)

lectotype F!

and 2 para I ectotypes FF!

in Australia Jpo conferta Kirk.

DISTRIBUTlm,: Sandhi II

spurs 1,1,2.

Bundaberg

Australian region:

(Queensland),

(R.C.L.

AUSTRALIA:

in B.

Perkins 1906c).

Bundaberg

OL!

4. Anteon reticuiaticeps Dodd

(Oueensland),

B!

532

Mem. Amer.

Fig.

339 -

Chela of

Ent.

Ins!.,

no.

37,

1984

Anteon reticulaticeps Dodd from Finisterre Mts. (New Guinea)

Anteon reticuiaticeps Dodd 1914a: 131.

FEMALE: fully winged; length 2,12-2,18 mm; black; mandibles testaceous, with teeth brown; antennae brown, with segments 1 or 1-2 testaceous; tegulae brown-testaceous; legs brown; sometimes tarsi testaceous; antennae distally thickened; antenna I segments in following proportions: 10: 5:

5:

4:

longitudinal 3;

TL

~

3;

4:

fully

between dorsal

vein

with one bristle

with basal

part much

frons with a

OL

~

3;

smooth,

transversal

keel

7:

transversal

proximal 1,5:

a

1,5:

part 1,5:

proximal

bands;

(4:12); 6;

fore

enlarged

prominence;

with some bristles and 2 apical

longer than distal

part;

tibial

spurs

1,1,2. MALE:

unknown

LOCUS TYP ICUS:

Nelson

TYPICAL MATERIAL:

(Austral ia)

holotype F!

HOSTS: unknown DISTRIBUTlm,: Australian

region:

~

keels and

with a

without dark

located on 339)

OOL

posterior surface without

much shorter than

(Fig.

6;

irregular longitudinal

in following proportions: tarsus

~

POL

scutellum and metanotum shiny,

fore-wing hyal ine,

segments

segment 5 of front

reticulate rugose; keels;

pronotum very short and reduced;

and posterior surface;

keels;

339)

lateral

propodeum reticulate rugose,

tarsal

lamellae,

head fully

sculptured by

part of radial (Fig.

6;

without

carina complete;

distal claw

4:

notaulices invisible;

without sculpture; longitudinal

4: 4:

occipital

scutum dull, granulated;

4:

median keel,

in AD. NEW GUINEA:

Finisterre IMs.

(Madang

Anteoninae

Fig.

Dist.),

340 -

BM!

Wau

Chimbu div., AUSTRALIA:

Chela of Anteon aulicum n.

(Morobe Distr.,

NE New Guinea),

Nelson

533

sp.

from Taui

NE New Guinea),

B!

(N Queensland),

Mt.

Kaindi

B!

(New Guinea)

Daulo Pass

(NE New Guinea),

AD!

5. Anteon aulicum n. sp.

(AsaroOL!

534

""em.

FEMALE:

fully

winged;

antennae testaceous, with

Amer.

Ent.

Inst.,

length 2,75 mm;

shiny,

in following proportions:

fully

keels;

reticulate rugose;

occipital

black;

16:

5:

POL ~ 5;

surface of pronotum shorter than scutum tegulae;

reaching

approximately 0,3

keel

rugose;

between

bristle 340)

8:

2:

2,5: 4:

located on

bands;

(3:10); 13;

a proximal

6:

length 2,18-2,31

distal

half light;

light;

antennae not distally

proportions:

antennae black;

TL ~ 5;

keels;

posterior

tubercles

notaul ices

incomplete,

without a strong

transversal

posterior surface without

transversely striate; distal

fore-wing

part of radial

segments

12: 6:

7:

8:

mm;

8:

8:

8:

8:

tarsus with basal (Fig.

340)

black;

12;

head dull,

part

with one

tarsus

(Fig.

tothe apex;

mandibles black,

with fore

with

tibiae and

antenna I segments

8:

vein

in following

segment 5 of front

legs brown,

thickened;

head

and 2 lateral

with 2 rows of nearly 16 lamellae extending continuously

tibial spurs 1,1,2. MALE: fully winged;

antennal 8;

OPL ~ 4;

enlarged claw

prominence;

5:

scutellum and metanotum

fore tarsal

(10:3);

5:

pronotal

segment 5 of front

part

legs testaceous,

keel

OOL ~ 6;

(7:18);

reticulate rugose and

longer than apical

5:

median

length of scutum;

part

5:

transversal

and posterior surface;

proximal

proportions:

5:

propodeum reticulate rugose,

keels,

mandibles testaceous;

reticulate rugose;

without dark transversal

shorter than much

scutum shiny,

dorsal

longitudinal hyal ine,

7:

reticulate rugose and with

reaching shiny,

1984

antennae distally thickened;

frons with a

carina complete;

pronotum dull,

37,

with segments 2-3 and 9-10 brown;

hind coxae proximally brown;

segments

no.

tarsi

in following

fully

reticulate

rugose; frons with a com,,,:ete median keel; occipital carina complete; ~ 6; OL ~ 3; OOL ~ 6; OPL ~ 2; TL ~ 2; scutum dull, with anterior surface fully by

reticulate rugose and with posterior third weakly sculptured

irregular striae;

sh i ny,

weak I y

notaul ices invisible;

punctate,

scutellum and metanotum smooth,

without scu I pture among punctures;

reticulate rugose,

without a strong

posterior surface;

posterior surface without

hyaline,

without dark transversal

shorter than proximal inner pointed process;

part

(3:8);

tibial

transversal bands;

keel

longitudinal

distal

gonoforceps

in B;

HOSTS:

3 paratypes MM!

propodeum

between

dorsal

keels;

part of radial

(Fig.

341)

and

fore-wing vein

without distal

spurs 1,1,2.

LOCUS TYP ICUS: Mt. Gi luwe (New Guinea) TYPICAL MATERIAL: holotype F! and 4 paratypes MM! F!

POL

in TW;

1 paratype

in OL.

unknown

DISTRIBUTION: Guinea),

B!

OL!

Gap

Tari

Australian

Mt. Giluwe, (Mt.

region: OL!

Hagen),

TW!

NEW GUINEA: Daulo Pass,

Taui OL!

(Mt.

TW!

Mt.

Kaindi,

TW!

6. Anteon sedlaceki n. sp.

FEMALE:

fully

winged;

length 3 mm;

black;

NE

Wilhelm,

mandibles testaceous;

New TW!

Anteoninae

Fig.

341

535

Male genitalia of Anteol1 aulicum n.

-

sp.

from Mt.

Giluwe

(New

Guinea)

with segment 1 proximally

antennae brown, antennae distally

thickened;

antennal

testaceous;

segments

legs testaceous;

in following proportions:

16: 6: 6: 6: 5: 5: 5: 5: 5: 9; head fully reticulate rugose, with a complete median keel;

4; OOl

=

5; OPl

=

4; Tl

=

occipital

carina complete;

5; pronotum strongly rugose,

dull; frons POL = 5,5; Ol = with numerous

transversal keels, with posterior surface shorter than scutum (5:18); pronotal tubercles reaching tegulae; scutum shiny, punctate, without sculpture among punctures, keels mainly on incomplete,

lateral

smooth,

with a

very

weak

faintly

visible among

length of scutum;

without sculpture;

transversal

irregular longitudinal

and near posterior margin;

approximately 0,5

reaching

metanotum shiny,

with numerous weak

regions

keel

between

the other keels;

notaul ices

scutellum and

propodeum reticulate rugose, dorsal

and posterior surface,

posterior surface of propodeum

without longitudinal keels; fore-wing hyal ine, without dark transversal bands; distal part of radial vein shorter than proximal part (4:10); fore tarsal

segments

(Fig.

342)

front

tarsus

in following proportions:

with

1 bristle

(Fig.

342)

located on

lOCUS TYPICUS:

(New Guinea)

HOSTS:

4:

13;

enlarged claw

prominence;

segment 5 of

14 lamellae extending

in TW.

unknown

DISTRIBUTIOi~:

TW!

Gap

holotype F!

3:

spurs 1,1,2.

tibial

Tari

2:

with 2 rows of nearly

continuously to the apex; MALE: unknown TYPICAL MATC:RIAl:

9:

a proximal

Australian

region:

NEW GUINEA:

Tari

Gap

(Mt.

Hagen),

536

Mem. Amer. Ent.

Inst.,

no. 37,

1984

Fig. 342 - Chela of Anteon sedlaceki n. sp.

NOTES:

(type)

the species is named in honor of Mr. J. Sedlacek.

7. Anteon anxium n. sp.

Anteoninae

Fig.

FEMALE:

fully

with proximal testaceous

343 -

winged;

5:

4:

3:

rugose,

3:

thickened;

4,5: 4,5: OL ;

legs black,

antennal

4:

median keel,

POL; 6;

black;

4:

7;

without

2,5;

segments

with

lateral

OOL ;

5;

fully

keels;

OPL ;

smooth,

tegulae;

scutum,

approximately 0,3

or

testaceous;

5;

reticulate rugose;

occipital TL ;

carina

4;

pronotum dull,

(3:16);

pronotal

scutellum and metanotum shiny,

without sculpture or weakly punctate;

reaching

tarsi

in following proportions:

head shiny,

with posterior surface shorter than scutum;

tubercles reaching

mandibles testaceous,

antennae brown with segment 1 testaceous,

with segments 2-3 brown;

frons with a complete;

Anteon anxium n. sp. (type)

length 2,00-2,50 mm;

half black;

antennae distally 14:

Chela of

537

length of scutum;

notaul ices

incomplete,

propodeum reticulate rugose,

wi th a strong transversa I keel between dorsa I and posterior surface; posterior surface without longitudinal keels, rugose, with transversal keels;

fore-wing

hyaline,

without dark transversal

bands;

distal

part of

538

Mem.

radial

Amer.

Ent.

Ins!.,

vein much shorter than proximal

in following proportions: basal

part

6:

2:

longer than apical

one bristle

located on

2:

3:

part

a proximal

tibial

MALE: unknown LOCUS TYPICUS:

(2:7);

fore tarsal

(8:2);

enlarged

claw

segments

tarsus with

(Fig.

343)

segment 5 of front

with

tarsus

lamellae extending continuously

to

spurs 1,1,2. Wamena

TYPICAL MATERIAL:

(New Guinea)

holotype F!

in B;

HOSTS: unknown DISTRIBUTION: Australian region: Numbu

1984

segments 5 of front

prominence;

343) with 2 rows of nearly 11

(Fig.

the apex;

part

10;

37,

no.

(Upper Chimbu V.,

NE

1 para type F!

NEW GUINEA:

New Guinea),

in OL.

Wamena

(Irian

Barat),

B!

OL!

8. Anteon australe n. sp.

FEMALE: fully

fully

winged;

coxae partly brown; partly brown; proportions:

occasionally

5:

6:

median

OOL = 6;

5:

5:

5:

TL = 3;

mandibles brown;

legs testaceous,

hind coxae, thickened;

5:

longitudinal

OPL = 3;

mm;

1 testaceous;

antennae distally

10:

frons with a = 5;

length 2,37-2,81

brown or with segment

5:

5:

keel,

hind femora and

antennal

7;

segments

head dull,

without

occipital

antennae

with hind hind tibiae in following

reticulate rugose;

lateral

keels;

carina complete;

POL = 9;

OL

pronotum dull,

reticulate rugose, with transversal keels, almost smooth on posterior surface; posterior surface shorter than anterior surface and shorter than scutum (4:16); scutum shiny, smooth, without sculpture or very weakly granulated; notaulices incomplete, reaching anterior third of scutum; scutellum and metanotum shiny, smooth, without sculpture; propodeum reticulate rugose,

with a

posterior surface;

posterior surface without

rugose;

with a

fore-wing

distal

part of radial

tarsal

segments

tarsus with basal (Fig.

MALE:

tibial

fully

tarsus

7,5:

located on 344)

with

keels,

and

reticulate

band beneath the pterostigma; proximal 2:

longer than apical

(Fig.

between dorsal

15

3:

part

3:

part

15;

(3:8);

(10:5);

a proximal

fore

segment 5 of enlarged

prominence;

lamellae and numerous

spurs 1,1,2.

winged;

antennae brown;

part

with one bristle

segment 5 of front

keel

longitudinal

dark transversal

in following proportions:

front

bristles;

transversal

vein much shorter than

claw

344)

strong

length 1,62-1,81

legs brown,

mm;

black;

mandibles testaceous;

with fore tibiae and tarsi

light;

antennae

not distally thickened; antennal segments in following proportions: 6: 4: 5: 5: 5: 5: 5: 5: 5: 7; head dull, granulated, sometimes with irregular keels on ocellar region;

frons with a short median

front of the anterior ocellus; OOL = 5;

OPL = 2;

TL = 3;

occipital

scutum,

keel

carina complete;

visible only POL

=

5;

OL

in

=

scutellum and metanotum shiny,

smooth, weak I y punctate, wi thout scu I pture among punctures; notau I ices incomplete, reaching only 0,25 length of scutum; ;:->ropodeum reticulate

3;

Anteoninae

Fig.

344 -

539

Chela of Anteon australe n.

rugose,

with a strong transversal

surface;

posterior surface without

keel

sp.

between dorsal

longitudinal

keels;

(type)

and posterior fore-wing

hyaline,

without dark transversal bands; distal part of radial vein shorter than proximal part ( 2:5); gonoforceps (Fig. 345) without distal inner pointed process; tibial spurs 1,1,2. LOCUS TYP ICUS: Dongarra (Western Austral ia) TYPICAL MATERIAL: F,

2 MM)!

HOSTS:

in TW;

holotype F! 3 paratypes

and (1

F,

1 paratype F! 2 MM)!

in BM;

3 paratypes

in OL.

unknown

DISTRIBUTION: Austral ia), (Tasmania),

Australian

BM! TW!

Yuendumu Waldheim

region:

AUSTRALIA:

Dongarra

(Northern Territory), (Tasmania),

TW!

OL!

TW!

(Western Roseberry

OL!

9. Anteon parvulum (R.C.L. Perkins)

nee Anteon pallidieornis Ashmead 1893: 97. Neoehelogynus parvulus R.C.L. Perkins 1905: 65. Neoehelogynus pallidieornis R.C.L. Perkins 1905: 65 (n. syn.); locus typieus Bundaberg (Austra I i a) nec Anteon parvulus Kieffer in Kieffer et Marshall 1905: 193. Anteon laetieornis Kieffer 1907b: 26; new name.

(1

540

Mem.

Fig.

345 -

Amer.

Ent.

Inst.,

no.

37,

1984

Male genitalia of Anteon australe n. sp. (Austral ia): left half removed.

from Yuendumu

Anteon parvulus (R.C.L. Perkins): Kieffer 1907b: 27. Chelogynus pallidicornis (R.C.L. Perkins): Kieffer 1914b: 191. Chelogynus parvulum (R.C.L. Perkins): Kieffer 1914b: 192.

FEMALE:

fully

winged;

with teeth brown; brown with

length

with segments

1-2 testaceous;

hind femora brown;

sides)

of body;

long as broad;

1,83-2,38 mm;

black;

antennae yellow or slightly body shiny,

head granulated;

tegulae and hairy

reaching

occasionally

legs yellow,

sometimes

(mainly on head and on

pronotum granulated,

scutum without sculpture or very

notaulices incomplete,

mandibles yellow,

darkened;

nearly twice as

weakly granulated;

approximately 0,25

length of scutum;

Anteoninae

Fig.

346 -

Chela of

541

Anteon parvulum (R.C.L. Perkins) from Mapleton (Australia)

scutell um and metanotum smooth;

propodeum rugose,

ma i n I y

near

metanotum, with a transversal keel between dorsal and posterior surface; posterior surface with 2 longitudinal keels; median area rugose; forewing hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal proportions:

4,5:

1,5: 2:

part;

3,5:

bristle located on a proximal 346)

with basal

fore tarsal

prominence;

(Fig.

346)

(Queensland,

lectotype F!

with one

segment 5 of front part

(3,5:5,5),

apex with a group of 10 lamellae;

Cairns

TYPICAL MATERIAL:

segments in following

enlarged claw

part shorter than apical

nearly 25 lamellae; MALE: unknown LOCUS TYPICUS:

9;

tarsus

(Fig.

with two rows of

tibial

spurs 1,1,2.

Australia)

(designated by Olmi

1982) of

N. parvulus

in B; lectotype F! (designated by Olmi 1982) of N. pallidicornis in B. HOSTS: in Australia Euleimonios sp. (R.C.L. Perkins 1906c) for parvulus;

Deltocephalus sp. luteotactella. DISTRIBUTION: Bundaberg

Australian

(Queensland),

Queensland) Roseberry

(R.C.L.

,

UQ !

Perkins 1906c) region: B!

TW

AUSTRALIA:

Eurimbula Ck.

Mapleton,

(T asman i a),

for

BR

! Mt.

pallidicornis; N. Cairns

(Queensland),

(Via Round Hi II

Hotham

(Victoria),

!

10. Anteon orientale n. sp.

Head, OL!

TW!

B!

542

Mem.

Fig.

FEMALE:

fully

testaceous;

347 -

Amer.

En!.

no.

37,

1984

Anteon orientale n. sp.

Chela of

winged;

Ins!.,

length 3,18 mm;

antennae testaceous,

black;

with dorsal

(type)

mandibles and

legs

side of segments 4-10 brown;

antennae distally thickened; antennal segments in following proportions: 14: 6: 8: 6: 6: 6: 5: 6: 7: 7; head shiny, strongly punctate, without sculpture among punctures, with anterior half of frons reticulate rugose; frons with a

longitudinal

orbits directed complete;

towards ~

POL

8;

median

~

Ol

keel

and with 2

inner side of antennal 4;

strongly

punctate,

pronotal

tubercles reaching

OOl

~

8;

OPl

~

5;

shiny,

smooth,

punctate,

tegulae;

scutum shiny,

only partly distal

transversal part

6:

longer than

apical

located on a with

2:

bands;

(3,5; 2,5:

10);

4:

part

proximal

1 row of 9

carina

pronotum shiny,

smooth,

(6:21);

strongly

notaul ices incomplete, scutellum and metanotum

without scul;:>ture among punctures;

without dark than proximal

5;

propodeum

with a strong transversal keel between dorsal and posterior surface with 2 longitudinal keels; median

area mostly smooth and shining,

proportions:

~

Tl

keels around

occipital

with posterior surface shorter than scutum

punctate, without sculpture among punctures; reaching approximately 0,3 length of scutum; reticulate rugose, posterior surface;

lateral

sockets;

15;

(9:6);

fore tarsal

segments

segment 5 of front enlarged claw

prominence;

lamellae;

rugose;

fore-wing

part of radial

in following

tarsiusi

(Fig.

347)

segment 5 of front

apex with a

group of 4

hyaline,

vein much shorter with basal

part

with one bristle

tarsus

lamellae;

(Fig. tibial

347) spurs

1,1,2. MALES:

fully

testaceous;

winged;

samet i mes

length 1,56-3,75 mm; legs brown;

black;

mandibles and

legs

antennae brown or testaceous-darkened;

Anteoninae

Fig.

Male genitalia of Anteon orientale n.

348 -

(Queensland):

antennae not distally thickened; proportion: 7: punctate,

543

4:

5:

5:

4,5: 5:

antennal 5:

sp.

from Tambourine Mts.

left half removed.

5:

5:

segments 8;

in following

head shiny,

without sculpture among punctures;

occipital

and malar space granulated and reticulate rugose;

smooth, region,

occipital

strongly

temples

carina

complete; frons with a median keel and without lateral keels; POL ~ 6; OL ~ 4; OOL ~ 4; OPL ~ 2; TL ~ 2; scutum, scutellum and metanotum shiny,

smooth,

weakly punctate,

notaulices incomplete, of scutum; between

longitudinal

and posterior surface;

keels;

hyaline,

median area dull, part

membranous process,

process; tibial spurs 1, 1, 2. LOCUS TYP ICUS: Tambourine Mts. TYPICAL MATERIAL: TW;

in BR;

holotype F!

1 paratype M!

2 paratypes MM!

HOSTS:

with

approximately 0,25

length

a strong transversal

keel

posterior surface with 2 nearly as rugose as

without dark transversal

vein much shorter than proximal without proximal

M!

reaching

propodeum reticulate rugose,

dorsal

fore-wing

without sculpture among punctures;

very reduced,

(3:7);

bands;

distal

gonoforceps

without distal

lateral

areas;

part of radial (Fig.

348)

inner pointed

(Austral ia) and 2 paratypes MM!

in AD;

1 para type F!

in UQ;

in BM;

1 paratype

2 para type I,jf;i!

in

in OL.

unknown

DISTRIBUTION: Queensland), Round Hi II

Australian region:

OL!

Head,

Gordonva!e

AUSTRALIA:

(NE Queensland),

Queensland),

UQ!

Loxton

(32 mls. N of Perth, Western Australia), TW! Mt. Giluwe, T'.\'!

Tambourine Mts. SR!

(South Austral ia),

BM!

(SE

Eurimbula Ck.

NEW GUINEA:

AD!

(via Yanchep

Baiyer R.,

OL!

544

Mem. Amer. En!.

Fig. 349 -

Inst.,

no.

37,

1984

Chela of Anteon aculeatum n. sp.

(type)

Anteoninae

545

11. An rcon aculeatum n. sp.

FEMALE:

fully

black;

antennae distally thickened;

following proportions: with

length 1,87 mm;

winged;

legs testaceous;

mandibles,

antennae and

antenna I segments

in

8: 4: 5: 4: 6: 5: 4,5: 4,5: 4,5: 7; head shiny,

vertex and posterior half of frons smooth and

without sculpture;

anterior half of frons reticulate rugose; ocellar region, region behind ocelli and regions on sides of ocelli with irregular keels; frons with a median kell; occipital carina complete; POL = 4; OL = 3; OOL = 3; OPL 2; TL = 3; ;:>ronotum strong I y rugose, with posterior surface shorter than scutum (5:10); pronotal tubercles reaching tegulae; scutum, scutellum and metanotum shiny, punctures; scutum;

smooth,

notaul ices

propodeum

between dorsal

weakly

punctate,

incomplete,

reticulate rugose,

with a strong

and posterior surface;

longitudinal keels; transversal bands;

without sculpture among

I ittle visible near anterior margin of transversal

posterior surface with

keel

two

median area rugose; fore-wing hyaline, without dark distal part of radial vein much shorter than proximal

(2:7); fore tarsal segments in following proportions: 3,5: 2,5: 2: 5: 12; enlarged claw (Fig. 349) with one bristle located on a proximal prominence; segment 5 of front tarsus (Fig. 349) with 2 rows of nearly 23 lamellae; apex with a group of nearly 19 lamellae; tibial spurs 1,1,2.

part

MALE:

unknown

LOCUS TYP ICUS:

Mt.

TYPICAL MATERIAL: HOSTS:

Glorious

(Austral ia)

holotype F!

in TW.

unknown

DISTRIBUTION: TWI

Australian

region:

AUSTRALIA:

Mt.

Glorious

(Queensland),

12. Ilnteon corruptum n. sp.

Ft:MALE:

fully

with proximal testaceous; proportions: keel

with coxae,

antennae distally

15: 6:

8:

9:

granulated on

and with 2

6:

7:

internal

lateral

sockets;

occipital

TL = 5;

pronotum shiny,

region

antennae brown,

legs testaceous,

partly brown; rugose,

length 2,81-3,00 mm;

winged;

half brown;

wi th a strong

9;

fully

head shiny,

reticulate

=

6;

OL

=

reticulate rugose,

and without sculpture;

reticulate rugose, shiny,

POL

(5:18); shiny;

pronotal

fully

frons with a

median

towards antenna I 3;

OOL

=

8;

OPL

=

4,5;

except for a short posterior surface of

tubercles reaching

notaulices invisible;

without sculpture;

transversa I keel

tibiae in following

surface of areolae;

carina complete;

near scutum smooth

metanotum smooth,

7:

side of segment

antenna I segments

keels near orbits directed

pronotum shorter than scutum scutum fully

7:

mandibles testaceous,

ventral

clubs of femora and

thickened; 7:

black; with

tegulae;

scutellum and

propodeum reticulate rugose,

between dorsa I and posterior surface;

546

Mem.

Fig.

350 -

Chela of

posterior surface with 2 as

lateral

areas;

part of radial

tarsal

segments

front

tarsus

En!.

Ins!.,

hyal ine,

keels; without

37,

1984

median

area nearly as rugose

dark transversal

vein much shorter than proximal

in following proportions:

with basal (Fig.

no.

Anteon corruptum n. sp. (type)

longitudinal

fore-wing

distal

enlarged claw

Amer.

350)

6:

3:

3:

9:

part much shorter than apical with one bristle

part 22;

bands;

(4:12);

fore

segment 5 of

part

(8:14);

located on a proximal

Anteoninae

prominence;

segment 5 of front

tarsus

547

(Fig.

350)

lamellae extending continuously to the apex; MALE:

spurs 1,1,2.

unknown

LOCUS TYPICUS:

Murray Pass

TYPICAL MATERIAL: F!

with 2 rows of nearly 31

tibial

(New Guinea)

holotype F!

in B;

1 paratype F!

in OL;

1 paratype

in TW.

HOSTS: unknown DISTRIBUTION: Australian region: NEW GUINEA: Murray Pass (W slope, New Guinea), B! Mt. Kaindi (NE New Guinea), OL! L. Sirunki, TW!

SE

13. Anteon miles n. sp.

FEMALE:

fully

winged;

length 2,87-3,12 mm;

anterior margin of clypeus reddish;

black;

mandibles testaceous;

antennae testaceous or brown,

with

segment 1 testaceous; legs testaceous, with hind coxae partly brown; antennae distally thickened; antenna I segments in following proportions: 10: 5:

10:

9:

9:

9:

with a median keel, = 5;

OL = 3;

rugose,

8:

8:

8:

without

OOL = 6;

10;

head dull,

lateral

OPL = 5;

keels;

TL = 5;

fully

reticulate rugose;

occipital carina complete;

pronotum dull,

except for a short region near scutum which

surface of pronotum shorter than scutum reaching tegulae; complete,

(5:16);

scutum fully reticulate rugose,

is smooth;

dull;

POL

reticulate

pronotal

reaching approximately 0,75 length of scutum;

frons

posterior

tubercles

notaul ices almost scutellum with

strong areolae on central region, with weak areolae laterally; metanotum reticulate rugose; propodeum reticulate rugose, with a strong transversal keel

between dorsal

longitudinal transversal part 17;

(4:13);

keels;

and posterior surface; median area rugose;

bands;

distal

fore tarsal

segment 5 of front

(7:10);

enlarged claw

prominence;

part of radial

(Fig.

351)

Angoram

DISTRIBUTION: Guinea),

B!

Kainde,

3:

part

with one bristle located on a proximal (Fig.

351) tibial

with 2 rows of 23 spurs 1,1,2.

(New Guinea)

holotype F!

Australian region:

Mt.

2,5:

part shorter than apical

lamellae; apex with a group of 6 lamellae; MALE: unknown TYPICAL MATERIAL: HOSTS: unknown

without dark

vein much shorter than proximal

segments in following proportions: 7,5: tarsus with basal

segment 5 of front tarsus

LOCUS TYPICUS:

posterior surface with 2

fore-wing hyal ine,

in B;

1 paratype F!

NEW GUINEA:

in TW.

Angoram

TW!

14. Anteon destructor (R.C.L. Perkins)

Neochelogynus destructor R.C.L. Perkins 1905: 64.

(Sepik,

NE New

7:

548

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

\

Fig.

351

-

Chela of

Anteon miles n. sp. (type)

Neochelogynus cognatus R.C.L. Perkins 1905: 64 (n. syn.); locus typicus Bundaberg (Austra I i a) •

549

Anteoninae

Neochelogynus ignotus R.C.L. Perkins 1906a: 493 (n. syn.); locus typicus Sydney (Austra I i a) • Anteon cognatus (R.C.L. Perkins): Kieffer 1907b: 25. An teon destructor (R. C. L. Perk i ns): Kieffer 1907b: 25 Chelogynus ignotus (R.C.L. Perkins): Kieffer 1914b: 192. Chelogynus destructor (R.C.L. Perkins): Kieffer 1914b: 192. Chelogynus cognatus (R.C.L. Perkins): Kieffer 1914b: 193.

FEMALE:

fully

winged;

with teeth brown;

length 2,50-3,11

sometimes only

mm;

distal

black;

mandibles yellow,

half of mandibles yellow

(destructor); antennae brown with segment 1 testaceous (ignotus); (cognatus) or segments 1-3 testaceous (destructor); tegulae testaceous; legs testaceous, sometimes with coxae

occasionally segments 1-2 testaceous brown; 5:

5:

keel

antennal 6;

segments in following proportions:

head fully

and

reticulate rugose;

with 2 l.:lteral

sockets;

POL = 7;

complete;

frons

11: 5:

with a

4:

3:

4:

longitudinal

4:

5:

median

keels near orbits directed towards antenna I

OL = 5;

OOL = 7;

OPL = 5;

pronotum re-ticulate rugose,

short,

posterior surface shorter than scutum

TL = 4;

occipital

carina

much broader than

(9:17);

pronotal

long,

with

tubercles reaching

tegulae; scutum shiny, smooth, weakly punctate, without sculpture among punctures; notaul ices very short, visible only near anterior margin of scutum;

scutellum and metanotum shiny,

among punctures; between

dor5al

longitudinal hyaline,

and posterior surface;

keels;

without dark transversal proximal

proportions:

6:

2:

4:

enlarged claw

prominence;

part 8:

20;

(Fig.

bands;

(4:10):

lateral

distal

segment 5 of front

transversa I keel

fore tarsal

2

areas;

fore-wing

part of radial

segments

segment 2 of front

352)

without sculpture

wi th a

posterior surface with

median area as rugose as

shorter than hook;

punctate,

propodeum ret i cu I ate rugose,

vein much

in following

tarsus produced

into a

with one bristle located on a proximal tarsus

(Fig.

352)

with 2 rows of 18

lamellae; apex with a group of 13 lamellae; tibial spurs 1,1,2. MALE: fully winged; length 1,43-2,68 mm; black; mandibles partly black or with

teeth brown;

testaceous; brown;

antennae brown or testaceous-dark,

tegulae testaceous;

rarely fully brown;

legs testaceous,

head reticulate rugose

reticulate rugose that head of female); keel

and

sockets;

with 2 lateral POL = 6;

complete; punctures; scutum;

OOL = 6;

smooth,

notaulices incomplete,

propodeum reticulate rugose, area as rugose as transversa I bands; part

OPL = 2;

weakly

(3:8);

longitudinal

TL = 3;

punctate,

reaching 0,3

scutellum and metanotum shiny,

posterior surface;

(less strongly

with a

median

keels near orbits directed towards antenna I

OL = 3;

scutum shiny,

frons

with segment 1

with fore coxae partly

(rarely 0,5)

smooth,

with a transversal

occipital

areas;

fore-wing

length of

without sculpture; keel

between dorsal

posterior surface with 2 longitudinal lateral

carina

without sculpture among

hyal ine,

keels;

without dark

d i sta I part of rad i a I vei n much shorter than

gonoforceps without dorsal

proximal

and

median prox i ma I

membranous process,

550

Fig.

Mem.

352 -

Amer.

En!.

Ins!.,

Chela of Anteon destructor

no.

(R.C.L.

37,

1984

Perkins)

from Bundaberg

(Queensland)

without distal process

(Fig.

inner process 354)

LOCUS TYP ICUS:

(Fig.

Bundaberg

lectotype F!

(designated by Olmi

1982)

1982) of N.

or with an tibial

indistinct rounded

spurs 1,1,2.

(Austral ia)

TYPICAL MATERIAL: by Olmi

353)

extending medially;

of N.

and 2 paralectotypes

(1

F,

1 M)!

destructor in B; lectotype F!

cognatus in 3; lectotype F!

(designated

(designated by Olmi

Anteoninae

551

B

Fig. 353 - Male genitalia of Anteon destructor (R.C.l. Perkins) from Tulley Falls (Queensland) (A: left half removed); B: gonoforceps (laterally viewed) •

1982) of N.

ignotus in B.

HOSTS: in Australia Eurinoscopus sp. (R.C.l. Perkins 1906c)(destructor) and Idiocerus sp. or ftiacropsis sp. (R.C.l. Perkins 1906c) (cognatus). DISTRIBUTION: Australian region: AUSTRALIA: Bundaberg (Queensland), B! Gordonvale (NE Queensland), SR! Tulley Falls (N Queensland), B! Mt. Glorious (Queensland), TIV! Wi Ison 's Peak (via Teviot Gap, SE Queensland), Ol! UQ! longreach (Queensland), Dl! Sydney (New South Wales), B! Cabramatta (New South Wales), BM! Yuendumu (Northern Territory), TW! Ol! Mt. Baw Baw (E Victoria), UQ! Strahan (Tasmania), Ol! TW! Bronte Park (Tasmania), TW! NEW BRITAIN: Gaulim (Gazelle Pen.), B! NEW GU INEA: Wau (Morobe Distr., NE New Guinea), B! Ambunti (Sepik R., NE New Guinea), B! Mi Ine Bay (SE New Guinea), B! Swart Val. (Irian), B! Karimui (NE New Guinea), S! Nabire (S Geelvinck Bay, NW New Guinea), B! Malgi (Mt. Giluwe, SE t<ew Guinea), Ol! lake Habbema (3250-3300 m), LEI Baiyer R., Ol! TW! Mt. Kainde, Ol! TW! Nadzab, OL! TW! Busu R. (lae), TW! Gent R. (Jimmy Valley), TW! Tari Gap (Mt Hagen), TW! SOLOMON ISLANDS: Soaika Hi II (Hutuna, Rennell Island),

15. Anteon caledonianum n. sp

CO!

552

Mem.

Fig.

354 -

Amer.

FEMALE:

fully

winged;

Ins!.,

no.

37,

1984

Anteon destructor (R.C.L. Perkins): variety

Male genital ia of with

Ent.

weak

rounded process.

length 2,50 mm;

antennae with segments 1-3 testaceous;

black;

mandibles testaceous;

other segments missing

in the only

known female specimen; legs testaceous, with hind coxae proximally infuscate; antennae damaged in the only known female specimen, with only first

three segments;

first

three segments

in following proportions:

9: 4,5: 6; head shiny, fully reticulate rugose; frons with a median keel and with 2 lateral keels near orbits; occipital carina complete; POL = 5; OL = 3; on I y

OOL = 2,5;

TL = 4;

pronotum shiny,

near posterior marg in;

scutum

(5: 12);

puncta te, reaching

pronotal

reticulate rugose,

tubercles reaching

tegulae;

without scu I pture among punctures; approximately 0,3

smooth

posterior surface of pronotum shorter than

length of scutum;

scutum shiny,

notau Ii ceS

weakly

i ncomp I ete,

scutellum and metanotum

shiny, smooth, weakly punctate, without sculpture among punctures; propodeum reticulate rugose, with a strong transversal keel between dorsal

and posterior surface;

posterior surface with 2

longitudinal

keels;

median area as rugose as lateral areas; fore-wing hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part

(2:8);

fore tarsal

segment 5 of front (6,5: 1,5); proximal lamellae; MALE:

segments

tarsus

enlarged claw prominence;

fully

winged;

coxae proximally

(Fig.

355)

group of 2

length

infuscate;

tarsus

lamellae;

(Fig.

tibial black;

1 testaceous;

5:

2:

3:

longer than apical

with one bristle

1,25-2,00 mm;

with segment

proportions:

part much

segment 5 of front

apex with a

antennae brown,

in following

with basal

8;

part

located on a 355)

with

1 row of 9

spurs 1,1,2. mandibles testaceous;

legs testaceous,

antennae not distally

4:

thickened;

with hind antenna I

553

An teon i nae

355 - Chela of Anteon caledonianum n. sp.

Fig.

segments shiny,

in following proportions:

reticulate rugose,

7:

4:

4:

3:

4:

4: 4:

4:

(type)

4:

6;

head

with ocellar region and a median frontal

less reticulate rugose or without areolae,

punctate,

among punctures; frons with a median keel, without lateral keels; occipital carina complete; POL = 6; OL = 3; OOL = 3; OPL = 2; TL scutum,

scutellum and metanotum shiny,

sculpture among punctures;

smooth,

weakly punctate,

notaulices incomplete,

0,3 length of scutum; propodeum reticulate rugose,

region

without sculpture

= 2; without

reaching approximately with a strong

transversal keel between dorsal and posterior surface; posterior surface with 2 longitudinal keels; meciian area as rugose as lateral areas; forewing hyal ine,

without dark transversal

bands;

distal

part of radial

vein

(2:6); gonoforceps (Fig. 356) with a long and narrow dorsal process (Fig. 357), without distal inner pointed process; tibial spurs 1,1,2.

much shorter than proximal

LOCUS TYPICUS:

Col

TYPICAL MATERIAL:

part

de la Pirogue (New Caledonia) holotYiJe F!

and 2 paratypes MM!

in B;

1 paratype

M! in OL. HOSTS: unknown DISTRIBUTION: Col

Australian

des Roussettes,

OL!

region:

Plum,

B!

NEW CALEDm
Col

B!

16. Anteon typicum (R.C.L. Perkins)

de la Pirogue,

B!

554

Mem.

Fig.

Amer.

En!.

Ins!.,

no.

37,

1984

Male genitalia of Anteon caledonianum n. sp.: A: viewed; 3: ventrally viewed (left half removed).

356 -

laterally

Neochelogynus typicus R.C.l. Perkins 1905: 61. Anteon typicus (R.C.l. Perkins): Kieffer 1907b: 27. Anteon doddi Kieffer 1911b: 232 (n. syn.); locus typicus Townsville (Australia).

Liodryinus doddi (Kieffer): Kieffer 1914b: 57. Chelogynus typicus (R.C.L. Perkins): Kieffer 1914b: 189. Anteon doddi Kieffer: Richards 1953: 54.

FEMALE:

fully

winged;

with teeth brown; distal

length 3,75-4,87 mm;

antennae testaceous,

part of 7 brown;

testaceous;

legs

11:

10: 9:

10: 9:

mandibles yellow,

sometimes also segments 1-4 brown;

testaceous,

femora and tibiae brown; 8:

black;

with segments 8-10 and sometimes

8:

with coxae brown;

antennal

8:

11;

tegulae

sometimes also clubs of

segments in following proportions:

sometimes reticulate rugose only on frons, wi thout scu I pture among punctures;

fronta I

whereas vertex line comp lete;

is punctate and frons

without

lateral keels; occipital carina complete; POL = 9; Ol = 4; OOl = 5; = 6; Tl = 5; pronotum strongly punctate and rugose, hairy, with posterior surface shorter than scutum tegulae;

scutum shiny,

punctures; scutum

smooth,

(12:22);

pronotal

weakly punctate,

notaulices incomplete,

(in doddi)

22:

head dull, fully reticulate rugose;

OPl

tubercles reaching

without sculpture among

visible only near anterior margin of

or reaching approximately 0,5 length of scutum

(typicum); scutell um and metanotum sh i ny, smooth, weak I y punctate, without sculpture among punctures; transversal

keel

between

dorsal

propodeum reticulate rugose,

and posterior surface;

with a

posterior surface

Anteoninae

555

.. '.

Fig.

Male of Anteon caledonianum n.

357 -

gonoforceps

with 2 longitudinal

keels;

median area shiny,

only near anterior margin; beneath

the pterostigma;

proximal 3:

4: 9:

part 22;

proximal

(5:13);

(Fig.

358)

Bundaberg

TYPICAL MATERIAL:

smooth,

punctate,

vein

rugose

band

much shorter than

with one bristle located on tarsus

lamellae extending continuously

1,1,2. MALE: unknown lOCUS TYP ICUS:

process of

segments in following proportions:

segment 5 of front

prominence;

dorsal

with 1 dark transversal

part of radial

tarsal

enlarged claw

of nearly 55

fore-wing

distal

fore

sp.:

(laterally viewed).

(Fig.

358)

to the apex;

10:

a

with some rows tibial

spurs

(Austral ia)

holotype F!

of N.

typicus in B; holotype F! of A.

doddi in 8M. HOSTS:

unknown

DISTRIBUTION:

Australian

Eurimbula Ck.

(via Round Hill

region:

AUSTRALIA:

Head,

Bundaberg

Queensland),

(Queensland),

Bri,!

Tamborine Mt.

(Queensland),

(Queensland),

Ol!

TW!

UQ!

Canberra,

Clare

Ol!

UQ!

(Queensland),

B!

Townsville

Mt.

Glorious

(South Australia),

TW!

17. Anteon bougainvillei n. sp.

FEMALE:

fully

winged;

antennae black; thickened; 9:

8:

8:

8;

length 3,75 mm;

legs black,

partly

black;

testaceous;

mandibles testaceous; antennae distally

antenna I segments in following proportions: antenna I segment 10 missing

in

17:

7:

12:

10: 9:

the only kmown specimen;

head shiny, rugose; frons with numerous longitudinal keels and with a median keel; vertex and region between ocellar triangle and occipital

556

Mem.

Fig.

358 -

Amer.

En!.

Ins!.,

Chela of Anteon typicum

no.

(R.C.L.

37,

1984

Perkins)

from Townsville

(Queens I and)

carina strongly sculptured by obi ique keels and with some areolae; occipital

POL = 7;

carina complete;

pronotum hairy,

dull,

OL = 5;

scutum shiny,

among punctures; length of scutum;

smooth,

strongly punctate,

transversal

surface with 2 transversal bands,

keel

between

longitudinal

keels,

one on

keels,

TL = 8; with

without sculpture

notaulices incomplete, reaching approximately 0,25 scutellum and metanotum shiny, smooth, punctate,

without sculpture among punctures; strong

OPL = 10;

transversal

(11 :20); pronotal tubercles reaching

posterior surface shorter than scutum tegulae;

OOL = 7;

strongly sculptured by

as

keels;

lateral

the basal

propodeum

dorsal

reticulate rugose,

and posterior surface;

with a

posterior

median area strongly sculptured by

areas;

fore-wing

with 2 dark

transversal

cells and another beneath the pterostigma;

fore

8: 2,5: 3: 8: 21; segment 5 of front tarsus with basal part shorter than apical part (8: 13); enlarged claw (rig. 359) with one bristle located on a proximal prominence; segment 5 front tarsus (Fig. 359) with 2 rows of nearly 27 lamellae; apex with a group of 4 lamellae; tibial spurs 1,1,2. tarsal

segments

MALE:

unknown

in following proportions:

LOCUS TYPICUS: Kokure (Bougainville I., TYPICAL MATERIAL: holotype F! in B. HOSTS:

Solomon

Islands).

unknown

DISTRIBUTION: Prince Ra.,

Australian

Bougainvi lie

region: Island),

SOLOMON

ISLANDS:

B!

18. Anteon curradoi n. sp.

Kokure

(near Crown

Anteoninae

Fig.

FEMALE:

fully

359 -

Chela of

winged;

antennae brown, brown;

legs

testacous,

with

tarsi

antennal 10;

segments

head shiny,

longitudinal 6,5;

1 testaceous;

black;

with hind coxae proximally brown;

in following proportions: occipital

TL = 5;

mandibles testaceous;

sometimes antennae fully

and fore tibiae testaceous;

reticulate rugose,

keels;

OPL = 4;

Anteon bougainvillei n. sp. (type)

length 3,12-4,18 mm;

with segment

brown,

557

sometimes

antennae distally

13:

7:

11:

10:

10:

with frons sculptured by

carina complete;

pronotum rugose,

POL = 6;

9:

9:

strong

OL = 3;

transversely striate,

legs

thickened;

OOL

with

9:

9:

558

Mem. Amer.

Fig.

Inst.,

no.

37,

1984

360 - Chela of Anteon curradoi n. sp.

posterior region smooth;

(5:20); pronotal

Ent.

(type)

posterior surface of pronotum shorter than scutum

tubercles reaching tegulae;

without sculpture among punctures;

scutum strongly punctate,

notaulices

incomplete,

reaching only

0,3 length of scutum; scutellum and metanotum shiny,smooth, punctate,

without sculpture among punctures;

with a strong

transversal

keel

between

weakly

propodeum reticulate rugose,

dorsal

and posterior surface;

posterior surface with two longitudinal keels; median area rugose; forewing hyaline,without dark transversal bands; distal part of radial vein much shorter than proximal

part

(5:11)

fore tarsal

segments in following

16; enlarged claw (Fig. 360) with one bristle located on a proximal prominence; segment 5 of front tarsus (Fig. 360) with 2 rows of nearly 14 lamellae; apex with a group of 4 lamellae; tibial spurs 1,1,2. proportions:

MALE:

7: 2: 3: 6:

unknown

LOCUS TYPICUS:

Tari

TYPICAL MATE::IAL:

Gap

(New Guinea)

holotype F!

and

1 paratype F!

in TW;

1 paratype F!

in OL. HOSTS:

unknown

DISTRIBUTION: TW!

Australian

region:

NEW GUINEA:

Tari

Gap

(Mt.

Hagen),

OL!

NOTES:

the species is named

in honor of Mr.

Italo Currado.

19. Anteon firmum n. sp.

FEMALE:

fully

winged;

length 4,56 mm;

black;

legs reddish,

with mid

559

Anteoninae

Fig.

361

-

Chela of Anteon firmum

and hind coxae blackish;

n.

sp.

(type)

mandibles and antennae black;

antennae

distally thickened; antennal segments in following proportions: 18: 9: 16: 14: 11: 12: 13: 11: 10; last antennal segment missing in the only known

560

Mem.

specimen;

head dull,

and with 2 OL = 5,5;

lateral

Amer.

fully

Ent.

Ins!.,

reticulate rugose;

keels near orbits;

OOL = 8;

no.

OPL = 11;

37,

frons

occipital

TL = 8;

1984

with a median

carina complete;

pronotum dull,

transversely striate,

except for posterior surface which

posterior surface of pronotum shorter than

(14:20);

pronotal

tubercles reaching

reticulate rugose, scutellum dull,

with notaulices faintly

reticulate rugose;

posterior area

and

latera!

reticulate rugose, posterior surface; area smooth, beneath

shiny,

with a

4:

5:

13:

(5:14);

31;

shorter than

scutum dull,

scutum

hairy,

fully

visible near anterior margin;

metanotum reticulate rugose, smooth

few

bands,

the pterostigma; part

areas

is weakly

and

without

with

sculpture;

a

propodeum

with a strong transversal keel between dorsal and posterior surface with 2 longitudinal keels; median

with 2 dark transversal proximal

tegulae;

8·,

strongly

punctate and

hairy;

keel

POL

distal

fore

irregular keels near margins; one on

part of radial

tarsal

segments

segment 5 of front apical

part

the basal

tarsus

(13:18);

fore-wing

cells ar,d another

vein much shorter than

in following proportions:

(Fig.

361)

enlarged claw

with basal

(Fig.

361)

10:

part

with one

bristle located on a proximal prominence; segment 5 of front tarsus with 2 rows of 39 lamellae; apex with a group of 7 lamellae; tibial spurs 1,1,2. MALE:

unknown

LOCUS TYPICUS:

Bulolo R

TYPICAL MATERIAL: HOSTS:

(NE New Guinea)

holotype F!

in B.

unknown

DISTRIBUTION:

Australian

region:

NEW GUINEA:

(NE New Guinea),

Bulolo R

8!

20. Anteon leiosomum (R.C.L. Perkins)

Neochelogynus leiosomus R.C.L. Perkins 1905: 62. Anteol1 leiosomus (R.C.L. Perkins): Kieffer 1907b: 26. Chelogynus leiosomus (R.C.L. Perkins): I
FEMALE:

fully

teeth brown; brown;

winged;

length 3,11

antennae yellow,

legs yellow,

punctate,

weakly

with keels on

punctate;

transversal with 2

pronotum nearly

scutum smooth,

approximately 0,5 keel

mandibles reddish,

propodeum rugose, between keels;

near anterior margin;

body shiny,

sides and on anterior surface;

dorsal

hairy;

long,

posterior surface

incomplete,

reaching

scutellum and metanotum smooth, mainly near metanotum,

and posterior surface;

median

fore-wing

notaul ices

with

tegulae yellow-

twice as broad as

weakly punctate;

length of scutum;

longitudinal

black;

with mid and hind coxae brown;

head reticulate rugose; smooth;

mm;

with segments 7-10 brown;

area

smooth,

with 2.dark

with

a

posterior surface

punctate,

transversal

rugose only bands,

one on

561

Anteoninae

Fig.

the basal rad i a I

362 -

Chela of Anteon leiosomum

cells and another beneath

Perkins)(type)

the ;:oterostigma;

vei n much shorter than proxi ma I part;

nearly as long as segment 1; chela: ,\1ALE:

(R.C.L.

fig.

362;

distal

part of

segment 4 of fron t tibial

tarsus

spurs 1,1,2.

unknown

LOCUS TYP I CUS:

Bundaberg

TYPICAL MATERIAL: HOSTS:

(Austra I i a)

holotype F!

in B.

in Australia Eurinoscopus sp.

DISTRIBUTlm,:

Australian region:

(R.C.L.

AUSTRALIA:

Perkins 1906c) 8undaberg

(Queensland),

21. Anteon nitidurn (r<'C.L. Perkins) n. comb.

Neochelogynus nitidus R.C.L. Perkins 1905: 62. nec Anteon nitidus I
FEMALE:

fully

winged;

with teeth brown;

length 2,25-4,02 mm;

yellow-reddish with segments 7-10 brown; yellow-brown,

black;

mandibles yellow,

antennae brown with segments 1-2 testaceous, tegulae yellow-brown;

or legs

sometimes with coxae dark or mid and hind femora and

tibiae brown; body shiny, hairy; head reticulate rugose; pronotum punctate, with keels on sides and on anterior surface; pronotum nearly

8!

562

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

~. . • . . .

Fig.

363 -

Chela of

Anteon nitidum (R.C.L. Perkins) from Mt. Lindesay Hawy.

twice as broad as

long,

notaulices incomplete,

(New South Wales)

with posterior surface smooth,

reaching

approximately 0,25-0,50

scutell um and metanotum smooth, mostly near metanotum, posterior surface;

with a

weak I y

punctate;

transversal

posterior surface without

keel

weakly punctate; length of scutum;

propodeum rugose,

between

longitudinal

dorsal

and

keels or with

Anteoninae

Fig.

364 -

Male genitalia of

Anteon nitidum (R.C.L. Perkins) from Tathra

(New South Wales):

only

tracks of

longitudinal

keels;

fore-wing

distal

part of radial

segments

with

keels,

1 dark

563

left half removed.

faintly visible among the

transversal

vein much shorter than

in following proportions:

5:

irregular

band beneath the pterostigma;

2,5:

3:

proximal 6:

15;

part;

fore

tarsal

enlarged claw

(Fig.

363) with one bristle located on a proximal prominence; segment 5 of front tarsus (Fig. 363) with basal part shorter than apical part (5:10), with 2 rows of nearly 70 tibial

spurs 1,

MALE:

fully

lamellae extending continuously

winged;

length 1,81-2,81

antennae brown or testaceous dark; proximally broxn Guinea; rugose,

(in holotype);

antennae not distally

proportions: dull;

7:

to the apex;

2.

1,

5:

frons

6:

5:

5:

6:

mm;

black;

fully brown thickened; 5:

5:

mandibles testaceous;

legs testaceous,

5:

8;

with or without median

with hind coxae

in some para types from New

antenna I segments

in following

head fully or partly keel,

with or without

reticulate lateral

564

,\'lem.

keels;

head

sometimes

Amer.

not

carina complete;

punctate, reaching

rugose,

=

POL

OL

=

37,

1984

punctate and

5;

=

OOL

6;

without

=

OPL

3;

punctures;

notaul ices

length of scutum;

strong transversal

keel

posterior surface without

=

TL

scutellum and melanotum shiny,

without sculpture among

a

no.

rugose,

6;

scutum,

approximately 0,3

Wilh

surface;

Inst.,

reticulate

sculpture among punctures; occipital

Ent.

3;

weakly

incomplete,

propodeurn retuculate

between dorsal

longitudinal

keels;

and posterior fore-wing

hyal ine,

without dark transversal bands; distal part of radial vein much shorter than proximal part (3:8); gonoforceps (Fig. 364) without dorsal proximal membranous process and 1,1,2. LOCUS TY!) I CUS:

without distal

Bundaberg

TYPICAL MATERIAL:

inner pointed process;

tibial

spurs

(Austra I i a)

holotype F!

N. nitidus in B; holotype F! of i'';.

of

dimidiatus in B. HOSTS:

Eurinoscopus sp.

in Australia

(".C.L.

Perkins 1906c:

under name

N. dimidiatus). DISTRIBUTIO:--J:

Australian

region:

AUSTRALIA:

Bundaberg

(Queensland),

B!

Yangaborra (NE Queensland), OL! Mt. :--Jebo (Queensland), TW! 1I1t. Tamborine (Queensland), TV/! Tathra (I~ew South Wales), OL! TIV! Mt. Lindesay

Hawy.

(3 Km S Border Gate,

(Northern Territory), :--Jew Guinea),

B!

OL!

Jimmi

rA'!

New South Wales),

NEIV GU IIJEA:

Valley -

Baiyer R.,

r.lt.

Kaindi

BR!

Yuendumu

(Morobe Clistr.,

NE

TW!

22. Anteon bouceki n. sp.

FEMALE: with

fully

winged;

teeth brown;

clubs of femora,

length 3,25-3,68 mm;

antennae and hind coxae,

tegulae

black;

mandibles testaceous,

testaceous;

legs testaceous,

mid and hind tibiae brown;

with

sometimes clubs

of fore and mid femora testaceous; antennal segments in following proportions: 11: 6: 7: 7: 6: 5: 5: 5: 6: 8; head shiny, strongly punctate,

with numerous irregular keels,

longitudinal

median keel

and with 2

towards antenna I sockets; OOL

=

5,5;

OPL

=

7;

TL

occipital

=

5,5;

also among ocell i;

lateral

carina complete;

pronotum shiny,

POL = 3;

hairy,

(20:18);

pronotal

scutellum and metanotum shiny, sculpture among punctures;

tubrcles reaching

smooth,

notaul ices

posterior surface tegulae;

weakly punctate,

incomplete,

OL = 3;

with anterior half

rugose and posterior half smooth and without sculpture; longer than scutum

frons with a

keels near orbits directed

scutum,

without

reaching

approximately

0,5 length of scutum; propodeum reticulate rugose, with a transversal keel between dorsal and posterior surface; dorsal surface with areolae much wider than areolae of posterior surface; longitudinal basal

keels;

fore-wing

cells and another beneath

the pterostigma;

vein much shorter than proximal following proportions:

8:

3:

posterior surface without

with 2 dark transversal

5:

part

19:

28;

(4: 10);

fore

bands,

distal

one on

the

part of radial

tarsal

segment 2 of front

segments

in

tarsus produced

565

Anteoninae

Fig.

into a

hook;

proximal

365 -

enlarged claw

prominence;

much shorter than

apical

Kinglake

TYPICAL M;\TERIAL: r!

HOSTS:

365)

with one bristle tarsus

(Fig.

located on a

365)

with 2 rows of nearly 31

part,

lamellae;

tibial

with basal lamellae;

part

apex

spurs 1,1,2.

(Australia)

holotype F!

1 paratype F!

in QU;

(Fig.

segment 5 of front

group of nearly 5

with a

MALE: unknown LOCUS TYPICUS:

Anteon bouceki n. sp. (type)

Chela of

1 paratype F in BM;

in CB;

paratype

in OL.

unknown

DISn,IBUTION:

Australian

region:

(Melbourne,

NOTES:

honor of Mr.

the species

is named

in

Mt.

AUSn,ALIA:

Oueensland), aU! OL! Kinglake (I'ew South Wales), B,\i!

Glorious

Victoria), Zdenek

CB!

(SC: Cabramatta

Boucek.

23. Anteon mundum n. sp.

rE!,'!ALC::

fully

proximal

third black;

winged;

length 3,81

slightly distally

thickened;

16:

8:

5:

12:

keels, OL

~

8:

8:

8:

mm;

antenna I segment 8:

9;

smooth, scutum

DOL

~

5;

OPL

~

weakly ;:>unctate, (10:20);

and rnetanotum

pronotal shiny,

5;

TL

head shiny, ~

3;

mandibles testaceous,

1 testaceous beneath;

antenna I segments

6:

without sculpture among keels; 5;

:,Iack;

in following

without sculpture arT'ong

smooth,

POL

~

5;

with anterior surface

punctures,

tegulae;

weakly punctate,

longitudinal

carina complete;

pronotum shiny,

tubercles reaching

pro;:>ortions:

with numerous

occipital

with

antennae

scutum,

shorter than scutellum

without sculpture among

566

Mem.

Amer.

En!.

Inst.,

no.

37,

\i

/lI

I

I l /

\

\I,

I

\

1

\

\ \

1

\

\

\

\

Fig.

punctures; scutum;

366 -

notaulices incomplete,

bands;

strong

part of radial

fore-wing

sp.

(type)

approximately 0,5

transversal

except for a central

is without sculpture; distal

reaching

posterior surface without

transversely striate, which

Chela of Anteon mundum n.

propodeum with a

posterior surface;

/

i /

,

I

1984

keel

length of

between dorsal

longitudinal

keels,

and

strongly

smooth area near dorsal

hyal ine,

without dark

surface

transversal

vein much shorter than proximal

part

567

Anteoninae

(3: 12);

fore tarsal

segment 5 of front enlarged claw prominence; 55

segments

in following proportions:

tarsus with basal

(Fig.

366)

with one bristle

segment 5 of front

tarsus

lamellae extending continuously

MALE:

part

6:

3:

6:

longer than apical

Fig.

Kassam

367 -

(12:9);

located on a proximal

(Fig.

366)

to the apex;

with some rows of nearly tibial

spurs

unknown

LOCUS TYPICUS:

12: 31; part

(NE New Guinea)

Chela of Anteon bellator n.

sp.

(type)

1,1,2.

568

Mem.

TYPICAL MATERIAL: HOSTS:

Amer.

En!.

holotype F!

Ins!.,

no.

37,

1984

in B.

unknown

DISTRIBUTION: Kainantu,

Australian

region:

NE New Guinea),

NEW GUINEA:

Kassam

(48 Km E of

B!

24. Anteon bellator n. sp.

FEMALE: fully

fully

thickened; 8:

8:

7:

OL = 3;

striate,

10;

length 3 mm;

abdomen

antennal

8:

longitudinal 5;

winged;

testaceous;

mandibles,

petiole black;

antennae and

without

OOL = 6;

fully retuculate rugose;

lateral

OPL = 7;

keels;

TL = 4;

occipital

(8:17);

13: 8:

pronotal

10:

10:

10:

frons with a median

carina complete;

pronotum shiny,

except for a smooth area near scutum;

pronotum shorter than scutum

legs

antennae not distally

segments in following proportions:

head shiny,

keel,

black;

testaceous;

POL

transversely

posterior surface of tubercles reaching

tegulae;

scutum shiny, smooth, weakly punctate, without sculpture among punctures or very weakly granulated; notaulices incomplete, reaching only approximately 0,25 without sculpture; keel

between dorsal

longitudinal distal

length of scutum;

scutellum and metanotum shiny,

propodeum reticulate rugose,

keels;

and posterior surface; fore-wing hyaline,

part of radial

vein

with a strong

transversal

posterior surface without

without

dark transversal

much shorter than proximal

tarsal segments in following proportions: 10: 4: 4: 7: front tarsus with basa! part shorter than apical part

part

bands;

(5:10);

fore

17; segment 5 of (7: 10); enlarged

claw (Fig. 367) with one bristle located on a proximal prominence; segment 5 of front tarsus (Fig. 367) with 2 rows of nearly 42 lamellae extending continuously MALE: unknown LOCUS TYPICUS:

U.

Watut

TYPICAL MATERIAL: HOSTS:

to the apex;

tibial

spurs 1,1,2.

(NE New Guinea)

holotype F!

in B.

unknown

DISTRIBUTION: Guinea),

Australian

region:

NEW GUlrjEA:

U.

Watut SW

(NE New

B!

25. Anteon nigricorne (R.C.L. Perkins)

Neochelogynus nigricornis R.C.L. Perkins 19Q5: 63. Anteon nigricornis (R.C.L. Perkins): Kieffer 1907b: 27 Chelogynus nigricornis (R.C.L. Perkins): Kieffer 1914b: 191.

FEMALE:

fully

winged;

wi th teeth brown;

length 2,62-2,93 mm;

antennae brown;

black;

mandibles testaceous,

tegu I ae testaceous;

legs redd i sh-

569

Anteoninae

368 -

Fig.

Anteon nigricorne (i'.C.L. Perkins) from Sandhill

Chela of

(Austral ia)

testaceous, with coxae brown; occasionally brown, with tarsi light; antennal segments in following proportions: 11: 5: 5; 5: 5: 5: 5: 5: 6;

head fully

reticulate rugose;

frons with a

median

longitudinal

5:

keel

and with 2 lateral keels near orbits directed towards antenna I sockets; occipital carina complete; POL ~ 7; OL ~ 3; OOL ~ 4; OPL ~ 4; TL ~ 4; pronotum reticulate rugose,

(8:12);

scutum

pronotal

metanotum smooth,

hairy,

shiny,

weakly

punctures;

notaulices incomplete,

of scutum;

propodeum

dorsal

reticulate

and posterior surface;

fore-wing

hyaline,

with posterior surface shorter than

tubercles reaching

without

punctate,

tegulae;

reaching only rugose,

scutum,

with

a

approximately 0,25 transversal

posterior' surface without

dark

scutellum and

without sculpture among

transversal

bands;

keel

length

between

longitudinal

distal

keels;

part of radial

(3:8); fore tarsal segments in 12; enlarged claw (Fig. 368) with one bristle located on a proximal prominence; segment 5 of front tarsus (Fig. 368) with 1 row of 21 lamellae; apex with a group of 7 lamellae; tibial spurs 1,1,2. vein much shorter than proximal

following proportions:

MALE: unknown LOCUS TYPICUS:

Bundaberg

TYPICAL MATERIAL: Olmi

1982)

HOSTS:

(Australia)

lectotype F!

and 3 paralectotypes FF!

(designated by

in B.

in Austral ia

DISTRIBUTION: Sandhi II,

part

8: 2: 2: 5:

B!

Eutettix sp. (R.C.L. Perkins 1906c).

Australian

region:

AUSTRALIA:

Bundaberg

(Queensland),

B!

OL!

26. Anteon fljianum n. sp.

FElj,ALE:

fully

tegulae and

winged;

length 1,25-1,56 mm;

legs yellow;

antenna I segments

black;

mandibles,

antennae,

in following proportions:

7:

570

Mem.

Fig.

Amer.

Ent.

Inst.,

no.

369 - Chela of Anteon fijianum n. sp.

37,

1984

from Lautoka

3,5: 2: 2: 2: 2: 2: 2: 2,5: 4;

antennae distally thickened;

granulated,

area

except for a

is without sculpture; carina complete; shiny,

( 3,5:7);

with

a

OL = 3;

pronotal

without sculpture,

incomplete,

reaching

metanotum shiny, wi th a

frons

POL = 5;

with numerous transversal

than scutum smooth,

small

smooth,

longitudinal OOL = 5; keels,

head dull,

median

OPL = 4;

keel;

occipital

TL = 5;

pronotum

with posterior surface shorter

tubercles reaching

with sides very

tegulae;

scutum shiny,

weakly granulated;

length of scutum;

without sculpture;

between

Islands)

in front of the anterior ocellus which

approximately 0,3

transversa I keel

(Fiji

notaulices

scutellum and

propodeum reticulate rugose,

dorsa I and posterior surface;

posteri or

surface wi thout long i tud i na I keel s; areol ae of dorsa I surface wider than those of posterior surface; fore-wing hyaline, without dark transversal bands;

distal

part of radial

vein much shorter than

proximal

part

(1,5:4); fore tarsal segments in following proportions: 4: 1: 2,5: 2,5: 8; enlarged claw (Fig. 369) with one bristle located on a proximal prominence; segment 5 of front tarsus (Fig. 369) with 1 row of 6 lamellae; apex with a group of 3 lamellae; tibial spurs 1,1,2. MALE:

unknown

LOCUS TYPICUS:

Lautoka

TYPICAL MATERIAL: F!

Islands) and 3 paratypes FF!

in 8M;

1 paratype

in OL.

HOSTS:

in Fiji

DISTRIBUTION: BM!

(Fiji

holotype F!

AUSTRAL IA:

Islands

Nesosteles sanguinescens Kirkaldy.

Australian Narrabri

region:

FIJI

ISLANDS:

(New South Wales),

Lautoka

8M!

27. Anteon dubium Fouts

(Viti

Levu),

DL!

Anteoninae

571

Anteon dubius Fouts 1935: 157.

FEMALE:

fully

winged;

with teeth brown; with coxae brown; granulated; broad

length 1,70 mm;

body shiny,

scutum granulated;

approximately 0,5 without

longitudinal

bands;

distal

very short,

head

twice as long as

reaching

scutellum and metanotum smooth;

fore-wing

part of radial

segment 4 of front

more than

mainly near scutellum;

keels;

legs yellow,

mainly on head;

notaulices incomplete,

length of scutum;

propodeum strongly rugose,

mandibles yellow,

tegulae yellow;

with sparse hairs,

pronotum granulated,

(3:1);

brown-black;

antennae yellow-brown;

hyal ine,

posterior surface

without dark transversal

vein much shorter than

proximal

part;

tarsus slightly shorter than segment 1 (17:22);

tibial

spurs 1,1,2. MAL:::

unknown

LOCUS TYPICUS:

Teivipakeka

TYPICAL MATERIAL:

(Marquesas

holotype F!

HOSTS: unknown DISTRIBUTION: Australian region: (Nukuhiva),

Islands)

in B. MARQUESAS

ISLANDS: Teivipakeka

B!

28. Anteon coriaceum (R.C.L. Perkins)

Neochelogynus coriaceus R.C.L. Perkins 1905: 65. Anteon coriaceus (R. C. L. Perk i ns): Kieffer 1907b: 25. Anteon apterus Dodd 1913: 181 (n. syn.); locus typicus Nelson (Austral ia)

Chelogynus coriaceus (R.C.L. Perkins): Kieffer 1914b: 192.

FEMALE: with and

fully

winged;

teeth brown; legs yellow;

length 2,01-2,12 mm;

antennae brown or yellow, legs sometimes brown;

without median and

lateral

keels;

body

black;

mandibles yellow,

weakly darkened; dull;

pronotum granulated,

surface shorter than

scutum

incomplete,

anterior third or fourth of scutum;

reaching

(6: 11 );

tegulae

head granulated;

scutum granu I ated;

frons

with posterior notau Ii ces scutellum and

metanotum sh i ny and smooth; propodeum rugose, ma i n I y near metanotum, with a transversal keel between dorsal and posterior surface; posterior surface fully

rugose,

without

longitudinal

keels;

fore-wing

hyal ine,

without dark transversal bands; distal part of radial vrin much shorter than proximal part; fore tarsal segments in following proportions: 6: 1,5: 2:

4:

10; enlarged claw

(Fig.

prominence;

segment 5 of front

than

part

apical

(4,5:5,5),

group of 6 lamellae;

tibial

370)

with one bristle located on

tarsus

(Fig.

370)

with basal

with one row of 12 lamellae; spurs 1,1,2.

a proximal

part shorter

apex with a

572

Fig.

Mem.

370 -

Amer.

En!.

Chela of ;jnteon coriaceum Creek

MALE:

Ins!.,

unknown

no.

(FLC.L.

(Queensland)

37,

1984

Perkins)

from Marlborough

Anteoninae

LOCUS TYPICUS:

Cairns

TYPICAL MATERIAL:

(Queensland,

lectotype F!

573

Australia) 1982) of N.

(designated by Olmi

coriaceus in S; hoiotype F! of A. apterus in AD. HOSTS: in Australia Euleimonios sp. (R.C.L. Perkins 1906c). DISTRIBUTION: Marlborough Cent.

Australian region:

(Queensland),

Queensland),

OLl

SR!

SR!

AUSTRALIA:

Cairns

Marlborough Creek

Nelson

(Cairns,

(Queensland),

S!

(2 Km N Marlborough,

N Queensland),

AD!

NOTES: A. apterus Dodd was described as wingless. I bel ieve that the single female specimen is damaged and the wings have been broken off.

29. Anteon sucklingi n. sp.

FEMALE:

unknown

MALE: fully winged; length 1,68-1,87 mm; black; mandibles testaceous; antennae brown with segments 1 or 1-2 testaceous; legs testaceous with hind coxae proximally brown; with all

thickened; 7:

7:

7:

antennal

9;

2;

=

OOL

5;

posterior half, incomplete,

OPL

lateral

=

2;

371

-

TL

keels;

=

2;

10: 5:

occipital

carina complete;

scutum shiny,smooth,

8:

gonoforceps

7:

7:

7:

frons

POL = 4;

OL

notaulices

length of scutum;

scutum without

weakly granulated in specimen

Male genital ia of Anteon sucklingi n. S:

also

weakly punctate on

strongly punctate near anterior margin;

reaching approximately 0,65

removed);

Wilhelm)

antennae not distally

with weak areolae or irregular keels;

sculpture among punctures in holotype;

Fig.

(specimen from Mt.

segments in following proportions:

head shiny,

without median and

=

sometimes

coxae and clubs of femora partly brown;

sp.

(type) (A:

(laterally viewed).

right

half

574

Mem.

from Mt.

Wilhelm;

sculpture; keel

Amer.

between

much shorter than proximal

with part

TYPICAL MATERIAL:

Fig.

smooth and

without

longitudinal

bands;

distal

gonoforceps

with an

keels;

fore-

part of radial

(Fig.

371)

inner subdistal

vein

without pointed

(New Guinea)

holotype M!

in B;

1 para type M!

in OL.

unknown

DISTRIBUTION: Wi Ihelm

(3:9);

1984

posterior surface with

two distinct

membranous process,

process; tibial spurs 1,1,2. LOCUS TYPICUS: Mt. Suckling HOSTS:

37,

without a distinct transversal

and posterior surface;

without dark transversal

proximal

no.

scutellum and metanotum shiny,

dorsal

hyal ine,

dorsal

Inst.,

propodeum reticulate rugose,

irregular areolae and keels, wing

Ent.

Australian

(NE New Guinea),

372 -

region:

NEW GUINEA:

Mt.

Suckling,

B!

Mt.

OL!

Male genitalia of

Anteon bismarckense n. half removed.

sp.

(type):

right

Anteoninae

575

30. Anteon bismarckense n. sp.

FEMALE: MALE:

unknown

fully

testaceous;

winged;

length 2,80 mm;

antennae brown,

propodeum black;

head black,

with mandibles

wi th segments 1-3 testaceous;

abdomen brown;

legs testaceous;

thorax and

antennae ha i ry;

antennal segments in following proportions: 13: 7: 10: 7: 8: 9: 9: 8: 9: 12; head shiny, fully reticulate rugose; frons with a complete median keel; = 4;

occipital

carina complete;

scutum smooth,

broad,

=

POL

=

8; OL

very strongly punctate;

similar to areolae;

5; OOL

=

7; OPL

=

6; TL

punctures of scutum very

notaulices incomplete approximately reaching

0,5 length of scutum; scutellum and metanotum strongly puctate; propodeum reticulate rugose, with 2 longitudinal keels on posterior surface;

median area of posterior surface rugose;

without dark proximal

transversal

part

(3:15);

bands;

distal

gonoforceps

fore-wing

part of radial

(Fig.

372)

with a

hyal ine,

vein shorter than

dorsal

process not

curved at apex; tibial spurs 1,1,2. LOCUS TYPICUS: Yalom (Bismarck Islands) TYPICAL MATERIAL:

holotype M!

HOSTS: unknown DISTRIBUTION: Australian

CO!

in CO;

region:

1 paratype M!

BISMARCK

ISLANDS:

in OL. Yalom

(New Britain),

OL!

31. An teon gioriosum n. sp.

FEMALE: MALE:

unknown

fully

winged;

length

1,25-2,00 mm;

mandibles testaceous,

with

proximal half black; antennae brown; legs brown, with tarsi light; antennae not distally thickened; antennal segments in following proportions:

6:

4:

3:

4:

4:

4:

smooth;

occipital

or with

incomplete median keel

3; OOL smooth,

4:

4:

carina complete;

4:

6;

head dull,

frons without

and

without

fully granulated,

longitudinal

lateral

keels;

median keel

POL = 5;

OL =

=

4; OPL = 2; TL = 3; scutum, scutellum and metanotum shiny, weakly punctate, without sculpture among punctures; notaulices

incomplete,

reaching

reticulate rugose, posterior surface; area shiny, hyaline,

approximately 0,3

slightly

rugose,

nearly as rugose as

without dark transversal

shorter than

length of scutum;

proximal

part

(2:6);

bands;

lateral

areas;

distal part of radial

gonoforceps

(Fig.

proximal

membranous process and without distal

process;

tibial

LOCUS TYPICUS:

propodeum

with a strong transversal keel between dorsal and posterior surface with 2 longitudinal keels; median

373)

fore-wing vein much

without dorsal

inner pointed or rounded

spurs 1,1,2. Mt.

TYPICAL MATERIAL:

Glorious

(Australia)

holotype ~'.!

in 8;

1 paratype M!

in 81-::;

1 paratype

576

Mem.

Amer.

Ent.

Ins!.,

no.

37,

1984

B Fig.

Male genitalia of Anteon gloriosurn n.

373 -

removed);

M!

in TV/;

HOSTS:

B:

paratype M!

gonoforceps

sp.

(type)(A:

left half

(laterally viewed).

in OL.

unknown

DISTRIBUTION:

Australian region:

SE Queensland), Queensland),

8!

BR!

AUSTRALIA:

2nd Palen Ck.

Mt.

Nebo

(SE

Mt.

Glorious

(Rain Forest,

Crossing from Rathdowney(SE

Queensland),

OL!

Strahan

(Tasmania),

TW!

32. Anteon vulsum n. sp.

FEMALE: unknown t.\ALE: fully winged; with

teeth brown;

testaceous,

with coxae and fore and

antennae not proportions: rugose;

distally 11:

frons

OL

=

rugose,

4;

6:

thickened;

7:

6:

with a

orbits directed 6;

length 2,06-2,25 mm;

antennae testaceous,

6:

antenna I segments 5:

6:

6:

10;

keel

towards antenna I sockets;

OOL

=

5,5;

OPL

approximately

=

mandibles testaceous,

4,5;

visj~le

legs

hind clubs of femora partly brown;

track of median

with areolae less

incomplete,

5:

black;

with segments 4-10 darkened;

=

TL

in following

head shiny,

and with 2 occipital

3,5;

reticulate keels near

carina complete;

scutum shiny,

near posterior margin;

reaching 0,5

fully

lateral

POL

reticulate

notaulices

length of scutum;

scutellum and

metanotum shiny, smooth, without sculpture; propodeum reticulate rugose, with a strong transversal keel between dorsal and posterior surface; posterior surface with 2 areolae smaller than dark

transversal

proximal

part

bands;

(2:9);

longitudinal

areolae of distal

keels;

lateral

median

areas;

part of radial

gonoforceps

(Fig.

374)

area rugose,

fore-wing

hyaline,

with without

vein much shorter than

without dorsal

proximal

577

Anteoninae

A

Fig.

Male genitalia of Anteon

374 -

B:

removed);

membranous process and

without

Mt. Suck ling

LOCUS TYP I CUS:

TYPICAL MATERIAL:

vulsum n. sp.

gonoforceps

distal

(I a tera II y

(type)(A:

right

half

viewed).

inner process;

tibial

spurs 1,1,2.

(Nevi Gu i nea)

holotype M!

and

1 paratype M!

in B;

1 paratype Iv'i!

in OL. HOSTS:

unknown

DISTRIBUTIO;":

Australian

(NE ~,ew Guinec.),

OL!

~,EW

region:

rill.

I
GUINEA:

11''.1. Suckling, B! Karimui

U"E New Guinea),

3!

33. Anteon 1uctuosum n. sp.

F::'MALE: IvIALE:

unknown

fully

testaceous;

winged;

length 2,37-2,75 !;om;

legs testaceous,

antennae not distally proportions: rugose;

frons

19:

6:

with

6:

thickened; 5:

5:

black;

mandibles and antennae

with coxae and clubs of femora partly brown; 5:

antennal 5:

5:

5:

segments 9,5;

a complete median keel

near orbits directed towards antennal

and

sockets;

in following

head fully with

two

occipital

reticulate lateral

keels

carina complete;

578

Mem.

Fig.

POL

Amer.

En!.

Ins!.,

no.

37,

Male genitalia of Anteon luctuosum n. removed.

375 -

1984

sp.

(type):

right half

= 6,5; OL = 4,5; OOL = 5; OPL = 4; TL = 3; scutum, scutellum and

metanotum shiny, punctures;

smooth,

weakly punctate,

notaulices incomplete,

scutum;

propodeum reticulate rugose,

between

dorsal

and posterior surface;

longitudinal

keels;

transversal

bands;

part tibial

(3,5:9);

with a strong

gonoforceps

part of radial (Fig.

length of

transversal

posterior surface with

median area smooth; distal

without sculpture among

reaching approximately 0,5

fore-wing hyal ine,

keel

two without dark

vein much shorter than proximal

375) without distal inner pointed process;

spurs 1,1,2.

LOCUS TYP ICUS:

Baiyer R.

TYPICAL MATERIAL: MM! in OL.

(New Guinea)

holotype M!

HOSTS: unknown DISTRIBUTION: Australian

region:

and 6 paratypes MM!

NEW GUINEA:

in TW;

Baiyer R.,

34. Anteon hageni n. sp.

TWI

3 paratypes

OLI

Anteoninae

Fig.

376 -

Male genital ia of

579

Anteon hageni n. sp. (type): right half removed.

FErviALE: MALE:

unknown

fully

winged;

antennae brown,

length 2,00-2,50 mm;

with

segmePits

coxae proximally brown;

1-2

black;

testaceous;

mandibles testaceous;

legs

testaceous,

antennae not distally thickened;

with

hind

antennal

segments in following proportions: 11: 5: 8: 7: 7: 7: 6: 7: 7: 9; head shiny, reticulate rugose and with a few or numerous longitudinal keels on frons;

frons with a median keel

and 2 lateral

directed towards antenna I sockets;

occipital

=

scutum,

3;

DOL

shiny,

=

6;

smooth,

OPL

=

3,5;

TL

=

4;

weakly punctate,

notaul ices incomplete,

smooth,

POL = 7;

OL

scutellum and metanotum

without sculpture among punctures;

reaching approximately 0,25 length of scutum;

propodeum with a strong transversa I keel surface;

keels near orbits

carina complete;

between dorsa I and posterior

posterior surface with 2 longitudinal without sculpture,

keels;

median area shiny,

except for a few short keels near margins;

fore-wing hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part (3:9); gonoforceps (Fig. 376) without dorsal

proximal

process;

spurs 1,1,2.

tibial

LOCUS TYP I CUS:

M t.

TYPICAL MATERIAL: MM!

in TW;

membranous process and without distal

Hagen

(New Gu i nea)

holotype M!

4 paratypes MM!

HOSTS: unknown DISTRIBUTION: Australian

inner

and 1 paratype M!

in Bj

5 paratypes

in OL.

region:

AUSTRALIA:

Mt.

Glorious

(Queensland),

580

Mem.

Fig.

377 -

Amer.

En!.

Ins!.,

no.

37,

II,ale genitalia of Allteon haustum n.

1984

sp.

(type):

left half

removed.

OL!

TV/!

NEW GUIf'JEA:

Mt.

Distr.,

NE I'Jew Guinea),

Valley,

TW!

Hagen

OLi

[}!

(NE

New Guinea),

Bulolo,

TW!

B!

Wau

8aiyer R.,

(Morobe

OLi

TW!

Jimmy

35. Anteon haustum n. sp.

FEr"ALE: unknown MALE: fully winged; length 1,37 mm; reddish; mandibles, antennae and legs fully testaceous; antennae not distally thickened; antenna I segments in following proportions: 8: 3,5: 4: 4: 4: 5: 4: 5: 4: 6,5; head shiny, fully reticulate rugose; frons with a median keel, without lateral keels; occipital scutum,

carina complete;

"'OL ~ 4;

OL ~ 2;

scutellum and metanotum shiny,

without sculpture among punctures;

notaul ices

approximately 0,3

length of scutum;

strong

keel

transversal

surface with 2

keels;

very

OPL ~ 2;

TL ~ 2;

weakly punctate,

incomplete,

reaching

propodeum reticulate rugose,

between dorsal

longitudinal

OOL ~ 3,5;

smooth,

and posterior surface;

with

median area slightly rugose,

nearly

rugose as lateral areas; fore-wing hyal ine, without dark transversal bands; distal part of radial vein much shorter than proximal part (2,5:7);

gonoforceps

process and

LOCUS TYP ICUS:

Mt.

TYPICAL HATERIAL: HOSTS:

(Fig.

without distal

unknown

377)

without dorsal

proximal

inner pointed process;

Glorious

(Austral ia)

holotype M!

in OL.

tibial

a

posterior

membranous spurs 1,1,2.

as

Anteoninae

Fig.

378 -

581

Anteon corallinum n. sp. (type):

Male genital ia of

left half

removed.

DISTRIBUTION: Forest,

Australian

SE Queensland),

region:

AUSTRALIA:

Mt.

Glorious

(Sclerophyll

OL!

36. Anteon corallinum n. sp.

FEMALe:; MALE:

unknown

fully

testaceous; testaceous; segments

winged;

length 1,56-2,62 mm;

black;

mandibles and clypeus

antennae fully testaceous or brown with segments 1-2 legs testaceous; antennae not distally thickened; antennal

in following proportions:

8: 4:

5:

5:

5:

5: 5: 5:

5: 7;

head

shiny, smooth, strongly punctate, without sculpture among punctures; occipital carina complete; frons without median and lateral keels; POL

4; OL = 2; OOL = 4; OPL = 1,5; TL = 2; scutum, scutellum and metanotum shiny,

smooth,

notaul ices

weakly punctate,

incomplete,

propodeum with

a

reaching

transversal

posterior surface with 2 few

irregular keels,

without

dark

than proximal

keel

as rugose as (2:5);

bands;

keels;

lateral

distal

gonoforceps

membranous process and without

length of scutum;

between dorsal

longitudinal

transversal part

without sculpture among punctures; approximately 0,5

distal

and posterior surface;

median

areas;

area shiny,

fore-wing

part of radial (Fig.

