MEMOIRS of the AMERICAN ENTOMOLOGICAL INSTITUTE Number 37, Part 1 Pages I-XII, 1-946
A REVISION OF THE DRYINIDAE (HYMENOPTERA) by Massimo Olmi Universit::i degli Studi della Tuscia Viterbo, Italy
The American Entomological Institute 5950 Warren Road, Ann Arbor, Michigan 48105, U. S. A. 1984
TABLE OF CONTENTS
Introduction . . . . . .
............ 1
'.Iaterial
.. . . . . . . . . . 1 .4
examined.
~cknowledgments
3 i
0
.•
.5 .7 .. ... 7 .. .. 10 · .31 . . . . 31
logy . . . . . . . . . .
~osts
attacked ••..•
::'conomic
importance.
',',orphology
and
terminology.
::ollection . • . . . . . • . . • • . • • . • ::; reserva t ion . . . . . . . . . . . . . . . "ami Iy
size and distribution . . . . . . . . .
<ey
the
to
subfami lies
of
Sdbfami Iy Aphelopinae.. :;enus Aphelopus . . . . . . . . Palaearctic species. 1.
me I a I eu cu s .•..
2.
atratus ..
3.
serra tus.
4.
camu s ••..
5. 6.
nigriceps. n epa I en sis.
7. querceus ... Ethiopian species . . . . 1.
mediocarinatus.
2.
wittei . . . . . . .
3.
incisus . . . .
Oriental
species.
. .. 43 , •. 44
5. 6.
leucopus ..
.78
di ffusus ..
. . . 46
7.
t r in ita tis .
· .47 · .50
8.
alvarenganus . . . . . .
.79 .80 .80
9.
jamaicanus . . . .
.52 .54 .56 .57
Australian
1.
townesi . . . .
2. 3.
guineensis. australis ..
4.
ca I ed on i cu s .
.60 . ... 60
5.
a I bop i c t us ..
1. 2.
nearctica.
.. .. 62 .64 . ... 64
borneanus .. maculiceps.
.81 .. .81 · .. 82
species.
.59
.. .. 65 .66 .66 3. malayanus •.. 4.ochreus . . . . . .67 . 67 5. 0 r i en t a lis • • • . . . . . . .68 6. penanganus .. 1. 2.
.32 .34
Dry i n i dae •.
· .82 · .. 83 . . . . . 83
· .• 84 Genus Crovettia . . . . . . . . . . . . . . . 84 Nearctic species . . . . . . . . . . . 84
.... 85 · .87
the I i ae ..••.•
Neotropic species • . . . . 1. barbara . . . . • . . 2.
· .89 .89
4.
neotropica •••••••.••• 92 plaumanniana. · .92 brasiliana . . . . . . . . . . . 93
5.
3.
colombiana.....
..94
7.
birmanus . . . . . • . . . .
.69
Subfami Iy Biaphelopinae.
..94
8.
phi I ippinus ..
.69
Genus Biaphelopus.......
. .95
\Iearctic
species ...
.69
1.
ru f i v en t r is.
2.
bicolor . . . .
.70 .71
3. 4.
varicornis.
.72
a I bop i c t us. species.
.73 .76 .76
~eotropic
1.
tropicalis • . . . • • • .
2.
diaphanus . . . .
· .77 . .. 77
3.
su r i namen 5 is.
4.
fen t on i . . . . • . • . . . • • . . 7 B
Palaearctic 1.
species...
.96
masneri.........
.96
Subfami Iy Conganteoninae. Genus Conganteon . . . . . . .
Palaearctic species. 1.
nepalense . . . . .
Ethiopian
species.
1.
townesi .•..
2.
vulcanicum.
Genus Chelanteon . . . .
.96
.98 .. . . . 98 .. . . . 98 · .99
.100 .103 . . . . . . 105
II
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
species . • • • . • • . . . 105
22.
insignis . . . . . . . . . . . . 193
richardsi . . . . . . . . . . . 105
23.
pseudobilobus ..••••• 193
Genus Fiorianteon ••.••••••••. 108 speci es . . . . . . . 1 09
24. 25.
rubrolobatus . • • • . . . • 197 nigrolobatus • . • • . . . . 199
junonium . . . . . . . . . . . . . 109
26.
vagans . . . . . . . . . . . . . . 201
Subfami Iy Anteoninae . . . . . . . . . 110
27.
pilosifrons . . . . . . . . . 204
Oriental 1.
Pa I aearct i c 1.
28.
politus . • . . . . • • • . • • . 206
species •••••..•• 114
29.
claripes . . . . . . . . . . . . 206
1.
aerias . . . . . . . . . . . . . . 115
30.
chiapasi • • . . . . . • • . . . 209
2.
fuscum . . . . . . . . . . . . . . . 116
31.
n i 9 r i can s ••••••••••. 21 1
Genus Deinodryinus . . . . . . . . . . . 118
32.
actuosus • • • . . . • • . . . • 211
Palaearctic species . . . . . . . 121 1. areolatus . . . • • • • • . . . . 121
33.
incaicus . . . . . . . . . . . . 214
34.
bol ivianus ••...••••• 217
Ethiopian
eminens . • . • . • . . . . . . • 217
Genus Metanteon . . . . . . . . . . . . . . 113 Neotropic
species . . . . . . . . . 123
35.
1. 2.
insulanus . . . . . . . . . . . 124 madecassus ••..•••.•• 125
36.
trinidadi •••...•••.• 220
37.
rapax . . . • • • • • . . . . . • . 223
3.
madagascariensis .••• 128
38.
croceus . • • • • . . . . . • . • 223
4.
ben 0 i t i . . • • • • • • • • . . . 1 29
39.
5.
mon t i co Ius . • • • • . • . . • 130
40.
asper ••••.•••••••... 226 schl ingeri .•.••••.•• 230
sp e c i e s •••••••••. 1 3 1
41.
alexandrae . . . • • • . . . . 230
as i at i cu s •.•.•••••••. 131
42.
carpens .••••••.•.••• 232
species . . . . . . . . . . 133
43.
per I u c en s . . • • . • . • . . . 233
1. 2.
at r i v en t r is ••••••••• 1 33 quercicolus . . . . . . . . . 143
44. 45.
i nermi s .••••••••••.• 233 paranus . • . . . • • • • • • . . 236
3.
paradoxus . . . . . . . . . . . 145
46.
hi r tic 0 rn is •••..•••. 237
Or i en t a I 1.
Nearctic
species . . . . . . . . . 147
47.
rivularis • • • . . . . . . . . 239
e I egan s • • • • . . . . . . • • • 154
48.
aequa lis . • • . . . . . . . • • 240
Neotropic 1.
2•
sp e c i 0 su s ••••••••••• 1 57
49.
minor ••.••••••..•••• 241
3.
diaphanus . . . . . . . . . . . 158
50.
ben i anus • • . . . . . • • . • • 243
4.
n i grorufus ••••...••. 159
51.
i t en e z i . . . • • • • • • . . . . 245
5.
iphias ••....•••••••• 160 bicolor • • • • • • . . • . • . . 163
52.
b i I ob 0 ide s .••••••••• 245
53. 54.
f I uv i at i i i s . . • • . • . . . 248 cuzcanus ••••••.••.•• 249
6. 7.
maximus ••••••••••••• 165
8. 9.
melanocephalus . . . . • . 168
10.
11.
a t I an t i cu s .•.••••••• 172 achterbergi .•.•••••• 174
12.
peruvianus . . • • . • • . . • 174
13. 14.
hymenaeus . . . . . . . . . . . 178 colombianus . . . . . . . . . 180
15. 16. 17.
cascus . . . . . . . . . . . . . . 182 albopictus . . . . . . . . . . 183 amoenus •••••••••••.. 185
18.
pegnai .•.••••••••••• 186
19.
pseudoamoenus •••••.• 188
noyesi •••••••••••••• 170
55. panamensi s . . . • • . . . . . 251 Genus Lonchodry i nus . . . . . . • . . . 253 Palaearctic
1. 2. 3.
ruficornis • • . . • • . • • . 258 nepalensis ..•••••••• 267
Ethiopian
1. 2.
species . . . . . . . 255
subapterus . . • . • • • • . . 256
species • . . • • • . . . 269
seyrigi •••••••••••.• 269 paul iani . . • • • • . • . . . . 271
Oriental
species •.••.••••. 271
1.
sin en sis ••••••••.... 273 Nearctic species . . . . . . . . . . 273
20.
townesi . . . . . . . . . . . . . 190
1.
f I a vu s • • • • . . • . . • . • • • 274
21.
b i I obu s . . . . . . . . . . . . . 191
2.
masneri . . . . . . . . . . . . . 277
Table of contents
bakeri . . . . . . . . . . . . . . 278 3. Neotropic species • . . . . . . . . 282 1. tricolor . . . . . . . . . . . . 282 Australian species . . . . . . . . 284 1. guineensis . . . . . . . . . . 284 2. notogeicus . . . . . . • . . . 286 :;enus Anteon . . . . . . . . . . . . . . . . . 286 Palaearctic species . . . . . . . 290 1. jurineanum . . . . . . . . . . 294 2. brachycerum . . . . . . . . . 300 3. reticulatum . . . . . . . . . 304 4. nepalense . . • . . . . . . . . 304 5. arcuatum . . . . . . . . . . . . 305 6. flavicorne . . . . . . . . . . 309 7. ephippiger . . . . . . . . . . 313 8. pubicorne . . . . . . . . . . . 316 9. scapulare . . . . . . . . . . . 322 10. infectum . . . . . . . . . . . . 325 11. tripartitum . . . . . . . . . 329 12. crassifrons . . . . . . . . . 332 13. gaul I ei . . . . . . . . . . . . . 334 14. fulviventre . . . . . . . . . 337 15. hilare . . . . • . . . . . . . . . 342 16. ingenuum . . . . . . . . . . . . 342 17. corax . . . . . . . . . . . . . . . 345 18. muni tum . . . . . . . . . . . . . 345 19. japonicum . . . . . . . . . . . 347 Ethiopian species . . . . . . . . . 348 1. brachypterum . . . . . . . . 353 2. agile . . . . . . . . . . . . . . . 354 3. maritimum . . . . . • . . . . . 357 4. kasaicum . . . . . . . . . . . . 362 5. fiorii • . . . . . . . . . . . . . 362 6. kivuanum . . . . . . . . . . . . 363 7. taylori . . . . . . . . . . . . . 366 8. dayi . . . . . . . . . . • . . . . • 367 9. canabense . . . . . . . . . . . 370 10. ugandanum . . . . . . . . . . . 372 11. kawandanum • . . . . . . . . . 374 12. afrum . . . . . . . . . . . . . . . 375 13. turneri . . . . . . . . . . . . . 378 14. townesi . . . . . . . . . . . . . 379 15. decellei . . . . . . . . . . . . 380 16. anthereon . . . . . . . . . . . 381 17. namibiense . • . . . . . . . . 383 18. gutturnium . . . . . . . . . . 384 19. arab i cum . . . . . . . . . . . . 387
20. 21 . 22. 23. 24. 25. 26. 27. 28. 29. 30. 31. 32. 33. 34 . 35. 36 . 37. Or i en 1. 2. 3. 4. 5. 6. 7. 8.
III
natalense . . . . . . . . . . . 388 hoyoi . . . . . . . . . . . . . . . 390 mingoae . . . . . . . . . . . . . 391 long i scapum . . . . . . . . . 392 madagascolum . . . . . . . . 393 medleri . . . . . . . . . . . . . 395 hova . . . . . . . . . . . , . . . . 397 compressicoxale . . . . . 398 inf latrix . . . . . . . . . . . 399 rufonigrum . . . . . . . . . . 402 oriphi lum . . . . . . . . . . . 403 fabulosum • . . . . . . . . • . 404 abruptum . . . . . . . . . . . . 404 zairense . . . . . . . . . . . . 406 mo sse Ii . . . . . . . . . . . . . 408 capei . . . . . . . . . . . . . . . 409 eme r i tum . . . . . • . . . . . . 41 a garambanum . . . . . . . . . . 411 t a I sp e c i e s . . • . . . . . . . 41 2 borneanum . . . . . . . . • . . 416 thai . . . . . . . . . . . . . . . . 416 s i I v i co I um . . . . . . . . . . 419 indicum . . . . . . . . . . • . . 419 mysorense . . . . . . . . • • . 422 yasumatsui . . . . . . . . . . 424 sarawaki . . . . . . . . . . . . 425 bengalense . • . . . . . . . . 428 9. k r omb e i n i . . . . . . . . . . . 429 10. mui ri . . . . . . . . . . . . . . . 432 11. laotianum . • . . . . . . . . . 435 12. fyanense . . . . . . . . . . . . 435 13. javanum . . . . . . . . . . . . . 437 14. mun roe i . . . . . . . . . . . . . 439 lankanum . . . . . . . . . . . . 441 15. nemora Ie . . . . . . . . . . . . 442 16. si Ivestre . . . . . . . . . . . 444 17. expol itum . . . . • . . . . . . 444 18. pet ersen i . . . . . . . . . . . 445 19. bauense . . . . . . . . . . . . . 446 20. debi Ie . . . . . . . . . . • . . . 447 21. 22. phi I ippinum • . . . . . . . . 448 Nearct ic species •.•... '" •. 450 1. popenoe i . . . . . . . • . . . . 452 2. arizonense . . . . . . . . . . 455 3. canadense . . . . . . • • . . . 457 4. nebulosum . . . . . . . • . . . 461 5. funestum • . . . . . . . • • • . 462
IV
Mem. Amer.
En!.
6. wasbaueri . . . . . . . . . . . 464 7. minimum • . . . . . . . . . . . . 467 8. masoni . . . . . . . . . . . . . . 469 9. rugosiceps . . . . . . . . . . 471 10. puncticeps . . . . . . '" .474 11. xanthothorax . . . . . . . . 479 12. osborni . . . . . . . . • . . . . 481 13. durangoi . . . . . . . . . . . . 484 14. ciudadi . . . . . . . . . . . . . 486 15. menkei . . . . . . . . . . . . . . 486 16. hirtifrons . . . . . . . . . . 487 Neotropic species . . . . . . . . . . 488 1. clavatum . . . . . . . . . . . . 491 2. mo I Ie . . . . . . . . . . . . . . . 492 3. nigrorubrum . . . . . . . . . 494 4. pan amen se . . . . . . . . . . . 497 5. jamaicanum . . . . . . . . . . 497 6. mexicanum . . . . . . . . . . . 499 7. oliveirai . . . . . . . . . . . 502 8. noyesi . . . . . . . . . . . . . . 503 9 . I ob a tum . . . . . . . . . . . . . 506 10. propodeale . . . . . . . . . . 506 11. chi riquense . . . . . . . . . 508 12. nycteum . . . . . . . . . . . . . 510 13. annul icorne . . . . . . . . . 511 14. albitarse . . . . . . . . . . . 512 15. v i vax . . . . . . . . . . . . . . . 514 16. p iii corn e . . . . . . . . . . . 51 5 17. conterminum . . . . . . . . . 516 pi aumann i . . . . . . . . . . . 517 18. 19. bol ivianum . . . . . . . . . . 518 micros . . . . . . . . . . . . . . 519 20. 21. surinamense . . . . . . . . . 521 Austral ian species . . . . . . . . . 522 1. gi luwense . . . . . . . . . . . 526 2. chelogynoides . . . . . . . 527 myrmecophi lum . . . . . . . 529 3. 4. reticulaticeps . . . . . . 531 au I i cum . . . . . . . . . . . . . 533 5. sedlaceki . . . . . . . . . . . 534 6. 7. anxium . . . . . . . . . . . • . . 536 8. australe . . . . . . . . . . . . 538 9. parvu I um . . . . . . . . . . . . 539 10. orientale . . . . . . . . . . . 541 11. aculeatum . . . . . . . . . . . 545 12. corruptum . . . . . . . . . . . 545 13. miles . . . . . . . . . . . . . . . 547
Ins!.,
no. 37,
1984
14. destructor . . . . . . . . . . 547 15. caledonianum . . . . . . . . 551 16. typicum . . . . . . . . . . . . . 553 17. bougainvi I lei . . . . . . . 555 18. curradoi . . . . . . . . . . . . 556 19. firmum . . . . . . . . . . . . . . 558 20. leiosomum . . . . . . . . . . . 560 21. ni t idum . . . . . . . . . . . . . 561 22. bouceki . . . . . . . . . . . . . 564 23. mundum . . . . . . . . . . . . . . 565 24. bel lator . . . . . . . . . . . . 568 n i gri corne . . . . . . . , .. 568 25. fij ianum . • . . . . . . . . . . 569 26. dub i um . . . . . . . . . . . . . . 570 27. cor i aceum • . . . . . . . . . . 571 28. su ck lin 9 i . . . . . . . . . . . 573 29. bi smarckense . . . . . . . . 575 30. gloriosum . . . . . . . . . . . 575 31 . vulsum . . . . . . . . . . . . . . 576 32. 33. I uctuosum . . . . . . . . . . . 577 34. hagen i . . . . . . . . . . . . . . 578 haustum . . . . . . . . . . . . . 580 35. coral I inum . . . . . . . . . . 581 36. 37. tasmanianum . . . . . . . . . 582 permirum . . . . . . . . . . . . 583 38. involutum . . . . . . . . . . . 584 39. 40. superbum . . . . . . . . . . . . 585 41. rufiscapum . . . . . . . . . . 586 42. gi raul t i . . . . . . . . . . . . 587 Genus Prioranteon . . . . . . . . . . . . 589 Palaearctic species . . . . . . . . 589 1. biroi . . . . . . . . . . . . . . . 591 Ethiopian species . . . . . . . . . . 593 1. richardsi . . . . . . . . . . . 594 2. casalei . . . . . . . . . . . . . 596 Subfami Iy Bocchinae . . . . . . . . . . 599 Genus Bocchoides . . . . . . . . . . . . . 601 Ethiopian species . . . . . . . . . . 602 1. beki Iyensis . . . . . . . . . 602 Genus Chelothel ius . . . . . . . . . . . 604 Palaearctic species . . . . . . . . 605 1. gryps . . . . . . . . . . . . . . . 607 2. berlandi . . . . . . . . . . . . 609 Genus Bocchus . . . . . . . . . . . . . . . . 611 Palaearctic species . . . . . . . . 613 1. ita lieu s . . . . . . . . . . . . 6 1 4 2. umber . . . . . . . . . . . . . . . 614
v
Table of contents
3.
szelenyi . . . . . . . . . . . . 617
1.
4.
slovacus . • • . • • . . . . • • 618
2.
mi rabi lis . . • • . . • . . • . 687 infuscatus • . . . . . . . . . 690
5. 6.
pagl ianoi • • . . . . . . • . . 620 scaramozzinoi . . . . . . . 621
3.
dentatus . . . . . . . . . . . . 690
4.
townesi . . . • . . . . . . . . . 692
7. 8.
scobiolae • . • • . . . • . • • 622 europaeus . . . . . . . . . . . 624
5.
migratorius . . . . . . • . . 694
Oriental
9.
parthicus . . . . . . . . . . . 624
1.
species . . . . . . . . . . . 695
phi I ippinus . . . . . . . . . 697
species . . . . . . . . . . 627
2.
asiaticus . . . . . . . . . . . 697
1.
richardsi . . . . . . . . . . . 628
3.
malayanus . . . . . . . . . . . 699
2.
bini . . . . . . . . . . . . . . . . 629
Nearctic
3.
s ey rig i . • . . . . . . . . . . . 629
1.
:::thiopian
Jriental
species . . . . . . . . . . . 632
species . . . . . . . . . . . 700
perkinsi . . . . . . . . . . . . 700
Neotropic 1.
species . . . . . . . . . . 702
variegatus . . • . . • • . . . 703
1. 2.
laotianus . . . . . . . . . . 635 mu I u en sis . . . . . . . . . . . 635
2.
minimus . . . . . . • . . • . . . 704
3.
pedunculatus •.•••••• 636
3.
macilentus . . . . . . . . . . 706
4.
rubricus . . . . . . . . • . . • 639
'<earctic
species . . . . . . . . . . . 641
1.
mirabilis . • . • . . . . . . . 643
2. 3. 4. 5.
testaceus . . . . . • . . . . . 643 wasbaueri . . . • . . . • . . . 644
a r i z on i cu s . . . . . . . . . . 645
4.
rufus . . . . . . • . . • . . • . . 710
5.
bruchi . . . . • . . . . . . . . . 714
6.
clarus . • . . . • . • • . . • • . 715
Australian
species . . . . . . . . . 717
1.
flavus . . . . . . . . . . . . . . 718
2.
koebelei ••.••••••••• 721
f lavipes . . • . . . . • . . . . 650
Subfami Iy Dryininae . . . . . . . . . . 723
6. 7.
flavicoll is ••••.•••. 650
Genus Dryinus . . . . . . . . . . . . . . . . 728
I at i c ep s . . . . . . . . . . . . 652
8.
dubius • • . . . • • . . . . . . . 653
Palaearctic species . . . . . . . . 729 1. sanderi . . . . . . . . . . . . . 731
species . . . . . . . . . 655
2.
canariensis . . . . . . . . . 734
austral iae . . . . . . . . . . 656
3.
collaris . . . . . . . . . . . . 735
guineensi s ••••..••.. 658
4.
nepalensis . . . . . . . . . . 740
minimus . . . . . . . . . . . • . 658
5.
tarraconensis . . . . . . . 742
6.
koreanus . . . . . . . . . . . . 747
7.
an t i quu s . . • . . . • • . . • • 751
>\ustral ian
1. 2. 3. 4.
robustus . . . . . . . . . . . . 660
5.
bouceki . . • • . . . . • . . . • 661
:=""us Mirodryinus . . . . . . . . . . . . 663 "alaearctic species . . . . . . . . 664 1. gobiensis . . . . . . . . . . . 666
Ethiopian 1.
species . . . . . . . . . . 751
bisulcatus . . . • . . . . . . 754
2.
seyrigi . . . . . . . . . . . . . 756 alticolus . . . . . . . . . . . 758 spangleri . . . . . . . . . . • 762
3.
ungulatus . . . . . . . . . . . 671
3. 4.
4.
atlanticus . . . . . . . . . . 672
5.
deceptor . . . • • . . . . • • . 762
5.
ponomarenkoae . . • . . . . 674
6.
mahensis . . . . . . . . . . . . 764
:=enus Radi imancus . . . • • • • . . . . . 675
7.
sierranus . . . . . . . . . . . 766
species . . . . . . . . 676
8.
pretorianus . . . . . . . . . 767
1. olmii . . . . . . . . . . . . . . . 676 :::enus Mystrophorus . . . . . . . . . . . 678
9.
saussurei . . . . . . . . . . . 770
10.
hova • • • . . . . . . . . • . . . • 772
species . . . . . . . 678
11.
basilewskyi . . . . . . . . . 774
formicaeformis . . . • • . 678 Iy Thaumatodryininae .. 682
12.
chamaeleo . . • • . . . • • • . 775
13. 14.
schoutedeni . . . . . . . . • 778 mayogo . . . . • . . . . • . . . . 780
15.
orophi Ius • • . . . . • . . . • 781
2.
xerophi Ius • • . . . . • . . . 668
Palaearctic
Palaearctic 1. 5'~bfami
:=enus Thaumatodryinus • • • . . . . • 683 Ethiopian
species . . . . . . . . . 686
VI
Mem. Amer. Ent.
16 . mod est us . . . . . . • • . . . . 785 17. i ncertus . . . . . . . . . . . . 785 18. burgeoni . . . . . . . . . . . . 786 19. afer . . . . . . . . . . . . . . . . 787 Oriental species . . . . . . . . . . 791 1. irregularis . . . . . . . . . 794 2. kiefferi . . . . . . . . . . . . 795 3. longipes . . . . . . . . . . . . 799 4. pulcher . . . . . . . . . . . . . 799 5. ex iii s . . . . . . . . . . . . . . 802 6. k r omb e i n i . . . . . . . . . . . 802 7. stantoni . . . . . . . . . . . . 806 8. trifasciatus . . . . . . . . 807 9. latus . . . . . . . . . . . . . . . 808 10. indi cus . . . . . . . . . . . . . 810 11. praeclarus . . . . . . . . . . 814 12. fulvus . . . . . . . . . . . . . . 815 13. lucens . . . . . . . . . . . . . . 817 14. browni . . . . . . . . . . . . . . 817 15. lankanus . . . . . . . . . . . . 821 Nearctic species . . . . . . . . . . 822 1. mexicanus . . . . . . . . . . . 823 2. crawfordi . . . . . . . . . . . 825 americanus . . . . . . . . . . 826 3. alatus . . . . . . . . . . . • . . 828 4. Neotropic species . . . . . . , .. 831 1. constans . . . . . . . • . . . . 833 citricolus . . . . . . . . . . 835 2. f I a von i 9 e r . . . . . . . . . . 838 3. well ingensi s . . . . . . . . 839 4. surinamensi 5 • • • • • • • • 839 5. striatus . . . . . . . . . . . . 845 6. argentinus . . . . . . . . . . 846 7. 8. caraibicus • . . . . . . . . . 849 9. napensi s . . . . . . . . . . . . 850 opaci frons . . . . . . . . . . 851 10. 11. grand is . . . • • . . . . . , .. 852 a I va renganu s . . . . . . . . 857 12. 13. sin op en sis . . . . . . . . . . 858 14. bel i zen sis . . . . . . . . . • 858 anti I leanus . . . . . . . . . 860 15. 16. t eu t on i a e . . • . . . . . . . . 862 17. catarinae . . . • • • . . . . . 864 18. kimseyae . . . . . . . . . . . . 864 19. palaeodominicanus ... 866 Austral ian species . . . . . . . . 868 1. pallidus . . . . . . . . . . . . 871
Inst.,
no. 37,1984
2. speciosus . . . . . . . . . . . 874 3. subtilis . . . . . . . . . . . . 875 4. pacificus . . . . . . . . . . . 876 5. montanus . . . . . . . . . . . . 878 6. p seudophanes . . . . . . . . 878 7. insularis . . . . . . . . . . . 880 8. bouceki . . . . . . . . . . . . . 880 9. k 0 eb e lei . . . . . . . . . . . . 882 10. guineensis . . . . . . . . . . 884 11. gigas . . . . . . . . . . . . . . . 885 12. bi smarcki . . . . . . . . . . . 887 13. aterrimus . . . . . . . . . . . 888 14. biaki . . . . . . . . . . . . . . . 890 15. punctatus . . . . . . . . . . . 893 16. areolatus . . . . . . . . . . . 894 17.leptias . . . . . . . . . . . . . 895 18. quatei . . . . . . . . • . . . . . 897 19. dahmsi . . . . . . . . . . . . . . 897 20. glaber . . • . . . . . . . . . . . 900 21. austral ianus . . . . . . . . 900 22. venator . . . . . . . . . . . . . 903 23. papuanus . . . . . . . . . . . . 904 Genus Richardsidryinus . . . . . . . 907 Palaearctic species . . . . . . . 908 1. albrecht i . • . . . . . . . . . 909 maroccanus . . . . . • • . . . 910 2. corsicus . . . . . . . . . . . . 913 3. Ethiopian species . . . . . . . . . 914 1. e r ra t i cu s . . . . . . . . . . . 91 6 ca r i n i c ep s . . . . . . . . . . 91 8 2. 3. undu I a t us . . . . . . . . . . . 919 Ori ental speci es . . . . . . . . . . 922 1. indianus . . . . . . . . . . . . 922 2. pyri Ilae . . . . . . . . . . . . 925 Nearctic species . . . . . . . . . . 927 1. nearcticus . . . . . . . . . . 927 2. wetmorei . . . . . . . . . . . . 929 3. canadensis . . . . . . . . . . 930 Austral ian species . . . . . . . . 931 1. t r iii n ea t us • . . . . . . . . 932 Genus Tridryinus . . . . . . . . . . . . . 932 Ethiopian species . . . . . . . . . 934 1. ugandanus . . . . . . . . . . . 935 2. ampuliciformis . . . . . . 937 Nearctic species . . . . . . . . . . 940 1. serratus . . . . . . . . . . . . 941 2. testaceus . . • . . . . . . . . 941
VII
Table of contents
oculatus • . . • . . . • . • . . 945 3. Neotropic species •••.•••.• 947 1. maxiexophthalmicus .. 948 2.
ruficauda ..•••••.•.• 951
3.
poecilopterae ••••••• 952
4.
maximus . . • • • • . . . . . • • 954
5 •
s t ria tic ep s ••••••••• 956
6.
ru f i c ep s •••.•.•.•.•. 958
7. 8.
4.
barbarus ••.••••••.• 1018
Nearctic
species .•••.•.•. 1018
1.
inconsul tus . . . . . • . • 1019
2. 3.
do r sa lis ••••.•••••• 1 021 solaris . . . . . . • . • . . • ;f)24
4.
favreauae •••••••••. 1024
Neotropic
species ••.•.•.. 1027
1.
forestalis . • . • . . . • . 1027
nigroflavus ••.••••.. 961
2.
caraibicus .•.•.•.•• 1029
exophthalmicus •••••• 962
3.
ruber • • . . . . . . . . . . • . 1031
9.
quechuanus • . . . . • . • • • 965
4.
aterrimus • . • . • . • . • . 1031
10.
pi cescen s . • . • . • . . • . . 965
11.
andinus ..•.•.•••••.• 967
gibbosus • . • • . • . . • . . • 969 12. Genus Megadryinus .•••••.•.•.• 971
1.
species .•.•.•.•• 1033
amoenus .•.•••••.•.• 1 034
Neotropic
species • . . . . . • . 1037
1.
autumnalis •.•.•.•.• 1041
magnificus ••••.••... 971
2. 3.
bol ivianus . . • . . • • . • 1041 neotropicus •••....• 1044
4.
bicolor •••••••••.•. 1045
Genus Alphadryinus .•••••.•••• 973 Palaearctic
species . . • . . . • 973
1.
b ru e s i ••••••••••..•• 975
2.
balticus • . • . • • . • . . . • 978
Ethiopian
species ••••.•... 980
1. aberrans .•.••••••••• 980 Oriental species . . . • • • . • • . 982 1. asiaticus •••.•.•.••• 982 2.
Nearctic
species ..•.•.••• 971
Neotropic 1.
Genus Perodry i nuS . . • . • . • . . . • 1033
gibbosus . . . • . • . • • . . . 985
Neotropic
Palaearctic 1.
species . • . . . . 1046
zherichini •..•.•••. 1046
Subfami ly Transdryininae . . . . 1048 Genus Transdryinus •••••.•.•• 1048 Australian species . . • . • . . 1048 1.
notogeicus • . . . . • . . . 1048
species • • . • . • . . . 985
Subfami Iy Gonatopodinae ..•.• 1050 Genus Neodryinus • . . . • • . . • . • . 1065
1.
lamellatus •.•.•.•.•• 986
2.
panamensis . . . • . • . • . • 989
3.
parvus • • . • . • . . . . . . • • 989
4. 5.
pegnai . • . . • • • . • . • • . • 991 brasi I ianus . . • • • . . . . 992
6. 7.
rufus •••.••••••.•••• 995 ferrugineus •..•.•••• 995
8.
piceus • . • . • . • • • • . . . . 996
Australian
Genus Cretodryinus • . . . . • . . . • 1046
species ••.••••• 998 1. planus . . • . • . • • . • . • • . 998 Genus Mesodryinus ••.••.•••.• 1000 Palaearctic species .••••. 1001 1. dayi • • . • . • . . • . . . • . . 1003 2. niger •••••••••••••• 1005 Ethiopian species . • . . . . • . 1007 1. aethiopicus ••..•.•• 1008 Or i en t a I sp e c i e s . • . • . • • . • 1 01 1 1. 0 r i en t a lis • . • • . . . • . 1 0 1 1 2. peterseni •••••••••. 1015 3. viet. • . . • • . • . . • . . . • 1016
Ethiopian
species ••••.•.• 1067
1.
an t i quu s . . • . • • • • . • . 1069
2.
gigas • • . • . • . • • . . . . . 1071
Oriental species •.••.•.•. 1073 1.leptopus . . . . . . • . • • . 1076 2.
reticulatus . . • . . • . • 1076
3.
ch e I at us . • . • • . • • . . . 1 079
4.
rubescens . • • • . . • . . . 1081
5.
dolosus . • . • • • • . . • . • 1081
6.
robustus . • . • • . . . . • . 1082
7.
javanus . . • • • . . • . . . . 1085
8.
pseudodiffusus .•••• 1086
9.
diffusus . . . . • • . • • . . 1089
Nearctic species ••.•..•.• 1091 1.
typhlocybae • . . . . . . . 1091
2.
ball i • . • . . • . • • . . . . • 1095
Neotropic
species .•.•..•• 1097
1.
acuticol I is . • . • . . . . 1098
2.
dominicanus •.•..•.• 1101
Mem.
VIII
Amer.
Ent.
Inst.,
no.
37,
1984
albosignatus . . . . . . . 1 101 radialis . . . . . . . . . . . 1104
E t h i op i an
5.
peruvianus •••••.•.• 1107
Oriental
6.
maculicornis . . . . . . . 1107
3. 4.
7.
incaicus . . . . . . . . . . . 1109
8.
brachycerus ••••••.. 11 11
9.
trinitatis . . . . . . . . . 1111
Australian 1.
species •.•.•.. 1115
cheesmani . . . . . . . . . . 1116
2.
nelsoni . . . . . . . . . . . . 1118
3.
koebelei . . . . . . . . . . . 1120
4.
amboni . . . . . . . . . . . . . 1123
Genus Adryinus . . . . . . . . . . . . . . 1126 Ethiopian species . . . . . . . . 1126 1. cerrutii . . . . . . . . • . . 1128
1. 1.
sp e c i e s . . . . . . • . 1 1 77
ugandanus . . . . . . . . . . 1177 species . . . . . . . . . 1180
sinensis . . . . . . . . . . . 1180
Nearctic
species . . . . . . . . . 1180
1.
americanus . . . . . . . . . 1182
2.
mimus . . . . . . . . . . . . . . 1183
Neotropic 1.
species . . . . . . . . 1186
cubensis . . . . . . . . . . . 1188
Austral ian 1.
species . . . . . . . 1189
vitiensis . . . . . . . . . . 1191
Genus Pseudogonatopus . . . . . . . 1191 Palaearctic species . . . . . . 1193
1. 2.
fulgori . . . . . . . . . . . . 1194 camelinus . . . . . . . . . . 1197 rosellae •••...••.•• 1199
2.
nigerianus • . . • • . . . . 1130
3.
3.
bellicosus . . . . . . . . . 1132 owen i . . . • • . . . . . . . . . 1 135
4.
flavifemur . . . . . . . . 1202
5.
ortholabi s . . . . . . . . . 1203
4. 5.
6.
f
species . . . . . . . . . 1138
7.
dromedarius . . . . . . . . 1209
1.
gracilis . . . . . . . . . . . 1138
8.
albosignatus . . . . . • • 1209
2.
meridionalis . . . . . . . 1139
9. 10.
augustae . • • • • . . . . . . 1212 distinctus . . . . . . . . . 1212
1 1. 12.
P r i e sn e r i •••..•.••• 1 2 1 6 septemdentatus . . . . . 1217
rugidorsal is •••••.. 1135
Nearctic
Genus Echthrodelphax • . . . . . . . 1140 Palaearctic species . . . . . . 1140
0
ca r i lei . . . . • • • . . . 1 206
1. 2.
fa i r ch i I d i i . . . . . . . . 1 1 41 ba e t i cu s • . . . . . . . . . . 1 142
3.
longicollis ....•••• 1144
4.
hortusensis . . . . . . . . 1147
1.
pilosus . . . . . . • • . . . . 1222
5.
italicus . . . . . . . . . . . 1150
2.
perkinsiellavorus •• 1224
Ethiopian species . . . . . . . . 1152 1. migratorius . . . . . . . . 1152
3.
seyrigi . . . . . . . . . . . . 1225
4.
polli . . . • • . . . • • . . . . 1226
5.
ruens . . . . . . . . . . . . . . 1228 acutus . . • . . . . . . . • . . 1230
2. afer . . . . . . . . . . . . . . . 1155 3. obscurus . . . • . . . . . . . 1155 Ori enta I speci es • • • . . . . . • 1157 1. ru f us . . . . • . • . . . . . . . 1 1 57 2. fairchildii . . • . . . . . 1158 3. laotianus •...•••••• 1158 Nearctic species . . . . . • . . . 1160 1. krombeini . . . . . . . . . . 1160 Australian species . . . • . • • 1161 1. fa i r ch i I d i i . • • • . . . . 1 1 61 2. nigricoll is . . . • • . . • 1166 bi fasciatus . . . . . . • . 1169 4. pel op s . • . . . . . • • • . . • 1 1 69 Genus Acrodontochelys . . . . . . . 1172 Palaearctic species . . . . . . 1174 1. b ou c ek i . . . . . . . . . . . . 1 1 74
13.ligusticus •••..••.. 1219 Ethiopian
6.
species . . . . . . . . 1221
7 •
sp i ra cu I a r is .•••••. 1 233
8.
similis • • . . . . • . • • . . 1234
Oriental 1.
species . . . . . . . . . 1235
sarawaki . • . • . . • • . • • 1238
2.
sarawakensis . . . . . . . 1241
3.
nudus •.•••.•.••..•• 1245
4. 5.
malesiae . . . . . . . . . . . 1248 validus . . . . • • . . • . • • 1248
6. 7.
lankae • • . . . • • . . . . . . 1249 muiri . . . . . . . . . . . . . . 1251
8.
hospes . . . . • • . . . . . . . 1254
9.
sogatae . . . . . . . . . . . . 1254
10.
pusanus . . . . . . . . . . . . 1255
Nearctic
species .•.•••••• 1257
IX
Table of contents
1. 2. 3. 4. 5.
1258 freytagi canadensis . . . • • • . . . 1260 s joe s t ed t i . . . . . . . . . 1 263 ari zon i cus . . . • . . . . . 1266 ontarianus . . • • . . . . . 1269
6.
stenocrani . . . . . . . . . 1271
7. 8.
au t ox en ob ius .••.••. 1273 magnus . • . . . . . . . . • . • 1275
Neotropic
species . . . . . • . . 1276
1.
variistriatus . • . . . . 1276
2.
flavus . . . . . . . . • . . . . 1277
3.
delphacidi s . . . • • • . . 1279
4.
muesebeck i . . . . . . . . • 1280
5.
morenoi . . . • . . . . . . . . 1284
Australian species •••••.. 1284 1. nigricans . . . . . . . . . . 1287 2.
hospes . . . . . . . . . . • . • 1 290
3.
dichromus . . . . . . . . . . 1293
Neotropic 1.
species . . . . . • • . 1342
longichelatus
1342
Genus Apterodryinus . . . . . . . . • 1342 Palaearctic 1.
species . . . . • • 1344
gobiensis . . . • . • . . . . 1344
Ethiopian
species . . . . . . . . 1344
1.
somerset i . • . . . • • • . . 1345
2.
richardsi . . . . . . . . . • 1346
3.
rufulus • . • . . . . . . . • • 1347
Oriental 1. 2.
species . . . • • • • . . 1348
as i at i cu s . . . . . . . . • • 1 350 insularis . . . • . • • • • . 1350
Nearctic species . . . . • . . . . 1350 1. torvus . . . . . . . . . . . . . 1353 2.
secundus . . . . . . . • . . . 1355
3.
cal ifornicus . . . . . . . 1357
Neotropic
species ••••.••. 1358
1.
citrinus . . . . . . • • . . • 1359 arnaudi • . . . . . . . . • . . 1362
4.
saccharetorum . . . . . . 1294
2.
5.
kurandae •.••••••••. 1296
3 .
t est a c eu s ••••••..•• 1 362
6.
juncetorum . . . . . . . . . 1298
4.
menkei . . . . . • . • . . . • . 1364
7.
plurisulcatus . . . . . . 1299
5.
tijucanus . . . . . . • . . . 1365
8. opacus . . . . . . . . . . . . . 1302 '::;enus Donisthorpina . . . . . . . . . 1304
6. 7.
longicornis . . . . . . • . 1367 bellottii . . . . . . . . . . 1369
Palaearctic 1.
Oriental 1.
species •••..• 1305
pa I I ida . . . . . . • . • . • • 1306 species . . . . . . . . . 1308
asiatica • . . • • • • . . • . 1309
::;enus Agonatopoides . . . . . . . . . 1310 Palaearctic 1.
2. 3.
species •••.•• 1312
felix . . . . . . . . . . • . . 1312 sol idus . . . . . . . . . . . . 1316
canariensis . . . . . . • . 1318 Ethiopian species . . . . . . . . 1321
Austral ian species •••.••• 1371 1. not ogei cus . . . . . . . . . 1372 2.
insularis ••.••••••• 1374
Genus Dicondylus . . . . . • . . . . . . 1374 Palaearctic
dichromus . • • . • • . . . . 1376
2.
bicolor . . . . . . . . . . . . 1378
Ethiopian 1.
Oriental
fuscus • . . . . . . . . . . 1321
2. 3.
acuminatus . . . . . . . . . 1324 cafer . . . . . . . . . . . . . . 1325
Nearctic
4.
ruber . • . • • • . . . . . • • • 1327
Neot rop i c
5.
ceres . . . . . . . . . . . . . . 1328 species . . . . . . . . . 1330
1. 1.
1.
species . • . . . . . . 1383
pallidior . . . . . . . . . . 1383
1.
Oriental
species . . . . . . 1375
1.
species . . . . . . . . . 1384
indianus . . . • . • • • . • . 1384 species . . . . . . . . • 1387
americanus • . . . . . . . . 1388 spec i es •.••••.. 1390
nigrithorax . . . . . . . . 1390
Austral ian
species . . . . . . . 1392
1.
bengalensis . . . . . . . . 1333
2.
borneanus . . . . . . . . • . 1333
2.
rufus . . . . . . . . . . • . . . 1398
3.
insularis . . . . . . . . . . 1334
3. 4.
dubius . . . . . • . • . . • • . 1399 kiefferi . . . . . . . . . . . 1399
5.
primitivus . . . • . . • . . 1401
4. pyri Ilae • . • • . . . . . . . 1336 Nearctic species . . . . . . . . • 1338
1.
perkinsi . . . . . . . . . . • 1394
1.
synchromus . . . • . . • • . 1339
6.
oceanicus . . . . . • . . . . 1403
2.
calidus . . . . . . . . . . . . 1339
7.
alpinus . . . . . . . . . . . . 1404
x
Mem.
Amer.
En!.
Genus Haplogonatopus . • . • . . • . 1408 Palaearct i c
speci es ...••• 1408
1. 2.
apicalis ••.•••••.•• 1409 oratorius . . . . . . . . . . 1409
3•
at ra t us ••..••••.. • '. 1 41 3
Ethiopian
species . . . . . . . . 1415
Ins!.,
no. 37,
1984
Austral ian species . . . . . . . 1473 1. anomalus . . . . . . . . . . . 1473 Genus Trichogonatopus . . . . . . . 1473 Nearctic species . . . . . . . . . 1475 1. simulator . . . . . . . . . . 1475 Neotropic species . . . . . . . . 1475 1.
rich a rd s i • . . . • • . . . . 1 476
species . . . . . . . . . 1416
2.
hi sp i du s •..••••.••• 1478
ap i ca lis .•••••••••. 1 41 7
3.
pall idi tarsi s . . . . . . 1481
species . . . . . . . . 1421
4.
raptor . . . . • • • . . . • • • 1485
1.
insularis . . . . . . . . . . 1423
5.
albomarginatus . . . . . 1485
2.
hernandezae . . . . . . . . 1423
6.
neotropicus . . . • . . • . 1487
Australian species . . . . . . . 1424 1. v i t i en sis . . . . . . . . . . 1 425
7.
rubriceps . . . . . . . ... 1488
1.
katangae . . . . . . . . . . . 1415
Oriental 1.
Neotropic
2.
apicalis . . . . . . . . . . . 1427
Genus Pentagonatopus . . • • . • . • 1427 Nearctic 1.
species . . . . . . . . . 1428
borealis •••...••••. 1428
Genus Epigonatopus • . . . . • . . • • 1490 Austral ian species . . . . . . . 1490 1.
fallax . . . . . . . . . . . . . 1491
2.
sol i tarius . . . . . . . . . 1493 Genus Esagonatopus . . . • • • • . . • 14 93
montanus . . . . . . . . . . . 1430
Nearctic species . . . . . . . . . 1494 1. niger . . . . . . . . . . . . . . 1495
Genus Plectrogonatopoides ... 1431 Palaearctic species . . . . . . 1431
Genus Gonatopus . . • • • . . . • • • . . 1498
1. ch 0 r tin is •...•••••• 1431 Nearctic species . . . . . . . . . 1433 1. nearcticus . . . . . . . . . 1433
Palaearctic species . . . . . . 1502 1. vistosus . . . . . . . . . . . 1508 2. horvathi . . . . . . . . . . . 1509
Austral ian 1.
species . . . . . . . 1429
Genus Tetrodontochelys . . . . . . 1434 Palaearctic
species . . . . . . 1436
2.
perdebilis . . . . . . . . . 1497
3•
dod e r
4.
bi I ineatus .••...••• 1513 distinguendus . . . . . . 1513
0
i .••..••••••• 1 51 1
1.
p ed est r is ..••••••.. 1 437
5.
2.
sakai i . . . . . • • • • • . • . 1441
6.
striatus . . . . . . . . . . . 1518
3.
pulicarius . . . . . . . . . 1442
7.
vollenhoveni
4. 5. 6.
pseudoaudax •••••••. 1446 aegyptiacus . . . . . . . . 1446 audax ..•.••••••••.• 1 447
8. 9 •
iranicus . . . . . . . . . . . 1523 pop ov i • • . . . . • • • • . • . 1 523
10.
a Ib
Ethiopian species . . . . . ... 1449 1. ochreus . . . . . . . . . . . . 1450
11. 12.
suarezi . . . . . . . . . . . . 1528 dimidiatus . . . . . . . . . 1528
2.obscurus . . . . . . . . . . . 1452
13.
atlanticus . . . . . . . . . 1530
3.
14.
uni I ineatus . . . . . . . . 1532
rubripes . . . . . . . . . . . 1454
0
1521
lin ea t us . . . . • • . 1 527
4.
zai rensi s ••.•••...• 1454
15.
aegyptiacus . . . . . . . . 1533
5.
nigrior •.••.••.•••• 1458
16.
plumbeus . . . . . . . . . . . 1535
6.
afer . . . . . • • • • . . . . . . 1458
17.
barbatel Ius .•..•••. 1538
7.
varipes . . . . . . . . . . . . 1459
18.
ater •...•••..•••••• 1538
Oriental species . . . . . . . . . 1461 1. lucens . . . . . . . . . . . . . 1461
19.
I indbergi •••.•..••• 1541 lunatus • . . . • . . . • . . . 1541
20.
species . . . . . . . . . 1463
21 •
t en e r i f e i •.•••••..• 1 548
1.
plesius . . . . . . . . . . . . 1465
canari ensi s .•••••.. 1550
2.
unicus ..••••••••••• 1468
22. 23.
wit t me r i •...••••••• 1 551
3.
pecul iaris . . . . . . . . . 1470
24.
nigrosignatus . . . . . . 1554
Nearctic
Table of contents
25. planiceps •••••••••. 26. campestris •.•....•• 27. subtilis •••..•.•.•. 28. spectrum .••.• •.•• •• 29. bernard i . • . . . • . . . • . 30. formicarius ••.•.••. 31. mongolicus • . . • . . . . . 32. longus . • . . • . • . • . . . . 33. nivosus • . . • . . . . . . • . 34. brunneicollis .•.•.. 35. chersonesius •...••. 36. sepsoides . • . . . . . • • . 37. rhaensis • . . . . . . . . • . 38. barbieri •••.••....• 39. schenklingi . . . • . . . . 40. graecus • . . . • . . . . • . . Ethiopian species . . . . . . . . 1. meridionalis .•.••.. 2. longiusculus ••••... 3. austral is •• .•• ..•.. 4. pilosipes . . . . . . . . . . 5. guigliae ••.••.••••. 6. seyrigi . • . . . . . . . . • . 7. ankaratrus . . . . . . . . . 8. capensis •.••.••.... 9. mo sse Ii . . . • . . . . . . . • 10. rubrithorax . . . . . . . . 11. hyalinus ••.•.•••••. 12. ridens . . . . . -. . . . . . . . 13 • c ommu n is ••..••.•••. 14. luteipes . . . . . . . . . . . 15. bekilyanus . . . . . . . . . 16. incogni tus . . • . . . . . . 17. I eptothorax . . . . . . . . 18. a e t hi op i cu s ..•.••.. 19. tananarivei . . . . . . . . 20 . a rma t us . . . • . • • • . . . . 21. okahandjae . . . . . . . . . 22 . j oh n s i . . . . . . . . . . . . . 23. taylori . . . . . . . . . . . . 24. upembanus . . . . . . . . . . 25. benoi t i • • • . . . . . . . • . 26. trochantericus ...•. 27. opacus . . . . . . . . . . . . . 28. patrizii .••••••••.. 29. turneri . . . . . . . . . . . . 30. ericeti . . . . . . . . . . . .
1554 1556 1558 1560 1562 1564 1567 1567 1570 1570 1572 1573 1582 1585 1587 1587 1587 1590 1590 1593 1595 1596 1598 1601 1601 1 603 1605 1607 1607 16 10 1611 1612 1613 1615 1619 1620 1 623 1623 1 626 1628 1628 1631 1631 1634 1634 1637 1637
XI
Oriental species . . . . . . . . . 1. yasumatsui •..••.•.• 2. attenuatus . . . . . . . . . 3. schenkl ingi . . . . . • . • 4. mysorensis . . . . . . . . . 5. plebeius . . . . . . . . . . . 6. perpolitus . . . . . . . . . 7. javanus . . . . . . . . . . . . 8. cristatus . . . . . . . . . . 9. abjectus . . . . . . . . . . . 10. daunus . . . . . . . . . . . . . 11. phi I ippinus •.••..•• 12. maurus . . . . . . . . . . . . . 13. medius . . . • • . . • • . . . • 14. lucidus . . . . . . . . . . . . 15. magnus . . . • . • . . . • . . . 16. sitae . . . . . . . . . . . . . . 17. indianus . . . . . . . . . . . 18. thai . • • . . • . • . . . . . . . Nearctic species . . . . . . . . . 1. clavicornis . . . . . • . . 2. cyphonotus . . . . . . . . . 3. mayori . . . . . . . . . . . . . 4. portalensis . . . . . . . . 5. translucidus . . . . . . . 6. echo . . . . . . . . . . . . . . . 7. elongatus •••.••••.. 8. freytagi . . . . • . • • • . . 9. contortulus . . . . . . . . 10. planipes . . . . . . . . . . . 1 1. m i mo ide s . . . . . • . . • . . 12. agropyrus . . . . . . . . . . 13. frequens . . . . . . . . . . . 14. areolatus . . . . . . . . . . 15. harpax ••.•••..••... 16. solitarius . . . . . . . . . 17. herbarum . . . . . . . . . . . 18. paraleptias . . . . . . . . 19. pseudochromus . . . . . . 20. flavifrons . . . . . . . . . 21. rap ax .•..••.....••• 22. ashmeadi . • . . . . • • . . . 23.0mb rodes . . . . . . . . . . . 24. moni lis ••..••..•••. 25. argyrias . . . . . . . . . . . 26. curriei . . . . . . . . . . . . 27. innitidus . . . . . . . . . .
1639 1641 1641 1645 1647 1647 1650 1652 1652 1655 1656 1656 1660 1662 1662 1665 1665 1667 1668 1670 1673 1676 1679 1679 1682 1683 1684 1685 1688 1689 1 693 1695 1697 1700 1 703 1703 1704 1709 1712 1712 1715 1715 1720 1723 1724 1724 1 727
XII
Mem.
Amer.
En!.
28.
brooksi . . . . . . . • . • • . 1728
29.
pal I idiceps •••••••. 1729
Neotropic
species .•••.•.. 1730
1.
fernandinae • . . . . . . . 1732
2.
breviforceps .•••••• 1735
Ins!.,
no. 37,
1984
cont ortus . . . • • • . . . . 1768
22. 23.
bol ivianus . • • . . . . . . 1768
24.
autumnalis .•••••••• 1770
Australian
species . . . • . . . 1771
1.
melanias . . • • . . . . . . . 1772 williamsi . . . . . • • • . • 1774
3.
a9ilis • • . . . . • • . . . . • 1737
2.
4.
peruvianus . • . . • . . . . 1738
3.
zealandicus . . . . • • • . 1776
5. 6.
sp i no I a i . . . . . • • • • • • 1 739 wi II inki . . . . . . • • . . . 1783
4. 5.
9i9as •...•••••••••• 1779 p I i ca t us . • . . . . . . . . . 1 783
7.
pseudorbitalis . . . . • 1742
6.
rossi • . • . • . . . . . . . . . 1783
8.
orbitalis .••••••... 1745
7. 8.
naumann i • . . . . . . • • • • 1785 papuanus . . . . . • . . . . . 1786
9.
opacithorax . . . . • • • • 1745
10.
purpurascens •..•••• 1746
1 1. 12.
town e 5 i . . . . . . . . • . . . 1 748 apicalis . . . . . • • • • • • 1750
13. 14.
bartletti ..•.•••••• 1751 flavipes . . . . . • . • • . . 1752
9.optabilis . • . • • • . . . . 1786 10. 11.
decoratus . . . . . • . . . • 1788 austral iae . . . • • . . . • 1790
12.
pulcherrimus ..••••• 1791
Genus Gynochelys . . . . . • . . . • . . 1792 Ethiopian
species . . . . • • . . 1793
15.
campbelli .•••.••••• 1755
16.
tristis . • • • • . . . . • . . 1757
17.
doellojuradoi ••...• 1757
Subfami Iy Apodryininae . . • . . . 1796
18.
si Ivestri i •.••••..• 1759 lacualis . • . . • • . . . . • 1762
Genus Apodryinus .•••••••••.. 1796
1.
braunsi . . . . • . . • • • • • 1793
Neotropic species . . • . • . . • 1796 19. 1. masneri •..••••••••. 1796 20. arlei ••••.•..•••••• 1764 cilipes . • . . • . . • • . . • 1764 21 • Nomina dubia and incertae sedis species • • . . • . . . • . . . . . . . . . . • • . . 1798 Nomina nuda • • • • • • • • • • • • • . • • • • • • • • • • • • . . . . . . . . . • • • • . . . . . • • . • • • • 1829 F0 ss i I
dry i n ids . . . . . . . . . . • • • • . . . . . • . • . . . • • . . . . . . • . . . • • . . • . • . . . 1 830
Parasites of Host
hopper
dryinids . . . . . . • . • • . . . . . . . . . . . • . • . • • • . . . • . • . . . . • . • 1831 records • . . • . . • . • . . • • • . . . . . . . . . . . . • • . . • • . • . . • . . • • • • 1834
Ref e r en c e s . . . • . . . • . . . . . . . . . . . . . . . . • • . . . . . . . . . . . . . . . . . . . . . . . . . . 1 847 Abb rev i at i on s . . . . • • • • • • • • • . . . . • . . • • • • • • • • • • . . . . . . . . • • • • • • • • . . . 1869 Index to hosts . . . • . . . . . . • . • • • . . • . . . . . . . . . . • . . . . . . . . • • . . . . . . . . • 1869 Index to the parasites and predators of dryinids . . . . . . . . . • • • • . 1875 Index
to dryinids . . • • • . • • . • . . . . . . • . . . . . • . . . • • • • . • . . . . . • . . . . . • • l877
A REVISION OF THE
DRYINIDAE
(HYMENOPTERA)
by f.iassimo Olmi
Institute of Plant Protection, Via 5.
Camillo de Lellis,
University of Tuscia, 01100 Viterbo,
Italy
I iHRODUCT I ON
The Dryinidae, one of the smaller fami I ies of the acu leate ,:jymenoptera, is included in the superfamily Chrysidoidea (sensu Day 1977). The
literature on dryinids is wide.
or unrel iable because of
inc'lr rect
Jnrel iable taxonomic descriptions. :roublesome. ~evision
of
::>y Kieffer .J,
c),
a
The more useful
key
.c,ustralian,
however,
(1939),
the family,
is a
to the European
is useless
identifications or
incorrect and
Desides Richards'
monograph of the world species
treatment of the I~earctic dryinids by
a
Oriental
it,
The nomenclature is often
literature on
the British species (1914b),
Much of
or questionable
Fenton
(1918a,
Aphelopus by Jervis (1977), some studies on
and Nearctic dryinids by
R.C.L.
Perkins
(1903,
1906a,
1906b, 1907, 1912), another revision of the British species by J.F. Perkins (1976), a study on the classification of the dryinids by Richards (1953), a -<ey
to
the Russian species by
c!ryinids by rreytag useful
Freytag
(1982).
(1977,
Kieffer's work
because of
Ponomarenko
1978,
1980),
is unreliable for
its completeness.
><earctic species by Ashmead
(1978)
and studies on Nearctic
Freytag and Back
The rudimentary
(1893)
(1977),
identifications,
Giri
but
it
and is
revision of the
is not recommended.
The object of this paper is to revise the species already t.iescribed, to assemble the biological information about them and to describe the new species represented
in
the collections.
world species reasonably Little
The user wi II
find
treatment of the
complete.
is known of the males and the keys are therefore
unsatisfactory.
The classification
association of the sexes this problem.
Only
is based mostly on
the females.
is difficult and much further work
breeding experiments can
finally
The
is needed on
answer these
questions.
MATERIAL EXAMINED
The keys and the descriptions are based on numerous collections.
Those
individuals and
material
examined
institutions from which
in
Mem.
2
Amer.
Ent.
Inst.,
material was borrowed are listed below abbreviation used throughout the text: AD
South Austral ian Museum,
AM
American Museum of Natural
AS
Academia Sinica,
37,
1984
together with
Adelaide
Zoological
no.
the appropriate
(South Austral ia)
History,
New York
Institute,
Peking
(N.Y.)
(China)
B
Bernice P. Bishop Museum,
BA
Museo Argentino de Ciencias Naturales "Bernardino Buenos Aires
Honolulu
(Hawaii) Rivadavia",
(Argentina)
BE
Department of Entomology,
University of Wageningen
BI
Ferdinando Bin collection,
Perugia
BM
British Museum
BN
Zoologisches Forschungsinstitut und Museum Alexander Koenig, (German Federal
(Natural
(Holland)
(Italy)
History),
London
(England)
BO BR
Istituto di Entomologia dell'Universita, Bologna (Italy) Department of Primary Industries, Indooroopilly (Queensland)
BU
Nat iona I Museum of Natura I History,
BY
Department of Bio!ogy,
CA
Cal ifornia Academy of SCiences,
CB
Australian National Entomology,
Bonn
r,epublic)
Internacional
(Hungary)
Waco
San Francisco
Collections of
Canberra
Budapest
Baylor University,
(Texas) (Cal ifornia)
Insects (ANIC),
CSIRO,
Division of
(Austral ia)
CI
Centro
CM
Museum of Comparative Zoology, (Massachusetts)
de Agricultura Tropical,
CO
Zoologisk Museum,
DA
Department of Entomology,
KObenhavn
Cali
(Colombia)
Harvard UniverSity,
Cambridge
(Denmark)
University of Cal ifornia,
Davis
(California) DL
Jiri
DU
National
Dlabola collection,
EB
Institut fur Pflanzenschutzforschung
Museum of
Praha
Ireland,
(Czechoslovakia) Dubl in
Landwirtschaftswissenschaften,
(I reland) der Akademie der
Eberswalde (O.O.R.)
ED
South African Sugar Association (South Africa)
Experiment Station,
EN
Istituto di Entomologia agraria ed Apicoltura dell'Universita, (I taly)
FO
Alessandro Focarile collection,
GE
Museo Civico di Storia i~aturale "Giacomo Doria",
Aosta
GV
Museum d'Histoire Naturelle,
Mount Edgecombe Torino
(Italy) Genova
(Italy)
Geneve (Switzerland)
HA
Martin-Luther-Universitat,
HE
Zoological Museum of the University,
Halle-Wittenberg
(D.D.R.)
HU IF~
Zoologisches Museum an der Humboldt-Universitat, Berlin (O.D.R.) Institut Royal des Sciences Naturelles de Belgique, Bruxelles
IT I(E
Cornell University, Ithaca (I"ew York) College of Agriculture, University of Kentucky,
KY
Entomological
LA
~Jatural
Helsinki
(Finland)
(Belgium)
Laboratory,
Kyushu University
History Museum of Los Angeles County,
(Cal ifornia)
Lexington (Kentucky)
(Japan) Los Angeles
3
Introduction
LE
Rijksmuseum
LI
Linnean Society,
van Natuurlijke Historie,
LN
Zoological
LO
(U.S.S.R. ) Museum d'Histoire naturelle,
LU
Zoological
MA
Department of Entomology,
London
Institute,
U.S.S.R.
Institute,
Leiden
(Holland)
(England)
Lund
Academy of Sciences, Lausanne
Leningrad
(Switzerland)
(Sweden) University of Wisconsin,
Madison
(Wisconsin) MD
Instituto Espanol
ME
National
de Entomologia,
MI
Museo Civico di Storia Naturale,
ML
Milwaukee Public Museum,
Museum of Victoria,
Madrid
Melbourne
(Spain)
(Victoria)
Milano (Italy)
Milwaukee (Wisconsin)
MO
Zoological
MP I: MR
Moscow Palaentological Institute, Moscow (U.S.S.R.) Claud Morley collection (part kept in Ipswich h'luseum,
Museum,
Moscow
(U.S.S.R.) Ipswich,
England) MT
Ecole Nationale Superieure Agronomique,
Montpell ier
MU
M.
(Japan)
Miura collection,
Shimane University
NA
Museo di
ND
Division of Entomology, Delhi
Zoologia dell'Universita,
Napoli
(France)
(Italy)
Indian Agricultural
Research
Institute,
New
(I ndia)
NI
Entomological
NZ
Department of Scientific and
Laboratory,
Kochi
University,
I ndustrial
Nangoku-City (Japan)
Research,
Auckland
(New
Zealand) OL OT
Massimo Olmi collection, Viterbo (Italy) Biosystematics Research Institute, Ottawa
OX
Hope Museum,
Oxford
(Canada)
(england)
P
Museum National
d 'Histoire ,',aturelle,
PG
National
\I'"atural
PH
Academy of r,atural
PK
:,adezdha Ponomarenko collection,
PL PO
Guido Pagl iano collection, Torino (I taly) Istituto di entomologia agraria dell 'Universita,
PT
Facultad de Ciencias Naturales y '''1useo,
QU RY
Queensland r'l,useum, Fortitude Valley (Queensland) Biological and Chemical Research Institute, Rydalmere
Museum
History),
Sciences,
Paris
Praha
Philadelphia r"oscow
(France)
(Czechoslovakia) (Pennsylvania) (U.S.S.R.) Portici
La Plata
(I taly)
(Argentina) (,',ew South
Wales) SA
Friedrich Sander collection,
SC
Department of Food and Agriculture,
SP
Spinola collection,
c/o
Jena
r..1useo
(D.
D.
R.)
Sacramento
f,egionale di
(California)
Scienze Natural i,
Torino
(I taly) ST SU
Naturhistoriska Riksmuseet, Stockholm (Sweden) Instituto de Aclimatacion, Almeria (Spain)
SW
Museo Nacional
SZ
Pier Luigi
TE
Department of Entomology, Station
de Historia Natural,
Scaramozzino collection,
(Texas)
Texas A.
Santiago Torino
(Chi Ie)
(I taly)
and M.
University,
College
4
Mem.
Amer.
Ent.
Inst.,
TN
Museo de Ciencias Naturales,
TO
Istituto e Museo di (I taly)
TR
Museo Regionale di Scienze Naturali,
TU
Instituto de Zoologia,
no.
S.ta Cruz
37,
1984
(Canary
Islands)
Zoologia sistematica dell'Universita,
Torino
Torino (Italy)
Universidad Nacional
de Tucuman,
Tucuman
(Argentina) TV
Musee Royal
TW
American
de I 'Afrique centrale,
UQ
Department of Entomology,
Entomological
Institute,
Tervuren
(Belgium)
Ann Arbor
(Michigan)
University of Queensland,
SI.
Lucia
(Queensland) VE
Museo Civico di Storia Naturale,
VI
Naturhistorischen Museum,
Wien
VR
"Iuseo Civico di Storia Naturale,
WA
National
WS
~jatuurhistorisch
Museum of Natural Museum,
Venezia
(Italy)
(Austria) Verona
History,
(Italy)
Washington
Maastricht
(D.C.)
(Holland)
ZO
Zoolog ische Staatssamm lung,
Munchen
YA
Keizo Yasumatsu collection,
Oonojo-shi
(German Federa I Repub I i c) (Japan)
ACKNOWLEDGMENTS
I have been much indebted to the following persons who helped with the collections, either by sending dryinids on loan, or acting as host when thei r museums were vis i ted: Kees van Achterberg (Le i den), Anders Albrecht (Helsinki), Paul H. Arnaud Jr. (San Francisco, California), Donald Azuma
(Philadelphia,
S. Baldridge (Waco, Texas), (Tervuren), Claude Besuchet Ferdinando Bin (Montpellier), A.J.M.
(Perugia),
G.R.
Brown
Pennsylvania),
Marcos Baez
(Tenerife),
Robert
Tony Bellotti (Cali), P. L. G. Benoit (Geneve), C. F. M. den Bieman (Wageningen),
Zdenek Boucek (Rydalmere),
Carnegie (Mount Edgecombe),
(London),
Ernst-G.
A.
Bournier
Burmeister (Munchen),
Achille Casale
(Torino),
Lennart
Cederholm (Lund), Raul Cortes (Santiago), I talo Currado (Torino), E.C. Dahms (Fortitude Valley), Roy Danielsson (Lund), Michael Day (London), Jean Decelle (Tervuren), (Bruxelles), Dorn
(Halle),
N.
F.
(New Delhi),
(Wien),
Ian Galloway
Delhi),
I
Mark A.
(Davis),
V.
Marjorie Favreau (Indooroopilly),
Antonio Giordani Soika
Karl-Johan Hedqvist (Praha),
(La Plata),
(Maastricht),
Christophe Dufour (Lausanne),
Farooqi (I~ew
Luis de Santis
Dingemans-Bakels
I
(Stockholm),
Jervis
Frank Koch
(Kobenhavn),
Matthews
(Adelaide),
(College Station,
N.
Arnold Menke
Y.),
(Gent),
M.K.
Giri
Steven Krauth F.
Leclant
(Praha),
(Praha),
(Santiago), Max Fischer
(Katmandu), J. Jelinek
(Paris),
(Madison,
(Montpellier),
Lynn S.
Wisconsin), Ole
Lubomir Masner (Ottawa),
(Washington,
Elvira Mingo Perez
D.C.),
(Madrid),
M. S.I.
Swaraj Ghai
(Fukuoka),
Simone Kelner-Pillault
D.C.),
Jan Macek
Texas),
(Venezia),
(Berl in),
(Washington,
Lomholdt
Geeraert
Yoshihiro Hirashima
(Cardiff),
Dessart
Dlabola
Mario Elgueta
(New York, E.
Paul
Jiri
Sammie J.
E.G.
Merritt
Laszlo Moczar
I,ltroduction
(Budapest),
Gordon
(Nangoku-City), O'Connor
(Dubl in),
Ibaraki), cheol
M.
Nishida
Gerald R.
Giuseppe
(Suweon,
Korea),
d'Entreves
(Torino),
L.
(Santiago), Ponomarenko (Pretoria),
(Moscow),
(Verona),
(Oxford),
Pechuman
Scaramozzino
(Ithaca,
(Oxford),
California),
C.N.
Francisco J.
(Ann Arbor,
(Auckland),
(Sydney),
Suarez
Michigan),
Carlo Vidano
Roy R.
Ulrich
(Torino),
Luis E.
(Bonn),
G.L.
Sander
(Gottingen),
(Jena),
Lucia,
Pier Luigi
Brisbane),
Snell ing
Gennaro Viggiani
Henry
Valentine
(Portici),
C.
Vogt
(Cambridge), Annette K. Walker (Auckland), Marius S. V,asbauer (Sacramento, California), Ingrid Wegener (Berlin), V. B. Whitehead Town),
S.C.
(Oonojo-shi, Prof.
Willemstein Japan), Paul
H.
(Leiden),
K.IV.R.
A.
Zwart
Freytag
Willink helpful
in sending
The scann i ng electron photographs were I wish especially disposal
to thank Prof.
the facilities of the
The
(Tucuman),
(Cape
Keizo Yasumatsu
(Wageningen).
has been
problems about Gonatopodinae and
M.J.
(Los Angeles,
(Sapporo),
E..IV.
Pena Nadezdha
Prinsloo
Ritzkowsky
S. Takagi
(Almeria), H.
S.
(St.
Jong-
(Genova),
(Bologna),
(London),
Schneider
(Torino),
Pietro Passerin
New York),
Friedrich VI.
rvrargaret A. Smithers
(Yatabe,
Roberto Poggi
Maria Matilde Principi
(Torino),
James P.
Akio Otake
(Budapest),
(Kobenhavn),
(Buenos Aires),
Toshihiko I,ishioka
Wisconsin),
Guido Pagliano
Jeni:i Papp
Owain Westmacott Richards
Arturo Roig Aisina
Townes
L.
Borge Petersen
Hawaii),
(Milwaukee,
Osella
Christopher O'Toole
Paik
Scobie
(Honolulu,
Noonan
5
in discussing
taxonomic
material. taken
at Pi acenza Un i vers i ty.
Giorgio Domenichini
for putting at my
Institute of Agricultural
Entomology.
illustrations of whole insects were drawn mostly
by Dr.
Falchi, partly by Mr. Luciano Couvert, the remainder by myself. indebted to Dr. Falchi and Mr. Couvert for their figures.
Nicolo I am
BIOLOGY
Most of what
is known of the biology of the Dryinidae
the research of R.C.L. Perkins (1903, and North America, Fenton (1918a, b,
is based on
1905, 1906a, 1906b, 1907) in Hawaii c,) in the U.S.A., Abdul-~
in France, Chandra (1978b, 1980a, 1980b, 1980c) in the Phi I ippines, Barrett, Westdal and Richardson (1965) in Canada, Behrisch (1952) and Strubing
(1956)
in
the German Federal
(1930,
1931,
1932,
1940),
Esaki
and Mochizuki
1975)
and Jervis
1933,
1934,
(1979a,
1935,
(1941)
1979b,
Republic,
1936),
Esaki
in Japan,
1980b,
Esaki
Richards
1980c)
and Hashimoto
and Sameshima (1939),
(1939,
Waloff
(1974,
in Great Britain,
Ponomarenko (1971, 1975b) in the u.s.s.n., Olmi (1976) and Currado and Olmi (1972) in Italy, Lindberg (1950) in Finland. Fragmentary notes have been
added by other workers. The dryinids are all parasitoids of iiolllopter<1 Auchenorrhyncha.
her chelate forelegs, She stings
it,
the female catches an
causing
insensibi I ity,
and
With
adult or nymph of the host.
lays an egg between
its two
Mem.
6
Amer.
Ent.
Inst.,
no.
37,
1984
overlapping thoracic or abdominal sclerites •• She may also bite the hopper with her mandibles (Richards 1939). Only the female of the Aphelopinae does not have chelate foretarsi, her front
and middle pairs of
and must catch and hold the hopper with
legs (Jervis 1980c).
The hopper soon
recovers and runs off. The egg
is usually
invisible.
I t may be regarded as morphologically
external. The larva brown,
is at first
green or yellow
composed of cast off added
to the sac.
invisible,
but within a
sac begins to appear.
larval
skins,
few
days a black,
This spheroidal
dark
sac is
and with each moult an extra piece is
There are normally five moults.
The host remains active,
but does not moult while its parasitoid
is
develop i ng. The more
Aphelopinae is rather different. The egg is much
life history of
internal
and at
least
the first-stage larva
pecul iar hypertrophied mass of host persists until consumed by
the parasite larva is in the
larva must pass
is surrounded by a
(trophamnion).
tissue
its final
through
it.
instar,
The
The
trophamnion
and all
food
larva breaks out of the
trophamnion in its late second instar to project from the host in a sac composed of
larval
skins.
The sac
is cyl indrical
its distal
in other dry in ids.
end
and of a
host,
trophamnion continues to isolate the larva from the host's
the
much softer tex ture than
with
rounded,
I nsi de the
haemocoel.
Just before the larva is fully grown, it eats out all the contents of the host, which usually dies firmly fixed to the food-plant. On completion of feeding, ground. soil.
the
larva splits open
I t moves quite actively
The white cocoon
the sac and crawls out or drops on and pupates on
is of dense silk,
lining. The manner of development of different from all other dryinids in is passed within the external
the host body,
larval
polyembryonic,
sac on
one egg
often with
an
inner closely-fitting
that
the entire
larval
feeding period
the consequent complete absence of
parasitized hosts.
In addition
to produce up
the species
to seventy
young
(Kornhauser 1919). The mature larvae bore their way through abdominal sternites of the host and pupate in the ground. There seems to be ei ther one or two generations a species wich are found the cocoon as mature
in
the the
Crovettia theliae (Gahan) is strikingly
with
dividing
the food-plant or in
temperate regions,
larvae or prepupae.
is
larvae
the
year of the
the winter being passed
in
Some dryinids overwinter as
first instar larvae (Jervis 1980b). Crovettia be in the pupal stage during the winter. I n
theliae (Gahan) is believed to tropical
regions
to be several
overlapping generations without diapause.
the dryinids,
however,
is considerably minimized during
there appear
The activity of the dry season
(Chandra 1980c). Many Authors have recorded
Gonatopodinae) this
in association with
is an example of mimicry,
the occurrence of several ants.
Donisthorpe
(1927)
species
(mostly
suggests that
because dryinids obtain protection from
Introduction
:heir likeness instance,
to the ants.
Anteon myrmecophilum (R.C.L. Perkins), for
appears to be definitely myrmecophi lous.
noppers which
frequently
resemblance
in
its actions:
"if a
vial,
are attended by
appearance to the ants.
four hind
legs,
face
to face,
soliciting food"
dryinids,
ants,
It parasitises
This resemblance
according
in pairs,
(R.C.L.
where ants and Gonatopodini
in a
glass
standing sub-erect on
Perkins 1905). (1939),
is enhanced by
together
stroking one another,
to Richards
leaf-
and bears some sl ight
number of the parasites be bred
they may be seen gathered
mouth,
7
their
licking each other's
In other species of
"it appears more probable that
have been found
together
it
is due merely
to
some overlapping of their habitats". The fema les of the dry i n i ds are parthenogenet i c as well In
(1919)
that mated females produce female progeny
progeny.
According
to Waloff
(1974)
female or male. Many fema I es are predatory feeding hosts
as sexua I.
Crovettia theliae (Gahan) it is assumed by Kornhauser
the case of
purposes only.
Adult
and virgin
females male
parthenogenetic offspring can be
as adu I ts and kill
leaf-hoppers for
Aphelopus, however, are not predacious upon
(Jervis 1980c). The hosts frequently
exhibit changes,
The reproductive organs are often pigmentation, reproduction
often
toward
both
internal
greatly modified,
that of the opposite sex,
usually ceases shortly
and external.
and changes in cuticle occur.
Host
after attack.
HOSTS ATTACKED
The dryinids are all
parasitoids of
fJomoptera Auchenorrhyncha
(Tables 1a,lb). Many published host records the host or the parasitoid was probably
ECONOMIC
IMPORTANCE
From the economic point of view reducing
the hopper popUlation by that a
parasitization.
greater mortality Only
adult
the value of the dryinids in
parasitization
the predaceous habits of the females. opinion
are unreliable because either
misidentified.
Several
is effected
in
is greatly enhanced by
Authors have expressed this way
than
Aphelopus are not predaceous upon hosts.
A number of species of dryinids exert an appreciable effect natural
control
the
by
of their hosts,
which
in
the
is often of considerable economic
importance.
fJaplogonatopus vitiensis R. C. L. Perk ins an d Pseudogona topus hos{Jes R.C.L.
Perkins were unsuccessfully
China,
respectively,
in connection
introduced with
into Hawaii
the biological
from Fiji
control
of the
and
8
Memo
Amero
Ent. Ins!., no.
37,
1984
(])
(])
(]) CO CO CO U U
QJ
u
._
0-
C
ro .- :0
0-
U
(])
co
U ._
~
6
~~~E-~
~ u 0 QJ :J -O:ZOLL
APHELOPINAE
x
x x
Aphelopus Crovettia
x
B I APHELOP I NAE Biaphelopus CONGANTEONINAE Conganteon Chel anteon Fiorianteon
ANTEONINAE
x x x x
Metanteon Deinodryinus
x x
Lonchodryinus Anteon
x x
x
Prioranteon BOCCHINAE
x
x
Bocchoides
Chelothel ius Bacchus
x
Mirodryinus Rad i i mancus Mystrophorus
x x
THAUMATODRYININAE Thaumatodryinus DRYININAE Dryinus Richardsidryinus Tri dry i nus Tabl e lao Hosts af Dryinidae
x xxxxxxxx
xx
xxxx x x x x
x x
x x x
9
Introduction
I
I
I
I
GJ
GJ
ro ro c ,-
GJ
C ,-
'GJ
U
0
ro C
,-
.D
'-
I
»
if)
0. 0
I
U
-0 L
GJ
ro
u 0. v ro » v - ::;;: f- -- roGJ GJ V GJ GJ GJ ro ro ro v v,- v,- v,- uro '-ro ,-
,-
GJ
I~
GJ
-
-
'- L U 0. 0
-
L GJ U ro :J
ro
GJ
V
'ro"
GJ ro ro v ,~I~ V
,,-
v
GJ
c
GJ 0. ro GJ GJ v v o ~ ro ro c v v v '- » ,v ro C ,,o.L ro ro GJI E u u ro '" u x 0 0. ro ro <Jl u ,,u if) '- 0 ,-
0
,-
,-
.D
'"
::;;:
U
U
G5
,-
0
U
~IGJ
GJ
ro
v,-
,-
f- --.J l1..
~egadryi nus
«
-
cr:
c u ,_ ,-
ro
V
L 0.
0
-
01 0
'-
0 01
-
GJ :J 0 l1..
z
x
Alphadryinus x
Mesodry i nus Perodryinus
Ix
Cretodry i nus TRANSDRYININAE Transdryinus
x
GONATOPODINAE Neodryinus
x x x x x
x x x x x
Adryinus Echthrodel phax
I
x
x
Acrodon toche I y s
x x
Pseudogonatopus
x x x
Don i sthorp i na
I
Agon a topo ides
x
Apterodry i nus
x x x
D i condyl us Haplogonatopus Pentagonatopus P lectrogona topoi des
x
Tetrodon tochel ys
I
Tr i chogona top us Epigonatopus Esagona topus
x
Gonatopus
x
Gynochelys APODRYININAE Apodryinus Table lb.
I
Hosts of
Dryinidae
I
I
10
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
Perkinsiella saccharicida K i rka I dy. rlphelopus albopictus Ashmead (= typhlocybae Muesebeck) was
sugprcane leaf-hopper, unsuccessfully with
introduced into New Zealand from
the biological
control
of the yellow apple
the U.S.A. leaf-hopper,
in connection
Typhlocyba
australis Frcgg.
MORPHOLOGY AND TERMINOLOGY
The terminology Reid
(1941)
except
in this work follows
that
that of Richards
(1977)
and
the naming of the wing cells and veins follows
system proposed by Townes (1969). SCULPTURE. The surface of the body
in
Dryinidae
is often sculptured
the in a
variety of ways. The principal terms used in describing the sculpture are listed as follows: without sculpture (smooth); punctate; granulate (Fig. 1); rugose; Eady
(1968)
reticulate rugose is useful
(Fig.
2);
striate
(Fig.
3).
The paper by
in explaining the descriptive terms.
HEAD AND I TS APPENDAGES. The head of Dryinidae is characteri st i ca II y orthognathous with its long axis vertical and the mouthparts directed downwards. head.
The romnoU:-Jrl e'J,c, occupy
the greater part of the side of the
Sometimes numerous hairs arise from between
the head between
the eyes
lower face and upper face.
is termed
face;
it
the facets.
The front of
is made up of the c I ypeus,
The antennae are articulated
in sockets
(antennal sockets) situated very low down, near the clypeus. The antenna I socket is surrounded by a sulcus defining a narrow antennal sclerite. The clypeus includes the greater part of the face below the antenna I sockets. Laterally and dorsally the clypeus is defined by the curved epistomal suture. On each side, near the top of the epistomal suture, is a small pit, the anterior tentorial r'it. The pit marks the point where an arm of the tentorium, the internal skeleton of the head, is invaginated. The ventral margin of the clypeus appears rounded, bidentate or shallowly emarginated as follows: 1) bidentate clypeus: only in females of Dryininae (except for Alphadryinus and jljesodryinus) , Prioranteon (Anteoninae), part of Gonatopus, Trichogonatopus, Apterodryinus, Adryinus, iVeodryinus (Gonatopodinae) ; 2) shallowly emarginated clypeus: in females of Alphadryinus and jljesodryinus (Dryininae), Mirodryinus and l?adiimancus (Bocchinae), Thaumatodryininae, TransdryininiJe,. in males of Dryininae (except for part of Dryinus and Mesodryinus) , Thaumatodryininae, Anteoninae (except for part of Anteon and Lonchodryinus), part of iVeodryinus (Gonatopodinae),. 3) rounded c I ypeus: in fema les of Apodryininae, Aphelopinae, Biaphelopinae, Conganteoninae, Gonatopodinae (except for part of Gonatopus, Trichogonatopus, Apterodryinus, Adryinus, Neodryinus), Anteoninae (except for Prioranteon), Bocchinae (excep t for J1lirodryinus and Radiimancus),. in ma I es of Aphelopinae, Conganteoninae, Bocchinae, part of
"'~,
Fig,
1- Variety
of sculpture:
............
'-"'-~,
granulate
, ,
...... "
(male of Gonatopus lunatus
Klug).
I/i ern.
12
.
Fig.
'
Ent.
I nst.,
no.
37,
1984
, ...
2 -
Variety of sculpture:
oratorius Westwood).
,
Amer.
reticulate rugose
(male of
IJaplogonatopus
13
Introduct i on
Fig.
3 -
Variety of sculpture:
Westwood) .
s triate (female of
Gonatopus s epsoides
14
Mem.
Amer.
En!.
Inst.,
no.
37,
1984
Dryinus and 1\1esodryinus (Dryininae), Gonatopodinae (except for part of Neodryinus) , part of Anteon and Lonchodryinus (Anteoninae). The ocelli I ie near the top of the face; they are three in number. They vary
much
in position;
the distance from the outer edge of a
lateral
ocular-ocellar line, OOL; the distance between the inner edges of the two I atera I ocell i is the postocellar line, POL; the distance from the inner edge of the anterior ocellus to the inner edge of a lateral ocellus is the ocellar line, OL; the distance of the lateral ocell i from the occipital margin is the ocellar-occipital line, OPL. In Apodryininae the ocell i are absent. The labrum ish i nged to the ventra I marg i n of the cl ypeus and is a transverse fl ap. In Dryinidae there is a ocellus to the compound eye
tendency inner)
for
the
is the
I ine of attachment to move upwards on the under (or
side of the clypeus so that much or all
The part of the face under the antennal
lower face.
The epistomal
sclerites but antennal
it
in
labrum
is hidden. as the
suture sometimes runs close to the antenna I
is often well
sclerites
of the
sockets may be known
separated from
the following
them.
It
is close to
the
dryinids:
females of Thaumatodryininae, Lonchodryinus (Anteoninae), Aphelopus (Aphelopinae) ; b) ma I es of Anteoninae and Aphelopus (Aphelopinae); a)
The epistomal
suture
is separated from
the antennal
sclerites in
the
following dryinids: a) fema I es of Dryininae,
Gona topodinae, Anteoninae (except for Lonchodryinus), Conganteoninae, Transdryininae, Biaphelopinae, Crovettia (Aphelopinae), Bocchinae, Apodryininae; b) males of Dryininae, Thaumatodryininae, Gonatopodinae, Bocchinae, Conganteoninae, Crovettia (Aphelopinae); The part of
the face above the antennal
upper face.
It
area between
is possible to define the the antennal
sockets and
sockets may be known as the
frons as the central part of the the median ocellus.
In
various
Dryinidae keel s or impressed lines may be developed on parts of the frons. Sometimes frontal
keels and
I ines are found
to be taxonomically
important.
The main central area of the upper face is often divided by a central keel (the frontal line). In many Dryinidae (males in particular) the frons is more or
less excavated
excavations are the which bears
the
to receive the basal
part of
the antennae;
these
antennal scrobes. The top of the head is the vertex
lateral
ocelli
and
is usually
limited posteriorly
by
the
occipital carina. The occipital carina is absent in the following dryinids: a) females of Transdryininae, part of Gonatopodinae, part of Dryinus (Dryininae), part of Tridryinus (Dryininae), Apodryininae; b) males of Gonatopodinae (except for Echthrodelphax and part of Neodryinus) ; The occipital
carina
is incomplete in
the following
dryinids:
Chelothelius (Bocchinae) , part of Thaumatodryininae, part of Gonatopodinae, part of Tridryinus, Alphadryinus, Mesodryinus, Dryinus (Dryininae) ; b) males of Mesodryinus, i2ichardsidryinus, part of Tridryinus and Dryinus a)
females of
Introduction
(Dryininae),
part of
The occipital
carina
15
Neodryinus (Gonatopodinae); is complete
in
the following
dryinids:
Anteoninae, Conganteoninae, Aphelopinae, Biaphelopinae, Bocchinae (except for Chelothelius), part of Thaumatodryininae, )jegadryinus, Richardsidryinus, Perodryinus, part of Tridryinus, Alphadryinus, fllesodryinus, Dryinus (Dryininae); b) ma I es of Aphelopinae, Anteoninae, Conganteoninae, Bocchinae, Thaumatodryininae, Perodryinus, part of Tridryinus and Dryinus (Dryininae) ; The foramen is an approximately circular gap in the skeleton of the back a)
fema les of
of the head
through
which
thorax.
If the occipital
off from
the
the
carina
internal
organs pass from
is effaced
the vertex
will
the head
to
the
not be marked
occiput which I ies between the occipital carina and the
foramen. The area between the compound eyes and the occipital carina the gena; the upper part of the gena is known as the temple. Viewed
is
dorsally the distance from the posterior edge of a compound eye to the occipital carina is the ocular-occipital line, TL. Sometimes the occipital carina
1s absent
region
between
and
vertex,
occiput
and
gena
are
the bottom of the compound eye and
then
continuous.
The
the articulation of the
mandibles is the malar space. In many Dryinidae a well-marked subocular sulcus (Fig. 4) connects the lower corner of the eye to the mandibular articulation.
The subocular sulcus
is always present
in males.
Only
in
Gonatopodinae, Apodryininae, Conganteon (Conganteoninae) it absent. The mouthparts of Dryinidae are found to be taxonomically important. The mandibles (Fig. 5) stand apart from the labio-maxi Ilary
females of
complex. cocoon, insects.
Perhaps because of their' use for such purposes as opening killing or handling prey, Within
the
they
Dryinidae, however,
"nay be greatly reduced
have not been reduced as
IS
the
in other
the number of mandibular teeth
as follows:
:;onatopodinae: female: 4; male: 3; Jryininae: female: 4; male: 3; Anteoninae: female and male: 4; Conganteoninae: female and male: 3 + 1 smaller; 30cchinae: female and male: 1, 2, 3, 3 + 1 smaller; Transdryininae: female:
1;
male unknown;
.-'iphelopinae: female and male: 3 irregular; 3iaphelopinae: female: 3 + 1 smaller; male unknown; Thaumatodryininae: female and male: 4; .~podryininae:
female:
The max ill ae and
4
irregular;
lab i um are fi rm I y
male unknown; un i ted
to one another by
membranes
labio-maxillary c0:!,,:lex. Each maxilla consists of a ::oasal piece or cardo which articulates with a process of the hypostoma. At and form a
single
tne distal (ventral) end of the cardo a stipes its distal end, the stipes bears the maxillary six segments but frequently
have fewer
is articulated.
Externally
at
palpi which primitively have
in special ized genera or species.
(" e rn . A me r.
16
:: n t.
Inst.,
!l0 .
3 7,
1984
•
•
•
•
f
1P' F
-4 .. -
",,_
Fig.
4 -
. _.
Subocular su lcus
the mandi bu lar art i culation
(S)
connecting
(male of
the
l ower corner of the eye to
Anteon fulviventre
Haliday).
Introduction
- l c,
'of
Fig.
5 -
posterior
Mandible with (female of
four
>
41
,
teeth progressing
Anteon gaullei
17
Kieffer).
larger from anterior on e to
18
Mem.
I nternally and a
Amer.
Ent.
Inst.,
no.
37,
1984
two lobes, a more proximal one, the lacinia, galea. The labium consists of a submentum and The distal end of the prernenturn bears externally the labial
the stipes bears
more distal
one,
the
a prementum. palpi, which characteristically have three segments though the number may be reduced
in particular genera and species.
palp-segments may
be concisely
maxi Ilary palp-segments;
The data as to reduction of
summarised as follows
second
number =
labial
(first number
palp-segments):
Dryininae: female and male: 6/3; ThaUluatodryininae: female and male: 6/3; Anteoninae: female and male: 6/3; Bocchinae: female and male: 6/3; Conganteoninae: female and male: 6/3, 5/3; Transdryininae: female: 6/3; male unknown; Biaphelopinae: female: 6/3, 5/3; male unknown; Aphelopus (Aphelopinae): fema I e and ma Ie: 5/2; Crovettia (Aphelopinae): fema I e and ma Ie: 5/3; Gonatopodinae: Gynochelys: female: 6/3; male unknown; Epigonatopus: female: 2/2; male unknown; Esagonatopus: female and male: 6/2; Gonatopus: female and male: 3/2, 4/2, 4/3, 5/2, 5/3, 6/3; Tetrodontochelys: female and male: 2/2, 3/2, 4/2, 5/2; Trichogonatopus: female: 6/3; male unknown; Haplogonatopus: female and male: 2/1; Dicondylus: female and male: 2/2, 3/2, 4/2; Pentagonatopus: female: 5/2; male unknown; Plectrogonatopoides: female: 5/3; male unknown; Pseudogonatopus: female and male: 2/2, 3/2, 4/2; Donisthorpina: fema Ie and ma Ie: 5/2; Agonatopoides: female and male: 5/3; Apterodryinus: female and male: 6/3; Acrodontochelys: female and male: 2/2, 3/2, 4/2; Adryinus: female: 6/3; male unknown; Echthrodelphax: fema Ie: 3/2, 4/2, 5/2, 5/3, 6/3; rna Ie: 4/2; Neodryinus: female and male: 6/3; Apodryininae: female: 4/3; male unknown. The maxillary
and
labial
palpi
are usually borne on special
stipes and prementum respectively. like a
basal
segment of the palp.
segment of the palpi
really
there can be no doubt that watching a
narcosis)". palpi
In
The question
is a palpiger or a
in the present paper. According seen by
These lobes,
to R.C.L.
it is a
the
as to whether the basal true joint
Perkins
the present paper,
(1905)
true moveable joint
species under the microscope,
is always considered
the first small
a true joint.
lobes of the
palpigers, may look
as
is not discussed "in many forms
(as can easi Iy be
it recovers from
basal
The maxillary
segment of the and
labial
palpi
Introduction
have been
largely used in
generic differentiation. palpal
segments
distinguishing pal pal
variable
segments
in
however,
Moreover,
the
last
labium shows
it now
lobes,
in
the genus.
longer of prime appears that palpi
The
importance in
the number of
except
two segments of the maxillary three central
the number of
the species and
is therefore no
the genera.
that
segments agrees between males and females,
particular, The
Gonatopodinae for
the classification of the
I t now appears,
is usually
number of palpal
19
that
in males
in
may be fused.
the outer ones being
the
paraglossae and the inner one the glossa. The Dryinidae have fi I iform or distally
thickened antennae,
segment or is small
and
with
segments. when
10 segments
the scape forms a
part of the antenna, of various
with
in both sexes.
The first
scape is articulated in a socket. The second segment or pedicel flagellum.
the
The surface of types.
ball
and socket joint for
the antennae
is closely covered
These are sometimes taxonomically
large enough
Thaumatodryininae,
the distal
The flagellum consists of eight with
important,
to be visible without great magnification. the females have tufts of
long hairs on
sense-organs especially In
the antennal
Gonatopodinae the females have longitudinal groove-I ike structures, the rhinaria (Fig. 6), on the distal
segments 5-10. segments. in
In some genera of
The number of distal
the genus
(2,
3,
4 or 5),
segments showing
but constant
in
the rhinaria
the species.
is variable
Rhinaria are
also present in the females of Apodryininae (on antenna I segments 3-10), Transdryininae (on the I as t three antenna I segmen ts) and Dryininae (on the last six antenna I segments: except for fiiesodryinus, which seem to be always without rhinaria). distal
The data as to distribution of rhinaria on
antenna I segments may
be concisely summarised as follows
(within
brackets number of rhinaria per segment):
Gonatopodinae Agonatopoides: 9th (1), 10th (2); Haplogonatopus: 9th (1), 10th (2); Acrodontochelys: no rhinaria; Tetrodontochelys: no rhinaria; Gonatopus: no rhinaria; Dicondylus: 9th (1), 10th (2); Pseudogonatopus (A): 9th (1), 10th (2); (8):
8th
(1),
9th
(1),
10th
(C):
7th
(1),
8th
(1),
9th
(2); (1),
10th
(2);
Trichogonatopus: no rh i nari a; Apterodryinus: 7th (1), 8th (1), 9th (1), 10th (2); Donisthorpina: 9th (1), 10th (2); Esagonatopus: no rhinaria; Plectrogonatopoides: no rhinaria; Echthrodelphax: 9th (1), 10th (2); Adryinus: 6th (1), 7th (1), 8th (1), 9th (1), 10th (2); Neodryinus: 6th (1), 7th (1), 8th (1), 9th (1), 10th (2); Pentagonatopus: 7th (1), 8th (1), 9th (1), 10th (2);
""
•
20
Mem . Amer. En!.
I ns!.,
no. 37,
1984
Gynochel ys : 6th ( 1 ), 7th ( 1 ), 8th (1), 9 th ( 1 ) , 10th (2); Tr an sdryininae Transdryinus : 8th ( 1) , 9th ( 1 ), 10th (2); Dry ininae Dryinus : 5th ( 1 ), 6th (1), 7th ( 1 ), 8 t h ( 1 ), 9th (1), 10 th (2); Richardsidryinus: 5th ( 1 ), 6 th ( 1) , 7th ( 1 ), 8 t h (1) , 9th ( 1 ), 10th (2); Al phadryinus : 5th (1), 6th (1), 7th (1), 8th (1), 9th (1), 10 th (2); Tr i d r y i nus: 5 t h ( 1), 6 t h ( 1), 7 t h ( 1 ), 8 t h ( 1 ), 9 t h ( 1 ), l Ot h ( 2 ) ;
Fig. 6 -
Last antennal segme nt s showing rh in ar i a Haplogo natopus oratorius Westwood).
(fema le of
Perodryinus : 5th ( 1 ) , 6 th ( 1 ) , 7th ( 1 ) , 8 th ( 1 ) , 9th ( 1 ) , 10 t h ( 2) ; Me sod ryinus: n o rh i nar i a j Megadryinu s : 5th ( 1 ) , 6th ( 1 ) , 7th (1 ) , 8th ( 1 ) , 9th ( 1 ) , 10th ( 2) ; Apodryininae Apodryinus : 3th ( 1 ) , 4th ( 1 ) , 5th ( 1 ) , 6 t h ( 1 ) , 7 th ( 1 ), 8 t h ( 1 ) , 9th ( 1 ) , lath THORAX.
( 2) ;
T h e thor ax consists of t h ree segm ent s (pr',ihorax,
mesothorax ,
I ntroduct ion
21
meta thorax} ; the two hinder ones normally bear wings. Some females of Gonatopodinae are apterous. In Gonatopodinae, Anteoninae and Bocchinae, females may be brachypterous or micropterous.
The first
abdominal
segment
(propodeurn) has become firmly attached to the thorax and is connected to the rest of the abdomen by a narrow petiole. Some authors propose the term mesosoma for the thorax + propodeum. Usually the segments of the mesosoma are easily visible. fused
(only
propectus
PROTHORAX.
less
is distinct).
The dorsal
the mesonotum, the
recognizable because the sutures are more or
Apodryininae, however, the mesosoma sclerites are completely
In
sclerite
whi Ie the
lateral
(pronotum) is rather closely attached to sclerites (propleura, composed entirely of
proepisterna) are loosely attached to their notum. The propleura form the prothorax and are termed propectus. They meet
the ven tra I part of along
the median
I ies the small
ventral
I ine.
Quite often hidden between
the first coxae,
prosternum. Anteriorly the propectus supports the head,
sharing this duty with the pronotum; posteriorly it is attached to the first coxae. The pronotum is commonly divided by an anterior transverse furrow into an a
anterior part
rather higher level
(anterior collar) and a more posterior part (disc) at (in
the plane of the mesonotum).
Sometimes a
posterior collar. Gonatopodinae no transverse furrow is visible. I n the ma les of Dryinidae as well as the fema I es of Aphelopinae, Biaphelopinae and Conganteoninae, the p ronotum is very reduced dorsa II y posterior transverse furrow
is also visible,
definining a
In some females of apterous
and
is almost
invisible from above.
This condition
is primitive.
In
the
more specialized females of Dryinidae the pronotum is distinctly visible dorsa II y. It is produced backwards and swoll en into pronotal tubercles. They may
reach back as far as the tegulae or be widely
separated from
Gonatopodinae are the pronotal tubercles absent. In females of the most specialized forms (Dryininae, Gonatopodinae, Transdryininae) the pronotum is remarkably elongated, them.
Only
in
the females of
further forward extension
is caused
of other prothoracic elements is unusually mobile. the prothorax adaptive,
in
by
the dorsal
(propleura
According
to Reid
in particular). (1941)
and
position and development The pronotum
itself
the peculiar modifications of
Gonatopodinae and Dryininae are in a large measure
being concerned
in
the raptorial
habits of the females.
In
Apodryininae the pronotum is fused with the posterior sclerites of the mesosoma. MESOTHORAX.
contains
The second
thoracic segment
the principle flight-muscles.
Only
is usually in
the
largest,
since
it
the apterous females of
Gonatopodinae is it more slender than the other thoracic segments. Dorsa II y the segment is covered by the mesonotum which is divided into two principa I sc I eri tes, the scutum and the scutellum; taxonomi sts often reserve the term mesonotum for the scutum alone. scutum
two sulCi,
the
From the front margin of the
notaulices, often converge backwards, defining a V-
shaped prescutal area. The sulci may be deep and strong and meet at about the centre of the posterior margin of the scutum. In other dryinids they extend back
to the hind margin of the scutum without meeting;
in
22
Mem.
many
groups,
they
are
weak
Amer.
and
very
Gonatopodinae median scutal line
apterous fema I es of
Aphclopinae bisects less
scutum. in
each
and
are the
37,
1984
i ncomp I ete or even
absen t.
the notau I ices are absen t. is often
each
present,
si de of
about
sulcus may
half way
connected
aspect of
anterior,
which
forwards
sternal that
It
very
narrow
are
the
it
scutum.
areas
and
and
They
I a tera I
pleural
to
are absent
regions of
is convenient
to
treat
mesopleuron
is
the
pleural sulcus, lateral
reduced,
the pleural
narrow and wh i ch
articulation Since
sulcus runs
near and
the mesopleuron
the mesopleura.
are partly
visible.
marked off by
a
Just
the
the mesepisternum
an
the
is
tegula
cutting
function
of fl ight,
forms
long
entirely of
anterior
very
iJryininae. marked.
much
while
dorsum of
the
scutellum
to Reid
(1941
be seen
l,
Gonatopodinae,
among there
prothorax,
In
meta thorax
METATHORAX.
Apodryininae and
runs from of
the
subserv ience and
the
It
the bulky,
is formed
reduced. by
In
to
and special ised.
female
the
The
almost
lateral
the prepectus,
stages of thoraces
this
among
various modifications of
is one
this
non-adapta t i ve one.
is
It
region
prepectus. The Gonatopodinae.
from
that
the
be obsol ete or strong I y
This condition
scutellum.
area and
antero-dorsal
visible.
The early
the reduction
the
the
are formed
some phylogenetic significance; of
At
or
is much
suture may
median,
transverse
the mesonotum
in
a
trochantin
the pronotum
importance.
the
the
apterous
the mesothoracic waist
The meso-metap I eura I apterous
is a
modified
The
the prepectus Can
According in
in
from
is
to
and continuous
antero-ventral
rei eased
parallel
parts by
is
epicnemium
developed
hO's assumed considerable
special isation of
area
waist connecting
thorax.
walls of
the
the mesothorax,
slender the
the scutum,
ventral
off an
has become
and
posterior part of
this
the mid
sterno-pleural sutures
trochantinal sulcus. epicnemial sulcus is towards
well
lateral
spec ies the
that
down
particularly
Gonatopodinae
developed
of each mid coxa
is
of
into two
many
in front
it off
apterous
thorax
In
seems
mesepisternum,
which
divided
It
the region prepectus
is
sulcus.
the sulcus cutting corner of
It
sternal sulcus.
vis ib I e
runs
of
the mesepimeron
well
suture, which divides mesosternum is relatively
The
are
The
into an
very
the
the forewings.
together.
divided
posterior,
d i v i ded by beneath
in
the mesothorax
them
the nieso-I~etapleural
and
two more or
the paraps i da I furrows
metapleuron.
a
the
Gonatopodinae. Arising on parapsidal furrows which
upwards from
is
and
l"ing-process
to
with
In
the
scutellar sulcus.
pair of
across the
The scuta I
the
apterous
be a
the mesothorax or
mesepimeron.
coxa
part I y
In
longitudinally
the scu tell um
with
in
large mesepisternum and a
very
obliquely
coincident
sulcus are absent
Gonato;Jodinae. parapsides. The
intimately
reduced
no.
the
the scutal
forward
apterous
lateral
On
is centrally
scutal
side of
extend
so
areas,
axi Ilae and
Axillae
area.
Ins!.,
axillae, which belong morphologically to the The scutellar sulcus runs behind the axi Ilae and, on its central front of the scutellum. The scutal sulcus divides the scutum from
triangular
part, the
a
the prescuta I
En!.
that
may
possibly
the enlargement of may
the scuturn
perhaps
the mesothorax
the
have and
be regarded
is fused
as
with
propodeum.
The metathorax,
which bears
the relatively
small
hind wings,
Introduction
is smaller and segment area,
less differentiated
is covered
usually
Gonatopodinae between
the
a
suture from In
region
metasternum fused
Vv
i th
The
tergite.
the propodeum
to
The
the a
the dorsal
is a
small
muscle of the
Dryinidae
together on
have
each
front edge of
micropterous. is
the
of
the
In
wings When
"hich
articulate
the subfamilies.
'.'eins
and
Irig.
7)
I
text
I
term
in front
of
as
the
inclined
No suture between
Apodryininae
In
the meta thorax
is
to
the first
abdominal
spiracles
lie near the
segment
is articulated
and
above
The
the point
lateral by
margins
its petiole
i nterna I organs where
they
leave
the mesosoma.
a
row
of
wings,
the fore
hooks or
hamuli,
and engaging
Dryinidae, 3-5 wing
the
follow
hamuli
margin
thorax
the
by
is
the
a
system
the
hind margin
some females of
are brachypterous or margin
dorsal margin, the
by
a
of
the
and
series of
axillary sclerites. the tegula.
(1969)
for for
wing
the
large plate,
single characteristics Townes
In
held
arise from
downfolded
is supported by
hind ones,
which
the front
the
means of
hidden
and
are found.
Some females
The membrane
imoortant
with
is outstretched
hind
are normally
.\ ing-veins provide
Two
are absent.
with
sclerites
the
and
and
is
orifice of the propodeum, through which passes petiole. In Apodryininae the propodeum is fused
costal margin, the wing is the apex.
axillary
In
wh ich
separated
metepisternum
into
the metap I euron
the name given
two pairs of
the
is not
the meSQsoma.
ventrally,
the hind wing
the
divided
groove
the metanotum,
doub Ie ba II-and-socket joi n t.
side by
the fore-wing.
:Jryinidae
is visible. is
the other segments of
NII~GS.
not
is what
+ propodeum. The metasternum is a very
abdominal
the petiole somewhat
the propodeum
transverse
wetapleuron
The
is
A
area
apterous
is probab I y
the scutellum
thorax.
second
with
the
two posterior pairs of coxae.
propodeum
attached
to
of
the
and
In
the
single central
This central
defined.
the propodeum.
behind
the other segments of
the
with
with
area
between
well
Gonatopodinae
the meta thorax
of
enter
not
the propodeum.
and metapleura
PROPODEUM. segment
is
Dorsally
consists of a
depressions.
the propodeum
ap terous
fused
the mesothorax. which
postscutellum.
the
the propodeum
the flat
surface of
narrow
with
the
indistinguishably fore
lateral
the metanotum fused
metepimeron. metanotum
by
termed
scutell um and
is more often by
has
than
metanotum.
the
defined
(1939)
Richards
by
23
tip
veins The The
the classification labelling
of
the wing-
cells which they enclose. The fore-wing of the Dryinidae pterostigma, 1-3 basal cells enclosed by pigmented veins, 2 costal vein, the metacarpus joining the pterostigma to the apex, and in SOme species an enclosed radial cell. The radial cell is open when the -adial
the
shows a
vein
is
incornplete.
In I~adiimancus
Moczar
the radial
vein
is
Gonatopodinae, Dryininae, Anteoninae, Bocchinae, -::lauIllatodryininae, Transdryininae have three basal cells (costal, median 3"ld submedian) enclosed by pigmented veins (costa, subcosta, medius, s:1!Jmea'ius, basal, nervulus). COI1ganteoninae and iliaphelopinae have two :casal cells {costal and median} enclosed by pigmented veins (costa, .o:..;:'costa, medius, basal). A;;helopinae have basal cell (costal) enclosed by ::igmented veins (costa and subcosta). The first brachial cell rarely 38sent.
Vvinged
, '"
24
Mem.
appear s
En!.
Ins!.,
no.
37,
1984
Th e male of My strophorus formicaeformis Ruthe has
to be enclosed.
fore-wings reduced,
Amer.
spoon-shaped;
the hind-wings a re absent.
"
Fig.
7 -
Fore-wing of Gonatopodinae : C
submedian cell;
=
ra di al
ve in,"
basal vein;
LEGS.
There
F cs
=
costal
cell;
M
=
median cell;
5 -
= first brachial cell; R = radial cell; t = metacarpus; r = costa," b = subcosta", e = medius", h = submedius", k =
= nervulus.
0
is a pair of
legs for each
thoracic segment.
The
legs consist
of a coxa,
trochanter, femur, tibia and a segmented tarsus. The females of the Dryinidae (except for Aphelopinae and Biaphelopinae) have relatively
elongated fore-coxae.
segments.
The
l ast tarsal
the foot-pad or arolium
T he tarsus always consists of five
segment bears the claws.
(Fig.
8).
Its dorsal
Between the claws
surface contains a
plate,
li es the
orbicula. The f ron t tarsi of most fema l e dryinids (excep t for Aphelopinae and Biaphelop inae) are formed into a chela (Fig. 9) for gripping the homopterous prey.
The chela
is composed of the 5th segment,
and an
enlarged claw. A second claw, the rudimentary claw (Fig. 8), i s reduced (in Thaumatodryininae, Bocchinae, Dryininae, Gonatopodinae) or absent (in Anteoninae, Conganteoninae and Transdryinillae) The trochanter is usua II y a srnall segment. I n the females of Dryininae, Transdryininae, Thaumatodryininae and Gonatopodinae it is very elongated. The femur usua l ly consists of a prox i ma l swol len part {club of femur} and a distal slender part {stalk of femur}. The tibiae are provided with articulated spurs o n
the ventral
apex of the tibia
in
s ide.
Typically,
the following
the
three legs have spurs at the
numbers:
1,
1,
2
(males,
females of
25
Introduction
Aphelopinae, Biaphel opin ae, Conganteoninae, Anteoninae, Thaumatodryininae, some Bocchinae, most of Dryininae ) ; 1, 1, 1 (females of Apodryininae, some Dryininae, most of Bocchinae); 1, 0, 1 (females of Transdryininae, most of Gonatopodinae) or 1, 0, 2 (some females o f Gonatopodinae). The fore-tibia always has only one apical spur. This spur i s mod ifi ed, tarsus,
Fig.
together with
into an
8 -
Arolium
the ve ntral
s i de of the first segment of the
antenna-cleaner ( Fig. 10). The first segment of the fore-
(A)
and rudimentary
claw
(C)
of
Haplogonatopus oratorius
(Westwood) .
(bas itarsus ) may be short or very elo ngated. The modified fifth segmen t of the fore-tarsus con s ist s of a proximal part (basal part, tarsus
d irec tl y distal m ay
art icul ated
part
form a
with
the fourth
proximal
anterior
segmen t of the fore-tarsus ABDOMEN.
and a more or
less
long,
free
(ap ical part) . Segments 2 or 3 (rarely 1) of the fore-tarsus (inner)
fore-tarsus may be short or very bris tles,
segment)
and
hook.
e longa ted.
The fourth The
seg men t o f
the
inner side of the fifth
the enlarged claw may bear
lamellae,
ha i rs or peg-I ike ha i rs. The abdomen consists of
ten
segments,
but this number may
not
~iem.
26
Fig. N ;
9 5th
Arner.
En!.
I ns!.,
Chela o f Anteon g aullei Kieffer t a r s al
segment).
(A ;
no.
37,
1984
aro liu m;
E ;
enlarged claw;
Introduction
be visible without careful of a dorsal
27
study or dissection.
Each
is normally
made up
tergite and a ventral plate or sternite. T he
plate or
....
;'1' ,-
""
j" "
\ ,
j
•
, •
•
,
L F i g.
10 -
Antenna-cleaner of
Jiaplogonatopus or atorius (Westwood) (female):
T = tib i a l spur.
•
spiracular openings
lie at the sides of the
segmen t
into the propodeum wh ic h
is modi fied
tergites.
The first abdominal
is fu nctiona ll y
part of t he
gaster is used f or all the abdominal segments beh in d the propodeum. Some authors propose the term metasoma for t he post-propodeal part of the abdomen. Th e petiole is the narrowed, anterior, mor e or less long stalk. Accord i ng to R ich ards (197 7 ) t he Dryinidae have 8 pairs of thorax.
The
abdom in al
ter m
spiracles.
are normall y
T erg it e 10 is
to which
ov ipos itor, segment
which
is reduced the
only 8 abdom inal Th e ninth
t he seventh
tergites
(7 gastral)
tergite is entire ly
indistinguishably fused
side of the abdomen
The eighth sternite
val vifer,
the female,
developed and v i s i ble.
and depigmented. the vent ral
In
is the
with
retracted
tergite 9.
On
last vis ible sternite.
and membranous and on each s i de is a
first
first valvula is attached. They are parts of the
is adapted for sti nging.
T he sternite of the ninth
is also redu ced and on each s ide has a
second valvifer with the
,
28
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
second valvula attached to its anterior end. The valvifers described as appendage bases. the ovipositor,
forming
immovably joined first The
valvulae.
the
The first
can be
valvulae are the active part of
lancets of the sting.
to one another and form
a
The second valvulae are
sheath
(sting-sheat)
for
the
terebra.
The two pairs of valvulae together constitute the
third valvulae each arise fror,; the posterior end of the second
valvifer.
The genital
orifice
is anterior to the first
longer act as an ovipositor.
The sting and
plates have not so far proved as useful corresponding parts of the male. gastral)
In
are exposed and normally
are available.
The first
next six sternites The external
(up
sternite
in separating
the male developed.
No data on
is not visible as a
to sternite 7 = sixth
last abdominal
species as have the
the first eight
tergites
(7
tergites 9 and
definite sclerite.
gastral)
genital ia of the male are attached
ninth segment.
valvifers which no
the modified
are fully
10
The
developed.
to the hind margin of the
The shape of the ninth sternite may
have taxonomic
importance in Dryininae, Anteoninae and Aphelopinae in separa t i ng species. The male genital ia (Fig. 11) consist of a basal ring, two lateral a median penis (consisting of two penis valves) and two volsellae on the sides of the penis. Each volsella consists of a proximal basivolsella and a d i sta I distivolsella. I n the Gonatopodinae the dorsa I end of each paramere forms an appendage termed the dorsal process of gonoforceps (Fig. 12). I ts shape has taxonomic importance in separating species. In the males of Anteoninae the parameres may bear proximally more or less large dorsal membranous lobes. LENGTH OF THE BODY. The adu I ts of the Dryinidae range in size from 1,00 mm (Gonatopus ashmeadi Kieffer) to 10,93 mm (i'legadryinus magnificus
parameres,
Richards).
Measurements of total
the front of the head (excluding
the an tennae)
to
length from
the apex of the abdomen
the sting).
COLOUR.
Most males are black.
gasters,
but head,
black,
length of adults refer to the
(exc Iud i ng
thorax
Most females
have black
to reddish-black
and propodeum are highly variable
in colour -
reddish or yellowish.
EGG AND POSTEMBRYON I C STAGES. The morpho logy of the egg has been described by Fenton (1918a). The egg appears oval or kidney-shaped and varies .in color from sculpturing on
light yellow
the chorion.
of the egg were added by Westdal,
Richardson
has been described by (1965).
The
larva
third and fourth and serves each molt, usually the oral process
Fenton
(1918a)
is apterous. instars.