378)

with a

hyal ine,

vein much shorter

without dorsal

inner pointed process;

proxirnal

tibial

spurs

1,1,2. LOCUS TYP ICUS:

t;:1.

TYPICAL MATERIAL:

I\i!

in OL.

HOSTS:

unknown

Sinewit

(New Britain)

holotype M!

in [3;

1 paratype M!

in UQ;

1 paratype

582

Mem.

DISTRIBUTION: Pen.),

B!

Australian

AUSTRALIA:

(via Darwin,

Amer.

::Ont.

~EW

region:

I
I nst.,

no.

37,

BRITAIN:

1984

,V,t.

Sinewit

(Northern Territory),

Northern Territory),

Ol!

(Gazelle

11,t.

Bundey ?d.

UQ!

37. Anteon tasmanianurn n. sp.

FErvlALE: MALE:

unknown

fully

segment fully

winged;

length 2,75-3,37 mm;

1 testaceous;

black,

legs

partly brown;

antennae not distally

strongly punctate,

12:

8:

10:

10:

thickened; 10:

ventral

side of antenna I

hind coxae black,

10:

mid and

hind

hind femora tibiae

antenna I segments

10:

10:

10:

13;

in

head shiny,

almost reticulate rugose on anterior half of frons;

frons without median and Ol = 4;

black;

with

fore and mid femora partly black,

following proportions:

7;

testaceous,

OOl = 6,5;

lateral

OPl = 5;

keels;

occipital

Tl = 4;

scutum,

carina complete;

POL =

scutellum and metanotum

shiny, smooth, weakly punctate, without sculpture among punctures; ,ootaul ices incomplete, reaching approximately 0,3 length of scutum; propodeum ret icu I a te rugose, dorsal

Fig.

and posterior surface;

379 -

Male genitalia of

wi th a

strong

transversa I keel

posterior surface with

Anteon tasmanianum n. sp. removed.

between

two longitudinal

(type):

left half

Anteoninae

keels;

median area smooth and shiny;

transversal (6:12);

bands;

gonoforceps

spurs 1,1,2. lOCUS TYPICUS: MM!

part of radial

(Fig.

379)

without dark

vein shorter than proximal

without distal

inner pointed process;

part tibial

(Tasmania)

holotype M!

and 3 paratypes MM!

in TW;

3 paratypes

in Ol.

HOSTS: unknown DISTRIBUTION: Australian TW!

fore-wing hyaline,

distal

Mount Barrow

TYPICAL MATERIAL:

583

NEW GU INEA:

region:

Baiyer R.,

Ol!

AUSTRALIA: TW!

Mount Barrow

Jimmy Valley -

(Tasmania),

Baiyer R.,

TV/!

38. Anteon permirum n. sp.

FEMALE: MALE:

unknown

fully

winged;

antennae brown,

length 2,0-2,87 mm;

black;

with segments 1-2 testaceous;

mandibles testaceous;

legs testaceous,

with coxae

and clubs of femora partly brown; antennae not distally thickened; antennal segments in following proportions: 10: 5: 8: 6: 6: 6: 6: 6: 9;

head shiny,

Fig.

380 -

strongly sculptured by

Male genitalia of

longitudinal

keels;

6:

vertex with

Anteon permirum n. sp. (type): right half removed.

584

Mem.

areolae and 5,5;

~

OPl

keels;

4;

Tl

Arner.

occipital ~

En!.

Ins!.,

carina complete,

4; scutum shiny,

sculpture among punctures;

no.

1984

~

POL

smooth,

notaulices

37,

7; Ol

~

4;

OOl

strongly punctate,

incomplete,

reaching

~

without

approximately

0,5 length of scutum; scutellum and metanotum shiny, smooth, without sculpture; between

propodeum

dorsal

longitudinal

reticulate rugose,

and posterior surface;

keels,

1,1,2. lOCUS TYP ICUS:

a strong transversal

with numerous transversal

without dark transversal bands; distal than proximal part (3:10); gonoforceps membranous process,

with

with

keels;

fore-wing

pointed process;

Ol!

spurs

in TVI;

1 paratype

unknown

DISTRIBUTION: Mts.,

tibial

(New Guinea)

TYPICAL MATERIAL: holotype M! and 9 paratypes /VIM! tAl in B; 4 paratypes lAM! in Ol. HOSTS:

hyaline,

part of radial vein much shorter (Fig. 380) without dorsal proximal

inner subdistal

Baiyer R.

keel

posterior surface without

Australian

West New Guinea),

nV!

Kougel

R.,

NEW GUI~jEA:

region: 8!

Bulolo,

TW!

Archbold lake

Saiyer R.,

Ol!

TW!

(Central

Jimmi

Valley,

TW!

39. Anteon involutum n. sp.

FEf'0AlE: MALE: brown,

Fig.

unknown

fully

winged;

length 2 mm;

with segment 1 testaceous;

381

-

Male genital ia of

black;

mandibles testaceous;

legs testaceous,

antennae

with coxae and clubs

Anteon involutum n. sp. (type): left half removed.

585

Anteoninae

of femora partly brown; segments shiny,

reticulate rugose;

occipital reaching

POL = 5;

weakly granulated;

0,5

length of scutum;

without sculpture; between

propodeum

dorsal

9:

4:

5:

thickened;

5:

5:

frons with a median

carina complete;

scutum shiny,

keel

antennae not distally

in following proportions:

OL = 2;

5:

keel,

notaulices

antenna I

5:

5:

without

OOL = 5;

8;

head

lateral

OPL = 2;

incomplete,

scutellum and

keels;

TL = 2;

approximately

metanotum shiny,

reticulate rugose,

and posterior surface;

5:

with a strong

smooth, transversal

posterior surface without

longitudinal keels, with numerous transversal keels; fore-wing hyal ine, without dark transversal bands; distal part of radial vein much shorter than proximal

part

(3:8);

membranous process and LOCUS TYP I CUS:

1,1(.

Gi I uwe

(Fig.

381)

without dorsal

inner process;

tibial

proximal

spurs 1,1,2.

(New Gu i nea)

holotype I'/I!

TYPICAL i\lATERIAL: HOSTS:

gonoforceps

without distal in B.

unknown

DISTRI3UTI0t-l: Guinea),

Australian

region:

I'~EW

GUINEA:

h1t.

Giluwe

(Malgi,

sE

~Iew

8!

40. Anteon superbum Dodd

Anteon Anteon Anteon typicus

FC::MALE:

superbus Jodd 1913: 181. superbus Dodd: Kieffer 1914b: 171. australis Dodd 1914a: 131 (n. syn.); locus r-lel son

fully

(Austra I i a) •

winged;

length 2,4 mm;

head black;

thorax and propodeum black or brown; antennae distally

thickened;

antennal

abdomen

antennae yellow;

testaceous;

segments

legs yellow;

in following proportions:

10: 6: 10: 6: 6: 6; 7: 6: 5: 7; head dull, granulated and reticulate rugose; frontal line complete; occipital carina complete; POL = 6; OL = 4,5;

OOL = 4,5;

keels,

POL = 4;

TL = 3;

pronotum rugose,

with anterior surface approximately

(13:13);

posterior surface shorter than

and metanotum shiny, punctures; scutum;

notaul ices

propodeum

posterior surface; hyal ine,

smooth,

scutum

punctate,

incomplete,

long

(6:14);

with

transversal

bands;

transversal

scutum,

approximately 0,5 two

longitudinal

median surface as rugose as

without dark

with

as posterior surface scutellum

without sculpture among

reaching

reticulate rugose,

as

distal

lateral

length of

keels on

areas;

fore-wing

part of radial

vein

shorter than proximal part (4:8); fore tarsal segments in following proportions: 22: 7: 8: 14: 42; enlarged claw (Fig. 382) with one bristle located on with

a proximal

prominence;

two rows of nearly 20

tibial spurs 1,1,2. i,1AL>:: unknown

segment 5 of front

larnellae;

apex with a

tarsus

group of 7

(Fig.

382)

lamellae;

586

Mem. Amer.

Fig.

En!.

Ins!.,

no.

37,1984

Chela of Anteon superbum Dodd from Nelson

382 -

LOCUS TYPICUS: Kuranda (Australia) TYPICAL MATERIAL: holotype F! of A.

(Queensland)

superbus in AD; holotype F! of A.

australis in AD. HOSTS:

unknown

DISTRIBUTION: Queensland),

Australian

AD!

Nelson

region:

AUSTRALIA:

(N Queensland),

Kuranda

(Cairns,

N

AD!

41. Anteon rufiscapum Dodd

Anteon rufiscapus Dodd 1913: 180 Anteon rufiscapus Dodd: Kieffer 1914b: 171.

FEMALE:

fully

winged;

length 2,16 mm;

black;

mandibles testaceous;

Anteoninae

Fig.

antennae brown,

383 -

Chela of Anteon rufiscapum Dodd

with segment 1 yellow;

antennae distally thickened; 14: 6:

5:

3:

5:

I ine complete;

5: POL

carina complete;

5:

587

5:

4:

antennal 8;

legs brown,

(type)

with tarsi

light;

segments in following proportions:

head dull,

fully reticulate rugose;

frontal

= 8; Ol = 4; OOl = 5; OPl = 5; Tl = 3; occipital

pronotum short,

long as anterior surface;

with posterior surface approximately

posterior surface shorter than scutum

as

(5:15);

scutum, scutellum and metanotum smooth, shiny, punctate, without sculpture among punctures; notaul ices incomplete, reaching approximately 0,20 length of scutum; transversal

keel

propodeum reticulate rugose,

between

without

longitudinal

beneath

the pterostigma;

dorsal

keels;

fore-wing

radial

with a strong

and posterior surface; with a dark

vein with distal

posterior surface

transversal

band

part shorter than

proximal part (2:9); fore tarsal segments in following proportions: 56: 15: 14: 16: 63; enlarged claw (Fig. 383) with one bristle located on a proximal prominence; segment 5 of front tarsus with one row of 5 lamellae, in ilddition to some bristles; tibial spurs 1,1,2. MALE:

unknown

lOCUS TYP ICUS:

Kuranda

TYPICAL lviAT"RIAL: HOS TS:

(Austral ia)

holotype F!

in AC>.

unknown

DISTRIBUTIOr,: Queensland),

Australian region:

AUSTRALIA:

Kuranda

AD!

42. An teon gira u Iti Dodd

Anteon giraulti Dodd 1914d: 251.

(Cairns,

N

588

Mem.

Fig.

384 -

Amer.

En!.

Inst., no. 37,

1984

Chela of Anteon giraulti Dodd

(type)

Anteoninae

FEMALE:

fully

winged;

abdomen brown;

antenna I segments 8;

head shiny,

shiny,

I"OL

reticulate rugose, ~

4;

~

3,5;

is smooth;

(6:11);

scutum,

mandibles testaceous;

antennae distally 10:

except for a

is smooth;

OL

reticulate rugose,

wh i ch

black;

legs yellow;

in following proportions:

ocellar triangle which complete;

length 2,4 mm;

antennae and

589

frontal ~

OOL

3;

8:

9:

7:

wide area

I ine complete; ~

OPL

6:

2,5;

6:

6:

6:

in fr'ont of

occipital ~

TL

thickened; 6:

2,5;

carina

pronotum

except for a short area near posterior margin

posterior surface of pronotum shorter than

scutellum

and metanotum shiny,

smooth,

scu tum

without

sculpture; notaulices incomplete, faintly visible near anterior margin of scutum; propodeum reticulate rugose, with a strong transversal keel between dorsal longitudinal distal

posterior surface; fore-wing

part of radial

segments 384)

and

keels;

front

in following proportions:

tarsus

(Fig.

1\-1ALE:

without dark

vein shorter than

with one bristle located on 384)

group of nearly 30

posterior surface without

hyaline,

with

7:

proximal

3: 4:

11:

a proximal

21;

tibial

(2:10);

bands;

fore tarsal

enlarged claw

prominence;

two rows of nearly 33

lamellae;

transversal

part

(Fig.

segment 5 of

lamellae;

apex with a

spurs 1,1,2.

unknown

LOCUS TYP I CUS:

Austra I i a

TYPICAL MATERIAL: HOSTS:

holotype F!

in AD.

unknown

DISTRI8UTIO~I:

Australian

,I Queensland),

region:

AUSTRALIA:

Yungaburra

(Cairns distr.,

AD!

5. Prioranteon n. gen.

TYPE SPEC I ES: Prioranteon casalei n. sp. FEMALE (Figs 385, 386): micropterous; thoracic structure simi lar to that of

Gonatopodinae: pronotum remarkably elongate; scutum forming a stalk

between

the pro- and metanotum;

propodeum segments;

humped;

located further' distally 390)

MALE:

than

i nv i si b Ie;

with 6 segments;

inner side of enlarged claw

prominence not bearing bristles; 388,

metanotum

maxillary palpi

(Fig.387)

metathorax + labial

tibial

prominence;

chela

spurs 1,1,2.

unknown

Palaearctic,

with 3

1-2 short bristles or peg-like hairs

the proximal

without rudimentary claw;

DISTRI8UTIO~:

palpi

with a proximal

Ethiopian

HOSTS: unknown SPECIES: 3

PALAEARCT I C SPEC I ES OF PRIORANTEON

(Figs 387,

590

Mem. Amer. En!.

Fig.

385 -

Ins!.,

no. 37,

1984

Female of Prioranteon casalei n.

sp.

(type)

Anteoninae

Fig.

386 -

One species

Female of

591

l'riora"teon casalei n. sp.

(type)

is known.

Prioranteon biroi n. sp.

FEMALE:

micropterous;

scutellum;

fore

testaceous,

with

teeth brown;

segments 1-3 testaceous; testaceous; black, 10:

8:

10:

10;

punctures;

segments

occipital

frons

mandibles ';,ith 4 teeth progressing

posterior; weakly

vertex flat;

punctate,

posterior

transversal

narrower than tegulae,but nearly as reduced; long as scutum humped,

disc;

very

without median

and

smooth,

tarsi

13:

13:

11:

keel,

but

POL = 7;

with Tl =

anterior one to

without sculpture or very

very

posterior lobe directed

humped,

scute II um; 389 8);

shiny,

smooth,

tegulae and

long anterior disc,

towards

carinate;

reduced

tegulae;

hollow,

without fore

reaching scutum

rugose,

sculpture,

wings on

very

nearly

as

sides of

fore wi ngs not reachi ng transversa I furrow beh i nd metanotum

invisible;

separated from scutellum by

transversely

hind 18:

posterior margin of collar almost straight,

as broad;

(Fig.

OOl = 11;

larger from

long as posterior collar of pronotum, scutellum

5:

without sculpture among

Ol = 5;

humped and

10:

impression se;oarating disc from a posterior collar,

without a

long

scutellum

pronotum shiny,

with a

with

with coxae almost fully

proportions:

POL = 4;

as

mandibles

antennae brown,

last segment of mid and

weakly punctate,

furrow;

long

black;

with posterior collar whitish-

legs testaceous,

carina complete;

median

nearly as

testaceous;

in following

head shiny,

longitudinal

12;

clypeus

pronotum black,

tegulae testaceous;

antennal

reduced,

length 3,25-3,80 mm;

clubs of femora darkened,

brown;

a

wings very

hind wings invisible;

a

metathorax + propodeum

transversal

hollow

impression

anterior surface of metathorax + propodeum shiny,

592

Mem. Amer.

Ent.

Inst.,

no. 37,

1984

i

I

j

!

\\\. /' -.

~~/

Fig. 387 -

/

Chela of Prioranteon biroi n. sp.

(type)

593

Anteoninae

partly granulated and smooth; disc of metathorax + propodeum with a distinct longitudinal median furrow; posterior surface of propodeum dull, granulated and with some weak smooth,

shiny,

complete;

fore tarsal

enlarged claw

(Fig.

the proximal

prominence; lamellae;

long);

meso-metapleural

with one bristle

into a

small

unknown

hook;

LOCUS TYPICUS:

tibial

11:

3:

6:

13:

25;

tarsus

a group of 5

387)

(Fig.

lamellae

with 2 rows

(among which

segment 2 of front

1

tarsus produced

spurs 1,1,2.

SzSztMiklos

TYPICAL MATERIAL:

pleura

located further distally than

segment 5 of front

apex with

rudimentary claw absent;

MALE:

striae;

suture distinct and

segments in following proportions:

387)

of nearly 29 very

irregular longitudinal

without sculpture;

(Hungary)

holotype F!

and 1 paratype F!

1 paratyep F!

in BU;

in OL. HOSTS:

unknown

DISTRIBUTION: NOTES:

Palaearctic region:

the species

HUNGARY:

is named in honor of the

SzSztMiklos, late Mr.

L.

BU! Biro.

ETHIOPIAN SPECIES OF PRIORANTEON

Two species are known.

KEY TO THE SPECIES

FEMALES

Posterior

surface of

propodeum strongly

. . • • • • • • . • • . . . . • • . . • . . . . • • . . • . . • . .. -

Posterior

surface of

transversely striate .•• sp.

1. richardsi n.

propodeum granulated,

kee Is. • • • • • • • . • • • • • . • . . • • • • • • • • • . •.

2.

without

casalei n.

transversal

sp.

MALES

Unknown

In the key to the Ethiopian species, the Palaearctic species PI'. biroi shou I d be near PI'. casalei, because the posterior surface of the propodeum is granu I ated and not transverse I y str i ate. Casalei and biroi NOTES:

can be distinguished as follows:

594

{v,em.

Amer.

Ent.

Inst.,

no.

t,letathorax

+ propodeum with a distinct

disc;

with

head

reaching

frons

transversal

fully

black;

furrow behind

frons black, reaching

with

a

central

furrow behind

winus

scutellum

longitudinal

yellowish

scutellum

furrow on

shorter, (Fig.

biroi n;

+ propodeum without

Metathorax

1984

longitudinal

fore

. .. . .. . .. . . . . . . . . .. . . . .. . . . . .. . -

37,

not

tl).

389

sp.

furrow on disc;

area;

fore

wings

longer,

389 A}......

(Fig.

casalei n.

.. . . . . .

sp.

1. Prioranteon richardsi n. sp.

FEMALE:

rnicropterous;

scutellum;

fore

hind wings

testaceous with

teeth brown;

segments 1-2 and proximal pronotum black, legs brown, tibiae and B:

5:

17:

with

with tarsi

12:

wings

reduced,

clypeus

testaceous;

part of coxae,

testaceous; 10:

8:

7:

antenna I segments 7:

9;

TL =

mandibles

carina complete;

larger from anterior one to posterior; smooth,

weakly punctate,

only on

sides;

pronotum

posterior transversal narrower than tegulae, nearl y

but

as

reduced;

disc;

a

behind

posterior

humped,

nearly

as

and

shiny,

scutellum;

(Fig.

24;

distally

weakly

fore

proximal

tarsal (Fig.

and

hook;

MALE:

unknown

LOCUS TYPICUS:

tibial

388)

absent;

spurs

Van Reenen

TYPICAL MATERIAL:

12;

very

OPL

=

6;

granulated

long anterior disc;

towards

posterior collar, reaching

tegulae;

hollow,

scutum

rugose,

without sculpture,

very

nearly

as

tegulae and reduced fore wings

wings

not

reaching

invisible; a

transversal

metathorax +

transversal

hollow

posterior surface strongly striate;

with

in following 2 bristles

apex

with

a

11:

tarsus

(Fig.

3BB)

group of nearly 4

segment 2 of front

South Africa)

in 8M.

pleura strongly proportions:

located further

segment 5 of front

1,1,2.

(Natal,

holotype F!

punctate,

teeth progressing

disc from a

metanotum

segments

lamellae;

rudimentary claw

small

=

pronotum shiny,

strong and complete;

prominence;

with 2 rows of nearly 43 lamellae;

OOl

anterior surface of metathorax +

granulated;

suture distinct,

striate;

into a

proportions:

strongly

3;

with 4

separated from scutellum by

enlarged claw

than

fore

389 C);

transversely carinate;

transversely

smooth,

long as broad;

impression

12:

humped

lobe directed

humped,

meso-metapleural

=

OL

vertex flat;

impression separating

scutellum

propodeum dull,

testaceous;

sculpture among punctures,

very

propodeum

7:

smooth,

long as posterior coil ar of pronoturn, scutellum

sides of scutum

furrow

with

with

stalk of femora,

posterior margin of collar almost straight,

without a

long as scutum, on

without

4;

as

antennae brown,

in following

head shiny,

=

long

mandibles

tegulae testaceous;

trochanters,

POL

occipital

black;

half of 3 testaceous;

without sculpture among punctures; 11;

nearly as

length 4,68 mm;

anterior margin and posterior collar

distal

10:

very

invisible;

tarsus produced

3:

595

Anteoninae

('ig.

388 -

Chela of Prioranteon richardsi n.

sp.

(type)

596

Mem.

Fig.

389 -

Amer.

Inst.,

no.

37,1984

Prioranteon casalei n. sp. (A), biroi n. richardsi n. sp. (C).

Reduced fore wings of sp.

HOSTS:

Ent.

(B),

unknown

DISTRIBUTION:

Ethiopian

Natal),

8M!

NOTES:

the species

region:

is named

SOUTH AFRICA:

in honor of Mr.

Van Reenen

Owain W.

(Drakensberg,

Richards.

2. Prioranteon casalei n. sp.

FEMALE long

(Figs 385,

386):

as scutellum;

mandibles testaceous, longitudinal

segments shiny,

=

with

genae,

=

OPL

5;

pronotum shiny, with a

TL

=

very

5:

clypeus and a

abdomen reddish-brown;

tarsi

13:

8;

occipital

9:

8:

testaceous;

7:

carina complete;

humped and very

than disc;

posterior margin of collar almost straight, posterior

disc from

antennal

5:

9;

head

POL = 3,5;

mandibles with 4 vertex flat;

long anterior disc;

impression separating a

6:

weakly punctate only on

trasversal but without

5:

anterior one to posterior;

without sculpture,

with

with posterior collar and

without sculpture among punctures;

larger from smooth,

8:

and

nearly as

black;

antennae brown,

testaceous;

junctions

in following proportions:

teeth progressing sides,

length 3,25-4,00 mm;

pronotum black,

tegulae

trochanters,

weakly punctate,

9;

teeth brown;

testaceous;

margins testaceous;

legs brown,

OL

with

fore wings very reduced,

invisible;

median area on frons yellowish;

segments 1--2 partly lateral

micropterous;

hind wings

a

lobe directed towards

posterior

posterior collar, tegulae;

reaching

narrower tegulae,

scutum nearly as

Anteoninae

Fig.

390 -

Chela of

597

Prioranteon casalei n. sp. (type)

long as posterior coil ar of pronotum,

hollow,

rugose,

very reduced;

scutellum humped, shiny, smooth, without sculpture, nearly as long as scutum, nearly as broad; tegulae and reduced fore wings on sides of scutum and scutellum; scutellum humped,

(Fig.

389 A);

fore w;ngs reaching metanotum

transversal

invisible;

separated from scutellum by a

furrow

behind

metathorax + propodeum

transversal

hollow

impression

598

Mem.

transversely carinate;

Amer.

En!.

Ins!.,

no.

37,

1984

anterior surface of metathorax + propodeum shiny,

smooth, without sculpture; posterior surface dul', granulated, without transversal striae; meso-metapleural suture distinct, strong and complete; pleura dull,

granulated,

with weak transversal

segments in following proportions: 390)

with 2 bristles

prominence; lamellae; absent;

9:

2: 5:

keels;

12: 21;

located further distally than

segment 5 of front

tarsus

(Fig.

390)

tarsus produced

(Fig.

the proximal with 1 row of nearly 33

apex with a group of nearly 3 lamellae;

segment 2 of front

fore tarsal

enlarged claw

rudimentary claw

into a small

hook;

tibial

spurs 1,1,2. MALE:

unknown

LOCUS TYPICUS:

Milnerton

TYPICAL MATERIAL: FF!

(South Africa)

holotype F!

and 7 paratypes FF!

in BM;

4 paratypes

in OL.

HOSTS: unknown DISTRIBUTION: Ethiopian region:

SOUTH AFRICA:

BM! OL! NOTES: the species is named in honor of Mr.

Fig. 391

-

Milnerton

(Cape Town),

Achi lie Casale.

Female of Bocchoides bekilyensis Benoit

(holotype)

Bocchinae

599

5. SUBFAMILY BOCCHINAE RICHARDS

Bocchini Richards 1939: 189. Bocchini Muesebeck and Walkley 1951: 1039. Bocchinae Nagy 1967b: 334.

TYPE

Bocchus Ashmead 1893

GENUS:

FEMALE:

fully

brachypterous

winged (Fig.

(Fig.

424);

391)

or micropterous (Fig.

fore-wing

(in fully

452)

or

winged species)

with costal,

median and submedian cells clearly enclosed by pigmented veins (Fig. 391); maxi Ilary palpi with 6 segments; labial palpi with 3 segments; front with

tarsus chelate; 1,

2,

chela

3 or 4 teeth

three

large teeth and a

(Fig.

396 A);

rarely

(Fig.

(Fig.

405)

396);

with rudimentary claw;

rudimentary

tooth between

quadridentate mandibles with

larger from anterior one to posterior; of strong

teeth

(Fig.

405);

mandibles

quadridentate mandibles usually teeth progressing

enlarged claw usually with a row

segment 5 of front

tarsus

(Fig.

405)

only with a preapical lamella, without other lamellae; pronotal directed towards the tegulae present; occipital carina complete, incomplete;

petiole usually very

long,

with

the two posterior teeth

rarely

short;

usually tubercles rarely

antennae without

tufts of long hairs; pterostigma usually very reduced, sometimes wide; tibial spurs 1,1,1, sometimes 1,1,2. MALE: fully winged (Fig. 393) or micropterous (Fig. 454); fore-wing (in fully

winged species)

enclosed by

Fig.

with costal,

pigmented veins)(Fig.

392 -

Female of

median and submedian cells clearly 393);

maxillary palpi

with 6 segments;

Bocchoides bekilyensis Benoit (holotype)

600

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

/

Fig.

393 -

Male of Bocchoides bekilyensis Benoit

(allotype):

antennal

segments 7-10 missing.

labial

palpi

with 3 segments;

mandibles with 1, 2,

3 or 4 teeth;

tridentate or quadridentate mandibles with teeth not progressing from anterior one to posterior;

occipital

carina complete;

tibial

larger spurs

1,1,2. DISTRIBUTION: Palaearctic, Ethiopian, Oriental, Nearctic, Australian HOSTS: Issidae; Cicadellidae (exc Iud i ng I diocerinae, Macropsinae and

Typhlocybinae) GENERA: 6 SPECIES:

39

KEY TO THE GENERA OF BOCCHINAE

FEMALES

Occipital head;

carina

enlarged

incomplete,

claw with

only

visible on dorsal

one preapical

lamella

(Fig.

side

of

399) .•

Bocchinae

...........................

2

Occipital

carina

preapical

lamella;

preapical

lamella

Segment with tooth

5 of

other

enlarged Enlarged front

most

segment

tarsus with (Fig.

front

of

lamella,

claw at

never

least

with

one

lamellae

preapical

(Figs

395,

lamella, 442,

451,

sometimes 456);

claw without teeth (Figs 395,442,451,456) . . . . . . . . . 3 claw wi th numerous bristles (Fig. 395 A); segment 4 of

Enlarged

least

claw withoit

451,456);

segment

segment

Fore-wing

as

long as

segment

front

1

(Fig.

395 B) . . . . . . . .

Bocchoides Beno i t

1.

brist les or

4 of

other appendixes

tarsus

than

0,5

(Figs 442, times as

1

(Figs

442,

(Fig.

452) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ..

complete

Fore-wing with

451,

less

reduced

........................... Fore-wing 5

one

tarsus with

Bocchus Asmead

tarsus without

group

tarsus at

long as

front

enlarged

3.

• • • • • • • • . • • • • • • • • • • • • • • • • ••

4

claw never with 5 of

one preapical

405);

405) •••••••••••

5 of

Chelothelius Reinhard

2.

enlarged

(Figs 395,405) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2

front

wi th a di stal 3

at

lamellae

(Fig.

Segment

complete;

601

radial

f\1ystrophorus Forster

6.

(Figs 438,

456) •••••••••••••••••••••••• 4

449) ••••••.••••••••••••••••••••• 5

vein distinct

(Fig.

438) . . . . . . . . . . . . . . . . .

••••••••• • ••••••••• • • •••• •• 4. Mirodryinus Ponomarenko Fore-wing with no radial vein (Fig. 449) . . . . . . . . . . . . . . . . . . . . . . . • ••••••••••••••••••••••••••

Nadiimancus Moczar

5.

MALES

Temp I e s not

vis i b I e

(F i g s

393,

• • • • • • • • • • • • • • • • • • • • • • • • • •• 2

Temples distinct (Figs 402,403,440,441,454,455) . . . . . . . . . . 2 Fore-wi ng reduced (F i g. 454) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ........................... Fore-wing

3

394) . . . . . . . . . . . . . . . . . . . . . . . . . . ..

Bocchoides Ben 0 i t

1.

compl ete

Mystrophorus Forster

6.

(Figs 402,

440) . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3

Propodeum with areolae very

small,

440) ••.••••••••••••••••••••

4.

Propodeum wi th at

I east

least

tegulae

as

large as

part

of

(Fig.

• • • . • • • • • • • • • • • • • • • • • • • • • ••

3.

less wide

than

tegulae

(Fig.

Mirodryinus Ponomarenko the areolae very

large,

at

402) . . . . . . . . . . . . . . . . . . . . . . . . . . .

Bocchus Asmead

The ma les of Chelothelius Rei nhard and Radiimancus Moczar are unknown.

1. Bocchoides Benoit

602

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

Bocchoides Benoit 1953b: 424.

Bocchoides bekilyensis Benoi t 1953, monotyp ic.

TYPE SPEC I ES: FEMALE labial

(Figs 391,392): palpi

tridentate

(Fig.

396 B);

segment 5 of front front

tarsus at

not reaching MALE

least as

palpi

visible;

394):

HOSTS:

(Fig.

lamellae (Fig.

395);

1 (Fig.

pronotal

395);

395);

segment 4 of tubercles

spurs 1,1,2. winged;

with 3 segments;

mandibles tridentate,

DISTRIBUTION:

mandibles

with numerous bristles

long as segment

fully

with 6 segments;

carina complete;

with numerous

tibial

anterior one to posterior

maxillary palpi

occipital

enlarged claw

tarsus

tegulae;

(Figs 393,

labial

fully winged;

with 3 segments;

(Fig.

maxillary palpi

occipital with

with 6 segments;

carina complete;

teeth not progressing

396 D);

tibial

temples not larger from

spurs 1,1,2.

Ethiopian

unknown

SPECIES:

1

13occhoides bekilyensis Benoi t

Fig.

394 -

Male of

Bocchoides bekilyensis Benoit (allotype): antenna I segments 7-10 missing.

603

Bocchinae

Fig.

395 -

Female of Bocchoides bekilyensis Benoit fore tarsa I segmen ts.

(holotype):

A:

chela;

B:

604

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

Bocchoides bekilyensis Benoi t 1953b: 424.

FEMALE

(Figs 391,392):

fully

winged;

clypeus and mandibles testaceous; darkened;

thorax,

testaceous,

propodeum, 14:

7:

33:

distally thickened;

14:

12:

head dull,

10:

8:

hairy,

7:

with

legs brown-

antennal

8:

black,

with segments 8-10

pet iole and abdomen black;

with coxae and clubs of femora dark;

following proportions: slightly

length 6,25-6,50 mm;

antennae testaceous,

10;

rugose,

segments

in

antennae

sculptured by short

irregular keels; occipital carina complete, reaching posterior margin of eyes; temples invisible; thorax and propodeum shiny, hairy; pronotum and scutum smooth,

without sculpture;

without sculpture;

propodeum reticulate rugose; median

area rugose;

five times as as broad

enlarged claw

for", tarsal

(Fig.

segment 5 of front

395)

tarsus

extending continuously MALE

(Figs 393,

mandibles yellow;

394):

(14:2,5);

(Fig.

and

hind

to the apex; fully

4:

14:

rugose;

mandibles tridentate,

396 D);

occipital

temples

invisible;

scutum;

scutellum granulated; without

(25:7);

tibial

yellow;

9;

legs yellow,

antennal

with

segments

1-6

intermediate tooth

in

(Fig.

visible only on anterior half of

metanotum reticulate rugose;

longitudinal

propodeum

keels on posterior surface;

long and ovoidal,

more than

fore-

three times as

long as

spurs 1,1,2. in P.

unknown

DISTRIBUTION: Ethiopian region: MADAGASCAR: 8ekily, P! NOTES: B. bekilyensis is very different from the other Bocchinae. general

spurs

(1,1,2)

are similar to tibial

tridentate mandibles the species

spurs

is included

arrangement of thi s genus prQbab I y

in

is between

The

Anteoninae or as in Dryininae. Female of Dryininae. Only for the Bocchinae. The exact Bocchinae and Dryininae.

structure of the chela is simi lar to that of

Dryininae. The pterostigma is long and narrow tibial

with

in allotype);

posterior margin of eyes;

LOCUS TYP ICUS: Beki Iy (Madagascar) TYPICAL MATERIAL: holotype F! and allotype M! HOSTS:

head black,

head and scutum reticulate

reaching

notaul ices weakly

wings with pterostigma broad

tegulae

with one rudimentary

carina complete,

lamellae

spurs 1,1,2.

length 3,12 mm;

10:

long

(11:14);

without other appendixes;

with 2-3 rows of 63

(segments 7-10 missing

10:

keels;

nearly

fore trochanters nearly twice as

abdomen brown-black;

tibiae darkened;

longitudinal

1 shorter than segment 4

tibial

winged;

antennae yellow

following proportions: 5:

reticulate rugose,

395)

two

long and narrow,

with numerous bristles,

thorax and propodeum black; femora

with pterostigma

segment

scutellum smooth,

with some areolae near scutellum;

posterior surface with

fore-wing

long as broad

(8:4);

notaul ices invisible;

metanotum smooth,

2. Chelothelius Reinhard

Bocchinae

605

A

Fig.

396 -

E

Mandibles of Bocchinae:

rudimentary tooth;

B:

from anterior one to posterior; mandible; E:

quadridentate mandible with one

C:

mandible with one tooth;

tridentate mandible with one rudimentary

Chelothelius Chelothelius Chelothelius Chelothelius

Reinhard

1863:

D:

larger

tridentate

intermediate tooth.

409.

Reinhard:

Kieffer

Reinhard:

Kieffer 1914b: 56.

in Kieffer et Marshall

Reinhard:

Berland

1928:

1905:

125.

143.

Chelothelius gryps Reinhard 1863, monotypic.

TYPE SPECIES: FEMALE

A:

tridentate mandible with teeth progressing

(Figs 397,

398):

fully

winged,

maxillary palpi

with 6 segments;

labial palpi with 3 segments: occipital carina incomplete, only visible on dorsal side of head; mandibles tridentate: enlarged claw with one preapical with

lamella and some bristles

lamellae

(Fig.

399);

pronotal

(Fig.

399);

segment 5 of front

tubercles not reaching

spurs 1,1,2. MALE: unknown DISTRIBUTION: Palaearctic HOSTS: unknown SPECIES:

2

PALAEARCTIC SPECIES OF

Two species are known.

CHELOTHELIUS

tegulae;

tarsus tibial

606

Mem.

Fig.

397 -

Amer.

Ent.

Ins!.,

no.

37,

1984

Female of Chelothelius gryps Reinhard from South France

KEY TO THE SPECIES

FEMALES

Head

fully

granulated,

scutellum dull,

wi th numerous keel s,

granulated and

•••• •••• •••••••• ••• 1. gryps Rei nhard Head shiny, granulated and with keels, wide areas

on

sides

of

ocell i

except

whi ch are

sculpture;

scutum and

sculpture,

weakly granulated,

without

berlandi

Bernard

•••••••••••••••••• • 2.

dull;

scutum and

with keels . . . . . . . .

scutellum shiny,

for

two

smooth and wi thout smooth,

almost

without

keels . . . . . . . . . . . . . . . . . . .

607

Bocchinae

MALES

Unknown

1. Chelothelius gryps Reinhard

Chelothelius gryps Rei nhard 1863: 410. Dryinus gryps (Reinhard): Dalla Torre 1898: 544. Chelotelius gryps Reinhard: Kieffer in Kieffer et Marshall 1905: 127. Chelothelius gryps Reinhard: Kieffer 1914b: 57. Chelothelius gryps Rei nhard: Ber I and 1928: 143. Chelothelius gryps Rei nhard: Bernard 1935a: 41.

FEMALE

(Figs 397,

398):

fully

winged;

length 5,0-5,4 mm;

antennae brown,

with segments 1-2 testaceous; head black, with mandibles, clypeus, a narrow frontal region near clypeus, malar space and a region around orbits reddish-testaceous;

Fig.

398 -

propectus black;

pronotum black,

with margins

Female of Chelothelius gryps Reinhard from South France

608

Mem.

Fig.

reddish;

399 -

Amer.

Ent.

Inst.,

no.

37,

1984

Chela of Chelothelius gryps Reinhard from South France

scutum,

scutellum,

metanotum,

mesopleura,

metapleura and

propodeum black; petiole black; abdomen black-brown; legs testaceous, with coxae partly black and clubs of femora brown; tegulae testaceous; antenna I segments in following proportions:

12: 7:

24:

13:

13: 8:

6:

6:

6:

9; head dull, granulated, with numerous keels, mainly longitudinal; mandibles with 3 teeth; ocellar triangle and occiput smooth; frons with incomplete median

longitudinal

carina visible only on dorsal temples almost transversal transversal

invisible;

keel;

= 5; Ol = 4; OOl = 10; occipital

POL

region of head,

pronotum shiny,

but not reaching eyes;

granulated,

with weak

keels; pronotal turcles not reaching tegulae; pronotum with a impression between anterior collar and posterior humped disc;

scutum dull,

granulated,

with numrous irregular keels or areolae;

notaulices invisible; scutellum as sculptured as scutum; metanotum fully reticulate rugose; propodeum fully reticulate rugose, with 2 incomplete longitudinal bands,

keels on posterior surface;

one on

the basal

fore wings with 2 dark

transversal

cell s and another beneath the pterost i gma;

pterostigma long and narrow;

distal

part of radial

vein

nearly as

long as

proximal part, not reaching wing margin; fore tarsal segments in following proportions: 13: 3: 5: 9: 16; segment 5 of front tarsus (Fig. 399) with some rows of nearly 26 lamellae extending continuously to the apex; enlarged claw

(Fig.

segment 3 of front tibial

spurs 1,1,2.

MALE:

unknown

lOCUS TYPICUS:

399)

with 3 proximal

tarsus produced

Bolzano

(Italy)

into a

teeth and one preapical hook,

with no very

lamella;

long bristle,

Bocchinae

TYPICAL MATERIAL: HOSTS:

probably

609

lost.

unknown

DISTRIBUTION: Goudargues

Palaearctic region:

(Gard),

OL!

ITALY:

FRANCE: Southern France,

Bolzano

P!

Ussel-

(Reinhard 1863).

2. Chelothelius berlandi Bernard

Chelothelius berlandi Bernard 1935a: 41. Chelothelius berlandi Bernard: Bernard 1935b: 53.

FEMALE:

fully

winged;

and mandibles

length 3,75 mm;

testacous;

head black,

antennae testaceous;

segment 10 distally testaceous;

thorax fully black,

(black,

petiole black;

with margins reddish);

testaceous,

with clypeus brown

with segments 6-10 brown; except for pronotum

abdomen black-brown;

with coxae and clubs of femora partly brown;

testaceous;

antennal 6:

9:

ocelli;

frons granulated and with

Fig.

9;

segments in following proportions:

10: 9:

head shiny,

400 -

smooth and

legs

tegulae

10: 5:

23:

14:

12:

without sculpture on sides of

irregular keels;

ocellar triangle rugose;

Chela of Chelothelius berlandi Bernard

(type)

610

Mem. Amer.

occipital

carina

reaching eyes;

incomplete,

Ent.

Ins!.,

smooth,

weakly

tegulae;

without keels;

granulated,

propodeum fully longitudinal basal

proportions:

long and narrow; 12:

and without

3:

5:

9:

very

16;

(Fig.

Fig.

401

400)

-

part

transversal pronotal

smooth,

(9:8),

impression

tubercles not

weakly granulated,

reticulate rugose;

posterior surface with 2 transversal

region;

fore tarsal

distal

bands,

the

vein

wing margin;

in following

tarsus produced

segment 5 of front

tarsus

lamellae extending continuously

with 7 bristles and one preapical

Female of Bocchus umber n.

incomplete

one on

part of radial

not reaching segments

segment 3 of front

long bristles;

with some rows of nearly 30 enlarged claw

dull;

the apical

longer than proximal

pterostigma

disc;

metanotum fully

fore wings with 2 dark

cells and another on

slightly

hook

reticulate rugose,

keels;

region of head,

with a

scutum and scutellum shiny,

notaulices invisible;

1984

POL = 5; Ol = 3; OOl = 8;

between anterior collar and posterior humped reaching

37,

visible only on dorsal

mandibles with 3 teeth;

pronotum shiny,

no.

sp.

to

into a

(Fig.

lamella;

(type)

400)

the apex; tibial

Bocchinae

611

spurs 1,1,2. MALE: unknown LOCUS TYP ICUS:

Plage de Saint-Raphael

TYPICAL MATERIAL: HOSTS:

(France)

in P.

unknown

DISTRIBUTION: Var),

holotype F!

Palaearctic region:

FRANCE:

Plage de Saint-Raphael

(Frejus,

P!

3. Bocchus Ashmead

Bocchus Ashmead 1893: 91. Phorbas Ashmead 1893: 90 (preoccup i ed by Phorbas Duchassa i ng and Michelotti

Fig.

402 -

1864;

syn.

proposed by Richards 1939);

type species

Male of Bocchus pedunculatus Nagy from Sarawak

612

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

Phorbas laticeps Ashmead 1893, orig. desig. Eukoebeleia R.C.L. Perkins 1905: 59 (syn. proposed by Fenton 1918b); type species Eukoebeleia mirabilis R.C.L. Perkins 1905, monotypic.

Tetradryinus Kieffer 1913c: 325 (n. syn.); type species Tetradryinus flavipes (Kieffer 1906), orig. desig. Anteonella Dodd 1913: 181 (n. syn.); type species Anteonella robusta Dodd 1913, orig. desig. Phorbasia Kieffer 1913b: 58 (syn. proposed by Richards 1939); type species Phorbasia laticeps (Ashmead 1893), orig. desig. Neoanteon Fouts 1922: 633 (syn. proposed by Nagy 1969); type species Neoanteon rubrica Fouts 1922, monotypic. Hymenodryinus Benoit 1953b: 427 (n. syn.); type species flymenodryinus seyrigi Benoit 1953b, orig. desig. Bocchus Ashmead: Nagy 1969: 321.

TYPE SPEC IES: FEMALE

Bocchus flavicollis Ashmead 1893, orig. desig.

(Figs 24,

401,

424,

425):

fully

winged

brachypterous or micropterous (Figs 424, segments;

labial

palpi

with 3 segments;

quadridentate mandibles usually

425);

(Figs 24,

401)

maxi Ilary palpi

mandibles with 1,

with a rudimentary

2,

or with 6 3 or 4 teeth;

tooth between

the two

posterior teeth (Fig. 396 A); rarely quadridentate mandibles with teeth progressing larger from anterior one to posterior; occipital carina complete; segment 5 of front tarsus with one preapical lamella, never with other lamellae (Fig. 404); enlarged claw with at least one tooth (Fig. 404);

pronotal

Fig.

tubercles reaching

403 -

tegulae;

tibial

spurs 1,1,1.

Male of Bocchus pedunculatus Nagy from Sarawak

Bocchinae

MALE

(Figs 402,

403):

fully

winged;

613

mandibles with 1, 2,

3 or 4 teeth;

quadridentate and tridentate mandibles with teeth not progressing from anterior one to posterior; wide as tegulae

(Fig.

segments;

palpi

labial

402);

larger

areolae of propodeum at least partly as

temples distinct;

with 3 segments;

cnaxillary palpi

fore wings complete;

with 6 tibial

spurs

1,1,2. DISTRIBUTION: HOSTS:

Palaearctic,

Ethiopian, Oriental,

Nearctic,

Australian

Issidae

SPECIES:

29

PALAEARCTIC SPECIES OF

BOCCHUS

Nine species are known.

KEY TO THE SPECIES

FEMALES

Species brachypterous, wi th fore-wing extending apex of propodeum ••••.•••••• 1. italicus n. sp. Species

fully

winged

(Fig.

wi thout

only

to

the

401) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 transversal bands ••••••••••••••

2

Fore-wing hyal ine,

dark

3

umbel' n. sp. 1-2 dark transversa I bands ••••••••••••••••••••• 3 Fore-wing wi th one dark transversal band •••••••••••••••••••••• 4 · . • . . • . . . . . . . . . . . . . . . . . . . . .. 2.

Fore-wi ng wi th

bands . . . . . . . . . . . . . . . . . . . . . 5 much shorter than scutellum; head black, with mandibles yellow; scutellum and propodeum black; pronotum, scutum and metanotum reddish • ••• · . . . . . . . . . . . . . . . . . . . . . . . . . .. 3. szelenyii Mo c za r Head with OOL more than twice as long as POL; scutellum nearly twice as long as metanotum; head reddish-testaceous; thorax and Fore-wing with

4

two dark

Head wi th POL not

transversal

shorter

than OOL;

propodeum black . . . . . . . . . . . . .

5

Thorax

fully

Thorax at

6

reddish ••••••••

least

8

4. slovacus Strejcek 9. parthicus Ponomarenko

partly black . . . . . . . . . . . . . • . . • . . . . . . . . . . . . . . . . • 6

5. paglianoi n. sp. reddish . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 Head mostly reddish; frons and vertex at least partly reddish · . • . . • . . . • . . • . . • . . . . . . . . . . .. 6. scaramozzinoi n. sp. Head fully black, at most only with mandibles and and clypeus not black ••••••••••••••••••••••••••••••••••••••••••••••••••••• 8 Pronotum reddish; scutum, scutellum and metanotum black . . . . . . . . Thorax

fully black •••••••.••

Thorax partly black and

7

metanotum not

614

Mem.

Amer.

Ent.

Inst.,

7. Pronotum and

scutum reddish;

no.

37,

1984

scobiolae Nagy

scutellum and metanotum black ..•.•

8. europaeus Bernard

1v1AlES

Only known.

B. scobiolae Nagy and B. slovacus Strejcek are

the males of

I cannot propose a key

to these

two ma les because

I

have not seen

B. scobiolae.

the male of

1. Bocchus italicus n. sp.

FEMALE:

brachypterous

propodeum); occipital distally 7:

7:

carina complete; thickened;

6,5:

6:

scutum dull, pronotal dull,

(fore wings reaching

length 2,56 mm;

6:

head dull,

granulated; antennae

antenna I segmentes in following proportions:

6:

9;

POL

granulated,

=

6;

=

Ol

3;

=

OOl

10;

OPl

with notaulices complete, tegulae;

scutellum

propodeum reticulate rugose,

posterior surface;

only posterior apex of

testaceous;

frons with a median complete keel;

tubercles reaching

rugose;

fully

fore-wing

short,

11:

=

TL

7:

metanotum

longitudinal

keels on

reaching only posterior

apex of propodeum, without pterostigma; mandibles tridentate, with a rudimentary tooth between the two posterior teeth; fore tarsal segments following proportions: 46: produced

6:

into a short hook;

10:

35:

42;

segment 3 of front

enlarged claw

(Fig.

404)

with

teeth and one bristle; segment 5 of front tarsus (Fig. 404) bristles and one preapical lamella; tibial spurs 1,1,1. MALE:

in

tarsus a row of 5 with

two

unknown

lOCUS TYPICUS:

lido (Venezia,

TYP ICAl MATER IAl: HOSTS:

11:

9;

granulated;

two

not complete,

8;

posteriorly separated;

dull,

with

=

holotype F!

Italy) in CO.

unknown

OISTRIBUTION:

Palaearctic region:

ITALY:

Lido

(Venezia),

CO!

2. Bocchus umber n. sp.

FEMALE

(Figs 24,401):

testaceous; with

tergite

hairy;

winged;

almost hairless;

occipital

distinct and complete,

length 2,50-2,62 mm;

thorax and propodeum black;

1 and apex brownish;

abdomen

granulated;

fully

antennae yellow;

tegulae yellow;

head,

carina complete;

pronotum, POL = 4;

posteriorly separated;

head reddishabdomen black,

legs yellow;

body dull,

scutum and scutellum OOl = 5;

notaulices

metanotum smooth,

without

Bocch i nae

615

Ii I

I

) 1,\ I \

\

\

Fig. 404 -

sculpture;

Chela of Bocchus italicus n.

scutellum nearly

reticulate rugose,

with 2

twice as

longitudinal

sp.

long as metanotum;

(type)

propodeum

keels on posterior surface;

fore-wing

616

Mem.

Amer.

Ent.

Inst.,

no.

37,1984

,, " \"1;: - - - - - >

f

,

Fig.

I~ :.-~

,

405 -

Female of Bacchus umber n.

claw of a specimen from Sesto D:

without dark teeth;

apophysis of front

transversal

mandibles tridentate, fore tarsal

(except for distal

bands;

(type):

tarsal

A: C:

chela;

tarsus with a distal 6,1

enlarged segments;

ovoidal,

large;

tooth between the two posterior apophysis

segment 1-4 in following proportions:

apophysis):

B:

fore tarsal

segment 3.

pterostigma brown,

with a rudimentary

segment 3 of front

segment;

sp.

Fiorentino (Italy);

longer than 8:

; segment 5 of front tarsus

1,5:

1,2

(Fig.

405)

Bocchinae

with one preapical

lamella and 2 bristles;

row of 7-8 teeth and 2 bristles; MALE:

617

tibial

enlarged claw

(Fig.

405)

with a

spurs 1,1,2.

unknown

LOCUS TYP ICUS:

Bivio Castel

TYPICAL MATERIAL:

Rigone

holotype F!

(I taly)

and 1 paratype F!

in OL;

1 para type F!

in MI. HOSTS:

unknown

DISTRIBUTION: OL!

Palaearctic region:

Sesto Fiorentino (Firenze),

ITALY:

Bivio Castel

Rigone

(Perugia),

M I!

3. Bocchus szelenyii Moczar

Bocchus szelenyii Moczar 1974: 131. Bocchus szelenyii Moczar: Moczar 1983b: 206.

FEMALE:

fully

winged;

Fig.

406 -

length 3,12 mm;

head black,

with mandibles yellow;

Chela of Bocchus szelenyii Moczar (type)

618

Mem.

antennae testaceous;

black;

abdomen black; OOL = 7;

visible,

pronotum

Inst.,

scutellum shiny,

with one transversal

mandibles quadridentate, posterior teeth; 10;

punctate;

fore tarsal

enlarged claw

segment 5 of front

with some

hairy;

POL

notaulices weak,

metanotum dull,

pterostigma short and narrow;

rudimentary

406)

with a

(Fig.

406)

but

reticulate

tooth between

the two

segments 1-4 in following proportions:

(Fig.

tarsus

swollen,

carina complete;

propodeum reticulate rugose;

dark band;

with a

scutellum

petiole reddish;

near posterior margin of scutum;

not much shorter than scutellum;

fore-wing

hairy,

occipital

weakly punctate,

weakly

1984

reddish-testaceousj

head shiny,

obsolete only

smooth,

37,

propodeum black;

without sculpture;

scutum shiny,

no.

and scutum

legs testaceous;

almost complete,

rugose,

3:

En!.

metanotum reddish-testaceous;

short keels on frons, 5;

Amer.

14:

2:

row of 5 teeth and 1 long bristle;

with one preapical

lamella and 2

long

bristles; tibial spurs 1,1,1. MALE: unknown LOCUS TYP ICUS: Spring "Talyn Bi Igech" TYPICAL MATERIAL:

holotype F!

(Mongol ia)

in BU.

HOSTS: unknown DISTRIBUTION: Palaearctic region: Km E guard-house "Caganbu lag",

MONGOLIA:

Spring

"Talyn Bilgech"

Bajanchongor a imak),

(47

BU!

4. Bocchus slovacus Strejcek

Bocchus slovacus Strejcek 1964: 325. Bocchus slovacus Strejcek: Nagy 1969: 322. Bocchus slovacus Strejcek: Ponomarenko 1978: 23.

FEMALE:

fully

winged;

antennae testaceous;

length 2,48-2,75 mm;

thorax and

brown-reddish nuances; head dull, than

hairy,

twice as

long as POL;

tooth between

swollen;

twice as

rarely pronotum with

abdomen black;

occipital

scutum dull,

posteriorly separated;

long as metanotum;

narrow;

fore

407)

with a

tarsus

(Fig.

row of 6 teeth and 407)

with one apical

bristle; tibial spurs 1,1,1. MALE: fully winged; length 2,20-2,67 mm; testaceous;

tarsi

thickened,

hairy;

9:

9:

9:

8:

7:

and fore tibiae

10;

granulated,

antennal

black;

testaceous;

segments

head dull,

hairy;

granulated,

pterost i gma short

segments 1-4 in following proportions:

(Fig.

segment 5 of front

OOL more

rudimentary

metanotum and propodeum reticulate

fore-w i ngs wi th one transversa I dark band;

enlarged claw

with a

scutellum dull,

rugose;

tarsal

legs testaceous;

carina complete;

mandibles quadridentate,

the two posterior teeth;

notaulices complete, nearly

petiole testaceous;

granulated,

head reddish-testaceous;

propodeum black;

13:

3:

and

9;

1 long bristle; lamella and 2

long

mandibles brown-

antennae not distally

in following proportions:

granulated;

2:

12:

6:

8:

frons with a median complete

9:

619

Bocchinae

Fig.

keel;

=

8;

407 -

Che I a of

occipital carina complete; scutum dull,

granulated;

Bocchus slovacus Strejcek {type}

POL

=

6;

OL

=

3,5;

notaul ices complete,

OOL

=

7;

OPL

=

4;

TL

posteriorly separated;

scutellum and metanotum dull, granulated; propodeum reticulate rugose, with two longitudinal keels on posterior surface; areolae of propodeum partly broader than hyaline,

tegulae;

without dark

lanceolate;

radial

LOCUS TYPICUS:

vein curved;

Cenkov

TYPICAL MATERIAL:

abdomen

transversal

with petiole very short;

bands;

radial

pterostigma

cell

open;

long

tibial

and

spurs

fore wings narrow,

not

1,1,2.

{Czechoslovakia}

holotype F!

in PG;

3 paratypes FF

in Strejcek's

collection. HOSTS: unknown DISTRIBUTION:

Palaearctic region:

U. S. S. R.: Papanga {Russian

Romania}, PK! HUNGARY: Keleshalom, BU! CZECHOSLOVAKIA: Cenkov, PG!

BU!

Topart

{Kelebia},

BU!

Budapest,

620

Mem.

Fig.

408 -

Amer.

Chela of

En!.

Inst.,

no.

37,

1984

Bocchus paglianoi n. sp. (type)

5. Bocchus paglianoi n. sp.

FEMALE:

fully

winged;

antennae brown, black;

length 3,75 mm;

head black,

with segments 1-3 testaceous;

petiole testaceous;

abdomen black;

with mandibles yellow;

thorax and propodeum

legs testaceous,

with mid and

Bocchinae

hind coxae partly occipital

brown;

head swollen,

carina complete;

tooth between

hairy,

shiny,

mandibles quadridentate,

the two posterior teeth;

notaulices complete,

621

scutum dull,

posteriorly separated;

with posterior third rugose;

segments 1-4 in following proportions:

claw

(Fig.

408)

front

tarsus

tibial

spurs 1,1,1.

MALE:

unknown

LOCUS TYPICUS:

15:

3: 10; enlarged

2:

teeth and one long bristle;

with one preapical

Montemitro

TYPICAL MATERIAL: HOSTS:

row of 4

408)

granulated;

pterost i gma short and narrow;

fore tarsal

(Fig.

hairy,

scutellum shiny and smooth,

metanotum and propodeum reticulate rugose;

fore-wi ng with 2 transversa I dark bands; with a

granulated;

with a rudimentary

lamella and 2

segment 5 of

long bristles;

(Italy)

holotype F!

in OL.

unknown

DISTRIBUTION: NOTES:

Palaearctic region:

the species

is named

in

ITALY:

Montemitro (Campobasso),

honor of Mr.

OL!

Guido Pagliano.

6. Bocchus scaramozzinoi n. sp.

FEMALE:

fully

testaceous,

winged;

length 3,6-4,2 mm;

anterior part of frons,

head black,

with mandibles

temples and region between ocelli

and

occipital margin reddish; ventral part of head partly black and reddish; sometimes head fully reddish, except for ocellar triangle black; antennae brown,

with segments 1-2 and partly 3 testaceous;

scutum and scutellum reddish; mesopleura, tegulae basal

propectus,

testaceous;

segments

sometimes

metapleura and posterior surface of propodeum reddish; petiole testaceous;

abdomen black;

legs testaceous,

part of hind coxae and clubs of hind femora brown;

shiny,

pronotum,

metanotum and propodeum black;

in following proportions:

almost

hairless,

2 keels around orbits;

15:

8:

with a median

14:

12:

11: 9: 9: 9: 9:

longitudinal

keel

with

antennal 12;

head

on frons and with

head with numerous irregular keels among which

the surface is granulated;

only a central

region of frons

is smooth and

without areolae, keels or sculpture; occipital carina complete; ocellar triangle convex, nearly equilateral; OOL = 12; OPL = 9; TL = 12; mandibles quadridentate, shiny, keels;

with a strong pronotal

with one rudimentary

transversal

impression

tubercles reaching

tegulae;

numerous areolae or irregular keels; separated; rugose; surface;

scutellum with numerous

propodeum reticulate rugose,

pronotum hairy,

scutum shiny,

notaul ices complete, longitudinal with 2

keels;

one on basal

long and narrow,

20:

3:

4:

prox i ma I 13:

19;

with

posterior keels on posterior fore-wing

with

with 2 dark transversal

cells and another beneath the pterostigma;

part of rad i a I ve in shorter than following proportions:

irregular

hairy,

metanotum reticulate

longitudinal

median area of posterior surface almost smooth;

pterostigma very reduced, bands,

tooth;

and with numerous

(5: 7);

distal

fore tarsa I segmen ts

enlarged claw

(Fig.

409)

in

with a

622

Mem.

Fig.

Amer.

En!.

Ins!.,

no.

37,

1984

Chela of Bocchus scaramo2zinoi n.

409 -

row of 6 teeth and 2 bristles; bristles and one preapical

segment 5 of front

lamella;

tibial

tarsus

(Fig.

409)

with 2

in OL;

1 paratype F!

unknown

DISTRIBUTION:

Palaearctic region:

Sardinia), OL!

Musei

NOTES:

(type)

spurs 1,1,1.

MALE: unknown LOCUS TYPICUS: Tempio Pausania (Italy) TYPICAL MATERIAL: holotype F! and 1 paratype F! in PL. HOSTS:

sp.

the species

(Cagliari),

is named

ITALY: OL!

Tempio Pausania

(Sassari,

PL!

in honor of Mr.

Pier Luigi Scaramozzino.

7. Bocchus scobiolae Nagy

Bocchus scobiolae Nagy 1967b: 334. Bocchus scobiolae Nagy 1968: 1029.

Bocchinae

Fig. 410 -

623

Chela of Bocchus europaeus Bernard from Saint-Paul-et-Valmalle (France)

624

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

Bocchus scobiolae Nagy 1969: 322, 323.

I

haven't seen the three only known specimens of this species

females and 1 male).

I have included

Palaearctic Bacchus only on

the basis of the original

typicus: Agigea (Dobroudja, Romania).

(2

the species in the key to the

Locus

description.

I don't know where is the holotype.

8. Bacchus europaeus (Bernard)

Eukoebeleia europaea Bernard 1939: 109. Bacchus europaeus (Bernard): Strejcek 1964: 325. Bacchus europaeus (Bernard): Nagy 1969: 322. Bacchus europaea (Bernard): Abdul-Nour 1976: 277.

FEMALE: clypeus

fully

winged;

testaceous;

reddish;

pronotum and scutum reddish;

propodeum black; legs yellow; 7:

5:

6:

Bernard

5:

abdomen black,

antenna I segment

1939);

head smooth, along

sides of median keel 2 or 3 teeth;

rugose; rugose,

scutellum,

weakly brown; 1 without

hairy,

internal

weakly

occipital

metanotum and tegulae yellow-brown;

proportions: teeth

10: 5,5:

(with teeth,

granulated,

margin of eyes,

of frons;

with mandibles and

with segments 1-4 yellow-

10:

OOl more than

reticulate rugose;

carina,

long as POL;

longitudinal

keels;

on

mandibles with 1,

notaul ices distinct and complete,

scutellum with numerous

7: to

with some areolae

near occipital

carina complete;

twice as

7:

according

tridentate mandibles with a rudimentary tooth between

two posterior teeth; separated;

head black,

antenna I segments in following 7;

near clypeus,

shiny,

length 3,25 mm;

antennae reddish-brown,

the

scutum hairy, posteriorly

metanotum reticulate

scutellum nearly twice as long as metanotum; propodeum reticulate with 2 longitudinal keels on posterior surface; fore-wing with 2

dark transversa I bands;

pterost i gma short and narrow;

fore tarsa I

segments 1-4 in following proportions: 14: 2: 3: 10; segment 5 of front tarsus (Figs 410, 411) with a preapical lamella and 2 bristles; enlarged claw 1,1,1. MALE:

(Figs 410,

411)

with a row of 4-5

teeth and 1-2 bristles;

tibial

unknown

lOCUS TYPICUS:

St-Aygulf

TYPICAL MATERIAL:

(France)

holotype F!

in P.

HOSTS: in France Caliscelis sp. (Abdul-No'Jr 1976). DISTRIBUTION: Palaearctic region: FRANCE: St-Aygulf Paul-et-Valmalle (Herault),

(Var),

P!

9. Bacchus parthicus Ponomarenko

P!

Saint-

spurs

Bocchinae

B

625

A / /

/

/

/

Fig.

411

-

Bacchus eurapaeus Bernard (type): A: chela; B: fare tarsal segments.

Bacchus parthicus Ponomarenko 1979b: 166.

FEMALE:

fully

winged;

length 3,60 mm;

fully

reddish,

triangle brown,

posterior margin of scutellum brown,

abdomen brown;

antennae brown,

with occipital propodeum darkened,

with segments 1-2 yellow;

antennae

626

Mem.

Fig.

412 -

Amer.

Ent.

Ins!.,

no.

37,

1984

Chela of Bocchus parthicus Ponomarenko

(type)

weakly distally thickened; antenna I segments in following proportions: 10: 20: 15: 15: 15: 14: 13: 13: 15; mandibles quadridentate, with a rudimentary

tooth between

the two posterior teeth;

head shiny,

20:

punctate;

Bocch i nae

frons with a occipital

median complete keel

carina complete;

pronotum shiny, reaching

hairy,

tegulae;

and

= 7;

POL

smooth,

627

with 2 keels around orbits;

= 7;

OL

=

OOL

weakly punctate;

pronotum crossed by

13;

a transversal

=

OPL

pronotal

16;

TL

=

18;

tubercles

impression;

scutum

shiny, hairy, weakly punctate, with notaulices complete and posteriorly separated; scutellum and metanotum shiny, smooth, weakly punctate; propodeum reticulate rugose, strongly one on long

the basal

and

tarsal

(Fig.

1,1,2. MALE:

with 2 brown

fore-wing

cells and another beneath

narrow,

segments

front tarsus claw

with posterior surface not reticulate rugose,

transversely striate; not

lanceolate;

radial

vein

in following proportions:

(Fig.

412)

412)

with a

regularly curvilinear;

lamella and 2 bristles;

with a row of 6 teeth and

bands,

pterostigma fore

5: 8: 25: 34; segment 5 of

33:

preapical

transversal

the pterostigma;

1 long bristle;

tibial

enlarged spurs

unknown

LOCUS TYPICUS:

Tashauz

TYPICAL MATERIAL: HOSTS:

(Turkmenistan,

holotype F!

U.

S.

S.

S.

R.:

R.)

in LN.

unknown

DISTRIBUTION:

Palaearctic region:

U.

S.

Tashauz

(Turkmenistan),

LN!

ETHIOPIAN SPECIES OF

BaCCHUS

Three species are known

KEY TO THE SPECIES

FEMALES

Micropterous;

fore-wing

nearly

as

long as

• • . . • . . . . . . . . . . • • • • . • • . . . • . • . . • . .. -

Fully

2

Segment

1 of

414 B);

antennae

least -

tegulae . . . . . . . . . . . . . . .

richardsi n. sp.

winged . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2

front

twice as

Segment

of

antennae nearly

1.

as

tarsus nearly

longer and more

long

front

shorter,

1,5

times as

slender,

as broad ••.•.• tarsus nearly as

2.

MALES

3.

4

segments

(Fig. 5-9 at

bini n. sp.

long as

distally more broadened,

long as broad •••••••••••

long as

wi th 4

(Fig.

with

415 A);

segments

seyrigi (Benoi t)

5-9

628

Mem.

Amer.

En!.

Inst.,

no.

37,

1984

Unknown

1. Bocchus richardsi n. sp.

FEMALE: 3,31

15: 6:

9:

areolae, lateral 10;

micropterous,

mm;

with

fore-wing nearly

fully reddish-dark; 6:

6:

6:

with a

5:

5:

median

5:

8;

keel

keels around orbits;

occipital

antennal

as

segments

head shiny,

long as tegulae;

length

in following proportions:

granulated and with

weak

joining anterior ocellus to clypeus and with 2 POL = 5;

carina complete;

OL = 3;

pronotum shiny,

OOL = 12; hairy,

OPL = 8;

weakly

TL =

granulated,

with a transversal impression between anterior and posterior collar; pronotal tubercles reaching tegulae; scutum dull, granulated, with weak areolae and strong

longitudinal

with

longitudinal

keels;

fully

reticulate rugose;

keels;

notaulices invisible;

metanotum very short and reduced; posterior surface without

scutellum dull, propodeum

longitudinal

keels;

fore

tarsal segments in following proportions: 15: 3: 3: 11: 14; enlarged claw (Fig. 413) with a row of 9 teeth and one bristle; segment 5 of front tarsus

(Fig.

413)

with a

preapical

lamella and a

row of 8 bristles;

spurs 1,1,1.

(\

Fig.

413 -

Chela of

Bocchus richardsi n. sp. (type)

tibial

Bocchinae

MALE:

unknown

LOCUS TYPICUS:

Milnerton

TYPICAL MATERIAL: HOSTS:

(South Africa)

holotype F!

in BM;

1 paratype F!

in WA.

unknown

DISTRIBUTlmJ: BM!

629

Ethiopian region:

Cape Town,

NOTES:

SOUTH AFRICA:

Milnerton

(Cape Town),

WAr

the species

is named

in

honor of Mr.

Owain W.

Richards.

2. Bocchus bini n. sp.

FEMALE: black

fully

winged;

length 3,12-4,06 mm;

head fully reddish or fully

(with mandibles and sometimes also clypeus testaceous);

occasionally

head black with region between ocell i and occipital carina reddish; thorax fully reddish; propodeum fully reddish or fully black; occasionally partly black

and reddish;

antennae fully

testaceous or brown,

legs testaceous;

reticulate rugose;

sometimes not reticulate rugose on frons on sides of

median keel,

where the surface is granulated;

ocellar triangle equilateral; rugose,

abdomen black;

with segments 1-2

or 1-3 testaceous;

hairy,

shiny;

pronotum hairy;

notaulices

head du II,

occipital

ha i ry,

fu II y

carina complete;

scutum fully reticulate

invisible or slightly visible;

scutellum

and metanotum reticulate rugose; propodeum reticulate rugose, with 2 longitudinal keels on posterior surface; fore-wing with 2 dark transversal bands; pterostigma short and narrow; fore tarsal segments 1-4 in following proportions: preapical

13: 2:

2:

8;

5-6 teeth and one bristle; MALE:

segment 5 of front

lamella and 2 bristles; tibial

tarsus

enlarged claw

(Fig.

(Fig.

414)

414)

with one

with a row of

spurs 1,1,1.

unknpwn

LOCUS TYP ICUS:

Afgoi

TYPICAL MATERIAL: in BM;

(Somal ia)

holotype F!

1 para type F!

and 3 para types FF!

in OL;

1 paratype F!

in OT.

HOSTS: unknown DISTRIBUTION: Ethiopian region:

SOMALIA:

Afgoi

(Benadir),

OL!

Ibadan, OT! Ikenne (\'I-State), BM! GHANA: Kwadaso (Kumasi), NOTES: the species is named in honor of Mr. Ferdinando Bin.

NIGERIA: OL!

3. Bocchus seyrigi (Benoit) n. comb.

Hymenodryinus seyrigi Benoi t 1953b: 427.

FEMALE:

fully

antennal

segments 3-10 brown,

winged;

reticulate rugose,

length 4,62 mm;

hairy;

reddish,

with ocellar spot brown,

abdomen black-brown;

antennae short,

head dull,

distally broadened,

fully

with

630

Mem.

Amer.

Ent.

Ins!.,

no.

37,

1984

, >

'

\ !

,

'

~

'

,

'0\ \\ \

\

B

"

\

\

Fig.

414 -

Bocchus bini n. sp. B:

segments 5-9 nearly proportions:

18:

7:

as 15:

(paratype from

fore tarsal

long as broad; 7:

5,5:

5:

6:

6:

Ibadan,

Nigeria);

A:

chela;

segments.

antenna I segments 5:

7;

occipital

in following

carina complete;

Bocchinae

Fig.

415 -

Bocchus seyrigi

(Benoit) (type);

631

A:

fore tarsal

segments;

B:

chela.

ocellar triangle equilateral; fully

reticulate rugose;

with 2

longitudinal

scutum,

scutellum and metanotum dull,

notaulices invisible;

hairy,

propodeum reticulate rugose,

keels on posterior surface;

fore-wing

with 2 dark

632

Mem.

transversa I bands;

Amer.

12: 2:

LOCUS TYPICUS:

3:

12;

37,

Behara

1984

fore tarsa I segments 1-4

segment 5 of front

teeth and one bristle;

TYPICAL MATERIAL: HOSTS:

no.

lamella and 2 bristles;

with one preapical

with a row of 11 MALE: unknown

Inst.,

pterost i gma short and narrow;

in following proportions: 415)

En!.

tibial

tarsus

enlarged claw

(Fig.

(Fig.

415)

spurs 1,1,1.

(Madagascar)

holotype F!

1 paratype F in TV.

in P;

unknown

DISTRIBUTION: 1953b) •

Ethiopian region:

MADAGASCAR:

Behara,

P!

3ekily

(Benoit

OR I ENTAL SPEC I ES OF BOCC/JUS

Four species are known.

KEY TO THE SPECIES

FEMALES

-

Fore-wing with Fore-wing with

2 Fore-wing of A);

front

with

1 dark

transversal transversal

band •••••

2 dark

radial

vein

curved

tarsus without

enlarged

weakly

a distinct

claw with a

row of

thorax and propodeum black

some

(except

(Fig.

proximal teeth

for

Fore-wing with segment

5 of

radial

front

(Figs 419,422); 422);

head,

vein more

tarsus

enlarged

wi th

strongly a

416 A);

prominence (Fig.

2.

418);

(Fig.

di st inct proximal

claw with

segment (Fig.

5

416

head,

muluensis n.

curved

1 only

thorax and propodeum reddish

tooth

sp.

416 B);

prominence

(Figs

419,

or brown-testaceous

••..••.••••••••••.•••...••••....•.....•........•..........•..•• 3 much longer than POL; head, thorax and propodeum redddish-brown •• •••••••••••••••••......•..• 3. pedunculatus Nagy OL more than twi ce as long as POL; head, thorax and propodeum reddi sh-I ight .••.•••••••.••••••••.•••..•••• 4. rubricus (Fouts)

3 OL not -

laotianus n. sp.

the yellow mandibles) .•••

. . . • • . • • . . . . • . . . • • • • • • • . • . • . . • • • • . . . . . . . . .. -

1.

bands • • • . . • . . . . . . . . . . . • . . . . . • 2

fAALES

Only

the male of B.

pedunculatus Nagy is known.

Bacchinae

633

A

,

,

E

Radial veins of females of Bacchus muluensis n. sp. (type)(A), pedunculatus Nagy from Tagembung (Phi I ippines) (B), laatianus n. sp. (type)(C), guineensis n. sp. (type)(D), australiae n. sp. (type)(E).

Fig. 416 -

634

Mem.

Amer.

Ent.

lnst.,

no.

37,

1984

i

i~

() /

/J 1/ I /

Fig. 417 -

Chela

0

f Bocchus laotianus n.

sp.

( type)

i

Bocchinae

635

1. Bocchus laotian us n. sp.

FEMALE:

fully

winged;

length 3,43-3,87 mm;

with segments 1-2 or 1-3 yellow; legs yellow;

head dull,

hairy,

thorax,

weakly granulated,

except for the sides of the ocelli rugose;

occipital

tooth between

equilateral;

scutum shiny,

granulated,

scutellum shiny,

punctate,

twice as

with

1 dark

transversal

posteriorly separated;

reticulate rugose only near posterior margin,

long as metanotum; with 2

ocellar triangle

reticulate rugose near

notaulices distinct and complete,

nearly

with a

mandibles quadridentate,

anterior margin;

reticulate rugose,

reticulate rugose,

the two posterior teeth; hairy,

antennae brown,

where the surface is not reticulate

carina complete;

rudimentary

head black;

propodeum and abdomen black;

metanotum reticulate rugose;

longitudinal band;

propodeum

keels on posterior surface;

pterostigma short and narrow;

fore-wing

radial

vein

weakly curved (Fig. 416 C); fore tarsal segments 1-4 in following proportions: 20: 3: 3: 14; segment 5 of front tarsus (Fig. 417) with a preapical

lamella and 2 bristles;

10 teeth and MALE:

1 bristle;

enlarged claw

(Fig.

417)

with a

row of

spurs 1,1,1.

unknown

LOCUS TYPICUS:

Ban Van Eue

TYPICAL MATERIAL: HOSTS:

tibial

(Laos)

holotype F!

1 para type F!

in B;

in OL.

unknown

DISTRIBUTION:

Oriental

region:

LAOS:

Ban Van Eue

(Vientiane),

B!

OL!

2. Hocchus muluensis n. sp.

FEMALE:

fully

winged;

segments 9-10 brown; legs testaceous,

length 3,25 mm;

head black;

thorax and propodeum black;

with hind coxae brown;

head dull,

antennae yellow, abdomen hairy,

with

black-brown; reticulate

rugose; frons narrow, very converging towards the clypeus; in front of the anterior ocellus the frons is nearly as broad as each eye;ocellar triangle equilateral;

occipital carina complete;

mandibles with 4 teeth,

larger from

anterior one to posterior;

scutum reticulate rugose,

and with numerous metanotum hairy, 2

longitudinal

transversal short and 3:

12;

bristle;

longitudinal

dull,

keels;

notaul ices invisible;

weakly punctate;

keels on posterior surface;

bands; narrow;

radial

vein

fore tarsal

segment 5 of front enlarged claw

propodeum reticulate rugose, fore-wing

weakly curved

(Fig.

(Fig.

418)

418)

with a

with a

Gunong Mulu National

TYPICAL MATERIAL:

holotype F!

and

416 A); preapical

row of 7 teeth and

Park

(Sarawak,

1 paratype F!

hairy with

with 2 dark pterostigma

1 long

Malaysia)

in 8M.

14:

lamella and

bristle; tibial spurs 1,1,2. MALE: unknown LOCUS TYPICUS:

dull,

scutellum and

segments 1-4 in following proportions:

tarsus

(Fig.

progressing

3: 1

636

Mem.

Fig.

HOSTS:

418 -

Amer.

Ent.

Inst.,

no.

37,

1984

Chela of Bocchus muluensis n.

sp.

(type)

unknown

DISTRIBUTION: District,

Oriental

Sarawak),

region:

MALAYSIA:

Gunong Mulu Nat.

Park

(Baram

BM!

3. Bocchus pedunculatus Nagy

Bocchus pedunculatus Nagy 1969: 323. Bocchus rubricus Fouts 1922: Nagy 1969: 324.

FEMALE: fully winged; length 2,18-3,20 mm; reddish-brown, with antennae yellow (with segments 9-10 or only 10 brown); head shiny, smooth, weakly punctate,

sometimes with

complete;

mandibles quadridentate,

two posterior teeth; weakly

punctate,

and distinct,

transversal

OL not much

keels on frons;

longer than

POL;

carina

tooth between

scutum shiny,

without sculpture among punctures;

posteriorly separated;

occipital

with a rudimentary

the

smooth,

notaulices complete

scutellum and metanotum shiny,

smooth, weakly punctate; propodeum reticulate rugose, with 2 lateral points and with 2 longitudinal keels on posterior surface; fore-wing with 2

Bocchinae

Fig.

419 -

637

Chela of Bocchus pedunculatus Nagy from Gunong Mulu (Sarawak)

dark transversal strongly curved proportions:

bands; (Fig.

13: 3:

3:

pterostigma short and narrow;

416 B);

fore tarsal

radial

vein

segments 1-4 in following

11; segment 5 of front tarsus (Fig. 419) with a

638

Mem. Amer.

Fig.

420 -

Ent.

Inst.,

no.

37,

1984

Male of Bocchus pedunculatus Nagy from Gunong Mulu

(Sarawak): A:

ninth abdominal sternite;

B:

genitalia.

preapical lamella, 2 bristles and a proximal distinct prominence; enlarged claw (Fig. 419) with 1 tooth and 1 long bristle; tibial spurs 1,1,2. MALE (Figs 402, 403): fully winged; length 2,60-3,00 mm; head black, with mandibles and sometimes clypeus testaceous; antennae yellow, with segments 6-10 brown; thorax, propodeum and petiole black or brpwn;

legs

testaceous, with coxae blackish; head hairy, dull, fully reticulate rugose; occipital carina complete; ocellar triangle equilateral OPL = OOL; mandibles quadridentate, wite. a rudimentary tooth between the two posterior teeth; scutum dull, hairy, fully reticulate rugose; notaulices distinct and complete, posteriorly separated; scutellum and metanoturn dull, weakly punctate, hairy; propodeum reticulate rugose, with 2 longitudinal keels on posterior surface and with 2 lateral points (as in female); forewing with 2 transversal darkened bands; radial vein strongly curved; pterostigma short and narrow, lanceolate, 420; abdomen with petiole very long. LOCUS TYPICUS: Pinigisan (Philippines) TYPICAL MATERIAL: holotype M! in CO. HOSTS: unknown

hyaline;

male genitalia:

fig.

DISTRIBUTION: Oriental region: MALAYSIA: Gunong Mulu (Baram District, Sarawak), OU BM! PHILIPPINES: Pinigisan (Mantalingajan, Palawan), CO! Tagembung (Mantalingajan, Palawan)(Nagy 1969: under name Bocchus rubricus Fouts), CO!

Bocchinae

Fig. 421

-

Female of

639

Bocchus rubricus (Fouts)(type)

4. Bocchus rubricus (Fouts)

Neoanteon rubrica Fouts 1922: 634. Neoanteon rubrica Fouts: Baltazar 1966: 205. nec Bocchus rubricus (Fouts): Nagy 1969: 324 (see Bocchus pedunculatus Nagy).

FEMALE

(Fig. 421):

fully winged;

length 3,0-4,0 mm;

reddish,

with

antenna I segments 1, 2, 3, 7, 8 and part of 9 yellow; head almost hairless, shiny, smooth, without sculpture, weakly punctate; OL more than twice as long as POL; occipital carina complete; mandibles quadridentate, with a rudimentary tooth between the two posterior teeth; scutum weakly punctate, almost hairless, smooth, shiny; notaulices weak, incomplete, not visible in posterior third of scutum; scutellum and metanotum smooth, shiny,

weakly punctate;

propodeum reticulate rugose,

with 2 longitudinal

640

Fig.

Mem.

422 -

Amer.

Ent.

Inst.,

Female of Hocchus rubricus tarsal

no.

(Fouts)(type):

strongly curved; 3:

12;

bands;

fore tarsal

segment 5 of front

Mt.

points on sides;

tibial

Makiling

(Fig.

422)

prominence;

with a preapical

unknown

fore

fore-wing radial

enlarged claw

lamella, (Fig.

(Philippines)(Los Banos according in WA.

vein

15:

422)

spurs 1,1,1.

original description; Mt. Makiling on type label). TYPICAL MATERIAL: holotype F! and 1 paratype F! HOSTS:

B:

segments 1-4 in following proportions:

bristles and a distinct proximal

LOCUS TYPICUS:

chela;

pterostigma short and narrow;

tarsus

1 tooth and 1 long bristle; MALE: unknown

A:

segments.

keels on posterior surface and with 2 lateral with 2 dark transversal

37,1984

to the

4: 2 with

641

Bocchinae

DISTRIBUTION: Los Banos

Oriental

(Luzon),

region:

PHILIPPINES:

Mt.

Makiling

(Luzon),

WA!

WA!

NEARCTIC SPECIES OF

BaCCHUS

Eight species are known

KEY TO THE SPECIES

FEMALES

Species micropterous -

Species

fully

(Figs

424,

425) .•••••••••••••••••••••••••• 2

winged . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3

2 Fore-wing less reduced, reaching approximately the keel between anterior and posterior surface of propodeum; notaul ices complete, sometimes slightly visible •••••••••••••••••••••••••••• 1. mirabilis (R.C.L. Perkins - Fore-wing more reduced, reaching approximately anterior margin of propodeum (Figs 424,425); notaulices invisible . . . . . . . . . . . . . . . • • . . . • . • • • . . . . . . . • . . . . • . .. 2. testaceus n. sp. 3 Species fully testaceous . . . 3. wasbaueri n. sp. - Species at least partly black . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 4 Thorax partly black . . . . . . . . 6. flavicollis Ashmead - Thorax fully black ••••••••••••••••••••••••••••••••••••••••••••• 5 5 Scutum dull, granulated . . . . 4. arizonicus (R.C.L. Perkins) - Scutum shiny, weakly punctate, wi thout sculpture among punctures . . . • . . . . . . . . . . . . . . . . . . • . . . . 5. flavipes Kieffer

The females of B.

laticeps (Ashmead) and dubius n. sp. are unknown

MALES

Scutum dull, -

fully granulaterJ;

head granulated,

rugose . . . . . . . . . . . . . . . . . . . . . 7. laticeps (Ashmead) Scutum shiny, without sculpture; head reticulate g ranu I at ed. • • • • • • • • • • • • • • ••

The males of

8.

not

reticulate

rugose and

dubius n. sp.

B. mirabilis (R.C.L. Perkins), testaceus n. sp.,

642

Fig.

Mem. Amer.

423 -

Ent.

Inst.,

no.

37,

Chela of Bocchus mirabilis (R.C.L. (Ohio)

1984

Perkins) from Columbus

wasbaueri n. sp. flavicollis Ashmead, arizonicus (R.C.L. Perkins) and flavipes Kieffer are unknown. Probab I y B. laticeps (Ashmead) and dub ius n. sp.

are the opposite sex of the other six species.

Bocchinae

643

1. Bocchus mirabilis (R.C.L. Perkins)

Eukoebeleia mirabilis R.C.L. Perkins 1905: 59. Eukoebeleia mirabilis R.C.L. Perkins: Kieffer 1914b: 47. Phorbas mirabilis (R.C.L. Perkins): Fenton 1918b: 261. Bocchus mirabilis (R.C.L. Perkins): Richards 1939: 189. Bocchus mirabilis (R.C.L. Perkins): Nagy 1969: 322.

FEMALE:

micropterous,

with fore-wing reduced and extending beyond the

keel

between anterior and posterior surface of propodeum;

mm;

usuallyfully testaceous with antenna I segments 4-10 black or brown;

occasionally head reddish fully black;

with brown ocellar spot;

occasionally antennae brown,

pronotum and scutum usually reddish, scutellum,

head dull,

rudimentary tooth between complete;

rarely head almost

with segments 1-2 or 1-3 yellow;

rarely

metanotum and propodeum black;

tegulae testaceous;

granulated;

almost fully black;

abdomen brown-black;

the two posterior teeth;

notaulices distinct and complete,

scutellum dull,

as sculptured as scutum;

scutellum nearly twice as

legs and

mandibles quadridentate, occipital

OOL nearly three-four times as long as POL;

granulated;

length 2,50-2,92

with a

carina

scutum dull,

posteriorly separated;

metanotum reticulate rugose;

long as metanotum;

propodeum reticulate rugose,

with 2 longitudinal keels on posterior surface; fore reduced wings without pterostigma; fore tarsal segments 1-4 in following proportions: 23: 4: 6: 17; segment 5 of front tarsus (Fig. 423) with a preapical lamella and 2 bristles; enlarged claw (Fig. 423) with a row of 6-7 teeth and 1 bristle; tibial

spurs 1,1,1.

MALE:

unknown

LOCUS TYPICUS:

Ohio

(U.

S. A.)

TYPICAL MATERIAL: lectotype F! and 1 para lectotype F! in B. HOSTS: in U. S. A. Bruchomorpha oculata Newm. (Issidae) (Fenton 1918b). DISTRIBUTION: Place (Ont.),

Nearctic region: Ol!

OT!

Ont.),

OT!

U. S. A.:

Refugio

Tex.),

BY!

Ol!

B!

Ohio,

CANADA:

Grenadier

I.

(Bee Co.,

Columbus

Foremost

Centre (St. Tex.),

(Ohio),

(Alta.),

Lawrence BY!

Waco

OT! Is.

Carleton Nat.

Park,

(Mc lennan Co.,

BM!

2. Bocchus testaceus n. sp.

FEMALE

(Figs 424,

425):

micropterous,

only anterior margin of propodeum;

with fore-wing reduced and reaching

length 2,01

mm;

fully

testaceous;

head

dull, granulated; occipital carina complete; mandibles quadridentate, with a rudimentary tooth between the two posterior teth; OOL nearly four times as long as POL;

scutum shiny,

scutellum shiny,

without sculpture,

weakly granulated;

notaulices invisible;

nearly twice as long as metanotum;

644

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

I 424 - Female of Bocchus testaceus n. sp.

Fig.

metanotum reticulate rugose;

propodeum reticulate rugose,

longitudinal

keels on posterior surface;

pterostigma;

fore tarsal

enlarged claw

with 2

fore reduced wings without

segments 1-4 in following proportions:

6; segment 5 of front tarsus (Fig. 426) with a preapical bristles;

(type)

(Fig.

426)

11:

1,5: 2:

lamella and 2

with a row of 8 teeth and 2 bristles;

tibial spurs 1,1,1. MALE: unknown LOCUS TYPICUS:

Nogales (U.

TYPICAL MATERIAL: HOSTS:

S.

holotype F!

A.) in OL.

unknown

DISTRIBUTION:

Nearctic region:

U. S. A.: Nogales (Arizona), OL!

3. Bocchus wasbaueri n. sp.

FEMALE:

fully

winged;

length 3,12 mm;

fully

testaceous,

ocellar spot and antenna I segments 4-10 darkened; occipital

carina complete;

tooth between

mandibles quadridentate,

the two posterior teeth;

with brown

head dull,

granulated;

with a rudimentary

OOL nearly five times as long as

Bocchinae

Fig.

POL;

425 -

scutum dull,

separated,

645

Female of Bocchus testaceus n.

granulated;

scutellum dull,

transversa I band;

2:

3:

8;

fore-wing

tibial

with a

dark

fore tarsa I segments 1-4

segment 5 of front

lamella and 1 bristle;

row of 7 teeth and 1 bristle; MALE:

propodeum reticulate rugose,

pterost i gma short and narrow; 14:

posteriorly

metanotum reticulate rugose;

keels on posterior surface;

in following proportions: with a preapical

(type)

notaulices complete and distinct,

granulated;

scutellum nearly twice as long as metanotum; with 2 longitudinal

sp.

enlarged claw

tarsus

(Fig.

(Fig.

427)

427)

with a

spurs 1,1,1.

unknown

LOCUS TYPICUS:

Uvalde (Texas,

TYPICAL MATERIAL: HOSTS:

U.

S.

A.)

in SC.

unknown

DISTRIBUTION: NOTES:

holotype F!

Nearctic region:

the species is named

U.

S.

A.:

Uvalde (Tex.),

in honor of Mr.

4. Bacchus arizonicus

Marius S.

(R.C.L.

SC!

Wasbauer.

Perkins)

Eukoebeleia arizonica R.C.L. Perkins 1907: 44. Eukoebeleia arizonica R.C.L. Perkins: Kieffer 1914a: 47. Piwrbas arizonica (R.C.L. Perkins): Fenton 1918b: 263. Bacchus arizonica (R.C.L. Perkins): Nagy 1969: 322.

FEMALE: yellow;

fully

winged;

length 3,11-3,31

antennae brown,

mm;

head black,

with mandibles

with segments 1-2 or 1-3 testaceous;

thorax and

propodeum black; legs and tegulae testaceous; head dull, granulated, numerous areolae near clypeus, along orbits, near occipital carina; occipital carina complete; tooth between

mandibles quadridentate,

the two posterior teeth;

with

with a rudimentary

OOL nearly three times as long as

646

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

, ,

,-, I

(

(

\

'-

\

\

\

\

\

\

\

\

..i

"

\

\:/.;I--~;~)

s:;:::..- --.. . 'S. I

.-//

Fig.

426 -

Female of

Bocchus testaceus segments;

POL;

scutum dull,

granulated;

B:

n.

sp.

(type):

A:

fore tarsal

chela.

notaulices distinct and complete,

posteriorly

separated; scutellum shiny, punctate, nearly twice as long as metanotum; metanotum reticulate rugose; propodeum reticulate rugose, with 2 longitudinal band;

keels on posterior surface;

pterostigma short and narrow;

proportions:

31:

5:

6:

18;

fore-wing

fore tarsal

segment 5 of front

with 1 dark transversal segments 1-4 in following

tarsus

(Fig. 428)

with a

Bocchinae

647

"'-"- .....

\ \

Fig. 427 -

Chela of Bacchus wasbaueri n. sp.

(type)

preap ica I I ameli a and 1-2 brist les; en I arged cl aw (F i g. 428) wi th a raw of 6-8 teeth and no or 1 bristle; tibial spurs 1,1,1. MALE: unknown LOCUS TYP ICUS: Nogales (Ariz., U. S. A.) TYPICAL MATERIAL: holotype F! in B. HOSTS: in U.S.A. Bruchamorpha sp. {Issidae} (Fenton 1918b). DISTRIBUTION: Nearctic region: U. S. A.: Nogales (Ariz.), B! Uvalde

648

Mem. Amer.

Fig.

428 -

Ent.

Inst.,

no. 37,

1984

Chela of Bocchus arizonicus

(R.C.L.

Perkins)

(Texas)

from Uvalde

Bocchinae

Fig. 429 -

Chela of

649

Bocchus flavipes Kieffer from Almanor (California)

650

Mem.

(Tex.),

Amer.

En!.

Ins!.,

no.

37,

1984

OL!

5. Bocchus flavipes Kieffer

Bocchus flavipes Kieffer 1906a: 237. Tetradryinus flavipes (Kieffer): Kieffer 1913c: 325. Tetradryinus flavipes (Kieffer): Kieffer 1914b: 48. Tetradryinus flavipes (Kieffer): Richards 1939: 189. Tetradryinus flavipes (Kieffer): Muesebeck and Walkley 1951: 1035. Tetradryinus flaviceps (Kieffer): Krombein 1979: 1243.

FEMALE:

fully

winged;

antennae brown, testaceous; proportions:

length 3,37-3,56 mm;

antennae distally thickened; 10: 8:

12:

11:

rugose and with numerous with two OL = 3; strong

lateral

black;

mandibles testaceous;

with segments 1-2 or 1-5 testaceous; 9:

8:

7,5:

OOL = 10;

6:

6:

irregular keels;

keels around orbits; POL = 10;

transversal

antennal

head dull,

carina complete;

pronotum shiny,

pronotal

legs

reticulate

frons with a median keel

occipital

TL = 10;

impression;

11;

abdomen brown;

segments in following

hairy,

tubercles reaching

and

POL = 3; with a

tegulae;

scutum

sh i ny, smooth, weak I y punctate, wi thoul scu I pture among punctures, weakly rugose near anterior margin; notaulices complete, posteriorly separated; scutellum shiny, smooth, weakly punctate, without sculpture among punctures, weakly rugose near posterior margin; metanotum dull, granulated, with some keels; propodeum with broad areolae; posterior surface of propodeum with

two longitudinal

keels;

among numerous broad areo I ae;

fore-wi ng

beneath

part of radial

part

the pterostigma;

(6:9);

fore tarsal

enlarged claw front

tarsus

(Fig.

(Fig.

distal

segments in following

429)

429)

with a

small

areolae are visible

wi th 1 dark transversa I band vein shorter than proportions:

20:

4:

row of 6 teeth and 1 bristle;

with a preapical

proximal 5:

17: 25;

segment 5 of

lamella and 8 bristles;

tibial

spurs 1,1,1. MALE: unknown LOCUS TYPICU5: Ormsby Co. (Nevada, U. TYPICAL MATERIAL: holotype F! in CA. HOSTS:

S.

A.)

unknown

DISTRIBUTION: Ormsby Co.

Nearctic region:

(Nev.),

U.S. A.: Almanor

{Plumas Co.,

Calif.),OT!

CA!

6. Bacchus flavicollis Ashmead

Bocchus flavicollis Ashmead 1893: 91. Bocchus flavicollis Ashmead: Kieffer in Kieffer et Marshall 1905: 67.

Bocchinae

Fig.

430 -

Chela of

651

Bocchus flavicollis Ashmead (type)

Bocchus flavicollis Ashmead: Kieffer 1914b: 45. Bocchus flavicollis Ashmead: Muesebeck and Walkley 1951: 1039. Bocchus flavicollis Ashmead: Krombei n 1979: 1243.

FEMALE:

fully

winged;

length 2,80-2,81

mm;

head reddish,

with a broad

brown spot on frons and vertex; mandibles and clypeus testaceous; prothorax reddish; scutum reddi sh or reddi sh-brown; scutell um, metanotum, pleura and propodeum black; testaceousj

sometimes scutellum partly reddish;

abdomen brown-testaceousj

legs testaceousj

petiole

antennae missing in

652

Mem.

holotype;

head dull,

rudimentary

flat,

complete;

separated;

12:

5:

12: 9:

scutellum smooth,

transversal

9:

8:

19: 3:

4:

1984

shiny,

10;

part;

carina

pronotum dull,

pronotal

tubercles

notaulices complete,

weakly granulated; fore-wing

distal

fore tarsal

12:

16; enlarged claw

lamella;

segment 5 of front tibial

frons with a

occipital

TL = 9;

impression;

very

with a

antenna I segments in

8:

OPL = 7;

granulated;

preapical lamella and 2 bristles; MALE: unknown

posteriorly

metanotum

with 1 dark

part of radial

vein

segments in following (Fig.

430)

tarsus

with a row of 7

(Fig.

430)

with a

spurs 1,1,1.

LOCUS TYPICUS: Marquette (Michigan, U. TYPICAL MATERIAL: holotype F! in WA. HOSTS:

9:

the pterostigma;

nearly as long as proximal teeth and 1 long

9:

propodeum reticulate rugose;

band beneath

proportions:

37,

keels around orbits;

transversal

scutum dull,

reticulate rugose;

no.

mandibles quadridentate,

OL = 2,5; OOL =7;

with a strong

tegulae;

Ins!.,

the two posterior teeth;

and with 2 lateral

POL = 2;

granulated, reaching

Ent.

granulated;

tooth between

following proportions: median keel

Amer.

S.

A.)

unknown

DISTRIBUTION:

Nearctic region:

Marquette (Mich.),

CANADA:

Nerepis (N.

B.),

u.

WA!

S.

A.:

WA!

7. Bocchus laticeps (Ashmead)

Phorbas Bocchus Bocchus Bocchus

FEMALE: MALE:

laticeps laticeps laticeps laticeps

Ashmead 1893: 90. (Ashmead):

Muesebeck and Walkley 1951:

(Ashmead):

Nagy 1969: 323.

(Ashmead):

Krombein

1979:

1040.

1243.

unknown

fully

testaceous;

winged;

length 2,32-2,50 mm;

antennae brown;

thorax,

head black,

propodeum,

with mandibles

pet iole and abdomen

black; legs black; tegulae yellow; head dull, almost hairless, granulated; antennal segments in following proportions: 11: 5: 7: 7: 7: 7: 7: 7: 7: 9; POL = 6;

Ol = 3;

OOl = 8;

OPl = 3;

Tl = 5;

notaulices longer than breadth of ocelli hairless; smooth,

notaulices complete, weak I y

punctate;

scutum dull,

posteriorly separated;

smooth and

propodeum reticulate rugose,

keels on posterior surface;

the

granulated,

scutellum shiny,

metanotum rugose on anterior ha If,

weakly punctate on posterior half; longitudinal

minimum distance between

(7:3);

median surface smooth,

with 2 without

sculpture, except for regions near keels which are rugose; internal surface of areolae rugose; fore-wing hyal ine, without dark transversal bands; pterostigma hyaline, oval, with distal genital ia:

part fig.

with only posterior margin brown;

longer than proximal

431;

tibial

spurs 1,1,2.

part;

radial

vein

abdomen with petiole short;

Bocchinae

Fig.

431

-

Bocchus laticeps (Ashmead}(type): A: ninth abdominal

Male of sternitej

LOCUS TYPICUS:

DISTRIBUTION: (Tex.), OLl

B:

genitalia

(right gonoforceps removed).

Jacksonville (Florida,

TYPICAL MATERIAL: HOSTS: unknown

653

holotype M!

Nearctic region:

U. S.

A.)

in WA. U. S. A.: Jacksonville

(Fla.),

WA!

Uvalde

8. Bocchus dubius n. sp.

FEMALE: MALE:

unknown

fully

wingedj

length 2,0 mmj

antennae not distally thickened; proportions:

10: 6:

8: 8:

7:

7,5:

black,

with antennae and

legs brownj

antenna I segments in following 7:

7:

7:

9;

head dull,

granulated,

654

Fig.

Mem.

432 -

Amer.

Male genitalia of

Ent.

Inst.,

no.

37,

Bocchus dubius n.

1984

sp.

(type)(right half

removed)

reticulate rugose;

frons with a complete median keel;

occipital

carina

complete; POL = 6; OL = 4; OOL = 7; OPL = 4; TL = 4; scutum shiny, without sculpture; notaul ices complete, posteriorly separated; minimum distance between the notaulices slightly

longer than

temples

(5:4);

655

Bocchinae

scutellum shiny,

without sculpture;

reticulate rugose; mostly granulated; distal fig.

fore-wing hyaline,

part of radial

432;

propodeum

without dark transversal

vein shorter than proximal

part

(6:8);

bands;

genitalia:

t i b i a I sp u rs 1, 1 , 2.

LOCUS TYPICUS:

Green River

TYPICAL MATERIAL: HOSTS:

metanotum reticulate rugose;

posterior surface of propodeum weakly reticulate rugose,

(Wyoming,

holotype M!

U. S.

A.)

in AM.

unknown

DISTRIBUTION:

Neotropic region:

U.

S.

A.:

Green River

AUSTRALIAN SPECIES OF

(Wyoming),

AM!

BOCCHUS

Five species are known.

KEY TO THE SPECIES

FEMALES

Segment

4 of

front

tarsus

shorter

than

segment

1 ••••••••••••••• 2

Segment 4 of front tarsus longer than segment 1 ••••••• •••• '" 2 Fore-wing wi thout dark transversal bands; radial vein less

-

curved

(Fig.

416 E);

scutum reticulate

•• 3

rugose; notaulices 1. australiae n. sp. - Fore-wing wi th 2 dark transversal bands; radial vein more strongly curved (Fig. 416 D); scutum smooth, wi thout sculpture; notaulices incomplete . . . . . . . . . . 2. guineensis n. sp. 3 Head granulated, reticulate rugose only behind ocelli, near occipi tal carina . . . . . . . . . . . . . . . 3. minimus n. sp. - Head fully reticulate rugose ••••••••••••••••••••••••••••••••••• 4 4 Enlarged claw and segment 5 of front tarsus regularly curved (Fig. 436); head, thorax and propodeum black (with mandibles testaceous) . . . . . . . . . • . . . . . . . . . . 4. robustus (Dodd) - Enlarged claw and segment 5 of front tarsus less curved, partly rect i I inear (Fig. 437); head, pronotum and scutum fully or partly reddish; scutellum, metanotum and propodeum black . . . . . . . . . . • . . • . . • . . . • . . • • . . • . • . . . . . . .. 5. bouceki n. sp. complete . . . . . . . . . . . . . . . . . . . . . . .

MALES

Only the male of

B. guineensis n. sp. is known.

Mem.

656

Amer.

Ent.

Inst.,

no.

37,

1984

1. Bacchus australiae n. sp.

FEMALE:

fully

winged;

length 3,62-3,87 mm;

yellow;

antennae brown dark or

thorax,

propodeum and abdomen black;

fully

reticulate rugose;

occipital

with 2-3 rudimentary teeth; reticulate rugose;

POL

weak

with 2

longitudinal

transversal

tarsal front

band;

(Fig.

433)

distinct prominence; long bristle;

tibial

MALE: unknown LOCUS TYPICUS:

=

5;

OL

weakly

=

4;

Mt.

TYPICAL MATERIAL:

oval,

proportions:

with 1 preapical

433)

hairy,

propodeum reticulate brown 3:

fore-wing (Fig. 12;

Tibrogargan

2 bristles and

1 paratype F!

in QU.

I

433 -

1 proximal

with a row of 4 teeth and 1

(f

Fig.

fore

segment 5 of

(Australia) in CB;

with 1

416 E);

spurs 1,1,2.

holotype F!

fully

posteriorly separated;

14: 3:

lamella,

(Fig.

ha i ry,

mandibles quadridentate,

scutum dull,

punctate;

pterostigma short,

enlarged claw

with mandibles head du II,

legs testaceous;

carina comlete;

keels on posterior surface;

segments 1-4 in following tarsus

head black,

with segments 1-4 testaceous;

notaul ices distinct and complete,

scutellum and metanotum dull, rugose,

light,

Chela of Bocchus australiae n.

sp.

(type)

Bocchinae

Fig. 434 -

657

Chela of Bacchus guineensis n.

HOSTS: unknown DISTRIBUTION: Australian region:

AUSTRALIA:

Mt.

sp.

(type)

Tibrogargan

658

Mem.

(Queensland),

CB!

Amer.

Woogaroo,

Ent.

Inst.,

37,

no.

1984

QU!

2. Boccus guineensis n. sp.

FEMALE:

fully

winged;

length 3,00-3,50 mm;

head black,

with mandibles

and sometimes clypeus yellow; antennae fully testaceous; thorax, propodeum and abdomen black; legs testaceous, with hind coxae black; head shiny or dull, some transversal

without sculpture or weakly granulated,

mandibles with 3 teeth;

occipital

smooth,

shiny,

only on

anterior third of scutum;

sculpture;

except for

weak keels on frons and some areolae behind ocell i; carina complete;

without sculpture hairless;

scutellum smooth,

metanotum weakly punctate;

POL = 2;

Ol = 3;

notaulices incomplete, shiny,

scutum

visible

without

propodeum reticulate rugose,

with 2

longitudinal keels on posterior surface; fore-wing with 2 dark transversal bands; pterostigma short and narrow; radial vein strongly curved (Fig.

416 D); fore tarsal segments 1-4 in following proportions: 12: 4: 3: 9; segment 5 of front tarsus (Fig. 434) with a preapical lamella, one proximal prominence and one bristle; enlarged claw (Fig. 434) with one teeth and one long bristle; tibial spurs 1,1,1. MALE: fully winged; length 2,00-2,62 mm; black; mandibles testaceous; antennae brown or wi th segments 1-2 testaceous; abdomen brown; legs brown or testaceous,

with coxae and fore clubs of femora brown;

antennae

antennal segments in following proportions: 9: 5: 6: 6: 8: 7: 7: 7: 6: 9; head dull, granulated; frons with a median keel; occipital carina complete; POL = 3; Ol = 2,5; OOl = 6,5; OPl = 4; Tl = not distally thickened;

2,5; scutum dull, granulated, with numerous

longitudinal

reaching approximately 0,5 shiny,

weakly granulated;

longitudinal

with numerous areolae on anterior half and

keels on posterior half; length of scutum;

notaulices incomplete,

scutellum and metanotum

propodeum reticulate rugose,

keels on posterior surface;

fore-wi ng wi th a dark transversal

with distal

lOCUS TYPICUS: Baiyer R. (New Guinea) TYPICAL MATERIAL: holotype F! and 7 paratypes MM! in B;

HOSTS:

1 paratype M!

in OT;

two

band beneath the pterost i gma;

pterostigma lanceolate; radial vein curvi I inear, than proximal part; tibial spurs 1,1,2.

F!

with

median area shiny and smooth;

4 paratypes

(1

M,

in TW;

3 FF)!

part shorter

1 paratype

in Ol.

unknown

DISTRIBUTION:

Australian region:

B!

TW!

Baiyer R.,

BRITAIN:

Kerrvat,

Jimmi

Valley,

NEW GUINEA: Ol!

Wau,

Ol!

Tifalmin

(NE New Guinea),

Kassam Pass,

OT!

3. Bocchus minimus n. sp.

Ol!

NEW

80cchinae

FEMALE:

fully

and mandibles

winged; light;

length 2,81

mm;

antennae fully

659

head reddish-brown,

testaceousj

with clypeus

thorax and propodeum

reddi sh, part I y darkened on scutum and propodeum; abdomen brown; iegs testaceous; head dull, granulated, reticulate rugose only behind ocelli, near occipital POL

~

4;

OL

~

anterior third;

carina; 3;

mandibles with 3 teeth;

scutum dull,

granulated,

notaulices distinct and complete,

scutellum and metanotum shiny, reticulate rugose, fore tarsal

band;

(Fig. (Fig.

435) 435)

carina complete;

posteriorly separated;

weakly punctate;

propodeum

keels on posterior surface; pterostigma short and oval,

segments 1-4 in following proportions:

tarsus

enlarged claw MALE:

smooth,

with 2 longitudinal

with 1 weak dark transversal of front

occipital

reticulate rugose only on

with a preapical

13: 2:

2:

14;

fore-wing brown; segment 5

lamella and 2 bristles;

with a row of 14 teeth;

tibial

spurs 1,1,2.

unknown

LOCUS TYPICUS: Spargoville (Australia) TYPICAL MATERIAL: holotype F! in 8M. HOSTS: unknown DISTRIBUTION: Australian region: Western Australia),

AUSTRALIA:

Spargoville

(Coolgardie,

BM!

L L

•.......•.....

Fig. 435 -

Chela of

~ ~~ ....•..•....

Bacchus minimus

n.

....

sp.

(type)

660

Mem. Amer.

Fig. 436 -

Ent.

Inst.,

Chela of Bocchus robustus

no.

(Dodd)

37,

1984

from Townsville

(Australia)

4. Bacchus robustus (Dodd) n. comb.

Anteonella robusta Dodd 1913: 182. Anteonella bicolor Dodd 1913: 182 (n. syn.); locus typicus Townsville (Australia).

Anteonella robusta Dodd: Kieffer 1914b: 213. Anteonella bicolor Dodd: Kieffer 1914b: 214. Dicondylus sulciscutellum Girault 1926: 135 (n. syn.); locus typicus Townsville

(Australia).

Bocchinae

FEMALE:

fully

testaceous;

winged;

length 4,87-5,12 mm;

antennae testaceous,

propodeum and abdomen black; clubs of femora black;

661

head black,

with mandibles

with segments 4-10 or 6-10 dark;

thorax,

legs fu Ily testaceous or wi th coxae and

head swollen,

dull shiny,

fully reticulate rugose,

with a median keel between the antennal sockets; ocellar region swollen; temples distinct; occipital carina complete; POL = OL; OOL = OPL; mandibles tridentate,

with an

intermediate rudimentary tooth;

pronotum

shiny, with a transversal impression, reticulate rugose and with transversal keels; pronotal tubercles reaching tegulae; scutum shiny, reticulate rugose; posteriorly

notaul ices complete and distinct,

separated;

scutellum shiny,

median furrow composed of areolae;

smooth,

fully

composed of areolae,

weakly punctate,

with a

metanotum shiny and smooth,

reticulate rugose;

propodeum fully reticulate rugose;

transversa I band;

pterost i gma broad and ova I,

fore-wing

brown;

laterally

with 1 dark

segment 3 of front

tarsus without distal apophysis longer than segment 3; segment 4 of front tarsus longer than segment 1; segment 5 of front tarsus (Fig. 436) with an apical

lamella and 1 long proximal

a row of 16 teeth and 3 proximal MALE: unknown LOCUS TYPICUS:

Townsville

bristle;

enlarged claw

bristles;

tibial

(fig.

436)

with

spurs 1,1,1.

(Australia)

of A. robusta in AD; holotype F! of A. bicolor in AD; holotype F! of D. sulciscutellum in QU. TYPICAL MATERIAL: HOSTS:

holotype F!

in Australia Fulgorids.

DISTRIBUTION: Australian region: AUSTRALIA: AD! Blunder Cr. (Brisbane, Queensland), UQ! Australia),

WA!

Sir Graham Moore

Island

Townsville (Queensland), Anjo Peninsula (Western

(Western Australia),

QU!

OL!

5. Bocchus bouceki n. sp.

FEMALE:

fully

reddish

winged;

in holotype;

length 3,37-4,87 mm;

head black

head,

pronotum and scutum

(with clypeus and mandibles

testaceous),

prothorax reddish, scutum black (with notaulices reddish) in paratype; antennae brown with segments 1-2 testaceous or fully testaceous; scutellum, metanotum,

propodeum and abdomen black;

hind coxae brown; = 4;

occipital

rudimentary rugose,

head hairy,

carina complete;

tooth between

hairy,

shiny;

shiny,

legs testaceous,

fully

sometimes with

reticulate rugose;

mandibles quadridentate,

the two posterior teeth;

POL = 5;

OL

with a

scutum fully reticulate

notaulices complete and distinct,

posteriorly

separated; scutellum and metanotum smooth, hairy, weakly punctate; propodeum reticulate rugose, with 2 longitudinal keels on posterior surface; brown;

fore-wing fore tarsal

segment 5 of front bristle; bristles;

with 1 dark transversal tarsus

enlarged claw tibial

band;

pterostigma short,

segments 1-4 in following proportions: (Fig.

(Fig.

spurs 1,1,2.

437)

417)

with a preapical

18:

3:

3:

oval, 26;

lamella and no

with a row of 17 teeth and without

662

Mem. Amer. Ent.

Ins!.,

no. 37,

1984

Fig. 437 - Chela of Bocchus bouceki n. sp.

MALE: unknown LOCUS TYPICUS: Mt. Tibrogargan

(Australia)

(type)

Bocchinae

TYPICAL MATERIAL:

holotype F!

in CB;

663

1 paratype F!

in AD.

HOSTS: unknown DISTRIBUTION: Australian region: AUSTRALIA: Mt. Tibrogargan (Queensland), CB! Landrigan Cliffs (North Western Australia), AD! NOTES:

the species

is named

in honor of Mr.

Zdenek Boucek.

4. Mirodryinus Ponomarenko

Mirodryinus Ponomarenko 1972: 673.

Fig. 438 -

Female of Mirodryinus xerophilus

(Benoit)

from Wassiya

(Egypt)

664

Mem. Amer.

Fig. 439 -

Ins!.,

no.

Female of Mirodryinus xerophilus

TYPE SPECIES: Mirodryinus FEMALE

En!.

(Figs 438, 439):

37,

1984

(Benoit) from Wassiya

(Egypt)

ungulatus Ponomarenko 1972, orig. desig.

fully winged;

maxi Ilary palpi

with 6 segments;

labial palpi with 3 segments; occipital carina complete; mandibles quadridentate, with a rudimentary tooth bet wen the two posterior teeth; segment 4 of front tarsus less than 0,5 times as long as segment 1 (Fig. 442); segment 5 of front tarsus without lamellae, with bristles or peg-like hairs (Fig. 442); enlarged claw without teeth, lamellae, bristles or peglike hairs (Fig. 442); fore-wing with radial vein distinct (though sometimes it is very short); pronotal tubercles reaching tegulae; tibial spurs 1,1,1. MALE (Figs 440, 441): fully winged; mandibles quadridentate, with a rudimentary tooth between the two posterior teeth; maxillary palpi with 6 segments; labial palpi with 3 segments; temples distinct; propodeum with areolae very small, less wide than tegulae (Fig. 440); tibial spurs 1,1,2. DISTRIBUTION: Palaearctic HOSTS: unknown SPECIES: 5

PALAEARCT I C SPEC I ES OF

MIRODRYINUS

Five species are known.

KEY TO THE SPECIES

Bocchinae

Fig.

665

Male of Mirodryinus xerophilus

440 -

(Benoit)

from

Ikingi

Mariout

(Egypt)

FEMALES

Fore-wing distal

with

parts;

radial

vein

curved,

with distinct

proximal

and

thorax and propodeum fully black . . . . . . . . . . • . . . . • .

1. gobiensis Ponomarenko -

Fore-wing wi th proximal pa r t I y

part

radial

vein

straight,

(Figs 438,439);

only wi th a distinct

thorax and propodeum fully

or

redd i sh •••••••••••••••••••••••••••••••••• ••' ••••••••••••• 2

2 Thorax and propodeum fully reddish . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . •••••••••••••••••••••••••••••• 2. xerophilus (Benoi t) -

Thorax and propodeum partly

3 Scutum and

or black • . . . . . . . . 3 ungulatus Ponomarenko

reddish and brown

scutellum brown . . . .

3.

666

Mem.

Fig.

441

-

Male of

Amer.

En!.

Ins!.,

37,

no.

1984

Mirodryinus xerophilus (Benoit) from Ikingi Mariout (Egypt)

- Scutum and scutellum reddish •••••• ••••••••••••••••••••••••••••• 4 4 Metapleura reddish; radial vein longer . . . . . . . . . . . . . . . . . . . . . . . . . . . • . . . . . . . . . . . . • . . . . . . . . . . . . . .. 4. atlanticus n. sp. - Metapleura black; radial vein shorter (Fig. 448 A) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. 5. ponomarenkoae n. sp.

MALES

Only

the male of M.

xerophilus is known.

1. Mirodryinus gobiensis Ponomarenko

Mirodryinus gobiensis Ponomarenko 1972: 676.

FEMALE:

fully

legs fully

winged;

testaceous;

length 2,62 mm;

head,

ocellar region brown;

mandibles,

antennae and

thorax and propodeum black;

abdomen black; antennae distally thickened; antennal segments in following proportions: 7: 3: 6: 5,5: 5: 5: 5: 4,5: 5: 5; head shiny, weakly

punctate,

or lateral 4;

keels;

Ol = 2,5;

without sculpture among punctures; temples very prominent;

OOl = 10; OPL = 4;

Tl = 9;

occipita!

frons without median

carina complete;

pronotum very short,

POL

weakly

Bocchinae

667

B

Fig.

442 -

Female of Mirodryinus gobiensis Ponomarenko (type):

B: fore tarsal segments.

A: chela;

668

Mem.

transversely striate;

pronotal

and metanotum shiny, punctures; scutum;

Amer.

Ent.

Inst.,

weakly punctate,

notaulices incomplete,

longitudinal

keels;

fore-wing

pterostigma;

radial

vein curved,

segments in following proportions: 442)

1 of front

LOCUS TYP ICUS:

DISTRIBUTION:

442);

and distal part

18: 29:

hollow

long

the

parts;

(2,5:4,5);

65;

segment 5

apex with a

lamella);

(antenna-cleaner)

fore

enlarged claw

bristles or other appendixes;

tibial

Bordson-Gobi

TYPICAL MATERIAL: HOSTS: unknown

15:

(among which one very

tarsus with a proximal

a big prominence (Fig. MALE: unknown

aymag,

teeth,

82:

or

bands beneath

with 2 rows of nearly 15 bristles;

group of nearly 5 lamellae

scutellum

without transversal

vein much shorter than proximal

without subapical (Fig.

dull,

with distinct proximal

part of radial

tarsus

scutum,

without sculpture among

with a dark transversal

tarsal

442)

1984

reaching approximately 0,3 length of

distal (Fig.

37,

tubercles reaching tegulae;

smooth,

propodeum fully reticulate rugose,

of front

no.

segment

del imitated by

spurs 1,1,1.

(Mongol i a)

holotype F!

in LN

Palaearctic region:

South-East Mongolia),

MONGOLIA:

Bordson-Gobi

(South Gobi

LN!

2. Mirodryinus xerophilus (Benoit) n. comb.

Anteon xerophilus Benoit 1951a: 301. Anteon xerophilum Benoit: Olmi and Currado 1979a: 346.

FEMALE

(Figs 438,

439):

fully

petiole and abdomen black; testaceous,

winged;

length 2,68 mm;

with segments 5-10 brown;

head shiny,

smooth,

with antennae

antennae distally thickened;

antenna I segments in following proportions: 4,5: 7;

reddish,

sometimes propodeum partl y brown; 6:

4:

strongly punctate,

11: 7:

5,5:

4,5:

4:

5:

without sculpture among

punctures; mandibles quadridentate, with a rudimentary tooth between the two posterior teeth; frons without medial keel; occipital carina complete; POL = 6,5; dorsal

OL = 4;

view;

punctate,

scutum,

OOL = 7;

with a

TL = 10; pronotum

without sculpture among punctures;

propodeum reticulate rugose, fore-wing

OPL = 8;

scutellum and metanotum shiny,

track of dark spot on the basal

proximal part, six times as

without the distal

long as segment 4,

part;

weakly

notaulices invisible;

without transversal

with a broad dark transversal

invisible in

smooth,

and

longitudinal

keels;

band beneath the pterostigma and cells;

radial

vein only

segment 1 of front

with a proximal

delimitated by a big prominence (Fig. 443);

hollow

with the

tarsus nearly

(antenna-cleaner)

enlarged claw

(Fig.

443)

without bristles or teeth or lamellae; segment 5 of front tarsus (Fig. 443) with a long row of small peg-like hairs; apex with a group of 4 lamellae; tibial

spurs 1,1,1.

669

Bocchinae

B

Fig. 443 -

Female of

Mirodryinus xerophilus (Benoit) from Egypt: A: chela; B:

fore tarsal

segments.

670

Mem. Amer.

Ent.

Ins!.,

no.

-1/

/

, \

II

\I

/

\

/ /

I /

I

II

I

II 1\

I

I

)

/

I

I

1/

/'

1984

I

\

/ \

37,

'\

I

1\

I I

I

I

\

I

I \ I \

/

I

1/ "-

I

I

\ I v

II

I / /

Fig. 444 - Male genitalia of Mil'odl'yinus xel'ophilus (Benoit)(type)

Bocch i nae

MALE

(Figs 440,

441),

fully

winged;

clypeus yellow and a small sockets and eyes; testaceous; scutellum,

671

length 2,5 mm;

head black,

reddish spot on region between

mandibles yellow,

with teeth brown;

antennae fully

pronotum and propectus reddi sh-testaceous or black; metanotum,

with

antennal

mesopleura and metapleura brown-black;

scutum propodeum

black; abdomen brown; legs fully testaceous; tegulae testaceous; antenna I segments in following proportions: 6: 2: 10: 9: 8: 7: 7: 6,5: 5: 7; head sh i ny,

swoll en,

punctures; complete;

smooth,

POL = 7;

wi th sparse punctures,

OL = 4;

scutum shiny,

smooth,

sculpture among punctures; approximately 0,3 smooth,

without sculpture, almost pointed;

propodeum dull,

hairy,

with sparse punctures,

notaulices distinct,

length of scutum;

convex,

wi thout scu I pture among

= 8; OPL = 7; TL = 7; occipital carina

OOL

swollen;

weak,

without

incomplete,

reaching

scutellum and metanotum shiny, laterally viewed metanotum very

mesopleura and metapleura shiny and smooth;

granulated,

with some irregular keels or areolae,

without

a visible transversal keel between dorsal and posterior surface; dorsal surface very reduced, shorter than breadth of metanotum, as scu I ptured as posterior surface; surface;

pterostigma, part,

propodeum almost fully composed of the posterior

fore wing hyal ine, with distal

nearly

invisible;

and without dorsal

with a dark transversal

part of radial gonoforceps

proximal

TYPICAL MATERIAL:

Ikingi

Mariout,

444)

without

BM!

proximal

inner distal

dorsally viewed

process long and

(Libya)

holotype M!

in GE.

HOSTS: unknown DISTRIBUTION: Palaearctic region: EGYPT:

(Fig.

membranous process,

broad; tibial spurs 1,1,1. LOCUS TYP ICUS: Giarabub oasis

band beneath the

vein much shorter than

LIBYA:

Giarabub Oasis (Cirenaica),

Fayoum-Tamiya,

WA!

Wassiya,

GE!

WA!

3. Mirodryinus ungulatus Ponomarenko

Mirodryinus ungulatus Ponomarenko 1972: 674.

FEMALE:

fully

winged;

length 2,31

part of frons darkened;

protorax redd ish-testaceous; brown-b lack; black,

mm;

head reddish,

antennae brown,

scutum brown;

propodeum black;

pleura,

scutell um and metanotum

mesosternum and metasternum

except for a reddish spot on mesopleura;

testaceous-dark;

antennae distally thickened;

following proportions,

7:

3,5, 9,

6:

5,

with vertex and

wi th segments 1-3 testaceous;

4,5:

abdomen brown;

antennal 4,5:

4,5:

segments 4,5:

6;

legs in

head shiny,

puncta te, wi thout scu I pture among punctures; frons wi thou! med i an and lateral keels; occipital carina complete; temples very prominent; POL = 6; OL = 3;

OOL

=

9;

tubercles reaching

OPL = 7,5; tegulae;

TL

=

scutum,

10;

pronotum shiny,

short;

pronotal

scutellum and metanotum shiny,

672

Fig.

Mem.

Amer.

Ins!.,

no.

37, 1984

445 - Female of Mirodryinus ungulatus Ponomarenko (type): A: chela; B:

smooth,

weakly punctate,

incomplete,

fore tarsa I segments.

without sculpture among punctures;

reaching approximately 0,5

reticulate rugose, wing

Ent.

dull,

without transversal

with a dark transversal

without a distinct distal

length of scutum;

part;

or longitudinal

band on distal fore tarsal

half;

segments

radial

notaulices

propodeum keels;

fore-

vein straight,

in following

proportions: 85: 13: 15: 16: 41; segment 1 of front tarsus with a proximal hollow ("antenna-cleaner") distally del imitated by a big prominence (Fig.

445); enlarged claw (Fig. 445) without subapical tooth, bristles or other appendixes; segment 5 of front tarsus (Fig. 445) with 2 rows of nearly 26 peg-like lamellae; apex with a group of nearly 5 lamellae; tibial spurs 1,1,1. MALE:

unknown

LOCUS TYPICUS: Husseyn-Dey (Algeria) TYPICAI_ MATERIAL: holotype F! in LN. HOSTS:

unknown

DISTRIBUTION:

Palaearctic region:

ALGERIA:

Husseyn-Dey,

4. Mirodryinus atlanticus n. sp.

LN!

Bocchinae

Fig. 446 -

FEMALE:

Fore tarsal

fully winged;

673

segments of Mirodryinus atlanticus n.

length 2,62 mm;

head reddish;

sp.

(type)

antennae brown,

674

Mem.

Amer.

with segments 1-3 testaceous;

En!.

pronotum,

metanotum and propodeum black; abdomen brown-dark; punctate, 10;

with

occipital

legs and

keels;

dark

straight,

band on

not curved;

scutum shiny,

(Fig.

446)

lamellae; segment

=

5;

head shiny, OL

1 of front

shiny,

distal

3:

LOCUS TYPICUS:

radial oval,

3,5:

7;

without

with a proximal

hairy,

(Canary

holotype F!

9;

TL

=

with smooth,

propodeum reticulate

vein

fore-wing

brown;

fore tarsal

bristles,

segments

tarsus

in addition

lamellae,

(Fig.

with a

very short and

segment 5 of front

hollow

big prominence

Gran Tarajal

TYPICAL MATERIAL:

446)

strongly

=

OOL

weakly punctate,

hairless;

region;

2,5:

(Fig.

tarsus

3;

rugose,

with some bristles and 3 peg-I ike hairs, enlarged claw

=

keels on posterior surface;

17:

distally del imitated by a MALE: unknown

HOSTS:

and scutellum reddish;

visible only on anterior third;

pterostigma short,

in following proportions:

1984

and metap leura redd i sh;

POL

notaul ices incomplete,

longitudinal

transversal

scutum

pronotum short,

scutellum and metanotum smooth, without

37,

tegulae testaceous;

carina complete;

without sculpture;

no.

mesop leura

temples very prominent;

numerous transversal

rugose,

Ins!.,

to 4 distal

hairs;

("antenna-cleaner")

446);

tibial

spurs 1,1,1.

Islands)

in HE.

unknown

DISTRIBUTION:

Palaearctic region:

(Fuerteventura),

CANARY

ISLANDS:

Gran Tarajal

HE!

5. Mirodryinus ponomarenkoae n. sp.

Mirodryinus ungulatus Ponomarenko partim 1972: 674.

FEMALE:

fully

winged;

on ocellar region; reddish,

length 3,20 mm;

antennae missing

with anterior surface brown;

propodeum and abdomen black;

head reddish,

in holotype;

with a brown spot

pronotum reddish;

scutellum reddish;

mesopleura reddish,

scutum

metanotum,

with sutures blackish;

metapleura black; legs testaceous; head shiny, weakly alutaceous, strongly punctate on frons; occipital carina complete; frons without median keel; POL

=

=

10;

9;

OL

=

TL

=

10;

? (anterior ocellus damaged and not visible); OOL temples very prominent;

with numerous transversal punctate;

keels;

notaulices incomplete,

scutellum and metanotum shiny, rugose, dark

without

transversal

straight tarsal (Fig.

longitudinal

(Fig.

segments 447)

a proximal

scutum shiny, smooth, region;

not curved;

hollow

lamellae,

bristles,

("antenn-cleaner")

20:

hairs;

3:

=

8;

OPL

hairy, not

length of scutum; propodeum reticulate

vein

fore-wing

3: 4:

segment

with a

very short and

pterostigma short and oval,

in following proportions:

without

hairless; radial

rugose,

weakly alutaceous,

reaching only 0,3

keels on posterior surface;

band on distal

448 A),

pronotum short,

10;

brown;

fore

enlarged claw

1 of front

distally delimitated by a

tarsus with big

Bocchinae

Fig.

447 -

Fore tarsal

segments of female of Mirodryinus ponomarenkoae n. sp.

prominence (Fig.

447);

nearly

17 lamellae;

1,1,1. MALE:

unknown

675

(type)

segment 5 of front

tarsus

LOCUS TYPICUS: Western part of Mount Zamryn-Ula TYPICAL MATERIAL: holotype F!in LN. HOSTS;

Zamryn-Ula

Palaearctic region:

(Central

with 1 row of tibial

spurs

(Mongolia)

Aymag),

MONGOLIA:

Western part of Mount

LN!

the holotype of this species was designated by Ponomarenko

as para type of M. of Mrs.

447)

unknown

DISTRIBUTION: NOTES:

(Fig.

apex with a group of nearly 8 lamellae;

(1972)

ungulatus Ponomarenko. The species is named in honor

Nadezdha Ponomarenko.

5. Radiimancus Moczar

Radiimancus Moczar 1983b: 203.

TYPE SPECIES: Radiimancus olmii Moczar 1983b: 204, FEMALE labial

(Figs 449, palpi

tooth between 0,5 times as

450):

ful!y

with 3 segments;

winged;

long as segment 1 (Fig. tarsus without

segment 4 of front 451);

lamellae,

occipital

desig.

with 6 segments;

mandibles quadridentate,

the two posterior teeth;

segment 5 of front

orig.

maxi Ilary palpi

with a rudimentary tarsus

less than

carina complete;

with bristles or peg-like hairs

Mem.

676

Amer.

En!.

Ins!.,

no.

37,

1984

A~

B~ Fig. 448 - Pterostigma and radial vein of females of Mirodryinus ponomarenkoae n. sp. (type)(A) and Radiimancus olmii Moczar (type)(B)

(Fig.

451);

bristles

enlarged claw without teeth,

(Fig.

451);

radial

vein

(Fig.

MALE:

unknown

DISTRIBUTION: HOSTS:

pronotal

448 B);

lamellae,

peg-I ike hairs or

tubercles reaching tegulae;

tibial

fore-wing with no

spurs 1,1,1.

Palaearctic

unknown

SPECIES:

1

Radiimancus olmii Moczar

Radiimancus olmii Moczar 1983b: 204.

FEMALE

(Figs 449,

450):

fully winged;

anterior surface of frons,

length 2,80 mm;

head black,

clypeus and mandibles testaceous;

with

antennae

testaceous, with last 5 segments darkened; pronotum and prosternum reddish; scutum reddish, with central surface darkened; scutellum, metanotum and propodeum black; mesosternum reddish, with black spots; metasternum and metapleura black; mesopleura reddish; abdomen brown; legs testaceous;

antennae weakly distally thickened;

following proportions: quadri dentate, head shiny, occipital

8:

3:

12: 8,5:

with a rud imentary

smooth,

6:

6:

6:

6:

antennal 8;

segments in

mandibles

tooth between the two posterior teeth;

weakly punctate,

carina complete;

7:

without sculpture among punctures;

frons without median keel;

POL = 8;

OL = 4;

OOL = 10; OPL = 8; TL = 8; pronotum short, hairy, rugose, pronotal tubercles reaching tegulae; scutum shiny, weakly punctate, without sculpture among punctures;

notaulices incomplete,

reaching only 0,3 lenth

of scutum; scutellum and metanotum shiny, smooth, without sculpture; propodeum reticulate rugose, without longitudinal or transversal keels;

677

Bocchinae

Fig. 449 -

Female of Hadiimancus olmii Moczar

(type)

fore-wing with a dark transversal band on distal surface; pterostigma broad and oval; radial vein absent (Fig. 448 B); fore tarsal segments following proportions:

17: 2:

2,5:

3:

9;

in

segment 1 of front tarsus with a

proximal hollow ("antenna-cleaner") distally del imitated by a big prominence (Fig. 451); segment 5 of front tarsus (Fig. 451) with 2 rows of nearly 27 bristles extending continuously to the apex; enlarged claw (Fig. 451)

without

lamellae, bristles or hairs;

tibial

spurs 1,1,1.

MALE: unknown LOCUS TYP I CUS: Bu I gan (Mongol i a) TYPICAL MATERIAL: holotype F! in BU. HOSTS: unknown DISTRIBUTION: Palaearctic region:

MONGOLIA:

Bulgan

(10 Km SSW Somon,

678

Mem.

Fig.

Chovd

aimak),

450 -

Amer.

Ent.

Inst.,

no.

37,

1984

Female of Radiimancus olmii Moczar (type)

BU!

6. Mystrophorus Forster

Mystrophorus Forster 1856: 91. Mystrophorus Forster: Richards 1937: 119.

TYPE SPECIES: Mystrophorus FEMALE labial

(Figs 452, 453): palpi

with 3 segments;

tooth between tarsus

less than 0,5 times as

or peg-like hairs

456);

I ike hairs or bristles (Fig. tibial spurs 1,1,2. (Figs 454,

rudimentary

less wide than tegulae;

with 6 segments;

396 A);

tarsus without

with a rudimentary

segment 4 of front 456);

456);

pronotal

occipital

lamellae,

enlarged claw without teeth,

455): micropterous;

tooth between

(Fig.

long as segment 1 (Fig.

segment 5 of front (Fig.

maxillary palpi

mandibles quadridentate,

the two posterior teeth

carina complete;

MALE

formicaeformis Ruthe 1859, monotypic.

micropterous;

with bristles lamellae,

mandibles quadridentate,

the two posterior teeth;

maxi Ilary palpi

peg-

tubercles reaching tegulae; with a

areolae of propodeum

with 6 segments;

labial

palpi

with

3 segments; temples distinct. DISTRIBUTION: Palaearctic HOSTS:

Cicadellidae (excl ud i ng I diocerinae, Macropsinae,

SPECIES:

1

Mystrophorus formicaeformis Ruthe

Typhlocybinae).

Bocchinae

679

A

Fig. 451 - Female of Radiimancus oimii Moczar (type): A: fore tarsal segment 1; B: fore tarsal segments 1 (partly)-4 and chela

Mystrophorus iormicaeiormis Ruthe 1859: 120. Mystrophorus iormicaeiormis Ruthe: Ponomarenko 1978: 23.

FEMALE (Figs 452, 453): micropterous, with fore wings as long as tegulaej hind wings rudimentary, rounded; length 1,75-2,50 mm; black, with

680

Mem. Amer. Ent.

Fig. 452 -

Inst.,

no.

37,

1984

Female of Mystrophorus formicaeformis Ruthe

Bocchinae

Fig.

453 -

antennae brown

Female of

681

Mystrophorus formicaeformis Ruthe

(except for segments 1-2 testaceous),

tegulae brown-testaceous,

legs yellow

mandibles yellow,

(with hind coxae almost

clubs of femora and hind tibiae brownish);

body dull,

hairy,

fully black, except for

abdomen hairless; head without median keel between antenna I sockets; occipital carina complete; head, thorax and propodeum granulated, without striae or keels; invisible;

some keels are visible near apex of propodeum;

fore tarsal

segments 1-4 in following proportions:

notaulices

5,5:

1,2:

2,3:

2,3; en I arged cl aw (Fig. 456) without I ameli ae or brist les, wi th apex rounded; segment 5 of front tarsus (Fig. 456) without lamellae, with numerous rows of bristles or peg-like hairs. MALE

(Figs 454,

455):

micropterous,

with fore wings narrow and broadened

at apex; hind wings rudimentary, pointed; length 1,87-2,75 mm; black, with antennae brown(except for segments 1-2 testaceous), tegulae yellowbrown,

fore-wing

femora blackish, for abdomen sockets;

almost hairless;

occipital

granulated; propodeum;

yellow-brown,

legs yellow

(with coxae and clubs of

mid and hind tibiae brownish);

body dull,

head without median keel

carina complete;

head,

without striae or keels,

hairy,

between

except

antenna I

thorax and propodeum

except for some keels near apex of

notaulices invisible.

LOCUS TYPICUS:

Boden

TYPICAL MATERIAL: HOSTS: in Germany

(Germany)

not found

(not

in BM and

in Gent,

Belgium).

Deltocephalus sp. (Haupt 1938); in U. S. S. R. Diplocolenus abdominalis F. (Ponomarenko 1978). DISTRIBUTION: 1978); det.

Palaearctic region:

HUNGARY: Or Sz.

Miklos,

U.

BU!

S. Vacz

S.

R.:

Moscow

(Tudosdomb),

(Ponomarenko BU!

Hungary

(male

Mystrophorus cursor Forster type), VI! CZECHOSLOVAKIA (Ogloblin

1924b);

GERMANY:

E I steraue (Ha lie),

HA!

BM!

Boden

(Ruthe 1859);

Ha II e-a-

5., HA! Bellinchen (Oder), HA! KUpka (Th.), EB! Mallertshofer Holzb. (MUnchen), EB! OT! Saale-Ufer (Haupt 1938); Schl inzheim (Mallertshofer Holz), GV! ENGLAND: Hayling Island (Hampshire), BM! GREECE: Kerkyra Island,

BU!

682

Mem.

Fig.

454 -

FEMALE

En!.

Inst.,

Male of Mystrophorus

no.

37,

1984

formicaeformis Ruthe from Hungary

5UBFAMILY THAUMATODRYININAE N.

6.

TYPE GENUS:

Amer.

Thaumatodryinus R.C.L. Perkins 1905

(Figs 19,

457):

fully

winged;

fore-wing

with costal,

submedian cells clearly enclosed by pigmented veins also first brachial labial

palpi

SUBF.

cell

clearly enclosed;

with 3 segments;

front

(Fig.

maxillary palpi

tarsus chelate;

median and

457);

usually

with 6 segments;

chela with rudimentary

claw (Fig. 459); antennae with tufts of long hairs on segments 5-10 (last segment with two tufts) (Fig. 19); occipital carina complete; mandibles with 4 teeth progressing spurs 1,1,2.

larger from anterior one to posterior (Fig.

5);

tibial

Thaumatodry i n i nae

Fig.

MALE

Male of Mystrophorus

455 -

(Figs 28,458):

fully

winged;

683

formicaeformis Ruthe from Hungary

fore-wing

with costal,

submedian cells clearly enclosed by pigmented veins palpi

with 6 segments;

teeth progressing carina complete; pterostigma

28);

GENERA:

with 3 segments;

with metacarpus as

tibial

Palaearctic

Oriental, Nearctic, HOSTS: Flatidae SPECIES:

palpi

median and 28); 5);

occipital

long as or longer than

spurs 1,1,2.

(?)(fossil

Neotropic,

maxillary

mandibles with 4

larger from anterior one to posterior (Fig. fore-wing

(Fig.

DISTRIBUTION:

labial

(Fig.

species:

see notes),

Ethiopian,

Australian

1 17

NOTES: Brues described three species of Thaumatodryinus from Palaearctic region (Baltic amber): filicornis (Brues 1923), deletus Brues 1933 and

gracilis Brues 1933. Though the original descriptions are incomplete and the typical correct.

material

is

In the original

lost,

I think that the Brues determination

description of Th.

is

deletus in fact Brues wrote

about the presence on the I ast two antenna I segments of tufts of long hairs: this character is recognizable only in Thaumatodryinus. In living

Thaumatodryinus, however, the tufts are on antennal segments 5-10. Probably

the three Palaearctic fossi I species are belonging

extinct genus of Thaumatodryininae,

to another

perhaps Harpactosphecion

Haupt 1944

(described by Haupt for one of the three above-mentioned species,

filicornis) .

Thaumatodryinus R.C.L. Perkins

Thaumatodryinus R.C.L. Perkins 1905: 58.

684

Mem. Amer.

En!.

Ins!.,

no.

37,

1984

Ilflll III/flilltilf

Fig. 456 -

Female of Nystrophorus tormicaeiormis Ruthe: tarsal

A:

chela;

B:

fore

segments 1-4.

Thaumatodryinus R.C.L. Perk i ns: Kieffer in Kieffer et Marshall 1906: 490.

Thaumatodryinus Thauma todryin us Thaumatodryinus Thaumatodryinus

TYPE SPECIES: FEMALE

R.C.L. R.C.L. R.C.L. R.C.L.

Perk i ns: Perkins: Perkins: Perkins:

Kieffer 1914b: 16. Krombein 1952: 97. Krombein 1958: 98. Krombein 1979: 1243.

Thaumatodryinus koebelei R.C.L. Perkins 1905, orig. desig.

(Figs 19, 457):

fully winged;

fore-wing with costal,

median and

submedian cells clearly enclosed by pigmented veins (Fig. 457); usually also first brachial cell clearly enclosed; maxi Ilary palpi with 6 segments; labial palpi with 3 segments; front tarsus chelate; chela with rudimentary claw (Fig. 459); antennae with tufts of long hairs on the segments 5-10 (last segment with two tufts) (Figs 19, 471); pronotal tubercles reaching tegulae; occipital carina complete; mandibles with 4 teeth progressing

Thaumatodry i n i nae

685

/

Fig.

457 -

Female of Thaumatodryinus Guinea)

flavus n. sp. from Waris (New

686

Fig.

Mem.

458 -

Amer.

En!.

Ins!.,

no.

37,

1984

n.

sp.

from Kira-Kira

Male of Thaumatodryinus flavus

(Solomon

Islands)

larger from anterior one to posterior MALE

(Figs 28,

458):

fully

winged;

(Fig.

5);

fore-wing

tibial

submedian cells clearly enclosed by pigmented veins palpi

with 6 segments;

teeth progressing carina complete; pterostigma

(Fig.

DISTRIBUTION: fossil

labial

palpi

spurs 1,1,2.

with costal,

with 3 segments;

(Fig.

28);

larger from anterior one to posterior (Fig. fore-wing 28);

with metacarpus as

tibial

Ethiopian,

maxillary

mandibles with 4 5);

occipital

long as or longer than

spurs 1,1,2.

Oriental,

Nearctic,

Neotropic, Australian

Palaearctic species see note on page 683).

HOSTS: Flatidae GENERA: 1 SPECIES:

median and

17

ETH lOP I AN SPEC I ES OF THAUMATODRYINUS

Five species are known.

KEY TO THE SPECIES

(for

Thaumatodry i n inae

687

FEMALES

Fore and hind wings with a dark

transversal

band ••••••••••••••• 2

bands •.••.•••.•••• 3 2 Head mostly black; mesothorax, metathorax and propodeum black ••• • • • • • • • • • • • • • • • • • • • • • • • • • • •. 1. mirabilis n. sp. Fore and

-

hind wings without

dark

transversal

Head fully reddi sh-testaceous; mesothorax, metathorax and propodeum reddi sh-testaceous, wi th scutellum black ••••••.••••.••

· . . . . • . . . . . . . . • . . • • . . . . . . . . . 2. infuscatus n. sp. 3 Head wi th OPL longer than OL; thorax and propodeum partly black • . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3. dentatus Benoi t -

Head wi th OPL as

long

as or

short er

than OL;

thorax and

4 4 Occiput more strongly excavated; fore trochanters nearly as long as fore femora; posterior surface of propodeum wi th 2 complete propodeum fully

longitudinal

reddish-testaceous •••••••••••••••••.•••••••••••

keels;

notaulices more converging . . . . . . . . . . . . . . . . ..

· . . . . . . • . . . . . . . . . . . . . . . . . . .• -

Occiput

not

long as

fore

tracks

of

or weakly femora;

incomplete

townesi n.

4.

excavated; posterior

sp. trochanters nearly

fore

surface

longitudinal

of

keels;

0,5 as

propodeum only wi th notaulices

2

less

5. migratorius Beno i t

converg i ng . . . . . . . . . . . . . . . . . .

MALES

Only

the supposed male of

Th.

tOlvnesi n. sp. is known.

1. Thaumatodryinus mirabilis n. sp.

FEMALE:

fully

winged;

length 6,25-7,00 mm;

clypeus and mandibles yellow;

head black,

with malar space,

sometimes occiput partly brown-light;

antennae testaceous; pronotum and propectus yellow ; mesothorax, metathorax and propodeum black, with propodeal spiracles brown; black;

abdomen reddish;

legs testaceous,

petiole

with fore clubs of femora black,

mid and hind clubs of femora brown; antenna I segments in following proportions: 20: 7: 40: 45: 44: 45: 35: 25: 19: 15; head dull, reticulate rugose; hairy,

occipital

carina complete;

crossed by a

transversal

striae;

reticulate rugose;

transversal pronotal

OPL

notaul ices complete, metanotum smooth,

reticulate rugose;

fore-wing

part of radial

vein

tegulae;

with weak

scutum dull,

posteriorly separated; transversal

longer than

proximal

fully

scutellum

with some anterior furrows;

with a brown

much

pronotum shiny,

weakly punctate,

tubercles reaching

reticulate rugose; distal

longer than OL;

impression,

propodeum

band and with part;

radial

cell

688

Mem.

Fig.

459 -

Chela of

Amer.

Ent.

Inst.,

no.

37,

1984

Thaumatodryinus mirabilis n. sp. from Ankasoka (Madagascar)

closed;

pterostigma brown-I ight;

transversal

band;

segment

hind wings with a

1 of front

distal

tarsus nearly as

brown

long as segment 4;

Thauma todry i n i nae

689

(

Fig. 460 -

Chela of Thaumatodryinus infuscatus n.

sp.

(type)

segment 3 produced into a hook; enlarged claw with 2 subapical teeth and with 32 lamellae (Fig. 459); segment 5 of front tarsus (Fig. 459) with 2 rows of 59 lamellae; apex with a group of 5 lamellae; tibial MALE: unknown LOCUS TYPICUS: Ankasoka (Madagascar) TYPICAL MATERIAL: holotype F! and 1 paratype F! in P.

spurs 1,1,2.

690

Mem.

HOSTS:

Amer.

Ent.

Inst.,

no.

37,

1984

unknown

DISTRIBUTION:

Ethiopian region:

MADAGASCAR:

Ankasoka

(Route Lakato),

P!

2. Tha umatodryinus infuscatus n. sp.

FEMALE:

fully

winged;

length 4,68 mm;

and scutellum black;

head shiny,

carina complete;

longer than OL;

POL

granu I a ted,

with weak

impression;

pronotal

rugose; radial

tubercles reaching

scutellum

fore-wing vein much

pronotum shiny,

transversa I stri ae,

sculptured by numerous separated;

reddish-testaceous,

weakly reticulate rugose;

longitudinal

tegulae;

and metanotum shiny,

with a

brown

longer than

weakly

transversa I

scutum shiny,

fully

notaulices complete, punctate;

transversal

proximal

occipital

hairy,

crossed by a

keels;

with petiole

posteriorly

propodeum reticulate

band and with distal

part;

radial

cell

part of

closed;

pterostigma brown-I ight; hind wings with a brown distal transversal band; segment 1 of front tarsus nearly as long as segment 4; segment 3 produced into a (Fig.

hook; 460);

enlarged claw with 2 sUbapical segment 5 of front

tarsus

lamellae;apex with a group of 6 MALE:

(Fig.

lamellae;

teeth and with 32 460)

lamellae

with 2 rows of 43

tibial

spurs 1,1,2.

unknown

LOCUS TYPICUS:

St.

TYPICAL MATERIAL:

Augustin

(Madagascar)

holotype F!

in BM.

HOSTS: unknown DISTRIBUTION: Ethiopian region:

MADAGASCAR:

St.

Augustin

(Tulear),

BM!

FEMALE: fully winged; length 4,50 mm; head yellow-testaceous, with occiput partly black, ocellar region black, frons with 4 black spots

(two

3. Thaumatodryinus dentatus Benoit

Thaumatodryinus dentatus Benoit 1953b: 404.

in front of the antennal antennae testaceous, pronotum

sockets and two behind the antennal

with segments 1 and 4-10 dark;

yellow-testaceous,

testaceous,

with sides partly brown;

with anterior margin black;

between scutellum and scutum brown; and metanotum black; petiole black;

clubs of femora brown; 28:

33:

41:

incomplete, OPL

40:

28:

scutellum yellow-testaceous;

13: 9:

11;

with anterior third reddish;

legs testaceous,

head shiny, the ocell i

pronotum sh i ny,

furrow

sides of scutellum with fore and hind

antenna I segments in following proportions:

visible only behind

longer than OL;

propectus yellow; scutum yellow-

metanotum black;

propodeum black-brown,

abdomen brown-yellow;

sockets);

granulated; and on

granu I a ted,

occipital

15: 9:

carina

the sides of the vertex; ha i ry,

crossed by

a

Thaumatodryi n i nae

Fig.

461

-

Chela of

691

Thaumatodryinus dentatus n.

sp.

(type)

transversal impression; pronotal tubercles reaching tegulae; scutum shiny, hairy, granulated, with weak longitudinal striae; notaulices almost complete, sculpture;

posteriorly separated; metanotum rugose;

third almost smooth; distal closed;

part of radial

fore-wing

without dark

vein much

pterostigma brown-light;

than segment 4

(26:19);

with 2 subapical tarsus

scutellum shiny,

(Fig.

461)

transversal

longer than proximal segment

1 of front

segment 3 produced

teeth and with 36 with 2 rows of 63

apex; tibial spurs 1,1,2. MALE: unknown

smooth,

without

propodeum reticulate rugose,

into a

lamellae (Fig.

with anterior

bands and with

part;

radial

tarsus much hook; 461);

cell

longer

enlarged claw segment 5 of front

lamellae extending continuously

to the

692

Mem.

LOCUS TYPICUS:

Bekily

TYPICAL MATERIAL: HOSTS:

Amer.

Ent.

Ins!.,

37,

no.

1984

(Madagascar)

holotype F!

in P;

2 paratypes FF!

in TV.

unknown

DISTRIBUTION:

Ethiopian region:

MADAGASCAR:

Bekily,

P!

TV!

4. Thaumatodryinus townesi n. sp.

FEMALE: black;

fully

length 3,43-5,62 mm;

winged;

slender,

weakly distally thickened;

7: 5: 50: 40;

proportions:

complete; shiny,

6; OL

tubercles

smooth,

transversal

with 2 cell

smooth,

granulated,

logitudinal

keels;

=

8; OPL

tegulae;

lateral

scutum,

occipital

=

1,5; TL a

notaulices complete,

with a

carina

7; pronotum

transversal

scutellum

with

keels,

impression;

and

metanotum

very converging;

with some areolae or tracks of areolae, and posterior surface;

median

area dull,

without dark

closed and with distal

part;

weakly alutaceous,

crossed by

between dorsal

hind wings hyal ine,

=

3,5; OOL

reaching

keel

flat,

frons without median or

without sculpture;

propodeum dull, a

=

with petiole

antennae very

in front of the anterior ocellus;

without sculpture,

pronatal

shiny,

=

POL

testaceous,

segments 1-4 in following

antennal

head shiny,

occiput strongly excavated; short median furrow

fully

with segments 1-2 testaceous;

antennae brown,

part of radial

fore trochanters nearly as

smooth,

transversal long

with

posterior surface

granulated;

fore and

fore-wing

with radial

bands;

vein much

as fore femora

longer than proximal

(44:46) and nearly as

(44:46) and as fore tibiae (44:42); fore tarsal segments 16: 4: 8: 31: 41; enlarged claw (Fig. 462) with 2 subapical teeth and with 6 lamellae; segment 5 of front tarsus (Fig. 462) with a row of 9 lamellae; apex with a group of 10 lamellae; tibial spurs 1,1,2. MALE (not reared: from Sokoke Forest, Kenya): fully winged; length 2,68 long as fore coxae

in following proportions:

mm;

head black,

with mandibles testaceous;

segments 3-10 darkened; partly

testaceous

antennae testaceous,

prothorax testaceous;

thorax black,

(notaulices and region between

with

wi th scutum

the notaulices);

abdomen

brown; legs testaceous, antennae not distally thickened; antennal segments in following proportions: 6: 6: 9: 12: 11: 12: 11: 10: 8: 10; head dull, granulated and reticulate rugose; median

keel;

granulated, separated;

POL

=

weakly

6; OL

occipital

2; OOL

=

reticulate rugose;

reticulate rugose,

keels;

fore-wing hyaline,

radial

vein

longer than proximal

LOCUS TYPICUS:

Kampala

TYPICAL MATERIAL: unknown

part;

=

1; TL

=

smooth,

tibial

posteriorly

without sculpture;

transversal

transversal

frons without

4; scutum dull,

notaul ices complete,

without

without dark

carina complete;

6; OPL

scutellum and metanotum shiny,

propodeum dull,

in OL. HOSTS:

=

or

bands;

longitudinal distal

part of

spurs 1,1,2.

(Uganda)

holotype F!

and 1 paratype F!

in TW;

1 paratype F!

Thaumatodry in inae

Fig.

462 -

Chela of Thaumatodryinus

693

townesi n. sp. (type)

694

Mem. Amer.

Ent.

Inst.,

no.

37,1984

DISTRIBUTION: Ethiopian region: UGANDA: Kampala, TW! Forest, TW! SOUTH AFRICA: Tzaneen (Transvaal), TW! NOTES:

the species is named in honor of Mr.

KENYA:

Sokoke

Henry Townes.

5. Thaumatodryinus migratorius Benoit

Fig. 463 -

Chel a of Thaumatodryinus migratorius n.

Thaumatodryinus migratorius Benoi t 1953b: 404.

sp.

(type)

Thaumatodry i n i nae

FEMALE: black;

fully

winged;

ocellar region

length 3,5 mm; weakly brownish;

4-10 darkened;

antennal

22:

10: 8;

23:

20:

incomplete, OPL = OL; transversal

10:

reddish-testaceous,

with petiole

antennae testaceous,

segments in following proportions:

head shiny,

weakly punctate;

visible only behind the ocelli pronotum shiny, impression;

695

and on

weakly punctate,

pronotal

with segments 7:

occipital

5:

14:

15:

carina

the sides of the vertex;

hairy,

tubercles reaching

crossed by tegulae;

a

scutum shiny,

hairy, punctate, notaulices almost complete, posteriorly separated; scutellum and metanotum shiny, weakly punctate; propodeum reticulate rugose,

with anterior third almost smooth;

transversal proximal

bands and with distal

part;

radial cell

fore-wing

part of radial

closed;

pterostigma brown-light;

front tarsus nearly as long as segment 4;

teeth and with 21

segment 5 of front

463)

(Fig.

a group of 4 lamellae; MALE: unknown

tibial

LOCUS TYP I CUS:

(Madagascar)

Beki I y

TYPICAL MATERIAL:

segment 1 of

segment 3 produced

enlarged claw with 2 subapical tarsus

without dark

vein much longer than

lamellae

with 2 rows of 38

into a hook;

(Fig.

lamellae;

463); apex with

spurs 1,1,2.

holotype F!

in P.

HOSTS: unknown DISTRIBUTION: Ethiopian region: 'MADAGASCAR:

Bekily,

P!

THAUMATODRYINUS

OR I ENTAL SPEC I ES OF

Three species are known

KEY TO THE SPECIES

FEMALES

segment 3 less than 1,5 times as long as segment ........................... " 2. asiaticus n. sp.

Antennal -

Antennal

segment

3 more

2 Scutum with numerous

than

1,5

than POL . . . . . . . . . . . . . . . . . . . . . -

Scutum without

times as

longitudinal

longitudinal

1.

striae;

3.

segment

slightly

1 •••• 2

longer

philippinus n. sp.

striae;

long as POL .••••••••• ••••••••

long as OOL

POL more

than

malayanus n. sp.

MALES

three

times as

696

Mem.

One only male

Amer.

is known.

Ent.

It

Ins!.,

no.

37,

1984

is not possible to know

if it's a

philippinus or an asiaticus.

Fig.

464 -

Chela of

Thaumatodryinus philippinus n. sp. (type)

Thaumatodryi n i nae

697

1. Thaumatodryinus philippinus n. sp.

FEMALE:

fully

winged;

suture darkened; darkened;

length 4,80 mm;

legs yellow,

reddish-testaceous;

segment 3 more than

antennal

epistomal

with clubs of fore and hind femora 1,5 times as

long

as segment

(30: 19); head swollen, dull, granulated; frons with tracks of irregular striae; occipital carina complete; frons without median keel; POL = 9; OL = 4;

OOL = 12; OPL = 3; TL = 3; pronotum dull, hairy, granulated, strong transversal impression and with posterior tubercles reaching tegulae;

scutum granulated and with numerous

and areolae;

notaulices complete,

metanotum shiny,

granulated;

transversal

or

transversal

bands;

part 37;

(30:15);

longitudinal distal

fore tarsal

enlarged claw

23 41

lamellae; lamellae;

MALE:

(Fig.

unknown

(see key

to

Dumaguete

TYPICAL MATERIAL:

vein

without

without dark

twice as

long as proximal

in following proportions:

with 2 preapical

32:

9: 26:

5:

teeth and a row of nearly

the males) (Philippines)

holotype F!

in AM.

unknown

DISTRIBUTION: Negros

part of radial segments

hyal ine,

segment 5 of front tarsus (Fig. 464) with two rows of nearly apex with a group of 4 lamellae; tibial spurs 1,1,2.

LOCUS TYPICUS: HOSTS:

fore-wing

weak striae

scutellum and

propodeum reticulate rugose,

keels;

464)

longitudinal

posteriorly separated;

with a

Oriental

Island),

region:

PHILIPPINES:

Dumaguete

(Camp Lookout,

AM!

2. Thaumatodryinus asiaticus n. sp.

FEMALE:

fully

winged;

length 4,00-5,00 mm;

someti mes wi th brown spot on frons; testaceous;

propodeum yellow-testaceous,

yellow-testaceous;

head reddish-testaceous

antennae testaceous;

propectus

with sides partly brown;

scutum

scutell um and metanotum yellow-testaceous or reddish-

brown; propodeum fully or partly black; petiole black; abdomen reddish; legs yellow, sometimes with clubs of femora brown; head swollen, granulated,

dull,

antennal

segment 3 sl ightly

longer than segment 1 (23:20

or 24:18); occipital carina incomplete, visible only behind the ocelli and on the sides of the vertex; POL = 7; OL = 3,5; OOL = 9; OPL = 2; TL = 2,5;

propodeum shiny,

weakly

granulated,

impression;

pronotal

granulated;

notaulices complete,

almost

invisible;

reticulate rugose;

tubercles reaching

hairy,

tegulae;

crossed by a scutum dull,

posteriorly separated,

scutellum and metanotum shiny, fore-wing

with posterior parts

granulated;

without dark transversal

bands,

pterostigma brown dark and with distal

part of radial

than

segment 1 of front

as

proximal

long

part;

as segment 4

radial

cell

(24:22);

closed;

segment 3 produced

transversal

hairy, propodeum with

vein much

into a

longer

tarsus nearly hook;

enlarged

698

Mem.

Fig.

465 -

Chela of

claw with 2 subapical front

tarsus

lamellae; MALE

(Fig.

tibial

(supposed:

head testaceous,

Amer.

465)

Ent.

Inst.,

teeth and with 26

lamellae

see key

to the males):

fully

antennae slender,

proportions:

11: 6:

10:

granulated,

weakly

rugose;

OPl

2;

Tl

12:

=

4;

13:

(type)

segment 5 of

apex with a group of 5

13:

12:

abdomen and

12;

head hairy,

carina complete;

scutum hairy,

dull,

scutellum dull,

metanotum shiny,smooth,

with

legs

segments in following

10:

11:

length 2,40 mm;

antennae testaceous;

light;

antennal

occipital

posteriorly separated;

and punctate;

margin;

=

sp.

(Fig. 465);

winged;

with occiput and vertex brown; with scutum

7;

n.

spurs 1,1,2.

hairy;

=

1984

with 2 rows of 48 lamellae;

testaceous;

OOl

37,

Thaumatodryinus asiaticus

thorax and propodeum brown,

complete,

no.

POL = 7;

granulated; rugose,

dull, Ol = 5;

notaulices

reticulate rugose

some areolae near anterior

propodeum dull,fully reticulate rugose,

without transversal or

longitudinal keels; fore-wing hyaline, without dark transversal bands, hairy; pterostigma long and narrow; distal part of radial vein longer

Thaumatodry i n i nae

Fig.

than

466 -

proximal

Chela of Thaumatodryinus malayanus n.

part

LOCUS TYPICUS:

699

(19:14);

radial

Gunung Mulu Nat.

cell

open;

Park

tibial

(Sarawak,

sp.

(type)

spurs 1,1,2. Malaysia)

TYPICAL MATERIAL:

holotype F! in BM; 1 paratype F! in B; 1 para type F! 1 paratype M! in CO. HOSTS: unknown DISTRIBUTION: Oriental region: MALAYSIA: Gunung Mulu Nat. Park (Sarawak), BM! Lake Area (Bau, Sarawak), B! PHILIPPINES: Mt. Data, TW! N of Batu Batu (Tarawakan, Tawi Tawi), CO! in TW;

3.

Thaumatodryinus malayanus n. sp.

FEMALE: fully winged; length 4,60 mm; head testaceous, with ocellar triangle and a median frontal region brown; occiput and ventral side of head partly brown;

antennae brown,

with segments 1-3 testaceous and with

regions bearing tufts of hairs whitish; testaceous,

with a

lateral

propectus brown;

streak brown;

scutum,

pronotum

scutellum and metanotum

testaceous;

mesopleura and metapleura black,

testaceous; testaceous;

propodeum black, with anterior region of dorsal surface abdomen brown; legs testaceous, with cl ubs of femora brown;

antenna I segment 3 more than

1,5 times as long as segment 1 (39:20);

head swollen,

dull,

median keels;

POL = 3;

shiny,

without sculpture,

hairy,

and with a complete,

granulated;

transversal

with posterior apexes

occipital

carina complete;

OL = 1; OOL = 15; OPL = 3;

pronotum

with posterior tubercles reaching tegulae

impression;

posteriorly separated;

frons without

TL = 4;

scutum dull,

granulated;

notaulices

scutellum and metanotum smooth,

shiny,

700

Mem. Amer.

very weakly granulated; transversal or transversal part 53;

distal

fore

tarsal

enlarged claw

Ins!.,

no.

keels;

fore-wing

part of radial segments in

(Fig.

466)

with 2 preapical

segment 5 of front

apex with a group of 4

HOSTS:

hyaline,

vein much

nearly 38 lamellae;

Templer Park

TYPICAL MATERIAL:

1984

tarsus

(Selangor,

holotype F!

dull,

without

without dark

longer than proximal

following proportions:

nearly 54 lamellae; MALE: unknown LOCUS TYP ICUS:

37,

propodeum reticulate rugose,

longitudinal

bands;

(50:22);

En!.

38:

6:

13: 34:

teeth and with a row of (Fig.

466)

lamellae;

Malaya,

with

tibial

two rows of

spurs 1,1,2.

Malaysia)

in CO.

unknown

DISTRIBUTION: Malaya),

Oriental

region:

MALAYSIA:

Templer Park

(Selangor,

CO!

NEARCT I C SPEC I ES OF THAUMATODRYINUS

One species is known.

Thaumatodryinus perkinsi Krombein

Thaumatodryinus perkinsi Krombei n 1952: 99.

FEMALE:

fully

winged;

petiole black; occipital

carina

of the vertex; impression, shiny,

length 4,37-5,62 mm;

pterostigma brown incomplete,

granulated;

granulated; fore-wing

fully

yellow-testaceous,

head swollen,

shiny,

visible only behind the ocell i

with

granulated;

and on

the sides

= OL; pronotum sh i ny, ha i ry, crossed by a transversa I

OPL

pronotal

tubercles reaching

notaulices almost complete,

scutellum and metanotum shiny, rugose;

dark;

with distal

weakly punctate;

part of radial

tegulae;

scutum hairy,

posteriorly separated; propodeum reticulate

vein much

longer than

proximal part and without dark transversal bands; segment 1 of front tarsus longer than segment 4 (23:21); segment 3 produced into a hook; enlarged claw

with 2 subapical

segment 5 of front a group of 5 MALE:

lamellae;

(Fig.

tibial

teeth and with 467)

19 lamellae

(Fig.

with 2 rows of 28 lamellae;

467); apex with

spurs 1,1,2.

unknown

LOCUS TYP ICUS:

Westmoreland State Park

TYPICAL MATERIAL: HOSTS:

tarsus

holotype F!

(U.

S.

A.)

in WA.

unknown

DISTRIBUTION:

Nearctic region:

Westmoreland State Park (Oaxaca),

TE!

U.

S.

A.:

Greenville

(Westmoreland Co.,

Va.),

(S.

WAr

C.),

LE!

MEXICO:

Miltepec

Thaumatodryininae

Fig.

467 -

Chela of

701

Thaumatodryinus perkinsi n. sp.

(type)

Th. perkinsi Krombein is very similar to Th. koebelei R.C.L. Th. perkinsi, however, has pterost i gma brown dark and transversal impression of pronotum deeper. Th. perkinsi is also similar to Th. rufus Richards from Neotropic region. Th. rufus, NOTES:

Perk ins from Austral i an region.

however,

has transversa I

impression of pronotum

less deep.

702

Mem.

Amer.

En!.

Inst.,

no.

37,

1984

THAUMATODRYINUS

NEOTROPIC SPECIES OF

Six species are known.

KEY TO THE SPECIES

FEMALES

-

Not a u I ice s

c omp let e •••••••••••••••••••••••••••••••••••••••••••• 2

Notaulices

incomplete ••••••••••••••••••••.•••••••••••••••••••.. 4

2 Antennal

segment

3

. . . . . . . . . . . . . . . . . .. -

Antennal

I ittle

longer

than

2. minimus n.

segment

1 •••••••••••••••••

sp.

segment

3 more than 1,5 times as long as segment 1 . . . . 3 long as breadth of ocell i; radial cell wide (Fig. 468 B} ••••••••••••••••••••••••.•••••••••••••••••.•••• . . . . . . . . . . . . . . . . . . . 1. variegatus De Santis and Vidal Sarmiento - Head wi th POL nearl y three times as long as breadth of ocel I i ; radial cell narrow (Fig. 468 A} . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . • . . . . . . • . • • • • • • •• 6. clarus n. sp. 4 Segment 1 of front tarsus nearly as long as segment 4; head black or brown-b lack •••••••••••••••••••••••••••••••••••••••••••• . . . . . . . . . . . . . . . . . . . 3. macilentus De Santis and Vidal Sarmiento - Segment 1 of front tarsus much longer than segment 4; head fully reddi sh or reddi sh-testaceous •••••••••••••••••••••••••••••••••• 5 5 Head with OL shorter than OPL ••••••••••••••••••••••••••••••••••• . . . . .. .. .. .. .. .. ... 4. rufus Richards - Head wi th OL IOr)ger than OPL . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. 5. bruchi De Santis and Vidal Sarmi en t 0

3 Head wi th POL nearly as

MALES

Antennal

segment

head with OOL

4

less

than

less

than

three

................... -

Antennal

segment

2 Head wi th OL as

5.