It
is
but soon darkens in
the second,
(1961),
process
Westdal,
is present
located near the anterior end the host.
light brown.
pink or green.
third and fourth
The
The first-instar
the
larva
Richardson in
the second,
the
larva
larva is white after
The mature larvae are A pair of fleshy oral
instars.
In
the fifth
lobes are replaced with biting-chewing mouthparts and has disappeared.
is no
Barrett,
The structure of
and Barrett,
in
There
notes on the morphology
Raatikainen
(1980c).
A ventral
to a
but occasionally
grey or dark brown.
fragmentary
(1940),
and Jervis
to anchor the parasitoid
white,
are present
Clausen
(1965)
to dark
Additional
lobes
instar the ventral
larva has a sacciform or
-
Introduct i on
29
•
•
, .
1
"
•
Fig. ring;
,
-
11
- Ma Ie gen i ta I i a of Deinodryinus atriventris (C resson):
N -
penis;
P ;
parameres;
S ;
basivolsella;
V ;
B ;
distivolsella.
basa I
30
Mem.
embryo n ic shape. or U-shaped.
12 -
En!.
third -
T he f i fth -i nstar larva
w i der posteriorly
F i g.
The second - ,
Amer.
I ns!.,
no.
is straigh t ,
and more pointed anteriorly.
N = penis;
1984
and fourth - ins t ar l arvae are curved d i stinctly
segmen t ed,
Other notes on
Male genitalia of Gonatopus lunatus Klug :
gonoforceps;
37,
0 -
the structure
dorsa l
process of
V = dist i vo l sella. •
of the ( 1905) ,
larva were published by Mik Jeanne l
( 19 13),
(1882),
Keilin and Thompson
G i ard
(1889),
( 1915),
Haupt
R.C.L. ( 19 16,
Perk i ns 1932),
Fenton ( 1918b), Maneval (1935), Esaki and Sameshima (1939, 1940), Clausen (1940), Pruthi and Mani ( 1942), L i ndberg (1950), Behrisch (1952), Heik in heimo (1957), Raa t ika i nen ( 1961), Guagliumi (1962), Chaudhary and Ramzan
( 1968),
Ponomarenko
(1975b)
and Jerv i s
(1980c).
Introduction
31
COLLECTION
Best collecting Townes
1972;
of flying
will
be from
Malaise traps
(classical
or
for
Dryinids.
near the ground, collecting
results
instance Townes ' and rVl asner l s models: 1\1alaise 1937; Masner and Goulet 1981). They are used mostly for sampling
modified models,
Pan
i.
traps may
e.
the
methods such
be used
normally
as sweeping or Malaise trapping
communication).
Apterous
Sweeping
recommended for collecting
is not
to sample Dryinids
spectrum of species not
Dryinids are best obtained
oy
active on or
obtained
by
(Masner personal pan
adult Dryinids,
traps. as
the results
are unsatisfactory. Collecting satisfactory Dryinids, obtained rearing
nor by
their hosts.
sweeping.
parasitized
parasitization Chandra's rearing
as
pantrapping
Dryinids
hoppers collected 1980c).
hoppers
Dryinids may
will by
be studied
Chandra
parasitized
or Malaise trapping
the behavior and
For these purposes,
Adult
abilities may
models:
Adult from
methods such
techniques for studying
in
resul ts
be obtained
sweeping. in
Good
glass
best
cages
may
tubes plugged
also be obtained by
wi II
from
be
the field
Biology,
rearing
results
are not
development of by
behavior and
(for
instance
also be obtained with
holding
by
cotton.
cocoons collected
plants. 'ilinged
Dryinids or parasitized
hosts may
be attracted
oy
light
traps. P,c<ESERVATION
Small
Dryinids are best mounted on
are pushed using
through
stainless steel
pinned
pins.
away
to obscure are relaxed genitalia
from
the body
the sides and it
with
is as a
rectangular cards.
and
to open needle.
identify
many males,
prepare genitalia relaxed capsule should be removed
examination
in
Faure
glycerol
20 cc,
and clearing
GOlla topodillae, identify
many
Arabic
and
the
the mouth
a
hooked
w hi ch have very
and
c I ear
dorsal
pin,
genitalia
be
wings
so as not
the
male
be glued it
to
has the
and
in
is essential.
To
The entire male genital
50 cc,
for
dorsa I the
the
specimens
pUllout
Dryinids Can
the
Maceration
Gonato:.JoJinae,
should
have
Vihi Ie the
which
if dropped.
water
recommended
the adul ts should
is recommended;
the specimen
gum 30 g).
the
parts
examination of
(distilled
pins
pinned
antennae positioned
specimen.
Alternatively,
using
are not
males of
easy examination of
legs
they
specimens are required.
liquid
solution
is used,
thorax,
This method of mounting
advantage of protecting To
the
the
underside of
well
hooked
minute stainless steel
Larger Dryinids are best
Whichever method
to one side of the centre of
pulled
added
Plastazote strips.
10
mounted
Chloral
for
microscope
hydrate 50 g,
% potassium
hydroxide
male genitalia of processes of gonoforceps.
volsellae should
process of gonoforceps.
In
be removed many
To for
species of
Dr/ininae, Anteoninae and Aphelopinae the n i n th stern i te shou I d a I so be
32
Mem.
removed from To
Amer.
the male genital
identify females,
En!.
removed
using
Maceration
a
pin
and
in
37,
1984
capsule for microscope examination. are required.
mounted
is essential.
The fore-tarsi
cleared
This method of mounting
the examination of which
To
should be
for microscope examination
10% potassium hydroxide solution
The chela should be dehydrated, examination.
no.
examination of the chela
prepare chelae relaxed specimens balsam.
Ins!.,
in
Canada
is not necessary.
and mounted for microscope
is recommended
is important for
also for the palpi,
identifying females and males of
Gonatopodinae. The palpi should be removed using a hooked pin or two fine needles. 1939)
Richards'
is too complicated
boi ling of the speci men amount of boiling, which
method for removing and in
the mouth-parts
is not recommended.
water or very
the mouth-parts can
weak
potash.
Dryinid
larvae should be preserved
acetic acid
(absolute alcohol
After
be pulled out with
must be made to pierce the gula area and
in Carnoy's fluid
then
is based on the proper
a
fine needle,
tear forwards.
in 90% alcohol
60 ml,
(Richards
This method
after being
chloroform 30 ml
fixed
and glacial
10 ml).
FAMILY SIZE AND DISTRIBUTION
This revision records 844 species grouped
probably comprises approximately distribution of subfami I ies and Krombein,
Hurd,
North of Mexico)
(1979:
Nearctic;
Neo.
=
tables 2a and 2b. names
Catalog of Hymenoptera
in America
Eth.
Ori.
Nea.
Neo.
46 37 5 2
24 22 1 1
22 16 3 3
79 55 1
-
-
2 3 3
-
2
1 7 7
-
-
8 8
-
-
-
6 4 2
-
-
1 1
-
-
-
-
-
-
I
Geographic distribution of Pal.
in
geographical
25 20 1 3
Pal.
2a -
world fauna
The geographical
have been follows.
Anteoninae Anteon Deinodryinus Lonchodryinus Metanteon Prioranteon Aphelopinae Aphelopus Crovettia Apodryininae Apodryinus Biaphelopinae Biaphelopus
per region:
is shown
and Canadian
Smith and Burks
ncmina dubia and nUda)
The actual
1,000 species.
genera
For abbreviations of U.S.
Tab.
(except for
into 46 genera and 10 subfamilies.
=
Palaearctic;
Neotropic;
Aus.
Eth. =
I
Dryinids and = Ethiopian;
Australian.
21
Aus.
44 42 2
14 9 5 1 1
-
-
-
5 5
-
number of known Ori.
=
Oriental;
species
Nea.
=
33
Introduction
Bocchinae Bocchoides Bocchus Chelothelius Mirodryinus Mystrophorus Radiimancus Conganteoninae Chelanteon Conganteon Fiorianteon Dryininae Alphadryinus Cretodryinus Dryinus Megadryinus Mesodryinus Perodryinus Richardsidryinus Tridryinus Gonatopodinae Acrodontochelys Adryinus Agonatopoides Apterodryinus Dicondylus Donisthorpina Echthrodelphax Epigonatopus Esagonatopus Gonatopus Gynochelys Haplogonatopus Neodryinus Pentagonatopus Plectrogona topoides Pseudogonatopus Tetrodontochelys Trichogonatopus Thaumatodryininae Thaumatodryinus Transdryininae Transdryinus Tab.
2b -
Eth.
Ori.
Nea.
Neo.
18
4 1 3
4
8
-
-
-
9 2
-
5
-
1 1 2
-
-
-
1 1 15 2 1 7
2
-
2
Pal.
Neo.
~
~
8
-
1 1
-
-
26 1
23 2
-
-
-
-
15
48 8
-
4
-
-
-
-
2
1
4
-
-
-
3
3 2 67 1 5 5 3 1
4 1 3 3 59 2 2 2 3 1
3 1 2 1 5
3
-
-
-
40
3 -
1 13 6
-
Aus.
Eth. ~
51 1
4 2 1 1 3
-
30 1 1 2
18
-
-
8 7
10 1
-
-
5 5
-
1 9
-
-
-
-
-
-
5 -
-
15
76 1
5
-
-
-
2
Aus.
-
19
Palaearctic;
Neotropic;
4
-
-
Geographic distribution of
per region: Nearctic;
Pal.
1
2 29
-
-
25 1
-
-
19 1 4 4
23
-
1
12 57 1
43 1
-
-
-
-
1 7 1
-
-
-
24
12
2 7 4 2
-
-
2 9
1 4 1
3 1 1
-
8
5 -
3 3
3 1 1 1
-
-
7 6 6
-
-
-
8 1
2 2 1 1
Dryinids and number of known species ~
Ethiopian;
Australian.
Ori.
~
Oriental;
Nea.
~
34
;"em.
Amer.
t<EY TO THE
En!.
Ins!.,
no.
37,
SUBFAi\', I LIE 5 Or=-
1984
DNYldI JA~'
F Er'IALE 5
Front
tarsus not
Front
tarsus
Fig.
13 -
Female of
2 Fore-wing
chelate
chelate
wi th
(Fig.
(Fig.
13).
.2 .3
16)
Aphelopus atratus (Dalman) from Caluso (Italy). only
costal
cell
enclosed by
pigmented
veins
(Fig.
13)
1. AphelopinaeR.C.L. Perkins -
Fore-wing vei ns
with
(F i g.
3 Mesosoma
costal
composed by
completely apparent
and median
cells
14)..................... fused
obliterated,
pronotum and
segments
except
a
for
lateral
pronotum and rnesopleura; -
Mesosoma not pronoturn,
because 4 Chela
or
scutum,
the
partly
composed
metanotum and
rudimentary
rudimentary
claw
claw
mesosoma
of
• 1 O. by
ocelli
15) . . . . . . . . . . . . . . . . .
Apodryininae n. fused
segments
visible
easily
(Fig.
subf. (Fig.
16);
recognizable,
16).
.4
9)
••...•.••••• 5
8)...
• .•..••..••. 6
(Fig.
(Fig.
sutures
suture between apparent
(Fig.
propodeum always
sutures are visible;
wi thout
Chela wi th
only
15);
pigmented
subf.
suture between propectus and
absent
. .•. • . . . • • ••• •. • ••• . • •• . • • . • • , ••• '"
enclosed by
13iaphelo,;;inae n.
(Fig. the
track
ocelli
clearly 2.
35
Introduction
f'i9.
g.
14 -
Female of BiaphelopliS Inasneri
n.
15 ~ Female of ArJoc--{r/inus n,d~neri n.
sp.
sp.
(type)
from
from Godavari
t,lalalcahuello
(I'epal).
(Chile).
36
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
Fig.
16 -
Female of
Conganteon vuic2nicum Benoit (type) from Ruanda.
Fig.
17 -
Fema I e of
Transdryinus notogeicus n. sp.
(type)
from
Queens I and.
Introduction
Fig.
18 -
Female of
37
j'letanteon aerias (Walker) from EI Coigo (Chile).
5 Median leg without tibial spur (tibial spurs 1,O,1)(Fig. 17); occipi tal carina absent • . . . . . • • . . • . • . 8. Tra.rJsdryininae n. subf. -
Median
leg with
tibial
spur
(tibi",1
spurs
1,1,2)(Fig.
18);
occipital
carina complete • . • . . • . . • • • . . . . . . • . . . . 4. Anteoninae ."I.C.L. Perkins 6 Antennae with tufts of long hairs on segments 5-10 (Fig. 19) .•.• ..• • . . .• . . . .••. . • .. . . . . . '" -
Antennae wi thout
7 Median
tufts
leg without
of
•....•.... long hai rs
tibial
spur
Thaumatodryininae n.
6.
(Fig.
(tibial
spurs
-
Median
leg with
tibial
four
posterior
(Fig.
teeth,
at
most
tarsus
(Fig.
5);
(tibial
spurs
progressing
enlarged
with
21)
spur
teeth,
claw
one preapical
1,0,1
1,1,1
larger
(Fig.
21)
tooth;
or
1,0,2)(Fig.
20)
00natopodinae K i ef fer
. . • • • . . . . . . . • • • • . • . • • . . . . • . • • . . . . • • . • . 9. 8 Mandible with
subf.
16) . • • • . . . . • • . . . . . • . . . . . 7
or
1,1,2)(Fig.
from anterior never with a
fifth
segment
16).8
one
to
row of
of
front
always wi th more
than one preapical lamella . . . . . . . . • . • • • . . . • • • • • • . . . . . . • . . . . . • . . . . . . • . . . . 7. Dryininae K i ef f er
-
Mandible with
1-4
large
teetn and a
(Fig.
22);
teeth
(Fi;j.
23)
front
tarsus
(Fig.
wi th veins
quadridentate mandible with
tooth between
winged; costal, (Fig.
23)
has
(Fig.
rarely
to posterior,
row of
fore-wing
pigmented veins
Fully
one
always has a
9 Fully winged; -
22);
the
two posterior
occasionally quadridentate mandible with
larger from anterior
by
(Fig.
rudimentary
strong
teeth
one preapical
with
costal
submedian
in
in
and
24) . . . . . . . . . . . . . . . . . . . . . . . 5.
the
lamella
progressing
enlarged
fifth
segment
cells
winged
cl early
claw of
only . . . . . . . . . . . . . . 9 clearly
Conganteoninae n.
fully
cell s
teeth
this case
and median
16) . . . . . . . . . . 3.
micropterous;
median and
but
three teeth
forms
enclosed
subf. fore-wing
enclosed by oigrnented
lJoccninae ,
38
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
Fig.
19 -
Female of
Thaumatodryinus flavus n. sp. from New Guinea.
Fig.
20 -
Female of
Trichogonatopus neotropicus n. sp. from Brazil.
Introduction
Fig.
Fig.
22 -
21
-
Typical
Varieties of mandibles
39
chela of
in
jjryininae.
Bocchinae and Conganteoninae.
MALES
Fully winged; veins -
(Fig.
fore-wing with
25);
enclosed, 2 Fully by
but
winged;
pigmented
Usually
(Figs
in
this
costal
cell
enclosed
by
pigmented
complete . . . . . . . . . . . . . . . . . . . . . . . . .
costal,
pi grnen t ed ve ins
(F i g.
four
occasionally
case without with
(Fig.
winged,
with
3 Mandible with
26,27);
fore-wing veins
fully
fore-wing
-
only
carina
........... .. . . . . . . . . . . . . . . • 1. ilphelopinae R.C.L. Perkins Usually fully winged, rarely micropterous; fully winged forms wi th fore-wing with costal and 1-2 basal cells clearly enclosed by pigmented veins
-
occipital
costal
26) . . . . . . . . . 3.
only
costal
cel
carina.........
and median
cells
clearly
..2 enclosed
Conganteoninae n. subf.
rarely micropterous; median and
with
occipital
submedian
fully
winged
cells
forms with
clearly
enclosed by
27) . . • . . . . . . . . . • . • • • . . . • • • • • • . . • . • . . • . . • • • • • • 3
teeth progressing
larger
from anterior one
to
posterior (Fig. 5) • . . . . • . • • • . . . . • . • . • • • . • . . • . • . • . . • . . . . . . . . . • • • • • . • 4 Mandible with 1-4 teeth (Fig. 22); quadridentate mandibles with three large
teeth and a
teeth
(Fig.
rudimentary
tooth between
the
two posterior
22) • • • . . . . . . . . . . . • . . • . . . . . . . . . . . . • . . . . • • . • • . • • . • . . . . . • • 5
40
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
/~
/
~ ~~~
/// /
Fig.
Fig.
24 -
23 -
Typical
chela of Bocchinae.
Female of Bacchus uwber n.
sp.
(type).
Introduction
Fig.
25 -
Fig.
Male of
26 -
Crovettia nearctica n. sp. from Portal (Arizona).
Male of
Fiorianteon junonium n. sp. (type).
41
42
Mem.
Fig.
Fig.
27 -
28 -
Male of
iviale of
Amer.
Ent.
Inst.,
no.
37,
1984
Esagonatopus niger (Fenton) from Lexington (Kentucky).
Thaumatodryinus flavus n. sp. from f
4 Fore-wi ng wi th metacarpus as long as or longer than the pt erost i gma (F i g. 28) . . . . . . . . . . . . • . . . . . • . . . . . . • . . . 6. Thaumatodryininae n. subf. -
Fore-wing
with
metacarpus
shorter
than
.................•...•.•............. 5 Mandible with not
1-4
progressing
teeth
larger
(Fig.
22);
the pterostigrna
4.
tridentate mandibles with
from anterior
one
. . • . . . . . • • . . . . • . . . . • . • • . • . . • . . • . • . . • . • 5. -
Mandible with p
0
s t e rio r
three
( rig.
teeth
(Fig.
progressing
29) . . .
Anteoninae R.C.L. Perkins to posterior
(Fig.
teetn 220) •.
ilocchinae R i cha rd s
larger
from anterior
one
to
2 2A ) • . • . . . . . . . • . . • . • . . • • . . . . . . . . . . . • . . • • • . • • • • . . . . • . 6
Introduction
6 Occipital -
carina
complete or
43
incomplete;
dorsal
7.
Occipital
complete,
dorsal
carina absent;
process
of
occasionally
gonoforceps present
(Fig.
•..•.•••. . . • • . • • . • • • • • • • • . . . . . . • . • • • •• 9. The males of Transdryininae,
process
of
Dryininae Kieffer
gonoforceps absent • • • • • • • • • • . . . . . • • • •
but
in
this
case
12) . . . . . . . . . . . . . . . . . . . . .
Gonatopodinae Kieffer
Apodryininae and Biaphelopinae are
unknown.
Fig.
29 -
Male of Anteon
tripartitum Kieffer from Italy.
1. SUBFAr/,ILY APHELOPIN!l8 R.C.L.
PeRKINS
Dryininae partim Kieffer in Kieffer et ~'larshall 1905: 215. Anteoninae partim Kieffer in Kieffel' et Marshall 1906: 510. Aphelopinae R.C.L. Perkins 1912: 7. Aphelopini Kieffer 1914b: 214. Aphelopinae Richards 1939: 278. Aphelopinae Muesebeck and Walkley 1951: 1042. Aphelopinae J. F. Perk i ns 1976: 11. Aphelopinae Krombe in 1979: 1250.
TYPE GC:NUS: Aphelopus Da I man 1823. FEMALE: fully winged; fore-wing with only costal veins;
front
tarsus not chelate;
occipital
cell
enclosed by
carina complete;
pigmented
maxi Ilary palpi
,~',em,
44
5-segmented;
labial
MALE:
fully
veins;
occipital
witn
DISH
with
Ent.
tibial
I nst.,
2-3 segments;
fore-wing
no,
spurs
maxillary
37,
tiClial
with only costal
carina complete;
2-3 segments;
HOSTS:
palpi
winged;
Amer,
cell
palpi
1984
spurs
1,1,2.
enclosecJ
oy
5-segmented;
pigmented
labial
palpi
1,1,2.
worldwide
i',eworacidae, Cicadellidae T/phloc/Jinae.
Gc"E"A: 2 SI"ECIC:S: 42
KEY TO TH:=' GEf\lEi=<:A OF APHELOPLVA2
FEt.~ALE5
Labial large
palpi teeth
radial
vein
with and
2
segments;
1-2
regularly
-
clypeus
I ess broad,
antennal
sci eri tes
curved
fully
wi th (Fig.
MA~~S
mandibles quadridentate,
rudimentary
and C1etanotuf1l usually
AND
teeth
(Fig.
(Fig.
30);
granulatecJ,
epi stomal
22Cl);
head,
not
with
fore-wing
2-3 with
scutum,
scutellum
ret iculate
rugose;
suture cJel1
separated
31A) . • • • . . . . . . • • • • . . . .
1.
from
the
ilphelopus Dalman
Labial
palpi with 3 segments; mandibles tricJentate, witn teeth progressing larger from anterior one to !-,osterior (Fig. 22A); fore-wing with radial vein less regularly curved (Fig. 43); usually distal and proximal part of radial vein forming an angle; at least the
anterior
broader, sclerites
with
surface
of
epistalnal
(Fig.
the
scutum
suture
reticulate
running
close
rugose;
to
313).,.,.,., . . . . . . . . . . . . . . . . . . .
clypeus
the antennal 2.
Crovettia n. gen.
1. Aphelopus Ualman
Aphelopus Oa I man 1823: 8.
Ceraf)hron parti", Katzeburg 1848: 141 (syn. proposed by r
Aphelopinae
45
AtJhelojJus Dalman; r(rombein 1979; 1250.
TYPt:: SPt::C I t::S; FEr!,ALt::
(Fig.
segments; teeth; and
Drylnus a tra tus Da I man 1823, des i 9. by Viestwood 1840. 30);
maxillary palpi
mandibles quadridentate,
fore-wing
metanotuDi
with radial usually
fully
vein
with 5 segments; with 2-3
regularly
granulated,
labial
large teeth cCJrved;
not
head,
reticulate
palpi
and
with 2
1-2 rudimentary
scutum,
rugose;
scutellum
epistornal
suture well separated from the antennal sclerites; antennae distally thickened. w:ALE; structurally resembles the female; antennae not distally thickened. DISTkIBUTIO:,; HOSTS;
worldwide
Clcadelllche Typ/llocyblnae
SPC:C I t::S;
Fig.
35
30 -
Female of Aphelo,)us atratus
(Dalman)
from
Caluso
(Italy).
46
Mem.
NOTES:
Amer.
Ent.
Inst.,
no.
in some species of Aphelopus a medial
prescutal
area,
is often present.
37,
1984
scutal
I ine,
which
bisects the
This characteristic has no specific value.
PALAEARCTIC SPECIES OF APEfELOPU5
Seven species are known.
KEY TO THE SPECIES
FEMALES AND MALES
Head with mandibles, surface white -
Head with
at
2 Notaulices
or
clypeus and a more or
most
mandibles and
reaching at
basivolsella with
one
most
Notaul ices
length
two
0,?5
strong bristles
only mandibles white or
uniformly dark; with -
1-2
frontal
males with
scutum;
inner part
of.the.
32A) . . . . . . . . . . . . . . . . . . . . .
of
scutum;
(Fig.
inner part
of
the
39A) . . . . . . . . . . . . . . . . . . ..
querceus n.
7.
testaceous;
inner part
testaceous . . . . . . . . 3
melaleucus (Da I man)
1.
length
· .. , . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. 3 Head wi th
of
(Fig.
•••••••••.•
reaching beyond
basivolsella with
clypeus white or
0,6
strong bristle
•• ••• ••••• .•••••• • ••• • . '" -
less conspicuous
testaceous • • • • • • . . • • • • • . • • . • • • • . • . . . • • . . . . • • • • • . . • 2
of
sp. clypeus black
or
the basivolsella without
or
strong bristles . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4
Head wi th
clypeus and mandibles white
somet i mes part I y
dark
the basivolsella without 4 Notaulices
reaching
or
testaceous;
(never un i f orml y);
at
or with
most
0,5
ma I es wi th
clypeus inner part
of
1 strong bristle . . . . . . . . . . . . . . . . . . ? length
of
scutum . . . . . . . . . . . . . . . . . . . 5
- Notaulices reaching beyond 0,65 length of scutum . . . . . . . . . . . . . . . . . . . 6 5 Males with penis distally tridentate! (Fig. 33A) •..••••••••••••••• -
• ••••••••.•••••••.•.•••••••••.•••••.. 2. atratus (Da I man) Males with penis distally trumpet-shaped! (Fig. 38A) . . . . . . . . . . . . ..
· . . . . . . . . . . . . . . . • • . . . . . . . . . . . . . . . . . .. 6 Notaulices
reaching beyond
the posterior margin long
0,75
of
the
scutum;
straight
rod
(Fig.
form of
a
narrow,
distally pointed
(Fig.
of
scutum;
sp. sometimes
reaching
males with distivolsella
34A);
basivolsella
in
the
long and
34A) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
• • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • ••
! The females of A. distinguish.
nepalensis n.
6.
length
3.
serratus R i cha rd s
atratus and A. nepaiensis are difficult to
Aphelopinae
-
Notaulices
reaching about
distivolsella differently (Fig.
reaching beyond
the posterior margin
of
straight
width
equal
to,
(Fig.
35A);
or greater
distivolsella
basivolsella -
length
formed
in
0,75
the
of
the
form of
long and narrow,
medially narrowest Fig.
curved point
(Fig.
35C);
distinctly
of
scutum;
epistomal
height a
long
width less
of
than
space at of
sometimes
reaching
its narrowest
clypeus
straight
(Fig. rod
(Fig.
(Fig.
males
34A);
34A) ••••••••••.
serratus Richards scutum;
epistomal
the malar space at medial 4.
point
35A);
height
of
suture
its
clypeus
males with distivolsella and basivolsella differently 36A) . . . . . . . . . . . . . . . . . . . . . . . . . . . .
broad
suture medially
distally pointed 3. of
males with
basivolsella very
nigriceps Kieffer
5.
the malar
than medial
scutum;
37);
length
scutum;
........... ........ .................. Notaulices reaching about 0,65 length
35C);
of
(Fig.
37) . . • . . . • • • . . . • . . . . . . . . • • . • • . .
7 Notaulices
with
0,65
47
(Fig.
formed
camus Richards
1. Aphelopus melaleucus (Dalman)
Gonatopus melaleucus Dalman 1818: 82. Dryinus (Aphelopus) melaleucus (Dalman): Dalman 1823: 14. Aphelopus melaleucus (Dalman): Hal iday 1833: 273. Ceraphron albipes Ratzeburg 1848: 141 (n. syn.); locus typicus Rugen
Island
(Germany).
Aphelopus melaleucus (Dalman): Giard 1889b: 708. Aphelopus albipes Kieffer in Kieffer et Marshall 1905: 217 (n. syn.); locus typicus forest near B i tche and Lemberg (Germany). Aphelopus melaleucus (Dalman): Kieffer in Kieffer et Marshall 1905: 218.
Aphelopus trisulcatus Kieffer 1914b: 216 (n. syn.) ; nomen novum for A. albipes Kieffer. Aphelopus melaleucus (Dalman) : Kieffer 1914b: 217. Aphelopus albipes (Ratzeburg) : Kieffer 1914b: 217. Aphelopus melaleucus (Dalman) : Berland 1928 : 169. Aphelopus melaleucus (Dalman) : Richards 1939: 280. Aphelopus melaleucus (Dalman) : Cavro 1950: 6. Aphelopus melaleucus (Dalman) : J.F. Perkins 1976: 11. Aphelopus melaleucus (Dalman) : Jerv i s 1977: 301. Aphelopus melaleucus (Dalman) : Ponomarenko 1978: 18. Aphelopus melaleucus (Dalman) : Jervis 1979b: 191.
FEMALE: fully winged; length 1,50-2,00 mm; black; mandibles, clypeus, malar space, lower face and a U-shaped mark (embracing bases of antennae with arms brown,
with
lying along
segments 8-10
I ight,
inner margins of eyes) or fully
testaceous;
white;
legs
antennae
testaceous-
48
Mem.
whitish,
Amer.
with hind coxae,
Ent.
Inst.,
no. 37,
1984
hind clubs of femora and hind tibiae partly
brown;
sometimes also mid coxae and mid and fore clubs of femora partly
brown;
antennae distally
proportions:
4:
3;5:
4,5:
thickened; 5:
5:
5:
antennal
4:
4,5:
4:
segments in following 7;
head dull,
smooth,
fully
31 - Head of females of Aphelopus melaleucus (Dalman)(A), Crovettia theliae (Gahan)( B), Biaphelopus masneri n. sp. (C) and Conganteon vulcanicum Benoit (D).
Fig.
granulate; 4;
=
OL
dull,
frontal
2;
line present or absent;
OOL = 2,6; OPL
smooth,
granulate;
=
2;
TL
=
2;
occipital scutum,
notaulices incomplete,
carina complete;
=
POL
scutellum and metanotum
reaching about 0,5 length
of scutum; propodeum reticulate rugose, with 2 longitudinal keels on the posterior surface; median area shiny, less rugose than the part outside the area,
almost smooth,
fore-wings hyaline,
with few
irregular weak keels,
without dark transversal
bands;
without areolae;
tibial
spurs 1,1,2.
MALE: fully winged; length 1,75-2,37 mm; black; mandibles, clypeus and some small spots between the antennal sockets and on the malar space yellowish or whitish; 9-10 light; distally 4,5:
5:
thickened; 5,5:
present; = 2.
antennae brown or black,
legs brown, 6:
sometimes with segments
and fore tibiae testaceous;
antenna I segments in following proportions:
7:
occipital
scutum,
with tarsi
7:
7:
10;
head dull,
carina complete;
smooth,
POL = 7;
scutellum and metanotum dull,
granulate;
OL = 3; smooth,
antennae not 3,5:
frontal
OOL = 4; granulate;
4: line
OPL = 2;
TL
notaulices
incomplete, reaching about 0,5 length of scutum; sometimes median scutal line present; propodeum reticulate rugose with 2 longitudinal keels on the posterior surface; the area, hyaline, (Fig.
median area shiny,
almost smooth,
with few
without dark transversal
32B)
less rugose than the parts outside
weak
irregular keels;
bands;
posteriorly strongly emarginated;
with 1 strong bristle (Fig.
32A);
32A); tibial spurs 1,1,2. LOCUS TYPICUS: Vastergotland
fore-wings
ninth sternite of the abdomen inner part of the basivolsella
apex of the penis not
tridentate (Fig.
(Sweden)
of G. melaleucus in STj types of C. albipes Ratzeburg and A. albipes Kieffer (= A. trisulcatus) probably lost. HOSTS: in England Edwardsiana crataegi (Edw.){Richards 1939; J.F. TYPICAL MATERIAL:
holotype F!
Aphelopinae
49
Perkins 1976); according to Jervis (1980c): Fagocyba carri and cruenta, Ossiannilssonola callosa, Edwardsiana avellanae, flavescens, geometrica, hippocastani, plabeja and rosae, Ribautiana ulmi, Typhlocyba quercus, Alnetoidia alneti; in France Edwardsiana hippocastani (Edw.) (Giard
B
Fig. 32 - Male of Aphelopus melaleucus (Dalman) from Vastergotland (Sweden): A: genitalia (viewed ventrally); B: ninth abdominal sternite.
1889b), Typhlocyba douglasi Edw. (Giard 1889b), Ribautiana ulmi (l.) (G i ard 1889b), Edwardsiana rosae; in Germany Fagocyba cruenta var.
douglasi (Edw.); in U.S.S.R. Typhlocyba douglasi Edw., Edwardsiana hippocastani (Edw.), Ribautiana ulmi (l.) (Ponomarenko 1978). BIOLOGY:
informations in Jervis
(1979b,
1980b,
species; overwintering stage: prepupa. DISTRIBUTION: Palaearctic region: U.S.S.R.
c);
in England bivoltine
(Ponomarcnko 1978); FINLAND:
Aminne (Ab, Pojo), HE! Dyrehaven (Sjall.), HE! SWEDEN: Vastergotland, ST! Skane, TW! Messaure, TW! Ol!; HUNGARY: Ugod, BU! Nagyvisnyo (BUkk), BU! Szigetsztmiklos, BU! SzSztMiklos, BU! Nagymezo, BU! Visegrad, BU! Oserdo (BUkk), BU! Szar-Fanniv., BU! Mecsek hgys., BU! Nagykovacsi,
50
BU!
Mem.
Fenyofo,
BU!
Amer.
Matra,
BU!
Ent.
Csakvar,
Inst.,
BU!
no.
37,
Budapest,
1984
BU!
Vertesh.
(Fanier
vgy.), BU! ROMANIA: Ba Victoria (Nagy 1967b; 1972); CZECHOSLOVAKIA (Ogloblin 1924b); YUGOSLAVIA: Kastel Novi (Dalmacija), BU! Prespa Geul (Otesevo,
Makedonija),
BM!
GERMANY:
Bot.
Garten
(Berl in),
HU!
Nymphenburg (MUnchen), EB! Hamburg, HE! Rhondorfer Tai (Siebengebirge), OT! Lohrberg-Fuss (Siebengebirge), OT! RUgen Island (Ratzeburg 1848); AUSTRIA: Huben (0 Tirol), HU! Wien, BU! Laaerb, P! DENMARK: Almindingen (Bornholm), CO! Espe, CO! Korsor, CO! Vicmou, CO! Bcrumaa, CO! Basnaes, CO! Grevind Scov, CO! Langeland I., CO! HOLLAND: Putten (Gld.), LE! Wijster (Dr.), BM! BELGIUM (Crevecoeur et Marechal 1933); SWITZERLAND: Schuls (Vulpera), GV! Epalinges (Vaud), LO! Martigny (Valais), GV! Engadine Nat. Park (Grigioni), BM! FRANCE: Montpellier (Herault), P! Foret de Clenay (Cote-d'Or), P! Maisons-Laffitte, P! Wimereux (Pas-de-Calais) (Berland 1928); Le Puy, P! Foret de Fontainebleau (Seine-et-Marne), P! Lille (Nord), P! Huelgoat (Fl.), BM! Meudon (Seine)(Berland 1928); Foret de Dabo (Moselle), VI! ENGLAND: Minsmere Reserve (Suffolk), BM! Spratton (Northants), BM! Oxford, BM! TW! Wight Island, GE! Dawlish Warren (Devon), OT! Leicester, TW! Esher Common (Surrey), BM! Bix (Henley, Oxfordshire), BM! Hounslow (Mddx.), BM! Burnham Beeches (Bucks.), BM! Dorking (Surrey), BM! OL! Hanworth (Mddx.), BM! Bald Hill (Lewknor, Oxfordshire), BM! Bedford Park (London), BM! Crave I Pits (Theale, Berks.) BM! Horsley (Surrey), BM! Radcliffe-on-Trent (Notts.), BM! Beaconsfield (Bucks.), BM! Epping Forest, B! numerous other localities in Richards 1939; SCOTLAND: Beinn Eighe Forest (R.W.), BM! Tentsmuir Nat. Res. (Fife), BM! Aviemore (EI.), OL! other localities in Richards 1939; IRELAND: Donaghadee (Co. Down), BM! Agher (Co. Meath), BM! other localities in Richards 1939; SPAIN: Valladolid (Ceballos 1959); ITALY: Mt. Bracco (Barge, Cuneo), OL! Oropa (Vercelli), GE! Pizzighettone (Cremona), TW! Fori) (Zangheri P., 1969); CYPRUS: Kilani-Krios R., BM! Pera Pedi-Krios R., BM! LEBANON: Becharre, BM!
Nearctic region
(error):
CANADA:
Port Hope (Ottawa)(Kieffer 1914b).
2. Aphelopus atratus (Dalman)
Dryinus (Aphelopus) atratus Dalman 1823: 15. Aphelopus atratus (Dalman): Hal iday 1833: 273. Aphelopus melalecus (Dalman) var. atratus (Dalman): Kieffer in Kieffer et Marshall
1905: 219.
Aphelopus atratus (Dalman): Kieffer 1914b: 218. Aphelopus holomelas Richards 1939: 289 (n. syn.); locus typicus Lustleigh
(England).
Aphelopus holomelas Richards: J. F. Perkins 1976: 11, 12. Aphelopus holomelas Richards: Jervis 1977: 301.
Aphelopinae
FEMALE:
fully
antennae fully legs fully
winged; black,
black;
segment
mandibles testaceous;
1 or segments 1-2 testaceous;
sometimes with hind coxae and hind clubs of femora
partly brown or only thickened;
length 1,5-2,0 mm; sometimes with
testaceous,
51
with clubs of hind femora brown;
antenna I segments
in following proportions:
antennae distally 5:
4,5:
4,5:
6:
6:
6: 5: 5: 5: 8; head dull, fully granulate; frontal line present; occipital carina complete; POL = 7; OL = 3; OOL = 4; OPL = 4; TL = 3; scutum, scutellum and metanotum dull, about 0,5 lengt of scutum; reticulate rugose, median area rugose, MALE:
fully
proportions:
weak
bands;
notaulices scutal
keels on
length
5:
6:
propodeum
not reticulate without dark
spurs 1,1,2. 1,50-2,25 mm;
black or brown; 4:
reaching
the posterior surface;
fore-wings hyaline, black;
legs brown,
with
antennae not distally thickened; 4:
incomplete, line present;
the parts outside the area,
irregular keels;
tibial
winged;
antennae fully testaceous;
with 2 longitudinal
less rugose than
with few
transversal
granulate;
sometimes median
6,5:
7:
7:
8:
8:
mandibles testaceous; tarsi
and fore tibiae
antenna I segments in following
12;
head dull,
granulate;
frontal
line present or absent; occipital carina complete; POL = 8; OL = 5; OOL 5; OPL = 3; TL = 3; scutum, scutellum and metanotum dull, granulate; notaul ices incomplete,
reaching
about 0,5 length of scutum;
sometimes
B
Fig.
33 -
Male of
Aphelopus atratus abdominal
median scutal longitudinal
line present;
(Dalman):
sternite
genitalia
(A)
and ninth
(B).
propodeum reticulate rugose with 2
keels on the posterior surface;
median area smooth,
less
rugose than the parts outs i de the area, wi th few weak i rregu I ar keel s; fore-wings hyaline, without dark transversal bands; ninth sternite of the abdomen not posteriorly emarginated
(Fig.
33B);
apex of the penis
52
Mem.
tridentate (Fig.
(Fig.
33A);
33A);
tibial
lOCUS TYPICUS:
Amer.
Ent.
Inst.,
no.
37,
1984
inner part of the basivolsella with 2 strong bristles
spurs 1,1,2.
Vastergotland
TYPICAL MATERIAL:
(Sweden)
lectotype M!
and 1 paralectotype M!
(hereby
of D. atratus in ST; holotype M, allotype F and 66 paratypes (29 FF, 37 MM) of A.holomelas in BM; 6 para types (4 MM, 2 FF)! of A. holomelas in OX; 1 para type F! of A. holomelas in DU. HOSTS: in England Typhlocyba opaca (Richards 1939; J.F. Perkins 1976); accord i ng to Jerv i s (1980c) Alebra albostriella and wahlbergi, Fagocyba cruenta, Edwardsiana hippocastani, Ribautiana tenerrima and ulmi, Typhlocyba bifasciata and quercus, Eupteryx aurata, cyclops, melissae and urticae, Zygina sp.; in I ta I y Eupteryx stachydearum, Typhlocyba rosae and Typhlocyba froggatti; in France Typhlocyba rosae. designated:
BIOLOGY:
to melaleucus)
the para lectotype actually belonging
informations in Jervis
overwintering
stage:
DISTRIBUTION:
(1980b,
c):
in England bivoltine species;
prepupa.
Palaearctic region:
FINLAND
(Hellen
1953);
SWEDEN:
Degaberga (Sk.), BM! Skane, TW! Ol! Vastergotland, ST! HUNGARY: Szod, BU! ReyfUlop, BU! Vacz (Tudosdomb), BU! Kelebia, BU! SzSzT Miklos, BU! Gezahaza
(Bakony),
ROMANIA
(Nagy
BU!
1972);
Fej ervarcsurgo,
YUGOSLAVIA:
Bosna),
BU!
Postojne (Slovenija),
a. Rh.,
OT!
AUSTRIA:
(Wien),
P!
BM!
Semmeringgebeit
Eisenzicken,
VI!
DENMARK:
LEI SWITZERLAND: Auvernier, lO! (Vaud), BM! FRANCE: Mt. Mezenc, BM!
Morlaix
(Fl.),
Maisons-laffitte, Rhinefield (Bx.),
BM!
P!
(Hants),
BM!
lustleigh
(Dartmoor),
OT!
leicester,
TW!
Korsor,
(Suffolk),
BM!
BM!
l ill afUred, Krupa
Heiligensee,
(Reichenau
distr.),
CO!
BU!
(Pazaric,
HU!
BM!
HOllAND:
BM!
ENGLAND: Spratton
Kew
Silwood Park
(D.T.),
GERMANY:
BU! BU!
Ingelheim
Prater
Putten
(Gld.),
lutry, lO! Nyon, GV! les Pleiades P! Arles-sur-Tech (Pyrenees Orientales),
Thames Ditton
Ol!
BM! 8M!
P!
Minsmere Reserve
Studland Heath
Budapest, (Bosna),
Mouleydier (Oordogne),
le Puy, BM!
BU!
Trebevic
(Surrey), BM!
Huelgoat
8M!
Oxford,
Esher Common
(Sy.),
BM!
(Berks.),
BM!
Watbarrow Point
Wimbledon Castle,
BM;
Com.
BM!
BM! Brickhill
(Surrey),
Bedford Park
(Wray
(Fl.),
(Northants),
BM!
(london),
(london),
BM!
lancashire),
Dartmoor, BM! Monks Wood (Cambs.), BM! SCOTLAND: Aviemore (EI.), Tentsmuir Nat. Res. (Fife), BM! Rannoch Forest (Perths.), BM!
IRELAND: Drumchory (Co. Wd.), BM! Ol! ITALY: 5. Benedetto Belbo (Cuneo), Pl! SZ! Peveragno (Cuneo), SZ! Caluso (Torino), EN! Pizzighettone (Cremona),
TW!
CYPRUS:
Pera Pedi-Moniatis R.,
3. Aphelopus serratus Richards
Aphelopus serratus Richards 1939: 284. Aphelopus serratus Richards: J.F. Perkins 1976: 11, 12. Aphelopus serratus Richards: Jervis 1977: 302.
BM!
Aphelopinae
FEMALE:
fully
winged;
testaceous or whitish; legs testaceous,
length
=
OOL
frontal
3,5;
OPL
antennae distally thickened; 6: 4: 6:
6:
6:
6:
I ine present;
occipital
height of clypeus;
=
epistomal
3;
=
TL
scutum,
notaulices reaching
3;
5:
6:
reticulate rugose,
beyond 0,75
fully
tibial
winged;
black;
legs brown,
Fig.
34 -
distally
thickened; 6:
6:
6:
6:
9;
median scutal
epistomal
fully
7;
OL
or greater than medial granulate;
line present;
keels on
=
width
sometimes reaching
the
propodeum
the posterior surface;
fore-wings hyaline,
1,75-2,12 mm;
black;
without dark
mandibles and clypeus
and fore tibiae testaceous;
and ninth abdominal
granulate;
England:
carina complete;
POL
space at
its narrowest point equal
clypeus;
scutum,
to,
genitalia
sternite (8).
frontal
suture medially straight
antennae
antennae not
antenna I segments in following proportions: head dull,
incomplete; occipital TL = 3;
to,
Male of Aphelopus serratus Richards from
6:
=
POL
sometimes clypeus uniformly dark or black;
with tarsi
(A)
head dull,
spurs 1,1,2.
length
testaceous or whitish;
8;
scnure medially straight;
length of scutum;
with 2 longitudinal
median area reticulate rugose; bands;
5:
sometimes
antenna I segments
scutellum and metanotum dull,
posterior margin of the scutum;
MALE:
antennae black;
carina complete;
of the malar space at its narrowest point equal
transversal
mandibles and clypeus
with hind coxae and hind femora partly brown;
in following proportions: 4;
black;
sometimes uniformly black or dark;
mid femora partly brown; granulate;
1,5-2,5 mm;
53
=
6;
(Fig.
OL
=
4;
35A);
4:
5:
6:
OOL
=
4;
OPL
=
3;
width of the malar
or greater than medial
scutellum and metanotum dull,
5:
line complete or
granulate;
height of
notaulices
54
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
complete and posteriorly separated or incomplete and reaching length of scutum;
median scutal
with 2 longitudinal
Fig.
35 -
line present;
keels on the posterior surface;
median area
Heads of males of Aphelopus serratus Richards and camus Richards
(Dalman)(B)
beyond 0,75
propodeum reticulate rugose, shining,
atratus
(A),
(C).
smooth, little rugose; fore-wings hyaline, without dark transversal bands; ninth sternite of the abdomen posteriorly strongly emarginated (Fig. 34B); distivolsella in the form of a long straight rod (Fig. 34A); basivolsella (Fig. 34A) long and narrow and distally pointed; basivolsella with two bristles distally; tibial spurs 1,1,2. LOCUS TYPICUS: Norwood (England) TYPICAL MATERIAL:
holotype M,
in BM;
(2 FF,
5 para types
allotype F and 8 paratypes (5 FF,
3 MM)!
in DU;
6 para types MM!
3 MM)
in OX.
according to Jervis (1980c), Alebra albostriella Kybos smaragdula (Fallen), Fagocyba cruenta (H.-S.), Edwardsiana geometrica (Schrank), Lindbergina aurovittata (Doug I as), Ribautiana tenerrima (H .-5.), Eupterycyba jucunda (H .-5.), Typhlocyba quercus (F.), Alnetoidia alneti (Dah I born), Zygina sp. HOSTS:
in England,
(Fallen),
BIOLOGY: informations in Jervis (1980b, c); in Egland univoltine species; overw i nteri ng stage: prepupa and first instar larva. DISTRIBUTION: HE!
HUNGARY:
Bellinchen
Palaearctic region: TiszasUly,
(Oder),
HA!
FINLAND:
BU! Szabohegy
Lohrbery-Fuss.
Aminne
(Koszeg),
(Ab.,
BU!
Pojo),
GERMANY:
(Siebengebirge),
OT!
HE!
Lojo,
Kiel,
DENMARK,
HU! CO!
HOLLAND: Putten (GLD.), LE! SWITZERLAND: Besazio (Tessin), GV! Follaterres (Valais), LO! ENGLAND: Norwood (London) (Richards 1939); Minsmere Reserve (Suffolk),
BM!
Rhinefield
(Hants),
BM!
Esher Common
(Surrey), BM! Leicester, TW! OL! Watey End (Boxmoor, HT.), BM! Bedford Park (London), BM! Slough (Bucks), BM! Staunton (GW.), BM! Wye (Kent), BM!
Kingsteingnton
SCOTLAND Agher
(SD.),
(Richards 1939);
(Co. Meath),
BM!
BM!
other localities in Richards
IRELAND:
D'dee (DO.),
Donaghadee (Co.
(1939);
Down),
BM!
OL!
BM!
4. Aphelopus camus Richards
Aphelopus heidelbergensis Richards 1939: 286 (syn. proposed by J.F.
Aphelopinae
55
Perkins 1976); locus typicus Heidelberg (Germany). Aphelopus camus Richards 1939: 287 (heidelbergensis has page priority over camus, but J.F. Perkins 1976, the first reviser, placed the former as a synonym of the latter); locus typicus Storeys Way
(Cambridge,
England).
Aphelopus camus Richards: J.F. Perkins 1976: 11, 12. Aphelopus camus Richards: Jervis 1977: 302.
FEMALE:
fully
winged;
clypeus whitish; legs testaceous, tibiae
length
light brown;
granulate;
mm;
black;
mandibles testaceous;
with segment 1 partly
with mid and hind femora dark brown and with antennae distally thickened;
following proportions: 4: = 3;
1,68-1,81
antennae black or brown,
frontal
3,5: 5:
6: 7:
I ine incomplete;
occipital carina complete;
6:
6:
POL = 6;
scutum,
7:
antennal 6:
7;
OL = 3;
yellow hind
segments in
head dull, OOL = 3;
OPL = 3;
TL
scutellum and metanotum dull,
granulate; notaulices incomplete, reaching 0,65 length of scutum; propodeum reticulate rugose, with 2 longitudinal keels on the posterior surface;
fore-wings hyaline,
1,1,2. MALE: fully
winged;
length 1,75-2,25 mm;
testaceous or whitish;
Fig.
36 -
"9nt;
=
bands;
tibial
spurs
mandibles and clypeus legs brown,
with tarsi
Male of Aphelopus camus Richards from Carlsbad genital ia
antennae not distally 3:
black;
antennae black or brown;
(Czechoslovakia):
::-oportions: 4:
without dark transversal
5:
6:
6:
(A)
and ninth abdominal
thickened; 6:
6:
6:
6:
antennal 8;
segments
head dull,
sternite (6).
in following
granulate;
frontal
-e incomplete; occipital carina complete; POL = 6; OL = 4; OOL = 3; OPL 3; TL = 3; scutum, scutellum and metanotum dull, granulate; notaulices
-~omplete,
reaching about 0,65
length of scutum;
propodeum reticulate
56
Mem.
rugose,
with 2 longitudinal
slightly
less rugose than
without
(Fig.
36A);
spurs 1,1,2.
tibial
LOCUS TYPICUS:
keels on
bands;
36B);
hoi otype M!
Inst.,
no.
37,
1984
the posterior surface;
median area
fore-wings hyaline,
ninth sternite of the abdomen posteriorly
inner part of the basivolsella with
Storeys Way
TYPICAL MATERIAL:
Ent.
the parts outside the area;
dark transversal
emarginated
BM;
Amer.
(Cambridge,
holotype M!
England) of A.
and allotype F!
and 4 para types MM
(1
1 bristle (Fig.
heidelbergensis in camus in BM.
!) of A.
seen
in Germany Chlorita sp.
HOSTS:
DISTRIBUTION: Miklos,
BU!
Palaearctic region:
Vacz
CZt=:CHOSLOVAKIA: Heidelberg,
BM!
HUNGARY:
(Tudosdomb),
BU!
Carlsbad,
Bohemia,
Bellinchen
P!
FenyOfb,
Rbjtbkmuzsaj,
(Oder)
TW!
BU!
Bartfa,
BU!
Szod,
BU!
Hoi ics., OL!
SzSzt
BU!
GERMANY:
(holotype and two para types of
Aphelopus chloritae Haupt, spec i es never descr ibed), HA! SW I TZERLAND: Saas-Fee (Valais), Casale Roletto
BM!
ENGLAND:
(Mongrando,
Storeys Way
Vercelli),
OL!
(Cambridge),
BM!
TURKEY: Ankara,
BU!
ITALY:
5. Aphelopus nigriceps Kieffer
A,0,':elopus melaleucus (Da I man) var. nigriceps Kieffer in Kieffer et Marshall
Aphelopus Aphelopus Aphelopus Aphelopus
FEMALE:
fully
1905:
nigriceps nigriceps nigriceps nigriceps
winged;
antennae black,
219.
Kieffer: Kieffer:
Kieffer 1914b: 216. Richards 1939: 291.
Kieffer: Kieffer:
J.F. Perkins 1976: Jervis 1977: 301.
length 1,75-2,25 mm;
black;
with segments 1-2 testaceous;
legs
12.
11,
mandibles
testaceous;
testaceous,
with hind
coxae and hind femora partly brown;
antennae distally thickened;
antennal
segments in following proportions:
5,5:
head
fully POL
granulate,
=
8;
OL
=
dull;
4;
OOL
frontal
=
4;
6:
5:
6:
I ine incomplete;
OPL
=
3,5;
=
TL
6:
6:
occipital
3;
scutum,
6:
5:
5:
7;
carina complete; scutellum and
metanotum smooth, granulate; notaul ices incomplete, reaching about 0,65 length of scutum; median scutal I ine sometimes present; propodeum reticulate rugose,
with 2
median area shiny, wings hyaline, MALE:
fully
longitudinal
less rugose than
without dark transversal
winged;
legs brown,
distally thickened;
antennal
7:
head dull,
7:
8:
8:
11;
segments
carina complete; POL = 9; OL scutellum and metanotum dull, about 0,65 length of scutum;
the posterior surface;
bands;
length 1,75-2,37 mm;
antennae brown or black; 7:
keels on
the parts outside the area; black;
with
tarsi
tibial
mandibles testaceous; light;
antennae not
in following proportions:
granulate;
frontal
fore-
spurs 1,1,2.
line present;
4: 4: 6: 6: occipital
=
4; OOL = 4,5; OPL = 3; TL = 4; scutum, granulate; notaulices incomplete, reaching
median scutal
line sometimes present;
57
Aphelopinae
propodeum reticulate rugose,
with 2 longitudinal
surface;
almost smooth,
median area shiny,
keels on
the posterior
slightly rugose;
fore-wings
8
Fig.
Male of Aphelopus nigriceps Kieffer:
37 -
abdominal
hyal ine,
without dark transversal
genitalia
(A)
and ninth
sternite (B).
bands;
ninth sternite of the abdomen
posteriorly emarginated (Fig. 37B); inner part of the basivolsella with 2 strong bristles (Fig. 37A); tibial spurs 1,1,2. LOCUS TYP ICUS:
Dumfries
(Scotland)
TYPICAL MATERIAL: holotype F! in BM. HOSTS: in England Eurhadina concinna (Germar)(Jervis 1980b, BIOLOGY:
informations in Jervis
JISTRIBUTION: Pulchauki
Palaearctic region:
(Katmandu),
Lemland
(AI.),
Tihany,
BU!
(1980b,
HE!
Vacz.
OT!
OL!
HUNGARY: Cscrcg.,
c);
NEPAL:
FINLAND:
Fenycfc,
BU!
Godavari Lojo,
BU!
c).
in England bivoltine species. (Katmandu),
HE!
Kangasala,
OT!
Istvinkut (Hegyalja),
SzSzt Miklos,
BU!
ROMANIA
OL!
HE! BU!
(Nagy
1972);
CZECHOSLOVAKIA: Donovali (Zvolan), OL! YUGOSLAVIA: Prespa Geul (Otesevo, Makedon ij a), BM! GERMANY: Aachen, P! AUSTR I A: Huben (0 Tirol),
HU!
Semmeringgebeit
(Gld.),
LE!
FRANCE:
=oret de Dabo (Moselle), JL!
Wimbledon Com.
'.:--iorthants),
BM!
'.Surrey),
BM!
iRELAND:
Co.
VI!
(London),
Norfolk,
SCOTLAND: Kerry,
(Reichenau
Le Puy,
TW!
BM!
P!
ENGLAND: OT!
(Fl.),
Rhinefield
Wicklow,
BM!
HOLLAND:
BM!
BM!
Putten
Maisons-Laffitte,
Minsmere Reserve
Bedford Park
Bonar Bridge, Co.
distr.),
Huelgoat
(Suffolk),
(Hants),
BM!
Spratton
(London),
BM!
Horsley
Claddich,
BM!
Dumfries,
TW!
6. Aphelopus nepalensis n. sp.
P!
BM!
BM!
58
Mem.
FEMALE:
fully
winged;
Amer.
Ent.
Ins!.,
no.
length 1,81- 1,87 mm;
37,
black;
1984
mandibles
antennae fully brown or with segments 1-2 testaceous; antennae distally 5:
6,5:
7:
POL; 6;
6:
6:
thickened; 5:
6:
Ol = 3,5;
5:
antennal
9;
segments in following proportions:
head dull,
OOl = 5;
OPl ;
3;
testaceous;
legs testaceous;
granulate; Tl ;
4;
frontal
scutum,
5:
I ine incomplete;
scutellum and
B \ ) (
/ /
/
A Fig.
Male of Aphelopus nepalensis n.
38 -
genitalia
(A:
metanotum dull,
penis and
granulate;
length of scutum; on
MALE:
median area shiny
without dark transversal
fully
winged;
from Bokaihunde (Nepal):
notaulices incomplete,
propodeum reticulate rugose,
the posterior surface;
hyaline,
sp.
left paramere), ninth abdominal left volsella (e)
length
bands;
1,87-1,93 mm;
sternite (B)
reaching
about 0,5
with 2 longitudinal
and smooth;
tibial black;
and
keels
fore-wings
spurs 1,1,2. mandibles testaceous;
antennae brown, sometimes with segment 1 testaceous; legs testaceous, with base of hind coxae black; sometimes also hind femora and hind tibiae
59
Aphelopinae
brown;
antennae not distally
proportions: 4: frontal ;
3,5;
4:
6: 6: 7:
line incomplete; OPL ;
granulate;
2,5;
TL ;
thickened;
6,5:
occipital 3,5;
7:
9;
head dull,
carina complete;
scutum,
notaulices incomplete,
propodeum reticulate rugose,
antenna I segments in following
6,5: 6,5:
POL; 6;
granulate; OL ;
3,5;
OOL
scutellum and metanotum dull,
reaching
about 0,5 length of scutum;
with 2 longitudinal
keels on
the posterior
surface; median area shiny and smooth; fore-wings hyaline, without dark transversal bands; apex of the penis not tridentate (Fig. 38A); inner part of the basivolsella with 2 bristles LOCUS TYPICUS:
TYPICAL MATERIAL: paratype F! HOSTS:
(Fig.
38C);
tibial
spurs 1,1,2.
Bokaihunde (Nepal) holotype M!
and 2 paratypes
(1
F,
1 M)!
in B;
~T.
in
unknown
DISTRIBUTION: Trisuli),
Palaearctic region:
NEPAL:
Bokaihunde (20 Km N of
B! Syabrubens (35 Km N of Trisuli),
B!
27° 58'
N 85° 00'
E,
OT!
7. Aphelopus querceus n. sp.
FEMALE: MALE:
unknown
fully
winged;
length 2,18 mm;
black;
mandibles and clypeus
B
Fig.
39 -
Male of Aphelopus querceus n. ninth abdominal
:estaceous;
head black,
sp.
(type):
genitalia
(A)
and
sternite (B).
with some whitish
spots between
the antennal
sockets; malar space white; antennae black; legs testaceous, with hind "ernora darkened; antennae not distally thickened; antennal segments in
60
Mem.
following proportions: granulate; 3,5; OOL
frontal
Amer.
5:
5:
7:
Ent.
8:
line complete;
9:
Inst.,
9:
9:
no.
9,5:
37,
1984
9,5:
14;
head dull,
occipital carina complete;
POL = 6;
OL
= 5; OPL = 2,5; TL = 2,5; scutum dull, granulate; notaulices
incomplete,
reaching about 0,75
length of scutum;
scutellum granulate;
metanotum shiny, without sculpture; propodeum reticulate rugose, without transversal or longitudinal keels, with a median shiny and smooth area on the posterior surface;
fore-wings hyaline,
without dark transversal
bands;
ninth sternite of the abdomen posteriorly weakly emarginated (Fig. 39B); inner part of the basivolsella with 2 bristles (Fig. 39A); apex of the penis not
tridentate
LOCUS TYPICUS:
TYPICAL MATERIAL: HOSTS:
(Fig.
39A);
Bhurumche
tibial
spurs 1,1,2.
(Nepal)
holotype M!
in OT.
unknown
DISTRIBUTION:
Palaearctic region:
NEPAL:
Bhurumche (Katmandu),
ETHIOPIAN SPECIES OF
OT!
APHELOPUS
Three species are known.
KEY TO THE SPECIES
FEMALES AND MALES
Notaulices margin
of
invisible or the
slightly
visible
scutum •••••••••••••••••
only near
the anterior
l.mediocarinatus (Benoi t)
Notaulices distinct, reaching at least 0,5 2 Notaulices reaching approximately 0,5-0,65
length length
of of
scutum ••••••• 2 scutum •••..•••
2.wittei Benoi t -
Notaulices
reaching
approximately
the posterior margin
scu tum. • • • • • • • • • • • • • • • • • • • • • • • • . • • • ••
3. incisus n.
of
the
sp.
1. Aphelopus mediocarinatus (Benoit) n. comb.
Antaphelopus Antaphelopus locus typicus Antaphelopus
FEMALE:
fully
mediocarinatus Beno it 1951 d: 23. madagascariensis Benoi t 1953b: 429 (n. syn.); Esira
(Madagascar).
mediocarinatus Benoit: Benoit 1953b: 430.
winged;
length 1,81-1,87 mm;
black;
mandibles testaceous,
Aphelopinae
with
teeth reddish;
white;
antennae distally 4:
5:
clypeus black or white;
antennae brown, 6:
5,5:
incomplete;
5:
5:
occipital
face fully
with segments 1-3 testaceous;
thickened;
5:
61
5:
antenna I segments
8;
head dull,
carina complete;
fully
black or partly legs fully
testaceous;
in following proportions: granulate;
POL = 6,5;
frontal
OL = 4;
4:
line
OOL = 3,5;
OPL
=
3; TL = 3; scutum dull, fully granulate; notaulices invisible; scutellum and metanotum shiny, without sculpture or very sl ightly granulate; sometimes dull and distinctly granulate; propodeum reticulate rugose; dorsal surface with wide areolae; 2 longitudinal keels on the posterior
surface;
median
area usually
separated from
transverse keel,
occasionally
reticulate rugose,
rugose along
the margins with central
propodeum reticulate rugose; granulate;
the dorsal
surface by
usually shiny,
area smooth;
lateral
areas of the
mesopleura and metapleura smooth,
fore-wings hyaline,
without dark
transversal
a
reticulate
bands;
fully tibial
spurs 1,1,2. MALE: fully winged; length 1,25-1,62 mm; black; with teeth reddish; clypeus black or light; fully antennae brown,
with segments 1-2 testaceous;
antennae not distally
thickened;
proportions:
5:
frontal
3:
3:
4:
line absent;
OPL = 0,5;
6:
7:
occipital
TL = 2;
scutum,
legs fully
antenna I segments
6,5:
7:
6:
10;
carina complete;
POL = 6;
granulate or reticulate rugose;
usually not reticulate rugose, tibial
transversal
bands;
fully
lateral
granulate;
ninth abdominal
granulate;
fully
OOL = 4; granulate;
2 longitudinal keels on keel; median area dull areas of the propodeum
fore-wings hyaline,
sternite variable
without
(Fig. 40);
spurs 1,1,2.
rig.
40 -
sternite
Male of
Aphelopus mediocarinatus (Benoit): ninth abdominal
(specimens from Garamba National Esira on
_=:US TYPICUS:
Paulis
Park on
the
left and from
the right).
(Zaire)
of A. mediocarinatus in TV; holotype M! of madagascariensis in P; 1 para type M! of A. madagascariensis in TV.
-v"ICAL MATERIAL: ~.
fully
OL = 3;
scutellum and metanotum dull,
ices invisible; propodeum reticulate rugose; the posterior surface and an anterior transversal
dark
testaceous;
in following
head dull,
~otaul
or shiny,
mandibles testaceous, black or partly white;
holotype F!
62
Mem.
HOSTS:
Entebbe, Butahu,
OT! Mt.
Ethiopian region: ZAIRE:
Paulis
Ruwenzori,
Hoyo (Albert Nat.
Ile-ife
(W.
State),
SOUTH AFRICA: Natal), BM!
Ent.
Inst.,
no.
37,
1984
unknown
DISTRIBUTION:
Mt.
Amer.
BM!
Albert Nat.
Park),
BM!
OL!
TV!
Mossel
Bay
Afgoi
TV!
Park), OT!
(Benadir),
Kalonge
TV!
Rwankwi
Ibadan,
Pietermaritzburg
MADAGASCAR:
SOMALIA:
(Haut-Uele),
(N of Kivu Lake),
Umuahia CRIN
Drakensberg
(Cape Province),
BM!
OL!
Fianarantsoa,
Park, TV!
Aff.
TV!
NIGERIA:
(EC State),
TW!
P!
UGANDA:
Katauleko,
Garamba Nat.
(Natal),
Esira,
OL!
(Riv.
BM!
(Van Reenen,
Ceres (Cape Province),
TV!
2. Aphelopus wittei Benoit
Aphelopus wittei Benoit 1951c: 16. Gymnaphelopus wittei (Benoit): Benoit 1951d: 23.
FEMALE: fully winged; length 1,62-2,50 mm; black; head usually with a more or less wide whitish frontal spot; sometimes face black; clypeus white;
antennae fully
legs fully
testaceous;
testaceous or brown,
with segments 1-2 testaceous; is usually black:
in
specimens with wide frontal spots the ventral side of head and body fully whitish; antennae not distally thickened; antennal segments in
is
following proportions:
the ventral
5:
5:
6:
side of the body
8:
8'
7:
5:
5:
granulate; frontal line incomplete; occipital OL = 4; OOL = 5; OPL = 4; TL = 5; scutum, fully
granulate;
notaulices
incomplete,
scutum.
sometimes reaching only 0,5
rugose,
usually with 2 longitudinal
area usually smooth and shiny, dorsal
surface and on
metapleura smooth,
5:
7;
head dull,
usually reaching 0,65 length of
length of scutum; keels on
propodeum reticulate
the posterior surface;
not reticulate rugose;
the sides of the posterior surface;
fore-wings with only
with segments 1-2 testaceous and
legs fully testaceous (at most with stalks of hind femora darkened); some specimens head with a more or less wide frontal whitish spot; sometimes this spot
is very
ocellar region brown; some specimens only
wide:
occasionally head fully
the antennae in the clypeus
is whitish;
antenna I segments
in following proportions:
7:
fully granulate;
occipital scutum,
head dull,
carina complete;
POL
=
7,5;
scutellum and metanotum dull,
incomplete,
OL
=
4;
fully
frontal
in
OOL
approximately 0,5
=
5,5:
5:
5,5:
6:
7:
line incomplete; 5;
granulate;
usually reaching approximately 0,65
occasionally reaching only
testaceous;
antennae not distally
thickened;
10;
in
whitish with
this case are fully
7,5: 7:
7:
the
mesopleura and
without sculpture or sl ightly granulate;
antennae brown
median
areolae only on
hyaline, without dark transversal bands; tibial spurs 1,1,2. MALE: fully winged; length 1,68-2,25 mm; black; head usually mandibles testaceous,
fully
carina complete; POL = 5,5; scutellum and metanotum dull,
OPL
=
4;
TL
=
5;
notaulices
length of scutum;
length of scutum;
propodeum
Aphelopinae
reticulate rugose, shiny
usually
and smooth,
with 2 longitudinal
not reticulate rugose,
occasionally posterior surface of and fully
reticulate rugose;
reticulate rugose, hyaline,
tibial
keels;
median area usually
granulate or without sculpture;
the propodeum without
longitudinal
mesopleura and metapleura smooth,
sl ightly granulate or without sculpture;
without dark transversal
(Fig. 41);
63
bands;
ninth abdominal
keels
not
fore-wings sternite variable
spurs 1,1,2.
B
"'-9.
41 - Male of Aphelopus wittei Benoit: specimens from Ngesho (A), Kirungu
ninth abdominal sternite in (B) and Burunga (C).
_~=~S TYPICUS:
-.=
Kashwa (Zaire) :::AL MATERIAL: holotype F!,
:~-=.:ype
-':::-5:
M!
and 90 paratypes!
in TV;
unknown
::: 5-:;IBUTION: :::- ,,-awasha
=_"',:A.:
allotype M!
in BM. Ethiopian region:
(Salisbury),
Ruhengeri
(Kirii
BM!
ANGOLA:
NIGERIA:
sources),
',',agera Lake (Albert Nat.
TV!
Park),
Gabela,
BM!;
ZIMBABWE:
Ile-ife (W State), ZAIRE:
TV!
BM!
Garamba Nat.
Yolohafiri
Cave
(Mt.
OL! Park, Hoyo,
TV!
1
64
Mem.
Albert Nat. Nord,
Park),
TV!
Albert Nat.
Kivu), (Riv.
TV!
Mt.
Amer.
Hoyo
Inst.,
TV!
(Albert Nat.
Albert Nat.
Park),
Park),
Park),
TV!
OL!
Kalonge
(Riv.
LE!
TV!
TV!
37,
(Riv.
Lumumbashi,
Albert Nat. Park), TV! Mt. Musimba Park), TV! Kalonge (Riv. Kamahoro, Nat.
no.
Secteur Tshiaberimu
Park),
Talia Nord,
Ent.
1984
Kalivina,
OL!
(Secteur Tshiaberimu,
(Secteur Tshiaberimu, Albert Nat. Aff. Butahu, Mt. Ruwenzori, Albert Kamusonge,
aff.
Butahu,
Albert Nat.
Park),
TV!
Kiondo ya Kwanza
Ruwenzori,
Albert Nat.
Park),
TV!
Riv.
Ruwenzori,
Ruwenzori, Nat.
Albert Nat.
Park),
Park),
Albert Nat.
TV!
TV!
Riv.
Park),
Park),
Burunga
(Mokoto,
Mt.
Ruwenzori,
Tshiaberimu,
TV!
Albert Nat.
OL!;
Riv.
Kyanyamu,
(Aff. Mt.
Mt.
Park),
Park),
Park),
TV!
TV!
TV!
Musabaki
Kalonge,
Bombi,
Ruwenzori,
Albert Nat. Park),
(Kyandolire,
TV!
(Aff.
Mt.
Albert
Albert Nat.
Bomboka
Elisabethville,
Riv.
(Aff.
Ruwenzori,
BM!
Mt.
Butahu,
Kakalari
(Kalonge,
Mt.
(Kalonge,
Mt. Sesero (Bitashimva,
Albert Nat.
Albert Nat.
Nyamwamba
Kikyo
Lusilube (Mt.
Ngesho (Kivu),
TV!
TV!
TV!
Talia
(Munagana,
Secteur Tshiaberimu
Kirungu
Ruwenzori, Mt.
aff.
Tshengalero
Hintumo (Secteur dr.
Talia Nord,
Secteur Tshiaberimu, Albert Nat. Park), TV! OL! Rutshuru (Kivu), TV! Mt. Ibale (Kyandolire, Mt. Ruwenzori, Albert Nat. Park), TV! N'Dalage (Kivu), TV!
Rweru
(1951c);
(Volc.
Mikeno,
MADAGASCAR:
Matjesfontein
Albert Nat.
Mandraka,
(Cape Province),
Park),
TV!
BM!
OL!
BM!
Bekily,
other localities in Benoit P!
SOUTH AFRICA:
OL!
3. Aphelopus incisus n. sp.
FEMALE: MALE:
unknown
fully
winged;
teeth brown; testaceous,
with hind coxae,
distally
thickened;
6:
6:
6:
6:
occipital scutum,
length 1,68 mm;
antennae brown,
6:
9;
POL = 6;
granulate; OL = 3;
almost reaching
OOL = 3;
tibial spurs 1,1,2. LOCUS TYPICUS: Umuahia CRIN
6:
antennae 4:
4:
line almost complete; OPL = 2,5;
granulate;
TL = 2,5;
notaulices almost
the posterior margin of the scutum;
fore-wings hyal ine,
TYPICAL MATERIAL:
frontal
with
legs
hind femora and hind tibiae brown;
head dull,
complete;
reticulate rugose;
mandibles testaceous,
antenna I segments in following proportions:
scutellum and metanotum dull,
complete,
HOSTS:
black;
with segments 1-2 testaceous;
without dark transversal
propodeum bands;
(Nigeria)
holotype M!
in BM.
unknown
DISTRIBUTION:
Ethiopian region:
NIGERIA:
OR I ENTAL SPEC I E5 GF
Eight species are known.
Umuahia CRIN
APHELOPUS
(EC State),
BM!
5:
65
Aphelopinae
KEY TO THE SPECIES
FEMALES AND MALES
Species -
fully
testaceous,
with part
of
the antennae brown . . . . . . . . . . .
• • . • • • • • • • . • • • • • • • • • • • • • • • • • • • • • • • • •• 1. borneanus n. sp. Species partly or fully black or brown •••••••••••••.•••••••••..•••• 2
2 Head testaceous, at most with ocellar region or vertex dark •••••••• 3 Head mostly or fully black or brown • • . • • • • • . • • • • • . • • • . • • . . • • . • • • • • • 5
-
3 Notaulices
invisible •••••••••••••••••
2. maculiceps Bergman
Notaulices distinct ••.••••••••.••••••..•.•••••••••••••••••••••••••• 4
4 Notaulices short, reaching at most 0,5 length of scutum . . . . . . . . . . . .. · . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. 3. malayanus n. sp. - Notaul ices reaching approximately 0,80 length of scutum . . . . . . . . . . . .. · . . . . . . . . • . . • . . . • . . . • . . . . . . . . . • . . . . .. 4. ochreus n. sp. 5 Notaulices short, reaching at most 0,5 length of scutum . . . . . . . . . . . . 6 - Notaulices long, reaching the posterior margin of scutum . . . . . . . . . . . 7 6 Notaulices reaching approximately 0,5 length of scutum . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5. orientalis n.sp. - Notaulices reaching approximately 0,3 length of scutum . . . . . . . . . . . . ..
6. penanganus n. sp. 7 Metapleura separated from the propodeum by a strong distinct longitudinal keel (Fig. 42 A) . . . . . . . . 7. birmanus n. sp. -
Metapleura distinctly areolae
Fig.
(Fig.
separated
from
the propodeum by
42 B) . . . . . . . . . . . . . . . . . . 8.
a
series
philippinus n. sp.
42 - Metathorax + propodeum and abdomen of Aphelopus birmanus n.
sp.
(A)
and
philippinus n. sp. (B).
1. Aphelopus borneanus n. sp.
::::::MAlE:
fully
winged;
length 2,37 mm;
:-10 brown;
antennae distally
::;roportions:
5:
'ne incomplete;
4,5:
6:
7:
POL = 7;
5:
testaceous,
thickened; 6:
6:
Ol = 3;
6:
5:
antennal 8;
OOl = 4;
with antennal segments
head dull, OPl = 5;
segments
in following
granulate;
Tl = 5;
frontal
occipital
of
66
Mem.
carina complete; notaulices
scutum,
Amer.
En!.
Ins!.,
no.
37,
scutellum and metanotum dull,
reaching approximately 0,5
incomplete,
1984
propodeum reticulate rugose,
with 2 longitudinal
surface;
without dark transversal
1,1,2. MALE:
fore-wings hyaline,
granulate;
length of scutum; keels on
the posterior
bands;
tibial
spurs
unknown
LOCUS TYPICUS:
Tenompok
TYPICAL MATERIAL: HOSTS: unknown DISTRIBUTION:
(Sabah,
holotype F!
Oriental
Malaysia)
in B.
region:
MALAYSIA:
Tenompok
(Sabah),
B!
2. Aphelopus maculiceps Bergman
Aphelopus maculiceps Bergman 1957: 9.
FEMALE
and MALE:
fully
winged;
with ocellar region brown; testaceous;
4:
4:
= 3;
not
4:
absent;
1,25-1,37 mm;
thorax and propodeum black,
abdomen brown-black; female,
length
antennae brown,
thickened
4:
3,5:
3:
occipital
scutum,
3:
legs testaceous; in male; 3:
2,5:
sometimes with scutum brown;
antennae distally thickened
antenna I segments 5;
carina complete;
head granulate, POL
head testaceous,
with segments 1-3 or 1-4
=
5; OL
scutellum and metanotum dull,
=
in following dull;
frontal
3; OOL
granulate;
=
in
proportions: line
4; OPL
=
3; TL
notaulices
invisible; propodeum reticulate rugose, with 2 longitudinal keels on the posterior surface; median area partly smooth; fore-wings hyaline, without dark transversa I bands; LOCUS TYPICUS:
t i b i a I spurs 1,1,2.
Bogor (Java)
holotype M and allotype F in LE; 1 paratype F! in 1 paratype F! in BM; paratypes in Bogor (Division of Entomology, Faculty of Agriculture). HOSTS: in Java Empoasca sundaica Bergman (Bergman 1957). DISTRIBUTION: Oriental region: INDONESIA: Bogor (W Java), BM! WA! TYPICAL MATERIAL:
WA;
3. Aphelopus malayanus n. sp.
FEMALE: occiput in
fully dark;
length
1,80 mm;
the only female specimen known);
with dorsal granulate; 4;
winged;
OOL
=
regions brown; frontal
5;
incomplete,
head
testaceous,
antennae with segment 1 testaceous
OPL
=
5;
TL
=
5;
vertex and
thorax and propodeum testaceous,
abdomen brown;
line complete;
with
(other segments missing
occipital
scutum dull,
reaching approximately 0,5
legs testaceous;
carina complete; granulate;
head dull, POL = 6;
OL
notaulices
length of scutum;
scutellum dull,
Aphelopinae
granulate;
metanotum shiny,
reticulate rugose,
with 2
fore-wings hyal ine, MALE:
fully
lengt
keels on
transversal
abdomen
propodeum
the posterior surface;
bands;
tibial
1,43-1,56 mm; head testaceous,
and part of vertex dark;
three segments dark; or brown; propodeum or black;
without sculpture;
longitudinal
without dark
winged;
ocellar region
smooth,
67
spurs 1,1,2. sometimes with
antennae testaceous,
with
last
thorax testaceous, usua II y with dorsa I reg ions dark testaceous, with dorsal and posterior surface brown
brown-testaceous;
legs testaceous;
antennae not d ista II y
4: 4: 5: 5: 5: 5: 5: 5: 5: 5; head dull, granulate; frontal line complete; ocipital carina complete; POL = 6; Ol = 3; OOl = 3; OPl = 4; Tl = 3; scutum, scutellum and metanotum dull, granulate; notaulices incomplete, reaching at most 0,5 thickened;
antennal
length of scutum;
segments
without dark
_OCUS TYPICUS:
TYPICAL MATERIAL:
with 2 longitudinal
median area smooth and shiny;
transversal
Klang Gates
:0; 1 paratype F! ..,OSTS:
proportions:
propodeum reticulate rugose,
on the posterior surface; -,yal ine,
in following
bands;
(Malaya,
holotype M! in OL.
tibial
keels
fore-wings
spurs 1,1,2.
Malaysia)
and 1 paratype F!
in B;
1 paratype M!
unknow n
JISTRIBUTION: Oriental region: MALAYSIA: Klang Gates (Malaya, Kuala _,-,mpur), B! Gomak For. Res. (Malaya, Kuala lumpur), Ol! Kuala Tahan "',alaya,
Pahang),
B!
PHiliPPINES:
Pinigisan
(Mantalingajan,
Palawan),
:O!
=:::':AlE:
fully
winged;
-=;ion and vertex dark; :==taceous and
dorsal
=:;:;omen brown; ==;'71ents
=_ :, c.::
n,
length 1,81
testaceous ,
mm;
head
antennae testaceous;
regions brown;
legs testaceous;
sp.
thorax
with
with ocellar ventral
side
propodeum brown or blackened;
antennae distally
thickened;
antenna I
3,5: 3,5: 4: 5: 5: 6: 6: 6: 6: 8; head line complete; POL = 5; OL = 3; OOl = 4; OPL =
in following proportions:
granulate; Tl = 3;
__
4. Aphelopus ochreus
frontal
occipital
granulate;
carina complete;
notaulices reaching
scutum,
:-::)odeum reticulate rugose,
with 2 longitudinal
o_-;ace;
without dark
fore-wings hyaline,
scutellum and metanotum
approximately 0,8
length of scutum;
keels on
transversal
the posterior
bands;
tibial
spurs
.. 2. ~_:::
unknown
_::.,;S TYPICUS: Fraser's Hill (Malaya, -.:; CAL MATERIAL: holotype F! in B.
-:::':::-5: :
Malaysia)
unknown
:-=
Oriental
region:
MALAYSIA:
Fraser's Hill
5. Aphelopus orientalis n. sp.
(Malaya),
B!
in
68
Mem.
FEMALE:
fully
winged;
antennae brown, distally 4:
4:
4:
4:
9;
antennal
no.
black;
~
POL
6;
approximatelyO,5-0,65
granulate;
Ol
1984
mandibles testaceous; legs testaceous;
~
3,5;
frontal ~
OOl
granulate;
antennae
length of scutum;
6:
line incomplete;
3,5;
~
OPl
2;
~
Tl
4:
5:
4:
occipital 2;
notaulices incomplete,
scutum,
reaching
propodeum reticulate rugose,
keels on the posterior surface;
fore-wings hyaline,
37,
segments in following proportions:
scutellum and metanotum dull,
shiny;
Inst.,
length 1,56 mmj
head dull,
carina complete;
2 longitudinal
Ent.
with segments 1-2 testaceous;
thickened;
4:
Amer.
with
median area smooth and
without dark transversal
bands;
tibial
spurs
1,1,2. MALE:
fully
winged;
antennae brown,
length 2,68 mm;
black;
mandibles testaceous;
with segment 1 testaceous;
hind femora brownj
legs testaceous,
antennae not distally thickened;
following proportions:
5:
6:
6:
7:
7:
8:
8:
8:
8:
9;
with clubs of
antennal
segments in
head dull,
granulate;
line complete; occipital carina complete; POL ~ 10; Ol ~ 5; OOl ~ 5; Tl ~ 4; scutum, scutellum and metanotum dull, granulate;
frontal 4; OPl
notaul ices incomplete, reaching approximately 0,5 length of scutum; propodeum ret icu I ate rugose, with a strong transversa I keel between and posterior surface; median area shiny, transversal
bands;
lOCUS TYPICUS:
not rugose;
two
longitudinal
fore-wings hyaline,
dorsa I
keels;
without dark
spurs 1,1,2. (Taiwan)
holotype F!
in B;
paratype M!
in WA.
unknow n
DISTRIBUTION: SRI
smooth, tibial
Fenkihu
TYPICAL MATERIAL: HOSTS:
posterior surface with
lANKA:
Oriental
region:
Adams Peak Trail
TAIWAN: (4,5 mi.
Fenkihu
(Chiayi Hsien),
W Maskeliya,
Kan.
B!
Dist.),
WA!
6. Aphelopus penanganus n. sp.
FEMALE:
fully
winged;
length
1,68 mm;
black;
mandibles testaceous;
antennae testaceous, with segments 7-10 dark; legs testaceous; antennae distally thickened; antenna I segments in following proportions: 4: 4: 5: 5: 4,5:
4,5:
occipital scutum,
4:
4:
4:
5;
head dull,
carina complete;
POL
~
granulatej 6;
Ol
scutellum and metanotum dull,
~
smooth; 1,1,2. MALE:
line incomplete; ~
3;
OPl
~
Penang
(Malaya,
holotype F!
bands;
tibial
Malaysia) in B.
unknown
DISTRIBUTION:
Oriental
region:
MALAYSIA:
Tl
median area shiny
without dark transversal
unknown
TYPICAL MATERIAL:
2,5;
~
3;
notaul ices incomplete,
propodeum reticulate rugose,
keels on the posterior surface;
fore-wings hyaline,
lOCUS TYP ICUS: HOSTS:
OOl
granulate;
reaching approximately 0,3 length of scutumj with 2 longitudinal
frontae
3,5;
Penang
(Malaya),
B!
and
spurs
Aphelopinae
69
7. Aphelopus birmanus n. sp.
FEMALE:
fully
winged;
1,87 mm;
length
black-brown;
mandibles testaceous;
antennae brown, with segments 1-2 testaceous; legs testaceous, with hind femora and hind tibiae dark; antennae distally thickened; antenna I segments complete; TL = 2;
in
occipital scutum,
reaching
the holotype);
= 5;
POL
= 3;
OL
the posterior margin of the scutum;
fore-wings hyaline,
without dark
two
= 4;
OOL
granulate;
line OPL
= 2;
notaulices
propodeum reticulate rugose,
keels on the posterior surface; strong distinct
(last frontal
metapleura separated
longitudinal
transversal
keel
bands;
42 A);
(Fig.
tibial
spurs 1,1,2.
unknow n
LOCUS TYP ICUS:
Kambatti
TYPICAL MATERIAL: HOSTS:
granulate;
scutellum and metanotum dull,
the propodeum by a
MALE:
head dull,
carina complete;
with 2 longitudinal from
5: 4: 5: 5: 5: 5: 4: 4,
in following proportions:
segments missing
(Burma)
holotype F!
in BM.
unknown
DISTRIBUTION:
Oriental
region:
BURMA:
Kambatti
(NE Burma),
BM!
8. Aphelopus philippinus n. sp.
FEMALE: unknown MALE: fully winged;
length 1,60 mm;
and ma 1ar space testaceous; ::lropodeum brown; "airy, 4: 4:
abdomen brown-testaceous;
not distally 5:
6:
:occipital
6:
7:
7:
thickened; 7:
7:
carina complete;
scutum dull,
head brown,
granulate;
with mandibles,
antennae testaceous-brown;
7;
antennal
head dull,
POL
=
6;
4; OOL
notaulices complete,
scutellum and metanotum shiny,
smooth,
antennae
in following proportions:
granulate;
=
OL
legs testaceous;
segments
clypeus
thorax and
=
frontal
3;
OPL
=
I ine complete;
4; TL
=
very
sl ightly granulate;
propodeum
-eticulate rugose, with 2 longitudinal keels on the posterior surface; -etapleura separated from the propodeum by a series of areolae (Fig. 3); fore-wings hyal ine, . , 1,2. _JCUS TYPICUS:
Pinigisan
-vPICAL MATERIAL: -JSTS:
without dark
transversal
(Philippine
holotype M!
bands;
tibial
spurs
Islands)
in CO.
unknown
= STRIBUTION: ::alawan),
Oriental
region:PHILIPPINES:
4;
posteriorly separated;
Pinigisan
CO!
NEARCT I C SPEC I ES OF APHELOPU5
(Mantalingajan,
42
70
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
Four species are known.
KEY TO THE SPECIES
FEMALES AND
Notau Ii ces -
MALES
i nv is i b Ie ••••.•••••.••.•••
2 Fore-wing with a dark
transversal
band;
testaceous ••••••••••••••••••••••••••• -
rufiventris Ashmead
1.
Notaulices distinct •••••••••••••••••••.••••••••.••.••.••••••••.••.• 2
Fore-wing hyaline,
without
dark
at
least
transversal
thorax and propodeum mostly black
or
the
scutum mostly
bicolor Fenton
2.
band;
dorsal
reddish-brown;
side of
occasionally
scutum partly testaceous • • • • • • . . • . • • • • . . • . . • • • • . • • • . • . • . • • • • • • . • . • • 3 3 Notaulices reaching beyond 0,65 length of scutum •••••••••••••••••••. ••••••••••••••••.••.••.••••••••.••••. -
Notaulices
4 Notaulices
reaching at
most
0,5
reaching approximately
0,5
. • • • • • • • • . • • • • • • • • • • • • • • • • • • . • • • • . • •. -
Notaulices
reaching
at
most
0,25
of
scutum •••.••••••••.•••••. 4
length
of
scutum .••••••••.•.••
albopictus Ashmead
4.
length
• • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • ••
varicornis Brues
3.
length
of
1.
scutum •••••••••••.••••.••
rufiventris Ashmead
1. Aphelopus rufiventris Ashmead
Aphelopus rufiventris Ashmead 1893: 100. Aphelopus affinis Ashmead 1893: 102 (no syn.); locus typicus Canada.
Aphelopus rufiventris Ashmead : Fenton 1918b: 276. Aphelopus rufiventris Ashmead: Muesebeck and Walkley 1951: 1042. Aphelopus rufiventris Ashmead: Krombei n 1979: 1250.
FEMALE AND MALE:
fully
winged;
length
with antennae brown or testaceous; segments 1-2 or 1 testaceous; reddish or black, testaceous;
hind
1,31-1,81
mm;
dark-red or black,
sometimes antennae brown,
clypeus black,
reddish or white;
with face
sometimes with a more or less wide white spot; legs with coxae,
antennae distally thickened
legs
clubs of femora and tibiae brown;
in female,
not thickened
in male;
antennal
segments in following proportions: (F) 3: 3,5: 4,5: 4,5: 5: 5: 5: 6: 6: 10; (M) 3: 4: 4: 4: 4,5: 5: 6: 6: 6: 8; head dull, fully granulate; frontal I ine absent; scutum,
occipital
carina complete;
POL=8;
OL=5;
scutellum and metanotum sl ightly granulate;
reaching only approximately 0,3
length of scutum;
OOL=S;
OPL=3;
TL=4;
notaul ices invisible or
propodeum reticulate
Aphelopinae
71
rugose, with or without longitudinal keels and median area on the posterior surface; fore-wings hyaline, without dark transversal bands; tibial
spurs 1,1,2.
lOCUS TYPICUS: Jacksonville (Florida,
U.S.A.)
TYPICAL MATERIAL: holotype F! of A. rufiventris in WA; lectotype F! (hereby designated) of A. affinis in WA (in WA a female labelled Aphelopus tibialis Ashmead Type agrees with the original description of A. affinis and is hereby designated as lectotype of A. affinis: A. tibialis was never described). HOSTS: unknown DISTRIBUTION: Nearctic region: CANADA: Gagnon Rd. (Terrace, B. C.), Mc Murray (Alta.), OT! Ottawa (Ont.), OT! Ol! St. lawrence Is. Nat.
OT!
Park (Ont.), OT! Ol! Kouchibouguac N. P. (N. B.), OT! Ol! U.S.A.: Portal (Chiricahua Mts., Cochise Co., Ariz.), SCI Speir Rch. (Uvalde, Uvalde Co., Tex.), SC! Jacksonville (Fla.), WA! Wattacoo (Pickens Co., C.),
TW!
OLl
Pink Beds (N. C.),
TW!
Washington
(D.
C.),
WA!
S.
IT!
2. Aphelopus bicolor Fenton
Aphelopus bicolor Fenton 1924: 192. Aphelopus bicolor Fenton: Muesebek and Walkley 1951: 1042. Aphelopus bicolor Fenton: Krombein 1979: 1250.
FEMALE AND MALE: fully winged; length 1,37-1,59 mm; testaceous or reddish dark, with propodeum, pet iole and abdomen reddi sh-brown; antennae brown, with segments 1-2 or 1-3 testaceous; sometimes propodeum fully black or with only the dorsal surface reddish-brown; antennae distally thickened in female, not thickened in male; antennal segments in following proportions: (F) 3,5: 3: 4: 4: 4: 4: 4: 4: 4: 6; (M) 5: 5: 7: 7: 7: 7: 6,5: 6,5: 6: 10; head dull, granulate; frontal line absent; occipital carina complete; POL = 6; Ol = 3,5; OOl = 3; OPl = 3,5; Tl = 3; scutum, scutellum and metanotum dull, granulate; notaulices incomplete,reaching approximately 0,5 length of scutum; propodeum reticulate rugose, with two longitudinal keels on the posterior surface; median area partly smooth; fore-wings with a dark transversal band beneath the pterostigma; tibial s p u rs 1, 1 , 2 • lOCUS TYPICUS: Spring Hill (Iowa, U.S.A.) TYPICAL MATERIAL: holotype F! in WA. HOSTS: in U.S.A. Hymetta trifasciata (Say) (Fenton
1924;
Krombein
1979).
ClISTRIBUTION: Nearctic region: CANADA: Kouchibouguac N. P. (N. B.), OT! Jl! U.S.A.: Spring Hill (Iowa), WA! Arkansas (Muesebeck and Walkley 1951; Krombein 1979); Cleveland (S. C.), TW! Yorklin (Del.), Ol!
72
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
3. Aphelopus varicornis Brues
Aphelopus varicornis Brues 1906b: 143. Aphelopus varicornis Brues: Fenton 1918b: 277. Aphelopus microtomus Wh i ttaker 1928: 389 (n. sy n. ); locus typicus Chilliwack (Canada). Aphelopus pilicornis Whittaker 1928: 389 (n. syn.); locus typicus Ch i II iwack (Canada). Aphelopus microtomus Whittaker: Muesebeck and Walkley 1951: 1042.
Aphelopus varicornis Brues: Muesebeck and Walkley 1951: 1043. Aphelopus microtomus Wh i ttaker: Krombei n 1979: 1250. Aphelopus varicornis Brues: Krombei n 1979: 1250.
FEMALE AND MALE:
fully
winged;
length
1,25-2,20 mm;
black;
mandibles
testaceous; usually clypeus also testaceous or whitish; face sometimes fully black, usually with a more or less wide whitish or testaceous spot; antennae fully occasionally
brown or with segments 1-2 or only
scutum partly
testaceous;
and stalk of hind femora brown; antennae distally
thickened
absent or present;
in female,
occipital
TL = 5;
scutum,
notaulices incomplete,
not thickened
testaceous;
in male;
antennal
(F) 5: 4: 5: 5: 5,5: 5: 5,5: 4: 4: head dull, granulate; frontal line
carina complete;
POL = 5;
OL = 3;
scutellum and metanotum dull,
5;
OOL = 4;
granulate;
reaching approximately 0,65-0,75 length of scutum;
propodeum reticulate rugose,
with 2 longitudinal
surface;
smooth;
median area partly
LOCUS TYPICUS:
with hind tibiae
sometimes hind tibiae fully
segments in following proportions: (M) 5: 4: 5: 6: 6: 6: 6: 5: 5: 8; OPL = 4;
1 testaceous;
legs testaceous,
tibial
keels on
the posterior
spurs 1,1,2.
Woods Hole (Massachusetts,
U.S.A.).
varicornis in ML; holotype M! and 1 paratype F! of A. microtomus in BM; 1 para type M! of A. microtomus in LA; holotype M! and 8 para types (6 FF, 2 MM)! of A. pilicornis in BM; 3 paratypes MM! of A. pilicornis (two very damaged) in LA. TYPICAL MATERIAL:
HOSTS:
of A.
holotype F!
unknown
DISTRIBUTION:
Nearctic region:
ALASKA:
OT!
Mile 140 Alaska Steese Hwy.,
OT!
Hixon
(B.C.),
(B.
OT!
C.),
OT!
Oliver (B.
OT!
Kleanza Cr. C.),
OT!
M.
289 Richard Hwy.
CANADA: (Terrace,
Spring Creek
Norman Wells B.
C.),
OT!
(Terrace,
(Shaw Crk.), (N.
T.),
W.
Burnaby
B.
C.),
OT!
Terrace (B. C.), OT! Cultus Lake (B. C.), OT! Robson (B. C.), OT! Chilliwack (B.C.), BM! LA! Galiano (B. C.), BM! Quadra Is. (B. C.), Chowichan L. C.),
OT!
OT!
Hatzic Lake (B.
Diamond Head Trail
(Banff Nat. OT!
(B. C.),
(Squamish,
B.
C.),
OT!
C.),
OT!
Pitt Meadows
Johnston Canyon
Park,
Alta.),
OT!
Medicine Hat.
(Alta.),
Elkwater Lk.
(Alta.),
OT!
Oldman River
(Lethbridge,
Jumping Pd.
Cr.
(Calgary,
Alta.),
OT!
SC!
(B.
OT!
Kenosee (Sask.),
Waterton Alta.), OT!
(Alta.), OT!
La Ronge
Aphelopinae
73
(Sask.), AM! Int. Peace Gardens (Turtle Mtn. For. Res., Man.), OT! Innisville (Ont.), OT! Ottawa (Ont.), OT! Ol! One Sided lake (Ont.), Marmora
(Ont.),
OT!
Ol!
Finland
(Ont.),
OT!
Bell's Corners
(Ont.),
OT!
OT!
Oxford Mills (Ont.), OT! North Branch (Ont.), OT! Ol! Maynooth (Ont.), OT! Tillsonburg (Ont.), OT! Dundas (Ont.), OT! Hull (Que.), OT! Ol! Cap Rouge (Que.), OT! Summit of King Mt. (Old Chelsea, Que.), OT! Kouchibouguac N. P. (N. B.), OT! U.S.A.: Corvallis (Oreg.), TW! Ochoco Creek (Oreg.), TW! Selma (Oreg.), TW! Mt. Hood (Oreg.), TW! Pinehurst (Oreg.), TW! H)'att Reservoir (Oreg.), TW! lowman (Idaho), TW! Stanley (Idaho), TW! OLl Salmon Creek Campground (Sierra City, Sierra Co., Calif.), CAl Bilby (Alta, Calif.), CAl Arroyo Seco Campgr. (Monterey Co., Calif.), SCI Oynard (Ventura Co., Calif.),SC! Crescent City (Calif.), TW! Somerset (EI Dorado, Calif.), TE! Clark Fork River (26 mi. NE Strawberry, Alpine Co.,
Calif.),
AM!
S.l.O. Reservoir (S.l.O.
Co.,
Calif.),
TE!
Atescadero (S.l.O. Co., Calif.), TE! Portal (Cochise Co., Chiricahua Mts., Ariz.), TW! SCI lee Creek (1,5 mi. SW lolo Hot Springs, Missoula Co., Mont.), AM! Doolittle Ranch (Mt. Evans, Colo.), OT! Speir Rch. (Uvalde, Uvalde Co., Tex.), SCI Heep Farm (Austin, Travis Co., Tex.), TE! lawrence (Kans.), TW! Williamsville (Mo.), OT! Ann Arbor (Mich.), TW! Ol! Indian Gap (Tenn.), OT! Dade Co. (Fla.), Ol! Pine Mountain (Rabun Co., Ga.), OT! Wattacoo (Pickens Co., S.C.), Ol! TW! Greenville (S.C.), TW! Cleveland (S.C.), TW! Highlands (N.C.), TW! OT! Clingman's Dome (Grt. Sm. Mt. Nat. Park, N.C.), OT! Pink Beds (N.C.), TW! Pisgah Mt. (N. C.), TW! Reed's Gap (Augusta Co., Va.), OT! Mt. Jackson (Shenandoah Co., Va.), OT! Takoma Park (Md.), TW! High Point St. Pk. (N.J.), TW! Moorestown (N.J.), TW! Farmingdale (N.Y.), TW! Ithaca (N.Y.), TW! Pughkeepsie (N.Y.), TW! Shokan (N. Y.), TW! Milford Center (N.Y.), TW! Hancock (N.Y.), TW! Kalbfleisch Field Station (Huntington, long Island, N.Y.), AM! lake Placid (N.Y.), OT! Cornell University (Ithaca, N.Y.), IT! Woods Hole (Barnstable Co., Mass.), Ml! Mt. Ktaadn (Maine), WA! Lakes of the Clouds (Mt. Washington, N. H.), OT! MEXICO: EI Saito (Durango), OT! la Ciudad (Durango), OT! EI Palmito (Sinaloa), OT! Concordia (Sinaloa), OT! Toluca (Mexico), OT! Ixtapan de la Sal (Mexico), OT!
4. Aphelopus albopictus Ashmead
Aphelopus albopictus Ashmead 1893: 101. nec Aphelopus albopictus Ashmead: Ashmead 1895: 787 Aphelopus arizonicus R.C.l. Perkins 1907: 53 (n. syn.); locus typicus Nogales (U.S.A.). Aphelopus viduus R. C. l . Perk i ns 1907: 54 (n. syn.); locus typicus Nogales (U.S.A.). Aphelopus pulcherrimus R.C.l. Perkins 1907: 54 (n. syn.); locus typicus Nogales (U .S.A.). Aphelopus microleucus R.C.l. Perkins 1907: 55 (n. syn.); locus
74
Mem. Amer.
Ent.
Inst.,
no.
37,
1984
typicus Nogales (U.S.A.). Aphelopus dikraneuri Fenton 1918b: 276 (n. syn.); locus typicus Columbus (U.S.A.).
Aphelopus Aphelopus Aphelopus Aphelopus
microleucus R.C.L. Perkins: Fenton 1918b: 276. arizonicus R.C.L. Perkins: Fenton 1918b: 277. viduus R.C.L. Perkins: Fenton 1918b: 277. comesi Fenton 1918b: 277 (n. syn.); locus typicus Columbus
(U.S.A.) •
Aphelopus albopictus Ashmead: Fenton 1918b: 278. Aphelopus pulcherrimus R.C.L. Perkins: Fenton 1918b: 278. Aphelopus typhlocybae Muesebeck 1935: 167 (n. syn.); locus typicus Poughkeepsie (U.S.A.).
Aphelopus typhlocybae Muesebeck: Dumb I eton 1937: 869. Aphelopus albopictus Ashmead: Muesebeck and Walkley 1951: 1042. Aphelopus arizonicus R.C.L. Perkins: Muesebeck and Walkley 1951: 1042.
Aphelopus comesi Fenton: Muesebeck and Walkley 1951: 1042. Aphelopus dikraneuri Fenton: Muesebeck and Walkley 1951: 1042. Aphelopus microleucus R.C.L. Perkins: Muesebeck and Walkley 1951: 1042.
Aphelopus pulcherrimus R.C.L. Perkins: Muesebeck and Walkley 1951: 1042.
Aphelopus typhlocybae Muesebeck and Walkley 1951: 1042. Aphelopus viduus R.C.L. Perkins: Muesebeck and Walkley 1951: 1043. Aphelopus albopictus Ashmead: Krombein 1958: 100. Aphelopus albopictus Ashmead: Krombei n 1979: 1250. Aphelopus arizonicus R.C.L. Perkins: Krombein 1979: 1250. Aphelopus comesi Fenton: Krombei n 1979: 1250. Aphelopus dikraneuri Fenton: Krombein 1979: 1250. Aphelopus microleucus R.C.L. Perkins: Krombein 1979: 1250. Aphelopus pulcherrimu sR. C. L. Perk i ns: Krombe i n 1979: 1250. Aphlopus typhlocybae Muesebeck: Krombei n 1979: 1250. Aphelopus viduus R.C.L. Perkins: Krombein 1979: 1250.
FEMALE AND MALE: fully winged; length 1,31-2,25 mm; black or reddishbrown; mandibles testaceous or whitish; clypeus black, brown, testaceous, or whitish; face fully black or with a more or less wide whitish spot; antennae fully brown or with segments 1-2 or only 1 testaceous; legs fully testaceous or with some parts more or less brown; sometimes legs almost fully brown, with only tarsi testaceous; sometimes only hind tibiae and stalk of hind femora brown; antennae distally thickened in female, not thickened in male: antenna I segments in following proportions: (F) 4: 4: 4: 5: 4,5: 5: 4: 4: 4: 6; (M) 5: 4,5: 5,5: 6: 6: 6: 6,5: 6: 6: 8; head dull, granulate; frontal I ine absent or present; occipital carina complete; POL = 6; OL = 4; DOL = 4; OPL = 3; TL = 3; scutum, scutellum and metanotum dull, granulate; notaulices incomplete, reaching approximately
Aphelopinae
0,5 length of scutum;
propodeum reticulate rugose,
75
with 2 longitudinal
keels on the posterior surface and with median area rugose or partly smooth; fore-wings hyaline, without dark transversal bands; tibial spurs 1,1,2. LOCUS TYPICUS: Washington
(D.C.,
U.S.A.).
TYPICAL MATERIAL: holotype F! of A. albopictus in WA; lectotype F! and 2 para lectotype FF! (designated by Olmi 1982) of A. arizonicus in B; lectotype M! and 1 para lectotype M! (designated by Olmi 1982) of A. viduus in B; holotype F! of A. pulcherrimus in B; lectotype M! (designated by Olmi 1982) of A. microleucus in B; holotype M! and 2 para type FF! of A. dikraneuri in WA; holotype M! of A. comesi in WA; holotype F! and 49 paratypes (3 seen !) of A. typhlocybae in WA; 2 paratypes (IF, 1M)! of A. typhloc~Tbae in BM •• HOSTS: in U. S.A. Typhlocyba pomaria Mc Atee (Muesebeck 1935), Dikraneura sp. (R.C.L. Perkins 1907), Typhlocyba sp. (R.C.L. Perkins 1907), Erythroneura sp. (R.C.L. Perkins 1907), Forcipata sp. (probably loca De L.
and C.)
(Krombein 1979); according to Fenton 1918b the host of A.
dikraneuri is Dikraneura fieberi; Erythroneura spp. (Krombein 1979); according to Fenton (1918b) the host of A. comesi is Erythroneura comes; according to Fenton (1924) also Erythroneura vulnerata Fitch; in New Zealand (imported: see Dumbleton 1937) Typhlocyba froggatti Baker. DISTRIBUTION: Nearctic region: CANADA: Rampart House (Yukon), aT! Chilliwack (B.C.), BM! Ketchum L. (B. C. ), aT! Gagnon Rd. (Terrace,
B.
C.), aT! Hixon (B. C.), aT! Terrace (B. C.), aT! Robson (B. C.), aT! Morley (Alta.), aT! Ninette (Man.), aT! Ottawa (ant. ), aT! Merivale (ant.), aT! Vineland Sta. (Ont.), aT! Marmora (Ont.), aT! Rondeau Provo Pk. (Ont.), aT! Finland (Ont.), aT! Brighton (ant.), aT! Blackburn (Ont.), aT! Cumberland (ant.), TW! Stittsville (Ont.), TW! Old Chelsea (Que.), aT! Summit King Mt. (Old Chelsea, Que.), aT! Mistassini (Que.), aT! Hull (Que.), aT! aLI Abbotsford (Que.), aT! Kouchibouguac N. P. (N. B.), aT! Saulok (N. 5.), aT! U.S.A.: Selma (Oreg.), TW! OL! Corvallis (Oreg.), TW! Seaside (Oreg.), TW! Needles (Calif.), IT! Alameda (Calif.), B! OL! Ogilby Rd. (Chocolate Mts., Imperial Co., Calif.), SCI Somerset (EI Dorado, Calif.), TE! Nogales (Ariz.), B! Portal (Ariz.), TW! Speir Rch. (Uvalde, Uvalde Co., Tex.), SCI Ames (Iowa), WA! Columbia (Boone Co., Mo.), WA! Bentonville (Ark.), WA! Lake Oachita S. P. (Ark.), aT! Ann Arbor (Mich.), BM! Columbus (Ohio), WA! Elm-Maple For. (Nashville, Tenn.), WA! Greenville (S. C.), TW! Wattacoo (Pickens Co., S. C.), TW! Cleveland (S.C.), OL! TW! Highlands (N. C.), TW! OL! Virginia (Krombein 1958); Mt. Lk. Bioi. Sta. (Va.), TW! Spring Br. (Pa.), TW! Maryland (Muesebeck and Walkley 1951); Poughkeepsie (N. Y.), TW! WA! BM! New York Agr. Exp. Stn. (N. Y.), NZ! Ithaca (N. Y.), aT! Otsego L. (N. Y.), TW! Canajoharie (N. Y.), TW! Shokan (N. Y.), TW! Milford Center (N. Y.), TW! Lewisboro (Westchester Co., N. Y.), AM! South Haven (Long Island, N. Y.), AM! Washington (D. C.), WA! Mt. Washington (N. H.), aT! MEXICO: 6 Km S of Maolinco (Veracruz, 1700 m), AM! Neotropic region: the record from Grenada I. (Ashmead 1895) is referable
76
Mem.
En t.
Amer.
I nst.,
to the new species Aphelopus diffusus Austra I i an reg ion: probably
NEW ZEALAND
unsuccessfully
no.
37,
1984
(see Neotropic species).
(imported as A.
typhlocybae: Dumb I eton 1937;
introduced).
APHELOPUS
NEOTROPIC SPECIES OF
Nine species are known.
KEY TO THE SPECIES
FEMALES AND MALES
-
Frontal
line complete ••••••••••••••••
Frontal
I ine absent
2 Scutum testaceous, -
Scutum black,
or
whi te
brown
or
3 Species fully whitish, antennae almost
1.
tropicalis n. sp.
incompl ete •••••••••••••••••••••••••••.•••••• 2 or yellow •••••••.•.••••••••••••••••••••••• 3
reddish-black ••••••••••••••••.•.•.••••••.••. except
completely
dorsal
dark ••••••
surface of
2.
7
the propodeum brown;
diaphanus n.
sp.
Species at least partly dark, differently colored •••••••••••••••••• 4 4 Prop odeum redd i sh • • . • • • • . • • • • • . . • • • •• 3. surinamensis n. sp. -
-
Propodeum black •••.••••••••••••••••••••••••••••••••.••••••••••••..• 5
5 Scutellum testaceous ••••.••••.•..•••• 4. -
6 Notaulices reaching approximately 0,5
I ength
8. -
fentoni n. sp.
Scu tel I um brown •••••••••.•••••••••••••••••••••••••••••••••••••.•••• 6
Notaulices
reaching
approximately
0,3
of
scutum •••••••••••.•.
alvarenganus n.
length
of
9. jamaican us n.
7 Notaulices almost
invisible,
slightly
sp.
scutum .•••••••••••••
visible near
sp.
the anterior
margin of the scutum •••••••••••••••••• 5. leucopus Kieffer Notaulices reaching at least 0,5 length of scutum •.•••••••••••••••• 8
8 Notaulices reaching about
0,5
length
.. . . •. . .. . . .. . . . . . . . . . . . .. . . .. . . . . . " -
of
6.
scutum •••••••••••••••••.••••
diffusus n.
sp.
0,65-0,75 length of scutum •••••••• . . . . . . . . . . . . . . . . . . . • . . . . . . • . . • . . • • • .• 7. trinitatis n. sp.
Notaulices
reaching approximately
1. Aphelopus tropicalis n. sp.
FEMALE AND MALE:
fully
winged;
clypeus and mandibles whitish; whitish;
antennae brown;
length 1,81-2,25 mm;
black;
in male malar space and
sometimes antennal
in female
lower face also
segment 1 testaceous;
legs
Aphelopinae
77
testaceous; sometimes coxae, clubs of femora and tibiae of mid and hind legs brown; clubs of fore femora dark; antennae distally thickened in female, (F)
not
thickened
in male;
antennal
5: 5: 7: 8: 9: 9: 9: 7: 7:
dull,
granulate,
frontal
2,5; TL = 3; scutum,
11;
segments in following proportions:
5: 5: 6: 9: 8: 8: 8: 7: 7: 9; head
(M)
line complete;
scutellum
POL = 7;
OL = 4;
and metanotum dull,
OOL = 4;
granulate;
incomplete,_ reaching approximately 0,65
length of scutum;
reticulate rugose,
keels on
with
median area smooth transversal
bands;
LOCUS TYPICUS: in AM;
HOSTS:
propodeum
the posterior surface;
fore-wings hyal ine,
without dark
spurs 1,1,2.
Chimborazo (Ecuador) holotype F!
1 paratype M!
2 paratypes MM!
in CA;
1 paratype
in TW;
in OL ..
unknown
DISTRIBUTION:
Neotropic region:
Chimborazo (30 mi. Puerto
longitudinal
and shiny; tibial
TYPICAL MATERIAL: M!
two
OPL =
notaulices
Ibanez
TRINIDAD:
SW of Alausi),
(Aysen),
CAl
Arima Valley,
Ascazubi-Pito,
TW!
AM!
ECUADOR:
OL!
CHILE:
TW!
2. Aphelopus diaphanus n. sp.
FEMALE:
fully
winged;
length
1,68 mm;
whitish,
with dorsal
surface of the
propodeum and antennal segments 5-8 brown; antennae distally thickened; antennal segments in following proportions: 4: 4: 4,5: 5: 5: 5: 5: 4,5 (last 2 segments missing line absent;
occipital
3; TL = 2,5; scutum, incomplete,
in
the holotype);
with 2 longitudinal
median area smooth;
granulate;
Ol = 4;
scutellum and metanotum dull,
reaching approximately 0,5
reticulate rugose,
head dull,
POL = 5;
carina complete;
granulate;
length of scutum; keels on
fore-wings hyaline,
frontal
OOl = 3;
OPl =
notaulices
propodeum
the posterior surface;
without dark transversal
bands;
tibial spurs 1,1,2. MALE: unknown lOCUS TYPICUS:
Gamboa
TYPICAL MATERIAL:
(Panama)
holotype F!
HOSTS: unknown DISTRIBUTION: Neotropic region: Zone),
in LEo PANAMA:
Gamboa
(Pipeline Road,
Canal
LE!
3. Aphelopus surinamensis n. sp.
FEMALE: unknown MALE: fully winged; and vertex and
lateral
dark;
length 1,56 mm;
antennae brown,
head whitish,
with
side of the thorax whitish;
with ocellar region
segments 1-3 testaceou5; scutum,
ventra I
scutellum and propodeum
78
Mem.
testaceous-dark;
abdomen
distally thickened; 4,5: 5:
5:
occipital scutum,
5:
Amer.
4,5:
En!.
Inst.,
testaceous-brown;
antenna I segments 4:
4:
6,5;
carina complete;
head dull,
POL; 5;
granulate;
OL ;
3,3;
DISTRIBUTION:
Neotropic region:
3;
4:
3,5:
line absent;
OPL ;
2,5;
TL ;
propodeum reticulate rugose, median area smooth;
bands;
tibial
spurs 1,1,2.
in LE. SURINAME:
Billiton,
LE!
4. Aphelopus fen toni n. sp.
FEMALE:
fully
region brown;
winged;
length 2,06 mm;
antennae brown,
and clypeus testaceous; metap leura,
scutum and scutellum
legs testaceous;
5;
Ol ;
granulate;
3,5;
OOl ;
metanotum dull,
3;
frontal OPl ;
granulate;
0,5 length of scutum;
6:
testaceous;
4,5: 5,5:
line absent; 2,5;
Tl ;
5:
mandibles
mesopleura,
propectus testaceous; 4,5: 4,5:
occipital
2,5;
hyaline,
without dark transversal
tibial
4:
6; POL
scutellum and
with 2
longitudinal
median area smooth and shiny; bands;
4:
reaching approximately
propodeum reticulate rugose,
the posterior surface;
antenna I 4:
carina complete;
scutum,
notaulices incomplete,
keels on MALE:
with ocellar
antennae distally thickened;
segments in following proportions: head dull,
head testaceous,
with segments 1-2 testaceous;
metanotum and propodeum brown-b lack;
abdomen brown;
fore-wings
spurs 1,1,2.
unknown
lOCUS TYPICUS:
Nova Teutonia (Brazil)
TYPICAL MATERIAL: HOSTS:
holotype F!
in BM.
unknown
DISTRIBUTION:
Neotropic region:
BRAZil:
Nova Teutonia (S.ta Catarina),
BM! NOTE:
the species is named
in honor of the
late special ist Mr.