4 more

1,5

times as

times as

long as

bruchi De Santis and Vidal than

1,5

segment

times as

long as

Sarmiento segment

Head wi th OL

3 .... 2

long as OPL •••••••••••••••••••••••••••••••••••••

. . . . . . . . . . . . . . . . . . . 3. macilentus De Santis and Vidal -

3;

long as OL . . . . . . . . . . . . . . .

longer

Sarmiento

than OPL ••••••••••••••••••••••••••••••••••••

4. rufus Ri chards

703

Thaumatodry i n i nae

\

\ \

\

\ \

\

\ \

\

:.:

...

Fig. 468 - Radial cells of females of Thaumatodryinus clarus n. sp. (type) (A) and variegatus De Santis and Vidal Sarmiento (type) (B)

The males of Th. n.

sp.

and minimus n.

variegatus De Santis and Vidal Sarmiento, clarus sp.

are unknown

1. Thaumatodryinus variegatus De Santis and Vidal Sarmiento

704

Mem.

Amer.

Ent.

I nst.,

no.

37,

1984

Thaumatodryinus variegatus De Santis and Vidal Sarmiento 1974: 25.

FEMALE:

fully

reddish;

antennae brown,

medial

winged;

length 7,18 mm; with joints,

region of segment 7,

head,

thorax and propodeum half of segments 5-6,

proximal

segments 8-10 testaceous;

petiole black;

abdomen reddish-testaceous; legs testaceous, with clubs of femora brown, distal half of tibiae black; antennal segment 3 more than 1,5 times as long as segment 1 (22:13); head dull, granulated; frons with a track of median keel;

occipital

POL = 5;

carina complete;

3; POL nearly as long as breadth of ocelli pronotum dull,

hairy,

with a strong

OL = 3;

(5:5);

transversal

OOL = 10;

temples

OPL

invisible;

impression,

with numerous

irregular keels; pronotal tubercles reaching tegulae; scutum shiny, granulated and with numerous irregular weak keels; notaul ices complete, posteriorly separated; reticulate rugose, wing

longitudinal

with first recurrent vein

fore-wing hyaline, vein

scutellum and metanotum shiny,

without

longer than distal

without dark transversal

longer than proximal

part

punctate;

propodeum

keels on posterior surface;

(30:12);

bands;

radial

fore-

part of discocubitus;

cell

distal open

part of radial (Fig.

468 B);

26: 3,5: 5,5: 19: 26; enlarged claw (Fig. 469) with 2 subapical teeth and with 26 lamellae; segment 5 of front tarsus (Fig. 469) with 2 rows of 53 lamellae; apex with a group of 4 lamellae; tibial spurs 1,1,2. fore tarsal

segments

MALE: unknown LOCUS TYPICUS:

in following proportions:

Dos de Mayo (Argentina)

TYP ICAL MATER IAL:

holotype F!

in PT.

HOSTS: unknown DISTRIBUTION: Neotropic region:

ARGENTINA:

Dos de Mayo (Misiones),

PT!

2. Thaumatodryinus minimus n. sp.

FEMALE:

fully

winged;

reddish-testaceous; black;

length 4,12 mm;

mesothorax;

head,

antennae and prothorax

metathorax and propodeum black;

abdomen brown-testaceous;

legs testaceous,

petiole

with tibiae and stalk of

fore femora brown; antenna I segment 3 sl i ght I y longer than segment 1 (11:9); read dull, granulated, with tracks of weak irregular striae, convex; frons without median and lateral keels; occipital carina complete; POL = 6; OL = 4; OOL = 10; OPL = 2; TL = 3; pronotum dull, granulated, with a strong scutum dull,

transversal granulated;

impression;

scutellum and metanotum dull,

distal open;

fore-wing

vein

segments

tegulae;

without sculpture among

without transversal

with a dark transversal

part of radial fore tarsal

tubercles reaching

posteriorly separated;

weakly punctate,

punctures or very reticulate rugose, keels;

pronotal

notaulices complete,

or longitudinal

band beneath the pterostigma;

longer than proximal

part

in following proportions:

(26:15); radial cell 23:

3:

7:

19:

30;

Thaumatodry i n i nae

Fig.

469 -

Chela of Thaumatodryinus variegatus De Santis and Vidal Sarmiento (type)

705

706

Mem.

Fig.

(Fig.

segment 5 of front

Ent.

470)

no.

with two subapical

tarsus

(Fig.

470)

apex with a group of 3 lamellae; MALE:

Ins!.,

37,

1984

Chela of Thaumatodryinus minimus n.

470 -

enlarged claw

Amer.

sp.

(type)

teeth and with 25

with 2 rows of nearly 44

tibial

lamellae; lamellae;

spurs 1,1,2.

unknown

LOCUS TYP I CUS: Tucuru i (Brazi I ) TYPICAL MATERIAL: holotype F! in TW HOSTS:

unknown

DISTRIBUTION:

Neotropic region:

BRAZIL:

Tucurui

(Para),

TW!

3. Thaumatodryinus macilentus De Santis and Vidal Sarmiento

Thaumatodryinus macilentus De Santis and Vidal Sarmiento 1974: 24.

FEMALE: space,

fully

winged:

clypeus and

4-10 darkened;

length 6,25-6,87 mm;

ma~dibles

reddish;

brown;

antenna I tufts: incomplete, OPL

transversal tegulae;

petiole black;

with segments mesothorax,

abdomen reddish;

legs

with clubs of femora and partly clubs of hind femora black-

carina vertex;

with malar

propectus and pronotum redd ish-testaceous;

meta thorax and propodeum black; testaceous,

head black,

antennae testaceous,

fig.

471;

head shiny,

granulated,

visible only behind the ocelli

longer than OL; impression,

scutum hairy,

pronotum sh i ny,

weakly punctate; shiny,

reaching at

and metanotum shiny,

weakly punctate;

ha i ry,

pronotal

granulated,

notaulices incomplete,

least 0,5

and on

swollen;

occipital

the sides of the

crossed by

a

tubercles reaching

with numerous irregular keels; length of scutum;

scutellum

propodeum reticulate rugose;

fore-

T ha uma todryi n i nae

Fig.

471

707

Tuft of long hairs on antenna of female of Thaumatodry inus macilentus De Santis and Vidal Sarmiento from Brazi I -

708

Mem. Amer.

Fig.

472 -

En!.

Inst.,

no.

37,

1984

Chela of Thaumatodryinus macilentus De Santis and Vidal Sarmiento from Nova Teutonia

(Brazil)

709

Thaumatodry in inae

Fig. 473 - Male genitalia of Thaumatodryinus macilentus De Santis and Vidal Sarmiento from Nova Teutonia (Brazi I): right half removed.

wing with a brown transversal distal part of radial

vein much

band

(rarely band very

longer than proximal

light)

part;

and with

radial

cell

710

Mem.

Amer.

En!.

closed;

pterostigma brown dark;

bands;

segment 1 of front

produced into a hook; lamellae

(Fig.

lamellae; MALE:

472);

Ins!.,

no.

winged;

1984

hind wings without dark transversal

tarsus nearly as

long as segment 4;

enlarged claw with 2 subapical

segment 5 of front

tarsus

apex with a group of 6 lamellae;

fully

37,

length 4,87;

(Fig.

tibial

head reddish,

segment 3

teeth and with 40

472)

with 2 rows of 67

spurs 1,1,2.

with ocellar region black;

antennae testaceous, wi th segments 3-10 dark; thorax and propodeum reddish, with scutellum and metanotum black, scutum dark; abdomen brown,

with segment

tibiae and tarsi

1 testaceous;

brown;

in following

proportions:

granulated;

smooth;

carina complete; dull,

legs testaceous,

8:

frons

7:

13: 23:

19:

OL = 2;

OOL = 7;

notaulices complete,

distance between the notaulices nearly as smooth, without

19:

without median and

POL = 7;

fully granulated;

notau Ii ces and

with mid and hind

antennae not distally thickened;

the paraps i da I furrows;

17:

18:

lateral

antennal

14:

17;

keels;

OPL = 2;

segments

head,

fully

occipital

TL = 4;

scutum

posteriorly separated;

minimum

long as distance between

the

scutell um and metanotum sh i ny,

without sculpture, finely hairy; propodeum reticulate rugose, dull, transversal or longitudinal keels; fore-wing hyaline, without dark

transversal bands; distal part of radial vein longer than proximal (36:21); genital ia: fig. 473; tibial spurs 1,1,2. LOCUS TYP ICUS:

Loreto (Argentina)

TYPICAL MATERIAL: HOSTS:

part

holotype F!

and

1 paratype F!

in PT.

unknown

DISTRIBUTION: ECUADOR:

Neotropic region:

Napo

(Muyuna,

Tena),

PANAMA: BM!

Barro Colorado

Island,

SURINAME: Sumatraweg

LE!

(Lelydorp),

OL! BRAZIL: Campsite (Mato Grosso), BM! Jatai (Goias), TW! OT! Nova Teutonia (S.ta Catarina), BM! 12° 50' S 51° 47' W (Mato Grosso), BM! OL! ARGENT INA:

Loreto

(Misiones),

PT!

4. Thaumatodryinus rufus Richards

Thaumatodryinus rufus Richards 1951: 815. Thaumatodryinus rufus Richards: Krombein 1952: 99.

FEMALE:

fully

winged;

length 3,0-4,0 mm;

reddish-testaceous,

with petiole

black, clubs of fore and hind femora brown; antennae testaceous, with segments 4-10 darkened; mesothorax, metathorax and propodeum reddish dark; antennal segment 3 more than 1,5 times as long as segment 1 (25:14); head swollen, shiny, granulated, with frontal weak longitudinal striae;

occipital

carina

incomplete,

the sides of the vertex; shiny,

hairy,

pronotal numerous

crossed by

POL = 4; a

tubercles reaching

visible only benhind the ocell i and on

OL = 2;

transversal tegulae;

irregular weak striae;

OOL = 8,5;

impression,

scutum hairy,

OPL = 3;

pronotum

weakly punctate; shiny,

notaul ices incomplete,

granulated,

reaching

at

with

least

Thaumatodry i n i nae

711

\

Fig.

474 -

Chela of Thaumatodryinus rufus Richards from St. Augustine (Trinidad)

0,5 length of scutum; scutellum and metanotum hairy, shiny, weakly punctate; propodeum reticulate rugose; fore wings with a brown transversal band and with distal part of radial vein much longer than proximal part; radial cell closed; pterostigma brown dark; hind wings without dark transversal bands; segment 1 of front tarsus longer than segment 4 (27:21); segment 3 produced into a hook; enlarged claw (Fig. 474) with 2 subapical teeth and with 29 lamellae; segment 5 of front tarsus (Fig. 474) with 2 rows of 45 lamellae; apex with a group of 5 lamellae; MALE:

tibial

fully

spurs 1,1,2.

winged;

length 2,50-2,87 mm;

head reddish-testaceous,

with

712

Mem. Amer.

Fig.

475 -

Ent.

Ins!.,

no.

37,

1984

Male genitalia of Thaumatodryinus rufus Richards from St. Augustine

(Trinidad):

right

half removed.

ocellar region black;

antennae testaceous;

with scutellum black,

metanotum and propodeum dark;

testaceous;

testaceous;

legs fully

thorax and propodeum reddish, abdomen brown-

antennae not distally thickened;

antennal 11: 10: 9: 9: 7: 10; head dull, fully granulated; frons without median and lateral keels; occipital carina complete; POL = 5; Ol = 2; OOl = 5; OPl = 1,5; Tl = 2; scutum dull, granulated; notaulices complete, posteriorly separated;minimum distance between the notaulices nearly as long as distance between the notaulices and the parapsidal furrows; scutellum and metanotum shiny, smooth, weak I y punctate, without SCu I pture among punctures; propodeum reticulate rugose, without transversal and longitudinal keels; fore-wing hyaline, without dark transversal bands; distal part of radial vein longer than proximal part (19:12); genitalia: fig. 475; tibial spurs 1,1,2. lOCUS TYPICUS: St. Augustine (Trinidad) segments in following proportions: 7:

5:

8:

14:

Thaumatodry i n i nae

Fig. 476 -

713

Chela of Thaumatodryinus bruchi De Santis and Vidal Sarmiento from Las Cesas

(Argentina)

TYPICAL MATERIAL: holotype F! in BM. HOSTS: in Trinidad Ormenis sp. (Richards 1951). DISTRIBUTION: Neotropic region: TRINIDAD: 51. Augustine (51. BM!

Curepe

(Sta.

Margarita,

Circular Rd.),

OT!

OLl

George),

OL!

714

Mem.

Amer.

En!.

Inst.,

no.

37, 1984

5. Thaumatodryinus bruchi De Santis and Vidal Sarmiento

Thaumatodryinus bruchi De Santis and Vidal Sarmiento 1974: 22.

FEMALE:

fully

winged;

length 6,56-8,62 mm;

head reddish;

antennae

testaceous, wi th segments 3-10 darkened; propectus and pronotum redd i sh; thorax (except for scutum which is reddish) and propodeum reddish-brown or black, reddish;

sometimes with

irregular brown spots;

legs testaceous,

clubs of hind femora brown-black; as

long as segment 1 (38:20);

longitudinal behind = 13;

keels on frons;

the ocelli OPL = 2;

impression,

and on

petiole black;

abdomen

with clubs of fore femora and distal

antenna I segment 3 more than

head swollen,

occipital

carina

dull,

granulated,

weakly punctate;

sh i ny,

pronotal

1,5 times

with some

incomplete, only visible

the sides of the vertex;

pronotum ha i ry,

part of

POL = 5;

crossed by

OL = 3,5;

OOL

a transversa I

tubercles reaching tegulae;

scutum

hairy, dull, reaching at least 0,5 length of scutum; scutellum shiny, smooth; metanotum with lateral keels; propodeum reticulate rugose forewing with a brown transversal much

longer than proximal

dark;

band and with distal part of radial

part;

radial

cell

hind wings without dark transversal

closed; bands;

vein

pterostigma brown segment

1 of front

(37:30); segment 3 produced into a hook; enlarged claw with 2 subapical teeth and with 29 lamellae (Fig. 476); segment 5 of front tarsus (Fig. 476) with 2 rows of 44 lamellae extending continuously to the apex; tibial spurs 1,1,2. MALE: fully winged; length 3,37 mm; head reddish-testaceous; antennae brown, with segments 1-2 testaceous; ocellar region usually dark; tarsus

longer than segment 4

prothorax reddish-testaceous; brownish;

scutellum,

propodeum testaceous; testaceous,

scutum reddish-testaceous,

metanotum,

petiole black;

darkened;

head dull,

granulated;

7: 7:

11 :16:

legs

mid and hind tibiae,

antennae not distally thickened;

segments in following proportions:

sometimes

abdomen brown-testaceous;

with mid and hind stalks of femora,

and hind tarsi

with sides

propodeum and pleura black;

mid

antennal

14: 14: 12: 12: 11: 14;

frons without median and

lateral

keels;

occipital

carina complete; POL = 7; OL = 3; OOL = 6; OPL = 1,5; TL = 4; scutum dull, granulated; notaul ices complete, posteriorly separated; scutellum and metanotum shiny, punctate, very weakly granulated; propodeum reticulate rugose, without transversal or longitudinal keels; fore-wing hyaline, without dark transversal proximal

part

bands;

LOCUS TYPICUS:

Alta Gracia

TYPICAL MATERIAL:

(Roraima),

Horco Molle

part of radial

vein

longer than

(Argentina)

holotype F!

HOSTS: unknown DISTRIBUTION: Neotropic region: Surumu

distal

(26-15); genitalia: fig. 477; tibial spurs 1,1,2. and 1 paratype F! BRAZIL:

in PT.

Encruzilhada (Bahia),

OL!

TW!

ARGENTINA:

Las Cejas

(Tucuman),

TU!

Alta Gracia

(Cordoba),

(Tucuman), PT!

TW! OL!

La Paz

TU!

(Cordoba),

715

Thaumatodry i n i nae

'/

Fig.

477 -

Male genital ia of Thaumatodryinus

Sarmiento from La Plata

PT!

La Plata,

TW!

(Argentina):

bruchi De Santis and Vidal right half removed

OU

6. Thaumatodryinus clarus n. sp.

716

Mem. Amer. En!.

Ins!., no. 37,

1984

Fig. 478 - Chela of Thaumatodryinus clarus n. sp.

(type)

Thaumatodry i n i nae

FEMALE:

fully

winged;

length 4,52 mm;

testaceous,

antennae with segment 3 more than 1,5 times as head dull, complete; times as strong dull,

granulated; POL

= 9;

OL

impression;

granulated;

=

or longitudinal

transversal

bands;

proportions:

cell:

radial fig.

= 3;

OPL

(9:3);

keels;

occipital carina

= 5;

POL nearly three

pronotum hairy,

dull,

with a

tegulae;

posteriorly separated;

fore tarsal

hyaline,

scutum

scutellum and

propodeum reticulate rugose,

fore-wing

vein with distal

468 A;

TL

posterior tubercles reaching

weakly granulated;

transversal radial

12;

notaulices complete,

metanotum shiny,

part;

OOL

with petiole black;

long as segment 1 (25: 15);

frons without median keel;

= 3;

long as breadth of ocelli

transversal

717

without

without dark

part much longer than proximal segments in following

5: 12: 28: 43; enlarged claw (Fig. 478) with 2 subapical segment 5 of front tarsus (Fig. 478)

3:

teeth and with nearly 24 lamellae;

with two rows of nearly 409 lamellae extending continuously to the apex; tibial spurs 1,1,2. MALE: unknown LOCUS TYPICUS: Yepocapa (Guatemala) TYPICAL MATERIAL: holotype F! in AM. HOSTS:

unknown

DISTRIBUTION:

Neotropic region:

Chimaltenango),

GUATEMALA:

Yepocapa

(Dep.

AM!

AUSTRAL I AN SPEC I ES OF THAUMATODRYINUS

Two species are known

KEY TO THE SPECIES

FEMALES

Fore-wing with

shorter than proximal 1. flavus n. sp. Fore-wing with first recurrent vein longer than proximal part of cubi tus (Fig. 479 A) ••••••••••••• 2. koebelei R.C.L. Perkins

part -

of

cubitus

first

(Fig.

recurrent

vein much

479 B) •••••

MALES

-

brown . . . . . . . . . . . . . . . flavus n. sp.

Head,

thorax and propodeum most Iy or 1.

fully

Head,

thorax and propodeum mostly

fully

or

testaceous ••••••••••

718

Mem.

Fig.

479 -

Nadzab,

Amer.

Ent.

Ins!.,

no.

37,

1984

Females of Thaumatodryinus koebelei R.C.L. and flavus

New Guinea) (A)

CD = first recurrent vein;

n.

sp.

Perkins

(from Waris,

DE = proximal

(from

New Guinea) (B):

part of cubitus.

1. Thaumatodryinus flavus n. sp.

FEMALE

(Figs 19, 457):

yellow-testaceous,

fully winged;

clypeus and mandibles yellow; 5-10 darkened;

length 6,30-6,87 mm;

head brown or

with ocellar spot brown and occiput partly brown: antennae testaceous,

propectus black or brown;

or with sides and partly anterior collar brown; yellow-testaceous or brown-b lack;

with segments 4-8 or

pronotum fu II y

yellow-testaceous

scutum and scutellum

metanotum brown-reddish or brown-b lack;

propodeum and pet iole brown-b I ack or black;

abdomen fu II y

redd ish or

with tergite 1 partly black;

legs yellow-testaceous or with fore clubs of

femora brown;

dull,

head swollen,

visible only behind the ocelli than OL; weakly shiny,

pronotum shiny,

granulated; granulated;

hairy,

pronotal

granulated; occipital

and on

carina

the sides of the vertex;

crossed by

a transverse

tubercles reaching tegulae;

notaul ices complete,

incomplete, OPL

longer

impression,

scutum hairy,

posteriorly separated;

scutellum

and metanotum shiny, weakly punctate; propodeum reticulate rugose; forewing with first recurrent vein much shorter than proximal part of cubitus (Fig.

479 B);

fore-wing

without dark transversal

brown dark and with distal part of radial part;

radial

cell

closed;

bands,

with pterostigma

vein much longer than proximal

segment 1 of front

tarsus

longer than segment 4

(35:32); segment 3 of front tarsus produced into a hook; enlarged claw (Fig. 480) with 2 subapical teeth and with 38-41 lamellae; segment 5 of front

tarsus

(Fig.

of 5-6 lamellae; MALE

480)

tibial

(Figs 28,458):

with 2 rows of 55-57

fully

a more or less wide frontal testaceous;

lamellae;

apex with a group

spurs 1,1,2. winged;

length 1,56-2,87 mm;

area testaceous;

occiput testaceous;

thorax and propodeum brown;

antennae brown, abdomen brown,

head brown,

with

clypeus and mandibles with segment 1 testaceous; wi th anterior region

T haumatodryi n i nae

Fig.

480 -

Chela of Thaumatodryinus

719

flavus n. sp. (type)

testaceous; legs testaceous; antennae not distally thickened; antennal segments in following proportions: 6: 6: 10: 13: 13: 13: 13: 11: 9: 10; head dull, hairy, granulated; frons without median and lateral keels occipital carina complete; POL = 5; OL = 2,5; OOL = 6; OPL = 1; TL = 3; scutum dull,hairy,

granulated;

min imum distance between

notaulices complete,

posteriorly separated;

the notau I ices of scutum shorter than distance

720

Mem.

Fig.

481

-

Islands) (A);

the notaulices and

metanotum shiny,

smooth,

distal

B:

Ins!.,

no.

1984

flavus n. sp. from Kira-Kira

furrows;

scutellum and

without sculpture among punctures;

without

transversal

posterior surface without

without dark transversal

37,

gonoforceps laterally viewed.

the parapsidal

punctate,

propodeum reticulate rugose, posterior surface; hyaline,

En!.

Male genitalia of Thaumatodryinus (Solomon

between

Amer.

bands;

keel

between dorsal

longitudinal

keels;

pterostigma lanceolate,

part almost reaching apex of radial

vein;

radial

and

fore-wing with

vein with distal

part longer than proximal part (29:13), almost reaching margin of wing; gonoforceps (Fig. 481) without proximal or distal processes, laterally viewed with a medial lOCUS TYPICUS:

TYPICAL MATERIAL: MM!

in TW;

prominence;

Tangtalau

holotype F!

6 para types

tibial

(Solomon

(1

F,

spurs 1,1,2.

Islands)

and 5 paratypes MM! 5 MM)!

in B;

4 paratypes

in Ol.

HOSTS: unknown DISTRIBUTION: Australian

region: NEW GUINEA: Wau, TW! all Jimmi Valley, all TW! Dumpu, TW! Waris (Hollandia, Western Irian), Ol! SOLOMON ISLANDS: B!

Ol!

Tangtalau

(Malaita

Island),

B!

Kira-Kira

(San Cristobal

Island),

Thaumatodryininae

Fig.

482 -

721

Chela of Thaumatodryinus koebelei R.C.L. Namatasopa

Perkins from

(New Hebrides)

2. Thaumatodryinus koebelei R.C.L. Perkins

Thaumatodryinus koebelei R.C.L. Perkins 1905: 58. Thaumatodryinus koebelei R.C.L. Perkins: Krombein 1952: 99.

FEMALE:

fully

testaceous,

winged;

length 3,47-6,00 mm;

lectotype fully

reddish-

with petiole black and antenna I segments 5-10 darkened;

other specimens head with ocellar spot brown, brown,

spots on si des of pronotum,

brown;

si des of scu tum brown ish;

transversal

in

impression

furrow between scutum and scutell um

furrow between scutell um and metanotum

722

Fig.

Mem.

483 -

Amer.

Ins!.,

no.

37,

1984

Male genital ia of Thaumatodryinus koebelei R.C.l. Nadzab

brown;

Ent.

(New Guinea):

right

propodeum brown or black abdomen brown;

femora brown;

head dull,

granulated,

Perkins from

half removed

swollen;

legs with clubs of fore

occipital

carina

incomplete,

visible only behind the ocelli and on the sides of the vertex; OPl longer than Ol; pronotum not humped, crossed by a transversal impression, hairy,

granulated;

pronotal

tubercles reaching

tegulae;

scutum dull,

hairy, granulated; notaulices incomplete (reaching at least 0,5 length of scutum) or complete (in a specimen from New Britain); scutellum and metanotum shiny,

weakly

punctate;

propodeum reticulate rugose fore-wing

with first recurrent vein longer than proximal part of cubitus (Fig. 479 A); fore-wing without dark transversal bands, with pterostigma yellow dark or brown and with distal part of radial vein much longer than proximal part; radial cell closed; segment 1 of front tarsus slightly longer than segment 4

(27:22);

with 2 subapical tarsus

(Fig.

lamellae;

482)

tibial

segment 3 produced

teeth and with 30 with 2 rows of 48

into a

lamellae lamellae;

(Fig.

hook; 482);

enlarged claw segment 5 of front

apex with a

group of 5

spurs 1,1,2.

MALE:

fully

brown

and antennal

winged;

length 2,25-3,62 mm;

testaceous,

segments 3-10 darkened;

with ocellar region

antennae not distally

I

Dry i n i nae thickened; 17:

16:

antennal

14:

12:

13;

segments in following proportions: head dull,

frons without median and 7;

=

OL

3;

=

OOL

7;

lateral

=

OPL

2;

keels;

=

TL

4;

keels;

fore-wing hyaline, vein

6:

10:

occipital

13:

carina complete;

scutum dull,

granulated;

scutellum and metanotum shiny,

propodeum reticulate rugose,

radial 1,1,2.

6:

15:

granulated and weakly reticulate rugose;

complete posteriorly separated; punctate;

723

without transversal

without dark transversal

longer than proximal

part;

genital ia:

bands; fig.

POL =

notaulices strongly

or longitudinal distal

483;

part of

tibial

spurs

LOCUS TYPICUS: Cairns (Australia) TYPICAL MATERIAL: lectotype F! in B. HOSTS:

in Australia

DISTRIBUTION: GUINEA:

Siphanta sp.

Australian

BUlolo, OL!

Lae-Zenag Road,

TW!

TW!

(R.C.L.

region: Bogia

(Om.

Busu R.

Perkins 1906c).

AUSTRALIA:

Cairns

New Guinea),

(Lae),

OL!

TW!

SUBFAMILY

Nadzab,

NEW HEBRIDES:

(Anelgaunat, Aneityum Island), BM! Namatasopa OL! NEW BRITAIN: Vunabakan (Keravat), B!

7.

(Queensland),

TW!

B!

TW!

NEW

OL!

Red Crest

(Espiritu Santo

Island),

DRYININAE KIEFFER

Dryininae partim Kieffer in Kieffer et Marshall 1906: 495. Dryininae partim Kieffer 1907b: 3. Lestodryinini partim Kieffer 1914b: 11. Dryinini partim Richards 1939: 189. Dryinini partim Muesebeck and Walkley 1951: 1034. Dryininae Richards 1953: 51. Dryininae partim Krombein 1979: 1243.

TYPE GENUS: FEMALE

Dryinus Latreille 1804

(Figs 495,

wing with costal,

I

pigmented veins palpi

529,

611,

632,

(Figs 611,

with 3 segments; front

612);

spurs 1,1,2 or 1,1,1MALE (Figs 531, 532, 613,

posterior veins

labial

occipital

fully

winged;

396 B);

with 6 segments;

fore-

lamella;

634,

palpi

segment 5 of front

680,

pronotal

705):

fully

with 3 segments;

dorsal

incomplete

(Fig.

tarsus

winged:

485);

(Fig.

tubercles present;

occipital

labial

larger from

carina complete or

mandibles with 3 teeth progressing

(Fig.

with costal,

703):

chela with rudimentary claw

long hairs;

never with one only preapical

incomplete;

678,

maxillary palpi

5);

tarsus chelate;

antennae without tufts of

with 6 segments;

662,

mandibles with 4 teeth progressing

anterior one to posterior (Fig. or absent;

659,

median and submedian cells clearly enclosed by

485)

tibial

maxi Ilary palpi carina complete or

larger from anterior one to

process of gonoforceps absent;

fore-wing

median and submedian cells clearly enclosed by pigmented

(Figs 532,613);

tibial

spurs 1,1,2.

724

Mem.

DISTRIBUTION:

Amer.

Ent.

Inst.,

no.

37,

1984

worldwide

HOSTS: Dictyopharidae,

Cixiidae, Lophopidae, Flatidae, Acanaloniidae, Issidae, Ricaniidae, Delphacidae, Tropiduchidae, Fulgoridae. GENERA:

8

SPEC I ES:

149

I<EY TO THE GENERA OF DRYININAE the fossi I genus Cretodryinus Ponomarenko)

(excluding

FEMALES

Enlarged

659, -

Enlarged

611,

claw more

than

three

660) . . . . . . . . . . . . . . . . . . . 632,

claw as

662,

long as

front

tibia

(Figs

Megadryinus Richards

or shorter than front tibia (Figs 528, 703) . . • . . . . • . . • . • • • • . • • . . • . • • . • • . . . . . . . . . . . 2

long as

678,

2 Enlarged claw very than arol ium

times as

4.

(Fig.

reduced,

nearly as

711) •••••

7.

long as or

sl ightly

longer

Perodryinus R.C.L. Perkins

reduced, much longer than arolium (Fig. 485).3 1 , 1 , 1 . . . . . . . . . . 2. Richardsidryinus Mocza r - Tibial spurs 1,1,2 •••••..••••••••.•••••••••••••.•••••••••.•.••. 4 4 Enlarged claw with one subapical tooth (Fig. 485) . . . . . . . . . . . . . . 5 -

Enlarged

3 Tib ia I

-

claw not

spu rs

Enlarged claw without teeth near apex (Figs

subapical

tooth

or with a

series of

small

669,691) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6

5 Notaulices at least partly visible . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1. Dryinus Lat rei I Ie Notaul ices invisible! . . . . . . 3. Tridryinus Kieffer 6 Notaulices at

least

partly

visible . . . . . . . . . . . . . . . . . . . . . . . . . . . . ..

. . . . • . . . . . • . . . • • . . . . . . • . • . .. 5. Alphadryinus n. g en. - Notaul ices invisible 1 . . . . . . 6. Mesodryinus Kieffer

NOTES:

the key doesn I t

inc I ude the fossi I genus Cretodryinus Ponomarenko,

because the only known specimen

is in

too bad conditions for

distinguishing generic differences.

! If the scutum

is sculptured by

longitudinal

keels,

two of the furrows

del imitated by the keels are considered notaul ices and the species is included in Dryinus Latrei lie or Alphadryinus n. gen. In some species with scutum reticulate rugose or smooth two keels at first sight can seem notaulices. I have considered, however, the notaulices invisible, the true notau I ices are furrows and not keel s.

because

Dryininae

725

MALES

I

Dryininae. The

was not able to find generic differences in males of

number of known males is too small

for drawing conclusions.

In addition

perhaps some males were associated by mistake with some genera. For the above mentioned reasons keys to males of species living

I think that actually only general

in a zoogeographic region are proposable,

as follows.

DRYININAE

KEY TO THE MALES OF THE KNOWN PALAEARCTIC

Scutum strongly as -

reticulate

rugose;

numerous areolae are as wide

ocelli •.•••••••••••••••••••••••••••••••••••••••••••••••••••• 2

Scutum granulated or weakly than

reticulate

rugose;

areolae

less wide

ocelli •••••••••••••••••••••••••••••••••••••••••••••••••••• 3

2 Posteri or

ocell i

not

areolae as wide as longer

than

touching

those of

segment

4,

less

the

occipi tal

scutum; than

carina;

antennal

three

times as

2; gonoforceps slightly shorter than penis

head wi th

3 slightly

segment

(Fig.

long as

segment

686) ••••••..••

· ............••.....•.••...•••. Mesodryinus niger (K i ef f er) -

Posterior ocell i areolae much approximately

touching

smaller

than

twice as

the

occipi tal

those of

long as

carina;

scutum;

segment

head wi th

antennal

segment

3

4, approximately four

long as segment 2; gonoforceps much shorter than penis (F i g. 684) •••••••.•••••••.••••• Mesodryinus dayi n. sp. 3 Fore wing with distal part of radial vein nearly as long as proximal part (Fig. 490 B) ••••• Dryinus tarraconensis Marshall times as

-

Fore wing wi th distal

part of radial vein much longer than 490 A) ••••••••••••••••••••••••••••••••••••• 4 4 Notaulices reaching approximately 0,5 length of scutum •••••.••• 5 - Notaulices reaching approximately 0,75 length of scutum •••••••• 7 5 Distivolsella farther from apex of penis (Fig. 491) . . . . . . . . . . . . . . . . . . . . . . . . • . . . . . . . . . . . . . . . . . . . Dryinus collaris (Linnaeus) - Distivolsella nearer apex of penis (Figs 501,618) . . . . . . . . . . . . . 6 6 Head more or less retiCUlate rugose ••••••.••.••••••••••••••••••• proximal

part

(Fig.

· . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Richardsidryinus corsicus (Ma rsha I I ) -

Head granulated,

not

reticulate

rugose ••.•••••••••••••••••••••••

· .•.........•...•.•..•.•..•..•. Dryinus koreanus (Moe za r) 7 Antennal

segment

3 nearly

twice as

long as

segment

2 . . . . . . . . . . ..

· ...•.••..•.••• , •.••.•.•....... Dryinus canariensis (Ceba I los) -

Antennal

segment

3 more

than

twice as

long as

segment

· •..........•..•..•............ Dryinus nepalensis n. sp.

2 .........

726

Mem.

NOTES: sex of

Amer.

Ent.

Ins!.,

no.

37,

1984

the male associated with D. canariensis could be also the opposite Richardsidryinus albrechti n. sp. On I y breed i ng experiments can

answer the question.

Dryinus sanderi n. sp., Nichardsidryinus maroccanus n. R. albrechti n. sp. (for the last species, however, see note

The males of sp.

and

above)

are unknown.

The fossil

Dryinus antiquus (Ponomarenko) was not included

species

into the key because the

type

(only known specimen)

is in

too bad

conditions for distinguishing differences with other males of Palaearctic

Dryininae.

KEY TO THE MALES OF THE KNOWN ETH I OP IAN

Notaulices

invisible;

scutum strongly

DRYININAE

reticulate

rugose,

with

numerous areolae as wide as

· . . . . . • . . . . . . . j\1esodryinus -

ocelly . . . . . . . . . . . . . . . . . . . . .. ethiopicus n. sp.

Notaulices visible; scutum granulated, not reticulate rugose; sometimes more or less weakly reticulate rugose, but in this case areolae much

less wide

than

ocel Ii •••••••••.•••••••••••••• 2

2 Occipital carina complete; dorsal part of occipital carina not touching eyes ••••••.••••••••••••••••••••••••••••.•••.••••.••••• 3 Occipital carina incomplete; dorsal part of occipital carina usually touching eyes ••••••••••••••••.•••••••.••••.•••••••••••• 4 3 Ninth abdominal sternite with apex rounded (Fig. 623} . . . . . . . . . . · . . . . . . . . . . . . . Richardsidryinus erraticus (Tu rn e r) - Ninth abdominal sterni te wi th apex emarginated . . . . . . . . . . . . .

· .....•....•.. Dryinus pretorianus n. 4 Head wi th OL

sp.

= OOl . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5

- Head wi th Ol more or less shorter than OOl ••••••. '" .•••••••••• 6 5 Fore-wing with radial cell closed; head with posterior ocelli not

touching

occipi tal

carina ••••••••••••••••••••••••••••••

••........•.• . Dryinus mahensis (Ki effer) -

Fore-wing with touching

radial

occipi tal

cell

open;

head with posterior ocelli

carina • . . • • • . • • • • . . • • •

· ...•....•••.. Dryinus saussurei (C eba I I os) 6 Fore-wing

wi th

radial

cell

open . . . . . . . . . .

. . . . . . • . . . . . . . Dryinus orophilus (Benoi t) -

Fore-wing with

radial

7 Frons with a median

cell

closed . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7

longitudinal

hump

from anterior ocellus

to

antennal

sockets ••••••••••••••.••••••••••••. . . . . . . . . . . . . . . Dryinus deceptor (Turner)

-

Frons

flat

in

the antennal 8 Breadth

of

front sockets

of

the anterior ocellus,

only

convex between

•..•.••••.••••.••••••••••••••••••••.••... 8

anterior ocellus

longer

than POL . . . . . . . . . . . . . . . . . . ..

.•....•••....• Lestodryinus luweli Benoi t

(see

incertae sedis nomi na)

Dryininae

- Breadth of

anterior ocellus

727

shorter

than POL . . . . . . . . . . . . . . . . . . ..

.......•• . Dryinus afer (Olmi)

KEY TO THE MALES OF THE KNOWN OR lENT AL DRYININAE

Dorsal

part

of

occipi tal

carina

touching

eyes •••••••••••••••••••

. . . . . . . . . . Dryinus indicus (Kieffer) -

Dorsal

part

of

2 Head wi th TL

occipital

shorter

carina not

than breadth

touching

of

eyes . . . . . . . . . . . . . . 2

ocel Ii •••••••••••••••••••••

· .•...•... Dryinus browni Ashmead -

Head with TL as

long as breadth

of

ocel Ii •••••••••••••••••••••••

· . . . . . . . . . Richardsidryinus pyrillae (K i ef fer)

KEY TO THE MALES OF THE KNOWN ~JEARCT I C DRYININAE

Head with breadth

of

ocelli

more

· •........ Tridryinus testaceus n. 2 Head wi th

ocell i

than

twice as

long as OPL . . . . . .

sp.

sl ight Iy broader

than OPL . . . . . . . . . . . . . . . . . . . . ..

· .......•. Perodryinus amoenus R. C • L. Perk ins

The male of Dryinus alatus check again

the

only known

(Cresson)

is known:

I,

however,

must

specimen.

KEY TO THE MALES OF THE KNOWN NEOTROP IC DRYININAE

Notaulices -

invisible •••••••••••••••••.••••••••••••••••••••••••• 2

Notaulices at

least

2 Posterior ocelli

partly

touching

visible . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 occipital

carina . . . . . . . . . . . . . . . . . . . . . .

· . . . . . . . . . Tridryinus ruficauda R i cha rd s Posterior ocell i

not

touching

occipital

carina . . . . . . . . . . . . . . . . . .

· . . . . . . . . . Tridryinus poecilopterae Richards 3

Scutum fully granulated,

not

reticulate

rugose •.•••••.••••••••••

· .......•. Dryinus antilleanus (Evan s ) -

Scutum fully

reticulate

4 Head with OPL

less

than

rugose . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 0,5 as

long as breadth

of

ocelli . . . . . . ..

· . . . . . . . . . Dryinus striatus (F en t on) -

Head

wi th OPL

5 Distal

part

of

sl ight Iy radial

shorter vein

than breadth

longer

than

· ....•.... Tridryinus striaticeps (K i ef fer)

of

proximal

ocell i . . . . . . . . . . 5 part . . . . . . . . . . . .

728

Mem. Amer. Ent.

Distal

part

of

radial

Ins!.,

no. 37, 1984

vein shorter than proximal

part ••••••.••••

. • • . . . • . . • • . . . . • . • • • • • . • • • • . • Dryinus surinamensis n. sp.

KEY TO THE MALES OF THE KNOWN AUSTRALIAN

DRYININAE

Notaul ices complete .•...•.... Dryinus areolatus n. sp. - Not a u I ice s i n c omp let e •••••••••••••••••••••••••.•••••.•••.••.•.• 2 2 Head granulated, not reticulate rugose ••••.••..•••••.••.••.•.••. •..•.•...•..•......••........ Dryinus pallidus (R.C.L. Perkins) - Head more or less reticulate rugose •••••••.••••••.•.•.••••..•• 3 3 Dorsal surface of propodeum approximately as long as metanotum .• .•.....••....•.••..•.•..•.•. . Dryinus aterrimus (Dodd) - Dorsal surface of propodeum longer than metanotum .•.••••••...•. 4 4 Scutellum granulated and punctate, not ret iculate rugose •.•••.•• .......•.....•.•..•.• .•••.•. . Dryinus australianus n. sp. - Scutellum granulated and reticulate rugose . . . . . . . . . . . . . . . . . . . . . 5 5 Scutum reticulate rugose, very rugose; areolae wide . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Dryinus koebelei (R.C.L. Perkins) - Scuum reticulate rugose, less rugose; areolae less wide ••••••••• . . .• .. • . . . . • , ...•..•..•..•... Dryinus venator (R.C.L. Perk i ns)

1. Dryinus Latrei lie

Dryinus Latrei lie 1804: 176. nee Dryinus Fabricius 1805: 200. Dryinus Latreille 1805: 228. Campylonyx Westwood 1835: 52 (n. syn.); type species Campylonyx ampuliciformis Westwood 1835, monotypic. Paradryinus R.C.L. Perkins 1905: 53 (n. syn.); type species Paradryinus koebelei R.C.L. Perkins 1905, orig. desig. Chlorodryinus R.C.L. Perkins 1905: 57 (n. syn.); type species Chlorodryinus pallidus R.C.L. Perkins 1905, orig. desig. Dryinus Latreille partim: Kieffer in Kieffer et Marshall 1905: 69 Campylonyx Westwood: Kieffer in Kieffer et Marshall 1905: 84. Plastodryinus Kieffer in Kieffer et Marshall 1906: 496 (syn. proposed by Moczar 1965); type species Plastodryinus szepligetii (Kieffer 1905), orig. desig.

Dryinus Latreille partim: Kieffer in Kieffer et Marshall 1906: 497. Hesperodryinus R.C.L. Perkins 1907: 40 (n. syn.); type species Hesperodryinus arizonieus R.C.L. Perkins 1907, orig. desig. Lestodryinus Kieffer 1911a: 108, new name for Dryinus Latreille (syn. proposed by Richards 1937); lype species Lestodryinus formiearius (Latreille 1804)(first species included in Dryinus

729

Dryininae

Latreille).

Lestodryinus Kieffer: Kieffer 1914b: 20. Campylonyx Westwood: Kieffer 1914b: 43. Dryinus Latrei lie: Richards 1935: 144. Dryinus Latreille: Richards 1937: 118. Dryinus Latrei lie: Richards 1939: 229. Mesodryinus Kieffer: Muesebeck and Walkley 1951: 1034. Hesperodryinus R.C.L. Perkins: Muesebeck and Walkley 1951: 1034. Hesperodryinus R. C. L. Perk i ns: Krombe i n 1979: 1242. Mesodryinus Kieffer: Krombei n 1979: 1243.

Dryinus iormicarius Latreille 1804, first included species

TYPE SPECIES: (Latreille 1805) FEMALE

(Figs 495,

segments;

labial

528,

palpi

529):

as or shorter than front subapical visible;

tooth

(rarely

pronotal

fully

tibia, two)

much

(Figs 531,

labial

palpi

532):

fully

tibial

winged;

with 6

(Fig.

longer than arol ium,

and some lamellae;

with 3 segments;

DISTRIBUTION:

maxi Ilary palpi

enlarged claw

tubercles reaching or not

complete or incomplete or absent; MALE

winged;

with 3 segments;

485)

notaulices at

tegulae;

as long

with one

occipital

least partly carina

spurs 1,1,2.

maxi Ilary palpi

with 6 segments;

for morphology see fami Iy characteristics.

worldwide

Dictyopharidae, Cixiidae, Flatidae, Acanaloniidae, Issidae, Ricaniidae, Delphacidae, Fulgoridae. HOSTS:

SPECIES: NOTES:

87.

on a different I

(Chlorodryinus for instance) were based

some synonym ized genera

have seen,

variable with

length of occipital however,

that the

carina

(complete or incomplete).

length of occipital

intermediate cases from occipital

Generic differences so are not proposable.

carina can be very

carina absent to complete.

The shape of posterior margin of

head also is variable, with intermediate cases from occiput concave to straight or convex (Hesperodryinus). In addition the posterior ocelli can be located on an front or behind,

imaginary

I ine connecting

with numerous

As the type of D.

hind margins of eyes or in

intermediate cases.

iormicarius Latreille seems lost and the original

description is unreliable, the diagnosis of Dryinus is based on classical opinions of Kieffer (1914b) and Richards (1939).

PALAEARCTIC SPECIES OF DRYINUS

Seven species are known.

KEY TO THE SPECIES

the

730

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

FEMALES

Occipital

margin

incomplete,

only dorsally visible

............................ -

Occipital

margin

complete

6.

(Fig.

2 Pronotum with weak posterior short

posterior collar

propodeum nearly as -

a

long as dorsal

long posterior collar

transversal

shorter

impression and wi th a

posterior surface of

surface;

fore-wing

hyal ine or

(Fig.

484 A,B,C);

than dorsal

posterior surface of

surface;

fore-wing

wi thout

dark

transversal

bands;

head and

sanderi n. sp.

prothorax mostly black . . . . . . 1. Fore-wing wi th dark transversal

bands;

redd ish •••••••••••••••••••••

canariensis (Ceba I los)

4 Transversal 488 A); furrow

(Fig.

488 B);

2.

head and prothorax

fully

furrow between metanotum and propodeum narrow

metanotum at

Transversal

wi th dark

bands ••••••••••.•••...••••.•••..••••••.•••.•••••••• 4

3 Fore-wing hyal ine,

-

4840);

wi th dark transversal bands •••••••••••••••••••••••••••••••••••• 3 Pronotum with a strong posterior transversal impression and with propodeum much

-

499 A) •

492) . . . . . . . . . . . . . . . . . . . . . . . . . . . 2

transversal

(Fig.

(Fig.

koreanus (Moczar)

least

five

488 A) . . . . . . . . .

3.

times as

long as breadth

this

collan's (Linnaeus)

furrow between metanotum and propodeum broad

metanotum nearly

(Fig. of

three-four

times as

(Fig.

long as breadth

of

thi s furrow (Fig. 488 B) •••.••••••••••••••••••••••••••••••••••• 5 5 Temples more prominent (Fig. 492 A); TL nearly twice as long as OL -

(Fig.

Temples (Fig.

492 A) . . . . . . . . . . . . . less prominent

(Fig.

492 B) ••••••••••••••••

The female of the fossil

4.

nepalensis n.

492 B); 5.

TL

sp.

slightly

longer

than OL

tarraconensis Marshall

species D.

antiquus (Ponomarenko) is

unknown.

MALES (excluding the fossi I Dryinus antiquus Ponomarenko)

Fore-wing with distal part of radial vein nearly as long as proximal part (Fig. 490 B) •• 5. tarraconensis Marshall -

Fore-wing proximal

with distal part

2 Notaulices -

Notaulices

(Fig.

part

of

radial

vein much

reaching approximately

0,5

reaching

0,75

3 Oistivolsella

longer

than

490 A) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 approximately

farther

from apex

length

of

scutum . . . . . . . . . 3

length of

of penis

(Fig.

scutum . . . . . . . . 4

491) . . . . . . . . . . . ..

3. collaris (Linnaeus) -

Oistivolsella nearer apex

of penis

(Figs.

487,

494) . . . . . . . . . . . ..

Dryininae

731

. . . . . . . . . . . . . . . . . . . . . . . . . . . . 6. koreanus (Moczar) segment 3 nearly twice as long as segment 2 • . . . . . . . • . . . . • . . . . . . • . . . . . . . •. . . . . • . . . .. 2. canariensis (Ceba I los) - Antennal segment 3 more than twice as long as segment 2 . . . . . . . . . 4. nepalensis n. sp.

4 Antennal

NOTES:

the male of D.

The

fossil

sanderi n. sp. is unknown. antiquus (Ponomarenko) isn't present in the

species D.

key because the only known specimen

is in

too bad conditions for

distinguishing differences with I iving species. The male associated in the key with D. canariensis could be also the opposite sex of Richardsidryinus albrechti n.

sp.

Only breeding

experiments could answer the question.

1. Dryinus sanderi n. sp.

FEMALE: fully winged; length 3,75 mm; head black, with mandibles, clypeus and malar space testaceous; antennae fully testaceous; thorax and propodeum black,

with posterior collar of pronotum reddish-testaceous;

A

B

c Fig. sp.

(laterally viewed) of females of Dryinus nepalensis n. collaris (Linneaus) from France (B), tarraconensis Marshall from Montecatini (Italy)(C) and sanderi n. sp. (type)(D).

484 -

Pronotum

(type) (A),

732

Mem.

Fig.

485 -

abdomen black-brown;

Amer.

Chela of

Ent.

no.

37,

legs testaceous,

with coxae partly black or brown

antennae distally

in following proportions:

8:

6:

23:

14:

punctate,

frons,

where there are numerous irregular striae;

complete;

thickened;

10:

shiny,

incomplete keel,

1984

lJryinus sanderi n. sp. (type)

and with clubs of femora brown; segments

Inst.,

8:

6:

antennal

5:

4,5:

8;

head

weakly granulated and rugose mainly on anterior half of frons with a

not visible on anterior half of frons;

POL = 4;

Ol = 3;

OOl = 8;

OPl = 1,5;

median

occipital

Tl = 3;

carina

pronotum

(Fig.

484 0) with a very weak anterior transversal impression and with a strong posterior transversal impression; disc humped; posterior collar very short; pronotal

tubercles not reaching

granulated; numerous,

scutum shiny,

short,

posterior margin; of scutum; smooth;

weak,

very

tegulae;

pronotum shiny,

weakly

weakly granulated and rugose,

longitudinal

with

keels near anterior margin and near

notaulices incomplete,

reaching

scutellum and metanotum shiny,

propodeum reticulate rugose,

very

approximately 0,65

length

weakly granulated,

with dorsal

surface nearly as long as

posterior surface; posterior surface without longitudinal keels; fore-wing hyaline, without dark transversal bands; distal part of radial vein longer than proximal 17: with

3:

6:

14:

part 24;

1 row of 7

of 19 lamellae; 1,1,2. MALE:

unknown

(14:7);

fore tarsal

enlarged claw lamellae;

(Fig.

segments 485)

segment 5 of front

apex with a

in following proportions:

with a

group of nearly

tarsus

subapical (Fig.

12 lamellae;

tooth and

485) tibial

with 2 rows spurs

Dryininae

LOCUS TYPICUS: Melnik (SW Bulgaria) TYPICAL MATERIAL: holotype F! in OL. HOSTS: in Bu I gari a Hysteropterum latifrons DISTRIBUTION: Palaearctic region: BULGARIA:

Fig. 486 -

Chela of

733

Melnik, OL!

Dryinus canariensis n. sp. from Tahodio (Canary Islands)

734

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

2. Dryinus canariensis (Ceballos) n. comb.

Paradryinus canariensis Ceba Ilos 1927: 101.

FEMALE:

fully

legs reddish;

winged;

length 4,25-4,50 mm;

mesothorax,

metathorax,

head,

antennae,

prothorax,

propodeum and petiole black;

abdomen brown, with segment 1 reddish; tegulae testaceous; antennae not distally thickened; antenna I segments in following proportions: 13: 5: 20: 9:

9:

6:

6:

6:

5:

8;

head dull,

smooth,

granulated;

frons with a median

longitudinal keel, without lateral keels; occipital carina complete; POL = 4; OL = 3; posterior ocelli not touching occipital carina; OPL nearly as long as breadth of ocell i; granulated, impression

partly and

collar short,

temples prominent;

without sculpture,

with

a

strong

nearly 0,5 as

pronotum shiny,

with an

posterior transversal

long as pronotal

partly

anterior weak transversal impression;

tubercles;

posterior

pronotal

tubercles

not reaching tegulae; scutum dull, hairy, granulated and with numerous weak areolae; notaul ices incomplete, reaching approximately 0,65 length of scutum; weakly

scute II um and metanotum sh iny, granulated;

strongly transversely striate, dark transversal

smooth,

wi thout scu I pture or very

propodeum reticulate rugose,

bands,

without

two on

with posterior surface

longitudinal

the basal

keels;

pterostigma;

distal

part of radial

(13:7);

tarsal

segments in following proportions:

fore

enlarged claw

(Fig.

segment 5 of front same length; MALE

testaceous;

(Fig.

incomplete; carina;

legs brown,

with

=

granulated;

proximal

part

broadened;

winged;

tibial

5;

OL

=

3;

OOL

with ventral

light;

=

3;

scutum dull,

(12:7);

granulated;

part of radial

gonoforceps (Fig.

ninth abdominal

sternite:

black;

side of segment 10: 8:

487 A)

fig.

8:

occipital

posterior ocelli

length of scutum;

distal

part

15: 24;

antennae not distally thickened;

propodeum reticulate rugose; bands;

7:

spurs 1,1,2.

length 2,50 mm;

frons with a median keel;

temples invisible;

dark transversal

tarsi

15: 4:

tooth and with 12 lamellae;

in following proportions: 8,5: 4:

granulated;

POL

fully

with 3

with 2 rows of nearly 27 lamellae

antennae black,

reaching approximately 0,75 dull,

486)

longer tha proximal

apex with a group of 12 lamellae;

antenna I segments head dull,

vein much

with a subapical

(see note on page 731):

mandibles testaceous;

9;

486)

tarsus

fore-wing

cells and one beneath the

8:

8:

6:

6:

carina

touching occipital

notaulices incomplete,

scutellum and metanotum fore-wing

hyaline,

vein much

with distal

487 B;

tibial

without

longer than region

spurs 1,1,2.

LOCUS TYPICUS: La Cuesta (Canary Islands) TYPICAL MATERIAL: holotype F! in MD. HOSTS: unknown DISTRIBUTION: Palaearctic region: OL!

TN!

B.co de Marchar

(Tenerife) (F), Tenerife) (F),

MO! TN!

CANARY

(San Sebastian,

Tahodio

(Tenerife) (F),

ISLANDS:

B.co Santos

Gomera)(F), MD! MO!

(F),

MD!

La Cuesta

Los Calderones

(Medano,

Dryininae

735

Fig. 487 - Male genital ia of Dryinus canariensis n. sp. from Los Calderones (Canary Islands) (A: left half removed; B: ninth abdominal sternite) .

3. Dryinus collaris (Linnaeus)

Sphex collaris Linnaeus 1767: 946. Dryinus formicarius Latreille 1805: 228 (syn. proposed by Fitton, Graham,

Boucek, Fergusson,

Huddleston, Quinlan, Richards 1978);

locus typicus Southern France. Campylonyx ampuliciformis Westwood 1835: 52 (n. syn.); locus typicus Lyon (France)

Dryinus formicarius Latrei lie var. : Kieffer in Kieffer et Marshall 1905: 78.

Dryinus formicarius Latreille var. : Kieffer in Kieffer et Marshall 1905: 79.

Lestodryinus formicarius (Latreille): Kieffer 1914b: 20. Lestodryinus corsicae Kieffer 1914b: 21 (n. syn.); locus typicus Corse (France).

Lestodryinus formicarius (Latrei lie): Berland 1928: 140. Lestodryinus corsicae Kieffer: Berland 1928: 141.

736

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

488 - Metanotum of Dryinus collaris (L.) from Le Puy (France )(A) and tarraconensis Marsha II from Orleans (France) (B). In metanotum of tarraconensis the anterior half is rugose and inclined. 5 ; transversal

Fig.

furrow between metanotum and propodeum.

Campylonyx ampuliciformis Westwood: Berl and 1928: 142. Lestodryinus formicarius (Latreille): Picard 1932: 29. Lestodryinus formicarius (Latrei lie): Maneval 1935: 4. Dryinus formicarius Latrei lie: Richards 1937: 118. Dryinus formicarius Latrei lie: Richards 1939: 229. Dryinus formicarius Latrei lie: Berland 1940: 121. Dryinus formicarius Latrei lie: Abdul-Nour 1974: 1015. Dryinus formicarius Latreille: J.I'. Perkins 1976: 27. Dryinus formicarius Latrei lie: Olmi 1976: figs 10-14. Dryinus collaris (L i nnaeus): F i !ton, Graham, Boucek, Fergusson, Huddleston,

Quinlan,

Richards 1978:

127.

Dryinus formicarius Latreille: Currado and Olmi 1979b: 4. Dryinus collaris (Linnaeus): Day 1979: 60.

FEMALE: fully winged; length 5,12-6,25 mm; head reddish-testaceous, with a brown small or wide spot on frons and vertex; antennae testaceous; propectus, pronotum, scutum, scutellum, metanotum, mesopleura, mesosternum, metasternum and metapleura reddish-testaceous; sometimes brown

spots on mesosternum,

metasternum,

between mesopleura and metapleura;

mesop leura and on furrow

rarely pronotum mostly black;

scutell um and metanotum someti mes reddi sh-dark;

mesosternum,

metasternum, mesopleura and metapleura sometimes almost fully brownblack; rarely scutum, scutellum and metanotum fully black; propodeum black,

with apex reddish;

fully reddish-testaceous, head dull,

granulated;

petiole black;

ocellar region swollen;

OPL shorter than breadth of ocelli; B)

crossed by

abdomen brown-reddish;

legs

sometimes with mid and hind coxae brownish;

two transversal

collar usually smooth and

impressions,

shiny;

without sculpture;

impression with short longitudinal

occipital carina complete;

temples prominent;

keels;

pronotum

(Fig.

484

disc and posterior

posterior transversal

anterior collar and sides weakly

Dryininae

Fig.

489 -

granulated; reaching dull;

Chela of

sides with a

tegulae;

Dryinus collaris (L.) from Dornbach (Austria)

few

longitudinal

scutum shiny,

notaul ices distinct and

of scutum;

737

keels;

pronotal

tubercles not

weakly punctate and granulated,

incomplete,

reaching

posterior third of scutum with

weak

rarely

approximately 0,5

longitudinal

length

short keels;

scutellum usually flat, shiny, smooth, weakly punctate; metanotum usually weakly punctate, smooth, always without transversal furrow in posterior half (Fig.

488 A);

propodeum with transversal proximal

tarsus

segment 1 of front with subapical (Fig.

489)

reticulate rugose;

longitudinal keels;

bands and with distal

part;

enlarged claw of front

propodeum fully

two short

part of radial tarsus slightly

tooth and

11

vein much

longer than

longer than segment 4;

lamellae

with 2 rows of 27

posterior surface of

fore-wing with 2 brown

(Fig.

lamellae;

489);

segment 5

apex with a

group

Mem.

738

Amer.

Ent.

Inst.,

no.

37,

1984

A

B Fig.

490 -

Radial

(Italy)(A)

of 19 lamellae; MALE:

fully

and

veins of males of Dryinus collan"s (L.) from Rosta tarraconensis Marshall from Orleans (France)(B).

tibial

spurs 1,1,2.

winged;

length 5,00 mm;

black,

with

tegulae and

legs

testaceous; mandibles with 3 teeth; scutum dull, hairy, granulated; notaulices incomplete, reaching approximately 0,5 length of scutum; scutellum and metanotum shiny,

smooth,

weakly punctate;

propodeum fully

reticulate rugose, without longitudinal keels on posterior surface; forewing with distal part of radial vein much longer than proximal part (20:10)(Fig.

490 A),

without dark transversal

the apex of penis (Fig. LOCUS TYPICUS:

491);

tibial

bands;

distivolsella far from

spurs 1,1,2.

Spain

holotype F of S. collaris in Ll; holotype F of D. formicarius probably lost; lectotype F! (hereby designated) and 1 para lectotype F! of C. ampuliciformis in P; lectotype F! (hereby designated) of L. corsicae in MT. HOSTS: in France Cixius nervosus (L.)(Maneval 1935); in England Cixius contaminatus Fl. (Capron 1885); in Italy Issus coleoptratus (F.) (Olmi 1976); in Germany Cixius nervosus L. (Haupt 1932) (quotation of Dictyophara europaea L. in Richards 1939 valid for D. tarraconensis Marshall); in addition Tachycixius pilosus (OI.)(Thompson 1955). TYPICAL MATERIAL:

DISTRIBUTION:

Palaearctic region:

U.

S.

S.

R.

? (Ponomarenko 1978);

Dryininae

Fig.

491

HUNGARY:

-

Male genitalia of

Zelenika,

BU!

739

Dryinus collaris (L.) from Rosta (Italy)

YUGOSLAVIA:

Opatija

(Istra), BU!

Krapina

(Hrvatska), BU! GERMANY: Aix-Ia-Chapelle (Kieffer in Kieffer et Marshall 1905); AUSTRIA: Werfen (Slzb.), LE! Dornbach (Wien), P! BELGIUM (Kieffer in Kieffer et Marshall 1905); SWITZERLAND: Peney, GV! Geneve, SP! FRANCE: Mont-de-Marsan (Abdul-Nour 1974); Lyon (Rhone), P! St. Gely-duFesc (Herault), P! St. Guilhem (Herault), P! Digne (Basses-Alpes), P! Hyeres (Var), P! Le Puy, P! Toulon (Var), P! Les Guerraux (Saone-etLoire) (Berland 1928); Corse, MT! ENGLAND: Cobham (Surrey) (Richards 1939); Shere (Surrey)(Richards 1939; Capron 1885); SPAIN (Linnaeus 1767; Day 1979); ITALY: Rosta (Torino), OL! Gaibola (Bologna), OL! S. Olcese (Genova), BM! Lucca (Kieffer in Kieffer et Marshall 1905); Napoli (Kieffer

740

Mem.

Amer.

Ent.

Ins!.,

no.

37,

1984

A o Fig.

492 -

B

o

Heads

0

o

of females of Dryinus nepalensis n. sp. tarraconensis Marshall (lectotype)(B)

and

in Kieffer et Marshall the cocoon

1905); S.

is oval

elongate. According by

0

(dorsally viewed)

(type)(A)

NOTES:

o

(Fig.

to Berland

Basilio

(Mottola,

498 A),

whereas

(1940),

Taranto)(Moczar 1965).

D. tarraconensis it's

in

occasionally the female

is attracted

light traps.

4. Dryinus nepalensis n. sp.

FEMALE: fully winged; length 6,25-6,55 mm; head black, wit>, ",andibles and anterior margi n of c I ypeus testaceous; antennae brown, wi th segments 1-2 testaceous and segments 7-10 whitish; black;

legs reddish,

distally 26:

thickened;

18:

broad

13:

11:

(26:5),

complete;

10:

8:

12;

POL

=

2,5;

Ol

around disc;

pronotum

collar

long,

flat,

distinct;

=

granulated and with numerous

median complete keel;

2,5;

OOl

=

11;

OPl

=

484 A)

with a

pronotal

keels;

metanotum nearly keels;

fore-wing

Tl

=

3;

6:

50:

long as

longitudinal carina temples:

fig.

impression; tegulae;

notaulices

scutellum dull,

posterior scutum dull,

incomplete, granulated;

wi th a strong transversa I furrow near scutell um and wi th

furrow near propodeum; transversal

occipital

tubercles not reaching

transversa I furrow near propodeum;

granulated;

15:

weak anterior transversal

almost reaching posterior margin of scutum; a strong

2;

strong posterior transversal

granulated and with some areolae near margins; metanotum du II,

antennae

in following proportions:

weakly granulated and with numerous striae (Fig.

impression and with a

propodeum and abdomen

antenna I segment 4 nearly five times as

frons with a

pronotum shiny,

thorax,

tibiae and part of coxae brown;

antenna I segments

head dull,

keels on frons; 492 A;

with

propodeum dull,

posterior surface with

with

and another beneath

centra I reg ion short and

three-four times as two short

two dark transversal the pterostigma;

long as breadth of

reticulate rrgose,

distal

bands,

without

incomplete longitudinal one on the basal

part of radial

vein

cells

longer

Dryininae

Fig.

than proximal

Chel a of Dryinus nepalensis n.

493 -

part

741

(19:10);

fore tarsal

sp.

(type)

segments in following proportions:

16: 3: 6: 15: 24; segments 2 and 3 of front tarsus produced into hooks; enlarged claw (Fig. 493) with a subapical tooth and with a row of 15 lamellae;

segment 5 of front

lamellae;

apex with a group of nearly 18 lamellae;

MALE:

fully

winged;

(partly black

in

tarsus

(Fig.

length 2,00-3,56 mm;

long specimens);

segments 1-2 testaceous; partly black or brown;

carina

(not touching occipital invisible;

antennae fully

testacoeus or brown,

legs testaceous, 7:

6:

12:

incomplete;

10:

with all

10:

11:

10: 9: 7:

in very

9;

segments

head dull,

touching occipital

long specimens);

scutum dull,

reaching approximately 0,75

sometimes with a strong

antennal

frons with a median complete

scutellum and metanotum as sculptured as scutum; rugose,

with

coxae or only hind coxae

posterior ocelli

carina only

spurs 1,1,2.

mandibles testaceous

POL = 5; Ol = 2; OOl = 4;

notaul ices incomplete,

tibial

black;

granulated and weakly reticulate rugose; occipital

with 2 rows of nearly 37

antennae not distally thickened;

in following proportions: keel;

493)

transversal

keel

carina

temples

as sculptured as head; length of scutum; propodeum reticulate

between dorsal

and

742

Mem.

Fig.

494 -

Amer.

Male genital ia of

Ent.

Ins!.,

no.

37,

1984

Dryinus nepaiensis n. sp. from Godavari

(Nepal) (left half removed)

posterior surface;

fore-wing

hyal ine,

without dark transversal

bands;

distal part of radial vein longer than proximal part (13:9 or 9:5 or 14:9); gonoforceps (Fig. 494)with distivolsella located near apex of penis; tibial spurs 1,1,2. lOCUS TYPICUS:

lothar

TYPICAL MATERIAL: (1

F,

2 MM)!

HOSTS:

(Nepal)

holotype F!

and 5 paratypes MM!

in OT,

3 paratypes

in Ol.

unknown

DISTRIBUTION: Godavari

Palaearctic region:

(Katmandu), OT!

Ol!

r,EPAl:

Pulchauki

lothar (Birganj), (Katmandu),

OT!

OLi

OTt

5. Dryinus tarraconensis Marshall

Dryinus tarraconensis Marshall 1868: 204. Dryinus szepligetii Kieffer in Kieffer et Marshall 1905: 77 (n. syn.);

Dryininae

Fig.

495 -

Female of Dryinus tarraconensis Marshall from Montecatini ( Italy)

743

744

Mem.

Fig. 496 -

Amer.

Ent.

Inst., no. 37,

1984

Chela of Dryinus tarraeonensis Marshall from Orleans (France)

locus typieus Novi (Yugoslavia) (nee Budapest: see Moczar 1965). Dryinus tarraeonensis Marshall: Kieffer et Marshall 1905: 82. Plastodryinus szepligetii (Kieffer): Kieffer in Kieffer et Marshall 1906: 496.

Dryinus tarraeonensis Marsha II: Kieffer in Kieffer et Marsha II 1906: 497.

Lestodryinus tarraconensis (Marshall): Kieffer 1914b: Plastodryinus szepligetii (Kieffer): Kieffer 1914b: 43. Lestodryinus tarraconensis (Marshall): Berl and 1928: Lestodryinus gregori Hoffer 1936: 164 (syn. proposed 1965); locus typicus Pouzdranyi (Czechoslovakia). Lestodryinus bidens Haupt 1937: 70 (n. syn.); locus Bellinchen

21. 141. by Moczar

typicus

(Germany).

Lestodryinus tarraconensis Marshall: Pi Ilault 1951: 67. Dryinus szepligetii Kieffer: Moczar 1965: 380. Dryinus tarraconensis Marshall: Moczar 1965: 381. Dryinus tarraconensis Marshall: Currado and 01 mi 1979b: 4. nee Dryinus szepligetii Kieffer: Ponomarenko 1981 b: 879 (see Dryinus stantoni Ashmead and Dryinus browni Ashmead).

FEMALE (Fig. 495): fully winged; length 5,12-8,25 mm; head usually black, with frons, part of vertex (more along orbits), clypeus and mandibles yellow-reddish; sometimes also partly temples reddish; occiput

Dryininae

Fi g.

745

Ma I e gen i ta I i a of Dryinus tarraconensis Marsha II from Orl eans

497 -

(France)

always reddish; and vertex; testaceous;

rarely head reddish-testaceous with brown spot on frons

antennae brown,

with segments 1-2 and sometimes also 3

segments 9-10 or only 10 yellow;

sometimes reddish-testaceous; and sides reddish;

with posterior collar

sometimes fully reddish-testaceous;

and metanotum usually black, mesosternum,

propectus usually black,

pronotum usually black, sometimes fully

scutum,

mesopleura and metapleura usually black,

less reddish-testaceous or fully reddish;

scutellum

reddish-testaceous; sometimes more or

propodeum usually black,

sometimes with apex reddish, rarely fully reddish; petiole black; abdomen brown-reddish or black-brown; legs usually testaceous, with coxae partly brown,

sometimes also with clubs of femora brown,

testaceous; longitudinal

head dull, keels;

granulated;

vertex swollen;

rarely fully reddish-

frons and vertex with tracks of POL

= 4; Ol

= 3;

OOl = 13;

OPl = 2,5;

746

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

A

C _______B-~ Fig.

498 -

Italy)(A)

Dryinus collaris (L.) (female from Rosta, tarraconensis Marshall (female from Orleans, France)(B)

Shapes of cocoons of and

TL = 7; occipital carina complete; OPL nearly as long as breadth of ocell i; temp I es: fi g. 492 B; pronotum (F ig. 484 C) crossed by 2 transversal impressions, shiny, granulated, with strong longitudinal keels on sides; posterior collar sometimes smooth and without sculpture; anterior impression sometimes weakly tegulae;

scutum shiny,

longitudinal

pronotal hairy,

keels on posterior third;

approximately 0,5 strong

visible;

granulated,

length of scutum;

longitudinal

metanotum with a propodeum fully

keels;

deep

rarely

with numerous short

notaulices incomplete, scutellum dull,

almost smooth,

transversal

reticulate rugose;

tubercles not reaching

rugose,

reaching

with numerous and

with weak keels;

furrow on posterior half (Fig.

488 B);

posterior surface with 2 short

longitudinal keels; fore-wing with 2 brown transversal bands and with distal part of radial vein much longer than proximal part; segment 1 of front

tarsus sl ightly

longer than segment 4;

tooth and 13 lamellae (Fig. 2 rows of 38

lamellae;

496);

enlarged claw with subapical

segment 5 of front

apex with a group of at

tarsus

least 15

(Fig.

lamellae;

496)

spurs 1,1,2. MALE:

fully

winged;

length 3,37-3,87 mm;

black,

with

tibial

with mandibles yellow

Dryininae

and tarsi carina

light;

fore tibiae yellow;

incomplete;

almost complete;

scutellum and metanotum shiny,

vein

dark transversal

granulated,

scutum dull,

occipital

hairy,

granulated;

reaching posterior third or fourth of scutum,

fully reticulate rugose, part of radial

head dull,

temples not visible;

notaul ices incomplete,

747

without

longitudinal

keels;

nearly as long as proximal

bands;

distivolsella

granulated; fore-wing

part

(Fig.

rarely

propodeum with distal

490 B),

located near apex of penis

without

(Fig.

497); tibial spurs 1,1,2. lOCUS TYPICUS, Torla (Spain) TYPICAL MATERIAL; holotype F! of D. tarraconensis in BU; holotype F! of L. gregori in Hoffer coli.; holotype F! of D. szepligetii in BU; holotype F! and allotype M! of L. bidens in HA! in VI there is the type M! of Labeo rufipes Forster (never described) without local ity label: it's a male of D. tarraconensis. HOSTS; Dictyophara europaea (l.) in France (Pi Ilault 1951) and in Germany (Richards 1939; under name Dryinus formicarius latrei lie; Haupt 1937: under name Lestodryinus bidens Haupt). DISTRIBUTION;

Palaearctic region;

U.

S.

S.

? (Ponomarenko 1978);

R.

HUNGARY; Kenese, BU! Crkvenica, Ol! ROMANIA: Nagyenyed (Transilvania)(under name Plastodryinus sziladyi n. sp.), BU! CZECHOSLOVAKIA; BU!

Sutomore

FRANCE;

Orleans

(Madrid), ITALY:

BM!

Sarre

(L.)

BM!

Parma,

BO!

P!

GERMANY; SPAIN;

S. NA!

Perugia,

(Caltanissetta),

Baghdad, NOTE:

GE!

(Hoffer 1936;

BU!

(Tarragona),

(Aosta),

(Pistoia),

Cataldo

(Loiret),

Salou

(Alessandria), Terme

Pouzdranyi

(Dalmacija),

CO!

Torla

OL!

Moczar 1965);

Bell inchen

BM!

(Huesca),

BU!

Benicasim (Cuneo),

Pietra Ligure

(Savona),

Maiella

GREECE;

(Abruzzo),

Kremasti

Hills

P!

Escorial

PL!

BM!

Gavi

PL!

NA!

Novi,

HA!

(Castellon),

Benedetto Belbo GE!

YUGOSLAVIA;

(Oder),

Montecatini

Messina,

(Rhodes),

BM!

CO!

WA!

the cocoon it's ovale.

(Fig.

498 B)

is very elongate,

Notes on biology by Pillault

whereas in

D. collaris

(1951).

6. Dryinus korean us (Moczar) n. comb.

Chlorodryinus koreanus Moczar 1983a; 192.

FEMALE:

fully

winged;

length 3,84 mm;

testaceous-light,

with petiole

black; antennae distally thickened; antenna I segments in following proportions: 10: 6; 33: 18; 15; 11: 9; 7; 7; 10; head (Fig. 499 A) swollen,

dull,

granulated;

frons with a

weak median complete keel;

occipital carina incomplete, visible only behind posterior ocelli and on sides of posterior ocelli; POL = 3; OL = 4; OOL = 8; OPL = 3; temples prominent;

pronotum shiny,

striae on sides,

without sculpture,

crossed by an

with some longitudinal

anterior and a posterior transversal

S.

IRAQ;

748

Mem.

Amer.

Ent.

A

Inst.,

no.

37,

B

o

1984

0

DQll) Fig.

499 -

Heads of Dryinus koreanus

from De-sang san, Wanuma,

impression;

Korea)

(Moczar) (A:

and pallidus

New Guinea;

D:

(R.C.L.

type female;

Perkins) (B:

C:

male

female from

male from Mulgrave, Australia)

anterior collar very short;

disc very

long;

posterior collar

short; pronotal tubercles not reaching tegulae; scutum granulated, with anterior surface of median area smooth and without sculptur0; notaulices incomplete,

reaching approximately 0,65

metanotum dull,

granulated;

transversal or longitudinal

length of scutum;

propodeum reticulate rugose, keels;

dorsal surface sl ightly

scutellum and without longer than

(27:20); fore-wing hyaline, without dark transversal bands; distal part of radial vein longer than proximal part (20:8); fore tarsal segments in fOllowing proportions: 24: 3: 6: 20: 29; enlarged claw (Fig. 500) with a subapical tooth and with a row of 11 lamellae; segment 5 of front tarsus (Fig. 500) with two rows of 17 lamellae; apex with a posterior surface

group of 7 lamellae; tibial spurs 1,1,2. MALE: fully winged; length 2,72 mm; head black,

with mandibles,

anterior region of frons and malar space testaceous; 1 testaceous

(other segments missing

and propodeum black; 499 C)

dull,

carina

incomplete,

in two only known specimens);

abdomen brown;

granulated;

legs yellow-testaceous;

frons with a median complete keel;

fully visible dorsally,

clypeus,

antennae with segment head

thorax (Fig.

occipital

partly visible on sides and

Dryininae

Fig.

500 -

Chela of

Dryinus koreanus (Moczar) (type)

749

750

Fig.

Mem.

501

-

Amer.

Ent.

Inst.,

no.

Male genital ia of Dryinus koreanus (Korea) ( I eft ha I f

37,

1984

(Moczar)

from De-sang san

removed)

occipital carina dorsally touching eyes; POL = 5; Ol = 2; OOl = = 1; scutum dull, granulated; notaulices incomplete, reaching

ventrally; 3;

OPl

approximately 0,5 shiny, without

smooth,

length of scutum;

without sculpture;

transversal

scutellum dull,

propodeum dull,

or longitudinal

keels;

granulated;

metanotum

reticulate rugose,

fore-wing hyal ine,

without dark

transversal bands; distal part of radial vein longer than proximal (16:7); genitalia: fig. 501; tibial spurs 1,1,2. lOCUS TYPICUS:

Pyongyan

TYPICAL MATERIAL: HOSTS:

part

(Korea)

holotype F!,

allotype M!

and 1 paratype M!

in BU.

unknown

DISTRIBUTION:

Palaearctic region:

Pyongan),

BU!

lyong-ak san

Pyongan),

BU!

De-sang san

KOREA:

Pyongyan

(Prov.

South

(14 Km IV from Pyongyan,

Prov.

South

(12 Km NE from Pyongyan,

Prov.

South

Dryininae

Pyongan),

751

BU!

7. Dryinus antiquus (Ponomarenko) n. comb.

Laberius antiquus Ponomarenko 1981 a: 139.

FEMALE: MALE:

unknown

fully

winged;

length 3,20 mm;

black,

with

legs testaceous;

palpi

not visible; antennae not distally thickened, densely hairy; antennal hairs less short than breadth of antennal segments; antennae la-segmented; antennal

segments

in following proportions:

7;

breadth of antenna I segments

4:

4:

3;

mandibles not visible;

by short dense hairs; posterior ocelli occipital

scutum dull,

posteriorly separated;

5:

head dull,

11:

10:

10:

9:

carina;

TL = a

bands;

three basal

cells enclosed by pigmented veins;

covered

(eyes almost touching notaulices complete,

fore-wing hyaline,

transversal

7:

eyes apparently bare;

metanotum and propodeum weakly

apparently propodeum reticulate rugose;

7:

4: 4: 4: 4: 4,5: 4: 4;

apparently granulated;

scutellum,

pterostigma

11:

apparently granulated,

occipital carina complete;

not touching occipital

carina);

8:

(greatest breadth):

long and narrow;

visible;

without dark

apparently fore-wing

distal

part of radial

with

vein

longer than proximal part; tibial spurs 1,1,2. LOCUS TYPICUS: Taimyr amber (Sibiria) TYPICAL MATERIAL: HOSTS:

holotype M!

in MPI.

unknown

DISTRIBUTION:

only

known

from Taimyr amber

(Sibiria,

U.

S.

S.

R.)

(Laberius Kieffer) is not correct, because Laberius is a genus of Gonatopodinae (it's a junior synonym of Dicondylus Hal iday). As its occipital carina is complete, L. antiquus is belonging to Dryininae. The new generic name Dryinus is provisionally proposed. The only known specimen of L. antiquus is in too bad conditions to permit to associate it for a certainty to a genus of Dryininae. D. antiquus is the oldest fossil dryinid known (nearly 100 millions NOTES:

the original

generic status

years old).

ETHIOPIAN SPECIES OF

Nineteen

species are

DRYINUS

known.

KEY TO THE SPECIES

752

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

FEMALES

Posterior collar of

2

3

4

5

6 7

8

pronotum invisible . . . . . . . . . . . . . . . . . . . . . . . .. . . . . . . . . . . . . . . . . . . . . . . . . . . . 1. bisulcatus (Benoi t) Posterior collar of pronotum visible (Figs 503 B, 519, 522) ... 2 Enlarged claw much shorter than segment 5 of front tarsus and wi th subapical tooth very small (Fig. 504) . . . . . . . . . . . . . . . . . . . .. · . . . . . . . . . . . . . . . . . . . . • • . . .. 2. seyrigi Ben 0 i t Enlarged claw nearly as long as segment 5 of front tarsus and usually with subapical tooth big (Figs 506,508) . . . . . . . . . . . . . . 3 Head (dorsally viewed) with posterior ocelli placed behind the imaginary straight line joining posterior edges of eyes (Figs 505, 507) ••••.••••••••••••••••••••••••••••••••.••••••••••••••• 4 Head (dorsally viewed) wi th posterior ocell i placed in front of the imagina;"y straight line joining posterior edges of eyes or on t his lin e (F i g s 51 4, 524) •••••.•••••••••••..•••••••••••• 8 Head wi th posterior margin of vertex excavated (Fig. 505 A); head excavated; posterior ocell i touching occipi tal carina (Fig. 505 A) . . . . . . . . . . . . . . . 3. alticolus (Benoit) Head wi th posterio margin of vertex convex (Figs 505 B, 507 B); head flat; posterior ocelli not touching occipital carina (Figs 505 B, 507) ••••••••••••••••••••••••••••••••••••••••••••.•••••• 5 At I east the propodeum black . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6 Species fully testaceous or reddish-testaceous, with petiole black ••••••••••••••••••••••••••••••••••••••••••••••••••••••••• 7 Head almost fully black . . . . 7. sierranus n. sp. Head almost fully reddish •• 8. pretorianus n. sp. Posterior margin of vertex more convex (Fig. 507 B); antennal segment 3 nearly ten times as long as segment 2 . . . . . . . . . . . . . . .. · . . . . . . . • • . . • . . . . . • • . • . . . .. 4. spangleri n. sp. Posterior margin of vertex less convex (Fig. 507 A); antennal segment 3 nearly six-seven times as long as segment 2 •••••••••• . . . . . . . . . . . . . . . . . . . . . . . . . . . 5. deceptor (Turner) Species fully testaceous, with petiole black and sometimes abdomen brownish; scutum shiny, weakly punctate ••••••••••.••••• . . . . . . . . . . . . . . . . . . . . . . . . . . . 6. mahensis Ki effer Species at least partly black; scutum shiny or dull, punctate or granulated, usually with more or less short irregular keels

· •.••••••..•..•..••.. " .••.•••.•.••.••••••.•• " •.••••. " •••••. 9 9

Posterior ocelli

touching

Posterior ocelli

not

524) ••••••••••••••••• " 10 Notaulices

occipital

touching

•••••••••• "

reaching approximately

........................... Notaulices

9. 10.

(Fig.

carina

514 A) . . . . . . 10

(Figs

514 B,

•.••.••••••••.•••••••••••. 11 0,65

length of

scutum . . . . . . . .

saussurei (Ceballos)

reaching approximately

• ••••••••••••••••••••••• "

carina

occipital

0,3

length of

hova (C eba I I os)

scutum . . . . . . . . .

753

Dryininae

11 Temples prominent prominent,

but

(Figs 514 B,

in

this

524 A);

rarely

temples weakly

case head with posterior margin

of

514 B, 524 A) . • . . . . . . . . . • • • • • • • • • • • • . . 12 Temples invisible (Fig. 524 B, C); head wi th posterior margin of vertex convex (Fig. 524 B, C) . • . • • . . . • . • • • . . . . . • • • • . • • • • • . 18 vertex

excavated

(Figs

12 Fore-wing with distal as

long as proximal

Fore-wing with distal times as

part

of

part ••

11.

part

of

radial

vein nearly

three

times

basilewskyi Benoit radial

vein

less

than

three

part ••••••••••••••••••••••••••••••• 13

long as proximal

13 Scutum punctate, not granulated, wi thout keel s ••••••••••••••••• •••••••••••••••••.•••••••• 12. chamaeleo Benoi t Scutum granulated or with keels ••••••••••••••••••.•.••••.•••• 14 14 Head granulated, with irregular or regular keels . . . . • • . • • • . . • 15 Head granulated,

without

keels ••••••••••••••••••••••••••••••• 17

15 Pronotum wi th anterior impression stronger (Fig. 519 A); head mostly reddish •••••••••••• 13. schoutedeni (Benoit) Pronotum with anterior impression weaker (Fig. 519 B); head mostly black ••••••.•••.••••••••••••••••.••......•.••••••••••• 16

16 Propodeum long, wi th dorsa I surface more than twi ce as long as posterior surface ••••••••• 14. mayogo (Benoit) Propodeum short, with dorsal surface slightly longer than posterior surface ••••••••• 15. ol'ophilus (Benoit) 17 Scutum granulated, not reticulate rugose; subapical tooth of enlarged claw very near apex (Fig. 525) •••••..•••••.••••••••••• 16. modestus n. sp. Scutum granulated, with posterior half reticulate rugose; subapical tooth of enlarged claw farther from apex (Fig. S 2 6) • • • • • • • • • • • • • • . • . . • . .• 1 7. incel'tus n. sp. 18 Anterior impression of pronotum very weak (Fig. 522 B) •••••••••

18. bUl'geoni Beno it of pronotum stronger (Fig.

•••••••••••••••••••••••••, Anterior

impression

••••••••••••••••••••••••••

19.

522 A) ••••••••••

aiel' (Dlmi)

MALES

Head wi th OL

= OOL •••••••••.••••••.•.•.••••••••••••••••••....• 2

Head wi th OL more or 2

Fore-wing with

radial

less

shorter

cell

than OOL •••••••••••••••••••• 3

closed;

head with posterior ocelli

not touching occipital carina . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. . . . . . . . . . . . . . . . . . . . . . . . . . . . 6. mahensis (Kieffer) Fore-wing with touching

radial

occipi tal

cell

• • • • • • • • •• •• • •• • • • • • • •• • • •• 3

open;

head with posterior ocelli

carina •••••••..••...••••••••••••.....••••••• 9.

saussul'ei (Ceba I los)

Fore-wing wi th

radial

cell

closed . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4

Fore-wing with

radial

cell

open . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5

754

4

Mem.

Amer.

Ent.

Ins!.,

no.

37,

1984

Frons flat in front of the anterior ocellus, convex only between the antennal sockets • • . . • • • • • • • • . . • • • • • • • • • . • . • • • • •

19. afer (Olmi) Frons wi th a

5

median

antennal

sockets •••

Vo I s e I I a

n ea r I y

as

longitudinal •••••••

shorter

from anterior

long as gonoforceps

• • • • • • • . • • • • • • • • • • • .. Volsella

hump

(Fig.

to

513 B) . . . . . . . . . . .

(Fig.

pretorianus n.

8.

513 A)

ocellus

deceptor (Turner)

5.

sp.

••••••••.

15. orophilus (Benoit)

The males of other species are unknown. The description of the male allotype of (1953b)

is not reliable.

addition

the

D. chamaeleo by Benoit

This male was not obtained by breeding;

in

locality of the holotype is different.

Dryinus bisulcatus (Benoit) n. comb.

1.

Mercetia bisulcata Benoit 1953b: 400.

FEMALE:

fully

winged;

length 5,62 mm;

head black,

with clypeus and

mandibles testaceous; antennae testaceous; propectus black; pronotum black, with sides reddish; mesothorax, metathorax and propodeum black; abdomen black; black;

legs testaceous,

antennae distally

proportions:

11: 6;

shiny,

smooth,

behind

the ocelli);

21:

weakly

with coxae and clubs of femora brown or

weakly

13:

11:

thickened; 7,5:

alutaceous;

OL

6,5:

antenna I segments

4,5:

occipital

longer than POL;

6:

6;

carina

in following

head excavated, invisible

pronotum shiny,

(no track

weakly

granulated, crossed by a transversal impression on anterior third; the pronotum so is divided into 2 collars, without disc; the posterior collar is weakly

humped;

tegulae;

pronotal

scutum shiny,

converging,

reaching

surface with 2 other wing

proximal

part;

with subapical

median keels;

spurs 1,1,2. unknown

tarsus produced

tooth and

transversal

strongly

scutellum shiny propodeum not

keels;

anterior

sides of anterior surface with 2

posterior surface with 2

segment 1 of front

with 2 rows of 41 MALE:

keels;

with 3 brown spots and with distal

segment 3 of front

not reaching

incomplete,

metanotum very reduced and short;

with numerous parallel

longitudinal

longitudinal

notaul ices

almost posterior margin of sctum;

and without sculpture; reticulate rugose,

tubercles not very prominent,

granulated;

longitudinal

keels;

fore-

vein as

long as

tarsus shorter than segment 4

(18:22);

into a

17 lamellae;

part of radial hook;

enlarged claw

segment 5 of front

lamellae extending continuously

tarsus

to the apex;

(Fig. (Fig. tibial

502) 502)

I

Dryininae

Fig.

I

502 -

LOCUS TYPICUS: HOSTS:

Chela of Dryinus bisulcatus

Behara

TYPICAL MATERIAL:

755

(Benoit)(type)

(Madagascar)

holotype Fl

in P.

unknown

DISTRIBUTION:

Ethiopian region:

MADAGASCAR:

Behara,

Pl

D. bisulcatus is species of transition from Dryininae to Gonatopodinae. For the tibial spurs (1,1,2) and the pronotal tubercles directed towards the tegulae it is belonging to Dryininae; for the absence NOTES:

of occipital

carina,

metanotum very

the clypeus ventrally rounded

reduced

it

is near Gonatopodinae.

(not bidentate)

and

the

756

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

A

B Fig. 503 - Pronotum (laterally viewed) of females of Tridryinus ampuliciformis (Turner) from Afgoi (Soma I i a )(A) and Dryinus seyrigi Benoi t (type) (B)

2. Dryinus seyrigi Benoit

Dryinus seyrigi Benoit 1953b: 407.

FEMALE:

fully

winged;

length 3,75 mm;

head black,

with front part of

vertex, clypeus and mandibles testaceous; antennae testaceous, with segments 6-9 brown; propectus black; pronotum black, with sides and posterior collar reddish;

mesothorax,

metathorax and propodeum black;

abdomen missing in holotype (only specimen known); legs testaceous, with coxae and clubs of femora brown; antennae distally weakly thickened; antennal 8;

segments

head shiny,

prominent;

in following proportions:

granulated;

pronotum

posterior transversal

(Fig.

occipital 503 B)

impresssion

10:

5:

21:

carina complete;

shiny,

granulated,

and by a

10:

9:

8:

OL = POL;

6:

6:

4,5:

temples

crossed by a

strong

weak anterior transversal

impression; disc humped; posterior collar short, distinct; anterior collar weak; pronotal tubercles not reaching tegulae; scutum dull, granulated, with short keels near posterior margin; approximately 0,5 short, without

length of scutum;

with numerous longitudinal

transversal

longitudinal

notaulices incomplete,

scutellum dull,

keels;

metanotum

propodeum reticulate rugose,

keels on posterior surface;

bands and with distal

reaching

granulated;

fore-wing

part of radial

with 3 brown

vein much

longer than

D'ry i n i nae

Fig.

504 -

Chela of

Dryinus seyrigi Benoit (type)

757

758

Mem. Amer.

Ent.

Inst.,

no.

37,

1984

A

Fig.

Heads of females of Dryinus alticolus (Benoit) (type) (A) and deceptor (Turner) from Port SI. Johns (South Africa) (B)

505 -

proximal part; segment 1 of front tarsus nearly twice as long as segment 4; segment 3 of front tarsus produced into a hook; enlarged claw (Fig. 504) with a small subapical tooth and 3 lamellae; segment 5 of front tarsus (Fig. 504) with 2 rows of 22 lamellae extending continuously to the apex; segment 5 of front tarsus apically without prominence and much longer than enlarged claw; tibial spurs 1,1,2. MALE: unknown LOCUS TYPICUS: Ranomafana (Madagascar) TYPICAL MATERIAL: holotype F! in P. HOSTS: unknown DISTRIBUTION: Ethiopian region MADAGASCAR:

Ranomafana,

3. Dryinus alticolus (Benoit) n. comb.

Lestodryinus alticola Benoit 1951c: 4.

P!

Dryininae

Fig.

FEMALE:

fully

506 -

winged;

Chela of Dryinus alticolus

length 5,0 mm;

side of head;

(Benoit) (type)

head ferruginous,

brown spot on ocell ar reg ion and part I y visible on ventral

759

on frons;

with an elongate

another brown spot is

antennae testaceous;

propectus black,

wi th a dorsal reddish spot; pronotum ferrug i nous; mesothorax, meta thorax and propodeum black; abdomen brown; legs testaceous, with hind and mid coxae partly brown;

head

(Fig.

505 A)

excavated,

dull,

granulated,

with

760

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

oa

o Fig.

507 -

Heads of females of

Johns

temples

(South Africa) (A)

0

Dryinus deceptor (Turner) from Port St. and spangleri n. sp. (type) (B)

very prominent and posterior margin of vertex excavated;

occipital

carina complete; TL nearl,' three times as long as POL; OL weakly longer than POL; posterior ocell i touching occipital carina; pronotum dull, granulated;

anterior collar and

posterior collar distinct;

anterior transversal

posterior transversal

tubercles not reaching

tegulae;

near posterior margin;

notaulices incomplete,

scutum dull,

length of scutum;

scutellum dull,

reticulate rugose,

without

wing

with 3 brown

much

longer than proximal

granulated;

longitudinal

transversal part

impression

granulated,

weak;

pronotal

with some areolae

reaching approximately 0,75 metanotum rugose;

keels on segment

propodeum

posterior surface;

bands and with distal (17:4);

very

impression strong;

fore-

part of radial

1 of front

vein

tarsus nearly

as long as segment 4; segment 3 of front tarsus produced into a hook; enlarged claw (Fig. 506) with subapical tooth and 4 lamellae; segment 5 of front of 10 MALE:

tarsus

lamellae; unknown

(Fig. tibial

506)

with 2 rows of 24

spurs 1,1,2.

lamellae;

apex with a

group

761

Dryininae

Fig.

508 -

Chela

0

f Dryinus spanglen. n.

sp.

(type)

762

Mem.

LOCUS TYPICUS:

Kilinga

TYPICAL MATERIAL: HOSTS: unknown

Amer.

Ent.

Inst.,

no.

37,

1984

(Zaire)

holotype F!

in TV;

1 paratype F!

DISTRIBUTION: Ethiopian region: ZAIRE: CAMEROON: N I Kongsamba (Beno i t 19S 1c) •

Kilinga

in P.

(Rutshuru),

TV!

4. Dryinus spangleri n. sp.

FEMALE:

fully

winged;

length 5,62 mmj

triangle and petiole black; white,

6-8 brown;

antennae distally weakly

in following

proportions:

507 B)

flat,

dull,

10:

OOL = 13;

OPL = 2j

imaginary straight of vertex convex;

57:

testaceous,

36:

24:

weak I y

antennal

14:

7:

11: 8:

frons

vis i b Ie on si des of occi put;

TL = 7;

posterior ocelli

POL = 3;

OL

posterior margin

impression;

disc humped;

longer than pronota I tubercles;

pronota I

pronotum granulated and with numerous

longitudinal

striae on sides;

longitudinal

striae on

scutum dull,

lateral region; fore-wing

granulated and with numerous

notaulices weak,

scutellum and metanotum dull,

fully reticulate rugose;

(Fig.

crossed by an anterior weak transversal

much

tubercles reaching tegulae;

segments

head

placed behind the

a posterior strong transversal long,

10;

with a median keel;

I ine joining posterior edges of eyes;

impression and by

with ocellar

with segments 4-5 partly

thickened;

weakly rugose;

pronotum du II,

posterior coil ar very

complete;

7:

granulated,

occipi ta I cari na comp I ete, = 2;

fully

antennae testaceous,

granulated;

but distinct and

propodeum shiny,

with 2 dark transversal

bands,

one on

the basal cells and another beneath the pterostigma; distal part of radial vein much longer than proximal part; fore tarsal segments in following proportions:

27:

3:

8:

22:

tooth and 12 lamellae; Clearly 33 lamellae;

33;

enlarged claw

segment 5 of front

(Fig.

tarsus

apex with a group of 23

508)

(Fig.

lamellae;

with a subapical 508) tibial

with 2 rows of spurs 1,1,2.

MALE: unknown LOCUS TYPICUS: Mombasa (Kenya) TYPICAL MATERIAL: holotype F! in WA. HOSTS:

unknown

DISTRIBUTION:

Ethiopian region:

KENYA:

Mombasa,

WAr

5. Dryinus deceptor (Turner) n. comb.

Lestodryinus deceptor Turner 1928: 147.

FEMALE:

fully

winged;

length 6,0 mm;

and abdomen reddish-brown; granulated;

occipital

head

yellow-testaceous,

(Figs 505 B,

carina complete;

507 A)

with petiole black

flat,

OPL shorter than POL;

dull, OL = POL;

Dryininae

763

Fig. 509 - Chela of Dryinus deceptor (Turner) from Port SI. Johns (South Africa)

764

Mem.

temples short;

Amer.

pronotum dull,

posterior transversal

Ent.

Ins!.,

granulated,

impression strong;

anterior transversa I impression weak; tegulae;

scutum shiny,

approximately 0,5 granulated;

granulated;

no.

with numerous keels on sides;

pronota I tuberc les not reach i ng incomplete,

longitudinal

reaching

scutellum and metanotum shiny,

propodeum weakly reticulate rugose;

propodeum without

1984

posterior collar long and visible;

notaul ices

length of scutum;

37,

keels;

posterior surface of

fore-wing

with 2 brown

transversal

bands and with distal part of radial vein much longer than proximal part; segment 1 of front tarsus shorter than segment 4 (18:22); segment 3 of front

tarsus produced

subapical

tooth and

2 rows of 34

into a

hook;

12 lamellae;

lamellae;

enlarged claw

segment 5 of front

(Fig.

509)

tarsus

apex with a group of 19 lamellae;

with

(Fig. tibial

509)

with

spurs

1,1,2. MALE: fully winged; length 2,43 mm; black; mandibles testaceous; antennae brown, with segments 1-2 testaceous; legs testaceous, with coxae, clubs of femora, mid and hind tibiae brown; antennae not distally thickened; antennal segments in following proportions: 5: 3: 11: 7: 6:

6:

5:

7;

head dull,

with a median frons

longitudinal

without median keel;

OOL = 4; dull,

granulated,

OPL = 1;

granulated,

reaching 0,5 transversal transversal

with weak tracks of areolae;

hump from anterior ocellus to antennal occipital

temples

invisible;

with weak

carina

incomplete;

scutum,

6:

sockets;

= 5,6; OL = 2;

scutellum and metanotum

tracks of areolae;

length of scutum;

POL

6:

frons

notaul ices incomplete,

propodeum reticulate rugose,

dull,

without

and longitudinal keels; fore-wing hyal ine, without dark bands; distal part of radial vein longer than proximal part;

radial cell closed; tibial spurs 1,1,2. LOCUS TYPICUS: Port SI. Johns (South Africa) TYPICAL MATERIAL:

holotype F!

and 1 paratype F!

HOSTS: unknown DISTRIBUTION: Ethiopian region:

UGANDA:

Port St.

St.

Johns

(Pondoland),

BM!

in SM.

Kampala,

TW!

SOUTH AFRICA:

Lucia Estuary,

TW!

OL!

6. Dryinus mahensis (Kieffer) n. comb.

Mesodryinus mahensis Kieffer 1912a: 45. Labeo saxetanus Kieffer 1912a: 46 (n. syn.); locus typicus Mahe Island

(Seychelles

Islands).

Mesodryinus mahensis Kieffer: Kieffer 1914b: 31. Laberius saxetanus (Kieffer): Kieffer 1914b: 64.

FEMALE:

fully

winged;

abdomen darkened; complete, slightly

length 4,75 mm;

head shiny,

testaceous,

weakly punctate,

almost invisible on sides of occiput; longer than POL;

pronotum shiny,

with petiole black and

hairy;

occipital carina

OPL shorter than OLi

smooth,

without sculpture;

OL

Dryininae

Fig.

510 -

posterior transversal

765

Dryinus mahensis (Kieffer)(type)

Chela of

impression strong;

anterior transversal

impression

and anterior collar weakly visible; posterior collar visible; pronotal tubercles not reaching tegulae; scutum shiny, smooth, weakly punctate; notaul ices incomplete, reaching approximately 0,65 length of scutum; scutellum and metanotum shiny, smooth, weakly punctate; propodeum reticulate rugose,

without

wing

with 2 brown

much

longer than proximal

longitudinal

transversal

part;

segment

segment 4;

segment 3 of front

(Fig.

510)

with subapical

(Fig.

510)

with 2 rows of 30 lamellae;

tibial

spurs 1,1,2.

·.lALE:

fully

winged;

1 of front

tarsus produced

hook;

vein

long as

enlarged claw

segment 5 of front

apex with a head black,

fore-

part of radial

tarsus as

into a

tooth and 14 lamellae;

length 3,43 mm;

antennae yellow-brown;

keels on posterior surface;

bands and with distal

tarsus

group of 15 lamellae;

with mandibles yellow;

thorax and propodeum bl ack;

abdomen brown;

legs

yellow, with hind coxae partly black; head dull, granulated; occipital carina incomplete; temples invisible; POL; 8; OL ; 3; OOL ; 3; OPL ; -~

;

0;

scutum dull,

granulated,

with

weak

areolae on

lateral

regions;

2;

766

Mem.

Fig.

511

-

Amer.

Chela of

Ent.

Inst.,

no.

37,

1984

Dryinus sierranus n. sp. (type)

notaulices incomplete, reaching approximately 0,65 length of scutum; scutellum shiny, weakly granulated; metanotum dull, rugose; propodeum reticulate rugose; fore-wing without dark transversal bands and with distal part of radial vein much longer than proximal part; tibial spurs 1,1,2 •• LOCUS TYP ICUS: Port Glaud (Seychelles TYPICAL MATERIAL: holotype F! of M.

Islands)

mahensis in BM; holotype M! of L.

saxetanus in BM. HOSTS: unknown DISTRIBUTION: Ethiopian region:

SEYCHELLES

ISLANDS: Port Glaud

Island), BM! Mount Sebert (Mahe Island), BM! (Silhouette Island)(Kieffer 1912a).

Mare aux Cochons

7. Dryinus sierranus n. sp.

(Mahe

Dryininae

767

FEMALE: fully winged; length 8,75 mm; head black, with occiput, anterior region of frons, clypeus and mandibles reddish; antennae black, with segments 1-2 reddish, segment 4 yellow,

junction between segments 3-4 testaceous, half of segment 4 yellow,

proximal

prothorax reddish-brown-dark; mesosternum reddish-dark; metasternum black;

scutum bl ack,

scutellum,

abdomen black,

with junctions and tarsi

reddish;

apex of

segment 10 yellow;

wi th sides redd ish-brown;

metanotum,

propodeum,

pleura,

with some reddish regions;

legs black,

antennae distally weakly thickened;

antennal segments in following proportions: 10: 7: 58: 33: 24: 14: 11: 8: 8: 10; head dull, granulated, with tracks of irregular keels on frons; frons with a median keel; occipital carina complete; POL = 5; OL = 3,5; OOL = 11; OPL = 2; TL = 7; posterior ocelli placed behind the imaginary straight line joining posterior edges of eyes; posterior margin of vertex convex; pronotum crossed by strong transversal impression and with a posterior collar long and distinctly visible; pronotum dull, granulated and with numerous longitudinal keels around disc; pronotal tubercles not reaching tegulae; scutum dull, granulated, with numerous weak longitudinal striae; notaul ices incomplete, almost reaching posterior margin of scutum, numerous

but weakly visible; longitudinal striae;

fully reticulate rugose,

scutellum dull, metanotum dull,

granulated and with reticulate rugose;

with 2 incomplete longitudinal

propodeum

keels on posterior

surface; fore-wing with 2 dark transversal bands, one on the basal cells and another beneath the pterostigma; distal part of radial vein longer than proximal 10: 23:

38;

lamellae;

part;

fore tarsal

enlarged claw

(Fig:

segment 5 of front

segments in following proportions: 511)

tarsus

apex with a group of 14 lamellae; MALE:

with a subapical (Fig.

tibial

511)

28:

5:

tooth and with 20

with 2 rows of 49 lamellae;

spurs 1,1,2.

unknown

LOCUS TYP ICUS:

Freetown

TYPICAL MATERIAL: HOSTS:

(Sierra Leone)

holotype F!

in TW.

unknown

DISTRIBUTION:

Ethiopian region:

SIERRA LEONE:

Freetown,

TW!

8. Dryinus pretorianus n. sp.

FEMALE:

fully

winged;

brown spot on frons; whi t ish;

prothorax,

propodeum black;

length 6,25 mm;

antennae brown,

head reddish-testaceous,

scutum and scutell um redd ish-dark;

abdomen brown,

with a

with segments 1-2 reddish,

with segment 1 partly reddish;

with chelae and segments 2 and partly

1 of hind tarsi

10

metanotum and whitish;

legs,

antennae

distally thickened; antenna I segments in following proportions: 9: 7: 49: 29: 21: 14: 10: 9: 7: 8; head dull, flat, granulated; frons with a median keel;

POL = 4,5;

complete;

DL = 3,5.

DOL = 11; DPL = 2;

posterior margin of vertex convex;

the imaginary straight

TL = 7;

occipital

carina

posterior ocelli placed behind

line joining posterior edges of eyes;

pronotum with

768

Mem. Amer. Ent.

Inst., no. 37, 1984

Fig. 512 - Chela of Dryinus pretorianus n. sp.

(type)

Dryininae

Fig.

513 -

769

Dryinus orophilus (Benoit) from Ile-ife

Male genitalia of

(Nigeria) (A: right half removed) and pretorianus n. sp. from Pretoria (South Africa) (8: right half removed; C: ninth abdominal sternite)

a strong

transversal

dull,

hairy,

dull,

granulated,

impression and with a

granulated;

pronotal

posterior collar

tubercles not reaching

with numerous irregular weak keels;

long;

tegulae;

pronotum scutum

notaul ices

incomplete, reaching approximately 0,5 length of scutum; scutellum and metanotum as sculptured as scutum; propodeum reticulate rugose, dull, with

two median

longitudinal

keels on dorsal

surface;

fore-wing

with 2

dark transversa I bands, one on the basa I cell s and another beneath the pterostigma; distal part of radial vein longer than proximal part (15:7); fore tarsal

segments

claw

(Fig.

512)

front

tarsus

to the apex; MALE:

fully

in following proportions:

with a subapical

(Fig.

512)

tibial winged;

with 2 rows of 45

4:

10:

legs brown,

3;

occipital

scutum dull,

17:

thickened; 8:

8:

8:

black;

28;

enlarged

segment 5 of

antennal 8:

carina complete; granulated;

mandibles and antennal

with joints and tarsi

6:

6:

segments 8;

with some weak areolae on sides of frons; keel;

8:

lamellae extending continuously

length 2,37-2,62 mm;

antennae not distally 4:

3,5:

spurs 1,1,2.

segments 1-2 testaceous; proportions:

20:

tooth and with 10 lamellae;

POL = 6;

notaul ices

head dull,

granulated and

frons with an OL = 3;

incomplete,

testaceous;

in following incomplete median

OOL = 4; reaching

OPL = 1;

TL =

approximately

770

Mem.

Amer.

En!.

Inst.,

no.

37,

1984

\) 0

Fig.

514 -

Heads of females of Dryinus saussurei (Madagascar) (A)

(Ceballos)

and basilewskyi Benoit

from Beki Iy

(type) (B)

0,65 length of scutum; scutellum dull, granulated; metanotum dull, granulated and with numerous areolae near anterior margin; propodeum reticulate rugose,

without transversal

or longitudinal

keels;

fore-wing

hyaline, without dark transversal bands; distal part of radial vein longer than proximal part (9:7); distivolsella (Fig. 513 B) with apex located nearly on the same level with the apex of gonoforceps; sternite: fig. 513c; tibial spurs 1,1,2. LOCUS TYP ICUS:

Pretoria

TYPICAL MATERIAL: HOSTS:

ninth abdominal

(South Africa)

holotype F!

and 2 paratypes MM!

in TW.

unknown

DISTRIBUTION:

Ethiopian region:

SOUTH AFRICA:

Pretoria

(Transvaal),

TVY!

9. Dryinus saussurei (Ceballos)

Lestodryinus saussurei Ceballos 1935: 48. Dryinus saussurei (Ceballos): Benoit 1953b: 412.

FEMALE:

fully

winged;

mandibles yellow;

length 4,00-5,93 mm;

rarely clypeus black;

with segments 7-8 or 3-10 brown;

head black,

propectus black;

margins and posterior collar reddish;

with clypeus and

antennae testaceous,

mesothorax,

sometimes

pronotum black,

with

metathorax and

propodeum black; abdomen black-brown; legs testaceous, wi th coxae and clubs of femora partly brown; head (Fig. 514 A) shiny or du", fully or partly

granulated;

occipital

carina complete;

posteior ocell i

touching

Dryininae

Fig.

515 -

Chela of

771

Dryinus saussurei (Ceballos) from Bekily (Madagascar)

772

Mem.

occipital

carina;

Amer. En!.

OL = POL;

Ins!.,

pronotum dull,

with numerous weak striae around disc; nearly 0,5 times as

no.

37,

1984

partly or fully granulated,

posterior collar of pronotum short,

long as distance from

tegulae to pronotal

tubercles;

pronotal tubercles not reaching tegulae; scutum dull, granulated, with short keels or areolae near posterior margin; notaulices incomplete, almost reaching posterior margin of scutum; dull,

rugose;

posterior surface; proximal twice as into a claw

hook;

with

segment

515)

(Fig.

lamellae; MALE:

fore-wing

with 2

granulated;

longitudinal

with 3 brown transversal

bands

metanotum

keels on

(the two

bands partly joint) and with distal part of radial vein nearly long as proximal part (18:8); segment 3 of front tarsus produced

(Fig.

tarsus

scutellum dull,

propodeum reticulate rugose,

515)

tibial

fully

1 of front tarsus as

with subapical

tooth and

with 2 rows of 39

long as segment 4; 13

lamellae;

lamellae;

enlarged

segment 5 of front

apex with a

group of 11

spurs 1,1,2.

winged;

teeth brown;

length 2,60-2,84 mm;

antennae brown,

black;

mandibles testaceous,

with segments 1-2 testaceous;

palpi

testaceous; abdomen brown; legs testaceous, with coxae, clubs of hind femora and hind tibiae brown; antennal segments in following proportions: 8:

7:

17:

12:

13:

13:

11:

incomplete median keel, between

10:

10:

flat

the antenna I sockets;

DOL = 3; posterior ocelli scutum dull, granulated;

11;

head dull,

granulated;

frons with an

in front of the anterior ocellus and humped occipital

carina complete;

POL = 9;

OL = 3;

touching occipital carina; temples invisible; notaulices incomplete, reaching approximately

0,65 length of scutum; scutellum almost fu!ly dull, granulated, with a narrow posterior region smooth and shiny; metanotum with anterior half rugose and with posterior half shiny and smooth;

propodeum reticulate

rugose,

fore-wing hyal ine,

without

longitudinal

without dark

transversal

part slightly

longer than

segments;

labial

LOCUS TYPICUS:

palpi Rogez

TYPICAL MATERIAL:

or transversal

bands;

proximal

MD!

Behara,

part

with 3 segments;

cell open; (14:13); tibial

radial

vein

with distal

maxillary palpi

with 6

spurs 1,1,2.

(Madagascar)

holotype F!

HOSTS: unknown DISTRIBUTION: Ethiopian region: Rogez,

radial

keels;

TV!

in MD. MADAGASCAR:

Fort Dauphin,

TV!

Bekily,

TV!

P!

Mandraka,

TV!

Ranomafana

Ivondro,

P!

(Benoit 1953b).

10. Dryinus hova

(Ceballos)

n.

comb.

ft;esodryinus hova Ceballos 1936: 53. fiJesodryinus hova Ceballos: Benoit 1953b: 414.

FEMALE:

fully

winged;

mandibles testaceous;

length 5,31

mm;

propectus black;

head black,

with clypeus and

pronotum black,

with margins and

773

Dryininae

Fig.

516 -

Chela of

Dryinus hova

(Ceballos)(type)

posterior collar reddish; mesothorax, metathorax, propodeum and abdomen black; legs testaceous, with coxae and clubs of femora brown; antennae distally weakly 5:

24:

10: 9:

transversal

thickened;

7:

6:

6:

5:

antennal 7,5;

segments

head dull,

keels on anterior half of frons;

posterior ocelli edges of eyes;

located on

in following proportions:

granulated,

occipital carina complete;

the imaginary straight

posterior ocelli

9:

with numerous short

touching occipital

line joining posterior carina;

OL = POL;

pronotum dull, granulated, with anterior transversal impression weak; anterior collar weak; posterior transversal impression strong; posterior collar short,

distinct;

pronotal

tubercles not reaching

tegulae;

scutum

dull, granulated, with some strong punctures; notaulices incomplete, reaching approximately 0,3 length of scutum; scutellum dull, granulated; metanotum rugose;

propodeum reticulate rugose,

on posterior surface;

fore-wing

half of fore wings missing species); of front

without

longitudinal

transversal

bands;

in holotype (only known specimen of the

tarsus produced into a hook; lamellae;

with

tarsus

(Fig.

516)

apex with a group of 12 lamellae;

tibial

spurs

segment 5 of front

unknown

LOCUS TYP I CUS:

Bek i I Y (Madagascar)

TYPICAL MATERIAL:

holotype F!

enlarged claw

in P.

(Fig.

segment 3

516)

tooth and 8 lamellae;

2 rows of 21

keels

distal

segment 1 of front tarsus nearly as long as segment 4;

subapical 1,1,2. MALE:

with 2 brown

with

774

Mem. Amer.

Fig. 517 -

Ent.

Ins!., no. 37,

1984

Chela of Dryinus basilewskyi Benoit

HOSTS: unknown DISTRIBUTION: Ethiopian region:

MADAGASCAR:

Bekily, P!

11. Dryinus basilewskyi Benoit

(type)

Dryininae

775

Dryinus basilewskyi Benoi t 1953a: 143.

FEMALE: vertex

fully

winged:

length 7,50 mm;

head black,

testaceous;

propectus black;

coil ar redd i sh; yellow-brown; sculpture,

mesothorax,

pronotum black, head

posterior ocelli

514 B)

very near occipital

than OL;

OL = POL;

except for posterior collar very

impression strong; distinct,

tubercles

reaching

tegulae;

abdomen

shiny,

without

(4:9);

granulated;

length of scutum; fore-wing

part of radial

vein

segment 3 of front

posterior collar

weak;

tegulae to

pronotal

tubercles not

notaulices incomplete,

without

longitudinal

transversal

three times as

tarsus nearly as

tarsus produced

weak;

reaching

scutellum and metanotum dull,

with 2 brown

more than

segment 1 of front

occipital

OPL shorter

posterior transversal

long as distance from

propodeum reticulate rugose,

posterior surface;

hairy;

impression

anterior collar very

scutum dull,

carina,

with numerous striae around disc,

anterior transversal

approximately 0,65 granulated;

pronotum shiny,

nearly 0,5 times as

pronotal

(17:5);

(Fig.

except for numerous weak striae on frons and vertex;

hairless,

distal

antennae

with sides and posterior

meta thorax and propodeum black;

legs yellow-testaceous;

carina complete;

short,

with front part of

(more along orbits), clypeus and mandibles yellow;

into a

long as proximal

long as segment 4

hook;

keels on

bands and with

enlarged claw

part

(17:16); (Fig.

517)

with subapical tooth and 11 lamellae (lamellae near subapical tooth very long and simi lar to peg-I ike hairs); segment 5 of front tarsus (Fig. 517) with 2 rows of 30 lamellae; 1,1,2. MALE:

apex

with

a group of 13 lamellae;

tibial

spurs

unknown

LOCUS TYPICUS:

Thysville

TYPICAL MATERIAL: HOSTS:

(Zaire)

holotype F!

in TV.

unknown

DISTRIBUTION:

Ethiopian region:

ZAIRE:

Thysville,

TV!

12. Dryinus chamaeleo Benoit

Dryinus chamaeleo Benoit 1953b: 409.

FEMALE: frons;

fully

winged;

length 6,0 mm;

antennae testaceous;

metanotum redd ish-brown; ferruginous;

propectus,

head reddish, pronotum,

propodeum black;

with a brown spot on

scutum,

scutell um and

abdomen reddi sh-brown;

antennae distally weakly thickened;

antennal

segments

legs in

following proportions: 10: 5: 48: 27: 22: 15: 12: 10: 8: 10; head shiny, weakly punctate, with numerous longitudinal keels on median region of frons; carina;

occipital

carina complete;

OPL = OL;

OL = POL;

posterior ocelli

pronotum shiny,

not touching occipital

without sculpture,

except

776

Memo Amero En!.

Ins!.,

noo 37,

1984

Fig. 518 - Chela of Dl'yinus chamaeleo Benoit (type)

777

Dryininae

A

B

Fig.

519 -

Pronotum of females of Dryinus schoutedeni and mayogo (Benoit) (type) (B).

for the sides which show

numerous

longitudinal

impression and anterior collar weak; strong;

posterior collar

not reaching notaulices,

tegulae;

reaching

rugose; distal

fore-wing

part of radial

1 of front produced

into a

14 lamellae; lamellae;

vein

tarsus nearly hook;

without

with 2 brown

nearly

as

keels;

long

twice as

segment 5 of front

tarsus

scutellum shiny,

longitudinal

keels on

bands and with

long as proximal

(Fig. (Fig.

apex with a group of 12 lamellae;

tubercles

punctate;

metanotum punctate,

transversal

as segment 4;

enlarged claw

impression

pronotal

length of scutum;

longitudinal

propodeum reticulate rugose,

posterior surface;

disc humped;

without sculpture,

approximately 0,65

punctate and with numerous

anterior transversal

posterior transversal

long and distinct;

scutum shiny,

keels;

(Benoit) (type) (A)

part;

segment 3 of front

518)

with subapical

518) tibial

segment tarsus

tooth and

with 2 rows of 42 spurs 1,1,2.

(in P!) of D. chamaeleo by Benoit (1953b) is not rei iable, because this male specimen MALE:

unknown;

the description of the male allotype

was not obtained by breeding. LOCUS TYPICUS:

Behara

TYPICAL MATERIAL:

(Madagascar)

holotype F!

in P.

778

Mem.

Fig.

HOSTS:

520 -

Amer.

Chela of

Ent.

Inst.,

no.

37,

Dryinus schoutedeni

1984

(Benoit) (type)

unknown

DISTRIBUTION:

Ethiopian region:

1953b);

(Beno i t 1953b ) •

Bek i I Y

MADAGASCAR:

13. Dryinus schoutedeni

Lestodryinus schoutedeni

FEMALE:

fully

winged;

Benoi t

(Benoit)

1950:

n.

P!

Ivondro (Benoit

comb.

224.

length 8,43-9,37 mm;

with clypeus and mandibles light;

Behara,

head reddish or reddish-dark,

antennae black,

with segments 1-2 and

8-10 testaceous; propectus and pronotum brown-reddish or reddi sh; mesothorax, meta thorax and propodeum black; abdomen brown; legs brownblack,

with tarsi

occipital

light;

head dull,

carina complete;

granulated,

posterior ocelli

weakly reticulate rugose;

located on

the imaginary

779

Dryininae

Fig.

521

-

Chela of

Dryinus mayogo

(Benoit) (type)

straight line joining posterior edges of eyes, not touching occipital carina; OPL = OL; OL = POL; TL nearl y twice as long as POL; pronotum (Fig.

519 A)

dull,

granulated,

with numerous keels around disc;

transversal

impression strong;

anterior collar distinct;

transversal

impression very strong;

posterior collar

anterior

posterior

long;

pronotal

780

Mem.

tubercles not reaching numerous

longitudinal

approximately 0,65 dull;

tegulae;

Ins!.,

rugose;

notaul ices

part;

part of radial

segment

with subapical

1 of front

with 2 rows of 34

18

lamellae;

1984

granulated,

hairy,

incomplete,

with

reaching

scutellum as sculptured as scutum, fore-wing

vein

hook;

long

as segment 4;

enlarged claw

segment 5 of front

apex with a

transversal

nearly twice as long as

into a

lamellae;

with 2

with 2 brown

tarsus nearly as

tarsus produced

tooth and

37,

propodeum reticulate rugose,

keels on posterior surface;

segment 3 of front

no.

scutum dull,

weak keels;

bands and with distal proximal

En!.

length of scutum;

metanotum dull,

longitudinal

Amer.

group of 13

(Fig.

tarsus

lamellae;

520)

(Fig. tibial

520) spurs

1,1,2. MALE:

unknown

LOCUS TYP ICUS:

Banana

TYPICAL MATERIAL: HOSTS;

(Zaire)

holotype F!

in TV.

unknown

DISTRIBUTION: Ethiopian region: ZAIRE: Banana ZIMBABWE: Chishawasha (Salisbury), BM!

(Low-Zaire),

TV!

14. Dryinus mayogo (Benoit) n. comb.

Lestodryinus mayogo Benoit 1950: 225.

FEMALE:

fully

winged;

part of vertex

length 8,12-10 mm;

(more along orbits),

sometimes with some small pronotum black, tarsi

light;

with genae,

reddish spots along orbits;

head dull,

abdomen black-brown;

weakly reticulate rugose;

front head

propectus black;

with sides and posterior collar reddish;

metathorax and propodeum black; with

head black,

clypeus and mandibles yellow;

mesothorax, legs brown-black,

occipital

carina

complete; posterior ocelli not touching occipital carina; OPL = OL; OL shorter than POL; TL little shorter than POL (8:9); pronotum (Fig. 519 B) dull, granulated, with numerous striae around disc; anterior transversal impression very weak; anterior collar almost invisible; posterior transversal

impression very strong;

tubercles not reaching some weak 0,65

irregular keels;

length of scutum;

rugose;

tegulae;

notaulices

hairy,

incomplete,

long;

fore-wing

with 2 short

with 2 brown

pronotal

granulated,

reaching

scutell um as scu I ptured as scutum;

propodeum reticulate rugose,

posterior surface;

posterior collar

scutum very

with

metanotum

longitudinal

transversal

dull,

approximately keels on

bands and with

distal part of radial vein nearly twice as long as proximal part; 1 of front tarsus nearly as long as segment 4; segment 3 of front

segment tarsus

produced into a hook; enlarged claw (Fig. 521) with subapical tooth and 19 lamellae; segment 5 of front tarsus (Fig. 521) with 2 rows of 38 lamellae; MALE:

apex with a group of

unknown

14 lamellae;

tibial

spurs 1,1,2.

Dryininae

781

A

B

c Fig.

522 -

Pronotum of females of Dryinus afer

Benoit from Kankunda

(Zaire) (B)

LOCUS TYP I CUS: Pau lis (Za ire) TYPICAL MATE,qIAL: holotype F! HOSTS:

burgeoni

(Benoit) (type) (C).

in TV.

unknown

DISTRIBUTION: LEONE:

(Olmi)(type)(A),

and orophilus

Ethiopian region:

Freetown,

ZAIRE:

Paulis

(High-Uele),

TV!

SIERRA

TW!

15. Dryinus orophilus (Benoit) n. comb.

Lestodryinus orophilus Benoit 1950: 226. Lestodryinus orophilus Benoit: Benoit 1951c: 7.

FEMALE:

fully

winged;

length 3,75-5,31

mm;

head black,

with clypeus and

782

Mem.

Fig.

523 -

mandibles testaceous; 3,

8-10 testaceous;

Amer.

Chela of

abdomen black;

antennae fully

occipital pronotum

(Fig.

around disc,

522 C)

crossed by

1984

testaceous or brown with segments 1with margins and

metathorax and propodeum black;

longitudinal

OPl = Ol;

shiny,

37,

with coxae and clubs of femora brown;

with some

carina complete;

no.

pronotum black,

mesothorax,

legs testaceous,

granulated,

Ins!.,

Dryinus orophilus (Benoit)(type)

propectus black;

posterior collar reddish; head dull,

Ent.

keels on frons and vertex;

Ol = POL;

granulated,

Tl

longer than POL;

with numerous weak striae

a strong posterior transversal

impression and by

a weak anterior transversal impression, posterior collar short, nearly 0,5 times as long as distance from tegulae to pronotal tubercles; anterior collar weak;

pronotal

tubercles not reaching

tegulae;

scutum dull,

granulated, with some irregular short keels on posterior half; notaulices incomplete, reaching almost posterior margin of scutum; scutellum dull, granulated,

metanotum reticulate rugose;

propodeum reticulate rugose,

without longitudinal keels on posterior surface; fore-wing with 3 brown transversal bands and with distal part of radial vein less than twice as long as proximal segment 4

enlarged claw of front tarsus of 10 lamellae; MALE:

part

(21 :15);

fully

(13:9);

segment

segment 3 of front

(Fig.

523)

1 of front

with subapical

tooth and

(Fig. 523) with 2 rows of 24 tibial spurs 1,1,2.

winged;

tarsus

tarsus produced

length 2,81-3,62 mm;

11

lamellae;

head black,

longer than

into a

hook;

lamellae;

segment 5

apex with a

group

with mandibles

Dryininae

o Fig.

524 -

783

0

Heads of females of Dryinus modestus n. sp.

(Olmi) (type) (B)

and burgeoni Benoit from Kankunda

yellow;

antennae yellow-brown;

brown;

legs

yellow,

thorax and propodeum black;

with coxae partly brown;

head dull,

afer

(type) (A), (Zaire) (C).

abdomen

granulated,

weakly reticulate rugose; occipital carina incomplete; OL shorter than OOL; OPL shorter than POL; OL shorter than POL; scutum du II, as scu I ptured as head;

notaulices incomplete,

reaching approximately 0,65 length of scutum;

scutell um and metanotum as scu I ptured as scutum; rugose; radial

fore-wing vein

without dark transversal

longer than proximal

part;

propodeum ret icu I ate

bands and with distal part of

radial

cell

open;

genitalia:

fig.

513 A; tibial spurs 1,1,2. LOCUS TYPICUS: Rutshuru (Zaire) TYPICAL MATERIAL:

holotype F!

designated after the original

HOSTS; unknown DISTRIBUTION: Ethiopian region: National Park,

TV!

Lusinga

in TV;

allotype M!

invalid,

because

description. ZAIRE:

Rutshuru

(Upemba National

(Kivu),

Park),

TV!

TV!

Garamba

Kalinga-Bitshumbi

784

Mem.

Amer.

(Albert National

Park),

GHANA:

(Kumasi),

Kwadaso

OT!

Ile-ife

(W State),

BM!

ZIMBABWE:

Fig.

TV! BM!

Ent.

Pweto OL!

Chishawasha

no.

37,1984

(Mweru Lake,

BM!

Akure

Ins!.,

NIGERIA:

(W State),

(Salisbury),

Shaba)(Benoit 1951c)j

Kabba OL!

BM!

(Lokoja),

BM!

ANGOLA:

Ikenne

CA!

Vila Luso,

525 - Chela of Dryinus modestus n. sp.

Ibadan,

(W State),

(type)

CA!

Dry i n i nae

CAMERON:

Victoria,

785

CAl

16. Dryinus modestus n. sp.

FEMALE:

fully

winged;

partly clypeus yellow; black,

length 5,0 mm;

head black,

antennae testaceous;

with margins and posterior collar reddish;

and propodeum black;

abdomen black-brown;

and clubs of femora brown; occipital carina complete; OPL shorter than OL; granulated,

OL = POL;

impr"ssion strong;

without

reaching

granulated;

longitudinal

flat,

dull,

granulated;

without sculpture;

posterior collar short; scutum dull,

approximately 0,75

metanotum rugose;

carina;

pronotum shiny, pronotal

posterior transversal granulated;

length of scutum;

propodeum reticulate rugose,

keels on posterior surface;

fore-wing

with 3 brown

transversa I bands and wi th di sta I part of radi a I ve in much proximal

metathorax wi th coxae

not touching occipital

TL shorter than POL;

anterior collar weak;

incomplete,

scutellum dull,

tegulae;

pronotum

mesothorax,

legs testaceous,

524 A)

(Fig.

with anterior collar smooth and

tubercles not reaching notaul ices

head

posterior ocelli

with mandibles and

propectus black;

longer than

(11 :5); segment 1 of front tarsus longer than segment 4

part

(12:9); segment 3 of front tarsus produced into a hook; enlarged claw (Fig.

525)

with subapical

tooth and 10 lamellae;

(Fig. 525) with 2 rows of 20 tibial spurs 1,1,2. MALE:

segment 5 of front

apex with a

group of 11

tarsus

lamellae;

unknown

LOCUS TYP I CUS:

Bek i I y

TYPICAL MATERIAL: HOSTS:

lamellae;

(Madagascar)

holotype F!

in TV.

unknown

DISTRIBUTION:

Ethiopian region:

MADAGASCAR:

Bekily,

TV!

17. Dryinus incertus n. sp.

FEMALE:

fully

winged;

length 3,75 mm;

head black,

with clypeus and

mandibles reddish; propectus black; pronotum black, with sides and posterior collar reddish; mesothorax, metathorax and propodeum black; abdomen brown; granulated;

legs testaceous;

occipital

imaginary straight

antennae testaceous;

carina complete;

carina;

shorter than

POL;

pronotum shiny,

around disc;

anterior transversal

posterior transversal pronotal

head shiny, located on

line joining posterior edges of eyes;

touching occipital

distinct;

posterior ocelli

OPL shorter than OL;

impression

granulated,

OL shorter than POL;

not

TL

with numerous weak striae

impression and anterior collar weak;

strong;

tubercles not reaching

reticulate rugose on

the

posterior ocelli

posterior half;

posterior collar short, tegulae;

notaul ices

scutum dull,

incomplete,

but granulated,

reaching

786

Mem.

Fig.

526 -

Amer.

Ent.

Inst.,

no.

37,

1984

Dryinus incertus n. sp. (type)

Chela of

approximately 0,65 length of scutum; scutellum and metanotum granulated; propodeum reticulate rugose without longitudinal keels on posterior surface; radial

fore-wing

vein

much

with 3 brown

tarsus

longer than segment 4

into a

hook;

lamellae;

transversal

longer than proximal

enlarged claw

segment 5 of front

(9:7);

(Fig.

tarsus

MALE: unknown LOCUS TYPICUS:

Grahamstown

TYPICAL MATERIAL: HOSTS:

bands and with distal (7:3);

with subapical

(Fig. tibial

526)

part of

segment 1 of front

segment 3 of front

526)

apex with a group of 13 lamellae;

part

tarsus produced tooth and 11

with 2 rows of 24

lamellae;

spurs 1,1,2.

(South Africa)

holotype F!

in BM.

unknown

DISTRIBUTION:

Ethiopian region:

SOUTH AFRICA:

Grahamstown,

BM!

18. Dryinus burgeoni Benoit

Dryinus burgeoni Benoi t 1953a: 142.

FEMALE: fully winged; length 6,00-8,43 mm; head black, with clypeus and mandibles reddish; sometimes clypeus black; antennae fully testaceous or

Dryininae

Fig.

527 -

Chela of Dryinus burgeoni Benoit

black with segments 8-10 light; brown-reddish; mesothorax,

metathorax and propodeum black; with

tarsi

light;

carina complete;

OPl shorter than Ol; with

propectus black;

(type)

pronotum fully black or

occasionally black with posterior collar brownish;

reddish-brown, occipital

787

(Fig.

posterior ocelli

Ol ;

longitudinal

numerous

head

and anterior collar weak;

POL;

not

pronotum

keels on sides;

abdomen brown;

524 C)

dull,

legs

granulated;

touching occipital

(Fig.

522 B)

dull,

anterior transversal

posterior transversal

carina;

granulated, impression

impression strong;

posterior collar long and distinct; pronotal tubercles not reaching tegulae; scutum dull, granulated, fully sculptured by numerous longitudinal keels; notaulices almost complete,

reaching almost posterior margin of scutum;

scutellum and metanotum dull, reticulate rugose, wing

without

with 2 brown

curv iii near, nearly as

much

with numerous

longitudinal

transversal

bands and with distal

longer than prox i ma I part;

long as segment 4;

longitudinal

keels;

propodeum

keels on posterior surface; segment 1 of front

segment 3 of front

fore-

part of radial

tarsus produced

vein

tarsus into a

hook; enlarged claw (Fig. 527) with subapical tooth and 13 lamellae; segment 5 of front tarsus (Fig. 527) with 2 rows of 37 lamellae; apex with a group of 10 lamellae; MALE: unknown lOCUS TYP ICUS:

holotype F!

in TV.

unknown

DISTRIBUTION: CAMERON: TV!

spurs 1,1,2.

Moto (Zaire)

TYPICAL MATERIAL: HOSTS:

tibial

Moto

Ethiopian region:

Nkoemvon, (I turi),

Ol!

BM!

NIGERIA:

ZAIRE:

Ibadan,

Kankunda

BM!

Ile-Ife,

(Upemba National

TV!

19. Dryinus afer (Olmi) n. comb.

TW! Park),

788

Mem.

Fig.

528 -

Amer.

Ent.

Inst.,

no.

37,

Female of Dryinus afer

1984

(Olmi)(type)

Paradryinus afer Olmi 1975: 221.

FEMALE

(Figs 528,

529):

fully

winged;

length 5,02 mm;

head black,

with

Dryininae

Fig.

529 -

front part of vertex antennae brown,

789

Female of Dryinus afer

(more along orbits),

clypeus and mandibles yellow;

with segments 1-2 yellow;

posterior collar reddish; abdomen brown;

mesothorax,

legs brown,

(Olmi)(type)

pronotum black,

with sides and

metatorax and propodeum black;

with trochanters and part of fore coxae

yellow; head (Fig. 524 B) dull, granulated; occipital carina complete; posterior ocell i located in front of the imaginary straight line joining posterior edges of eyes; pronotum crossed by 2 transversal impressions (anterior impression very weak; posterior impression strong); pronotum (Fig.

522 A)

dull,

granulated,

and on posterior transversal nearly as

with numerous

impression;

long as distance from

tegulae to pronotal

tubercles not reaching tegulae; longitudinal

longitudinal

keels on sides

posterior collar of pronotum

scutum dull,

short keels on posterior third;

tubercles;

granulated,

with

pronotal numerous

tracks of incomplete notaulices

(reaching approximately 0,5 length of scutum) are visible; scutellum and metanotum dull, granulated and reticulate rugose; propodeum reticulate rugose,

without

longitudinal

keels on posterior surface;

fore-wing

with 2

brown transversal bands and with distal part of radial vein rectilinear, much longer than proximal part (18:8); enlarged claw (Fig. 530) with subapical

tooth and 14 lamellae;

2 rows of nearly 38 of front MALE

tarsus nearly as

(Figs 531,

532):

mandibles yellow; abdomen brown; tarsi

light;

frons flat antennal

lamellae;

segment 5 of front

long as segment 4;

fully

winged;

legs brown,

with tibiae

scutum dull,

invisible;

reticulate rugose,

530)

with

segment 1

spurs 1,1,2. head black,

(except for the hind tibiae)

fully reticulate rugose;

temples

(Fig.

with

thorax and propodeum black;

in front of the anterior ocellus, sockets;

tibial

length 2,71-3,07 mm;

antennae yellow-brown;

head dull,

tarsus

apex with a group of 13 lamellae;

occipital

carina

only convex between

and

incomplete; the

POL = 6; Ol = 3; OOl = 4; OPl = 1;

hairy;

notaul ices weakly visible,

790

Mem.

Amer.

Ent.

Ins! .,

no. 37, 1984

Dryininae

Fig.

incomplete,

-

Male of

Dryinus afer (Olmi) (allotype)

reaching approximately 0,5 length of scutum;

metanotum dull, wing without much

531

791

granulated,

punctate;

dark transversal

longer than proximal

tibial spurs 1,1,2. LOCUS TYP ICUS: Canavial

bands and with distal

part;

radial

cell

closed;

part of radial genitalia:

(Sao Tome)

TYPICAL MATERIAL:

holotype F!

HOSTS:

Ricania insularis lall. (Olmi 1975)

in Sao Tome

nISTRIBUTION:

Ethiopian reclion:

and 3 paratypes MM! SAO TOME':

OR lENT AL SPEC I ES OF

Fifteen

scutellum and

propodeum reticulate rugose;

Canavial.

DI2YINUS

species are known.

KEY TO THE SPECIES

in OL. Ol!

fig.

forevein 533;

792

Mem.

Fig.

Amer.

Ent.

Inst.,

no.

37,

1984

532 - Male of Dryinus afer (Olmi)(allotype)

FEMALES

i

\~

Posterior excavated

collar of pronotum invisible (Fig. 534 A); pronotum (Fig. 534 A) ••••••••••• 1. irregularis n. sp.

Posterior collar of more

2

or

Segment

1 of

long

segment

as

Segment

pronotum visible

less humped,

1 of

front

rarely

flat

tarsus nearl y

534 B); pronotum 534 8) •••.•••.••••••••• 2

(Fig.

(Fig.

twi ce or more

than

twi ce as

4 •••.••••••••••••••.•••••••••••..••••••••••••• 3

front

tarsus

slightly

longer

than

segment

4 or

shorter ••••••••••••••••••••••••••••••••••••••••••••••••••••••• 4

3

Posteri or ocell i black;

fore

touching

coxae

shorter

occipi tal than

carina;

head;

prothorax

fore

trochanters kiefferi (F ou t s )

less

than

three times as long as broad . . . . . 2. Posteri or ocell i not touching occipi tal carina; prothorax redd ish; fore coxae nearl y twi ce as long as head; fore trochanters nearly

five

times as

. . . . . . • . . . . . . . . . . . . . . . . . • . . . . . . .•

4

Posterior ocell i (Fig.

541

touching

long

3.

or almost

as broad •••••••••••••••••

longipes n. touching

sp.

occipi tal

carina

A) ••••••••••••••••••••• ~ •••••••••••••••••••••••••••• 5

Posterior ocelli

farther

from occipital

carina

(Fig.

541

B) . . . .

••••••• •••••••••••• ••••••••••••••••••• ••••••••••••••••••••••• 10

5

Head and pronotum fully

testaceous •••••••••••••..••••••••••••••

• • • • • • . . . . . • • . • • • • • • . . . . . • • • . . . .. 6

4.

pulcher n.

sp.

Head and pronotum mostly black . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6 Head shiny, smooth, hairless, without sculpture, very excavated

Dryininae

Fig.

8

-

Male genitalia of

Dryinus afer (Olmi)(allotype)

5.

sculpture,

shiny

or dull,

granulated,

never without usually

flat,

sometimes

or without

krombeini Ponomarenko

excavated •••••••••••••••••••

Head

f I at ••••••••••••••.•••••••••••...••.••••..••...•••••••.•• 8

Head with

longitudinal

keels

or

Head without

6.

with

excavated ••••••••.• 7

Head

tracks

• •••..••••••.••••.••••••••••..•••

9

exilis n. sp.

• • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • •• Head

keels, 7

533

793

7.

of

longitudinal

keels •••

stantoni Ashmead

longitudinal keels or tracks of longitudinal keels • ., ••••••••••••••••••.•.•• , •••••..•••••.•••.•••••••.•••..•••.• 9 Head longer, with sides of frons more converging (Fig 543 A). • •••••••••••••••••••••••••••••••• 8. trifasciatus K i ef f er

794

Mem.

Head

transverse,

Amer.

wi th

En!.

sides

Inst.,

of

frons

B) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 10 Species full:,Species at

testaceous,

least

less

long as

segment

dorsal

surface; times as

slender,

10.

surface more

antennae more long as

543

3

segment

less

twi ce as

six

long as

3 more 2 ........................ . with

segment

1 1. praeciarus n. 12.

than

indicus (Kieffer)

than

segment

fully black,

long as posterior

slender,

reddish . . . . . . . . . .

Pronotum almost

as

with

2 ......

• • • • • • • . • • • • • • • • • • • . . . . • • • • • . . .. 12 Pronotum fully

(F i g.

converging

• .12

surface nearly

antennae

eight

less

latus n. sp. .. . . . . . . . 11

times as

than

1984

with petiole black.

surface;

Propodeum wi th

9.

37,

partly black ••••••••••••••••

11 Pronotum with dorsal

posterior

no.

sp.

fulvus n. sp.

sometimes partly

reddish . . . . . . . 13

13 Metapleura wi th a narrow region smooth, shiny, hairless, wi thout sculpture ••••••••••••••• 13. lucens n. sp. Metapleura wi thout without

narrow region

smooth,

shiny,

hairless and

sculpture •••••••••••••••••••••.••••••..•••••••••••••• 14

14 Antennae wi th segments 4-5 not broader

than

other

segmen t s

cyl indri c,

(F i g.

but

flat,

much

551) . . . . . . . . . . . . . . . . . . . . . . . ..

14. browni Ashmead· Antennae

with

segments 4-5 as

cylindric as

other

segments,

not

broader than other segments . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15 15 Dorsal surface of propodeum nearly twice as long as posterior surface ••••••.••••••••••••••••••• 7. Dorsal surface of propodeum sl ightly

stantoni Ashmead

15.

lankanus n. sp.

surface •••••••••••••••••••••••••

longer

than posterior

MALES

Dorsal

part

of

occipi tal

carina

laterally

touching

eyes . . . . . . .

10. indicus (Kieffer) Dorsal

part

of

occipital

carina

laterally not touching 14. browni Ashmead

eyes . . . .

The males of the other species are unknown.

1. Dryinus irregular-is n. sp.

FEMALE: fully winged; length 5,00 mm; head fJlack, with mandibles yellow; antennae brown, with segments 1-2 and partly 3 yellow; propectus black; pronotum black, mesothorax,

with sides reddish and posterior margin

metathorax and propodeum black;

weakly reddish;

abdomen brown;

legs

795

Dryininae

A

Fig.

(laterally viewed) of females of Dryinus irregularis n. and st2ntoni Ashmead from Sandakan (Sabah) (B)

534 - Pronotum sp. (type) (A)

testaceous,

with mid and

antennae short

than

hind coxae black;

body,

with segment 3

head shiny,

reticulate rugose;

longer than segment 4;

occipital

carina complete; pronotum (Fig. 534 A) with a median transversal impression, shiny, with deep punctures; anterior surface and sides without sculpture, fully 0,5

smooth;

reticulate rugose;

length of scutum;

pronotal

tubercles not reaching

notaulices almost

invisible,

tegulae;

scutum

approximately reaching

scutellum and metanotum reticulate rugose;

propodeum reticulate rugose,

with 2

longitudinal

keels on

posterior

surface; fore-w ing without dark transversa I bands, with dista I part of radial vein nearly as long as proximal part; segment 1 of front tarsus twice as long as segment 4; segment 3 produced into a hook; enlarged claw with subapical tooth and 6 lamellae (Fig. 535); segment 5 of front tarsus

(Fig.

group of 6 MALE:

535)

with 2 rows of 3

lamellae;

(proximal)

+ 11

lamellae;

spurs 1,1,2.

unknown

LOCUS TYPICUS:

Chiangdao

TYPICAL MATERIAL: HOSTS:

tibial

(Thailand)

holotype F!

in B.

unknown

DISTRIBUTION:

Oriental

region:

THAILAND:

Chiangdao,

2. Dryinus kiefferi (Fouts)

n.

8!

comb.

apex with a

796

Mem.

Fig.

535 -

Amer.

Chela of

Ent.

Inst.,

no.

37,

1984

Dryinus irregularis n. sp. (type)

Lestodryinus kiefferi Fouts 1922: 630. Lestodryinus kiefferi Fouts: Ba I tazar 1966: 203.

FEMALE:

fully

testaceous;

winged;

propodeum black; and tarsi

length 6,87 mm;

antennae brown, light;

head black,

with mandibles

with segments 8-10 yellow;

abdomen black-reddish; head hairy,

flat,

dull,

legs brown, granulated,

thorax and with coxae black

fully sculptured by

numerous longitudinal keels; occipital carina incomplete, visible only behind ocelli and on sides of vertex, where the occipital carina is touching eyes; on sides of occiput the occipital carina is invisible; occipital carina visible on ventral face of head; temples invisible; pronotum dull, hairy, granulated, with some keels on sides; pronotal tubercles not reaching tegulae; transversal

impression strong;

posterior collar short, notaulices complete, dull,

almost invisible among keels;

reticulate rugose;

into 3 regions;

posterior

impression

weak;

longitudinal

keels;

scutellum and metanotum

propodeum reticulate rugose,

fore-wing vein

humped;

fully sculptured by numerous

keels on posterior surface; part of radial

disc not very

anterior transversal

with 2 longitudinal

posterior half of posterior surface is so divided

with 2 brown

transversal

longer than proximal part;

bands and with distal

fore coxae shorter than

head; fore trochanters less than three times as long as broad; segment of front tarsus nearly twice as long as segment 4; segment 3 of front tarsus produced into a hook;

enlarged claw

(Fig.

536)

spatulate,

with

797

Dryininae

Fig.

subapical 536)

535 -

Chela of Dryinus kiefferi

tooth and 13 very

long

with 2 rows of 39 lamellae

other lamellae)

lamellae;

(Fouts)(type)

segemnt 5 of front

(among which 5 lamellae much

extending continuously

to the apex;

tibial

tarsus

(Fig.

longer than

spurs 1.1,2.

798

Mem. Amer.

Fig.

537 -

Ent.

Inst.,

no.

37,

1984

Chela of Dryinus longipes n. sp.

MALE: unknown lOCUS TYPICUS: Mt. Makiling (Philippines) TYP ICAl MATER IAl: holotype F! in WA. HOSTS: unknown DISTRIBUTION: Oriental region: PHiliPPINES: Mt.

(type)

Makiling

(luzon),

WA!

Dryininae

NOTES:

for the morphology of the chela

of the enlarged claw very

long;

lamellae

lamellae)

Moczar.

longer than other The tibial

799

(enlarged claw spatulate;

segment 5 of front

lamellae

tarsus with some

this species is near Nichardsidryinus

spurs however are 1,1,2.

3. Dryinus longipes n. sp.

FEMALE:

fully

winged;

length 6,43 mm;

mand ib I es testaceous; darkened;

prothorax redd i sh;

abdomen black, trochanters,

head black,

antennae testaceous, mesothorax,

meta thorax and propodeum black;

with some regions testaceous;

clubs of fore femora,

with clypeus and

with segments 3 and 4 legs testaceous,

with fore

stalks of mid and hind femora,

distal

apexes of tibiae brown; antennae distally thickened; antenna I segments in following proportions: 14: 7: 49: 30: 26: 19: 15: 11: 9: 9; antenna I segment 4 nearly fifteen

times as long as broad

reticulate rugose and with keel; = 5;

occipital

irregular keels;

carina complete;

pronotum shiny,

POL = 3,5;

without sculpture on

(30:2);

frons

head shiny,

with a median complete

OL = 3;

OOL = 12;

OPL = 3;

reticulate rugose and punctate on sides and on posterior collar; with strong long;

anterior and posterior transversal

pronotal

tubercles not reaching

TL

anterior collar and on disc, impressions;

tegulae;

scutum,

pronotum

posterior collar

scutellum and

metanotum reticulate rugose; notaulices incomplete, reaching approximately 0,65 length of scutum; propodeum reticulate rugose, without transversal keels, with two incomplete longitudinal keels on posterior surface; forewing with two dark transversa I bands, one on the basa I cell s and another beneath part 28;

the pterostigma

(14:11);

segment 3 of front

and slender,

with a subapical

tarsus 7

(Fig.

lamellae;

distal

part of radial

537) tibial

MALE: unknown LOCUS TYPICUS:

vein

longer than proximal

segments in following proportions:

tarsus produced

into a

with coxae nearly twice as

trochanters nearly five 537)

;

fore tarsal

times as

hook;

fore

long as head

long as broad

(30:6);

tooth and with 12 lamellae;

with 2 rows of 25 lamellae;

46:

4:

legs very

(50:25)

Nilgira Hills

and with

HOSTS: unknown DISTRIBUTION: Oriental

region:

apex with a

group of nearly

in OT. INDIA:

(Fig.

segment 5 of front

(India)

holotype F!

18:

long

enlarged claw

spurs 1,1,2.

TYPICAL MATERIAL:

7:

Nilgira Hills

(S

India),

4. Dryinus pulcher n. sp.

Dryinus formicarius Latreille: Ponomarenko 1981b: 879.

OT!

800

Mem. Amer. Ent.

Ins!.,

no. 37,

1984

Fig. 538 - Chela of Dryinus pulcher n. sp.

(type)

FEMALE: fully winged;length 3,25-5,00 mm; head, antennae, propectus a'lG propodeum fully testaceous; mesothorax; metathorax and propodeum black; abdomen brown-testaceous; legs testaceous; head shiny, weakly granulated; occipital carina complete; posterior ocelli almost touching occipital carina;

Dryininae

Fig.

539 -

Chela of

OL slightly

longer than POL;

granu I a ted;

anterior transversa I

posterior transversal pronotal

801

Dryinus exilis n.

temples prominent;

and with some irregular keels;

(type)

pronotum shiny,

impress ion and anterior coil ar weak;

impression strong;

tubercles not reaching

sp.

tegulae;

posterior collar short, scutum dull,

notaulices incomplete,

hairy,

distinct;

granulated

reaching

802

Mem.

approximately 0,5 granulated and short

Amer.

Ent.

Ins!.,

length of scutum;

with

longitudinal

no.

37,

1984

scutellum and metanotum dull,

irregular keels;

propodeum reticulate rugose,

keels on posterior surface;

fore-wing

transversa I bands and wi th di sta I part of rad i a I vei n much proximal

part;

segment

1 of front

tarsus as

with 2

with 2 brown longer than

long as segment 4;

segment 3

of front tarsus produced into a hook; enlarged claw (Fig. 538) with subapical tooth and 9 lamellae; segment 5 of front tarsus (Fig. 538) 2 rows of 18 1,1,2. MALE:

lamellae;

apex with a

group of 6 lamellae;

tibial

with

spurs

unknown

LOCUS TYPICUS:

Labugama

TYPICAL MATERIAL: HOSTS:

(Sri

Lanka)

holotype F!

in BM;

2 paratypes FF!

in WA.

unknown

DISTRIBUTION:

Oriental

Gilimale

Dist.)(Ponomarenko 1981b:

(Rat.

Latrei lie), IVA!

region:

SRI

LANKA:

Udawattakele Sanc!.

Labugama

under name

(Kandy,

Kan.

(Colombo),

BM!

Dryinus formicarius

Dis!.),

WA!

5. Dryinus exilis n. sp.

FEMALE:

fully

winged;

length 3,75 mm;

clypeus and mandibles yellow; brown; shiny,

legs yellow, smooth,

head black,

with clubs of femora and

excavated,

with malar space,

thorax and propodeum black;

without

sculpture,

abdomen

hind coxae brown; with

temples

very

head prominent;

occipital carina complete; posterior ocell i touching occipital carina; pronotum shiny, smooth, without sculpture, with longitudinal keels on sides;

pronotum crossed by a posterior transversal

tubercles not reaching sculpture,

smooth;

weakly granulated; distance between

tegulae;

lateral

scutum shiny,

region

impression;

with central

without sculpture,

notaulices complete,

smooth;

smooth,

posteriorly separated; without sculpture;

without

lateral

the notaulices longer than breadth of ocelli

scutellum and metanotum shiny,

pronotal

region

regions

minimum (6:3);

propodeum

reticulate rugose, without longitudinal keels on posterior surface; forewing with 3 brown transversal bands and with distal part of radial vein much longer than proxi ma I part; segment 1 of front tarsus s light I y longer than segment 4; enlarged claw with subapical tooth and 12 lamellae (Fig. 539);

segment 5 of front

apex with a MALE:

group of 6

(Fig.

539)

tibial

with 2 rows of 28

lamellae;

spurs 1,1,2.

unknown

LOCUS TYPICUS:

IVapikhamthong

TYPICAL MATERIAL: HOSTS:

tarsus

lamellae;

holotype F!

(Laos) in B.

unknown

DISTRIBUTION:

Oriental

region:

LAOS:

Wapikhamthong

(Wapi),

6. Dryinus krombeini Ponomarenko

B!

Dryininae

Fig.

540 -

Chela of

803

Dryinus krombeini Ponomarenko from Culion I. (Philippines)

Dryinus krombeini Ponomarenko 1981 b: 879.

FEMALE: vertex,

fully

winged;

malar space,

length 4,37-6,00 mm;

head black,

clypeus aC'd mandibles yellow;

thorax and propodeum black,

with front part of

antennae testaceous;

sometimes with posterior collar of pronotum

reddish; abdomen brown-reddish; legs reddish-testaceous, with coxae, clubs of femora and tibiae brownish; head excavated, dull, granulated, with some

longitudinal

keels on frons;

occipital

carina complete;

posterior

ocell i almost touching occipital carina; pronotum dull, granulated, with numerous keels around disc; pronotal tubercles not reaching tegulae; scutum granulated, areolae on central

with numerous areolae on region

lateral

near posterior margin;

regions

lateral

and with some

regions usually

804

Mem.

Fig.

541

-

Amer.

Ent.

and fulvus

with areolae on posterior half; metanotum granulated;

37,

part of radial tarsus slightly

n.

sp.

1984

(type) (8)

notaulice almost complete;

propodeum reticulate rugose,

keels on posterior surface; front

no.

Heads of females of Dryinus stantoni Ashmead from Sandakan (Sabah) (A)

distal

Ins!.,

fore-wing

vein much

with 3 brown

longitudinal

transversal

longer than proximal

longer than segment 4;

scutellum and

without part;

bands;

segment 1 of

enlarged claw with subapical

tooth and 13 lamellae (Fig. 540); segment 5 of front tarsus (Fig. 540) 2 rows of 31 lamellae; apex with a group of 11 lamellae; tibial spurs 1,1,2. MALE:

unknown

LOCUS TYPICUS:

Udawattakele

TYPICAL MATERIAL: HOSTS:

with

(Sri

holotype F!

Lanka)

in WA.

unknown

DISTRIBUTION:

Oriental

region:

VIET NAM:

Phan Rang,

B!

LAOS:

Ban Van

Eue (Vientiane), OL! B! Wapikhamthong (Wapi), B! THAILAND: Mae Sa (Chiang Mai), B! Konthathan waterfall area (Doi Suthep-Pui natn. Park), CO! PHILIPPINES: Dis!.), WA!

Culion

Island,

OL!

SRI

LANKA:

Udawattakele

(Kan.

Dryininae

Fig.

805

542 - Chela of Dryinus stan toni Ashmead from Forest Camp (Sabah)

806

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

7. Dryinus stan toni Ashmead

Dryinus stan toni Ashmead 1904: 134. Prodryinus stantoni (Ashmead): Fouts 1922: 629. Paradryinus javanus R.C.L. Perkins 1912: 11 (n. syn.); locus typicus Pekalongan (Java). Lestodryinus luzonicus Kieffer 1913a: 189 (n. syn.); locus typicus Mt.

Maki I ing

(Phi I ippine

Islands).

Lestodryinus stan toni (Ashmead): Fouts 1922: 629. Neodryinus stantoni (Ashmead): Baltazar 1966: 204. Dryinus szepligettii Kieffer partim: Ponomarenko 1981 b: 879.

FEMALE:

fully

winged;

length 3,12-5,62 mm;

head black,

with malar space,

clypeus and mandibles yellow; antennae testaceous, sometimes brownish; rarely with last segment whitish; pronotum black, with sides and posterior collar reddish;

sometimes posterior collar black;

meta thorax black; A)

dull,

abdomen brown-reddish;

granulated,

complete;

with some longitudinal

posterior ocell i

touching occipital

mesothorax and

legs testaceous; keels;

located very near occipital

carina);

head

occipital

(Fig.

541

carina

carina

(but not

pronotum crossed by 2 transversal

impressions

(anterior impression weak; posterior impression strong); pronotum (Fig. B) du II, ha i ry, granu I a ted; pronota I tuberc I es not reach i ng tegu I ae; scutum

dull,

granulated,

hairy;

notaul ices

incomplete,

534

reaching

approx i matel yO, 5 length of scutum; scute II um and metanotum as scu I ptured as scutum; propodeum rugose, without longitudinal keels on posterior surface; radial

fore-w i ng

vein

much

with 3 brown

slightly

longer than segment 4;

lamellae

(Fig.

lamellae;

transversa I bands and with di sta I part of

longer than proximal

542);

part;

segment

1 of front

enlarged claw with subapical

segment 5 of front

tarsus

apex with a group of 14 lamellae;

(Fig.

tibial

542)

tarsus

tooth and 11

with 2 rows of 27

spurs 1,1,2.

MALE: unknown LOCUS TYPICUS: Manila (Philippines) TYPICAL ,'vlATERIAL: lectotype F! and 1 para lectotype F!

(designated by

javanus in B; holotype F! of D. stantoni in WA; holotype F! of L. luzonicus in WA. HOSTS: in Java Thanatodictya sp. (Dictyopharidae) (R.C.L. Perkins 1912). PARASITES: in Java probably Cheiloneurus javanus R.C.L. Perkins (R.C.L.

Olmi

1982) of P.

Perkins 1912). DISTRIBUTION: Oriental region: INDONESIA: Pekalongan (Java), B! PHILIPPINES: Manila (Luzon), WA! La Trinidad (Luzon), B! Mt. Polis (Luzon),

MD!

Nueva Vizcaya

(Imugan,

Luzon),

B!

Mt.

Makiling

(Luzon),

WA! Zamboanga (Mindanao), B! Wa-wa Dam (Rizal, Mt. Montalban), B! Pinigisan (Mantalingajan, Palawan), CO! Tagembung (Mantalingajan, Palawan), TAIWAN:

CO!

THAILAND:

Klapan,

8!

Chiangdao

VIET NAM:

(Chiangmai),

Pleiku,

B!

Kontum

OL!

B!

Banna,

(Pleiku),

B!

B!

M'Drak

Dryininae

Fig.

543 -

Heads of females of JJryinus

Wapikhamthong

(Ban Me Thuot),

B!

LAOS:

Sayaboury,

Sion,

B!

MALAYSIA:

8M!

Forest Camp

(Kalabakan,

(Sabah),

Kanneliya Jungle (Induruwa Jungle,

OL!

8!

Phou-kow-kuei,

Port Dickson

Sabah),

B!

Singkor Sabah),

(Gal.

Dist.),

Rat.

from

Ban Van Eue (Vientiane),

B!

Vientiane),

trifasciatus Kieffer from

(Laos)(A) and latus n. sp. (Phi I ippines) (B).

(Phou-kow-kuei),

Sandakan

807

Dist.),

WA!

(Negeri SRI

Lahugala

WA!

B! 8!

LANKA:

(Amp.

HONG KONG:

Ban Van

Heue

De Tha Ngone

Sambi lan,

Gunong Mulu B!

Irawan Riv.

(Sarawak), Kandy,

Dist.),

Taipokau,

WA!

(Gi

Malaya), OL!

B~1!

BM! Gilimale

B!

8. JJryinus trifasciatus Kieffer

Dryinus trifasciatus Kieffer 1906b: 335.

FEMALE:

fully

winged;

length 3,75-4,37 mm;

head black,

with clypeus and

mandibles reddish; antennae testaceous; pronotum black, with sides and posterior collar reddish; mesothorax, meta thorax and propodeum black; abdomen brown, legs fully testaceous or with clubs of fore femora and hind coxae brown;

head

(Fig.

543 A)

flat,

shiny,

smooth,

weakly

808

Fig.

Mem. Amer.

544 -

Ins!.,

no.

37,

1984

Chela of Dryinus trifasciatus Kieffer from Wapikhamthong

granulated; ocelli

Ent.

sides of frons converging;

touching occipital

carina;

occipital

carina complete;

impression,

with anterior collar smooth and without sculpture; pronotal

granulated;

notaul ices more or less visible,

propodeum

surface; radial

tubercles not reaching

reticulate

rugose,

544);

without

with 3 brown

extending continuously MALE: unknown

part;

tarsus

to the apex;

Matheran

region:

MALAYSIA:

(Kalabakan,

(Maharashtra),

BU!

tibial

LAOS:

part of

tarsus sl ightly

tooth and 6 lamellae

with 2 rows of 37 lamellae

spurs 1,1,2.

Wapikhamthong

Sabah),

Cheramgode (Nilgiri

Pinigisan

544)

and 1 para lectotype F!

HOSTS: unknown DISTRIBUTION: Oriental

PHILIPPINES:

segment 1 of front

(Fig.

posterior

(India)

TYPICAL MATERIAL: lectotype F! Moczar 1965) in BU.

Forest Camp

keels on

bands and with distal

enlarged claw with subapical

segment 5 of front

LOCUS TYPICUS:

disc weakly

scutum dull,

reaching approximately 0,5

longitudinal

transversal

longer than proximal

longer than segment 4; (Fig.

tegulae;

scutell um and metanotum as scu I ptured as scutum;

fore-wing

vein

posterior

pronotum with a posterior transversal

granulated;

length of scutum;

(Laos)

(Mantalingajan,

B!

Hills,

(Wapi),

INDIA: S.

Palawan),

(designated by

OLl

Matheran

India),

TW!

CO!

9. Dryinus latus n. sp.

FEMALE:

fully

winged;

length 2,68-3,12 mm;

head black,

with clypeus and

Dryininae

Fig.

545 -

Chela of Dryinus latus n. sp.

809

(type)

810

Mem.

mandibles yellow;

antennae fully

and 10 testaceous;

Amer.

Ent.

Ins!.,

no.

37,

1984

testaceous or brown with segments 1-2

pronotum black,

with sides and posterior collar

redd i sh; mesothorax, meta thorax and propodeum black; abdomen brown; legs fully testaceous or brown with tarsi light; head (Fig. 543 B) dull, weakly excavated,

granulated;

touching occipital around disc;

carina;

pronotal

occipital

carina complete;

pronotum shiny,

granulated,

tubercles not reaching

tegulae;

posterior ocelli

with numerous keels scutum dull,

hairy,

granulated, with tracks of keels on posterior half; notaul ices incomplete, reaching approximately 0,5 length of scutum; scutellum as sculptured as scutum;

metanotum rugose;

longitudinal bands and

propodeum reticulate rugose,

keels on posterior surface; with distal

part of radial

fore-wing

vein much

without

with 3 brown

transversal

longer than proximal

part; segment 1 of front tarsus sl ightly longer than segment 4; enlarged claw (Fig. 545) with subapical tooth and 9 lamellae; segment 5 of front tarsus

(Fig.

545)

with 2 rows of 14 lamellae;

lamellae; tibial MALE: unknown

spurs 1,1,2.

LOCUS TYP ICUS:

Zamboanga

TYPICAL MATERIAL: FF! in OL. HOSTS:

apex with a group of 7

(Phi I ippines)

holotype F!

and 2 paratypes FF!

in B;

2 paratypes

unknown

DISTRIBUTION: Irawan Riv.

Oriental

(Mt.

Wapikhamthong

region:

Beaufort,

(Wapi),

B!

PHILIPPINES:

Palawan), VIET NAM:

OL!

Zamboanga

THAILAND:

Dalat,

(Mindanao),

Banna,

B!

B!

LAOS:

OL!

10. Dryinus indicus C
Nesodryinus indicus Kieffer 1914c: 311. Mesodryinus indicus (Kieffer): Pruthi and Mani 1942: 425. Chlorodryinus pallidus R.C.L. Perkins: Pruthi and Mani 1942: 425. i\,'esodryinus indicus Kieffer: Kurian 1954a: 256. Chlorodryinus pallidus R.C.L. Perkins: Kurian 1954a: 257. Mesodryinus indicus Kieffer: Kelner-Pillault 1958: 146.

FEMALE:

fully

antennal

segments

8:

11;

winged;

length 3,56-5,00 mm;

in following proportions:

testaceous, 11:

5:

41:

proportions of antenna I segments 2-3 different

specimens

(in small

granulated;

on sides of vertex; without sculpture,

impression; distinct;

specimens 5:22;

occipital carina POL

13:

10:

10:

and big

head dull,

visible only behind the ocelli

and

= 3; OL = 3; OOL = 7; OPL = 1,5; pronotum shiny,

smooth,

anterior collar

21:

in small

in big specimens 5:41);

incomplete,

crossed by a posterior strong

anterior transversal

scutum shiny,

with petiole black; 25:

weak;

impression weak; pranatal

strongly punctate;

tubercles

transversal

posterior collar short, not

reaching

notaulices reaching 0,5

tegulae;

length of scutum

Dryininae

Fig.

546 -

Chela of

(posterior half of scutum punctate;

longitudinal

Dryinus indicus (Kieffer)(type)

not visible,

metanotum rugose,

811

du II;

hidden by pin);

scutellum shiny,

propodeum reticu I a te rugose,

keels on posterior surface;

fore-wing

without dark

wi thout transversal

812

Mem.

Fig.

547 -

Amer.

Male genitalia of

En!.

Ins!.,

no.

37,1984

Dryinus indicus (Kieffer) from Pusa (India): left half removed.

bands,

with distal

segment front

1 of front

tarsus produced

subapical

tooth and 9

2 rows of 16 1,1,2. MALE:

part of radial tarsus

fully

lamellae; winged;

vein much

longer than proximal

longer than segment 4

into a

hook;

lamellae;

(15: 12);

enlarged claw

segment 5 of front

apex with a group of length 2,20-3,60 mm;

head with anterior part of frons,

testaceous;

antennae not distally

546)

with

tarsus

(Fig.