F.
A.
Fenton.
5. Aphelopus leucopus Kieffer
Aphelopus leucopus Kieffer 1906a: 239.
FEMALE:
fully
winged;
antennae brown,
length
1,25-1,62 mm;
black;
with segments 1-2 testaceous;
2;
notaulices incomplete,
keels on the posterior surface;
holotype M!
antennae not
frontal
OOL ;
granulate;
fore-wings hyaline, without dark transversal LOCUS TYP I CUS: Bill i ton (Suri name) TYPICAL MATERIAL: HOSTS: unknown
1984
legs testaceous;
approximately 0,65 length of scutum;
with 2 longitudinal
37,
in following proportions:
scutellum and metanotum dull,
reaching
no.
mandibles testaceous;
legs testaceous,
with hind
Aphelopinae
coxae,
hind femora and hind tibiae brown;
antennal 3,5:
79
6;
segments
in following proportions:
head dull,
complete;
=
POL
granulate;
=
6; OL
3;
frontal
invisible,
slightly
=
OOL
scutellum and metanotum dull,
4:
4,5: 4:
2,5;
OPL
granulate;
=
4: 4:
3,5:
occipital
carina
=
scutum,
3,5;
TL
notaulices
3;
incomplete,
3,5:
almost
visible near the anterior margin of the scutum;
propodeum reticulate rugose,
Managua
TYPICAL MATERIAL: HOSTS:
4:
line absent;
with 2 longitudinal
surface; median area smooth and shiny; transversal bands; tibial spurs 1,1,2. MALE: unknown LOCUS TYPICUS:
antennae distally thickened;
keels on
the posterior
fore-wings hyaline,
without dark
(Nicaragua)
holotype F!
in CA.
unknown
DISTRIBUTION: (Tabasco),
Neotropic region:
NICARAGUA:
Managua,
CAl
MEXICO:
Teapa
BM!
6. Aphelopus diffusus n. sp.
;"ec Aphelopus albopictus Ashmead 1893: 101. Aphelopus albopictus Ashmead: Ashmead 1895: 787. Aphelopus albopictus Ashmead: Ashmead 1900: 235.
FEMALE AND MALE: reddish-dark;
fully
winged;
face usually
sometimes head almost fully testaceous or whitish; testaceous;legs fully tibiae brown; male; 8:
antennal
7,5:
frontal OPL =
10;
1,56-2,12 mm;
testaceous;
testaceous,
TL = 2;
4:
5:
6:
occipital
scutum,
notaulices incomplete,
reddish or
clypeus and mandibles black or
sometimes with hind femora and hind thickened
in female,
segments in following proportions:
(M) 4,5:
black,
antennae fully brown or with segments 1-2
antennae distally
line absent; 1;
length
with a more or less wide whitish spot;
6:
7:
6:
6:
6:
carina complete;
7;
not
(F) 4:
4:
5:
head dull,
POL
=
7;
scutellum and metanotum dull,
reaching
thickened
approximately 0,5
6:
in 7:
8:
8:
granulate;
=
OL
3;
OOL
=
4;
granulate;
length of scutum;
propodeum reticulate rugose, with 2 longitudinal keels on the posterior surface; median area smooth and shiny; fore-wings hyaline, without dark transversal
bands;
tibial
spurs 1,1,2.
LOCUS TYPICUS: Nova Teutonia (Brazil) TYPICAL MATERIAL: holotype M! and 1 para type F!
in BM;
2 paratypes
MM! in WA; 5 para types (4 MM, 1 F)! in OT; 1 paratype M! paratypes MM! in AM; 5 paratypes (2 FF, 3 MM)! in OL. HOSTS: unknown D I STR I BUT I ON: Neotrop ic reg ion:
in TW;
2
GRENADA I.: SI. John's River (Aphelopus albopictus Ashmead misident.: Ashmead 1895), WA! Balthazar (Aphelopus albopictus Ashmead misident.: Ashmead 1895), WA! ANTIGUA I.: 51. John's,
80
Mem.
AM!
COLOMBIA:
ECUADOR: OT!
Amer.
Ol!
Insl.,
Municipio la Cumbre
Portete de Tarqui,
BM!
Enl.
TW!
Estacao Forestal
no.
(Hda.
BRAZil:
37,
1984
la Maria,
Valle),
Nova Teutonia
Cabeca do Veado
Ol!
(S.ta Catarina),
(Distrito Federal),
OT!
Ol!
7. Aphelopus trinitatis n. sp.
FEMALE AND MALE:
fully
winged;
more or less wide whitish
spot;
length 1,62-1,87 mm;
black;
face with a
clypeus and mandibles whitish or
testaceous; antennae brown, with segmen ts 1-2 or 1-3 testaceous, somet i mes with mid and hind coxae and femora brown; sometimes also hind tarsi brown;
antennae distally thickened
antennal 5:
7;
(M)
absent; Tl
=
in female;
segments in following proportions: 6: 8: 8:
6:
occipital
8:
9:
9:
not
(F)
thickened
4:
3,5:
8: 9; head dull,
8:
carina complete;
POL; 6;
Ol ;
4:
in male;
5:
6:
granulate;
3;
6:
7:
6:
frontal
OOL = 3,5;
line
OPL = 3;
3; scutum, scutellum and metanotum dull, granulate; notaulices
incomplete, reaching approximately 0,65-0,75 length of scutum; propodeum reticulate rugose, with 2 longitudinal keels; median area smooth and shiny;
fore-wings hyaline,
1,1,2. lOCUS TYP ICUS:
without dark transversal
Chaguaramas Bay
bands;
tibial
spurs
(Trinidad)
TYPICAL MATERIAL: holotype M! and 8 paratypes (1 F, 7 MM)! in BM; paratypes (8 MM, 7 FF)! in aT; 5 paratypes (4 MM, 1 F)! in TW; 1 paratype M! in AM; HOSTS: unknown DISTRIBUTION:
4 paratypes (1
Neotropic region:
3 MM)!
F,
TRINIDAD:
15
in Ol.
(51.
Chaguaramas Bay
George),
BM! Arima Valley (51. George), BM! Ol! Curepe (S.ta Margarita), BM! GRAND CAYMAN: Georgetown, OT! MEXICO: Teapa (Tabasco), BM! S. Cristobal OT!
de las Casas (Chiapas),
COLOMBIA:
ECUADOR: Quito-S.
TW!
Ol!
Yerba Buena
(Hda.
la Maria,
(Bochil,
Valle),
Chiapas),
AM!
Tulcan,
TW!
loja,
Tarqui,
Ol!
Vn.
Chiles,
TW!
Troya,
Domingo,
OL!
Portete de Tarqui,
Ol!
Papallacta,
TW!
BRAZil:
Nova Teutonia Veado,
OT!
Municipio La Cumbre
(S.ta Catarina),
Distrito Forestal),
OT!
OT!
BM!
Ol!
ARGENTINA:
Estacao Forestal
TW!
(Cabeca do
Horco Molle (Tucuman),
OT!
8. Aphelopus alvarenganus n. sp.
FEMALE:
fully
of the face,
winged;
1-2 testaceous;
scutum testaceous;
propodeum black;
abdomen brown;
distally thickened; 6:
6:
5:
5:
length 2 mm;
head testaceous,
clypeus and mandibles whitish;
5:
9;
carina complete;
antennal head dull,
POL = 5,5;
with anterior region
antennae brown,
with segments
scutell um and metanotum brown; legs fully
testaceous-whitish;
segments in following proportions: granulate; Ol = 4;
frontal
OOl = 3;
line absent;
OPL = 2;
antennae 5:
4:
6:
occipital
Tl = 2;
scutum,
6:
Aphelopinae
scutellum and metanotum dull, approximately 0,5 central
granulate area on
without MALE:
dark transversal fully
winged;
OPL = 1,5;
4,5: 4: TL = 2;
LOCUS TYP I CUS:
tibial
with
a
spurs 1,1,2.
structurally resembles the female;
antenna I segments in following
6: 6: 6: 6: 6: 9;
POL
=
5; OL
=
4;
OOL
=
4;
spurs 1,1,2.
holotype M!
in TW;
Braz i I)
1 paratype F!
in OL.
unknown
DISTRIBUTION: TW!
thickened;
reaching
fore-wings hyal ine,
Represa Rio Grande (Guanabara,
TYPICAL MATERIAL: HOSTS:
bands;
5: 6: tibial
notaul ices incomplete,
propodeum reticulate rugose,
the posterior surface;
length 2 mm;
antennae not distally proportions:
granulate;
length of scutum;
81
Neotropic region:
BRAZIL:
Represa Rio Grande (Guanabara),
OL!
9. Aphelopus jamaican us n. sp.
FEMALE: MALE:
unknown
fully
winged;
region of the face, scutum
length
testaceous;
thickened;
antennal 8,5;
legs testaceous;
head dull,
granulate;
with 2 longitudinal
median area rugose;
frontal
propodeum black; 3:
4: 4,5: 5:
line absent;
slightly granulate;
keels on
fore-wings hyaline,
Hardwar Gap
TYPICAL MATER IAL: HOSTS:
6:
occipital
without dark
transversal
(Jamaica)
holotype M!
Neotropic region:
in OT. JAMAICA:
Hardwar Gap,
AUSTRAL I AN SPEC I ES OF APllELOPUS
Five species are known
(one introduced).
KEY TO THE SPECIES
FEMALES AND MALES
propodeum
the posterior surface;
unknown
DISTRIBUTION:
anterior
antennae not di sta II y
spurs 1,1,2.
LOCUS TYPICUS:
with
antennae testaceous;
segments in following proportions:
scutellum and metanotum dull,
reticulate rugose, tibial
testaceous-dark,
POL = 5,5; OL = 4; OOL = 3,5; OPL = 1; TL = 2; scutum, notaulices incomplete, reaching approximately 0,30 length
carina complete; dull, granulate; of scutum;
head
scutellum and metanotum brown;
abdomen brown-testaceous; 6,5: 6,5: 7: 7:
1,56 mm;
clypeus and mandibles whitish;
OT!
bands;
82
Mem.
Head and
thorax
Amer.
Ent.
no.
37,
1984
testaceous . . . . . . . . . . . . . . . . .
townesi n.
• • . • • • • • • • • • • • • • • • • • • • • • • 1. -
Inst.,
sp.
thorax mostly black . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2
Head and
2 Notaulices reaching approximately 0,65-0,75 length of -
Notaulices
3 Frontal
reaching
line absent
0,5
or
(invisible near
incomplete
oce I Ius) .••.•••••.••••••• 3.
length
australis n.
of
scutum . . . . . . . 3
scutum . . . . . . . . . . . . . 4
approximately
the anterior
sp.
- F ron t a I lin e comp let e ••.• 4. caledonicus n. sp. 4 Posterior surface of the propodeum with median area shiny and smooth, not rugose . . . . . . . 2. guineensis n. sp. -
Posterior
surface of
the propodeum with median area dull
and
albopictus Ashmead (see Nearct ic species)
rugose . . . . . . . . . . . . . . . . . . . 5.
1. Aphelopus townesi n. sp.
FEMALE: MALE:
unknown
fully
winged;
length
segments 4-10 dark, antennae not
distally
proportions: 5: I ine complete;
5:
7:
thickened; 8:
= 8;
POL
carina complete;
1,75-2,43 mm;
metanotum brown,
scutum
8:
9:
antennal
8:
= 6;
OL dull,
testaceous,
7:
7:
OOl
segments
9; head dull, granulate; frontal OPl = 6; Tl = 5; occipital
granulate;
notaulices incomplete,
length of scutum;
sl ightly
propodeum reticulate rugose,
granulate;
between
shiny dark
dorsal
and smooth,
lOCUS TYP I CUS:
Atherton
with a strong
transversal
posterior surface centrally
laterally reticulate rugose;
TYPICAL MATERIAL:
reaching
scutellum and metanotum shiny,
and posterior surface;
transversa I bands;
abdomen dark;
in following
= 4;
approximately 0,65 keel
with antennal
propodeum black,
fore-wings hyal ine,
without
t i b i a I spurs 1,1,2. (Austra I i a)
holotype M!
and 1 paratype M!
in TW;
1 paratype M!
in Ol. HOSTS:
unknown
DISTRIBUTION: NEW GUINEA: NOTE:
Australian region: Baiyer R.,
the species
TW!
is named
AUSTRALIA:
Atherton
(Queensland),
TW!
OL! Henry K.
in honor of Mr.
Townes.
2. Aphelopus guineensis n. sp.
FEMALE AND MALE: testaceous;
fully
winged;
length 1,56-1,87 mm;
clypeus brown or testaceous-dark;
black;
antennae brown,
mandibles with
segments 1-2 testaceous and segment 3 partly or fully testaceous; legs testaceous; antennae distally thickened in female, not thickened in male; antennal 5;
(M)
3:
segments 4:
4,5:
in following proportions: 6:
6:
7:
7:
6,5:
6,5:
9;
(F)
5:
4:
head dull,
5:
5:
5:
4:
granulate;
4:
4:
frontal
4:
Aphelopinae
I ine absent; 2,5;
occipital
carina complete;
83
=
POL
5;
=
Ol
3;
=
OOl
3;
OPl
= 2,5; scutum, scutellum and metanotum dull, granulate;
Tl
notaul ices
incomplete,
reaching
propodeum reticulate rugose, suerfacej
approximately 0,5 length of scutum;
with 2 longitudinal
median area smooth and shiny;
transversal
bands;
lOCUS TYPICUS:
tibial
keels on
without
dark
spurs 1,1,2.
Kundiawa
(New Guinea)
TYPICAL MATERIAL: holotype F! and 1 paratype F! in TW; 4 paratypes (1F, 3MM)! in Ol. HOSTS:
the posterior
fore-wings hyaline,
in B;
6 paratypes MM!
unknown
DISTRIBUTION:
Australian
NE New Guinea),
B!
(NE New Guinea),
region:
Hospital
B!
Tari
NEW GUINEA:
Sk.
Gap
(Wau,
(Mt.
Kundiawa
NE New Guinea),
Hagen),
TW!
(East Highlands, 8!
Kassam Pass
Ol!
3. Aphelopus australis n. sp.
FEMALE AND MALE:
fully
testaceousj
whitish
clypeus
1-2 testaceous;
winged; or
legs testaceous;
femora brown;
length 1,43-1,75 mm;
testaceousj
antennae distally thickened
antenna I segments
in following
4:
4:
6:
frontal
7;
(F)
5:
line absent;
4:
5:
mandibles
or with
segments
somet i mes with mid and hind coxae and
male; 4:
black;
antennae brown
5:
5:
in female,
proportions:
4,5:
4,5:
4,5:
occipital carina complete;
not
thickened
(M) 4,5: 7;
POL
4:
4:
head dull,
=
5,5;
5:
5:
granulate;
=
Ol
in
5:
3;
=
OOl
4; OPl = 2,5; Tl = 2; scutum, scutellum and metanotum dull, granulate; notaulices incomplete, reoaching approximately 0,65-0,75 length of scutum; propodeum reticulate rugose, surface;
transversal
bands;
tibial
lOCUS TYP ICUS: Casula TYPICAL MATERIAL: in B;
spurs
keels on
the posterior
fore-wings hyaline,
without dark
1,1,2.
(Austral ia)
holotype M!
9 paratypes MM!
HOSTS:
with 2 longitudinal
median area smooth and shiny;
in TW;
and 1 para type F! 4 paratypes
(1F,
in BM;
3MM)!
1 paratype F!
in Ol.
unk now n
DISTRIBUTION:
Australian
BM!
(Victoria),
Warburton
Yangaborra
region: TW!
(NE Queensland),
Nebo (Queensland),
TW!
AUSTRALIA:
Casula
Foot of Black Mtn. B!
Mt.
Port Arthur
Glorious
(New South Wales),
(Canberra),
(Queensland),
(Tasmania),
TW!
BM!
TW!
Ol!
Mt.
Ol!
4. Aphelopus caledonicus n. sp.
FEMALE:
fully
winged;
length
1,43-1,68 mm;
black;
mandibles testaceous;
clypeus and part of the face or part of the malar space whitish; br·own, distally
with segments 1-2 or 1-3 testaceousj thickened;
antennal
legs testaceous;
segments in following
antennae
antennae
proportions:
3,5:
3:
4:
84
5:
Mem.
5:
5:
5:
5:
5:
carina complete;
7;
Amer.
head dull,
POL; 5;
OL ;
scutellum and metanotum dull, approximately 0,65-0,75 with 2
longitudinal
Ent.
Inst.,
granulate; 3;
OOL ;
granulate;
length of scutum;
keels on
smooth or slightly rugose;
no.
37,
1984
frontal
3;
line complete;
OPL ;
2;
TL ;
4;
occipital
scutum,
notaulices incomplete,
reaching
propodeum reticulate rugose,
the posterior surface;
fore-wings hyaline,
median area shiny,
without dark
transversal
bands; tibial spurs 1,1,2. MALE: unknown LOCUS TYPICUS:
Col
TYPICAL MATERIAL: HOSTS:
d'Amieu
(New Caledonia)
holotype F!
in B;
1 para type F!
in OL.
unknown
DISTRIBUTION: NEW GUINEA:
Australian region: Umboi
I.
(1
NEW CALEDONIA:
Km N of Awelkom,
Col
d'Amieu,
NE New Guinea),
B!
DL!
5. Aphelopus albopictus Ashmead (see Nearctic species)
2. Crovettia n. gen.
TYPE SPEC I ES:
Crovettia theliae (Gahan 1918)
FEMALE: maxillary pal,:" with 5 segments; labial palpi with 3 segments; mandibles tridentate, with teeth progressing larger from anterior one to posterior; fore-wing with radial vein forming a very pronounced curve; usually distal and proximal part of radial vein forming an angle; at least the anterior surface of the scutum reticulate rugose; running close to the antennal MALE:
sclerites;
structurally resembles the female;
thickened
(in Cr.
antennae slightly DISTRIBUTION:
epistomal
suture
antennae distally thickened. antennae not usually distally
barbara from the Neotropic region, the male has distally
Nearctic,
thickened).
Neotropic
HOSTS: Membra cidae SPECIES: 7
NEARCTIC SPECIES OF CROVETTIA
Two species are known.
KEY TO THE
SPECIES
85
Aphelopinae
FEMALES AND MALES
Head,
scutum,
scutellum and metanotum fully
maxillary palpi
with
segment
1 much
longer
reticulate than
rugose; 2 ................. .
1. nearctica n. -
Head,
scutellum and metanotum not
granulate;
scutum granulate,
with
reticulate
sp. fu I I y
the anterior surface
rugose; maxillary palpi with segment •• • • • • •• • • • • •• • • • • • • • • •• • • • •• . . • • •• •
Fig. 43 -
rugose,
1 nearly as 2.
Male of Crovettia nearctica n.
theliae
sp.
reticulate
long as (Gahan)
from Forsyth
2
....... .
(Georgia).
1. Crovettia nearctica n. sp.
=EMALE:
fully
:eeth reddish;
winged;
length 2,25-3,12 mm;
antennae brown,
black;
mandibles testaceous,
with segment 1 testaceous;
coxae and clubs of femora brown;
legs testaceous,
antennae distally thickened;
antennal
with with
86
Mem.
segments palpi
Ent.
in following proportions:
with segment
1 much
and propodeum dull, carina comlete; invisible;
fully
=
POL
10;
Inst.,
8:
6:
no.
6:
longer than 2;
6:
=
5;
=
OOL
5:
9;
1984
4:
head,
reticulate rugose; OL
37,
4:
without dark transversal
shorter than proximal part
(2:8),
4:
scutum,
frontal
=
OPL
bands;
4:
7;
maxi Ilary
scutellum,
metanotum
line incomplete;
8;
TL
posterior surface of the propodeum without
wings hyaline, much
Amer.
=
10;
longitudinal
distal
occipital
notaul ices keels;
part of radial
fore-
vein
the two parts forming an angle;
tibial
spurs 1,1,2. MALE
(Figs.
testaceous;
25,
43):
fully
winged;
antennae testaceous;
Fig.
44 -
length 2,50-2,81
legs testaceous,
mm;
black;
mandibles
with coxae black and femora
Male genitalia of Crovettia nearctica n. sp.
(type).
brown; antennae not distally thickened; antennal segments in following proportions: 7: 6: 5: 7: 6: 6: 7: 6: 6: 9; maxi Ilary palpi with segment longer than 2; occipital scutum,
carina complete;
fully POL
=
reticulate rugose; 9;
OL
scutellum and metanotum dull,
invisible;
propodeum reticulate rugose,
longitudinal
keels;
part of radial forming
head dull,
fore-wings hyaline,
=
4,5 OOL
fully dull,
=
8;
OPL
without
=
8;
TL
transversal (2:7);
=
6;
notaulices
without dark transversal part
much
line incomplete;
reticulate rugose;
vein much shorter than proximal
an angle;
frontal
or bands:
distal
the two parts
gonoforceps with apex not externally emarginated
(Fig.
44);
87
Aphelopinae
spurs 1,1,2.
tibial
LOCUS TYPICUS:
Forsyth
TYPICAL MATERIAL:
(Georgia,
holotype M!
U.S.A.)
and 3 para types
paratype F!
in lO;
1 paratype F!
(1
in AM;
4 para types MM!
F,
1 M)!
HOSTS:
in SC;
(2 MM,
F)!
3 paratypes MM!
in OT;
in TW;
1
2 paratypes
in OL.
unknown
DISTRIBUTION: Ariz.),
SCi
Cleveland
Nearctic region:
Forsyth (S.
(Ga.),
C.),
TW!
Conn.), AM! South (Sinaloa), OT!
OL!
Haven
U.S.A.:
OT!
OLl
Portal
Wattacoo
Wilmington
(Long
(Chiricahua Mts., (Pickens Co.,
lO!
(Del.),
Island,
N.
Y.),
S.
Westport
AM!
OL!
Cochise Co.,
C.),
TW!
OLl
(Fairfield Co.,
MEXICO:
EI
Palmito
2. Crovettia theliae (Gahan) n. comb.
Aphelopus theliae Gahan 1918: 151. Aphelopus theliae Gahan: Muesebeck and Walkley 1951; 1042. Aphelopus theliae Gahan: Krombei n 1979: 1250.
FEMALE: teeth with
fully
winged;
reddish-brown; tarsi
light;
length 2,06-2,20 mm;
antennae distally
thickened;
7: 6: 5: 4: 4: 4: 4: 4: 4: 6;
proportions:
maxi Ilary palpi
with segment
black;
mandibles testaceous,
with segments 1-;2
antennae brown,
1 nearly
as
testaceous;
antenna I segments
head dull, long
fully
as 2;
with
legs brown;
in following
granulate;
frontal
I ine
incomplete;
occipital carina complete; POL = 9; OL = 5; OOL = 5; OPL = 3; TL = 4; scutum dull, granulate; with anterior third reticulate rugose; notaulices invisible; scutellum fully
granulate;
metanotum reticulate rugose;
propodeum fully
reticulate rugose, without longitudinal keels on the posterior surface; forewings hyaline, without dark transversal bands; distal part of radial vein
(2:4),
much shorter than proximal
part
sp u rs 1, 1 , 2. MALE: fully winged;
1,93-2,18 mm; black; mandibles testaceous,
teeth brown; light;
antennae not distally
proportions: segment
occipital
scutum dull,
invisible;
legs brown,
thickened;
two parts forming
with fore
an
angle;
tibiae and fore
antenna I segments
tibial with
tarsi
in following
8: 5: 4,5: 4: 4: 3,5: 3,5: 3,5: 3,5: 6; maxillary palpi with
1 nearly as
incomplete; = 4;
length
antennae brown;
the
long
granulate,
scutellum dull,
reticulate rugose,
as 2;
head dull,
carina complete;
without
with
POL
fully
=
9;
granulate;
OL
=
5;
OOL
frontal
=
4;
anterior third reticulate rugose;
granulate;
metanotum shiny,
longitudinal
keels on
punctate;
line
OPL
=
3;
propodeum
the posterior surface;
fore-
wings hyal ine, without dark transversal bands; distal part of radial vein much shorter than prox i ma I part (2:4), the two parts form i ng an ang Ie; gonoforceps with apex externally emarginated LOCUS TYPICUS:
Cold Spring Harbor
TYPICAL MATERIAL:
holotype F!
(New
(Fig.
York,
45);
tibial
spurs 1,1,2.
U.S.A.)
and 4 paratypes
(1
F,
3 MM)!
TL
notaulices
in WA.
88
Mem.
Fig. 45 -
Amer.
Ent.
Ins!.,
no. 37,
1984
Male genitalia of Crovettia theliae (Gahan) (New York)
from Cold Spring Harbor
HOSTS: in U.S.A. Thelia bimaculata Fabr. (Membracidae) (Gahan 1918; Kornhauser 1919). BIOLOGY: informations in Kornhauser (1916, 1917, 1919); the species is polyembryonic; the external larval sac is absent. DISTRIBUTION: Nearctic region: CANADA: Richter Pass (OsoyOOS, B. C.), OT! OL! Rockport (Ont.), OT! U.S.A.: Glenville (Kern Co., Calif.), TE! lone (Amador Co.,
Calif.),
SCI
5 mi.
W Portal
(Cochise Co.,
Ariz.),
AM!
Alexandria
(La.),
89
Aphelopinae
aT!
Ol!
Harbor Haven
Roscommon (Long
(long
(Mich.),
Island, Island,
N.
OL!
Y.),
N.
Y.),
TW!
WA!
Washington
lewisboro
C.),
Ol!
Cold Spring N.
Y.),
Ol!
South
AM!
NEOTROPIC SPECIES OF
Five species
(D.
(Westchester Co.,
CROVETTIA
are known.
KEY TO THE SPECIES
FEMALES AND MALES
Fore-wing with distal
part
of
radial
vein much
shorter
than proximal
••••• 2 part • • . . • . . .• .•. . . . •. . • . . .••••••. ••• 4 2 Head, scutum, scutellum and metanotum fully ret iculate rugose. 3. plaumanniana n. sp. - At most head, anterior surface of the scutum and metanotum reticulate rugose; scutellum never reticulate rugose .•••..••.•••••• 3 3 Head only granulate, without irregular keels; pal much longer than Ol. • • • • • • • • • • • • • • • • • . • • • • . • . • • • • • • . • 4. brasiliana n. sp. - Head granulate with irregular keels; pal shorter than Ol . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . " 5. colombian a n. sp. 4 Head ret iculate rugose . . . . . . . . . . . . . . . 1. barbara n. sp. - Head not reticulate rugose, fully granulate •••••••••. 2. neotropica n. sp. part .• '" •••...•••••••••••••.••••••• Fore-wi ng wi th d i sta I part of rad i a I
vein
longer
than
proximal
1. Crovettia barbara n. sp.
FEMALE:
fully
winged;
antennae brown,
length
and fore tibiae testaceous; in following
1,50-1,87 mm;
black;
with segments 1-2 testaceous;
proportions:
antennae distally
8:
4:
3:
2,5:
3:
3:
mandibles testaceous;
legs testaceous with thickened;
:J: 3:
3:
antennal
4;
tarsi
segments
head dull,
::wanulate and fully reticulate rugose, less strongly reticulate rugose than 'C1 male; frontal line incomplete; occipital carina complete; POL = 6; Ol 3;
OOl = 4,5;
-..Jgose only ,;-anulate, -.Jgose;
OPl = 1,5;
Tl = 2;
scutum dull,
near the anterior margin; not reticulate rugose;
granulate,
notaulices
reticulate
invisible;
scutum dull,
metanotum and propodeum reticulate
propodeum without transversal
or longitudinal
keels;
fore-wings
90
Mem.
hyaline,
fully
part
antennal
Fig.
head dull,
length
invisible;
genitalia:
hyaline,
46;
tibial
part of radial
black;
vein
longer
mandibles testaceous; 6:
4:
with
Ol = 3;
3:
3:
3:
3:
frontal
3:
3:
4,5;
line complete;
OOl = 4; OPl = 2;
Tl = 2,5;
numerous areolae on the anterior half; granulate;
metanotum and propodeum
longitudinal
keels on
without dark transversal
twice as long as proximal spurs 1,1,2.
Nova Teutonia
3:
barbara n. sp. type.
reticulate rugose;
scutellum dull,
vein more than fig.
mm;
propodeum without
fore-wings
lOCUS TYPICUS:
distal
1984
spurs 1,1,2.
POL = 6;
granulate and
37,
antennae slightly distally thickened;
granulate and fully
reticulate rugose;
bands;
no.
Male genitalia of Crovettia
46 -
scutum dull,
of radial
1,81
Ins!.,
in following proportions:
carina complete;
notaulices
tibial
legs brown;
segments
occipital
(6:4);
winged;
antennae brown;
surface;
En!.
without dark transversal
than proximal MALE:
Amer.
(Brazil)
the posterior
bands;
part
distal
(7,5:3);
part
91
Aphelopinae
TYPICAL MATERIAL: F!
HOSTS:
in BM;
1 paratype F!
in OT;
1 paratype
unknown
DISTRIBUTION: BM!
holotype M!
in OL.
Caraca
Sao Paulo),
Fig.
Neotropic region:
(S. OT!
47 -
Barbara,
BRAZil:
Minas Gerais),
Nova Teutonia OT!
Bocaina
(S.ta Catarina),
(S.
Jose Barreiro,
Ol!
Male genital ia of Crovettia neotropica n.
sp.
type
92
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
2. Crovettia neotropica n. sp.
FEMALE AND MALE: testaceous-brown; testaceous; antennal
fully
length 1,43-1,56 mm;
winged;
antennae brown;
legs brown,
antennae distally thickened
segments
with
in female,
in following proportions:
black;
tarsi
not
mandibles
and fore tibiae
thickened
in male;
5,5: 4: 4: 4: 4: 4: 4: 3: 3:
5,5; head dull, granulate; frontal line absent; occipital carina complete; POL = 5; Ol = 2,5; OOl = 3; OPl = 2,5; TL = 3; scutum dull, granulate and with nUmerous areolae near the anterior margin; scutellum and metanotum dull, without
longitudinal
without
dark
than proximal
LOCUS TYPICUS:
distal
holotype M!
Neotropic region:
invisible;
fore-wings hyaline,
part of radial
(4,5:4); male genital ia: fig. 47;
Nova Teutonia
TYPICAL MATERIAL: HOSTS: unknown DISTRIBUTION:
bands;
notaulices
propodeum reticulate rugose,
keels on the posterior surface;
transversal part
granulate;
vein sl ightly tibial
longer
spurs 1,1,2.
(Brazil) and 1 paratype F! BRAZil:
in BM.
Nova Teutonia
(S.ta Catarina),
BM!
Fig.
48 - Male genitalia of Crovettia plaumanniana n. sp. paratype
3. Crovettia plaumanniana n. sp.
93
Aphelopinae
FEMALE:
fully
winged;
length 2,18 mm;
black;
antennae brown,
with segments 1-5 testaceous;
tarsi
antennae distally
testaceous;
following
proportions:
granulate,
fully
8:
6:
5:
5:
thickened; 5:
reticulate rugose;
3,5:
antennal
4:
frontal
mandibles testaceous; legs brown,
3,5:
I ine
4:
with
tibiae and
segments in
6,5;
head dull,
incomplete; occipital
complete; POL = 9; DL = 5; DOL = 8; OPL = 8; TL metanotum dull, fully reticulate rugose; notaulices
carina
=
5; scutum, scutellum, invisible; propodeum
reticulate rugose, without transversal or longitudinal keels; fore-wings hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part MALE:
fully
winged;
antennae testaceous-dark; partly brown;
(2:5);
length 2,31
tibial mm;
legs testaceous,
antennae not distally
following proportions: reticulate rugose;
8:
spurs 1,1,2.
black;
5:
frontal
5:
6:
mandibles testaceous;
with coxae and cl ubs of femora
thickened;
5,5:
5:
5:
I ine incomplete;
antenna I segments
5:
5:
in
8; head dull, fully
face without
lateral
keels;
occipital carina complete; POL = 9; OL = 6; OOL = 9; OPL = 6; TL = 6; scutum, scutellum and metanotum dull, fully reticulate rugose; notaulices invisible;
propodeum reticulate rugose,
between dorsal longitudinal
and posterior surface;
keels,
reticulate rugose;
transversal
bands;
part
genitalia:
(2:6);
LOCUS TYPICUS:
distal
HOSTS:
48;
tibial
transversal
keel
without dark
vein much shorter than
proximal
spurs 1,1,2.
(Brazil)
holotype F!
in OT;
paratype M!
in BM.
BRAZIL:
Nova Teutonia
unknown
DISTRIBUTION: OT!
fig.
a strong
fore-wings hyaline,
part of radial
Nova Teutonia
TYPICAL MATERIAL:
with
posterior surface without
Neotropic region:
(S.ta Catarina),
BM!
Cr. plaumanniana structura II y resemb les Cr. nearctica (Nearct ic
NOTES: region).
It
is named in honor of Mr.
Fritz Plaumann.
4. Crovettia brasiliana n. sp.
FEMALE:
fully
winged;
antennae brown, light;
length
antennae distally
proportions:
8:
1,75-1,81
mm;
black;
with segments 1-2 testaceous; 5:
3:
3:
thickened;
antennal
4:
3,5:
4:
3,5:
mandibles testaceous;
legs brown,
segments
3,5:
5;
with
tarsi
in following
head dull,
granulate;
frontal line incomplete; occipital carina complete; POL = 7; OL = 4; OOL 4,5; OPL = 2; TL = 2; scutum dull, granulate, reticulate rugose only near the anterior margin;
notaul ices
incomplete,
visible only
near the anterior
Clargin of the scutum; scutellum dull, granulate; metanotum shiny, smooth, .\ ithout sculpture; propodeum reticulate rugose, without transversal and :ongitudinal distal
keels;
fore-wings hyal ine,
part of radial
S::lurs 1,1,2. '!ALE: unknown
without dark transversal
vein much shorter than
proximal
part
bands;
(2:4);
tibial
94
Mem.
LOCUS TYP ICUS:
Nova Teutonia
TYPICAL MATERIAL: HOSTS:
En!.
Ins!.,
no. 37,
1984
(Brazi I)
holotype F!
in OT;
paratype F!
in OL.
MEXICO:
Yerba Buena
(Bochil,
unknown
DISTRIBUTION: OL!
Amer.
BRAZIL:
Neotropic region:
Nova Teutonia
(S.ta Catarina),
Chiapas),
OT!
5. Crovettia colombiana n. sp.
FEMALE: brown,
fully
thickened; 4,5:
4,5:
frontal ;
8;
winged;
antennal 4,5:
6;
3;
TL ;
5;
reticulate rugose; longitudinal
with
very
carina complete;
scutum dull,
granulate,
scutellum dull,
fore-wings hyal ine,
part of radial
9:
weak
6:
8:
7:
6:
5:
irregular keels;
POL; 7,5;
OL ;
4;
OOL near
reaching approximately metanotum dull,
fully
without transversal
without dark
vein much shorter than
legs
antennae distally
reticulate rugose only
granulate;
propodeum reticulate rugose,
keels;
mandibles testaceous;
notaulices almost invisible,
length of scutum;
distal
granulate,
line incomplete; occipital
OPL ;
black;
antennae black;
segments in following proportions:
head dull,
the anterior margin; 0,5
length 2,50 mm;
with fore tibiae testaceous;
transversal
proximal
or
bands;
part (4:8);
tibial
spurs 1,1,2. MALE:
unknown
LOCUS TYPICUS:
1°10'
TYPICAL MATERIAL: HOSTS: unknown DISTRIBUTION:
N,
7°5'
1°10'N,
7°5'W
SUBFAM I L Y BIAPHELOPINAE N.
SUBF.
TYPE GENUS: Biaphelopus n. fully
winged;
tibial
maxi Ilary palpi
COLOMBIA:
(Putumayo),
OT!
gen.
fore-wing
enclosed by pigmented veins; complete;
Colombia)
in OT.
Neotropic region:
2.
FEMALE:
W (Putumayo,
holotype F!
with costal
front
tarsi
and median cells clearly
not chelate;
with 5-6 segments;
labial
occipital palpi
with
carina 3 segments;
spurs 1,1,2.
MALE: unknown DIST.RIBUTION: Palaeartic HOSTS:
unknown
GENERA:
1
SPECIES: NOTES:
1
the new subfami I y Biaphelopinae structura II y
resemb les
Conganteoninae (ma in I y due to the scu I pture of head and thorax and venation of the fore-wing). In females of Biaphelopinae, however, the fore tarsi are not chelate. Differences regarding the males of Biaphelopinae (as yet unknown)
probably
wi II
concern
the radial
vein
(in females and males
95
Biaphelopinae
of
Conganteoninae the distal part is as long as or longer than the in females of Biaphelopinae the distal part is shorter than
proximal part; the proximal
part).
Biaphelopus n. gen.
TYPE SPEC I ES:
Biaphelopus masneri n. sp.
FEMALE
14,
(Figs.
with 3 segments; wing
with
head,
proximal
scutum,
49):
maxi Ilary palpi
scutellum and metanotum fully sclerites
vein forming
an
reticulate rugose; (Fig.
31
epistomal
C).
Palaearctic
1
Fig.
49 -
Female of
fore-
angle;
unknown
SPEC I ES:
palpi
teeth;
unknown
DISTRIBUTION: HOSTS:
labial
with 2 rudimentary
and distal parts of radial
suture running close to the antennal MALE:
with 5-6 segments;
mandibles quadridentate,
Biaphelopus masneri n. sp. (type).
96
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
1. Biaphelopus masneri n. sp.
FEMALE:
fully
winged;
antennae brown,
coxae basally black;
antennae distally
following proportions: reticulate rugose, complete;
=
POL
invisible;
8:
4:
10: 8:
granulate;
5;
OL
and metanotum dull, keels;
length 2,43 mm;
black;
with segments 1-2 testaceous;
=
3;
fully
8:
frontal
=
OOL
thickened; 6,5:
5:
5:
7;
OPL
=
propodeum reticulate rugose,
6;
5:
=
TL
vein shorter the right, MALE:
than
proximal
maxillary palpi 6 on
the
(4:6);
fore tarsi
left);
labial
palpi
(in
hind
8;
fully
carina
scutum,
scutellum
notaul ices
bands, distal
or transversal with costal
and
part of radial
not chelate;
tibial
spurs
the holotype 5 segments on
with 3 segments.
unknown
LOCUS TYPICUS:
Godavari
TYPICAL MATERIAL: HOSTS:
(Nepal)
holotype F!
in OT.
unknown
DISTRIBUTION: NOTE:
part
with 5 or 6 segments
head dull,
longitudinal
without dark transversal
median cells clearly enclosed by pigmented veins; 1,1,2;
7;
occipital
granulate;
without
with
antenna I segments in
I ine absent;
reticulate rugose,
fore-wing hyal ine,
mandibles testaceous; legs testaceous,
Palaearctic region:
the species
is named
3.
NEPAL:
Godavari
in honor of Mr.
SUBFAM I L Y
OT!
CONGANTEONINAE N. SUBF.
TYPE GENUS: Conganteon Benoit 1951. FEMALE: fully winged; fore-wing with costal enclosed by pigmented veins
(Katmandu),
Lubomir Masner.
(Fig.
16);
and median cells clearly
maxi Ilary palpi
with 5-6 segments;
labial palpi with 3 segments; front tarsus chelate (Fig. 16); mandibles quadridentate, with an intermediate rudimentary tooth (Fig. 22 B); chela with rudimentary claw carina complete; MALE:
fully
tibial
winged;
(Fig.
fore-wing
enclosed by pigmented veins labial
palpi
pronotal
with costal
(Fig.
with 3 segments;
intermediate rudimentary tibial
53);
tubercles present;
tooth
26);
and median cells clearly
maxillary
palpi
mandibles quadridentate, (Fig.
22 B); occipital
with 5-6 segments; with an
carina complete;
spurs 1,1,2.
DISTRIBUTION: HOSTS:
Palaearctic,
Ethiopian,
Oriental
unknown
GENERA: SPECIES:
occipital
spurs 1,1,2.
3 5
KEY TO THE GENERA OF
CONGANTEONINAE
97
Conganteoninae
FEMALES
Enlarged
claw wi th
••• • • • •• • • • . • • • • . " -
subapi cal
tooth
•••.••.••• 2.
Enlarged
claw without subapical . • . . . . . . . . . . . . . . • . . • • . . • . • • • . 1. The females of Fiorianteon
n.
(Fig.
59) . . . . . . . . . . . . . . . . .
Chelanteon n. gen. tooth
(Fig.
56) . . . . . . . . . . . . ..
Conganteon Benoit
gen.
are unknown.
MALES
Distal
part
of
radial
vein
longer
• • • • • • • • • • • • • • • • • • • • . • • . • • •• -
Distal
part
of
I'adial
1.
vein much
60) ••••••.•••••.•••••.••.••. The males of Chelanteon
Fig. 50 -
n.
3.
gen.
than
proximal
part
(Fig.
50).
Conganteon Ben 0 i t shorter
than proximal
Fiorianteon n. gen. are unknown.
Male of Conganteon nepalense n.
sp.
(type)
part
(Fig.
98
Mem.
Amer.
Ent.
Inst.,
no. 37,
1984
1. Conganteon Benoit
Conganteon Benoit 1951c: 11.
TYPE SPECIES:
Conganteon vulcanicum Benoit 1951, orig. desig.
FEMALE:
winged;
fully
segmented;
maxi Ilary palpi
mandibles quadridentate,
with 5-6 segments;
with
an
labial
palpi
3-
intermediate rudimentary
22 B); fore-wing with short and oval pterostigma (Fig. 55); vein longer than proximal part (Figs. 16, 55); enlarged claw without subapical tooth, lamellae, bristles or hairs (Fig. 53); pronotal tubercles reaching tegu lae. MALE: fully winged; maxillary palpi with 5-6 segments; labial palpi 3segmented; mandibles quadridentate, with an intermediate rudimentary tooth (Fig. 22 B); fore-wing withshort and oval pterostigma (Fig. 50); distal part of radial vein longer than proximal part (Fig. 50); occipital carina complete. DISTRIBUTION: Palaearctic, Ethiopian HOSTS: unknown SPECIES: 3 tooth
(Fig.
distal
part of radial
PALAEARCTIC SPECIES OF
One species
CONGANTEON
is known.
Conganteon nepalense n. sp.
FEMALE: MALE
unknown
(Fig.
50):
fully
tegulae brown; OOL
=
7;
OPL
winged:
legs brown;
=
4;
reticulate rugose;
TL
=
7;
reticulate rugose; longitudinal distal
proximal
carina complete;
reticulate rugose;
with central
a curve;
genitalia:
figs.
51,
holotype M!
52 A;
(Nepal)
in B.
OL
2;
invisible;
and smooth;
metanotum
without transversal
without dark transversal
without distal
35 Km N of Trisuli
TYPICAL MATERIAL:
POL = 4;
temples fully
notaul ices
area shiny
longer than proximal
membranous process,
mandibles black;
keels
posterior surface without
fore-wings hyal ine, vein
black;
reticulate rugose;
and posterior surface;
keels;
spurs 1,1,2. LOCUS TYPICUS:
1,87 mm;
propodeum reticulate rugose,
part of radial
forming
occipital
scutum fully
scutellum reticulate rugose, between dorsal
length
head fully
part;
bands;
the two parts
gonoforceps without dorsal inner pointed process;
tibial
99
Conganteoninae
HOSTS:
unknown
DISTRIBUTION: Syabrubens),
Fig.
51
Palaearctic region:
NEPAL:
35 Km N of Trisuli
(Nawakot,
B!
-
Conganteon nepalense n. sp. (type)
Male genitalia of
ETH lOP I AN SPEC I ES OF
COiv'GANTEON
Two species are known.
KEY TO THE SPECIES
FEMALES
Segment
of
fore
tarsi
I ess
than
twi ce as
. •. . . •. . . . •. •. . . . . ••. . . •. •. . . •. ••. . . •. " -
Segment
1 of
fore
tarsi
more
than
1.
twi ce as
........................................
2.
long
as
segment
townesi n. long as
4 ...
sp.
segment
4 ...
vulcanicum Benoit
100
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
MALES
Vertex
of
the head,
reticulate
rugose,
near with
the
occipi tal
longitudinal
carina,
. . . . . . . . . . . . . . . . . . . . . . . . • . . . . . . . . • . . . . •• Vertex
of
rugose,
the head,
without
near
the occipital
longitudinal
shiny,
not
keels •.••.••••..•.••
townesi n.
1.
carina,
sp. reticulate
vulcanicum Benoit
2.
keels ••••••
dull,
B
/
52 - Gonoforceps (laterally viewed) of Conganteon nepalense (A: type) and vulcanicum Benoit (B: specimen from Mt. Hoyo)
Fig. n.
sp.
1. Conganteon townesi n. sp.
FEMALE:
fully
winged;
length 2,81
anterior part of face testaceous; coxae basally black; following proportions: reticulate rugose; shiny
and with
complete;
POL
very short;
=
black;
mandibles, testaceous,
antennae distally thickened; 10:
5:
12: 9:
9:
8:
6:
6:
5:
antennal 5:;
clypeus and with
hind
segments
head dull,
posterior region of the vertex near the occipital
longitudinal 7;
OL
=
4;
scutum shiny,
scutellum shiny,
mm;
antennae fully
smooth,
keels; OOL
fully finely
=
frontal 8;
OPL
line present;
=
8;
TL
reticulate rugose; punctate,
=
9.
occipital
in
fully carina carina
pronotum hairy,
notaul ices invisible;
without sculpture among
the
Conganteon i nae
Fig. 53 -
Chela of
Conganteon townesi n. sp.
101
(type)
102
Mem.
punctures;
metanotum sh i ny,
the punctures; longitudinal distal
(Fig.
53)
Inst.,
vein
a curve;
no.
puncta te,
longer than
tooth,
without
without dark proximal
in following proportions:
15:
bristles,
1984
wi thout scu I pture among
segment 3 of fore tarsi
without subapical
37,
reticulate rugose,
fore-wings hyal ine,
part of radial segments
Ent.
strong I y
propodeum fully
keels;
parts forming tarsal
Amer.
part
(13:10);
produced
3,5:
3:
transversal
transversal
10:
the
into a 15;
and
bands; two
hook;
fore
enlarged claw
hairs or lamellae;
segment 5
of fore tarsi (Fig. 53) with 1 row of nearly 52 small lamellae extending continuously to the apex; apex with 3 long lamellae; rudimentary claw present; MALE:
tibial
fully
spurs 1,1,2.
winged;
length 2,75-2,87 mm;
anterior part of face testaceous; hind coxae basally segments dull,
antennae not distally
in following proportions:
fully
reticulate rugose;
carina shiny,
with
carina complete;
Fig.
black;
54 -
=
9;
Ol
mandibles,
10:
5:
10:
8:
thickened; 8:
8:
clypeus and
legs testaceous, 6:
6:
with
antenna I 5:
7;
head
posterior region of the vertex near occipital
longitudinal
POL
black;
antennae testaceous;
=
Male genitalia of
keels; 5;
OOl
frontal
=
8;
line present;
OPl
=
9;
occipital
TL = 10;
pronotum
Conganteon tOlvnesi n. sp. (paratype)
Conganteon i nae
103
hairy, very short; scutum shiny, fully reticulate rugose; notaulices invisible; scutellum and metanotum strongly punctate, without sculpture among
the punctures;
longitudinal
keels;
part of radial
vein
forming a curve;
propodeum reticulate rugose,
fore-wings hyal ine,
longer than proximal
genitalia:
fig.
54;
LOCUS TYP ICUS:
as in vulcanicum;
Pietermaritzburg
TYPICAL MATERIAL: M!
holotype F!
part
without
dark
transversal
transversal
(13:11);
or
bands;
the two parts
gonoforceps without proximal
membranous process and without distal (laterally viewed)
without
inner pointed process;
tibial
spurs
gonoforceps
1,1,2.
(South Africa) and 2 para types MM!
in TW;
paratype
in OL.
HOSTS:
unknown
DISTRIBUTION: NOTE:
Ethiopian region:
the species is named
SOUTH AFRICA:
in honor of Mr.
Pietermaritzburg,
Henry K.
TW!
OL!
Townes.
2. Conganteon vulcanicum Benoi t
Conganteon vulcanicus Benoit 1951c: 12.
FEMALE
(Figs.
16,
55):
fully
winged;
mandibles testaceous;
antennae brown,
and propodeum black;
legs testaceous,
black;
head hairy,
complete; = 8;
length 2,37 mm; with
head black,
segments 1-2 testaceous;
shiny,
fully
reticulate rugose;
occipital
fully
reticulate rugose;
POL = 2,5;
antenna I segments in following
Fig.
55 -
thorax
with coxae and hind femora partly
temples distinct,
OPL = 6;
with
Female of Conganteon
proportions:
9:
carina OL = 5; 5:
vulcanicum Benoit
10:
7:
OOL 7:
104
6:
Mem.
6:
6:
5,5:
8;
pronotal
Amer.
En!.
Ins!.,
tubercles reaching
no.
37,
tegulae;
1984
scutum shiny,
fully
reticulate rugose, hairy; notaulices invisible; scutellum shiny, with anterior half smooth and without sculpture; posterior half reticulate rugose;
metanotum reticulate rugose;
without keels;
fore wings hyal ine,
pterostigma short and oval,
brown;
enclosed by pigmented veins; than proximal fore tarsi
part
produced
propodeum fully
without dark
radial
costal vein
reticulate rugose,
transversal
bands;
and median cells clearly with
distal
(9:7);
the two parts forming
into a
hook;
fore tarsal
part slightly
an 2ngle;
segments
longer
segment 3 of
in following
proportions: 11: 2: 2: 3: 7; enlarged claw (Fig. 56) as long as segment 5; segment 5 of fore tarsi (Fig. 56) with a long row of 27 lamellae extending continuously very hairs;
long;
rudimentary
Fig.
MALE:
fully
antennae not
56 -
distally 3:
8:
apex with 5
length
tibial
thickened; 7:
one of these
lamellae,
bristles or
spurs 1,1,2.
1,81 mm; black; mandibles black,
antennae brown; 7:
lamellae,
tooth,
Conganteon vulcanicum Benoit (type)
Chela of
winged;
6:
the apex;
without subapical
claw present;
half testaceous-brown; proportions:
to
enlarged claw
7:
6:
legs black-brown,
antenna I segments 5:
6:
7;
with distal
with
tarsi
in following
head dull,
fully
reticulate
light;
Conganteon inae
=
4; Ol = 2; OOl = 5; OPl = 4; Tl scutellum and metanotum dull, fully
rugose; frontal I ine present: POL occipital carina complete; scutum, reticulate rugose; without
notaulices invisible;
transversal
surface without transversal
bands,
the
keels;
with costal
distal
an
vein
angle;
Volcano Muhavura
TYPICAL MATERIAL: HOSTS:
hyaline,
posterior
without dark
holotype F!
longer than proximal
part
gonoforceps without proximal inner pointed process;
tibial
spurs
(Ruanda)
in
TV.
unknown
DISTRIBUTION: ZAIRE: Hoyo,
7;
and median cells clearly enclosed by
membranous process and without distal 1,1,2. lOCUS TYP ICUS:
and posterior surface;
fore-wings
part of radial
two parts forming
=
propodeum reticulate rugose,
between dorsal
longitudinal
pigmented veins;
(6:5);
keel
105
Ethiopian region:
Bomboka TV!
Ol!
(Kyandolire,
MADAGASCAR:
RUANDA:
Mt.
Volcano Muhavura
Ruwenzori,
Ankaratra,
Albert Nat.
(Burambi),
Park),
TV!
TV!
Mt.
P!
2. Chelanteon n. gen.
TYPE SPEC I ES:
Chelanteon richardsi n. sp.
FEMALE
57,58):
(Figs.
fully
winged;
maxillary palpi
with 5 segments;
labial palpi with 3 segments; mandibles quadridentate, with an intermediate rudimentary tooth (Fig. 22 B); fore-wing with short and oval pterostigma
(Figs.
part
57,
(Figs.
57,
58);
bristles or hairs (Fig. MALE:
58);
distal
part of radial
enlarged claw
vein
with subapical
longer than proximal
tooth,
without
lamellae,
59); pronotal tubercles reaching tegulae.
unknown
DISTRIBUTION: HOSTS:
Oriental
unknown
1
SPECIES:
Chelanteon richardsi n. sp.
FEMALE
(Figs.
57,
58):
fully
winged;
anterior part of face (particularly
length
2,81
along orbits),
mm;
head black,
with
clypeus and mandibles
yellow; antennae brown, with segments 1-2 yellow; thorax, propodeum, petiole and abdomen black; legs yellow, with hind coxae, hind clubs of femora
and hind
and shiny,
two areas on where
brown;
the sides of the frontal
the surface
temples distinct, sculpture;
tibiae partly
tegulae yellow;
anterior part of face hairy,
POL
=
I ine
is smooth and finely reticulate rugose, 5;
Ol
=
2;
OOl
=
fully
head slightly
reticulate rugose,
swollen
except for
in front of the anterior ocellus,
punctate;
occipital
carina complete;
except for a wide smooth area without OPl;
antenna I segments
in following
106
Mem.
Fig.
proportions: with
57 -
8:
transversal
dull,
fully
finely nearly
brown; radial
14:
10:
keels;
10:
Ins!.,
no.
37,
1984
10:
7:
6:
6:
8;
pronotum rugose,
tubercles reaching
notaulices invisible;
the tegulae;
metanotum shiny,
reticulate rugose, transversal
scutum
fully
smooth,
scutellum
reticulate rugose;
hairy,
without keels;
bands;
pterostigma short and oval,
and median cells clearly enclosed by
vein curvilinear,
hairy,
scutellum shiny,
with some areolae near the posterior margin;
without dark costal
En!.
Chelanteon richardsi n. sp. (type)
pronotal
long as metanotum;
propodeum fully hyaline,
Female of
reticulate rugose;
punctate, as
4:
Amer.
with distal
part
fore-wings
pigmented veins;
longer than proximal
part;
Conganteon i nae
segmen t 3 of fore
tars i produced
following proportions:
Fig.
shorter than
lamellae,
the apex;
8:
hook;
59 -
(12:13);
segment 5 of fore tarsi
enlarged claw
Dalat
with a
rudimentary
Chela of
fore tarsa I segments
13; enlarged claw
(Fig.
Chelanteon richardsi n. sp.
present;
(Viet Nam)
in slightly
(type)
(Fig.
to the apex;
subapical
claw
59)
59)
with
a
apex with 5
tooth and without tibial
spurs 1,1,2.
Chelanteon richardsi n. sp. (type)
unknown
_OCUS TYPICUS:
into a
4:
lamellae extending continuously
bristles or hairs;
Fig.
\1ALE:
3,5:
Female of
segment 5
long row of 44 lamellae at
58 -
17:
107
108
Mem.
TYPICAL MATERIAL: HOSTS:
holotype F!
En!.
Inst.,
no.
37,
1984
in B
unknown
DISTRIBUTION: NOTE:
Amer.
Oriental
the species
region:
is named
VIET NAM:
Dalat,
B!
in honor of Mr. Owain W.
Richards.
3. Fiorianteon n. gen.
Fiorianteon junonium n. sp.
TYPE SPEC I ES: FEMALE:
unknown
Fig.
MALE labial
(Figs. palpi
26,
60 -
60):
Male of
fully
pterostigma proximal
winged;
with 3 segments;
intermediate rudimentary (Figs.
part
26,
(Figs.
tooth
60); 26,
Fiorianteon junonium n. sp. (type)
(Fig.
distal
60);
maxillary palpi
with 6 segments;
mandibles quadridentate, 22 B);
fore-wing
part of radial
occipital
with an with short and oval
vein much shorter than
carina complete.
Conganteon i nae
109
DISTRIBUTION: Palaearctic HOSTS: unknown SPECIES: NOTES:
1
the genus
is named
in honor of the
late Mr. Giorgio Fiori.
Fiorianteon junonium n.
FEMALE: MALE black;
unknown
(Figs.
c I ypeus
sp.
26,
60):
testaceous;
fully
winged;
antennae and
antennae not distally
proportions: punctate,
Fig.
11:
5:
18:
14:
thickened;
13:
11:
without sculpture among
61
-
length 3,81
Male genitalia of
mm;
legs testaceous, 9:
black;
antenna I segments 9:
8:
10;
the punctures;
mandibles and
with hind coxae basa II y in following
head shiny, frontal
Fiorianteon junonium
n.
strongly
I ine complete;
sp.
(type)
110
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
occipital carina complete; POL = 6; OL = 4; OOL = 11; OPL = 6; TL = 12; scutum shiny, very strongly punctate, without sculpture among the punctures; scutum;
notaulices incomplete,
scutellum punctate,
metanotum rugose, keel
between
longitudinal distal
du II;
dorsal keels;
genital ia:
fig.
wi thou t
61;
without dark
much shorter than
gonoforceps without
transversal
proximal
dorsal
inner pointed process;
Yamada
transversa I
posterior surface without part
and median cells clearly enclosed by
TYPICAL MATERIAL: HOSTS:
vein
length of
the punctures;
propodeum ret i cu I ate rugose,
fore-wings hyal ine,
and without distal LOCUS TYPICUS:
approximately 0,5
and posterior surface;
part of radial
wings with costal
reaching
without sculpture among
fore-
pigmented veins;
proximal
tibial
bands;
(4: 10);
membranous process
spurs 1,1,2.
(Japan)
holotype M!
in OT.
unknow n
DISTRIBUTION:
Palaearctic region:
4.
JAPAN:
Yamada
(Nagano),
OT!
ANTHONINAE R. C. L. PERKINS
SUBFAMILY
Dryininae partim Kieffer in Kieffer et Marshall 1905: 130. Anteoninae partim Kieffer in Kieffer et Marshall 1906: 510. Anteonini R. C. L. Perkins 1912: 7. Anteonini Kieffer 1914b: 137. Anteonini Richards 1939: 190. Anteonini Muesebeck and Walkley 1951: 1040. Anteoninae J. F. Perkins 1976: 13. Anteoninae partim Krombei n 1979: 1240.
TYPE GENUS: Anteon Jurine 1807. FEMALE: fully winged, brachypterous or micropterous; winged species)
with costal,
by pigmented veins palpi
(Fig.
with 3 segments;
rudimentary (Fig.
62);
claw
(Fig.
62);
anterior one to posterior MALE:
fully
winged;
labial
teeth progressing carina complete; 29);
tibial
enlarged claw
(Fig.
5);
fore-wing
with
palpi
with
pronotal
teeth
(Fig.
median 29);
labial
without
and
lamellae
larger from
tibial
occipital spurs 1,1,2.
and submedian cells
maxi Ilary palpi
with 6
mandibles quadridentate,
larger from anterior one to posterior fore-wing
(in fully
chela
tubercles present; long hairs;
with costal,
3 segments;
without
18);
teeth progressing
tufts of
pigmented veins
with 6 segments;
(Fig.
(Fig.
5);
with
occipital
with metacarpus shorter than pterostigma
spurs 1,1,2.
DISTRIBUTION: HOSTS:
maxi Ilary palpi tarsus chelate
antennae without
enclosed by
segments;
18); front
fore-wing
and submedian cells clearly enclosed
mandibles quadridentate,
carina complete; clearly
median
worldwide
Cicadellidae (i nc Iud i ng I diocerinae, Macropsinae and
(Fig.
Anteoninae
occasiona II Y Typhlocybinae) GENERA: SPEC I ES:
5 239
Fig.
62 -
Chela of Metanteon
fuscum n. sp. (type)
111
112
Mem.
Amer.
Ent.
Ins!.,
no.
37,
1984
ANTEONINAE
KEY TO THE GENERA OF
FEMALES
62) •..•..•.........••..•••.••......••
of at least 4 bristles 1. Metanteon n. gen.
Inner
1-2 bristles
Inner
2
side of
enlarged
side of
claw with
enlarged
a
claw with
row
only
(Fig.
or peg-like hairs
located proximally (Fig. 63) •.•••.•••••••••.•••••••••.••••••..••••• 2 Inner side o-f enlarged claw wi th a proximal prominence bearing 1
b r i s tie (F i g. 64) •••••••••.•••••••••••••••••••••.••••.••••.••.••••• 3 Inner side of enlarged claw wi th a proximal prominence not bearing bristles (Fig. 63); 1-2 short bristles or peg-like hairs located further distally than the proximal prominence (Fig. 63) •••••••.••.• 6 3
Species brachypterous
-
Species
fully
winged
4 Propodeum wi th posterior -
a
strong
5 Fore-wi ng proximal
transversal
keel
a
st rong
4.
transversa I
surface •••••••••••••.•.•••• wi th d i sta I part
(Fig.
part
158);
case propodeum without -
162) .••••••••••••••••••••••••••••••••.• 4 158) ••••••••••.•••..••••.••••.•••••••.•• 5
surface . • • • • • • • • . • • . • . • . • . •
Propodeum wi thou t posterior
(Fig. (Fig.
a
of
rad i a I
vei n
strong
and
Anteon Jurine
kee I 3.
occasi onally
between dorsal
between dorsa I
and
Lonchodryinus Kieffer as
long as
sl ight Iy
transversal
or
longer
shorter,
keel
but
between
in
dorsal
than that and
posterior surface ••••.••••••••••••••• 3. Lonchodryinus Kieffer Fore-wing wi th di stal part of radial vein much shorter than proximal or
longer
strong
part
(Fig.
178),
than proximal
transversal
keel
occasionally
part,
but
between
in
sl ightly
that
dorsal
shorter,
as
long as,
case propodeum with a
and posterior
surface ••.•••••
.••••.•••.••.•..•.•••...••••.•....••• 4. Anteon Jurine 6 Micropterous
species with
thoracic
structure
Gonatopodinae (pronotum remarkably elongated; between pro- and metanotum; propodec:m humped)(Figs. -
Fully
winged
species with
Anteoninae (pronotum not the pronotum;
metanotum
385,386) •••• thoracic
metanotum visible;
invisible; 5.
of stalk
+
to
that
scutum broader
propodeum not 2.
that
metathorax
similar
elongated;
.•••••••••••••••••••••••••.••••••••••
to
Prioranteon n. gen.
structure
remarkably
similar
scutum forming
humped)(Fig.
of than 67) •••••
Deinodryinus R.C.L. Perkins
MALES
Pterostigma
four
or more
times as
long as broad
• • • • • • • • • • • • • • • • • . • • . • •• • . • • • • • • • • • •• -
Pterostigma
less
2 Fore-wing with
than
four
times as
distal
part
of
radial
2.
(see fig.
176) ••..•
Deinodryinus R.C.L. Perk ins
long as broad vein much
(see
shorter
fig.
176) ••• 2
than proximal
Anteoninae
113
.,/
Fig.
part
(Figs.
longer
Chela of Deinodryinus atriventris
63 -
29,
180);
than proximal
occasionally part,
but
in
slightly that
(Cresson)
shorter,
case with a
as
long as,
or
strong
transversal
-
keel between dorsal and posterior surface . . . . . . . . . . . . .. . . . . . • . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4. Anteon Jurine Fore-wing wi th distal part of radial vein as long as or longer than proximal part (Fig. 160); occasi onally sl ight Iy shorter, but in that case propodeum without a strong transversal keel between dorsal and posterior surface . . . . . . . . . . . . . . . . . . . . 3. Lonchodryinus Kieffer
The males of Metanteon
n.
gen.
and Prioranteon
n.
gen.
are unknown.
1. Metanteon n. gen.
TYPE SPEC I ES: Metanteon FEMALE palpi
(Figs.
18,
65):
with 3 segments;
proximal
part
(Fig.
aerias (Wa I ker 1839). fully
winged;
fore-wing
65);
maxi Ilary palpi
with distal
with 6 segments;
part of radial
vein
labial
longer than
inner side of enlarged claw with a row of at
least 4
114
Mem.
Fig.
bristles (Fig. 62); MALE: unknown D I STR I BUT I ON: HOSTS:
64 -
Amer.
Ent.
Inst.,
no.
37,
1984
Chela of Anteon jurineanum Latreille
occipital
carina complete.
Neotrop ic
unknown
SPECIES:
2
NEOTROP I C SPEC I ES OF NETANTEON
Two species are known.
KEY TO THE SPECIES
FEMALES
Head and pronotum fully -
testaceous-reddish . . . . . . . . . . . . . . . . . . . . . . ..
•••••••••••••••••••••••••••••••••••••
1.
Head and pronotum partly black . . . . . . .
2.
MALES
Unknown
aerias (Walker) fuscum n. sp.
Anteoninae
115
1. Metanteon aerias (Walker) n. comb.
Dryinus aerias Walker 1839: 92 Anteon aerias (Walker): Dalla Torre 1898: 537.
FEMALE
(Figs.
18, 65): fully winged;
length 2,81-3,00 mm;
pronotum and scutum
testaceous-reddish
propectus,
metanotum,
Fig.
scutellum,
(last
mesopleura,
head,
antenna I segment metapleura,
antennae,
slightly dark);
propodeum,
petiole,
65 - Female of Metanteon aerias (Walker) from EI Coigo (Chile)
abdomen black;
legs testaceou5,
with hind coxae proximally
brown;
tegulae
116
Mem.
testaceous; 7: = 7;
8;
antenna I segments
head dull,
OPl = 5;
with a
weak
reaching
smooth,
Tl = 5;
En!.
Ins!.,
frontal
1984
9: 5: 11: 9: 9: 8: 7: 7:
line absent;
carina complete;
impression,
scutum,
37,
no.
in following proportions:
granulate;
occipital
transversal
tegulae;
punctate, reaching
Amer.
stronger on
POl= 3;
pronotum dull, the sides;
scutellum and metanotum shiny,
without sculpture among punctures; approximately 0,65
without transversa I keel
pronotal smooth,
notaulices distinct,
length of scutum;
propodeum fully
OOl
tubercles
weakly incomplete,
reticulate
rugose,
du II,
without rugose;
longitudinal keels on the posterior surface; metapleura reticulate mesopleura smooth, weakly granulated; fore-wings hyaline, with distal
part of radial parts formi ng
vein slightly a curve;
between dorsa I and
Ol = 2;
granulate,
longer than proximal
pterost i gma
10: 2: 3:
part;
long and narrow;
posterior surface and
distal
fore
and proximal
tarsa I segments
with
lamellae in addition
row
of 6 short bristles MALE: unknown LOCUS TYPICUS:
11:
in
17; segment 5 of fore tarsi (Fig. 66) some bristles and a row of nearly 26 lamellae-like bristles; apex with 3
following proportions:
to some bristles;
(Fig.
66);
inner side of enlarged claw
Chile
TYPICAL MATERIAL:
holotype F!
in BM.
HOSTS: unknown DISTRIBUTION: Neotropic region:
CHILE:
(Cord.
Nahuelbuta,
Arauco),
OTt
Chile,
EI
Coigo
fully
winged;
length 3,00-3,87 mm;
half of face yellow-testaceous; side somet imes testaceous; wide black spot on
(Curico),
OT!
OL!
n. sp.
head fully
black or with
anterior
malar space and anterior half of the ventral
dorsa l s i de of head somet i mes testaceous,
vertex;
Pichinahuel
BM!
2. Metanteon fuscum
FEMALE:
with a
spurs 1,1,2.
tibial
mandibles and clypeus
with
antennae brown, with segments 1-2 or only 1 testaceous; prothorax black, the posterior margin of the pronotum I ight or testaceous; mesothorax, metathorax, basally distally
propodeum and abdomen black;
black
occipital
thickened;
reaching
antennal
punctate and granulc:te;
POL = 4;
OL = 3;
granulate and rugose,
tegulae;
scutum,
with
black;
with
frontal
OOl = 8;
OPL = 7;
transversal
keels;
scutellum and metanotum shiny,
with
hind coxae antennae
10: 7:
segments in following proportions:
10; head dull,
carina complete;
pronotum dull,
legs testaceous,
and sometimes with clubs of femora partly
12: 11: 9: 7: 7:
a
testaceous or black;
15:
15:
line absent; TL = 7;
pronotal
tubercles
punctate and
granulate; notaulices incomplete, reaching approximately 0,65 length of scutum; propodeum dull, reticulate rugose, without transversal and longitudinal keels; fore-wings hyal ine, without dark transversal bands; distal part of radial longer than proximal part (17:10); fore tarsal segments in following
vein
15: 3: 5: 16: 24; inner part of enlarged claw (Fig. 62) with a segment 5 of fore tarsi (Fig. 62) with 2 rows of nearly 55 lamellae and a group of 5 lamellae at apex; tibial spurs 1,1,2.
proportions:
row of 6 bristles;
Anteoninae
Fig.
MALE:
66 -
Chela of Metanteon
117
aerias (Walker)(type)
unknown
LOCUS TYPICUS:
Cobquecura
TYPICAL MATERIAL: TW; 4 paratypes FF! HOSTS: unknown JISTRIBUTION:
(Chile)
holotype F!
and 3 paratypes FF!
in OT;
1 paratype F!
in OL.
Neotropic region:
CHILE:
Cobquecura
(Nuble),
OT!
Butamalal
in
118
Mem.
(Araueo),
Ol!
Piehinahuel
las Cabras
(Cord.
Amer.
Ent.
Ins!.,
(Chillan Vuleain,
Nahuelbuta,
Araueo),
aT!
no.
37,
1984
Nuble), EI
aT!
TW!
Ol!
Coigo (Curieo),
Ol!
2. Deinodryinus R.C.l. Perkins
Fig.
67 -
Female of
Deinodryinus paradoxus R. C. l . Perkins from Nogales
(Arizona)
Anteoninae
119
Deinodryinus R.C.L.Perkins 1907: 45. Trisanteon Kieffer 1913b: 300 (n. syn.); type species Trisanteon hirticornis (Kieffer 1911b), monotypic and orig. desig. Deinodryinus R.C.L. Perkins: Kieffer 1914b: 138. Trisanteon Kieffer: Kieffer 1914b: 196. Electrodryinus Ponomarenko 19 75c: 126 (n. syn.); type speci es Electrodryinus areolatus Ponomarenko 1975c, monotypic.
Fig.
68 -
Female of Deinodryinus paradoxus R.
C.
L.
Perkins from Nogales
(Arizona)
TYPE SPECIES: Deinodryinus paradoxus R.C.L. Muesebeck and Walkley 1951. FEMALE labial
(Figs. palpi
67,
68):
fully
with 3 segments;
winged;
Perkins 1907,
maxi Ilary palpi
fore-wing
longer or slightly shorter than proximal
with part
distal (Fig.
desig.
by
with 6 segments;
part of radial
vein
67)(the length of the
distal and proximal parts of the radial vein in females is not a specific characteristic: in laboratory bred females the length is variable in the same species);
inner side of enlarged claw with a proximal
bearing bristles
(Fig. 63);
further distally than complete;
the proximal
prominence
(Fig.
63);
elongated;
scutum broader than pronotum;
propodeum not humped);
to occipital
carina (pronotum
metanotum
vertex of head frequently
oblique keels connecting posterior ocelli
located
occipital
thoracic structure not simi lar to that of Gonatopodinae
not remarkab I y viSible;
prominence not
1-2 short bristles or peg-like hairs
carina.
with
two strong
120
MALE
Mem.
(Figs.
69,
labial
palpi
broad
(see fig.
slightly
70):
Amer.
fully
winged;
with 3 segments; 176);
En!.
wrapping
37,
maxillary palpi
fore-wing with distal
shorter than proximal branch
no.
HOSTS:
part
penis
(Fig.
(Fig. (Fig.
69);
79); 83);
Ethiopian,
vein
long as longer or
antenna I hairs usually much
gonoforceps usually
with
vertex of head frequently
Oriental,
to occipital
Nearctic,
an with
carina.
Neotropic.
Cicadellidae
SPEC I ES:
Fig.
Palaearctic,
with 6 segments;
part of radial
two strong oblique keels connecting posterior ocelli DISTRIBUTION:
1984
pterostigma four or more times as
longer than breadth of segments internal
Inst.,
65
69 -
Male of
Deinodryinus paradoxus R. C. L. Perkins from Nogales (Arizona)
Anteoninae
Fig.
70 -
121
Deinodryinus paradoxus R. C. L. Perkins from Nogales
Male of
(Arizona)
PALAEARCT I C SPEC I ES OF
One species
(fossil)
DEINODRYINUS
is known.
Deinodryinus areolatus (Ponomarenko) n. comb.
Electrodryinus areolatus Ponomarenko 1975c: 128
FEMALE
(Fig.
71):
fully
winged;
anterior part of face brown; spot;
mandibles testaceous,
antennae testaceous; testaceous-brown;
with
thorax, 17:
(length of head
bidentate;
30:
distally 22:
18:
less than
15:
three
from occipital
157:63;
antennal
clypeus
with
11:
brown
surface brown; black;
thickened;
antenna I hairs very
view
the apex of the mandibles): emarginated,
10:
antennae
in dorsal
head black, with a central
propodeum and abdomen
densely hairy;
breadth of segments;
testaceous,
teeth and proximal
antennae slightly
in following proportions: 10-segmented,
length 4,48 mm;
clypeus
11:
short,
times as
legs
antennal 10:
13;
segments antennae
much shorter than long
carina behind
as head the ocelli
to
with anterior margin weakly
sclerites well
separated from epistomal
122
Mem.
Fig.
suture;
Ent.
occipital
Ins!.,
no.
37,1984
Deinodryinus areolatus (Ponomarenko)(type)
Female of
teeth progressing
eyes apparently bare;
protruding; carina;
-
mandibles with 4
posterior; sulcus;
71
Amer.
carina complete;
posterior ocelli
larger from anterior one to
malar space with a temples prominent;
not easily
visible,
well-marked subocular eyes normally
not reaching
sculpture of vertex and frons not easily
visible;
the occipital
pronotum not
crossed by transversal impressions; thoracic structure simi lar to that of living Deinodryinus; pronotal tubercles reaching tegulae; posterior margin of pronotum visible;
longer than
scutum shiny,
posteriorly separated; than pronotum very
broad;
(20: 15);
dorsal
anterior margin;
finely punctate,
sculpture of pronotum not easily
with
notaul ices complete and
scutellum and metanotum
invisible;
propodeum reticulate rugose;
surface of propodeum nearly
as
scutum
longer
areolae of propodeum long
as posterior
Anteoninae
surface;
123
posterior surface not easi Iy visible;
posterior surface of
propodeum very steep; fore-wings hyaline, without dark transversal bands, with the usual venation of Anteoninae; pterostigma narrow, more than four times as
long as broad
(40:8);
radial
nearly as long as proximal part wings with cells);
the usual
three cells enclosed
hind wings hyal ine,
typical of Deinodryinus; slender stalk,
slightly
cell
(18:17);
open;
distal
radial (costal,
without dark bands; (10:6);
vein
fore-
median and sUbmedian shape of wings and
fore trochanters not slender, longer than broad
part of radial
vein not s-shaped;
legs
without proximal
segment 3 of fore
tarsi
produced into a hook; rudimentary claw absent; enlarged claw much longer than arolium (32:7), without subapical tooth and other teeth; segment 5 of fore tarsi visible;
nearly as long as enlarged claw,
segment 1 of fore
tarsi
nearly as
abdomen without a distinct petiole; labial
palpi
MALE:
unknown
with 3 segments;
LOCUS TYPICUS:
tibial
with
lamellae not easily
long as segment 4
maxi Ilary palpi
(15:17);
with 6 segments;
spurs 1,1,2.
Baltic amber
TYPICAL MATERIAL:
holotype F!
in MPI.
HOSTS: unknown DISTRIBUTION: only known from Baltic amber. NOTES:
this species seems
to have the structure of the
living Deinodryinus,
and therefore belongs to the genus Deinodryinus. There is no known species of living Deinodryinus from the Palaearctic region.
ETH lOP I AN SPEC I ES OF DEINODRYINUS
Five species are known.
KEY TO THE SPECIES
FEMALES
Posterior segment
surface
(Fig.
fore
of
pronotum with
tarsi
bearing
sharp
lateral
lamellae of
margins;
same 1. insulanus (Benoit) - Posterior surface of pronotum with rounded lateral margins .••.• 2 2 Antennal segment 3 nearly twice as long as segment 2; segment 5 of fore tarsi bearing small lamellae, some longer than others (Fig. 73) •••••••••••••••••••••••••••• 2. madecassus (Benoit) - Antennal segment 3 more than three times as long as 2 •••••••••• 3 3 Segment 5 of fore tarsi bearing lamellae of approximately same length
5 of
72) ••••••••...•••.•••..•
approximately
124
Mem.
Amer.
Ent.
Ins!.,
no.
37,
1984
length (Fig. 76); species mostly black . . . . . . . . . . . . . . . . . . . . . . ... . • • • • • • • • • . • • • • • • • • . • • • • • • • • • • • • • • •• 4. benoiti n. sp. -
Segment
5 of
others
(Fig.
fore
tarsi
75)
species mostly
bearing
smal
lamellae,
some
longer
than
reddish-testaceous • • . . . • . . • • • • •
3. madagascariensis (Ben 0 i t )
The female of
D. monticolus n. sp. is unknown.
MALES
Distal
part
of
reticulate
radial
rugose,
vein
punctures.. . . . . . . . -
Distal
part
of
ret i cu I ate
longer
strongly
than
punctate,
. ...............
radial
vein much
proximal without 2.
shorter
rugose ••••••••••••••••••••
part;
head not
sculpture among
madecassus (Benoi t) than proximal
5.
part
monticolus n.
head
sp.
The males of D. insulanus (Benoit), D. benoiti n. sp. and D. madagascariensis (Benoi t) are unknown.
1. Deinodryinus insulanus (Benoit) n. comb.
Prenanteon insulanum Benoi t 1953b: 419. FEMALE:
fully
winged;
length 5,00 mm;
triangle and petiole black,
reddish-testaceous,
abdomen partly
brown,
with ocellar
last 3 antenna I
segments dark; antennal segments in following proportions: 18: 5: 8: 8: 9: 9: 9: 11; head shiny, strongly reticulate rugose; frontal
18: 10: line
present;
=
=
3;
ocellar triangle convex,
=
OOL
shiny,
10;
OPL
strongly
=
10;
TL
=
with keels among
8;
occipital
punctate and with
impression separating an
transversal
anterior collar and a
margins of posterior surface sharp; scutum shiny, punctures;
scutum;
smooth,
notaul ices
incomplete,
scutellum shiny,
the punctures; without a
smooth,
metanotum sh i ny,
margin very convex and transversal
surface with
a central
raised
keel
distal part of radial
approximately 0,65
OL
lateral tegulae; the
length of
without sculpture among
without sculpture,
with
anterior
propodeum reticulate rugose,
and posterior surface;
reticulate rugose, with
5;
transversal
tubercles reaching
into a carina;
vein
with a
posterior disc;
weakly punctate, smooth,
between dorsal
area r,ot
keels,
POL
pronotum
without sculpture among
reaching
mesopleura and metapleura rugose, darkened;
pronotal
strongly punctate,
the ocelli;
carina complete;
smooth,
weak keels;
posterior
without keels;
fore-wings fully
longer than proximal
part
(11:7);
Anteoninae
fore tarsal
segments
claw
72)
(Fig.
prominence;
with
in following
segment 5 of fore tarsi same
segment 2 produced
Fig.
MALE:
proportions:
72 -
10:
4:
6:
1 bristle located further distally
lamellae of approximately lamellae:
125
Chela of
(Fig.
length;
into a
72)
26;
enlarged
the proximal
with 2-3 rows of nearly 27
apex with
hook;
11:
than
tibial
a
group of nearly 6
spurs 1,1,2.
Deinodryinus insulanus (Benoit)(type)
unknown
LOCUS TYP I CUS:
I vondro
TYPICAL MATERIAL: HOSTS: unknown
(Madagascar)
holotype F!
DISTRIBUTION: Ethiopian region: (Benoit 1953b).
in P;
1 para type F
MADAGASCAR:
in TV.
Ivondro,
P!
La Mandraka
2. Deinodryinus madecassus (Benoit) n. comb.
Hirtanteon madecassum Benoi t 1953b: 422. FEMALE: fully winged; length 3-5 mm; reddish-dark; mandibles testaceous, with teeth brown; antennae testaceous, with segments 7-10 brown; propodeum black; abdomen brown; an tenna I segments in follow i ng proportions: 11: 5: 11: 6: 6: 6: 6: 6: 5: 7; head shiny, strongly punctate, = 7;
especially anterior part,
OL = 5;
OOL = 9;
pronotum shiny, punctures,
smooth,
with a
OPL = 7; strongly
transversa I
without sculpture among
TL = 7;
occipital
punctate,
impress i on
punctures;
POL
carina complete;
without sculpture among
between
the anteri or coil ar and
126
Mem.
Amer.
En!.
Ins!.,
no.
37,1984
the posterior disc; lateral margins of posterior surface rounded; pronotal tubercles reaching tegulae; scutum shiny, smooth, finely punctate, without sculture among punctures; 0,65
length of scutum;
puncta te, convex
incomplete,
without scu I pture among punctures;
and raised
transversal with
notaul ices
keel
a central
reaching
scutellum and metanotum shiny,
into a carina;
between
dorsal
approximately
smooth,
finely
anterior marg i n of metanotum
propodeum reticulate rugose,
and posterior surface;
smooth area not reticulate rugose;
without a
posterior surface
mesopleura shiny,
strongly punctate, with a tuft of hairs on the anterior third; metapleura partly smooth, without sculpture; fore-wings with 2 dark transversal bands, one on the basal cells and another beneath the pterostigma; distal part of radial vein longer than proximal part (14:8); fore tarsal segments in following proportions: bristle
8:
3: 4:
located further distally
fore tarsi
(Fig.
73)
8:
than
with 2 rows of
19;
enlarged claw
the proximal lamellae
(about 8
shorter);
apex with a group of nearly 7 lamellae;
produced
into a
Fig.
MALE
hook;
73 -
(supposed;
Chela of
it
mandibles testaceous,
D. madecassus
to as fully
winged;
teeth brown;
tegulae testaceous;
and hind tibiae testaceous;
with
1
segment 5 of
longer and 21
no segment of fore
Deinodryinus madecassus
breeding): with
73)
tarsi
spurs 1,1,2.
was referred
was not obtained by testaceous;
tibial
(Fig.
prominence;
(Benoit) (type)
by
antennae brown,
abdomen brown;
Benoi t
length 2,37 mm;
1953b;
it
black;
with segments 1-2
legs brown,
antennae with hairs at most as
with
tarsi,
long as
mid
Anteoninae
127
general breadth of segments; head shiny, strongly punctate, without sculpture among punctures, hairy; POL; 8; OL ; 4; OOL ; 6; OPL ; ;
4;
occipital
carina complete;
scutum shiny,
without sculpture among punctures; approximate:y 0,50 smooth, fully
finely
punctate,
reticulate rugose,
Fig.
74 -
notaul ices
length of scutum;
smooth,
incomplete,
Male genital ia of
reaching
scutellum and metanotum shiny,
without sculpture among dull;
5;
finely punctate,
punctures,
propodeum
mesopleura and metapleura smooth,
without
Deinodryinus madecassus (Benoit) (allotype)
TL
128
Mem.
scu Ipture;
propodeum without a
posterior surface; slightly without
Amer.
Ins!.,
proximal
wrapping penis;
tibial
LOCUS TYPICUS:
Bekily
TYPICAL MATERIAL:
no.
37,
transversa I keel
fore-wings hyaline,
longer than proximal dorsal
En!.
part
with
between dorsa I and
distal
(10:9);
1984
part of radial
genitalia:
membranous process,
fig.
74;
with an apical
vein
gonoforceps branch
spurs 1,1,2. (Madagascar)
holotype F!,
allotype M!
and 2 para types FF!
in P;
2
paratypes FF! in TV. HOSTS: unknown DISTRIBUTION:
Ethiopian
region:
MADAGASCAR:
Bekily,
P!
TV!
3. Deinodryinus madagascariensis
(Benoi t)
n.
comb.
Hirtanteon madagascariensis
FEMALE:
fully
winged;
length 5,0 mm;
with segments 6-10 brown; segments shiny,
1953b: 423.
testaceous;
petiole black;
in following proportions:
with face fully
Benoi t
12:
antennae testaceous,
abdomen partly
7:
reticulate rugose;
25:
10:
8:
8:
vertex and
brown;
8:
8:
8:
antennal 11;
head
temples not reticulate
rugose, but strongly punctate; frontal I ine present; occipital carina complete; ocellar triangle convex; POL = 5; Ol = 4; OOl = 13; OPL = 12; Tl = 8;
pronotum shiny,
punctures;
smooth strongly punctate,
pronotum with
a strong
transversa I
anterior collar and the posterior disc; rounded;
pronotal
strong I y
punctate,
incomplete,
tubercles reaching
approximately 0,5
smooth,
scutum shiny,
smooth,
notau I ices
length of scutum;
without sculpture;
the
margins of posterior surface
without scu I pture among punctures;
reaching
metanotum shiny,
lateral
tegulae;
without sculpture among
impress ion between
scutellum and
anterior margin of the
metanotum very convex and raised into a carina; propodeum shiny, reticulate rugose, without a transversal keel between dorsal and posterior surface; posterior surface with a central smooth area not reticulate rugose; mesopleura not reticulate rugose, strongly punctate, with the anterior third bearing a rugose, on
strong I y
dense tuft of hairs;
punctate;
the basa I cell s and another beneath
segments
in following proportions:
75)
1 bristle
with
metapleura not reticulate
fore-w i ngs with 2 dark
transversa I bands,
the pterost i gma;
11: 4:
located further distally
7:
16:
than
30;
one
fore tarsa I
enlarged claw
the proximal
(Fig.
prominence;
segment 5 of fore tarsi (Fig. 75) with 2 rows of lamellae, many small (nearly 30), some much longer (nearly 9); apex with a group of nearly 9 lamellae
(one of these very
long);
no segment of fore tarsi
a hook; tibial spurs 1,1,2. MALE: unknown lOCUS TYPICUS:
Tananarive
TYPICAL MATERIAL:
(Madagascar)
holotype F!
in TV.
produced
into
129
Anteoninae
Fig.
HOSTS:
Chel a
75 -
Deinodryinus madagascariensis (Benoi t) (type)
of
unknown
DISTRIBUTION:
Ethiopian region:
MADAGASCAR:
Tananarive,
TV!
4. Deinodryinus benoiti n. sp.
FEMALE:
fully
the face
(mainly
testaceous,
winged;
length 6,62 mm;
along orbits),
head black,
with anterior part of
clypeus and mandibles testaceous;
with segments 9-10 brown;
margins of pronotum and apex of propodeum testaceous; abdomen segment
with segment 1 mostly
reddish-testaceous;
1 and other segments black;
testaceous,
with distal
tegulae
posterior part of
testaceous;
legs reddish-
region of fore and hind clubs of femora brown;
anterior region
among punctures; frontal
pronotum shiny,
smooth,
punctures,
lateral
with
of face strongly
occipital
I ine incomplete;
with
petiole black;
antennae not distally thickened; antenna I segments in following proportions: 18: 7: 30: 19: 16: 14: 13: 12: 12: 17; head smooth, punctate;
antennae
thorax and propodeum black,
POL
punctate,
carina complete;
=
finely
6;
OL
=
5;
punctate,
frons without
OOL
=
12;
shiny,
without sculpture OPL
lateral
=
13;
TL
keels;
=
9;
without sculpture among
margins of posterior surface rounded;
pronotal
130
Mem.
Fig.
76 -
tubercles reaching smooth,
finely
incomplete,
Amer.
Chela of
tegulae;
punctate,
reaching
reticulate rugose, region,
one on
Inst.,
no.
37,
1984
Deinodryinus benoiti n. sp. (type)
scutum,
scutellum and metanotum shiny,
without sculpture among punctures;
approximately 0,65
without
transversal
fore-wings with 3 dark
surface;
Ent.
the basal
length of scutum;
keel
between
transversal
dorsal
notaul ices
propodeum fully and posterior
bands,
on the proximal
cells and another beneath
the pterostigma;
part of radial vein longer than proximal part (18:13); fore tarsal segments in proportions: 15: 5: 10: 32: 48; enlarged claw (Fig. 76) bristle
located further distally
fore tarsi same
(Fig.
length;
MALE:
76)
Perinet
TYPICAL MATERIAL:
2 rows of about 90
a
prominence;
lamellae;
tibial
(Madagascar)
holotype F!
Ethiopian
the species
region:
is named
in BM. MADAGASCAR:
in honor of Mr.
Perinet,
P.L.G.
unknown
BM!
Benoit.
5. Deinodryinus monticolus n. sp.
FEMALE:
with
1
segment 5 of
lamellae of approximately
group of nearly 8
unknown
DISTRIBUTION: NOTE:
the proximal
unknown
LOCUS TYPICUS: HOSTS:
with
apex with
than
distal
spurs 1,1,2.
Anteoninae
MALE:
fully
winged;
antennae brown; brown;
length 2,43 mm;
legs testaceous,
antennae not distally
general
breadth of
9: 5: 6: 6: 6: 6: rugose;
frontal
OOl = 5; hairy,
segments;
line present;
without
TL = 3;
finely
with
head dull,
occipital scutum
without
posterior surface;
posterior surface with
transversal part
bands;
distal
proportions:
smooth,
POL = 7;
finely
notaulices
transversal
areas;
Ol = 3;
punctate and
incomplete,
sculpture among punctures;
strong
lateral
following
reaching
scutellum and metanotum shiny,
with
as rugose as
in
keel
between
2 longitudinal
fore-wings
part of radial
hyal ine,
vein
propodeum
dorsal
keels; without
much shorter
and
median
branch
lOCUS TYPICUS:
wrapping
Kayove
TYPICAL MATERIAL: HOSTS:
penis;
tibial
spurs
area
dark
than proximal
(3,5: 8,5); gonoforceps without proximal membranous process,
wide apical
partly
long as
granulate and reticulate
carina complete;
shiny,
reticulate rugose, nearly
a
hairs at most as
antenna I segments
length of scutum;
punctate,
mandibles testaceous;
hind coxae and hind femora
sculpture among punctures;
approximately 0,5 smooth,
black;
with
thickened,
5,5: 5: 6: 8;
OPl = 2;
131
with a
1,1,2.
(Ruanda)
holotype M!
in TV.
unknown
DISTRIBUTION:
Ethiopian
region:
RUANDA:
Kayove
OR I ENTAL SPEC I ES OF
One spec ies
(Kisenyi),
TV!
DEINODRYINUS
is known.
Deinodryinus asiaticus n. sp.
FEMALE:
fully
winged;
antennae testace0us, testaceous; with
lateral
keels;
the sides of
and orbits;
the
occipital
temples
surrounded by invisible;
around orbits; transversal punctate, shorter
keels;
without
than
temples
furrow
triangle convex;
wide furrow
POL = 4;
sculpture among
the scutum
(15:22);
this keel
OOl
= 11;
pronotal
with
sculpture;
OPl
the
short the
behind posterior ocelli, carina; with
occipital short keels;
a
strong
crossed by
hairy on
punctures;
without
crossed by
side of head
smooth,
tarsi
keels and between
to occipital
transversely
Ol = 3;
shiny,
frons
longitudinal
and ventral
with
in following
starts at anterior ocellus;
carina complete;
between eyes and
pronotum
legs brown,
head shiny,
triangle;
2 oblique keels connect posterior ocelli carina
mandibles testaceous;
antenna I segments
median keel
at sides of ocellar
keels on
keels
thickened;
6: 4: 4: 6: 6: 6: 5: 9;
7:
longitudinal
keels start
transversal lateral
13:
3
black;
segments 7-10 brown;
antennae distally
proportions: frons
length 3,75 mm;
with
=
6;
Tl
the sides,
keel
short
=
4;
ocellar
very
finely
posterior surface of pronotum
tubercles
reaching
the
tegulae;
132
Mem.
scutum
shiny,
notaulices
smooth,
incomplete,
scutellum very punctures;
Amer.
dark
humped,
fore
bands,
distal
humped, with
one on
segments tarsi
length of scutum;
without sculpture among
without sculpture, longitudinal
keels;
shiny;
smooth,
without
fore-wings with 2
cells and another beneath
vein much
shorter than proximal
in following proportions:
produced
smooth,
posterior surface shiny,
the basal
part of radial
tarsal
segment 3 of fore
37,1984
without sculpture among punctures;
punctate,
posterior surface without
transversal
no.
approximately 0,25
strongly
metanotum very
pterostigma; (3:8);
Inst.,
finely punctate, reaching
propodeum reticulate rugose, sculpture;
Ent.
into a hook;
7:
1:
6:
enlarged claw
12: (Fig.
the
part 25; 77)
with
1 bristle located further distally than the proximal prominence; segment 5 of fore tarsi with a few rows of nearly 39 lamellae; apex with a group of 7
lamellae;
tibial
Fig.
MALE: unknown LOCUS TYP I CUS:
77 -
spurs 1,1,2.
Chela of
Deinodryinus asiaticus n. sp. (type)
Phou Kou Khouei
TYPICAL MATERIAL:
holotype F!
HOSTS: unknown DISTRIBUTION: Oriental
region:
(Laos) in B.
LAOS:
Phou
Kou Khouei
(Vientiane),
B!
Anteoninae
133
NEARCT I C SPEC I ES OF DEINODRYINUS
Three species are known.
KEY TO THE SPECIES
FEMALES
Propodeum strongly
reticulate rugose, mainly on dorsal surface .. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1. atriventris (Cresson) - Propodeum not reticulate rugose; dorsal surface mainly smooth wit h on I y weak irregular keels ....•••••••••••••••••••••.••••••• 2 2 Face with a few irregular keels near the clypeus . . . . . . . . . . . . . . .. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2. quercicolus R.C.L. Perkins - Face fully sculptured by irregular keels •...••••••••••••..••.••• . . . . .. . . . . . . . . . . . . .. . . . . . . . . . . . . . . . 3. paradoxus R.C.L. Perk i ns
MALES
Gonoforceps without head
strongly
long apical
reticulate
branch
rugose ..••
wrapping
penis
(Fig.
86);
3. paradoxus R.C.L. Perkins
long apical branch wrapping penis (Figs. 81, 84) . . • • • . • • . . • • . . • • . . . • . . . . . . . • . • . . . • • . . . . • • . . • • • . . . . • . • 2 2 Head strongly reticulate rugose (Fig. 78) . . • . . . . . . . • . . • • . . . . • . • . • • . . . . . • • • • • . . • . . • • • . • . . . • . . • • • . . . • 1. atriventris (Cresson) -
Gonoforceps wi th a
82,
-
83,
Head punctate,
without
sculpture among punctures,
rugose . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
2.
not
reticulate
quercicolus R.C.L. Perkins
1. Deinodryinus atriventris (Cresson)
Dryinus atriventris Cresson 1872: 193. Chelogynus henshawi Ashmead 1893: 93 (n. syn.); locus typicus Mil ton (Massachusetts,
U.S.A).
Anteon rugosus Ashmead 1893: 96 (n. syn.); locus typicus III i noi s (U.S.A) .
Bocchus atriceps Brues 1904: 118 (n. syn.); locus typicus Moshol u (New York,
U.S.A).
Chelogynus ferrugineus Brues 1905: 183 (n. syn.); locus typicus Texas (U.S.A) •
-
134
Mem. Amer.
Fig.
78 -
Male of
En!.
Ins!. ,
37,
1984
Deinodryinus atriventris (Cresson): sculpture of the head
•
no .
Anteon i nae
Fig. 79 - Male of
•
,
135
Deinodryinus atriventris (Cresson): antennal hairs
136
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
Chelogynus atriceps (Brues): Brues 1905: 183. Chelogynus grandis Brues 1905: 184 (n. syn.); locus typicus Riverside (Massachusetts,
U.S.A).
Anteon atriventris (Cresson): Kieffer 1907b: 25. Anteon ferrugineus (Brues): Kieffer 1907b: 26. Anteon grandis (Brues): Kieffer 1907b: 26. Anteon henshawi (Ashmead): Kieffer 1907b: 26. Chelogynus schaefferi Brues 1907a: 101 (n. syn.); locus typicus Huachuca Mts
(Arizona,
U.S.A).
Deinodryinus variabilis Fenton 1927: 10 (n. syn.); locus typicus Falls Church
(Virginia,
U.S.A).
Deinodryinus variabilis Fenton var. carinatus Fenton 1927: 11 syn.); locus typicus Flatbush (New York, U.S.A). Deinodryinus pilosus Fenton 1927: 12 (n. syn.); locus typicus Chiricahua Mts
(Arizona,
(n.
U.S.A).
Deinodryinus atriceps: (Brues): Muesebeck and Walkley 1951: 1041. Deinodryinus atriventris (Cresson): Muesebeck and Walkley 1951: 1041. Deinodryinus ferrugineus (Brues): Muesebeck and Walkley 1951: 1041. Deinodryinus grandis (Brues): Muesebeck and Walkley 1951: 1041. Deinodryinus henshawi (Ashmead): Muesebeck and Walkley 1951: 1041. Deinodryinus pilosus Fenton: Muesebeck and Walkley 1951: 1041. Deinodryinus schaefferi (Brues): Muesebeck and Walkley 1951: 1041. Deinodryinus variabilis Fenton: Muesebeck and Walkley 1951: 1041. Trisanteon rugosus (Ashmead): Muesebeck and Walkley 1951: 1042. Deinodryinus atriceps (Brues): Krombein 1979: 1242. Deinodryinus atriventris (Cresson): Krombein 1979: 1242. Deinodryinus ferrugineus (Brues): Krombei n 1979: 1242. Deinodryinus grandis (Brues): Krombein 1979: 1242. Deinodryinus henshawi (Ashmead): Krombein 1979: 1242. Deinodryinus pilosus Fenton: Krombein 1979 : 1242. Deinodryinus schaefferi (Brues): Krombei n 1979: 1242. Deinodryinus variabilis Fenton: Krombein 1979: 1242. Trisanteon rugosus (Ashmead): Krombei n 1979: 1242. r
FEMALE:
fully
the varieties); fully 15: 8:
winged;
length 5-7 mm;
body fully
iestaceous or blackened; 17:
13: 9:
10: 9:
colour very
variable
testaceous or almost fully
9:
7:
antennal 9;
black;
(see key
to
legs almost
segments in following proportions:
head shiny,
with sculpture very
variable; head fully reticulate rugose or with numerous irregular keels, sometimes with areolae and keels; frons with 3 median and parallel longitudinal keels or without visible keels; the old specific taxonomy based on the presence or absence of these keels is not recommended, because of the wide variability of sculpture in
the same species;
always with a keel
carina complete;
shiny,
hairy,
with
strongly punctate;
around a
strong
pronotal
the orbits;
occipital
transversal
anterior
tubercles reaching
impression;
tegulae;
frons pronotum
pronotum
scutum shiny,
Anteon i nae
Fig.
80 -
Male of Deinodryinus atriventris
137
(C re sson):
notaulices
Mem. Amer . En!.
138
Ins!. ,
no. 37,
1984
:. _>C."
":-, . ',¥ :
Fig .
81
-
Male of
Deinodryinus atriventris (Cresson): genital ia (A: apical branch)
Anteon i nae
139
-••
A
!
•
I ,
Fig.
82 -
Male of
Deinodryinus atriventris {Cresson}: genitalia {A: apical branch}
140
Mem.
smooth,
finely
usually
almost complete,
punctate,
only 0,65-0,75
Amer.
En!.
Inst.,
no.
37,
1984
without sculpture among punctures; reaching posterior margin;
length of scutum;
scutellum and metanotum shiny,
without
sculpture or finely
rugose;
areolae always present on dorsal
surface;
usually
with
without areolae;
2 dark
an excavated median
pterostigma; (10:14);
distal
one on
sometimes slightly
hook;
distally
3:
enlarged claw
than
7:
tibial
spurs 1,
usually
(13:14); 28;
63)
1,
posterior surface fore-wings
with
shorter than proximal
fore tarsal
segments
segment 3 of fore tarsi
with
prominence;
with 2-3 rows of nearly 30-51 lamellae;
vein
12:
(Fig.
the proximal
smooth,
less reticulate
the basa I cell s and another beneath
shorter
11:
propodeum more or
region
part of radial
following proportions: into a
punctate;
transversa I ban ds,
notaul ices
sometimes reaching
1 bristle
apex
with
in produced
located further
segment 5 of fore tarsi
lamellae;
the part
(Fig.
63)
a group of nearly 9-16
2.
MALE: fully winged; length 3,0-4,5 mm; black; antennae fully testaceous or brown, with segment 1 testaceous; mandibles testaceous, with teeth brown; fully
clypeus black or testaceous;
testaceous,
and clubs of femora black, shiny,
hairy,
strongly
antennae not distally proportions:
15:
7:
in addition
to occipital connecting invisible; smooth, (Fig.
thickened;
15:
12:
carina
carina;
with frons
the ocell i
12:
occipital
scutum)
11:
(Fig.
79);
11:
areolae
tibiae;
in following
14;
antennal
hairs much
region of head between ocell i
with 2 strong
longitudinal
sometimes
temples prominent;
and smooth,
keels
scutum shiny,
punctures;
notaul ices
the posterior margin of
sometimes reaching only 0,65
scutellum and metanotum shiny
parallel
these keels are almost
without sculpture among
incomplete (reaching approximately
length of scutum;
without sculpture or finely
punctate; propodeum strongly reticulate rugose, without a transversal between dorsal and posterior surface; posterior surface without longitudinal
keels;
fore-wings hyaline,
without dark
transversal
(Fig.
LOCUS TYPICUS:
Texas
holotype F! holotype holotype para type
wrapping penis;
tibial
(U.S.A)
holotype F of
of
WA. HOSTS:
of
spurs 1,1,2.
D. atriventris not found (not in PH); Ch. hensha"ri in WA; holotype M! of A. rugosus in WA; F! of B. atriceps in CM; holotype F! of Ch. ferrugineus in PH; F! of Ch. grandis in PH; holotype F! of Ch. schaefferi in WA; F! and 2 para types FF of D. variabilis in WA; hoi otype F! and F! of D. variabilis carinatus in WA; hoi otype F! of D. pilosus in
TYPICAL MATERIAL:
holotype
83),
keel
bands;
distal part of radial vein sl ightly or much shorter than proximal part (9: 10 or 10: 14); gonoforceps with a long apical branch (Figs. 81, 82), variable shape
head
very deep;
2 strong obi ique keels connecting posterior ocell i
carina complete;
or complete;
legs with coxae
mid and hind
(Fig.78); 11:
legs sometimes
sometimes
antenna I segments 12:
to the clypeus;
strongly punctate,
80)
testaceous;
to black
reticulate rugose
longer than breadth of segments and occipital
tegulae
with hind cOXae basally black;
unknown
DISTRIBUTION:
Nearctic region:
CANADA:
Bothwell
(Ont.),
OT!
Dresden
Anteoninae
141
\
'
,, , '\ ,
~j!) Fig.
83 -
Male genitalia of
shape of the apical Arbor,
branch
Michigan
(B),
Deinodryinus atriventris (Cresson): variable (specimens from Cochise Co.,
Uvalde Co.,
Texas
(C),
Arizona
Salmon,
Texas
(A), (D).
Ann
142
Mem.
Amer.
(Ont.),
aT!
(Ont.),
aT!
Hamilton
(Ont.),
aT!
Innisville (Ont.),
Cumberland
(Ont.),
Stittsville (Ont.), Que.), OL!
aT!
aT!
Hull
Lytle Creek
Co.,
Calif.),
TW!
IT!
SCi
(S.
Salmon
(Anderson Co.,
Bentsen
Bdo.
Huachuka Mts
Chiricahua Mts
(Ont.),
OL!
(Uvalde Co.,
Tex.),
N.P.
(Tex.),
Co.,
Tex.),
Whitewood
aT!
(5.
TE!
Alfalfa Co. Wisconsin BM!
PH!
WA!
Genesee Co. aT!
Mo.),
(Ont.),
CAl
S.
Portal
(Old Chelsea, (N.
(Tex.),
WA!
Tex.),
Tex.),
TE!
TW!
WA!
Tex.),
TE! WA!
TE!
Speir Rch. WA!
Big Bend
Elkhart
(Brazos Co.,
(Kans.),
Ariz.), TW!
(Tex.),
(Tex.),
aT!
Diego
(Ariz.),
Presidio
Hidalgo Co.
B.),
Marcos (5.
(Cochise Co.,
Hidalgo Co.,
aT!
aT!
(Anderson
Tex.),
TE!
Marysville (Kans.),
Big Fork
(Minn.),
TW!
Milwaukee Co.
(Mich.),
TW!
BM!
OL!
Lake Odessa
County 4 (Iowa),
WAr
Illinois,
(Mich.),
WA!
(Boone Co.,
Golden Pond P.
TW!
(Lexington,
WAr
Pine Heath
Tenn.),
aT!
(Gatlinburg,
Rapides Parish Ol! TW!
Mathews Co.
Bethesda
(Rabun Co.,
C.),
TW!
lost River State Park
(Va.),
WA!
Glencarlun
Falls Church Hummelstown
Pine Mountain
WA!
Montevallo
aT!
(N.
aT!
(Ky.),
Mo.),
(Wis.),
Cleveland (5. C.), TW! Ol! Wattacoo (Pickens Co., S. C.), TW! Mc Clellanville (S. C.), TW! Greenville (S. C.), TW! Wake Co. (N. C.), Highlands
(Ga.),
Ancaster
Ann Arbor
Natchez Trace S. Forsyth
WAr
(Erath Co.,
Columbia
TE!
One Sided Lake
Kouchibouguac N.P.
College Station
WA!
aT!
Foxboro (Ont.),
Stephenville (Erath Co.,
WAr
Lawrence
(Okla.),
(Mo.),
(Vernon Co., TW!
Bluffdale
(Brues 1905);
Williamsville WA!
Tex.),
aT!
Parker Canyon Lk.
Brownsville
Texas,
Oak.),
TW!
Tenn.),
TW!
TE!
ML!
(Mich.),
SCI
(Brownsville,
(Ont.),
Calif.),
(Rio Grande Valley State Park,
Esprza Rch.
1984
Summit of King Mt.
(Ariz.), WA!
37,
(On!.), CAl
OL!
Co.,
(Ariz.), Tex.),
aT!
aT!
no.
Park
Ottawa
(Que.),
(Que.),
Inst.,
Rondeau
aT!
Wainfleet
Hemmingford
U.S.A.:
(la.),
TW!
Ent.
(Va.), (Pa.),
WAr WAr
Pk.
Md.),
(N.
J.),
(Va.),
(Pa.),
laurel
(Montg'y Co.,
High Point 51.
Dupont
TW!
CAl
WAr
(Md.), WAr
(W.
Va.)(Krombein
Ol!
(Pa.),
Plummers
Takoma Pk.
Moorestown
1979);
Martinsville (Va.),
Spring Br.
aT!
Ga.),
(Md.),
(N.
J.),
lsi.
TW! TW!
aT!
TW! (Md.),
WAr
New Jersey, WAr
Princeton
WA! (N.
J.), 'IVA! (N. Y.),
Brown's Mills Jc (N. J.), WAr Westerly (R. I.), TW! Poughkeepsie TW! Farmingdale (N. Y.), TW! Millwood (N. Y.), TW! Armonk (N.
Y.),
Milford Center
AM!
TW!
Flatbush
(N.
Y.),
(N.
AM!
Y.),
TW!
Ithaca
South Haven
(N.
Y.),
AM!
Station (Huntington, N. Y.), AM! Huntington Kerhonkson (Cherry town, Ulster Co., N. Y.), N.
Y.),
AM!
Lewisboro
(Westchester Co.,
N.
(long
Island,
Y.),
AM!
Newburgh
N.
Y.),
lO!
Six-Mile (Ithaca,
N.
(Tompk Co.,
N.
Y.),
ZO!
lake Ridge Road
(Tompkins Co.,
(N.
West Point Riverside
Y.),
(N.
WA!
Y.),
(Mass.),
Mosholu
WAr PH!
(N.
Y.),
CM!
Wash'n
(N.
H.),
Y.),
lO!
Me.
Mt Whiteface
Pigeon Cove (Mass.), Mt.
Y.),
(N. Y.), AM! 4 mi. NNW AM! Brainard (Rensselaer Co.,
(Fostertown, Flatbush
N.
Kalbfleisch Field Res.
CM!
AM!
Milton
MEXICO:
lean Bogs
N. (N.
Y.),
aT!
Y.),
WAr
(Mass.),
'IVA!
Iturbide (Nuevo
Leon), TE! Colima (Manzanillo), SC! EI Palmito (Sinaloa), aT! Valle Nacional (Oaxaca), TW! Miltepec (Oaxaca), TE! Tehuacan (Puebla), OL! NOTES:
the colour of the female of D.
variable. Though
atriventris (Cresson) is very
Some varieties were described
there are intermediate colours
in
the past as good species.
I am proposing
the following key
to
Anteoninae
the main colour varieties
Body
fully
(without
testaceous with
143
taxonomic
value):
petiole black . . . . . . . . . . . . . . . . . . ..
· . . • . . . . . • . . . . . . . . . . . . . . . . ferrugineus (B ru es) - Body at least partly dark or black . . • . . . . . • . . . . . . . . . . . . . . . . ... 2 2 Thorax and propodeum fully or almost fully black . . . . . . . . . . . . . . 3 -
Thorax and propodeum at
3 Legs
fully
testaceous,
least at
partly
most
wi th
testaceous hind
or
reddish . . . . . 5
coxae basally
black . . . .
· . . . . • . . . . . . . . . . . . . . . . . . . . henshawi (Ashmead) (= variabilis Fen t on) - Legs most Iy black • • • • . • • • . • • • • . • • • • • • • • • • • • • • • . • • . • • • . • • • . . • • . . 4 4 Frons with three distinct longitudinal and parallel median keels
. grandis (Brues) -
Frons without
three distinct
longitudinal
and parallel
median
keel s •••..••••••••••..••• . pilosus Fenton 5 Head,
thorax and propodeum testaceous;
on propodeum;
abdomen
occasionally brown brown . . . . . . . . . . . . . . . . . .
spots
· . . • . . . . . . . . . . . • • . . . . . . . . . atriventris (C res son) -
Thorax and propodeum part Iy
6 Prothorax
testaceous;
testaceous . . . . . . . . . . . . . . . . . . . . . . . . . 6
mesothorax,
metathorax and propodeum black
. . • . . . . . . . . . . . . • . . • . . . . . . . schaefferi (Brues) -
Thorax and propodeum mostly
testaceous,
spots . . . . . . . . . . . . . . . . . . . . • atriceps
with
(Brues)
irregular dark
(= carinatus Fenton)
The presence or absence of keel s on the frons (see grandis and pilosus) has no taxonomic value: this characteristic is very variable,
with
intermediate cases.
2. Deinodryinus quercicolus R.C.L. Perkins
Deinodryinus quercicola R.C.L. Perkins 1907: 47. Deinodryinus quercicola R.C.L. Perk i ns: Muesebeck and Walkley 1951 : 1041.
Deinodryinus quercicola R.C.L. Perkins: Krombein 1979: 1242.
FEMALE:
fully
winged;
length 4,57 mm;
near the tegulae and with segments 4-10 dark; face with
a few
complete;
OPL
segment 3 internal keels;
body
petiole black; shiny,
hairy;
longer than OL;
head shiny,
antennae thin,
organs visible;
(10:6);
body
impress ion,
tegulae;
weakly
occipital
distally
granulate; carina
thickened,
the head,
with
as
long
with
but without
pronotum crossed by
approx ima te I y
spot
with
almost transparent,
pronotum as sculptured as
tubercles reaching
anterior transversa I
with a black
antennae testaceous,
irregular keels near the clypeus;
longer than segment 4
pronotal
yellow-reddish,
as broad;
an scutum
144
Mem.
as sculptured as
the pronotum;
approximately 0,65 propodeum with with 2 dark beneath distally
weak
than
1,1,2. MALE:
fully
8:
black;
distally
7:
7:
7:
occipital
shorter than 4 1 bristle
mandibles,
with coxae, thickened;
7:
6:
without sculpture among punctures;
vertex;
carina complete;
fore-wings
cells and another
6:
8;
(22:36);
located further
segment 5 of fore tarsi
legs testaceous, 5:
reaching
group of about 15 lamellae;
antennae not 8:
19B4
not reticulate rugose; the basal
1 of fore tarsi
length 2,37 mm;
following proportions: than
a
37,
incomplete,
tooth and with
prominence;
apex with
testaceous;
tibiae brown;
punctate,
keels,
segment
no.
scutellum and metanotum smooth;
one on
without sUbapical
winged;
antennae fully hind
bands,
the proximal
Ins!.,
notaul ices
transversal
the pterostigma;
of 13+26 lamellae;
En!.
length of scutum;
transversal
enlarged claw
Amer.
with 2 rows
tibial
spurs
clypeus and clubs of femora
antenna I segments head shiny,
and in
smooth,
frons more strongly punctate
POL = 5;
OL = 3;
OOL =
7;
OPL
5; TL = 6; scutum, scutellum and metanotum shiny, smooth, very finely puncta te, without scu I p ture among the punctures; notau Ii ces strong,
/
Fig.
BL, - Male genitalia of Deinodryinus quercicolus R. C. L. Perkins from Nogales,
Arizona
(right half removed).
Anteoninae
incomplete,
reaching
reticulate rugose,
approximately 0,65
with
a
keel
between
posterior surface without
longitudinal
without
bands;
dark
as proximal ( Fig.
84);
transversal part
(7:7);
length of scutum; dorsal
keels,
distal
rugose;
fore-wings hyal ine,
part of radial
gonoforceps with
propodeum
and posterior surface;
an apical
vein nearly
branch
as
long
wrapping penis
t i b i a I s p u rs 1, 1 , 2 •
LOCUS TYPICUS:
Nogales
TYPICAL MATERIAL: HOSTS:
145
(Arizona,
holotype F!