546)

lamellae;

tibial

spurs

black;

testaceous;

thickened;

12

(Fig.

part;

segment 3 of with

legs and antennae

clypeus and mandibles antennal

segments

in following

Dryininae

Fig.

548 -

Chela of

Dryinus praeclarus n. sp. (type)

813

814

Mem.

proportions:

7:

7:

19:

Amer.

14:

En!.

14:

14:

Ins!.,

14:

frons with a complete median keel; part

laterally reaching

eyes;

scutellum and metanotum dull, transversal

transversal

bands;

or

HOSTS:

in

10:

11;

head dull,

carina

OL = 4;

granulated;

granulated;

incomplete,

OOL = 4;

with dorsal

OPL = 1,5;

holotype F!

propodeum reticulate rugose,

keels;

part of radial

(17:10); genitalia: fig. 547; tibial LOCUS TYPICUS: Pusa (India) TYPICAL MATERIAL:

1984

granulated, with posterior third rugose; approximately 0,65 length of scutum;

longitudinal

distal

37,

occipital

POL = 7;

temples invisible; scutum dull, notaul ices incomplete, reaching without

12:

no.

fore-wing vein

hyal ine,

without dark

longer than proximal

part

spurs 1,1,2.

and 4 paratypes FF!

in P.

Phromnia marginella (Kieffer 1914b); Pyrilla sp. (I(urian

India

1954a) . DISTRIBUTION:

Oriental

region:

Pusa

(Bihar),

P!

ND!

11. Dryinus praeclarus n. sp.

FEMALE:

fully

winged;

antennae slender, proportions: weakly

10:

length 7 mm;

distally

5:

45:

granulated;

30:

and on with a

shiny,

occipital

flat,

10:

8:

10;

in

and without sculpture;

carina

incomplete,

POL =2;

following

head flat,

shiny,

frons with a

visible behind the ocell i

OL = 4,5;

OOL = 8,5;

transversal

posterior collar very short;

pronotum

without sculpture,

keels on

impression;

incomplete,

weak

visible only

and metanotum sh i ny,

punctate,

tubercles not reaching tracks of areolae;

and

except for

tegulae;

near anterior mar"in of scutum;

propodeum reticulate rugose,

longitudinal

dorsal

scutum

notaulices weakly

without scu I pture among

weakly alutaceous; keels;

OPL = 2,5;

impression

sides and for some short keels on posterior

pronotal

granulated and with

visible,

a

11:

posterior

longitudinal

with

with petiole black;

antenna I segment

weak

transversal dull,

15:

sides of posterior ocelli;

pronotum

some

26:

temples long

median complete keel;

testaceous,

thickened;

scutellum

punctures or

without transversal

surface of propodeum more than

twice as

or long as

posterior surface (27: 10); fore-wing hyal ine, without dark transversal bands; distal part of radial vein longer than proximal part (26:9); fore tarsal (Fig. tarsus

segments 548)

(Fig.

lamellae;

in following proportions:

with a 548)

tibial

MALE: unknown LOCUS TYPICUS:

subapical

tooth and

with 2 rows of 21

12

32:

3,5:

lamellae;

lamellae;

5:

20:

26;

enlarged claw

segment 5 of front

apex with a group of 7

spurs 1,1,2. Los Banos

TYPICAL MATERIAL:

(Philippines)

holotype F!

HOSTS: unknown DISTRIBUTION: Oriental

region:

in TW. PHILIPPINES:

Los Banos

(Luzon),

TW!

Dryininae

815

/r~

I

!

/cf '

cf=

t

\ \

Fig.

549 -

Chela of

Dryinus fulvus n. sp. (type)

12. Dryinus fulvus n. sp.

FEMALE: vertex, reddish,

fully

winged;

length 5,62 mm;

clypeus and mandibles yellow; with brown

spots on

head black,

with front part of

antennae testaceous;

anterior weak

collar;

pronotum

propectus reddish;

816

Mem.

Fig.

mesothorax; testaceous,

Ent.

Chela of

Inst.,

no.

(Fig.

keels;

541

B)

occipital

dull,

around disc;

pronotal

1984

abdomen brown-reddish;

granulated,

carina complete;

carina pronotum dull,

37,

Dryinus lucens n. sp. (type)

metathorax and propodeum black; head

longitudinal occipital

550 -

Amer.

with a few

posterior ocelli

weakly granulated,

tubercles not reaching

legs

weak not touching

with numerous keels

tegulae;

scutum dull,

hairy,

Dryininae

granulated; 0,5

notaul ices almost

length of scutum;

reticulate rugose, transversal proximal

of 8

incomplete,

reaching

longitudinal

keels;

fore-wing

part of radial

approximately

granulated;

vein

propodeum

with 3 brown longer than

longer than segment 4;

tarsus slightly

549) with subapical tooth and 9 lamellae; segment 5 549) with 2 rows of 21 lamellae; apex with a group tibial spurs 1,1,2.

(Fig.

tarsus

(Fig.

lamellae;

MALE:

without

segment 1 of front

enlarged claw of front

invisible,

scutellum and metanotum dull,

bands and with distal

part;

817

unknown

LOCUS TYP ICUS:

Kontum

TYPICAL MATERIAL: HOSTS:

(Viet Nam) in B.

holotype F!

unknown

DISTRIBUTION:

Oriental

region:

VIET NAM:

Kontum

(Plateau),

B!

13. Dryinus lucens n. sp.

FEMALE:

fully

winged;

mandibles reddish; abdomen

reddish;

length 6,25 mm;

legs reddish;

carina complete;

impression strong);

numerous

keels

around

smooth,

regions punctate; of scutum;

di sc;

notaul ices

reticulate rugose,

without

with 3 brown

much

longer than proximal

550);

Ranau

TYPICAL MATERIAL: HOSTS:

tarsus

group of 15

MALE: unknown LOCUS TYPICUS:

part;

enlarged claw

segment 5 of front

apex with a

incomplete,

bands,

reaching

(Sabah,

part of radial

550)

tibial

forevein

tarsus slightly

tooth and

length

propodeum

keels on posterior surface;

segment 1 of front (Fig.

weak; with

approximately 0,5

metanotum dull,

with distal

with subapical

lamellae;

holotype F!

without sculpture,

not reach i ng tegu I ae; without sculpture; lateral

weakly punctate;

transversal

pronotum

tuberc I es

region

longitudinal

wing

than segment 4;

pronota I

occipital

carina;

(anterior impression very

pronotum shiny,

with central

scutellum shiny,

reticulate rugose;

not touching occipital

impressions

posterior

with clypeus and

thorax and propodeum black;

head dull,

posterior ocelli

crossed by two transversal

scutum shiny,

head black,

antennae testaceous;

longer

12 lamellae (Fig.

with 2 rows of 28

lamellae;

spurs 1,1,2.

Malaysia) in

B.

unknown

DISTRIBUTION:

Oriental

region:

MALAYSIA:

Ranau

(Sabah),

B!

14. Dryinus browni Ashmead

Dryinus browni Ashmead 1905: 109. Paradryinus terryi R.C.L. Perkins 1912: 10 (n. s)-n.); locus typicus Hong Kong

(China).

818

Mem.

Amer.

Ent.

Ins!.,

no.

37,

1984

5

3

4

Fig.

551

-

Dryinus browni Ashmead: antenna I segments 3-6.

Female of

Lestodryinus browni (Ashmead): Kieffer 1914b: 24. Paradryinus terryi R. c. L. Perk i ns: Ki effer 1914b: 35. Dryinus szepligettii Kieffer partim: Ponomarenko 1981 b: 879.

FEMALE:

fully

winged;

mandibles yellow; yellow;

antennae brown,

propodeum black; light;

length 6,25-7,68 mm;

antennal

head black,

with clypeus and

sometimes also malar space and front part of vertex with

abdomen

last 2-4 segments whitish;

brown;

thorax and

legs testaceous or brown,

with

tarsi

much broader than other segments

segments 4-5 flat,

(F i g.

551}; head dull, granulated, with some longitudinal keels on sides of frons; occipital carina complete; posterior ocell i not touching occipital carina;: pronotum dull, granulated, hairy, with numerous keels around disc; pronotal tubercles not reaching tegulae; pronotum crossed by two

transversal

impressions {anterior impression very weak;

posterior

impression strong}; scutum hairy, dull, granulated, rugose on posterior third; notaulices complete, posteriorly separated; scutellum as sculptured as scutum;

propodeum reticulate rugose,

posterior surface; distal

fore-w i ng

part of radial

wi th

vein much

without

three brown longer than

longitudinal

keels on

transversa I bands and with proximal

part;

segment

1 of

front tarsus slightly longer than segment 4; enlarged claw (Fig. 552) with subapical tooth and 14 lamellae; segment 5 of front tarsus (Fig. 552) with 2 rows of 40 1,1,2.

lamellae;

apex with a

group of 19 lamellae;

tibial

spurs

Dryininae

Fig.

MALE:

552 -

fully

Dryinus browni

Chela of

winged;

antennae yellow-brown;

brown;

legs testaceous,

=

head black,

with hind coxae partly brown; 9;

OL

with mandibles

thorax and propodeum black;

granulated and reticulate rugose; POL

Ashmead from De Tha Ngone (Laos)

length 3,25-3,75 mm;

yellow;

antenna I sockets;

819

=

frons with a median keel 4;

OOL

=

5;

OPL

=

2;

abdomen

head dull, between the

TL = 3;

occipital

Mem.

820

Amer.

Ent.

Inst.,

no.

37,

1984

\

\

\ \

I I /

Fig.

553 -

Male genitalia of Dryinus browni Ashmead from Kalutara Lanka):

right

half removed.

(Sri

Dryininae

carina complete;

temples

reticulate rugose; shiny,

scutum dull,

notaulices complete,

granulated;

longitudinal

invisible;

metanotum rugose;

bands and with distal

part of radial

(15:9);

553;

lOCUS TYPICUS:

fig.

Manila

TYPICAL MATERIAL: terryi in B. HOSTS:

PHiliPPINES: GV!

(Gi Sion,

Oriental Manila

Vientiane),

(Kanneliya Jungle,

scutellum

propodeum reticulate rugose, fore-wing

vein much

without dark

without

transversal

longer than proximal

part

spurs 1,1,2.

of D.

browni in WA; holotype F! of P;

(=Pyrops) dohrni (Fulgoridae).

region:

(luzon),

Pinigisan

weakly

(Philippines)

holotype F!

in Sri lanka Zanna

DISTRIBUTION: Island,

tibial

granulated,

posteriorly separated;

keels on posterior surface;

genitalia:

821

CHINA:

Ta Hian

(Hainan

Id.),

WA!

los Banos

(luzon),

(Mantalingajan,

Palawan),

CO!

B!

Cal.

SRI

Ol!

lANKA:

Kalutara,

BM!

lAOS:

Ol!

B!

TW! WA!

Bilaran

De Tha Ngone

Hiniduma

under name Dryinus

Dist.)(Ponomarenko 1981b:

szepligettii Kieffer), WA! MALAYSIA: Gn. Mulu (Sarawak), BM! HONG KONG, B!

15. Dryinus lankanus n. sp.

Dryinus trifasciatus Kieffer: Ponomarenko 1981b: 879.

FEMALE: fully winged; length 3,60-4,88 mm; head black, with clypeus, malar space, mandibles and part of ventral side testaceous; antennae testaceous with segments 9-10 light or brown thorax and propodeum black, of pronotum

testaceous or reddish;

testaceous or black, antennal 7:

4:

with tarsi

10: 8:

6:

5:

5:

4:

abdomen black or brown;

light;

segments 4-5 not flat;

21,5:

with segments 9-10 testaceous,

with posterior collar of pronotum and sides antennal

antenna I segments in following proportions: 5;

head flat,

irregular keels on frons and vertex;

frons

dull,

not reaching

tegulae;

around disc;

scutum dull,

impression;

pronotum dull,

=

10: OPl = 1,5; Tl impression and by a

disc humped;

granulated,

granulated,

keels near posterior margin;

granulated and with

with a complete median keel;

occipital carina complete; POL = 5; Ol = 4; OOl pronotum crossed by a weak anterior transversal strong posterior transversal

legs

segments cylindric;

pronotal

tubercles

reticulate rugose or with weakly

irregular visible

scutell um and metanotum du II,

granulated;

propodeum reticulate rugose,

longitudinal

keels;

fore-wing

with 3 dark transversal

bands beneath the

pterostigma and on

the basal

cells;

distal

vein much longer than proximal following proportions: 28: 5: 9: subapical

5;

with numerous striae

notaul ices almost complete,

near posterior marg i n of scutum;

=

without transversal or

radial

cell

open;

part of radial

part (10:5); fore tarsal segments in 21: 33; enlarged claw (Fig. 554) with a

tooth and with a row of 11

lamellae;

segment 5 of front

tarsus

822

Fig.

Mem. Amer.

Ent.

Inst.,

no.

37,

1984

554 - Chela of Dryinus lankanus n. sp. from Kandy

(Sri

lanka)

(Fig. 554) with two rows of nearly 31 lamellae; apex with a group of nearly 15 lamellae; tibial spurs 1,1,2. MALE: unknown lOCUS TYPICUS: Kibissa (Sri lanka) TYPICAL MATERIAL: holotype F! and 9 paratypes FF! in WA; 1 paratype F! in Ol. HOSTS: unknown DISTRIBUTION: Oriental region: SRI lANKA: Kibissa (W of Sigiriya, Mate Dist.), WA! Angunakolapelessa (Mon. Dist.), Ol! WA! Kandy (Kan. Dist.), WA! Parayanalankulam (25 mi. NW Medawaehehiya, Vav. Dist.), WA! Hasalaka (Kan. Dist.), WA! Ekgal Aru Palatupana tank (Ham. Dist.), WA!

(Sanet. Jungle, Amp.

NEARCTIC SPECIES OF

DRYINUS

Dist.), WA!

823

Dryininae

Four species are known.

KEY TO THE SPECIES

FEMALES

Segment

1 of

enlarged Segment 4; 2

1 of

enlarged

tarsus nearly

front

tarsus at

claw not

Head with TL

twi ce as

long as

segment

4;

longer

most

1,5

times as

long as

segment

spatulate •••••••••••••••••••••••••••••••• 3

propectus and pronotum 1. mexicanus (R.C.L. Perkins) Head with TL shorter than POL; head (except for yellow mandibles), propectus and pronotum black ••••••••••••••••••••••• . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2. crawfordi (Krombei n) Head (dorsally viewed) wi th posterior ocell i located in front of the imaginary straight line joining posterior edges of eyes (Fig. 557 A) •••••••••••••• 3. ameMcanus (Ashmead) Head (dorsally viewed) wi th posterior ocell i located behind the imaginary straight I ine joining posterior edges of eyes (Fig. 557 B) ••••••••••••••••••••••••• 4. alatus (Cresson) fully

3

front

claw spatulate ••••••••••••••••••••••••••••••••••••••• 2

than POL;

head,

reddish-testaceous . . . . . . .

MALES

Only the male of

D. alatus

(Cresson)

1. Dryinus mexicanus

Paradryinus mexicanus

FEMALE:

fully

winged;

abdomen reddish-brown; very prominent;

Perkins)

head,

mesothorax,

comb.

antennae,

propectus and

meta thorax and propodeum black;

legs testaceous-reddish;

occipital carina complete;

pronotum dull,

n.

Perkins 1907: 39.

length 4,68 mm;

pronotum reddish-testaceous; reticulate rugose;

R.C.L.

(R.C.L.

is known.

head swollen,

dull,

POL shorter than OL;

crossed by a strong transversal

temples

posterior

impression, granulated; anterior transversal impression weak; disc humped; posterior collar very visible; anterior collar weak; pronotal tubercles not reaching tegulae; on

lateral

scutum

dull,

regions and near posterior margin;

granulated,

reticulate rugose

notaulices almost complete,

824

Mem.

Fig.

reaching

555 -

Amer.

Ent.

Ins!.,

no.

Chela of Dryinus mexicanus

almost posterior margin of scutum;

37,

1984

(R.C.L.

Perkins)(type)

scutellum and metanotum dull,

granulated; propodeum reticulate rugose, without longitudinal keels on posterior surface; fore-wing with 2 brown transversal bands and with distal front hook;

part of radial

vein much

longer than

proximal

tarsus nearly twice as long as segment 4; enlarged claw

lamellae; (among

(Fig.

555)

with subapical

segment 5 of front tarsus

which 6 lamellae much

continuously to the apex; MALE:unknown LOCUS TYPICUS:

tibial

(Fig.

555)

part;

tooth and with 5 very

holotype F!

HOSTS: unknown DISTRIBUTION: Nearctic region:

longer than other lamellae) spurs 1,1,2.

in WA. MEXICO:

long

with 3 rows of 36 lamellae

Patzcuaro (Mexico)

TYPICAL MATERIAL:

segment 1 of

segment 3 produced into a

Patzcuaro,

WA!

extending

Dryininae

------------------Fig.

556 -

Chela of

825

-

Dryinus crawfordi (Krombein) (type)

2. Dryinus crawfordi (Krombein) n. comb.

Mesodryinus crawfordi Krombein 1962: 3.

FEMALE:

fully

winged;

antennae brown, black;

abdomen brown-reddish;

head dull,

granulated,

of vertex;

occipital

granulated, distinct; notaul ices

carina complete;

posterior

granulated,

incomplete,

notaulices reticulate; without

10 yellow;

legs testaceous,

with mandibles yellow;

thorax and propodeum

with coxae partly black;

OPL shorter than OL;

anterior transversal

impression weak;

disc not very swollen;

scutum dull,

head black,

with numerous areolae or short keels on front part

not very humped;

collar invisible;

rugose,

length 4,37 mm;

with segments 1-3 and

pronotal

posterior collar short,

tubercles not reaching

with numerous areolae on

lateral

reaching posterior margin of scutum; scutellum and metanotum dull,

longitudinal

pronotum dull,

impression and anterior

furrow of

granulated;

keels on posterior surface;

tegulae;

regions; reticulate

fore wings with 2

826

Mem. Amer.

Ent.

Inst.,

no.

37,

1984

o

Fi g.

557 -

Heads of fema les of lJryinus american us

0

(Ashmead) (type) (A)

and

alatus (Cresson) from Nogales (Arizona) (B).

brown

transversal

than proximal segment 4; subapical

bands and with distal part of radial

part;

segment 1 of front

segment 3 produced into a hook; tooth and 9 lamellae;

3 rows of 45

lamellae (among

tarsus

lamellae much

to the apex;

LOCUS TYP ICUS: Plummers Island (Maryland, TYPICAL MATERIAL: holotype F! in WA. HOSTS:

enlarged claw

segment 5 of front

which 5

lamellae) extending continuously MALE: unknown

vein much

tarsus nearly twice as

tibial U.

S.

longer

long as

(Fig. (Fig.

556) 556)

with with

longer than other spurs 1,1,2.

A.)

unknown

DISTRIBUTION: Tompkins Co.

Nearctic region: (S.

Lansing,

N.

U. Y.),

S.

A.:

Plummers

Island

(Md.)

WA!

OLl

3. Dryinus american us (Ashmead)

Mystrophorus american us Ashmead 1887b: 128. Dryinus americanus (Ashmead): Ashmead 1893: 87. Prodryinus american us (Ashmead): Kieffer 1907b: 9. Mesodryinus americanus (Ashmead): Muesebeck and Walkley 1951: 1034.

FEMALE: black

fully

winged;

length 5,0 mm;

and abdomen brown-black;

head

reddish-testaceous, (Fig.

557 A)

shiny,

with petiole weakly

granulated, weakly rugose on frons; occipital carina incomplete, only visible behind the ocell i and on sides of vertex; posterior ocell i almost touching occipital

carina;

OL sl ightly

longer than POL;

vertex flat;

827

Dryininae

Fig.

558 -

pronotum shiny, disc not very

Chela of

granulated,

humped;

Dryinus american us

crossed by 2 weak

anterior collar invisible;

(Ashmead) (type)

transversal

impressions;

posterior collar

invisible,

because the posterior transversal impression is just on posterior margin of pronotum; pronotal tubercles not reaching tegulae; scutum shiny,

828

Mem.

granulated,

Amer.

Ent.

with weak areolae on

not very visible,

reaching

I nst.,

lateral

no.

regions;

approximately 0,65

and metanotum as scu I ptured as scutum, propodeum reticul a te rugose;

37,

1984

notaul ices incomplete,

length of scutum;

scutellum

but wi thout weak areol ae;

posterior surface wi th transversa I keel sand

with 2 longitudinal keels; the posterior surface so is divided into 3 regions; fore wings with 3 brown transversal bands and with distal part of radial

vein slightly

tarsus nearly as into a

hook;

lamellae;

enlarged claw

segment 5 of front

extending continuously MALE:

longer than proximal

long as segment 4; (Fig.

558)

tarsus

part;

segment 1 of front

segment 3 of front with subapical

(Fig.

to the apex;

tibial

558)

tarsus produced

tooth and 8

with 2 rows of 35

lamellae

spurs 1,1,2.

unknown

LOCUS TYP ICUS: Jacksonvi lie (Florida, U. S. A.) TYPICAL MATERIAL: holotype F! and 1 paratype F! HOSTS: unknown DISTRIBUTION: Nearctic region:

U. S.

A.:

in WA.

Jacksonville (Fla.),

WA!

4. Dryinus alatus (Cresson)

Gonatopus alatus Cresson 1872: 193. Dryinus alatus (Cresson): Patton 1879: 65. Dryinus alatus (Cresson): Ashmead 1893: 86. Hesperodryinus amphiscepae R.C.L. Perkins 1907: 41 (n. syn.); locus typicus Nogales (Arizona, U. S. A.). Hesperodryinus arizonicus R.C.L. Perkins 1907: 42 (n. syn.); locus typicus Nogales (Arizona, U. S. A.). Hesperodryinus audax R. C. L. Perk ins 1907: 43 (n. syn.); locus typicus Nogales (Arizona, U. S. A.). Prodryinus alatus (Cresson): Kieffer 1907b: 9. Mesodryinus alatus (Cresson): Kieffer 1914b: 32. Hesperodryinus amphiscepae R. C. L. Perk i ns: Kieffer 1914b: 39. Hesperodryinus arizonicus R.C.L. Perkins: Kieffer 1914b: 40. Hesperodryinus audax R. C. L. Perk ins: Kieffer 1914b: 41. Mesodryinus alatus (Cresson): Muesebeck and Walkley 1951: 1034. Hesperodryinus amphiscepae R.C.L. Perkins: Muesebeck and Walkley 1951:

1034.

Hesperodryinus arizonicus R.C.L. Perkins: Muesebeck and Walkley 1951:

1035.

Hesperodryinus audax R.C.L. Perkins: Muesebeck and Walkley 1951: 1035.

Hesperodryinus amphiscepae R.C.L. Perkins: Krombein 1979: 1242. Hesperodryinus arizonicus R. C. L. Perk i ns: Krombei n 1979: 1243. Hesperodryinus audax R. C. L. Perk i ns: Krombei n 1979: 1243. Mesodryinus alatus (Cresson): Krombein 1979: 1243.

Dryininae

Fig.

559 -

Chela of

829

Dryinus alatus {Cresson} from Nogales {Arizona}

FEMALE: fully winged; length 5,00-7,87 mm; yellow-testaceous, with petiole black; sometimes with black spots on head, scutum, scutellum, propodeum and abdomen; sometimes in addition frons almost fully black, propectus black, mesothorax, meta thorax and propodeum almost fully black; head {Fig. 557 B} dull, granulated; occipital carina incomplete, only visible behind the ocell i and on sides of vertex; pronotum as sculptured as head,

830

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

A

c Fig.

Varieties of enlarged claws of Dryinus alatus

560 -

from Nogales

(Arizona);

B:

from Ann Arbor (Michigan); Co. (Michigan)

with a strong posterior transversal transversal

impression;

posterior collar and a

impression and a

the pronotum so has a central less visible anterior collar;

as head or sculptured by more or less numerous notaulices complete,

posteriorly separated;

as sculptured as scutum;

prcpodeum shiny,

with a series of transversal anterior surface smooth, median

series

from

parallel

(Cresson):

C:

weak anterior humped disc,

a

scutum as sculptured

longitudinal

keels,

shiny;

scutellum and metanotum shiny, more or less reticulate rugose,

weak keels on posterior surface;

with more or less numerous areolae forming a

anterior margin

A:

from Midland

to posterior surface;

occasionally

831

Dryininae

anterior surface with strong keels; bands,

with distal

part of radial

trochanters more than twi ce as

fore wings with 2 brown vein

transversal

longer than proximal

part;

fore

(15: 6); segment 4 of front

long as broad

tarsus nearly twice as long as segment 1 or sl ightly shorter than segment 1 (33:41); segment 3 of front tarsus produced into a hook; enlarged claw (Fig. 559) with subapical tooth (big or small) and 5-11 lamellae or pegI ike hairs,

without series of teeth;

subapical

tooth more or less far from

560); segment 5 of front tarsus (Fig. 559) with 2 rows of 18-30 lamellae; apex with a group of nearly 20 lamellae; tibial spurs 1,1,2. MALE: fully winged; length 2,85-4,00 mm; head black; with mandibles the apex

reddish;

(Fig.

antennae brown;

legs brown, i ncomp lete;

with tarsi

thorax and propodeum black;

light;

head dull,

abdomen brown;

reticulate rugose;

OPL shorter than POL and than OL;

scutum,

occipital

carina

scute II um and

metanotum dull, as reticulate rugose as head; notaulices almost invisible; propodeum reticulate rugose, with areolae broader than areolae of head; fore wings without dark transversal longer than proximal part; tibial LOCUS TYPICUS: Texas (U. S. A.) TYPICAL MATERIAL:

alatus in WA;

bands,

with distal part of radial

alatus in AE; 1 para type F! of G.

holotype F of G.

and 2 paralectotypes

lectotype F!

vein

spurs 1,1,2.

(1

F,

1 M)!

(designated

amphiscepae in B; 1 paratype F! of H. amphiscepae in BM; lectotype F! (designated by Olmi 1982) of H. arizonicus in B; holotype F! of H. audax in B. HOSTS: in U.S.A. Acanalonia (=Amphiscepa) bivittata (Say) (Acanaloniidae) and Mistharnophantia sonorana Kirkaldy (Flatidae) (R.C.L. Perkins 1907). by Olmi

1982) of H.

DISTRIBUTION: A.: Co.,

Nogales Tex.),

Midland Co. Wattacoo

Nearctic region:

(Ariz.), TE!

OLl

B!

Waukesha Co.

(Mich.),

(Pickens Co.,

WA! S.

CANADA:

BM!

Texas,

(Wis.),

SI.

Catharines

AE!

WA!

WA!

Ann Arbor

Columbia

(Mo.),

C.),

Cleveland

TW!

OL!

WA! (S.

(ant.),

aT!

College Station (Mich.),

Forsyth

C.),

TW!

U.

S.

(Brazos

TW!

(Ga.),

OL! OT!

Moorestown

OLl (N.

J.), OL! TW! Horicon Lake (Lakehurst, Ocean Co., N. J.), AM! Westfield (N. J.), OL! Kalbfleisch Field Station (Huntington, N. Y.), AM! Shokan (N. Y.),

TW!

NEOTROP I C SPEC I ES OF

Nineteen species,

including a fossil

DRYINUS

species,

are known.

KEY TO THE SPECIES (excluding

the fossi I species Dryinus palaeodominicanus Currado and Olmi)

FEMALES

832

2

Mem.

Occipital

carina

Occipital

carina

Amer.

Ent.

no.

37,

invisible ••••••••••••

1.

complete

or

Thorax and propodeum fu I I y Head with POL

longer

constans n. sp.

fu I I y

t estaceous •••••••••• 3

black ••••••••.•••••••••••• 4

or fully

than OL;

1984

incomplete ••••••••••.•••••••••••• 2

or a I most

Thorax and propodeum mostly

3

Inst.,

occipital

carina

complete •••••••••

••••••••••••. •••. •••. . ••••••••. •. . . . " 2. citricolus n. sp. Head with POL almost 0,5 as long as OL; occipital carina incomplete •.•.•.•••••••••••••••••••••• 3. flavoniger n. sp. 4

Scutum fully

sculptured by more

or

less numerous

longitudinal

keels •••••••••••••••••••••••••••••••••••••••••••••••••••••••••• 5 Scutum without keels, fully reticulate rugose or partly g ranu I at ed •••••••••••••••••••••••••••••••••••••••••••••••••••• 16

5

Pronotum fully

granulated

keels on posterior

,wi thout

transerversal

keel s,

except

impression;

for

some

occipital

4. wellingensis n.

comp let e ••.•••••••••••••••••••••

short

carina

sp.

Pronotum differently sculptured, with numerous irregular keels; occipital carina incomplete •••••.••.••••••••••••••••••••••••••• 6

6

Posterior ocelli more distant from occipital carina (Fig. 566 A); head with OPL nearly as long as POL (Fig. 566 A) . . . . . . . . . . ..

. . . . . . . . . . . . . . . . . . . . . •. . •. . . . . . . . " Posteri or ocell i OPL

shorter

nearer occipi tal

than POL

(Fig.

Segment Segment

8

Prothorax mostly b l a c k . . . . . . . . . . . . .

6.

Prothorax

7.

front front

tarsus tarsus

(Fig.

566 B);

sp.

head wi th

566 B) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7

7

1 of of

surinamensis n.

5.

carina

shorter than segment 4 ••••••••••••••• 8 slightly or much longer than segment

4 ••••••••••••••••••.•••.••••••••••••••••••••••••••••••••••••••• 9

9

Antennae broad;

fully less

slender,

prothorax

Antennae more long

fully

slender,

as broad;

10 Dorsal

or mostly

wi th

segment

wi th

segment

fully

striatus (Fenton) argentinus n. sp.

4 nearly

testaceous..

prothorax

surface of

reddish..

4 nearly

or partly

twice as

long

as

caraibicus n. sp.

8.

five-nine

times as

reddish . . . . . . . . . . . . . . 10

propodeum sculptured by numerous

longitudinal

keel s ••••••••••••••••••••••••••••••••••••••••••••••••••••••••• 11 11

Dorsal surface of propodeum fully reticulate rugose . . . . . . . . . . . 13 Head with OL nearly two-three times as long as POL; head fully redd ish. • . • • • • • • • • • • • • • • • • • • • • • • • • • • 9. napensis n. sp. Head with OL as long as or slightly longer than POL; head partly black and

12 Pronotum

testaceous •••••••••••••••••••••••••••••••••••••••••• 12 (laterally

viewed)

with disc nearly

as anterior collar . . . . . . . . . . . . . . . . . . . Pronotum

(laterally

viewed)

with disc

anterior collar . . . . . . . . . . . . . . . . . . . . . . 13 Fore

tibiae nearly

Fore 14 Head

tibiae less than twice as fully reddish-testaceous;

by

longitudinal

twice as

keels;

long as

less 10.

fore

three

times as

long

kimseyae n. sp.

18.

than

twice as

long as

opacifrons n. sp. trochanters . . . . . . . . . . 14

long as fore trochanters ••••••• 15 scutum more strongly sculptured

notaulices weakly

• • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • ••

1 1.

visible among keels

grandis (Og I ob lin)

Dryininae

Head partly black and keels;

833

testaceous;

notaulices clearly

distinct

scutum wi th weak longitudinal among kee Is ••••••••••

alvarenganus n • sp. sinopensis n. sp. 14. belizensis n. sp.

...................................

12.

15 Head part Iy black •••••••.••••••••••• Head

fully

13.

reddish-testaceous . . . . . .

16 Scutum granulated, wi th areolae on lateral posterior half of median region ••••.

regions and

on

15. antilleanus (Evans) Scutum fully

17 Antennal

reticulate

4

segment

less

rugose •••••.•••••••••.••••..•••••••••• 17 slender,

near I Y

twi ce as

8. Antennal

long as broad ••

caraibicus n.

sp.

4 more slender, nearly five times as

segment

long as

broad ••••••••••••••••••••••••••••••••••••••••••••••••••••••••• 18

18 Posterior ocelli

more distant

from occipital

• • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • •• Posteri or ocell i

nearer

occipi tal

16.

carina

carina

(Fig.

580 A)

teutoniae n. sp. 580 B)

(Fig.

17. catarinae n. sp.

MALES

Frons wi thout

2

median keel . . . . . . . . . . . .

Frons with a

complete or

Frons with a

complete median keel;

5.

surinamensis n. sp.

incomplete median keel . . . . . . . . . . . . . . . . 2 notaul ices

reaching

approximately 0,5 length of scutum; scutum granulated, not reticulate rugose . . . . . . . . . . . . . . . . . . . 15. antilleanus (Evans) Frons wi th an half

of

incomplete median keel

frons:

notaulices complete.

scutum reticulate

onlv visible on posterior posteriorlv

ruCJose . . . . . . . . . . . . .

6.

separated:

striatus (Fenton)

The males of other species are unknown.

D. palaeodominicanus n. sp., fo=sil species from Dominica, isn't present

in

through

the amber,

the key because the only known so that

its morphology

specimen

is weakly

visible

isn't very distinct.

1. Dryinus constans n. sp.

FEMALE:

fully

winged;

broad median frontal 9-10 testaceous;

head testaceous,

antennae brown,

thorax and propodeum black,

pronotum reddish; testaceous,

length 6,87 mm;

stripe black;

abdomen black,

with posterior margin of

with some regions testaceous;

with hind coxae partly black;

antenna I segments

with vertex and a

with segments 1-2 and

in following proportions:

legs

antennae distally thickened; 16: 6:

44:

24:

20:

20:

14:

14:

834

Mem.

Fig.

561

-

Amer.

En!.

Ins!.,

no.

37,

1984

Chela of Dryinus cons tans n.

sp.

(type)

12: 15; head dull, granulated and with numerous weak irregular striae; frons with a complete median keel; occipital carina invisible; POL = 4; OL = 2,5; OOL = 8,5; pronotum with a strong anterior transversal impression and with a strong posterior transversal

impression;

disc humped;

posterior

835

Dryininae

A

Fig.

562 -

and

collar

Dryinus striatus (Fenton) (type) (A) and citricolus n. sp. from Manati (Puerto Rico) (8)

Pronotum of females of

invisible;

pronotal

by numerous parallel separated;

tubercles not reaching

longitudinal

keels;

scutellum and metanotum dull,

tegulae;

scutum sculptured

notaul ices complete, reticulate rugose,

posteriorly with 2

incomplete longitudinal keels on posterior surface; fore wings with two broad dark transversa I bands, one on the basa I cell s and another beneath the pterostigma; distal part of radial vein longer than proximal part (21:15); fore tarsal segments in following proportions: 21: 4,5: 11: 20:

36;

proportions of fore tibiae and trochanters: 37:23; enlarged claw (Fig. 561) with a subapical tooth and 20 lamellae; segment 5 of front tarsus (Fig.

561) with two rows of nearly 24 lamellae; apex with a group of nearly 12 lamellae; tibial spurs 1,1,2. MALE:

unknown

LOCUS TYP I CUS:

Represa do Rio Grande

TYPICAL MATERIAL: HOSTS:

holotype F!

(Braz i I)

in TW.

unknown

DISTRIBUTION: Neotropic region: (Guanabara), TW!

BRAZIL:

Represa do Rio Grande

2. Dryinus citricolus n. sp.

FEMALE: fully winged; length 3,12-3,50 mm; fully testaceous, with petiole black; head dull, granulated; occipital carina complete; OPL ; OL; OL shorter than

POL;

pronotum

(Fig.

562 B)

dull,

granulated,

not humped,

836

Mem.

Fig.

563 -

Amer.

Ent.

Inst.,

no.

37,

Chela of Dryinus citricolus n.

with 2 weak anterior and posterior transversal posterior collar weakly visible; scutum shiny,

granulated;

I ength of scutum; shiny,

1984

pronotal

sp.

(type)

impressions;

anterior and

tubercles not reaching

tegulae;

notaulices incomplete,reaching approximately 0,5

scutell um and metanotum du II,

not reticulate rugose or weakly

granu I a ted;

reticulate rugose;

propodeum

areolae more

visibi Ie on anterior surface; posterior surface smooth, without sculpture and without longitudinal keels; fore wings with 2 brown transversal bands and with distal front hook;

part of radial

enlarged claw

segment 5 of front

(Fig.

tarsus

group of 9 lamellae; MALE:

vein

longer than proximal

tarsus nearly twice as long as segment 4;

tibial

563) (Fig.

with subapical 563)

part;

segment 1 of

segment 3 produced tooth and without

with 1 row of 10 lamellae;

into a

lamellae; apex with a

spurs 1,1,2.

unknown

LOCUS TYPICUS: Lajas (Puerto Rico) TYPICAL MATERIAL: holotype F! and 1 paratype F! 1 paratype F!

in WA;

in OL.

HOSTS: unknown DISTRIBUTION: Neotropic region:

PUERTO RICO:

Lajas,

WA!

Manati,

OL!

Dryininae

Fig.

564 -

Chela of

Dryinus flavoniger n. sp. (type)

837

838

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

3. Dryinus flavoniger n. sp.

FEMALE: fully winged; length 4,56-5,00 mm; testaceous, with a central black spot on propodeum; petiole black; internal side of stalks of hind femora black; antennae distally thickened; antennal segments in following proportions: 8: 6: 28: 15: 12: 8: 6,5: 5,5: 5: 7; head flat, dull, granulated; incomplete,

frons without median and visible behind the ocelli

occipital carina OPl = 3;

keels;

invisible behind the temples;

pronotum dull,

POL

as sculptured as head,

anterior and posterior transversal pronotal

lateral

and on sides,

tubercles not reaching

impressions;

tegulae;

occipital

carina

but not reaching eyes;

=

3;

=

Ol

5,5;

weakly humped, posterior collar

scutum,

=

OOl with

7;

weak

invisible;

scutellum and metanotum

dull, as sculptured as head; notaulices incomplete, weakly visible, reaching 0,65 length of scutum; propodeum reticulate rugose, with two longitudinal

median keels joining metanotum

to petiole;

fore wings with

two

dark transversa I bands, one on the basa I cell s and another beneath the pterostigma; distal part of radial vein longer than proximal part (27:8); fore tarsal

segments in following proportions:

2 and 3 of front tarsus produced into hooks; a subapical 564)

tooth and with 10 lamellae;

with 1 row of 15 lamellae;

21:

4:

5,5:

18: 27;

enlarged claw

(Fig.

segment 5 of front tarsus

apex with a

group of nearly

segments 564)

14 lamellae;

tibial spurs 1,1,2. MALE: unknown lOCUS TYPICUS: Faz. Taperinha (Para, TYPICAL MATERIAL: holotype F! in OT; HOSTS: unknown DISTRIBUTION: Neotropic region: Encruzilhada

(Bahia),

Fig.

565 -

Brazil) 1 paratype F!

BRAZil:

Faz.

in OL.

Taperinha

(Para),

OL!

Chela of

with

(Fig.

Dryinus wellingensis n. sp. (type)

OT!

839

Dryininae

4. Dryinus wellingensis n. sp.

FEMALE:

fully

testaceous; black,

length 5,62 mm;

winged;

with tarsi

ligth;

following proportions:

black;

mandibles and clypeus

with segments 1-3 and 10 testaceous;

antennae brown,

antennae distally thickened;

antennal

legs

segments in

11: 6: 22: 11: 11: 9: 8: 7,5: 6,5: 9; antenna I (11 :3); head dull, flat,

segment 4 nearly four times as long as broad granulated,

with numerous areolae on frons and vertex around eyes;

OPL

=

=

1,5; TL

occipital

on posterior transversal

carina complete;

impression;

transversa I impression weak;

= 4;

POL

pronotum weakly

= 4;

frons

= 11; 3; pronotum dull, granulated, with numerous short keels

with a median keel;

OL

humped,

OOL

with anterior

posterior transversa I impression stronger;

posterior collar almost invisible;

pronotal

tubercles reaching tegulae;

scutum dull, granulated, with numerous longitudinal parallel keels; notaulices weakly visible among keels; scutellum and metanotum dull, granulated;

propodeum with dorsal surface sculptured by

longitudinal

parallel keels; posterior surface reticulate rugose and without longitudinal keels; propodeum without transversal keels; fore wings with two dark transversal

bands,

one on

the basal

cells and another beneath

the

(16:7); 19: 2: 5: 13: 19; segments 2 and 3 of front tarsus produced into hooks; enlarged claw (Fig. 565) with a subapical tooth and with 10 lamellae; segment 5 of front tarsus (Fig. 565) with 2 rows of nearly 36 lamellae extending continuously to the apex; tibial spurs 1,1,2. pterostigma;

fore tarsal

MALE:

distal

vein

longer than proximal

part

segments in following proportions:

unknown

LOCUS TYPICUS:

Middlesex

TYPICAL MATERIAL: HOSTS:

(British Honduras)

holotype F!

in OT.

unknown

DISTRIBUTION: NOTE:

part of radial

Neotropic region:

the species is named

in

BRITISH HONDURAS: honor of Mr.

E.C.

Middlesex,

OT!

Welling.

5. Dryin us surinamensis n. sp.

FEMALE: vertex

fully

reddish,

with

testaceous, testaceous;

length 6,87-10,12 mm;

head black,

clypeus and mandibles yellow;

irregular black spots on vertex;

sometimes head

occiput yellow;

meta thorax ,

propodeum and abdomen black;

light or testaceous with brown spots on coxae;

shiny or dull,

granulated,

with ocellar region swollen; the ocelli

with front part of antennae

with segment 3 brown; propectus black, with dorsal region pronotum black, with margins and posterior collar reddish;

mesothorax, tarsi

winged;

(more along orbits),

with numerous occipital

and on sides of vertex;

longitudinal

carina

legs brown

head

wi th

(Fig. 566 A)

keels;

vertex flat,

incomplete, only visible behind

pronotum shiny,

with numerous keels

840

Fig.

Mem. Amer. Ent.

566 -

Ins!., no. 37,

1984

Heads of females of Dryinus surinamensis n. striatus (Fenton) (type) (B).

sp.

(type)(A)

and

Dryininae

Fig. 567 -

around disc;

keels;

impression and anterior collar weak;

impression strong;

tubercles not reaching

longitudinal of keels;

Dryinus surinamensis n. sp. (type)

anterior transversal

posterior transversal pronotal

Chela of

841

tegulae;

notaulices,complete,

posterior collar short, scutum shiny, but almost

scutellum and metanotum punctate;

distinct;

with numerous

invisible among furrows

propodeum reticulate rugose,

842

Mem.

Fig.

568 -

Amer.

Ent.

no.

37,

1984

Male genitalia of Dryinus surinamensis n. (Brazi I):

with 2 short brown

Ins!.,

longitudinal

transversal

than proximal

keels on posterior surface;

bands and with distal

part

proportions: 70:35;

(25:18);

with subapical

567)

with 2 rows of 35 lamellae;

Itapetinca

vein much longer

trochanters in following

tarsus nearly as long as segment 4;

tarsus produced

567)

from

fore wings with 2

part of radial

fore tibiae and

segment 1 of front

segments 2 and 3 of front

sp.

right half removed.

tooth and 26 lamellae;

into hooks;

enlarged claw

segment 5 of front

tarsus

apex with a group of 20 lamellae;

(Fig. (Fig. tibial

spurs 1,1,2. MALE: fully winged; length 3,50-4,25 mm; black; clypeus and mandibles testaceous; antennae brown, wi th segments 1-2 testaceous; legs testaceous,

Dryininae

Fig.

with hind coxae, thickened; 10: 9:

9:

569 -

Dryinus striatus (Fenton)(type)

hind femora and hind tibiae brown;

antennal 10;

Chela of

843

antennae not distally

segments in following proportions: 7:

head dull,

fully reticulate rugose;

frontal

7:

20:

11:

line absent;

12:

12:

844

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

/ ./ 1 - - - - - 1 / ___ I

/

/

/

Fig.

570 -

Male genitalia of Dryinus striatus (Brazil):

(Fenton)

left half removed.

from Encruzilhada

Dryininae

occipital

carina

of head,

on

incomplete,

not visible on

sides and on ventral

side;

head directed towards the eyes and 4;

OOL = 5,5;

dull,

OPL = 3;

complete,

temples,

occipital

laterally

temples invisible;

reticulate rugose;

apparently

845

visible on dorsal

carina on dorsal

touching eyes;

scutum,

side

side of

POL = 7;

OL =

scutellum and metanotum

notaulices almost invisible near posterior margin,

incomplete, but reaching

posteriorly separated;

almost posterior margin of scutum;

propodeum;

transversal or longitudinal keels; pterostigma; distal part of radial

dull,

reticulate rugose,

if

without

fore wings weakly darkened beneath the vein shorter than proximal part (15:18);

genitalia: fig. 568; tibial spurs 1,1,2. LOCUS TYPICUS: Charlesburg (Suriname) TYPICAL MATERIAL: (1

F,

2 MM)!

HOSTS:

holotype F!

in TW;

in LE;

3 paratypes

(1

1 paratype F!

M,

Neotropic region:

Raleigh Falls

(Coppename River),

ECUADOR:

Ongota,

TW!

BRAZIL:

SURINAME: OL!

Lestodryinus striatus

Itapetinca

FEMALE:

fully

winged;

(more along orbits),

with a

median frontal specimen);

mesothorax,

brown-reddish;

legs testaceous;

incomplete,

13;

strong

a

collar short, long

head

(Fig.

with 2

transversal

long as proximal

proportions: fore

25:

5:

part 12:

37:

carina;

56;

an

tegulae,

anterior

weakly

fore tarsal

visible among

keels;

part of radial

segments

twice

fully propodeum

keels on posterior surface; longitudinal

posterior

nearly

metanotum granu la ted;

dorsal

fore wings vein nearly

in following

segments 2 and 3 of fore tarsi

tibiae and trochanters

pronotum pronotum

scutum shiny,

notaul ices complete,

bands and with distal (19:20);

carina POL =

anterior collar visible;

tubercles not reaching

longitudinal

abdomen

occipital

transversa I impress ion and by

keels;

(only

granulated and

sides without sculpture;

disc humped;

pronotal

longitudinal

holotype

and on sides of vertex;

surface reticulate rugose or sculptured by with 2 brown

dull,

keels on frons;

scutell um as scu I ptured as scutum;

reticulate rugose,

hooks;

566 B)

as distance between tegulae and tubercles;

sculptured by

sometimes face in

with sides and posterior

posterior ocell i touching occipital

impression;

distinct;

with front part of

antennae missing

longitudinal

posterior strong

trasversal

comb.

head black,

pronotum black,

wih numerous keels around disc;

crossed by

OL!

meta thorax and propodeum black;

with some

COL =

n.

LE!

WA!

2.

only visible behind the ocelli

1,5; OL = 3; shiny,

1927:

stripe testaceous;

propectus black;

reticulate rugose,

(Fenton)

TW!

(Loreto),

clypeus and mandibles yellow;

collar redd i sh;

as

3 paratypes

(Paramaribo),

San Antonio

(Bahia),

length 6,0-7,5 mm;

vertex known

Fenton

Charlesburg

PERU:

6. Dryinus striatus

keel s;

in WA;

in OL.

unknown

DISTRIBUTION:

as

2 FF)!

in following proportions:

produced

44:28;

into

enlarged

846

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

claw (Fig. 569) with subapical tooth and 11 lamellae; segment 5 of front tarsus (Fig. 569) with 2 rows of 27 lamellae; apex with a group of 28 lamellae; MALE:

tibial

fully

spurs 1,1,2.

winged;

length 4,44 mm;

and hind coxae black;

black;

hind femora brown;

antenna I segments in following proportions:

mandibles testaceous,

with mid

antennae not distally thickened; 9:

8:

23:

12:

12:

12:

12:

10: 9:

10; head dull, reticulate rugose; frontal I ine incomplete, only visible on posterior half of frons; occipital carina incomplete, invisible on temples; POL = 7; Ol = 4,5; OOl = 4,5; OPl = 1; scutum, scutellum and metanotum reticulate rugose, dull; notaulices complete, posteriorly separated, weakly visible near posterior margin of scutum; a transversal

strong keel

surface reticulate rugose; of radial tibial

vein

nearly as

and posterior surface;

fore wings with distal long as proximal

part

with

posterior

half darkened;

distal part

(17: 17); genital ia:

fig.

570;

38'

W 40

spurs 1,1,2.

lOCUS TYPICUS:

Santarem

TYPICAL MATERIAL: HOSTS: unknown DISTRIBUTION: 33'

propodeum reticulate rugose,

between dorsal

(Brazil)

holotype F!

in WA.

Neotropic region:

S (Amazonas),

TW!

BRAZil:

Encruzi Ihada

Santarem

(Bahia),

(Para),

TW!

WA!

71

0

Ol!

7. Dryinus argentinus n. sp.

FEMALE:

fully

winged;

length 4,37-5,62 mm;

head

testaceous,

with vertex

and posterior ha If of frons black; antennae brown-testaceous; prothorax reddish, sometimes with two black spots on sides; mesothorax, metathorax and propodeum black; brown;

abdomen brown;

antennae distally thickened;

proportions:

10: 7:

32:

15:

14:

with numerous longitudinal occipital

10: 9:

striae;

legs testaceous,

antennal 9: 7:

wi th coxae part I y

segments in following 10;

head flat,

granulated and

frons with a median complete keel;

carina

incomplete,

visible behind the ocelli

posterior ocelli;

POL = 2,5;

Ol = 2,5;

and on sides of

OOl = 10; Tl = 2;

pronotum shiny,

with two strong transversal impressions; anterior collar distinct and long; posterior collar invisible; pronotum with numerous longitudinal keels around disc; pronotal tubercles not reaching tegulae; scutum dull, sculptured by numerous parallel longitudinal keels; notaul ices invisible among longitudinal

keels;

scutellum sculptured by numerous parallel

longitudinal

keels; metanotum dull, granulated; propodeum with dorsal surface sculptured by numerous parallel longitudinal keels; posterior surface reticulate rugose and with two incomplete longitudinal without transversal

keels between dorsal

with two dark transversal beneath the pterostigma; proximal

part

fore tarsal

(14:16);

bands, one on distal

keels;

propodeum

and posterior surface; the basal

part of radial

vein

fore wings

cells and another little shorter than

proportions of fore tibiae and trochanters:

segments in following

proportions:

18: 4:

9:

23:

38;

30:22;

enlarged

Dryininae

Fig. 571

claw

(Fig.

front

tarsus

571)

continuously MALE:

-

Chela of

Dryinus argentinus n. sp.

with a subapical

(Fig.

571)

tibial

(type)

tooth and with 7 lamellae;

with 2 rows of nearly 45

to the apex;

847

segment 5 of

lamellae extending

spurs 1,1,2.

unknown

LOCUS TYP ICUS:

Horco Molle (Argentina)

TYPICAL MATERIAL:

holotype F!

and 5 paratypes FF!

in TW;

3 paratypes

848

Mem. Amer.

Fig.

FF!

in AM;

572 -

Ent.

Inst.,

no.

37,

1984

Chela of Dryinus caraibicus n.

5 paratypes FF!

sp.

(type)

in Ol.

HOSTS: unknown DISTRIBUTION: Neotropic region: ARGENTINA: Horco Molle (Tucuman), TW! BRAZil: Caruaru, OL! Silva Jardin (Rio de Janeiro) TW! Sinop (Mato Grosso),

TW!

Represa do Rio Grande (Guanabara),

Ol!

BOLIVIA:

Rio

Ichilo

Dryininae

Fig.

573 -

Chela of Dryinus napensis n.

849

sp.

(type)

(Santa Cruz), AM! 2 Km N mouth of Rio Chapare (Rio Mamore, Cruz), AM! OLl ECUADOR: Napo and Coca Rivers, TW!

Dept.

Santa

8. Dryinus caraibicus n. sp.

FEMALE:

fully winged; length 4,56-4,62 mm; head testaceous, with region of vertex near occipital carina dark; antennae testaceous-dark; prothorax testaceous; mesothorax, meta thorax and propodeum black; abdomen browntestaceous, with fore and mid coxae almost fully black, hind coxae apically black, clubs of femora brown; antennae distally thickened; antennal segments in following proportions: 14: 7: 22: 7: 7: 8: 7: 6: 6: 9; antenna I segment 4 nearly twice as long as broad; head dull, very weakly reticulate rugose or sculptured by irregular keels; frons with a median keel; occipital carina incomplete, only visible behind the ocelli and on sides of ocelli; occipital carina not visible on temples; POL = 3; OL = 2,5; OOL = 9; OPL 2; TL = 0; pronotum dull, broader than long (22:17); pronotum as sculptured as head, with some keels around disc, with an anterior strong transversa I impress ion; disc very humped; posterior transversa I impression strong; posterior collar invisible; pronotal tubercles reaching tegulae; scutum dull, sculptured by numerous paC'allel longitudinal keels; sometimes (in Bolivian specimen) keels weakly visible and sculpture partly reticulate rugose; notaul ices complete, posteriorly separated; scutum punctate, dull, with some irregular striae near anterior margin; metanotum without sculpture, smooth; propodeum reticulate rugose, dull; fore wings with two dark transversa I bands, one on the basa I cell s and another beneath the pterostigma; distal part of radial vein longer than proximal part (17:11); fore tibiae and trochanters in following proportions: 21 :10; fore tarsal segments in following proportions: 18: 3: 6: 14: 24; enlarged claw (Fig. 572) with a subapical tooth and with 6 lamellae; segment 5 of front tarsus

850

Mem.

(Fig.

572)

lamellae;

with

Amer.

Ent.

Inst.,

1 row of 19 lamellae;

segments 2 and 3 of front

no.

37,

1984

apex with a

group of nearly 10

tarsus produced

into hooks;

tibial

spurs 1,1,2. MALE:

unknown

LOCUS TYPICUS:

Simia

TYPICAL MATERIAL:

(Trinidad)

holotype F!

1 para type F!

in OT;

1 para type F!

in TW;

in AM. HOSTS:

unknown

DISTRIBUTION: 31'

S 55°

37'

Neotropic region: W (Sinop,

(Rio Mamore,

Dept.

TRINIDAD:

Mato Grosso),

Seni),

TW!

Simia

(Arima),

SOL IVIA:

OTt

BRAZIL:

12°

10 Km E San Antonio

AM!

9. Dryinus napensis n. sp.

FEMALE:

fully

winged;

margins of clypeus testaceous; partly

thorax,

testaceous;

propodeum,

abdomen

tarsus nearly seven

in following proportions:

excavated"

dull,

sculptured by

visible among ocell i;

frons

visible behind the ocelli OOL =

13;

OPL =

head reddish,

antennae brown, and

1;

11:

numerous

with a median

52:

pronotal

15:

11:

longitudinal keel;

9:

antennal

8:

7:

parallel

occipital

9;

head

keels,

carina

also

incomplete,

POL = 1;

OL = 3;

with a strong anterior transversal

impression and a strong posterior transversal invisible;

trochanters

long as broad;

14:

and on sides of posterior ocelli;

pronotum humped,

fore

antennae distally thickened;

times as 8:

with mandibles and

with segments 1-2

legs black;

hind trochanters testaceous;

segment 4 of front segments

length 6,87 mm;

testaceous;

tubercles not reaching

impression;

tegulae;

posterior collar

pronotum with numerous

keels around disc; scutum and scutellum dull, sculptured by numerous longitudinal parallel keels; notaul ices not clearly visible among keels; metanotum dull, dorsal

weakly sculptured by

surface sculptured by

surface reticulate rugose and with propodeum wi thout

two dark

another beneath

the pterostigma;

proximal

part

(24:12);

fore tarsal

longitudinal

two complete

transversa I keel s between

fore wings with

37:18;

irregular keels;

numerous

transversal

bands,

distal

fore tibiae and

segments

segments 2 and 3 of front

keels;

longitudinal

keels;

posterior

dorsa I and posterior surface; one on

the basal

part of radial trochanters

vein

cells and

longer than

in following proportions:

in following proportions: tarsus produced

propodeum with

parallel

25:

into hooks;

5:

11:

23:

42;

enlarged claw

(Fig.

573) with 2 subapical teeth and with 22 lamellae; segment 5 of front tarsus (Fig. 573) with 2 rows of nearly 50 lamellae extending continuously to the apex;

tibial

spurs 1,1,2.

MALE: unknown LOCUS TYPICUS:

Limoncocha

TYPICAL MATERIAL: HOSTS:

(Ecuador)

holotype F!

in OT.

unknown

DISTRIBUTION:

Neotropic region:

ECUADOR:

Limoncocha

(Napo),

OTt

Dryininae

Fig.

574 -

Chela of

851

Dryinus opaciirons n. sp. (type)

10. Dryinu5 opaciirons n. sp.

FEMALE:

fully

space brown frons;

winged;

and a

length 5,00-5,62 mm;

antennae testaceous,

pronotum

head testaceous,

with malar

broad black spot on vertex and on posterior half of

testaceous,

with

somet imes darkened;

two lateral

propectus testaceous;

black or brown spots;

pronotal

margins testaceous; mesothorax, metathorax and propodeum black; abdomen brown-testaceous; legs testaceous; antennae distally thickened; antenna I segments

in following proportions:

12:

5:

34:

15:

12:

12:

10:

9:

11;

8:

head

dull, flat, granulated, with some weak longitudinal keels on frons; frontal line complete; occipital carina incomplete, invisible on sides of head, only visible on dorsal

side of head,

where the occipital

carina

is touching

posterior edges of eyes; POL = 2; OL = 3; OOL = 10; OPL = 1; temples invisible; pronotum shiny, with two strong transversal impressions; anterior collar

long and distinct;

pronotum collar;

posterior collar invisible;

with numerous keels around disc,

pronotal

tubercles not reaching

disc very humped;

less than

tegulae;

twice as

scutum,

long

as

scutellum and

metanotum dull, sculptured by numerous parallel longitudinal keels; notaul ices weakly visible among keels; propodeum with dorsal surface sculptured by

numerous parallel

surface reticulate rugose; longitudinal basal

keels and with posterior

fore-wing

with

two dark

nearly as with

tarsus

long as proximal

part

in following proportions: two subapical

(Fig.

574)

(16:16); 26:

4:

teeth and with 8

spurs 1,1,2.

distal

radial

cell

9:

38;

24:

lamellae;

with 2 rows of nearly 40

to the apex; tibial MALE: unknown

two incomplete

transversal

cells and another beneath the pterostigma;

segments 574)

keels;

longitudinal

posterior surface also with

bands,

one on

part of radial

open;

fore

the vein

tarsal

enlarged claw

(Fig.

segment 5 of front

lamellae extending continuously

852

Mem.

Fig.

lOCUS TYPICUS:

575 -

Amer.

Ent.

no.

37,

1984

Dryinus grandis (Ogloblin)(type)

Chela of

Jacareacanga

TYPICAL MATERIAL:

Ins!.,

(Brazil) and 1 para type F!

holotype F!

in TW;

1 para type F!

in

Ol. HOSTS:

unknown

DISTRIBUTION: Teutonia

Neotropic species:

(Santa Catarina),

TW!

BRAZil:

Jacareacanga

(Para),

TW!

Nova

Ol!

11. Dryinus grandis (Og lob lin) n. comb.

Nesodryinus grandis Ogloblin 1938: 35.

FEMALE:

fully

winged;

length 8,12 mm;

thorax and propodeum black, pronotum reddish; trochanters, tarsi

abdomen brown-reddish;

femora

testaceous,

antennae distally weakly

following proportions:

carina

16: 8: 50: 20:

OPl =

longitudinal

incomplete,

1,5; Tl

transversa I

=

keels;

impression and by distinct;

longer than broad;

thickened;

with coxae,

legs black,

with

antenna I segments in

18: 16: 14: 12: 10: 14; antennal

frons

with a

head dull,

humped,

granulated and

median keel;

POL = 2,5;

temples;

2,5; pronotum shiny,

legs testaceous,

mid and hind

long as broad;

reaching only

posterior collar short, pronotum

fore

and tibiae partly brown;

segment 4 nearly five times as with numerous

head and antennae testaceous;

except f0r posterior margin and sides of

crossed by a

a strong posterior transversa I pronotal

occipital

Ol = 2,5;

anterior

impression;

tubercles not reaching

scutum shiny,

OOl = 14;

strong

tegulae;

strongly sculptured by numerous

longitudinal keels; notaul ices invisible among keels; scutellum and metanotum dull, reticulate rugose,; propodeum reticulate rugose with 2

Dryininae

Fig.

576 -

Chela of Dryinus alvarenganus n.

853

sp.

(type)

854

Mem.

Fig.

incomplete

577 -

longitudinal

transversa I bands, pterostigma;

Amer.

segments

Inst.,

37,

keels on posterior surface;

one on

1984

sp.

(type)

fore-wing

with 2 dark

the basa I cell s and another beneath the

distal part of radial

vein

longer than proximal

in following proportions:

in following proportions: 40:

with a subapical

no.

Chela of Dryinus sinopensis n.

fore tibiae and trochanters 575)

Ent.

4:

60:30;

part

(15:8);

fore tarsal

11: 27: 45; enlarged claw (Fig. segment 5 of front tarsus

tooth and 16 lamellae;

Dryininae

Fig.

(Fig.

575)

578 -

Chela of

Dryinus belizensis

with 2 rows of nearly 40 lamellae;

20 lamellae;

tibial

spurs 1,1,2.

855

n.

sp.

(type)

apex with a group of nearly

856

Mem. Amer.

Fig.

579 -

Che-Ia of

En!.

Ins!.,

no.

37,

1984

Dryinus antillean us (Evans) (type)

MALE: unknown LOCUS TYPICUS: Jujuy (Argentina) TYPICAL MATERIAL: holotype F! in BA. HOSTS: unknown DISTRIBUTION: Neotropic region: ARGENTINA:

Jujuy,

BA!

Dryininae

Fig.

857

Heads of females of Dryinus teutoniae n.

580 -

catarinae n. sp.

sp.

(type) (A)

and

(type) (B).

12. Dryinus alvarenganus n. sp.

FEMALE:

fully

winged;

antennae testaceous;

length 6,25 mm;

propectus black;

head

testaceous,

for sides of pronotum testaceous;

abdomen brown;

distally thickened,

very slender;

antennal

proportions:

60:

12: 5:

irregularly striate;

25:

frons

21:

14:

with vertex black;

thorax and propodeum black,

10: 9:

legs testaceous;

except antennae

segments in following 7:

9;

with a median keel;

head dull,

occipital

granulated and

carina

incomplete,

only visible behind the ocelli and on sides of ocellar triangle; temples invisible; POL = 3; OL = 4; OOL = 10; OPL = 2,5; pronotum with a strong anterior transversal impression; reaching complete, scutum;

tegulae;

posterior collar short;

scutum dull,

posteriorly separated, scutellum punctate,

reticulate rugose;

with numerous

pronotal

tubercles not

longitudinal

with areolae along margins;

fore-wing

cells and another beneath

keels;

notaulices

weakly distinct near posterior margin of

propodeum reticulate rugose,

on posterior surface; basal

impression and with a strong posterior transversal

disc humped;

metanotum

with two

longitudinal

with two dark transversal the pterostigma;

bands,

keels

one on

distal part of radial

longer than proximal part (25:15); fore tarsal segments in following proportions: 32: 5: 10: 29: 45; enlarged claw (Fig. 576) with a subapical tooth and with 25

lamellae;

segment 5 of front

rows of nearly 33

lamellae;

apex with a group of nearly 16 lamellae;

proportions of fore tibiae and trochanters: MALE:

60:28;

(Fig.

tibial

576)

Represa do Rio Grande

TYPICAL MATERIAL:

holotype F!

HOSTS: unknown DISTRIBUTION: Neotropic region: (Guanabara),

(Brazil)

in TW. BRAZIL:

Represa do Rio Grande

TW!

the species is named

in honor of Mr.

M.

with two

spurs 1,1,2.

unknown

LOCUS TYPICUS:

NOTES:

tarsus

Alvarenga.

the

vein

858

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

13. Dryinus sinopensis n. sp.

FEMALE:

fully

spots on si des;

head testaceous,

propectus testaceous;

mesothorax,

legs testaceous; proportions:

length 5 mm;

winged;

antennae testaceous;

pronotum

metathorax and propodeum black;

antennae distally thickened;

12: 7:

33:

with vertex black;

testaceous,

16:

12:

11:

10: 9:

antennal

8:

12,5;

wi th

two

abdomen

and

segments in following

head dull,

granulated,

with numerous longitudinal keels on vertex and frons; frontal I ine complete; occipital carina incomplete, only visible behind the ocell i and on sides of ocellar triangle;

temples

pronotum shiny, transversal invisible;

invisible;

impressions; pronotal

2;

anterior collar

=

OL

long;

tubercles not reaching

sculptured by numerous parallel among keels;

=

POL

3;

with numerous striae around disc,

metanotum dull,

longitudinal

=

disc humped;

tegulae;

granulated;

OOL

=

10; OPL

crossed by

1;

two strong

posterior collar

scutum and scutellum

keels;

notaul ices not distinct

propodeum fully reticulate rugose,

with two longitudinal keels on posterior surface; fore-wing with two dark transversal bands, one on the basal cells and another beneath the pterostigma; (16:15); tarsal

distal

fore

part of radial

vein slightly

segments

in following proportions:

segment 4 more than five times as (Fig.

577)

tarsus

577)

with 2 rows of nearly 35

to the apex; tibial MALE: unknown LOCUS TYPICUS:

3,5:

30:20;

part fore

9: 22: 36; antennal (16:3);

enlarged claw segment 5 of front

lamellae extending continuously

spurs 1,1,2.

12° 31'

TYPICAL MATERIAL: HOSTS:

27:

long as broad

tooth and with 9 lamellae;

with a subapical

(Fig.

longer than proximal

tibiae and trochanters in following proportions:

S 55° 37'

holotype F!

W (Sinop,

Mato Grosso,

Brazil)

in TW.

unknown

DISTRIBUTION: Grosso),

Neotropic region:

BRAZIL:

12° 31'

S,

55° 37'

W (Sinop,

Malo

TW!

14. Dryinus belizensis n. sp.

FEMALE: brown,

fully

winged;

length 5,62 mm;

with segments 1-2 testaceous;

meta thorax and propodeum black;

head reddish-testaceous;

prothorax reddish;

abdomen and

antennae

mesothorax,

legs redd i sh-testaceous;

antennae distally thickened; antennal segments in following proportions: 13: 7: 38: 20: 16: 13: 12: 10: 9: 12; antennal segment 4 nearly six times as long as broad;

head dull,

the sides of ocelli; and on POL

=

flat,

occipital

the sides of ocelli; 2;

OL

=

2,5;

OOl

=

granulated,

carina

occipital 12; OPL

with some

incomplete,

=

carina 1,5;

longitudinal

keels on

visible behind the ocelli

invisible behind the temples;

pronotum humped,

shiny,

with

numerous keels around disc,

with strong anterior and posterior transversal

impressions;

invisible;

posterior collar

pronotal

tubercles not reaching

Dryininae

Fig.

tegulae; clearly

581

-

Chela of

Dryinus teutoniae n. sp. (type)

scutum sculptured by numerous visible among keels;

859

longitudinal

keels;

notaulices not

scutellum sculptured by numerous

longitudinal

keels; metanotum smooth, dull, granulated; propodeum reticulate rugose; posterior surface of propodeum with two incomplete longitudinal keels; forewing with two dark transversal beneath part

the pterostigma;

(19:17);

fore tarsal

fore

subapical

one on the basal vein

in following proportions:

tarsus produced into hooks;

tooth and with

with 2 rows of nearly 51

13 lamellae;

29:

4:

10: 24:

enlarged claw

segment 5 of front

lamellae extending continuously

spurs 1,1,2. MALE: unknown LOCUS TYPICUS:

Middlesex

TYPICAL MATERIAL: HOSTS:

(Belize)

holotype F!

in OT.

unknown

DISTRIBUTION:

cells and another

longer than proximal

tibiae and trochanters in following proportions:

segments

and 3 of front

bands,

distal part of radial

Neotropic region:

BELIZE:

Milddlesex,

OT!

40;

(Fig. tarsus

34:23;

segments 2 578)

with a

(Fig.

to the apex;

578) tibial

860

Mem. Amer. En!.

Ins!., no. 37,1984

Fig. 582 - Chela of Dryinus catarinae n. sp.

15. Dryinus antillean us (Evans) n. comb.

(type)

Dryininae

861

Mesodryinus antilleanus Evans 1969: 11.

FEMALE;

fully

winged;

mandibles testaceous;

length 3,60-4,00 mm; antennae testaceous;

with sides and posterior collar reddish; propodeum black; thickened; (11:1,5);

abdomen brown-reddish;

head dull,

hairy,

pronotum du II,

transversal

POL

granu I a ted,

impression and by

posterior collar short, pronotal

granulated,

carina complete; a

distinct;

tubercles not reaching

=

region;

pronotum black,

metathorax and antennae distally

10: 6:

23:

11:

9:

9;

times as long as broad

with some 4,5;

OL

=

longitudinal

3; OOL

=

keels on

7,5; OPL

=

1;

crossed by a posterior strong

weak anterior transversal

impression;

anterior collar less visible; tegulae;

scutum dull,

numerous areolae on posterior half of lateral margin of central

with clypeus and

legs testaceous;

antenna I segment 4 nearly seven

occipital

TL = 4;

mesothorax,

antenna I segments in following proportions;

8: 6: 5: 7; frons;

head black,

propectus black;

disc humped;

granulated,

with

regions and near posterior

notaul ices incomplete,

distinctly visible on

anterior half of scutellum, weakly visible on posterior half, because hidden among areolae; scutellum and metanotum dull, granulated; propodeum reticulate rugose, surface very

without

inclined;

longitudinal

fore-wing

distal

part of radial

tarsal

segments in following proportions:

front

vein much

keels on posterior surface;

with 2 brown transversal longer than proximal

tarsus produced into a hook;

18: 2:

5:

enlarged claw

part

13: (Fig.

(13:7);

fore

segment 3 of

579) with a

(Fig. 579) with 2 rows of 40 lamellae extending continuously to the apex; fore tibiae and trochanters in following proportions: 31:18; tibial spurs 1,1,2. MALE: fully winged; length 3,00 mm; head black, with mandibles yellow; antennae brown; abdomen black; legs yellow, wi th fore and hi nd coxae partly black; antennal segments in following proportions: 6: 4,5: 12: 9: 10: 10: 8: 8: 7: 7; head dull, granulated; occipital carina incomplete; POL = 5,5; OL = 3; OOL = 5; OPL = 1; TL = 1,5; frontal line complete; scutum dull, granulated; notaulices incomplete, extending to beyond 0,65 length of scutum; scutellum dull, granulated, except for a narrow region near metanotum which is shiny, smooth and without sculpture; metanotum smooth and without sculpture; propodeum reticulate rugose, without longitudinal keel s on posterior surface; fore-w i ng wi thout dark transversa I bands and with distal part of radial vein much longer than proximal part (11 :6); tibial spurs 1,1,2. LOCUS TYP ICUS: Mouth of Layou River (Dominica) TYPICAL MATERIAL: holotype F! and allotype M! in WA; 1 paratype F! in CM. HOSTS: unknown DISTRIBUTION: Neotropic region: DOMINICA: Mouth of Layou River, WA! South Chiltern, CM! subapical

tooth and 13 lamellae;

segment 5 of front

19;

posterior

bands and with

tarsus

862

Mem. Amer.

Fig.

583 -

En!.

Inst.,

no.

37,

1984

Chela of Dryinus kimseyae n.

sp.

(type)

16. Dryinus teutoniae n. sp.

FEMALE fully winged; length 4,50 mm; head black, with mandibles brown; antennae testaceous; thorax and propodeum black; abdomen black-brown; legs testaceous, with coxae partly brown and clubs of hind femora black; head (Fig. 580 A) dull, hairy, convex, reticulate rugose; occipital carina complete; posterior ocell i not touching occipital carina; OPl = Ol; Ol

Dryininae

Fig.

584 -

863

Female of Dryinus palaeodominicanus Currado and Olmi

(type)

shorter than POL; pronotum dull, granulated, crossed by a weak anterior transversa I impression; anterior coil ar weak; posterior transversa I impression strong; posterior collar short, distinct; scutellum fully reticulate rugose; notaulices weakly visible, incomplete, reaching approximately 0,5 length of scutum; scutell um and metanotum ret icu I ate rugose; propodeum reticulate rugose, without longitudinal keels on posterior surface; fore-wing without dark transversal bands and with distal part of radial vein longer than proximal part (15:7); segment 1 of front tarsus longer than segment 4 (20:12); segments 2 and 3 produced into hooks; enlarged claw (Fig. 581) with subapical tooth and 16 lamellae; segment 5 of front tarsus (Fig. 581) with 2 rows of 37 lamellae extending continuously to the apex; tibial spurs 1,1,2. MALE: unknown LOCUS TYPICUS: Nova Teutonia (Brazil) TYPICAL MATERIAL: holotype F! in BM. HOSTS: unknown DISTRIBUTION: Neotropic region:

BRAZIL:

Nova Teutonia

(S.ta Catarina),

864

Mem.

Amer.

En!.

Inst.,

no.

37,

1984

BM!

17. Dryinus catarinae n. sp.

FEMALE:

fully

winged;

length 4,50 mm;

head black,

with clypeus and

mandibles reddish; antennae testaceous; propectus black; pronotum black, with posterior collar reddish; mesothorax, metathorax and propodeum black; abdomen brown; B)

shiny,

carina complete; than OL;

legs testaceous,

granulated,

with coxae partly brown;

with some longitudinal

posterior ocelli

OL shorter than

POL;

head

keels on frons;

not touching occipital pronotum dull,

carina;

granulated;

(Fig.

580

occipital POL shorter

anterior

transversa I impress ion and anterior coil ar weak; posterior transversa I impression strong; posterior collar short, distinct; pronotal tubercles not reaching tegulae; scutum shiny, fully reticulate rugose; notaulices weakly visible, incomplete, reaching approximately 0,5 length of scutum; scutellum and metanotum shiny, granulated; propodeum reticulate rugose, without longitudinal keels on posterior surface; fore-wing with 2 brown transversal bands and with distal part of radial vein longer than proximal part (13:8); segment 1 of front tarsus longer than segment 4 (18:12); segments 2 and 3 of front tarsus produced into hooks; enlarged claw (Fig. 582) with subapical tooth and 16 lamellae; segment 5 of front tarsus (Fig. 582) with 2 rows of 42 lamellae extending continuously to the apex; tibial spurs 1,1,2. MALE: unknown LOCUS TYPICUS: Nova Teutonia (Brazil) TYPICAL MATERIAL: holotype F! in BM. HOSTS: unknown DISTRIBUTlmj: Neotropic region: BRAZIL: Nova Teutonia (S.ta Catarina), BM!

18. Dryinus kimseyae n. sp.

FEMALE: spot on

fully

winged;

ventral

length 10,8 mm;

head black,

with a broad yellow

side near mouth-parts and with anterior region of frons

near clypeus testaceous; clypeus testaceous with a brown spot; mandibles testaceous, with teeth brown; antennae brown-reddish, with segment 1 yellow;

thorax and propodeum black,

testaceous; brown;

abdomen brown;

antennae distally

proportions:

24:

11: 90:

nearly nine times as

weakly 33:

keel;

occipital

carina

30:

thickened; 21:

long as broad

with numerous areolae and

with posterior margin of pronotum

legs testaceous, 16:

15:

(35:4);

irregulae keels;

incomplete;

POL

= 4;

with anterior tibiae partly antennal 12:

17;

segments in following antenna I segment 4

head flat, frons OL

=

dull,

granulated,

with a complete median 4;

OOL

=

17;

OPL

=

2;

Dryininae

Fig.

Dryinus palaeodominicanus Currado and Olmi (type)

585 -

occipital

carina not visible behind the eyes;

transversal collar

(in

strong;

numerous weakly

impression, lateral

with disc nearly

view:

50:14);

posterior collar short,

reaching

longitudinal tegulae;

scutell um dull,

distinct;

keels,

keels; basal

cells

keels;

pronotum shiny,

hairy,

fore-wing

is very

with

posterior tubercles not

longitudinal

keels;

notaul ices

except for the posterior half not metanotum dull,

granulated;

surface sculptured by numerous strong

posterior surface with

transversal

long as anterior impression

but complete and posteriorly separated;

reticulate rugose,

with dorsal

times as

keels on collar and disc;

scutum with numerous

visible among

pronotum crossed by a strong

three

a posterior transversal

reticulate rugose and granulated; shiny,

865

two with

longitudinal

propodeum

longitudinal

keels and with numerous

two dark transversal

and another beneath the pterostigma;

bands,

distal

one on

the

part of radial

vein longer than proximal part (43:22); radial cell open; fore tarsal segments in following proportions: 50: 7: 14: 45: 72; enlarged claw (Fig. 583) front

with a tarsus

subapical (Fig.

583)

tooth and a with

row of nearly 28

two rows of nearly 45

lamellae; lamellae;

segment 5 of apex with a

866

Mem. Amer.

group of nearly 25 MALE:

lamellae;

Ent.

Ins!.,

tibial

no.

37,

1984

spurs 1,1,2.

unknown

LOCUS TYP I CUS:

Barro Co lorado

TYPICAL MATERIAL: HOSTS:

I.

holotype F!

(Panama)

in DA.

unknown

DISTRIBUTION:

Neotropic region:

PANAMA:

Barro Colorado

I.

(Canal

Zone),

DA! NOTE:

the species

is named in honor of Mrs.

Lynn S.

Kimsey.

19. Dryinus palaeodominicanus Currado and Dlmi

Dryinus palaeodominicanus Currado and Olmi 1983: 330.

FEMALE

(Figs 584,

585):

brown-testaceous and weakly

thickened;

fully winged;

legs

light;

length 6 mm;

black,

antennae 10-segmented,

densely hairy;

with mandibles

slender,

antenna I hairs very short,

distally

much shorter

than breadth of segments, weakly visible through the amber; antennal segments in following proportions (length): 12: 7: 39: 19: 18: 13: 8: 8: 6,5:

6,5;

aantennal

segments in following proportions

(breadth):

3:

2:

2:

2,5: 3: 3: 3: 2,5: 2: 2; antennae more than three times as long as head (length of head in dorsal view from occipital margin behind the ocelli to apex of mandibles):

137:40;

antennal

sockets touching margins of clypeus;

clypeus with ventral margin strongly bidentate; mandibles with 4 teeth progressing larger from anterior one to posterior; eyes apparently bare, except for some

long hairs;

of eye to base of mandibles; head,

where

it

malar space with a furrow joining occipital

is directed towards

carina

distinct only on

the upper corners of eyes;

carina not visible on sides of head and on ventral

lower corner dorsal

side of

occipital

side because of the

thickness of the amber; temples not prominent, absent; eyes normally protruding; POL; 5; OL ; 3; DOL; 9; OPL ; 1; frons with a complete median

keel;

vertex and frons apparently rugose,

pronotum crossed by an

anterior weak

anterior collar and the disc; strong posterior transversal

disc humped;

posterior collar very short,

(3:11);

pronotum fully with

granulated;

reticulate rugose; impression between

the

pronotum also crossed by a

impression between

collar; (36:40),

transversal

the disc and the posterior

much shorter than anterior collar pronotum slightly shorter than

posterior angles produced

into

lobes directed towards

head the

tegulae, but not reaching tegulae; sides of pronotum contracted where they meet the scutum; posterior transversal impression and posterior collar with a

distinct dorsal

median carina,

granulated and reticulate rugose, reaching

approximately 0,65

apparent I y scutell um,

scutum shorter than pronotum with

notaul ices incomplete,

lenth of scutum;

as scu I ptured as scutum,

scutellum and metanotum

shorter than scutum;

metanotum and pronotum usua I for

(22:36),

distinct,

shape of scutum,

Dryininae; propodeum not

Dryininae

Fig.

easily

586 -

867

Dryinus paJaeodominicanus Currado and Olmi (type)

Chela of

distinct through

the amber,

because of the thickness of the amber

and because the abdomen

is hiding

dorsal

reticulate rugose;

surface apparently

long as posterior' surface; on basal

cells and on distal

venation of the (30:4);

radial

proximal basal

part

closed;

(50:8);

very slender,

legs

in proportions:

(tarsal

dorsal

surface apparently as

two broad dark transversal

half of wings;

distal

radial

fore-wing

part of radial

vein s-shaped; (costal,

without dark bands,

wing

(tarsal

with

with

bands

the normal

Dryininae; pterostigma narrow, much longer than broad cell

cells clearly enclosed

wings hyaline,

21

fore-wing

the posterior surface of propodeum;

with shape usual 35

segment 1):

segment 4):

34

(coxae): 3

(tarsal

(tarsal

segments 2 and 3 of front

median

with

longer than the usual

and submedian cells);

three

hind

for Dryininae; foreDryininae; segments of the fore

with shape usual for

(trochanters):

segment 2):

segment 5):

trochanters nearly four times as fore tronchanters with a

40

vein much

fore-wing

8

34

long as broad

long proximal

62

(tarsal

56

24

fore

(greatest breadth):

into hooks;

(tibiae):

segment 3):

(enlarged claw);

slender stalk,

tarsus produced

(femora):

40:

9,5;

distally broadened; rudimentary

claw

868

Mem.

apparently present

31

lamellae;

central (tarsal

(34:8),

(tarsal

proportions:

25 (coxae):

segment 1): 11

(Fig.

1984

30

segment 2):

segment 5):

maxillary palpi

tibial

labial

43

49

(tibiae):

segment 3):

for

(femora):

spurs 1,1,2; segment very for

28 5

(tarsal

legs in following

15 (tarsal

length and breadth of the wings usual

morphology of the body usual

row

legs in following

(femora):

74 (tibiae): segment 3): abdomen

apparently with 6 segments;

last

much

lamellae located around a

segments of hind

segment 2):

586)

with 2-3 rows of nearly

10 (tarsal

10 (trochanters):

19 (tarsal

(Fig.

tooth and with a

586)

segments of mid

(trochanters):

apparently with 3 segments;

MALE:

37,

enlarged claw

tarsus

segment 5);

(tarsal

distinct petiole:

5

15 (tarsal

8

shape,

distinct);

lamellae;

14 (coxae):

segment 1):

segment 4):

no.

apex with a group of nearly 30

segment 4): (tarsal

Inst.,

with a strong subapical

segment 5 of front

hollow without

proportions:

Ent.