U.S.A)
in B.
unknown
DISTRIBUTION:
Nearctic region:
U.S.A.:
Nogales
(Ariz.),
B!
3. Deinodryinus paradoxus R.C.L. Perkins
Deinodryinus paradoxus R.C.L. Perkins 1907: 46.
FEMALE: small
fully
on segment 10 dark; 6:
6:
5:
keels; 12;
winged;
black spot
finely
1 of the abdomen;
antenna I segments 7;
head shiny,
face fully ~
OPL
length 4,02-5,30 mm;
near tegulae and a
8;
TL
punctate,
antennae yellow-testaceous
weakly
alutaceous,
10;
hairy,
tubercles reaching
the
with a
weak
without sculpture among punctures; reaching 0,5- 0,65 finely
punctate,
shiny,
almost
unsculptured,
13:
3;
8:
7:
7:
irregular weak ~
OL
impression;
smooth,
2;
~
OOL
smo')th,
pronotal
finely
punctate,
incomplete,
scutellum and metanotum smooth,
without sculpture among with
6:
pronotum shiny,
notaulices distinct,
length of scutum;
shiny,
transversal
scutum shiny,
~
POL
carina complete;
tegulae;
with segments 9-
11:
'with numerous
irregular keels;
occipital
with a
the petiole and partly
in following proportions:
sculptured by ~
yellow-testaceous,
black spot on
weak
the punctures;
transversal
keels,
propodeum
without a strong
transversa I keel between dorsa I and posterior surface; posterior surface without longitudinal keels; fore·-wings with 2 dark transversal bands, one on
the basal
radial
vein
proximal narrow;
cells and another beneath longer or sl ightly
parts forming fore tarsal
(Fig.
85)
radial
proximal
cell
open;
into a hook;
lamellae;
apex with
with a medial
distal
part;
part of
distal
pterostigma
in following proportions:
produced
with 2 rows of 30-39
enlarged claw
curve;
segments
segment 3 of fore tarsi 85)
a
the pterostigma;
shorter than
9:
3:
and
long and
7:
14:
30;
segment 5 of fore tarsi
(Fig.
a group of 10-15 lamellae;
peg-like bristle;
tibial
spurs 1,1,2.
MALE: fully winged; length 2,56-2,81 mm; black; antennae yellow; mandibles testaceous with teeth reddish; clypeus and tegulae testaceous; legs testaceous with coxae, antennae very segment 3; 10: keel
10:
10:
hairy;
clubs of femora
hairs very
antenna I segments 9:
11;
head shiny,
around the orbits;
long,
and
nearly
tibiae partly
twice as
in following proportions: strongly
long 14:
reticulate rugose,
frons with 2 parallel
longitudinal
6:
brown; as breadth of 14:
with
9:
10:
a strong
keels and
146
Mem.
Fig.
85 -
Chela of
Amer.
En!.
Inst.,
no.
37,
1984
Deinodryinus paradoxus R. C. l . Perkins from Nogales (Arizona)
frontal
I ine;
connecting
vertex behind posterior ocell i with 2 short obi ique keels
the posterior ocelli
to the occipital
there are 2 areolae and the occipital
carina
carina;
between
is excavated;
these keels
POL
=
7;
=
Ol
4; OOl = 11; OPl = 7; Tl = 11; scu tum, scu tell um and metanotum smooth, shiny, finely punctate, without sculpture among punctures; notaul ices distinct,
incomplete,
reaching
propodeum reticulate rugose, posterior surface, surface; of radial and
with 2
incomplete
fore-wings hyaline, vein slightly
narrow;
approximately 0,75 without a
longitudinal
without dark
(Fig.
86)
without
keel
part
long apical
and 2 paralectotypes
(designated by Olmi
1 paralectotype F!
in B;
dorsal
and
posterior
bands;
(8:10);
penis; tibial spurs 1,1,2. lOCUS TYPICUS: Nogales (U.S.A) TYPICAL MATERIAL: lectotype F! 1982)
between
keels on
transversal
shorter than proximal
gonoforceps
length of scutum;
transversal
distal
part
pterostigma
long
branch wrapping
(IF,
in BM.
1M)!
Anteoninae
147
B Fig.
86 -
Nogales,
Male genitalia of Deinodryinus paradoxus R. Arizona
(A);
left gonoforceps
(B)
C.
L.
Perkins from
of a specimen from EI
Naranjo
(Mexico)
HOSTS:
in U.S.A.
Cicadell ids
(R.C.L.
Perkins 1907).
DISTRIBUTION: Nearctic region: U.S.A.: Nogales (Ariz.), B! BM! OL! Portal (Ariz.), TW! OL! Fripp Island (Beaufort Co., S.C.), WA! MEXICO: EI I~aranjo
(San Luis Potosi),
TE!
San Pedro
Iturbide
(Linares,
TW!
NEOTROP I C SPEC I ES OF DEINODRYINUS
Fifty five species are known.
KEY TO THE SPECIES
Nuevo Leon),
148
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
FEMALES
Head
fully
rugose,
smooth,
wi thout
without
areolae
sculpture,
or keel s,
not
except
granulated, for
the
not
frontal
line
• ••••••••••••••••••••••••••••••••••••••••••••••••••••••••••••• 2
Head more
2 3
or
less
rugose,
0,5
reaching
approximately posterior margin of scutum .. 4 transversal keel between dorsal and
Species
a
st rong
1. elegans n.
transversa I
testaceous,
at
most
Head,
least
pronotum and
scutum fully
pronotum and
scutum fully
or almost
dark
fully
..5
nigrorufus n.
4.
or almost
fully
bands,
sp.
black . . .
iphias n.
5.
transversal
reddi sh-
sp.
fully
hyal ine
or
1-2 distinct
dark
transversal
bands . . . . . . . . . . 9
two
strong
obi ique keels
connecting posterior
ocell i
to occipi tal carina • • • . • . • • . . . • • • • • • • • • • • . . • • • • • • • • • . • • . • • • . . • 8 Frons with two median longitudinal keels; notaulices short, reaching
less
0,5
than
length
of
scutum . . . . . . . . . . . . . ..
.................................. Frons with almost
three median
reaching posterior margin
of
Antennal
segment
3
sl ight Iy
shorter
bicolor (Olmi
6.
longitudinal
· . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. 9
sp.
Head without two strong oblique keels connecting posterior ocell i to occipi tal carina . . . . . . . . 49. minor n. sp. Head with
8
and
darkened . • • • • • • • • • • • • • • • • • . . • • • • • • • • • • • • . • . • • • • • • • • • • • • 7
Fore-wings with
7
diaphanus n.
3.
· . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. Fore-wings wi thout fully
dorsa I
sp.
with propodeum b l a c k . . . . . . . . . . . . . . . . .
testaceous . . . . . . . . . . . . . . . . . . . . . . . . Head,
sp.
between
with petiole black.
· . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. Species at
keel
speciosus n.
2.
surface . . . . . . . . . . . . . . . . .
fully
of
strong
surface ••.••••••••••••••
posterior
6
smooth.6
Notaulices
Propodeum wi thout
5
never
approximately
Propodeum with a
length
scutum . . . . . . . . 3
reaching
posterior
4
with areolae or keels,
Notaulices
keels;
surface of
smooth median
7.
finely
maximus n.
than
propodeum almost
region,
longer,
scutum . . . . . .
••• • • ••• . •••••. . •. . •••••• •••• ••• •. 8. Antennal segment 3 longer than segment 10 Posterior
and Currado)
notaulices
punctate,
segment
sp. 1 •••••••••••••
melanocephalus (Cameron) 1 • . • . . • • • • • • . • • • . . • • • • 10 fully
smooth
without
or wi th
keels
a
or
areolae ••••••••••••••••••••••••••...••••••••••••.•••••••..••• 11 Posterior keels 11
surface
of
Vertex of head wi th po s t e rio roc e I l i t 0 Vertex
propodeum rugose,
dul I,
sculptured by
or areolae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ..
of
.. 27
two strong obi ique keel s connect ing ') C c i pit a I ca r ina •..•••••••••••••••..••••. 1 2
head without
two obi ique keels
connecting posterior
ocelli to occipital carina . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 20 12 Posterior surface of pronotum rugose, wi th numerous irregular transversal
keels ••••••••••••••••••••••••••••••••••••..•••••• 13
Anteoninae
Posterior
surface
sculpture,
or
of
finely
149
pronotum smooth,
without
keels,
without
punctate . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 14
13 Thorax and propodeum black . . • • • . . • Thorax and propodeum fully
9.
or almost
noyesi n. fully
sp.
reddish-testaceous ••
• ........••....•......••.••••... " 10. atlanticus n. sp. 14 Posterior surface of propodeum wi th two strong longitudinal keels and a shiny
strong
transversal
keel
median area • • • . • • . • • • • . . • • • .
Posterior smooth
surface
area
of
surrounding
propodeum without
surrounded by
a
smooth and
11. achterbergi n.
sp. keels;
longitudinal
irregular keels •••••••• 15
areolae and
15 Head mainly b l a c k . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Head
fully
or mainly
16 Thorax and propodeum fully black .• Thorax and propodeum mainly
17 Pronotum with
two
strong
18 Vertex of head reticulate Vertex
of
punctate,
two
transversal
strong
(behind and
not
(behind and reticulate
dorsal
• •• 17
lobes ••••••••
fluviatilis n.
53.
on
the
sides
on
the
of
the ocell i)
fully
atlanticus n. sp.
10. sides
rugose .••
sp.
lobes . • • . . • • . . • 18
transverse dorsal
rugose •.••••.•••••••••• head
peruvianus n. sp.
12.
testaceous...... .••••..•
· ............................... " Pronotum without
. ••• 16 • ••• 19
reddish-testaceous. ••••••..••••
of
the
ocell i)
13. hymenaeus n.
sp.
19 Pronotum, scutum, scutellum and metanotum fully black ••. • •••.••••••••..•••..•••.•••.•••• " 14. colombian us n. sp. Pronotum,
scutum,
scutellum and metanotum mainly
testaceous ••••••••••••••••••••••••
20 Region
of
head behind
ocell i
15.
smooth,
punctate,
of
head behind
longitudinal
ocell i
rugose,
wi thout
sculptured by
numerous
..25 ..22 • .24
keels..............
21 Prothorax fully or mainly black. Prothorax
reddishsp.
keels ••••••••••.••••••. 21
sculpture among punctures and wi thout Region
cascus n.
fully
testaceous
22 Frons wi thout
areolae, pronotum with two very
pronotum black,
with
•••••••
.••••••••.•••.••••••
wi th only a few I ongi tudinal prominent dorsal transversal
dorsal
lobes and pronotal
wh i t i sh ••••••••••••••••.•••••••• "
16.
keel s; lobes;
tubercles
albopictus n.
sp.
Frons reticulate rugose, sometimes with longitudinal keels; pronotum without dorsal lobes or with only slightly prominent dorsal
23 Frons
transverse reticulate
claw with
lobes;
pronotum fully
rugose and with
one peg-like hair
black •••••••.•••••••. 23
longi tudinal
(Fig.
keel s;
enlarged
110) . . . • • . • • . . . • . . • • • • . • . • . . .
17. amoenus n. sp. Frons without very broad,
longitudinal
keels,
reserrbl ing areolae;
strongly
enlarged
punctate;
claw wi th
punctures
two peg-I ike
hairs (Fig. 111) ....•••••..•••.•.• 18. pegnai n. sp. 24 Mesothorax, metathorax and propodeum black ••••••.••••••.••• . ••••• . ••••••••••• . • . . ••••••. . . . " 19. pseudoamoenus n. sp. Mesothorax,
metathorax and propodeum
testaceous ••••••••••••••••
150
Mem.
Amer.
Ent.
Inst.,
· ....•••........••...•.........• " 25 Notaulices scutum;
thin,
reaching
pronotal
Notaulices deep, pronota I
dorsa I
of
reaching
posterior margin
reddish;
22.
notaulices
transversal
21.
of
scutum;
52.
furrow
about
approximately
0,50
biloboides n. sp. edges
pseudobilobus n.
of
sp.
Head without
a
of
ocell i • • • • • . • • . • • • • • • . • • • • • • • • • . • • • • • • • . . • • . • • • . 28
posteri or
furrow
0,75
bilobus (Fenton)
joining posterior 23.
transversal
sp.
reaching
reaching
ocell i . . . . . . . . . . . . . . . deep
insignis n.
notaul ices
scutum . . . . . . . . . . . . . . . . . .
posterior
of
less promi nen t . . . . . . . . . . . . . . . . . . . . . . . . ..
fully
27 Head with a deep
length
almost
scutum . . . . . . . . . . . . . . . . . .
of
sp.
0,50-0,75
lobes more prominent . . . . . . . . . . . . . . . . . 26
Pronotum mostly black; length
1984
townesi n.
20.
approximately
· . . . . . •• . . . . • •• . . •• . . ••. •• . . . . length
37,
dorsal lobes
26 Pronotum almost
no.
joining posterior
28 Posterior surface of pronotum approximately long l and with two strong transverse dorsal Posterior
surface
less
than
twice as broad as
wi th
weak
29 Region
of
paral lei
of
pronotum approximately
(not
longl,
keels;
ocell i
pronotum fully
of
rugose,
head behind posterior
without
longitudinal
as
as broad as
without
dorsal
long
lobes
or or
with
numerous
reddish-testaceous.
· .•.•....••..•.•...............•.• Region
as broad
lobes . . . . . . . . . . . . 29
lobes • • . . . • • • . . • . . • • • . . • 31
transversal)
head behind posterior
longitudinal
and
twi~e
edges
24.
ocel I i
rubrolobatus n. punctate
or
sp.
reticulate
keels...................
.. .. 30
nigrolobatus n. sp. Prothorax fully reddish . . . . . . . . . . . 26. vagans n. sp.
30 Prothorax
fully
31
surface
Posterior rugose,
or with
or mainly of
Posterior
surface of wi thout
32 OOL approximately 33 Head
than
or
pronotum fully
twice as long
rugose,
posterior
long
rugose, to
reticulate wi thout
occipital
34 Head almost keels
on
Head more
or
longitudinal 35 Posterior
two
smooth, not
le~s
with
ocell
two to
or almost
strong
fully
smooth,
rugose,
wi th
a
.38 .33
27.
granulate,
median
reticulate
ret iculate
carina . . . . . . . . .
pilosifrons Fen t on almost
smooth,
weakly
connect ing posteri or 28. keel
.34
oblique keels
occipital
obi ique keel s
ocell i
politus n. sp. and
two weak
lateral
rugose . . . . . . . . . . . . . . . . . . . . . . . . 35
rugose
or
sculptured by
numerous
keels • • • . • • . . • • . . • • • • • • • . • • . . • • • • . • • • • • . • . . • . . . . 36
surface
of
pronotum slightly
Breadth measurements refer surface.
reticulate
keels . . . . . . . . . 32
as POL......... ••••••. •••.
carina . . . . . . . . . . . . . . .
frons,
or
as POL.............
· . . . . . • . . . . . . . • . . . . . . . . • • • . • • • . . .. Head not
rugose
longitudinal
sculpture among punctures........
twice as
reticulate
connecting
25.
pronotum fully
transversal
punctate, OOL more
black ..
to
broader
the anterior margin of
than
long,
the posterior
Anteoninae
smooth,
with
margin,
without
Posterior long,
of
numerous
and with
short
longitudinal
transversal
surface
with
margin
numerous
keels.
short
some
less
than
three
times as
than
three
least
long
as POL... • • . . • . . . . •
reddish
or
31.
keels
5 of
chiapasi n.
tarsi
wi th
a very
long
• •• 37
nigricans (Cameron) actuosus n.
of head behind posterior ocel I i without connecting posterior ocelli to occipital
fore
sp.
testaceous •••.•.•••••••
32. 38 Region
sp.
as POL ••••••. 30.
times as
partly
keels . . . . . . . . .
rivularis n.
long
37 Species fully black •••••.••••.•••• Species at
47.
sp.
times as broad as
keels near posterior
transversal
. . • . • . • • • • • . • . • • . • . • • • . . • OOL more
1,5
longitudinal
strong
keels near posterior
claripes n.
29.
pronotum nearly
.............................. 3600L
151
I ame I I a
on
sp.
two oblique carina;
segmen t
(F i g •
129) •..
ap ex
33. incaicus n. Region
of
head behind posterior ocelli
connect ing posterior
ocell
fore
very
39 Head
tarsi
without
sculptured by
a
to long
with
occipi tal lamella
longitudinal
keels,
• •• . . • • • • • • • • . • • • . . . . . . . • . . . . • .
34.
on
sp. oblique keels
two
carina; apex
segment
(Figs.
5 of
131,
132)
.•. 39 not reticulate rugose ••• bolivianus n. sp.
Head fully or mainly reticulate rugose ••••••••••.•••••••••••• 40 40 Vertex of head behind ocell fully reticulate rugose . . . . . . . . . 41 Vert ex 41
of
Prothorax
head beh i nd
oce I
not
ret i cu I ate
reddish . . . . . . . . . . . . . . . . .
35.
rugose...
• ...•••• 42
Prothorax b l a c k . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
42 Head fully
or almost
fully
testaceous
• • . • • . • • • • • • • • • . • • • • • • • • . . • • • . . •• , Head ma i n I y 43 Vertex among
of
b I a ck • • • • • • • • • • • • • • . ••
head behind
punctate,
of
head behind
long i t ud i na I ful Iy
ocel I i
strongly
kee Is • . . . • . • . . . . • . . • • • . . .
reddish-testaceous.
Head most I y black •••.•••..•••.••• 45 Prothorax black . . . . • • • • • • • . . . . • Prothorax
reddi sh-testaceous •••
sp.,
sp.,
reddish ..
trinidadi n. sp. rapax n. sp. and wi thout
sculpture •••. 44
sculptured by . . . . . . . . . 45
croceus n. sp. asper n. sp. 40. schlingeri n. sp. 41. alexandrae n. sp. 38. 39.
D. hirticornis (K i ef f er), inermis n. sp., cuzcanus aequalis n. sp., paranus n. sp., perlucens n. sp., carpens n. benianus n. sp., itenezi n. sp., panamensis n. sp. are unknown. The females of
n.
or
punctures . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
Vertex 44 Head
ocell i
36. ;J 7.
.. . . . 43
eminens n. sp.
152
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
MALES
Antennal
hairs
Antennal
hairs as
shorter
segments
(see
than breadth
long as
or much
of
segments ••••••••.•••••• 2
longer
than breadth
2
Head
3
Head at least partly rugose, reticulate rugose Gonoforceps wi th reduced internal branch (Fig.
smooth,
not
rugose,
wi thout
· . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. 1 17, 4
5
large
internal
26.
122) . . . . . . . . . . ..
vagans n.
sp. (Figs.
1 4 2 ) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. 4 Internal branch of gonoforceps subapical (Fig. 141.) . . . . . . · ............................... " 42. carpens n. sp. Internal branch of gonoforceps apical (Figs. 117,142) . . . . . 5
Posterior
surface of
propodeum smooth
surface
of
43.
Gonoforceps much Gonoforceps
shorter
slightly
than penis
23.
than penis
shorter,
94,
(Figs.
and
shiny ••••••••
perlucens n.
sp.
propodeum rugose and dul I ••••••••••..
· ............................... " as
pseudobilobus n. 96,
(Figs.
long as,
or
137,
sp.
156) . . . . . . 7
slightly
longer
143) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9
Head not reticulate rugose, with longitudinal keels, punctate and slightly rough; very strong and distinct oblique keels connecting posterior
ocell i
to
occipi tal
....... Head
fully
reticulate
connecting posterior
8
sp.
or with keels .. 3
141,
Posterior
7
.. . . . . ..
branch wrapping penis
· ............................... "
6
.. . . . . . 6
areolae or keel s.
4. nigrorufus n.
Gonoforceps wi th
of
79)............................
fig.
Gonoforceps without
rugose; ocelli
internal
indistinct branch
internal
branch
carina . . . . . . . . . . . . . . .. and Currado)
bicolor (Olmi
to occipital
· ............................... " Gonoforceps with an
6.
oblique keels carina . . . . . . . . . . . . . . . 8 (Fig.
137) ..
sp. wrapping penis (Fig.
156) ..
wrapping penis
38.
croceus n.
· . . . . . • . . . . . . . . . . . . . . . . . . . . . . . . . . . 55. panamensis n. 9
Gonoforceps wi thout (Figs.
94,
branch
keels,
large
internal
branch
except
fully for
wrapping penis
granulate and punctate, few
short
rugose,
wi th
without
keels near antennal
. . . . . . . . •. . . . . . . . . . . . . . . . •"
11
very
(Figs.
144) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .' . . . . . . . . 14
10 Head smooth,
Head
or wi th
143) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 10
Gonoforceps wi th
130,
internal
sp. reduced branch
5.
iphias n.
areolae or
sockets . . . . . . . . sp.
areolae and keel s . . . . . . . . . . . . . . . . . . . . . . . . . . 11
reaching approximately 0,80 length of scutum . . . . . . .. · ............................... " 44. inermis n. sp. Notaulices reaching at most 0,50 length of scutum . . . . . . . . . . . . 12 Notaulices
12 Posterior surface of propodeum without longitudinal keels . . . . .. · ............................... " 1. elegans n. sp. Posterior
surface
of
propodeum with
strong
longitudinal
keels
153
Anteoninae
· . . . • . . . • . . . • • . • • . . . • • • • . • • . . . . . . . . . • • • • . • • . . . . . . . . • • . . • • • • . . 13 13 Gonoforceps with distal inner pointed process (Fig. 155) . . . . . . . · •.....••.••...••••••••••..•.•..•. 54. cuzcanus n. sp. Gonoforceps without distal inner pointed process (Fig. 152) ••.. • • • • . • . • . . . . • • . . • . . • • . . . . • • • . . . . •• 51. itenezi n. sp . 14 Head without two oblique keels connecting posterior ocelli to occipital
carina;
vertex
of
head behind
ocelli
never
reticulate
rugose •••••••••••••••••••••.••••••••••••••••••••••••••••••••• 15 Head with
two
to occipital
15 Head fully
strong
smooth,
punctures,
not
region
keels
sometimes
finely
reticulate
Head with anterior posterior
oblique
carina,
region
connecting posterior
indistinct
punctate,
among
without
45.
of
strongly
frons
sculpture among
paranus n. sp.
rugose ..
and vertex
ocelli
areolae . . . . . . 16
reticulate
rugose;
smooth . . . . . . . . . . . . . . . . . . . . . . . . . . ..
• ••.•••••.•••...••••.•••.•••...••. 33. incaicus n. sp. 16 Gonoforceps with internal branch on proximal region (Fig. · . . . . . . . . . . . . . . . . • • . • • • • . . . . . . . . •. 25. nigrolobatus n. sp. Gonoforceps with (Figs.
104,
17 Frons with
internal
branch
on
apical
or
subapical
two median Some
Frons wi th
or wi thout
longitudinal
transversal
folds,
keels; not
region between
reticUlate
region
of
segment
two di st inct
frons always
reticulate
3 approximately as
....................••.....•...... Antennal
segment
19 Posterior
surface
3
longer of
excavated
than
Posterior
surface of
than
lateral
20 Distal
part
than
segment
propodeum wi th
lateral
areas . . . . . .
propodeum wi th
1 .... .
segment
hirticornis (Kieffer) 1 •••..•••••••••.•••••• 19
median
14.
keel s;
rugose . . . . . . . . . . . . . . 18
long as
46.
sp.
I ongi tudinal
median
these
rugose . . . . . .
12. peruvianus n.
18 Antennal
region
145) • • • • • . . . . • • . . . . . • . • • • . . . . • . . • . • • • . . . . • . . . . . . . 17
keels with
median
120)
area more deeply
colombianus n.
median
area
sp.
less
excavated
areas . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 20 of
radial
vein
shorter
or at
most
as
long
as
proximal p a r t . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . • .21 Distal part of radial vein longer than proximal part . . . . . . . . . 22
21
Notaulices
reaching
approximately posterior margin
(F i g.
157 B ) . . . . . . . . . . . . . . . . . . . Notaulices shorter (Fig. 157 A) . . . 22 Posteri or surface of
propodeum wi thout
· . . . . . . . . . • • . . . . . • • . . . . . • • . . . . . . •. Posterior
surface
of
propodeum with
· . . • . . • . . . . . . . • . . . . . . . . . . . . . . . . . ..
of
scutum
9. noyesi n. sp. 36. trinidadi n. sp. longitudinal
50. benianus n. two
48.
keels . . . . . .
sp.
longitudinal
aequalis n.
keels . . . . .
sp.
The males of D. achterbergi n. sp., schlingeri n. sp., eminens n. amoenus n. sp., asper n. sp., diaphanus n. sp., bolivianus n. sp., melanocephalus (Cameron), nigricans (Cameron), pegnai n. sp., alexandrae n. sp., bilobus (Fenton), pilosifrons Fenton, maximus n. sp., insignis n. s::>., townesi n. sp., albop'-ctus n. sp., hymenaeus n. sp., fluviatilis n. sp.,
154
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
sp., biloboides n. sp., rivularis n. sp., minor n. sp., politus n. sp., c1aripes n. sp., rubrolobatus n. sp., rapax n. sp., speciosus n. sp., atlanticus n. sp., pseudoamoenus n. sp., chiapasi n. sp., cascus n. sp.,
actuosus n. sp. are unknown.
1. Deinodryinus elegans n. sp.
FEMALE: region
fully
winged;
blackish,
length 3,0 mm;
sides of pronotum
reddish-testaceous,
with
a
black
scutum
and metanotum blackish,
black,
abdomen and petiole brown-testaceous;
antenna I segments head
dull,
complete;
fully
=
POL
3;
=
OL
behind
posterior ocelli
strong
anterior
granulated and
with
tegulae;
scutum
shiny,
reaching
approximately 0,5
shiny,
smooth,
with
2 dark
beneath part
nearly
strong
without
fore
(Fig.
produced
into a
winged;
partly
legs
brown
in following fully OL
=
3;
=
7;
punctate,
reaching
5:
=
6;
tibial
scutum,
without
10:
6,5: some
vein
nearly
branch 88);
hyal ine, as
6;
penis,
spurs
LOCUS TYPICUS:
(Fig.
dark
part a
keel
fore-wings
longer than
87)
6:
with
4:
7:
7:
7:
7:
segment
and
19; the
3 of
hind
6,5:
hairs much
tibiae
antenna I segments 8,5;
head dull,
region
behind
carina complete;
punctures;
(7:7);
12:
than
mandibles and antennae
thickened;
POL = 5;
longer than
notaul ices
smooth, incomplete,
propodeum reticulate rugose, without bands;
longitudinal distal
keels;
part of radial
gonoforceps without
short apical
(Trinidad)
proximal
2:
2 rows of
lamellae;
irregular keels;
transversal
with only
a
1,1,2.
black;
length of scutum;
Augustine
transversal
keels;
clubs of femora
antennal
1,1,2.
St.
strong
scutellum and metanotum shiny,
long as proximal
wrapping
tibial
without
with
incomplete,
located further distally
with posterior surface reticulate rugose and fore-wings
a
vein
occipital
sculpture among
approximately 0,5
8;
tubercles reaching
proportions:
spurs
coxae,
=
6:
of head
hairy,
notaulices
group of nearly 6
with
TL
region
longitudinal
following
2 obi ique keels;
OPL
breadth of segments; finely
10: frons
7;
6:
carina
and posterior surface fully
antennae not distally
proportions:
OOL
in
with
6:
the basa I Ce II s and another
length 2,31-2,50 mm;
reticulate rugose; with
with
dorsal
part of radial
a
thickened;
6:
scutellum and metanotum
propodeum
hook;
testaceous,
or black;
posterior ocell i
with
=
TL
6:
occipital
pronotal
segment 5 of fore tarsi
fore tarsi
7:
pronotum dull,
keels;
with one bristle
apex
11:
granulated;
one on
distal
lamellae;
fully
7;
length of scutum;
segments
87)
prominence;
testaceousj
=
posterior surface without
tarsal
6:
posterior surface of pronotum
weakly
sculpture;
54
MALE:
OPL
transversal
smooth,
the pterostigma;
enlarged claw
7;
furrow;
transversa I bands,
(13:9);
proximal
=
OOL
and posterior surface;
reticulate rugose;
10:
without oblique keels;
transversal
between dorsal
proportions:
propodeum
antennae distally
without areolae or keels;
3;
with ocellar
anterior margins of
mesopleura distally blackened,
in following
granulated,
spot,
an apical
membranous band
(Fig.
Anteoninae
Fig. 87 -
Chela of
Deinodryinus eJegans n. sp.
155
(type)
156
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
v
Fig.
88 -
Male genitalia of Deinodryinus elegans n.
(Trinidad);
A:
left half (dorsally viewed); (ventrally
B:
and 1 paratype M!
DISTRIBUTION:
TRINIDAD:
BOLIVIA:
Neotropic region:
Mouth of Rio Baures
(Rio
from St.
Augustine
viewed).
TYPICAL MATERIAL: holotype F! in AM; 1 para type M! in OL. HOSTS: unknown OL!
sp.
right gonoforceps
st.
Itenez,
in BM;
Augustine Dept.
1 paratype F!
(St.
Beni),
George),
AM!
BM!
157
Anteoninae
2. Deinodryinus speciosus n. sp.
FEMALE:
fully
not black;
winged;
length 3,50 mm;
antennae distally
proportions:
9:
6:
14:
7:
frons without median or Ol
~
3;
~
OOl
7;
OPL
numerous transversal scutum
shiny,
incomplete,
6,5:
lateral 8;
keels,
TL
6:
6:
keels; ~
7;
hairy;
reaching
5,5:
transversa I keel
posterior surface small;
petiole testaceous,
pronotal
in following
head dull,
granulate;
carina complete;
pronotum dull,
granulated,
tubercles reaching
weakly granulated; length of scutum;
without sculpture; between
6,5;
occipital
approximately 0,5
smooth,
testaceous;
antenna I segments
without sculpture or very
metanotum shiny, without
~
6:
fully
thickened;
POL
~
3;
with tegulae;
notaulices
scutellum and
propodeum reticulate rugose,
dorsa I and posteri or surface;
posterior surface without
longitudinal
areol ae on keels;
fore-
wings with 2 dark transversal bands, one on the basal cells and another beneath the pterostigma; distal part of radial vein longer than proximal part 21;
(14:8);
fore
tarsal
enlarged claw
(Fig.
segments 89)
in following
proportions:
with one bristle
the proximal
prominence;
segment 5 of fore tarsi
41
apex with a
group of 7
lamellae;
Fig.
MALE:
89 -
2,5:
5:
lamellae;
(Fig.
tibial
89)
15: than
with 2 rows of
spurs 1,1,2.
Deinodryinus speciosus n. sp. (type)
unknown
lOCUS TYPICUS:
Teodoro Sampaio
TYPICAL MATERIAL: HOSTS:
Chel a of
7:
located further distally
holotype F!
(Sao Paulo,
Brazil)
in OT.
unknown
DISTRIBUTION:
Neotropic region:
BRAZIL:
Teodoro Sampaio
(Sao Paulo),
OT!
158
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
3. Deinodryinus diaphanus n. sp.
FEMALE:
fully
petiole black;
Fig.
winged;
length 3-4 mm;
antennae slightly
90 -
fully
testaceous;
distally thickened;
sometimes with
antennal
Chel a of Deinodryinus diaphanus n.
sp.
segments
(type)
in
159
Anteoninae
following proportions: finely
punctate,
granulated, POL
~
3;
without
OL
~
12:
2;
tegulaej
transversal
proximal 2:
4:
part
15:
than
10:
9:
and
lateral
OPL
~
8;
furrow,
8;
weakly
scutellum
granulated;
7:
7:
10;
~
bands; 11);
distal fore
enlarged claw
the proximal
and
lamellae;
carina complete;
pronotum shiny,
granulated; metanotum
pronotal shiny,
tarsal
radial with
vein
in following
with
without
shorter than
proportions:
9:
located furter distally
segment 5 of fore tarsi
apex
without
weakly
sl ightly
1 bristle
with
almost
fore-wings hyaline,
segments 90)
smooth, tubercles
smooth,
propodeum shiny,
weak;
part of
(Fig.
prominence;
rows of nearly 40
occipital
8;
head shiny,
frons weakly
notaulices almost complete,
with sculpture very
(10:
24;
7:
keels;
TL
the posterior margin of scutum;
reticulate rugose, dark
11:
~
scutum,
sculpture or weakly reaching
15:
median
OOL
an anterior transversal reaching
6:
without sculpture among punctures;
(Fig.
a group of nearly 5
90)
with 2
lamellae;
tibial
spurs 1,1,2. MALE: unknown LOCUS TYPICUS:
Nova Teutonia
TYPICAL MATERIAL: HOSTS:
(Brazil)
holotype F!
and
1 paratype F!
in 8M.
unknown
DISTRIBUTION:
Neotropic region:
BRAZIL:
Nova Teutonia
(Sta Catarina),
BM!
4. Deinodryinus nigrorufus n.
FEMALE:
fully
winged;
length 3,12-3,25 mm;
testaceous;
antennae fully
4-10 dark;
scutum reddish,
anterior margin;
testaceous,
head and prothorax
occasionally
without or with 2
scutellum fully
black,
=p.
reddish-
with segments 3-10 or
lateral
brown spots near the
occasionally fully
or partly
reddish; metanotum and propodeum black; mesosternum, metasternum, mesopleura reddish; metapleura black, with reddish central spot; abdomen brown; femora
legs testaceous,
with
hind coxae brown;
and extremities of hind tibiae brown;
occasionally
hind clubs of
antenna I segments
in
following proportions: 10: 5: 13: 8: 7: 7: 7: 5: 6: 9; head dull, smooth, granulated, without areolae or keels; occipital carina complete; frons without median or ~
8;
lateral
pronotum dull,
impression; smooth,
pronotal
very
weakly
with
a
tubercles reaching granulated;
posterior margin of scutum; without sculpture;
POL ~ 4;
keels;
granulated,
OL ~ 2;
weak
tegulae;
notaul ices
scutellum
propodeum dull,
OOL ~
10;
OPL ~ 7;
TL
anterior transversal scutum hairless,
incomplete,
and metanotum shiny,
reticulate rugose,
shiny,
almost reaching
hairy;
the
smooth and dorsal
surface
with wide areolae; posterior surface with small areolae; no longitudinal keel on posterior surface; fore-wings hyal ine, without dark transversal '::lands; tarsal (Fig.
distal
part of radial
segments 91)
with
vein
longer than
in following proportions: 1 bristle
8:
proximal
3:
located further distally
5:
13:
than
part 21;
(11 :9);
fore
enlarged claw
the proximal
160
Mem.
Fig.
prominence; of nearly 31 spurs
1,
MALE:
1,
fully
91
-
Chela of
Amer.
Ent.
no.
37,
1984
Deinodryinus nigrorufus n. sp.
segment 5 of fore tarsi lamellae;
Inst.,
apex with
a
(Fig.
91)
(type)
double-S shaped,
with 2 rows
group of nearly 4 lamellae;
tibial
2. winged;
length 2,81
mm;
black;
mandibles and antennae
black; legs brown; antennae not distally thickened; antenna I hairs shorter than breadth of segments; antenna I segments in following proportions: 8: 6:
9,5:
10:
9,5:
9:
9:
9:
without sculpture among occipital scutum,
8:
9;
head shiny,
punctures;
carina complete;
POL
=
7;
=
Ol
scutellum and metanotum shiny,
sculpture among punctures;
smooth,
finely
punctate,
frons without median or 2;
=
OOl
smooth,
6,5;
finely
notaulices complete,
lateral
OPl
=
5;
punctate,
posteriorly
Tl
keels;
=
5;
without
separated;
propodeum dull, reticulate rugose, without transversal or longitudinal keel s; fore-w i ngs hya line, without dark transversa I bands; d i sta I part of radial
vein slightly
shorter than proximal
92) with long apical branch wrapping lOCUS TYPICUS: Puyehue (Chile) TYPICAL MATERIAL:
3 para types FF!
HOSTS:
unknown
DISTRIBUTION:
in OT;
3paratypes
Neotropic region:
Santiago Province), OT!
Pichinahuel
OT!
Ol!
TU!
(Cord.
Cobquecura
Puyehue
CHilE:
(2 FF,
Maipu
(Ososno),
Nahuelbuta,
(Nuble),
(9:10);
tibial
and 1 paratype F!
holotype F!
in TU;
part
penis;
in CA;
1 M)!
Ol!
OT!
(Fig.
5. Deinodryinus iphias n. sp.
Que.
Tregualemu
Ol!
OT!
1 paratype F!
in Ol.
(Rinconada,
CAl
Arauco),
gonoforceps
spurs 1,1,2.
la Reina
la Plata, (Maule), (Santiago),
Anteoninae
Fig.
92 -
Male genitalia of Pichinahuel
FEMALE:
fully
testaceous;
winged;
Deinodryinus nigrorufus n. sp. from
(Chile):
right half removed
length 3,75 mm;
antennae brown,
161
black;
with hind coxae and clubs of femora partly margin of pronotum testaceous; segments
clypeus and mandibles
with segments 1-3 testaceous; antennae distally
in following proportions:
12:
7:
legs
brown or black;
17:
12:
testaceous,
posterior
thickened;
antenna I
10:
10:
10:
9:
9:
10;
head dull, punctate and granulated, without areolae or keels, smooth; frons without median and lateral keels; occipital carina complete; region behind ocelli
without
occipital carina; shiny,
with
reaching
a
strong
tegulae;
transversal nearly
as
weakly
granulated;
transversal
smooth
as broad;
margin of scutum; sculpture;
OL = 4;
OOL = 8;
impression,
posterior ocelli
OPL = 8;
TL = 7;
with pronotal
posterior surface of pronotum rugose,
striae, long
two oblique keels connecting
POL = 4;
near. the posterior margin; scutum shiny,
notaulices
smooth,
incomplete,
punctate,
almost reaching
without
tubercles
with some
posterior surface
finely
scutellum and metanotum shiny,
propodeum reticulate rugose,
to
pronotum
smooth,
transversal
very
the posterior without or
longitudinal
keels; posterior surface fully reticulate rugose; fore-wings with two weak dark transversal bands, one on the basal cells and another beneath the distal
part of radial
fore tarsal
pterostigma;
segments
in following
claw
93)
(Fig.
prominence;
with
vein shorter than proportions:
11:
3:
1 bristle located further distally
segment 5 of fore tarsi
(Fig.
93)
with
proximal 7:
20:
than
part
27;
(12:15);
enlarged
the proximal
two rows of 39
162
Mem.
lamellae;
apex with a
fully
winged;
antennae brown,
group of 5
no.
37,
11:
thickened;
antenna I segments
11:
not reticulate rugose,
tibial
black;
1984
11:
13;
spurs 1,1,2.
legs testaceous,
without keels,
brown;
antennal dull,
with
fore and
hind tibiae
hairs
in following
head smooth,
(type)
mandibles testaceous;
clubs of femora partly
antennae not distally
10: 11:
lamellae;
segment 1 testaceous;
brown;
breadth of segments;
17: 9:
Ins!.,
length 2,68-3 mm;
with
hind coxae partly darkened;
En!.
93 - Chela of Ueinodryinus iphias n. sp.
Fig.
MALE:
Amer.
longer than
proportions:
punctate,
13:
7:
granulated,
except for a short keel
between eyes
and antenna I sockets; frons without median and lateral keels; region behind ocell i without two obi ique keels connecting posterior ocell i to occipital = 8;
carina;
TL = 7;
incomplete,
occipital
without
smooth,
distal (Fig.
94)
with
or
sculpture;
longitudinal
fore-wings hyal ine,
part of radial very
vein
keels;
reduced
apical
branch;
unknown
OPL
scutellum and reticulate
posterior surface fully
without dark
shorter than proximal
OOL = 8;
notaul ices
propodeum dull,
transversal
part
tibial
LOCUS TYPICUS: Campina Grande (Brazil) TYPICAL MATERIAL: holotype F! and 1 para type M! in OL. HOSTS:
OL = 3;
punctate;
the posterior margin of scutum;
without
transversal
reticulate rugose;
POL = 6;
weakly granulated,
almost reaching
metanotum shiny, rugose,
carina complete;
scutum shiny,
(9:14);
bands;
gonoforceps
spurs 1,1,2. in TW;
1 paratype M!
Anteoninae
DISTRIBUTION:
Neotropic region:
Serra do Caraca
Fig.
94 -
(5.
Barbara,
BRAZIL:
163
Campina Grande
Minas Gerais),
(Curitiba),
TW!
OL!
Male genitalia of Deinodryinus iphias n. sp. Grande (Brazi I): right half remov",C;.
from
Campina
6. Deinodryinus bicolor (Olmi and CurradoJ n. comb.
Lonchodryinus bicolor 0 I m i and Currado 1979a: 340.
FEMALE: fully winged; length 3,75-5,81 mm; head black, with mandibles black or yellow; antennae yellow, with segments 6-10 or 7-10 brown-black; segment 3 brownish; black;
pronotum,
segment
scutum,
1 with a brown
scutellum,
sometimes anterior collar of pronotum black; propodeum black; brown or black;
dorsal
metanotum
mesopleura,
somet i mes mesop I eura part I y legs testaceous,
with a
streak;
dorsal
propleura
reddish-testaceous; red;
metapleura and
pet io I e black;
basal
abdomen
black spot on
hind
164
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
I
Fig.
coxae;
95 -
Chela of Deinodryinus bicolor (Olmi
tegulae testaceous;
antennal
and Currado)(type)
segments in following proportions:
12:
7: 15: 7: 7: 7: 7: 6: 5: 8; head shiny, smooth, hairy on frons and clypeus, without sculpture; frons with 2 longitudinal parallel keels connecting ocellar region region
to clypeus and with keel
behind ocellar triangle with
around the orbits;
two obi ique keels connecting posterior
Anteoninae
ocelli
to occipital
very 15;
strong
carina;
and keel
pronotum shiny,
scutum,
smooth,
scutum;
=
3;
small
longer than proximal segments
claw
95)
(Fig.
with
approximately 34
testaceous; distally
legs
segments
part;
with
distal
distal
apex
with
length 3,5 mm; with
with
hairs
frons with
longitudinal
reticulate rugose;
two
keels;
vertex
a
length of
2,5:
with
5:
8:
11:
lateral
an
19;
than
the proximal
lamellae;
tibial
mandibles and antennae brown;
antennae not
11:
10:
11:
11:
10:
antenna I
10:
keels around the orbits and
frons weakly
angle;
enlarged
than breadth of segments; 7:
without
rectilinear,
three rows of
hind coxae basally
longer
smooth,
vein
group of nearly 40
black;
TL
(areolae very
parts forming
9:
95)
15;
the sides;
without sculpture;
located further distally
in following proportions:
head shiny;
hairy on
part of radial
(Fig.
carina
=
OPL
approximately 0,30
and proximal
two bristles
testaceous,
thickened,
strongly
in following proportions:
lamellae;
spurs 1,1,2. MALE: fully winged;
15;
surface reticulate rugose
segment 5 of fore tarsi
prominence;
occipital
=
OOL
mesopleura and metapleura shiny,
fore-wings hyaline,
fore tarsal
hollow; 3;
smooth,
reaching
dorsal
posterior surface,
=
OL
finely punctate,
incomplete,
propodeum hairy;
sculpture;
median
with a
POL
scutellum and metanotum shiny,
notaulices distinct, wide);
vertex
shaped;
165
rugose among
not reticulate rugose,
12;
8: with
the keels,
two
not
punctate and without
sculpture among punctures; region of head behind ocell i with two obi ique keels connecting posterior ocelli to occipital carina; vertex with some deep furrows
= 9;
around
OPL
finely
the ocell i;
= 9;
TL
punctate,
complete,
surface smooth distal
(12:11); broad
10;
without
and shiny;
transversal vein
longer than
holotype F!
Quincemil
(Marcapata),
sl ightly
fully
part of frons,
winged;
with
96);
posterior
transversal
longer than proximal (Fig.
OOL
smooth,
propodeum
keels,
without dark
part
gonoforceps with
spurs 1,1,2.
in GE. COLOMB I A:
TW!
BRAZIL:
Meta
length 5,31-7,50 mm;
(Cord.
Itajtuba
7. Deinodryinus maximus n.
FEMALE:
OL = 3;
notaul ices almost
longitudinal
gonoforceps
(Fig. 96); tibial I tajtuba (Braz i I)
HOSTS: unknown 01 STR I BUT I ON: Neotrop ic reg ion: PERU:
or
fore-wings hyal ine,
branch
TYPICAL MATERIAL:
punctures;
the posterior margin of scutum;
part of radial
LOCUS TYP I CUS:
POL = 5;
carina complete;
scutellum and metanotum shiny,
without sculpture among
penis much
apical
occipital
scutum,
almost reaching
reticulate rugose, bands;
=
8M!
GE!
sp.
head black,
clypeus and mandibles testaceous;
Macarena),
(Para),
with
anterior
antennae brown,
with
segmen ts 1-3 or 1-5 testaceous; thorax and propodeum black; tegu I ae testaceous; abdomen black; legs with coxae black (distally testaceous), trochanters
testaceous,
clubs of femora black,
stalks of femora,
tibiae and
166
Mem.
Fig.
96 -
Male genitalia of Quincemil
tarsi
testaceous
segments
in
head shiny,
finely
punctate,
clypeus;
clypeus with the orbits;
a
=
smooth,
right
5;
OL
=
finely
5;
=
OOL
25:
10:
27:
punctate,
impression;
tubercles reaching
without
14:
19;
keel;
scutum
TL
=
shiny,
head
with
17;
strong
with
to keels
carina
to occipital carina; lateral
20;
longitudinal
pronotum
smooth,
incomplete,
punctures;
13: frons
anterior ocellus
hairy,
punctures, finely
with
furrow;
shiny, anterior
pronotal
punctate,
reaching
scutellum and metanotum shiny,
without sculpture among
13:
and occipital
sculpture among
notaulices
antennal 14:
posterior disc centrally excavated;
tegulae;
length of scutum;
=
17:
keels connecting
triangle with
OPL
brownish);
20:
posterior ocell i
ocellar
sculpture among punctures; punctate,
1984
sculpture among punctures;
longitudinal
between
18;
37,
half removed.
tibiae distally
without
median
carina complete;
transversal
0,8
no.
2 oblique keels connecting posterior ocelli
occipital POL
(Peru):
and parallel
region
Ins!.,
Deinodryinus bicolor (Olmi and Currado) from
(except for hind
longitudinal
with
En!.
following proportions:
3 median around
Amer.
without
approximately
smooth,
finely
propodeum with posterior
Anteoninae
surface smooth,
shiny,
strongly
with 2-3 rows of areolae anterior surface dull finely dark
punctate;
dorsal
surface reticulate rugose,
in front of posterior surface;
and
strongly punctate;
metapleura dull
transversa I bands,
punctate;
167
fu II y
part of radial
vein
nearly
Fig.
97 -
Chela of
and strongly
darkened;
as
mesopleura with
posterior surface shiny and punctate;
hind
long as proximal
fore-wings without
wings a I so darkened; part;
fore tarsal
Deinodryinus maximus n. sp. (type)
d i sta I
segments
168
Mem.
in following proportions:
Amer.
12: 4:
6:
En!.
Ins!.,
15:
26;
one bristle located further distally than 5 of fore tarsi lamellae;
(Fig.
97)
LOCUS TYPICUS:
Gourdonville
DISTRIBUTION: Tingo Maria
1984
enlarged claw
(Fig.
97)
prominence;
tibial
with segment
with one row of nearly 23
with a group of nearly 22 lamellae;
with a group of nearly 9 lamellae; MALE: unknown TYPICAL MATERIAL: HOSTS: unknown
37,
the proximal
double-S shaped,
intermediate region
no.
apex
spurs 1,1,2.
(French Guyana)
holotype F!
Neotropic region:
(Monzon Valley),
in P;
1 paratype F!
FRENCH GUYANA:
in CA.
Gourdonville,
P!
PERU:
CA!
8. Deinodryinus melanocephalus (Cameron) n. comb.
Dryinus melanocephalus Cameron 1888: 446. Bocchus melanocephalus (Cameron): Kieffer 1914b: 46.
FEMALE:
fully
winged;
mandibles testaceous;
length 3,75 mm; antennae brown,
head black, with
with clypeus and
segments 1-4 testaceous;
thorax
and propodeum testaceous, with brown spot near tegulae; petiole and abdomen bl ack-brown; fore legs testaceous, with coxae and basa I part of clubs of femora brown; mid legs brown; hind legs brown, with coxae and proximal
half of femora
testaceous;
in following proportions: fully
reticulate rugose,
longitudinal keels
there
parallel is a
13: 6: with
tegulae testaceous;
11: 7:
7:
7:
a strong keel
6:
around
keels connecting ocell i
6:
segments
head dull,
the orbits;
frons
between
carina;
hairy, with 2
these
vertex behind ocelli
to occipital
carina complete and strongly carinate; POL 9; TL = 11; pronotum shiny, smooth, hairy,
antennal 8;
to clypeus;
track of another median keel;
2 obi ique keels connecting posterior ocell i
6:
without
occipital
=
5; OL = 3; OOL = 12; OPL finely punctate, with a
=
transversal impression; pronotal tubercles reaching tegulae; scutum, scutellum and metanotum shiny, smooth, finely punctate, without sculpture among punctures;
notaulices distinct,
0,75 length of scutum; transversal without on
between
longitudinal
the basal
radial
keel
incomplete,
propodeum shiny, dorsal
keels;
reaching approximately
with weak keels,
and posterior surface;
without a
posterior surface
fore-wings with 2 dark transversal
cells and another beneath
vein nearly as long as proximal
the pterostigma; part
(8,5:9);
distal
bands,
one
part of
pterostigma
long and
narrow; fore tarsal segments in following proportions: 12: 5: 9: 14: 30; segment 5 of fore tarsi (Fig. 98) with two rows of 34 lamellae; apex with a group of 8 lamellae;
enlarged claw
bristle; segment 3 of fore MALES: unknown LOCUS TYPICUS:
tarsi
(Fig.
98)
with a hook;
Volcan de Chiriqui
(Panama)
with one medial
tibial
spurs 1,1,2.
peg-like
169
Anteoninae
Fig.
98 -
Chela of Deinodryinus melanocephalus
TYPICAL MATERIAL: FF! in BM. HOSTS:
lectotype F!
(hereby
designated)
(Cameron) (lectotype)
and 2 paralectotypes
unknown
DISTRIBUTION: 3M!
Neotropic region:
PANAMA:
Volcan de Chiriqui,
BM!
Bugaba,
170
Mem.
Fig.
99 -
Amer.
Chela of
En!.
Ins!.,
no.
37,
1984
Deinodryinus noyesi n. sp. (type)
9. Deinodryinus noyesi n. sp. FEMALE:
fully
wingedj
and clypeus
testaceousj
6-10 brownj
legs brown,
length 4,12-5,40 mmj antennae testaceous, with
blackj
mandibles,
malar space
with segments 8-10,
fore tibiae and fore tarsi
7-10 or
testaceous;
An teon i nae
sometimes also fore coxae segments
ha irs much
longer
numerous areolae, behind ocell i particularly carina;
testaceous;
in following proportions:
171
antennae distally
13:
7:
20:
8:
than breadth of segmen ts; in addition
to 3 median
not reticulate rugose,
with
8:
thickened;
8:
7:
7:
head sh i ny;
and 2 lateral
numerous
with 2 oblique keels connecting
7:
frons keels;
longitudinal
posterior ocelli
antenna I
9;
antennal
with region keels,
to occipital
temples finely punctate and without sculpture among punctures;
occipital
carina complete;
pronotum hairy,
shiny,
=
POL
with an
3;
posterior surface of pronotum weak I y transversal tegulae;
=
OL
3;
scutellum
pronotal
and metanotum shiny,
without sculpture among punctures;
12;
OPL
=
transversal
10;
TL
=
11;
furrow;
granu I a ted and with numerous
keels mainly on anterior half;
scutum,
=
OOL
anterior strong
notaul ices
tubercles reaching
smooth,
finely
incomplete,
punctate,
reaching
approximately 0,65 length of scutum; propodeum reticulate rugose; posterior surface with 2 incomplete longitudinal keels; median area smooth; forewings with 2 dark beneath proximal 15:
34;
than
transversal
the pterostigma; (10,5:8);
fore tarsal
enlarged claw
the proximal
rows of nearly 30
(Fig.
fully
segments
prominence;
winged;
one on
the basal
part of radial
99)
lamellae;
segment 3 of fore tarsi MALE:
bands,
distal
with
vein
in following proportions:
1 bristle a
into a
99)
(Fig.
group of nearly 26 hook;
length 3,20-3,37 mm;
9: 2: 9:
located further distally
segment 5 of fore tarsi
apex with
produced
cells and another
I ittle longer than
tibial
black;
with 2
lamellae;
spurs 1,1,2.
mandibles and
antennae
fully testaceous; legs testaceous, with hind coxae proximally black; antennae not distally thickened; antenna I segments in following proportions:
14:
6,5:
14:
10:
10:
10:
10:
10:
10:
12;
antenna I hairs much
longer than breadth of segments; head ret icu I ate rugose; frons with 3 median and 2 lateral keels around the orbits; occipital carina complete; frons
(in front of anterior ocellus)
than breadth of ocellus;
with
a
transversal
furrow
region behind posterior ocelli
connecting posterior ocelli
to occipital
carina;
POL
=
less broad
with 2 oblique keels
5;
OL
=
3;
OOL
=
10;
OPL = 9; TL = 7; scutum, scutellum and metanotum shiny, smooth, strongly punctate, without sculpture among punctures; notaul ices almost complete, almost
reaching
the posterior margin of scutum
reticulate rugose, surface;
few
median
area
irregular keels;
distal
part of radial
proximal tibial
part;
(Fig.
with 2 incomplete longitudinal almost
fully
smooth,
fore-wings hyaline, vein
gonoforceps
nearly
as
(Fig.
100)
not
157 B);
reticulate
without dark
rugose,
transversal
long as or sl ightly with apical
propodeum
keels on posterior
branch
shorter
with
a
bands; than
wrapping penis;
spurs 1,1,2.
LOCUS TYP ICUS:
SI. Augustine (Trinidad) paratyoe M! in TYPICAL MATERIAL:holotype F! and 1 paratype M! in BM; LE; 1 paratype F! in WA; 6 paratypes (5 FF, 1 M)! in AM; 3paratypes (2 FF, 1 M)! in OL. HOSTS: unknown DISTRIBUTION: Curepe
Neotropic region:
(Sta Margarita),
BM!
TRINIDAD:
SURINAME:
SI.
Krepi
Augustine
(SI.
(Charlesburg,
George),
BM!
Paramaribo),
172
Mem.
Fig.
100 -
Amer.
Male genitalia of
Ent.
VENEZUELA:
no.
37,
1984
Deinodryinus noyesi n. sp. from Curepe
(Trinidad):
LE!
Inst.,
left half removed.
Ocumare de La Costa
(Araqua),
WA!
BOLIVIA:
Baures (Rio Itenez, Beni), AM! Ol! laranjeiras (Dept. Beni), NOTE: the species is named in honor of Mr. J.S. Noyes.
Mouth of Rio AM!
10. Deinodryinus atlanticus n. sp.
FEMALE:
fully
winged;
testaceous with ctypeus
and
a
anterior region
segments 8-10 dark; with black spots on pronotum
and
length 4,37-5,00 mm;
head fully
large black spot on frons and vertex; of frons
testaceous;
mandibles,
antennae
testaceOU5,
with
thorax and propodeum testaceous-reddish, sometimes the sides of pronotum, on pleura, between scutum and
between metanotum and propodel1m;
black-testaceous;
black or reddish-
legs testaceous;
petiole black;
antennae distally
thickened;
abdomen antennal
Anteoninae
Fig.
segments
101
-
173
Deinodryinus atlanticus
Chela of
in following proportions:
16:
8:
20:
12:
10:
n.
sp.
9:
9:
(type)
9:
8:
11;
head
dull, fully reticulate rugose; frons with a complete median keel; region behind ocelli with 2 slightly visible oblique keels connecting posterior ocelli
to occipital
OOL = 13; pronotal
occipital
TL = 10;
tubercles reaching
weak dorsal shi ny,
carina;
OPL = 9;
lobes parallel
smooth,
notaulices
fi nel y
incomplete,
carina complete;
pronotum shiny,
tegulae;
OL = 3;
not rugose,
with
posterior surface of pronotum with
to the sides;
punctate,
POL = 3,5;
smooth,
scutum,
two
scutellum and metanotum
wi thout scu I pture among punctures;
almost reaching
the posterior margin of scutum;
propodeum reticulate rugose, without transversal or longitudinal keels; posterior surface with median area smooth, not rugose; fore-wings with dark
transversal
pterostigma;
bands,
distal
one on
the basal
part of radial
vein
cells and another beneath
longer than proximal
part
two
the
(13:9);
fore tarsal segments in following proportions: 12: 3: 9: 13: 30; segment 3 of fore tarsi produced into a hook; enlarged claw (Fig. 101) with one bristle
located further distally
fore tarsi
(Fig.
lamellae; tibial MALE: unknown
101)
than
the proximal
with 2 rows of 28
prominence;
segment 5 of
apex with a
group of 14
spurs 1,1,2.
LOCUS TYPICUS: Nova Teutonia (Brazil) TYPICAL MATERIAL: holotype F! in OT;
."!
lamellae;
1 para type F!
in TW;
1 paratype
in OL.
'-IOSTS:
unknown
JISTRIBUTION:
Neotropic region:
BRAZIL:
Nova Teutonia
(Sta Catarina),
174
OLl
Mem.
OT!
S.
Amer.
En!.
Ins!.,
no.
J. Barreiro (Serra da Bocaina),
TW!
37,
1984
11. Deinodryinus achterbergi n. sp.
fully
mandibles and almost fully frons
black;
winged;
length 4,18-4,31
FEMALE: clypeus,
antennae testaceous,
propodeum black; black,
abdomen
brown;
thickened;
head black,
testaceous;
legs
testaceous,
and distal
antennal
with malar space,
ventral
segments 8-10 brown;
with
stalk of hind femora brown
antennae distally
mm;
with coxae partly
apex of hind
segments
face of head
thorax and
in following
tibiae brown; proportions:
14: 7: 19: 13: 9: 9: 8: 8: 7: 11; head fully reticulate rugose shiny; with 3 median longitudinal keels; 2 lateral keels around the orbits;
frons
occipital carina complete; region behind posterior ocelli reticulate rugose and with 2 oblique keels connecting posterior ocelli to occipital carina; POL = 3;
OL = 3;
DOL = 11;
OPL = 10;
strong anterior transversal punctate, pronotal shiny,
furrow;
without sccdpture among punctures, tubercles reaching
smooth,
tegulae;
finely punctate,
notaul ices almost complete, median
area
with
pronotum hairy,
scutum,
reaching
anterior
finely lobes;
scutellum and metanotum the punctures;
the posterior margin of scutum;
with 2 longitudinal
an
with a
smooth,
2 weak parallel
with
without sculpture among
almost
propodeum reticulate rugose, surface;
TL = 9;
posterior surface shiny,
keels on posterior
transversal
keel,
smooth;
fore-wings
with 2 dark transversal bands, one On the basal cells and another beneath the pterostigma; distal part of radial vein longer than proximal part (16:11); fore tarsal segments in following proportions: 11: 3: 7: 15: 30; enlarged claw (Fig. 102) with 1 bristle located further distally than the proximal prominence; segment 5 of front tarsus (Fig. 102) with 2 rows of nearly 29 lamellae; apex with a group of 12 lamellae; tibial spurs 1,1,2. MALE:
unknown
LOCUS TYPICUS:
Gamboa
TYPICAL MATERIAL: HOSTS:
NOTE:
and 1 para type F!
in LE.
unknown
DISTRIBUTION: Zone),
(Panama)
holotype F!
Neotropic region:
PANAMA:
Gamboa
(Pipeline Rd.,
Canal
LE! the species
is named
in
honor of Mr.
Kees van Achterberg.
12. Deinodryinus peruvianus n. sp.
FEMALE: and
fully
somet i mes
winged;
length
4,68-5,12 mm; black;
anterior reg ion of frons
with segments 7-10,
6-10 or 3-5;
with coxae fully black,
testaceous j
mandibles,
antennae
segments 9-10 brown;
clypeus
testaceous,
legs testaceous,
clubs of femora and mid and hind tibiae partly
175
Anteoninae
Fig.
102
-
Chela of Deinodryinus achterbergi n.
sp.
(type)
176
Mem.
black; distally 13:
11:
Amer.
Ent.
Inst.,
sometimes fore clubs of femora thickened; 10:
8:
with 3 median
Fig.
8:
antennal 7:
10;
head shiny,
longitudinal
103 -
segments keels;
2
fully
no.
37,
1984
testaceous;
antennae slightly
in following proportions:
14:
8:
reticulate rugose and punctate; lateral
keels around
Chela of Deinodryinus peruvian us n.
the orbits;
sp.
(type)
18: frons
Anteoninae
occipital
carina complete;
posterior ocelli Tl = 16;
vertex with 2 obi ique strong keels connecting
to occipital
pronotum hairy,
punctures;
pronotal
transversal
furrow;
carina;
dull,
punctures;
margin of scutum;
tubercles reaching smooth,
dorsal finely
one on
part of radial
front
distal
tarsus
proximal
in following proportions:
(Fig.
103)
(Fig.
prominence;
with basal
103)
apex with a
group of 12
tibial
spurs 1,1,2.
MALE:
fully
Fig.
winged;
lamellae;
with
blaCk;
7:
13:
(Peru):
strong scutum,
the posterior between dorsal
posterior surface
another benea th 30;
the
part; part
(13:17); than
with 2 rows of 28
mandibles,
the
lamellae;
produced
antennae and
antennae not distally
right half removed.
fore
segment 5 of
apical
104 - ·Male genitalia of Deinodryinus peruvianus n. Maria
13;
1-2 dark
segment 2 of fore tarsi
length 3,0 mm;
with hind coxae brown;
keel
located further distally tarsus
=
without sculpture
long as proximal 5:
OPl
lobes;
areolaej
part shorter than
1 bristle
segment 5 of front
hook;
testaceous,
with
dorsal
punctate,
wide
11:
12;
pronotum with a
fore-wings as
=
OOl
almost reaching
the basa I cell s and vein
4;
without sculpture among
transversal
surface with punctate;
transversa I bands,
enlarged claw
finely
propodeum with a strong
almost smooth,
segments
=
Ol
tegulae;
notaul ices almost complete,
pterostigma; tarsal
3;
posterior surface with 2 small
and posterior surface; hairy,
=
POL
finely punctate,
scutellum and metanotum shiny, among
177
into a legs
thickened;
sp.
from Tingo
178
Mem.
antennal 11;
segments
head shiny,
median
Ent.
Ins!.,
no.
37,
in following proportions:
10:
6:
reticulate rugose or with
longitudinal
carina complete; ocelli
Amer.
keels;
vertex
to occipital
2
keels around
= 5,5;
POL
longer than
and metanotum shiny,
smooth,
propodeum with a
punctate,
longitudinal with
transversal
(8:
10);
bands;
spurs 1,1,2. lOCUS TYPICUS:
104)
Tingo Maria
8:
8:
with 2
occipital posterior
scutum,
=
Tl
11;
scutell um
the posterior margin
between dorsal
and
posterior surface with shiny,
fore-wings hyaline,
finely
without dark
vein shorter than proximal
with apical
branch wrapping penis;
part tibial
(Peru)
TYPICAL MATERIAL:
holotype F!
paratype F!
1 para type F!
HOSTS:
9:
= 9;
OPl
area almost smooth,
hairy;
part of radial
(Fig.
8;
keel
wide areolae;
median
keels,
distal
in OT;
9:
frons
without sculpture among
strong transversal
keels;
=
almost reaching
surface with
transversal
gonoforceps
9:
the orbits;
OOl
finely punctate,
of scutum; 2 complete
9:
keels;
breadth of segments;
notaul ices almost complete, dorsal
= 3;
Ol
punctures;
posterior surface;
10:
transversal
with 2 obi ique strong keels connecting
carina;
antenna I ha irs much
lateral
1984
and 2 paratypes in AM;
(1M,
IF)!
1 para type F!
in
BM;
in OL.
unknown
DISTRIBUTION:
Neotropic region:
Circular Rd.),
OT!
Tingo Maria,
BM!
(Dept.
AM!
Beni),
COLOMBIA: BOLIVIA:
TRINIDAD:
la Hormiga
Rurenabaque
Curepe
(Sta.
(Mocoa,
(Beni),
Margarita,
Putumayo),
BM!
Ol!
BM!
PERU:
laranjeiras
13. Deinodryinus hymenaeus n. sp.
FEMALE: frons,
fully
vinged;
length 3,87 mm;
head black,
cl ypeus and mand i b les testaceous;
1-2 testaceous;
with anterior region of
antennae brown,
thorax and propodeum reddish-testaceous;
wi th segments petiole black;
abdomen brown, with segment 1 testaceous; legs testaceous; antennae distally thickened; antenna I segments in following proportions: 14: 7: 10:
9:
9:
9:
9:
7:
10;
head shiny,
19:
with frons sculptured by numerous
areolae and some longitudinal keels; frontal line present; regions behind and on the sides of ocell i not reticulate rugose or keeled, punctate and without sculpture among punctures, except for two short obi ique incomplete keels connecting posterior ocelli to occipital carina; occipital carina complete; smooth, nearly
POL
=
3;
Ol
with strong as
long
=
3;
as broad,
tubercles reaching transverse dorsal
lobes, and
=
11;
=
10;
Tl
=
10;
pronotum shiny,
posterior surface of pronotum
without sculpture,
with
pronotal
posterior surface of pronotum without
only
with
metanotum
notaulices almost complete,
OPl
impression;
not rugose,
tegulae;
scutum,
scutellum
OOl
transversal
two dorsal
shiny,
reaching
propodeum reticulate rugose,
without
posterior surface with a central
weak
smooth,
almost
lobes;
the posterior margin of scutum;
longitudinal
smooth
longitudinal
without sculpture;
region;
or transversal
fore-wings with
keels; two dark
179
Anteoninae
Fig.
105 -
Chela of Deinodryinus hymenaeus n.
sp.
(type)
180
Mem.
Amer.
En!.
transversa I bands,
one on
pterostigma;
part of radial
fore
tarsal
distal segments
enlarged claw proximal 26
lamellae;
(Fig.
HOSTS:
105)
37,
1984
the basa I ce II s and another benea th vein nearly as
with
1 bristle
group of 8
Quatro Barros
the
long as proximal
11:
2,5:
5,5:
11:
part;
26;
located further distally
than
tarsus
two rows of
segment 5 of front
apex with a
TYPICAL MATERIAL:
no.
in following proportions:
prominence;
MALE: unknown lOCUS TYPICUS:
Ins!.,
lamellae;
(Fig. tibial
105)
with
the
spurs 1,1,2.
(Brazil)
holotype F!
nv.
in
unknown
DISTRIBUTION:
Neotropic region:
BRAZil:
Quatro Barros
(Curitiba),
TW!
14. Deinodryinus colombian us n. sp.
FEMALE:
fully
testaceous;
winged;
propodeum black; partly
black;
abdomen
with 3 median
testaceous-dark;
15:
and
7:
17:
keels;
temples not
10:
8:
keels,
occipital rugose,
2
antennal
8:
8:
8:
lateral
mandibles
thorax and
legs testaceous,
thickened;
13:
longitudinal
and numerous transversal posterior ocell i
head reddish-testaceous;
with segments 8-10 dark;
antennae not dis'tallY
following proportions: rugose,
length 4 mm;
antennae testaceous,
with coxae
segments
10;
in
head shiny;
keels around
carina complete;
region behind
punctate and without sculpture
among punctures; behind posterior ocelli two oblique keels connect posterior ocell i to occipital carina; POL ~ 3; Ol ~ 3; OOl ~ 12; OPl TL
~
10;
furrow;
pronotum hairy,
shiny,
posterior surface finely
punctures, pronotal
with 2 weak dorsal
tubercles reaching
smooth,
shiny,
with a
strong
punctate,
tegulae;
transversal
smooth,
lobes parallel
finely punctate,
frons
the orbits
scutum,
~
10;
anterior
without sculpture among
to the sides of pronotum; scutellum and metanotum
without sculpture among punctures;
notaul ices almost complete, almost reaching the posterior margin of scutum; propodeum with dorsal surface reticulate rugose; posterior surface almost fully
smooth and finely punctate;
bands;
distal
part of radial
vein
fore-wings
with 2 dark
transversal
little longer than proximal
part
(15:14);
fore tarsal segments in following proportions: 12: 3: 8: 13: 33; enlarged claw (Fig. 106) with 1 bristle located further distally than the proximal prominence;
segment 5 of front
lamellae;
apex with
produced
into a
tarsus
(Fig.
a group of nearly
hook;
tibial
18
106)
with 2 rows of nearly 26
lamellae;
fore tarsal
segment 3
spurs 1,1,2.
MALE: fully winged; length 2,81-3,12 mm; black; mandibles and part of c I ypeus testaceous; antennae fu II y testaceous; legs testaceous, wi th coxae partly
black;
antennae not distally
following proportions:
11:
7:
15:
9:
thickened; 9:
much
longer than breadth of segments;
frons
with 3 median
behind ocelli
longitudinal
keels;
9:
9:
antennal
9:
9:
head shiny, 2 lateral
11;
segments antennal
fully
reticulate rugose;
keels around
two oblique keels connect posterior ocelli
in hairs
the orbits;
to occipital
carina;
Anteoninae
Fig.
106 -
Chela of Deinodryinus colombianus n.
181
sp.
(type)
182
Mem.
occipital scutum,
Amer.
carina complete;
En!.
POL = 5;
Ins!.,
OL = 3;
scutellum and metanotum shiny,
no.
37,
1984
OOL = 11;
smooth,
OPL = 9;
sculpture among punctures;
notaul ices
posterior margin of scutum;
propodeum reticulate rugose,
longitudinal
keels on posterior surface;
an anterior transversal hyaline,
without dark
shorter than
proximal
keel,
transversal part
branch wrapping penis; LOCUS TYPICUS:
Fig.
107 -
Tame
strongly
(Roraima),
TW!
(Fig.
107)
vein
little
with an apical
left half removed.
and 1 paratype M!
in BM;
1 para type M!
in OL.
HOSTS: unknown DISTRIBUTION: Neotropic region: Surumu
fore-wings
part of radial
Deinodryinus colombianus n. sp. from Tame
holotype F!
1 paratype M!
area clearly del imited by
spurs 1,1,2.
(Colombia):
in TW;
gonoforceps
the
with 2 complete
(Colombia)
Male genitalia of
TYPICAL MATERIAL:
distal
without
almost reaching
transversely striate;
bands;
(10:11);
tibial
incomplete, median
TL = 10;
finely punctate,
COLOMBIA:
Tame
(Arauca),
OL!
15. Deinodryinus cascus n. sp.
BM!
BRAZIL:
Anteoninae
FEMALE:
fully
ocellar spot,
winged;
antennae testaceous, testaceous,
length 5,0 mm;
occipital
carina black,
with segment
183
head reddish-testaceous,
ventral
10 darkened;
black;
propodeum black,
posterior surface reddish-testaceous;
black;
pronotum reddish-
with anterior surface and sides partly
and metanotum reddi sh-testaceous;
with brown
face almost fully
scutum,
with med i an
abdomen black;
scutellum area of
legs testaceous,
with
mid and hind coxae and mid and hind clubs of femora partly black; antennae sl ightly proportions:
16:
distally 8:
reticulate rugose; and with
20:
two
antenna I segments
10:
9:
12;
lateral
occipital
in following
head shiny,
weakly
keels around
carina complete;
the orbits region
with 2 obi ique keels connecting posterior ocell i
=
POL
10:
tracks of
tracks of keels;
behind posterior ocell i carina;
12:
11:
frons with
two median
occipital
thickened;
15:
5;
=
OL
4;
=
OOL
13;
=
OPL
12;
TL
=
14;
to
pronotum
hairy, shiny, smooth, finely punctate, without sculpture among punctures; posterior surface of pronotum nearly as broad as long; pronotal tubercles reaching
tegulae;
punctate,
scutum,
scutellum and metanotum shiny,
without sculpture among punctures;
reaching approximately 0,65
length of scutum;
with posterior surface smooth, longitudinal basal vein
keels;
part
the pterostigma; (14:
10);
distal
fore tarsal
9: 25: 44; enlarged claw
4:
11:
posterior surface without
fore-wings with 2 dark transversal
longer than proximal
proportions:
finely
propodeum reticulate rugose,
without areolae;
cells and another beneath
smooth,
notaulices incomplete,
(Fig.
bands,
segments
108)
one on
the
part of radial in following
with one bristle
located further distally than the proximal prominence; segment 5 of front tarsus (Fig. 108) with 2 rows of 43 lamellae; apex with a group of 8 lamellae; tibial MALE: unknown
spurs 1,1,2.
LOCUS TYPICUS:
San Cristobal
TYPICAL MATERIAL: HOSTS:
de
holotype F!
las Casas
(Mexico)
in OT.
unknown
DISTRIBUTION: (Chiapas),
Neotropic region:
MEXICO:
San Cristobal
de
las Casas
OT!
16. Deinodryinus albopictus n. sp.
FEMALE:
fully
winged;
mandibles testaceous; Clrothorax black,
length 6 mm;
:7:
15:
thickened; 13:
12:
,ongitudinal
antennal
10:
keels;
10:
11;
JOL
=
to occipital 12;
OPL
=
head shiny,
carina; 13;
TL
=
without occipital 13;
mesothorax,
legs testaceous;
in fai lowing proportions:
antennae 16:
8:
29:
with frons sculptured by some
region of head behind ocell i
sculpture among punctures, Gcell i
lobes whitish;
abdomen and
segments
with clypeus and
with segments 1-3 testaceous;
with posterior and dorsal
Cletathorax and propodeum black; distally
head black,
antennae brown,
punctate,
without
two obi ique keels connecting carina complete;
pronotum shiny,
POL = 4,5;
smooth,
posterior OL = 2,5;
finely punctate,
184
Mem.
Fig.
108 -
Amer.
Chela of
En!.
no.
twice as broad
prominent dorsal
and transverse
lobes;
tegulae;
scutellum and metanotum shiny,
without sculpture among punctures; approximately 0,80
1984
pronotum with a strong
posterior surface nearly
scutum,
37,
Deillodryillus cascus n. sp. (type)
without scu I pture among punctures; impression;
Ins!.,
pronotal
notaul ices
length of scutum;
as
transversa I
long,
with
two
tubercles reaching smooth,
finely punctate,
incomplete,
reaching
propodeum reticulate rugose,
without
transversal or longitudinal keels, with posterior surface smooth and shiny; fore-wings with two weak dark transversal bands, one on the basal cells and another beneath the pterostigma;
distal
part of radial
vein shorter
185
Anteoninae
part (14:20); fore tarsal segments in following proportions: 14: 3,5: 10: 17: 37; enlarged claw (Fig. 109) with 1 bristle located
than proximal
further dis ta II y
than
the prox i ma I prom i nence;
segment 5 of front
109) with two rows of approximately 45 lamellae; of approximately 10 lamellae; tibial spurs 1,1,2. (Fig.
Fig.
'.1ALE:
109 -
sp.
(type)
unknown
_OCUS TYP I CUS:
Encruzi I hada
-YPICAL MATERIAL: --1JSTS:
Chela of Deinodryinus albopictus n.
tarsus
apex with a group
(Brazi I )
holotype F!
in TW.
unknown
JISTRIBUTION:
Neotropic region:
BRAZIL:
Encruzilhada
17. Deinodryinus amoenus n. sp.
(Bahia),
TW!
186
Mem.
FEMALE:
fully
winged;
Amer.
mid and
legs black;
17:
occipital
not reticulate rugose,
pronotum shiny,
9:
black;
distally
27:
20:
carina;
hairy,
15:
longitudinal
antenna I
15:
13:
16;
keels and by
region behind ocell i
OOl = 11;
Ol = 3;
OPl = 10;
with a strong anterior transversal punctate, lobes;
with 2 dorsal
transverse
without sculpture among punctures;
tubercles reaching
smooth,
finely
notaulices incomplete,
length of scutum;
Tl = 12;
furrow;
without sculpture among
pronotal
scutellum and metanotum shiny,
approximately 0,5
16:
without oblique keels connecting posterior
POL = 4;
punctures,
scutum,
antennae occasionally
thickened;
18:
temples and
posterior surface of pronotum finely tegulae;
occasionally anterior testaceous;
legs testaceous;
numerous
carina complete;
1984
punctate and without sculpture among punctures;
behind posterior ocelli to occipital
37,
mand i b les
antennae slightly
with frons sculptured by
some areolae; region
testaceous;
in following proportions:
head shiny,
ocell i
no.
with segments 3-10 or 6-10 dark;
hind
segments
Ins!.,
length 5,00-5,62 mm;
reg ion of frons and cl ypeus testaceous,
En!.
punctate,
reaching
propodeum reticulate rugose,
except
for the posterior surface almost fully smooth, finely punctate, without scu I pture among punctures; fore-w i ngs darkened benea th the pterost i gma; distal
part of radial
segments 110)
vein
shorter than
in following proportions:
with
1 bristle
segment 5 of front apex with a
12:
proximal 3:
9:
located further distally tarsus
(Fig.
group of nearly
110)
16: than
part 35;
(15:19);
the proximal
with 3 rows of nearly
16 lamellae;
fore tarsal
enlarged claw 52
(Fig.
prominence; lamellae;
segment 3 of fore tarsi
produced
into a hook; tibial spurs 1,1,2. MALE: unknown LOCUS TYP ICUS: Nova Teutonia (Brazi I) TYPICAL MATERIAL: F! in Ol. HOSTS:
in BM;
1 paratype F!
in TlV;
1 paratype
unknown
DISTRIBUTION: BM!
holotype F!
Azul
Neotropic region:
(Minas Gerais),
TW!
BRAZil:
Nova Teutonia
Alto da Serra
(Sta Catarina),
(Morretes),
Ol!
18. Deinodryinus pegnai n. sp.
FE:MAlE:
fully
winged;
length 4,5 mm;
and part of malar space yellow; thickened; 9:
9:
9:
9:
antenna I segments 11;
head du II,
black;
mandibles,
chelae testaceous;
in following proportions:
strong I y
punctate;
sides of clypeus
antennae distally 14:
8:
punctures very
20:
12:
11:
dense and
deep, resembl ing areolae; frons without median and lateral keels; occipital carina complete; region of head behind ocelli without two oblique keels connecting posterior ocell i to occipital carina POL = 7; Ol = 5; OOl = 10; OPl = 10; Tl = 14; region of head behind ocelli punctate, not reticulate rugose or keeled;
pronotum hairy,
shiny,
with a
weak
transversal
impression and with pronotal tubercles reaching tegulae; posterior surface of pronotum smooth, finely punctate, without sculpture among punctures,
Anteoninae
187
I
/
Fig.
110 -
Chela of Deinodryinus amoenus n.
sp.
(type)
Mem.
188
nearly
as
long as broad,
metanotum shiny,
smooth,
Amer.
Ent.
Inst.,
without dorsal finely
lobes;
punctate,
punctures;
notaul ices almost complete,
of scutum;
propodeum reticulate rugose,
no.
37,
1984
scutum,
scutellum and
without sculpture among
almost
reaching
without
the posterior margin
transversal
or
longitudinal
keels, with posterior surface almost fully smooth and shiny; fore-wings with one dark transversal band beneath the pterostigma; distal part of radial vein longer than proximal part (16:11); fore tarsal segments in following proportions: 11: 3: 8: 14: 29; enlarged claw (Fig. 111) with 2 bristles
located further distally
of front
tarsus
a
(Fig.
group of nearly
MALE:
11
111)
with
than
the proximal
lamellae;
tibial
segment 5
lamellae;
apex with
spurs 1,1,2.
unknown
LOCUS TYPICUS:
EI
Canelo
TYPICAL MATERIAL:
(Chile)
holotype F!
in TW.
HOSTS: unknown DISTRIBUTION: Neotropic region:
CHILE:
NOTE:
prominence;
two rows of nearly 38
the species
is named
EI
in honor of Mr.
Canelo
(Santiago),
Luis E.
TW!
Pegna.
19. Deinodryinus pseudoamoenus n. sp.
Fig.
111
-
Chela of
Deinodryinus pegnai n. sp.
(type)
189
Anteoninae
FEMALE:
fully
winged;
length 3,87-5 mm;
testaceous;
clypeus black,
testaceous,
with segments 7-10 dark;
meta thorax and testaceous;
propodeum black;
antennae distally
proportions:
17:
10:
25:
head black;
with anterior margin
12:
prothorax
testaceous;
antennae mesothorax,
abdomen brown-testaceous;
thickened;
antenna I segments
12:
11:
9:
reticulate rugose and with some
(at
least one median)
vertex not reticulate rugose,
mandibles
testaceous;
strongly
9:
9:
11;
punctate;
legs
in
head dull,
following strongly
longitudinal
frontal
keels;
I ine complete;
occipital carina complete; POL = 7; Ol = 3,5; OOl = 13; OPl = 11,5; Tl = 11; pronotum hairy, shiny, weakly rugose, with two dorsal lobes; pronotal tubercles reaching half,
tegulae;
scutum shiny,
without sculpture among
approximately 0,75 sculpture,
punctures;
length of scutum;
punctate mainly on anterior notaulices
incomplete,
scutellum shiny,
smooth,
reaching without
only with SOme punctures near the anterior margin;
metanotum
shiny, smooth, without sculpture; propodeum reticulate rugose, with posterior surface rugose and centrally smooth; propodeum without transversal beneath part 37;
or
longitudinal
the pterostigma;
(15:23);
fore
keels;
distal
tarsal
fore-wings
with a
part of radial
segments
112)
with one bristle
dark
transversal
into hooks;
located further distally
than
11:
3:
9::15:
en I arged claw
the proximal
prominence; segment 5 of front tarsus (Fig. 112) with 2 rows of 33 lamellae; apex with a group of 14 lamellae; tibia! spurs 1,1,2. MALE: unknown LOCUS TYPICUS:
Fig.
112 -
Nova Teutonia
Chela of
band
shorter than proximal
in following proportions:
segments 2 and 3 of fron t tarsus produced
(Fig.
vein
(Brazil)
Deinodryinus pseudoamoenus n. sp.
(type)
190
Mem.
TYPICAL MATERIAL: HOSTS: unknown
holotype F!
DISTRIBUTION: OT!
Amer.
Neotropic region:
Campina Grande
(Curitiba),
Ent.
Ins!.,
no.
37,
1984
in OT;
paratype F!
BRAZil:
Nova Teutonia
in TW. (Sta Catarina),
TW!
20. Deinodryinus townesi n. sp.
FEMALE:
fully
testaceous; yellow;
winged;
mesothorax,
testaceous-brown; coxae;
15:
legs testaceous, 7:
22:
frons without
with small
thickened; 13:
10:
9:
longitudinal
without sculpture among posterior ocelli
head black,
with mandibles
with segments 1-2 testaceous;
prothorax
meta thorax and propodeum redd i sh-testaceous;
antennae distally
proportions: rugose;
length 5 mm;
antennae brown,
antennal 9:
8:
keels;
punctures,
to occipital
brown spots on
8:
in following
head dull,
reticulate
region behind ocell i
without
carina;
segments
9;
occipital
abdomen
mid and hind
punctate,
two obi ique keels connecting carina complete;
POL = 5;
= 3,5;
OOl = 13; OPl= 11; Tl= 10; pronotum shiny, finely punctate, without scu I pture among punctures; pronotum wi th a strong transversa I impression;
posterior surface nearly
prominent dorsal
Fig.
transverse
113 -
lobes;
Chela of
twice as broad as
pronotal
long,
tubercles reaching
with
two
tegulae;
Deinodryinus tOI.rnesi n. sp. (type)
OL
Anteoninae
scutum,
scutellum and metanotum shiny,
sculpture among punctures; 0,75
length of scutum;
longitudinal wi th
two dark
beneath part 31;
keels,
113)
prominence;
tibial
spurs
MALE:
unknown
one on
punctate,
reaching without
transversal
1 bristle
apex with a
proportions:
proximal
12:
3:
9:
located further distally
segment 5 of front
lamellae;
or
fore-wings
vein shorter than
in following
with
without
approximately
the basa I cell s and another
part of radial
segments
(Fig.
rows of nearly 31
tarsus
(Fig.
group of nearly
113) 12
15:
than
with
two
lamellae;
1,1,2.
LOCUS TYP I CUS:
Encruzi I hada
TYPICAL MATERIAL: HOSTS:
distal
fore tarsal
enlarged claw
finely
posterior surface smooth and shiny;
transversa I bands,
(15:23);
smooth,
incomplete,
propodeum reticulate rugose,
with
the pterostigma;
the proximal
notaul ices
191
(Brazi I )
holotype F!
in TW.
unknown
DISTRIBUTION: NOTE:
Neotropic region:
the species
is named
in
BRAZIL:
Encruzilhada
honor of Mr.
(Bahia),
Henry K.
TW!
Townes.
21. Deinodryinus bilobus Fenton
Deinodryinus bilobus Fenton 1927: 11.
FEMALE:
fully
winged;
length 6,87 mm;
clypeus and mandibles testaceous;
black,
with anterior part of frons,
antennae testaceous,
brown;
propectus and pronotum redd i sh-testaceous,
black;
mesothorax,
tegulae testaceous; coxae;
propodeum black;
legs testaceous,
11;
segments head dull,
mid and
in following proportions: hairy,
rugose,
with 3 incomplete longitudinal
tubercles
abdomen brown-redd i sh;
with black spots on mid and
stalks of hind femora black;
antennal 10:
meta thorax,
with segments 7-10
with pronota I
20:
10:
with numerous
median keels;
hind
hind tibiae and tarsi 30:
14:
14:
longitudinal
occipital
brown;
14:
11:
keels;
11:
frons
carina complete;
head without keels around the orbits; POL = 5; OL = 3; OOL = 15; OPL 13; TL= 10; pronotum shiny, smooth, hairy, without sculpture, with an anterior transversal impression; posterior disc with 2 lateral distinct dorsal
lobes;
pronotal
metanotum shiny,
tubercles reaching
smooth,
finely
punctures;
notaulices incomplete,
of scutum;
propodeum reticulate rugose,
smooth,
finely
punctate,
without sculpture; basal
hairy;
vein shorter than proximal 7:
3:
weak,
scutum,
7:
12:
part
26;
located further distally than
scutellum and
without sculpture among
reaching with
approximately 0,65
mesopleura and metapleura shiny, the pterostigma; (15:23);
the proximal
(Fig.
bands,
distal
fore tarsal
enlarged claw
length
posterior surface shiny,
fore-wings with 2 dark transversal
cells and another beneath
proportions:
tegulae;
punctate,
prominence;
the
part of radial
segments
114)
smooth,
one on
with
in following
1 bristle
segment 5 of front
192
Mem.
Fig.
tarsus
(Fig.
114 -
Amer.
En!.
Ins!.,
114) double-S shaped,
Santarem
TYPICAL MATERIAL: HOSTS:
unknown
37,1984
Chel a of lJeinodryinus bilobus Fenton
with 2 rows of nearly
apex with a group of nearly 10 lamellae; MALE: unknown LOCUS TYPICUS:
no.
(Brazil)
holotype F!
in VIA.
tibial
(type)
33 lamellae;
spurs 1,1,2.
Anteoninae
DISTRIBUTION:
Neotropic region:
BRAZIL:
193
Santarem
(Para),
IVA!
22. Deinodryinus insignis n. sp.
FEMALE:
fully
of frons,
winged;
clypeus,
testaceous,
petiole black;
antennae distally
proportions:
15:
8:
rugose and with keels;
ventral
with segments 7-10 brown;
testaceous-reddish; black;
length 4,03-4,62 mm;
mandibles and
two
24:
9:
9:
lateral
keels around
7:
7:
10;
frons with
=
3;
=
OOL
transversal
12;
OPL
=
12;
striae on median
surface; pronotal metanotum shiny, almost reaching
=
TL
10;
region,
in following
head dull,
three median
the orbits;
occipital
fully
reticulate
longitudinal
carina complete;
longitudinal
pronotum shiny,
with
antennae
with hind coxae partly
segments
region behind posterior ocell i with numerous OL
with anterior half
propodeum and abdomen
antennal
7:
irregular keels;
thorax,
legs testaceous,
thickened;
13:
head black,
side testaceous;
two dorsal
keels;
smooth,
lobes on
POL = 3; with
weak
posterior
tubercles reaching tegulae; scutum, scutellum and smooth and without sculpture; notaulices almost complete, the posterior margin of scutum;
propodeum with dorsal
surface reticulate rugose and with posterior surface smooth and without fore-wings with 2 dark
sculpture;
transversal
cells and another beneath
the pterostigma;
longer than
proximal
(24:17);
proportions:
12:
4:
8:
part 12:
27;
bands,
distal
fore tarsal
enlarged claw
one on
segments
(Fig.
the basal
part of radial
vein
in following
115)
with
1 bristle
located further distally than the proximal prominence; segment 5 of front tarsus (Fig. 115) with 2 rows of 26 lamellae; apex with a group of 12 lamellae; MALE:
tibia!
spurs 1,1,2.
unknown
LOCUS TYPICUS:
Sinop
TYPICAL MATERIAL: HOSTS:
(Mato Grosso,
holotype F!
Brazil)
in OT;
1 paratype F!
in AM.
unknown
DISTRIBUTION: BOLIVIA:
Neotropic region:
BRAZIL:
Sinop
(Mato Grosso),
OT!
5 km NIV mouth of Rio Grande (Rio Mamore, Dept. Beni), AM!
23. Deinodr:linus pseudobilobus n. sp.
FEMALE:
fully
winged;
length 4,50-5,62 mm;
head black,
with clypeus and
mandibles testaceous; antennae brown, with segment 1 and partly 2 testaceous; prothorax reddish-testaceous, occasionally black with reddish margins
(in
a
paratype from Horco Molle);
mesothorax,
metathorax and
propodeum black; abdomen black and partly testaceous; legs fully testaceous, occasionally with hind coxae and stalks of hind femora partly black; antennae distally thickened; antennal segments in following :lroportions: 12: 5: 15: 9: 9: 9: 7: 6: 6: 7; head shiny, rugose; frons
194
Mem.
Fig.
115 -
sculptured by strong
Amer.
Chela of
Ent.
no.
37,
1984
Deinodryinus insignis n. sp. (type)
longitudinal
sculpture among keels;
Inst.,
keels;
posterior third of frons without
anterior and median
vertex and region between ocelli
third granulated among keels;
and occipital
carina punctate,
without
keels and without sculpture among punctures; posterior edges of posterior ocelli joint by a transversal furrow; occipital carina complete; POL ~ 5; OL ~ 3,5;
OOL ~ 15;
without sculpture;
OPL ~
TL ~ 10;
13;
pronotum shiny,
sides transversely striate,
on posterior surface;
pronotal
tubercles reaching
scutellum and metanotum shiny, sculpture among punctures;
smooth,
notaul ices,
very
hairy,
dorsally
with 2 prominent dorsal tegulae;
finely punctate,
reaching
lobes
scutum, without
approximately 0,75
length
Anteoninae
of scutum;
propodeum reticulate rugose,
between dorsal longitudinal
Fig.
and posterior surface;
keels,
116 -
with
strong
Chela of
195
without strong
transversal
keels
posterior surface without
transversal
keels,
dull;
fore-wings with 2
Deinodryinus pseudobilobus n. sp. (type)
196
Mem.
dark
transversal
pterostigma; fore
tarsal
claw
(Fig.
prominence; 60
lamellae;
bands,
distal
one on
Ent.
with
Inst.,
the basal
part of radial
segments 116)
Amer.
no.
37,1984
cells and
another beneath the
vein shorter than proximal
in following proportions:
14:
3:
9:
13:
1 bristle arising further distally than
segment 5 of front apex with a
tarsus
(Fig.
group of nearly
24
116)
part
34;
(6:15);
enlarged
the proximal
with some rows of nearly
lamellae;
tibial
spurs 1,1,2.
\
\ \ !
\
II
\ \
d
\ I
I
\1
Fig.
117 -
I J
I
Male genitalia of Deinodryil1us pseudobilobus n. Molle (Argentina):
left half removed.
sp.
from Horco
197
Anteoninae
MALE:
fully
testaceous, brown,
with
th ickened; segments
tarsi
and
trochanters
testaceous;
14: 7:
19:
13:
with frons strongly sculptured by punctate
and
without
carina complete;
scutum shiny, notaulices,
~6;
POL
13:
12:
strongly punctate,
reaching
metanotum shiny,
~
4;
OOL
~
antenna I
10: 9:
punctures;
vertex
to occipital
12; OPL
12;
keels and with
~
12;
carina;
TL
~
11;
without sculpture among punctures;
approximately 0,80
punctate,
10:
longitudinal
sculpture among
OL
legs
antennae not di sta II y
strong oblique keels connecting posterior ocelli
occipital
antennae
1 brown;
an tenna I ha i rs much shorter than breadth of segmen ts;
strongly
without
mandibles testaceous;
sides of segments 2-3-4 and partly
in following proportions:
head shiny, vertex
length 3,87-5,00 mm;
winged;
with dorsal
length of scutum;
scutellum and
without sculpture among punctures;
propodeum
reticulate rugose, without a transversal keel between dorsal and posterior surface; posterior surface reticulate rugose; fore-wings weakly crossed by a transversal dark band beneath the pterostigma; distal part of radial vein shorter than proximal part (9:18); gonoforceps (Fig. 117) with apical branch
wrapping penis;
LOCUS TYPICUS:
TYPICAL MATERIAL: in B;
spurs
1,1,2.
(3 FF,
and 1 paratype F!
holotype F!
2 paratypes FF!
paratypes
tibial
Horco Molle (Argentina)
4 para types
in TU;
4 ,'I,M)!
(3 FF,
in BM;
1 '''1)!
1 paratype F! 7
in TW;
in OL.
HOSTS: unknown DISTRIBUTION: Neotropic region:
BOLIVIA:
Rurenabaque
(Beni),
B~'.!
BRAZIL: Pitfall (Cerrado), OL! Rezende (Estado de Rio), B! Campina Grande (Curitiba), OL! Teodoro Sampaio (Sao Paulo), TW! Serra do Caraca (S. BM!
Barbara, TW!
Minas Gerais),
Las Cejas
OL!
(Tucuman),
ARGaHINA:
OL!
TU!
Horco ,'.,101Ie (Tucuman),
Los Puestos
(Tucuman),
OL!
OL!
24. Deinodryinus rubrolobatus n. sp.
FEMALE:
fully
winged;
length 5,25 mrn;
head
black,
with mandibles and
c I ypeus testaceous; antennae brown, with segment 1 testaceous; prothorax reddi sh; mesothorax, meta thorax and propodeum black; abdomen black, with some regions testaceous; femora, hind tibiae and hind distally
13: 13: keels;
thickened; antenna I segments in following proportions: 17: 8: 24: 12: 9: 8: 7: 9; head shiny, sculptured by numerous longitudinal
region
behind posterior ocell i
temples without keels,
4; OOL
legs testaceous, with hind coxae, hind tarsi partly brown or black; antennae
~12;
OPL ~
punctate;
10; TL
~
8;
with numerous
occipital
pronotum shiny,
without sculpture among punctures;
longitudinal
carina complete; smoooth,
keels;
POL ~ 4;
finely
OL ~
punctate,
posterior surface of pronotum with
two
dorsal transverse lobes; pronotal tubercles reaching tegulae; scutum, scutellum and metanotum shiny, smooth, finely puntate, without sculpture among punctures; of scutum;
notaul ices
incomplete,
propodeum reticulate rugose,
reaching dull,
approximately 0,65
without transversal
or
length
198
Mem.
Fig.
longitudinal rugose;
118 -
keels;
Amer.
Chela of
En!.
Ins!.,
no.
37,
1984
Deinodryinus rubrolobatus n. sp.
posterior surface of propodeum dull,
fore-wings with
two dark
cells and another beneath
transversal
the pterostigma;
bands,
distal
(type)
fully reticulate
one on
the basal
part of radial
vein
Anteoninae
shorter than
proximal
part
proportions:
13: 4:
14:
8:
located further distally tarsus
(Fig.
118)
(12:21); 31;
than
spurs 1,1,2.
LOCUS TYPICUS:
Pompeya
TYPICAL MATERIAL: HOSTS: unknown
tarsal
enlarged claw the proximal
with 2-3 rows of 53
lamellae; tibial MALE: unknown
DISTRIBUTION:
fore
199
segments in following
(Fig.
118)
prominence;
lamellae;
with
1 bristle
segment 5 of front
apex with a
group of
18
(Ecuador)
holotype F!
Neotropic region:
in OT. ECUADOR:
Pompeya
(Napo R.,
Pastaza),
OT!
25. Deinodryinus nigrolobatus n. sp.
FEMALE:
fully
winged;
pronotum fully testaceous,
length 3,75-4,75 mm;
black or with margin
darkened;
legs testaceous,
hind clubs of femora partly antennal 14;
segments
head with
keels;
brown;
black;
testaceous;
occasionally 14:
reticulate rugose,
vertex behind ocell i
with hind coxae and
antennae slightly distally
in following proportions:
frons fully
mandibles testaceous;
antennae brown or
7:
18:
10:
9:
without median
not reticulate rugose,
thickened; 10:
and
9:
9:
strongly punctate,
without sculpture among punctures, without obi ique keels connecting posterior ocell i to occipital carina; occipital carina complete; POL = 4: =3; OOL = 10; OPL = 9; TL = 9 pronotum shiny, with an anterior transversal
furrow;
posterior surface with 2 strong
anterior half strongly
transversely striate;
surface finely punctate, scutum,
not striate;
punctures; propodeum
reaching
reticulate rugose,
lobes;
tubercles reaching
punctate,
posterior surface of pronotum nearly
notaulices incomplete,
transversal
OL
posterior half of the posterior
pronotal
scute II um and metanotum fi nel y
9:
lateral
tegulae;
without scu I pture among
twice as broad as
approximately 0,5
long;
length of scutum;
with posterior surface partly
reticulate rugose
and transversely striate; fore-wings darkened beneath the pterostigma; distal part of radial vein little shorter than proximal part (14:16); fore tarsal (Fig.
segments 119)
with
prominence; lamellae;
in following proportions: 1 bristle
segment 5 of front
apex with a
10:
tarsus
group of 12
(Fig.
or brown;
legs testaceous,
rugose, around ocelli
15:
shiny;
8:
15:
14:
13:
thickened; 13:
frons with 3 median
the orbits;
occipital
10:
than
25;
enlarged claw
the proximal
with 2 rows of nearly 29
segment
black;
3 of fore
tarsi
mandibles and
antennae
with coxae and clubs of femora partly black
antennae not distally
proportions:
6:
119)
lamellae;
produced into a hook; tibial spurs 1,1,2. MALE: fully winged; length 3,75-4,31 mm; testaceous;
3:
located further distally
11:
11:
antennal 9:
12;
longitudinal
carina complete;
segments
head fully
keels;
region
with 2 oblique keels connecting posterior ocelli
2
in following reticulate
lateral
behind
keels
the posterior
to occipital
carina;
200
Mem.
Fig.
POL =7;
119 -
OL = 3,5;
metanotum shiny, antennal complete,
Amer.
Chela of
hairs much
OPL = 6;
punctate,
longer than
almost reaching
Ins!.,
no.
37,
1984
Deinodryinus nigrolobatus n. sp. (type)
OOL = 11; smooth,
En!.
TL = 10;
scutum,
scutellum and
without sculpture among punctures;
breadth of segments;
notaul ices almost
the posterior margin of scutum;
propodeum fully
Anteoninae
201
/
Fig.
120 -
Male genitalia of Teutonia
reticulate rugose, wings hyaline,
without
subapical
proximal
holotype F!
in TU;
2 paratypes MM!
HOSTS:
unknown
in
TW;
DISTRIBUTION:
Neotropic region:
:'Jova Teutonia
(S.ta Catarina),
(Tucuman),
TU!
TW!
las Cejas
bands;
(13:14);
part
Nova Teutonia
TYPICAL MATERIAL:
keels on
transversal
branch wrapping penis;
lOCUS TYP ICUS:
right half removed.
longitudinal
without dark
little shorter than
Deinodryinus nigroiobatus n. sp. from Nova
(Brazi I):
tibial
posterior surface; distal
gonoforceps
fore-
part of radial (Fig.
120)
with
vein a
spurs 1,1,2.
(Brazi I) and 1 para type M! 2 paratypes
PERU: BM!
Ol!
(1
in BM;
F,
M)!
Machu Picchu, ARGENTINA:
(Tucuman),
TW!
1 paratype F!
in Ol. Ol'
BRAZil:
Horco Molle
Ol!
26. Deinodryinus vagans n. sp.
202
Mem.
FEMALE:
fully
winged;
mandibles and clypeus partly 3 testaceous; mesothorax,
Amer.
Ent.
Inst.,
length 4,18-6,00 mm; testaceous;
no.
prothorax reddish-testaceous,
antennae distally
following proportions:
12:
with frons sculptured by region behind ocelli
8:
19:
thickened; 16:
13:
occipital OL
=
=
OOL
puncta te, broad as reaching punctate,
OPL
impression;
long,
with
tegulae;
9;
TL
=
9;
du II;
occipital
two strong
or
transverse dorsal
length of scutum;
longitudinal
vein shorter than proximal
-
carina complete; hairy,
lobes;
region
keels;
two dark
notaulices
part
Chela of
to POL
=
8;
finely strong
twice as
pronotal
scutellum and metanotum shiny,
fore-w i ngs with
121
areolae;
pronotum with a
tubercles
smooth,
finely
incomplete,
propodeum reticulate rugose,
posterior surface fully
transversa I bands,
cells and another beneath the pterostigma;
Fig.
in
head shiny,
keels and with
posterior surface of pronotum nearly
approximately 0,65
rugose,
10;
resembling
pronotum shiny,
without sculpture among punctures;
transversal
basal
line complete;
=
scutum,
without
10:
two obi ique keels connecting posterior ocell i
frontal
13;
10:
longitudinal
wi thout scu I pture among punctures;
transversal
reaching
without
carina;
4;
testaceous-brown; segments
punctate and without sculpture among punctures;
occasionally punctures very strong and broad, behind ocell i
2 and
occasionally darkened;
abdomen 10:
with malar space,
with segments 1,
antennal
13:
nUmerous parallel
1984
head black,
antennae black,
meta thorax and propodeum bl ack;
legs testaceous;
37,
(18:22 or 18:24);
distal
one on
reticulate the
part of radial
fore tarsal
segments
Deinodryinus vagans n. sp. (type)
in
Anteoninae
following proportions: bristle
fore tarsi
(Fig.
of nearly MALE:
12:
located further 15
fully
121)
winged; ventral
9:
20:
37;
than
enlarged claw
the proximal
with 2 rows of nearly
lamellae;
antennae with
3:
distally tibial
203
31
(Fig.
121)
prominence;
lamellae;
with
1
segment 5 of
apex with
a
group
spurs 1,1,2.
length 3,43 mm;
black;
side testaceous,
mandibles partly
dorsal
side brown;
testaceous;
legs testaceous,
with femora, hind coxae, mid and hind tibiae partly brown; antennae not distally thickened, with hairs shorter than breadth of segments; antennal segments
in following proportions:
head shiny,
and with region behind ocelli posterior ocelli
to occipital
sculpture among punctures; complete;
POL =7;
Ol = 4;
and metanotum shiny, punctures;
Fig.
122 -
13:
7:
16:
10:
10:
10:
with frons sculptured by numerous parallel
notaul ices
without
carina; frontal
region behind ocelli OPl = 8:
punctate,
incomplete,
9:
12; keels
Tl = 6;
without
carina
scutum,
scutellum
without sculpture among
almost reaching
gonoforceps
punctate,
occipital
the posterior margin of
Male genitalia of Deinodryinus vagans n. (Brazil)(A);
10:
two oblique keels connecting
I ine complete;
OOl = 10:
smooth,
10:
longitudinal
sp.
from
(laterally viewed)(B).
Teresopolis
204
Mem.
scutum;
Ent.
propodeum reticulate rugose,
posteriorsurface; distal
Amer.
vein
no.
with small
fore-wings hyal ine,
part of radial
Inst.,
37,
1984
smooth area on
without dark transversal
shorter than
proximal
part
(12:
(Fig. 122) with small apical branch \Vrapping penis; LOCUS TYPICUS: Teresopolis (Brazil)
tibial
TYPICAL MATERIAL: holotype F! and 4 paratypes FF! in AM; 4 paratypes (3 FF, 1 M)! in OL. HOSTS: unknown
in TW;
DISTRIBUTION:
Neotropic region:
(Romansos,
Dept.
Teresopolis,
OL!
Jatai
Serra do Caraca ARGENTINA:
Beni), (S.
AM!
(Goias),
Barbara,
Horco Molle
BOLIVIA:
Rio
Ichilo
OU
S.
Junction Rio
(Santa Cruz), J.
Barreiro
Minas Gerais),
(Tucuman),
TW!
bands;
14);
gonoforceps
spurs 1,1,2. paratype F!
Itenez-Rio Paraguay OL!
(Serra
BRAZIL: da Bocaina),
Caruaru,
OL!
TW!
TW!
27. Deinodryinus pilosifrons Fen ton
Deinodryinus pilosifrons Fenton 1927: 11.
FEMALE: frons,
fully
winged;
length 5,0 mm;
head black,
clypeus and mandibles testaceous;
segment weak I y
dark;
with anterior part of
antennae testaceous,
propectus and pronotum redd i sh;
with
last
mesothorax,
meta thorax and propodeum black; abdomen reddish; legs testaceous, with mid and hind clubs of femora brown; tegulae testaceous; antennal segments in following proportions: reticulate rugose; connecting 10;
~
TL
18:
anterior ocellus
10;
8:
23:
17:
frons with 3 median
occipital
punctate and rugose,
to clypeus;
14:
with
13:
POL
~
4;
OL
~
12:
14;
OOL
hairy,
impression;
tubercles reaching
head fully
irregular keels
3;
pronotum dull,
an anterior transversal pronotal
13:
longitudinal
carina complete;
disc centrally excavated;
14:
and
tegulae;
~
10;
OPL
strongly posterior scutum
shiny, smooth, finely punctate, without sculpture among punctures; notaulices not fully visible because hidden by ,he pin (in the only known specimen); scutellum and metanotum shiny, smooth, finely punctate, without sculpture among punctures; propodeum reticulate rugose; posterior surface not smooth, reticulate rugose, without longitudinal keels; fore-wings with two dark transversa I bands, one on the basa I cell s and another beneath the pterostigma; radial cell: fig. 147 B; distal part of radial vein longer than proximal part (21:15); fore tarsal segments in following proportions: 12: 4:
7:
distally
14: than
26;
enlarged claw
the proximal
double-S shaped,
123)
with one bristle
tibial
Semacoch
TYPICAL MATERIAL:
spurs 1,1,2.
(Gua tema I a)
holotype F!
in WA.
located further
segment 5 of fore tarsi
with some rows of nearly 40
of nearly 4 lamellae; MALE: unknown LOCUS TYP I CUS:
(Fig.
prominence;
lamellae;
(Fig.
apex with a
123) group
Anteoninae
Fig.
HOSTS:
123 -
205
Chela of Deinodryinus pilosifrons Fenton
(type)
unknown
DISTRIBUTION:
Neotropic region:
GUATEMALA:
Semacoch
(Alta Vera,
Paz),
206
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
WA!
28. Deinodryinus politus n. sp.
FEMALE: brown
fully
pronotum margin a
winged;
and ventral reddish,
brown;
reddish
10: 9:
with hind coxae and
9: 9: 8:
9:
weak short complete; TL = 6;
with sides partly brown; abdomen
head dull,
irregular striae,
pronotum ha i ry,
mainly
du II,
very
antennae
strong
tegulae;
14:
weakly rugose,
near the antennal POL = 4;
OL = 3;
sockets; OOL = 8;
transversa I
with
legs
in following proportions:
with a
tubercles reaching
testaceous-brown;
granulated,
occipital carina complete;
with pronotal
with anterior
mesopleura black,
hind clubs of femora brown;
antenna I segments 11;
with ocellar region
propectus black;
scutum reddish,
propodeum black;
metapleura black;
thickened;
head reddish,
antennae testaceous;
scutellum reddish;
spot;
testaceous, distally
length 4,87 mm;
side black;
8:
18:
with
frontal
line
OPL = 9;
impression and
posterior surface of pronotum
nearly as long as broad, flat, strongly transversely striate, rugose; scutum dull, granulated; notaul ices incomplete, almost reaching the posterior margin of scutum; granulated;
propodeum dull,
longitudinal
keels;
scutellum and metanotum shiny, reticulate rugose,
very
weakly
without transversal
posterior surface of propodeum fully
or
reticulate rugose,
dull; fore-wings with two dark transversal bands, one on the basal cells and another beneath the pterostigma; distal part of radial vein little shorter than proximal part (12: 14); fore tarsal segments in following proportions: 12: 3: 6: 14: 25; enlarged claw (Fig. 124) with 1 bristle located further distally tarsi
(Fig.
nearly 6 MALE:
124)
lamellae;
the proximal
tibial
prominence; lamellae;
segment 5 of fore
apex with a
group of
spurs 1,1,2.
unknown
LOCUS TYPICUS:
Serra do Caraca
TYPICAL MATERIAL: HOSTS:
than
with 2 rows of nearly 50
holotype F!
(Brazil) in TW.
unknown
DISTRIBUTION: Neotropic region: Minas Gerais), TW!
BRAZIL:
Serra do Caraca
(S.
Barbara,
29. Deinodryinus c1aripes n. sp.
FEMALE: pronotum,
fully
winged;
length 4,38 mm;
sides of pronotum,
scutum,
testaceous,
propodeum,
with collar of
partly
metapleura,
dorsal
side of abdomen black; metanotum brown; antennae not or slightly distally thickened; antennal segments in following proportions: 9: 5: 15: 13: 11: 10: 9: 10: 9: and with two
11; head shiny, granulated; frons with a weak frontal line lateral longitudinal keels; occipital carina complete; POL =
207
Anteoninae
Fig.
124 -
Chela of Deinodryinus politus n.
sp.
(type)
208
3,5;
Mem.
OL = 4,5;
OOL= 11;
anterior surface rugose; numerous short
Amer.
Ent.
OPL = 10;
TL = 8;
lobes;
scutum;
37,
1984
pronotum partly
hairy,
pronotal notaul ices
as
tubercles reaching long
incomplete,
long
(26:22);
almost
scutellum and metanotum shiny,
as broad,
tegulae;
flat,
scutum shiny,
reaching smooth,
without
posterior surface of weakly
the posterior margin of
finely punctate,
sculpture among punctures; propodeum dull, reticulate rugose, longitudinal keels on anterior surface and without transversal
Fig.
with
granulated and with
long i tud i na I keel s near the posterior marg i n of pronotum;
pronotum slightly broader than granulated;
no.
posterior surface smooth,
posterior surface of pronotum nearly dorsal
Inst.,
125 -
Chela of
Deinodryinus c1aripes n. sp.
without
with some or
(type)
Anteoninae
longitudinal fully
beneath
posterior surface; dull;
the pterostigma;
proximal 85:
keels on
reticulate rugose, part;
117;
fore
tarsal
enlarged claw
posterior surface of propodeum
fore-wings
distal
with one dark
part of radial
segments
(Fig.
209
125)
vein
transversal
about as
in following proportions:
with
1 bristle
band
long as 45:
9:
21:
located further distally
than the proximal prominence; segment 5 of fore tarsi (Fig. 125) with two rows of nearly 29 + 21 lamellae; apex with a group of nearly 8 lamellae (one of these very MALE: unkonwn LOCUS TYPICUS:
long);
Essequibo
TYPICAL Mf\TERIAI: HOSTS:
tibial
spurs
1,1,2.
(Guyana)
holotype F!
in CO.
unknown
DISTRIBUTION:
Neotropic region:
GUYANA:
Essequibo,
CO!
30. Deinodryinus chiapasi n. sp.
FEMALE:
fully
of frons,
head black,
with anterior region
clypeus and mandibles reddish-testaceous;
winged;
length 5,62 mm;
antennae testaceous,
with segments 7-10 brown; pronotum and posterior testaceous, black;
thorax and propodeum black,
lobes of pronotum testaceous;
with hind coxae,
hind clubs of femora and hind
antennae slightly distally
proportions:
15:
rugose;
frons
median
keels;
9:
21:
with
two
occipital
14:
12:
lateral
~
with
4;
OOL~
14;
OPL~
irregular keels,
with pronotal shiny,
12;
with
punctate,
reaching
reticulate rugose,
area rugose;
10:
10:
keels around
TL
~
15;
tegulae;
located further distally
fore tarsi
(Fig. to
MALE:
unknown
LOCUS
TYP ICUS:
126)
tibial
Yerba Buena
TYPICAL MATERIAL:
OT!
POL
as broad as
long and
scutellum and metanotum notaulices
propodeum dull, median
bands, one on the basal
fore tarsal
segments (Fig.
prominence;
lamellae
in
126)
with one
segment 5 of
(extending
spurs 1,1,2.
in OT. MEXICO:
5;
reticulate rugose and
enlarged claw
the proximal
with ~
(Mexico)
holotype F!
HOSTS: unknown ::JISTRIBUTION: Neotropic region: Chiapas),
than
32;
with 2 rows of nearly 38
the apex);
carina;
keels on posterior surface;
the pterostigma;
continuously
to occipital
length of scutum;
4:
15:
two
behind posterior ocelli
scutum,
following proportions:
7:
in following reticulate
the orbits and with
fore-wings with 2 dark transversal 12:
segments
without sculpture among punctures; longitudinal
legs
tibiae partly
head shiny,
posterior surface nearly
cells and another beneath bristle
10;
pronotum dull,
approximately 0,75
with 2
antennal 8:
region
posterior ocell i
tubercles reaching
smooth,
incomplete,
thickened;
11:
carina complete;
two obi ique keels connecting OL
with sides of
abdomen bl ack;
Yerba Buena
(20 mi.
N Bochil,
210
Mem. Amer.
Fig.
126 -
En!.
Ins!.,
no. 37,
1984
Chela of Deinodryinus chiapasi n. sp.
(type)
Anteoninae
211
31. Deinodryinus nigricans (Cameron) n. sp.
Dryinus nigricans Cameron 1888: 446. Plastodryinus nigricans Cameron): Kieffer 1914b: 43.
FEMALE: fully winged; length 4,5 mm; head black, with clypeus and mand i b les testaceous (wi th teeth brown); antennae testaceous-brown, wi th segments 7-10 brown;
thorax,
propodeum,
of abdomen
light;
testaceous;
antenna I segments
8:
7:
8:
7:
longitudinal
10;
head dull,
keels,
with 3 median POL = 3; a
weak
legs brown,
hairy;
pronotal
OPL = 11;
tubercles reaching
scutum;
with
tegulae;
incomplete,
numerous
reticulate rugose, surface;
without a
around
fore-wings
smooth,
keel
transversal
12:
9:
to clypeus; hairy,
with
keels;
granulated;
approximately 0,75
granulated;
transversal
with 2 dark
22:
the orbits and
irregular short
length of
propodeum shiny,
between dorsal
posterior surface with 2 irregular longitudinal
smooth;
9:
pronotum dull,
scutum dull,
reaching
scutellum and metanotum dull,
14:
apex
tegulae
keels connecting ocell i
TL = 12;
with
black;
light;
numerous irregular
strong keel
and parallel
impression,
notaulices distinct,
following proportions:
frons with a
OOL= 12;
transversal
in
pet iol e and abdomen
last segments of tarsi
granulated,
longitudinal
OL = 3;
with
bands,
and
keels;
one on
posterior
median
the basal
area
cells
and another beneath the pterost i gma; d i sta I part of rad i a I vei n much longer than proximal part (20:12); radial cell very wide (Fig. 147); pterostigma (9:
15);
long and
narrow;
segment 5 of fore
lamellae;
apex with
127)
1 peg-I ike bristle
with
prominence; MALE:
fore tarsal
tarsi
(Fig.
a group of at
segment
127)
least
16
lamellae;
located further
segment 3 of fore
tarsi
1 shorter than
segment 4
with some rows of nearly 44 distally
produced
enlarged claw than
into hook;
(Fig.
the proximal tibial
spurs 1,1,2.
unknown
LOCUS TYP ICUS: Volcan de Chiriqui (Panama) TYPICAL MATERIAL: holotype F! in BI.i. HOSTS:
unknown
DISTRIBUTIOI'j:
Neotropic region:
PANAMA:
Volcan
de Chiriqui
BM!
32. Deinodryinus actuosus n. sp.
FEMALE:
fully
winged;
length 4,37-4,68 mm;
less wide testaceous frontal occasionally
two reddish
area;
spots on
head black,
with a
more or
clypeus and mandibles testaceous; temples;
antennae
testaceous,
with
segments 7-10 brown; prothorax reddish-testaceous; scutum reddishtestaceous, usually with region near the anterior margin darkened; sometimes almost fully
darkened;
propodeum redd i sh-testaceous,
scutellum,
metanotum,
with furrows between
pleura and
scutum and scutell um,
212
Mem.
Fig.
127 -
Chela of
scutell um and metanotum, brown-reddish; brown, distally 13:
10:
hind
Ent.
Inst.,
propodeum and metanotum
9:
antennal 8:
10;
37,
1984
darkened;
abdomen
with clubs of hind and mid femora partly
tibiae and sometimes also mid 9:
no.
Deinodryinus nigricans (Cameron)(type)
legs testaceous,
thickened; 10:
Amer.
segments
head dull,
tibiae brown;
antennae slightly
in following proportions:
fully
reticulate rugose;
13:
frons
6: with
18: two
Anteoninae
lateral region ocelli
keels around
the orbits and with
behind posterior ocell i to occipital
OOl =12;
carina;
OPl = 10;
tracks of three median keels;
with 2 obi ique keels connecting
occipital
Tl = 13;
213
carina complete;
pronotum dull,
posterior
POL = 3,5;
rugose,
Ol =3;
with numerous
transversal keels on posterior surface; posterior surface nearly as broad as long; pronotal tubercles reaching tegulae; scutum, shiny, smooth, finely
punctate,
without sculpture among punctures;
reaching approximately 0,65 length of scutum; shiny,
smooth,
without sculpture;
posterior surface rugose, dark transversal pterostigma;
bands,
distal
without one on
the basal
part of radial
vein
keels;
with
two the
fore-wings
longer than proximal
claw
128)
13: 3:
8:
segment 5 of fore tarsi
apex with a
group of 12 lamellae;
(Fig. tibial
128)
(20: 16);
enlarged
the proximal
with 2 rows of 26 lamellae;
spurs 1,1,2.
MALE: unknown lOCUS TYPICUS: San Cristobal de las Casas (Mexico) TYPICAL MATERIAL: holotype F! and 1 paratype F! in OT;
128 -
part
13: 30;
with one bristle located further distally than
prominence;
with
cells and another beneath
segments in following proportions:
Fig.
incomplete,
propodeum reticulate rugose, longitudinal
fore tarsal (Fig.
notaul ices
scutellum and metanotum
Chela of Deinodryinus actuosus n.
sp.
2 paratypes FF!
(type)
214
Mem.
Amer.
En!.
Inst.,
no.
1984
37,
in OL. HOSTS:
unknown
DISTRIBUTION: (Chiapas),
Neotropic region:
OL!
MEXICO:
San Cristobal
de
las Casas
OT!
33. Deinodr:linu5 incaicU5 n. sp.
FE,'-'lALE:
fully
winged;
length 4,37-4,81
sometimes with anterior half of frons testaceous;
antennae fully
7-10 brown;
legs fully
mm;
head
testaceous;
testaceous;
testaceous or black, clypeus and mandibles
occasionally
testaceous or with coxae,
with segments 6-10 or
mid and hind
and hind clubs of femora black;
occasionally only mid and
partly
and abdomen
black;
thorax,
propodeum
of pronotum testaceous;
antennae distally
black;
thickened;
tibiae,
occasionally margins antenna I segments
following proportions: 14: 7: 21: 11: 9: 10: 8: 8: 7: 10; head shiny; in front of anterior ocellus with a central region sculptured by longitudinal regions on finely
keels;
frons
without
lateral
the sides of the ocell i
punctate,
temples smooth;
keels;
and behind
surface smooth, dorsal
flat, lobes,
with
finely
tegulae;
areolae smaller, bands,
pterostigma; fore tarsal claw
with
without sculpture among punctures,
scutellum
rugose;
surface with longitudinal
the basal
areolae wide;
keels;
propodeum
posterior surface
fore-wings with 2 dark
longer than proximal
in following proportions: 1 bristle
notaulices
cells and another beneath
vein
pronotal
and metanotum shiny,
the posterior margin of scutum;
part of radial
segments
(Fig - ,29)
without
one on
distal
11; OPl = 10; TL = 9; anterior furrow; posterior
without sculpture among punctures; dorsal
11:
2:
6:
located further distally
20:
the
part 33;
than
(21:17);
enlarged
the proximal
prominence; segment 5 of fore tarsi (Fig. 129) with 2 rows of nearly lamellae; apex with a group of nearly 5 lamellae (one of these very long);
tibial
MALE:
fully
52
spurs 1,1,2. winged;
legs testaceous;
length 3,37 mm;
black
antennae not distally
following proportions: much
to
=
anterior surface weakly
scutum,
almost reaching
reticulate rugose;
transversal
punctate,
occasionally
finely punctate,
almost complete, dull,
carina complete;
posterior ocell i smooth,
without sculpture among punctures or weakly granulated;
tubercles reaching smooth,
occipital
in frons
vertex xithout oblique keels connecting posterior ocelli
occipital carina; POL = 2,5; Ol = 3; OOl pronotum shiny, with a strong transversal without
mid
hind coxae
14:
6:
15:
9:
9:
longer than breadth of segments;
reticulate rugose;
frontal
sides of the ocelli
I ine present;
without sculpture among punctures; carina complete;
10:
POL
=
Ol
=
3;
11:
10:
antennae,
segments in
11;
antennal
without keels,
and on
to occipital
=
7;
OPl
carina;
=
7;
the
punctate and
region behind ocell i without OOl
hairs
wi th frons fu II y
regions of head behind
posterior ocelli 5,5;
10:
clypeus,
antennal
head sh i ny,
not reticulate rugose,
oblique keels connecting
mandibles,
thickened;
Tl
two
occipital
=
5;
scutum,
Anteoninae
/
\
Fig.
129 -
215
r.! .1
~/
Chela of Deinodryinus incaicus n.
sp.
(type)
216
Mem.
Amer.
scutellum and metanotum shiny, among punctures;
internal
proximal
branch;
3 paratypes
para type M! HOSTS:
in AM;
without sculpture
almost reaching
the posterior
without transversal
part
tibial
(2 FF,
bands;
(12:14);
distal
gonoforceps
or
fore-
part of radial (Fig.
130)
vein
with
spurs 1,1,2.
1 M)!
1 para type F!
in TW;
2 paratypes FF!
in BM;
2 paratypes FF!
1 paratype F! in OT;
1
in OL.
unknown
DISTRIBUTION:
Fig.
1984
with posterior surface fully reticulate rugose;
LOCUS TYP ICUS: Mitu (Colombia) TYPICAL MATERIAL: holotype F! and in LE;
37,
finely punctate,
without dark transversal
little shorter than broad
smooth,
no.
propodeum reticulate rugose,
keels,
wings hyaline,
Ins!.,
notaul ices almost complete,
margin of scutum; longitudinal
Ent.
130 -
Neotropic region:
PANAMA:
Gamboa
(Pipeline Road,
Male genitalia of Deinodryinus incaicus n. (Brazil):
right
half removed
sp.
from
Canal
Vila Vera
217
Anteoninae
Zone),
LE!
Catarina), IV
(Sinop,
COLOMBIA:
Mitu
IV 50° 30'
BM!
Mato Grosso),
Tucuman,
OT!
Jujuy,
(Vaupes),
S 12° 30' TIV!
TIV!
BM!
BRAZIL:
Rio Guapore
Horco Molle
Nova Teutonia
(Sta
OL!
TIV!
12° 31' S 55° 37'
(Rondonia),
AM!
ARGENT INA:
(Vi la Vera), (Tucuman),
OT!
OL!
34. Deinodryinus bolivianus n. sp.
FEMALE:
fully
length 5,0 mm;
winged;
anterior region of frons
testaceous;
black;
10 black; abdomen and legs fully testaceous; thickened;
antennal
7: 7: 7: 7: 9; keels;
segments
head dull,
in
mandibles,
antennae slightly
sculptured by
occipital carina complete;
numerous
region behind ocell i
distally
15: 8: 20:
following proportions:
hairy,
clypeus and wi th segments 9-
antennae testaceous,
13:
10:
longitudinal
with 2 obi ique keels
connecting posterior ocelli to occipital 13: OPl ; 9; Tl ; 9; pronotum sh i ny,
carina; POL; 3; Ol ; 2,5; OOl ; crossed by a strong transversa I
furrow;
finely
posterior surface smooth,
among punctures;
pronotal
and metanotum shiny, punctures; scutum;
flat,
punctate,
tubercles reaching
smooth,
tegulae;
finely punctate,
notaulices incomplete,
reaching
propodeum reticulate rugose;
approximately 0,75
dorsal
transversa I bands,
scutellum
without sculpture among
surface with
posterior surface with areolae smaller and without fore-w i ngs with 2 dark
without sculpture scutum,
one on
length of
areolae wide;
longitudinal
keels;
the basa I cell sand
another beneath the pterostigma; distal part of radial vein longer than proximal part (20: 14); fore tarsal segments in fOllowing proportions: 12:
3: 8:
13: 32; enlarged claw (Fig.
131) with 1 bristle located further segment 5 of fore tarsi (Fig. 131) with 2 rows of nearly 32 lamellae; apex with a group of nearly 27 lamellae; segment 3 of fore tarsi produced into a hook; tibial spurs 1,1,2.
distally
MALE:
than
the proximal
unknown
LOCUS TYPICUS:
Rurenabaque
TYPICAL MATERIAL: HOSTS:
prominence;
(Bolivia)
holotype F!
in
BM.
unknown
DISTRIBUTION:
Neotropic region:
BOLIVIA:
Rurenabaque
(Beni),
8M!
35. Deinodryinus eminens n. sp.
FEMALE:
fully
winged;
surface of frons,
prothorax redd i sh; brown-black;
length 4,56-4,81
mm;
head black,
clypeus and mandibles testaceous; mesothorax,
legs testaceous;
metathorax and
with anterior
antennae testaceous,
propodeum black;
antennae slightly distally
abdomen
thickened;
segments in following proportions: 15: 8: 20: 18: 14: 13: 10: 13; head dull, fully reticulate rugose; frons with 3 weak median
antennal
10:
keels;
2
lateral
keels around the orbits;
occipital
carina complete;
10:
region
218
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
r
/
Fig.
131
-
Chela of Deinodryinus bolivianus n.
sp.
(type)
Anteoninae
219
/
/
/
II / 'Y~
,
If~. l'
/
Fig.
132 -
Chela of Deinodryinus eminens n.
of head behind posterior ocelli ocelli
to occipital
carina;
sp.
(type)
with 2 oblique keels connecting posterior
POL = 5; Ol = 4; OOl = 13; OPl = 12; Tl = 12;
220
Mem.
pronotum shiny,
hairy,
Amer.
En!.
Ins!.,
no.
37,
1984
with a strong anterior transversal
furrow;
posterior surface of pronotum nearly as long as broad,
finely
without
lobes parallel
sculpture among punctures,
the sides of pronotum;
pronotal
with 2 weak dorsal
tubercles reaching
scutellum and metanotum shiny,
smooth,
tegulae;
very finely
punctate, to
scutum,
punctate,
without
sculpture among punctures; notaul ices almost complete, almost reaching the posterior margin of scutum; propodeum reticulate rugose, with posterior surface dull,
reticulate rugose;
one on
the basal
radial
vein
little shorter than proximal
in following proportions: bristle
12:
3:
7:
fore tarsi
(Fig.
132)
the pterostigma;
part 30;
(13:14);
the proximal
tibial
(Fig.
part of
segments
132)
with 1
segment 5 of
prominence;
lamellae;
bands,
distal
fore tarsal
enlarged claw
with 2 rows of nearly 26
15 lamellae;
apex with a group
spurs 1,1,2.
unknown
LOCUS TYPICUS:
Nova Teutonia
TYPICAL MATERIAL:
(Brazil)
holotype F!
HOSTS: unknown DISTRIBUTION: Neotropic region: BM!
12:
located further distally than
of nearly MALE:
fore-wings with 2 dark transversal
cells and another beneath
Sta Barbara
in BM; BRAZIL:
(Minas Gerais),
paratype F!
in TW.
Nova Teutonia
(Sta Catarina),
TW!
36. Deinodryinus trinidadi n. sp.
FEMALE: fully winged; length 3,75-4,75 mm; head testaceous or reddishtestaceous, with ocellar region brown or dark; antennae testaceOU5, with segments 8-10 or 9-10 dark; legs testaceous;
thorax and propodeum black;
occasionally extremities of hind tibiae brown; mid tibiae and hind coxae partly brown; thickened; 8:
8:
8:
lateral
antennal
11;
abdomen brown;
stalk of hind femora and extremities of hind tibiae brown;
segments
head shiny,
keels around
reticulate rugose;
in following proportions:
reticulate rugose;
the orbits;
region
occasionally extremities of
antennae slightly distally 15: 7:
21:
13:
9:
frons with 3 median keels;
region behind ocelli
behind posterior ocell i
8: 2
weakly or strongly
with 2 obi ique keels
connecting posterior ocelli to occipital carina; occipital carina complete; POL = 3; OL = 3; OOL = 12; OPL = 12; TL = 12; pronotum shiny, hairy, with a strong finely dorsal
anterior transversal
punctate,
lobes parallel
tegulae;
scutum,
punctate, reaching
furrow;
to the sides of pronotum;
the posterior margin of scutum
one on
part of radial
the basa I cells and vein slightly
with 2 weak
tubercles reaching
smooth,
very finely
notaul ices incomplete,
(Fig.
157 A);
almost
propodeum with
fore-wings with 2 dark transversal
another beneath
longer than proximal
segments in following proportions:
smooth,
pronotal
scutellum and metanotum shiny,
without sculpture among punctures;
posterior surface almost fully rugose; bands,
posterior surface of pronotum
without sculpture among punctures,
12: 4:
9:
15:
the pterost i gma;
part 35;
(17:15);
fore
enlarged claw
di sta I tarsal (Fig.
Anteoninae
221
133) with 1 bristle located further distally than the proximal prominence; segment 5 of fore tarsi (Fig. 133) with 2 rows of nearly 34 lamellae; apex with a MALE:
group of nearly 15 lamellae; fully winged;
Fig.
133 -
tibial
length 2,60-3,56 mm;
Chela of
spurs 1,1,2. black;
mandibles and antennae
Deinodryinus trinidadi n. sp. (type)
222
Mem.
Amer.
with
Ent.
Inst.,
testaceous;
legs testaceous,
hairs much
longer than breadth of segments;
37, 1984
no.
hind coxae proximally black; antennae not
antennal
distally
antennal segments in following proportions: 15: 7: 15: 10: 12: 12: 12: 12: 12: 15; head shiny, fully reticulate rugose; frons with 3
thickened; median
keels;
region behind ocelli
2
lateral
to occipital
scutum,
keels around
posterior ocelli carina;
the orbits;
occipital
POL
=
4;
=
Ol
scutellum and metanotum shiny,
3;
OOl
smooth,
=
5;
finely
OPl
notaul ices almost complete,
posterior margin of scutum;
propodeum reticulate rugose,
surface with 2
complete keels;
longitudinal dull,
rugose;
bands or weakly darkened;
distal
part of radial
8;
Tl
=
7;
without
almost reaching
the
with posterior
area with a
without dark
transversal
transversal
vein nearly as
long as
(14:14); gonoforceps (Fig. 134) with apical branch wrapping tibial spurs 1,1,2.
proximal penis;
median
fore-wings hyal ine,
=
punctate,
sculpture among punctures;
anterior keel,
carina complete;
with 2 oblique keels connecting posterior
part
lOCUS TYPICU5:
5t.Augustine
TYPICAL MATERIAL:
Fig.
(Trinidad)
holotype F!
and 4 paratypes MM!
in 8M;
1 paratype
134 - Male genital ia of Deinodryinus trinidadi n. sp. from 51. Augustine
(Trinidad):
right
half removed.
Anteoninae
F!
in OT;
HOSTS:
2 paratypes MM!
in OL.
unknown
DISTRIBUTION: BM!
223
Simla
Neotropic region:
(Arima),
TRINIDAD:
St.
Augustine
(SI.
George),
OL!
OT!
37. Deinodryinus rapax n. sp.
FEMALE:
fully
length 6,56 mm;
winged;
anterior region of frons
testaceous;
black;
mandibles,
antennae testaceous,
clypeus and with segments 7-
10 brown; fore legs testaceous; mid and hind legs brown; antennae thickened; antenna I segments in following proportions: 19: 9: 29: 22: 20: 19: 17: 17: 16: 18; head shiny, fully reticulate rugose; frons distally
with 3 median keels;
2 lateral
keels around
the orbits;
region of head
behind posterior ocelli reticulate rugose and with 2 oblique keels connecting posterior ocell i to occipital carina; occipital carina complete; POL; 5; OL ; 2,5; OOL ; 12; OPL ; 13; TL ; 15; pronotum hairy, dull, crossed by
an
anterior strong transversal
pronotum smooth,
strong I y
punctate,
posterior surface of pronotum dorsal
weak parallel
lobes;
less than
pronotal
scutellum and metanotum shiny, among punctures; of scutum;
posterior surface of
twice as broad as
long,
tubercles reaching tegulae;
smooth,
finely
punctate,
with 2 scutum,
without sculpture
reaching approximately 0,80
notaul ices incomplete,
propodeum dull,
furrow;
wi thout scu I pture among punctures;
reticulate rugose;
length
posterior surface dull,
reticulate rugose, without longitudinal keels; fore-wings with 2 dark transversal bands, one on the basal cells and another beneath the pterostigma; fore tarsal
distal
part of radial
vein
longer than
segments in following proportions:
part (30:25); 19: 39; enlarged
proximal
14: 5:
10:
135) with one bristle located further distally than the proximal segment 5 of fore tarsi (Fig. 135) strongly S-curved, with some rows of at least 60 lamellae (extending continuously to the apex); tibial spurs 1,1,2.
claw
(Fig.
prominence;
MALE: unknown LOCUS TYP ICUS:
Chiriqui
TYPICAL MATERIAL: HOSTS:
(Panama)
holotype F!
in LE.
unknown
DISTRIBUTION: Panama),
Neotropic region:
PANAMA:
Chiriqui
(Fortuna,
North
LE!
38. Deinodryinus croceus n. sp.
FEMALE:
fully
winged;
length 5 mm;
reddish-testaceous,
with abdomen
black and antennal segments 9-10 brown; antennae distally thickened; antennal segments in following proportions: 14: 8: 17: 13: 9: 9: 8: 8:
8:
224
Mem. Amer. En!.
Ins!., no. 37,
1984
/
Fig.
135 -
Chel a of Deinodl'yinus l'apax n. sp.
(type)
Anteoninae
Fig.
136 -
Chela of Deinodryinus croceus n.
225
sp.
(type)
226
10;
Mem.
head shiny,
longitudinal
Amer.
with frons
keels;
and
occipital
En!.
Inst.,
37,
1984
vertex reticulate rugose and with some
carina complete;
oblique keels connecting posterior ocelli without
no.
sculpture among punctures,
region behind ocell i with
to occipital
carina,
two
punctate and
not reticulate rugose or keeled;
POL
4,5; OL = 3,5; OOL = 13; OPL = 12; TL = 13; pronotum shiny, with a strong transversal impression and with pronotal tubercles reaching tegulae; smooth,
posterior surface of pronotum nearly as with
metanotum
almost
shiny,
reaching
rugose, dull
two weak dorsal
dull,
smooth,
without
vein
sculpture;
without transversal
the basal
radial
lobes;
long as broad,
scutum,
or
with
shorter than
keels;
two dark
cells and another beneath
slightly
propodeum reticulate
longitudinal
fore-wings
proximal
shiny,
scutellum and
notaulices almost complete,
the posterior margin of scutum;
and reticulate rugose;
one on
parallel
posterior surface
transversal
the pterostigma; part
(14:15);
bands,
distal
part of
fore tarsal
segments in following proportions: 12: 3: 8: 15: 32; enlarged claw (Fig. 136) with 1 bristle located further distally than the proximal prominence; segment 5 of fore a
group of 17
MALE:
fully
tarsi
(Fig.
lamellae;
winged;
136)
tibial
length 3,31
testaceous;
antennae testaceous;
femora and
hind
hairs
tibiae brown;
proportions:
15:
8:
the orbits;
15:
12:
mm;
black;
lamellae;
with coxae,
11:
11:
11:
carina complete;
antenna I segments
POL
=
6;
=
OL
3;
=
OOL
clubs of
11:
9:
12;
keel,
region
head dull,
with
two
10;
=
OPL
antennal
in following fully
lateral
keels around
of head behind ocelli
tracks of two oblique keels connecting posterior ocelli
carina;
apex with
mandibles and clypeus
antennae not distally thickened;
frons without median
occipital
two rows of 30
legs testaceous,
longer than breadth of segments;
reticulate rugose; with
with
spurs 1,1,2.
9;
=
TL
13;
only
to occipital
scutum,
scutellum
and metanotum shiny, smooth, finely punctate, without sculpture among punctures; notaulices almost complete, almost reaching the posterior margin of scutum; longitudinal
propodeum reticulate rugose, keels,
for-ewings hyaline,
without dark
vein shorter than proximal apical
part
branch wrapping penis;
LOCUS TYPICUS:
Amaicha del
TYPICAL MATERIAL:
without transversal
with posterior surface dull transversal (11 :13); tibial
Valle
holotype F!
HOSTS: unknown DISTRIBUTION: Neotropic region:
and
and
reticulate rugose;
bands;
distal
part of radial
gonoforceps
(Fig.
137)
without
spurs 1,1,2.
(Argentina)
and
1 paratype M!
ARGENTINA:
in TW.
Amaicha del
Valle,
TW!
39. Deinodryinus asper n. sp.
FEMALE:
fully
winged;
length 3,56 mm;
and anterior half of frons mandibles testaceous;
head black,
(mainly along orbits)
antennae testaceous,
prothorax reddish-testaceous,
with
temples reddish
testaceous;
clypeus and
with segments 8-10 brown;
with sides partly black;
mesothorax and
Anteoninae
Fig.
137 -
Male genitalia of Valle
Deinodryinus croceus n. sp. from Amaicha del
(Argentina):
meta thorax reddish-testaceous; testaceous-brown;
227
right half removed.
propodeum black;
legs testaceous,
pet iol e black;
with hind coxae partly
abdomen
brown;
antennae
sl ightly distally thickened; antenna I segments in following proportions: 13: 7: 19: 13: 9: 9: 7,5: 7,5: 7: 10; head with frons reticulate rugose and with 3 median vertex,
longitudinal
keels;
temples and region
2
lateral
behind ocelli
and without sculpture among punctures; 2 ob I i que keel s connect i ng carina complete; shiny,
hairy,
POL
with
=
3;
OL
of pronotum without sculpture, ::Jronotum, scutum,
nearly
as
long
region
posteri or ocell i
=
3;
=
OOL
an anterior strong
keels around
10;
scutellum and metanotum shiny,
,-.ithout sculpture among punctures;
OPL
transversal
pronotal
punctate
behind posterior ocell i
to occi pita I cari na;
with 2 dorsal
as broad;
the orbits;
not reticulate rugose,
=
8;
TL
furrow;
=
11;
pronotum
posterior surface
lobes parallel
to the sides of
tubercles reaching
smooth,
with
occi pita I
very finely
tegulae;
punctate,
notaulices almost complete,
almost
228
Mem.
Amer.
Ent.
Inst.,
reaching the posterior margin of scutum; posterior surface reticulate rugose, transversal
bands,
pterostigma; (15:13);
distal
segments
(Fig.
138)
prominence;
nearly 36 lamellae;
37,
1984
propodeum reticulate rugose,
with
dull; fore-wings with 2 dark
the basal
part of radial
fore tarsal
enlarged claw proximal
one on
no.
cells and another beneath the
vein slightly
longer than proximal
in following proportions:
12: 3:
7:
with 1 bristle located further distally
segment 5 of fore tarsi
(Fig.
apex with a group of nearly
138)
9:
part
29;
than
the
with 3 rows of
10 lamellae;
tibial
spurs
~ \
Fig.
138 -
Chela of Deinodryinus asper n.
sp.
(type)
\
229
Anteoninae
Fig.
139 -
Chela of
Deinodryinus schlingeri n. sp.
(type)
230
1,1,2. MALE:
Mem.
Amer.
En!.
Inst.,
no.
37,
1984
unknown
LOCUS TYPICUS:
Nova Teutonia
TYPICAL MATERIAL: HOSTS:
(Brazil)
holotype F!
in BM.
unknown
DISTRIBUTION: BM!
Neotropic region:
BRAZIL:
Nova Teutonia
(Sta Catarina),
40. Deinodryinus schlingeri n. sp.
FEMALE:
fully
winged;
length 3,62 mm;
malar space testaceous; legs brown, testaceous;
with fore tarsi, 13:
6:
17:
8:
mandibles,
clypeus and
with segments 7-10 brown;
fore tibiae and stalks of fore femora
antennae distally
proportions:
black;
antennae testaceous, thickened;
8:
8:
6:
6:
antennal
6: 9;
segments in following
head shiny;
frons
with
numerous longitudinal keels and areolae; occipital carina complete; region of head behind ocell i with numerous longitudinal keels, among which two oblique keels connecting posterior ocelli
=
2,5; OOL transversal
to occipital
carina;
POL = 3,5;
OL
=
15; OPL = 11; TL = 11; pronotum shiny, with an anterior furrow; posterior surface nearly as long as broad, with 2
weak parallel
lobes,
tubercles reach i ng
hairy,
tegu I ae;
weakly scutum,
rugose,
almost smooth;
pronotal
scute II um and metanotum smooth,
shiny,
finely punctate,
without sculpture among punctures;
almost
complete,
reaching
almost
the
posterior
margin
of
notaulices
scutum;
propodeum
reticulate rugose; posterior surface reticulate rugose, without longitudinal keels; fore-wings with 2 dark transversal bands, one on the basal cells and another beneath the pterostigmaj distal part of radial vein as long as proximal part (15:15); enlarged claw (Fig. 139) with 1 bristle located further distally than the proximal prominence; segment 3 of fore tarsi produced nearly
into a
22
hook;
lamellae;
segment 5 of fore tarsi
apex with
a
(Fig.
group of nearly
139)
with 2 rows of
18 lamellae;
tibial
spurs
1,1,2. MALE:
unknown
LOCUS TYPICUS:
Tingo Maria
TYPICAL MATERIAL: HOSTS:
in CA.
unknown
DISTRIBUTION: NOTE:
(Peru)
holotype F!
Neotropic region:
the species
is named
in
PERU:
Tingo Maria
honor of Mr.
E.I.
(Monson Valley),
CAl
Schlinger.
41. Deinodryinus alexandrae n. sp.
FEMALE: frons,
fully
winged;
length 6 mm;
head black,
clypeus and mandibles testaceous;
1-4 testaceous; propodeum,
prothorax reddish-testaceous;
abdomen
black;
fore
with anterior region of
antennae black, mesothorax,
legs testaceous,
with
with segments metathorax,
tarsi
brown;
mid
Anteoninae
and hind
legs black;
antennae distally
following proportions;
18;
8:
30:
23:
231
thickened antenna I segments
16:
15:
13:
with frons reticulate rugose and sculptured by behind ocelli carina POL
=
and 5;
with
=
12:
15;
longitudinal
4;
=
OOl
longitudinal
16;
=
OPl
keels;
13;
=
Tl
occipital 15;
in
head shiny, keels;
two oblique keels connecting posterior ocelli
with numerous
Ol
13:
region
to occipital
carina complete;
pronotum shiny,
smooth,
with a strong transversal impression and with pronotal tubercles reaching tegulae; posterior surface of pronotum nearly as long as broad, with two dorsal
parallel
longitudinal
lobes;
scutum,
scutellum and metanotum shiny,
smooth, without sculpture; notaul ices almost complete, almost reaching posterior margin of scutum; propodeum reticulate rugose, without transversal
or
longitudinal
reticulate rugose; basal vein
keels;
fore-wings with
cells and another beneath longer than proximal
proportions;
11:
Fig.
4:
140 -
11:
(Fig.
140)
42;
Chela of
located further distally tarsi
part
17:
with
than
posterior surface of propodeum dull, two dark
(22: 19);
HOSTS;
bands,
distal
tarsal (Fig.
segments 140)
one on
in following
with
1 bristle
Deinodryinus alexandrae n. sp. (type)
the proximal
two rows of 29
holotype F!
Neotropic region:
the species
is named
the
part of radial
prominence;
lamellae;
segment 5 of fore
apex with a
is the male of
group of 13
D. alexandrae)
(Ecuador)
in TW.
unknown
DISTRIBUTION: NOTE;
fore
enlarged claw
Napo and Coca Ri vers
TYPICAL MATERIAL:
transversal
the pterostigma;
lameillae; tibial spurs 1,1,2. MALE: unknown (perhaps D. hirticornis LOCUS TYP I CUS;
the
ECUADOR:
Napo and Coca Rivers,
in honor of Mrs.
Alessandra Mucci.
TW!
232
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
42. Deinodryinus carpen,s n. sp.
FEMALE: MALE:
unknown
fully
winged;
length 3,68 mm;
black;
antennae not distally thickened antennal breadth of segments; 17:
11:
11:
11:
10:
frons with numerous OL = 3,5;
OOL = 9;
Fig.
141
9:
9:
11;
head shiny,
longitudinal OPL = 8;
keels;
TL = 7;
to occipital
mainly on anterior half,
-
much shorter than
antenna I segments in following proportions:
connecting posterior ocelli punctate,
fore tibiae testaceous;
hairs short,
Male genitalia of
punctate,
12: 8:
without sculpture;
occipital carina complete;
POL
6;
vertex without two oblique keels carina;
scutum shiny,
strongly
without sculpture among punctures;
Deinodryinus carpens n. sp. (type): left half removed.
Anteon i nae
notaulices
incomplete,
almost reaching
scutellum and metanotum shiny, among punctures;
233
the posterior margin of scutum;
smooth,
finely
propodeum reticulate rugose,
punctate, without
without sculpture
longitudinal
posterior surface of propodeum reticulate rugose and with small shiny central distal
area;
fore-wings hyaline,
part of radial
vein
without dark transversal
about as long as proximal
part;
keels;
smooth bands;
gonoforceps
(Fig. 141) with subapical branch wrapping penis; tibial spurs 1,1,2. LOCUS TYPICUS: S.J. Barreiro (Brazil) TYPICAL MATERIAL: holotype M! and 2 paratypes MM! in Ol; 2 paratypes MM!
in TW ••
HOSTS:
unknown
DISTRIBUTION: OL!
Neotropic region:
BRAZil:
S.J.
Barreiro (Serra da Bocaina),
TW!
43. Deinodryinus perlucens n. sp.
FEMALE: MALE:
unknown
fully
winged;
antennae fully testaceous, thickened,
length 3,75-4,12 mm;
testaceous,
black;
mandibles testaceous;
sometimes with segments 3-10 darkened;
with stalks of hind femora brown;
with hairs much shorter than breadth of segments;
segments in following proportions: head sh i ny,
15: 7:
20:
legs
antennae not distally 12:
12:
12:
11:
antennal 11:
with frons scu I ptured by numerous strong para Ilel
11:
14;
and
longitudinal keels; region behind ocelli strongly punctate, without sculpture among punctures, without two strong oblique keels connecting posterior ocelli complete; finely
to occipital
POL = 4;
punctate,
incomplete,
smooth,
occipital
carina complete;
OOl = 11; OPL = 11;
TL = 6;
without sculpture among punctures;
smooth,
finely
punctate,
propodeum with dorsal
surface smooth,
shiny,
fore-wings hyaline,
finely
lOCUS TYPICUS: HOSTS:
notaul ices
Avispas
scutellum and
without sculpture among
without sculpture among punctures;
transversal
bands;
distal
vein shorter than proximal part (12:19); gonoforceps (Fig. wide apical branch wrapping penis; tibial spurs 1,1,2. TYPICAL MATERIAL:
part of radial 142)
with a
(Peru)
holotype M!
in TW;
1 para type M!
in OL.
unknown
DISTRIBUTION:
Neotropic region:
PERU:
Avispas
(Marcapata),
44. Deinodryinus inermis n. sp.
FEMALE:
unknown
line
surface reticulate rugose and posterior
punctate,
without dark
frontal
scutum shiny,
almost reaching the posterior margin of scutum;
metanotum shiny, punctures;
carina;
OL = 2,5;
Ol!
TW!
234
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
"
/
I
Fig.
Male genitalia of Deinodryinus perlucens n.
142 -
ha I f
MALE:
fully
winged;
length 3,37 mm;
black;
antennae testaceous;
legs
with coxae,
tibiae partly black;
antennae not distally thickened;
following proportions: rugose; POL = 7;
frons
17:
testaceous, 8:
with 3 median
OL = 4;
OOL = 12;
15:
13:
sp.
(type):
left
removed.
14:
longitudinal OPL = 11;
oblique keels connecting posterior ocelli
14:
mandibles testaceous; clubs of femora and hind
12: 9:
keels;
TL = 11;
antenna I segments in 11:
occipital
11;
head reticulate
carina complete;
vertex with 2 strong
to occipital
carina;
antennae with
Anteoninae
ha i rs much
longer than breadth of segments;
metanotum sh i ny,
smooth,
punctate,
notaul ices almost complete, strong
transversal
keel
almost smooth;
between
fore-wings
scutell um and
approximately 0,80
with some transversal dorsal
posterior surface with 2 slightly visible shiny,
scutum,
without scu I pture among punctures;
reaching
propodeum reticulate rugose, and a
235
surface
and posterior surface
longitudinal
hyaline,
length of scutum;
keels on dorsal keels;
without dark
median area
transversal
bands;
distal part of radial vein nearly as long as proximal part (13:13); gonoforceps (Fig. 143) without apical branch wrapping penis, only with a small
Fig.
apical
143 -
lobe;
tibial
spurs 1,1,2.
Male genitalia of half removed);
lOCUS TYP I CUS:
las Cej as
TYPICAL MATERIAL: M! in TW. HOSTS:
Deinodryinus inermis n. sp. (type)(A: right
B:
gonoforceps
(laterally
viewed).
(Argen tin a)
holotype M!
in TU;
1 para type M!
in Ol;
paratype
unknown
DISTRIBUTION:
Neotropic region:
ARGENTINA:
las Cejas
(Tucuman),
Ol!
TU!
236
Mem.
Jujuy,
Amer.
En!.
Inst.,
no.
37,1984
TW!
45. Deinodryinus paranus n. sp.
FEMALE: unknown MALE: fully winged; antennae testaceous;
length 1,62-2,68 mm; black; mandibles, legs and stalk of hind femora and distal half of hind tibiae
dark; antennae not distally thickened; antenna I segments in following proportions: 9: 5: 7: 4: 5: 5: 5: 5: 5: 6,5; head shiny, smooth, punctate, without sculpture among punctures; occipital
carina complete;
keels connecting posterior ocelli = 4;
OPl = 4;
scutum,
Tl = 3;
frons without median and
region behind posterior ocelli to occipital
antenna I hairs almost reaching
keels;
POL = 4;
Ol = 2;
OOl
longer than breadth of segments;
scutellum and metanotum shiny,
notaul ices incomplete,
carina;
lateral
without oblique
smooth,
without sculpture;
the posterior margin of scutum;
propodeum reticulate rugose, without transversal or longitudinal keels; fore-wings hyal ine, without dark transversal bands; distal part of radial vein slightly shorter than proximal an apical
Fig.
branch wrapping penis;
144 -
part tibial
(7:8)
gonoforceps
Male genital ia of Deinodryinus paranus n. ha I f
lOCUS TYPICUS:
Faz.
Taperinha
removed.
(Para,
(Fig.
144)
with
spurs 1,1,2.
Brazil)
sp.
(type):
right
237
Anteoninae
TYPICAL MATERIAL: M!
holotype M!
in OT;
2 paratypes MM!
in TW;
1 paratype
in Ol.
HOSTS:
unknown
DISTRIBUTION: ARGENTINA:
Neotropic region:
Horco Molle
BRAZil:
(Tucuman),
Ol!
Faz.
Taperinha
(Para),
OT!
TW!
46. Deinodryinus hirticornis (Kieffer) n. sp.
/
I
/
I
\/
\
I
/
/
/
\
II \
\
I
,I '-
\ '-
/
Fig.
145 -
Male genital ia of
Deinodryinus hirticornis (Kieffer) (type):
gonoforceps removed.
238
Mem.
Fig.
146 -
Amer.
Male genital ia of de las Cenyas
Ent.
Inst.,
no.
37,
1984
Deinodryinus hirticornis {Kieffer} from Rio {Mexico}:
left half removed.
Anteon hirticornis Kieffer 1911b: 231. Trisanteon hirticornis {Kieffer}: Kieffer 1913b: 300. Trisanteon hirticornis {Kieffer}: Kieffer 1914b: 197.
FEMALE: MALE:
unknown
fully
winged;
antennae testaceous;
length 3,25-4,18 mm;
black;
occasionally clypeus black;
abdomen brown-black;
legs testaceous,with coxae,
mandibles,
clypeus and
tegulae testaceous; clubs of femora,
mid and
hind tibiae partly black or brown; antennal segments in following proportions: 13: 7: 13: 12: 12: 11: 10: 10: 8: 12; antennal hairs longer
Anteoninae
than breadth of segments; with
two median
head shiny,
longitudinal
keels;
keels connecting posterior ocell i complete;
=
POL
7;
=
OL
5;
notaul ices
margin of scutum;
strongly reticulate rugose;
region behind ocell i
to occipital
=
OOL
scutellum and metanotum se;iny, among punctures;
239
12;
smooth,
carina;
=
OPL
11;
finely
incomplete,
keels on
12;
punctate,
frons
two obi ique
occipital
=
TL
almost reaching
propodeum reticulate rugose,
incomplete longitudinal
with
carina
scutum,
without sculpture the posterior
without or with 2
posterior surface;
area between
these
two
keels rugose, dull, with strong transversal keels; fore-wings hyaline, without dark transversal bands; distal part of radial vein nearly as long as proximal proximal
part
(10:9);
gonoforceps
membranous process,
(Figs.
with apical
145,
146)
without dorsal
branch wrapping penis;
tibial
spurs 1,1,2. LOCUS TYPICUS:
Atoyac
TYPICAL MATERIAL: HOSTS:
(Mexico)
holotype M!
in
BM.
unknown
DISTRIBUTION:
Neotropic region:
MEXICO:
Atoyac
(Vera Cruz),
BM!
Rio de
las Cenyas (8 mi. NW of Acaponeta, Nayarit), CAl ECUADOR: Coca, TW! Napo and Coca Rivers, OL! TW! BRAZIL: Teodoro Sampaio (Sao Paulo), OL! TW!
Nova Conquista
NOTES:
(Bahia),
this species
is very
TW!
Surumu
(Roraima),
near the Nearctic
chief difference is concerning
the presence of two
posterior surface of propodeum
OL!
longitudinal
D. atriventris no keel
(in
TW!
D. atriventris (Cresson). The keels on
is visible).
47. Deinodryinus rivularis n. sp.
FEMALE: fully winged; length 3,80 mm; testaceous-reddish, with antennal segments 9-10 darkened; posteri or edges of eyes and part I y temp les blackened;
a
small
anterior marg in;
darkened median
sutu re between
mesosternum black; thickened;
antennal
10:
10:
9:
10:
frontal
=
clubs of femora
3,5;
scutum
brown;
antennae weakly distally
in following
proportions:
head dull,
granulate;
frons rugose,
two
lateral
weak
longitudinal
keels connecting ocellar triangle
OOL
=
is visible near the
segments
line and with
longitudinal OL
13;
area on
metanotum and propodeum blackened;
10;
OPL
=
10;
TL
=
6;
15:
keels;
to occipital
occipital
8:
with
19:
14:
11:
incomplete
vertex without carina;
POL = 3;
carina complete;
pronoturn
shiny, weakly granulated, with pronotal tubercles reaching tegulae; posterior surface of pronotum nearly 1,5 times as broad as long, with numerous short
long i tud i na I keel s near the posterior marg in;
surface sculptured by some strong metanotum shiny, reaching without
without sculpture;
approximately 0,75
radial
bands, cell
one beneath open;
distal
keels;
keels;
scutum,
posterior scutellum and
notaulices incomplete,
length of scutum;
longitudinal or transversal
transversal cells;
smooth,
transversal
propodeum reticulate rugose,
fore-wings with
two dark
the pterostigma and another on part of radial
vein
about as
the basal
long as
240
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
".::: :.:-
A
Fig.
147 -
cells of females of Deinodryinus nigricans
Radial
and pilosifrons Fenton
proximal 3:
7:
part
15:
27;
distally than with
(20:20);
fore tarsal
enlarged claw the proximal
lamellae
Rio
HOSTS:
Itenez
148)
lamellae;
(one of these very
TYPICAL MATERIAL:
segments in following proportions:
prominence;
two rows of nearly 40
MALE: unknown LOCUS TYPICUS:
(Fig.
long);
(Cameron) (A)
(B).
with
13:
1 bristle located further
segment 5 of fore
tarsi
(Fig.
148)
apex with a group of nearly 7
tibial
spurs 1,1,2.
(Bolivia)
holotype F!
in AM.
unknown
DISTRIBUTION:
Neotropic region:
BOLIVIA:
Rio
Itenez (Dept.
Beni),
AM!
48. Deinodryinus aequalis n. sp.
FEMALE: unknown MALE: fully winged; testaceous;
distally thikened, segments
length 3,25-4,25 mm;
antennae testaceous; with
black;
legs brown,
with
clypeus and mandibles tarsi
ligth;
antennae not
hairs longer than breadth of segments;
in following proportions:
15: 9:
20:
10:
11:
13:
12:
antenna I
12:
12:
14;
head dull, reticulate rugose; frons fully reticulate rugose, with frontal line; region behind ocelli punctate or reticulate rugose and with two obi ique keels connecting carina complete;
posterior ocell i
POL = 5;
OL = 3,5;
scutellum and metanotum shiny, among punctures; margin of scutum;
to occipital
carina;
OOL = 11; OPL = 6;
smooth,
finely punctate,
notaulices almost complete,
occipital
TL = 8;
scutum,
without sculpture
reaching almost the posterior
propodeum reticulate rugose;
posterior surface of
Anteoninae
241
propodeum with median area weakly rugose, two
lateral
longitudinal
keels and by
wings with one weak dark
Fig.
distal (Fig.
almost smooth,
vein
longer than
with broad apical
band beneath
proximal
branch wrapping
part penis;
LOCUS TYPICUS; S.J. Barreiro (Brazil) TYPICAL MATERIAL: holotype M! and 3 paratypes MM! MM! in OL. HOSTS:
surrounded by
anterior transversal
keel;
fore-
the pterostigma;
Chela of Deinodryinus rivularis n.
148 -
part of radial 149)
transversal
an
sp.
(type)
(17:15); tibial
gonoforceps
spurs 1,1,2.
2 paratypes
in TW;
unknown
DISTRIBUTION:
Neotropic region:
Bocaina), OL!
TW!
BRAZIL:
S.
J.
Barreiro (Serra da
49. Deinodryinus minor n. sp.
FEMALE: fully winged; length 3,60 mm; head black, with clypeus and mand i b les testaceous; antennae and legs testaceous; thorax, propodeum and abdomen
testaceous;
anterior half of pronotum brown;
antennae long,
weakly distally thickened;
antenna I segments in following proportions:
7:
11:
17:
14:
12:
head rugose; longitudinal
11:
11:
frons
11:
with
keels;
14;
head dull,
incomplete frontal
occipital
granulated;
I ine and with
carina complete;
two
vertex without
lateral
POL ~ 3;
OL ~ 4;
keels connecting ocellar triangle to occipital
carina;
an anterior transversal
OPL
furrow;
~
8;
TL
pronotal
~
5;
pronotum shiny,
lobes reaching
tegulae;
with
weak
longitudinal
~
7;
13:
anterior half of
pronotum weakly granulated,
OOL fully
242
Mem.
Fig.
149 -
Amer.
Male genitalia of
En!.
sculptured by strong than
long
(22:20);
sculpture;
transversal
scutum; keels; open;
(15:17); proximal
prominence;
nearly 41
lamellae;
very MALE:
long);
tibial
150)
almost reaching without
1 bristle
apex with a
Itenez
(Bolivia)
holotype F!
(Fig.
group of nearly 6
spurs 1,1,2.
Rio
TYP:CAL MATERIAL:
transversal or bands;
in AM.
without
longitudinal radial
14:
3:
6:
located further distally
segment 5 of fore tarsi
broader
smooth,
the posterior margin of
in following proportions:
with
unknown
LOCUS TYPICUS:
area near the
vein slightly shorter than proximal
segments
(Fig.
except for a small
without dark transversal
part of radial
fore tarsal
removed.
keels,
propodeum reticulate rugose,
enlarged claw
1984
scutellum and metanotum shiny,
incomplete,
fore-wings hyaline, distal
37,
posterior surface of pronotum slightly
scutum,
notaulices
no.
Deinodryinus aequalis n. sp. (type): right ha I f
posterior margin smooth;
Ins!.,
150)
cell
part 19: than
30; the
with 2 rows of
lamellae
(one of these
Anteoninae
150 -
Fig.
HOSTS:
243
Chela of Deinodryinus minor n.
sp.
(type)
unknown
DISTRIBUTION:
Neotropic region:
BOLIVIA:
Rio
Itenez
(Dept.
Beni),
AM!
50. Deinodryinus benianus n. sp.
FEMALE:
unknown
(probably
other species known from
this species
the same
is the male of one of the two
locality:
D. rivularis and minor; the
males of these two species are unknown; only breeding experiments can finally answer this question). MALE:
fully
winged;
length 3,20 mm;
and mandibles testaceous; hind femora partly brown;
black;
anterior margin of clypeus
antennae testaceous;
legs yellow,
hind coxae partly brown;
hairs nearly as long as breadth of segments;
with mid and
antennae hairy,
antennae not distally
with
Mem.
244
thickened; 11:
12:
antenna I segments
10:
12:
the occipital longitudinal
16;
En!.
frontal
parallel
posterior ocelli scutum,
keels;
scutellum
than distal
bands;
sl ightly
shorter than penis;
lOCUS TYPICUS:
Rio
TYPICAL MATERIAL:
Fig.
=
Itenez
areas;
151)
with
tibial
8:
21:
punctate;
track of two
4,5;
Ol
=
3;
OOl
smooth,
notaulices incomplete, median
12:
11:
occipital
incomplete
vein
8;
OPl
=
almost reaching
rugose,
without
longer than
without dark proximal
part
branch wrapping penis,
1,1 ;2.
(Bolivia)
holotype M!
in AM.
unknown
151
-
Male genitalia of
Deinodryinus benianus ha I f
removed.
8;
area of posterior surface
large internal
spurs
=
strongly punctate,
fore-wings hyaline,
part of radial
gonoforceps
16:
temples and vertex near
propodeum reticulate
lateral
(Fig.
with
and metanotum shiny,
(14:12);
1984
two oblique keels connecting
POL
keels on posterior surface;
not more excavated
HOSTS:
frons
with
cari'1a;
without sculpture among punctures;
transversal
37,
not reticulate rugose,
vertex
to occipital
no.
reticulate rugose;
line complete;
the posterior margin of scutum;
longitudinal
Ins!.,
in following proportions:
head shiny,
carina almost fully
carina complete;
Tl = 7;
Amer.
n.
sp.
(type):
left
Anteoninae
DISTRIBUTION:
Neotropic region:
BOLIVIA:
245
Rio
Itenez
(Beni),
AM!
51. Deinodryinus itenezi n. sp.
FEMALE:
unknown
(probably
this species
is the male of one of the two
other species known from the same locality: D. rivularis and minor; the males of these two species are unknown; only breeding experiments can finally MALE:
answer this question). fully
testaceous,
winged;
length 2,0 mm;
black;
with coxae little darkened;
distally;
antenna I hairs
segments
in following proportions:
segments 4-9 nearly
mandibles,
antennae and
antennae not distally
longer than breadth of segments;
as
antennal
7: 7: 5: 5: 5: 5: 5: 6:
12:
long as broad;
head shiny,
legs
thickened; 11;
antennal
reticulate rugose;
frons without longitudinal keels; occipital carina complete; vertex with little visible longitudinal keels connecting posterior ocelli to occipital carina; POL = 10; OL and metanotum shiny,
= 8;
punctures;
incomplete,
scutum; without
notaul ices
dark
median
transversal
than proximal penis,
OOL = 7; OPL = 6; TL = 5; scutum, scutellum smooth, finely punctate, without sculpture among reaching
propodeum reticulate rugose,
posterior surface; part
without
two
two
longitudinal
area shiny and smooth; bands;
(3:10);
internal
approximately 0,30
with
distal
branch
fore-wings hyal ine,
part of radial
gonoforceps
(Fig.
wrapping
152)
penis;
length of
keels on
vein much shorter
slightly
tibial
shorter than
spurs
1,1,2.
LOCUS TYPICUS: Rio Itenez (Bolivia) TYPICAL MATERIAL: holotype M! in AM. HOSTS:
unknown
DISTRIBUTION:
Neotropic region:
BOLIVIA:
Rio
Itenez
(Beni),
AM!
52. Deinodryinus biloboides n. sp.
FEMALE:
fully
winged;
length 5,80 mm;
cl ypeus and ma I ar space testaceous; 6-10 brown;
propectus black,
with anterior, whitish;
posterior and
pronotal
abdomen
mid and hind
12:
12:
11:
lateral
reddi sh;
13;
margins
testaceous;
legs testaceous, partly
dull;
without
to occipital
vertex behind posterior ocell i
but with numerous complete frontal
longitudinal
line;
POL
=
6;
keels; OL
=
lobes
meta thorax and hind coxae part I y
antennae distally 23:
occipital
7:
45:
20:
20:
carina complete;
keels connecting posterior ocell i not reticulate rugose,
frons reticulate rugose, 3;
segmen ts
pronotum black,
dorsal
with
black;
in following proportions:
head reticulate rugose, longitudinal
with
mesothorax,
region behind ocell i carina;
with mandibles,
side testaceous;
testaceous;
tibiae and femora
thiCkened antenna I segments 16:
with dorsal
tubercles partly
propodeum black; black,
head black,
antennae testaceous,
OOL
=
18;
OPL
=
13;
with TL
=
12;
246
Mem.
Fig.
152 -
Amer.
En!.
Ins!.,
no.
37,
1984
Deinodryinus itenezi n. sp. (type): right
Male genital ia of
half removed.
pronotum with pronotal
tubercles reaching
transverse impression and pronotum dull, punctate, reaching
alutaceous;
tegulae,
two dorsal
scutum,
longitudinal
the basal
radial
vein shorter than proximal
cells and another beneath
following proportions:
18:
8:
18:
part 24:
notaulices thin,
lobes;
smooth,
incomplete,
propodeum reticulate rugose;
keels and with median area shiny,
fore-w i ngs wi th
one on
anterior strong prominent
scutellum and metanotum shiny,
length of scutum;
not ret icu I ate rugose;
with an
transverse very
without sculpture among punctures; approximately 0,5
posterior surface without smooth,
with
two dark
(22:28);
54;
transversa I bands,
the pterostigma; fore tarsal
enlarged claw
distal
part of
segments
(Fig.
153)
in
with one
bristle located further distally than the proximal prominence; segment 5 of fore tarsi (Fig. 153) with two rows of nearly 40 lamellae; apex with a group of nearly 25 MALE:
lamellae;
tibial
spurs 1,1,2.
unknown
lOCUS TYPICUS:
Mouth of Rio Baures
TYP ICAl MATER IAl:
holotype F!
(Bolivia)
in AM.
Anteoninae
247
I
Fig.
HOSTS:
153 -
sp.
(type)
unknown
DISTRIBUTION: Dep t.
Chela of Deinodryinus biloboides n.
Ben i ),
Neotropic region:
AM!
BOLIVIA:
Mouth of Rio Baures (Rio
Itenez,
248
Mem.
Amer.
Ent.
Ins!.,
no.
37,
1984
53. Deinodryinus fluviatilis n. sp.
FEMALE:
fully
clypeus,
antennal
testaceous,
winged;
antennae distally
without
occipital
keels,
smooth,
OPl = 16;
Tl = 14;
tuberc I es reach i ng broad as
long,
with some punctures, to occipital
tegu I ae;
beneath
except
carina;
shiny,
smooth,
without transversal
transversal
the pterostigma;
frontal
bands, distal
line
two oblique keels
POL = 4;
Ol = 4;
without sculpture; lobes;
without sculpture;
OOl = 14;
pronotal tw i ce as
scutum, notaulices
the posterior margin of scutum; or
longitudinal
surface with median area shiny and smooth, with two dark
and
in
posterior surface of pronotum near I y
almost reaching
reticulate rugose,
antenna I segments
two prominent transverse dorsal
scutellum and metanotum shiny, almost complete,
antennae
abdomen
region of head behind ocell i partly
pronotum :1airy,
with
propodeum,
keels on frons and vertex;
carina complete;
connecting posterior ocelli
with mandibles,
13: 10: 10: 9, 9: 9: 11; head shiny,
with numerous longitudinal
complete;
thorax,
thickened;
17: 10: 28:
following proportions:
head black,
malar space testaceous;
wi th segments 7-10 brown;
legs testaceous; rugose,
length 4,40 mm;
sockets and partly
keels;
without areolae;
one on the basal
part of radial
propodeum
posterior fore-wings
cells and another
vein nearly
as
long as
(21 :21); fore tarsal segments in following proportions: 15: 4: 11: 17: 41; enlarged claw (Fig. 154) with 1 bristle located further distally than the proximal prominence; segment 5 of fore tarsi (Fig. 154) proximal
part
Fig.
with
154 - Chela of Deinodryinus fluviatilis n. sp.
two rows of nearly 23
lamellae; MALE:
tibial
lamellae;
spurs 1,1,2.
unknown
lOCUS TYPICUS:
Rio Mamore
(Bolivia)
apex with a
(type)
group of nearly
17
Anteoninae
TYPICAL MATERIAL: HOSTS:
holotype F!
249
in AM.
unknown
DISTRIBUTION: Rio Grande,
Neotropic region:
Dept.
Beni),
BOLIVIA:
Rio Mamore (5 km NW mouth of
AM!
54. Deinodryinus cuzcanus n. sp.
FEMALE: MALE:
unknown
fully
winged;
testaceous;
length 2,25 mm;
legs brown,
with
tarsi
black;
ligth;
antennae and mandibles
antennae not distally
with segments 3-5 triangular and with distal
Fig.
Male genitalia of Deinodryinus cuzcanus n.
155 -
ha I f
apex;
antennal
6:
6:
7:
10;
segments with
sp.
proximal
(type):
right
removed.
segments in following proportions:
antennal
thickened,
apex broader than
hairs
10: 6,5:
5,5:
5:
4:
5:
longer than breadth of
segments; antennal segments 3-5 not uniformly hairy, but with groups of hairs on distal apex of each segment; head dull, fully reticulate rugose; frontal
I ine absent;
occipital
frons with two
carina complete;
lateral
obi ique keels connecting posterior ocell i 4,5; OOL punctate,
=
5;
OPL
=
5;
TL
keels around
the orbits;
region of head beind posterior ocell i
=
5;
to occipital
scutum shiny,
without sculpture among punctures;
carina;
smooth,
without
POL = 7;
hairy,
finely
notaulices incomplete,
OL
250
Mem.
Fig.
156 -
Amer.
En!.
reaching
approximately 0,30
smooth,
fi nel y
median
area shiny,
transversal part
(4:11);
without
bands;
smooth,
gonoforceps
internal
1984
with a
strong
not rugose;
part of radial (Fig.
scutellum and metanotum
without scu I pture among punctures; transversal
posterior surface with
distal
37,
removed.
length of scutum;
punctate,
propodeum reticulate rugose, and posterior surface;
no.
Deinodryinus panamensis n. sp. (type): right
Male genitalia of
ha I f
sh i ny,
Inst.,
155)
two
fore-wings hyaline,
between dorsal without dark
vein much shorter than
with distal
branch wrapping penis;
keel
long i tud i na I keel s;
tibial
proximal
inner pointed process and spurs 1,1,2.
251
Anteoninae
LOCUS TYPICUS:
Quincemil
TYPICAL MATERIAL: HOSTS:
(Peru)
holotype M!
in OT;
1 paratype M!
in AM.
unknown
DISTRIBUTION: Turrialba,
Fig.
Neotropic region:
PERU:
Quincemil
(Cuzco),
OT!
COSTA RICA:
AM!
Males of Deinodryinus
157 -
(B):
trinidadi n. sp.
and noyesi n.
(A)
sp.
length of notaul ices.
55. Deinodryinus panamensis n. sp.
FEMALE: MALE:
unknown
fully
testaceous,
winged;
legs black,
18:
16:
without two
head shiny,
OPL = 16;
keels,
TL = 16;
occipital
hairy,
almost reaching punctate,
propodeum reticulate rugose,
part of radial
gonoforceps (Fig.
vein with distal
spurs 1,1,2.
~OCUS
TYPICUS:
Barro Colorado
TYPICAL MATERIAL:
holotype M!
-IOSTS: unknown JISTRIBUTION: Neotropic region: JA!
4;
carina;
or
longitudinal
part nearly as it's,
long as proximal
however, open,
because
is partly broader and partly effaced;
156) shorter than penis,
tibial
=
two
band beneath the pterostigma;
apparently closed: vein
OL
with
to occipital
without transversal
cell
the distal
frons 7;
the posterior margin of scutum;
radial
radial
=
without sculpture among
fore-wings with one dark transversal radial cell
vertex not
without sculpture among punctures;
keels;
open;
POL
region behind posterior ocelli
finely punctate,
notaul ices almost complete,
(10:10);
antennae not
carina complete;
fully reticulate rugose;
scutellum and metanotum shiny,
part
mandibles
last segment dark;
ligth;
oblique keels connecting posterior ocelli
scutum shiny,
punctures;
partly
with
strongly reticulate rugose;
strongly punctate;
longitudinal
longitudinal
clypeus testaceous;
strongly hairy, with hairs longer than breadth of segments in following proportions: 21: 10: 29: 17: 16:
15: 20;
reticulate rugose, OOL = 15;
black;
antennae testaceous,
with fore tibiae and fore tarsi
distally thickened, segments; antennal 18:
length 4,80 mm;
with teeth dark;
I.
with branch wrapping penis;
(Panama)
in DA. PANAMA:
Barro Colorado
I.
(Canal
Zone),
252
Fig.
Mem.
158 -
Amer.
En!.
Ins!.,
no.
Female of Lonchodryinus ruficornis (England)
37,
1984
(Dalman)
from
Point of Ayr
Anteoninae
253
3. Lonchodryinus Kieffer
Lonchodryinus Kieffer 1905a: 95. Prenanteon Kieffer 1913b: 301 (syn. proposed by Olmi 1983); type species Prenanteon melanocerum (Kieffer 1905), design. by Richards 1935.
Psilanteon Kieffer 1913b: 301 (n. syn.); type species Psilanteon aequalis (Kieffer 1905), orig. desig. Lonchodryinus Kieffer: Kieffer 1914b: 14. Prenanteon Kieffer: Kieffer 1914b: 199. Psilanteon Kieffer: Kieffer 1914b: 212. Prenanteon Kieffer: Berland 1928: 153. Prenanteon Kieffer: Richards 1937: 119. Psilanteon Kieffer: Richards 1937: 119. Prenanteon Kieffer: Richards 1939: 235. Prenanteon Kieffer: Muesebeck and Walkley 1951: 1041. Prenanteon Kieffer: J.F. Perkins 1976: 23. Prenanteon Kieffer: Krombein 1979: 1242. Lonchodryinus Kieffer: Olmi and Currado 1979a: 342. Prenanteon Kieffer: Moczar 1983a: 186. Lonchodryinus Kieffer: Olmi 1983: 81.
Fig.
159 -
Female of Lonchodryinus ruficornis (Dalman) from Point of Ayr (England)
254
Fig.
Mem. Amer.
160 -
Male of
Ent.
Inst.,
no. 37,
1984
Lonchodryinus ruficornis (Dalman) from Hunstanton (Norfolk, England)
Type species: FEMALE (Figs.
Lonchodryinus tricolor Kieffer 1905, monotypic. 158,
159):
fully winged
(Fig.
158) or occasionally
brachypterous (Fig. 162); maxillary palpi with 6 segments; labial palpi with 3 segments; fore-wing with distal part of radial vein as long as or longer than proximal part (Figs. 158, 159); occasionally slightly shorter; inner side of enlarged claw with a proximal
prominence bearing one
255
Anteoninae
Fig.
161
Male of Lonchodryinus ruficornis
-
(Norfolk,
bristle
(Fig.
strong
transversal
MALE (Figs. labial palpi
164); occipital keel
from Hunstanton
propodeum usually without a
and posterior surface.
160, 161): fully winged; maxj Ilary palpi with 6 segments; with 3 segments; fore-wing with distal part of radial vein
shorter;
proximal
pterostigma less
propodeum usually posterior surface; 167);
carina complete;
between dorsal
as long as or longer than slightly
(Dalman)
England)
part
(Figs.
160,
than four times as
without a strong
transversal
gonoforceps without apical
keel
161); occasionally long as broad; between dorsal
branch wrapping penis
and (Fig.
vertex of head without two obi ique keels connecting posterior ocell i
to occipital antennal
carina;
DISTRIBUTION: HOSTS:
antennal
hairs at most as
long as breadth of
segments. worldwide
Cicadellidae (i nc I ud i ng Typhlocybinae)
SPECIES:
12
PALAEARCT I C SPEC I ES OF LONCHODRYINUS
Three species are known.
KEY TO THE SPECIES
256
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
FEMALES
Fore-wing
-
reduced, short, without pterostigma (Figs. 162, 163) .. .••••.•••••••••••....•...••.. 1. subapterus (Kieffer) Fore-wing not reduced, with pterostigma (Figs. 158, 159) . . . . . . . . . • • • • • • • • • • • • • • . • • • • • • • • • • • •• 2. ruficornis (Da I man) The female of L.
nepalensis n. sp. is unknown
MALES
Temples nearly emarginated -
twice as
(Fig.
long as POL;
167) . . . . . . .
Temples nearly as
long as,
3.
gonoforceps apically
nepalensis n. sp.
slightly
longer or
shorter
than POL;
gonoforceps apically not emarginated (Fig. 166) . . . . . . . . . . . . . . . . . • • • • • . . • • • • • • • • . • • • . • • • • • • • •• 2. ruficornis (Da I man) The male of L.
subapterus (Kieffer) is unknown.
1. Lonchodryinus subapterus (Kieffer)
n.
comb.
Anteon subapterus Kieffer in Kieffer et Marshall 1905: 138. Mystrophorus subapterus (Kieffer): Kieffer 1914b: 195. Prenanteon ruficornis (Dalman) var. subapterus (Kieffer): Richards 1939:
243.
Prenanteon subapterus (Kieffer): J.F. Perkins 1976: 23.
FEMALE
(Figs.
reaching 0,5
162,
163):
micropterous,
length of propodeum;
and tegulae yellow,
antennae fully
with fore-wing reduced and not
length 3,81
mm;
yellow-reddish,
black,
with mandibles
legs yellow-reddish,
with proximal region of hind coxae and distal part of hind femora brown; head shiny, smooth, finely punctate; pronotum hairy, rugose, strongly punctate; scutum shiny, smooth, finely punctate; notaulices incomplete, extending distinctly beyond 0,5 length of scutum; scutellum shiny, smooth, without sculpture;
metanotum rugose,
dull;
propodeum reticulate rugose;
posterior surface with 2 longitudinal keels; another median keel is visible between these two keels; fore tarsal segment 1 twice as long as 4 (10: 5); enlarged claw
(Fig.
164)
segment 5 of fore tarsi lamellae; MALE:
with a
(Fig.
proximal
164)
prominence bearing one bristle;
with 2 rows of 3 (proximal)
apex with a group of 7 lamellae;
unknown
tibial
spurs 1,1,2.
+ 17
257
Anteoninae
Fig.
162 -
Female of Lonchodryinus subapterus ( Ireland)
LOCUS TYPICUS:
Aviemore
TYPICAL MATERIAL:
(Scotland)
holotype F!
in BM.
(Kieffer)
from Mullanx
258
Mem. Amer.
HOSTS:
Ent.
Inst.,
no.
37,
1984
unknown
DISTRIBUTION:
Palaearctic region:
U.
S. S.
R.:
Russian Lapland
(Hellen
1919); ENGLAND: Church Stretton (Caer Caradoc, SP.), BM! Salop (J. F. Perkins 1976); Hopton Fen (Suffolk), BM! SCOTLAND: Aviemore (Inverness East),
BM!
Wichen sedge Fen
Hawksmoor (Richards 1939); IRELAND: Mullanx, BM! NOTES:
(Cambs),
BM!
Corrie (Gretna,
Lichfield Dumfries)
probably this species is a variety of L. answer th i 5 quest ion.
(Staffs),
OX!
(Richards 1939);
ruficornis. Only breeding
experi ments can
Fig.
163 -
Female of Lonchodryinus subapterus (Kieffer) from Mullanx (Ireland)
2. Lonchodryinus ruficornis (Da I man)
Gonatopus ruficornis Dalman 1818: 83. Gonatopus frontalis Dalman 1818: 84 (n. syn.); locus typicus Vastergotland
(Sweden) (discussion on synonymy:
see page 267).
Gonatopus basalis Dalman 1818: 84 (syn. proposed by Richards 1939); locus typicus Smaland (Sweden). Gonatop us fuscicornis Da I man 1818: 87 (syn. proposed by Richards 1939); locus typicus Vastergotland (Sweden). Dryinus iongicornis Dalman 1823: 10 (syn. proposed by Richards 1939); locus typicus Smaland (Sweden).
Anteon i nae
259
Dryinus crassimanus Haliday in Curtis 1828: 206 (syn. proposed by Richards 1939); locus typicus Holywood (Ireland). Dryinus daos Walker 1837: 423 (syn. proposed by Dlmi 1983); locus typicus London (England) (the M! selected as type by Richards 1935 is very damaged and not identifiable: a second male of daos in Haliday collection is a ruficorne). Dryinus ilus Walker 1837: 424 (syn. proposed by Richards 1939); locus typicus Eng I and. Dryinus misor Walker 1837: 424 (syn. proposed by Richards 1939); locus typicus Eng I and. Dryinus lapponicus Thomson 1860: 176 (n. syn.); locus typicus South part of Lap I and
(Sweden).
Dryinus retusus Thomson 1860: 176 (n. syn.); locus typicus Norway. Chelogynus lepidus Forster 1861: 39 (n. syn.); locus typicus Roseggthal
(Switzerland).
Chelogynus frontalis (Dalman): Dalla Torre 1898: 540. Anteon aequalis Kieffer in Kieffer et Marshall 1905: 177 (n. syn.); locus typicus Glen Lyon (Scotland). Anteon frontalis (Dalman): Kieffer in Kieffer et Marshall 1905: 179. Anteon melanocera Kieffer in Kieffer et Marshall 1905: 179 (syn. proposed by Richards 1939); locus typicus Mugdock (Scotland). Anteon crassiscapus Kieffer in Kieffer et Marshall 1905: 180 (syn. proposed by Richards 1939); locus typicus Dieppe (France). Anteon vitellinipes Kieffer in Kieffer et Marshall 1905: 181 (n. syn.)j locus typicus Clober (Scotland). Anteon proximus Kieffer in Kieffer et Marshall 1905: 182 (n. syn.) jlocus typicus Svabhegy (Hungary). Anteon foersteri Kieffer in Kieffer et Marshall 1905: 183 (n. syn.)j locus typicus unknown. Anteon procericornis Kieffer in Kieffer et Marshall 1905: 184 (syn. proposed by Richards 1939); locus typicus Sutherland (Scotland). Anteon longicornis (Dalman) var. ruficornis (Dalman): Kieffer in Kieffer et Marshall
1905:
186.
Anteon luteipes Kieffer in Kieffer et Marshall 1905: 187 (syn. proposed by Richards 1939); locus typicus Meudon (France). Anteon declivis Kieffer in Kieffer et Marshall 1905: 188 (syn. proposed by Richards 1939); locus typicus Clydesdale (Scotland). Anteon pallidinervis Kieffer in Kieffer et Marshal 1905: 189 (syn. proposed by Richards 1939); locus typicus Kill i ngdock (Scot I and). Anteon dolichocerus Kieffer in Kieffer et Marshall 1905: 190 (syn. proposed by Richards 1939); locus typicus Sevres (france). Anteon integer Kieffer in Kieffer et Marshall 1905: 206 (n. syn.); locus typicus Mugdock (Scotland). Anteon parvicollis Kieffer in Kieffer et Marshall 1905: 206 (syn. proposed by Richards 1939); locus typicus Mesnil-Ie-Roi (France). Anteon pyrenaicus Kieffer in Kieffer et Marshall 1905: 207 (syn. proposed by Richards 1939); locus typicus Ax-Ies-Thermes (France).
260
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
Anteon curvinervis Kieffer in Kieffer et Marshall 1905: 208 (n. syn.); locus typicus Bonar Bridge (Scotland). Anteon fractinervis Kieffer in Kieffer et Marshall 1905: 208 (syn. proposed by Richards 1939); locus typicus Clober (Scotland). Anteon hyalinipennis Kieffer in Kieffer et Marshall 1905: 209 (syn. proposed by Richards 1939); locus typicus Dairy (Scotland). Anteon fuscipennis Kieffer in Kieffer et Marshall 1905: 209 (n. syn.); locus typicus Maroth (Hungary). Anteon longifilis I
inval id;
type in BU
according
to
Moczar 1967).
Anteon daos (Walker): Kieffer in Kieffer et Marshall 1906: 521. Anteon vitellineipes Kieffer: Cameron 1907: 32. Anteon pallidicornis Kieffer: Cameron 1907: 33. Anteon crassimanus (Haliday): Kieffer 1907b: 25. Anteon ilus (Walker): Kieffer 1907b: 26. Anteon lapponicus (Thomson): Kieffer 1907b: 26. Anteon lepidus (Foerster): Kieffer 1907b: 26. Anteon retusus (Thomson): Kieffer 1907b: 27. Antaeon morleyi Chitty 1908: 144 (syn. proposed by Richards 1939); locus typicus Foxha II (Eng I and). Antaeon luffnessensis Chitty 1908: 145 (syn. proposed by Richards 1939); locus typicus Luffness Links (Scotland). Antaeon beaumonti Chitty 1908: 145 (n. syn.); locus typicus Chobham (England) •
Prenanteon frontalis (Dalman): Kieffer 1913b: 301. Anteon daos (Walker): Kieffer 1914b: 164. Chelogynus walkeri Kieffer 1914b: 185 (syn. proposed by Richards 1939); locus typicus Holywood (Ireland). Prenanteon vitellinipes longifilis (Kieffer): Kieffer 1914b: 203. Prenanteon longicornis (Dalman): Kieffer 1914b: 205. Prenanteon pyrenaicus (K i effer) var parcepunctatus Kieffer 1914b: 209 (syn. proposed by Richards 1939); locus typicus Clober (Scotland). PrelJanteon palustris Oglobl in 1924a: 102 (n. syn.); locus typicus Strbske pleso
(Czechoslovakia).
Prenanteon filicornis Oglobl in 1924b: 127 (n. syn.); locus typicus Pisek
(Czechoslovakia). frontalis (Dalman):
Prenanteon PrelJanteon PrelJanteolJ PrelJanteon Prenanteon Prenanteon Prenanteon
Berland
1928:
154.
luteipes (Kieffer): Berland 1928: 155. crassiscapus (Kieffer): Berland 1928: 156. pyrenaicus (Kieffer): Berland 1928: 156. parvicollis (Kieffer): Berland 1928: 156. longicornis (Kieffer): Berland 1928: 157. luteipes (Kieffer): Maneval 1935: 6.
Anteoninae
Prenanteon Prenanteon Prenanteon Prenanteon
261
daos (Walker): Richards 1939: 236. ruficornis (Dalman): Richards 1939: 240. frontalis (Dalman): Richards 1939: 245. euscelisi Haupt 1941: 61 (n. syn.); locus typicus Ha II e
(Germany) •
Prenanteon luteiceps (lapsus}(Kieffer): Menozzi 1942: 37. Prenanteon longicornis (Dalman): Cavro 1950: 5. Prenanteon semenovi Ponomarenko 1970: 428 (n. syn.); locus typicus Gremyachka
(U.
S.
S.
R.).
Prenanteon foveatus Richards 1971: 48 (syn. proposed by Olmi 1983); locus typicus Si I wood Park (Eng I and) • Prenanteon frontalis (Dalman): J.F. Perkins 1976: 23. Prenanteon ruficornis (Dalman): J.F. Perkins 1976: 23. Prenanteon longicornis (Dalman): J.F. Perkins 1976: 24. Prenanteon foveatus Richards: J.F. Perkins 1976: 24. Prenanteon basalis (Dalman): J.F. Perkins 1976: 25. Prenanteon ruficorne (Dalman): Ponomarenko 1978: 21. Prenanteon semenovi Ponomarenko: Ponomarenko 1978: 21. Prenanteon ruficornis (Dalman): Moczar 1983a: 186. Prenanteon pektusanense Moczar 1983a: 187 (n. syn.); locus typicus Mt.
Pektusan
(Korea).
Prenanteon basale (Dalman): Moczar 1983a: 189. Prenanteon c1avatum Moczar 1983a: 189 (n. syn.); locus typicus Mt. Pektusan
(Korea).
Prenanteon Prenanteon Prenanteon Prenanteon
FEMALE
(Figs.
occasionally dark;
melanocera (Kieffer): Moczar 1983b: 200. basale (Dalman): Moczar 1983b: 201. longicornis (Da I man): Moczar 1983b: 202. c1avatum Moczar: Moczar 1983b: 203.
158,
159):
fully
winged;
length 2,25-4,68 mm;
head and thorax brown-reddish;
occasionally frons mainly reddish;
black;
mandibles yellow,
with
teeth
sometimes malar space and frons
frontalis Dalman); clypeus yellow, black or (var. longicornis. daos and ret usus ) or brown, with segments 1-2 testaceous (var. ruficornis and basalis); mainly
whitish or yellow
redd ish-dark;
(var.
antennae testaceous
occasionally
antennae brown
occasionally
also segments 6-10 or 7-10 testaceous;
fu II y
testaceous or with
brown-testaceous; 14:
12:
shiny,
=
7;
12:
10:
strongly
Ol
=
4;
deep,
very
=
10;
black;
tegulae
testaceous;
abdomen
black or
in following proportions:
antennae slightly distally
smooth,
OPl
1-2 or 1-3 testaceous;
=
impression;
8;
12:
thickened;
6:
13:
head
without sculpture among punctures; Tl
=
8;
frons without median
sometimes a
deep groove
legs
pal
groove on
is visible
(in
foveatus and retusus); frontal impression more or less
variable,
pronotum shiny,
15;
punctate,
OOl of
10:
1,
hind coxae basa II y
antenna I segments
10:
disc or with discal holotypes FF!
with segment
with no specific value;
strongly
punctate;
pronotal
occipital
carina complete;
tubercles reaching
tegulae;
the
262
Mem.
Fig.
164 -
Amer.
En!.
Inst.,
no.
37,
1984
Lonchodryinus subapterus (Kieffer) from Mullanx
Chela of
(Ireland)
scutum shiny, strong I y
smooth,
punctate,
without punctures;
occasionally scutum finely or
wi thout scu I pture among punctures;
incomplete, occasionally
long,
the holotype F!
as in
Perkins
1976 notaulices at most reaching mid
as in
the holotype F!
of the var.
basalis (according to J. F.
scutum,
occasionally notaul ices short,
length of scutum);
reaching approximately 0,5
of the var.
typ ica I finely
as in
the
lectotype F!
reaching and
length of scutum,
longicornis (according to J. F. Perkins
1976 notaulices extending distinctly beyond mid occasionally notaul ices shorter, scutum,
notau I ices
reaching approximately 0,65 length of
in
length of scutum);
approximately 0,3
length of
the paralectotypes FF!
of the
ruficornis (i n types the notau I ices are not sha Ilower, narrow and aciculate,
metanotum shiny,
as written by J. smooth,
F.
Perkins 1976);
scutellum and
without sculpture or punctate;
propodeum
Anteoninae
reticulate rugose,
posterior surface with 2 longitudinal
usually as rugose as lateral and
longicornis)
rugose
(as
hyaline;
in
and dull;
nearly
(9:
7);
(as in
part;
of
retusus
vein as
and
long as,
fore tarsal
segment 5 of front
15-20 lamellae;
keels;
the holotypes FF!
occasionally median area shiny,
part of radial
longer than proximal segment 4
areas
the holotypes FF!
distal
263
fore-wing
slightly shorter or slightly
(Fig.
165)
longer than
with 1 row of
apex with a group of nearly 3-5 lamellae;
2 of front tarsus produced into a hook; enlarged claw (Fig. bristle located on proximal prominence; tibial spurs 1,1,2.
Fig.
165 -
basalis
smooth and not
euscelisi);
segment 1 slightly
tarsus
median area of
Chela of
Lonchodryinus ruficornis
(Dalman)
segment
165)
with one
from Granarolo
( Italy)
MALE
(Figs.
160,
161):
mandibles testaceous, testaceous;
5;
winged;
length 2,12-3,75 mm;
teeth brown;
tegulae testac'eous;
occasionally punctate,
fully
with
legs testaceous,
POL = 5;
OL = 2,5;
head shiny, OOl = 8;
frons with or without a more or less deep central
carina complete; among punctures; of scutum,
scutum shiny,
smooth,
notaulices incomplete,
finely
smooth,
finely
finely
OPl
=
4;
punctate,
without sculpture
reaching approximately 0,5
punctate,
TL =
impression; occipital
broad and deep or narrow and finely crenulate;
metanotum shiny,
occasionally
with coxae partly brown;
also clubs of femora partly brown;
without sculpture;
black;
antennae fully brown;
length
scutellum and
without sculpture among
264
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
, I
!
/
Fig.
166 -
(Dalman)
punctures;
Gonoforceps
areas,
long as,
(Switzerland) (B).
keels on
without dark transversal
slightly
almost without sculpture;
bands;
shorter or slightly
lateral
distal
longer than
view variable,
part of radial proximal
narrow or broad
part;
(Fig.
166);
spurs 1,1,2.
LOCUS TYPICUS:
Vastergotland
TYPICAL MATERIAL: lectotype F!
basalis,
(Sweden)
the numerous types are in following collections:
holotype M!
of G.
paralectotypes FF! lectotYpe M! of
lectotype F!
frontalis; in D. daos, lectotype
of G.
of G.
ruficornis,
in ST:
C. fuscicornis, lectotype F! (hereby desi gnated) D. longicornis, holotype F! of D. lapponicus,
and 3 paralectotypes FF!
and 1 paralectotype F! of holotype F! of D. retusus, DU:
Lonchodryinus ruficornis
with 2 longitudinal
occasionally smooth and shiny,
shape of gonoforceps in tibial
ru~ose,
of
and Valetta
usually median area on posterior surface as rugose as
fore-wing hyal ine, vein as
(Ireland) (A)
propodeum reticulate
posterior surface; lateral
(laterally viewed)
from Avimore
/
(hereby PK: F!
designated)
and 2
Pro semenovi; D. crassimanus, lectotype
holotype F! of
holotype F! of
of
in F!
Anteoninae
of Ch.
walkeri; in VI:
lectotype M!
265
(hereby
designated)
and 3
of Ch.
lepidus; in MR: holotype M! of A. luffnessensis; in BM: holotype F! of A. aequalis, holotype F! of A. vitellinipes, holotype M! of A. curvinervis, holotype M! of A. integer, holotype M! of A. beaumonti, holotype F! allotype M! and 2 paratypes FF! of Pl'. foveatus, holotype M! of A. fractznervis, holotype F! of A. pallidinervis, holotype F! of A. dechvis, holotype F! of A. melanocera, holotype F! of A. procericornis, holotype M! of D. ilus, holotype M! of D. misor, holotype M! of A. morleyi, holotype M! of A. hyalinipennis, holotype M! of Pl'. parcepunctatus; in P: holotype F! of A. crassiscapus, holotype F! of A. luteipes, holotype F! of A. dolichocerus, holotype M! of A. pyrenaicus, lectotype M! (hereby designated) of A. parvicollis; in HA: holotype F! of Pl'. euscelisi; in BU: holotype F! of A. halidayi (type in BU, not in DU); types of A. proximus. A. foersteri. A. fuscipennis. Pl'. palustris. Pl'. filicornis. A. longifihs probably lost. HOSTS: in Italy Empoasca flavescens (Flor), Empoasca decipiens Paoli, Asymmetrasca decedens Paol i (= Empoasca decedens) (Menozzi 1942; specimen from Granarolo (BO!), leg. Menozzi, determined by Menozzi as Pl'. luteiceps Kieffer), Jassargus bisubulatus Theny; in England Psammotettix confinis (Dahlb.) (Waloff 1974, 1975; J.F. Perkins 1976), Psamn;otettix nodosus (Rib.) (Richards 1971; Waloff 1975); Psammotettix cephalotes (H.-S.) (Richards 1971), t-'uscelis obsoletus (Waloff 1974, 1975), Euscelis plebejus (Waloff 1974, 1975), Macrosteles laevis Rib. (Richards 1971), £lymana sulph urella and Arthaldeus pascuellus (Wa loff 1974, 1975); in Germany Streptanus sordid us (Zett.) (= lo'uscelis sordidus: Haupt 1941) and, for var. frontalis. Jassargus obtusivalvis (= lJeltocephalus picturatus Fieb.) and t-'rrastun us ocellaris F. (= lJeltocephalus ocellaris: Haup t 1941); in U.S.S.R. Nacrosteles laevis Rib. (Ponomarenko 1978). paralectotypes
(2FF,
DISTRIBUTION:
Palaearctic region:
Pay plateau
1M)!
(Sam-zi-yan,
Provo
JAPAN:
Kamikochi,
Ryang-gang),
BU!
TW!
KOREA:
Sam-zi-yan
Chann-
(Mt.
Pektusan), BU! Mu-do-bong (Mt. Pektusan, Chann-Pay plateau, Provo Ryang-gang), BU! MONGOLIA: Semon Bajanzogt (Central aimak), BU! Bogdo ul.
(Ulan-Baator,
Central
(Changaj Gebirge, Central
aimak),
Namnan
ul
(Chovsgol
S.
HE!
Metsapirtti,
HE!
Ekenas, (AI.),
(Tudosdomb),
HE!
SWEDEN:
(Lpl.),
Eksharad
OL!
Fokstua
Lojo,
(Lpl.),
Opheimsbakken
OT!
Gremyachka HE!
HE! HE!
Mausgnsbyn BM!
R.:
ovoo
OT!
(Dovre),
HE!
Vastergotland, Lapland,
ST!
Messaure,
(Vmld.),
OT!
NORWAY:
Vacz-Szod,
ST!
27°
BU!
(Trondhj.),
Abisko
(Lpl.),
BU!
ST!
OT!
Csevharaszt,
00'
HE!
OT!
~~orway,
Munkacs,
OL!
Sordavala, HE!
Foglo
OT!
Dalaro
BU!
E,
Kantalaks,
(Sjall.),
(Lpl.),
Smaland,
BU!
NEPAL:
N 85°
FINLAND:
Dyrehaven
Hatfjelddal,
Visegrad,
aimak),
Somon Burenchaan
58'
PK!
Riksgransen TW!
HUNGARY: BU!
Hell
HE!
BU!
aimak),
TW!
Somon Culuut
(Ulan-Baator,
Central
(Ryazan Province),
OT!
BU!
(Bulgan
Phakding,
Tammerfors,
CO!
Ichtamir -
Pass Zosijn
Bulgan aimak),
OL!
(Aurdal),
Somon BU!
(Ulan-Baater,
Chutag,
Somon Abzaga
(Katmandu),
U.
Jebrenjokk
BU!
BU!
aimak),
Tosgoni (Somon
aimak),
OT!
Bano
S.
BU!
Gebirge
Bhurumche
aimak),
Archangaj
Skane, ST!
Godollo, BU!
TW!
(Malmen), BU!
Tasnad,
Vacz BU!
266
Mem.
Felsohagi,
BU!
Amer.
Bukkszentielek,
Tihany,
BU!
Mecsek hgs.,
(Bukk),
BU!
Irsa pta.,
Budapest,
BU!
Nagykovacsi,
Ent.
BU!
BU!
BU!
BU!
Bukk h.
BU!
BU!
BU!
Budai
BU!
Unoko,
BU!
Mecsekh,
Ocsa,
Fulophaza,
BU!
Inarcs,
37,
BU!
BU!
H.,
BU!
O-Sebeshely,
BU!
BU!
BU!
BU!
Kunfeherto,
Kurtaberc
BU!
Valko,
Velem,
Hoi ics,
BU!
BU!
Nagyvisnyo,
Budakeszi,
BU!
1984
Csakvar park,
Barszentmihaly,
Borzsony, BU!
no.
Alp Kudsir.,
Dobogoko,
Balvany,
Tompa,
Inst.,
BU!
Szeged,
BU!
Kormochanya, BU!
BU!
SzSztMiklos,
BU!
Rev., BU! Bi harfured, BU! Koszeg, BU! Szod, BU! Tata tovaros, BU! Cs i k Szepvis, BU! Svabhegy (Budapest) (Kieffer in Kieffer et Marshall 1905); Szaohegy (Koszeg), BU! Boroszno, BU! ROMANIA (Nagy 1972); CZECHOSLOVAKIA: Bohemia, TW! M. Tatra, BU! Kobyli (Moravia), BULGARIA:
Mts Rila,
(Slovenija),
CO!
BU!
Gruz
YUGOSLAVIA:
(Dalmacija),
Mosbrucher Hochmoor (Eifel), (Bohmerwald), Berlin,
HU!
HU!
HU!
Weinberg
Beuerburg
Westpreussen),
CO!
HU!
Postojne (Slovenija),
VI!
GERMANY! Halle,
Bellinchen
(Gross-Machnow),
(Oberbayern),
CO!
Westf.),
HU!
HU!
HU!
Veldes
HA!
HA! Schwarzer See
Frechland
(Elbe),
Sachsen),
HU!
Aachen,
Berkesnvercher (Berlin),
HU!
Huben
HU! (0.
Niemegk, Tirol),
HU!
HU!
Kuhhornhatte
Igls
(Tirol),
(N.
OT!
Siebenburgen),
St.
Anton,
BM!
P!
LE!
Finkenkrug
(Berlin), HU! Mecklenburg, HU! Feldwege I. (Unna, Westf.), HU! (Hoheneck, Vogesen), HU! Misdrog, HU! Gipfel (Gr. Wasserkuppe, Gbg.),
HU!
Salesche (Tucheler Haide,
Mehlsheuer (Vagsland,
Saline (Unna,
(Oder) ,
BM!
BM!
HU!
Wien,
Fischbodle Rhon-
AUSTRIA: WA!
P!
Prater (Wien), P! Austria, P! Obergurgl (Tirol), OT! Semmeringgebeit (Reichenau), BM! Reichenau, P! DENMARK: Gentafte, CO! Tipperne (Jylland),
CO!
Ordrup Naes
(Odsherred,
Sjaell.),
CO!
Korsor,
CO!
Hoisteinborg, CO! Aarhus, CO! Vesterhede (Romo, Sdr. Jylland), CO! Espe, CO! Ulfshale, CO! Danmark, CO! HOLLAND: Wijster (Dr.), LE! Asperen, LE! SWITZERLAND: (Neuchatel), Valley
Auvernier, LO!
Duillier
(Engadin),
BU!
LO!
Les Ponts
(Vaud),
LO!
Suchet-Issom.
(Neuchatel),
Aclens (Vaud),
LO!
(Vaud), LO!
Montmollin
LO!
Nyon,
Morteratsch
GV!
Vercorin
(Valais), LO! Arolla (Valais), BM! Aarau, BM! Derborence (Valais), GV! Foret d'Aletsch (Valais), GV! Epalinges (Vaud), LO! Boussens (Vaud), OL! Pleiades
(Vaud),
Roseggthal Cluozza), P!
GV!
Fopps
Dieppe (Seine
(Seine-et-Oise), (Gard),
GV!
Mayens de Syon
(Grigioni),
GV!
(Val
Inf.), P!
Sevres
VI!
(Valais),
Roseg Valley Cluozza),
P!
Meudon
LO!
(Engadin),
GV!
FRANCE:
P!
Ax-Ies-Thermes
P! P!
(Vaud),
Valetta
Le Puy,
(Seine-et-Oise),
Maisons-Laffitte (Seine-et-Oise), (Seine),
Gryon BU!
P!
GV!
(Val
BO!
Bolquere,
Mesnil-Ie Roi Pont du Gard
(Ariege),
P!
Lille
(Nord),
P! Vincennes, P! La Frette, P! Foret de Marassan (Hautes-Alpes) , P! ENGLAND: Boxmoor (HT.), BM! Nunton (Moczar 1967); Eastbourne (Sussex), BM!
Wi mb ledon Com.
(London),
OT!
Cambri dge,
BM!
Sherford Bri dge (DT.),
BM! Chobham (Surrey), BM! Oxford, OT! Rhinefield (Hants.), BM! Ben Rhydding (Yorkshire), HA! Spratton (Northants), BM! Si Iwood Park (Berks.), Fowhall
8M!
Claygate
(Suffolk East),
(Dartmoor,
Devon),
Richmond Park,
OX!
OT!
(SR.), BM!
8M!
Wotton-under-Edge
White Downs
Aldbury
Hunstanton
(HT.),
(Dorking,
BM!
(Norfolk),
(Gloster West),
Surrey),
Point of Ayr
8M!
SCOTLAND:
BM!
(Flint),
BM!
Hay tor BM!
Wood Walton
Fen
Anteoninae
(Cambs.),
BM!
Glen lyon,
Bonar Bridge,
BM!
BM!
(Perths.),
Enochdhu
Killingdock, BM!
BM!
BM!
DU!
Avimore
DU!
Glenasmole
SZ!
SZ!
(Senales, BO!
Res.
(Fife),
BM!
(WX.),
BM!
BM!
(Torino),
Ol!
Co.
TW!
SZ!
(Genova),
MI!
Ol!
(Dublin),
Cavehill (in
OT!
trout's stomach),
1903);
(Aosta), (Cuneo),
OLl Pl!
(Bussoleno
Unserfrau
Granarolo
Monte Fascie (Genova), (Perugia),
BM!
(Belfast),
Pietrabianca
(Cobelli
BM!
(Kirkcudbrigth),
Benedetto Belbo
(Torino),
Monte Peglia
TW!
BM!
(Perths.),
ITALY: Jovencan
S.
Trentino
BO!
Clober,
(Perths.),
Dairy
(Wicklow)
TW!
BM!
BM!
Gollierstown
Naturno (Bolzano),
Rezzoaglio d'Aveto (Genova),
lavagna
BM!
Wicklow,
Kerry,
Noasca
OT!
Ballinluig
IRELAND: Co.
Aosta),
Condove (Torino),
Bolzano),
BM!
Bally Smutton
BM!
(Perths.),
Birks of Aberfeldy
Sutherland,
Ballyteige (Dublin),
BM!
Aviemore, BM!
(Valsavaranche,
la Cassa
Torino),
BM!
(Harwood),
Eaux Rousses
Rannoch Forest
(Stirling),
Clydesdale,
Tentsmuir Nat.
Athdown,
BM!
Mugdock
267
(Bologna), Ol!
Cavi
di
Ol!
NOTES: specimens from Japan are bigger than specimens from other countries. Synonymy questions about L. frontalis were defined in consequence of the discovery of males. Previously I observed in females intermediate forms with frontal spot very wide and very small. Doubts were removed in consequence of the discovery of males. I observed no difference between males of L. ruficornis and frontalis. Frontalis is only a variety with part of frons whitish or yellowish in females. This variety is not frequent. I know the var. frontalis from the following local ities: SWEDEN (Vastergotland), GERMANY (Aachen, Bellinchen, Schwarzer See), SWITZERLAND (Aarau, Mayens de Syon), FRANCE (Pont du Gard), ENGLAND (Oxford, Matley Bog), ITALY (La Cassa, Monte Fascie, Monte Peglia).
3. Lonchodryinus nepalensis n. sp.
FEMALE: MALE:
unknown
fully
winged;
mandibles whitish; testaceous;
legs testaceous,
distally thickened; 12:
11:
length 3 mm;
10: 9:
9:
8:
11;
~
5,5;
OL
~
2,3;
metanotum shiny,
notaul ices incomplete,
malar space, ventral
frontal ~
8;
smooth, ~
5;
TL
LOCUS TYPICUS:
28°
00'
N.
part;
scutum,
8:
6:
13:
without
carina complete; scutellum and
bands;
with
spurs 1,1,2.
85° 00'
E.
length of scutum; or longitudinal distal
keels;
part of radial
the two parts forming a curve;
inner process,
tibial
punctate,
occipital 10;
without transversal
vein nearly as long as proximal 167);
antennae not
without sculpture among punctures;
fore-wing hyaline without dark transversal
(Fig.
~
reaching approximately 0,5
propodeum reticulate rugose,
emarginated
finely
line absent;
OPl
punctate,
gonoforceps without distal
clypeus and
side of segment 1
segments in following proportions:
head shiny,
OOL
finely
with
with hind coxae basally black;
antennal
sculpture among punctures; POL
black;
antennae brown,
(Nepal)
internal
apical
margin
268
Mem. Amer.
En!.
Ins!.,
\ I I [\
Fig.
167 -
no.
37,
1984
/ ./
Male genitalia of Lonchodryinus nepalensis n. I eft ha I f removed.
TYPICAL MATERIAL: holotype M! in aT. HOSTS: unknown DISTRIBUTION: Palaearctic region: NEPAL:
28° 00'
N 85° 00'
sp.
E,
(type):
aT!
Anteoninae
ETHIOPIAN SPECIES OF
269
LONCHUDRYINUS
Two species are known.
KEY TO THE SPECIES
FEMALES
Fore-wing hyal ine, -
wi thout dark transversal bands •.•••••••••• . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1. seyrigi (Benoi t) Fore-wing wi th two dark transversal bands . . . . . . . . . . . . . . . . . . .. ••••••••..•••.•••••••.••••.•• 2. pauliani (Benoi t)
MALES Unknown
1.
Lonchodryinus seyrigi (Benoit) n. sp.
ifirtanteon seyrigi Benoi t 1953b: 421.
FEMALE:
fully
damaged,
winged;
without
length 3 mm;
head and fore
holotype (only
legs;
margins of pronotum reddish;
abdomen black;
testaceous;
punctate,
reaching
pronotum strong I y
tegulae;
scutum,
approximately 0,5 distal
notaulices
length of scutellum;
without keels; part of radial
fore-wing hyaline, vein
mid and hind
sh i ny,
incomplete,
propodcum fully
mandibles and clypeus yellow;
Bekily
TYPICAL MATERIAL:
part
description the head antennae yellow,
(10:7);
HOSTS:
Ethiopian region:
in P. MADAGASCAR:
spurs
with palpi,
with segments 7-10 black;
unknown
DISTRIBUTION:
bands;
tibial
is black,
(Madagascar)
holotype F!
punctate, reaching
reticulate rugose,
fore legs yellow. MALE: unknown LOCUS TYPICUS:
legs fully
without dark transversal
longer than proximal
1,1,2. According to the original
very with
with pronota I tuberc I es
scutellum and metanotum shiny,
without sculpture among punctures; dull,
known specimen)
thorax and propodeum black,
Bekily,
P!
270
Mem. Amer. En!.
Fig.
168 -
Ins!.,
no. 37,1984
Chela of Lonchodryinus pauliani (Benoit)(type)
271
Anteoninae
2. Lonchodryinus pauliani (Benoit) n. comb.
Prenanteon pauliani Benoit 1953b: 420.
FEMALE:
fully
testaceous,
winged;
length 5,50 mm;
with segments 8-10 brown;
head
yellow-testaceous;
propectus,
pronotum,
antennae
scutum,
scutellum and metanotum reddish-dark; tegulae testaceous; propodeum and abdomen black; mesopleura black, with reddish spot on anterior half; metapleura black;
fore
black,
ligth;
27:
with
12:
8:
tarsi 8:
8:
8:
8:
legs fully
reddish-testaceous;
11;
head shiny,
without sculpture among punctures; around
the orbits;
OPL = 13;
strongly
and hind
punctate,
pronotum shiny,
=
POL
smooth,
5;
finely
3;
punctate,
scu I pture among punctures,
with a strong transversa I an terior tegulae;
tubercles reaching
without sculpture among punctures; smooth,
keel
with central
smooth area,
metapleura shiny,
without sculpture;
pterostigma;
part of radial
distal
almost reaching long 7:
as broad
16:
29;
the basa I ce II s and
(19:4);
fore tarsal (Fig.
segment 5 of front
45 lamellae
(some of these very
hook;
segment 2 not produced
MALE: unknown LOCUS TYPICUS:
Mt.
TYPICAL MATERIAL: HOSTS:
with a (Fig.
long);
Tsaratanana
reticulate rugose;
fore-wing
with 2 dark
another beneath
longer than
proximal 168)
the
proximal
hook;
tibial
Ethiopian region:
group of nearly 7 tarsus produced
spurs 1,1,2.
(Madagascar) in P. MADAGASCAR:
OR lENT AL SPEC I ES OF
One species is known.
Mt.
4:
prominence bearing
unknown
DISTRIBUTION:
part,
with 2-3 rows of nearly
apex with a
segment 3 of front
into a
holotype F!
without
pterostigma more than four times as
tarsus
apex very prominent;
reaching
segments in following proportions:13:
168)
one bristle; lamellae;
vein much
the margin of wing;
enlarged claw
impress ion;
posterior surface
mesopleura
smooth and without sculpture; one on
14;
finely punctate,
reticulate rugose,
and posterior surface;
transversa I bands,
=
OOL
without
scutellum and metanotum shiny,
propodeum shiny,
between dorsal
smooth,
notaulices incomplete,
length of scutum;
without scu Ipture;
transversal
scutum shiny,
14: 7:
without keels
=
OL
pronotal
approximately 0,75
legs
mainly on frons,
frons with median keel,
occipital carina complete;
TL = 11;
mid
antenna I segments in following proportions:
Tsaratanana,
LONCHOD12Y INUS
P!
into a
272
Mem. Amer. En!.
Ins!.,
no. 37, 1984
Fig. 169 - Chela of Lonchodryinus sinensis n. sp.
(type)
Anteoninae
273
Lonchodryinus sinensis n. sp.
FEMALE:
fully
winged;
length 2,12 mm, except for the abdomen,
holotype (only known specimen); reg ion of frons testaceous;
black;
antennae brown;
femora and hind coxae partly brown; antennal
mandibles,
clypeus and
legs testaceous,
missing
with cl ubs of
antennae not distally thickened;
segments in following proportions:
10:
7:
13:
13:
10:
11: 9:
9:
12; head shiny, smooth, finely punctate, without sculpture among punctures; frons without median and lateral keels; POL = 4; OL = 2,5;
=
7;
OPL
=
4;
=
TL
strongly punctate, smooth,
occipital
carina complete;
with a strong
shorter than scutum shiny,
5;
(6:16);
finely punctate,
notaulices incomplete,
reaching
approximately 0,5 smooth,
with a strong
part
(10:9);
bands;
pterostigam
following proportions: proximal 169)
distal 3:
keel
lateral
areas;
part of radial
2,5:
6:
propodeum
between
dorsal
fore-wing vein
(25:6);
hyal ine,
longer than fore tarsal (Fig.
segment 5 of front
part shorter than proximal
tarsus with 2 rows of 44 lamellae;
length of scutum;
14; enlarged claw
prominence bearing one bristle;
with basal
scutum
surface
posterior surface with 2 longitudinal
longer than broad
11:
OOL
hairy,
posterior surface
without sculpture;
transversal
median area as rugose as
dark transversal
impression;
tubercles reaching tegulae;
and median area of posterior surface; keels;
pronotum shiny,
9:
without sculpture among punctures;
scutellum and metanotum shiny, reticulate rugose,
transversal
pronotal
in
anterior
part
(6:8);
without
proximal segments in
169) tarsus
with a (Fig.
segment 5 of front
apex with a group of 5 lamellae;
tibial
spurs 1,1,2. MALE: unknown LOCUS TYPICUS:
Tainan Hsien
TYPICAL MATERIAL: HOSTS:
(Taiwan)
holotype F!
in B.
unknown
DISTRIBUTION:
Oriental
region:
TAIWAN:
Tainan Hsien
NEARCT I C SPEC I ES OF
(Kwantzeling),
B!
LONCHODRY INUS
Three species are known.
KEY TO THE SPECIES
FEMALES
At -
least
the head
testaceous . . . • . .
1.
flavus n. sp.
Head most Iy black . . . . • . . . . . . • . . • • . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2
274
Mem.
2 Head dull, -
Amer.
granulated;
Ent.
Inst.,
antennae very
+ pronotum •••.•••.••••••••••• 2.
Head
shiny,
punctate,
without
antennae very
37,1984
short,
head
granulated;
no.
sl ight Iy
[{;asneri n.
longer
sculpture among punctures,
long,
much
longer
+ scutum . . . . . . . . . . . . . . . . . . . . . . . . . . 3.
than
sp.
than
not
+ pronotum
head
bakeri (Kieffer)
MALES
Posterior -
OPL at
ocell i
nearly
as
,5 times as
I east
long as
or
slightly
shorter
• . • • • ••
1.
flavus n.
sp.
long
as posterior
........
than
POL.
ocell i . . . . . . . . . . . . ..
bakeri (Kieffer)
3.
L. masneri is unknown.
The male of
1. Lonchodryinus flavus n. sp.
FEMALE:
fully
scutum brown
winged;
length 2,68-3,25 mm;
and pronotum dark;
abdomen black or darkened; not distally
thickened;
occasionally
antennal
fully
testaceous;
occasionally
in some specimens also propodeum and also metanotum dark;
segments
antennae
in following proportions:
8:
5:
7: 8: 8: 7: 6: 5: 5: 7; head shiny, smooth, finely punctate, without sculpture among punctures; frons without median and lateral keels; occipital
carina complete;
=
POL
5;
=
OL
3;
OOL = 7;
=
OPL
4;
=
TL
4;
pronotum very short, almost invisible, shiny, smooth; scutum, scutellum and metanotum shiny, smooth, finely punctate, without sculpture among punctures;
notaul ices incomplete,
scutum;
propodeum dull,
between
dorsal
and posterior surface,
poste,-ior surface; distal
fore-wing
part of radial
fore
tarsal
claw
(Fig.
5 of front
vein
hyaline,
more than
with 2
longitudinal
without dark three
lamellae;
winged;
170)
located on
a
as proximal
2,5:
4:
with 2 rows of nearly 25
tibial
bands;
long
proximal
(Fig.
2:
keel
keels on
transversal
times as
170)
with one bristle
length of
without transversal
6:
tarsus
fully
approximately 0,5
segments in following proportions:
a group of 11 MALE:
reaching
reticulate rugose,
11;
prominence; lamellae;
part;
enlarged segment
apex with
spurs 1,1,2.
length 3,00-3,75 mm;
black;
mandibles testaceous,
with teeth reddish; antennae brown, with segment 1 testaceous; legs testaceous; somet i mes anterior ha I f of frons testaceous; antennae not distally 13:
14:
thickened; 14:
punctures; 3,5;
OOL
=
14:
15:
antenna I segments 13:
15;
frontal
I ine absent;
5;
=
OPL
4;
TL
slightly shorter than POL;
in following proportions:
head shiny,
=
4;
occipital
punctate,
carina complete;
posterior ocell i
scutum shiny,
smooth,
10:
7:
14:
without sculpture among POL = 7;
nearly as finely
OL =
long as or
punctate,
without
An teon i nae
Fig.
170 -
Chela of
sculpture among punctures; 0,5
length of scutum;
sculpture;
275
Lonchodryinus flavus n. sp. (type)
notaulices
incomplete,
reaching
scutellum and metanotum shiny,
propodeum fully
reticulate rugose;
approximately
smooth,
without
posterior surface with
two
276
Mem.
longitudinal distal 171;
171
Ent.
fore-wing hyal ine,
part of radial
tibial
Fig.
keels;
Amer.
vein
Inst.,
no.
37,
1984
without dark transversal
nearly as long as proximal
part;
bands;
genital ia:
fig.
spurs 1,1,2.
Male genital ia of Lonchodryinus flavus n. (Canada): right half removed.
-
lOCUS TYPICUs:
Forsyth
TYPICAL MATERIAL: paratypes (2 FF, 3 MM)!
in Ol.
HOSTS:
unknown
DISTRIBUTION:
(Georgia,
holotype F!
4 MM)!
Nearctic region:
from Alberta
U.S. A.)
and 6 paratypes
in TW;
sp.
1 paratype F!
CANADA:
(1
F,
in AM;
5 MM)!
in OT;
6
6 paratypes (3 FF,
Rampart House (Yukon),
OT!
Jumping Pd. Cr. (Calgary, Alta.), Ol! OT! Bothwell (Ont.), Ol! OT! Hull (Que.), OT! Kirks Ferry (Que.), OT! OL! lac Mondor (Ste. Flore, Que.), OT!
U.S.A.:
(Md.),
TW!
Kerhonkson TW!
Ann Arbor
(Mich.),
High Point St. (Cherry town,
Bemus Pt.
(N.Y.),
Pk.
TW!
Ulster Co.,
Ol!
TW!
Forsyth
(N.J.),
TW!
N.Y.),
(Ga.),
Shokan AM!
Pinkham Notch
OT!
Takoma Pk.
(N.Y.),
TW!
Poughkeepsie (N.H.),
TW!
4 mi.
(N.Y.),
NNW Ol!
Anteoninae
277
2. Lonchodryinus masneri n. sp.
FEMALE:
fully
winged;
length 2,31-3,00 mm;
black;
mandibles testaceous;
occasionally also clypeus and malar space testaceous; with segments 1 and 7-10 brown; testaceous,
with coxae and clubs of femora partly brown;
hind coxae,
hind femora
distally
thickened;
7:
5:
7:
5:
frontal
4:
4:
half granulate; punctures; scutum; keels; dark 12:
occipital
pronotal smooth,
posterior surface with transversal part
19;
(7:8);
punctate,
172 -
(Fig.
reaching
Chela of
OL = 4;
scutum,
approximately 0,5
without
segments
segment 5 of front
transversal or
OOL = 8;
scutellum and length of longitudinal
fore-wing hyal ine,
without
vein sl ightly shorter than
in following proportions:
with one bristle
tarsus
(Fig.
lamellae;
13:
posterior
located on
172) tibial
with 2 rows of 40 spurs 1,1,2.
6:
2:
a proximal
Lonchodryinus masneri n. sp. (type)
apex with a group of 5
unknown
OPL = 4;
tegulae;
part of radial 172)
5:
without sculpture among
areolae smaller;
distal
fore tarsal
enlarged claw
prominence; lamellae;
bands;
9:
with anterior half rugose;
tubercles reaching finely
antennae slightly
with some punctures on frons;
carina complete;
pronotum dull,
legs
occasionally only
in following proportions:
granulate,
notaul ices incomplete,
Fig.
MALE:
head dull,
propodeum reticulate rugose,
proximal 3:
5;
TL = 6,5;
metanotum shiny,
and hind tibiae partly brown;
antenna I segments
line absent;
POL = 4,5;
antennae testaceous,
occasionally only segment 10 dark;
278
Mem.
LOCUS TYPICUS:
Cave Creek Cyn.
TYPICAL MATERIAL: FF!
in SC;
HOSTS:
holotype F!
1 paratype F!
En!.
Inst.,
(Arizona, in OT;
no.
37,1984
U.S.A.)
1 paratype F!
in TW;
2 paratypes
in OL.
unknown
DISTRIBUTION: OT!
Amer.
Nearctic region:
Chiricahua Mts.
Canyon
(Culberson
NOTES:
the species
(Portal,
Co.,
Tex.),
is named
U.S.A.:
Cave Creek Cyn.
Cochise Co.,
Ariz.),
OL!
(Portal,
SCI
Mc.
Ariz.),
Kittrick
TW!
in
honor of Mr.
Lubomir Masner.
3. Lonchodryinus bakeri (Kieffer) n. comb.
Anteon bakeri Kieffer 1906a: 239. f'renanteon bakeri (Kieffer): Kieffer 1914b: 212. f'renanteon micropunctatus Fenton 1927: 15 (n. syn.); locus typicus Nerepis
(N.
Prenanteon Prenanteon Prenanteon f'renanteon
FEMALE:
fully
black-brown;
B.,
U.
S.
A.).
bakeri (Kieffer): Muesebeck and Walkley 1951: 1041. micropunctatus Fenton: Muesebeck and Walkley 1951: 1042. bakeri (Kieffer): Krombein 1979: 1242. micropunctatus Fenton: Krombein 1979: 1242.
winged;
length 2,50-3,00 mm;
occasionally
antennae dark or testaceous, pleura,
scutum,
testaceous;
head and pronotum black or
pronotum partly reddish;
scutellum,
mandibles testaceous;
sometimes with segments 3-10 darkened;
metanotum and propodeum blaCk;
abdomen black or brown;
tegulae
legs brown-testaceous or testaceous,
with coxae basally black; occasionally only hind coxae basally black; antennal segments in following proportions: 11: 6: 15: 17: 14: 13: 10: 9: 9: 11; head shiny, smooth, finely punctate on vertex, temples, behind ocelli, on the sides of ocelli, on posterior half of frons; anterior half of frons more strongly or very keels; 7;
occipital
pronotum shiny,
scutum shiny,
strongly
finely
anterior margin; scutum;
finely punctate;
carina complete;
=
POL
punctate;
punctate,
more strongly
notaulices incomplete,
irregular keel
posterior surface with 2
between
longitudinal
=
3;
OOL
=
longitudinal 7;
OPL
=
tubercles reaching
7;
smooth,
0,50-0,65
length of
without SCUlpture;
propodeum reticulate rugose,
dorsal
TL
tegulae;
punctate only near the
reaching
scutellum and metanotum shiny,
transversal
frons without
OL
pronotal
occasionally metanotum weakly rugose; a
6;
without
and posterior surface;
keels sometimes weakly
visible;
median area nearly as rugose as lateral areas; fore-wing usually very long, hyaline, without dark transversal bands; distal part of radial vein longer than proximal proportions: hOOk;
12:
3:
enlarged claw
prominence;
part
3:
6:
(Fig.