(but weakly

longer than arolium of 16 lamellae;

Amer.

31

7 (tarsal

without a

labial

palpi

long and slender;

Dryininae; shape and

Dryininae.

unknown

LOCUS TYP I CUS:

Dom i n i ca amber.

TYPICAL MATERIAL: HOSTS:

DISTRIBUT ION: NOTES:

holotype F!

in TR.

unknown Neotropic region:

only known from Dominica amber.

Dryinus. It Dryinus. The only species of Dryinus known from

this species shows morphologic characteristics of living

is belonging to the genus

D. antillean us (Evans), clearly distinct. D. palaeodominicanus is the only fossi I Dryinid known from Dominica

Dominica is

amber and from

the Neotropic region.

DRYINUS

AUSTRAL I AN SPEC I ES OF

Twenty

three species are known.

KEY TO THE SPECIES

FEMALES

Fore wings without

dark

........................ Fore wings with dark 2

Propodeum not (transversal

bands •••••••••••••••••••••••

1. pallidus (R.C.L. Perk i ns)

transversal

reticulate or

transversal

rugose,

longitudinal)

bands ••••••••••••••••••••••••• 2 with numerous parallel

keels at

least

on anterior half;

sometimes with weak areolae among keels •••••••••••••••••••••••• 3 Propodeum reticulate 3

rugose at

least

Propodeum with numerous parallel

on anterior half •••••••••• 5

longitudinal

keels .••••••••••••

Dry i n i nae

869

2. speciosus (8odd) Propodeum with numerous parallel

4

Pronotum wi th posterior nearly

three

species

times as

transversal

transversal

long as pronotal

reddish-testaceous,

tubercles

6

subtilis n.

darkened •..•••••..••••.

3.

Pronotum with posterior

transversal

7

carina

incomplete ••••••••••••••••••••

carina

complete .••••..•••

excava t ed ••••••••• swol len ••••••.••••

Occipital

carina

sides

of

• ••• 7 6.

ocellar

triangle;

...................•... fully fully

7.

the posterior ocelli;

head

pseudophanes (R.C.L. Perkins)

carina visible behind the

.•••.••.••••.. 6 . ••••••••••.•• 8

montanus n. sp.

only visible behind

granulated ••••••••••••• Occipital

5.

nearly

species mostly

Occipital Head

589 A);

sp.

impression narrow,

Occipital Head

(Fig.

wi th petiole black and abdomen

as long as pronotal tubercles (Fig. 589 B); b I a ck • • • • • • . • • • • . • • • • •• 4. pacificus n. sp.

5

keels •••••..••••• 4

impression very broad,

the posterior ocell i head punctate,

not

and

on

the

granulated

insularis (Dodd)

8

Head Head

or mostly reddish • • • . . • • • • . . • • • • • . • • • . . . • • • . • • • • . • • 9 or mostly black •••••••••••••••.•••..••..•••.•.•••• 10

9

rugose, without longitudinal 8. bouceki n. sp. Scutum reddish, rarely black, not reticulate rugose, with numerous longitudinal keels ••..••••.•••••.••••.••• . . . • • . . • • • • . . . . • . . . . • . . 9. koebelei (R.C.L.Perkins) Frons and vertex almost fully sculptured by numerous strong longitudinal irregular keels •••.••••.••••••••••..••••..•••..• 11 Frons and vertex without longitudinal keels or with a few keels on a narrow surface •••••••••..••••.•••••.••••.••••••••••••••• 12 Posterior ocelli touching occipital carina ••••••••••••••••••••• •••••••••••••••••••• 10. guineensis n. sp. Posterior ocelli not touching occipital carina •••••• •.••....•••...••.•••.. 11. gigas (R.C.L. Perkins) Metapleura wi th a narrow region smooth, shiny and hairless, without sculpture; pronotum, sometimes also head and scutum, fully smooth, shiny, hairless, without sculpture, sometimes only with Some keels on sides of pronotum .••••••••••••••••••• 13 Metapleura without region smooth, shiny and hairless; pronotum, head and scutum usually not smooth, shiny and hairless ••••••• 15 Head ex cava ted. • • • • • •• 15. montanus n. sp. Head flat •••••••••••••••••••••••••••••••••••••••••••••••••••• 14 Scutum shiny, smooth, without sculpture •••••••••••••••••••••••• •••••••••••••••••••••• 13. aterrimus (Dodd) Scutum dull, with numerous longitudinal keels •••••••••••••••••• • • • • • • • . • • • • • • • • • • • • •• 14. biaki n. sp. Scutum almost fully reticulate rugose •••••••••••••••••••••••• 16 Scutum not reticulate rugose or with a few areolae (sometimes

Scutum black,

fully

reticulate

keels ••••••••••••••••••

10

11

12

13 14

15

870

Mem.

on I y

n ea r

Amer.

Inst.,

frons and wertex not

punctate;

no.

37,

1984

ma rg in) •••••••••••••••••••••••••••••••••• 1 8

po s t e rio r

16 Pronotum,

Ent.

pronotum wi th

15. punctatus n.

· . . . . . . . . . . . . . . . . . . . .. Frons and vertex as

reticulate

rugose;

f ron s

smoo t h ,

weak keels around disc ••

reticulate

sp.

rugose as

scutum •••••••••••••• 17

17 Pronotum as reticulate rugose as scutum ••••••••••••••••• • • • • • • • • • • • • • • • • • • • • ••

16.

areolatus n.

sp.

Pronoium with keels around disc, not as reticulate scutum •••••••••••••••• 17. leptias (R.C.L. Perkins) 18 Scutum without

keels

or with

very

short

keels near

rugose as scutellum

• •••••••••••••.•••.••••••.•••••••••.••••.•.•••••••••••••••••• 19 Scutum almost

fully

19 Posterior ocel I i

sculptured by

touching

• • • • • • . • . . . • • • . . • . . • •. Posterior

ocelli

not

1 8.

20 Pronotum smooth,

19.

shiny,

carina;

TL

keels ••••••••• 20 shorter

than POL

qua te i n . sp.

touching

POL. • . • • . • • • • • • • • • • • ••

longitudinal

occipital

occipital

carina;

TL

longer

than

dahmsi n. sp.

almost

hairless,

wi thout

sculpture or

wi th a few keel s around di sc ••••••••••••••••••••••••••••••••• 21 Pronotum not smooth and shiny, sculptured by numerous keels around d i s c . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

• ••••• 22

21 OPL = POL; posterior collar of pronotum yellow.... • • • • • • . • . . • • • . . • . • • • •. 20. glaber n. sp.

• •••••••

Posterior POL;

ocell i

posterior

very

near occipital

collar of

carina;

OPL

shorter

than

pronotum black • . • . . • • • . . • • • • • • . • • • • • . •

• .•.••.•.•..•..••••••• 21. australianus n. sp. 22 Frons and vertex reticulate rugose •••••••••••••.•••••••••••••••

. . . . . . . . . . . . . . . . . . . . . . 22. venator (R.C.L. Perkins) Frons and vert ex not

ret i cu I ate

rugose,

punctate and wi th

some

keels •••••••••••••••••••••••••••••••••••....•••.••••••••••.•• 23

23 Posteri or ocell i farther from the occipi tal carina (Fig. 598 B); frons and vertex smooth, hai rl ess, finely punctate, wi th weak

keel s

numerous

connect ing

ocell i

to

longitudinal

keels,

punctate ••••••••••••••••

• • • . • • • • • • • • • • • • • . . . •• Posterior ocelli and vertex connecting

nearer

1 2.

frons;

bismarcki n.

occipital

scutellum wi thout sp.

carina

(Fig.

598 A)

smooth, hairless, weakly punctate, without ocelli to frons; scutellum with numerous

longitudinal

kels •••••

23.

papuanus n.

frons keels

sp.

MALES

Notaulices complete ••• Notaulices 2

16.

areolatus n. sp.

incomplete ••••••••••••••••••••••••••••••••••••••••• 2

Head granulated,

not

reticulate

rugose •••••••••••••••••••••••••

. . . . . . . . . . . . . . . . . . . . . . . 1. pallidus (R.C.L. Perkins)

Dryininae

Head more or 3

Dorsal

less

reticulate

surface of

Dorsal

as

long as metanotum •

aterrimus (Dodd)

13.

surface of propodeum

longer

than metanotum ••••••••••••• 4

Scutellum granulated and punctate, • • • • • • • • • • • • • • • • • • • • ••

5

rugose . . . . . . . . . . . . . . . . . . . . . . . . . . . 3

propodeum approximately

... .. ... • •• .. .. .. .. . . . 4

871

not

reticulate

australianus n.

21.

rugose . . . . . . .

sp.

Scutellum granulated and reticulate rugose •••••••••••••••••••• 5 Scutum reticulate rugose, very rugose; areolae wide . . . . . . . . . . . . ....................... Scutum reticulate

••••••••••••••••••••••

koebelei (R.C.L. Perkins)

9.

rugose,

less

rugose;

areolae

smal I •••••••••••

venator (R.C.L. Perkins)

22.

The males of other species are unknown.

1. Dryinus pallidus (R.C.L. Perkins) n. comb.

Chlorodryinus pallidus R.C.L. Perkins 1905: 57. nec Chlorodryinus pallidus R.C.L. Perkins: Pruthi and Mani 1942: 425 (see Dryinus indicus Kieffer). nee Chlorodryinus pallidus R.C.L. Perkins: Kurian 1954a: 257 (see Dryinus indieus Kieffer).

FEMALE: black;

fully

winged;

length 3,66-4,20 mm;

punctate and granulated, behind

the ocelli

pronotum shiny,

and on

swollen;

transversa I pronotal

(F i g.

carina

with some

incomplete,

du II, visible only

OPL shorter than OL;

longitudinal

pronotum crossed by

with petiole

499 B)

keels on sides

a strong posterior

impress ion and a weak anterior transversa I impress ion; tegulae;

irregular keels;

length of scutum;

bands with distal

scute II um and metanotum sh i ny,

part of radial

segment 1 of front

fore-wing

vein much

tarsus sl ightly

587)

with subapical

(Fig.

587)

with 2 rows of 26 lamellae;

tibial

spurs 1,1,2.

MALE:

fully

brown;

legs

yellow,

almost touching occipital

segment 4;

occipital

carina;

part

enlarged claw

segment 5 of front

head black,

tarsus

with mandibles

thorax and propodeum black;

with fore coxae black;

temples invisible;

as scu I ptured as

apex with a group of 18 lamellae;

length 2,19-2,74 mm;

antennae yellow-brown;

with

to beyond

without dark transversal

tooth and 12 lamellae;

yellow;

granulated, extending

longer than proximal

longer than

(Fig.

winged;

scutum shiny,

notaulices incomplete,

propodeum reticulate rugose;

granulated;

head

the sides of the vertex;

tubercles not reaching

numerous weak scutum;

occipital

without sculpture,

and on posterior impression;

0,5

yellow-testaceous,

head somet i mes wi th vertex brown ish;

head

carina

scutum dull,

(Fig.

499 D)

incomplete;

abdomen dull,

posterior ocelli

as sculptured as head;

872

Fig.

Mem.

587 -

Chela of

Amer.

Ent.

Inst.,

Dryinus pallidus

no.

(R.C.L.

37,

1984

Perkins)

from Wanuma

(New

Guinea)

notaulices incomplete,

reaching

approximately 0,5 length of scutum;

scutellum with anterior half as sculptured as scutum and with posterior half smooth,

without sculpture;

propodeum reticulate rugose;

metanotum as sculptured as scutum;

fore-wing

without dark transversal

bands,

Dryininae

Fig.

588 -

Chela of

with distal part of radial

Dryinus speciosus (Dodd) (type)

vein much

longer than proximal

spurs 1,1,2. LOCUS TYPICUS: Mulgrave (Australia) TYPICAL MATERIAL: lectotype F! and 5 paralectotypes (designated by Olmi HOSTS: in Australia

1982)

873

part;

(4 MM,

in B; 3 paratypes (2 MM, 1 F)! Siphanta sp. (R.C.L. Perkins 1906c).

tibial

1 F)!

in BM.

DISTRIBUTION: Australian region: AUSTRALIA: Mulgrave (Queensland), B! BM! Cairns (Queensland), B! Kuranda (Queensland), B! NEW GUINEA: Wanuma (Adelbert MIs.), B! Finschhafen, CAl Tapini (Cen. Dist., Papua), TW!

Ba i yer R.,

TW!

OL!

874

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

A

B

Fig.

589 -

Pronotum of females of Dryinus subtilis n.

sp.

(type)(A)

and

pacificus n. sp. from Keravat (New Britain) (B).

2. Dryinus speciosus (Dodd) n. comb.

Paradryinus speciosus Dodd 1916: 21.

FEMALE: brown

fully

wionged:

spot on vertex;

on si des of pronotum; propodeum black;

dull,

antennae testaceous;

in following proportions: POL = 3;

Ol = 2,5;

not touching occipital disc,

with a

transversal reaching

weak

carina;

28:

17:

OOl = 9;

OPl = 1;

pronotum dull,

a

meta thorax and

thickened; 15: 9:

with stalk

antennal

8:

7:

7:

occipital

Tl = 4;

8;

head

carina

posterior ocell i

with numerous keels around

impression and a strong posterior

posterior collar distinct;

scutum,

with

two brown spots

legs testaceous,

frons with a median keel;

anterior transversal

impression;

tegulae;

10: 4:

with

mesothorax,

antennae distally weakly

fully reticulate rugose;

complete;

head reddish-testaceous,

abdomen reddish-testaceous;

of hind femora black; segments

length 5 mm;

prothorax reddish-testaceous,

pronotal

tubercles not

scutellum and metanotum fully sculptured by

longitudinal parallel keels; notaulices invisible among keels; propodeum with a transversal keel between dorsal and posterior surface; dorsal surface not reticulate rugose,

with numerous parallel

posterior surface reticulate rugose; bands on the basal radial

vein

fore-wing

cells and beneath the pterostigma;

longer than proximal

following proportions:

20:

3:

7:

part 15: 26;

(17:9);

longitudinal

keels;

with 3 dark transversal fore

distal

tarsal

enlarged claw

part of

segments in

(Fig.

588)

with a

Dryininae

590 - Chela of Dryinus subtilis n. sp.

Fig.

subapical 588)

tooth and a row of 10 lamellae;

with 2 rows of nearly 23

segment 3 of front MALE: unknown lOCUS TYPICUS:

lamellae;

(type)

segment 5 of front

tarsus

apex with a group of 25

tarsus produced into a

Cairns

TYPICAL MATERIAL: HOSTS:

875

hook;

tibial

(Fig.

lamellae;

spurs 1,1,2.

(Australia)

holotype F!

in AD.

unknown

DISTRIBUTION:

Australian

region:

AUSTRALIA:

Cairns

(Queensland),

AD!

3. Dryinus subtilis n. sp.

FEMALE: black

fully

longitudinal behind

winged;

length 8,12 mm;

and abdomen darkened; striae on frons;

the ocell i

and on

reddish-testaceous,

head shiny, occipital

weakly punctate,

carina

incomplete,

the sides of the vertex;

with petiole with weak

only

OPl ~ Ol;

visible

pronotum

876

Mem.

Fig. 591

(Fig.

589 A)

pronotal pronotal numerous

shiny,

-

Amer.

En!.

longitudinal

keels;

sp.

notaulices incomplete,

(type)

reaching

scutellum and metanotum shiny,

long as

part;

enlarged claw

bands,

with distal

segment 1 of front

(Fig.

segment 5 of front tarsus

apex with a group of 15 lamellae;

590) (Fig.

tibial

parallel

keels;

part of radial

tarsus slightly

with subapical 590)

approximately

weakly punctate;

with numerous transversal

with 2 brown transversal

than segment 4;

MALE:

1984

with posterior collar nearly three times as

longer than proximal

lamellae;

37,

Chela of Dryinus pacificus n.

propodeum not reticulate rugose, much

no.

tubercles; posterior collar granulated; disc without sculpture; tubercles not reaching tegulae; scutum shiny, fully sculptured by

0,75 length of scutum; fore-wing

Inst.,

vein

longer

tooth and 18

with 2 rows of 33 lamellae;

spurs 1,1,2.

unknown

LOCUS TYPICUS:

Ifar (New Guinea)

TYPICAL MATERIAL:

holotype F!

HOSTS: unknown DISTRIBUTION: Australian

region:

in B. NEW GUINEA:

Ifar (Western

Irian),

B!

4. Dryinus pacificus n. sp.

winged:

length 3,43 mm;

head black,

partly clypeus yellow;

FEMALE:

fully

antennae brown,

wjth segments 1-2 yellow;

and propodeum black,

with mandibles and

with sides of pronotum reddish;

thorax

abdomen brown-

877

Dryininae

Fig.

592 -

reddish;

legs brown,

occipital

carina

(anterior

Chela of

with

incomplete;

tarsi

Dryinus montanus

light;

n.

sp.

head excavated,

(type)

shiny,

pronotum crossed by 2 transversal

impression weaker than posterior impression);

punctate; impressions

pronotum

(Fig. 589 B) dull, hairy, weakly granulated; pronotal tubercles not reaching tegulae; scutum dull, hairy, strongly granulated; notaulices incomplete, extendi ng to beyond 0,5 length of scutum; scutell um and metanotum as sculptured as scutum; propodeum not reticulate rugose, fully sculptured by numerous parallel keels; fore-wing with 2 brown transversal bands and with distal part of radial vein longer than proximal part; segment 1 of front tarsus slightly longer than segment 4; enlarged claw with subapical tooth and 16 lamellae (Fig. 591); segment 5 of front tarsus (Fig. 591) with 2 rows of 30 lamellae; apex with a group of 13 lamellae; tibial spurs

1,1,2. MALE:

unknown

LOCUS TYPICUS:

Saiyer

TYPICAL MATERIAL: OL.

(New Guinea)

holotype F!

and 1 para type F!

in

B;

1 para type F!

in

878

Mem.

HOSTS:

Amer.

En!.

Ins!.,

no.

37,

1984

unknown

DISTRIBUTION:

Australian

NEW BRITAIN:

Keravat,

region:

OL!

NEW GUINEA:

Vunakanau

Baiyer

(W

(Gazelle Pen.),

Highiands),

B!

B!

5. Dryinus montanus n. sp.

FEMALE:

fully

winged;

and partly clypeus black,

with

abdomen

sides

reddish;

brown-reddish:

punctate;

occipital

apparently

POL;

length 4,37-5,00 mm;

yellow or reddish;

carina

incomplete;

keels;

complete;

smooth~

posterior transversal

impression dull,

very

fully

weak,

excavated,

sometimes

the carina

impression

rugose;

scutellum surface

without

longitudinal

hairless,

without

with distal front

tarsus

nearly

of front

tarsus

long

592)

(Fig.

lamellae;

MALE:

as

(Fig.

is

weakly

strong;

and

very

with

a

subapical

with 2 rows od 33

spurs

short

scutum

almost propodeum

transversal

narrow

and

and

some

tegulae;

reaching

region

keels

part;

15

lamellae;

and

smooth,

tranversal

proximal

tooth

and

anterior transverSal

with 2 brown

longer than

weak

OL : : :

punctate with

metanotum punctate;

numerous

black; weakly

carina;

into a

lamellae; apex

shiny,

bands

segment

as segment 4 segment 3 produced

with

592)

tibial

disc

incomplete,

with

fore-wing

vein

mandibles

pronotum

shiny,

posterior collar

very

metapleura

sculpture;

part of radial

enlarged claw of 13

keels;

occipital

tubercles not reaching

reticulate

posterior

with

notaul ices

posterior margin of scutumj rugose;

touching

sculpture;

pronotal

reticulate

head

pronotum shiny, without

with

testaceous;

metathorax and propodeum

testaceous;

posterior ocelli

temples prominent;

anterior collar

mesothorax,

legs

head black,

antennae

and

1 of hook;

segment 5

wi th

a

group

1,1,2.

unknown

LOCUS TYPICUS: TYPICAL HOSTS:

Bunya Mts

MATERIAL:

(Australia)

holo)ype F!

in QU;

1 paratype F!

in

OL.

Conondale,

OL!

unknown

DISTRIBUTION: (Queensland),

Australian

region:

AUSTRALIA:

Bunya MIs.

QU!

6. Dryinus pseudophanes (R.C.L. Perkins) n. comb.

Chlorodryinus pseudophanes R.C.L. Perkins 1906a: 492.

FEMALE: antennae

fully

winged;

yellow-brown:;

r-eddish;

mesothorax,

reddish;

legs

carina

length 4,75 mm; pronotum

meta thorax

brown~reddish;

incomplete,

only

and

head

head

black,

with

black, sides

with mandibles

propodeum black;

shiny,

visible behind

swollen,

the ocell i;

yellow;

and posterior collar abdomen

granulated;

pronotum

brownoccipital

without

Dryininae

Fig.

sculpture,

593 -

shiny,

Chela of

Dryinus insularis (Dodd) (type)

with two transversal

weaker than posterior impression); notaulices incomplete,

879

impressions

(anterior impression

scutum as sculptured as head;

reaching approximately 0,5 length of scutum;

scutellum and metanotum as sculptured as head; propodeum reticulate rugose; fore-wing with 2 brown transversal bands; enlarged claw with subapical

tooth and with some lamellae;

rows of lamellae; MALE:

unknown

LOCUS TYP I CUS:

segment 5 of front

apex with a group of lamellae;

Brisbane

TYPICAL MATERIAL:

(Austra I i a)

holotype F!

in B.

tibial

tarsus with

spurs 1,1,2.

880

Mem.

Amer.

En!.

HOSTS: unknown DISTRIBUTION: Australian region:

Ins!.,

no.

AUSTRALIA:

37,

1984

Brisbane

(Queensland),

B!

7. Dryinus insularis (Dodd) n. comb.

Paradryinus insularis Dodd 1913: 180. Paradryinus insularis Dodd: Kieffer 1914b: 38.

FEMALE: frons,

fully

winged;

propodeum black, fore

38:

head blaCk,

weakly

22:

18:

with anterior part of

antennae testaceous;

except for margins of pronotum reddish;

legs testaceous;

distally 6:

length 5,62 mm;

cl ypeus and mand i b I es testaceous; mid and hind

thickened; 12: 9:

legs brown,

antennal

with

tarsi

thorax and

abdomen brown; light;

antennae

segments in following proportions:

8 (last two antennal

segments missing

8:

in holotype);

head shiny, weakly swollen, strongly punctate, without sculpture among punctures; frons with a median keel; occipital carina incomplete, visible behind

the ocelli

and on

not touching occipital

the sides of the ocellar triangle;

carina;

POL

=

4;

OL

=

3;

OOL

=

10;

posterior ocelli OPL

=

0,5;

pronotum dull, hairy, with numerous longitudinal striae around disc, with a weak anterior transversal impression and a strong posterior transversal impression;

sides

posterior collar

of

pronotum

long;

pronotum very convex,; weakly

visible,

partly

pronotal

scutum dull,

incomplete,

smooth,

shiny

and

tubercles not reaching

without

tegulae;

fully reticulate rugose;

sculpture;

disc of notaulices

reaching approximately 0,5 length of scutum;

scutell um and metanotum du II, strong I y puncta te; propodeum du II, reticulate rugose, without longitudinal or transversal keels; fore-wing

with

2 dark transversa I bands, one on the basa I cell s and another beneath the pterostigma; distal part of radial vein longer than proximal part (14:6); fore tarsal

segments

in following proportions:

20:

4:

8:

21:

35;

enlarged

claw (Fig. 593) with a subapical tooth and a row of 13 lamellae; segment 5 of front tarsus (Fig. 593) with 2 rows of nearly 39 lamellae; apex with a group of 18

lamellae;

tibial

spurs 1,1,2.

r.-1ALE:

unknown

LOCUS TYPICUS:

Double

TYPICAL MATERIAL:

segment 3 of front

Island

Queensland),

into a hook;

(Australia)

holotype F!

H05TS: unknown DISTRIBUTION: Australian

tarsus produced

region:

in AD. AUSTRALIA:

Double

AD!

8. Dryinus bouceki n. sp.

Island

(Cairns,

881

Dryininae

Fig.

594 -

Chela of Dryinus bouceki n.

sp.

(type)

882

Mem.

FEMALE: reddish;

antennae testaceous,

Ins!.,

no.

impression;

37,

1984

head reddish,

mesothorax black,

abdomen

legs almost fully

occipital

brown;

testaceous;

carina complete;

propectus

with mesopleura partly reddish-

scutell um and metanotum black or reddish-brown;

metapleura black;

sometimes with

with segments 6-10 brown;

with 2 brown spots on sides of anterior

pronotum reddish,

transversal

fore

Ent.

blly winged;length 5,62-5,75 mm;

black spots;

brown;

Amer.

legs brown, head dull,

posterior ocelli

with

flat,

not

propodeum and

tarsi

light;

sometimes

reticulate rugose:

touch"lng occipital

carina;

OPL shorter than OL; pronotum dull, with numerous keels around disc; posterior transversal impression strong; anterior transversal impression weak;

posterior collar very

tubercles not reaching incomplete,

reaching

metanotum dull, longitudinal part;

anterior collar weak;

scutum reticulate rugose;

almost posterior margin of scutum;

hairy,

rugose;

segment

part of radial

vein much

tarsus

longer than segment 4

into a

hook;

2 rows of 26

12 lamellae;

lamellae;

enlarged claw

segment 5 of front

apex with a group of 24

without transversal

longer than proximal

1 of front

tooth and

scutellum and

with 3 brown

fore-wing

tarsus produced

subapical

pronotal notaul ices

propodeum reticulate rugose,

keels on posterior surface;

bands and with distal of front

visible;

tegulae;

(18:14);

(Fig. tarsus

lamellae;

segment 3

594)

with

(Fig. tibial

594)

with

spurs

1,1,2. MALE:

unknown

LOCUS TYPICUS:

Indooroopilly

TYPICAL f,IATERIAL: F!

(Australia)

holotype F!

in CB;

1 paratype F!

in AD;

1 paratype

in UO.

HOSTS: unknown DISTRIBUTION: Australian Oueensland), NOTE:

CB!

the species

region:

Ashgrove, is named

UO! in

AUSTRALIA: Adelaide

Indooroopilly

(Brisbane,

(South Austral ia),

honor of Mr.

AD!

Zdenek Boucek.

9. Dryinus koebelei (R.C.L. Perkins) n. comb.

Paradryinus koebelei R.C.L. Perkins 1905: 54.

FEMALE:

fully

winged;

length 5,12-6,22 mm;

head,

antennae,

pronotum,

propectus, scutum, scutellum reddish; rarely scutum black; rarely pronotum part I y black; transversa I furrow between scutum and scutell um brown;

rarely

scutellum black;

scutellum and metanotum brown; abdomen

reddish-brown;

OPL shorter than

metanotum brown or reddish; propodeum black or brown;

legs reddish;

with segment 3 much

longer than segment 4;

pronotum crossed by 2 transversal than

posterior

head dull,

length of ocellar triangle;

impression);

impression

pronotum dull,

sides of petiole black;

rugose and granulated;

antennae shorter than body, occipital

carina complete;

(anterior impression

weaker

with numerous keels around

Dryininae

Fig. 595 -

883

Dryinus koebelei (R.C.L. Perkins) from Bundaberg

Chela of

(Queens I and)

disc;

pronotal

tubercles not reaching

sculptured by numerous to beyond 0,5

longitudinal

length of scutum;

metanotum dull,

rugose;

tarsi

part of radial

tooth and 8

vein

1,1,2. MALE: fully yellow;

legs yellow;

length 2,56-3,66 mm;

part;

hairy,

longitudinal

brown bands;

segment 1 of fore

tarsus

head black,

thorax and propodeum black;

head dull,

(Fig. tibial

595)

with

spurs

with mandibles abdomen brown-

rugose and granulated;

length of ocellar triangle;

scutum as sculptured as head;

extending

enlarged claw with subapical

segment 5 of front

incomplete; OPL shorter than

invisible;

without

apex with a group of 15 lamellae;

antennae yellow;

reddish; carina

winged;

595);

(17:8,5);

fully

as scu I ptured as scutum;

with 3 transversal

longer than proximal

lamellae (Fig. lamellae;

scutell um du II,

fore-wing

twice as long as segment 4

2 rows of 20

scutum dull,

notaulices incomplete,

propodeum reticulate rugose,

keels on posterior surface; distal

tegulae; keels;

notaulices incomplete,

occipital temples extending

884

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

(1

.

'

~",~.'

...

'~

\ 0:: ' 'i

Fig.

to beyond 0,5

596 -

Chela of Dryinus guineensis n.

length of scutum;

except for posterior third smooth;

without

longitudinal

part;

tibial

bands,

propodeum reticulate rugose,

keels on posterior surface; with distal

part of radial

fore-wing

vein

1983)

without dark

longer than proximal

spurs 1,1,2.

LOCUS TYPICUS: Bundaberg (Australia) TYPICAL MATERIAL: lectotype F! and 4 paralectotypes FF! Olmi

(type)

scutell um and metanotum as scu I ptured as

scutum,

transversal

sp.

'

(designated by

in B.

HOSTS: in Australia Flatidae: Colgar sp., Siphanta sp., Massila sp., Phantiopsis (= Aphanophantia) sp. (?) (R.C.L. Perkins 1906c). DISTRIBUTION: Australian region: AUSTRALIA: Bundaberg (Queensland), OL! B! Brisbane (Queensland), QU! UQ! Acacia Ridge (Brisbane, Queensland), QU!

Southport

(Queensland),

OL!

Gold Creek, QU! Deception Bay, NEW GUINEA: Gent R., OLI TWI

Mt. BR!

Glorious

(SE Queensland),

Botany Bay

QU!

TW!

(New South Wales),

WA!

10. Dryinus guineensis n. sp.

885

Dryininae

FEMALE:

fully

winged;

mandibles yellow;

length 5,62-6,25 mm;

antennae brown;

thorax and propodeum black; hind coxae black;

head black,

abdomen brown-reddish;

legs brown,

trochanters and fore and mid coxae yellow;

granulated,

with frons and vertex fully sculptured by

longitudinal

keels;

occipital

with segment 3 much

carina complete; scutum dull,

as sculptured as head;

with

head dull,

numerous

antennae shorter than body,

longer than segment 4;

numerous keels around disc; keels,

with clypeus and

with segments 1-2 and 8-10 yellow;

pronotum shiny,

with

numerous

notaulices incomplete,

with

longitudinal

extending

to beyond

0,5 length of scutum; scutellum dull, granulated; metanotum rugose; propodeum reticulate rugose, without longitudinal keels on posterior surface; and

rows of 37

transversal

longer than segment 4;

18 lamellae

MAL::::

with 2 brown

fore-wing

tarsus slightly

(Fig.

lamellae;

596);

segment 5 of front

apex with a

segment 1 of front

bands;

enlarged claw with subapical

group of 15

tarsus

(Fig.

lamellae;

596)

tibial

tooth

with 2

spurs 1,1,2.

unknown

LOCUS TYPICUS:

Brown River

TYPICAL MATERIAL:

(Papua,

holotype F!

New Guinea).

in B;

1 para type F!

in LE;

1 paratype F!

in TW. HOSTS:

unknown

DISTRIBUTION: Wau,

LE!

Australian

region:

Lae-Zenag Road,

~<EW

GUINEA:

Brown River

(Papua),

B!

TW!

11. Dryinus gigas (R.C.L. Perkins) n. comb.

Paradryinus gigas R.C.L. Perkins 1905: 55. Neodryinus semipolita Girault 1925; 2 (n. syn.); locus typicus Brisbane

(Australia).

Paradryinus nigricorpus Girault 1925: 2 (n. syn.); locus typicus Gordonvale

FEMALE:

fully

winged;

pronotum black, mesothorax, reddish;

length 8,12-9,68 mm;

with anterior margin

head black;

shiny,

segment 3 much

hairy;

abdomen reddi sh-brown:

antennae distally

longer than segment 4;

keels on frons

antennae reddish;

and posterior collar reddish;

meta thorax and propodeum black;

body

longitudinal

(Australia).

weakly

thickened;

legs

with

head with numerous strong

and vertex;

pronotum with numerous keels

around disc; scutum fully reticulate rugose; notaul ices incomplete, extending beyond 0,5 length of scutum; scutellum as sculptured as scutum; metanotum rugose; enlarged claw of front of 20

tarsus

lamellae;

part of radial

propodeum reticulate rugose,

(Fig. (Fig.

597)

with subapical

597)

fore-wing vein

longer

with 2 rows of 33 with 3 brown than

without

tooth and 20 lamellae;

transversal

proximal

part;

longitudinal

lamellae;

keels;

segment 5

apex with a

group

bands and with distal

tibial

spurs 1,1,2.

886

Mem. Amer. En!.

Ins!.,

no. 37,

\

1984

\

1 ~ I,

Fig. 597 - Chela of Dryinus gigas (R.C.L. Perkins) from Brisbane (Queens I and)

Dryininae

Fig.

598 -

MALE:

887

Heads of fema les of Drylnus papuanus n. Guinea)(A)

and bismarcki n.

Bundaberg

(Australia)

sp.

sp.

from Kassam

(New

(type)(B)

unknown

lOCUS TYPICUS:

of P. gigas in B; holotype F! of N. semipollta in QU; holotype F! of P. nigricorpus in QU. TYPICAL MATERIAL:

HOSTS:

holotype F!

unknown

DISTRIBUTION:

Australian

Brisbane (Queensland), Townsville

region:

BR!

Ol!

(North Queensland),

Queensland),

AUSTRALIA: QU!

AD!

Bundaberg

(Queensland),

Gordonvale (Queensland), Glasshouse,

UQ!

B!

QU!

Enoggera (Brisbane,

Oll

12. Dryinus bismarcki n. sp.

FEMALE:

fully

winged;

length 5,60 mm;

head black,

with clypeus and

mandibles yellow; antennae testaceous; pronotum black, with anterior edges of si des redd ish; mesothorax, meta thorax and propodeum black; abdomen brown;

legs yellow,

brown;

a brown spot is visible on fore coxae;

hairless,

with mid and coxae,

mid and hind clubs of femora

weakly punctate and with some weak

posterior ocell i almost touching occipital

head

carina;

pronotum shiny,

without sculpture,

smooth;

tubercles not reaching tegulae;

fully

pronotal

sculptured by numerous

extending

to beyond 0,5

metanotum rugose; parallel

599)

with

shiny,

occipital carina complete;

longitudinal keels;

length of scutum;

scutum dull,

notaul ices incomplete,

scutellum shiny,

fore-wing

part of radial

tarsus slightly

subapical

tooth

vein

with 3 brown

punctate;

14 lamellae;

with

transversal

longer than proximal

longer than segment 4;

and

disc

granulated,

propodeum reticulate rugose on anterior surface,

bands and with distal (Fig.

598 B)

with numerous keels around disc;

keels on posterior surface;

segment 1 of front

(Fig.

longitudinal keels on frons;

part;

enlarged claw

segment 5 of front

tarsus

888

Mem.

Fig.

(Fig.

599)

599 -

Amer.

Ent.

Inst.,

no. 37,

1984

Chela of iJryinus bismarcki n.

with 2 rows of 33 lamellae;

apex with

sp.

(type)

a group of 17 lamellae;

tibial spurs 1,1,2. MALE: unknown LOCUS TYPICUS:

Momote (Manus

TYPICAL MATERIAL: HOSTS:

Island,

holotype F!

Bismarck Archipelago)

in B.

unknown

DISTRIBUTION: Island),

Australian region:

BISMARCK ARCHIPELAGO:

B!

13. Dryinus aterrimus (Dodd) n. comb.

Paradryinus aterrimus Dodd 1916: 21.

Momote (Manus

Dryininae

Fig.

600 -

FEMALE:

Chela of

fully

winged;

mandibles yellow; thorax and smooth,

889

Dryinus aterrimus (Dodd) from Laloki (New Guinea)

lengh 4,68-5,31

antennae brown,

propodeum black;

without sculpture;

mm;

head black,

with clypeus and

with segments 1-2 and

abdomen brown;

10 yellow;

I egs brown;

occipital carina complete;

head sh i ny,

posterior ocell i very

near occipital carina; pronotum crossed by 2 transversal impressions (anterior impression weaker than posterior impression); pronotum shiny, smooth,

without sculpture;

disc sometimes raised

scutum shiny,

smooth,

to beyond 0,5

length of scutum;

sculpture; surface,

without sculpture;

metanotum rugose;

with

parallel

into a strong carina;

notaulices incomplete,

scutellum shiny,

smooth,

extending

without

propodeum reticulate rugose on anterior

keels on

posterior surface;

fore-wing

with 3 brown

890

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

transversal bands; segment 1 of front tarsus sl ightly longer than segment 4; enlarged claw with subapical tooth and 11 lamellae (Fig. 600); segment

5 of front tarsus (Fig. 600) of 13 lamellae; MALE:

fully

tibial

winged;

antennae brown, proximal

with a

invisible;

occipital

scutum dull, reaching dull,

carina

OL = 3;

incomplete,

granulated and fully

as sculptured as scutum;

without than

POL = 5;

keel

proximal

keels;

OOL = 5;

(23: 11);

spurs 1,1,2. LOCUS TYPICUS:

Cairns

TYPICAL MATERIAL:

10:

OPL =

with

10:

9: 9: 8:

10:

areolae small;

frons

1,5; temples

reaching posterior edges of eyes;

reticulate rugose;

notaulices

incomplete,

scutellum and metanotum

and posterior surface;

fore-wing hyaline,

part of radial part

group

thickened;

propodeum reticulate rugose,

between dorsal

longitudinal distal

13:

reticulate rugose;

approximately 0,65 length of scutum;

transversal bands;

7: 4:

in following proportions:

keel;

legs testaceous,

antennae not distally

granulated and fully

median

apex with a

2,87 mm; black; mandibles testaceous;

length

half of hind coxae brown;

head dull,

lamellae;

with segments 1-2 testaceous;

antenna I segments 10;

with 2 rows of 29

spurs 1,1,2.

without

posterior surface

without dark

vein reaching margin of wing, gonoforceps without dorsal

transversal much

process;

longer

tibial

(Australia)

holotype F!

HOSTS: unKnown DISTRIBUTION: Australian

in AD.

region:

AUSTRALIA:

Cairns

(Wueensland),

AD!

I~EW GUINEA:

OL!

B!

Popondetta (Papua), OL! BM! Laloki (Port Moresby, Papua), Swart Val (Ridge), B! Wau (Morobe Distr.), BM! B! SOLOMON

ISLANDS:

Kira Kira

(San Cristobal),

B!

Gizo

Island

(New Georgia group),

OL!

14. Dryinus biaki n. sp.

FEMALE:

fully

winged;

mandibles yellow; abdomen brown;

length 4,68 mm;

brown-redd i sh; head shiny,

connecting ocell i posterior ocelli

to

very

with segment 3 much transversal

head black,

antennae reddish-brown; legs testaceous,

with cl ubs of femora and coxae

smooth,

without sculpture,

internal

margin of eyes;

near occipital carina;

with

some

occipital

longitudinal

impression

keels

carina complete;

antennae shorter than

longer than segment 4;

impressions (anterior

with clypeus and

thorax and propodeum black;

pronoturn humped, weaker than

body,

with 2

posterior

irnpression); pronoturn shiny, smooth, without sculpture; scutum dull, hairy, with numerous longitudinal keels; notaul ices incomplete, reaching approxirnately 0,5 rugose;

length of scutum;

scutellum dull,

granulated;

propodeum reticulate rugose on anterior half,

on posterior surface; part of radial tarsus sl ightly

vein

fore-wing much

with 3 brown

transversal

longer than proximal

longer than segment 4;

part;

enlarged claw

rnetanotum

with parallel bands;

segment

keels

distal

1 of front

with subapical

tooth

Dryininae

Fig.

601

and 10 lamellae (Fig. rows of 23 lamellae; MALE: unknown LOCUS TYPICUS: HOSTS:

Chela of

601);

Island

LJryinus biaki n. sp. (type)

segment 5 of front

apex with

Biak

TYPICAL MATERIAL:

-

891

tarsus

(Fig.

a group of 14 lamellae;

601)

tibial

(New Guinea)

holotype F!

in B.

unknown

DISTRIBUTION:

Australian region:

NEW GUINEA:

Biak

Island,

B!

with

2

spurs 1,1,2.

892

Mem. Amer. En!.

Inst.,

no. 37,

1984

I

(~: I

\ \

Fig. 602 -

\

Chela of Dryinus punctatus n. sp.

(type)

Dryininae

893

15. Dryinus punctatus n. sp.

FEMALE: vertex, black;

fully

winged;

length 8,12 mm;

clypeus and mandibles yellow; pronotum black,

propodeum black; coxae black;

head black,

with sides reddish;

abdomen brown-reddish;

head shiny,

posterior ocell i very

excavated,

with front part of

antennae testaceous; mesothorax, legs brown,

propectus

metathorax and with tarsi

light and

punctate; occipital carina complete;

near occipital carina;

pronotum dull,

hairy,

granulated, with weak keels around disc; pronotum crossed by 2 transversal impressions (anterior impression weaker than posterior impression);

pronotal

reticulate rugose; scutum;

tubercles not reaching

notaulices incomplete,

scutellum as sculptured as scutum;

Fig. 603 -

tegulae;

scutum dull,

extending to beyond 0,5 metanotum rugose;

Chela of Dryinus areolatus n.

sp.

fully length of

propodeum

(type)

894

Mem. Amer.

reticulate rugose,

without

wing with 2 brown much

bands, (Fig.

tarsus

apex with a group of 14 lamellae; MALE:

no.

37,

1984

keels on posterior surface; with distal

segment 1 front

part;

enlarged claw

segment 5 of front

lamellae;

I nst.,

longitudinal

transversal

longer than proximal

than segment 4;

Ent.

602) (Fig.

tibial

part of radial tarsus slightly

with subapical

tooth

longer

and

with 2 rows of 36

602)

forevein 18

lamellae;

spurs 1,1,2.

unknown

LOCUS TYPICUS: Kuranda (Australia) TYPICAL MATERIAL: holotype F! in OL. HOSTS:

unknown

DISTRIBUTION:

Australian

region:

AUSTRALIA:

Kuranda

(Queensland),

OL!

16. Dryinus areolatus n. sp.

FEMALE: vertex

fully

winged;

brown-reddish; with

length 5,62 mm;

(more along orbits),

tarsi

complete;

thorax and propodeum black;

light;

head dull,

OPL shorter than

than body,

head black,

with

posterior impression);

antennae

abdomen brown;

fully reticulate rugose;

legs brown,

occipital carina

length of ocellar triangle;

segment 3 much

crossed by 2 transversal

with front paret of

clypeus and mandibles yellow;

antennae shorter

longer than segment 4;

pronotum hairy,

impressions (anterior impression weaker than

pronotum reticulate rugose;

pronotal

tubercles not

reaching tegulae; scutum dull, fully reticulate rugose; notaul ices incomplete, extending to beyond 0,5 length of scutum; scutellum and metanotum reticulate rugose; surface,

with

transversal proximal

propodeum reticulate rugose on anterior

parallel keels on posterior surface;

bands,

part;

enlarged claw

with

distal

part of radial

segment 1 of front with subapical

tarsus sl ightly

tooth

and

fore-wing

vein much

longer than segment 4;

14 lamellae (Fig.

of front tarsus (Fig. 603) with 2 rows of 36 of 19 lamellae; tibial spurs 1,1,2.

with 3 brown

longer than 603);

segment 5

iamellae; apex with

a group

MALE: fully winged; length 3,06 mm; black; mandibles testaceous; antennae brown, with segments 1-2 testaceous; legs testaceous, with coxae proximally

infuscate;

antennae not distally

segments in folowing proportions: dull,

granulated,

longitudinal 3,5;

with

keels;

OPL: 1;

weak

occipital

temples

6:

13:

9:

9:

9:

8:

areolae mainly on frons; carina

invisible;

areolae on anterior half;

7:

thickened;

incomplete;

scutum dull,

notaulices complete,

8:

8:

11; head

frons without

POL: 7;

fully

hind

antennal

OL :

2,5;

granulated,

OOL

with weak

posteriorly separated;

scutellum and metanotum shiny, weakly punctate, very weakly granulated; propodeum reticulate rugose, without a transversal keel between dorsal and posterior surface; hyaline,

powterior surface without

without dark transversal

longer than proximal part gonoforceps (Fig.

604)

(20:11)

bands;

longitudinal

distal

keels;

part of radial

fore-wing vein much

and reaching margin of wing;

without dorsal membranous process;

tibial

spurs

Dryininae

Fig. 604 -

Male genitalia of Dryinus areolatus n. sp. from Mt. Caledonia): left half removed.

895

Koghi

1,1,2. LOCUS TYPICUS: Couli (New Caledonia) TYPICAL MATERIAL: holotype F! in B; 1 paratype M! in OL. HOSTS: unknown DISTRIBUTION: Australian region: NEW CALEDONIA: Couli (La Foa), Koghi, OLl

B!

(New

Mt.

17. Dryinus leptias (R.C.L. Perkins) n. comb.

Paradryinus leptias R.C.L. Perkins 1905: 56.

FEMALE: fully winged; length 4,02 mm; head black, with clypeus and mandibles yellow; antennae yellow-brown; pronotum black, with anterior margin yellow and sides reddish; mesothorax, metathorax and propodeum black; abdomen brown-reddish; legs reddish; antennae distally thickened, with segment 3 much longer than segment 4; head granulated; occipital carina complete;

OPL shorter than

length of ocellar triangle; ocellar

896

Mem.

Fig.

region

605 -

swollen;

Chela of

visible,

Ins!.,

Dryinus quatei

n. Guinea)

scutum granulated,

incomplete,

no.

sp.

37,

part of radial

without

1984

from Kulumadau Hill

pronotal

longitudinal

reaching approximately 0,5

propodeum reticulate rugose; with distal

En!.

pronotum with keels around disc;

reaching tegulae; weakly

Amer.

fore-wing

vein

with 2 brown

longer than

proximal

(New

tubercles not keels;

notaulices

length of scutum;

transversal part;

bands and

enlarged claw

Dryininae

with subapical rows of MALE:

segment 5 of front

tooth and some lamellae;

lamellae;

897

apex with a group of

lamellae;

tarsus with

spurs 1,1,2.

tibial

unknown

LOCUS TYPICUS:

Cairns

(Australia)

TYPICAL MATERIAL:

holotype F!

HOSTS: 1954) •

Hasta hastata rDictyopharidae); Coigar sp. (Thompson

in Austra I i a

DISTRIBUTION:

Australian

in B.

region:

AUSTRALIA:

Cairns

(Queensland),

1:3!

18. Dryinus quatei n. sp.

FEMALE:

fully

winged;

mandibles reddish;

length 3,12-4,68 mm;

antennae brown,

sometimes segments 8-10 yellow;

head black,

with clypeus and

segments 1-2 and

with

10 yellow;

thorax and propodeum black;

abdomen

brown-reddish; legs testaceous, with clubs of femora and sometimes hind coxae black; head dull, granulated, with some longitudinal keels on sides of frons; occipital carina complete; POL = 5; TL = 2; posterior ocell i touching occipital

OL = 2,5; OOL = 8,5; OPL = carina; pronotum crossed by

0;

2 transversal

impression);

impressions

(anterior impression

pronotum dull,

tubercles not reaching

weaker than

posterior

with numerous keels around disc;

tegulae;

scutum dull,

hairy,

pronotal

granulated;

notaulices

incomplete, extending to beyond 0,5 length of scutum; scutellum and metanotum dull, a sculptured as scutum; propodeum reticulate rugose on anterior surface, brown

with parallel

transversal

than proximal

keels on

bands and with

segment 1 of front

part;

posterior surface;

distal part of radial tarsus sl ightly

4; enlarged claw

(Fig.

605)

with

5 of front

(Fig.

605)

with 2 rows of

tarsus

of 14 lamellae; MALE: unknown LOCUS TYPICUS:

tibial

(New

NOTE:

1:3!

Kulumadau

the species

longer than segment

tooth and 9

lamellae;

19

apex with a

lamellae;

segment group

Guinea)

holotype F!

in B;

HOSTS: unknown lJISTRIBUTION: Australian region: Volgelkop),

with 3

longer

spurs 1,1,2.

Kebar Val

TYPICAL tvlATERIAL:

subapical

fore-wing

vein much

Hill

is named

1 paratype F!

NEVV GUINEA:

(Woodlark

Kebar Val

Island,

in honor of Mr.

in OL.

L.

Murua,

(1\IIanokwari, Papua),

OL!

Quate.

19. Dryinus dahmsi n. sp.

FEMALE:

fully

winged;

mandibles testaceous; margin,

length

3,43-4,25 mm; head black,

antennae testaceous;

posterior collar and sometimes

mesothorax,

with clypeus and

pronotum black,

lateral

meta thorax and propodeum black;

with

anterior

margins reddish; abdomen

black-brown;

legs

898

Mem.

Fig.

606 -

testaceous or brown, granulated; occipital

Amer.

Ent.

tarsi

light;

POL; OL;

TL

37,1984

head weakly

occipital carina complete;

carina;

no.

Dryinus dahmsi n. sp. (type)

Chela of

with

Inst.,

swollen,

posterior ocelli

longer than

POL;

not

shiny,

touching

pronotum shiny,

weakly

granulated; posterior transversal impression very strong; posterior collar distinct; anterior transversal impression weak; pronotal tubercles not reaching tegulae; scutum shiny, granulated, reticulate rugose on posterior third; notaulices incomplete, reaching almost posterior margin of scutum; scutellum dull,

granulated;

metanotum with

propodeum reticulate rugose, surface;

fore-wing

without

with 3 brown

longitudinal

short keels;

longitudinal keels on posterior

transversal

bands and with distal

part of

radial vein much longer than proximal part; segment 1 of front tarsus longer than segment 4 (14:10); segment 3 of front tarsus produced into a hook;

enlarged claw

segment 5 of front

(Fig.

tarsus

a group of 12 lamellae; MALE: unknown LOCUS TYPICUS: in AD;

tibial

Acacia Ridge

TYPICAL MATERIAL: F!

606) (Fig.

606)

tooth and

10 lamellae;

with 2 rows of 17 lamellae;

apex

with

spurs 1,1,2. (Australia)

holotype F!

2 paratypes FF!

with subapical

in QU;

in UQ;

1 paratype F!

1 paratype F!

in BR;

in OL.

1 paratype

899

Dryininae

Fig. 607 -

Chela of Dryinus glaber n.

sp.

(type)

HOSTS: unknown DISTRIBUTION: Australian region: AUSTRALIA: Acacia Ridge (Brisbane, Queensland), UQ! QU! Port Douglas (NE Queensland), BR! Brisbane (Queensland), AD! Blunder Cr. (Brisbane, Queensland), Ol! Eurimbula Ck. (via Round Hill, Queensland), UQ! NOTE: the species is named in honor of Mr.

E.C.

Dahms.

900

Mem.

Amer.

En!.

Ins!.,

no.

37,

1984

20. Dryinus giaber n. sp.

FEMALE: vertex

fully

winged;

length 4,0 mm;

(more along orbits),

testaceous;

head black,

with

front part of

clypeus and mandibles yellow;

pronotum black,

with

posterior collar yellow;

meta thorax and propodeum black;

abdomen

brown;

antennae

mesothorax,

legs yellow,

with clubs

of femora and hind coxae brown; head shiny, flat, punctate; occipital carina complete; OPL shorter than length of ocellar triangle; pronotum crossed by 2 transversal posterior impression); strong

longitudinal

reaching keels;

tegulae;

impressions

keels on

fully

fore-wing

with 2 brown

pronotal

sculptured by weakly

transversal

longitudinal

length of scutum;

punctate;

with parallel

with

tubercles not

numerous

to beyond 0,5

smooth,

anterior half,

weaker than

without sculpture,

impression;

extending

scutellum and metanotum shiny, surface;

smooth,

posterior

scutum shiny,

notaulices incomplete,

reticulate rugose on

(anterior impression

pronotum shiny,

propodeum

keels on posterior

bands and with

distal

part of

radial vein much longer than proximal part; segment 1 of front tarsus slightly longer than segment 4; enlarged claw (Fig. 607) with subapical and 8

tooth

segment 5 of front

tarsus

(Fig.

607)

19 lamellae; apex with a group of 13 lamellae; MALE: unknown

lamellae;

tibial

spurs 1,1,2.

Island,

New Guinea)

LOCUS TYPICU5: Bagiai Crater Trail (Kar Kar TYPICAL MATERIAL: holotype F! in B. HOSTS:

with 2 rows of

unknown

DISTRIBUTION: Kar Kar

Australian

Island),

region:

NEW GUINEA:

Bagiai

Crater Trail

(Kurum,

B!

21. Dryinus australianus n. sp.

FEMALE: and

fully

part I y

black;

winged:

length 4,00-7,50 mm;

c I ypeus redd i sh;

abdomen brown;

head black,

antennae testaceous;

legs brown,

with

tarsi

with mandibles

thorax and propodeum

and

tibiae

light;

head

shiny, punctate, flat; occipital carina complete; posterior ocelli almost touching occipital carina; UPL shorter than POL; OL ~ POL; temples very short;

pronotum shiny,

smooth,

without sculpture;

impression strong;

anterior transversal

long

pronotal

and distinct;

almost fully central

region

separated;

which

scutellum

reticulate rugose, wing

tubercles not reaching

sculptured by

with 3 brown

longitudinal

has no sculpture; shiny,

without

punctate;

longitudinal

transversal

posterior transversal

impression weak; keels,

posterior collar

tegulae;

scutum shiny,

except for anterior half of

notaul ices complete, metanotum

keels on

rugose;

psoteriorly

propodeurn

posterior surface;

bands and with distal

fore-

part of radial

vein

Dryininae

Fig. 608 - Chela of Dryinus australianus n. sp. (type)

901

902

Fig.

Mem. Amer.

609 -

Ent.

Ins!.,

no.

Chela of Dryinus venator (R.C.L. (Queensland)

37,

1984

Perkins) from Bundaberg

Dryininae

much

longer than proximal

segment 4

(23:20);

enlarged claw

(Fig.

608)

(Fig.

of 17 lamellae;

tibial

fully

yellow;

POL;

tooth and

with 2 rows of 32

longer than

13 lamellae;

lamellae;

segment 5

apex with a group

spurs 1,1,2. length 2,50-3,56 mm;

thorax and propodeum black,

incomplete;

tarsus

tarsus produced into a hook;

with subapical

608)

winged;

coxae partly black; than

segment 1 of front

segment 3 of front

of front tarsus MALE:

part;

903

head shiny,

posterior ocell i

head black,

abdomen brown;

reticulate rugose;

occipital

almost touching occipital

temples invisible;

scutum dull,

hairy,

with mandibles legs yellow,

with

carina

carina;

OL shorter

reticulate rugose;

notaulices incomplete, reaching approximately 0,65 length of scutum; scutellum and metanotum punctate; propodeum reticulate rugose, without longitudinal bands,

keels on posterior surface;

with distal

tibial

part of radial

longer than proximal

part;

spurs 1,1,2 .•

LOCUS TYPICUS:

Brisbane

(Australia)

TYPICAL MATERIAL:

holotype F!

paratype F!

4 paratypes

in UQ;

and 9 paratypes (2 FF, (1

HOSTS: in Australia Flatidae. DISTRIBUTION: Australian region: BM!

fore-wing without dark transversal

vein much

Lockerbie (Cape York,

F,

3 MM)!

AUSTRALIA:

N Queensland),

7 IvIM)!

in BIYI;

1

in OL. Brisbane (Queensland),

OL!

UQ!

22. Dryinus venator (R.C.L. Perkins) n. comb.

Paradryinus vena tor R. C. L. Perk i ns 1905: 55. Paradryinus threnodes R. C. L. Perk i ns 1905: 55 (n. syn.); locus typicus Bundaberg (Austra I i a) • Paradryinus varipes R.C.L. Perkins 1905: 56 (n. syn.); locus typicus lvIulgrave (Australia).

FEMALE: vertex,

fully

winged;

length 4,20-6,22 mm;

clypeus and mandibles yellow,

or 5-10 brown;

propectus black;

pronotum black,

sides and posterior coil ar redd i sh; black

with sides reddish;

posterior margins reddish; posterior th ird brown; reddish, rugose;

scutum black;

metanotum black;

scutellum black,

2 transversal

weaker than posterior impress ion); around disc;

pronotal

head dull,

length of ocellar

with segment 3 much impressions

pronotum du II,

longitudinal

keels;

legs

granulated and longer than

(anterior impression with numerous keel s

tubercles not reaching tegulae;

sculptured by numerous

sometimes with

abdomen brown-redd i sh;

OPL shorter than

antennae shorter than body, pronotum with

with anterior margin,

occasionally pronotum with anterior and

occipital carina complete;

segment 4;

with front part of with segments 3-10

somet i mes pronotum fu II y redd i sh or

with mid and hind coxae brown;

triangle;

head black,

antennae yellow,

scutum dull,

notaulices incomplete,

fully extending

904

fv',em.

to beyond 0,5 rugose;

Amer.

En!.

length of scutum;

Ins!.,

posterior surface;

1984

scutellum reticulate rugose;

propodeum reticulate rugose,

without

fore-w i ng with 3 brown

of rad i a I vei n much

37,

no.

longer than

longitudinal

metanotum

keels on

transversa I bands;

d i sta I part

segment 1 of front

prox i ma I part;

tarsus

(14:11); enlarged claw with subapical tooth and 14 lamellae (Fig. 609); segment 5 of front tarsus (Fig. 609) with 2 rows of 33 lamellae; apex with a group of 22 lamellae; tibial spurs 1,1,2. MALE: fully winged; length 2,56-3,29 mm; head black, with mandibles yellow; antennae yellow-brown or brown with segments 1-2 yellow; abdomen slightly

reddish-brown; carina

segment 4

longer than

legs yellow;

incomplete;

invisible;

head dull,

OPL shorter than

rugose and granulated;

scutum as sculptured as head,

notaulices incomplete,

and metanotum as sculptured as scutum; without

longitudinal

transversal

bands,

part; tibial spurs 1,1,2. LOCUS TYPICUS: Bundaberg

short keels

temples less visible;

length of scutum;

scutellum

propodeum reticulate rugose,

keels on posterior surface; with distal

TYPICAL MATERIAL:

but with

to beyond 0,5

extending

occipital

length of ocellar triangle;

part of radial

fore-wing

vein

without dark

longer than proximal

(Australia) and 3 paralectotypes FF!

lectotype F!

(designated by

venator in B; lectotype F! and 1 para lectotype M! (designated by Olmi 1982) of P. varipes in B; lectotype F! and 3 paralectotypes (1 F, 2 MM)! (designated by Olmi 1982) of P. threnodes in

Olmi

1982) of P.

B. in Austra I i a Privesa sp. (Ricaniidae), Massila sp. (Flatidae), 5iphanta sp. (Flatidae), Colgar sp. (Platidae) (R.C.L. Perkins 1906c), Privesa aphrophoroides Wa Ik. (Thompson 1954). HOSTS:

DISTRIBUTION: B!

National

Australian

Park

region:

(Queensland),

AUSTRALIA:

QU!

Bundaberg

(Queensland),

Mulgrave (Queensland),

B!

(Queensland), B! Blackall Ranges (Queensland), AD! Kurnell Wales), RY! Berowra Waters (Sydney, New South Wales), UQ! (New South Wales), (Port Glasgow),

TW!

NEW GUINEA:

Wau

(Morobe Distr.),

OL!

Cairns

(New South Blue Mts. B!

Mamai

Pltn.

B!

23. Dryinus papuanus n. sp.

FEMALE:

fully

winged;

length 5,00-6,00 mm;

sometimes clypeus reddish; testaceous; black;

sometimes only

abdomen brown;

antennae brown,

segment 1 testaceous;

legs brown,

with

tarsi

femora reddish; head (Fig. 598 A) shiny, occipital carina complete; posterior ocell i pronotum humped, weaker than

with 2 transversal

posterior impression);

keels on disc;

pronotal

sculptured by numerous

black,

thorax and propodeum

and sometimes clubs of

smooth, weakly punctate; touching occipital carina;

impressions

pronotum dull,

tubercles not reaching longitudinal

keels;

with mandibles and

with segments 1-2 and 9-10

(anterior hairy,

tegulae;

impression with numerous

scutum dull,

notaulices incomplete,

fully

weakly

Oryininae

905

Fig. 610 - Chela of Dryinus papuanus n. sp.

visible,

reaching

approximately 0,5

numerous longitudinal keels,

dull;

propodeum reticulate rugose on posterior surface; distal

length of scutum; metanotum dull,

anterior surface,

fore-wing with 3 brown

part of radial

vein

much

scutellum with

without sculpture;

with parallel

transversal

longer than

(type)

proximal

keels on

bands and with part;

segment

1 of

906

Mem.

Amer.

En!.

Ins!.,

no.

37,1984

front tarsus sl ightly longer than segment 4; enlarged claw (Fig. 610) with subapical tooth and 12 lamellae; segment 5 of front tarsus (Fig. 610)

with

two rows of 33

tibial

spurs 1,1,2.

MALE:

unknown

LOCUS TYPICUS:

lamellae;

Oaradae Pl'n

TYPICAL MATERIAL:

apex with

(Papua,

holotype F!

a group of 19 lamellae;

New Guinea)

and 1 paratype F!

in B;

1 para type F!

in OL. HOSTS:

unknown

DISTRIBUTION:

Australian

Moresby, Papua), Irian), B!

Fig.

611

B!

-

Wau

region:

NEW GUINEA:

(Morobe Distr.),

Fernale of

OL!

Daradae Pl'n

(Port

Kassam

(Kainantu,

f2ichardsidryinus corsicus

(Iviarshall)

Western

Dryininae

907

2. Richardsidryinus Moczar

Richardsidryinus Moczar 1965: 376. Avodryinus Ponomarenko 1981a: 143 (n. syn.); type species Avodryinus canadensis Ponomarenko 1981a, monotypic and orig. desig.

TYPE SPECIES: Richardsidryinus corsicus (Marshall 1874), orig. desig. FEMALE (Figs 611, 612): fully winged; maxillary palpi with 6 segments; labial palpi

with 3 segments;

shorter than front

tibia,

tooth and some lamellae;

enlarged claw

much

notaulices at

tubercles not reaching tegulae; 1,1,1. MALE (Figs 613,614): labial

palpi

fully

HOSTS:

Palaearctic,

least partly

as long as or

with one subapical visible;

maxillary palpi

pronotal

tibial

spurs

with 6 segments;

other generic morphologic characteristics:

see description of the subfamily; DISTRIBUTION:

617)

occipital carina complete;

winged;

with 3 segments;

(Fig.

longer than arolium,

tibial

Ethiopian,

spurs 1,1,2. Oriental,

Nearctic,

Australian

Lophopidae, Issidae, Delphacidae, Tropiduchidae

SPECIES:

12

Fig.

612 -

Female of

Richardsidryinus corsicus (Marshall)

908

Mem.

Fig.

613 -

Amero

En!.

Ins!.,

no.

37,

1984

Richardsidryinus corsicus (Marshall)

Male of

RICHARDSIDRYINUS

PALAEARCTIC SPECIES OF

Three species are known.

KEY TO THE SPECIES

FEMALES

Lamellae along

5 of

segment

front

tarsus nearly

same

length

(Fig. 615); segment 1 of front tarsus nearly as long as or slightly shorter than segment 4; occasionally slightly longer ••

-

0

0

•

•

•

•

Segment numerous

•

•

•

•

•

5 of

•

•

•

shorter

tarsus nearly

•

•

front

•

•

•

•

•

•

•

•

•

•

•

•

•

tarsus with

lamellae

twice as

(Figs

long as

•

•

•

•

•

••

616,

albrechti n. sp.

1.

some very

long

617);

segment

lamellae among

segment

1 of

front

4 ....................... 2

Dryininae

Fig.

2 Head

fully

614 -

909

Male of Richardsidryinus corsicus

reticulate

rugose;

head,

prothorax and mesothorax

mostly black . . . . . . . . . . . . . . . . . . . . . . . . . . -

Head

fully

granulated,

and mesothorax mostly

not

(Marshall)

reticulate

3. corsicus (Marshall) rugose;

reddish . . . . . . . . .

head,

prothorax

2. maroccanus n.

sp.

MALES

Only

the male of R.

corsicus is known.

1. Richardsidryinus albrechti n. sp.

FEMALE:

fully

testaceous;

winged;

length 3,62-4,00 mm;

antennae brown,

segment 10 white; testaceous-dark;

with

black;

clypeus and mandibles

segments 1-4 testaceous or dark and

margins and posterior collar of pronotum reddish; antennae distally thickened;

antennal

legs

segments in

following proportions: 10: 6: 24: 13: 10: 7: 6: 6: 5: 7; head dull, fully granulated; frons with a median keel, without lateral keels; POL: 4; OL :

3;

TL :

transversal shiny,

6;

occipital carina complete;

pronotum with a

impression and a strong posterior transversal

partly

smooth and without sculpture,

partly

collar of pronotum nearly as long as pronotal hairy, margin;

granulated,

weak anterior impression,

granulated;

tubercles;

posterior

ecutum dull,

with numerous short keels and areolae near posterior

notaulices incomplete,

almost reaching posterior margin of scutum;

910

Mem.

Fig.

615 -

Amer.

Chel a of

weakly

bands on

longitudinal

the basal

vein much

keels;

fore-wing

cells and beneath 18:

4:

6:

propodeum reticulate

transverse I y

stri ate;

16:

25;

posterior

with 3 dark transversal

the pterostigma;

distal

part of

(10:S); fore tarsal segments

longer than proximal part

in following proportions:

1984

no. 37,

granulated;

wi th posterior surface strong I y

surface without radial

Ins!.,

Richardsidryinus albrechti n. sp. (type)

scutellum and metanotum dull, rugose,

En!.

enlarged claw

(Fig.

615)

with

a subapical tooth and 11 lamellae; segment 5 of front tarsus (Fig. 615) with 2 rows of 22 lamellae; apex with a group of nearly 13 lamellae; segment 3 of front MALE:

unknown

tarsus produced into a hook;

(see ma Ie of

LOCUS TYPICUS:

Orotava

TYPICAL MATERIAL: F!

tibial

spurs 1,1,1.

Dryinus canariensis Ceba Ilos)

(Canary

holotype F

!

Islands in HE;

1 paratype F!

in MD;

1 paratype

in OL

HOSTS:

unknown

DISTRIBUTION:

Palaearctic region:

HE! EI Risco de Famara Fuerteventura), MD! NOTE:

CANARY

(Lanzarote),

OL!

ISLANDS:

Orotava

(Tenerife),

Las Penilas (Valle Granadillo,

the species is named in honor of Mr.

Anders Albrecht.

2. Nichardsidryinus maroccanus n. sp.

FEMALE: fully winged; length 3,25-3,43 mm; head, antennae, propectus, pronotum, scutum, scutellum, mesosternum, metasternum, mesopleura and metapleura reddish;

metanotum and propodeum black;

reddish; legs testaceous; complete; posterior ocelli OL;

OL shorter than POL;

abdomen brown-

head swollen, shiny, granulated, occipital carina not touching occipital carina; OPL shorter than temples prominent;

pronotum shiny,

granulated,

Dryininae

Fig.

hairy;

616 -

Chela of

anterior transversal

posterior transversal

Richardsidryinus maroccanus n. sp. (type)

impression and anterior collar very

impression strong;

tubercles not reaching tegulae; areolae near parapsidal approximately 0,25

911

scutum shiny,

furrows;

granulated,

notaulices incomplete,

length of scutum;

weak;

posterior collar distinct;

scutellum shiny,

pronotal

with some weak

reaching granulated;

metanotum reticulate rugose; propodeum reticulate rugose, with 2 short longitudinal keels on posterior surface; fore-wing with 2 brown transversal bands and with distal

part of radial vein much longer than proximal part (14:6); radial cell open; segment 1 of front tarsus nearly twice as long as segment 4; segment 3 of front tarsus produced into a hook; enlarged claw (Fig. 616) spatulate, with subapical tooth and 14 lamellae; segment 5 of front

tarsus

(Fig.

616)

with 2 rows of 8

extending continuously to the apex; MALE: unknown LOCUS TYPICUS: Tangeri (Morocco) TYPICAL MATERIAL:

holotype F!

tibial

(long) + 38 (short)

lamellae

spurs 1,1,1.

and 1 paratype F!

in GV;

1 para type F!

in OL. HOSTS:

unknown

DISTRIBUTION: GV!

OL!

Palaearctic region:

MOROCCO:

Tangeri,

GV!

ALGERIA:

Oran,

912

Fig.

Mem. Amer.

617 -

En!.

Ins!.,

no.

Chel a of Richardsidryinus corsicus (France)

37,

1984

(Marsha II) from seri gnan

Dryininae

Fig.

618 -

913

Male genital ia of Richardsidryinus corsicus seri gnan (France)

(Marshall) from

3. Richardsidryinus corsicus (Marsha II)

Dryinus corsicus Marsha II 1874: 207. Mesodryinus corsicus (Marshall): Kieffer 1907b: 10. Mesodryinus escorialensis Ceballos 1927: 102 (n. syn.); locus typicus EI Escorial (Spain). Mesodryinus corsicus (Ceballos): Berland 1928: 142. Richardsidryinus corsicus (Marsha II): Moczar 1965: 377. Richardsidryinus corsicus (Marshall): Abdul-Nour 1976: 276. Richardsidryinus corsicus (Marsha II): Currado and 01 mi 1979b: 5.

FEMALE (Figs 611, 612): fully winged; length 4,4-6,0 mm; head black, with mandibles and partly clypeus reddish; antennae reddish, with segments 5-9 brown;

propectus black;

pronotum black,

with anterior

914

Mem.

margin,

Amer.

En!.

Ins!.,

posterior collar and sides reddish;

propodeum black,

37,

no.

1984

mesothorax,

with apex of propodeum reddish;

metathorax and

petiole black;

abdomen

brown-redd i sh; legs testaceous, with t i b iae, femora, trochanters and coxae brown; head dull, granulated and reticulate rugose; occipital carina complete;

temples short,

region swollen;

distinct;

antennae shorter than

pronotum humped,

with 2 transversal

impression weaker than posterior impression);

body;

ocellar

impressions

pronotum shiny,

(anterior

granulated,

with anterior collar without sculpture and posterior impression with numerous

longitudinal keels;

scutum dull,

granulated,

pronotal

tubercles not reaching

incomplete and weakly visible;

scutellum shiny,

transversal

proximal

part;

without

metanotum rugose;

longitudinal

bands and with distal tarsus nearly

produced into a hook;

keels;

part of radial

fore trochanters nearly twice as

segment 1 of front

fore-wing vein

(10:5);

long as broad

enlarged claw spatulate,

with 2

longer than

long as segment 4;

twice as

notaulices

with anterior half

granulated and posterior half without sculpture; propodeum reticulate rugose, brown

tegulae;

with posterior half reticulate rugose;

segment 3

with subapical

tooth

and

5-9 very long lamellae (Fig. 617); segment 5 of front tarsus (Fig. 617) with 3 rows of lamellae extending continuously to the apex; two rows are composed of 26-27 I ameli ae per row; one row is composed of 5 very long lamellae and 5 bristles; tibial spurs 1,1,1. MALE (Figs 613, 614): fully winged; length 2,37-3,20 mm; black, with teeth of mandibles reddish, fully

reticulate rugose;

front

of

ocelli;

the

imaginary

occipital

straight

carina

sculptured as scutum; fore-wing

radial

vein

618);

head dull,

anterior margins of

hairy,

invisible,

in

posterior

incomplete,

scutellum and metanotum as

propodeum reticulate rugose,

without dark transversal

(Fig.

ligth;

anterior ocellus located

notaulices almost

length of scutum;

bands,

without

longitudinal

with distal

part of

(8:5); distivolsella located near tibial spurs 1,1,2.

longer than proximal

apex of penis

tarsi

line joining

incomplete;

reaching approximately 0,5 keels;

tegulae yellow,

ocellar region swollen;

part

lOCUS TYPICUS: Campoloro (Corse, France) TYPICAL MATERIAL: holotype F! of D. corsicus in BU;

holotype F!

of

M.

escorialensis in MD. HOSTS: in France Hysteropterum flavescens (01 i v. ) (=grylloides F.). DISTRIBUTION: 5erignan (Herault), Corse),

Palaearctic region:

(Vaucluse), BU!

P!

P!

HUNGARY:

Kapasvar,

la Figarede (Herault),

P!

Aix-en-Provence (Bouches-du-Rhone),

SPAIN:

Palamos

(Barcelona),

SZ!

BU!

FRANCE:

S!. Gely-du-Fesc BM!

Manzarron

Campoloro (Ajaccio, (Murcia),

BM!

Sierra de Espuna (Totana, Murcia), BM! Sierra Nevada, Ol! OT! EI Escorial (Madrid), MD! Cubillas (Granada), Ol! BM! Nerja (Granada), ITALY: Opicina (Trieste), (Zangheri 1969); GREECE:

EB! M. del Casino (Tossignano, Emilia) Ilia (Olympia), BM! CYPRUS: Limassol, BM!

ETHIOPIAN SPECIES OF

RICHARDSIDRYINUS

BM!

Dryininae

Fig.

619 -

915

Heads of females of Richardsidryinus erraticus

(Turner) (A)

and

cariniceps (Cameron) (B)

Three species are known.

KEY TO THE SPECIES

FEMALES

Head wi th posterior margin

of

vertex

concave

(Fig.

619 A);

posterior collar of pronotum short, nearly as long as ocellar triangle (Fig. 620 B); disc of pronotum weakly

humped

(Fig. 620 B); scutellum nearly twice as long as metanotum; propodeum short (Fig 621 B) ••.•• 1. erl"aticus (Turner) -

Head wi th posterior margin posterior collar of (Fig.

620 A);

of

pronotum

di sc of

vertex long;,

convex longer

pronotum very humped

(Fig. than (Fig.

619 B);

ocellar

triangle

620 A);

scutellum slightly longer than metanotum; propodeum long (Fig. 621 A) •••.••.••••••.•.•..•.•••.•.••••.•.••.•••.•••••••••••••••• 2 2 Head, propectus and pronotum fully ferruginous-testaceous ••••.•• • . • • . • • • . • • . • . • • • • . • • • . • • . • • • • • • 2. cariniceps (Came ron) - Head, propectus and pronotum mostly black .•• •••.•.•.••.•• ••••••• •••.•.••••••.•••••••••••••••••• • 3. undulatus (Benoi t)

i I

MALES

916

Mem.

Amer.

Ent.

Ins!.,

no.

37,

1984

A

Fig.

620 -

and

Only

Richardsidryinus cariniceps (Cameron) (A) erraticus (Turner) (B)

Pronotum of fema les of

R. erraticus is known.

the male of

1. Richardsidryinus erraticus (Turner) n. comb.

Neodryinus erraticus Turner 1928: 149. Neodryinus erraticus (Turner): Benoit 1951b: 157. Dryinus erraticus (Turner): Carneg ie 1975: 249.

FEMALE fully winged; length 5,0-6,0 mm; head black, with clypeus and mandibles yellow; antennae testaceous, with segments 7-9 weakly darkened; propectus black; collar reddish; black;

pronotum black,

mesothorax,

abdomen brown;

with

anterior margin,

sides and posterior

metathorax and propodeum black;

legs testaceous-reddish,

petiole

sometimes with coxae

darkened; head (Fig. 619 A) shiny, granulated, sometimes with some longitudinal keels on frons; occipital carina complete; pronotum (Fig. B)

shiny,

granulated,

with

longitudinal

keels on

transversa I impression;

pronotum with

impression

anterior transversal

and

scutum dull, notaulices

a

weak

as sculptured as head,

incomplete,

scutellum dull,

reaching

granulated;

620

sides and on posterior

a posterior strong transversa I impression

weakly

visible;

with posterior half reticulate rugose;

approximately 0,5 length of scutum;

metanotum rugose;

propodeum

(Fig.

621

B)

Dryininae

917

A

B

.~ Fig.

621

-

Propodeum of females of Richardsidryinus cariniceps (Cameron) (A) and erraticus (Turner) (B)

reticulate rugose, transversal proximal

without

longitudinal keels;

bands and with distal

part;

rarely

transversal

fore-wing

part of radial

vein

with 3 brown longer than

bands weakly visible;

segment

1 of front

tarsus nearly as long as segment 4; segment 3 of front tarsus produced into a hook; enlarged claw (Fig. 622) with subapical tooth and 6 lamellae;

segment 5 of front

tarsus

apex with a group of 16 lamellae; MALE: yellow;

fully

winged;

(Fig. tibial

length 2,18-2,75 mm;

antennae yellow-brown;

thorax

622)

with 2 rows of 24 lamellae;

spurs 1,1,1. head black,

with mandibles

and propodeum black;

abdomen

brown; legs testaceous, with hind coxae partly brown; head dull, granulated; occipital carina incomplete; temples invisible; posterior ocell i almost touching occipital carina; OL shorter than POL; scutum dull, granulated, with weak keels near posterior margin; notaulices incomplete, reaching approximately 0,65 length of scutum; granulated;

propodeum reticulate rugose,

posterior surface; distal

scutellum and metanotum

without

longitudinal keels on

fore wings without dark transversal

part of radial

vein

longer than

proximal

part

bands and with (12:9);

ninth

abdominal sternite: fig. 623; tibial spurs 1,1,2. LOCUS TYPICUS: Mossel Bay (South Africa) TYPICAL MATERIAL: holotype F! and 2 paratypes FF! in BM. HOSTS: in South Africa Numicia viridis Muir (Tropiduchidae) (Carnegie 1975) • DISTRIBUTION:

Ethiopian region:

Edgecombe (Natal), (Swaziland),

OL!

ED!

OL!

Warmbad

ED!

Mt.

Mossel

Bay

(Cape Province),

BM!

(Transvaal),

WA!

ZIMBABWE:

BM!

SOUTH AFRICA:

Pongola,

Big Bend

Chishawasha

918

Mem.

Fig. 622 -

Amer. Ent.

Inst.,

no.

37,

1984

Chela of Richardsidryinus erraticus

(Turner)

(Salisbury), BM! ANGOLA: R. Giraul (Mocamedes), BM! ZAIRE: (Albert Nat. Park), TV! SOMALIA: Afgoi (Benadir), Ol!

Mt. Hoyo

2. Richardsidryinus cariniceps (Cameron) n. comb.

Dryinus Carln1CepS Cameron 1906: 158. Lestodryinus cariniceps (Cameron): Turner 1928: 147.

FEMALE: fully winged; length 5,62 mm; head ferruginous-testaceous; antennae brown, with segments 9-10 yellow, distal part of segment 4 and proximal part of segment 5 white; propectus and pronotum ferruginoustestaceous; mesothorax, meta thorax and propodeum ferrug i nous or redd i shbrown; sometimes metanotum and propodeum black; petiole black; abdomen reddish-brown; legs reddish-brown; head (Fig. 619 B) flat, granulated, weak I y rugose; posterior margi n of vertex convex; occi pita I cari na complete, but weakly visible on sides of occiput; OPl slightly shorter than Ol; Ol ~ POL; pronotum (Fig. 620 A) dull, granulated, with weak keels around disc; posterior transversal impression very strong; anterior transversal impression invisible; posterior collar long, longer than length of ocellar region; pronotal tubercles not reaching tegulae; scutum dull, reticulate rugose; notaul ices weakly visible, incomplete, reaching approximately 0,5 length of scutum; scutellum dull, reticulate rugose,

Dryininae

Fig. 623 -

slightly

919

Ninth abdominal sternite of male of Richardsidryinus erraticus (Turner) from Mt. Edgecombe (South Africa)

longer than metanotum;

metanotum dull,

punctate;

propodeum

(Fig.

621 A) reticulate rugose, without longitudinal keels on posterior surface; fore-wing with 2 brown transversal bands and with distal part of radial vein much longer than proximal part; segment 1 of front tarsus longer than segment 4 (15:12); segment 3 of front tarsus produced into a hook; enlarged claw (Fig. 624) with subapical tooth and 11 lamellae; segment 5 of front tarsus (Fig. 624) with 2 rows of 37 of 16 lamellae; tibial spurs 1,1,1. MALE: unknown

lamellae;

apex with a group

LOCUS TYPICUS: Port SI. Johns (South Africa) TYPICAL MATERIAL: holotype F! in BM. HOSTS: unknown DISTRIBUTION: Ethiopian region: SOUTH AFRICA: Port SI. Johns (Pondoland), BM! TV! ZAIRE: Garamba Nat. Park, TV!

3. Richardsidryinus undulatus (Benoit) n. comb.

Lestodryinus undulatus Benoit 1950: 226.

FEMALE: fully winged; length 6,25-7,50 mm; head black, with front part of vertex (more along orbits), clypeus and mandibles yellow; antennae testaceous, with dorsal side of segments 1-2, segments 3 and 6-8 brown; propectus black; pronotum black, with margins weakly reddish; mesothorax and propodeum black; abdomen brown; legs brown-black, with tarsi light; head dull,

granulated,

with numerous weak

longitudinal keels on frons

920

Fig.

Mem.

624 -

Chela of

Amer.

En!.

Inst.,

no.

37,

1984

Richardsidryinus cariniceps (Cameron) from Port S!. johns (South Africa)

and vertex;

occipital

carina complete;

posterior ocelli

placed behind the

imaginary straight line joining posterior edges of eyes, not touching occipital carina; OPL shorter than OL; OL = POL; TL more than twice as long as POL;

pronotum dull, granulated, with numerous keels around disc; impression very weak; anterior collar invisible;

anterior transversal

posterior transversal

impression strong;

tubercles not reaching tegulae; longitudinal

weak

keels;

posterior collar long;

scutum dull,

granulated,

notaulices incomplete,

pronotal

with numerous

reaching almost posterior

Dryininae

Fig. 625 -

Chela of

Richardsidryinus undulatus (Benoit)(type)

921

922

Mem.

Amer.

Ent.

Inst.,

no.

37,

1984

margin of scutum; scutellum as sculptured as scutum; metanotum rugose; propodeum reticulate rugose, without longitudinal keels on posterior surface; radial

fore-wing with 2 brown

vein much

tarsus slightly

transversal

longer than proximal segment 4

longer than

part

bands and with distal (10:4);

part of

segment 1 of front

segment 3 of front tarsus 625) with subapical tooth and 9 lamellae; segment 5 of front tarsus (Fig. 625) with 2 rows of 39 lamellae; apex with a group of at least 60 lamellae; tibial spurs 1,1,l. MALE: unknown LOCUS TYPICUS: Nialopul (Zaire) TYPICAL MATERIAL: holotype F! in TV. HOSTS: unknown DISTRIBUTION: Ethiopian region: ZAIRE: Nialopul (Kivu), TV! UGANDA: Kampala, BM! BURUNDI: Bururi, CAl produced into a hook;

enlarged claw

(15:13);

(Fig.