(11:7); 15;
fore
tarsal
segments in following
segment 2 of front
173)
segment 5 of front
with one bristle tarsus
(Fig.
173)
tarsus produced located on a with basal
into a
proximal part shorter
Anteoninae
Fig.
173 -
Chela of Lonchodryinus bakeri (Kieffer) (N.
than distal
part,
of 4-7
lamellae;
MALE:
fully
279
with 2 rows of nearly 23-27 lamellae; tibial
winged;
from Good Hope
W. T., Canada)
apex with a group
spurs 1,1,2.
length
1,75-3,25 mm;
black;
mandibles testaceous;
280
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
antennae brown, with segments 1-2 or only 1 testaceous; legs testaceous, occasionally with hind coxae proximally black; antennae not distally thickened; 14:
13:
antennal
13:
12:
13;
seglcents
sculpture among punctures; deep median 3;
OOl
=
longitudinal
6;
=
OPl
posterior ocelli;
in following proportions:
head shiny,
3;
Tl
scutum,
smooth,
frons
furrow;
=
4;
without median and occipital
OPl at
transversal
keel
with 2 weakly hyal ine,
Fig.
visible
longitudinal
Male genital ia of Odessa
incomplete,
14:
keels, POL = 6;
long
keels;
fore-wing distal
14:
with a Ol
as
length of
smooth,
punctate,
reaching
propodeum reticulate rugose,
bands;
15:
without
lateral
1,5 times as
and posterior surface;
without dark transversal
174 -
least
notaulices
length of scutum;
between dorsal
6:
carina complete;
scutellum and metanotum shiny,
'without sculpture among punctures; approximately 0,5
10:
strongly punctate,
without a
posterior surface
usually very
part of radial
long,
vein
nearly
Lonchodryinus bakeri (Kieffer) from lake
(Michigan):
right
half removed.
Anteoninae
as long as proximal proximal
dorsal
part
(9:9);
281
gonoforceps
174) without membranous tibial
(Fig.
process and without distal
inner pointed process;
spurs 1,1,2. lOCUS TYPICUS:
Ormsby Co.
TYPICAL MATERIAL:
(Nev.,
holotype Fr
U.
S.
A.)
A. bakeri in CA; holotype Fr of Pro
of
micropunctatu5 in WA. HOSTS:
unknown
DISTRIBUTION:
Nearctic region:
Salmon (Naknek), Rich. Hwy.), aT! Big Delta, House
aT!
Hope
(N.
T.),
Pk.
Sandspit
aT!
(B.C.
W.
Stone Mt. aT!
Paxson,
(Yukon),
Chilkat Pass
(B.C.),
(Mi.
aT!
TW!
Unalakleet,
Tombstone C.G.
aT!
Atlin
(Hedley,
(Yukon),
Pro
Watson lake (Yukon),
aT!
(Dawson,
Good
(N.W.T.), (B.C.),
aT! TE!
Johnston Canyon
(Banff,
(Mi.
Yukon),
Kitimat
aT!
(B.C.),
Stewart
aT!
Alta.),
aT!
Gagnon Road
aT!
Battle River
(B.C.),
B.C.), (B.C.),
aT!
Terrace B.C.),
(Terrace,
(Alta.),
aT! aT!
B.C.),
aT!
(Terrace,
Spring Creek
TE!
Hixon
(Terrace,
(B.C.),
lakelse l .
aT!
aT!
(B.C.),
Ketchum l .
Cultus lake
(B.C.),
aT!
392 Alaska Hwy.,
B.C.),
aT!
B.C.),
Rupert
aT!
Rampart
TW!
(B.C.),
(Terrace,
aT!
TE!
lk.
Summit l .
B.C.),
King
aT!
Kitwanga
aT!
OTr
Gravel
496 Alaska Hwy.,
Shames
(B.C.),
aT!
Norman Wells
(B.C.),
(B.C.),
aT!
(Yukon),
Yukon),
OTr
151., Queen Charlotte Islands, B.C.), AM!
(Moresby
Nickle Plate City Robson
Dawson
CAl
Cowichan l .
Pete lake
CANADA:
-
Umiat,
Anchorage, OTr Shaw Crk. (M. 289 206 Richardson Hwy.), aT! Haynes,
TW!
(B.C.),
Liard Hot Spg.
ALASKA:
OTr Naknek, OTr Isabel Pass (Mi.
aT!
aT!
B.C.), Morley
(Alta.), aT! Frank (Alta.), aT! Mc Murray (Alta.), aT! Cameron l . Rd. (Waterton Nat. Park, Alta.), aT! White Fox (Sask.), aT! Churchill (Man.), aT!
Brandon
(Ont.), (Kam.
(Man.),
aT!
TW!
Co.,
Que.),
Whale River Chelsea, Co.,
Que.),
Cartwright
aT!
Mistassini
aT!
aT!
aT!
Warkworth Cr. (Ont.),
aT!
(Que.),
Que.),
aT!
Ottawa
Hull
(labrador),
(N.B.), aT!
(Que.),
(Que.),
Old Chelsea
Nerepis
(Churchill,
Merivale
(Que.), WA!
Hebron
aT!
aT!
(Oreg.),
TW!
Stanley
(Mt.
Shasta,
Ormsby Co. aT!
(Nev.),
Beaver Ck.
(Idaho),
TW!
CAl
aT!
aT!
Idaho City Cal if.),
(23 mi.
NNW West Yellowstone,
aT!
(labrador),
TW! U. S. A.: Westport (Oreg.), TW! Ochoco Creek
Utah),
TW! aT!
Gallatin Co.,
Chicago Cr.
(Mt.
Evans,
Colo.),
aT!
Dool ittle Ranch
aT!
(Mt.
Evans,
Colo.),
aT!
West Chicago Cr.
Echo l .
(Papineau
(N.B.),
Mc Bride
Crescent City
(Vernal,
Great
(Old
Goose Bay
(Idaho),
OTr
Stittsville
aT!
Buckingham Twp.
Kouchibouguac N.P.
Big Brush Cr.
aT!
lac Crescence
Chima (Que.),
(labrador),
Siskiyou Co.,
aT!
Summit of King Mt.
aT! Raleigh (labrador), TW! Doyles (labrador), (Wash.), TW! Mt. Ranier (Wash.), TW' Mt. Hood Springs
Man.),
(Ont.),
(Mt.
(Cal if.),
Vernal Mont.),
Evans,
(9800',
TW!
(Utah), AM!
Colo.),
Clear Cr.
Co., Colo.), aT! Estes Park (Colo.), aT! Phantom Vy (Colo.), TW! Gould (Colo.), TW! loveland Pass (Colo.), aT! Idaho Springs (Colo.), aT! Horosier Pass
(Summit Co.,
Colo.),
aT!
(Nebr.),
aT!
Polk Co.
(Mich.),
TW!
Washtenaw Co.
(Mich.),
Columbus
Crystal
Falls
(Mich.),
Ol!
Iron Co.
(Mich.),
BM!
Ol!
Ol!
aT!
(Black Hi 115,
Sylvan l .
(Wis.),
CAl
Iron River
(Mich.),
lake Odessa
Ol!
Oak.), TW!
Benzie Co.
(Mich.),
Yellow Dog Plains (Marquette Co.,
S.
(Mich.),
BM!
Mich.),
Ann Arbor
TW!
Rapides
282
Mem.
Parish
(La.),
Hi II
(Pr.
Pk.
(N.J.),
Placid Mt.
TW!
Pisgah
Georges Co., TW!
(N.Y.), OT!
NOTES: L. (Dalman).
Mt.
Md.),
Ent.
Inst.,
(N.C.), WA!
TW!
Troy
(N.Y.),
(6100'-6288',
EI
Saito
TW!
N.H.),
(Durango),
37,
1984
(N.C.),
(Md.),
TW!
(Huntington,
Dryden OT!
no.
Highlands
Cabin John
Kalbfleisch Field Station OT!
Washington
Durango,
Amer.
N.Y.),
(Maine),
Durham
TW!
TW!
(N.H.),
OT!
Oxon
High Point St. AM!
8M!
WA!
Lake
Summit of
MEXICO:
OT!
bakeri is very near the Palaearctic species L. ruficornis
LONCHOf)J(Y1NUS
NEOTRJP I C SPEC I ES OF
One species
is known.
Lonchodry in us tricolor Kieffer
Lonchodryinus Lonchodryinus Lonchodryinus Lonchodryinus
FEMALE:
fully
tricolor tricolor tricolor tricolor
winged;
Kieffer 1905a: 96. Kieffer:
Kieffer
Kieffer:
Olmi
and Currado 1979a:
Kieffer:
Olmi
1983:
length 6,0-7,5 mm;
1914b:
15.
head black,
anterior region of face and malar space testaceous; with
teeth reddish;
antennae fully
testaceous;
scutum usually reddish-testaceous; margins reddish,
with clypeus,
mandibles testaceous,
propectus,
pronotum and
occasionally pronotum black,
and scutum black;
metapleura black;
342.
81.
scutellum,
metanotum,
occasionally scutellum reddish;
with
propodeum and
mesopleura black,
distal region reddish, or fully reddish; petiole black; abdomen, and legs testaceous; antenna I segments in following proportions: 19:
18:
15:
numerous around
14:
13:
the orbits;
pronotum hairy, reaching
12:
longitudinal
14;
POL
shiny,
tegulae;
among punctures;
head dull,
keels
=
6;
granulated,
(obi ique on OL
smooth,
=
2,5;
finely
scutum shiny,
OOL
smooth,
notaulices distinct,
with frons hairy,
the sides of ocell i);
=
13;
punctate;
with
tegulae 20: 8: 25:
OPL
=
17;
pronotal
keel
TL
=
with is
12;
tubercles
finely punctate,
almost complete,
a
without sculpture
reaching
almost the
posterior margin of scutum; scutellum and metanotum shiny, smooth, without sculpture; mesopleura and metapleura dull, rugose; propodeum fully
reticulate rugose,
surface; with a vein
dull,
weak dark spot beneath
longer than proximal
forming
without keels between
posterior surface without
an
proportions:
angle; 12: 4:
6:
13:
22;
curvi I inear; fig.
176;
dorsal
keels;
the pterostigma,
part,
pterostigma:
longitudinal
with distal
distal
fore tarsal
segment 5 of front
and posterior
fore-wing hyal ine, part of radial
and proximal segments
tarsus
(Fig.
parts
in following 175)
with
Anteoninae
Fig.
175 -
Chela of
Lonchodryinus tricolor
283
Kieffer
(type)
284
Mem.
Amer.
Ent.
Ins!.,
no.
37,
1984
some rows of nearly 57 enlarged claw prominence;
Fig.
176 -
lamellae; apex with a group of 9 lamellae; 175) with one bristle located on a proximal spurs 1,1,2.
(Fig.
tibial
Pterostigma of
Lonchodryinus tricolor Kieffer (type): length of
pterostigma measured from A to B.
MALE:
unknown
lOCUS TYPICUS:
Goias
TYPICAL MATERIAL:
(Brazil)
holotype F!
HOSTS: unknown DISTRIBUTION: Neotropic region: TW!
Caruaru,
Ol!
TW!
in P. BRAZil:
Encruzilhada
Goias,
(Bahia),
AUSTRAL IAN SPEC I ES OF
P!
Ol!
Vilhena
(Rondonia),
TW!
LONCHODl2YINUS
Two species are known.
KEY TO THE SPECIES
FEMALES Unknown
MALES
Head black,
wi th mandibles,
clypeus and anterior
region of face 1. guineensis n. sp. Head fully black; only mandibles testaceous . . . . . . . . . . . . . . . . . . .. . . . . . . . . . . . . . . . . . . . . • . . • • . . • . . . . . . . . . . . 2. notogeicus n. sp. testaceous . . . . . . . . . . . . . . . . . . . . . . . . . . . .
-
1. Lonchodryinus guineensis n. sp.
Anteoninae
FEMALE: MALE:
285
unknown
fully
winged;
length 2,37 mm;
and anterior reg ion of frons testaceous;
head black,
testaceous;
thorax and propodeum black;
antennae not distally
thickened;
proportions:
9:
7:
5:
6:
9:
9:
sculpture among punctures,
abdomen brown;
antenna I segments
8:
8:
8:
with mandibles,
ma I ar space testaceous;
12;
I ittle rugose on
clypeus
antennae
legs testaceous;
in following
head shiny,
punctate,
without
anterior half of frons;
frons
without median and lateral keels; occipital carina complete; POL; 5,5; OL ; 2,5; OOL ; 5; OPL ; 5; TL ; 3; scutum, scutellum and metanotum shiny, smooth, finely punctate, without sculpture among punctures; notaul ices almost complete, almost reaching the posterior margin of scutum; propodeum dull,
reticulate rugose,
wing
hyaline,
longer than proximal genitalia missing 1,1,2. LOCUS TYP I CUS:
(14:10);
Ba i yer R.
the (only
or
bands;
longitudinal distal
keels;
fore-
part of radial
two parts forming
an angle;
known
tibial
specimen);
vein
spurs
(New Gu i nea)
holotype M!
in TW.
unknown
DISTRIBUTION:
Fig.
part
transversal
transversal
in the holotype
TYPICAL MATERIAL: HOSTS:
without
without dark
177 -
Australian region:
Male genitalia of
NEW GUINEA:
Baiyer R.,
Lonchodryinus notogeicus half removed.
n.
TW!
sp.
(type):
right
286
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
2. Lonchodryinus notogeicus n. sp.
FEMALE: MALE:
unknown
fully
winged;
length 2,37 mm;
black;
antennae testaceous;
legs testaceous,
with hind coxae proximally brown;
antennae not 8:
5:
7:
8:
distally 8:
8:
8:
thikened; 8:
7:
9;
sculpture among punctures; occipital
carina complete;
scutum shiny, notaul ices
strongly
incomplete,
antennal
between dorsal
frons without median POL = 6;
punctate,
and
OOL = 5;
smooth,
finely
OPL = 5;
length of scutum;
punctate,
without sculpture
without a
transversal
without dark
vein
much
dorsal
longer than
gonoforceps
process and without distal Kogh i
transversal
proximal
(Fig.
TL = 3;
punctures;
fore-wing hyal ine,
proximal
Mt.
without keels;
posterior surface without
the margin of wing;
TYPICAL MATERIAL:
lateral
without sculpture among
reaching approximately 0,75
reaching
HOSTS:
OL = 3;
punctate,
and
posterior surface;
part of radial
spurs 1,1,2. LOCUS TYP I CUS:
in following proportions:
strongly
propodeum reticulate rugose,
longitudinal 'keels; distal
segments
head shiny,
scutellum and metanotum shiny, among punctures;
mandibles testaceous;
177)
part
keel
bands;
(21 :8)
and
without membranous
inner pointed process;
tibial
(New Ca I edon i a)
holotype M!
in B.
unknown
DISTRIBUTION:
Australian
region:
fjEW CALEDONIA:
Mt.
Koghi,
B!
4. Anteon Jurine
Anteon Jurine 1807: 302. Antaeon Haliday 1833: 275, invalid emend. Chelogynus Hal iday 1838: 518 (syn. proposed by Kieffer in Kieffer et Marshall 1905); type species Chelogynus infectus (Haliday 1837), desig. by Muesebeck and Walkley 1951 (desig. of Ch. gaullei by Kieffer
1914b invalid because the
type was not an originally
included species)(discussion on synonymy:
see page 288).
Anteon Jurine partilli: Kieffer in Kieffer et Marshall 1905: 130. Neochelogynus R.C.L. Perkins 1905: 60 (syn. proposed by R.C.L. Perkins 1907); type species Neochelogynus typicus R.C.L. Perkins 1905, orig.
desig.
Prosanteon R.C.L. Perkins 1905: 66 (syn. proposed by R.C.L. Perkins 1907); type species Prosanteon chelogynoides R.C.L. Perkins 1905,
monotypic and orig.
desig.
Paranteon R.C.L. Perkins 1905: 67 (n. syn.); type species Paranteon rnyrrnecophilus R.C.L. Perkins 1905, orig. desig. Chelogynus Haliday: R.C.L. Perkins 1907: 48. Chelogynus Haliday: R.C.L. Perkins 1907: 48.
Anteoninae
287
Anteon Jurine: R.C.L. Perkins 1907: 50. Lasianteon Kieffer 1913b: 300 (syn. proposed by Muesebeck and Walkley 1951); type species Lasianteon rubrifrons (Kieffer 1905), desig.
by Muesebeck and Walkley
1951.
Xenanteon Kieffer 1913b: 300 (syn. proposed by Richards 1939); type species Xenanteon reticulatun, (Kieffer 1905), desig. by Richards 1939.
Liodryinus Kieffer 1913b: 300 (syn. proposed by Richards 1953); type species Liodryinus doddi (Kieffer 1911b), orig. desig. Anteon Jurine: Kieffer 1914b: 140. Chelogynus Haliday: Kieffer 1914b: 172. Xenanteon Kieffer: Kieffer 1914b: 193. Lasianteon Kieffer: Kieffer 1914b: 197. Allanteon Kieffer 1914b: 198 (syn. proposed by Berland 1928); type species Allanteon punctatus (Kieffer 1905), monotypic. Anteon Jurine: Berland 1928: 157. Chelogynus Haliday: Berland 1928: 162. Anteon Jurine: Richards 1937: 118. Chelogynus Haliday: Richards 1937: 118. Anteon Jurine: Richards 1939: 245. Anteon subg. Anteon Jurine: Richards 1939: 246. Anteon subg. Chelogynus Haliday: Richards 1939: 246. Hirtanteon Benoit 1951b: 163 (n. syn.); type species Hirtanteon canabensis Benoit 1951b, orig. desig. Anteon Jurine: J.F. Perkins 1976: 13. Chelogynus Haliday: J.F. Perkins 1976: 18. Chelogynus subg. Chelogynus Haliday: J.F. Perkins 1976: 18. Chelogynus subg. Neochelogynus R.C.L. Perkins 1976: 19. Anteon Jurine: Krombein 1979: 1240. Anteon subg.Anteon Jurine: Krombein 1979: 1240. Anteon subg. Chelogynus Haliday: Krombein 1979: 1241.
Anteon jurineanum Latreille 1809, first included species.
Type species: FEMALE palpi
(Figs.
178,
179):
with 6 segments;
distal
part of radial
fully
labial
winged; palpi
rarely brachypterous;
with 3 segments;
vein much shorter than
occasionally sl ightly shorter,
as
long as,
or
proximal
strong MALE labial
(Fig.
64);
occipital
transversal (Figs. palpi
29,
times as
carina complete;
between
180): fully proximal
dorsal
winged;
with 3 segments;
much shorter than as long as,
keel
part
(Fig.
or longer than proximal
long as broad
transversa I keel
between
(see fig.
178); part;
propodeum usually
with a
and posterior surface. with distal
with 6 segments; part of radial
vein
180); occasionally sl ightly shorter,
part;
176);
with
(Fig.
prominence bearing one
maxi Ilary palpi
fore-wing
part
maxillary
longer than proximal
inner side of the enlarged claw with a proximal bristle
fore-wing
pterostigma less than four
propodeum usually with
dorsa I and posterior surface;
a strong
gonoforceps without
288
Mem.
Fig.
apical
Amer.
178 -
branch wrapping
En!.
Female of
the penis;
Ins!.,
hairs at most as HOSTS:
1984
vertex of the head usually to occipital
long as breadth of antennal
carina;
without two
antenna I
segments.
worldwide
Cicadellidae (including Idiocerinae and Macropsinae)
SPECIES: NOTES:
37,
Anteon gaullei Kieffer
oblique keels connecting posterior ocelli DISTRIBUTION:
no.
156
whereas the females of
Chelogynus Hal iday and Anteon Jurine are
easily separable, the males show less distinct characteristics. The problem of the separation or synonymy of the two genera is resolved only after careful
study of the males.
main difference concerned
the
According
to J.F.
Perkins
length of the temples and
the scape ("head viewed dorsally
with
(1976)
the
the breadth of
temples shorter than breadth of
Anteon; "head viewed dorsally with temples at least as long as breadth of scape" in Chelogynus). I have seen that this difference is not
scape"
in
289
Anteoninae
Fig.
179 -
Female of
Anteon gaullei Kieffer
always visible: in some males of A. infectum (Hal iday) and A. scapulare (Haliday) for instance, the temples are shorter than the breadth of the scape.
Other differences concern
the petiolar area
and
the ninth sternite.
flavicorne, arcuatum), the petiolar area is usually broader and more shiny than in species of Chelogynus. The lucidity, however, seems unimportant since I have seen males of pubicorne and ephippiger with petiolar areas fully smooth and shiny, although these species are Chelogynus. The breadth In European species of
and
Anteon (for instance brachycerum,
the shape of the petiolar area usually
However, I have seen that males, for instance, of A.
are helpful
characteristics.
these characteristics also are variable. The inflatrix Benoit from the Ethiopian region, A.
canadense Ashmead from the Nearctic region, A. thai n. sp. from the Oriental
region have narrow petiolar areas that are rugose,
shape characteristics similar to
Chelogynus,
three species have segment 4 of the front the basitarsus,
proving they are true
dull,
and
whereas the females of the
tarsus at most 0,5 as
Anteon.
long as
In European species of
Anteon, the distal lobe of the ninth sternite is truncated, whereas in Chelogynus it is rounded. I have seen however, that the males of A. canadense (Anteon) have the distal lobe of the ninth sternite rounded. In conclusion I think that Anteon and Chelogynus are synonyms, even if in a single zoogeographic region the two groups can be separated easi Iy (for instance in Europe). The differences, however, are not val id for the world fauna.
290
Mem.
Fig.
180 -
Amer.
Ent.
Inst.,
no.
37,
1984
Male of Anteon tripartitum Kieffer
PALAEARCTIC SPECIES OF
ANTEON
Twenty species are known.
KEY TO THE SPECIES
FEMALES
(type)
Anteoninae
Segment
4
segment
5 of
front
tarsus at
front
most
tarsus
291
0,5 as
with basal
long part
as
segment
longer
1;
than
distal
(see fig. 183); front tarsal segments 3 or 4 produced into a hook • . • • • • • • • • . . • • • . . • • . • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • . • • • • • 2
part
Segment
4
of
front
tarsus
at
segment
5 of
front
tarsus
wi th
than
distal
produced
2
surface
tarsal
Posterior front
3
(see of
segment
surface
tarsal
Scutum shiny,
of
keel
on
into a
centrally
least
mid-length fully
of
keels;
longitudinal for
keels;
hook . . . . . . . . . . . . . 4
into a
except
arabicum n. or
some weak
tracks
of
sp.
reticulate
(see Ethiopian rugose;
never
species)
wi thout
1. jurineanum Latrei lie
2.
occasionally ocellus;
with
fore
a
short
tarsal
median
3
segment .. . . . . . .
brachycerum (Dalman)
carinate;
median keel
frons;
strongly
into a
Head not
fore
complete
tarsal
or
reaching
at
3 or 4 produced
segments
reticulate
reticulate
3.
hook
rugose;
fore
tarsal
segment
4
reticu12tum Kieffer
rugose;
fore
tarsal
3 produced
segment
into
.6
hook • • • • • • • • . • • . • • • . • • . • • . • . . • . . . • . . . • • . . . • . •
Notaulices
reaching
Antennae at
approximately
most
least
4.
fully
smooth
and
5. 6.
median median
rugose,
as
pronotum, and
thorax
scutum.
sp. of
scu tum.... .
...7
or brown . . .
flavicorne (Dalman)
area
of
area
posterior
scutum and 7.
fully
part Iy
of
rugose as
· • • • • • • . . . . . . • • • • .. Head
of
arcuatum Kieffer
SCUlpture,
Propodeum with Head,
length
yellow . . . . . . . . . . . . . . . . . . . ..
wi thout
nearly
nepalense n.
partly black
•••••••••••••••.•••• Propodeum wi th
0,65
reaching mid-length
•....••••.•.....•••• Antennae
9
2
sides . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ..
carinate;
· . . . . . . . . . . . . . . . . . ..
8
sculpture,
the anterior
produced
Notaulices at
7
1;
shorter
hook • • • • • • • • • • • • • • • • • . . • • . • • • • • • . . • • . . • • . • • . . • . • . . • • • . . 5
into a
6
segment
hook...........................
Frons
a
two
3 or 4 produced
20.
not
visible near
Head
or
longitudinal
the
......... ...... .....
5
segment as
hook . . . . . . . . . . . . . . . . . . . 3
granulated
centrally
produced
as
long
tarsal
into a
propodeum with
sculpture . . . . . . . . . . . Frons
front
long as
4 produced
without
................... Scutum dull,
191);
part
propodeum without
segments
irregular keel s
4
fig.
0,66 as
basal
hook . . • • . • • • . • • • . • • . . • • • • . . • • . • . . • • . . . • . . . • . . • 8
into a
Posterior front
part
least
posterior
lateral
surface
shiny,
rugose........ surface
most Iy
.. . . . . . . . 9
dull,
fully
areas • • . • . . . • • . • • • • • . . . • • 12
scutellum yellow or
reddish . . . . . . . .
ephippiger (Da I man)
black;
at
most
clypeus and
frons part Iy
testaceous . . • . . • • • . • • • • • • • • • • . . • • • . . • . • • • • • • . . • • . • • • • • • • • • • • • 10 10
Species very and
fully
segment
5
small,
smooth; of
front
• • • • . • • • • • • • • • • • • •
long
frons
less
tarsus 8.
than
smooth, with
2,5 mm;
without one
head
keels
row of
pubicorne (Da I man)
finely
punctate
or areolae;
lamellae
(Fig.
193) ..
292
Mem.
En!.
Species bigger,
long more
punctate;
with keels
frons
tarsus wi th 11
Amer.
Antennal
two
rows
segments
of
segments
least
(Figs
1,5
head and
13 Frons almost
1984
with
head
segment 195,
strongly
5 of
front
197) •••••••••
times as
long
as
• .11
1. .•.
as
long as
1 or
little
longer •••..
infectum (Ha I i day) joining
slightly visible;
the
ocelli;
posterior keel
lateral
always
thorax black ••••..•••••.•••••••••••••••.••••.
tripartitum K i ef f er
11.
triangle not
or differently
37,
scapulare (Ha I i day)
9. 10.
.. .. .. .. .. .. .. . . . . . Ocellar
3,0 mm,
triangle delimited by keels
keels occasionally visible;
no.
or areolae;
2+3 nearly
. . • • • • • • • • • • • • . •• .• 12 Ocellar
than
lamellae
2+3 at
• . • • • • • • • • • • • • • • • • .. Antennal
Ins!.,
delimited by
keels;
head and
thorax black
colored..............................
fully
reticulate
•••••••••• ••••• •••• Frons never almost
..13
rugose . . . . . . . . . . . . . . .
crassifrons (Moczar)
12. fully
reticulate
rugose;
at
most
anterior
half rugose • . . . . • • • • • • • . . . • • • • • • • . . . . . . . . • . . . • . . • • . • • • • • • • • • • 14 14 Head, thorax and propodeum fully black, occasionally with cupreous tints; head .. .. .. .. .. .. .. .. . . . . Head,
without sculpture among 8. pubicorne (Da I man)
thorax and propodeum partly
or
fully
the punctures . . . . .
yellow,testaceous,
reddish or reddish-dark; occasionally fully brown-dark, almost black or fully black, but in this case head granulated among punctures ••••••••••••••••••••••••••••••.••••..•..•••••••••••• 15 15 Head
fully
black;
scutum fully
reddish,
yellow or
two
of
rows
or mainly black;
testaceous;
lamellae
(Fig.
segment
5 of
pronotum
front
tarsus
with
202) . . . . . . . . . . . . . . . . . . . . . . . . . . .
13. gaullei Kieffer At
least
the head partly
occasionally
head and
yel low,
case pronotum brown-dark, testaceous of
lamellae
16 Head dul I, segment
or yel low; (Figs. always
1 nearly
shiny,
weakly 17 Head, frons
segment
clearly and twice as 14.
punctate,
granulated;
slightly
longer
black
or blackish,
5 of
front
long
testaceous; not
tarsus
but
in
that
reddish, wi th
1-2
rows
4
SCUlpture among punctures
segment
1 nearly as
long as
or very or
........................ .
Pronotum never black; head and body partly or fully reddish, testaceous 7.
antennal
as 4 . . . . . . . . . . . . . . . . . . . . . . . . . . . .
fulviventre (Ha I i day)
without
antennal
than
strongly granulated;
thorax and propodeum fully black, testaceous ... 15. hilare n. sp.
• • • • • • •• • . •
or
or black,
191,204,207) . . . . . . . . . . . . . . . . . . . . . . . . . . . . 16
• • • • • • • • • • • • • • • • • •• Head
reddish
scutum brown-dark
with anterior half
•• 17 of
variably colored, usually or yel low . . . . . . . . . . . . ..
ephippiger (Da I man)
The fema I es of A. ingenuum n. sp., corax n. sp., japonicum n. sp., munitum n. sp. are unknown. Probab I y A. ingenuum, corax or munitum are
Anteoninae
the males of
293
A.hilare and A. nepalense. Only breeding esperiments can
answer the question.
MALES
Posterior
surface
of
propodeum without
longitudinal
keels ••••••
1. jurineanum La t re i I Ie 2
Posterior surface of propodeum with two longitudinal keels ••.• 2 Gonoforceps with no distal inner process more or less pointed
(F i g.
181) • • • • • • • • • • • • • • • • • • • . . • • • • . • • • • • • • . . • • • . . • • • • • • • • • . . • 3
Gonoforceps wi th distal (Fig.
3
Posterior almost
inner process more
or
less pointed
192) •••••••.••••••••••••••••••••••••••.••..•••.•••.••••• 9 surface of
fully
Posterior
smooth,
surface of
propodeum wi th median
area
shiny
and
rugose . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4
not
propodeum with median area dul I
and
rugose
• ••••••••••••••••••••••••••••••••••••••••••••••••••••••••••••• 7
4
Head more
or
punctures;
less
finely
frons without
areolae . . . . . . . . . . . . . . . Head more not
or
less
reticulate
6
18. but
rugose;
with
sculpture among
punctures
resembling
sp. occasionally apparently
strong punctures
resembling
frons . . . . . . . . . . . . . . . . . . . . . . . . . . . 5 Hind femora fully black or brown..... . . . . . . . . . . ......... . . . . . . . . . . . . . . . . . . . . . . . 10. infectum (Hal iday) Hind femora fully testaceous . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6 Head partly
or
region
fully
Head partly
or fully
of
reticulate
9.
granulated •••••••••.•••
16.
rugose,
partly
or
fully
scapulare (Hal iday)
reticulate
· • . . • • . . • . • . . . • • • • • • .•
7
without
strong
munitum n.
reticulate
rugose,
areolae on anterior
5
punctate and areolae or
rugose,
ingenuum n.
not
granulated
sp.
Scutum finely punctate, wi thout sculpture among punctures, wi thout areolae, not granulated •••••••• .•••••••••••••••••.•
• • • • • . • • • • • • • • • • • • . . .. Scutum differently strongly
reticulate
1 7.
corax n.
sculptured, rugose
sp. granulated,
with anterior half
or with posterior half
strongly
punctate ••••••••••••••••••••••••••••••••••••••••••••.••••••••• 8
8
Scutum granulated;
antennae almost
fully
brown;
legs with
tibiae partly brown . . . . . . . . . . . . . . . . . . . . • . . . • • • • . . • . • . . • • . . . • •. 1. jurineanum La t re i I I e
coxae,
femora and
Scutum not granulated, but strongly anterior half and strongly punctate anrJ
legs
fully
testaceous,
. • . • • • • . . . • • • • • • •• 9
Gonoforceps wi thout head not
brown
or black . . . . . . . . .
for hind
1 9. japonicum n.
dorsal
184);
except
reticulate rugose on on posterior half; antennae
punctate,
proximal
granulated; 2.
coxae basally brown.
sp.
membranous process antennae and
brachycerum (Dalman)
(Fig.
legs mainly
294
Mem.
En!.
Gonoforceps wi th a
more or
membranous process
(Fig.
10 Distal
inner process
apical
11
Amer.
margin
of
excavated
Ins!.,
less
extended medially
(Figs.
190) •••••••••••••••••.••• 11
188,
inner process
of
gonoforceps
or
straight
testaceous,
at
Legs
testaceous,
most
wi th
proximal
gonoforceps
margin
tibiae partly brown . . . .
1984
large dorsal
Distal Legs
37,
192) ••.••••••..••••••..••••••••••••• 10
apical
convex
no.
hind 6.
extended apically
(Figs.
192,
legs with
hind
Coxae brown;
coxae,
femora
clubs
occasionally
of
femora
stalks
of
brown
coxae and
femora brown
13 Head not dorsal (Fig. Head
less brown
punctate,
femora brown
strongly granulated
206) . . . . . . . . . . . . strongly or
or
finely
14.
finely of
punctate,
strongly
or weakly
granulated,
15 Head
strongly
weakly
smooth,
granUlated,
205);
wi thout
without
long
proximal (Fig.
short
dorsal
203) . . . . . . . 14
sculpture among
without wi th
sculpture among punctures
short
or
long median keel
on
gaullei Ki effer occasionally with areolae and
tripartitum Kieffer
Head punctate,
sculpture among punctures 8.
proximal
very
sculpture among
granulated;
11.
granulated . . . . . . . . . . . .
hind
med ian kee I on frons. ephippiger (Da I man)
usually 13.
(Fig.
irregular keels . . . . . . . without
mid and
fulviventre (Hal iday)
punctate,
frons . . . . . . . . . . . . . . . . .
. . . . . . • • • • • 13 least
gonoforceps usually
gonoforceps very
punctures, usua I I y wi thou t • • • • • • • • . • • • • • • • • • . . • •. 7. Head more
of
punctate,
occasionally
membranous process
with at
or blackish . . . . . . . . . . . . . . . . . . . . . . . . . . . 15
membranous process
punctures 14 Head
or blackish,
or dark .
extremeties
hind
. • • • • • • . • • . . . • • •• . • . . . . . • . • • . • . • . • • • . • . . . . • • • or
and
flavicorne (Dalman)
coxae and
Legs more
and with
194) . . . . . . . . . . . . 12
. . . . . . . . . . . . . . . . . . . . . . . 5. arcuatum Kieffer 12 Legs fully yellow or testaceous, with proximal of
and with
or weakly
pubicorne (Dalman)
The males of A. reticulatum Kieffer, hilare n. sp., arabicum n. sp., nepalense n. sp. are unknown. Probably A. ingenuum n. sp., corax n. sp. and munitum n. sp. are the ma I es of A. hilare and A. nepalense. Only breeding experiments can answer the question.
1. Anteon jurineanum Latrei lie
Anteon jurineanum Latrei lie 1809: 35 Gonatopus brevicornis Dalman 1818: 85 (syn. proposed by Olmi 1983); locus typicus V astergot I and (Sweden). Dryinus cursor Haliday, in Curtis 1828: 206 (syn. proposed by Olmi 1983); locus typicus England. Dryinus otiartes Walker 1837: 425 (n. syn.); locus typicus England.
Anteoninae
295
Dryinus sisithrus Wa I ker 1837: 426 (n. syn.); locus typicus Eng I and. Dryinus nanus Haliday, in Walker 1837: 427 (n. syn.); locus typicus Wicklow
(Ireland).
Anteon crenulatus Kieffer in Kieffer et Marshall 1905: 141 {n. syn.)i locus typicus Bonar Bridge (Scotland). Anteon thomsoni Kieffer in Kieffer et Marshall 1905: 142 (syn. proposed by Richards 1939); locus typicus Sweden (new name for Dryinus brevicornis Thomson 1860 nec Gonatopus brevicornis Dalman 1818) •
Anteon vicinus Kieffer in Kieffer et Marshall 1905: 145 (n. syn.); locus typicus Dumfries (Scotland). Anteon marginatus Kieffer in Kieffer et Marshall 1905: 147 (syn. proposed by Olmi 1983); locus typicus Maisons-Laffitte (France). Anteon rectus Kieffer in Kieffer et Marshall 1905: 171 (n. syn.); locus typicus Dumfri es (Scot I and) • Anteon scoticus Kieffer in Kieffer et Marshall 1905: 174 (n. syn.); locus typicus Sutherland (Scotland). Anteon jurineanus Latreille: Kieffer 1907b: 26. Anteon nanus (Haliday): Kieffer 1907b: 27. Anteon otiartes (Walker): Kieffer 1907b: 27. Anteon sisithrus (Walker): Kieffer 1907b: 27. Antaeon barbatus Chitty 1908: 142 (n. syn.); locus typicus Bentley Wood
(England).
Xenanteon brunneipes Berl and 1928: 168 (n. syn.); locus typicus Maisons-Laffitte (France).
Dryinus sisithrus Walker: Richards 1935: 147. Anteon crenulatum Kieffer: Richards 1935: 147. Anteon scoticum I
296
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
FEMALE: fully winged; length 2,00-3,12 mm; black; mandibles testaceous, with teeth brown; antennae fully brown or with segments 1-2 or 1-5 testaceous;
legs brown,
testaceous,
with hind coxae partly brown,
with fore tibiae and
and tibiae partly brown or blackish; distally
thickened;
5:
4:
5:
5:
5:
5:
antennal 8;
lateral
tegulae testaceous;
granulate,
OL,
these
OOL,
lateral
OPL,
face with a frontal keels are incomplete;
variable;
posterior surface short,
temples visible;
shorter than scutum
reaching tegulae;
scutum dull,
occasionally only
granulate;
propodeum reticulate rugose,
181
-
longitudinal distal
and Tentsmuir Nat.
keels;
5:
carina complete;
pronotum dull,
metanotum shiny,
and posterior surface;
Res.
fore-wing hyaline,
part of radial
4:
sockets;
pronotal
rugose,
with
tubercles
reaching mid-length of smooth or rugose,
with a strong
transversal
posterior surface without
Male genitalia of Anteon jurineanum Latreille from
(England) (A)
12:
granulate and reticulate rugose;
without sculpture; between dorsal
clubs of femora antennae short,
with or without
occipital
notaul ices incomplete,
scutellum and
Fig.
line,
(4:18);
scutum or shorter;
keel
occasionally
with more or less numerous
keels near orbits directed towards the antennal
occasionally POL,
ligth;
segments in following proportions:
head dull,
areolae or irregular keels;
tarsi
or with coxae,
(Scotland) (8):
without dark transversal
vein much shorter than proximal
New Forest
right half removed.
part
bands;
(3:8);
fore
tarsal segments in proportions: 9: 2: 2: 2: 8; segment 4 of front tarsus produced into a hook; enlarged claw (Fig. 64) with one bristle located on a proximal
prominence;
segment 5 of front
tarsus
(Fig.
64)
with 1 row of
Anteoninae
1-4 lamellae; lamellae; MALE:
in addition
tibial
fully
winged;
Fig.
light;
182 -
apex with a
length 1,56-3,00 mm; legs more or
tegulae testaceous;
black;
dull,
in following proportions:
face with frontal directed
(laterally
viewed)
complete;
line,
6:
with or without
POL = 8;
OL = 4;
sockets;
7:
and fore
of
antennal
Anteon jurineanum
(Sweden).
8:
8:
weakly or strongly
lateral
8:
8:
8:
7:
12;
head
granulated;
keels near the orbits
occasionally
OOL = 7;
scutum dull or shiny,
sculptured;
12:
tarsi
thickened;
with more or less numerous areolae or irregular keels;
towards the antennal
incomplete;
with
antennae not distally
Shapes of gonoforceps
granulate,
mandibles testaceous;
less brown,
Latrei lie from Vastergotland
segments
group of 1-3
spurs 1,1,2.
antennae brown or black; tibiae
numerous bristles;
297
smooth,
OPL = 3;
these
TL = 4;
without sculpture;
lateral
keels are
occipital
carina
occasionally
anterior surface of scutum more strongly
notaul ices incomplete,
reaching mid-length of scutum or
298
Mem.
shorter;
Amer.
Inst.,
scutellum and metanotum shiny,
propodeum reticulate rugose, and posterior surface;
bands;
areas;
(Fig.
181)
without membranous proximal apex very rounded
long
(Fig.
smooth,
longitudinal
1984
without sculpture;
transversal
and narrow
keels;
fore-wing
distal of radial
gonoforceps
37,
keel
posterior surface usually without
area as rugose as lateral transversal
no.
with a strong
with 2 distinct
occasionally
(3:9);
Ent.
process;
keels;
this case with median
hyaline,
without dark
with
part
inner pointed process and
gonoforceps,
182);
182); occasionally
dorsal
vein much shorter than proximal
without distal
(Fig.
in
between
longitudinal
in
lateral
occasionally broad,
view,
with
pointed or
intermediate characteristics;
tibial
spurs 1,1,2. lOCUS TYPICUS:
Paris (France)
lectotype M! (designated by Olmi 1983) of A. jurineanum in SP; lectotype F! and 2 paralectotypes MM! (designated by Olmi 1983) of G. brevicornis in ST; holotype F of D. cursor in ME; holotype M! of D. otiartes in BM; holotype F! in of D. sisithrus in BM; lectotype M! of A. nanus in DU; holotype F! in of A. crenulatus in BM; holotype F! of A. vicinus in BM; lectotype M! (designated by Olmi 1983) of A. marginatus in P; holotype F! of A. rectus in BM; holotype M! of A. scoticus in BM; holotype M! of A. barbatus in BM; holotype F! of X. brunneipes in P; type of A. thomsoni not seen (probably in lU; not found TYPICAL MATERIAL:
in ST). HOSTS: in U. S. S. R. name A. brevicorne). DISTRIBUTION:
Oncopsis flavicollis l . (Ponomarenko 1978: under
Palaearctic region:
JAPAN:
Sapporo (male determined as
Anteon japonicum by Ashmead: undescribed species), WA! MONGOLIA: 3 km N form Somon Uenc aimak),
BU!
Ol!
lothar
OT!
Ignalina
(Chovd
NEPAL:
aimak),
Pulchauki
(Birganj),
(lietuva),
EB!
BU!
SO from Somon Bajanzogt
(Katmandu),
OT!
27° 58'
Ol!
OT!
N 85° 00'
other localities
Godavari
E,
Ol!
(Central
(Katmandu),
OT!
U.
S.
in Ponomarenko (1978);
S.
Ekenas, HE! lOjo, HE! SWEDEN: Vastergotland, ST! Hoor distr. (SK.), Ring Sjo, (SK.), BM! Degaberga (SK.), BM! Dagstorp sjo. (SK.), BM! Rostanga Duka BU!
(SK.),
BM!
(Csoroghegy), Rakos.,
BU!
Skaralid BU!
Nagykovacsi,
Duczfalva,
BU!
Nagyvisnyo
(Fenyofo),
BU!
ROMANIA:
Chodau Gora
(Bohemia),
(Karawanken,
AUSTRIA:
BM!
(SK.),
BM!
Mogyoroskert BU!
Rev.,
(BUkk),
Tasnad
Sornmeringgebeit
BU!
HU!
Engadine Nat.
Park
BU!
Kolasin
Tata,
BM!
BU! BU!
BU!
BM!
OT!
BM!
HUNGARY:
Bukk h. Omassa,
Asvany,
BU!
Kisszepalma
CZECHOSLOVAKIA:
Halle,
(Reichenau distr.),
(Grigioni),
BU!
(Crna Gora),
GERMANY:
Prater (Wien), P! Wien, ZO! DENMARK: CO! Ostiylland, CO! HOllAND: Utrecht, lO!
Narvik,
Budapest,
(Maramures),
YUGOSLAVIA:
Siovenija),
NORWAY:
(Bakony hgs.),
R.:
FINLAND:
HU!
Huben
BM!
Kranjiska
Mecklenburg, (0 Tirol),
HU!
HU!
Korsor, CO! Blldso, CO! Danmark, Ol! SWITZERLAND: Boussens (Vaud), FRANCE:
lille (Nord),
P!
Fontainebleau, P! Maisons-laffitte (Seine-et-Oise), P! Paris, SP! Morlaix (Fl.), BM! ENGLAND: Hengistbury Head (SH), Ol! Epping Forest (Essex), CA!
Coombe Wood
(Surrey),
BM!
(Surrey),
Southampton,
P!
Chobham Cornm.
BM!
New Forest,
(Surrey),
BM!
Ol!
Esher Cornmon
Farnham Cn.
(Bucks.),
Anteoninae
BM!
Bentley Wood
(Surrey),
BM!
Risborough Aldbury (SH),
(Suffolk East),
Rammamex Heath BM!
Whaddon
(Herts.),
BM!
Romsey
BM!
Gt.
Brickhi II
(BX.),
BM!
Lustleigh
(SO.,
Oartmoor,
BM!
Slough
(EK.),
BM!
(Romsey,
I<ew Gans.
BM!
(SH),
BM!
BM!
Ivinghoe
BM!
BM!
BM!
Boxmoor
Chase,
BM!
(HT.),
SH.),
Horsley
BM!
P. BM!
Lyndhurst
Beaconsfield (Exmoor), BM!
BM! Wye
Kent's Oak
Wendover
BM!
(HT),
(Surrey),
BM!
BM!
(New Forest,
(New Forest,
BM!
Porlock Oistr.
(Nairn),
BM!
Bricket Wood
(Bucks.),
BM!
Holland's Wood
Lyndhurst-Brockenhurst Rd.
BM!
Newton,
Wimbledon Common
Darnaway Forest
(London),
(Berks.),
BM!
Cannock
Oartmoor),
(HU.),
BX.),
Chase (BX.),
(Bucks.),
Woodwalton
SH.),
Silwood Park
(Brickhill,
(8X.),
(Bucks.),
BM!
299
(BX.),
SH.),
BM!
BM!
SCOTLAND:
Grantown
(Moray), BM! Tentsmuir Nat. Res. (Fife), BM! Inchnadamph N.R. (N.S.), Bf,~! Sutherland, BM! Aviemore (Inverness), BM! Loch Marech (Ross), BM! Kinlochewe
(R.W.),
BM!
Bonar Bridge,
(Inverness),
BM!
Lochinver.
N.R.
BM!
Gairloch
(RVV.),
Glen.,
8M!
Kincraig
Dowry
(Wicklow),
Glen
(Co. I'll),
DO),
BM!
S.
BM!
BM!
SPAIN:
(RW.),
(EI.),
Wicklow,
Manzarron
Limonetto (Cuneo), NOTES:
according
species without propodeum)
~~r.
Gretna,
~~ethy
DU!
(Co.
(Murcia), SZ!
BM!
Killin
TW!
BM!
Loch Morlich
Dumfries,
Bridge
Glenasmole
WI),
B/.l!
BM!
BM!
(EI.),
(Perthshire),
BM!
(Co.
Beinn Eighe
BM! DU),
Leixoes,
(Bussoleno),
Cavruber
IRELANO:
Tipperary Wa.
PORTUGAL:
Pietrabianca
BM!
Devilis
(~~ewcastle,
OLl
Torino),
ITALY: SZ!
GE! to Richards
longitudinal
(1939)
keels on
was composed of only
cursor Haliday) however,
BM!
BM!
Kippure Ho.
Benedetto Belbo (Cuneo),
BM!
(Sutherland),
the brevicorne group
(the complex of
the posterior surface of the
two species,
and marginatum Kieffer.
brevicorne (Dalman) (=
According
to J.F.
Perkins (1976),
the brevicorne group was composed of three spec i es,
cursor was considered a separate species.
Differences in
because
these species,
however,
were hardly recognizable with the keys of Richards and J.F.
Perk ins.
Nei ther author actua II y
saw
the type of marginatum,
and both
based their keys on unrel iable original descriptions. I found and studied the type of marginatum in the National Paris Museum and also studied numerous other specimens of the brevicorne group kept collections.
I
in European
have always found specimens exhibiting characteristics
intermediate to those I isted
in the keys for separating
the three species.
Variations are present in the proportions of the sides of the ocellar triangle, the frons, exist
I
breadth of the ocell i, frontal
think
keels,
sculpture of the scutum,
OPL,
sculpture of
colour and gonoforceps. Since intermediate forms
that the supposed brevicorne group
only one species (brevicorne Dalman)
which
is really composed of
is very variable.
As proposed
(1983), however, brevicorne (Dalman) is a junior synonym of jurineanum Latrei lie (nec jurineanum Latrei lie in Richards 1939 and J.F. by Olmi
Perkins 1976). This synonymy
is based on
the discovery of the type of
Anteon jurineanum Latrei lie in the Spinola collection. The classic jurineanum of Richards and J.F. Perkins has become Anteon arcuatum Kieffer. Another interesting variation
in the morphology of the brevicorne
300
Mem. Amer.
Ent.
Ins!.,
no.
37,
1984
group concerns the posterior surface of the propodeum. This is the main area used in identifying brevicorne - absence of longitudinal keels in this group, presence in the other species. I, however, have seen some males of the brevicorne group with tracks of two longitudinal keels, and modified my keys accordingly.
2. Anteon brachycerum (Dalman)
Dryinus brachycerus Dalman 1823: 12. Dryinus lyde Walker 1837: 427 (syn. proposed by Richards 1939); locus typicus Eng I and. Anteon nigricornis Kieffer in Kieffer et Marshall 1905: 172 (syn. proposed by Richards 1939); locus typicus Eccles (Scotland). Anteon triareolatus Kieffer in Kieffer et Marshall 1905: 175 (syn. proposed by Richards 1939); locus typicus Bonar Bridge (Scotland). Anteon brevicollis Kieffer in Kieffer et Marshall 1905: 176 (syn. proposed by Richards 1939); locus typicus Bonar Bri dge (Scot I and) • Anteon flavitarsis Kieffer in Kieffer et Marshall 1905: 176 (syn. proposed by Richards 1939); locus typicus Clober (Scotland). Anteon indivisus Kieffer in Kieffer et Marshall 1905: 199 (syn. proposed by Richards 1939); locus typicus Bishopton (Scotland). Anteon nigroclavatus Kieffer in Kieffer et Marshall 1905: 205 (syn. proposed by Richards 1939); locus typicus Kilsyth (Scotland). Anteon curvatus Kieffer in Kieffer et Marshall 1906: 517 (syn. proposed by Richards 1939); locus typicus Sutherland (Scotland). Anteon obscuricornis Kieffer in Kieffer et Marshall 1906: 523 (syn. proposed by Richards 1939); locus typicus Eccles (Scotland). Anteon brachycerus (Dalman): Kieffer 1907b: 25. Anteon nigroclavatus Kieffer: Cameron 1907: 33. Antaeon suffolciensis Chitty 1908: 144 (syn. proposed by Richards 1939); locus typicus Felden (England). Anteon curvus Kieffer 1914b: 155 (syn. proposed by Richards 1939); locus typicus Sutherland (Scotland). Anteon neglectus Kieffer var. suffolciensis (Chitty): Kieffer 1914b: 162.
Anteon Anteon Anteon Anteon Anteon Anteon
lyde (Walker): Richards 1935: 151. indivisum Kieffer: Richards 1935: 151. obscuricorne Kieffer: Richards 1935: 151. brachycerum (Dalman): Richards 1939: 250. brachycerum (Dalman): J.F. Perkins 1976: 16. brachycerum (Dalman): Ponomarenko 1978: 18.
FEMALE: fully winged; length 2,3-3,3 mm; black, with mandibles yellow (except for teeth brown) and legs brown (with tarsi and junctions light);
Anteoninae
Fig.
301
8
Chela of Anteon brachycerum
183 -
A:
tegulae testaceous;
apical
part;
antennae short,
(Dalman)
B:
basal
from Dowry
(Ireland);
part.
with segment 1 nearly twice as long
as 4; head dull, smooth, granulated; frontal line absent; scutum shiny, punctate, hairy, smooth; notaulices distinct, incomplete, reaching midlength of scutum; sculpture;
scutellum and metanotum smooth,
propodeum reticulate rugose,
between dorsal longitudinal
and posterior surface;
keels;
a
shiny,
transversal
without irregular keel
posterior surface with 2
median area very shiny
almost smooth or weakly rugose; radial
with
(more than
fore-wing hyal ine,
vein much shorter than proximal
part;
lateral
areas),
with distal
segment 1 of front
part of tarsus
much longer than segment 4; fore tarsal segment 3 produced into a hook; segment 5 of front tarsus (Fig. 183) with nearly 10-13 bristles; apex with a group of 4
lamellae; enlarged claw
(Fig.
183)
on a proximal prominence; tibial spurs 1,1,2. MALE: fully winged; length 2,3-3,3 mm; black, mandibles yellow and junctions
(except for teeth brown);
light;
tegulae testaceous;
with one bristle located with antennae brown
and
legs fully brown or with tarsi
head smooth,
dull,
granulated;
302
Fig.
Mem.
184 -
Amer.
Ent.
Ins!.,
no.
37,
Male genital ia of Anteon brachycerum
1984
(Dalman)
from Burnham
Beeches (Eng I and) •
frontal
line absent;
scutum shiny,
hairy,
punctate,
smooth;
notaulices
distinct, incomplete, reaching mid-length of scutum; scutellum and metanotum shiny, without sculpture; propodeum reticulate rugose, with transversa I i rregu I ar keel surface with 2
longitudinal
lateral
smooth;
areas),
keels;
fore-wing
much shorter than proximal inner pointed process,
between dorsa I and posterior surface; median area very shiny hyaline,
part;
with distal
gonoforceps
without proximal
dorsal
(Fig.
(more than
part of radial
184)
vein
with a distal
membranous process;
spurs 1,1,2. LOCUS TYPICUS: Smaland (Sweden) TYPICAL MATERIAL: lectotype M! and 1 para lectotype F!
a
posterior
tibial
(designated by
MO':Lar 1967) of D.
brachycerus in ST; holotype M! of A. suffolciensis in BM; holotype F! of A. nigricornis (= A. obscuricornis) in BM; lectotype M! of A. lyde in BM; holotype F! of A. triareolatus in BM; holotype F! of A. brevicollis in BM; holotype F! of A. flavitarsis in BM; holotype M! of A. indivisus in BM; lectotype M! of A. nigroc1avatus in BM; holotype M! of A. curvatus (= A. curvus) in BM. HOSTS: in England Oncopsis spp. (J.F. Perkins 1976); in U. S. S. R. Oncopsis flavicollis L. (Ponomarenko 1978). DISTRIBUTION:
Palaearctic region:
U.S.S.R.
(Ponomarenko 1978);
Smal and,
ST!
V I!
Umea.
NEPAL:
FINLAND:
meadow
L.
Godavari
(Katmandu),
Rago (Estl.),
(Vasterb.),
LE!
HE!
Stock hoi m,
OT!
SWEDEN: TW!
Ri ngsjon,
303
Anteoninae
Fig.
185 -
Chela of
Anteon reticula tum Kieffer from
M.
Peglia
(Italy).
304
TW!
Mem.
NORWAY:
ROMANIA
Laugesund,
HU!
En!.
Ins!.,
HUNGARY:
37,
no.
1984
Nagyszenas
(Budai
H.),
1972); GERMANY: Mecklenburg, HU! Annatal
(Nagy
Rheinland),
Amer.
OTt
DENMARK:
Biskelund,
CO!
Bildso,
CO!
BU!
(Siebengebirge,
FRANCE
(Richards
1939); ENGLAND: Felden (Herts), BM! Burnham Beeches (Bucks.), BM! Wimbledon Common Bishopton BM!
(Sussex),
(Renfrew),
BM!
BM!
VI!
Epping Forest
Kilsyth
Bonar Bridge (Sutherland South),
Sutherl and,
BM!
IRELAND:
Portmanock
(Dublin),
BM!
Dowry
BM!
BM!
CAl
SCOTLAND:
Clober
(Stirling),
Eccles (Dumfries),
(Wick low),
Woodbrook,
(Essex),
(Stirling), BM!
Athdown
BM!
(Wick low),
BM!
BM!
3. Anteon reticulatum Kieffer
Anteon reticulatus Kieffer in Kieffer et Marshall 1905: 140. Xenanteon reticulatus (Kieffer): Berland 1928: 168. Anteon reticulatum Kieffer: Richards 1939: 256.
FEMALE:
fully
testaceous
winged;
length 4,25-4,56 mm;
black,
(except for segments 6-10 brown);
coxae proximally black, femora brown;
with antennae
legs testaceous,
hind coxae almost fully black,
tegulae testaceous;
twice as long as segment 4;
antennae short,
head shiny,
with fore
clubs of hind
with segment
1 almost
fully strongly reticulate rugose;
frontal line present; pronotum rugose; scutum shiny, smooth, punctate, without sculpture, among punctures; notaulices distinct, incomplete, reaching approximately 0,3 length of scutum; punctate or without punctures; sculpture;
propodeum reticulate rugose,
between dorsal longitudinal
and posterior surface;
keels;
scutellum shiny,
metanotum shiny,
smooth,
smooth,
without
with a transversal
irregular keel
posterior surface with 2
median area of posterior surface strongly rugose,
with
numerous transversal keels; fore-wing hyaline, with distal part of radial vein much shorter than proximal; segment 1 of front tarsus nearly three times as long as segment 4;
segment 4 produced into a hook; 185) with 4 bristles; apex with a group of 6 lamellae; enlarged claw (Fig. 185) with one bristle located on a proximal prominence; tibial spurs 1,1,2. segment 5 of front
MALE:
tarsus
fore tarsal
(Fig.
unknown
LOCUS TYPICUS:
Maisons-Laffitte (France)
TYPICAL MATERIAL:
holotype F!
in P.
HOSTS: unknown DISTRIBUTION: Palaearctic region: Oise), NOTES:
P!
ITALY:
only
Monte Pegl ia
FRANCE:
(Perugia),
two known specimens
Maisons-Laffitte (Seine-etOLl
!
4. Anteon nepalense n. sp.
Anteoninae
FEMALE:
fully
testaceous, thickened; 6:
6:
5:
frontal
length 3,0 mm;
antennal
antennae brown;
punctate,
pronotum shiny,
shiny,
smooth and
scutum
(6:13);
POL
~
5;
two OL
~
without sculpture;
the orbits;
~
3,5;
OOL
fore-wing hyaline,
radial
vein much shorter than proximal
in following proportions:
tegulae;
MALE:
8:
3:
2:
unknown Godavari
TYPICAL MATERIAL: HOSTS:
4:
~
TL
scutum shiny,
part
with 2
area as rugose as bands;
(3: 11);
fore
enlarged claw
prominence;
finely
scutellum and metanotum
median
13;
with 1 row of 15 lamellae;
LOCUS TYPICUS:
OPL
notau I ices i ncomp lete,
without dark transversal
one bristle located on a proximal spurs 1,1,2.
4,5;
propodeum reticulate rugose,
areas;
186)
6:
posterior surface shorter than
keels on posterior surface;
(Fig.
5:
keels around
tubercles reaching
without sculpture;
tibial
6:
lateral
without scu I pture among punctures;
longitudinal
9:
with anterior surface rugose and posterior surface
pronotal
smooth,
6:
4;
reaching approximately 0,65 length of scutum; shiny,
10:
without sculpture among punctures;
frons also with
carina complete;
legs
antennae distally
segments in following proportions:
head shiny,
I ine present;
puncta te,
black;
with coxae and clubs of femora darkened;
6;
occipital 3;
winged;
305
lateral
distal tarsal
(Fig.
part of segments
186)
segment 5 of front
apex with a groc'p of 7
with
tarsus
lamellae;
(Nepal)
holotype F!
in OT;
1 paratype F!
in OL.
unknown
DISTRIBUTION:
Palaearctic region:
NEPAL:
Godavari
(Katmandu),
OT!
Ol!
5. Anteon arcuatum Kieffer
Anteon arcuatum Kieffer in Kieffer et Marshall 1905: 144. Anteon imberbis Kieffer in Kieffer et Marshall 1905: 149 (n. syn.); locus typicus Rannoch (Scotland). Anteon arcuatus Kieffer: Kieffer 1907b: 25. Anteon flavicorne (Dalman) var. bensoni Richards 1939: 255 (n. syn.); locus typicus Brickell Wood (England). Anteon jurineanum Latreille partim (only F): Richards 1939: 256 (discussion on synonymy:
see page 299).
Anteon jurineanum latreille: J.F. Perkins 1976: 16. Anteon arcuatum Kieffer: Olmi 1983: 80. Anteon arcuatum Kieffer: Moczar 1983b: 198.
FEMALE:
fully
winged;
length 2,18-3,00 mm;
black,
with antennae brown
(sometimes partly brown and testaceous); legs testaceous, with coxae partly brown and clubs of hind femora brown; tegulae testaceous; mandibles yellow, nearly
with teeth brown;
twice as long as segment 4;
antennae short, head shiny,
with segment
smooth,
1
without sculpture
306
Mem.
Fig.
186 -
or weakly alutaceous,
Amer.
Chela of
with
En!.
Ins!.,
no.
37,
1984
Anteon nepalense n. sp. (type)
irregular keels at
least near the occipital
triangle; frons with frontal I ine and 2 lateral longitudinal keels near orbits; pronotum rugose; scutum smoooth, shiny, punctate on anterior half,
almost without punctures on posterior half;
incomplete,
approximately
metanotum shiny,
smooth,
notaulices distinct,
reaching mid-length of scutum; without sculpture;
scutellum and
propodeum reticulate rugose,
with a transversal irregular keel between dorsal and posterior surface; posterior surface with 2 lateral longitudinal keels; median area dull, rugose, nearly as rugose as lateral areas; fore-wing hyal ine, with distal part of radial vein much shorter than proximal part; segment 1 of front tarsus much hook;
longer than segment 4;
segment 5 of front
apex with a group of 4
tarsus
fore tarsal
(Fig.
lamellae;
187)
tibial
segment 3 produced
into a
with 1 row of 8-18 bristles;
spurs 1,1,2.
Anteoninae
Fig.
187 -
MALE:
fully
with dorsal
Chela of
winged;
Anteon arcuatum Kieffer from Portmanock {Ireland}
length 2,31-2,50 mm;
part of the segmen ts brown;
brown;
mandibles yellOW,
brown;
femora brown;
testaceous;
307
with
teeth brown;
sometimes only
head dull or shiny,
black;
antennae testaceous,
somet i mes on I y
segments 1-3
legs testaceous,
hind femora brown;
granulated,
with coxae
tegulae
with numerous areolae or
irregular keels; frontal line present; frons without lateral longitudinal keels; scutum shiny, smooth, without sculpture; hairy, punctate on anterior half, distinct,
almost without punctures on
incomplete,
metanotum shiny, with a
transversal
posterior half;
reaching mid-length of scutum;
smooth,
without sculpture;
irregular keel
posterior surface with 2
propodeum reticulate rugose,
between dorsal
longitudinal
keels;
notaulices
scutellum and
and posterior surface;
median
area shiny,
less
308
Fig.
Mem. Amer.
188 -
Ent.
Inst., no.
37,
1984
Male genitalia of Anteon al'cuatum Kieffer from Esher Common (England)
rugose than lateral areas; fore-wing hyal ine, with distal part of radial vein much shorter than proximal part; gonoforceps (Fig. 188) with distal inner process pointed and extended medially; its apical margin excavated; dorsal proximal membranous process present; tibial spurs 1,1,2. LOCUS TYPICUS: Eccles (Scotland) TYPICAL MATERIAL: holotype F! of A.
al'cuatum in BM; holotype F! of A. flavicol'ne var. bensoni in BM; holotype F! of A. imbel'bis in BM. HOSTS: in I ta I y Rhytidodus decimusqual'tus (Schrk.). DISTRIBUTION: Palaearctic region:
MONGOLIA:
15 Km S from Dalanzadgad
(Gurban Sajchan ul., SLid gobi aimak), BU! U.S.S.R. (Ponomarenko 1978: under name jUl'ineanum); FINLAND (Hellen 1953); HUNGARY: Batorliget, BU! Tiszang, BU! Asvany, BU! Osborokas (Bugac), BU! GERMANY: Ferchland (Elbe), HU! Eisteraue, HA! AUSTRIA: Eisenzicken, VI! DENMARK: Legind, CO! Gejlbjerz, CO! HOLLAND: Den Haag, LE! SWITZERLAND: Brig (Valais), HE! ENGLAND: Esher Common (Surrey), BM! Brickett Wood (Herts), BM! Ham Common, BM! Bix (Henley, Oxfordshire), BM! Siapton (Linslade, BX.), BM! Starks Pond (Kingsteignton, SO.), OL! SCOTLAND: Eccles, BM! IRELAND: Portmanock
(Dublin),
BM!
ITALY: Alfiano Natta (Alessandria),
Anteoninae
309
EN!
6. Anteon flavicorne (Dalman)
Gonatopus flavicornis Dalman 1818: 83. Anteon sericeus Kieffer in Kieffer et Marshall 1905: 198 (n. syn.); locus typicus Maisons-Laffitte (France). Anteon flavicornis (Dalman): Berland 1928; 159. Anteon sericeus Kieffer: Berland 1928: 161. Anteon flavicorne (Dalman): Richards 1939: 253. Anteon levigatus Kieffer: Haupt 1941: 40. Anteon flavicornis (Dalman): Haupt 1941: 42. Anteon subflavicornis Haupt 1941: 43 (n. syn.); locus typicus Be II i nchen 312) •
Anteon Anteon Anteon Anteon Anteon
FEMALE:
(Germany)( discussion on synonymy:
flavicorne (Dalman) : subfl a vicorne Haupt: flavicorne (Dalman) : subflavicorne Haupt: flavicorne (Dalman) :
fully
winged;
see notes on page
J. F. Perkins 1976: 17. J. F. Perkins 1976: 18. Ponomaren ko 1978: Olmi
18.
and Currado 1979a:
Currado 1983:
length 2,18-4,00 mm;
345.
127.
black;
antennae yellow,
sometimes with fuscous dorsal markings; legs yellow, with hind coxae partly black and stalk of hind femora brown; tegulae testaceous; mandibles yellow,
with teeth brown;
nearly
twice as long as segment 4;
weakly
granulated,
frontal
line present;
with few
antennae short, head shiny,
with segment
punctate,
irregular keels near occipital
1
strongly or carina;
frons with 2 lateral complete or incomplete keels
near orbits directed towards the antenna I sockets; strongly punctate on anterior third,
without
scutum shiny,
smooth,
punctures on posterior
surface; notaulices distinct and incomplete, reaching approximately 0,30 length of scutum; scutellum and metanotum shiny, smooth, without sculpture;
Fig.
propodeum reticulate rugose,
189a -
Propodeum of Anteon
with a
transversal
keel
between
flavicorne (Dalman) (A) and subflavicorne
Haupt
(B).
310
Fig.
Mem. Amer.
189b -
Chela of
Ent.
Ins!.,
no. 37,
1984
Anteon flavicorne (Dalman) from Courmayeur (Italy)
Anteon inae
dorsal
and posterior surface;
311
posterior surface with 2 long i tudi na I keels;
median area smooth, rugose near the margins, projecting or not forward beyond anterior margin of lateral areas (Fig. 189a: this difference, used in
identifying A.
flavicorne and subflavicorne, is only individual
in Anteon;
variation
it
is present
also in other species,
for
instance A.
ephippiger); fore-wing hyaline, with distal part of radial vein much shorter than segment 4; tarsus
(Fig.
lamellae; proximal MALE:
proximal
part;
fore tarsal 189b)
prominence;
punctate;
apex with a group of 8
with one bristle located on
length 2,00-2,62 mm;
black;
legs testaceous,
tegulae testaceous;
granulated,
scutum shiny,
incomplete,
189b)
smooth,
finely
posterior surface with 2 longitudinal fore-wing
much shorter than proximal
mandibles yellow,
with teeth
keels;
median area shiny,
with distal
gonoforceps
(Fig.
its apical
dorsal proximal membranous process present; LOCUS TYP I CUS: Vastergot I and (Sweden) lectotype F!
(hereby
scutellum and
propodeum reticulate rugose, dorsal and posterior surface;
hyal ine,
part;
notaulices distinct,
length of scutum;
process pointed and extended medially;
TYPICAL MATERIAL: (4 MM, 1 F)! of G.
antennae testaceous,
with coxae and clubs of
punctate;
reaching approximately 0,3
partly rugose;
a
fully or partly reticulate rugose and
metanoturn shiny, smooth, finely punctate; with a transversal irregular keel between smooth,
longer than
segment 5 of front
spurs 1,1,2.
markings;
femora partly brown; head dull,
(Fig.
tibial
winged;
with fuscous dorsal brown;
into a hook;
with 1 row of bristles;
enlarged claw
fully
segment 1 of front tarsus much
segment 3 produced
almost
part of radial
190)
with distal
vein inner
margin excavated;
tibial
designated)
spurs 1,1,2. and 5 paralectotypes
flavicornis in ST; lectotype M! (hereby designated) of A. sericeus in P; holotype F! and 2 para types FF! of A. subflavicornis in HA. in England Idiocerus sp. (J.F. Perkins 1976), Populicerus laminatus (Flor) (~ Idiocerus laminatus) and P. populi (L.) (~ Idiocerus populi) (Chambers 1955); in Germany Populicerus confusus (Flor) (~ I diocerus confusus) (Haupt 1941); in I ta I y Rhytidodus decimusquartus (Schrk.), Populicerus albicans (Kbm.), Tremulicerus distinguendus (Kbm.) (~ Idiocerus distinguendus) , Idiocerus stigmaticalis Lewis (Currado 1983); in U. S. S. R. Populicorus laminatus (Flor) (~ Idiocerus laminatus) (Ponomarenko 1978). PARASITES: in England and Switzerland Ismarus flavicornis Thomson HOSTS:
(Nixon
1957;
Chambers 1955).
DISTRIBUTION:
Palaearctic region:
U.
S.
S.
R.
(Ponomarenko 1978);
FINLAND (Hellen 1953); SWEDEN: Vastergotland, ST! Sweden, VI! HUNGARY: Ujpest, BUI Szolnok, BU! SzSztMiklos, BU! CZECHOSLOVAKIA: Chodau (Bohemia),
BM!
GERMANY:
(Kurland),
HU!
Holzhausen
(Halle), AUSTRIA:
HA!
Thuringen,
Voslau,
Kobenhavn,
CO!
P!
Niederlehme (Kyswusterh), (Leipzig),
GE!
EB!
Mecklenburg,
Prater (Wien),
SWITZERLAND:
P!
Chancy
Halle, EB!
Graz,
HA!
HU!
Jelowka-Lepel
Dolauer Heide
Bellinchen P!
(Geneve),
(Oder) ,
DENMARK: GV!
Ecogia
Korsor,
HA! CO!
(Geneve),
GV!
312
Mem.
Amer.
\"\
Ent.
Inst.,
no.
37,
1984
: .
"\
Fig.
190 -
Male genitalia of Anteon (England):
Mies M.
(Vaud),
(Vaud),
Crespian
La!
La!
(Gard),
Moremont
FRANCE: LE!
flavicorne (Dalman) from Oxford
right half removed.
(Vaud),
La!
La Ferte-Milon
La Sauge (Vaud), (Aisne),
P!
Maisons-Laffitte (Seine-et-Oise),
in Kieffer et Marshall
1905);
ENGLAND:
Kings Wood
OL!
Romand s.
Lille (Nord), P!
(BD.),
Cretai I BM!
P! (Kieffer
Wood
Walton Fen (Cambs.), BM! Northwood, OX! Oxford, aT! BM! Monks Wood NNR (Cambs.), BM! SCOTLAND: Sutherland, BM! IRELAND (Richards 1939); ITALY:
S.
Benedetto Belbo
S. Giorio (Torino), in Olmi
EN!
(Cuneo),
Ol!
and Currado 1979a);
Pl!
Courmayeur ALGERIA,
Alfiano Natta (Aosta)
(= A.
(Alessandria),
EN!
subflavicorne Haupt
LN!
the flavicorne group was flavicorne (Dalman) and subflavicorne Haupt. About jurineanum see notes on page 299 (jurineanum sensu Richards 1939 and J. F. Perk i ns 1976 is rea II y Anteon arcuatum Kieffer). About subflavicorne, accord i ng to J. F. Perk i ns (1976), the mai n characteristic used in identifying flavicorne and subflavicorne was the NOTES:
according to J.F.
Perkins
(1976)
composed of three species: jurineanum,
shape of the petiolar area ("petiolar area projecting forward beyond anterior margin of lateral areas" in flavicorne; "petiolar area not projecting"
in subflavicorne). According to me the different shape of the
petiolar area is only an in other species;
variation in Anteon; it's present also ephippiger (Dalman) can show petiolar
individual
for instance A.
Anteoninae
313
area projecting or not forward beyond anterior margin of lateral An other interesting characteristic used in
concerns antenna I segment 3 ("segment 3 about three times as broad"
in flavicorne;
areas.
identifying the two species
"segment 3 about 2,5 times as
long as
long as broad"
in
subflavicorne). The proportions of the antennal segments, however, are very variable in Anteon: in bred series I have seen antennal segments variously long and broad (see also notes on page 322 about Anteon pubicorne). Other characteristics proposed by J. F. Perkins (1976) (sculpture of petiolar area and frons;
Anteon.
I
think that subflavicorne
colour)
are very variable
in
is a junior synonym of flavicorne.
7. Anteon ephippiger (Da I man)
Gonatopus ephippiger Dalman 1818: Gonatopus collaris Dalman 1818: 82 1939); locus typicus Vastergot I and Dryinus facialis Thomson 1860: 177
81. (syn.
proposed by Richards
(Sweden). (n. syn.);
locus typicus Got I and
(Sweden) •
Anteon albidicollis Kieffer in Ki effer et Marsha II 1905: 157 (n. syn. ); locus typicus Volosca (I ta I y) • Anteon rubrifrons Kieffer in Kieffer et Marshall 1905: 158 (n. syn.); locus typicus K is-Pose (Hungary). Anteon collan's (Dalman): Kieffer 1907b: 25. Anteon facialis (Thomson): Kieffer 1907b: 26. Chelogynus rufovariegatus Berl and 1928: 164 (syn. proposed by Richards 1939); locus typicus France. Anteon fulviventre (Haliday) partim (only M): Richards 1939: 265. Anteon ephippiger (Dalman) partim (only F): Richards 1939: 267. Anteon albidocolle Kieffer: Richards 1939: 270. Chelogynus ephippiger (Dalman) partim Haupt 1941: 47. Anteon flaviscapus Jansson 1950 partim (only M): 221 (n. syn.); locus typicus unknown. Anteon rubrifrons Kieffer: Moczar 1967: 297. Anteon albidicolle Kieffer: Olmi and Currado 1976b: 73. Chelogynus ephippiger (Dalman): J.F. Perkins 1976: 22, 23. Chelogynus ephippiger (Dalman): Ponomarenko 1978: 21. Chelogynus albidocollis (Kieffer): Ponomarenko 1978: 21. nec Chelogynus ephippiger (Dalman): Moczar 1979: 77. Anteon pyonganensis Moczar 1983a: 184 (n. syn.); locus typicus Pyongyan
(Korea).
Anteon ephippiger (Da I man): Moczar 1983b: 198.
FEMALE: fully winged; length 1,81-2,87 mm; colour variable; head yellow or reddish (light or dark); occasionally black or brown; antennae yellow
Mem.
314
Fig.
191
-
Chela of
Amer.
En!.
Ins!.,
Anteon ephippiger
no.
37,
(Dalman)
1984
from Benicasim
(Spain)
or testaceous; occasionally with segments 1-2 testaceous and 3-10 brownish; pronotum whitish or yellow; occasionally reddish or reddishdark with brown spots;
propectus yellow or reddish;
occasionally
blackish; scutum yellow or reddish; occasionally brown or black; scutell um and metanotum as colored as scutum; mesop leura and metap leura
Anteoninae
bl ack or redd i sh; reddish; slightly
abdomen black or brown;
tegulae testaceous;
longer than segment 4
punctures, shiny,
propodeum black;
legs yellow; hairless,
315
(8:6);
antennae long,
head shiny,
smooth,
without sculpture;
scutum,
smooth,
shiny,
without sculpture;
notaulices distinct,
0,3-0,5
length of scutum;
irregular keel smooth;
keels;
proximal
part;
and posterior surface;
median area rugose,
fore-wing hyaline,
with distal
segment 1 of front
fore tarsa I segment 2 produced 191)
with
prominence; MALE:
fully
testaceous,
(Fig.
tibial
winged;
keels;
Fig.
black;
head shiny,
without frontal
smooth,
antennae brown;
(F i g.
line;
notaulices distinct,
smooth,
without
frons without incomplete,
scutellum and metanotum shiny, with a
lateral
right half removed.
without
reaching
smooth,
transversal
Male genitalia of Anteon ephippiger (Dalman) (Spain):
legs
tegulae testaceous;
strongly punctate on anterior half,
propodeum reticulate rugose,
192 -
tarsus
apex with a group of 3 lamellae;
length 1,75-2,50 mm;
punctures on posterior half; sculpture;
shorter than segment 4;
segment 5 of front
with one bristle located on a proximal
with teeth brown;
mid-length of scutum;
shiny and
vein shorter than
1,1,2.
finely punctate,
scutum shiny,
reaching
transversal
posterior surface
at most with hind cOXae proximally black;
mandibles yellow, sculpture,
191)
spurs
with a
occasionally
tarsus slightly
1 row of 16-19 lamellae;
enlarged claw
incomplete,
part of radial
into hook;
pronotum
scutellum and metanotum
propodeum reticulate rugose,
between dorsal
with 2 longitudinal
1
without
without sculpture or weakly alutaceous;
smooth,
occasiona II y
with segment
vithout
irregular keel
from Nerja
316
Mem.
between
Amer.
Ent.
Inst.,
dorsa I and posterior surface;
no.
37,
1984
posterior surface with 2
longitudinal keels; median area rugose; fore-wing hyaline, with distal part of radial vein much shorter than proximal part; gonoforceps (Fig. 192)
with distal
inner pointed process and with proximal
membranous process; LOCUS TYPICUS:
tibial
dorsal
spurs 1,1,2.
Vastergotland
(Sweden)
of G. ephippiger in ST; holotype F! of G. collaris in ST; holotype F! of A. albidicollis in VI; holotype F! of A. rubrifrons in BU; holotype F! and 2 paratypes FF (one specimen from Prechaq is an A. fulviventre according to Richards 1939) of Ch. rufovariegatus in P; holotype F! of D. facialis in ST; holotype of A. flaviscapus not found. HOSTS: in Germany Macropsis sp. (Haupt 1941); in U. S. S. R. Psammotettix striatus L. (Ponomarenko 1978, under name Ch. albidocollis). TYPICAL MATERIAL:
DISTRIBUTION: Prov.),
BU!
holotype F!
Palaearctic region:
Ryesan
Somon Uenc
(Chovd aimak),
BU!
23 Km NW from Somon Chutag
BU!
Mongol ia
Nora BU!
Ekenas,
(Stockholm), Karansebes,
BM! BU!
HE!
U.S.S.R.
SWEDEN:
HUNGARY: Holies,
(South Pyongan
MONGOLIA:
(Namnan ul
(Ponomarenko 1975a),
HE!
Pyongyan BU!
SO from Somon Bajanzogt
BU!
Tvarminne,
KOREA:
(Ryang-gang Prov.),
BUkk h,
(Central
BU! BU!
ST!
Tabdi, N.
aimak),
Bulgan aimak),
(Ponomarenko 1978);
Vastergotland,
Budapest,
BU!
Gebirge,
3 Km N from
BU!
Enyed,
FINLAND:
Gotland, Mt.
BU!
ST!
Meszes,
Kis-Pose,
BU! Tatatovaros, BU! Bartia, BU! Orsz Miklos, BU! R. Szombat, BU! CZECHOSLOVAKIA: Carlsbad, P! ROMANIA (Nagy 1973); BULGARIA: KUstendil,
BU!
Dalmacija: VI!
GERMANY:
Neue MUhle, Langeland (Vaud),
YUGOSLAVIA:
I.,
OL!
Bellinchen
HU!
(Oder) ,
AUSTRIA:
CO!
St.
Kostanjika
(Istra),
BM!
of Anteon zernyanus Fahringer,
type F!
Wien,
Kobenhavn,
Giez (Vaud),
HA! P!
CO! GV!
Aachen, Prater
HU!
Mainz,
(Wien),
SWITZERLAND: FRANCE:
Gruz (= Gravosa; undescribed species), P!
OT!
Auvernier,
Le Puy,
P!
Halle,
HU!
DENMARK: LO!
Huelgoat,
La Praz BM!
Palavas (Herault), GV! ENGLAND: Spratton (Northants), BM! Wimbledon Comm. (London), OT! Esher Common (Surrey), BM! IRELAND (R i chards 1939);
SPAIN:
Giorio
(Torino),
Benicasim EN!
(Castellon),
Volosca
BM!
(Trieste),
Nerja
VI!
(Granada),
TURKEY:
BM!
Smyrna,
ITALY:
S.
WA!
8. Anteon pubicorne (Dalman)
Gonatopus pubicornis Dalman 1818: 87. Dryinus tenuicornis Dalman 1823: 13 (n. syn.); locus typicus Vastergotland
(Sweden).
Gonatopus cephalotes Ljungh 1824: 267 (n. syn.); locus typicus Smaland
(Sweden).
Dryinus lucidus Haliday, in Curtis 1828: 206 (n. syn.); locus typicus England. Dryinus penidas Walker 1837: 423 (n. syn.); locus typicus Britain.
Anteoninae
317
Dryinus alorus Walker 1837: 425 (n. syn.); locus typicus Britain. Chelogynus alorus (Walker): Haliday 1838: 518. Antaeon alorus (Walker): Marshall 1873: 7. Anteon azorus (Walker): Cameron 1907: 33. Anteon fuscoclavatus Kieffer in Kieffer et Marshall 1905: 196 (n. syn.); locus typicus Cadder (Scot I and) • Anteon triangularis Kieffer in Kieffer et Marshall 1905: 197 (n. syn.); locus typicus Bonar Bridge (Scotland). Anteon divisus Kieffer inK ieffer et Marsha II 1905: 200 (n. syn.); locus typicus Mugdock (Scotland). Anteon vulgaris Kieffer in Kieffer et Marsha II 1905: 202 (n. syn.); locus typicus Dumfries (Scotland). Antaeon breviventralis Ch i tty 1908: 144 (n. syn.); locus typicus Walmer (England).
Antaeon delicatulus Chitty 1908: 215 (n. syn.); locus typicus Ranworth Broad
(Eng I and).
Anteon subarcticus Hellen 1935: 7 (nomen nudum). Anteon alorus (Walker): Richards 1935: 151. Anteon fuscoclavatum Kieffer: Richards 1935: 151. Anteon penidas (Walker): Richards 1935: 151. Anteon vulgare Kieffer: Richards 1935: 151. Chelogynus serratus Maneval 1935: 6 (n. syn.); locus typicus Mont Mezenc
(France).
Anteon lucidus (Haliday) partim (only F): Richards 1939: 270. Chelogynus exiguus Haupt 1941: 52 (n. syn.); locus typicus Bellinchen
(Germany)(discussion on synonymy:
see notes on page
322).
Chelogynus ephippiger (Dalman) partim (only M): Haupt 1941: 47. Anteon flaviscapus Jansson 1950 partim (only F): 221 (n. syn.); locus typicus unknown (discussion on synonymy: see notes on page 322) •
Anteon subarcticus Hellen 1953: 96 (n. syn.); locus typicus Ivalo (Finland) •
Anteon pubicornis (Dalman); Moczar 1967: 299. Anteon cephalotes (Ljungh): Olmi and Currado 1975: 51. Chelogynus lucidus (Haliday): J.F. Perkins 1976: 22. Anteon cameroni Kieffer: Olmi and Currado 1979a: 347. Anteon lucidum (Haliday): Olmi and Currado 1979a: 347. Anteon pubicorne (Dalman): Moczar 1983a: 184. Anteon pubicorne (Dalman): Moczar 1983b: 198. Anteon mongolicum Moczar 1983b: 198 (n. syn.); locus typicus Bogdo ul.
(Mongolia).
FEMALE: fully winged; length 2,00-2,81 mm; black; antennae brown, with segments 1-2 or only 1 yellow; sometimes antennae testaceous, with segments 3-6 brown; legs fully testaceous; sometimes with clubs of hind
318
Mem. Amer.
Fig.
193 -
Chela of
Ent.
Inst.,
no.
37,
1984
Anteon pubicorne (Dalman) from Genova (Italy)
femora and clubs of femora brown; tegulae testaceous; mandibles yellow, with teeth brown; antennae long, with segment 1 slightly longer than segment 4; proportions of the antennal segments very variable (see notes on page 322); head shiny, smooth, finely or strongly punctate, without sculpture among punctures;
frons usually with a short frontal
line;
Anteoninae
sometimes
(in small
specimens)
pronotum strongly punctate,
without frontal
shiny;
reaching mid-length of scutum; keel
longitudinal shiny,
proximal;
with distal
fore tarsal
tarsus
(Fig.
5-6 lamellae;
193)
long as segment 4; (Fig.
193)
sometimes
segment 1 of front
segment 5 of front apex with a
with a proximal
bearing one bristle;
tibial
length 1,37-2,87 mm;
winged;
irregular
vein much shorter than
into a hook;
with one row of 20- 23 lamellae;
enlarged claw
smooth,
posterior surface with 2
part of radial
MALE:
fully
with a transversal
occasionally rugose near the margins;
segment 2 produced
tarsus approximately 0,66 as
punctate,
incomplete,
median area usually rugose and dull,
smooth and without keels;
fore-wing hyaline,
keels present;
finely
scutellum and metanotum shiny,
and posterior surface;
keels;
lateral
smooth,
notaul ices distinct,
propodeum reticulate rugose,
between dorsal
line;
scutum shiny,
without sculpture among punctures; finely punctate;
319
group of
prominence
spurs 1,1,2. black;
antennae brown or
black; sometimes with antennal segment 1 yellow; legs brown or black,with tarsi and junctions ligth; tegulae testaceous; mandibles yellow,
with teeth brown;
variable sometimes
Fig.
proportions of the antennal
(see notes on page 322); (in small
194 -
specimens)
Male genitalia of Ditton
head shiny,
smooth,
finely
segments very
with sculpture variable; punctate,
without sculpture
Anteon pubicorne (Dalman) from Thames
(England):
right half removed.
320
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
among punctures; sometimes (in large specimens) rugose or smooth, strongly or finely punctate, with areolae or weak keels, without sculpture or keels among punctures; notaulices distinct,
and metanotum shiny, rugose,
with a
surface; shiny,
scutum hairy,
incomplete, smooth,
transversal
shiny,
finely punctate;
irregular keel
fore-wing hyaline,
shorter than proximal
part;
with distal
gonoforceps
pointed process and with
proximal
spurs 1,1,2. LOCUS TYPICUS:
(Sweden).
Smaland
TYPICAL MATERIAL:
finely punctate;
holotype M!
(Fig.
dorsal
of G.
scutellum
propodeum reticulate
between dorsal
posterior surface with 2 longitudinal
rugose;
smooth,
reaching mid-length of scutum;
keels;
and posterior
median area dull
part of radial 194)
or
vein much
with distal
inner
membranous process;
tibial
pubicornis in ST; lectotype M! and
2 paralectotypes MM! (hereby designated) of D. tenuicornis in ST; holotype F! of G. cephalotes in LU; holotype M! of A. breviventralis in BM;
lectotype F!
of D.
BM;
holotype F!
of Ch.
lucidus in DU; holotype F! of A. delicatulus in serratus in P; holotype M! of D. penidas in BM; holotype M! of A. vulgaris in BM; holotype F! of Ch. exiguus in HA; holotype of A. flaviscapus not found; lectotype F! and 2 paralectotypes FF! (hereby designated) of A. subarcticus in HE; holotype M! of A. fuscoclavatus in BM; holotype M! of A. divisus in BM; lectotype M! of D. alorus in DU; holotype M! of A. triangularis in BM. HOSTS: in Germany Macropsis sp. (Haupt 1941: host of a male! determined by Haupt as Ch. ephippiger); in England Streptanus sordidus (Zett. ), Psammotettix con finis (Dah lb. ), Euscelis plebejus (F a II. ) , Macrosteles sp. (Waloff 1974, 1975), Psammotettix nodosus (Rib.) (Richards 1948: under name lucidum). D I STR I BUT I ON: BU!
Road
plateau, Central
Pa I aearctic region:
to Mt.
Pektusan
Ryang-gang Prov.), aimak),
BU!
KOREA:
Hyesan
(Ryang-gang Prov.),
(24 Km NW from Sam-zi-yan, BU!
MONGOLIA:
Bogdo ul.
SO from Somon Bajanzogt
(Central
Chann-Pay (Ulan-Baator, aimak),
BU!
8 Km
W from Somon Urdtamir (Changaj Gebirge, Archangaj aimak), BU! 7 Km NW from Somon Chanzargalant (Bulgan aimak), BU! U.S.S.R. (Ponomarenko 1978: under name lucidum); FINLAND: Landiskar (Ab Nagu), HE! Lovisa, HE! Paanajarvi, HE! Lulto (Lps.), HE! Ivalo (W.-H.), HE! SWEDEN: Ljungby
(Sm.),
OT!
Eksharad
(Vmld.),
OT!
Fjellfota sjo.
(SK.),
BM!
Messaure, TW! Mausgnsbyn, OT! Solna (Bergshamra, Up.), BM! Smaland, ST! LU! Genarp (SK.), BM! NORWAY: Opheimsbakken (Aurdal), CO! Hatfjelddal,
HU!
POLAND:
Koscieliska Valley
(Tatra Mts.),
BM!
HUNGARY:
Budapest, BU! Retyezat, BU! HUvosvolgy, BU! Lesencistvan, BU! Nagykovacsi, BU! Matrasztistvan, BU! Bakony, BU! Vacz (Tudosdomb),
BU!
Borzsony hgs., BU! SUmeg (Sarvaly), BU! Holies, BU! Budai H., BU! Kisret (Szabadszallas), BU! Galyateto (Matra), BU! Velem (Koszegy hgs.), BU!
Meszes vgy.
PUspokladany, Matrasztlaszlo, BU!
(Koszeg),
BU! BU!
Szolnok, Gal-ret
CZECHOSLOVAKIA:
BU! BU!
Parad,
BU!
Inares,
Tabd i,
BU!
(Borzsony hgs.),
Bohemia,
TW!
Carlsbad,
BU!
R-Szombat,
Koroscsatoma BU! P!
(Gyula),
Tatatovaros, Bartfa,
BU!
BU!
BU! BU! Szod,
BULGARIA:
Anteoninae
Madara,
BU!
GERMANY: Rostock
Kuleftse,
Bell inchen
(Mecklenburg),
(Rhon-Gbg.), (Munster i. BM!
BU!
HU!
HU!
HA!
HU!
Kurpark
W.),
Weinberg
YUGOSLAVIA:
(Oder),
HU!
(Unna,
Vaake a. OT!
Rh.,
HU!
d.
(Slovenija), Ol!
lohr (Romberg),
Westf.),
Ingelheim a.
Schliersee (Bayern),
Ratece
Mosbrucher Hochmoor (Eifel),
Sendelbach b.
(Gross-Machnow),
Kgswusterh.,
321
HU!
OT!
Weser,
HU!
1m hart bei
Korheide u.
Harz Mts.
Oderberg
i.
M.,
Irsingen,
Gersfeld
Kanal
(Brocken Dist.),
HU!
Paunsdorf
CO!
HU!
Niederlehme b.
(leipzig),
HU!
Moorlake,
HU! HU!
Finkankrg., HU! Eisenach, HU! Roshenberg, HU! Halle, HU! Tegel (Berlin), HU! Fangschleuse (Berlin), HU! Freienwalde (Berlin), HU! Berl in,
HU!
Bot.
Tirol),
HU!
Obergurgl
(Tirol),
(Wien),
DENMARK:
Schneeberg CO!
Ostiylland,
Garten P!
CO!
(Berl in),
HOllAND:
Bennekomsa Meent,
Ol!
lE!
Gryon
SWITZERLAND:
(Vaud), lO!
lO!
GV!
Derborence Mt.
Moremont
Peney,
P!
(Cote-d'Or),
P!
Sollested
(Vaud),
GV!
lO!
GV!
Lolland),
Biithoven
Ardou
(Valais),
BELGIUM:
Buzenol,
GV!
Ferreyres
lO!
(Valais),
GV!
BM!
Auvernier,
GV!
lutry,
),
Korsor,
lO!
FRANCE:
Gorges du Rhoin
(det. Anteon viduum Maneval,
(H.-l.)
Arles-sur-Tech
CO!
lE!
(Vaud),
(Pyrenees Or.
(0.
P!
(Utrecht),
BM!
Follaterres
Huben
Reichenau,
lausanne (Vaud),
GV!
Chambon-sur-lignon P!
(Dyreh,
lE!
AUSTR I A:
P!
S.te Catherine
(Valais),
Arles-Corsavy road
undescribed species),
ZO!
(Stekelhoekduin),
(Vaud),
Valais),
Munchen,
Prater (Wien),
Utrecht,
Rockaje
Bois Noir
(Ardou,
Meczenc,
HU!
OT!
(Pyrenees Or.),
BM!
Dijon
(Cote-
d'Or), P! Biscarosse, P! Maisons-laffitte (Seine-et-Oise), P! Touraine, IN! Brout-Vernet, P! ENGLAND: Byflaet (Surrey), BM! Brabourne (Kent), BM! Wye (Kent), BO! Whaddon (Bucks.), BM! Ranworth Broad (Norfolk East),
BM!
(Surrey),
Colne Point BM!
Walmer
Matfieldlings
(S.
Oxfordshire),
BM!
Spratton
(NE.),
BM!
Boxmoor
(Kent East),
Yorks.),
BM!
(HT.),
Craswall
Chobham Comm.
Thames Ditton
(Northants),
BM!
(Surr-2Y),
BM!
leicester,
Hothfield Bog
(Kent),
BM!
Puttenham,
BM!
Mapledurham
(Oxon),
BM!
Ham Street
(Kent),
(Cambs.),
BM!
Brickleigh Vale
TW!
BM!
BM!
(Surrey), Torquay
Rhinefield
(Devon),
Richmond Park
(Herefordshire) , BM!
(Devon),
(Hants.),
Wimbledon Com.
BM!
BM!
Bix
BM! (Henley, OT!
BM!
(london),
OT!
Monks Wood NNR
Walford's Gibbet
(Quantock
Forest, SS.), BM! Banstead (SR.), BM! Dancer's End (Aston Clinton, BM! Walland Marsh (EK.), BM! Dawlish Warren (Devon), OT! WALES: LI anddona
(A.),
(Perths.),
BM!
Aberfeldy
BM!
SCOTLAND:
Mugdock,
BM!
Cadder
(Perths.),
BM!
Dumfries,
(Dublin),
BM!
Co. Kerry,
Glenasmole
(lanark),
Tentsmuir Nat. BM!
Res.
BM!
TW!
Athdown
Rannoch Forest
(Fife),
Bonar Bridge,
BX.),
BM!
BM!
Birks of
IRELAND:
(Wicklow),
BM!
Royal
Canal (leixlip, KD.), BM! Portmarnock (Co. Dublin), DU! Dowry (Wick low) , BM! Ballyknockan (Wicklow), BM! Agherlow, BM! Merrion Square S.
(Dubl in),
BM!
Benedetto Belbo
Don Bosco (Torino), Genova,
GE!
1979a), GE!
M.
ITALY:
(Cuneo), Pl!
Antola
Aspromonte
Currado and Olmi
Eaux Rousses Pl!
SZ!
la Cassa (Genova)
(Valsavaranche,
Perevagno (Cuneo),
(Torino),
Ol!
Siusi
SZ!
Aosta),
BM!
Castelnuovo
(Bolzano),
BM!
(=A. cameroni K. in Olmi and Currado
(Reggio Calabria),
1972 under name A.
BU!
the following
localities
vulgaris K.) are doubtful:
(in
322
Mem.
Farazzano
(Fori))
Amer.
(Zangheri
Ent.
1969)
Inst.,
no.
37,
1984
and Trieste (Kieffer in Kieffer et
Marshall 1905); ALBANIA: Ipek, BU! GREECE: Kifisia, BM! NOTES: Chelogynus exiguus Haupt 1941 is a very small species with antennal segment 3 nearly as long as segment 2; it's the same species later described by Jansson (1950) as Anteon flaviscapus (nec Anteon
trivia lis Kieffer var. flaviscapus Kieffer 1905) and by Hellen (1953) as Anteon (Chelogynus) subarcticus. I n European collections I have seen many males and females belonging to these forms. specimens with antennal
They are small
segment 3 as long as segment 2.
In the typical
pubicorne the antennal segment 3 is much longer than segment 2. A. exiguus should be considered a valid species. The study of many collections of pubicorne, however, showed that the proportions of antennal segments 2 and 3 were variable. (segment 3 twice as
These proportions vary from 2,00
long as segment 2)
to 1,66;
1,60;
1,50;
1,33;
1,30;
1,25; 1,00 (segment 3 as long as segment 2). This variability is easily visible if a large population from the same local ity is studied. In collections of HU there is a population from Huben showing
separation of Hellen)
(0.
Tirol,
these different proportions of antenna I segments.
Austria)
I think that the
A. exiguus Haupt (= flaviscapus Jansson; = subarcticus
is not proposable,
the range of variability
although the specimens at opposite extremes of
seem very different.
specimens emerge from hosts parasitized by
I
two
think that these small larvae of dryinids.
Usually under these conditions the only emerging specimen (generally one only specimen completes its metamorphosis) or the two emerging specimens are very sma II
and with colour,
scu I pture and antenna I proportions very
different.
9. Anteon scapu lare (Ha I i day)
Dryinus scapularis Haliday, in Walker 1837: 419. Anteon longiforceps Kieffer in Kieffer et Marshall 1905: 151 (syn. proposed by Richards 1939); locus typicus C I amart (France). Anteon carinatus Kieffer in Kieffer et Marshall 1905: 152 (syn. proposed by Richards 1939); locus typicus Maisons-Laffitte (France). Anteon scapularis (Haliday): Kieffer 1907b: 27. Chelogynus carinatus (Kieffer): Berland 1928: 163. Anteon scapulare (Haliday): Richards 1939: 260. Chelogynus longiforceps (Kieffer): Haupt 1941: 53. Chelogynus lanionis Haupt 1941: 55 (n. syn.); locus typicus Bellinchen
(Germany).
Chelogynus scapularis (Haliday): J.F. Perkins 1976: 18.
FEMALES:
fully
winged;
length 3,5-5,0 mm;
part of the frons testaceous;
black;
antennae testaceous,
rarely clypeus and with fuscous dorsal
Anteoninae
Fig.
195,- Chela of Anteon scapulare (Haliday) from Clamart (France)
markings;
antenna I segment
and clubs of femora brown; testaceous;
antennae
long,
antenna I segments 2+3 at shiny,
323
punctate,
10 brown;
legs testaceous,
mandibles yellow, wi th segment
usually with coxae
teeth brown;
1 sl i ghtl Y longer than
least 1,5 times as
with numerous
with
long as segment 1;
irregular short keels,
tegulae segment 4; head
without sculpture
324
Mem.
among keels, and 2
Amer.
Ent.
with frons usually
lateral
Inst.,
no.
granulated;
37,
1984
frons with a
keels near orbits directed towards
median
the antennal
keel
sockets;
pronotum dull, strongly rugose and reticulate rugose, smooth only near the posterior margin; scutum shiny, strongly punctate on anterior half, finely
punctate on posterior half;
reaching mid-length of scutum; finely punctate; keel
notaul ices distinct,
pronotum reticulate rugose,
between dorsa I and posterior surface;
longitudinal
keels;
near the margins;
incomplete,
scutellum and metanotum shiny,
median area smooth, fore-wing hyaline,
with a
transversal
smooth, irregular
posterior surface wi th
shiny,
2
not rugose or rugose only
with distal
part of radial
vein
much shorter than proximal part; fore tarsal segment 2 produced into a hook; segment 5 of front tarsus (Fig. 195) with 2 rows of 30-70 lamellae extending continuously proximal
Fig.
bristle;
196 -
tibial
to
the apex;
enlarged claw
(Fig.
195)
with
1
spurs 1,1,2.
Male genitalia of
Anteon scapulare (Haliday) from Boussens
(Switzerland):
right half removed.
Anteoninae
325
MALE: fully winged; length 2,62-3,25 mm; black; antennae testaceous, with segments 1-6 or 1-5 dorsally fuscous; legs testaceous, with hind coxae almost fully balck; brown;
head shiny,
tegulae testaceous;
punctate,
mandibles yellow,
without sculpture among keels with frons granulated; vertex granulated; pronotum rugose, smooth, half;
frons
dull,
with a median keel
strongly punctate on anterior half,
scutellum shiny, with a
smooth,
transversal
incomplete,
finely punctate;
irregular keel
finely punctate on posterior
propodeum reticulate rugose,
between
dorsal
keels;
not rugose or rugose only near margins; part of radial
inner process,
and posterior surface;
median area shiny,
fore-w;ng
vein much shorter than proximal
with no distal
keels; scutum shiny,
reaching mid-length of scutum;
posterior surface with 2 longitudinal
196)
teeth
sometimes also
and 2 lateral
smooth only near posterior margin;
notaulices distinct,
with
with nUmerous short keels or areolae,
hyaline,
part;
with proximal
smooth,
with distal
gonoforceps
dorsal
(Fig.
membranous
process; tibial spurs 1,1,2. LOCUS TYPICUS: England TYPICAL MATERIAL: lectotype F! of D. scapularis in DU; holotype F! of A. longiforceps in P; holotype F! of A. carinatus in P; holotype F! of Ch. lanionis in HA. HOSTS: in Germany Oncopsis lanio L. (Haupt 1941). DISTRIBUTION: GERMANY:
Palaearctic region:
Halle,
HA!
Bellinchen
HUNGARY:
(Oder),
HA!
BU!
Koszeg,
BU!
Dolauer Heide
Budapest,
(Halle),
HA!
Wiesbaden, HU! Aachen, HU! AUSTRIA, P! HOLLAND: F. Willemsen bijstation Santpoort, LE! SWITZERLAND: Viay (Vaud), LO! Boussens (Vaud),
OL!
Auvernier, Laffitte
La Sarraz
LO!
(Vaud),
FRANCE:
(Seine-et-Oise),
Brickhi II,
BX.),
BM!
LO!
Aclens
Brout-Vernet, P!
Clamart,
Spratton
P!
P!
(Vaud),
Dijon
ENGLAND:
(Northants),
LO!
Peney,
(Cote-d'Or),
P!
GV! Maisons-
Rammamex Heath
(Lit.
BM!
10. Anteon infectum (Haliday)
Dryinus infectus Haliday in Walker 1837: 419. Dryinus inclytus Haliday in Walker 1837: 421 (syn. proposed by Richards 1935); locus typicus England. Dryinus lateralis Thomson 1860: 178 (syn. proposed by Richards 1939); locus typicus Bohuslan (Sweden). Anteon fusiformis Kieffer in Kieffer et Marshall 1905: 166 (syn. proposed by Richards 1939); locus typicus Dumfries (Scotland). Anteon punctatus Kieffer in Kieffer et Marshall 1905: 168 (n. syn.); locus typicus Ma isons-Laffi tte (France). Anteon inclytus (Haliday): I
326
Mem. Amer.
Fig.
197 -
En!.
Ins!.,
no. 37,
1984
Chela of Anteon infectum (Haliday) from Maisons-Laffilte (France)
Anteon ellimani Chitty 1908: 143 (syn. proposed by Richards 1939);
Anteoninae
327
locus typicus Chesham (Eng I and). Chelogynus punctatus (Kieffer): Berland 1928: 166. Anteon infectum (Haliday): Richards 1939: 258. Chelogynus infectus (Haliday): J.F. Perkins 1976: 18.
FEMALE:
fully
winged;
length 3,00-6,43 mm;
with segments 6-10 or 7-10 dorsally fuscous; partly brown,
hind clubs of femora brown,
femora brown;
tegulae testaceous;
antennae short,
black;
antennae testaceous,
legs testaceous,
with coxae
sometimes also clubs of fore
mandibles yellow,
with segment 1 nearly twice as
with teeth brown;
long as segment 4;
segments 2+3 nearly as long as, or sl ightly longer than segment 1; head shiny, smooth, punctate, with frons reticulate rugose; frons with a short median keel and with 2 lateral keels; pronotum rugose, dull, smooth only near the posterior margin; scutum shiny, smooth, punctate; notaulices distinct, incomplete, reaching mid-length of scutum; scutellum and metanotum shiny, smooth, punctate; propodeum reticulate rugose, with a transversal irregular keel between dorsal and posterior surface; posterior surface with 2 longitudinal keels; median area shiny, smooth, rugose only near the margins; fore-wing with a dark transversal band near the pterostigma, with distal part of radial vein much shorter than proximal part; tarsal segment 1 not quite as long as segment 4; fore tarsal segment 2 produced into a hook; segment 5 of front tarsus (Fig. 197) with 2 rows of 12-36 lamellae extending continuously to the apex; enlarged claw (Fig. 197) with one bristle located on a proximal prominence; tibial spurs 1,1,2. MALE: fully winged; length 2,30-3,43 mm; black; antennae testaceous, with segments 1-8 or 1-10 dorsally fuscous; legs testaceous, with coxae partly brown or black and clubs of femora brown; sometimes clubs of fore femora testaceous; hind ti b i ae brown; tegu I ae testaceous; mand ib les yellow, with teeth brown; head shiny, punctate, with numerous short keels; frons with a median short keel and with 2 lateral keels; pronotum dull, rugose; scutum shiny, smooth, strongly punctate on anterior half, finely punctate on posterior half; notaul ices distinct, incomplete, reaching mid-length of scutum; scutellum and metanotum finely punctate; propodeum reticulate rugose, with a transversal irregular keel between dorsal and posterior surface; posterior surface with 2 longitudinal keels; median area shiny, smooth, rugose only near the margins; fore-wing hyaline, with distal part of radial vein much shorter than proximal part; gonoforceps (Fig. 198) with no distal inner process and with proximal dorsal membranous process; tibial spurs 1,1,2. LOCUS TYPICUS: England or Ireland (?) TYPICAL MATERIAL: lectotype F! of D. infectus in DU; holotype F! of D. lateralis in ST; holotype M! of A. ellimani in BM; holotype F! of A. punctatus in P; holotype F! of A. fusiformis in BM; lectotype M! of D. inc1ytus in DU. HOSTS: unknown antennal
328
Fig.
Mem. Amer.
198 -
Ent.
Inst.,
no.
37,
1984
Male genitalia of Anteon infectum (Haliday) (Switzerland): right half removed.
from Boussens
DISTRIBUTION: Palaearctic region: JAPAN: Kamikochi, TW! SWEDEN: Bohusl an, ST! HUNGARY: Badacsony, BU! Harsbokorhegy, BU! Budapest, BU! Mogyoroskest (Bakony), BU! Sranazug (Gyula), BU! ROMANIA (Nagy 1972); GERMANY: Eschwege, TW! AUSTRIA: Graz, P! Konigswarth, GV! DENMARK: Roden Sko.v (Lolland), CO! Kobenhavn, CO! HOLLAND: Putten (Gld.), LE! Rys-Goosterland, LE! SWITZERLAND: Follaterres (Valais), LO! Vitznau, BM! Boussens (Vaud), OL! Aclens (Vaud), LO! La Sarraz (Vaud), LO! Auvernier, LO! Peney, GV! FRANCE: Maisons-Laffitte (Seine-et-Oise), P! Vincennes, P! Foret de Chatillon (Cote-d'Or), P! Morlaix (Fl.), BM! ENGLAND: Chesham, BM! Dorking (Sr.), OT! Rhinefield (Hants), BM! Minsmere Reserve (Suffolk), BM! Fedlen (Herts), BM! Monks Wood (Cambs.), BM! Capler Wood (Fownhope, HF.), BM! Bricket Wood (HT.), BM! Bagley, BM! SCOTLAND: Dumfries, BM! IRELAND: Athdown (Co. Wick low) , BM! ITALY: Mombarcaro (Cuneo), PL! S. Benedetto Belbo
Anteoninae
(Cuneo), NOTES:
Oll
Tempio Pausania
(Sassari),
Oll
the only known specimen from Japan
(length 6,43 mm) 3,00-4,50).
than
329
(a female)
is much bigger
the specimens from western Palaearctic region
A Japanese sUbspecies? probably only
(mm
the study of males can
answer the question.
11. Anteon tripartitum Kieffer
Anteon tripartitus Kieffer in Kieffer et Marshall 1905: 194. Anteon tricarinatus Kieffer in Kieffer et Marshall 1906: 513 (n. syn.); locus typicus Botusfleming (England). Antaeon kiefferi Chitty 1908: 143 (n. syn.); locus typicus Ipswich (England) •
Chelogynus angusticoliis Berl and 1928: 165 (syn. proposed by Richards 1939); locus typicus Maisons-laffitte (France). Chelogynus prehensor Maneva I 1935: 9 (n. syn.); locus typicus Chambon-sur-lignon
(France).
Chelogynus berlandi Richards 1936a: 169 (syn. proposed by Richards 1939); locus typicus Ma isons-laffi tte (France). Anteon kiefferi (Chitty): Richards 1939: 274. Chelogynus silvaticus Ponomarenko 1970: 429 (n. syn.); locus typicus Kamenka (U.S.S.R.). Chelogynus kiefferi (Chitty): J.F. Perkins 1976: 22, 23. Anteon flavicorne (Dalman): Olmi and Currado 1976b: 75. Chelogynus silvaticus Ponomarenko: Ponomarenko 1978: 21.
FEMALE:
fully
winged;
segments 1-2 or only
length 3,12-3,50 mm;
1 yellow;
black;
legs testaceous,
antennae brown,
with
with hind coxae and
clubs of hind femora partly brown; tegulae testaceous; mandibles yellow, with proximal half black; teeth brown; antennae short, with segment, 1 nearly
twice as
long as segment 4;
head shiny,
smooth,
granulated,
with
frons and occiput strongly punctate; vertex not punctate; frons and occiput with numerous irregular keels; ocellar triangle with keels joining ocelli;
frons
rugose,
with a median keel
dull,
scutum shiny,
with transversal smooth,
punctate;
notaulices distinct,
keels;
incomplete,
scutellum and metanotum shiny,
propodeum reticulate rugose,
between dorsal
lateral
pronotum
smooth near the posterior margin;
strongly punctate on posterior half,
sculpture among punctures; length of scutum;
and 2 weak
keels,
with a
surface and posterior surface;
smooth,
transversal
without reaching midfinely irregular keel
posterior surface with 2
longitudinal keels; median area rugose; fore-wing hyaline, sometimes distally darkened, with distal part of radial vein much shorter than proximal
part;
enlarged claw
bearing one bristle;
(Fig.
199)
with a proximal
segment 5 of front tarsus
(Fig.
199)
prominence with 2 rows of
330
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
/
Fig.
199 -
Chela of Anteon tripartitum Kieffer from S. ( Italy)
Giovanni
Ilarione
Anteoninae
27-30 lamellae; tarsus MALE
apex with a group of 7-12 lamellae;
little slightly than segment 1;
a hook;
tibial
(Figs.
331
fore tarsal
segment 4 of front
segment 2 produced
into
spurs 1,1,2.
29,
180):
fully
winged;
length 2,37-3,00 mm;
black;
antennae
brown or black; legs brown or black, with junctions, tarsi and fore tibiae light; tegulae testaceous; mandibles yellow, with proximal half black
and teeth brown;
head dull,
mainly punctate on frons, with keels joining ocelli;
weakly granulated or alutaceous,
with numerous frons
irregular keels;
directed towards the antenna I sockets; strongly punctate on anterior half,
scutum shiny,
notaul ices distinct,
length of scutum;
scutellum and metanotum shiny,
longitudinal
and posterior surface;
keels;
part of radial 200)
vein
with distal
200 -
smooth,
irregular keel
posterior surface with 2 fore-wing hyaline,
much shorter than proximal tibial
punctate
finely
with a transversal
median area rugose;
Trieste
keels
hairy,
reaching 0,30-0,50
part;
with distal
gonoforceps
inner pointed process and with proximal
membranous process; LOCUS TYPICUS:
Fig.
incomplete,
propodeum reticulate rugose,
between dorsal
smooth,
without punctures or finely
on posterior half; punctate;
ocellar triangle
with a median keels and 2 lateral
(Fig.
dorsal
spurs 1,1,2.
(Italy)
Male genitalia of
Anteon tripartitum Kieffer from Colnbrook
(England):
right half removed.
332
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
TYPICAL MATERIAL: holotype F! of A. kiefferi in BM; holotype F! of Ch. silvaticus in PK; holotype F! of Ch. angusticollis in P; holotype F! of Ch. prehensor in P; lectotype M! and 3 paralectotypes MM! (hereby designated) of A. tripartitus in VI; lectotype M! (hereby designated) of A. tricarinatus in BU. HOSTS:
unknown
DISTRIBUTION: PK!
Palaearctic region:
HUNGARY:
Or Sz.
Matrasztistvan, (Nagy
Fu i renda I,
CO!
Korsor,
CO!
CO!
Nyon,
Chambon-sur-Lignon
Pelsocs, BU!
Altbuchhrst,
(Oberfranken),
SWITZERLAND:
U.S.S.R.:
BU!
SzSztMiklos,
1971); GERMANY:
Muggendorf LE!
BU!
Miklos,
HU!
Prater
FRANCE:
(Haute-Loire),
CO!
BU!
(Csakvar),
Scheffau
AUSTRIA:
(Perm Province),
Budaors,
Hajduvagas
North Zea I and,
GV!
Kamenka
BU!
(Bayern),
(Wien),
P!
HOLLAND:
Retyezat, BU! OT! DENMARK:
Putten
(G I d. ),
Maisons-Laffitte (Seine-et-Oise),
P!
ENGLAND:
New Forest
BU!
ROMANIA
(SH.),
P!
BM!
Colnbrook (Bucks.), BM! Botusfleming (Cornwall), BU! Ashford (Kent), BM! Ipswich (Suffolk East), BM! SCOTLAND: Forres (Moray), BM! Gretna, BM! ITALY: S. Benedetto Belbo (Cuneo), PL! Pietrabianca (Bussoleno, Torino), SZ! S. Giovanni Ilarione (Verona), OL! Naturno (Bolzano), OT! Trieste, VI!
12. Anteon crassifrons (Moczar) n. comb.
Chelogynus crassifrons Moczar 1983a: 181.
FEMALE:
fully
winged;
length 3,60 mm;
black,
with mandibles testaceous,
antennae testaceous,with segments 4-10 brown; brown;
antennae weakly distally
following proportions:
17:
10:
thickened;
13:
12:
frons almost fully reticulate rugose,
11:
antenna I segments
10:
dull,
legs testaceous; 10: 9:
rugose;
9:
12;
abdomen in
head with
vertex and occiput
shiny, smooth, finely punctate, without sculpture among punctures; frontal line complete; occipital carina complete; POL = 7; OL = 5; OOL 9; OPL = 8; TL tegulae; scutum,
=
7; pronotum shiny, rugose; pronotal tubercles reaching scutellum and metanotum shiny, smooth, finely punctate,
without sculpture among punctures;
notaulices incomplete,
pronotum nearly as
reticulate rugose;
posterior surface of propodeum with 2 longitudinal
keels;
median area as rugose as
transversal
keel
between dorsal
without dark transversal proximal 4: 5: (Fig.
part
bands;
(6: 13); fore tarsal
long as scutum
reaching mid-
length of scutum;
lateral
areas; vein
propodeum
propodeum also with a
and posterior surface; radial
(20:21);
fore-wing
with distal
hyaline,
part shorter than
segments in following proportions:
12:
12: 32; segment 2 of fore tarsi produced into a hook; enlarged claw 201) with one bristle located on a proximal prominence; segment 5
of front
tarsus
(Fig.
201)
with 2 rows of nearly 23
with a group of nearly 6 lamellae;
tibial
long
spurs 1,1,2.
lamellae;
apex
Anteoninae
Fig. 201 - Chela of Anteon crassifrons (Moczar) (type)
MALE: unknown LOCUS TYPICUS: Za-mo san (Korea)
333
334
Mem.
TYPICAL MATERIAL: HOSTS:
Amer.
holotype F!
Ent.
Inst.,
no.
37,
1984
in BU.
unknown
DISTRIBUTION: Palaearctic region: KOREA: Za-mo san NE from Pyongyan, pyong-sung Prov.), BU!
(Bek-sung-li,
60 Km
13. Anteon gaullei Kieffer
Dryinus dorsalis Nees (the types of Nees are lost: the name Dryinus dorsalis Nees is best ignored; see also nomina dubia): Thomson 1860:
177 (syn.
proposed by Richards 1939).
Anteon gaullei Kieffer in Kieffer et Marshall 1905: 161. Anteon cameroni Kieffer in Kieffer et Marshall 1905: 162 (n. syn.); locus typicus Thornhi II (Scotland). Anteon maculipennis Kieffer in Kieffer et Marshall 1905: 164 (syn. proposed by Richards 1939); locus typicus Smaland (Sweden). Anteon trivialis Kieffer in Kieffer et Marshall 1905: 201 (n. syn.); locus typicus Claddich (Scotland). Antaeon rufulocollis Ch itty 1908: 143 (n. syn.); locus typicus Tubney
(England).
Anteon dorsalis (Nees): Berland 1928: 159. Chelogynus gaullei (Kieffer): Berland 1928: 163. Anteon rufulocolle Chitty: Richards 1935: 150. Anteon maculipenne Kieffer: Richards 1935: 151. Anteon rufulocolle (Chitty): Richards 1939: 262. Anteon cameroni Kieffer: Richards 1939: 275. Anteon gaullei Kieffer: Richards 1939: 277. Chelogynus ephippiger (Dalman) partim (3 FF): Haupt 1941: 47. Chelogynus gaullei (Kieffer): J.F. Perkins 1976: 22. Chelogynus rufulocollis (Chitty): J.F. Perkins 1976: 22. Chelogynus cameroni (Kieffer): J.F. Perkins 1976: 22. Chelogynus rufulocollis (Chitty): Ponomarenko 1978: 21. Chelogynus cameroni (Kieffer): Ponomarenko 1978: 21. Chelogynus gaullei (Kieffer): Ponomarenko 1978: 21. nec Anteon cameroni Kieffer: Olmi and Currado 1979a: 347.
FEMALE
(Figs.
178,
179):
fully
winged
(sometimes fore-wings slightly
shortened); length 2,37-4,00 mm; black; antennae testaceous with segments 7-10 brown; occasionally brown with segments 1-2 or 1-3 testaceous; mandibles yellow, with teeth brown; propectus yellow or black (in the type yellow); pronotum yellow or reddish; scutum black; occasionally partly reddish-dark; femora brown;
legs fully
testaceous,
abdomen brown or black;
antennae variable;
head shiny,
smooth,
strongly punctate on vertex and occiput,
sometimes with stalk of hind
tegulae testaceous;
length of the
strongly punctate on frons, without sculpture among
less
Anteoninae
335
Fig. 202 - 'Chela of Anteon gaullei Kieffer from Dieppe (France)
punctures; frons with a short median keel, without lateral keels; pronotum shiny, strongly punctate on anterior half, finely punctate or without punctures on posterior half; punctate,
incomplete,
reaching mid-length of scutum;
length of scutum; punctate; between
scutum shiny,
without sculpture among punctures;
and posterior surface;
finely
occasionally only
scutellum and metanotum shiny,
propodeum reticulate rugose,
dorsal
smooth,
notaul ices distinct,
with a
smooth,
transversal
reaching 0,3
finely irregular keel
posterior surface with 2
longitudinal keels; median area rugose; fore-wing sometimes shortened, hyal ine or apically infuscate, with distal part of radial vein much shorter than proximal or
part;
longer than segment 4
segment
1 of front
(this characteristic
tarsus as
is very
long as,
variable;
shorter
it cannot
be used as specific character in identifying gaullei, cameroni and rufulocolle); fore tarsal segment 2 produced into a hook; segment 5 of front
tarsus
(Figs.
group of 7-20 located on a
9,
lamellae; proximal
202)
with 2 rows of 18-30
enlarged claw prominence;
(Figs.
tibial
9,
lamellae; 202)
apex with a
with one bristle
spurs 1,1,2.
MALE: fully winged; length 2,12-2,50 mm; black; antennae brown or brown-light; mandibles yellow, with teeth brown; legs testaceous, with hind coxae proximally black and sometimes with stalk of hind femora brown;
tegulae testaceous;
scu I pture among punctures,
head shiny,
strongly punctate,
in some areas rugose;
without
scutum sh i ny,
smooth,
336
Mem. Amer.
Fig.
203 -
Ent.
Inst.,
(Germany):
dorsal
without punctures or finely punctate
notaulices distinct,
incomplete,
scutellum and metanotum smooth,
propodeum reticulate rugose,
with a
and posterior surface;
1984
right half removed.
strongly punctate on anterior half, scutum;
37,
Anteon gaullei Kieffer from Reichenau
Male genitalia of
on posterior half;
no.
shiny,
reaching mid-length of without sculpture;
transversal
irregular keel
posterior surface with 2
between
longitudinal
keels;
median area rugose; fore-wing hyaline, with distal part of radial vein much shorter than proximal part; gonoforceps (Fig. 203) with distal inner pointed process and with dorsal spurs 1,1,2. LOCUS TYPICUS:
S.
Germain
TYPICAL MATERIAL:
proximal
membranous process;
tibial
(France)
holotype F!
of
A. gaullei in P (the supposed type in holotype F! of A. cameroni
DU designated by Richards 1939 is not valid); in BM;
lectotype F!
(hereby designated)
and 3 paralectotypes FF!
of A.
maculipennis in ST; holotype F! of A. rufulocollis in BM; holotype M! of A. trivia lis in BM. HOSTS: in Germany Macropsis sp. (Haupt 1941, under name Ch. ephippiger) . DISTRIBUTION: FINLAND: Gotland,
Palaearctic region:
Nystad, ST!
HE!
HUNGARY:
SWEDEN:
U.
S.
S.
Degaberga
Igenpataka,
BU!
R.
(Ponomarenko 1978);
(SK.),
BM!
CSipkeskut
Smaland,
(Bukk
hgs),
ST! BU!
VI! Alp.
Anteoninae
Kudsir, BU!
BU!
Csevharaszt,
Horvatkimle,
Templomter
BU!
(Tompa),
BU!
Nadudvar (Hortobagy), (Kdezegi
hgs.),
BU!
BU!
Bartia,
Nagyvisnyo,
Emlekerdd
BU!
Kisret
Szabohegy
BU!
BU!
337
Vacz
(Tudosdomb),
Nagyszenas (Budai
(Szeged),
BU!
Oroszmezd,
(Szabadszallas),
(Kdszeg),
BU!
BU!
BU!
H.),
BU!
Pelsocs,
Meszes vgy.
Koszeg,
BU! BU!
Velem
(Koszeg),
BU!
Holics, BU! Budapest, BU! Retyezat, BU! CZECHOSLOVAKIA: Karlovy Vary (Bohemia), P! GERMANY: Bellinchen (Oder), HA! Aachen, HU! Mainz, OT! Naunhof
(Leipzig),
HU!
(Schleswig-Holstein), AUSTRIA: (Wien),
Turnau P!
GV!
B.,
Nyon,
Mesni I-Ie-Roi Gui Ihem
(Steiermark),
(Sealand),
Ulvenhout N.
(Valais),
Reichenau
Kaltenleutgeben
Ruderhegn LE!
Fangschleuse (Berl in),
CO!
(Wien), HOLLAND:
LE!
Wijster
GV!
P!
Huben VI!
WA!
P!
S.
Berl in,
(0.
Tirol),
Waidbruck.
Den Haag,
(Dr.),
Corcelettes
(Seine-et-Oise),
(Herault),
WA!
HU!
(Baden-Wurttemberg),
LE!
Pont du Gard
HU!
(Tirol),
LE!
LO!
HU!
Utrecht,
FRANCE:
P!
ENGLAND:
EB!
P!
Prater
DENMARK:
LE!
Asperen,
Vouvry
Dieppe,
(Seine-et-Oise),
(Gard),
Malente
Halle,
Wien,
SWITZERLAND:
(Vaud), Germain
HU! P!
P!
P!
51.
Bald Hi II
(Lewknor, Oxfordshire), BM! Boxmoor (Herts), BM! Hay tor (Dartmoor, Devon), OT! Finningley (5. Yorks), BM! Thames Ditton (Surrey), BM! Skotover,
BM!
Leicester,
(Berks.),
BM!
Ham Street
TW!
(Kent),
BM!
Wimbledon Com.
Yorks),
BM!
Norfolk,
(Argyll),
BM!
BM!
IRELAND:
Spratton
(Kent),
(Northants),
BM!
Tubney
(London),
SCOTLAND:
Murlough Ho.
OT!
Windsor Forest BM!
Faversham
Hatfield Lings (Dunsville,
Thornhill (S.
BM!
(Berks.),
DO.),
(Dumfries), BM!
BM!
5.
Claddich
Tollymore Park
(Co.
DO.), BM! Kerry, TW! Co. Wicklow, TW! ITALY: 5. Benedetto Belbo (Cuneo), PL! Pietrabianca (Bussoleno, Torino), SZ! S. Mauro (Torino), OL!
Oropa
(Bolzano),
(Vercelli), OT!
MI!
Sestola
Pizzighettone (Cremona),
(Modena),
GE!
BO!
TW!
Monte Peglia
Naturno (Perugia),
OL!
14. Anteon fulviventre (Hal iday)
Dryinus fulviventris Hal iday in Curtis 1828: 206. Dryinus fuscipes Thomson 1860: 177 (n. syn.); locus typicus Stockholm (Sweden).
Anteon ephippiger (Dalman) var. similis Kieffer in Kieffer et Marsha II 1905: 156 (syn. proposed by Richards 1935); locus typicus Ireland.
Anteon gracilicollis Kieffer in Kieffer et Marshall 1905: 165 (syn. proposed by Richards 1939); locus typicus Poss i I Marsh (Scot I and) • Anteon flavinervis Kieffer in Kieffer et Marshall 1905: 192 (n. syn.); locus typicus Dumfries (Scotland). Anteon parvulus Kieffer in Kieffer et Marshall 1905: 193 (n. syn.); locus typicus Dumfries (Scotland). Anteon xanthostigma Kieffer in Kieffer et Marshall 1905: 198 (n. syn. ); locus typicus Dumfries (Scot I and) • Anteon trivialis Kieffer var. flaviscapus Kieffer in Kieffer et
338
Mem.
Marshall
1905: 201
Amer.
(n.
En!.
no.
37,
1984
locus typicus Scotland (holotype with
syn.)
indecipherable local ity
Ins!.,
label).
Anteon parvus Kieffer in Kieffer et Marshall 1906: 523 (n. syn.); locus typicus Dumfries (Scotland). An teon fu sc ipes (T homson ): Kieffer 1907b: 26. Chelogynus alutaceus Richards 1935: 152 (syn. proposed by Richards 1939); locus typicus unknown ("British" in the locality label of the holotype) •
Anteon jurineanum Latreille partim (only M): Richards 1939: 257. Anteon fulviventre (Haliday) partim (only M): Richards 1939: 264. Anteon fulviventris (Haliday): J.F. Perkins 1976: 20, 22.
FEMALE:
fully
winged;
length 2,0-3,0 mm;
or reddish more or less dark; fully
colour variable;
head yellow
occasionally black or brown;
yellow or brown with segments 6-10 or 7-10 yellow;
antennae
pronotum and
scutum yellow or reddish light; occasionally dark or brown; scutellum and metanotum yellow (sometimes with anterior margin of metanotum black)
or reddish;
yellow or black; brown or reddish; with
teeth brown;
segment 4; short;
occasionally blackish;
propodeum black, legs testaceous; antennae short,
head dull,
pronotum
dull,
mesopleura and metapleura
sometimes with sides yellow; tegulae testaceous; with segment
not punctate, granulated;
1 nearly
granulated, scutum
twice as
smooth,
shiny,
abdomen
mandibles yellow frontal
smooth,
long as
line
without
sculpture or weakly alutaceous; notaulices distinct, incomplete, reaching mid-length of scutum; scutellum and metanotum shiny, smooth, without sculpture;
propodeum reticulate rugose,
between dorsal longitudinal distal front
and posterior surface;
keels;
with a
transversal
irregular keel
posterior surface with 2
median area rugose,
dull;
fore-wing
hyal ine,
vein much shorter than
proximal
tarsus s light I y
shorter than segment 4;
fore tarsa I segment 2
produced
into a
hook;
segment 5 of front
tarsus
(Fig.
part;
with
part of radial
204)
segment with
1 of
1 row of
20-21 lamellae; apex with a group of 3-6 lamellae; enlarged claw (Fig. 204) with one bristle located on a proximal prominence; tibial spurs 1,1,2. MALE: fully winged; length 1,87-2,50 mm; black; antennae brown-yellow; mandibles yellow, with teeth brown; legs fully yellow, sometimes with stalks of hind femora brown; granulated, keels;
tegulae testaceous;
without punctures;
scutum shiny,
smooth,
frontal
hairy,
head dull,
line present;
smooth,
frons without
lateral
wothout sculpture or weakly
granulated; notaulices distinct, incomplete, reaching mid-length of scutum; scutellum and metanotum shiny, smooth, without sculpture; propodeum reticulate rugose,
with a
transversal
irregular keel
between
dorsal surface and posterior surface; posterior surface with 2 long i tud i na I keel s; med i an area rugose; fore-wi ng hya line, with d i sta I part of radial 206)
vein much shorter than proximal
with distal
part;
gonoforceps (Fig.
inner pointed process and with proximal
very
short
339
Anteoninae
!
/
..
C~j
~ \_;;1 ..
\
Fig. 204 -
dorsal
Chela of
//
,
/ ~/
/
Anteon fulviventre (Haliday) from Spratton (England)
membranous process;
LOCUS TYPICUS:
/'
)
tibial
spurs 1,1,2.
England
TYPICAL MATERIAL:
lectotype F!
of D.
fulviventris in DU; lectotype F!
Mem. Amer.
340
En!.
Ins!. , no .
37,
1984
•
Fig.
205 -
Head of
Anteon fuiviventre (Haliday) from Bennekomsa Meent (Holland) •
Anteoninae
341
\\V':
I
I
'
\
\
j
I
II) \ \
Fig.
Male genitalia of Anteon fulviventre (Haliday) (England): right half removed.
206 -
from Spratton
(hereby designated) of D. fuscipes in ST; flavinervis in BM; holotype M! of A. xanthostigma in BM; holotype M! of A. parvulus (~ A. parvus) in BM; holotype F! of A. gracilicollis in BM; holotype M! of A. trivia lis var. flaviscapus in BM; holotype F! of Ch. alutaceus in DU; hoi otype F! of A. similis never and 3 paralectotypes FF! holotype M!
of A.
existed (see Richards 1935). HOSTS: in England Nocydia crocea
H.S.
(J.F.
Perkins 1976;
Chambers
1967) • DISTRIBUTION: Palaearctic region: U. S. S. R. ? (Ponomarenko 1978)(in Ponomarenko, f,:oscow and Leningrad collections I haven't seen specimens of fulviventre from U.
S.
s.
R.;
were gaullei or ephippiger); ST!
HUNGARY:
N.
Barszentmihaly,
Veracze, BU!
the specimens determined as fulviventre
FINLAND
BU!
Budapest,
(Hellen 1953); SWEDEN:
Barlangliget, BU!
Gyula
BU!
Nagyenyed,
(Karas Csat.),
BU!
Stockholm, BU! Tihany,
BU!
Mecsek hgs., BU! Tatatovaros, BU! Rev" BU! Visegrad, BU! Lasztonya, BU! CZECHOSLOVAKIA: Kytin (Bohemia), TW! GERMANY: Halle, HA! Mainz, OT! CO!
Niemegk, HU! Thuringen, GE! Finkenkrug (Berlin), HU! Glucksburg, AUSTRIA: Prater (Wien), P! DENMARK: Langeland I., CO! Kobenhavn,
CO!
HOLLAND:
(Oosteinde 34),
Bennekomsa Meent, OL! LE!
SWITZERLAND:
Pullen
Cudrefin
(G I d. ), (Vaud),
LE! CA!
Waarder FRANCE:
342
Mem.
Amer.
Ent.
Maisons-Laffitte (Seine-et-Oise),
Inst.,
P!
37,
no.
Le Puy,
P!
1984
Prechacq
(Landes)
(del.
Ch. l'uioval'iegatus by Berland), P! ENGLAND: Wytham (Berks), BM! Spratton
(Northants),
(Cambs.),
BM!
BM!
Ramsey
Rhinefield
(SH.),
BM!
(Hants),
BM!
Wood Walton
Castor Hanglands
Fen
(Cambs.),
BM!
St.
Catherine's Point (Watershoot Bay, IW.), BM! Torquay (Devon), OT! Burnham Beeches (Buckes.), BM! SCOTLAND: Rannoch Forest (Perths.), BM!
Possil
TW!
Ireland,
Marsh
(Vercelli), road,
(Lanark),
BM!
ITALY:
Benedetto Belbo
DU!
SZ!
S.
Dumfries,
Pizzighettone (Cremona),
High Atlas),
8M!
IRELAND:
(Cuneo),
TW!
MAROC:
Co.
PL! Asni
Wicklow,
SZ!
Mongrando
(Marrakesh
BM!
15. Anteon hilal'e n. sp.
FEMALE:
fully
winged;
antennae testaceousj
length 3,75 mm;
clypeus and
black;
mandibles
anterior half of frons and
testaceous;
legs
testaceOU5,
with stalks of hind femora and hind coxae partly black; antennae slightly distally thickened; antennal segments in following proportions 10:
7: 14:
10: 10: 11:
10: 9: 9: 14; head strongly punctate, without
sculpture among punctures; complete;
POL
=
4; OL
=
frontal
=
4; OOL
I ine present;
7,5; OPL
=
occipital
5,5;
TL
=
carina
8; pronotum
shiny, hairy, punctate, without sculpture among punctures; posterior surface of pronotum nearly as long as scutum (14:14); scutum shiny, punctate,
without sculpture among punctures;
notaul ices
reaching approximately 0,65 length of scutum; smooth,
shiny,
longitudinal areas;
without
sculpture;
propodeum
keels on posterior surface;
fore-wing hyaline,
incomplete,
scutellum and metanotum
reticulate rugose,
with
2
median are as rugose as lateral
without dark transversal
bands;
distal
part of
(9:14); fore tarsal segments in following proportions: 8: 2,5: 4: 16: 28; enlarged claw (Fig. 207) with one bristle located on a proximal prominence; segment 5 of front tarsus (Fig. 207) with 2 rows of 30 lamellae; apex with a group of 5 lamellae; tibial spurs 1,1,2. radial
vein shorter than proximal
MALE:
unknown
LOCUS TYPICUS:
Godavari
TYPICAL MATERIAL: HOSTS:
part
(Nepal)
holotype F!
in OT;
1 paratype F!
in OL.
unknown
DISTRIBUTION:
Palaearctic region:
NEPAL:
Godavari
(Katmandu),
OT!
OL!
16. Anteon ingenuum n. sp.
FEMALE: MALE:
unknown
fully
winged;
length 2,81-3,43 mm;
antennae brown or testaceous;
black;
legs testaceous,
mandibles testaceous;
with
hind coxae basally
Anteoninae
Fig. 207 - Chela of Anteon hilare n. sp.
343
(type)
344
Mem.
Fig.
208 -
Amer.
Male genitalia of
Ent.
Inst.,
no.
37,
1984
Anteon ingenuum n. sp. (type): right half removed.
black;
antennae not distally thickened;
proportions:
12: 7:
9:
7:
7:
8:
8:
without sculpture among punctures, mainly near the orbits; the orbits;
occipital
9:
antenna I segments in following
9:
10;
head shiny,
punctate,
with more or less numerous areolae
frons with frontal
carina complete;
POL
I ine and
=
7;
OL
two
=
3;
lateral OOL
=
keels near 7;
OPL
3; TL = 3,5; scutum, scutellum and metanotum shiny, smooth, finely punctate, without sculpture among punctures; notaulices incomplete, reaching mid-length of scutum; propodeum reticulate rugose, with 2 longitudinal keels on posterior surface; median area smooth, shiny; lateral distal (Fig.
areas rugose; part of radial
208)
without dorsal
inner process;
tibial
LOCUS TYPICUS:
hyaline,
without dark transversal
membranous proximal
part
(3:10);
bands;
gonoforceps
process and without distal
spurs 1,1,2.
Godavari
TYPICAL MATERIAL: MM! in OL. HOSTS:
fore~wing
vein shorter than proximal
(Nepal)
holotype M!
and 3 paratypes MM!
in OT;
2 paratypes
unknown
DISTRIBUTION:
Palaearctic region:
NEPAL:
Godavari
(Katmandu), OT!
OL!
Anteoninae
345
17. Anteon coI'ax n. sp.
FEMALE:
unknown
MALE: fully winged; length 2,5-3,0 mm; black; mandibles testaceous; antennae testaceous-dark; legs testaceous, with hind coxae basally black; antennae not distally
thickened;
antenna I segments
proportions: 12: 8: 11: 10: 10: 10: 10: 11: without sculpture among punctures; frontal complete;
POL = 5;
OL = 3;
OOL = 7,5;
OPL = 3,5;
scutellum and metanotum finely punctate, punctures; scutum;
notaul ices incomplete,
reaching
propodeum reticulate rugose,
posterior surface; hyaline,
shorter than proximal membranous proximal
part
(8:12);
scutum,
approximately 0,65
with 2
bands;
TL = 5;
without sculpture among
median area as rugose as
without dark transversal
in following
10: 12; head shiny, punctate, I ine present; occipital carina
longitudinal lateral
distal
gonoforceps
length of
keels on
areas;
fore-wing
part of radial (Fig.
process and without distal
209)
vein
with dorsal
inner process;
tibial
spurs 1,1,2. LOCUS TYPICUS:
Godavari
TYPICAL MATERIAL: MM!
(Nepal) and 5 paratypes MM!
holotype M!
in OT;
3 paratypes
in OL.
HOSTS:
unknown
DISTRIBUTION: Pulchauki
Palaearctic region:
(I(atmandu),
NEPAL:
Godavari
(Katmandu),
OT!
OL!
OT!
18. Anteon munitum n. sp.
FEMALE: MALE:
unknown
fully
winged;
length 2,12-3,43 mm;
antennae testaceous or brown; black;
black;
legs testaceous,
mandibles testaceous;
with
occasionally testaceous with all coxae and
partly brown,
antennae not distally
following proportions:
9:
6:
8:
8:
thickened;
7,5:
8:
8:
hind coxae basally
all
clubs of femora
antennal
8:
8:
10;
segments in head shiny,
punctate, without sculpture among punctures; frontal line present; occipital carina complete; POL = 7; OL = 4; OOL = 6; OPL = 3; TL = 4; scutum,
scutellum and metanotum finely punctate,
punctures;
notaulices incomplete,
propodeum reticulate rugose,
with 2
surface;
median area smooth,
hyaline,
without dark transversal
shorter than proximal membranous proximal
part
without sculpture among
reaching mid-length of scutum; longitudinal
shiny;
(3:11);
lateral
bands;
keels on posterior
areas rugose;
distal
gonoforceps (Fig.
process and without distal
fore-wing
part of radial 210)
vein
with dorsal
inner process;
tibial
spurs 1,1,2. LOCUS TYPICUS: Godavari (Nepal) TYPICAL MATERIAL: holotype M! and 31
paratypes MM!
in OT;
16
Mem.
346
Amer.
En!.
Ins!.,
~
no.
37,
1984
/
/
Fig.
209 -
Male genitalia of
Anteon corax n. sp. (type): right half removed.
paratypes MM! HOSTS:
in OL.
unknown
DISTRIBUTION: Pulchauki
Palaearctic region:
(Katmandu), OT!
OL!
NEPAL:
Godavari
(Katmandu),
OL!
OT!
347
Anteoninae
I
5'~~
~
,
\
"\
'\
\
\
!
Fig.
210 -
Male genitalia of
Anteon munitum n. sp. (type): right half removed.
19. Anteon japonicum n. sp.
FEMALE: MALE: and dull,
unknown
fully
winged;
legs fully strongly
complete;
POL
length 2,50 mm;
testaceous
reticulate rugose; ~
4;
Ol
~
black;
mandibles,
antennae,
(except for hind cOXae basally black); 3;
OOl
frontal ~
7;
OPl
line absent; ~
4;
occipital
Tl = 5;
scutum
tegulae head
carina dull,
strongly reticulate rugose on anterior half and strongly punctate on posterior half; notaulices incomplete, reaching only approximately 0,25 length of scutum;
scutellum and metanotum shiny,
smooth,
strongly
348
Mem. Amer.
En!.
Ins!.,
no.
37,
1984
B
Fig.
211
punctate; dorsal
-
Male genitalia of Anteon japonicum n. sp. (type): (A: dorsally viewed; B: laterally viewed).
propodeum reticulate rugose,
and posterior surface;
(4:12);
bands;
gonoforceps
and without distal
a transversal
areas;
distal
part of radial
(Fig.
211)
fore-wing
hyaline,
between
inner process;
holotype M!
gonoforceps
JAPAN:
proximal
(laterally viewed)
in B!
HOSTS: unknown DISTRIBUTION: Palaearctic region:
keels;
without dark
vein much shorter than proximal
with membranous dorsal
apex broad; tibial spurs 1,1,2. LOCUS TYPICUS: Tokyo (Japan) TYPICAL MATERIAL:
keel
posterior surface with 2 longitudinal
median area as rugose as lateral transversal
with
gonoforceps
Tokyo,
B.
ETH I OP IAN SPEC I ES OF ANTEON
Thirty seven species are known.
KEY TO THE SPECIES
process with
Anteoninae
349
FEMALES
Species brachypterous Species
2
(Fig.
212) .•
4 of front tarsus slightly shorter, nearly as long as, than basal part of segment 5 (Fig. 223) ••••••••••••• 3 Segment 4 of front tarsus at most 0,5 as long as basal part of segment 5 (Fig. 243) •••••.•••••••••.••.•.•••.••.••••••••••••• 16 Posterior surface of propodeum with two longitudinal keels ••••• longer
. . . . . • . . . . • . . . . • . . . • . . . . • . . • • . . . •• Posterior surface of
4
brachypterum n. sp.
1.
winged •••••••••••••••••..•••.••••.••.•••••••••.• 2
Segment
or
3
fully
Segment
5 of
without
a distinct
front
tarsus with basal
agile n. sp.
2.
propodeum without
tarsus with apical part
inner
region
longer
longitudinal
side almost
(Fig.
217);
than apical
segment
part
keels ••••• 4
straight,
(Fig.
5 of front 217) ••••••
...•........••..•.•...•..........• 3. maritimum (Turner) 5 of front tarsus wi th inner side curvi I inear, wi th a apical region (Fig. 223); segment 5 of front tarsus with basal part shorter than apical part (Fig. 223) ••••••••••• 5 5 Fore-wing wi th distal part of radial vein more than 0,5 as long as proximal part •••••••••••••••••••••••••••••••••••••••••.•••• 6 Fore-wing wi th distal part of radial vein 0,5 or less than 0,5 as long as proximal part . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 6 Pronotum wi th posterior surface longer than anterior surface, more than 0,5 as long as scutum . . . 4. kasaicum (Benoit) Pronotum wi th posterior surface shorter than anterior surface, less than 0,5 as long as scutum . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 Species mostly reddish-testaceous. 5. fiorii n. sp. 7 Species mostly black . . . . . . . . . . . . . . 6. kivuanum (Benoit) 8 Fore-wing with 1-2 dark transversal bands . . . . . . . . . . . . . . . . . . . . . 9 Fore-wing hyaline, without dark transversal bands . . . . . . . . . . . . 12 9 Posterior surface of propodeum sculptured by strong transversal keels, not reticulate rugose; pronotum humped, with posterior surface shorter than anterior surface; head fully granulated, with areolae or irregular keels on vertex and frons . . . . . . . . . . . . • • . • • • • • • • • • . • • • • • • . . • • • . . • • • • • • •• 7. taylori n. sp. Posterior surface of propodeum weakly reticulate rugose, wi th or without transversal keels; pronotum flat, with posterior surface longer than anterior surface; head fully or almost fully granulated, with only weak tracks of areolae . . . . . . . . . . . 10 10 Head and thorax almost fully black ••••••••••••••••••••••••••••• · . . . . . . . . . . . . . . . . . . . . • . . • • . . . . . . • , 8. dayi n. sp. Head and thorax fully or almost fully reddish-testaceous . . . . . 11 11 Propodeum fully reddish-testaceous ••••••••••••••••••••••••••••• • • . • • . . • • . • • • • • . . • • . . . • • . • • • . • • •. 14. townesi n. sp. Propodeum almost fUlly brown ••••• 15. decellei n. sp. 12 Head with at least the frons reticulate rugose ••••••••••••••• 13 Segment
distinct
350
Mem. Amer.
Head
fully
not
reticulate
En!.
not
with
only
reticulate
9.
frons
1984
wi thout
reticulate
rugose;
canabense (Benoit)
reticulate
rugose;
• . . . . . . . . . . . . • . . . . . . . . • . . . . . . . . .• 14 Scutum weakly punctate,
37,
scutum partly
reddish-testaceous.
Head granulated, granulated,
no.
rugose ••••.•••••••••••.•••••••••••• 14
13 Head fully reticulate rugose; species mostly
Ins!.,
rugose;
scutum
species mostly black •••
10.
ugandanum n.
sp.
sculpture among punctures,
not
granu I a t ed . . . . . . . . . . . . . . . . . . . . . . . 11. kawandanum n. sp. Scutum fully or almost fully granulated •••••••••••.••••••.••• 15
15 Species mostly black; nctaulices reaching approximately 0,5 length of
scutum . . . . . . . . . . . . . . . . .
Species mostly
12.
reddish-testaceous;
visible near anterior margin
of
airum n. sp.
notaulices only
faintly
scutum . . . . . . . . . . . . . . . . . . . . . . . . .
· . • . . . . . • . . . . . . . . . . . . . . . . . . • . . . .. 1 3. turneri n. sp. 16 Posterior surface of propodeum without longitudinal keels or with tracks of only two longitudinal keels . . . . . . . . . . . . . . . . . . . 17 Posterior surface of propodeum with two strong longitudinal keels •••••••••••••••••••••••••••••••••••••••••••••••••••••••• 26 0,5 as long as
17 Pronotum with posterior surface very short, scutum or
shorter •••••••••••••••••••••••••••••••••••••••••••• 18
Pronotum with posterior long as
18 Notaul ices weak, scutum;
surface very
long,
more
0,5 as
than
scutum ••••••••••••••••••••••.••••••••••••••••.••••••• 24 species
but fully
reaching approximately
0,5
length
of
testaceous,
with petiole black . . . . . . . . . .. .•....•••.•......•••••••.....•... 16. anthereon Benoit
Notaulices invisible or faintly visible near anterior margin of scutum; species almost fully black . . . . . . . . . . . . . . . . . . . . . . . . . . . 19 19 Fore-wing with
1-2 dark
Fore-wing hyal ine,
20 Head granulated, not
21
transversal
wi thout
dark
reticulate
bands •••••••••••••••••••• 20
transversal
bands . . . . . . . . . . . . 21
rugose •••••••••••••••••••••••••
• • • • • • • • • • . • . • • • • • • • • • . • • • • • • • • ••
17. namibiense n.
Head
18.
reticulate
Head granulated, reticulate Head fully
rugose . . . . . . . . . . . with weak
irregular
sp.
gutturnium (Benoit) short
keels,
not
rugose •••••••••••••••• 19. arabicum n. sp. reticulate rugose ••••••••••••••••••••••••••••••••• 22
22 Scutum fully
reticulate rugose . . . 20. Scutum punctate or weakly granulated,
23 Scutum punctate,
wi thout
fully
reticulate
• • • • • • . • • • • • • • • • • . • . • • • • • • • • • . • •• Scutum weakly
granulated, fully
reticulate
sculpture among punctures;
surface of pronotum almost
pronotum almost
natalense n. sp. not
not
smooth,
21.
rugose.
hoyoi n. sp.
punctate; without
rugose •• 23 posterior
posterior surface of
sculpture ••••.•••••••••••
. . . . . . . . • . . . . . . . . . . . . . . • . . . . . . . . . 22. mingoae n.
sp.
24 Fore-wing wi th a dark subapical transversal band; posterior surface of propodeum almost smooth, wi th weak irregular keels . . . . . . . . . . . . . . . . . . . . . . . . . . . . Fore-wing hyaline,
without
dark
23.
longiscapum (Ceballos)
transversal
bands;
posterior
Anteoninae
surface of
propodeum strongly
reticulate
25 Head fully reticulate rugose ••••• Head
fully
granulated,
weak areolae near 26 Posterior with keels
not
surface of
25.
rugose •••••••••••••• 25
madagascolum (Benoit) rugose
or
only
with
very
medleri n. sp.
propodeum with median area most Iy
only around
smooth,
the margins •••••••••••••••••••••••••••••
••••••••..••.•.••.••••••••.•••..• Posterior
24.
reticulate
the clypeus ••••
surface of
351
26.
hovaBenoit
propodeum with median
with numerous keels also
in
the central
area
fully
rugose,
region . . . . . . . . . . . . . . . 27
27 Head wi th OL longer than POL; fore-wing hyal ine, not darkened .. · • • . • . • • • • • • • . . • • . • • • • • . • • • • • • • .• 27. compressicoxale (B en 0 i t ) Head with OL shorter than POL; fore-wing with 1-2 dark transversal bands •••••••••••••••••••••••••••••••••••••••••••• 28 28 Scutum shiny, smooth, weakly punctate, without sculpture among punctures;
species mostly black ••••••••••.•••••••.•••••••••
. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ... Scutum dull,
granulated,
not
28.
inflatrix Benoi t
reticulate
redd ish. • • • • • • • • • • • • • • • • • • • • • • • ••
29.
rugose;
species mostly
rufonigrum n. sp.
The females of A. zair2nse Kieffer, oriphilum Kieffer, fabulosum n. abruptum n. sp. mosseli n. sp., capei n. sp., emeritum n. sp., garambanum n. sp. are unknown. sp.,
MALES
2
two
long i t ud i na I
Posterior
surface of
propodeum wi th
Posterior
surface of
propodeum wi thout
Posterior
surface of
propodeum wi th median area
long i t ud ina I
kee Is •••
2
keels ••••• 7
sh i ny,
at
I east
smooth, not rug ose •••••••••••••••••••••••••••••• . . . . 3 Posterior surface of propodeum wi th median area du I I , rugose, with numerous keels or areolae . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 cen t ra I I y
3
Head punctate, without distal
wi thout sculpture among punctures; gonoforceps inner pointed process (Fig. 252) . . . . . . . . . . . . . . ..
.................................. Head punctate, distal
inner pointed process
(Fig.
• •..•••..•••.•••••••••••••.••••• ,
4
Head
strongly
distal
30.
oriphilum Kieffer
granulated among punctures;
punctate,
not
inner pointed process
Head reticulate pointed process
reticulate (Fig.
gonoforceps with
253) . . . . . . . . . . . . . . . . . . . . . . .. 31. fabulosum n. sp. rugose;
gonoforceps with
216) •••••••••••••••••••••••. 2. agile n. sp.
rugose; gonoforceps without distal inner (Fig. 254) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ..
• •... " . • . • . . . . . • . . . • . . . . . . . . • . . . . . . . . . • . . . . . . . . . . . . • . . . . . . . . . 5 5
Posterior
surface of
propodeum with median area
sculptured by
352
Mem.
areolae much
smaller
Amer.
than
Ent.
Inst.,
those of
no.
I at era I
surface of
areolae as
6
large as
Gonoforceps
1984
areas •••••••••••••••
abruptum n.
32. Posterior
37,
propodeum with median area those of
slightly
shorter
sp.
sculptured by
areas ••••••••••••••••••.• 6
lateral
than penis
(Fig.
255) •••••••••••••
••••••••••••••••••••••••••••••••• 33. zairense Benoi t Gonoforceps much shorter than penis (Fig. 250) •••••••••••••••••
• •••••••• " 7
28. inflatrix Beno it inner pointed process (Fig.
••••••••••••••••••••••
Gonoforceps without
distal
Gonoforceps wi th distal
inner pointed process
(Figs.
259) •••••••••••••••••••••••• , •••••••••••••••• '"
8
Propodeum without posterior
a
strong
transversal
keel
227) ••• 8 258,
•.••••••••••• 17
between dorsal
and
surface ••••••••••••••••••••••••••••••••••••••••••••• 9
Propodeum wi th a strong transversal keel between dorsal and posterior surface •••••••••••••••••••••••••••••••••••••••••••• 13
9
Head granulated •••••••••••••••••••••••••••••••••••••••••••••• 10 Head punctate or without
10 Scutum dull, Scutum not only
11
near
fully
sculpture,
fully granulated,
most
granulated ••••••••••. 12
ugandanum n. sp. very
weakly granulated
and posterior margins •••••••••••••••••••••• l l
lateral
Scutum punctate,
at
not
10.
granulated ••••
without
sculpture among punctures,
not
kawandanum n.
sp.
granulated •••••••••••••••••••••••
11.
Scutum very weakly punctate, without sculpture among punctures, but very weakly granulated near lateral and posterior
1 2.
ma rg ins. • • • • • • • • • • • • • • • • • • • • • • • ••
afrum n. sp.
12 Head strongly punctate ••••••••••• 34. mosseli n. sp. Head without sculpture or very weakly punctate ••••••••••••••••• • •••••••••••••••••••••••••••••••• 13 Head Head
fully
granulated,
reticulate
rugose
35.
capei n. sp.
without
areolae or
or with
irregular keels,
irregular keels •••• 14 granulated
among areolae and keels •••••••••••••••••••••••••••••••••••••• 16 14 Scutum fully Scutum not lateral
granulated,
granulated,
dull ••••
at
most
10.
weakly
ugandanum n. sp. granulated
only near
and posterior margins •••••••••••••••••••••••••••••••• 15
15 Scutum punctate, wi thout sculpture among punctures, not granulated ••••••••••••••••••••••• 11. kawandanum n. sp. Scutum very weakly punctate, wi thout sculpture among punctures, weakly granulated near lateral and posterior margins •••••••••••
1 2. afrum n. sp. 18. gutturnium (Benoit)
• • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • •• 16 Scutum dull, strongly granulated. Scutum shiny,
without
sculpture or very
. . . • . . • . . •.•••. . . . • . . . . • . • . • . . . . . • 17 Apex of
the distal
6.
inner process of
apex
of
the gonoforceps
Apex
of
the distal
from
the apex
(Fig.
weakly granulated ••••••
kivuanum (Benoi t )
the gonoforceps nearer
36. emeritum n. of
the
258) •••••••••.•••••••••••••••••••
inner process of
the gonoforceps
sp.
the gonoforceps
(Figs.
218,
246,
farther
259) •••••••••
Anteoninae
353
••••••••.•••••••••••••••••••••••••••••••••••••••••••••••••••• 18 18 Head weakly granulated,
with numerous
•••••••••••••••••••••••••••••••••• Head granulated,
without
19 Head with TL nearly approximately
0,5
length
of
long as OPL;
notaulices
slightly
reaching
scutum . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
. . . . . . . . . . . . . . . . . . . . . . . . . . . • . . . . • 25. medleri n. Head with TL
keels ••••••.•
keels . . . . . . . . . . . . . . . . . . . 19
transversal
twice as
transversal
maritimum (Turner)
3.
longer
than OPL;
sp.
notaulices
reaching
0,3 length of scutum . . . . . . . . . . . . . . . . . . . . . . . . . . . .. • . . • . • • . • • • • . . . • • . . • • • . . . . • . . . . .. 37. garambanum n. sp.
approximately
The males of A. mingoae n. sp., brachypterum n. sp., fiorii n. sp., rufonigrum n. sp., namibiense n. sp., turneri n. sp., natalense n. sp., taylori n. sp., dayi n. sp., canabense (Benoit), anthereon Benoit, hova Benoi t, madagascolum (Benoi t), longiscapum (Ceba Ilos), compressicoxale (Benoit), townesi n. sp., arabicum n. sp., decellei n. sp., kasaicum (Benoit), hoyoi n. sp. are unknown.
1. Anteon brachypterum n. sp.
FEMALE
(Figs.
longer than
212,
tota I
213):
brachypterous;
length of scutum,
fore-wing
very reduced,
scutell um and metanotum:
sl ightly
hi nd-w ing
invisible; length 3,43 mm; black; mandibles testaceous-dark; antennae brown, wi th segments 1-3 testaceous; tegu I ae testaceous; legs testaceous, with coxae and clubs of femora partly brown; following proportions: granulated; TL = 5;
frontal
10:
6:
7:
7:
line present;
7:
=
POL
occipital carina complete;
pronotum
tubercles reaching tegulae; (5: 13), very reduced,
6: 6;
6: OL
antennal 6:
=
8;
3;
segments in
head dull,
OOL
=
5;
OPL
=
4,5;
mandibles with 4 teeth progressing
larger from anterior one to posterior; pronotal
6:
pronotum dull, scutum dull,
hairy,
granulated;
nearly 0,5 as
long as
granulated and reticulate rugose;
scutellum
shiny, small, without sculpture, slightly shorter than scutum (4:5); metanotum short, nearly 0,5 as long as scutellum (2:4); propodeum dull, granulated, with a transversal keel between dorsal and posterior surface; dorsal surface reticulate rugose; posterior surface almost smooth, with some irregular keels;
pleura transversely striate;
following proportions: 7: proximal lamellae;
prominence;
2:
2:
4:
segment 5 of front
apex with a group of nearly 4
rudimentary claw;
LOCUS TYPICUS:
tarsus with lamellae;
no segment of fore tarsi
spurs 1,1,2. MALE: unknown Ghinda
TYPICAL MATERIAL:
(Eritrea)
holotype F!
fore tarsal
segments in
10; enlarged claw with one bristle on a
in HE.
1 row of nearly 20
chela
produced
(Fig.
214)
into a hook;
without
tibial
354
Mem. Amer. En!.
Fig. 212 -
Ins!., no.
37,
1984
Female of Anteon brachypterum n. sp.
HOSTS: unknown DISTRIBUTION: Ethiopian region:
(type)
ETHIOPIA: Ghinda (Eritrea),
2. Anteon agile n. sp.
HE!
Anteoninae
Fig.
213 -
Female of
355
Anteon brachypterum n. sp. (type)
Chelogynus kivuanus Benoit 1951c (only M): 15.
FEMALE: fully winged; length 2,56-3,18 mm; black; mandibles testaceous; antennae brown, with segments 1-2 testaceous; pronotum reddish or brown; legs testaceous, sometimes with hind coxae partly brown; antennae not distally thickened; antenna I segments in following proportions: 10: 6: 9:
10:
8:
8:
8:
8:
8:
11;
head shiny,
sculpture among puncutures, without median and = 5;
OOL = 6;
OPL = 5;
posterior surface tegulae;
length of scutum;
weakly punctate,
carina complete;
pronotum shiny, (14:11);
weakly punctate, smooth,
pronotal
frons
POL = 5;
almost flat,
OL
with
tubercles reaching
without sculpture among
finely punctate,
notaulices incomplete,
without
punctate behind ocelli;
occipital
longer than scutum
scutum shiny,
punctures;
keels;
TL = 7;
pronotum smooth,
punctures; among
lateral
smooth,
more strongly
reaching
without sculpture approximately 0,65
scutellum and metanotum shiny,
smooth,
weakly
punctate, without sculpture among punctures; propodeum reticulate rugose, with a strong transversal keel between dorsal and posterior surface; posterior surface with 2 lateral
areas;
part of radial segments
longitudinal
fore-wing hyaline,
in following proportions: longer than basal
front
(Fig.
enlarged claw front 4
tarsus
lamellae;
median
vein much shorter than proximal
tarsus much tarsus
keels;
215)
9:
(Fig.
215)
proximal
with 2 rows of 28
spurs 1,1,2.
3,5:
12:
bands;
(3:6);
21;
prominence;
lamellae;
fore
distal
tarsal
segment 4 of front
(12:8);
segment 5 of
part shorter than apical
with one bristle on a
tibial
3:
part
part of segment 5
with basal
area as rugose as
without dark transversal
part
(8:13);
segment 5 of
apex with a
group of
356
Mem. Amer.
Fig.
MALE:
fully
winged;
legs testaceous, 10:
10:
length
1,68-2,68 mm;
9:
antennal 10:
10;
sculpture among punctures; 5; OL ; 3; DOL; 7; OPL ;
black;
1984
sp.
(type)
mandibles testaceous; tegu I ae testaceous;
head shiny strongly punctate,
12:
6:
without
frons without median and lateral keels; POL; 3; TL ; 5; ocellar triangle with weak keels
joining ocelli;
occipital
strong I y punctate,
notaulices incomplete,
antennae
segments in following proportions:
shi ny,
carina complete;
scutum,
scutellum and metanotum
wi thout scu I pture among punctures;
reaching approximately 0,65
propodeum reticulate rugose, dorsal
37,
sometimes with hind coxae proximally darkened;
10: 8:
smooth,
no.
somet imes wi th segment 1 testaceous;
not distally thickened; 10:
Inst.,
Chela of Anteon brachypterum n.
214 -
antennae brown,
Ent.
and posterior surface;
length of scutum;
with a strong transversal
keel
between
posterior surface with 2 longitudinal
keels;
median area nearly as rugose as lateral areas; fore-wing hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal
part
(4: 12);
gonoforceps (Fig.
membranous process and with distal 1,1,2. LOCUS TYPICUS:
216)
with a proximal
inner pointed process;
tibial
spurs
Mann's Quelle (Cameroon)
TYPICAL MATERIAL:
holotype F!
and
1 paratype M!
in BM;
4 paratypes (1
357
Anteoninae
Fig.
F,
3 MM)!
HOSTS:
in TV;
Chela of Anteon agile n.
2 paratypes (1
F,
1 M)!
sp.
(type)
in OL.
unknown
DISTRIBUTION: Cameroon), Kalonge
215 -
Ethiopian region:
BM!
(Mt.
SOUTH AFRICA:
Ruwenzori,
(Talia N River,
CAMEROON: Katberg
Albert Nat.
Tshiaberimu,
(E.
Park),
Albert Nat.
Mann's Quelle (Mt. Cape Prov.), TV!
Park),
Oll
BM!
ZAIRE:
Mbulikerere River
OL!
Magera Lake (Albert
Nat. Park), TV! Rweru (Volc. Mik<er,o), TV! Kashwa (Benoit 1951c, under name Chelogynus kivuanus: male); Rutshuru (Benoit 1951c, under name
Chelogynus kivuanus: male); Garamba Nat. Park, TV! NOTES: the supposed males of Ch. kivuanus Benoit (Benoit 1951c) have two longitudinal therefore,
keels on
actually belong
the posterior surface of the propodeum. to another species
(agile).
3. Anteon maritimum (Turner) n. comb.
Chelogynus maritimus Turner 1928: 151. Chelogynus maritimus Turner: Benoit 1951b: 162.
They,
358
Fig.
Mem.
216 -
FEMALE:
winged;
no.
37,
1984
reddish or black,
with mandibles
with segments 8-10 or
with dorsal
surface
with anterior and
posterior lobes of pronotum testaceous;
metanotum,
abdomen black;
head black,
antennae testaceous,
propectus reddish or brown,
pronotum fully
margi ns testaceous; scutellum,
Inst.,
length 2,50-3,25 mm;
(with teeth brown);
9-10 darkened; testaceous;
Ent.
Male genitalia of Anteon agile n. sp. from Mbulikerere River (Zaire): left half removed.
fully
testaceous
Amer.
mesopleura,
legs brown,
metapleura,
with fore coxae,
propodeum,
lateral
scutum, petiole and
fore tibiae and tarsi
testaceous; tegulae testaceous; antennae distally thickened; antennal segments in following proportions: 15: 5: 6: 5: 5: 5: 5: 5: 4: 5; head shiny; keel;
frons with numerous transversal two keels around orbits;
keels and a
longitudinal
vertex without sculpture,
oblique keels connecting posterior ocell i
to occipital
median
with only two
carina;
POL = 7;
OL
= 4;
OOL = 6; OPL = 5; TL = 5; pronotum shiny, hairy, smooth, without sculpture, with a raised transversal carina between anterior and posterior surface;
anterior surface nearly as
surface shorter than scutum smooth,
hairy;
notaulices distinct,
length of scutum; sculpture;
(5:15);
long as OPL
scutum shiny, incomplete,
reticulate rugose,
posterior
reaching approximately 0,3
scutellum and metanotum smooth,
propodeum dull,
(6:5);
without sculpture, shiny,
with a
without
transversal
keel
Anteoninae
Fig.
217 -
Chela of Anteon maritimum Africa)
(Turner)
359
from Mi Inerton
(South
360
Mem.
between
Ent.
dorsal and posterior surface;
longitudinal basal
Amer.
keels;
no.
37,
vein much shorter than
the pterostigma;
proximal
segment 4 of front
part;
1984
posterior surface without
fore-wing with 2 dark transversal
cells and another beneath
segment 4;
Inst.,
bands,
distal
fore tarsal
one on
the
part of radial
segment
1 as
long as
tarsus slinghtly shorter than basal
part of
segment 5; segment 5 with inner side almost straight, without a distinct apical region; basal part of segment 5 longer than apical part; segment 5 (Fig. long,
217)
with nearly 23 lamellae;
longer than segment 5,
segment 3 of front MALE:
fully
testaceous; weakly, with a
tarsus produced into a hook;
winged;
with teeth brown;
enlarged claw
with one bristle on
length 2,00-2,18 mm;
antennae brown,
legs brown,
granulated, longitudinal
with tarsi
with
black;
(Fig.
tibial
very
prominence;
spurs 1,1,2.
mandibles testaceous,
with segment 1 yellow; and fore tibiae light;
and with 2 lateral
tegulae head shiny,
numerous irregular transversal
median keel
217)
a proximal
keels;
frons
keels around orbits;
POL = 6; OL = 3; OOL = 5; OPL = 2; TL = 2; scutum, scutellum and metanotum shiny, smooth, without sculpture; notaulices distinct, incomplete,
reaching approximately 0,3 length of scutum;
a transversal
keel
reticulate rugose; rugose;
between dorsal
propodeum with
and posterior surface;
posterior surface without
longitudinal
areolae of posterior surface smaller than
dorsal keels,
surface reticulate
those of dorsal
surface;
mesopleura with posterior surface partly smooth and shiny; anterior surface dull, with numerous irregular keels; metapleura dull, with numerous irregular keels; fore-wing hyaline, with distal part of radial vein much shorter than proximal part (2:5); gonoforceps (Fig. 218)
Fig.
218 -
Male genitalia of Anteon maritimum (South Africa):
(Turner)
right half removed.
from Milnerton
Anteoninae
361
without dorsal proximal membranous process, with a dorsal distal process; gonoforceps much shorter than penis; tibial spurs 1,1,2. LOCUS TYPICUS:
Milnerton
(South Africa)
\
/
Fig. 219 -
Chela of
Anteon kasaicum (Benoit)(type)
pOinted
362
Mem.
TYPICAL MATERIAL: HOSTS:
Amer.
holotype Fl
Ent.
Inst.,
37,
no.
and 4 paratypes
1984
(2 MM,
2 FF)!
in BM.
unknown
DISTRIBUTION: AFRICA:
Ethiopian region:
Milnerton
ZAIRE:
(Cape Town),
TV!
Garamba Nat.
Ol!
BM!
Park, OLl
SOUTH
Ceres (Cape Province),
BM!
4. Anteon kasaicum (Benoit) n. comb.
Chelogynus kasaicus Benoit 1950: 222.
FEMALE: dark;
fully
length 3,75-5,62 mm;
winged;
black;
mandibles testaceous
with segments 1-4 or 1-6 testaceous;
antennae brown,
tegu I ae
testaceous; pronotum with posterior lobes and lateral margins reddish; abdomen brown-testaceous; legs testaceous, with femora and hind tibiae partly or fully brown,
with junctions,
trochanters and tarsi
light;
antennal segments in following proportions: 24: 8: 12: 8: 7: 7: 8: 7: 7: 10; head dull, granulated, with nUmerous irregular keels, frons with three strong keels, OOl = 12;
one median and
OPl = 8;
with an anterior transversal transversal
two around orbits;
Tl = 11; occipital
keels on
carina complete;
impression,
anterior surface;
granulated,
POL
= 6; Ol = 3;
pronotum dull,
with numerous
posterior surface of pronotum
shorter than scutum (13:18); pronotal tubercles reaching tegulae; scutum shiny, smooth, without sculpture or weakly alutaceous; notaulices almost invisible, reaching approximately 0,2 length of scutum; scutellum convex, without sculpture, smooth, shiny; metanotum shiny and smooth; propodeum reticulate rugose,
wi:h a strong
posterior surface;
posterior surface without
with 2 dark transversal beneath
the pterostigma;
bands, distal
transversal one on
keel
between dorsal
longitudinal
the basal
part of radial
keels;
and
fore-wing
cells and another
vein shorter than
proximal
(8: 13); pterostigma four times as long as broad; fore tarsal segments in following proportions: 7: 5: 6: 14: 39; enlarged claw (Fig. 219) with one bristle on a proximal prominence; segment 5 of front tarsus (Fig. 219) with 2 rows of nearly 34 lamellae, extending continuously to part
the apex;
apex weakly prominent;
a prominent hook; MALE:
segment 2 of front
tarsus produced
into
spurs 1,1,2.
unknown
lOCUS TYPICUS:
Kasai
TYPICAL MATERIAL: HOSTS:
tibial
(Zaire)
holotype F!
in TV.
unknown
DISTRIBUTION:
Ethiopian region:
ZAIRE:
Kasai,
TV!
Ol!
5. Anteon fiorii n. sp.
CAMEROON:
Victoria,
363
Anteoninae
FEMALE:
fully
winged;
petiole black;
length 3,00-3,62 mm;
sometimes ventral
region of thorax black;
with segments 7-10 brown; 6:
6,5:
5:
reddish-testaceous,
with
sometimes brown spots on dorsal surface of propodeum;
5:
5:
line and two
4:
5:
antennal
5:
7;
antennae reddish-testaceous,
segments
head dull,
lateral keels near orbits;
in following proportions:
reticulate rugose, POL = 8;
OL = 5;
13:
with a frontal OOL = 7;
OPL =
7; TL = 7; occipital carina complete; pronotum shiny, weakly granulated, with numerous transversal keels, hairy; pronotal tubercles reaching tegulae;
pronotum humped,
posterior surface (14:7); shiny,
smooth,
granulated;
0,3 length of scutum; humped; with a
with
anterior surface nearly twice as
pronotum shorter than scutum notaul ices incomplete,
scutellum smooth,
metanotum very short, transversal
surface without
keel
du II,
between
longitudinal
with 2 dark transversal
bands,
smooth;
dorsal
keels,
shiny,
(3:12);
reaching approximately
without sculpture,
very
propodeum reticu I ate rugose,
and posterior surface;
smooth,
long as
scutum
without
one on the basal
areolae;
posterior
fore-wing
cells and another
beneath the pterostigma; distal part of radial vein shorter than proximal part (6,5:9); fore tarsal segments in following proportions: 6: 3: 5: 9: 25;
enlarged claw
(Fig.
segment 5 of front apical
region
220)
tarsus
with one bristle on a proximal
(Fig.
distinct from
Port St.
TYPICAL MATERIAL: FF!
in OL;
HOSTS:
Johns
prominence;
inner side curvilinear,
and proximal
region;
with an
basal
part of
segment 5 of front tarsus with 2 rows
tarsus produced into a hook;
tibial
(South Africa)
holotype F!
1 paratype F!
and 2 paratypes FF!
in TW;
1 paratype F!
in BM;
2 paratypes
in AM.
unknown
DISTRIBUTION: (Pondoland), NOTES:
part;
segment 2 of front
spurs 1,1,2. MALE: unknown LOCUS TYPICUS:
with
the medial
segment 5 shorter than apical of 28 lamellae;
220)
Ethiopian region:
BM!
OL!
SOUTH AFRICA:
Magoebaskloof (Tzaneen),
this species is named in honor of the
Port St. TW!
Johns
Durban
late Mr.
(Natal),
AM!
Giorgio Fiori.
6. Atlteon kivuanum (Benoit) n. comb.
Chelogynus kivuanus Benoit 1951c (only F): 13; the male described by Benoit (1951c: 15) is not kivuanum, because the posterior surface of propodeum has 2 in females);
FEMALE:
fully
this male is agile n.
winged;
longitudinal
keels
(absent
sp.
length 2,68-3,00 mm;
black;
antennae black,
with
segments 1-5 brown or testaceous; fore legs brown or testaceous; mid and hind legs brown-black, with tarsi light and coxae partly light; antennae distally thickened;
antenna I segments in following proportions:
18:
7:
9:
364
Mem.
Fig.
220 -
Amer.
Chela of
Ent.
Inst.,
no.
37,
1984
Anteon fiorii n. sp. (type)
Anteoninae
Fig.
221
-
Chela of
365
Anteon kivuanum (Benoit) from Upemba Nat. Park (Zaire)
6:
6:
7:
7:
7:
6 (segment 10 missing
granulated and reticulate rugose;
scutum
(8:20);
incomplete, longitudinal
keels;
very reduced,
smooth,
weakly
granulated;
notaulices
visible only near anterior margin of scutum;
smooth,
keels;
head
frons with two
posterior surface of pronotum shorter than
scutum shiny,
scutellum shiny,
line present;
POL ~ 6; OL ~ 5; OOL ~ 8; OPL ~ 7; TL ~ 7; pronotum dull, granulated, punctate, with
lateral keels near orbits; occipital carina complete; strong transversal
in the only studied female);
frontal
without sculpture;
metanotum shiny,
propodeum with a strong
transversal
keel
with between
dorsal and posterior surface; dorsal surface reticulate rugose; posterior surface without longitudinal keels, reticulate rugose; fore-wing with 2 dark transversal
bands,
one beneath
the pterostigma and another on
the
basal cells; distal part of radial vein shorter than proximal part (4:6); fore tarsal segments in following proportions: 6: 3: 4: 8: 25; segment 5 of front
tarsus
(11:14);
segment 5 of front
enlarged claw
(Fig. (Fig.
tibial
spurs 1,1,2.
MALE:
fully
winged;
antenne brown,
221) 221)
with basal
part shorter than apical
tarsus curvilinear,
part
with two rows of lamellae;
with one bristle on a proximal prominence;
length 2,18-2,37 mm;
with segment 1 testaceous;
black;
mandibles testaceous;
legs testaceous,
wi th part of
hind coxae and part of hind clubs of femora brown; antennal segments in following proportions: 10: 4: 4: 4: 4: 4: 4: 4: 4: 5,5; head shiny, granulated, ~
3;
with weak areolae;
frons with a median keel,
POL
~
without
7;
OL
~
lateral
4;
OOL
keels;
~
5;
OPL
~
2,5;
scutum shiny,
TL
366
Mem.
Fig.
Amer.
Ent.
Inst.,
no.
37,
1984
222 - Male genitalia of Anteon kivuanum (Benoit) from Ceres (South Africa):
smooth,
left half removed.
without sculpture or weakly granulated;
notaulices incomplete,
reaching approximately 0,25 length of scutum; scutellum and metanotum shiny, smooth, without sculpture; propodeum with a transversal keel between dorsal
and posterior surface;
dorsal
surface reticulate rugosc;
posterior surface without longitudinal keels, fully reticulate rugose; areolae of posterior surface smaller than those of dorsal surface; forewing hyaline,
without dark transversal
much shorter than proximal
bands;
distal
part of radial
vein
(3:7); gonoforceps (Fig. 222) without
part
proximal membranous process and without distal
inner process;
tibial
spurs 1,1,2. LOCUS TYPICUS:
Rutshuru
TYPICAL MATERIAL: HOSTS: unknown DISTRIBUTION: River,
Mt.
River,
Tshiaberimu,
OL!
Ceres
ZAIRE:
(Upemba Nat.
Ruwenzori,
Albert Nat.
Albert Nat.
Rutshuru
(Cape Province),
BM!
BM! OL!
in TV. (Kivu),
TV!
TV!
Park),
Mbulikerere River
Park),
TV!
TV!
Mt.
Kakalari
OL!
Park), OL!
Bomboka (Kyandolire, Albert Nat.
(Cape Province), Town),
and 1 paratype F!
Ethiopian region:
Kiamokoto-Kiwakishi
TV!
(Zaire)
holotype F!
OL!
(Bombi (Talia N
Hoyo (Albert Nat.
Park),
TV! TW!
TW!
7. Anteon taylori n. sp.
Park),
SOUTH AFRICA:
Port St. Johns (Pondoland),
Pietermaritzburg,
River
BM!
Somerset East
Kirstenbosch
(Cape
Anteoninae
FEMALE:
fully
winged;
antennae testaceous, testaceous,
length 3 mm;
with hind coxae,
occipital
antennal
legs
segments in following proportions:
head dull,
granulated;
and with 2 lateral
carina complete;
tegulae testaceous;
hind femora and hind tibiae partly brown;
6: 6: 6: 6: 5: 5: 5: 8;
frons with a median keel
reticulate rugose;
keels around orbits;
region between occipital
with 2 oblique keels connecting the
ocelli
mandibles testaceous;
with segments 7-10 brown;
antennae distally thickened; 15: 6:
black;
367
carina and posterior
internal
side of posterior ocelli
to occipital carina; POL = 7; OL = 4; OOL = 7; OPL = 5; TL = 4; pronotum humped, dull, granulated, with numerous transversal keels; posterior surface nearly 0,5 as long as scutum anterior surface (7: 11); shiny,
smooth,
pronotal
granulated;
0,25 length of scutum;
notaulices incomplete,
scutellum shiny,
metanotum with some longitudinal a strong transversal surface without transversal
keel
smooth,
keels;
keels;
shorter than
tegulae;
scutum
reaching approximately
without sculpture;
propodeum reticulate rugose,
between dorsal
longitudinal
keels,
(8:16),
tubercles reaching
and posterior surface;
with
posterior
posterior surface with strong fore-wings with 2 dark
not reticulate rugose;
transversa I bands, one on the basa I cell s and another beneath the pterostigma; distal part of radial vein much shorter than proximal part (3:8);
fore tarsal
segments in following proportions:
segment 5 of front tarsus part
(8:12);
223)
(Fig.
enlarged claw
(Fig.
with basal
223)
5:
Kawanda
TYPICAL MATERIAL: HOSTS:
2,5:
20;
6:
apical
with one bristle on a proximal
prominence; segment 5 of front tarsus with 1 row of 11 vith a group of 8 lamellae; tibial spurs 1,1,2. MALE: unknown LOCUS TYPICUS:
3:
part shorter than lamellae;
apex
(Uganda)
holotype F!
in BM.
unknown
DISTRIBUTION: NOTES:
Ethiopian region:
UGANDA:
Kawanda,
this species is named in honor of Mr.
BM!
T .H.C. Taylor.
8. Anteon dayi n. sp.
FEMALE: fully winged; length 2,37-3,00 mm; black; mandibles testaceous; antennae brown, with segments 1-5 testaceous; legs testaceous, with hind coxae brownish; brown;
sometimes also mid and hind coxae,
antennae distally thickened;
proportions:
6:
5:
5:
4:
4:
4:
weak areolae near clypeus; occipital
carina complete;
pronotum flat, long as scutum
dull,
POL
=
granulated,
(10:14),
5: 7;
5:
7;
head dull,
line present; OL
=
femora and tibiae
segments in following
5;
OOL
granulated,
with
frons without keels;
=
5;
OPL
=
5;
TL
=
4;
with posterior surface more than 0,5 as
longer than anterior surface (10:5);
tubercles reaching tegulae; si des granu I ated;
5:
frontal
antennal
scutum shiny,
notau I ices weak,
smooth,
pronotal
without sculpture,
i nvi s ib Ie or fa int I y
v is ib Ie and
with
368
Mem.
Fig.
incomplete,
with
surface; keels;
Chela of
Ent.
Inst.,
no.
smooth and without sculpture;
a strong transversal
1984
keel
scutellum and
propodeum reticulate
between dorsal
posterior surface weakly reticulate rugose,
fore-wing
37,
Anteon taylori n. sp. (type)
almost reaching mid-length of scutum;
metanotum shiny, rugose,
223 -
Amer.
with 1 dark transversal
and posterior
without
longitudinal
band beneath the pterostigma;
distal part of radial vein much shorter than proximal part (2:6); fore tarsal segments in following proportions: 6: 2: 3: 8: 16; segment 5 of front
tarsus with basal
claw
(Fig.
224)
part shorter than apical
with one bristle on a proximal
part
(6:10);
prominence;
enlarged
segment 5 of
Anteoninae
Fig.
front tarsus 7 lamellae; MALE:
(Fig. tibial
224 -
224)
Chela of
369
Anteon dayi n. sp. (type)
with 2 rows of 26 lamellae;
apex with a group of
spurs 1,1,2.
unknown
----------
370
Mem.
LOCUS TYPICUS:
Regenstein
TYPICAL MATERIAL: HOSTS:
Amer.
En!.
Ins!.,
no.
37,
1984
(South West Africa)
holotype F!
and
1 paratype F!
in BM.
unknown
DISTRIBUTION:
Ethiopian region:
SSW Windhoek), NOTES:
BM!
SOUTH WEST AFRICA:
Okahandja,
Regenstein
(15 mls.
BM!
this species is named in honor of Mr.
Michael Day.
9. Anteon canabense (Benoit) n. comb.
Hirtanteon canabensis Benoit 1951b: 164.
FEMALE:
fully
winged;
length 2,75 mm;
reddish-testaceous,
with petiole
black; antennal segments in following proportions: 12: 4: 7: 6: 5: 6: 5: 8; antennae distally thickened; head shiny, granulated and reticulate rugose,
hairy;
frons
with a median
two keels around the orbits;
POL = 7;
4;
pronotum dull,
occipital carina complete;
transversal
keels;
granulated,
(7:14);
pronotal
incomplete,
keel
5:
and with
OPL = 4;
TL =
with numerous
longer than OPL
tubercles reaching tegulae;
reticulate rugose near lateral
notaulices distinct,
OOL = 5:
granulated,
posterior surface of pronotum
shorter than scutum dull,
longitudinal
OL = 5;
6:
(7:4) scutum
and posterior margins;
reaching mid-length of scutum;
scutellum
shiny, smooth, without scul"'cure; metanotum dull, not smooth, with numerous longitudinal keels; propodeum dull, with a transversal keel between dorsal and posterior surface; dorsal surface reticulate rugose; mesopleura and metapleura granulated and with short transversal keels; posterior surface without longitudinal keels, keels only near margins; fore-wing hyal ine, bands, (3:7)
with distal
part of radial
vein much shorter than proximal
(according to Benoit 1953b distal part
1 slightly
longer than segment 4
fully granulated, with some without dark transversal
(7:6);
longer);
fore tarsal
segment 5 of front
tarsus
part
segment (Fig.
225) with basal part shorter than apical part (7:8), with inner side curvilinear and with apical region distinct; segment 4 of front tarsus nearly as long as basal part of segment 5; segment 5 with 1 row of 23 lamellae; apex with a group of 4 lamellae; enlarged claw (Fig. 225) with one bristle on a proximal
prominence;
segment 3 produced into a hook;
tibial spurs 1,1,2. MALE: unknown LOCUS TYP ICUS:
Mundjungani
TYPICAL MATERIAL: HOSTS:
in TV.
unknown
DISTRIBUTION: TV!
(Zaire)
holotype F!
Ethiopian region:
ZAIRE:
Mundjungani
(Likimi,
Ubangi),
Anteon i nae
371
(1\ Ii \
I ,
1
Fig.
225 -
Chela of
\ '
Anteon canabense n. sp. (type)
Mem.
372
Amer.
Ent.
Inst.,
no.
37,
1984
10. Anteon ugandanum n. sp.
FEMALE:
fully
winged;
length 1,87-2,18 mm;
black;
mandibles testaceous;
antennae brown (in specimens from Uganda) or yellow (in specimens from Zaire); legs brown, with tarsi light (in specimens from Uganda) or yellow
(in specimens from Zaire);
segments in following proportions: 4,5;
head dull,
median keel, OL
=
3;
scutum
without
=
OOL
transversal
granulated; 4;
keels;
(5:9);
granulated;
lateral
OPL
=
3;
antennae distally thickened;
antennal
6:
3,5:
3,5:
3,5:
3:
2,5:
frons reticulate rugose, keels; TL
=
occipital
4;
3:
3,5:
with a
carina complete;
pronotum dull,
POL = 5;
granulated,
with some
posterior surface of pronotum more than 0,5 as
pronotal
tubercles reaching tegulae;
notaulices incomplete,
3,5:
longitudinal
scutum dull,
reaching approximately 0,25
long as
smooth, length of
scutum; scutellum and metanotum shiny, smooth, without sculpture; propodeum with a strong transversal keel between dorsal and posterior surface; dorsal surface reticulate rugose; posterior surface without longitudinal keels, with areolae smaller than those of dorsal surface; fore-wing hyaline,
without dark transversal
vein much shorter than proximal following proportions:
5:
2:
2:
3:
part sl ightly shorter than apical with one bristle on (Fig.
226)
tibial
spurs 1,1,2.
a proximal
part 9; part
Fig.
226 -
distal
fore tarsal
segment 5 of front (4:5);
prominence;
with 1 row of 10 lamellae;
bands;
(2:6);
part of radial
segments in tarsus with basal
enlarged claw
(Fig.
segment 5 of front
apex with a group of 4
Chela of Anteon ugandanum n.
sp.
(type)
226)
tarsus lamellae;
373
Anteoninae
Fig.
227 -
Male genitalia of Anteon ugandanum n. sp. from Ameib Farm (South West Africa): right half removed.
MALE:
fully
winged;
antennae fully
length
1,68-1,93 mm;
black;
mandibles testaceous;
testaceous or with segments 3-10 weakly darkened;
testaceous or with coxae,
femora and
tibiae brown;
legs
antennae not distally
thickened; antennal segments in following proportions: 6: 5: 5: 8; head dull, fully granulated; frons with a
9: 5: 5: 6: 5: 6: longitudinal median
keel, without lateral keels; occipital carina complete; POL = 6; OL = 4; OOL = 4; OPL = 2,5; TL = 2,5; scutum dull, granulated, with some weak areolae near anterior margin;
notaulices
incomplete,
reaching
nearly
mid-length of scutum; scutellum and metanotum shiny, smooth, without sculpture; propodeum reticulate rugose, with or without a strong transversal without
keel
between dorsal
longitudinal
surface;
and posterior surface;
posterior surface
keels and with areolae smaller than
fore-wing hyal ine,
without dark transversal
those of dorsal
bands;
distal
part
of radial vein much shorter than proximal part (3:7); gonoforceps (Fig. 227) without proximal membranous process and without distal inner pointed process; tibial spurs 1,1,2. LOCUS TYPICUS: Kawanda (Uganda) TYPICAL MATERIAL: holotype F! and 9 paratypes (1 para types HOSTS:
(7 MM,
in TV;
M,
9 paratypes (5 FF,
8 FF)!
4 MM)!
in BM;
9
in OL.
unknown
DISTRIBUTION: ZAIRE:
2 FF)!
Ethiopian region:
Paulis (Haut-Uele),
(Albert Nat. Albert Nat.
Park),
TV!
TV!
UGANDA:
Kakalari
River
Park),
TV!
OL!
Kiribata
Albert Nat.
Park),
OL!
TV!
Talya River
Park), BM!
TV!
OU
Kawanda
Garamba Nat.
SOUTH WEST AFRICA:
(Bombi
,
Park,
River,
(Moyenne Lume, (May
OL! TV!
BM! OL!
Mt. Mt.
BM!
Hoyo
Ruwenzori, Ruwenzori,
ya Moto River,
Ameib Farm (19 mls.
Serere, Mt.
Albert Nat.
NW Karibib),
374
Mem.
Amer.
Ent.
Inst.,
no.
11. Anteon kawandanum n.
FEMALE:
fully
antennae and thickened;
winged;
length 1,50-2,00 mm;
legs brown,
antennal
sometimes fully
37,
1984
sp.
black;
mandibles testaceous;
testaceous;
antennae distally
segments in following proportions:
6:
3,5:
2:
2:
2:
2:
2: 2: 2,5: 5; head dull, fully granulated; frons with a longitudinal median keel, without lateral keels; occipital carina complete; POL = 4; Ol = 3; OOl = 5; OPl = 3; Tl = 5; pronotum shiny, with anterior surface granulated and posterior surface smooth and without sculpture; posterior surface of pronotum shorter than scutum
(3:8);
pronotal
reaching tegulae;
finely
punctate,
scutum shiny,
sculpture among punctures; I ength of scutum; sculpture;
notaul ices incomplete,
dorsal
longitudinal
without
smooth and wi thout
keel
between
surface reticulate rugose;
keels,
tubercles
nearly reaching mid-
scutell um and metanotum sh i ny,
propodeum with a strong transversal
posterior surface; without
smooth,
dorsal
and
posterior surface
with areolae smaller than
those of dorsal
surface; fore-wing hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part (2:4,5); fore tarsal segments in following proportions: 4,5: tarsus
(Fig.
228)
enlarged claw
with basal
(Fig.
228)
1:
1: 3:
7;
segment 5 of front
part shorter than apical
with one bristle on a
part
proximal
segment 5 of front tarsus with 1 row of 10 lamellae; of 7 lamellae;
tibial
spurs 1,1,2.
Fig.
228 -
Chela of Anteon kawandanum n.
(3:4);
prominence;
apex with a group
sp.
(type)
Anteoninae
Fig.
MALE
Male genitalia of Anteon kawandanum n. (Uganda): left half removed.
229 -
(bred):
legs brown; antennal
fully
winged;
length 1,06-1,43 mm;
mandibles testaceous; smooth,
without sculpture; carina complete;
granulated;
black;
from Kawanda
antennae and
6: 4:
4:
4:
3:
4:
4:
4:
with a more or less wide frontal
frons without median and
POL = 4;
sp.
antennae not distally thickened;
segments in following proportions:
head shiny,
375
Ol = 2;
OOl = 3;
lateral
5:
6;
region
keels; occipital
OPl = 3 Tl = 4;
scutum,
scutellum and metanotum shiny, smooth, weakly punctate, without sculpture among punctures; notaulices incomplete, reaching mid-length of scutum;
propodeum reticulate rugose,
between dorsal longitudinal
and posterior surface;
keels,
fore-wing hyal ine,
with areolae smaller than those of dorsal
vein much shorter than proximal tibial
transversal
part
process,
(2:4);
bands;
distal
gonoforceps
without distal
surface;
part of radial
(Fig.
229)
inner pointed
spurs 1,1,2.
lOCUS TYPICUS:
Kawanda
(Uganda)
TYPICAL MATERIAL: holotype F! and 16 paratypes (11 4 paratypes (3 FF, 1 M)! in OL. HOSTS:
keel
posterior surface without
without dark transversal
without membranous proximal process;
with a strong
FF,
5 MM)!
in BM;
unknown
DISTRIBUTION:
Ethiopian region:
Kawanda,
Ol!
BM!
Serere,
BM!
SOMALIA:
Afgoi
(Benadir),
Ol!
UGANDA:
Ol!
12. Anteon air-um n. sp.
FEMALE:
fully
tegulae and
winged;
legs fully
length 2,00-2,68 mm; yellow-testaceous;
black;
antennae,
mandibles,
antenna I segments in following
376
Mem. Amer. Ent.
Inst.,
no. 37, 1984
Fig. 230 - Chela of Anteon afrum n. sp.
(type)
proportions: 8: 4: 2: 2: 2,5: 2: 2: 3: 2,5: 4,5; head dull, granulated, smooth; frons without median and lateral keels; POL = 4; OL = 3; OOL 4; OPL = 2,5; TL = 4; occipital carina complete; pronotum dull, granulated; pronotal tubercles reaching tegulae; posterior surface of
Anteoninae
377
pronotum
longer than anterior surface and more than 0,5 as
scutellum
(5:7);
scutum granulated,
reaching mid-length of scutum;
propodeum with a
posterior surface;
dorsal
longitudinal
transversal
keel
surface reticulate rugose;
keels,
long as
notaulices incomplete,
scutellum and metanotum shiny,
without sculpture; without
smooth;
not reticulate rugose,
between
smooth,
dorsal
granulated;
fore-wing
hyal ine, without dark transversal bands; distal part of radial vein shorter than proximal part (2:4); fore tarsal segments in following proportions: on a
rows of MALE: and
1,5:
16 lamellae;
fully
2:
3:
prominence;
winged;
legs fully
segments dull,
4:
proximal
length 1,43 mm;
line;
Tl = 4;
frons
which are very
-
7:
5:
5:
5:
keels,
carina complete;
scutum shiny,
scu I pture among punctures,
mandibles,
smooth,
POL
5:
5:
=
5;
5:
tegulae
antenna I 5:
7;
head
incomplete and Ol
=
3;
OOl
=
4,5;
without
latera I and posterior margi ns
notaulices
Male genitalia of Anteon afrum n. (Zaire):
6:
with 2
spurs 1,1,2.
antennae,
weakly punctate,
wi th areas near
weakly granulated;
230)
tibial
thickened;
with an
much
with one bristle
(Fig.
lamellae;
black;
lateral
230)
tarsus
antennae not distally
without
occipital
(Fig.
group of 3
in following proportions:
weak frontal
231
enlarged claw
segment 5 of front
apex with a
testaceous;
granulated;
OPl = 3;
Fig.
9;
and
posterior surface
incomplete,
sp.
left half removed.
reaching nearly
from Garamba Nat.
Park
378
Mem. Amer.
Ent.
Inst.,
no.
37,
1984
mid-length of scutum; scutellum and metanotum shiny, smooth, without sculpture; propodeum reticulate rugose, with a strong transversal keel between dorsal
and posterior surface;
posterior surface without
longitudinal keels, finely reticulate rugose; fore-wing hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part (1,5:5); gonoforceps (Fig. 231) without proximal membranous process and without distal inner pointed process; tibial spurs 1,1,2. LOCUS TYP I CUS: Okahandja (South West Africa) TYPICAL MATERIAL: holotype F! in BM; 1 paratype M! in TV; 1 paratype F! in OL. HOSTS:
unknown
DISTRIBUTION: Ethiopian region:
ZAIRE: Garamba Nat.
SOUTH WEST AFRICA:
BM!
Okahandja,
Park,
TV!
OL!
13. Anteon turneri n. sp.
FEMALE: fully winged; length 3,43 mm; reddish-testaceous, with petiole black, metanotum and dorsal surface of propodeum partly black; first abdominal segment partly black;
antenna I segments
in following
proportions: 10: 5: 6: 5: 5: 5: 5: 5: 4,5: 6; head dull, smooth, granulated; frons with a median longitudinal keel and with 2 lateral keels near orbits; POL = 4; OL = 5; OOL = 5; OPL = 5; TL = 5; occipital carina complete; pronotum shiny, hairy, weakly granulated; with
Fig.
232 -
Chela of Anteon turneri n.
sp.
(type)
Anteoninae
posterior surface much long as scutum shiny,
smooth,
379
longer than anterior surface,
more than 0,5 as
(10:11); pronotal tubercles reaching tegulae; scutum granulated;
notaul ices faintly
margin;
scutellum smooth,
shiny,
smooth,
without sculpture;
propodeum with a
visible near anterior
without sculpture;
metanotum shiny,
transversal
keel
between
dorsal and posterior surface; dorsal surface reticulate rugose; posterior surface without longitudinal keels, not reticulate rugose, granulated; fore-wing hyaline,
without dark
transversal
bands;
distal
part of radial
(2,5:7); fore tarsal segments in following proportions: 6: 2: 3: 7: 16; enlarged claw (Fig. 232) with one bristle on a proximal prominence; segment 5 of front tarsus (Fig. 232) with 2 rows of 30 lamellae; apex with a group of 4 lamellae; tibial spurs 1,1,2. vein much shorter than proximal
MALE:
part
unknown
LOCUS TYPICUS:
Mossel
Bay
(South Africa)
TYPICAL MATERIAL: holotype F! in BM. HOSTS: unknown DISTRIBUTION: Ethiopian region: SOUTH AFRICA: Province), NOTES:
Mossel
Bay
(Cape
BM!
this species is named in honor of Mr.
Rowland E.
14. Anteon townesi n. sp.
Fig.
233 -
Chela of Anteon townesi n.
sp.
Turner.
380
Mem. Amer.
FEMALE:
fully
proportions:
Inst.,
length 3-3,56 mm;
winged;
petiole black;
Ent.
antennae distally
no.
fully
thickened;
1984
37,
reddish-testaceous,
antennal
segments
10: 6: 8: 7: 6: 6: 6: 6: 6: 7; head shiny,
weakly reticulate rugose;
frons with an
with
in following
granulated and
incomplete median keel
and with
2 incomplete lateral keels directed towards antennal sockets; occipital carina complete; POL = 5,5; OL = 5; OOL = 6,5; OPL = 6; TL = 6; pronotum dull,
granulated,
smooth,
with a
weak
transversal
anterior
impression; posterior surface of pronotum nearly as long as scutum; pronotal tubercles reaching tegulae; pronotum flat; scutum shiny, without sculpture,
with weak
tracks of irregular keels on sides;
visible near anterior margin of scutum; smooth and without sculpture; between dorsal
little
propodeum with a strong transversal
and posterior surface;
posterior surface with small
notaul ices
scutellum and metanotum shiny,
dorsal
areolae and
irregular transversal
fore-w i ng with 2 dark transversa I bands,
keel
surface reticulate rugose;
one on
keels;
the basa I cell sand
another beneath the pterostigma; distal part of radial vein shorter than proximal part (4:9) fore tarsal segments in following proportions: 8: 2:
3,5: 10: 20; enlarged claw (Fig. 233) with one bristle on a proximal prominence; segment 5 of front tarsus (Fig. 233) with 2 rows of 40 lamellae; apex with a group of 4 lamellae tibial spurs 1,1,2. MALE:
unknown
LOCUS TYPICUS:
Royal
TYPICAL MATERIAL: HOSTS:
National
Park
in TW;
(South Africa)
1 paratype F!
in 0'-.
unknown
DISTRIBUTION: TW!
Natal
holotype F!
Ethiopian region:
Ken ton-on-Sea,
NOTES:
this species
SOUTH AFRICA:
Royal
Natal
Nat.
Park,
OL! is named
in
honor of Mr.
Henry K.
Townes.
15. Anteon decellei n. sp.
FEMALE:
fully
winged;
length 2,62 mm;
reddish-testaceous,
except for
metanotum and part of propodeum brown; antennae brown, with segments 1-2 testaceous; antennae distally thickened; antennal segments in following proportions:
9:
5:
5:
4:
4:
4:
4:
4
(last 2 segments missing
in
the honly known specimen); head dull, granulated, reticulate rugose only near clypeus; frons with a longitudinal median keel and with 2 lateral keels around orbits;
occipital
4;
pronotum shiny,
OPL = 5;
TL = 5;
some transversal
tubercles rc,6ching
metanotum shiny,
POL = 6;
OL = 4;
weakly granulated,
posterior surface very
OOL = with
long,
(12:10) and longer than anterior surface (12:3);
notaulices incomplete, with a
smooth,
keels on anterior surface;
longer than scutum pronotal
carina complete;
tegulae;
scutum shiny,
smooth,
reaching mid-length of scutum;
smooth,
strong transversal
posterior surface without
without sculpture; keel
between
longitudinal
propodeum reticulate rugose,
dorsal
keels,
granulated;
scutellum and
and posterior surface;
weakly
reticulate rugose;
381
Anteoninae
Fig.
234 -
Chela of Anteon decellei n.
fore-wing with 2 dark transversal another beneath the pterostigma; than proximal
part
(3,5:7);
bands, distal
fore tarsal
one on
sp.
the basal
part of radial segments
(type)
cells and
vein much shorter
in following proportions:
6: 2: 2,5: 8: 18; segment 5 front tarsus with basal part shorter than apical part (8: 10); enlarged claw (Fig. 234) with one bristle on a proximal
prominence;
29 lamellae; MALE:
tarsus
(Fig. tibial
234)
with 2 rows of
spurs 1,1,2.
unknown
LOCUS TYPICUS:
Garamba National
TYPICAL MATERIAL: HOSTS:
segment 5 of front
apex with a group of 3 lamellae;
holotype F!
Park
(Zaire)
in TV.
unknown
DISTRIBUTION: Ethiopian region: ZAIRE: Garamba Nat. Park, NOTES: this species is named in honor of Mr. Jean Decelle.
16. Anteon anthereon Benoit
TV!
382
Mem. Amer.
Anteon anthereon
FEMALE: black;
fully
winged;
antennal
Ent.
Inst.,
Benoit 1951b:
no.
37,
1984
160.
length 2,68 mm;
fully
testaceous,
segments in following proportions:
10:
with petiole
5:
5:
5:
5:
5:
5:
5: 5: 7; antennae distally thickened; head shiny, hairy, fully weakly reticulate rugose; POL = 5; OL = 4; OOl = 4; OPL = 4; TL = 2; frons with a weak longitudinal median keel, without lateral keels; pronotum shi ny, weak I y rugose, with posterior surface very short, much shorter than scutum
(5:15);
weakly punctate;
pronotal
length of scutum; dorsal
keel
between dorsal
surface strongly reticulate rugose; granulated,
incomplete,
scutellum and metanotum shiny,
propodeum with a transversal dull,
tubercles reaching tegulae;
notaulices weakly visible,
scutum shiny, reaching mid-
without sculpture; and posterior surface;
mesopleura and metapleura
weakly reticulate rugose;
posterior surface of
propodeum without longitudinal keels, granulated; fore-wing with a faint preapical dark transversal band; distal part of radial vein much shorter
Fig.
235 -
Chela of
Anteon anthereon
Benoit
(type)
Anteoninae
than proximal segment 4
part
(9:2,5);
(3: 10); fore tarsal segment 5 of front
almost rectilinear and with 4 basal
part of segment 5;
segment 1 much tarsus
lamellae;
(Fig.
longer than
235)
with
inner side
segment 4 nearly 0,5 as
enlarged claw
long as
with 1 bristle on a proximal
segment 4 of front tarsus produced into a
prominence; 1,1,2. MALE:
383
hook;
tibial
spurs
unknown
LOCUS TYPICUS: Port St. Johns (South Africa) TYPICAL MATERIAL: holotype F! in BM. HOSTS:
unknown
DISTRIBUTION: (Pondoland),
Ethiopian region:
SOUTH AFRICA:
Port St.
Johns
BM!
17. Anteon namibiense n. sp.
FEMALE:
fully
winged;
length 2-2,12 mm;
testaceous;
an tennae brown,
testaceous;
mid and hind
proportions: granulated; lateral
11:
4:
3:
frons with a
legs brown; 3,5: 3,5:
antennal
3,5:
mandibles and
1 tes taceous;
3,5:
fore
granu I a ted,
longitudinal median keel
POL; 8;
OL ;
4;
OOL ;
5;
4,5;
head dull,
and with
OPL ;
with posterior surface very short,
pronotal
Fig.
tubercles reaching tegulae;
236 -
tegulae
legs
segments in following
3,5:
keels near orbits directed towards antenna I sockets;
carina complete; (2:15);
4:
black;
with segmen t
3;
TL ;
tracks of 2 occipital 2;
pronotum
much shorter than scutum
scutum dull,
Chela of Anteon namibiense n.
sp.
granulated;
(type)
384
Mem.
notaul ices very short,
Amer.
Ent.
keel
37,
1984
without sculpture;
and posterior surface;
longitudinal
keels, bands,
granulated, one on
distal
sl ightly rugose;
the basal
part of radial
propodeum with
dorsal
posterior surface not reticulate rugose,
transversal pterostigma;
shiny,
between dorsal
reticulate rugose;
no.
faintly visible near anterior margin of scutum;
scutellum and metanotum smooth, a transversal
I nst.,
surface
without
fore-wing
with 2 dark
cells and another beneath
the
vein much shorter than proximal
part
(2:6); fore tarsal segments in following proportions: 6,5: 1,5: 1: 1,5: 5,5; enlarged claw (Fig. 236) with one bristle on a proximal prominence; segment 5 of front tibial
tarsus
(Fig.
236)
with 4
lamellae and some bristles;
spurs 1,1,2.
MALE:
unknown
LOCUS TYPICUS:
Aus
(South West Africa)
TYPICAL MATERIAL:
holotype F!
and 1 paratype F!
in BM;
1 para type F!
in OL. HOSTS: unknown DISTRIBUTION: Ethiopian region: AFRICA:
Glen
(0.
F. S.),
SOUTH WEST AFRICA: Aus,
BM!
SOUTH
BM!
18. Anteon gutturnium (Benoi t) n. comb.
Xenanteon gutturnium Benoit 1951b: 162. Xenanteon gutturnium Benoit ssp. rutshuricum Benoit 1951c: 13 (n. syn.); locus typicus Rutshuru (Zaire).
FEMALE: yellow
fully
(with
winged;
occasionally 5:
5:
6:
granulated, keel,
head black,
6:
light;
6:
6:
9;
tegulae testaceous;
antenna I segments in following proportions: antennae distally thickened;
fully reticulate rugose;
without
with mandibles
antennae brown with segment 1 testaceous;
with segments 1-3 or 1-5 testaceous;
brown with tarsi 5:
length 2,37-3,12 mm;
teeth brown);
lateral keels;
frons with a
POL ~ 8;
OL ~ 5;
head hairy,
longitudinal
OOL ~ 7;
legs 17: 4:
shiny,
median
OPL ~ 5;
TL ~ 4;
pronotum rugose, granu I a ted, wi th posterior surface very short, much shorter than scutum (4: 14); pronotal tubercles reaching tegulae; scutum shiny,
granulated,
scutellum shiny,
with weak
irregular keels;
weakly punctate,
smooth;
propodeum wi th a transversa I keel
notaulices invisible;
metanotum strongly rugose;
between dorsal
dorsal
surface reticulate rugose;
keels,
granulated and with numerous transversal
metapleura granulated, dark bands,
and posterior surface;
posterior surface without
with numerous transversal
keels; keels;
one beneath the pterostigma and another on
longitudinal
mesopleura and fore-wing with 2 the basal
cells
(only the band beneath the pterostigma in specimens from Uganda); distal part of radial vein much shorter than proximal part (2:8); fore tarsal segment 1 much longer than segment 4
(8:3);
segment 4 of front
tarsus
Anteoninae
Fig.
237 -
Chela of Anteon gutturnium Africa)
(Benoit)
385
from Mossel
Bay
(South
386
Mem.
produced into a lamellae;
hook;
Amer.
Inst.,
segment 5 of front
(Fig.
tibial
spurs 1,1,2.
MALE
(not reared):
237) fully
winged;
with
testaceous;
tegulae testaceous;
hind femora,
teeth brown;
tarsus
legs yellow, head dull,
frons with a short
keels around orbits;
scutum dull,
granulated;
POL
(Fig.
237)
black;
with hind coxae,
longitudinal
mandibles clubs of
granulated and weakly median keel,
incomplete,
without
dorsal keels,
3·,
reaching
scutellum and metanotum shiny,
propodeum wi th a transversal
and posterior surface;
prominence;
wi th segments 1-2
longitudinal
notaulices distinct,
wi thout scu I pture;
with 10
part of segment 5;
= 6; OL = 3; OOL = 5; OPL = 4; TL
approximately 0,3 length of scutum;
surface without
1984
length 2,18-2,50 mm;
antennae brown,
hind tibiae brown;
reticulate rugose;
dorsal
37,
with one bristle on a proximal
testaceous,
smooth,
no.
segment 4 nearly 0,5 as long as basal
enlarged claw
lateral
En!.
keel
between
surface reticulate rugose; reticulate rugose,
posterior
with areolae smaller
than those of dorsal surface; mesopleura and metapleura granulated, with irregular keels and areolae; fore-wing hyal ine, with distal part of radial vein much shorter than proximal part; gonoforceps (Fig. 238) with dorsal proximal
membranous process,
gonoforceps nearly as LOCUS TYPICUS:
Mossel
TYPICAL MATERIAL:
Fig.
238 -
without distal
long as penis; Bay
tibial
inner pointed process; spurs 1,1,2.
(South Africa)
holotype F!,
allotype M!
and 1 paratype F!
Male genitalia of Anteon gutturnium (South Africa):
(Benoit)
left half removed.
of X.
from Katberg
387
Anteoninae
gutturnium in BM; 1 paratype F! of X. gutturnium in TV; holotype F! of X. gutturnium rutshuricum in TV. HOSTS: unknown DISTRIBUTION: Ethiopian region:
UGANDA:
BM!
Beach,
Bushenyi,
BM!
KENIA:
Diani
Kawanda, BM!
ZAIRE:
Ol!
BM!
Kampala,
Rutshuru
(Kivu),
TV!
Ikonongi Cave (Mt. Hoyo, Albert Nat. Park), Ol! TV! SOUTH AFRICA: Grahamstown, TW! Hluhluwe Game Res., Ol! TW! Kenton-on-Sea, TW! Ol! Mossel
Bay
Katberg
(Cape Province),
(E Cape Province),
BM!
TV!
Port St.
Johns
(Pondoland),
BM!
BM!
19. Anteon arabicum n. sp.
FEMALE:
fully
winged;
length 2,43 mm;
black;
mandibles testaceous;
antennae black, with segment 1 brown; legs black, with tarsi brown; antennae distally thickened; antennal segments in following proportions: 10: 4: 3,5: 4: 3: 4: 4: 4: 4: 5; head dull, granulated, with weak irregular short keels; frons with a longitudinal median keel and with 2 lateral
incomplete keels near orbits directed towards antenna I sockets;
occipital
carina complete;
Fig.
239 -
POL = 7;
Chela of
Ol = 4;
OOl = 4;
OPl = 3;
Anteon arabicum n. sp. (type)
Tl = 3,5;
388
Mem.
pronotum dull,
Amer.
granulated,
Ent.
longitudinal
37,
1984
keels,
shorter than anterior surface;
surface also shorter than scutum scutum shiny,
no.
with numerous transversal
posterior surface very short, tegulae;
Inst.,
(5:14);
pronotal
very weakly alutaceous,
weak keels or furrows;
notaulices
with posterior
tubercles reaching
with numerous invisible;
scutellum and
metanotum shiny, smooth, without sculpture; propodeum reticulate rugose, with a strong transversal keel between dorsal and posterior surface; posterior surface without dark transversal proximal 1,5:
part
1,5:
apical
bands;
(2:8);
1,5: 7;
and some bristles; MALE:
fore tarsal
tarsus with basal
(Fig.
239)
segment 5 of front
tibial
without
vein much shorter than
segments in following proportions:
enlarged claw
prominence;
fore-wing hyaline,
part of radial
segment 5 of front
part (5:2);
proximal
longitudinal keels; distal
6:
part longer than
with one bristle on a
tarsus
(Fig.
239)
with 4
lamellae
spurs 1,1,2.
unknown
LOCUS TYPICUS: Jebel Jihaf (Southern Yemen) TYPICAL MATERIAL: holotype F! in BM. HOSTS: unknown DISTRIBUTION: Ethiopian Jihaf (Aden),
Palaearctic region:
SOUTHERN YEMEN:
Jebel
BM!
20. Anteon natalense n.
sp.
FEMALE: fully winged; length 2,56-2,81 mm; black; mandibles and legs testaceous; antennae brown, with segment 1 testaceous; antenna I segments in following proportions:
13:
4,5:
4:
reticulate rugose;
occipital
OPL = 4;
pronotum rugose,
short,
TL = 3;
shorter than scutum
scutum shiny, longitudinal smooth,
(5:16);
reticulate rugose,
keels;
4:
shiny,
5:
pronotal
5:
5:
5:
POL = 8;
7;
head shiny,
OL = 5;
OOL = 5;
with posterior surface very tubercles reaching
tegulae;
with or without numerous strong
notaulices invisible;
without sculpture;
4:
carina complete;
scutum and metanotum shiny,
propodeum with a strong transversal
keel
between dorsal and posterior surface; posterior surface without longitudinal keels and with areolae smaller than those of dorsal
surface;
fore-wing hyal ine, without dark transversal bands or spots; distal of radial vein much shorter than proximal part (3:9); fore tarsal segments in following proportions: 7: tarsus with basal tarsus
less than 0,5 as long as basal
enlarged claw
(Fig.
segment 5 of front tibial MALE:
1,5: 2:
2,5: 8;
part much longer than apical 240)
tarsus
part;
segment 5 of front segment 4 of front
part of segment 5
with one bristle on a proximal (Fig.
240)
with 8
part
(2,5:7); prominence;
lamellae and some bristles;
spurs 1,1,2. unknown
LOCUS TYPICUS: Van Reenen (South Africa) TYPICAL MATERIAL: holotype F! in BM; 1 paratype F!
in TV;
1 paratype
Anteoninae
Fig. 240 -
Chela of
389
Anteon natalense n. sp. (type)
F! in OL. HOSTS: unknown DISTRIBUTION: Ethiopian region: ZAIRE: Garamba Nat. SOUTH AFRICA: Van Reenen (Drakensberg, Natal), BM!
Park,
OL!
TV!
390
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
21. 11nteon hoyoi n. sp.
FEMALE:
fully
winged;
antennae brown,
length 2,50 mm;
black;
mandibles testaceous;
with segments 1-2 testaceous;
legs fully
with hind coxae and hind femora partly brown; thichened;
antenna I segments
testaceous or
antennae distally
in following proportions:
12:
5:
7:
5:
5:
6'
6: 6: 6: 8; head shiny, reticulate rugose, 'Nith a longitudinal median keel and with 2 lateral keels around orbits; occipital carina complete; POL
~
5;
rugose,
Ol
~
3;
OOl
~
5;
~
OPl
4;
posterior surface shorter than scutum tegulae;
Tl
with posterior surface nearly as scutum shiny,
punctures;
smooth,
notaulices faintly
transversal
keel
2;
pronotum dull,
reticulate
long as anterior surface
(7:15);
pronotal
weakly punctate,
tubercles reaching
without sculpture among
visible near anterior margin of scutum;
scutell um and metanotum sh i ny, sculpture among punctures;
~
smooth,
weak I y
punctate,
propodeum reticulate rugose,
between dorsal
and posterior surface;
without with a strong
posterior surface
without longitudinal keels; fore-wing hyal ine, without dark transversal bands; distal part of radial vein much shorter than proximal part (2:9); fore tarsal
segments
Fig.
241
in following proportions:
-
Chela of
10:
1,5:
2:
3:
Anteon hoyoi n. sp. (type)
8;
segment 5
Anteoninae
of front
tarsus with basal
enlarged claw
(Fig.
segment 5 of front lamellae; MALE:
tibial
241)
tarsus
391
part much longer than apical with one bristle on a proximal (Fig.
241)
part
(7:1);
prominence;
with some bristles and 4 apical
spurs 1,1,2.
unknown
LOCUS TYPICUS: ,
(1< i vu),
1 paratype F!
ZAIRE:
Kabambi
Cave
in OL. (Mt.
Hoyo),
TV!
OL!
22. Anteon mingoae n. sp.
FE\\ALE:
fully
winged;
length 1,87 mm;
black;
mandibles testaceous;
antennae testaceous, with segments 6-10 brown; legs testaceous, with hind coxae partly brown; antennae distally thickened; antennal segments in following proportions: reticulate rugose;
Fig.
10: 4:
3:
3:
3:
4:
4:
frons with a median keel
242 -
3,5:
4:
5;
head shiny,
and 2 lateral
Chela of Anteon mingoae n.
sp.
keels near
(type)
392
Mem.
Amer.
Ent.
Inst.,
orbits directed towards antenna I sockets; 4;
OL = 3;
OOL = 6;
OPL = 3;
TL = 3;
no.
37,
occipital
smooth and without sculpture;
shorter than scutum
(3: 12); pronotal
weakly granulated;
shiny,
smooth,
keel
POL
with posterior
posterior surface
tubercles reaching tegulae;
notaulices invisible;
without sculpture;
strong transversal
carina complete;
pronotum rugose,
surface almost fully shiny,
1984
scutum
scutellum and metanotum
propodeum reticulate rugose,
between dorsal posterior surface;
with a
posterior surface
without longitudinal keels; fore-wing hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part (1 :7); fore tarsal
segments in following proportions:
segment 5 of front (4,5:1,5);
enlarged claw
prominence;
(Fig.
242)
segment 5 of front
some bristles; MALE:
tarsus with basal
tibial
6:
part much
1,5:
1,5:
1,5: 6;
longer than apical
part
with one bristle on a proximal
tarsus
(Fig.
242)
with 6 lamellae and
spurs 1,1,2.
unknown
LOCUS TYPICUS: Ankaratra (Madagascar) TYPICAL MATERIAL: holotype F! in MD. HOSTS: unknown DISTRIBUTION: NOTES:
Ethiopian region:
this species is named
MADAGASCAR:
Ankaratra,
in honor of Mrs.
MD!
Elvira Mingo Perez.
23. Anteon longiscapum (Ceba Ilos) n. sp.
Xenanteon longiscapus Ceballos 1936: 63. Xenanteon longiscapus Ceballos: Benoit 1953b: 417.
FEMALE:
fully
teeth brown; testaceous;
winged;
length 3,75 mm;
antennae testaceous,
legs brown,
proportions:
17: 6:
7:
with tarsi 5:
5:
6:
6:
black;
mandibles testaceous,
with segments 9-10 brown; light; 6:
6:
antennal 11;
with
tegulae
segments in following
antennae distally thickened;
head shiny,
granulated and reticulate rugose;
median keel
and with 2 lateral
frons with a
keels around orbits;
longitudinal
POL = 6;
OL = 5;
OOL = 4; OPL = 6; TL = 4; pronotum shiny, smooth, weakly granulated, without sculpture near posterior margin; posterior surface very long, more than 0,5 as long as scutum tegulae;
scutum shiny,
smooth,
scutellum and metanotum shiny, a transversal
keel
reticulate rugose; granulated, smooth,
smooth,
between dorsal
pronotal
tubercles reaching notaulices invisible;
without sculpture;
and posterior surface;
propodeum with
dorsal
surface
posterior surface with weak track of longitudinal
with numerous weak
without sculpture;
fore-wing with an apical radial
(9:16);
weakly granulated;
transversal
keels;
metapleura with numerous wide dark transversal
vein much shorter than proximal
longer than segment 4 (7:4);
part
irregular keels;
band;
(3:10);
keels,
mesopleura shiny, distal
part of
fore tarsal
segment
segment 4 nearly 0,5 as long as basal
part
Anteoninae
Fig.
of segment 5; (Fi g.
243)
a proximal
243 -
Chela of
I ameli ae;
prominence;
MALE: unknown lOCUS TYPICUS:
tibial
Fanovana
TYPICAL MATERIAL: HOSTS: unknown DISTRIBUTION:
Anteon longiscapum (Ceballos)(type)
segment 4 produced into a
with 22
393
hook;
en I arged cl aw
segment 5 of front
(F i g;
243)
tarsus
wi th one brist Ie on
spurs 1,1,2.
(Madagascar)
holotype F!
in P.
Ethiopian region:
MADAGASCAR:
Fanovana,
P!
24. Anteon madagascolum (Benoit) n. comb.
Xenanteon madagascola Benoit 1953b: 418.
FEMALE:
fully
winged;
wi th teeth brown; brown;
tegulae testaceous;
brown with tarsi 4:
4:
length 2,50-3,00 mm;
4:
4:
5:
5:
light; 5:
7;
long,
segments in following proportions:
antennae distally
thickened;
Tl = 4;
pronotum shiny,
granulated; reaching
notaul ices invisible;
Ol = 4;
scutellum shiny,
occasionally visible:
5:
keel;
OOl = 5;
posterior surface very
(10:13);
approximately 0,3 length of scutum;
longitudinal
POL = 7;
granulated;
more than 0,5 as long as scutum
13:
head shiny,
frons with a median
keels around orbits present or absent;
OPl = 6;
mandibles testaceous,
legs testaceous with hind coxae brown or
antennal
granulated and reticulate rugose; lateral
black;
antennae brown wi th segments 1-6 testaceous or fu II y
smooth,
in this case
scutellum and metanotum
394
shiny,
Mem.
smooth,
Amer.
weakly granulated;
between dorsal
Inst.,
numerous irregular keels;
longitudinal
37,
with
irregular keels;
bands;
distal
part of radial
1984
keel
dorsal
surface reticulate rugose;
keels,
granulated and with
mesopleura granulated;
rugose,
segment
no.
propodeum with a transversal
and posterior surface;
posterior surface without
fore tarsal
Ent.
fore-wing hyaline,
metapleura strongly without dark transversal
vein much shorter than proximal
1 longer than segment 4
(8:3,5);
part
(3:9);
segment 4 of front
tarsus produced into a hook, nearly 0,5 as long as basal part of segment 5; segment 5 (Fig. 244) with 10 lamellae; enlarged claw (Fig. 244) with one bristle on a proximal MALE: unknown LOCUS TYPICUS:
Bekily
TYPICAL MATERIAL:
prominence;
244 -
spurs 1,1,2.
(Madagascar)
holotype F!
HOSTS: unknown DISTRIBUTION: Ethiopian region:
Fig.
tibial
Chela of
in P;
2 paratypes FF in TV.
MADAGASCAR:
Bekily,
Anteon madagascolum
P!
TV!
Ivondro,
(Benoit)(type)
395
Anteoninae
OL!
Ranomafana
(Benoit 1953b);
DIEGO SUAREZ
(Benoit 1953b).
25. Anteon medleri n. sp.
FEMALE: fully winged; length 2,37 mm; black; mandibles testaceous; antennae brown, with segments 1-2 testaceousj tegulae testaceous; legs brown, with tibiae and tarsi testaceous; antennae distally thickened; antennal segments in following proportions: 9: 5: 6: 5: 5: 5: 5: 5: 5: head dull, a
granulated,
longitudinal
complete;
median keel,
POL = 5;
granulated,
with some weak areolae near clypeus;
except for a
without sculpture;
without
Ol = 4; OOl = 5;
lateral
long,
occipital carina
Tl = 6;
narrow posterior region
numerous transversal
surface of pronotum
keels;
OPl = 6;
7;
frons with
which
pronotum dull, is smooth and
keels on anterior half;
more than 0,5 as long as scutum;
posterior
pronota I
tubercles reaching tegulae; scutum dull, granulated; notaul ices invisible; scutellum and metanotum shiny, smooth, without sculpture; propodeum reticulate rugose,
with a strong
posterior surface;
posterior surface without
Fig.
245 -
Chela of
transversal
keel
between dorsal
longitudinal
keels;
Anteon medleri n. sp. (type)
and
fore-wing
396
Mem.
Amer.
Ent.
Ins!.,
no.
37,
1984
hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part (2:5); fore tarsal segments in following proportions: basal
6:
2:
2:
2,5:
8;
segment 5 of front
part longer than apical
one bristle on a row of 8
proximal
lamellae;
part (5:3);
prominence;
apex with a
tarsus
(Fig.
enlarged claw
segment 5 of front
group of 6 lamellae;
245)
(Fig.
with
245)
with
tarsus with
tibial
1
spurs 1,1,2.
MALE: fully winged; length 1,37 mm; black; mandibles and antennae testaceous; legs testaceous, with hind coxae, hind femora and distal region of hind tibiae brownish;
antennae not distally thickened;
segments in following proportions: 4,5:
6;
occipital
head dull,
granulated;
carina complete;
scutum shiny, punctures;
smooth,
246 -
4,5: 4,5:
4,5:
4,5:
4,5:
frons without median and
POL = 4;
OL = 2,5;
weakly punctate,
notaulices incomplete,
scutellum and metanotum shiny,
Fig.
6:
OOL = 4;
antennal
4,5:
lateral
4,5:
keels;
OPL = 2;
TL = 4;
without sculpture among
reaching nearly mid-length of scutum; smooth,
without sculpture;
Male genitalia of Anteon medleri n. left half removed.
sp.
from
propodeum
Ile-ife
(Nigeria):
397
Anteoninae
reticulate rugose,
with a strong median transversal
and posterior surface; wing hyaline,
posterior surface without
without dark transversal
much shorter than proximal proximal
part
membranous process,
(2:4);
bands;
distal
gonoforceps
with distal
NOTES:
th i s species
is named
NIGERIA:
between dorsal keels;
fore-
part of radial
(Fig.
inner process;
1,1,2. LOCUS TYPICUS: Ile-ife (Nigeria) TYPICAL MATERIAL: holotype F! and 1 paratype M! HOSTS: unknown DISTRIBUTION: Ethiopian region:
keel
longitudinal
vein
246)
without
tibial
spurs
in BM.
Ile-ife (W State),
in honor of Mr. J. T.
BM!
Med I er
26. Anteon hova Benoit
Anteon hova Benoi t 1953b: 415.
FEMALE: fully winged; length 3 mm; black; mandibles yellow; antennae yellow; legs yellow; tegulae testaceous; head missing in the only known specimen;
pronotum shiny,
anterior surface; distinct,
incomplete,
metanotum shiny, keel
without sculpture,
scutum shiny,
with
transversal
without sculpture;
reaching mid-length of scutum;
smooth,
between dorsal
smooth,
without sculpture;
and posterior surface;
keels on
notaulices
scutellum and
prnpodeum with a transversal dorsal surface reticulate
rugose; posterior surface with 2 long i tud i na I keel s shorter than scutum (6:12); r·ledian area mostly smooth, with keels only around margins;
Fig.
247 -
Chela of Anteon hova Benoit
(type)
398
Mem. Amer.
lateral
areas reticulate rugose;
posterior surface smooth, hyaline, (2:7); front
with distal
fore tarsal
segment 5;
no.
37,
1984
mesopleura with anterior surface rugose;
without sculpture;
metapleura rugose;
into a hook,
segment 5 (Fig.
247)
one bristle on a proximal
fore-wing
vein much shorter than proximal part
segment 1 longer than segment 4
with numerous bristles;
MALE:
Inst.,
part of radial
tarsus produced
part,
Ent.
nearl yO,S as
without
segment 4 of
lamellae on proximal
apex with 4
prominence;
(7:2);
long as bas a I part of and medial
lamellae; enlarged claw with
tibial
spurs 1,1,2.
unknown
LOCUS TYPICUS:
Bekily
TYPICAL MATERIAL: HOSTS:
(Madagascar)
holotype F!
in Pi
1 para type F
in TV.
unknown
DISTRIBUTION:
Ethiopian region:
MADAGASCAR:
Bekily,
P!
TV!
27. Anteon compressicoxale (Benoi t) n. comb.
Xenanteon compressicoxale Benoit 1953b: 416.
FEMALE:
fully
teeth brown; testaceous;
winged;
legs testaceous,
partly brown; 4:
4:
4:
length 2,68 mm;
antennae brown,
4:
antennal
7;
black;
with
tegulae
with hind coxae and hind clubs of femora
segments in following proportions:
antennae distally thickened;
numerous transversal
mandibles testaceous,
with segments 1-4 testaceous;
irregular keels;
POL
head dull,
=
=
4; Ol
5;
11: 4:
5:
granulated, OOl
=
4;
4:
4:
with
OPl
=
5; Tl = 2; pronotum damaged in holotype, with transversal keels, except for the area near posterior margin which is smooth; scutum shiny, smooth,
weakly punctate;
scutum;
scutellum and metanotum shiny,
notaul ices reaching approximately 0,25
propodeum with a transversal
Fig.