OR lENT AL SPEC I ES OF

RICHARDSIlJRYINUS

Two species are known.

KEY TO THE SPECIES

FEMALES

Temples the

shorter,

external

0,5 times as long as distance between

nearly

margins of

posterior ocell i . . . . . . . . . . . . . . . . . . . . . . .

. • . . . • • . . . . . . . . • . . . . . . . . . . . . . . . . 1. indianus n. -

sp.

Temples longer, nearly as long as distance between the external margins of posterior ocell i . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2. pyrillae (Kieffer)

MALES

Only the male of

R. pyrillae is known.

1. Richardsidryinus indianus n. sp.

FEMALE:

fully

winged;

length 4,81

clypeus and mandibles yellow;

mm;

head black,

antennae with

with malar space,

segments 1-3 testaceous

Dryininae

Fig. 626 -

Chela of

(other segments missing dorsal

Richardsidryinus indianus n. sp. (type)

in only

sur-face testaceous;

known specimen);

pronotum black,

collar reddish;

mesothorax,

brown-reddish;

legs testaceous;

propectus black,

head dull,

the external

with

with margins and posterior

metathorax and propodE'um black;

longitudinal keels on posterior surface; long as distance between

923

granulated,

temples short,

abdomen

with some nearly 0,5 times as

margins of posterior ocell i; occipital

924

Mem.

Fig.

627 -

posterior ocell i

pronotum dull,

disc very humped; weak;

granulated,

Ins!.,

pronotal

almost

no.

37,

1984

touching occipital

from Lyallpur

carina;

OL =

with numerous weak keels around disc;

anterior transversal

posterior transversal

distinct;

En!.

Chela of Richardsidryinus pyrillae (Kieffer) (I ndia)

carina complete; POL;

Amer.

impression and anterior collar very

impression strong;

tubercles not reaching

posterior collar short,

tegulae;

scutum dull,

but

granulated,

with numerous irregular short keels on lateral regions; notaulices incomplete, almost reaching posterior margin of scutum; scutellum dull, granulated;

metanotum rugose;

longitudinal

keels on posterior surface;

bands and with distal

propodeum reticulate rugose,

part of radial

without

fore-wing with 3 brown transversal

vein much

longer than proximal

Dryininae

part;

segment

produced 12

lamellae;

lamellae; MALE:

1 of front

into a hook;

tarsus nearly

enlarged claw

segment 5 of front

apex with

925

as

long as segment 4;

(Fig. 626)

tarsus

(Fig.

a group of 16 lamellae;

segment 3

with subapical

626)

tooth and

with 2 rows of 29

tibial

spurs 1,1,1.

unknown

LOCUS TYPICUS:

Padegaon

TYPICAL MATERIAL: HOSTS:

(India)

holotype F!

in BM.

unknown

DISTRIBUTION:

Oriental

region:

INDIA:

Padegaon,

BM!

2. Richardsidryinus pyrillae (K ieffer) n. comb.

Lestodryinus pyrillae Kieffer 1911a: 108. Lestodryinus pyrillae Kieffer: Pruth i and Man i 1942: 423.

FEMALE: vertex

fully

winged;

length 3,20-4,40 mm;

(more along orbits),

testaceous;

head black,

with front part of

clypeus and mandibles yellow;

propectus black,

pronotum black,

marg i ns and posterior coil ar yellow-redd ish;

with

lateral

mesothorax,

antennae and anterior

meta thorax and

propodeum black; abdomen brown-reddish; legs testaceous; head flat, granulated, with some longitudinal keels on frons and with short

dull,

transversal keels on the sides of ocellar region; occipital carina complete; posterior ocelli located near occipital carina; OPL shorter than POL; OL ~ POL;

TL nearly

between with

twice as

the external

some longitudinal

strong;

long as POL and nearly as

margins of posterior ocelli; keels on sides;

posterior collar

long and

long as distance

pronotum dull,

posterior transversal

visible;

granulated

impression

anterior transversal

impression

weak; pronotal tubercles not reaching tegulae; scutum dull, granulated, with some areolae near posterior margin; notaulices complete, posteriorly separated; scutellum dull, granulated; metanotum rugose; propodeum reticulate rugose, without longitudinal keels on posterior surface; forewing

with 3 brown

much

longer than proximal

segment 4 claw

(18: 15);

(Fig.

tarsus

627)

(Fig.

lamellae;

transversal

segment

segment 3 of fore tars i

with subapical

627)

tibial

bands and with distal

part;

tooth and

with 2 rows of 24

part of radial

vein

1 of front

tarsus

longer than

produced

into a

hook;

10

lamellae;

lamellae;

en I arged

segment 5 of front

apex with a

group of

15

spurs 1,1,1.

MALE: fully winged; length 2,00-2,18 mm; head black, with mandibles yellow; antennae testaceous; thorax and propodeum black; abdomen brown; legs yellow;

antennal

10:

10;

9:

9:

8:

dorsal part ocelli POL;

not

segments

head dull,

in following proportions:

granulated;

laterally not reaching eyes;

touching occipital

scutum hairy,

dull,

carina;

granulated,

occipital

4:

temples weakly

15:

visible;

UPL shorter than OL; with short

6:

carina complete,

10:

10:

with

posterior

OL shorter than

irregular keels;

926

Mem.

Fig.

628 -

Amer.

En!.

Inst.,

no.

37,

1984

Male genitalia of l?ichardsidryinus pyrillae (Kieffer)

from Pusa

(India)(right half removed)

notaul ices

incomplete,

reaching almost posterior margin of scutum;

scutellum and metanotum dull,

granulated;

propodeum reticulate rugose,

with two longitudinal keels on posterior surface; fore-wing without dark transversal bands, with distal part of radial vein nearly twice as long as proximal part; genitalia; fig. 628; tibial LOCUS TYPICUS: Pusa (Bihar, India) TYPICAL MATERIAL: based on typical HOSTS: Mani

in

lost

spurs 1,1,2.

(?)(the proposed description of R.

material,

but on

India Pyrilla sp.

(Lophopidae) (Kieffer 1911a, 1914b; Pruthi and

1942); Pyrilla perpusilla

(Wlk.)(Rahman

1954a); Pyrilla aberrans and pusan a (Kurian BIOLOGY: notes in Rahman and Nath (1940). NATURAL ENEMIES;

in Punjab

Cheiloneurus sp. (Encyrtidae)

and Nath 1940;

Kurian

1954a).

(India) the species is parasitized by (Rahman and Nath 1940).

DISTRIBUTION:

Oriental region:

(Punjab),

Pusa (Bihar),

NO!

pyrillae is not

topotypic material present in NO).

INDIA:

NO!

Lyallpur (Punjab),

Bengal

(Kurian 1954a);

BM! Delhi

Karnal (Punjab)

Dryininae

(Kurian

1954a); South

India

(Kurian

1954a).

RICHARDSIDRYINUS

NEARCT I C SPEC I ES OF

Three species,

927

including a fossil,

are known.

KEY TO THE SPECIES

FEMALES

R. canadensis)

(excluding the fossil

Scutum sculptured by numerous parallel -

longitudinal

keels . . . . . .

• • • • • • • • • • . . • • • • • • • • • . • • • • • • • • • • 1.

nearcticus n. sp.

Scutum without

granulated and partly

reticulate

longitudinal

rugose,

most lyon

keels,

lateral

. . . • • . . . . • • • • . • . . • • • . . . • . . . . • • • . 2.

areas • •••••••••••• ••••••••

wetmorei n. sp.

MALES

Unknown

NOTES: the foss i I R. canadensis is not presen t i n the key because the on I y known specimen is weakly visible through the amber, so that its morphology

isn't very distinct.

1. Richardsidryinus nearcticus n. sp.

FEMALE:

fully

winged;

clypeus testaceous;

length 4,37 mm;

antennae testaceous,

head black,

with mandibles and

with segments 4-10 brown;

thorax,

propodeum and abdomen black; legs testaceous, with clubs of femora brown and coxae black; tegulae testaceous; head dull, fully reticulate rugose; posterior ocell i TL much

almost touching occipital

longer than OPL;

with posterior coil ar very tegulae;

POL

occipital

carina complete;

pronotum dull,

granulated,

short and pronota I tubercles not reach i ng

anterior transversal

granulated,

carina;

longer than OL;

impression

with numerous longitudinal

scutellum and metanotum shiny,

very

weak;

keels;

granulated;

scutum dull,

notaulices invisible;

propodeum fully reticulate

928

Mem.

Fig.

629 -

Amer.

Ent.

Ins!.,

no.

37,

1984

Chel a of Richardsidryinus nearcticus n.

sp.

(type)

rugose, without longitudinal keels on posterior surface; fore-wing missing in holotype; fore tarsal segments 1-4 in following proportions: 17: 2,5: 4: 10; segment 5 of front tarsus (Fig. 629) with 2 rows of 8 (long) + 23 (short) lamellae; apex with a group of 4 lamellae; enlarged claw (Fig. 629) with a subapical tooth and a row of 12 lamellae; tibial spurs 1,1,1.

Dryininae

Fig.

MALE:

Chela of Richardsidryinus wetmorei n.

630 -

sp.

(type)

unknown

LOCUS TYPICUS:

Linares (Mexico)

TYPICAL MATERIAL: HOSTS:

929

holotype F!

in CA.

unknown

DISTRIBUTION:

Nearctic region:

MEXICO:

Linares (N.

Leon),

CAl

2. Richardsidryinus wetmorei n. sp.

FEMALE:

fully

winged;

mandibles testaceous;

length 4,37 mm;

thorax and propodeum black, pronotum; black;

antennae distally 13:

7:

21:

legs testaceous,

thickened; 12:

with clypeus and

with segments 3-10 darkened;

with reddish nuances on posterior collar of

abdomen brown-reddish;

proportions:

head black,

antennae testaceous,

10:

9:

with hind coxae partly

antenna I segments in following 7:

6:

5:

7;

head dull,

fully

granulated

and with numerous areolae or irregular keels on lateral areas of frons and vertex; frons with a median keel; occipital carina complete; POL = 6; OL

=

3;

OOL

=

9;

=

OPL

1;

TL

=

4;

pronotum dull,

granulated,

with a

very weak anterior transversal impression and with a strong posterior transversal

impression;

than posterior lobes; granulated,

disc weakly

pronotal

humped;

with numerous areolae on

notaulices incomplete,

posterior collar short,

tubercles not reaching tegulae; lateral

shorter

scutum dull,

areas and on posterior half;

reaching approximately 0,65 length of scutum;

930

Mem.

Amer.

En!.

scutellum and metanotum weakly without keels;

fore-wing

Ins!.,

no.

granulated;

the pterostigma;

longer than proximal part

(11:6,5);

proportions:

23,5;

3:

5:

15:

fore tarsal

tarsus produced into a hook;

MALE:

tibial

one on

the basal vein

segments in following (Fig.

segment 5 of front

2 rows of 36 lamellae extending continuously front

bands,

distal part of radial

enlarged claw

tooth and a row of 15 lamellae;

1984

propodeum reticulate rugose,

with 2 dark transversal

cells and another beneath 20:

37,

630)

with

tarsus

a subapical

(Fig.

to the apex;

630)

with

segment 3 of

spurs 1,1,1.

unknown

LOCUS TYPICUS:

William

TYPICAL MATERIAL:

(Arizona,

holotype F!

HOSTS: unknown DISTRIBUTION: Nearctic region: NOTE:

the species is named

U.

S. A.)

in WA. U.

S.

A.:

William

in honor of Mr.

A.

(Ariz.),

WA!

Wetmore.

3. Richardsidryinus canadensis (Ponomarenko) n. comb.

Avodryinus canadensis Ponomarenko 1981 a: 143.

FEMALE: brown,

fully pa I p i

winged;

length 2,8 mm;

apparently black,

and antennae testaceous,

legs testaceous,

with abdomen

with femora part I y

black; antennae only partly visible, distally thickened; antennal hairs not visible; antennae about three times as long as head: 24: 8; dorsal side of head not visible; side of head; segments; visible; radial

temples distinct;

pronotal

hyaline,

mandibles not visible;

maxillary palpi

dorsal

without dark transversal part;

carina visible

labial

palpi

in

bands;

petiole short;

pterostigma

for

fore-wing

long and narrow;

with distal part approximately

forewing shape usual

trochanter more than five times as

as long as

Dryininae; fore legs slender, with

long as broad;

segment 1 of front

tarsus approximately as long as segment 4; enlarged claw with a subapical tooth and with a row of 17 lamellae; segment 5 of front with

two rows of nearly

segment 5 very slender, longer than arolium; 1,1,1; MALE:

18 (short) + 8 (very with an

apical

Canadian

TYPICAL MATERIAL: HOSTS: unknown DISTRIBUTION:

long)

lamella;

lamellae;

for

Nearctic region:

much

tibial

spurs

Richardsidryinus.

amber from Medicine Hat

holotype F!

tarsus

apex of

enlarged claw

rudimentary claw apparently present;

general morphology of body usual unknown

LOCUS TYPICUS:

ventral

with 3

side of thorax and propodeum not

tubercles not reaching tegulae;

vein curvilinear,

proximal

occipital

with 6 segments;

(Alta.,

Canada)

in OT. CANADA:

only known from

the typical

local ity. NOTES:

according to McAlpine and Martin

(1969)

the Canadian amber from

Dryininae

Fi g.

Medicine Hat

631

-

CheJ a of

931

Richardsidryinus trilineatus (Dodd) (type)

is retinite of Cretaceous Age.

AUSTRALIAN SPECIES OF

One species is known.

RICHARDSIDRYINUS

932

Mem.

Amer.

En!.

I nst.,

no.

37,

1984

Richardsidryinus trilineatus (Dodd) n. comb.

Neodryinus trilineatus Dodd 1914b: 129.

FEMALE:

fully

winged;

length 4,62 mm;

spot on frons and ocellar region; propodeum black, testaceous;

with margins of pronotum reddish;

legs testaceous,

distal

not distally thickened;

antennal

17:

14:

11:

9:

8:

7:

9;

posterior ocelli

thorax and

abdomen brown-

half of mid and hind tibiae brown;

head dull,

not

antennae

segments in following proportions:

sculptured by numerous longitudinal complete;

with a black central

with mid and hind coxae and mid and hind

clubs of femora brown; 40:

head reddish,

antennae testaceous;

excavated,

parallel

granulated,

keels;

touching occipital

10:

5:

with frons

occipital carina

carina;

POL = 3;

OL = 3;

OOL = 11; OPL = 1; TL = 7; pronotum du II, crossed by a posterior transversal impression; pronotal tubercles not reaching tegulae; posterior collar distinct;

scutum dull,

notaulices incomplete,

granulated and fully

scutellum and rnetanotum dull, fully

reticulate rugose;

reaching approximately 0,5 length of scutum;

reticulate rugose;

granulated;

propodeum dull,

posterior surface without

wing with 3 dark transversal

bands on

granulated and

longitudinal

keels;

the basa I cell s and beneath

forethe

pterostigma; distal part of radial vein longer than proximal part (12:6); fore tarsal segments in following proportions: 21: 3: 8: 16: 27; enlarged claw (Fig. 631) with a subapical tooth and 1 row of 10 lamellae; segment 5 of front tarsus (Fig. 631) with 2 rows of 28 lamellae; apex with a group of 15 lamellae; MALE:

tibial

spurs 1,1,1.

unknown

LOCUS TYPICUS:

Chindera

TYPICAL MATERIAL:

HOSTS: unknown DISTRIBUTION: Australian South Wales),

(Australia)

holotype F! region:

in AD. AUSTRALIA:

Chindera (Tweed River,

New

AD!

3. Tridryinus Kieffer

Tridryinus Kieffer 1913c: 325. Tridryinus Kieffer: Kieffer 1914b: 28.

TYPE SPECIES:

Tridryinus striaticeps (Kieffer 1909), orig. desig.

FEMALE (Figs 632, 633): fully winged; maxillary palpi with 6 segments; labial palpi with 3 segments; enlarged claw (Fig. 647) as long as front tibia or shorter, some lamellae;

much

longer than arolium,

notaulices inviSible;

pronotal

with one subapical

tooth

tubercles not reaching

and

933

Dryininae

Fig.

tegulae;

632 -

occipital

spurs 1,1,2. MALE (Figs 634,

Female of

carina complete or incomplete, 635):

fully

winged;

labial palpi with 3 segments; see family diagnosis. DISTRIBUTION: HOSTS:

Flatidae

SPECIES:

17

Tridryinus maximus

Ethiopian,

n.

sp.

(type)

sometimes absent;

maxillary palpi

tibial

with 6 segments;

other generic morphologic characteristics:

Nearctic,

Neotropic

934

Mem.

Fig.

633 -

Amer.

En!.

Ins!.,

no.

37,

1984

Female of Tridryinus maximus n. sp.

ETHIOPIAN SPECIES OF

(type)

TRIDRYINUS

Two species are known.

KEY TO THE SPECIES

FEMALES

Species almost

fully

black;

POL. • • • • • • • • • • • • • • • • • • • ••

head with OOl

1.

slightly

longer

than

ugandan us n. sp.

Species almost fully reddish-testaceous; head with OOl nearly two-three times as long as POL . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .

.........................

2.

ampuliciformis (Turner)

MALES

Dryininae

Fig.

634 -

Male of

935

Tridryinus poecilopterae (Richards)

Unknown

1. Tridryinus ugandanus n. sp.

FEMALE:

fully

wingedj

length 4,62-5,12 mmj

sometimes malar space testaceousj segments 1-2 testaceousj

legs testaceous,

fore trochanters partly brownj thickenedj

antenna I segments

blackj

antennae fully

mandibles,

clypeus and

testaceous or brown,

with coxae,

abdomen brown-testaceousj in following proportions:

with

clubs of femora and 13:

antennae distally 6:

28:

14:

10:

936

Mem.

Fig.

9: 6: 6: 6: 8; rugose; = 2;

635 -

Ent.

head dull, OPl = 3;

irregular striae,

Inst.,

no.

37,

1984

Male of Tridryinus poecilopterae (Richards)

weakly convex,

frons with a median keel;

OOl = 9;

humped;

Amer.

Tl = 4;

crossed by

posterior collar

granulated and fully

occipital

carina complete;

pronotum dull,

a transversal

invisible;

pronotal

granulated,

reticulate

POL = 4;

Ol

with weak

impression; disc weakly tubercles not reaching tegulae;

scutum, scutellum and metanotum dull, granulated and reticulate rugose; notaulices invisible; propodeum reticulate rugose, with 2 short incomplete longitudinal

keels on posterior surface;

transversal bands, pterostigma; distal fore tarsal claw

(Fig.

fore-wing with 2 very weak dark

one on the basal cells and another beneath the part of radial vein longer than proximal part (15:9);

segments in following proportions: 26: 4: 7: 11: 20; enlarged 636) with a subapical tooth and a row of 7 lamellae; segment 5

tarsus (Fig. 636) with a row of 11 lamellae; apex with a group of 9 lamellae; tibial spurs 1,1,2. MALE: unknown lOCUS TYPICUS: Kampala (Uganda) TYPICAL MATERIAL: holotype F! and 2 para types FF! in TW; 1 paratyPE'! F! in Ol. HOSTS: unk now n DISTRIBUTION: Ethiopian region: UGANDA: Kampala, TW! SOUTH AFRICA: Pietermaritzburg, DL! SIERRA LEONE: Freetown, TW! of front nearly

Dryininae

Fig.

937

Chela of Tridryinus ugandanus n. sp.

636 -

(type)

2. Tridryinus ampuliciformis (Turner) n. comb.

Lestodryinus ampuliciformis Turner 1928: 148. Mesodryinus ampuliciformis Benoit 1953a: 144.

FEMALE:

fully

winged;

length 5,12-6,56 mm;

reddish-testaceous,

ocellar region,

petiole,

tegulae black;

sometimes head almost fully black;

brown spot on propodeum;

antennae distally

segments in following proportions: dull,

with

region behind the tegulae and sometimes also

fully reticulate rugose,

9:

flat;

6:

29:

rarely

weakly 16:

14:

with median

thickened; 11: 9:

antenna I

8:

frons with a median keel;

6,5:

9;

head

occipital

carina complete; POL = 3,5; Ol = 2,5; OOl = 11; OPl = 2,5; Tl = 5; sometimes (in a specimen from Uganda) POL = 5 and Ol 9; posterior margin of vertex excavated; pronotum (Fig. 503 A) shiny, weakly rugose, with di sc weak I y humped and two transversa I impressions very weak; posterior collar very short; scutum dull,

metanotum dull,

without dark transversal vein

longer than

following proportions: subapical

bands, proximal

36:

4:

8:

scutellum and

propodeum reticulate rugose;

2 incomplete longitudinal fully 16:

distal

segments in

fore

tarsal

enlarged claw

segment 5 of front

(Fig.

a group of 10 lamellae;

Port SI.

TYPICAL MATERIAL:

Johns

holotype F!

(South Africa) and 1 paratype F!

in BM.

part of

637)

tarsus

unknown

LOCUS TYPICUS:

posterior

fore-wing

(1 :7);

26;

apex with

keels;

weakly darkened;

part

tooth and with 6 lamellae;

with 1 row of 12 lamellae; 1,1,2. MALE:

tubercles not reaching tegulae;

notaulices invisible;

reticulate rugose;

surface of propodeum with radial

pronotal

reticulate rugose;

with a

(Fig. tibial

637) spurs

938

Mem. Amer. Ent.

Fig. 637 -

Chela of

Ins!.,

no. 37,

1984

Tridryinus ampuliciformis {Turner}

Dryininae

939

\

Fig. 638 -

Chela of

Tridryinus serratus n. sp. (type)

HOSTS: unknown DISTRIBUTION: Ethiopian region: SOMALIA: Afgoi (Benadir), Ol! BOTSWANA: Gweta, BM! Kuke Pan, Ol! BM! SOUTH AFRICA: Port St. Johns (Pondoland),

940

Mem. Amer.

Fig.

BM!

639 -

Ent.

Inst.,

no.

37,

1984

Heads of females of Tridryinus oculatus n. sp. testaceus n. sp. from Sebring (Florida)(B)

Umtentweni

(Natal),

BM!

Sundays River (Addo Valley,

(type)(A)

and

Cape Province),

BM! SOUTH WEST AFRICA: Aus, Ol! BM! Kuiseb River Bed (Gobabeb), CO! ZAIRE: Zaziba (Upemba Nat. Park), TV! Garamba Nat. Park, TV! NIGERIA: Ikoyi Park (lagos), BM! KENYA: Karen (Nairobi), Ol! TW! UGANDA: Kampala, TW!

NEARCTIC SPECIES OF

TRIDRYINUS

Three species are known

KEY TO THE SPECIES

FEMALES

Head with Ol

shorter

than POl . . . . . . . . . . . . . 1.

- Head wi th Ol more than 2 Head almost flat (Fig. -

Head

strongly

excavated

serratus n. sp.

three times as long as POL •••••••••••••• 2 639 B) . . . . . . . . . . . . . 2. testaceus n. sp. (>'ig.

639 A) . . . . . . 3.

MALES

oculatus n. sp.

Dryininae

Only

the male of

941

Tr. testaceus is known.

1. Tridryinus serratus n. sp.

FEMALE:

fully

winged;

length 4,68 mm;

head ferruginous,

with brown spots

on ocellar region and on ventral side; antennae testaceous, with segments 5-10 brown; propectus and pronotum ferrug i nous-testaceous; mesothorax, metathoraw and propodeum black; testaceous;

legs testaceous,

abdomen brown,

1

with segment

with mid and hind tibiae fully or partly

brown and with stalks or also clubs of mid and hind femora brown;

head

dull, reticulate rugose, convex; occipital carina complete; temples short, distinct; posterior ocelli not touching occipital carina; OPL shorter than OL;

OL shorter than

striae around disc, collar weak; short,

POL;

pronotum shiny,

hairy;

posterior transversal

distinct;

pronotal

notaulices invisible;

reticulate rugose;

propodeum hairy,

keels on posterior surface; part of radial

vein

numerous

posterior collar

tegulae;

scutum fully

scutellum and metanotum fully

reticulate rugose, with 2 brown

fore-wing

with

impression and anterior

impression strong;

tubercles not reaching

reticulate rugose;

with distal

granulated,

anterior transversal

with 2

longitudinal

transversal

nearly as long as proximal

bands and

part;

segment

1 of front tarsus more than twice as long as segment 4 (22:9); segment 3 of front

tarsus produced into a hook;

subapical

1 row of 10 lamellae;

enlarged claw

segment 5 of front

tooth and 6 lamellae; apex with

(Fig.

tarsus

a group of 10 lamellae;

638)

with

(Fig. tibial

638)

with

spurs

1,1,2. MALE:

unknown

LOCUS TYP I CUS:

Temesca I tepec (Mex ico)

TYPICAL MATERIAL:

holotype F!

HOSTS: unknown DISTRIBUTION: Nearctic region: Temescaltepec

(Real

in BM; U. S.

de Arriba),

paratype F!

A.:

Portal

in TW.

(Ariz.),

TW!

MEXICO:

BM!

2. Tridryinus testaceus n. sp.

FEMALE:

fully

petiole black

winged; and a

length 5,62-6,00 mm;

black

spot between

sometimes also ocellar region black,

reddish-testaceous,

tegulae and parapsidal

mid coxae,

with furrows;

femora and tibiae partly

brown; head (Fig. 639 B) dull, reticulate rugose; occipital carina incomplete, only visible behind the ocell i and on the sides of vertex; posterior ocell i POL;

touching occipital

pronotum shiny,

hairy,

carina;

crossed by

impression and by an anterior weak posterior collar short because the

OL nearly

three times as

a posterior transversal

transversal

impression

impression;

long as

strong

disc humped;

is very near posterior margin

942

Mem. Amer. Ent.

Fig. 640 -

Inst.,

no. 37,

1984

Chela of Tridryinus testaceus n.

sp.

(type)

Dryininae

Fig.

641

-

Male genitalia of

943

Tridryinus testaceus

n. sp.

from Columbia

(South Carolina)(righ\ half removed)

of pronotum; tegulae;

anterior collar visible;

scutum shiny,

pronotal

fully reticulate rugose;

tubercles not reaching notaulices invisible;

scutellum reticulate rugose; metanoutm smooth; propodeum reticulate rugose, with posterior surface very inclined; posterior surface with 2 longitudinal keels;

fore-wing

very small) proximal

with 3 brown

and with distal

part;

transversal

part of radial

segment 1 of front

bands (median band sometimes vein nearly as

long as

tarsus nearly as long as segment 4;

944

Mem.

Fig.

642 -

segment 3 of front with sUbapical 1,1,2. MALE: fully

Inst.,

no.

37,

1984

Chela of Tridryinus oculatus n. sp.

and

14 lamellae;

lamellae;

winged;

antennae brown,

Ent.

tarsus produced into a hook;

tooth

with 2 rows of 31

Amer.

(type)

enlarged claw

segment 5 of front

tarsus

apex with a group of 15 lamellae;

length 3,87-4,18 mm;

black;

with segment 1 testaceous;

(Fig.

640)

(Fig. tibial

640) spurs

mandibles testaceous;

legs testaceous,

with coxae

brown, clubs of hind femora brown, apexes of hind tibiae brown; antennae not distally thickened; antenna I segments in following proportions: 8: 7:

Dryininae

21:

14:

15:

15:

13:

12:

11:

13;

945

head dull,

fully strongly reticulate rugose;

frons with an

incomplete weakly

visible median keel;

complete;

=

=

POL

9;

=

OL

4;

OOL

metanotum and propodeum dull, invisible;

=

OPL

fully

1;

without dark

part

3;

scutum,

strongly reticulate rugose;

transversal

as long as proximal

occipital carina

=

TL

posterior surface of propodeum without

wing hyaline, nearly

6;

bands;

(15:15);

scutellum, notaulices

longitudinal keels;

distal

genitalia:

fore-

part of radial

fig.

641;

tibial

vein spurs

1,1,2. LOCUS TYPICUS:

Lake Placid

TYPICAL MATERIAL: in WA; F!

(Florida,

1 para type F!

in TE;

U.

S.

A.)

and 1 paratype F!

holotype F!

3 para types (1

F,

1 paratype F! 1 paratype

in OT;

2 MM)!

in TW;

in OL.

HOSTS:

unknown

DISTRIBUTION:

Nearctic region:

TE!

Columbia

(Mo.),

WA!

TW!

Columbia

(5. C.),

U.S. A.:

Lake Placid

TW!

Dryden

Salmon

(Fla.),

(Maine),

(Anderson Co.,

OL! OT!

TW!

OT!

Tex.),

Sebring

(Fla.),

OL!

3. Tridryinus oculatus n. sp.

FEMALE:

fully

winged;

length 6,56 mm;

reddish-testaceous,

with

scutum,

scutellum, metanotum, propodeum, mesopleura and metapleura black; antennae testaceous, with segment 2, apexes of segments 3-4-5 and partly 6 darkened;

antennae distally thickened;

proportions:

10: 8:

dull,

strongly excavated,

hairy,

median keel; carina

36:

14:

12:

9:

frons and vertex

incomplete,

8:

antenna I segments in following

6,5:

6,5:

9,5;

with eyes very

irregularly striate,

visible only behind and on

head

bulging;

(Fig.

639 A)

frons without rugose;

occipital

the sides of ocell i;

weakly

posterior

ocelli touching occipital carina; temples invisible; POL = 2,5; OL = 8; OOL = 14; pronotum hairy, dull, weakly striate around disc, with an anterior weak

transversal

impression and with

a strong posterior transversal

impression; disc humped; posterior collar invisible; pronotal tubercles not reaching tegulae; scutum, scutellum and metanotum dull, fully strongly reticulate rugose;

notaul ices invisible;

posterior surface weak I y rugose; keels;

fore-wing with

a small

broad dark band beneath longer than

proximal

proportions:

21:

4:

part

9:

20:

dark spot on

the pterostigma; (22: 10); 35;

enlarged claw

and with a row of 14 lamellae;

with

two rows of nearly 27 lamellae; segments 2 and 3 of front

Portal

TYPICAL MATERIAL:

(Arizona,

holotype F!

part of radial

(Fig.

642)

with a subapical

U.S.

A.)

(Fig.

group of nearly

tarsus produced

in TW.

vein

segments in following

apex with a

unknown

LOCUS TYPICUS:

with

long i tud i na I

segment 5 of front tarsus

spurs 1,1,2. MALE:

two

the basal cells and with a distal

fore tarsal

tooth

lamellae;

propodeum reticulate rugose,

posterior surface with

into hooks;

642)

15

tibial

Mem.

946

Amer.

En!.

HOSTS: unknown DISTRIBUTION: Nearctic region:

U.

Ins!.,

S.

A.:

no.

37,

Portal

1984

(Ariz.),

TW!

A

c

Fig.

643 -

Heads of fema I es of Tridryinus maxjexophthalmicus n.

(type)(A),

exophthalmicus n. sp. (type)(B) and ruficauda (Richards) (type) (C)

sp.

Part 1: index

946 a

PART 1

INDEX TO DRYINID5

abruptum (Anteon) Olmi 404 achterbergi (Deinodryinus) Olmi 174 actuosus (Deinodryinus) Olmi 211 aculeatum (Anteon) Olmi 545 *aequalis (Anteon) Kieffer 259 aequalis (Deinodryinus) Olmi 240 *aequalis (Psilanteon) Kieffer 253 *aerias (Anteon) Walker 115 *aerias (Dryinus) Walker 115 aerias (Metanteon) Walker 113, 115 afer (Dryinus) Olmi 787 *afer (Paradryinus) Olmi 788 *affinis (Aphelopus) Ashmead 70 afrum (Anteon) Olmi 375 ag i I e (An t eon) 0 I mi 354 alatus (Dryinus) Cresson 828 *a I a tus (Gona t opus) Cresson 828 *alatus (Mesodryinus) Cresson 828 *alatus (Prodryinus) Cresson 828 *albidicolle (Anteon) Kieffer 313 *albidicoll is (Anteon) Kieffer 313 *albidocolle (Anteon) Kieffer 313 *albidocoll is (Chelogynus) Kieffer 313 *albipes (Aphelopus) Kieffer 47 *albipes (Aphelopus) Ratzeburg 47

*albipes (Ceraphron) Ratzeburg 47 albitarse (Anteon) Cameron 512 *albi tarsis (Dryinus) Cameron 512 *albi tarsi s (Mesodryinus) Cameron 512 albopictus (Aphelopus) Ashmead 73, 79, 84 albopictus (Deinodryinus) Olmi 183 albrechti (Richardsidryinus) Dlmi 909 alexandrae (Deinodryinus) Olmi 230 *Allanteon Kieffer 287 *alorus (Antaeon) Walker 317 *alorus (Anteon) Walker 317 *alorus (Chelogynus) Walker 317 *alorus (Dryinus) Walker 317 *alticola (Lestodryinus) Benoit 758 alticolus (Dryinus) Benoit 758 *alutaceus (Chelogynus) Richards 338 alvarenganus (Aphelopus) Olmi 80 alvarenganus (Dryinus) Olmi 857 *americanus (Aphelopus) Ashmead 475 americanus (Dryinus) Ashmead 826 *americanus (Mesodryinus) Ashmead 826 *americanus (Mystrophorus) Ashmead 826 *americanus (Prodryinus) Ashmead 826 amoenus (Deinodryinus) Olmi 185 *amphiscepae (Hesperodryinus) R.C.L. Perkins 828 *ampul iciformis (Campylonyx) 728, 735, 736

Mem. Amer. Ent. Ins!., no. 37, 1984

946 b

*afllJu I i c if ormi s (Lestodryinus) Turner 937

*afllJul iciformis (Mesodryinus) 937

afllJuliciformis (Tridryinus) Turner 937 *angust i col lis (Chelogynus) Berland 329

annulicorne (Anteon) Brues 511

*annul icornis (Anteon) Brues 511

*Antaeon Hal iday 286 *Antaphelopus Benoit 44 Anteon Jurine 110, 286, 287,

131

288

*Anteonella Dodd 612 Anteoninae R.C.L. Per~ins 110 anthereon (Anteon) Benoi t 381

anti Ileanus (Dryinus) Evans

asiaticus (Thaumatodryinus) 01 mi 697 asper (Deinodryinus) Olmi 226 aterrimus (Dryinus) Dodd 888 *aterrimus (Paradryinus) Dodd 888

860

*anti Ileanus (Mesodryinus) Evans 861 antiquus (Dryinus) Ponomarenko 751 *antiquus (Laberius) Ponomarenko 751 anxium (Anteon) Olmi 536 Aphelopinae R.C.L. Perkins 43 Aphelopus Dalman 44 *apterus (Anteon) Dodd 571 arabicum (Anteon) Olmi 387 arcuatum (Anteon) Kieffer 289,

Perkins 455 *arizonensis (Anteon) R.C.L. Perkins 455 *arizonica (Bocchus) R.C.L. Perkins 645 *arizonica (Eukoebeleia) R.C.L. Perk i ns 645 *arizonica (Phorbas ) R.C.L. Perkins 645 *arizonicus (Aphelopus) R.C.L. Perk ins 73 arizonicus (Bocchus) R.C.L. Perkins 645 *arizonicus (Hesperodryinus) R.C.L. Perkins 728, 828 ashmeadi (Gonatopus) Kieffer 28 asiaticus (Deinodryinus) Olmi

299,

305,

312

*arcuatus (Anteon) Kieffer 305

areolatus (Deinodryinus) Ponomarenko 121 areolatus (Dryinus) Olmi 894 *areolatus (Electrodryinus) Ponomarenko 119,121 argent inus (Dryinus) Olmi 846

arizonense (Anteon) R.C.L.

atlanticus (Gonatopus) Olmi 172

atlanticus (Mirodryinus) Olmi 672

atratus (Aphelopus) Dalman 50

*atratus (Dryinus) Dalman 50

*atriceps (Bocchus) Brues 133

*atriceps (Chelogynus) Brues 136 *atriceps (Deinodryinus) Brues 136

*atriventris (Anteon) Cresson 136

atriventri s (Deinodryinus) Cresson 133, 136 *atriventris (Dryinus) Cresson 133

*audax (Hesperodryinus) R.C.L. Perkins 828 aul icum (Anteon) Olmi 533 australe (Anteon) Olmi 538

Part 1:

australiae

bicolor bicolor

(Bocchus) Olmi

656 aust ra I i anus

(Dry i nus) 0 I mi

900 *australis (Anteon) Dodd 585 austral is (Aphelopus) Olmi 83 *Avodryinus Ponomarenko 907 *azorus (Anteon) Walker 317 *bakeri (Anteon) Kieffer 278 bakeri (Lonchodryinus) Kieffe 278 *bakeri (Prenanteon) Kieffer 278 barbara (Crovettia) Olmi 89 *barbatum *barbatus

84,

(Anteon) Chitty 295 (Antaeon) Ch it ty

946 c

index

(Aphelopus) Fenton 71 (Deinodryinus) Olmi and

Currado 163 *bicolor (Lonchodryinus) Olmi and Currado 163 *bidens (Lestodryinus) Haupt 744 bi loboides (Deinodryinus) Olmi 245 bi lobus (Deinodryinus)

295 *basale (Prenanteon)

Dal.

261

bisulcatus

*basal is

Dal.

258

Bocchinae Richards 599

(Gonatopus)

*basa lis (Prenan t eon) 261 basi lewskyi (Dryinus) 774, 775 bauense (Anteon) Olmi

Da I man Benoi t 446

*beaumonti (Antaeon) Chitty 260 beki Iyensis (Bocchoides) Ben.

602, 604 belizensis (Dryinus) Olmi 858 bellator (Anteon) Olmi 568 bengalense (Anteon) Olmi 428 benianus (Deinodryinus) Olmi

243 benoi t i 129 *bensoni

(Deinodryinus) Olmi (Anteon)

Richards

305

Fenton

191 bini (Bocchus) Olmi 629 birmanus (Aphelopus) Olmi 69 biroi (Prioranteon) Olmi 591, 593 bismarckense (Anteon) Olmi 575 bismarcki (Dryinus) Olmi 887 *bisulcata (Mercetia) Ben. 754 (Dryinus)

Ben.

754

*Bocchini Richards 599 Bocchoides Benoi t 601, 602 Bocchus Ashmead 611 bolivianum (Anteon) Olmi 518 bol ivianus (Deinodryinus) Olmi 217 borneanum (Anteon) Olmi 416 borneanus (Aphelopus) Olmi 65 bouceki (Anteon) Olmi 564 boucek i (Bocchus) 0 I mi 661 bouceki (Dryinus) Olmi 880 bouga i nv i I lei (An t eon) 0 I mi brachycerum (Anteon) Dalman 300 *brachycerus *brachycerus

555 289,

(Anteon) Dal. 300 (Dry i nus) Da I. 300

brachypterum (Anteon) Olmi 353 brasiliana (Crovettia) Olmi 93

*berlandi (Chelogynus) Rich. 329 berlandi (Chelothelius) Bern.

*brevicollis

609 biaki (Dryinus) Olmi 890 Biaphelopinae Olmi 94 Biaphelopus Olmi 94, 95 *bicolor (Anteonella) Dodd

295, 299 *brevicornis (Anteon) Dal. 295 *brevicornis (Dryinus) Dal. 295 *brevicornis (Gonatopus) Dalm.

660

300 *brevicorne

(Anteon)

Kieffer

(Anteon) Dalman

294, 295 breviventral is

(Antaeon) Ch.

317

946 d

Mem.

browni

(Dryinus)

*brown i

Amer.

Ashmead

(Lest odry i nus)

Ent.

817

Ashm.

(Thaumatodryinus)

Santis and V. *brunneipes

Sarm.

1984

Cameron carin i ceps

918

(R i chardsi dry i nus)

(De i nodry i nus)

714

casalei

(Prioranteon) Dlmi

593, 596 cascus (Deinodryinus) catarinae

Benoi t

786 (Anteon)

Dlmi

D I mi

Dlmi

(Dryinus) Dlmi

*cephalotes

(Anteon)

*cephalotes

(Gonatopus)

232 589, 182

864

Ljungh

317

Ljungh

316

551 caledonicus

(Aphelopus)

Dlmi

chamaeleo

(Dryinus)

Chelanteon Dlmi

83 *cameroni 334

(Anteon)

*cameroni

Kief.

(Chelogynus)

317,

Kief.

334 *Campylonyx Westwood (Aphelopus)

canabense

918

carpens

295

(Dryinus)

caledonianum

camus

37,

De

(Xenanteon)

Berland burgeoni

no.

Cameron

818 bruchi

Inst.,

(Anteon)

*canabensis

728,

729

Richards Benoit

54 370

chelogynoides

775

(Anteon)

R.C.L.

Perk ins 527 *chelogynoides (Prosanteon)

R.

Perk i ns 286, 527 *Chelogynus Haliday 286,287, 288 Chelothel ius Reinhard chiapasi

(Hirtanteon)

Ben.

105

604,

(Deinodryinus)

605

Olmi

209

Benoit 287, 370 canadense (Anteon) Ashmead

chiriquense (Anteon) Cam. 508 *chiriquensis (Dryinus) Cam.

289, 457 *canadensis (Anteon)

508 *chi riquensis

Ashmead

Cameron

459 *canadensi s *canadensis

458,

Ponomarenko

Ceba I

(Paradryinus)

Ceba I los 734 capei (Anteon) Dlmi

409

(Dryinus) Dlmi 849 (Anteon) Kieffer (Chelogynus)

Kief.

322 *carinatus (Deinodryinus) Fen t on 136 *cariniceps 918 * ca r i n i c ep s

(Dryinus)

Cameron

(L est od ry i nus)

Perkins

728,

(Chelogynus)

481 (Dryinus) Dlmi

(Anteon) Olmi

835

486

(Deinodryinus)

Dlmi

206 clarus (Thaumatodryinus) Dlmi 715 clavatum (Anteon) Currado 491 *clavatum

Olmi

(Prenanteon)

and Moczar

261 *cognatus

322 *carinatus

ciudadi claripes

930

(Dry i nus)

734 *canariensi s

caraibicus

Fen ton citricolus

459

(Richardsidryinus)

canari ens is

'~carinatus

*chlorotettixi

930

(Chelogynus)

Ashmead canadensis

*Chlorodryinus R.

(Avodry inus)

Ponomarenko

(Mesodryinus) 508

(Anteon)

R.

Perkins

549 *cogna tus

(Che I ogynus)

527, *cognatus

549 (Neochelogynus) R.

Perk i ns

R. Perk.

548

*collaris

(Anteon)

"collaris

(Gonatopus)

Dalman

313

Dalm.

313

Part 1:

collaris

(Dryinus) linn. (Sphex)

(Crovettia) Olmi

94 colombianus Olmi

(Aphelopus) Fenton

compressicoxale 398 *compressicoxale Benoi t

(Anteon)

74

Ben.

(Xenanteon)

(Dryinus)

Olmi

(Anteon)

571 *coriaceus

516 581

345 R.

Perkins

(Anteon) R.

Perk.

571 *coriaceus (Chelogynus) Perk i ns 571 *coriaceus

R.

571

corruptum

(Anteon) Olmi

*corsicae

(lestodryinus)

Kieffer

curradoi *cursor

545

(Anteon) Olmi

299 *cursor (Dryinus)

294 300

(Anteon)

(Anteon)

897 260

*daos

(Dryinus) Walker

*deceptor

(lestodryinus)

762 *declivis

(Anteon)

(Anteon)

Moczar

Moczar

(Chelogynus) 332

R.

Perkins

547, 549 *destructor (Chelogynus) *destructor Perk ins

*crassifrons

(Neochelogynus)

diaphanus

(Aphelopus)

diaphanus

(Deynodryinus)

Olmi

(Anteon) Haliday

158 diffusus

(Dryinus) Hal.

79 *dikraneuri

(Aphelopus)

74 *dimidiatus

(Anteon)

Kief.

259 *crassiscapus

(Prenanteon)

561

77

Olmi

(Aphelopus) Olmi

Perkins

R.

547

260 *crassimanus

(Anteon)

R.

549

*crassimanus

259 *crassiscapus

317

(Thaumatodryinus)

Perkins

332

259

118

(Anteon) Chit.

Benoit 690 destructor (Anteon)

crassifrons

Turner

Kieffer

Perkins

(Mesodryinus)

913

261

(Anteon) Olmi 380 (Dryinus) Turner 762

913 *corsicus

907,

259

*daos (Prenanteon) Walker dayi (Anteon) Olmi 367 debi Ie (Anteon) Olmi 447

*delicatulus

(Richardsidryinus)

Olmi

Olmi

(Anteon)

260

300

Walker

*daos

dentatus

Marshall

Kief.

Kieffer

(Deinodryinus)

249 dahmsi (Dryinus)

295,

Kieff.

(Dryinus) Marshall

913 corsicus

556

Hal iday

(Anteon)

*corsicus

Mars.

223

84

Deinodryinus R.

735

Olmi

(Anteon) Haliday

decellei deceptor

R.

(Neochelogynus)

Perk ins

295

(Deinodryinus)

cuzcanus

833

(Anteon) Olmi

(Anteon) Olmi

coriaceum

295

Kief.

*curvus

coral I inum (Anteon) Olmi corax

Kief.

(Anteon)

*curvinervis

Conganteon Benoit 98 Conganteoninae Olmi 96 conterminum

(Anteon)

*curvatus

398

825

*crenulatus

Crovettia Olmi

180

Krom.

(Mesodry i nus) Krom.

825 *crenulatum croceus

(Deinodryinus)

260

(Dryinus)

*crawf ord i

linnaeus

735 colombiana

constans

Kieffer

735,

crawfordi

736 *collaris

*comesi

946 e

index

76,

Fenton

R.C.l.

946 f

Mem. Amer.

*dimidiatus

En!.

(Chelogynus)

Ins!.,

no.

37,

R.

261

Perkins 561 *dimidiatus (Neochelogynus) R. Perkins 561 *divisus (Anteon) Kief. 317 *doddi (Anteon) Kieffer 554

*exiguus

*doddi

fabulosum *facialis *facial is

(Liodryinus) Kieffer

287, 554 *dolichocerus

(Anteon)

259 Dryininae Kieffer

Kief.

1984

(Anteon)

322

*exiguus (Chelogynus) Haupt 317, 322 exi lis (Dryinus) Dlmi 802 expol i tum (Anteon) Dlmi 444 (Anteon) Olmi 404 (Anteon) Thomson 313 (Dryinus) Thomson 313

fentoni (Aphelopus) Olmi 78 *ferrugineus ( Anteon) Brues

723

*Dryinus Fabricius 728 Dryinus Latrei lie 728, 729 dubium (Anteon) Fouts 570

*ferrugineus 133

*dubius (Anteon) Fouts 571 dubius (Bocchus) Dlmi 653 durangoi (Anteon) Dlmi 484 *Electrodryinus Ponomarenko 119

(Chelogynus)

136

Brues

*ferrugineus (Deinodryinus) Brues 136 fijianum (Anteon) Olmi 569 'filicornis (Prenanteon) Og I ob lin 260 Fiorianteon Olmi 108

elegans (Deinodryinus) Dlmi 154 *ell imani (Anteon) Chi tty 326 emeri tum (Anteon) Dlmi 410 eminens (Deinodryinus) Dlmi 217 ephippiger (Anteon) Dalman

*flaviceps (Tetradryinus) Kief. 650 *flavicollis (Bocchus) Ashmead 612, 650, 651 flavicorne (Anteon) Dalman 289,

289, 312, 313 *ephippiger (Chelogynus) Dalm. 313, 317, 334,

305, 309, 312, 313, 329 *flavicornis (Anteon) Dalman 309

*ephippiger

(Gonatopus)

439

Dalm.

313 *erraticus (Dryinus) Turner 916 *erra t i cus (Neodry i nus) Tur. 916 erraticus (Richardsidryinus) Turner 916 *escorialensis (Mesodryinus) Ceballos 913 *Eukoebeleia R. Perkins 612 europaea 624

(Bocchus)

*europaea 624 europaeus 624 *euscelisi

Bernard

(Eukoebeleia) (Bocchus)

Bern.

Bernard

(Prenanteon)

Haupt

f i 0 r i i firmum

(An t eon) 0 I m i (Anteon) Olmi

362 558

*flavicornis (Gonatopus) Dalm. 309 *flavinervis (Anteon) Kieffer 337 flavipes (Bocchus) Kief. 650 *flavipes (Tetradryinus) Kief. 612, 650 *flaviscapus (Anteon) Kieffer 322, 337 "flaviscapus (Anteon)

Jansson

313, 317, 322 *flaviscapus (Anteon) Whittaker 481 *flavitarsis (Anteon) Kief. 300 flavoniger (Dryinus) Olmi 838 flavus (Lonchodryinus) 274

Olmi

Part 1:

flavus (Thaumatodryinus) Olmi 718 fluviatilis (Deinodryinus) Olmi 248 *foersteri (Anteon) Kieffer 259 formicaeformis

(Mystrophorus) Latr.

729, 735, 736 *formicarius (Lestodryinus) Latreille 728,735,736 *foveatus (Prenanteon) Rich. 261 *fractinervis (Anteon) Kieff. 260 *frontalis

(Anteon)

*frontal is

(Chelogynus)

259 *frontal is

Dalm.

(Gonatopus)

259

Dalm. Dalman

258 *frontalis (Prenanteon) Dalm. 260, 261 fulviventre (Anteon) Hal iday 313, 337, 338 *fulviventris (Anteon) Hal. 338 *fulviventris (Dryinus) Hal 337 fulvus (Dryinus) Olmi 815 funestum (Anteon) R. Perkins 462 *funestus

(Anteon) R.

Perkins

463 *funestus (Chelogynus) Perkins 463

R.

*fuscicornis (Gonatopus) Dalm. 258 *fusciformis

(Anteon)

Kieffer

325 *fuscipennis

(Anteon)

Kieffer

260 *fuscipes *fuscipes

(Anteon) Thom. 338 (Dryinus) Thomson

337 *fuscoclavatum (Anteon) 317

*fuscoclavatus (Anteon) Kieff. 317 fuscum (Metanteon) Olmi 116 * f u s i form i s (An t eon) K i e f. 325 fyanense (Anteon) Olmi 435 garambanum (Anteon) Olmi 411 gaullei (Anteon) Kief. 334 *gaullei

Ruthe 678, 679 *formicarius (Dryinus)

Kief.

946 g

index

(Chelogynus)

286, 334 gigas (Dryinus) R.

Kieffer

Perkins

*gigas (Paradryinus) R. giluwense (Anteon) Olmi

885

Per. 526

885

9 i rau I t i (An t eon) Dodd 587 glaber (Dryinus) Olmi 900 gloriosum (Anteon) Olmi 575 gobiensis (Mi rodryinus) Ponom. 666 *graci I i coil is 337 *grandi s

(Anteon)

(Anteon)

Brues

Ki ef. 136

*grandis (Chelogynus) Bru. 136 *grandis (Deinodryinus) Brues 136 grandis (Dryinus) Oglob. 852 *grandis (Mesodryinus) Oglob. 852 *gregori

(Lestodryinus) Hoffer

744 gryps (Chelothel ius) Reinhard 605, 607 *gryps (Dryinus) Reinhard 607 guineensis (Aphelopus) Olmi 82 guineensis (Bocchus) Olmi 658 guineensis (Dryinus) Olmi 884 guineensi s (Lonchodryinus) Olmi 284 gutturnium (Anteon) Ben. 384 *gutturnium (Xenanteon) Benoi t 384 *Gymnaphelopus Benoi t 44 hageni (Anteon) Olmi 578 *halidayi (Anteon) Kieff. 260 haustum (Anteon) Olmi 580 *heidelbergensis (Aphelopus) Ri chards 54 *henshawi (Anteon) Ashmead 136 *henshawi (Chelogynus) Ash. 133

Mem. Amer.

946 h

Ent.

342

1984

(Anteon)

Kief.

300

325, 327 *infectus (Anteon) Haliday 325 *infectus (Chelogynus) Haliday 286, *infectus inflatrix

( Dei nod ry i nus)

Kieffer *h i rt i corn is

37,

*indivisus (Anteon) Kief. 300 inermi 5 (Deinodryinus) Olmi 233 infectum (Anteon) Haliday 289,

*Hirtanteon Benoit 287 '~hirticornis (Anteon) Kief. 238 h i r tic 0 rn i s

no.

810 *indivisum

*hen shawi (De i nod ry i nu s) Ashmead 136 *Hesperodryinus R. Perkins 728, 729 hilare (Anteon) Olmi

Ins!.,

237 (Tri sant eon)

327 (Dryinus) Hal iday 325 (Anteon) Benoit 289,

Kief. hirtifrons

119, 238 (Anteon) Whittaker

399 infuscatus

487, *holomelas

488 (Aphelopus)

Olmi 690 ingenuum (Anteon) Olmi 342 insignis (Deinodryinus) Olmi

Rich.

50 hospes (Pseudogonatopus) R. Perk ins 7 hova (Anteon) Benoi t 397 hova (Dryinus) Ceballos 772 *hova

(Mesodryinus) Ceb.

193 *insulanum 124 insulanus

772

hoyoi (Anteon) Olmi 390 *hyalinipennis (Anteon) Kief. 260 hymenaeus (Deinodryinus) Olmi 178 *Hymenodryinus Benoi t *ignotus (Chelogynus)

612 R.

Perkins 549 *ignotus (Neochelogynus)

(Thaumatodryinus)

(Prenanteon)

Benoi t

(Deinodryinus)

Ben.

124 insularis (Dryinus) Dodd 880 *insularis (Paradryinus) Dodd 880 *integer (Anteon) Kief. 259 involutum (Anteon) Olmi 584 iphias (Deinodryinus) Olmi 160 irregularis (Dryinus) Olmi 794 italicus (Bocchus) Olmi 614 i tenezi

R.

Perkins 549 *ilus (Anteon) Walker 260 *i Ius (Dryinus) Walker 259 *imberbis (Anteon) Kief. 305 incaicus (Deinodryinus) Olmi 214 incertus (Dryinus) Olmi 785 incisus (Aphelopus) Olmi 64 *inclytus

(Anteon) Haliday

*inclytus

(Dryinus) Hal iday

(Deinodryinus)

Olmi

245 jamaicanum (Anteon) Olmi 497 jamaicanus (Aphelopus) Olmi 81 japonicum (Anteon) Olmi 347 javanum (Anteon) Olmi 437 *javanus (Paradryinus) R.C.L. Perkins 806 junonium (Fiorianteon) Olmi

325

325 indianus (Richards!dryinus) Olmi 922 indicum (Anteon) Olmi 419 indicus (Dryinus) Kief. 810, 871 *indicus (Mesodryinus) Kief.

108,

109

jurinea~um (Anteon) La!. 287, 294,

295,

338 *jurineanus

299,

305,

312,

(Anteon) Latrei lie

295 kasaicum (Anteon) Benoi t 362 *kasaicus (Chelogynus) Ben. 362 kawandanum (An t eon) 0 I mi 374

Part 1:

*kiefferi

(Antaeon) Chitty (Anteon) Chitty 329 (Chelogynus) Chitty

329 kiefferi *kiefferi

(Dryinus)

Fouts 795

(Lestodryinus) (Dryinus)

kivuanum

(Anteon)

*kivuanus 355, koebelei

B64

Benoi t

(Chelogynus)

363

Benoi t

363 (Dryinus) R.

Perkins

(Paradryinus)

R. R.

721

R.C.L.

560

(Neochelogynus)

R.C.

Perkins 560 *Iepidus

(Anteon)

*Iepidus

(Chelogynus)

Forster 260 Forster

leptias

(Dryinus)

Ponom.

B02, 803 *Iaeticornis (Anteon)

Kieffer

539 (Chelogynus)

(Paradryinus)

* L est od ry i n i n i leucopus levigatus

(Dryinus)

Haupt

322

R.C.L.

K i e f fer

(Aphelopus)

723 728,

729

Kieffer

(Anteon) Kieffer

*Liodryinus Kieffer

78

309

287

lobatum (Anteon) Dlmi Lonchodryinus Kieffer

506 253

*Iongicornis

(Anteon)

Dalman

259 *Iongicornis

(Dryinus)

Dalman

258 *Iongicorni s

441 lankanus (Dryinus) Dlmi B21 laotianum (Anteon) Dlmi 435 laotianus (Bocchus) Dlmi 635 *Iapponicus (Anteon) Thomson 260 *Iapponicus (Dryinus) Thomson 259 *Lasianteon Kieffer 287 *Iateral is (Anteon) Thomson 325 *Iateralis (Dryinus) Thomson 325 laticeps (Bocchus) Ashmead 652 *Iaticeps (Phorbas) Ashmead 612, 652 *Iaticeps (Phorbasia) Ashmead 612 Dlmi

R.C.L.

895

*Lestodryinus Kieffer

747

lankanum (Anteon)

Perkins

Perkins 895

koreanus (Dryinus) Moczar 747 krombeini (Anteon) Dlmi 429

*Ianionis

(Chelogynus)

*Ieptias

(Chlorodryinus)

Moczar

krombeini

*Ieiosomus

Perk i ns

Perkins 684,

R.

259

Perkins 72B, 882 koebelei (Thaumatodryinus) *koreanus

(Anteon)

*Ieiosomus Dlmi

882 *koebelei

560 *Ieiosomus 560

Perk i ns

Fouts 796 kimseyae

946 i

latus (Dryinus) Dlmi 808 leiosomum (Anteon) R. Perkins

329 *kiefferi "kiefferi

index

(Prenanteon)

260, 261 *Iongifilis (Anteon) *Iongifilis

Dalman

Kieffer

260

(Prenanteon) Kieffer

260 *Iongiforceps

(Anteon)

Kieffer

322 *Iongiforceps Kieffer longipes

(Chelogynus) 322

(Dryinus) Dlmi

longiscapum

799

(Anteon) Ceballos

392 *Iongiscapus Ceba I los lucens

(Xenanteon) 392

(Dryinus)

Dlmi

817

*Iucidum (Anteon)

Hal iday

317

*Iucidus

(Anteon)

Hal iday

317

*Iucidus

(Chelogynus)

Hal iday

317 *Iucidus

(Dryinus) Hal iday

316

946 j

Mem. Amer. En!.

lucluosum (Anleon) Olmi 577 *Iuffnessensis (Anlaeon) Chilly 260 * I u su s (An I eon) R. C . L . Perkins 463 *Iusus (Chelogynus) R.C.L. Perkins 463 *Iuleiceps (Prenanteon) Kieffer 261 *Iuteipes (Anleon) Kieffer 259 *Iuleipes (Prenanleon) Kieffer 260 *Iuzonicus (Lestodryinus) Kieffer 806 *Iyde (Anteon) Walker 300 *Iyde (Dryinus) Walker 300 maci lentus (Thaumatodryinus) De San I. and Vida I Sa r • 706 maculiceps (Aphelopus) Bergman 66 *maculipenne (Anteon) Kieffer 334 *macu I i penn is (An I eon) Kieffer 334 *madagascariensis (Antaphelopus) Benoi I 60 madagascariensis (Deinodryinus) Benoi t 128 *madagascariensis (Hirlanleon) Benoit 128 *madagascola (Xenanleon) Benoi t 393 madagascolum (Anteon) Benoi t 393 "madecassum (Hirtanteon) Beno it 125 madecassus (Dei nodry i nus) Benoit 125 magnificus (Megadryinus) Richards 28 mahensis (Dryinus) Kieffer 764 *mahensis (Mesodryinus)

Ins!.,

no. 37,

1984

Kieffer 764 malayanus (Aphelopus) Olmi 66 malayanus (Thaumatodryinus) Olmi 699 *marginatum (Anteon) Kieffer 295, 299 *marginatus (Anteon) Kieffer 295 mari timum (Anleon) Turner 357 *mari timus (Chelogynus) Turner 357 maroccanus (Richardsidryinus) Olmi 910 masneri (Biaphelopus) Olmi 96 masneri (Lonchodryinus) Olmi 277 masoni (Anteon) Olmi 469 maximus (Deinodryinus) Olmi 165 mayogo (Dryinus) Benoi I 780 *mayogo (Lestodryinus) Benoi t 780 *mediocarinatus (Anlaphelopus) Benoi t 44, 60 mediocarinatus (Aphelopus) Benoi t 60 medleri (Anteon) Olmi 395 melaleucus (Aphelopus) Dalman 47 *melaleucus (Dryinus) Dalman 47 *melaleucus (Gonatopus) Dalman 47 *melanacrias (Anleon) R.C.L. Perk i ns 463 *melanacrias (Chelogynus) R.C.L. Perkins 463 *melanocephalus (Bocchus) Cameron 168 melanocephalus (Deinodryinus) Cameron 168 *melanocephalus (Dryinus) Cameron 168 *melanocera (Anteon) Kieffer 259

Part 1:

*melanocera

*misor

(Prenanteon)

Kieffer

molle

*melanocerum (Prenanteon) Kieffer 253 Perk i ns R.C.L. menkei

(Anteon)

Perkins 527

(Anteon) Olmi

Mesodryinus Kieffer Metanteon Olmi

mosseli

729

muiri

113

(Anteon) Olmi

mexicanus

(Dryinus)

499

R.C.L.

823

*microleucus

(Anteon)

Perk ins Fen t on

R.C.L.

(Antaeon)

Chitty

Olmi

(Anteon) Olmi

muluensis

Olmi

635

mundum (Anteon) Olmi 565 munitum (Anteon) Olmi 345 (Anteon)

Perkins

7,

Perkins

Olmi

439

R • C • L.

R.C.L.

529 (Anteon)

R.C.L.

529

*myrmecophi Ius

728

260

408

432

(Bocchus)

"myrmecophilus (Prenanteon)

micros (Anteon) Olmi 519 *microtomus (Aphelo,pus) Wh it taker

Moczar

myrmecophi lum (Anteon)

73

*micropunctatus

492

(Anteon)

(Anteon)

munroei

*mexicanus (Paradryinus) R.C.L. Perkins 823

Olmi

785

montanus (Dryinus) Olmi 878 monticolus (Deinodryinus) Olmi 130 *morleyi

486

mexicanum

Perkins

(Anteon)

259

Olmi

317

R.C.

(Prosanteon)

Walker

(Dryinus)

*mongol icum

527

*melanostigmus

(Dryinus)

modestus

261

*melanost igmus

946 k

index

(Paranteon)

Perk ins

286,

529

mysorense (Anteon) Olmi 422 Mystrophorus Forster 678

72

migratorius (Thaumatodryinus) Benoit 694

namibiense (Anteon) Olmi 383 '~nanum (Anteon) Haliday 295

miles (Anteon) Olmi 547 mingoae (Anteon) Olmi 391

*nanu s

(An t eon)

*nanus

(Dryinus)

mi n i mum

napensis

(An t eon)

*minimus minimus

(Anteon) (Bocchus)

*minimus 467 minimus

467

Fenton Olmi

(Chelogynus)

467

658 Fenton

241 *minutus

295

Olmi

850

natalense

(Anteon) Olmi

388

nearctica

(Crovettia)

nearcticus

nemorale

Olmi

Olmi

85

(Richardsidryinus)

(Anteon)

461

Kieffer

(Anteon) Olmi

300

442

*Neoanteon Fouts 612 (Anteon)

*Neochelogynus R.C.L.

Ashmead

286,

453 mirabilis

(Bocchus)

R.C.L.

Perk i ns 643 *mirabilis (Eukoebeleia) R.C.L. Perkins 612, *mirabilis

(Phorbas)

643

R.C.L.

Perkins 643 mirabilis (Thaumatodryinus) Olmi

(Dryinus)

neglectus

704

(Deinodryinus)

295

Hal iday

Olmi 927 nebulosum (Anteon) Olmi

(Thaumatodryinus)

Olmi minor

Fe" t on

Ha I i day

687

Mirodryinus Ponomarenko

663

neotropica

Perkins

287 (Crovettia)

nepalense

(Anteon)

nepalense

(Conganteon)

Olmi

Dlmi

92

304 Olmi

98

nepalensis nepalensis

(Aphelopus) Olmi 57 (Dryinus) Olmi 740

nepalensis

(Lonchodryinus) Olmi

267 nigricans 211

(Deinodryinus)

Cameron

9461

Mem.

*nigricans

37,

Perk. Kieffer

(Dryinus)

*oriphilus

Olmi

Olmi

851

541

(Aphelopus) Olmi

67 oriphi lum (Anteon) orophi Ius

(Anteon)

1984

(Anteon)

orientalis Kief.

(Anteon) R.

568 *nigricornis

no.

orientale

211

(Aphelopus)

56 nigricorne

Inst.,

676 opacifrons

211

(Plastodryinus)

Cameron nigriceps

Ent.

(Dryinus)

Cameron *nigricans

Amer.

Kieffer

403

(Anteon) Kieffer (Dryinus) Benoi t

*orophilus

(Lestodryinus)

403 781

Benoit

781

300 *nigricornis

(Anteon)

R.

Per.

568 *nigricornis

R.

(Chelogynus)

Perk ins 568 *nigricornis (Neochelogynus)

R. Perkins 568 *nigricorpus (Paradryinus) Giraul t

(Anteon)

Kief.

300 nigrolobatus

(Deinodryinus)

Olmi 199 nigrorubrum (Anteon) Olmi

(Anteon)

Fenton

(Chelogynus)

481

Fenton

*otiartes *otiartes

(Anteon) Walker 295 (Dryinus) Walker 294

pacificus

(Dryinus) Olmi

876

pagl ianoi (Bocchus) Olmi 620 palaeodominicanus (Dryinus) Cur.

885

*nigroclavatus

osborni "osborn i 481

and Olmi

866

*pa I lid i corn is

(An t eon)

Ashmead

459, 539 *pallidicornis

(Anteon)

Kieffer

n i grorufus (Dei nodry i nus) Olmi 159 nitidum (Anteon) R. Perkins

260 *pallidicornis (Chelogynus) R. Perk i ns 540 *pa I lid i corn is (Neoche I ogynus)

561 *nitidus

(Anteon)

R. Perk ins 539 *pallidinervis (Anteon)

*nitidus

(Chelogynus)

494

Kieffer R.

561 Per.

"pall idus

561 *ni tidus

R.

(Neochelogynus)

Perk i ns notogeicus

Olmi 286 noyesi (Anteon) Olmi 503 noyesi (Deinodryinus) Olmi 170 nycteum (Anteon) Olmi *obscuricorne 300 *obscuricornis

945 oliveirai olmi i

510

260 panamense

Kief.

251 papuanus

Olmi

Kief. 67

(Tridryinus) Olmi (Anteon) Olmi

502

(Radi imancus) Moczar

810, R.

Perkins

(Prenanteon)

Ogloblin 497

(Deinodryinus)

(Dryinus)

R.

871

(Anteon) Olmi

paradoxus (Anteon)

728,

(Dryinus)

871 *palustris

panamensis

(Anteon)

300 ochreus (Aphelopus) oculatus

(Chlorodryinus)

Perk ins pall idus

561

(Lonchodryinus)

Kieffer

259

Olmi

(Deinodryinus)

Perkins 119, 145 "Paradryinus R. Perkins

Olmi

904

R. 728

*Paranteon R. Perkins 286 paranus (Deinodryinus) Olmi 236 *parcepunctatus

K i ef.

260

(Prenanteon)

C.

Part 1:

parthicus

(Bocchus)

pilicorne

Ponom.

624, 625 "parvicollis (Anteon)

Kieffer

259 *parvicollis

(Prenanteon) K i ef. 260 parvulum (Anteon) R. Perkins 539 *parvulum (Chelogynus) R.C.L. Perkins 540 *parvulus (Anteon) Kief. 337, 539 *parvulus (Anteon) R. Perkins 540 *parvulus (Neochelogynus) R. Perkins 539 *parvus (Anteon) Kieffer 338 pauliani (Lonchodryinus) Ben.

Beno i t

(Bocchus)

636, 639 pegnai (Deinodryinus) 186 *pektusanense

(Deinodryinus) Olmi

204 ·pi losus

(Deinodryinus)

(Anteon)

Walker

·penidas

(Dryinus)

Whitt.

Fenton

136 *Plastodryinus Kieffer plaumanni

(Anteon)

plaumanniana 92 ·pol i tus

Olmi

728 517

(Crovettia) Olmi

(Anteon)

Ashmead

459 pol i tus (Deinodryinus)

458,

Ashmead

206 (Mirodryinus)

674 (An t eon)

praeclarus *prehensor

Olmi

Ashmead

(Dryinus)

452,

Ashmead

452

(Dryinus) Olmi 814 (Chelogynus) Maneval

329

(Prenanteon)

*penidas

68

317

*Prenanteon Kieffer 253 pretorianus (Dryinus) Olmi Prioranteon Olmi *procericornis

Walker

767

589

(Anteon)

Kieffer

259 propodeale (Anteon) Fenton 506 *propodealis (Chelogynus) Fenton

(Thaumatodryinus)

Kronnein 700 perlucens (Deinodryinus) (Anteon)

peruvianus

Olmi

Olmi

583

174 (Anteon) Olmi

pseudoamoenus 445 448

phi I ippinus

(Aphelopus) Olmi

69 ph i i i pp i nu s

(Thauma t od ry i nu s)

697

*Phorbas Ashmead

611

*Phorbasia Kieffer

612

506 *Prosanteon R. Perkins 286 *proximus (Anteon) Kieffer 259

(Deinodryinus)

phi I ippinum (Anteon) Olmi

Olmi

(Aphelopus)

72 pilosifrons

453 *popenoei

Nagy

Moczar 261 penanganus (Aphelopus Olmi

Olmi peterseni

(Anteon) Ogloblin

515 *pilicornis

popenoe i

271

pedunculatus

233 permirum

(Anteon) Ogloblin

Dlmi

(Prenanteon)

ill

515 *pilicornis

ponomarenkoae

271 ·paul iani

316 perkinsi

946

index

(Dei nodry i nus)

01 mi 188 pseudobi lobus (Deinodryinus) 01 mi 193 *pseudophanes (Chlorodryinus) R. Perk ins 878 pseudophanes (Dryinus) R. Perk. 878 *Psilanteon Kieffer ::~j

946 n

Mem.

pubicorne

317 *pubicornis

richardsi

(Prioranteon)

(Gonatopus)

Richardsidryinus Moczar 907 rivularis (Deinodryinus) Olmi

Olmi

322

Dalm.

(Anteonella)

R.

660 robustus

(Bocchus)

Kief.

*rubrica

(Neoanteon)

(Aphelopus)

(Allanteon) (Anteon)

*punctatus

(Chelogynus)

327 punctatus

Kief.

(Dryinus)

puncticeps

594

612,

660

Fouts 612,

(Bocchus)

(Anteon) Kief.

*rubrifrons

(Lasianteon) Kief.

287 rubrolobatus

(Prenanteon)

Kieffer

260

(Lestodryinus)

Kief.

Kieffer

925

(Dryinus)

Olmi

(Aphelopus)

(Prenanteon)

(Anteon) Dalman

259 "ruficornis

(Gonatopus)

258 ruficornis

(Prenanteon)

Olmi

59

rufiscapum

(Anteon)

Perk ins 143 quercicolus (Deinodryinus)

*rufiscapus rufiventris

R.

Ashmead 70 rufonigrum (Anteon)

143

Radi imancus Moczar 675 rapax (Deinodryinus) Olmi 223 "rectus (Anteon) Kieffer 295 reticulaticeps

(Anteon)

"reticulatum

Kief.

(Xenanteon)

*rufulocol Chitty

304 "reticulatus

(Anteon)

Kieff.

(Xenanteon)

304 "retusus

(Anteon)

*retusus

(Dryinus)

Kief.

Dlmi

586

402

(Chelogynus)

is

(Antaeon)

Chitty

is

(Chelogynus)

334

287 *reticulatus

586

334

304

Kief.

Dodd

Berland 313 rufulocolle (Anteon) Chitty '~rufulocol

(Anteon)

Dalm.

(Anteon) Dodd (Aphelopus)

*rufovariegatus

Dodd

532

reticulatum

Dalm.

261

897 R.

Perkins

Dalman

(Lonchodryinus)

(Deinodryinus)

*quercicola

Olmi

Dalman

*ruficornis

258 *ruficornis

(Ri chardsidryinus)

(Deinodryinus)

261

Kieffer

259 *pyrenaicus

313

197

Moczar

"ruficorne (Anteon)

Fouts 636,

893

Ashmead

(Anteon)

rubricus

639 *rubrifrons

313 *pyrenaicus

531,

Dodd

Dodd

105

Olmi

Kief.

Olmi

(Anteon)

325

475

*pyonganensis

querceus

628

Olmi

639

287

quatei

Olmi

239 *robusta

799

*punctatus

pyri Ilae

1984

(Anteon) Dalman

317,

Perk ins 73

*pyri Ilae 925

37,

(Bocchus)

*pulcherrimus

474,

no.

(Chelanteon)

Dalman

316,

316 pulcher (Dryinus)

*punctatus

Ins!.,

richardsi

313,

*pubicornis

En!.

richardsi

(Anteon)

289,

Amer.

rufus

334

(Thaumatodryinus)

710 rugos i ceps

(An t eon)

Thomson

260

471, 473 *rugosus (Anteon)

Thoms.

259

*rugosus

Rich.

K i ef f er

Ashmead

(Trisanteon)

133

Ashm.

136

Part 1:

':'rugulosus

(Anteon)

':'serratus

Fenton

"rugulosus

(Chelogynus)

Fent

481 "ru t shu r i cum

(Xenan t eon)

Ben.

serratus

Maneval

(Tridryinus) (Bocchus)

Benoi t

629

seyrigi

(Dryinus)

Benoit

756

(Dryinus)

Dlmi

731

269

(Anteon)

Dlmi

425

"seyrigi

sarawaki saussurei

(Dryinus)

Ceballos

*saxetanus

764

Kieffer

(Anteon)

289,

Haliday

(Dryinus)

*silvaticus

Dlmi

(Chelogynus)

766 Ponom.

329

(An t eon)

sinensi s

(Lonchodryinus)

(Chelogynus)

sinopensis ':'si si thrus

(Dryinus) (Dryinus)

"si sythrus

(Anteon)

444 419

K i e f fer

337 Dlmi

273 322

Hal iday "scapularis

(Dryinus)

Haliday

slovacus

322 scaramozzinoi

(Bocchus)

Dlmi

(Bocchus)

spangleri

Dlmi 858 Walker 295

295

Walker Strejcek

(Dryinus)

Dlmi

618

762

speciosu5

(Deinodryinus) Dlmi 157 speciosus (Dryinus) Dodd 874 "speciosus (Paradryinus) Dodd

621 (Chelogynus)

"schaef fer i

136

Brues

( Dei nod ry i nus)

"schaef f er i

874

136

Brues

stantoni

(Dei nodry i nus)

sch ling e r i

(Dryinus)

"stantoni

Olmi 230 schoutedeni (Dryinus)

"stantoni

"schou t eden i

':'stantoni Nagy

622,

Kieffer

':'scot i cus

(Anteon)

Kief.

sedlaceki ':'semenovi

(Anteon)

Dlmi

(Neodryinus)

Ashmead

(Prodryinus)

Ashmead

806 striatus

624 (Anteon)

806

Ashm.

806

(Lest odry i nus)

Benoi t 778 scobiolae (Bocchus)

Ashmead

(Lestodryinus)

806

Benoi

778

(Dryinus)

*striatus

845

Fenton

(Lestodryinus)

Fent.

845

295

(Prenanteon)

261 "semipol i ta

295

Pono.

(Neodryinus)

(Anteon)

Kief.

(Aphelopus)

"subapterus subapt erus

534

(Anteon)

Kief.

Kieffer 256 "subapterus (Mystrophorus) Kieffer 256 "subapterus (Prenanteon)

309

Kieffer ':'subarcticus

Rich.

52

256

(Lonchodry i nus)

Gi r.

885 "sericeus

Benoi t

" s i mil i s

322 ':'scapularis

"'scoticum

(Lonchodryinus)

(Anteon) Haliday

322

"scapularis

Benoi

629

silvestre (Anteon) Kief. silvicolum (Anteon) Dlmi

764 scapulare

(Hymenodryinus)

sierranus Kieffer

(Laberius)

Benoi t

269

(Lest odry i nus)

Ceba I los 770 *saxetanus (Labeo)

(Hi rtanteon)

612, seyrigi

770 *saussurei

941

Dlmi

seyrigi ':'seyrigi

384

serratus

(Chelogynus)

317

481

sanderi

9460

index

256 (Anteon)

Hellen

946 p

Mem.

317,

Amer.

En!.

322

*subflavicorne

(Anteon)

Haupt

Ins!.,

no.

37,

1984

84, 87 *thomsoni (Anteon) *threnodes

309, 312, 313 *subflavicornis (Anteon)

Kief.

295

(Paradryinus)

903 townesi (Anteon)

Olmi

R.

Per.

379

Haup t 309 subtilis (Dryinus)

Olmi

875

townesi

(Aphelopus) Olmi

sucklingi

Olmi

573

townesi

(Conganteon)

Chit.

townesi

(Deinodryinus)

townesi

(Thaumatodryinus) Olmi

(Anteon)

*suffolciensis

(Anteon)

300 * su f f

0

I c i en sis

(An t eon)

Ch i t .

300 *sulciscutellum

(Dicondylus)

G i rau I t 660 superbum (An t eon) *superbus

Dodd

(Anteon)

585

surinamense (Anteon) Olmi 521 surinamensis (Aphelopus) Olmi (Dryinus)

(Anteon)

Kief.

317

*triareolatus

(Anteon)

Kief.

300

*tricarinatus

(Anteon)

Kief.

329

(Lonchodryinus)

Kieffer

254, 282 Tridryinus Kieffer 932 trifasciatus (Dryinus) Kieffer 821

"tri I ineatus

Olmi

(Neodryinus)

szel eny i i

(Bocchus)

tri I ineatus

Moczar

Dodd

617 *szepligeti

(Dryinus)

Kieffer

trinidadi

(Deinodryinus)

220 trinitatis

Kieffer 728, 744 tarraconensi s (Dryinus)

80 tripartitum

742,

Mars.

* t rip art i

744

*tarraconensis Marsha I I tasmanianum

(Lestodryinus)

(Richardsidryinus)

932

742, 744, 806, 818 "szepl igeti i (Plastodryinus)

(Aphelopus) (Anteon)

tus

*trisulcatus

744

(Anteon) Olmi

582

Kieffer

Olmi

Kief.

(An t eon)

*Trisanteon Kieffer

Olmi

(Aphelopus) 47

316 *terryi (Paradryinus)

turneri (Anteon) Olmi 378 *typhlocybae (Aphelopus)

817, testaceus testaceus

818 (Bocchus) Olmi 643 (Tridryinus) Olmi

941 *Tetradryinus Kieffer teutoniae

Per.

553

862

682

R.

Perkins

682, 683 "theliae (Aphelopus) Gahan

*typicus

(Chelogynus)

554 *typicus

R.

(Neochelogynus)

Perk i ns 286, 554 ugandanum (Anteon) Olmi

Perkins

(Crovettia) Gahan

(Anteon)

554

612

416

Thauma t odry i n i nae 0 I mi Thaumatodryinus R.

(Anteon) Kief. 334 (Aphelopus) Olmi 76

Muesebeck 74, 76 typicum (Anteon) R. Perkins *typicus

(Dryinus) Olmi

(Anteon) Olmi

329

119

*trivialis tropicalis

R.

329

K i e f.

taylori (Anteon) Olmi 366 *tenuicornis (Dryinus) Dalman

theliae

Dodd

932

839

thai

190

692

807,

77 surinamensi s

100

Olmi

*triangularis

tricolor

585

82

Olmi

87 6,

ugandanus 935

(Tridryinus)

Perk. R. 372

Olmi

Part 1:

umber

(Bocchus) Dlmi

946 q

index

Perk ins 7

R.

614

*vivariensis

(Anteon)

Benoi t 919 undulatus (Richardsidryinus)

475 *vivariensis

(Chelogynus) Brad.

Benoit 919 ungulatus (Mirodryinus)

475 vivax (Anteon) Dlmi

*undulatus

(Lestodryinus)

664, 671, 674 *unifasciatus (Anteon)

Pon.

vulcanicum Ashm.

452, 453 vagans

(Deinodryinus) Dlmi

201 variabi lis

(Deinodryinus)

Fenton 136 varicornis (Aphelopus) 72 variegatus De

Brues

(Thaumatodryinus) and Vidal

703, 704 *varipes (Paradryinus)

Sarm.

903 *venator

(Dryinus)

R. Perkins

(Paradryinus)

R.C.L.

Kief. Kief.

(Aphelopus)

73 *virginiensis

260 *vi tell inipes

K i ef. *vitiensis

260

464

(Richardsidryinus) Dlmi

929 (Anteon) Muesebeck

and Walkley

481

(Aphelopus)

*wittei

Ben.

62

(Gymnaphelopus) (Anteon) (An t eon)

*xanthothorax

Benoit

337

Kief. Brad.

(Chelogynus)

479 Brad.

479 (Anteon)

Fenton

(Chelogynus) (Anteon) (Anteon)

Kief. Kief.

259 *vi tell inipes

317

wasbaueri (Bocchus) Dlmi 644 well ingensi s (Dryinus) Dlmi 839

xan thothorax

475

*vitellineipes

317

Kief.

(Anteon) Dlmi

295

475 *virginiensis

Ben.

vulsum (Anteon) Olmi 576 *walkeri (Chelogynus) Kief.

44, 62 *xanthostigma

R. Perk.

Kief.

(Anteon)

295

Perkins 903

Fen t on

(Anteon)

*vulgaris

wittei

*vicinum (Anteon) *vicinus (Anteon) *viduus

103 *vulgare

*whittakeri

Perkins 903 venator

Ben.

103 *vulcanicus (Conganteon)

wetmorei R.

514

(Conganteon)

98,

wasbaueri

San.

Bradl.

(Prenanteon)

260 (Haplogonatopus)

*Xenanteon Kieffer 287 *xerophilum i/.r.teon) Benoit 668 *xerophi Ius

(Anteon)

Benoi t

668 xerophi Ius

(Mi rodryinus)

668 yasumatsui

(Anteon) Dlmi

Benoi t

424 zairense 407

(Anteon)

Benoi t

406,