248 -
keel
smooth,
length of
without sculpture;
between dorsal
and posterior surface;
Chela of Anteon compressicoxale
(Benoit)(type)
Anteoninae
399
dorsal
surface reticulate rugose;
posterior surface with 2 longitudinal
keels;
median
lateral
area
metapleura dull,
as rugose as
rugose,
without dark transversal shorter than proximal segment 4 basal
(9:2);
areas;
with transversal bands,
part
with distal
(2:7);
into a
segment 5
(Fig.
and
fore-wing hyal ine,
part of radial
fore tarsal
segment 4 produced
part of segment 5;
mesopleura
keels;
segment
hook,
248)
vein
much
1 longer than
nearly 0,5 as
without
long as
lamellae on median
and proximal parts, with bristles; apex with 6 lamellae; enlarged claw (Fig. 248) with one bristle on a proximal prominence; tibial spurs 1,1,2. MALE:
unknown
LOCUS TYP I CUS:
Bek i I y
TYPICAL MATERIAL: HOSTS:
(Madagascar)
holotype F!
in P.
unknown
DISTRIBUTION:
Ethiopian region:
MADAGASCAR:
Bekily,
P!
28. Anteon inflatrix Benoi t
Anteon inflatrix Benoit 1951b: 161.
FEMALE:
fully
winged;
length 3,00-3,12 mm;
black;
mandibles yellow,
with teeth brown; antennae fully testaceous; occasionally with segments 6-10 fully brown or with dorsal surface brown; anterior margin of pronotum reddish-dark; occasionally
tegulae testaceous;
legs fully
with mid clubs of femora partly brown;
testaceous, hind coxae,
clubs of
hind femora and hind tibiae brown;
antenna I segments in following
proportions:
5:
head dull, median 5;
9:
5:
5:
4:
granulated,
longitudinal
OPL = 5;
4:
keel,
TL = 4;
5:
5:
without
posterior surface nearl y
(5:12);
scutum;
shiny,
6;
lateral
pronotum rugose,
smooth;
antennae distally thickened;
smooth,
keels;
POL = 7;
frons
OL = 4;
with a OOL =
wi th posterior surface sh i ny and
as long as OPL and shorter than scutum without sculpture,
notaul ices invisible occasionally length of scutum;
5:
weakly or strongly reticulate rugose;
incomplete,
weakly punctate;
reaching approximately 0,25
scutellum and metanotum shiny,
smooth,
without
sculpture; propodeum with a transversal keel between dorsal and posterior surface; dorsal surface reticulate rugose; posterior surface with 2 longitudinal
keels;
median area as rugose as lateral
and metapleura rugose; band beneath than proximal (7:2,5);
on medial MALE
the pterostigma; part (3:7);
distal part of radial
and proximal parts;
with one bristle on a (not reared):
fully
segment
tarsus produced
part of segment 5;
mesopleura
segment 5
vein much shorter
1 longer than segment 4
into a hook, (Fig.
249)
apex with 8 lamellae;
proximal winged;
areas;
with a more or less dark transversal
fore tarsal
segment 4 of front
long as basal 249)
fore-wing
prominence;
tibial
length 2,37 mm;
nearly 0,5 as
without
lamellae
enlarged claw spurs 1,1,2.
black;
mandibles
(Fig.
400
Mem. Amer. Ent.
I nst., no. 37,
1984
Fig. 249 - Chela of Anteon inflatrix Benoit from Port St. Johns (South Africa)
Anteoninae
testaceous;
antennae brown,
junctions and proportions:
tarsi
10: 6:
with a median 4; OOl = 6;
401
with segment 1 testaceous;
legs brown,
testaceous;
antennal
segments in following
7:
6:
7;
6:
6:
longitudinal
OPl = 5;
6: keel
Tl = 3;
6:
6:
head shiny,
and without
occipital
lateral
reticulate rugose,
keels;
carina complete;
wi th
POL = 7;
Ol
scutum shiny,
smooth, weakly punctate, without sculpture among punctures; notaul ices incomplete, reaching approximately 0,3 length of scutum; scutellum and metanotum shiny,
smooth,
without sculpture;
propodeum reticulate rugose,
with a transversal keel between dorsal and posterior surface; posterior surface with 2 longitudinal keels; median area as rugose as lateral areas;
fore-wing hyaline,
radial
vein
250)
without dorsal
process;
tibial
lOCUS TYP ICUS: in BM;
proximal
part
bands;
(3:10);
membranous process,
distal
part of
gonoforceps
(Fig.
without distal
inner
spurs 1,1,2. Port St.
TYPICAL MATERIAL: MM)!
without dark transversal
much shorter than proximal
Johns (South Africa)
holotype F!,
allotype M!
and 5 paratypes
(3 FF,
HOSTS: unknown DISTRIBUTION: Ethiopian region: SOUTH AFRICA: Port St. Johns (Pondoland), BM! Jonkershoek (Stellenbosch), TW! Eshowe (Zulu land) , Mi Inerton
Fig.
(Cape Town),
250 -
2
4 paratypes in TV.
OLl
BM!
Male genital ia of Anteon inflatrix Benoit from Port St. (South Africa):
left half removed.
Johns
402
Mem. Amer.
En!.
Inst.,
no.
37,
1984
29. Anteon rufonigrum n. sp.
FEMALE:
fully
winged;
length 4,37 mm;
brown-b I ack spot on vertex; reddish;
scutum,
head reddish,
antennae and
scutellum and metanotum reddish,
propodeum black;
with a
legs testaceous;
wide
pronotum
with brown nuances;
mesopleura and metapleura reddish;
abdomen brown-
testaceous; antennal segments in following proportions: 16: 7: 8: 6: 6: 8: 7: 7: 10; head dull, fully reticulate rugose, with frontal line; POL 10; Ol = 7;
OOl = 6; OPl = 8;
pronotum hairy,
shiny,
shorter than scutum
Tl = 3;
occipital
reticulate rugose;
(8:20);
pronotal
granulated,
fully reticulate rugose;
dull,
granulated,
with some punctures;
propodeum reticulate rugose, posterior surface;
251
keel
fore-wing
band beneath the pterostigma;
Fig.
-
scutum
scutellum
metanotum reticulate rugose; between dorsal
posterior surface with 2 longitudinal areas;
tegulae;
notaulices invisible;
with a transversal
area nearly as rugose as lateral transversal
carina complete;
posterior surface of pronotum
tubercles reaching
dull,
7:
distal
keels;
median
with a dark
part of radial
Chela of Anteon rufonigrum n.
sp.
(type)
vein
and
Anteoninae
much shorter than proximal
part
proportions:
14; enlarged claw
(Fig.
segment 5 of front
tarsus
12: 2,5:
on a proximal
3:
4:
prominence;
bristles and 10 lamellae, part;
(5:11);
403
with basal region much
segment 4 of front tarsus produced
as basal
part of segment 5;
MALE: unknown LOCUS TYPICUS:
Port St.
TYPICAL MATERIAL:
tibial
Johns
segments in following 251)
with one bristle
(Fig.
251)
with some
longer than apical
into a hook,
nearly 0,3 as
long
spurs 1,1,2.
(South Africa)
holotype F!
HOSTS: unknown DISTRIBUTION: Ethiopian region: (Pondoland),
fore tarsal
in BM. SOUTH AFRICA:
Port St.
Johns
BM!
30. Anteon oriphilum Kieffer
Anteon oriphilus Kieffer 1912a: 47. Anteon oriphilus Kieffer: Kieffer 1914b: 166.
FEMALE: MALE:
unknown
fully
teeth brown;
Fig.
252 -
winged;
length 2,25 mmj
antennae,
tegulae and
blackj
mandibles testaceous,
legs fully
testaceous;
Male genital ia of Anteon oriphilum Kieffer removed.
with
head shiny,
(type):
left half
404
Mem. Amer.
strongly punctate, longitudinal
En!.
Inst.,
no.
37,
1984
without sculpture among punctures;
median keel
and with 2 lateral
POL = 5;
antenna I sockets;
OL = 3; OOL = 4;
scutellum and metanotum shiny, sculpture among punctures; length of scutum;
smooth,
very
frons with a
keels directed towards OPL = 2;
TL = 2;
weakly punctate,
notaulices distinct,
propodeum with a transversal
incomplete, keel
scutum, without
reaching
between dorsal
midand
posterior surface; dorsal surface reticulate rugose; posterior surface with 2 longitudinal keels; median area shiny, smooth, with keels only near margins; lateral areas reticulate rugose; mesopleura and metapleura rugose, with posterior surface smooth and without sculpture; fore-wing hyaline, with distal part of radial vein much shorter than proximal part (3:6); gonoforceps (Fig. 252) without dorsal proximal membranous process and without distal inner pointed process; tibial spurs 1,1,2. LOCUS TYPICUS: Mount Sebert (Seychelle Islands) TYPICAL MATERIAL: holotype M! in BM. HOSTS: unknown DISTRIBUTION: Ethiopian region: SEYCHELLE ISLANDS: Mount Sebert (Mahe), BM!
31. Anteon fabulosum n. sp.
FEMALE: MALE:
unknown
fully
testaceous; black;
winged;
length 2,18 mm;
legs testaceous,
black;
mandibles and antennae
with hind coxae basally black;
antennae not distally thickened;
antenna I segments
femora partly in following
proportions: 9: 5: 5: 5,5: 5: 5: 5: 5: 5: 8; head dull, granulated and punctate; frons with a median keel; occipital carina complete; POL = 7; OL = 5;
OOL = 5;
OPL = 4;
TL = 4;
scutum shiny,
smooth,
weakly
punctate, without sculpture among punctures, granulated only near anterior margin: notaulices incomplete, reaching approximately 0,65 length of scutum; scutellum and metanotum shiny, smooth, without sculpture; propodeum reticulate rugose, with a strong transversal keel between dorsa I and posterior surface;
posterior surface wi th 2
longitudinal keels; median area shiny, with central region smooth, with some keels around margins; fore-wing hyaline, without dark transversal bands;
distal
gonoforceps
part of radial
(Fig.
LOCUS TYP I CUS:
253)
Jonkershoek
TYPICAL MATERIAL: HOSTS:
vein shorter than proximal
with a distal
inner process;
part
tibial
(South Africa)
holotype M!
in TW.
unknown
DISTRIBUTION:
Ethiopian region:
(Stellenbosch),
TW!
SOUTH AFRICA:
Jonkershoek
32. Anteon abruptum n. sp.
(3:6,5);
spurs 1,1,2.
405
Anteoninae
FEMALE: MALE:
unknown
fully
wingedj
antennae brown,
coxae partly blackj tibiae brown j
length 2,37-2,62 mmj
antennae not distally
granulated and fully
253 -
9:
5:
6:
6:
6:
thickened; 7:
reticulate rugose;
6:
6:
mandibles testaceous;
legs testaceous,
stalks of hind femora brown;
following proportions:
Fig.
blackj
with segments 1-2 testaceousj
distal
antennal 6:
10;
segments
removed.
n.
sp.
in
head dull,
frons with a short
Male genitalia of Anteon fabulosum
with hind
half of hind
incomplete
(type):
right half
406
Mem.
median keel
Amer.
Ent.
I nst.,
no.
in front of the anterior ocellus;
37,
1984
occipital
carina complete;
POL = 6; Ol = 4; OOl = 7; OPl = 4; Tl = 3; scutum shiny, granulated; posterior region punctate, without sculpture among punctures; notaulices incomplete,
reaching approximately 0,3
metanotum shiny, punctures;
smooth,
length of scutum;
weakly punctate,
propodeum ret i cu I ate rugose,
between dorsal
and posterior surface;
scutellum and
without sculpture among
wi th a strong
transversa I keel
posterior surface of propodeum with
2 longitudinal keels; median area reticulate rugose,
dull,
with areolae
smaller than those of lateral areas; gonoforceps (Fig; 254) without dorsal proximal membranous process and without distal inner pointed process; tibial
spurs 1,1,2.
lOCUS TYPICUS:
Pietermaritzburg
TYPICAL MATERIAL:
holotype M!
(South Africa) and 1 paratype M!
in TW;
1 paratype M!
in Ol. HOSTS:
unknown
DISTRIBUTION: TW!
Ethiopian region:
Hluhluwe Game Res.,
TW!
SOUTH AFRICA:
Pietermaritzburg
(Natal),
Ol!
33. Anteon zairense Benoit
Fig.
254 -
Male genitalia of Anteon abruptum n.
sp.
(type)
Anteoninae
407
Anteon zairense Benoit 1951d: 21.
FEMALE: MALE:
unknown
fully
winged;
teeth brown;
tegulae testaceous; of femora,
length 1,87-2,38 mm;
black;
mandibles yellow,
with
antennae fully brown or with segments 1-3 testaceous; legs brown or testaceous,
hind tibiae brown;
head shiny,
with
hind coxae,
hind clubs
reticulate rugose and
granulated; frons with a longitudinal median keel, without lateral keels; POL ~ 8; OL ~ 4; OOL ~ 6; OPL ~ 4; TL ~ 4; scutum shiny, strongly rugose near anterior marg in, half;
notaul ices distinct,
smooth and weak I y
incomplete,
scutell um and metanotum sh i ny, a transversal
keel
reticulate rugose;
smooth,
between dorsal
posterior surface with 2 longitudinal
area as rugose as lateral
255)
areas,
with numerous keels;
without dorsal
LOCUS TYPICUS:
255 -
part
(5:10);
propodeum wi th dorsal
keels;
surface
median
mesopleura and with distal
gonoforceps
part of (Fig.
proximal membranous process and without distal
with apex very
narrow;
tibial
inner
spurs 1,1,2.
Basoko (Zaire)
TYPICAL MATERIAL:
Fig.
fore-wing hyaline,
vein much shorter than proximal
pointed process,
weak I y punctate;
and posterior surface;
metapleura fully reticulate rugose; radial
punctate on posterior
reaching mid-length of scutum;
holotype M!
in TV.
Male genitalia of Anteon zairense Benoit removed.
(type):
left half
408
Mem. Amer.
HOSTS:
Ent.
Inst.,
no.
37,
1984
unknown
DISTRIBUTION: Jonkershoek
Ethiopian region:
(Stellenbosch),
OL!
ZAIRE: TW!
Basoko,
TV!
Grahamstown,
SOUTH AFRICA:
TW!
34. Anteon mosseli n. sp.
FEMALE:
unknown
MALE: fully winged; length 2,25-3,43 mm; black; mandibles and tegulae testaceous; antennal segments in following proportions: 8: 5: 10: 9: 9: 9: 9:
9:
9:
punctate, OPL
9;
antennae not distally thickened;
without sculpture among punctures;
head shiny, POL = 8;
strongly
OL = 4;
OOL = 7;
= 3; TL = 5; occipital carina complete; scutum shiny, strongly
punctate,
without distinct sculpture among punctures;
incomplete, reaching mid-length of scutum;
Fig.
256 -
Male genitalia of Anteon mosseli n. removed.
notaulices
scutellum and metanotum
sp.
(type):
right half
409
Anteoninae
smooth,
shiny,
without sculpture;
without transversal hyal ine,
keel
propodeum fully
between
dorsal
without dark transversal
shorter than proximal
part
(3:9);
membranous process and without
bands;
distal
gonoforceps
distal
reticulate rugose,
and posterior surface; part of radial
(Fig.
256)
fore-wing vein much
without proximal
inner pointed process;
tibial
spurs
1,1,2. lOCUS TYPICUS: Mossel Bay (South Africa) TYPICAL MATERIAL: holotype M! in BM; 10 paratypes MM!
in TW;
4
para types MM! in Ol. HOSTS: unknown DISTRIBUTION: Province),
Ethiopian region:
BM!
Jonkershoek
SOUTH AFRICA:
(Stellenbosch),
TW!
35. Anteon capei n.
FEMALE: MALE:
winged;
antennae brown;
Fig.
Bay
(Cape
sp.
unknown
fully
segments
Mossel Ol!
length 3,12 mm;
legs brown;
in following proportions:
257 -
black;
mandibles testaceous;
antennae not distally thickened;
Male genitalia of
8:
5:
10:
Anteon capei removed.
11,5:
n.
11,5:
sp.
11,5:
(type):
antennal 10:
10: 9:
right half
410
Mem.
9; head shiny, punctures; furrow;
En!.
Inst.,
weakly punctate,
no.
= 7;
= 4;
OL
scutum shiny,
punctures;
= 6;
OOL
smooth,
smooth,
= 3;
OPL
keels,
1984
with a weak median
= 5;
TL
weakly punctate,
notaulices incomplete,
and metanotum shiny,
37,
without sculpture among
frons without median and lateral
POL
complete;
smooth,
Amer.
occipital
carina
without sculpture among
reaching mid-length of scutum;
weakly
punctate,
scutellum
without sculpture among
punctures; propodeum reticulate rugose, without a transversal keel between dorsal and posterior surface; posterior surface without longitudinal keels; fore-wing hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part (3:9); gonoforceps distal
(Fig.
257)
inner process;
without proximal tibial
LOCUS TYPICUS: Matjesfontein TYPICAL MATERIAL:
(South Africa)
holotype M!
HOSTS: unknown DISTRIBUTION: Ethiopian region: Province),
membranous process and without
spurs 1,1,2. in BM. SOUTH AFRICA:
Matjesfontein
(Cape
BM!
36. Anteon emeritum n. sp.
FEMALE: MALE:
unknown
fully
winged;
length 1,87 mm;
antenna I segments 1-5 testaceous
black;
mandibles,
tegulae and
(other segments missing
in
the only
(
, \
/
j../
/1
\
I
\ I
Fig.
258 -
Male genitalia of Anteon emeritum n. removed.
sp.
(type):
left half
Anteoninae
known specimen); tibiae brown;
legs testaceous,
antennal keels;
with hind coxae,
hind femora and hind
segments in following proportions: 7:
(other segments missing irregular weak
411
in
the holotype);
occipital
head granulated,
carina complete;
4:
Tl ;
punctures;
without
lateral
keels,
frons with a
notaulices incomplete,
3; OOl ;
4:
4
longitudinal
2; 3; scutum shiny, smooth, finely punctate, without sculpture among
median keel,
POL; 6; Ol ;
4:
with numerous
5; OPl ;
reaching approximately 0,25 length of
scutum; scutellum and metanotum shiny, smooth, without sculpture; propodeum reticulate rugose, with a strong transversal keel between aorsal
and posterior surface;
with areolae smaller than without dark transversal than proximal
part
posterior surface without
those of dorsal bands;
distal
tibial
longitudinal
keels,
fore-wing hyaline,
part of radial
vein much shorter
(2:6); gonoforceps (Fig. 258) with dorsal proximal
membranous process and with a distal apex;
surface;
inner pointed process very
near
spurs 1,1,2.
lOCUS TYPICUS:
Okahandja
TYPICAL MATERIAL:
(South West Africa)
holotype M!
HOSTS: unknown DISTRIBUTION: Ethiopian region:
in BM. SOUTH WEST AFRICA:
Okahandja,
BM!
37. Anteon garambanum n. sp.
FEMALE: MALE:
Fig.
unknown
fully
259 -
winged;
length 1,62 mm;
black;
mandibles testaceous;
Male genitalia of Anteon garambanum n. removed.
sp.
(type):
left half
412
Mem.
Amer.
Ent.
Ins!.,
no.
antennae brown,
with segment 1 testaceous,
and hind tibiae,
hind coxae partly brown;
37,
1984
with clubs of femora,
mid
antennae not distally
thickened; antennal segments in following proportions: 7: 4: 3: 3,5: 3: 3,5: 3,5: 3,5: 3,5: 6; head dull, fully granulated; frons with median keel and without lateral keels; occipital carina complete; POL =5; OL = 3; OOL = 4; OPL = 1,5; TL = 2; scutum shiny, smooth, without sculpture; notaul ices incomplete, reaching approximately 0,3 length of scutum; scutellum and me\anotum shiny, smooth and without sculpture; propodeum reticulate rugose, with a strong transversal keel between dorsal and posterior surface;
posterior surface without
longitudinal
keels;
fore-wing
hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part (1 :4); gonoforceps (Fig. 259) without proximal membranous process and with a distal
inner pointed process;
tibial
spurs
1,1,2. LOCUS TYPICUS:
Garamba National
TYPICAL MATERIAL: HOSTS:
holotype M!
Park
(Zaire)
in TV.
unknown
DISTRIBUTION:
Ethiopian region:
ZAIRE:
Garamba Nat.
ORIENTAL SPECIES OF
Twenty
Park,
TV!
ANTEON
two species are known.
KEY TO THE SPECIES
FEMALES
Segment
4 of front
tarsus at
most
0,5 as
long as
segment
1
(Fig. 260) •••••.••••••••••••••••••••••••••••••••••••••••••••• 2 Segment 4 of front tarsus slightly shorter, as long as, or longer
than
1
segment
277) ••••••••.•••••••••••••••••••• 7 longitudinal keels ••••• 1. borneanum n. sp.
(Fig.
2
Posterior surface of propodeum without
3
Posterior surface of propodeum wi th two longi tudinal keels ••• 3 Head fully strongly reticulate rugose •••••••.•••••••••••••••• 4 Head at
most
reticulate
4
wi th
rugose;
irregular keels or wi th never fully
Scutum sculptured by
longitudinal
weakly punctate ••••••••• Scutum and
only vertex rugose ••••••••••••• 5
reticulate keels;
scutellum smooth,
22. philippinum n.
sp.
scutellum strongly punctate wi th very wide
impressions,
similar
to areolae,
without
. . . . . . . . . . . . . . . . . . . . . . . . . . 2. thai n.
sp.
longitudinal
keels •••
413
Anteoninae
5
Posterior lateral
surface of propodeum wi th median area as
3.
areas . . . . . . . . . . . . . . . . . . . . . . . . .
Posterior
rugose as
silvicolum n. sp.
surface of propodeum with median area
smooth,
not
rugose ••••••••••••••••••••••••••••••••••••.•••••••••••••••••••• 6
6
Head punctate,
without
sculpture among punctures ••••••••••••••••
· . . . . . . . . . . . . . . . . . . . . • . . . . . . . . . . . . . . •. 4. indicum n. Head punctate,
weakly granulated and
with
weak
sp.
irregular keels •.
· . . . . . . • . . . . . . . . . . . . . . . . . . • . . . . . . . . • .. 5. mysorense n. 7
Posterior surface of propodeum without Posterior surface of propodeum wi th
8
Head granulated, not for
the frontal
rugose,
line and
two
without
to
9
rugose,
the
not
two frontal
frontal
Segment
of
granulated,
line and
front
two
lateral
lateral
tarsus nearly as
Segment
of
front
scutum;
head not
keels .••• 11
6.
except
keels near orbits ..
yasumatsui n. sp.
wi th areolae or keels
(in addition
keels near orbits) . . . . . . . . . 9 long as
• • . • • . . • . . • . . . • . • . . • . . . . • . . . . • . . . . • . .. 10 Notaulices shorter,
longi tudinal
areolae and keels,
• . . . . . . . . . • . . . • . . • . . . • . • • . . . • . . • . . • . •• Head
sp. keels . . . . . . 8
longitudinal
7.
segment
4 . . . . . . . . . ..
sarawaki n. sp.
tarsus much shorter than segment 4 •••••••.• 10 faintly visible near anterior margin of
reticulate
rugose on
sides of
ocellar
triangle
bengalense n. sp. Notaulices longer, reaching approximately 0,5 length of scutum; head fully reticulate rugose . . . . . . . . . . 9. krombeini n. sp. · . . . . . . . . . . . • . . . . . . • . . • . . . • . . • • . . . . • ..
8.
11 Head smooth, more or less punctate, wi thout sculpture among punctures,
not
reticulate
rugose . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 12
Head rugose, fully reticulate 12 Head with OPL slightly longer
rugose •••••••••••••••••••••••••• 14 than breadth of posterior ocelli ..
13. Javanus n. Head wi th OPL nearly
twice as
long as breadth
of
sp.
posterior
oce I Ii ••••••••••••••••••••••••••••••••••••••••.••.••.••••••••• 12
13 Head with OL slightly longer than POL; species fully testaceous, with petiole black . . . . . . . . . . . . . . . . . . . 14. munroei n. sp. Head wi th OL nearly
twice as
long as POL;
species almost
fully
black •••••••••••••••••••••• '" ••••••• 15. lankanum n. sp. 14 Fore-wi ng hya line, wi thou t dark transversa I bands . . . . . . . . . . . . . . .
• • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • •• 10. muiri n. sp. 1-2 dark transversa I bands . . . . . . . . . . . . . . . . . . . . . 15
Fore-wi ng wi th
15 Pronotum fully smooth, without sculpture or weakly punctate . . . .. • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • . •• 11. laotianum n. sp. Dorsal
surface of pronotum strongly
• ••.•..••.•••••.••••••••••••••••.•. "
sp.,
reticulate
12.
rugose •••••••••••
fyanense n.
sp.
The females of A. nemorale n. sp., silvestre n. sp., expolitum n. peterseni n. sp., bauense n. sp. and debile n. sp. are unknown.
414
Mem.
Amer.
En!.
Inst.,
no.
37, 1984
MALES
2
Posterior
surface of
propodeum without
Posterior
surface of
propodeum with
Gonoforceps with a
longitudinal
two
keels . . . . . . 2 keels . . . . . 7
longitudinal
281) 16. nemorale n. sp. inner process (Figs. 270, 282) . . . . . . 3
distal
inner pointed process
(Fig.
• • • • • • • • • • • • . • . • • • • • • • • • • • • • • . • • • • • •• 3
Gonoforceps wi thout di stal Scutum fully reticulate rugose;
notaulices
• • • • • • • • • • • • • • • • • • • • • • • • • • • • . • • • • • • .• Scutum mostly
4
invisible . . . . . . . . . . ..
silvestre n. sp.
sculpture or punctate,
not
fully
ret iculate
rugose;
notaul ices visible . . . . . . . . . . . . . . • • • . . . . . . . . . 4
Head dull,
smooth,
granulated . . . . . . . . .
Head
5
or fully without
17.
shiny,
rugose,
6. yasumatsui n. sp.
irregular keels . . . . . . . . . . . 5
with areolae and
6
reaching approximately 0,65 length of scutum . . . . . . . . . · . . . . . . . . . . . . . . . . . . . . . . . . • . . . . . . . . • . .. 7. sarawaki n. sp. Notaulices reaching at most 0,5 length of scutum . . . . . . . . . . . . . . . 6 Notaul ices reaching approximately 0,5 length of scutum . . . . . . . . .. · . . . . . . . . . . . . . . • . . . . . . . . . . . . . . . . . . . . .. 9. krombeini n. sp. Notaulices reaching approximately 0,3 length of scutum . . . . . . . . . . · . . • • • . . . . . . . . . . . . . • • . . . . . . • . . . . . . . . .. 2. thai n. sp.
7
Posterior
surface of
propodeum wi th median
fully
not
reticulate
rugose . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8
Posterior
surface of
propodeum wi th median
Notaulices
I ate ra I
8
area
smooth,
area as
almost
rugose as
area s ••••••••••••••••••••••••••••••••••••••••••••••••• 1 2
(Fig. 266) ••••.• 5. mysorense n. sp. Gonoforceps without distal inner process (Figs. 283, 285) . . . . . . 9 Notaul ices reaching approximately 0,5 length of scutum . . . . . . . . 10 9 Notaulices reaching approximately 0,65 length of scutum . . . . . . . 11 10 Head fully or almost fully reticulate rugose . . . . . • • . . . . . . . . . . • • • • • . • • . . • • • • . • • . • • . . . . . . • • • • • • . • • . . . •• 18. expolitum n. sp. Gonoforceps wi th a
distal
inner
rounded process
· . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ..
Head
fully
or almost
fully
weakly punctate •••••••••• ••••••••••••
• • • • • • . • • • • • • • . • • • • • • • . . . • • • • • . • • • • •• 11 Gonoforceps nearly as strongly
punctate;
long as penis
notaulices
punctate;
12 Head fully
shorter
than penis
Head punctate,
not
rugose . . . . . . . . . .
reticulate
20.
(Fig.
notaul ices weak . . . . . . . . . . . . reticulate
peterseni n.
sp.
284); head more
strong . . . . . . . . . . . . . . . . . . . . . . . . . . ..
• ••••..••......•••••••••••••..•.••••• Gonoforceps much
1 9.
(Fig.
21. 2.
bauense n.
sp.
285); head weakly
debile n. sp. thai n. sp.
rugose . . . . . . . . . . . . . . . . . . . . . . . . . . ..
• • • • . • . . • • • • • • . • • • • • • • • • • • • • • • • • • • • ••
13. javanum n.
sp.
The males of A. philippinum n. sp., munroei n. sp., silvicolum n. lankanum n. sp.; bengalense n. sp., laotianum n. sp., borneanum n. sp., muiri n. sp., fyanense n. sp. and indicum n. sp. are unknown. sp.,
Anteoninae
Fig.
260 -
Chela of Anteon borneanum n.
415
sp.
(type)
416
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
1. Anteon borneanum n. sp.
FEMALE:
fully
testaceous;
winged;
length 1,87 mm;
legs testaceous,
black;
mandibles and antennae
with hind coxae partly brown;
clubs of femora partly brown;
mid and hind
antennae distally thickened;
antenna I
segments in following proportions: 12: 5: 5: 4: 4: 4: 4: 4: 4: 5; head shiny, granulated, with numerous irregular keels; frons with a median keel and 2 lateral keels around orbits; occipital carina complete; POL = 5; OL
3; OOL keel s;
= 5; OPL = 3; TL = 3; pronotum shiny, with numerous transversal posterior surface of pronotum smooth, short, shorter than scutum
(3:14);
pronotal
tubercles reaching tegulae;
scutum shiny,
except for the anterior region near pronotum which smooth;
notaulices incomplete,
reaching approximately 0,5
scutell um and metanotum sh i ny, reticulate rugose, posterior surface;
smooth,
granulated,
is not sculptured and
wi thout scu I pture;
length of scutum; propodeum
with a strong transversal keel between dorsal and posterior surface without longitudinal keels; fore-wing
hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part (3:7); fore tarsal segments in following proportions: a proximal than apical
11
1,5: 2:
5:
2: 9; enlarged claw (Fig. 260) with one bristle on
prominence;
segment 5 of front
lamellae extending continuously
MALE: unknown LOCUS TYPICUS:
to the apex;
Nanga Pelagus (Sarawak,
TYPICAL MATERIAL: HOSTS:
tarsus with basal
(7:2); segment 5 of front tarsus (Fig. 260)
part
holotype F!
tibial
part
longer
with 2 rows of
spurs 1,1,2.
Malaysia)
in B.
unknown
DISTRIBUTION: Sarawak),
Oriental
region:
MALAYSIA:
Nanga Pelagus (near Kapit,
B!
2. Anteon thai n. sp.
FEMALE:
fully
winged;
length 4,6 mm;
black,
with mandibles and
legs
testaceous; antennae testaceous, wi th segments 4-10 brown; antennae distally thickened; antennal segments in following proportions: 9: 5: 4,5: 5: keel; 4;
5:
5:
5:
7;
head shiny,
head fully reticulate rugose;
OOL
hairy,
5:
=
5;
OPL
=
5;
TL
=
3;
hairy;
occipital
carina complete;
pronotum dull,
nearly twice as broad as
long;
strongly punctate,
pronotal
with punctures very
without sculpture among punctures; approximately 0,3 length of scutum; metanotum,
shiny,
anterior margin;
smooth,
= 6; OL
tubercles reaching tegulae; (5:13);
broad,
scutum shiny,
similar to areolae,
notaul ices incomplete,
reaching
scutellum as sculptured as scutum;
without sculpture,
propodeum fully
POL
fully reticulate rugose,
posterior surface of pronotum shorter than scutum hairy,
6:
frons with a track of median
with some areolae only near
reticulate rugose,
dull,
with a strong
Anteoninae
417
Fig.
261
-
Chela of Anteon thai n.
sp.
(type)
418
Mem.
transversal with two
keel
keels;
proportions:
50:
on a proximal
Inst.,
10:
37,
lateral
with some transversal
11:
no.
distal
part of radial
fore-wing vein much
segments in following
enlarged claw
segment 5 of front
group of 2 lamellae;
posterior surface
areas of posterior surface
keels on median area;
bands;
(5: 18); fore tarsal
16: 45;
prominence;
1984
and posterior surface;
median and
without dark transversal
shorter than proximal part
with a
Ent.
between dorsal
longitudinal
reticulate rugose, hyaline,
Amer.
(Fig.
261)
with one bristle
tarsus with 2 lamellae;
segment 5 of front
tarsus
(Fig.
261)
apex
with basal
part longer than apical part (37:8); tibial spurs 1,1,2. MALE: fully winged; length 1,68-2,88 mm; black; mandibles and antennae testaceous;
legs testaceous,
with
hind coxae proximally black;
ditally thickened;
antenna I segments in following proportions:
5:
head shiny,
5:
keel
5:
4:
5:
9;
fully reticulate rugose;
antennae not 10: 7:
6:
5:
frons with a median
and with 2 lateral keels near orbits directed towards antenna I sockets;
occipital carina complete; scutum,
POL
= 7; OL = 4; OOL = 7; OPL = 4; TL = 4;
scutellum and metanotum shiny,
sculpture among punctures; strongly punctate;
smooth,
weakly punctate,
without
anterior surface of scutum reticulate rugose or
notaulices incomplete,
reaching approximately 0,3 length
of scutum; propodeum reticulate rugose, with a strong transversal keel between dorsal and posterior surface; posterior surface with 2 strong or weak
(almost invisible)
lateral
areas;
part of radial
Fig.
262 -
longitudinal
fore-wing hyaline,
keels;
median area as rugose as
without dark transversal
vein much shorter than proximal
Male genital ia of Anteon thai
n.
sp.
left half removed.
part
bands;
(3,5:9);
distal
gonoforceps
from Chawang
(Thai land):
419
Anteoninae
262) without proximal membranous process and without distal inner tibial spurs 1,1,2.
(Fig.
process;
LOCUS TYPICUS:
Mudigere (India)
TYPICAL MATERIAL:
holotype F!
2 paratypes MM!
in B; HOSTS:
in WA;
and 1 paratype M!
3 paratypes MM!
in CO;
2 para types MM!
in OL.
unknown
DISTRIBUTION:
Oriental
region:
THAilAND:
Chawang
(Nabon,
Banna),
Banna, B! HONG KONG: Taipokau (Kowloon), OL! INDIA: Mudigere (Karnataka), OL! CO! SRI LANKA: Udawattakele Sanct. (Kan. Dist.), Ol!
Kande Ela
(NE Dist.),
B! WA!
WA!
3. Anteon silvicolum n. sp.
FEMALE: fully winged; length 3,20 mm; black, with mandibles, antennae and legs testaceous; antennae distally thickened; antenna I segments in following proportions: punctate,
14: 7: 12: 10: 10: 9: 9: 8: 8:
wi thout scu I pture among punctures,
anterior half of frons; reticulate rugose; keel;
POL = 7;
rugose;
occipital
OL = 6;
pronotal
punctate,
vertex sculptured by
with a strong transversal areas;
part of radial
keel
two
smooth,
notaul ices incomplete,
scutellum and metanotum shiny,
without sculpture among punctures;
posterior surface with
pronotum reticulate
scutum shiny,
without sculpture among punctures;
punctate,
punctate on
frons with a complete median TL = 4,5;
tubercles reaching tegulae;
reaching 0,5 length of scutum;
lateral
OPL = 5;
head shiny,
irregular keels and partly
carina complete;
OOL = 8;
11;
more strong I y
between dorsal
longitudinal
fore-wing hyaline,
keels;
and posterior surface; median area as rugose as
without dark transversal
vein much shorter than proximal
segments in following proportions:
smooth,
propodeum reticulate rugose,
part
bands;
distal
(5:15); fore tarsal
12: 3: 3: 6: 15; enlarged claw (Fig. 263)
with one bristle on a proximal prominence; segment 5 of front tarsus (Fig. 263) with 1 row of 18 lamellae; apex with a group of nearly 6 lamellae; tibial
spurs 1,1,2.
MALE unknown lOCUS TYPICUS:
Kanda-ela Reservoir
TYPICAL MATERIAL:
holotype F!
HOSTS: unknown DISTRIBUTION: Oriental region: Nuwara Eliya,
N.E.
Dist.),
(Sri
lanka)
in WA. SRI
lANKA:
Kanda-ela Reservoir
(5.6 mi. SW
WA!
4. Anteon indicum n. sp.
FEMALE:
fully
testaceous;
winged;
legs fully
length 2,18-3,12 mm;
black;
mandibles and antennae
testaceous or with hind coxae and hind clubs of
420
Mem. Amer. En!.
Inst., no. 37, 1984
Fig. 263 - Chela of Anteon silvicolum n. sp.
(type)
Anteoninae
Fig. 264 - Chela of Anteon indicum n. sp.
421
(type)
422
Mem. Amer.
femora
brown;
En!.
Inst.,
antennae slightly distally
following proportions:
10:
4:
5:
6:
5:
no.
5:
5:
without scu I pture among punctures;
punctate;
frons without median
median keel; TL = 3; short,
occipital
and
carina complete;
pronotum shiny,
with
shorter than scutum
keels;
keels,
pronotal
segments
head shiny,
in
weakly
somet i mes more strong I y
= 7;
POL
antennal
9;
5:
lateral
transversal
(6:16);
1984
thickened;
5:
puncta te,
keel
37,
OL
sometimes with
= 5;
= 4;
OOL
5-,
OPL
with posterior surface
tubercles reaching
tegulae;
scutum shiny, smooth, weakly punctate, without sculpture among punctures; notaul ices incomplete, reaching approximately 0,3 length of scutum; scutellum and metanotum shiny,
without sculpture;
with a
posterior surface;
posterior surface with 2
shiny,
smooth,
also in central
area;
part of radial
tarsal
segments
fore-wing hyaline,
front
tarsus with basal (Fig.
front
tarsus
part
(Fig.
tibial
longitudinal
264)
and
median
area
occasionally with keels part
bands;
(3:7);
fore
9: 2: 2: 2,5: 12,5; segment 5 of
proximal
part
(10:2,5);
prominence;
1 row of 5 lamellae;
with
keels;
without dark transversal
longer than proximal
with one bristle on a
propodeum
between dorsal
near margins;
in following proportions:
264)
keel
vein much shorter than proximal
claw
MALE:
transversal
with a few keels only
distal
lamellae;
strong
smooth,
reticulate rugose,
enlarged
segment 5 of
apex with a group of 16
spurs 1,1,2.
unknown
LOCUS TYPICUS:
Bangalore
TYPICAL MATERIAL: HOSTS:
in
India
DISTRIBUTION:
(India)
holotype F!
and
paratype F!
INDIA:
Bangalore
in BM.
ldiocerus sp.
Oriental
region:
(Mysore),
BM!
5. Anteon mysorense n. sp.
FEMALE:
fully
testaceous; thickened; 3:
4:
keels;
6;
winged;
length 2 mm;
legs testaceous, antennal
with
mandibles and antennae fully
segments in following proportions:
head shiny,
punctate,
keels;
POL
9:
antennae distally 4:
3:
3:
weakly granulated and with weak
occipital carina complete;
incomplete lateral
black;
hind coxae partly black;
=
frons with a median keel 6;
OL
=
4;
OOL
=
4;
TL
=
3:
3:
3:
irregular
and with 2 4;
pronotum
shi ny, wi th SOme transversa I keel s, with posterior surface shorter than scutum (4:10); pronotal tubercles reaching tegulae; scutum shiny, smooth, weakly punctate, without sculpture among punctures; notaulices incomplete, reaching approximately 0,3 length of scutum; scutellum and metanotum shiny, smooth, between dorsal keels;
without sculpture; propodeum with a strong transversal keel and posterior surface; posterior surface with 2 longitudinal
median area shiny,
fore-wing hyal ine,
smooth with a few keels only near margins;
without dark transversal
vein much shorter than proximal following proportions: basal
part
6:
1,5:
longer than apical
part
1,5: part
bands;
(3:5);
1,5: 9; (6:3);
fore
distal
tarsal
part of radial
segments
segment 5 of front enlarged claw
(Fig.
in
tarsus with 265)
with
Anteoninae
423
265 - Chela of Anteon mysorense n. sp.
Fig.
(type)
one bristle on a proximal prominence; segment 5 of front tarsus with 4 lamellae; apex with a group of nearly 9 lamellae; tibial
(Fig. 265) spurs
1,1,2. MALE: fully
fully
length 2,18 mm;
winged;
testaceous;
following proportions: 9: punctate, rugose, without
black;
mandibles,
antennae not distally thickened;
antennae and
antennal
legs
segments in
5: 5: 5: 5: 5: 5: 5: 5: 8; head shiny, strongly
with vertex and region between ocelli
and occipital carina
without sculpture among punctures; frons with a median keel, lateral keels; occipital carina complete; POL = 7; Ol = 3,5; OOl
4; OPl = 4;
Tl = 4;
anterior half,
scutum shiny,
smooth,
weakly
without sculpture among punctures;
reaching approximately 0,3 length of scutum;
punctate,
mostly on
notaulices incomplete,
scutellum and metanotum
shiny, smooth, without sculpture; propodeum with a strong transversal keel between dorsa I and posterior surface; posterior surface wi th 2 longi tud i na I keels; lateral distal
median area shiny, areas rugose; part of radial
gonoforceps (Fig.
smooth,
with a few keels only near margins;
fore-wing hyaline,
without dark transversal
vein much shorter than proximal
part
bands;
(3:5);
266) without proximal membranous process and with distal tibial spurs 1,1,2.
inner rounded process; lOCUS TYPICUS:
Malleswaram
TYPICAL MATERIAL:
(India)
holotype F!
and 1 paratype M!
in BM;
1 paratype F!
INDIA:
(Mysore),
Ol. HOSTS: unknown DISTRIBUTION: Oriental
region:
Malleswaram
BM!
OL!
in
424
Mem.
Amer.
En!.
Inst.,
no.
37,
1984
266 - Ma Ie gen i ta I i a of Anteon mysorense n. sp. from Ma II eswaram
Fig.
(India):
right half removed.
6. Anteon yasumatsui n. sp.
FEMALE: fully 2,5:
fully
winged;
length 2,00 mm;
yellow-testaceous; 2,5:
2: 3:
longitudinal
3:
3:
antennal
3:
5;
median keel;
black;
antennae,
tegulae and
segments in fol/owing proportions:
head dull,
occipital
granulated;
frons
carina complete;
legs
8:
4:
with an incomplete
POL = 4;
OL = 3;
OOL
4; OPL = 3; TL = 5; pronotum granulated, with posterior surface slightly longer than scutum (8:6) and with transversal keels; pronotal tUbercles reaching tegulae; incomplete,
scutum shiny,
metanotum shiny, rugose,
smooth,
posterior surface without dark transversal proximal
part 9;
bands;
(2:4);
propodeum fully
between dorsal
longitudinal
keels;
fore-wing hyaline,
distal part of radial
fore tarsal
reticulate
and posterior surface; without
vein much shorter than
segments in following proportions:
segment 2 of front tarsus produced into a hook;
(Fig.
267)
nearly 4 lamellae; MALE:
with 2 rows of nearly 18 lamellae; tibial
fully winged;
tegulae and 5;
keel
notaulices
scutellum and
5:
1,5:
enlarged claw
267) with one bristle on a proximal prominence; segment 5 of front
(Fig. tarsus
=
without sculpture;
without sculpture;
with a transversal
2: 3,5:
smooth,
reaching approximately 0,5 length of scutum;
OL
=
3;
length 1,50-2,00 mm;
legs fully OOL
=
apex with a group of
spurs 1,1,2.
yellow-testaceous;
4; OPL
=
3;
TL
=
4;
black;
mandibles,
antennae,
head fully granulated, occipital
dull;
carina complete;
POL
pronotum
Anteoninae
Fig.
incomplete, smooth,
Chela of Anteon yasumatsui n.
267 -
short and rugose;
scutum shiny,
without
keel
distal
keels;
gonoforceps (Fig. without distal
268)
F)!
without dorsal
without dark transversal
proximal tibial
HOSTS:
2 paratypes
(1
F,
1 M)!
with a
posterior surface
vein much shorter than proximal
inner pointed process;
in BM;
notaulices
reticulate rugose,
and posterior surface;
part
(2:5);
membranous process and
spurs 1,1,2.
LOCUS TYPICUS: Phi bun Mangsahan (Thailand) TYPICAL MATERIAL: holotype F! and 1 paratype M! M,1
(type)
scutellum and metanotum shiny,
fore-wing hyal ine,
part of radial
sp.
without sculpture;
propodeum fully
between dorsal
longitudinal
bands;
smooth,
reaching 0,5 length of scutum;
without sculpture;
transversal
425
in VA;
2 paratypes (1
in OL.
unknown
DISTRIBUTION: Ratchathani),
Oriental VA!
region:
Non Sa-ard
THAILAND:
Phibun Mangsahan
(Maha Sarakham),
(Java), OL! INDIA: Delhi, BM! OL! NOTE: this species is named in honor of the
VA!
late Mr.
(Ubol
INDONESIA:
Pekalongan
Keizo Yasumatsu.
7. Anteon sarawaki n. sp.
"EMALE:
fully
2nd legs fully 3:
14: 9:
9:
9:
winged;
length 3,62 mm;
testaceousj 8:
8:
7:
blaCk;
antennae segments 9;
head shiny,
mandibles,
antennae,
tegulae
in following proportions:
rugose,
12:
with numerous areolae on
426
Mem.
Fig.
268 -
Amer.
Ent.
Inst.,
no.
37,
Anteon yasumatsui
Male genital ia of
1984
n.
sp.
from
Non Sa-ard
(Thailand)
frons and with
irregular keels on vertex and behind ocelli;
among areolae and keels; lateral
frons with a
keels around orbits;
POL = 6;
longitudinal OL = 4;
no sculpture
median keel
OOL = 6;
and 2
OPL = 5;
TL = 3;
occipital carina complete; pronotum hairy, rugose, shiny, strongly transversely striate and punctate, smooth only near posterior margin, posterior surface nearly as reaching
tegulae;
scutum shiny,
sculpture among punctures; margin of scutum;
long as scutum hairy,
(8:9);
smooth,
notaulices long,
pronotal
with
tubercles
weakly punctate,
without
almost reaching posterior
scutellum and metanotum shiny,
smooth,
weakly punctate,
without sculpture among punctures; propodeum reticulate rugose, with a transversal keel between dorsal and posterior surface; posterior surface without longitudinal keels; fore-wing hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part (3:12); fore tarsal
segments
in following proportions:
10:
3:
5:
10:
23;
enlarged
claw (Fig. 269) with one bristle on a proximal prominence; segment 5 of front tarsus (Fig. 269) with basal part much shorter than apical part, with 2 rows of nearly 35 1,1,2. MALE:
fully
tegulae and shiny, keel;
winged;
lamellae;
apex with a group of 4
length 2,12-2,50 mm;
legs fully
testaceous,
fully reticulate rugose, POL = 8;
OL = 4,5;
black;
lamellae;
mandibles,
tibial
spurs
antennae,
except for fore coxae basally black;
granulated;
OOL = 6;
head
frons with no visible median
OPL = 5;
TL = 3;
occipital
carina
complete; scutum hairy, shiny, smooth, strongly punctate, without sculpture among punctures; notaulices long, incomplete, almost reaching posterior margin of scutum;
scutellum and metanotum shiny,
propodeum reticulate rugose, posterior surface; hyal ine,
with a
transversal
posterior surface without
without dark transversal
bands;
smooth,
keel
longitudinal
distal
weakly punctate;
between dorsal keels;
part of radial
and
fore-wing vein much
Anteoninae
Fig.
269 -
shorter than proximal proximal
part
Gunong Mulu
TYPICAL MATERIAL: OL.
Anteon sarawaki n. sp. (type)
(3:11);
membranous process,
1,1,2. LOCUS TYPICUS:
HOSTS:
Chela of
427
gonoforceps
without distal (Sarawak,
holotype F!
(Fig.
270)
without dorsal
inner process;
tibial
spurs
Malaysia)
and 1 paratype M!
in BM;
1 paratype M!
in
unknown
DISTRIBUTION:
Oriental
region:
MALAYSIA:
Gunong Mulu
(Sarawak),
BM!
OL!
428
Mem.
Fig.
Amer.
Ent.
Inst.,
no.
37,
1984
270 - Male genitalia of Anteon sarawaki n. sp. from Gunong Mulu (Malaysia)
8. Anteon bengalense n. sp.
FEMALE:
fully
antennae
winged;
length
testaceous-dark;
5 mm;
legs
black;
mandibles
testaceous,
with
testaceous;
femora,
tibiae and
hind coxae partly brown; antennae distally thickened; antennal segments in following proportions: 20: 8: 13: 10: 8: 8: 8: 8: 8:
11; head shiny,
rugose,
frons wi th a median keel
wi thout and
also with numerous areolae; with
some areolae and
occipi tal
carina
(as
two
sculpture among areolae and keels; lateral
region
of
two oblique keels in
keels around
orbits;
from posterior ocelli
to
Deinodryinus); SOme areolae also near occipital
carina and on temples; POL = 5; OL = 3; OOL = 8; OPL = 8; TL = 7; pronotum shiny, with numerous transversal striae on anterior region, on posterior region,
frons
head behind posterior ocelli
without sculpture,
of pronotum shorter than scutum
with sides hairy;
(14: 16); pronotal
smooth
posterior surface
tubercles reachi ng
tegulae; scutum shiny, weakly punctate, without sculpture among punctures; notaulices incomplete, only visible near anterior margin of scutum; scutellum and metanotum shiny, among punctures; keel
between
smooth,
weakly
propodeum reticulate rugose,
dorsal and posterior surface;
punctate,
without sculpture
with a strong transversal
posterior surface rugose,
without
Anteoninae
Fig.
271
-
429
Chela of Anteon bengalense n.
longitudinal
keels;
fore-wing with 1 dark transversal
pterostigma;
distal
part of radial
(5:16);
fore tarsal
sp.
band beneath
vein much shorter than
segments in following proportions:
(type)
5,5:
proximal 4:
5:
the part
12: 26;
enlarged claw (Fig. 271) with one bristle on a proximal prominence; segment 5 of front tarsus (Fig. 271) with 2 rows of 25 lamellae; apex with a group of 8 I ameli ae; tibial
spurs 1,1,2.
MALE:
unknown
segment 2 of front
tarsus produced
into a hook;
LOCUS TYPICUS: Ammatti (India) TYPICAL MATERIAL: holotype F! in OT. HOSTS: unknown DISTRIBUTION: Oriental
region:
INDIA:
Ammatti
(S.
Coorg),
OT!
9. Anteon krombeini n. sp.
FEMALE: fully winged; length 4,52 mm; head black, with mandibles testaceous; antennae brown, with ventra I si de of segment 1 and segments 5-6 testaceous;
thorax,
propodeum and abdomen black;
mid tibiae and mid tarsi and fore tarsi specimen;
brown;
testaceous;
occasionally
28:
10:
18:
11:
10:
10:
antennal
10: 9:
9:
segments in following
14;
wi th
with trochanters
posterior legs missing in the only known
antennae distally thickened;
proportions:
legs testaceous,
legs black,
head shiny,
flat,
430
Mem. Amer.
Ent.
Inst.,
no.
37,1984
fully reticulate rugose; frons with a faintly visible median keel and two lateral keels around orbits; occipital carina complete; POL = 9; OL = 4; OOL
=
9;
OPL
=
by transversal smooth;
pronotal
irregular keels,
Fig.
13;
TL
=
10;
pronotum hairy,
shiny,
strongly sculptured
keels, except for a short area near the scutum which tubercles reaching tegulaej
scutum shiny,
except for anterior half of median area which
272 -
Chela of
is
sculptured by
Anteon krombeini n. sp. (type)
is smooth,
Anteoninae
431
punctate and without sculpture among punctures; reaching approximately 0,5 punctate, by
without sculpture among punctures;
longitudinal
transversal without
keel
keels;
(4:8);
dorsal
keels;
beneath the pterostigma; part
notaulices incomplete,
scutellum shiny, metanotum shiny,
propodeum reticulate rugose,
between
longitudinal
proximal
length of scutum;
fore
distal
part of radial
tarsal
sculptured
with a strong
and posterior surface;
fore-wing crossed by a
smooth,
posterior surface
dark
transversal
band
vein much shorter than
segments in following proportions:
10:
5:
8: 20: 48; enlarged claw (Fig. 272) with one bristle on a proximal prominence; segment 5 of fore tarsi (Fig. 272) with some rows of nearly 50 lamellae extending continuously MALE:
fully
winged;
antennae testaceous, partly
Fig.
black;
273 -
to the apex;
length 3,00-3,28 mm;
with segments 4-10 dark;
antennae hairy,
Male genitalia of
not distally
tibial
black;
spurs 1,1,2.
mandibles testaceous;
legs testaceous,
thickened;
antennal
with coxae segments
Anteon krombeini n. sp. from Induruwa
Jungle (Sri Lanka).
432
Mem.
Amer.
Ent.
15:
8:
in following proportions: fully
reticulate rugose;
carina complete;
POL
8:
punctures;
=
=
9; OL
without dark
8:
=
smooth,
8:
8:
1984
8:
transversal
band;
(4:12);
11;
head shiny,
punctate,
occipital
=
7; TL
4; scutum,
without sculpture among
reaching approximately 0,5 distal
part
without distal
inner pointed process;
hairy;
273)
(Fig.
tibial
length of
fore-wing hyaline,
part of radial
gonoforceps
LOCUS TYPICUS: Weddagala (Sri TYPICAL MATERIAL: holotype F!
=
8; OPL
scutellum and metanotum very
than proximal
F!
8:
8; OOL
notaulices incomplete,
scutum,
8:
37,
no.
frons with a median complete keel;
scutellum and metanotum shiny, scutum;
Inst.,
vein much shorter
long and narrow,
spurs 1,1,2.
Lanka) and 2 paratypes MM!
1 paratype
in WA;
in OL.
HOSTS:
unknown
DISTRIBUTION: Rat.
Dist.),
Jungle
Oriental
WA!
(Gi I imale,
NOTES:
region:
SRI
LANKA:
Weddagala
Kandy Reservoir Jungle (Kan. Rat.
this species
Dist.),
WA!
Hasalaka
Dist.),
(Kan.
is named in honor of Mr.
(Sinharaja Forest, OL!
Dist.),
Induruwa WA!
V. Krombein.
Karl
10. Anteon muiri n. sp.
FEMALE:
fully
winged;
length 4,43 mm;
black;
mandibles and antennae
testaceous; legs testaceous, with coxae and clubs of femora partly brown; antennae distally thickened; antennal segments in following proportions:
13: 6: 8: 6: 5: 6: 6: 6: 6: 9; head shiny, fully strongly reticulate rugose;
frons with a median keel
occipital
c:c;rina complete;
POL
=
and with 2 5;
OL
=
3;
lateral OOL
=
8;
keels around orbits; OPL
=
6;
TL
=
6;
pronotum shiny, strongly transversely striate, except for an area near posterior margin which is smooth and without sculpture; posterior surface
(12:19); pronotal tubercles reaching
of pronotum shorter than scutum tegulae;
scutum,
punctate,
scutellum and metanotum shiny,
without sculpture among punctures;
smooth,
weakly
notaul ices incomplete,
reaching approximately 0,65 length of scutum; propodeum reticulate rugose, with a strong transversa I keel between dorsa I and posterior surface;
posterior surface with 2 longitudinal
rugose as bands;
lateral
areas;
fore-wing hyal ine,
distal part of radial
keels;
median area as
without dark transversal
vein much shorter than proximal
part
(4:15); fore tarsal segments in following proportions: 6: 3: 4: 9: 25; enlarged claw (Fig. 274) with one bristle on a proximal prominence; segment 5 of front tarsus with basal part shorter than apical part (9:16); segment 5 of front tarsus (Fig. 274) with 2 rows of 22 lamellae; apex with a MALE:
group of 4
LOCUS TYP I CUS:
Mowong
TYPICAL MATERIAL: HOSTS:
lamellae;
tibial
spurs 1,1,2;
unknown
unknown
(Western Borneo,
holotype F!
in B.
I ndonesi a)
Anteoninae
DISTRIBUTION: NOTES:
Oriental
region:
INDONESIA:
433
Mowong
this species is named in honor of Mr.
Fig.
274 -
Chela of
Anteon muiri
F.
n.
(W Borneo),
Muir.
sp.
(type)
B!
434
Mem. Amer. Ent.
Fig. 275 -
Chela of
Ins!', no. 37, 1984
Anteon laotianum n. sp. (type)
Anteoninae
Anteon laotianum n. sp.
11.
FEMALE:
fully
brown;
length 4,37 mm;
winged;
and testaceous;
antennae testaceous,
antennae distally
proportions:
thickened;
black;
frons
with a
keels around orbits;
mandibles partly brown
with segments 6-10 brown; antennal
4,5: 5: 5: 5: 8: 8: 6: 7:
20:
reticulate rugose; lateral
435
median
occipital
segments
13;
head rugose,
longitudinal
keel
carina complete;
legs
in following fully
and with 2
furrow
around
occipital carina crossed by short keels; POL = 8; OL = 5; OOL = 7; OPL 10; temples invisible; pronotum shiny, smooth, weakly punctate, without sculpture among punctures,
and a
with
a
weak
weak posterior transversa I
tegulae;
scutum shiny,
punctures;
smooth,
anterior transversal
impress ion;
weakly punctate,
notaul ices incomplete,
without sculpture among
reaching approximately 0,25
scutum;
scutellum and metanotum dull,
rugose;
rugose,
with a
between dorsal
strong transversal
posterior surface with 2
surface;
by transversal
keels;
fore-wing
keel
longitudinal
with 2 dark
vein much shorter than proximal
pronotum shorter than scutum
keels;
proximal
rows of 27 MALE:
and posterior
median area crossed
transversal
bands,
distal
one on
part of
(4:9); posterior surface of segments in following
part
fore tarsal
9: 2,5: 3,5: 8: 22; enlarged claw (Fig. 275) with one bristle
proportions: on a
(17:21);
length of
propodeum reticulate
the basal cells and another beneath the pterostigma; radial
impression
pronota I tubercles reach i ng
prominence;
lamellae;
segment 5 of front
apex with a
group of 4
tarsus
(Fig.
lamellae;
275)
tibial
with 2
spurs 1,1,2.
unknown
LOCUS TYPICUS:
Ban Van Eue
TYPICAL MATERIAL: HOSTS:
(Laos)
holotype F!
in B.
unknown
DISTRIBUTION:
Oriental
region:
LAOS:
Ban Van Eue
(Vientiane),
B!
12. Anteon fyanense n, sp.
FEMALE:
fully
testaceous; distally 10:
9:
winged;
thickened;
9:
8:
median keel
length 3,68 mm;
antennae testaceous, 8:
8:
antennal 10;
mandibles and
segments in following
head shiny,
and with 2
black;
wi th segments 7-10 brown;
lateral
fully
proportions:
reticulate rugose;
keels around orbits;
legs
antennae 16:
6:
14:
frons with a
occipital
carina
complete; POL = 6; OL = 4; OOL = 9; OPL = 10; TL = 8; pronotum shiny, hairy, strongly reticulate rugose, with posterior surface shorter than scutum smooth,
(15:20);
pronotal
tubercles reaching
strongly punctate,
incomplete,
reaching
metanotum shiny, punctures;
approximately 0,65
smooth,
tegulae;
scutum shiny,
without sculpture among punctures; weakly
length of scutum;
punctate,
propodeum reticulate rugose,
notaulices
scutellum and
without sculpture among
with a strong transversal
keel
436
Mem. Amer. Ent.
Inst.,
no. 37, 1984
between dorsal and posterior surface; posterior surface with 2 longitudinal keels; median area rugose, but with a narrow, shiny,
Fig. 276 -
Chela of Anteon fyanense n. sp.
(type)
Anteoninae
437
central region which is smooth and without sculpture; fore-wing with dark transversal band beneath the pterostigma; distal part of radial much shorter than proximal proportions: on a
9:
4:
proximal
5:
13:
part
31;
(8:19);
fore
enlarged claw
segment 5 of front
prominence;
rows of 24 lamellae extending continuously 1,1,2. MALE:
tarsal (Fig.
segments
276)
tarsus
1 vein
in following
with one bristle
(Fig.
to the apex;
276)
with 2
tibial
spurs
unknown
LOCUS TYPiCUS:
Fyan
TYPICAL MATERIAL:
(Viet Nam)
holotype F!
HOSTS: unknown DISTRIBUTION: Oriental
region:
in B. VIET NAM:
Fyan,
B!
13. Anteon javanum n. sp.
FEMALE: brown,
fully
winged;
length 2,62 mm;
with segments 1-3 testaceous;
testaceous-dark;
scutell um,
and metasternum part I y antennae distally
thickened; 6:
8:
8:
7:
6:
6:
6:
abdomen brown;
antennae
scutum reddi shmesosternum
legs testaceous;
antenna I segments in following proportions:
9;
sculpture among punctures;
reddish-testaceous;
metanotum and propodeum black;
reddi sh-dark;
9:
5:
head
prothorax yellow;
head shiny,
strongly
punctate,
frons with a median keel
without
and with 2
lateral
weak keels near orbits; occipital carina complete; POL; 3; OL ; 3; OOL ; 4; OPL ; 3,5; TL ; 4,5; OPL slightly longer than breadth of posterior ocelli; pronotum shiny, rugose, with posterior surface flat, longer than scutum (13:11); pronotal tubercles reaching tegulae; scutum shiny, smooth,
weakly punctate,
almost complete,
without sculpture among punctures;
reaching
and metanotum shiny,
approximately 0,65
smooth,
without sculpture;
rugose,
with a strong transversal
surface;
posterior surface with 2
keel
notaul ices
length of scutum;
between dorsal
longitudinal
scutellum
propodeum reticulate
keels;
and posterior
median
area nearly
as rugose as lateral areas; fore-wing hyal ine, without dark transversal bands; distal part of radial vein much shorter than proximal part (3:11);
fore tarsal
segment 5 of front (5:14);
enlarged claw
prominence; lamellae; MALE:
segments in following proportions: tarsus with basal (Fig.
277)
fully
winged;
occipital 5;
OL ;
punctate,
carina complete; 3;
OOL ;
5;
(Fig.
lamellae;
length 1,87 mm;
fu II y testaceous; antennae brown, shiny, smooth, strongly punctate,
277) tibial
black;
2;
10:
19;
part
spurs 1,1,2. tegulae and
legs
with segments 1-2 testaceous; head without sculpture among punctures;
TL ;
3;
longitudinal
scutum shiny,
without scu I pture among punctures;
reaching approximately 0,5
2:
with 2 rows of 20
mandibles,
frons with a median
OPL ;
2:
witr, one bristle on a proximal
segment 5 of front tarsus
apex with a group of 4
6:
part shorter than apical
length of scutum;
keel;
smooth,
POL
weakly
notau I ices i ncomp I ete, scutellum and metanotum
438
Mem.
Fig.
shiny,
smooth,
transversal with 2
277 -
Ent.
between dorsal
longitudinal
Inst.,
no.
37,
Chela of Anteon javanum n.
without sculpture;
keel
wing hyal ine,
Amer.
keels;
sp.
(type)
propodeum reticulate rugose,
and posterior surface;
part
(5:10);
bands;
distal
gonoforceps
with a
posterior surface
median area as rugose as lateral
without dark transversal
much shorter than proximal
1984
areas;
part of radial (Fig.
278)
forevein
without
Anteoninae
Fig.
439
Male genitalia of Anteon javanum n.
278 -
sp.
from Tjibodas
(Java):
left half removed.
dorsal
proximal membranous process,
spurs 1,1,2. lOCUS TYPICUS:
Tjibodas
TYPICAL MATERIAL:
without distal
inner process;
tibial
(Java)
holotype F!
and 1 paratype M! in BM; 1 para type M! 1 paratype M! in OT. HOSTS: unknown DISTRIBUTION: Oriental region: INDONESIA: Tjibodas (Java), BM! lAOS: Ban Van Eue (Vientiane), B! INDIA: Kodaikanal (Pulney Hills, S. India), OT! in B;
14. Anteon munroei n. sp.
Chelogynus ephippiger (Da I man): Moczar 1979: 77.
FEMALE: black;
fully
winged;
length 2,20-2,40 mm;
antennae distally thickened;
proportions:
9:
5:
without sculpture, median keel;
5:
5:
5:
with small
=
POL
carina complete;
7:
4; Ol
=
5:
tegulae;
5;
OOl
testaceous,
8;
head flat,
notaulices incomplete,
with petiole
=
5;
smooth,
OPl
=
5;
shiny,
smooth,
smooth,
frons with a complete Tl
=
6;
without sculpture;
pronotum longer than scutum
scutellum and metanotum shiny, sculpture;
6:
punctures on vertex;
pronotum shiny,
tubercles reaching
6:
fully
antenna I segments in following
occipital pronotal
(17: 10);
weakly alutaceous,
scutum,
without
reaching approximately 0,80 length of
440
Mem.
Fig.
scutum;
Amer.
propodeum reticulate rugose,
37,
with a
1984
sp.
(type)
transversal
posterior surface wi th two
without dark transversal
shorter than proximal proportions:
5:
2:
3:
part 9:
a proximal prominence; segment 5 of front apex with
(3:9);
16;
Sq.
TYPICAL MATERIAL:
bands;
fore tarsal
enlarged claw
segment 2 of front
tarsus
(Fig.
279)
a group of 3 lamellae;
MALE: unknown LOCUS TYPICUS:
keel
Kabah
areas;
part of radial
forevein
segments in following
(Fig.
279)
with one bristle on
tarsus produced into a hook;
with 1 row of nearly 17 lamellae;
tibial
(Sarawak,
holotype F!
distal
between
long i tudi na I
spurs 1,1,2.
Malaysia)
and 1 paratype F!
in BU.
unknown
DISTRIBUTION: Oriental Sarawak), NOTES:
no.
median area of posterior surface as rugose as lateral
wing hyal ine,
HOSTS:
Inst.,
Chela of Anteon munroei n.
279 -
dorsa I and posterior surface; keels;
En!.
region:
MALAYSIA:
Sq.
Kabah
(Ng.
BU!
this species is named
in honor of Mr.
D.D.
Munroe.
Tada,
3d Div.,
Anteoninae
441
15. Anteon lankanum n. sp.
FEMALE: frons,
fully
winged;
clypeus,
length 2,40 mm;
head black,
mandibles and part of ventral
legs yellow;
antennae distally thickened;
proportions:
11: 7:
7:
7:
7:
7:
6:
7:
7:
antennal 9;
with anterior half of
side yellow;
antennae and
segments in following
head shiny,
smooth,
flat,
hairless, except for yellow anterior ha If of frons whic has dense long hairs; head punctate only along orbits and along occipital carina; frons and vertex without sculpture; occipital carina complete; POL = 3; Ol = 6; OOl
=
5;
=
OPl
posterior ocelli punctate and
4;
Tl
(4:2);
=
5;
OPl nearly twice as
pronotum shiny,
long as breadth of
rugose near anterior margin,
without sculpture among punctures on disc and near
Fig.
280 -
Chela of Anteon lankanum n.
sp.
(type)
442
Mem.
Amer.
En!.
Inst.,
no.
37,
1984
posterior margin; pronotal tubercles reaching tegulae; scutum shiny, smooth, wearkly punctate, without sculpture among punctures; notaulices incomplete,
reaching approximately 0,65 lenth of scutum;
metanotum smooth, punctures;
shiny,
weakly punctate,
pronotum s light I y
a strong transversal surface with two transversal
keel
longer than scutum
between dorsal
longitudinal
keels,
keels;
scutellum and
without sculpture among (14: 11);
propodeum wi th
and posterior surface;
posterior
median area sculptured by
as rugose as lateral
areas;
fore-wing hyaline,
without
dark transversal bands; distal part of radial vein much shorter than proximal part (4,5:10); fore tarsal segments in following proportions: 9:
3: 3: 9:
19;
prominence;
enlarged claw
(Fig.
280)
with one bristle on a proximal
segment 5 of front tarsus
lamellae extending continuously
(Fig.
280)
to the apex;
with 2 rows of nearly 14
tibial
spurs 1,1,2.
MALE unknown LOCUS TYPICUS:
Adams Peak Trail
TYPICAL MATERIAL:
holotype F!
HOSTS: unknown DISTRIBUTION: Oriental Maskeliya,
Kan.
Dis!.),
region:
(Sri
Lanka)
in WA. SRI
LANKA:
Adams Peak Trail
(4.5 mi.
W
WA!
16. Anteon nemorale n. sp.
FEMALE: MALE:
unknown
fully
testaceous;
winged;
distally thickened;
Fig.
281
-
length 1,68 mm;
antennae brown, antennal
black;
mandibles and
with segments 1-2 testaceous;
segments in following proportions:
Male genitalia of Anteon nemorale n. removed;
legs antennae not
B:
sp.
apex of gonoforceps).
(type)(A:
9:
4:
right
4:
half
Anteoninae
4:
5:
4:
4:
4:
5:
8;
head shiny,
443
granulated and fully reticulate rugose;
frons with a median keel and with 2 lateral keels near orbits directed towards antenna I sockets; occipital carina complete; POL; 6; OL ; 3; OOL ;
4;
OPL ;
2;
TL ;
3;
scutum shiny,
without sculpture among punctures; approximately 0,3 length of scutum; smooth,
without sculptur,e;
transversal without
keel
keels;
and posterior surface;
fore-wing hyal ine,
(1,5:5);
gonofoceps (Fig.
inner pointed process;
LOCUS TYPICUS:
281)
Forest Camp
TYPICAL MATERIAL:
Fig.
reaching
scutellum and metanotum shiny, with a strong posterior surface
without dark transversal
di sta I part of rad i a I vei n much shorter than prox i ma I part
a distal
HOSTS:
weakly punctate,
incomplete,
propodeum reticulate rugose,
between dorsal
longitudinal
bands;
smooth,
notaulices,
without proximal tibial
(Sabah,
holotype M!
membranous process,
with
spurs 1,1,2. Malaysia)
in B.
unknown
282 -
Male genitalia of
Anteon silvestre removed.
n.
sp.
(type):
right half
':'44
Mem.
Amer.
DISTRIBUTION: Oriental Kalabakan, Sabah), B!
En!.
region:
Inst.,
MALAYSIA:
no.
37,
1984
Forest Camp
(19 Km N of
17. Anteon silvestre n. sp.
FEMALE:
unknown
MALE: fully winged; length 1,62 mm; black; mandibles testaceous; antennae brown; legs testaceous, with coxae and fore and mid clubs of femora brown;
antennae not distally thickened;
granulated and fully with 2
lateral
occipital
reticulate rugose;
frons
keels near orbits directed POL = 6;
carina complete;
scutum dull,
with a strong transversal
keel
posterior surface without
OL = 3;
OOL = 4;
keels;
OPL = 1;
TL = 2;
notaulices invisible;
and posterior surface;
fore-wing hyaline,
distal part of radial
gonoforceps
and
sockets;
propodeum reticulate rugose,
between dorsal
longitudinal
bands;
(2:6);
part
in
with a median keel
granulated and fully reticulate rugose;
dark transversal
segments
towards antennal
scutellum and metanotum reticulate rugose;
proximal
antennal
6: 4: 3: 4: 4: 4: 4: 4: 4: 6; head dull,
following proportions:
without
vein much shorter than
282) without proximal membranous
(Fig.
process, without distal inner process; tibial spurs 1,1,2. LOCUS TYPICUS: Tenompok (Sabah, Malaysia) TYPICAL MATERIAL: HOSTS:
holotype M!
in B.
unknown
DISTRIBUTION:
Oriental
region:
MALAYSIA:
Tenompok
(Sabah),
B!
18. Anteon expolitum n. sp.
FEMALE:
unknown
MALE: fully winged; antennae testaceous,
length 2,28-2,81 mm; black; mandibles testaceous; with segments 7-10 brown; legs testaceous, with hind
coxae proximally brown; segments shiny,
reticulate rugose,
ocellus which keels, long,
antennae not distally
in following proportions:
10:
except for a
6:
8:
small
7:
complete;
incomplete median keel,
occipital carina complete;
7:
area
is smooth and without sculpture;
with a short,
thickened; 7:
7:
antennal 7:
7:
10;
head
in front of anterior
frons without
lateral
occasionally median
POL = 7;
OL = 4;
keel
OOL = 7;
OPL
= 5;
TL = 5; scutum shiny, smooth, weakly punctate, without sculpture among punctures; notaulices incomplete, reaching 0,5 length of scutum;
scutellum and metanotum shiny, sculpture among punctures; transversal with 2
keel
between
longitudinal
keels;
smooth,
weakly punctate,
propodeum reticulate rugose,
dorsal
and posterior surface;
median area shiny,
except for some keels near margins;
smooth,
fore-wing hyaline,
without with a strong
posterior surface without sculpture, without dark
Anteoninae
Male genitalia of Anteon expolitum n.
283 -
Fig.
445
sp.
(type):
right half
removed.
transversal
vein much shorter than proximal 283) without proximal membranous process and without distal inner process; tibial spurs 1,1,2. LOCUS TYPICUS: Ban Van Eue (Laos) TYPICAL MATERIAL: holotype M! in B; paratype M! in CO. HOSTS: unknown DISTRIBUTION: Oriental region: LAOS: Ban Van Eue (Vientiane), B! part
bands;
(5:12);
distal
part of radial
gonoforceps (Fig.
PHILIPPINES:
Pinigisan
(Mantalingajan,
Palawan),
CO!
19. An teon peterseni n. sp.
FEMALE: MALE:
unknown
fully
testaceous; hairy; 4,5:
winged;
antennal
4,5:
6;
length 1,84 mm;
antennae brown,
with mandibles and
segments in following proportions:
head smooth,
among punctures;
black,
with segmen ts 1-2 testaceous;
shiny,
8:
weakly punctate,
frons with a complete
longitudinal
4:
legs
antennae 5:
4:
4:
4,5:
without sculpture median keel;
5:
446
Mem.
Amer.
Ent.
Inst.,
no.
37, 1984
occipital carina complete; POL = 5; OL = 3; OOL = 7; OPL = 2,5; TL = 3; scutum shiny, smooth, very weakly punctate, without sculpture among punctures; shiny,
weakly punctate,
keels on posterior
median area of posterior surface partly smooth and shiny,
rugose; radial
scutellum
without sculpture among punctures;
with 2 longitudinal
propodeum reticulate rugose, surface;
reaching 0,5 length of scutum;
notaulices incomplete,
smooth,
fore-wing hyal ine,
without dark transversal
vein shorter than proximal
part (4:8);
damaged in the only known specimen;
bands;
distal
partly part of
gonoforceps and penis
gonoforceps without distal
inner
process; tibial spurs 1,1,2. LOCUS TYPICUS: Pinigisan (Philippines) TYPICAL MATERIAL: HOSTS:
holotype M!
in CO.
unknown
DISTRIBUTION: Oriental Palawan), NOTES:
region:
PHILIPPINES:
Pinigisan
(Mantalingajan,
CO!
this species is named
in honor of Mr.
Borge Petersen.
20. Anteon bauense n. sp.
FEMALE:
unknown
MALE: fully winged; length 1,87-2,37 mm; black; mandibles, antennae and legs fully testaceous; sometimes antennal segments 3-10 darkened; antennae not distally thickened; antennal segments in following proportions: punctate, occipital lateral
=
3; TL
10: 5:
7:
7:
7:
6:
6:
6:
6,5:
10;
head shiny,
without sculpture among punctures, carina;
keels;
=
4;
strongly
irregular keels near
frons with a short incomplete median keel,
occipital
carina complete;
scutum shiny,
among punctures;
with
smooth,
weakly
notaul ices incomplete,
= 6;
POL
OL
punctate,
= 3;
without
OOL
= 5;
OPL
without sculpture
reaching approximately 0,65
I ength of scutum; scutell um and metanotum sh i ny, smooth, weak I y punctate, without sculpture among punctures; propodeum reticulate rugose, with a strong
transversal
keel
between
posterior surface with 2 longitudinal
dorsal
keels;
and posterior surface;
median area shiny,
with some keels near margins; lateral areas shiny, fore-wing hyal ine, without dark transversal bands; vein much shorter than proximal as penis, (Fig.
with proximal
part
(3: 11); gonoforceps nearly as long
membranous process,
without distal
inner process
284); tibial spurs 1,1,2.
LOCUS TYPICUS: Bidi (Sarawak, Malaysia) TYPICAL MATERIAL: holotype M! in B; 1 paratype M! M!
smooth,
with a few keels; distal part of radial
in WA;
1 paratype
in OL.
HOSTS: unknown DISTRIBUTION: Oriental B! OL!
Forest Camp
Udawattakele Sanct.
region:
MALAYSIA:
(N of Kalabakan, (Kan.
Dis!.),
WA!
Bidi
Sabah),
(Bau District, B!
SRI
LANKA:
Sarawak),
Anteoninae
Fig.
447
Male genitalia of Anteon bauense n.
284 -
(Sabah):
sp.
from Forest Camp
left half removed.
21. Anteon debile n. sp.
FEMALE: MALE:
unknown
fully
legs fully
in following smooth,
winged;
weak,
weakly punctate,
9:
4:
6:
5:
5:
5:
mandibles, 5:
5:
5:
7;
antennae and antennal
lateral
segments
head shiny,
without sculpture among punct<.1res;
frons without median and
keels;
occipital
POL = 4;
OL
= 4;
OPL = 2; TL = 3; scutum, scutellum and metanotum shiny, weakly punctate, without sculpture among punctures; notaulices
incomplete,
reaching approximately 0,65 length of scutum;
propodeum reticulate rugose, dorsal
black;
antennae not distally thickened;
proportions:
carina complete; 2; OOL smooth,
length 1,56 mm;
testaceous;
with a strong
and posterior surface;
median area shiny,
smooth,
transversal
keel
between
posterior surface with 2 longitudinal
with a few keels near margins;
lateral
keels; areas
448
Mem.
En!.
Inst.,
no. 37,
Male genital ia of Anteon debile n.
285 -
Fig.
Amer.
1984
sp.
(type):
rigth half
removed.
shiny,
with a
few keels;
bands;
distal
part of radial
fore-wing hyaline,
without dark transversal
part (3:7); 285) shorter than penis, without proximal membranous process and without distal inner process; tibial spurs 1,1,2. LOCUS TYPICUS: Ban Van Eue (Laos) TYPICAL MATERIAL: holotype M! in B. HOSTS: unknown DISTRIBUTION: Oriental region: LAOS: Ban Van Eue (Vientiane), B! gonoforceps
vein much shorter than proximal
(Fig.
22. Anteon philippinum n. sp.
FEMALE:
fully
winged;
length 3,00 mm;
black;
mandibles testaceous;
antennae and legs fully testaceous; abdomen brown; antennae distally thickened; antennal segments in following proportions: 15: 7: 8: 7: 8: 7:
7:
7:
10;
median keel; complete;
head shiny,
two keels are present around orbits;
POL = 8;
OL = 4;
fully reticulate rugose, which
OOL = 6;
OPL = 5;
7:
frons with a complete occipital
TL = 5;
carina
pronotum shiny,
except for a narrow area near posterior margin
is smooth and without sculpture;
tegulae; shiny,
fully reticulate rugose;
posterior tubercles reaching
posterior surface of pronotum shorter than scutum
fully sculptured by weak
longitudinal
keels;
(7:20);
scutum
notaulices incomplete,
Anteoninae
Fig.
286 -
Chela of
449
Anteon philippinum
n.
sp.
(type)
reaching approximately 0,5 length of scutum; scutellum shiny, smooth, without sculpture; metanotum reticulate rugose; propodeum reticulate rugose, wi th a strong transversa I keel between dorsa I and posterior surface; posterior surface with two longitudinal keels; median area as rugose and dull transversal part
(4:11);
as lateral
bands;
distal
fore tarsal
areas;
fore-wing hyaline,
part of radial
vein
without dark
much shorter than proximal
segments in following proportions 10:
4:
4:
5:
450
Mem.
Amer.
Ent.
Inst.,
37,
no.
1984
17; enlarged claw (Fig. 286) with one bristle on a proximal prominence; segment 5 of front tarsus (Fig. 286) with two rows of nearly 25 lamellae; group of nearly 4
apex with a MALE:
lamellae;
tibial
spurs 1,1,2.
unknown
LOCUS TYPICUS:
Camp Lookout
TYPICAL MATERIAL: HOSTS:
(Philippines)
holotype F!
in AM.
unknown
DISTRIBUTION: Oriental Negros lsi.), AM!
region:
PHILIPPINES:
Camp Lookout
(Dumaguete,
ANTEON
NEARCTIC SPECIES OF
Sixteen species are known.
KEY TO THE SPECIES
FEMALES
5 of front tarsus with basal part longer than apical 291); segment 4 of front tarsus at most 0,5 as long front tarsal segments 3 or 4 produced into a hook ••••••• 2
Segment part as
1;
(Fig.
Segment
5 of
than apical 0,66 as
long as
produced 2
front part
tarsus wi th basal
303);
(Fig. segment
1;
usually
part
as
4 of
front
front
long as
or
shorter
tarsus at
tarsal
segment
least
2
into a hook ••••••••••••••.••••.•••••••••••••••••••••• 4
Posterior
surface of
propodeum without
• •.•••.••...••••••.•..••.•••• Posterior
3
segment
surface of
Notaulices
reaching
longitudinal
keels ••••••
1. popenoei (Ashmead)
propodeum with approximately
. . . . . . . . . . , . . • . . . . . . . . . . . . . . • 2.
two
longitudinal
keels •••• 3
0,5
length of scutum ••••••••• arizonense R.C.L. Perk i ns
reaching at most 0,3 length of scutum ••••••••••.•••• · . . • . • . . . . . . . . . • . • • • • . . . . . . . . 3. canadense (Ashmead)
Notaul ices
4 5
Posterior
surface of
propodeum without
Posterior
surface of
propodeum with
Prothorax black •••••••••••••• Prothorax
6
reddish •••..•••••••
Fore and hind wings
fully
darkened, 7
Antennal
hind wings hyal ine; but
longitudinal
keels ••••• 5 keel s •••• 6
iunestum (R.C.L. Perkins) wasbaueri n. sp.
dark •••••••••••••••••••••••••••••••••
. . . . . . . . . . . . . . . . . . . . . . . . . . . . .. Fore and
5. 6.
longitudinal
two
4. at
nebulosum n. sp. most
in
that
case hind-wing
segment
2 as
long as
• • • • • • • • • • • • • • • • • • • • • • • . • • • ••
segment 7.
fore-wing
51 ight Iy
always hyal ine ••••••••••• 7 3 ••••••••.••••••.••••••••
minimum (F en t on)
451
Anteoninae
Antennal
8
Segment
9
segment
4 of 4 of
Segment
2
shorter
than
segment
front
tarsus nearly
front
tarsus nearly as
3 ••••••••••••••••••••• 8
twice as
long as
long as or
segment
sl ight Iy
1. .. 9
longer
than
segment
1 •.•••.....••..•..••.•....••••..•••.•••.•••••••. 10
Head
smooth,
weakly
thorax mostly Head
rugose,
around
punctate,
testaceous ••••• punctate,
frons and
wi th
fully granulated,
Head
shiny,
punctate,
11 Pronotum fully punctures;
without
smooth,
head and
thorax black ••••••••••••••
rugosiceps K i ef f er
9.
punctate and wi th
or wi thout
10. puncticeps Ashmaead sculpture among punctures.aSW •. l 1
weakly punctate,
wi thout
sculpture among
thorax black wi th prothorax yellow •••••
• • • • • • • • • • • • • • • • • • • • • • • • • • •• Pronotum rugose,
head and
irregular keel s mostly
head and
irregular keels ••••••••••••.
irregular keels;
masoni n. sp.
8.
numerus
on vert ex;
• • • • . • • • • • • • • • • • • • • • • • • • • • • • ••
10 Head dull,
without
xanthothorax (B rad I ey )
1 1.
irregular keels on anterior half and on posterior half; sometimes more strongly rugose; thorax differently colored ••••••••••••••••••••••••••••
punctate head and
wi th
12. osborni (Fent on)
A. durangoi n. sp., hirtifrons Whittaker, ciudadi n. menkei n. sp. are unknown.
The females of sp.
and
MALES
2
Posterior
surface of
propodeum without
Posterior
surface of
propodeum wi th
Gonoforceps with distal
longitudinal
two
inner pointed
longi tudinal
process
(Fig.
keels ••••• 2 keels •••.
4
308 B) •••••
1 3. durangoi n. sp. inner pointed process
. . . • . . . . . . . . . . . . . . . . . . . . . . ..
(Figs. 288 297) •••••.•••••••••••••••••••.••••••••••••.••••••••••••• 3
Gonoforceps without
A, 3
B,
Propodeumwith straight
4
short
transversal
keel
line ••••••••••••••••
Propodeum with a
distal
transversal
Gonoforceps without
distal
forming
a
short
funestum (R.C.L. Perkins) 6. wasbaueri n. sp.
keel
line •••••••••••••••••
centrally
5.
centrally
forming
an
angle,
not
1. popenoei (Ashmead)
inner pointed process
(Figs.
302,
308 A) ••••••••••••••••••••••••.•••••.•••••••.••••••••••••••••• 5 Gonoforceps with distal inner pointed process (Fig. 293
A,
B, C) •••••••.••••••••••••••••.•••••••••.•••.••••••••••••••• 6
have been unable to find characteristics for separating males of the two species
452
5
Mem. Amer.
Head weakly punctate,
Ent.
smooth,
punctures ••••••••••••••••••• Head
6
7
8
9
10
11
12
13
smoot h ••••••••••••• "
The males of A.
dull,
no.
shiny, 14.
37,
1984
wi thout
ciudadi n.
sculpture among
sp.
or strongly granulated, 9. rugosiceps K i ef f er Gonoforceps much short er than pen is (F i g. 293 A, B) ••••••••••• 7 Gonoforceps as long as or longer than penis (Figs. 293 C, 309). • ., ••••••••••••••••••••••••••••••••••••••••••••• " •••••••••••• 8 Gonoforceps with small distal inner process (Fig. 293 A) ••••••• . • . . . • • . . . . . . . . . . . . . . . . . . . . . .. 3. canadense (Ashmead) Gonoforceps wi th large distal inner process (Fig. 293 B) ••••••• 16. hirtifrons Whi ttaker Posterior surface of propodeum with indistinct longitudinal keel s •••••••••••••••••••••••• 8. masoni n. sp. Posterior surface of propodeum with distinct longitudinal keels • •••••.••••••• " ••••••••••••••••••..••••.•• '" ••.••••.•••••••• 9 Head sparsely punctate, wi thout sculpture among punctures •••• 10 Head densely punctate, weakly or strongly granulated among punctures, wi th strong or weak areolae or irregular keels •••• 12 Face with anterior half testaceous; posterior half black ••••••• • • • • • • • • • • • • • • • • • • • • • • • • • • •• 15. menkei n. sp. Face fully black ••••••••••••••••••••••••••••••••••••••••••••• 11 Legs testaceous, at most with hind coxae and hind femora partly brown ••••••••••••••••••••••• 12. osborni (Fenton) Legs brown, with tarsi light and sometimes also tibiae testaceous •••••••••••••••••• 11. xanthothorax (Bradley) Scutum dull, rugose, fully strongly punctate, wi th some irregular keels near margins ••••••••••••••••••••••••••••••••••• • • • • • • • • • • • • • • • • • • • • • • • • • • •• 10. puncticeps Ashmead Scutum shiny, punctate, without sculpture among punctures, at most wi th anterior half rugose and dul I •••••••••••••••••••••• 13 Distal inner pointed process of gonoforceps extended medially and with apical margin excavated (Fig. 290) •••••••••••••••••••• . . • • . . . • • . • • . . . . . . . . . . . . . . . . . 2. arizonense R.C. L. Perkins Distal inner pointed process of gonoforceps extended apically and with apical margin convex or straight (Fig. 293C) •••••••••• • • • • • • • • • • • • • • • • • • • • • • • • • • •• 10. puncticeps Ashmead not
reticulate rugose,
Inst.,
weakly
••••
nebulosum n. sp. and minimum (Fenton) are
unknown.
1. Anteon popenoei (Ashmead)
Dryin us popenoei Ashmead 1888: i. Anteon unifasciatus Ashmead 1893: 95 (n. syn.); locus typicus
Anteoninae
Biscayne Bay
(Florida,
U.
S.
453
A.).
Anteon minutus Ashmead 1893: 97 (n. syn.); locus typicus Washington
Anteon Anteon Anteon Anteon
FEMALE:
fully
antennae fully legs fully
(D.
C.,
U.
winged;
3:
3:
testaceous;
4,5;
or areolae;
A.).
length 1,5-3,0 mm;
black;
mandibles testaceous;
testaceous or with segments 5-10 brown; antennae distally thickened;
following proportions: 9: 4: 3:
s.
popenoei (Ashmead): Ashmead 1893: 98. minutus Ashmead: Krombein 1979: 1241. popenoei (Ashmead): Krombein 1979: 1241. unifasciatus Ashmead: Krombein 1979: 1241.
head dull,
5:
2:
3:
granulated,
occipital carina complete;
keel
and with 2 strong or weak
Fig.
287 -
Chela of
3:
3:
3:
4:
tegulae testaceous;
antennal 6;
or 8:
with weak or strong frons with a median
lateral
keels;
POL = 5;
segments in
3:
3:
3:
3:
3:
irregular keels longitudinal
OL = 3;
OOL = 4;
Anteon popenoei (Ashmead) from Biscayne Bay (Florida)
454
OPL
Mem.
=
2;
rai sed
=
TL
into a
than scutum
no.
37,
1984
with posterior surface convex and
posterior surface of pronotum shorter
pronota I tubercles reach i ng tegu I ae;
notaul ices
incomplete,
reaching
scutellum and metanotum shiny,
posterior surface; hya line,
Inst.,
pronotum rugose,
propodeum reticulate rugose,
beneath
Ent.
transversa I cari na; (4: 11);
granulated; of scutum;
4;
Amer.
with a
smooth,
transversal
posterior surface without distal
part of radial
length
without sculpture;
keel
between dorsal
longitudinal
wi thout dark transversa I bands or with the pterostigma;
scutum du II,
approximately 0,25-0,30
keels;
and
fore-wing
1 dark transversal
band
vein shorter than proximal
part (2:4 or 2:5); fore tarsal segments in following proportions: 5: 2: 1,5: 2: 7; enlarged claw (Fig. 287) with one bristle on a proximal prominence; segment 5 of front
tarsus
longer than apical
part,
to the apex;
spurs 1,1,2.
MALE:
fully
testaceOU5 j
segment with
tibial winged;
287)
with basal
1 row of 7-8
length 1,31-2,00 mm; tegu I ae testaceous;
and fore tibiae
testaceous;
part sl ightly or much
lamellae extending continuously
black or brown;
antennae brown or brown-testaceous j
1 testaceous;
tarsi
(Fig. with
mandibles
somet i mes brown
legs fu II y
wi th
testaceous or brown
antennae not distally
thickened;
B
Fig.
288 -
Anteon funestum osborni (Fenton) (C):
Male genitalia of (Ashmead) (B),
(R.C.L.
Perkins)(A),
right half removed.
popenoei
Anteoninae
antennal 5,5;
segments in following proportions:
head dull,
areolae;
granulated,
8:
4,5:
complete;
kee!
and with or without
POL = 5;
Ol = 3;
4:
3:
3:
3:
3:
3,5:
4:
with weak or strong irregular keels or
sometimes strongly reticulate rugose;
longitudinal smooth,
455
frons with a median
lateral
OOl = 4;
keels;
OPl = 3;
occipital
Tl = 4;
carina
scutum shiny,
weakly punctate and without sculpture among punctures or weakly
granulated;
rarely scutum strongly granulated;
reachi ng approxima tel yO, 3 I ength of scutum; shiny,
smooth,
without sculpture or weakly
notaulices incomplete,
scute II um and metanotum
granulated;
propodeum
reticulate rugose, with a transversal keel between dorsal and posterior surface; posterior surface without longitudinal keels; transversal keel of propodeum centrally forming an angle; transversal
bands;
part
gonoforceps (Fig.
(2:5);
distal
tibial spurs 1,1,2. lOCUS TYPICUS: Marlatt
fore-wing hyaline,
part of radial 288 B)
(Kansas,
without distal
U.
S.
without dark
vein much shorter than proximal inner pointed process;
A.)
popenoei in WA; holotype F! of A. minutum in WA; holotype F! of A. unifasciatus in WA. TYPICAL MATERIAL:
HOSTS:
holotype M!
of D.
unknown
DISTRIBUTION:
Nearctic region:
CANADA:
Scandia
(Alta.),
OT!
Lethbridge
(Alta.), OT! Ottawa (Ont.), OT! Mer Bleue (Ottawa, Ont.), aT! Ol! Britannia (Ont.), aT! OL! Cayuga (Ont.), OT! Beech Grove (Que.), OT!
u.
Ol! Kouchibouguac N. P. (N.B.), OT! Plas~r Rock (N.B.), OT! S. A.: Mc Kittrick Canyon (Culberson Co., Tex.), TW! Nebraska, WA! Marlatt (Kans.), (Ga.),
WA!
OT!
Wattacoo OT!
Ol!
Rapides Parish
Rabun Bald
(Pickens Co., S.C.), Washington
Research Station
(D.
(la.),
TW!
(Rabun Co., C.),
(Huntington,
TW! WA!
Biscayne Bay
Ga.),
OT!
Greenvi lie Farmingdale
long
Island,
N.
(Fla.),
Cleveland (S.C.),
TW!
(N.Y.), Y,),
(S.
Ol!
Forsyth OL!
Di Ilwyn
TW!
AM!
WA! C.),
TW!
(Va.),
Kalbfleisch
Ol!
2. Anteon arizonense R.C.l. Perkins
Anteon arizonensis R.C.l. Perkins 1907: 51. Anteon arizonensis R.C.l. Perkins: Krombein 1979: 1240.
FEMALE: yellow,
fully
winged;
length 2,56 mm;
with teeth brown;
black;
abdomen brown;
antennae yellow;
legs yellow,
mandibles
with hind coxae
and hind clubs of femora brown; body shiny, hairy on sides of thorax and on front part of vertex; head rugose, strongly punctate; pronotum short, more than
twice as
long as broad
(5:1),
as sculptured as head;
scutum
smooth, without punctures, without sculpture; notaulices incomplete, reaching approximately 0,5 length of scutum; scutellum and metanotum smooth;
propodeum reticulate rugose,
between dorsal
and posterior surface;
with an
irregular transversal
keel
posterior surface with 2 straight
456
Mem. Amer.
Fig.
289 -
Ent.
Inst.,
no.
37,
Chela of Anteon arizonense R.C.L.
1984
Perkins from Vermilion Bay
(Ontario) •
longitudinal hyaline,
keels;
median area as rugose as lateral
without dark transversal
much shorter than proximal; segment 4
(35: 10);
enlarged claw
segment 5 of front
distal
part;
tibial
MALE:
fully
winged;
darkened; distal
(Fig.
289)
areas;
fore-wing
part of radial
tarsus much
vein
longer than
with one bristle on a proximal
tarsus with basal
part much
longer than
spurs 1,1,2. length 2,10-2,37 mm; black; mandibles testaceous; with segments 2-3 and apical region of segment 1
testaceous,
legs testaceous,
with hind coxae,
part of hind tibiae brown;
antennae not distally thickened; proportions:
with distal
segment 1 of front
prominence;
antennae
bands,
10: 5:
strongly punctate,
6:
6,5:
7:
antennal
6,5:
segments in following
6,5: 7:
mostly on frons,
to areolae; occipital
stalk of hind femora and
sometimes thorax and propodeum reddish; 6,5:
9;
head dull,
granulated,
where the wide punctures are similar
carina complete;
frons with a median keel,
hairy,
without lateral keels; POL = 7; Ol = 3,5; OOl = 5; OPl = 3; Tl = 3; scutum, scutellum and metanotum shiny, smooth, without sculpture or very weakly punctate, without sculpture among punctures; notaulices incomplete, reaching approximately 0,5 length of scutum; propodeum reticulate rugose, with a strong
transversal
keel
between dorsal
and posterior surface;
posterior surface with 2 longitudinal
keels;
as
without dark transversal
lateral
distal
areas;
fore-wing hyal ine,
part of radial
gonoforceps (Fig. with a distal
290)
median area nearly as rugose
vein much shorter than without dorsal
proximal
proximal
part
membranous process and
inner pointed process extended medially and with apical
margin excavated; tibial spurs 1,1,2. LOCUS TYPICUS: Nogales (Arizona, U. S.
A.)
TYPICAL MATERIAL: lectotype F! and 1 para lectotype M! Olmi 1982) of A. arizonense in B. HOSTS:
in U.
S.
A.
(designated by
Cicadellids.
DISTRIBUTION:
Nearctic region:
CANADA:
OT!
(B.C.), OT!
Cowichan Bay
Crowsnest
bands;
(4,5:7);
Ol!
Hixon
(B.C.), OT! (Vancouver
Terrace
Is.,
(B.C.),
B.C.),
OT!
457
Anteoninae
Sandspit (Moresby lsi., Queen Charlotte Islands, B.C.), AM! Twin Buttle (Alta.), OT! Waterton (Alta.), OT! Ol! Cooking lake (Alta.), OT! Elkwater (Alta.), OT! Jumping Pd. Cr. (Calgary, Alta.), OT! Maple Creek (Sask.), OT! Ol! Ninette (Man.), OT! Stittsville (On!.), OT! Vermilion Bay (On!.), OT! OLl Marmora (On!.), OT! Mistassini (Que.), OT! Wood R.-Wood l . (Baie Comeau, Que.), OT! U. S. A.: Brinnon (Jeff Co., Wash.), TE! Idaho City (Idaho), OL! TW! Crescent City (Calif.), TW! Tuscarora (Nev.), Ol! TW! OLl
Nogales (Ariz.),
B!
Cleveland
(S.
C.),
OLl
TW!
lockport
(N.Y.),
TW!
3. Anteon canadense (Ashmead)
/
1/
~
"-
I
I
"
,.I
I
1 / ! l-
Fig.
290 -
i
Male genitalia of Anteon arizonense R.C.l. lockport
(New York):
right half removed.
Perkins from
458
Mem. Amer. En!.
Inst.,
no. 37,
1984
basal part
Fig. 291 - Chela of Anteon canadense (Ashmead) (type)
Chelogynus canadensis Ashmead 1893: 93. Anteon politus Ashmead 1893: 96 (n. syn.); locus typicus Toronto (Ontario, Canada).
Anteoninae
459
Anteon pallidicornis Ashmead 1893: 97 (n. syn.); locus typicus Utah Lake
(Utah,
U.
S. A.).
Chelogynus canadensis Ashmead: Fenton 1918b: 272. Anteon canadensis (Ashmead): Muesebeck and Walkley 1951: 1040. Anteon (Anteon) canadensis (Fenton): Krombein 1979: 1240. Anteon (Anteon) pallidicornis Ashmead: Krombein 1979: 1241. Anteon (Anteon) politus Ashmead: Krombein 1979: 1241.
FEMALE:
fully
winged;
antennae fully
length 2,25-2,75 mm;
brown or testaceous,
black;
mandibles testaceous;
sometimes brown
with segment
1
testaceous; legs fully testaceous, with hind coxae basally black; antennal segments in following proportions: 10: 4,5: 5: 5: 5: 4: 4: 4: 4: 6; head shiny,
granulated or not granulated,
centra I area of the frons smooth frons,
region behind ocelli,
and granulated; ocell i; =
4;
occipital
fronta I
POL = 7;
pronotum du II,
pronotal
weakly punctate,
scutum;
sculpture among punctures;
keel
Fig.
292 -
also among
OPL = 4;
smooth on I y
tegulae;
TL
near
scutum shiny,
scutum sometimes with areolae near
keels;
smooth,
propodeum reticulate
between dorsal
Chela of
rugose,
OOL = 5;
reaching approximately 0,3-0,5
scutellum and metanotum shiny,
with 2 longitudinal
OL = 4;
tubercles reaching
notaulices incomplete,
transversal
less wide
anterior ha If of
strongly punctate only near anterior margin,
without sculpture among punctures; margins;
more
with numerous irregular keels,
carina complete;
posterior margin; smooth
with a
punctate;
sometimes also sides of frons weakly rugose
sometimes head
line present;
usually
and weak I y
length of
weakly punctate, rugose,
and posterior surface;
median area as rugose as,
with
an
without irregular
posterior surface or
less rugose
than
Anteon canadense (Ashmead) from Toronto (Ontario):
variety
with
lamellae on tarsal
segment 5.
460
Mem.
Fig.
293 -
Amer.
Male genital ia of Whittaker
lateral
areas;
part of radial segments
fore-wing
En!.
Ins!.,
no.
37,
Anteon canadense (Ashmead) (A), hirtifrons puncticeps Ashmead (C).
(B),
hyal ine,
without dark transversal
vein much shorter than proximal
in following proportions:
tarsus produced
into a hook;
1984
6:
1,5:
1,5:
enlarged claw
part 2:
(Figs.
9;
bands;
(3:10);
distal
fore tarsal
segment 3 of front
291,
292)
with one
bristle on a proximal prominence; segment 5 of front tarsus (Figs. 291, 292) without medial lamellae and with bristles only, or with 3 lamellae; apex with a than distal MALE:
group of 2-5 part;
fully
tibial
winged;
lamellae;
segment 5 with basal
part much
longer
spurs 1,1,2.
length 2,62 mm;
black;
mandibles testaceous;
antennae brown; legs testaceous, with hind coxae basally black; antennae not distally thickened; antenna I segments in following proportions: 10: 5: 7: 8: 8: 8: 8: 8: 8: 11; head shiny, strongly punctate, without sculpture among punctures, with numerous irregular keels mostly on vertex and among ocell i;
frons
with
an
incomplete median
keel
visible only
in front of
anterior ocellus; occipital carina complete; POL = 5; Ol = 3; OOl = 7; = 4; Tl = 5; scutum shiny, weakly punctate, without sculpture among punctures,
strongly
OPl
punctate and almost rugose only near anterior margin;
notaul ices incomplete, reaching approximately 0,5 length of scutum; scutellum and metanotum shiny, smooth, weakly punctate, without sculpture among punctures; keel
between
longitudinal
propodeum ret icu I ate rugose,
dorsal keels;
and posterior surface;
with a
strong transversa I
posterior surface with 2
median area as rugose as
lateral
areas;
fore-wing
Anteoninae
461
slightly darkened; hind-wing hyaline; distal part of radial shorter than proximal part (5:11); gonoforceps (Fig. 293 A) pointed process; gonofoceps much shorter than penis; LOCUS TYPICUS: Ottawa (Canada)
vein much with a distal
tibial
spurs 1,1,2.
TYPICAL MATERIAL: holotype F! of Ch. canadensis in WA; holotype F! of A. pallidicornis in WA; holotype F! of A. politus in WA. HOSTS:
unknown
DISTRIBUTION:
Nearctic region:
Terrace (B.C.), OT! Crowsnest
(B.C.),
Burnaby
OT!
C.),
Mac Gillivray
Zymoetz
(Terrace,
B.C.),
(Alta.),
OT!
View
Mtn.
CANADA:
(B.
OT!
Gagnon Road
OT!
Shames
(Game Reserve,
Upper Hay River Post
(Alta.),
OT!
Ninette
(Terrace,
(Terrace,
B.C.),
B.C.),
Chilliwack, (Alta.),
(Man.),
OT!
B.C.),
CA!
OL!
OT!
OT! OT!
Morley
Ottawa
(Ont.), WA! Innisville (Ont.), OT! Toronto (Ont.), WA! Stittsville (Ont.), OT! Merivale (Ont.), OT! Nominingue (Que.), OT! OL! Laniel (Que.), OT! U. S.
A.:
Lowman
(Calif.), TE!
OL!
OT!
TW!
Idaho City
(Calif.),
OT!
Washoe Lake (Washoe Co.,
Estes Park
(Colo.),
Co.
(Mich.),
OL!
Maryland
Long
(Idaho),
Victorville
OL!
Island,
OT!
Golden
I ndian Gap
Nev.),
(Colo.),
TE!
OT!
OL!
OL!
TW!
Mc Cann
(Placer Co.,
Utah Lake
(Great Smoky Mt.
(Krombein 1979); N.Y.),
(Idaho),
Goose Meadow
Bena
(Minn.),
N.P.,
Calif.),
(Utah),
N.C.
-
Kalbfleisch Research Station
WA!
OT!
Benzie
Tenn.),
OT!
(Huntington,
AM!
4. Anteon nebulosum n. sp.
FEMALE:
fully
reddi sh-dark; testaceous;
winged;
length 3,31
mm;
antennae testaceous,
black;
mandibles and clypeus
with segments 1-4 brown;
antennae distally sl ightly thickened;
following proportions:
11: 7:
8,5:
8,5:
7:
6,5:
antennal
6:
6,5:
legs
segments in
6,5:
8;
head dull,
granulated and punctate,
mostly near clypeus;
keel
keels around orbits directed towards antennal
and with two
lateral
frons with a short median
sockets; occipital carina complete; POL = 5; OL = 4; OOL = 6; OPL = 5; = 6; pronotum dull, strongly sculptured by transversal keels; pronotal tubercles reaching tegulae; scutum
(10:18);
punctures; scutum;
posterior surface of pronotum shorter than
scutellum shiny,
notaul ices incomplete,
punctate,
without sculpture among
reaching approximately 0,65 length of
scutellum and metanotum shiny,
propodeum reticulate rugose, posterior surface;
TL
smooth,
with a transversal
without sculpture; keel
between dorsal
posterior surface with 2 longitudinal
keels;
and
median area
as rugose as lateral areas; wings fully darkened; fore-wing with distal part of radial vein much shorter than proximal part (7: 14); fore tarsal segments in following proportions: 294)
with one bristle on a
(Fig.
294)
tibial
spurs 1,1,2.
MALE:
10:
proximal
4:
with 2 rows of 2+23 lamellae;
unknown
4,5:
13:
prominence;
25;
enlarged claw
segment 5 of front
apex with a group of 4
(Fig.
tarsus
lamellae;
462
Mem. Amer.
Fig. 294 -
En!.
Ins!.,
no.
37,
1984
Chela of Anteon nebulosum n.
sp.
(type)
LOCUS TYPICUS: Lyme (Connecticut, U. S. A.) TYPICAL MATERIAL: holotype F! in WA. HOSTS: unknown DISTRIBUTION: Nearctic region: U. S. A.: Lyme (Connecticut),
5. Anteon funestum (R.C.L. Perkins)
WA!
Anteoninae
463
Chelogynus funestus R.C.L. Perkins 1907: 48. Chelogynus melanacrias R.C.L. Perkins 1907: 49 (n. syn.); locus typicus Nogales (Ariz. U. S. A.). Chelogynus lusus R.C.L. Perkins 1907: 50 (n. syn.); locus typicus Tucson (Ariz., U. S. A.). Chelogynus funestus R.C.L. Perkins: Muesebeck and Walkely 1951: 1040.
Anteon (Chelogynus) funestus (R.C.L. Perkins): Krombein 1979: 1241.
Anteon (Chelogynus) lusus (R.C.L. Perkins): Krombein 1979: 1241. Anteon (Chelogynus melanacrias (R.C.L. Perkins): Krombein 1979: 1241.
FEMALE:
fully
winged;
antennae yellow, segment 10 brown; femora brown;
length 1,83-2,59 mm;
black;
mandibles yellow;
with segments 6-10 or 7-10 brown; tegulae testaceous;
sometimes all
legs yellow,
occasionally only with stalk of hind
coxae, clubs of femora and tibiae brown;
occasionally only clubs of femora and hind tibiae brown; segments in following proportions: shiny,
granulated,
frons with one
11:
5:
8:
3:
4:
4:
4:
antennal 4:
4:
6;
head
with a few or numerous irregular keels or areolae;
longitudinal
towards antenna I sockets;
median keel
POL
=
5;
OL
and 2 keels near orbits directed
=
3;
OOL
=
6;
OPL
=
4;
TL
=
6;
occipital carina complete; pronotum granulated, except for an area near posterior margin which is smooth and without sculpture; pronotal tUbercles
Fig.
295 -
Chela of Anteon funestum
(R.C.L.
(Arizona)
Perkins)
from Nogales
464
Mem.
reaching
tegulae;
Amer.
scutum shiny,
notaulices incomplete and approximately 0,20 metanotum shiny, with an
part
(2:5);
37,
1984
keel
distal
scutellum and
segments
and posterior surface;
fore-wing hyaline,
part of radial
fore tarsal
smooth;
propodeum reticulate rugose,
between dorsal keels;
granulated,
reaching only
of scutum;
without sculpture; longitudinal
bands;
no.
very reduced,
(rarely 0,30)
length
smooth,
irregular transversal
proximal
Ins!.,
weakly or strongly
usually
posterior surface without dark transversal
En!.
without
vein much shorter than
in following proportions:
5:
2:
2: 5:
14; enlarged claw (Fig. 295) with one bristle on a proximal prominence; segment 5 of front tarsus (Fig. 295) with 2 rows of nearly 16 lamellae extending continuously
to the apex;
shorter than distal
spurs 1,1,2.
MALE:
fully
part;
winged;
teeth brown;
tibial
length
antennae fully
coxae,
clubs of femora and
shiny,
granulated,
longitudinal
TL = 3;
scutum weakly granulated;
keel
with
tibiae
tegulae testaceous;
legs yellow,
(except for the fore tibiae)
occipital
smooth,
not
notaulices incomplete,
head
frons with a
POL = 5;
OL = 3;
granulated; very
OOL
occasionally
reduced,
reaching
scutellum and metanotum shiny,
propodeum reticulate rugose,
between dorsal
brown;
keels near orbits directed
carina complete;
length of scutum;
without sculpture;
transversal
with
brown;
scutum shiny,
only approximately 0,25 smooth,
1,64-1,90 mm; black; mandibles yellow,
and with 2 lateral
median keel
OPL = 2;
part of segment 5
with some irregular keels or areolae;
towards antenna I sockets; = 4;
basal
and posterior surface;
with an
irregular
posterior surface
without longitudinal keels; propodeum with transversal keel centrally forming a short straight line; fore-wing hyaline, without dark transversal bands;
distal
part of radial
vein much shorter than proximal
288 A) without dorsal membranous process, not pointed distal inner process; tibial spurs 1,1,2. gonoforceps
(Fig.
LOCUS TYPICUS:
Tucson
TYPICAL MATERIAL:
(Arizona,
U.
(2:5);
S. A.)
and 5 paralectotypes
lectotype F!
part
with a wide,
(3 FF, 2 MM)!
iunestus in B; lectotype F! and 1 paralectotype F! (designated by Olmi 1982) of Ch. melanacrias in B. lectotype F! and 1 paralectotype M! (designated by Olmi 1982) of Ch. lusus in B. (designated by Olmi
1982) of Ch.
HOSTS: in U. S. A. Jassids. DISTRIBUTION: Nearctic region: OT!
U.
S.
(Ariz.),
A.:
OL!
Tucson
TW!
(Ariz.),
Ramsey Cyn.
CANADA: B!
CAl
(Sierra Vista,
Cochise Co.
(Portal,
Co.,
TE!
Gainesville (Fla.),
TW!
Monterey
Tex.),
(S.C.),
OL!
Kouchibouguac N.P.
Nogales (Ariz.),
Chiricahua Mts., (Va.),
OT!
Ariz.),
OT! OU
OL!
B!
Huachuca Mts., SCI
(N.B.),
WA!
Ariz.),
College Station
Forsyth
Takoma Pk.
(Ga.), (Md.),
OL!
Portal
OT! OL!
OT!
(Brazos Cleveland TW!
BERMUDA IS.: Bermuda, OT! MEXICO: La Ciudad (Durango), OT! OL! EI Saito (Durango), OT! OU Monterrey (Nuevo Leon), OL! TE! Teziutlan ·(Puebla),
OT!
6. Anteon wasbaueri n. sp.
Anteoninae
Fig.
296 -
winged;
465
Chela of Anteon wasbaueri n.
FEMALE:
fully
reddish;
clypeus,
mandibles,
length 2,06 mm;
distally
thickened;
antennal
black,
antennae and segments
sp.
(type)
with prothorax fully
legs fully
testaceous;
in following proportions:
antennae
6:
3,5:
4,5: 4: 4: 3,5: 3,5: 3,5: 3,5: 5; head dull, granulated, with frons weakly sculptured by longitudinal keels; frons also with an incomplete median keel; occipital carina complete; POL ~ 4,5; Ol ~ 3; OOl ~ 4,5; OPl ~ 3;
Tl ~ 4; pronotum dull, granulated, weakly transversal striate, with posterior surface shorter than scutum (7:10); pronotal tubercles reaching tegulae;
scutum shiny,
sculpture; scutum;
notaul ices
smooth,
very
incomplete,
weakly
faintly
scutellum and metanotum shiny,
propodeum reticulate rugose, and posterior surface;
granulated or almost without
visible near anterior margin of smooth,
without sculpture;
with a strong transversal
posterior surface without
keel
longitudinal
between keels;
dorsal fore-
wing hyaline, without dark transversal bands; distal part of radial vein shorter than proximal part (1,5:8); fore tarsal segments in following proportions:
5,5:
bristle on a
proximal
with
1 row of 25
1:
2,5:
5:
10,5; enlarged claw
prominence;
lamellae;
(Fig.
segment 5 of front
apex with a
group of 5
296)
tarsus
lamellae;
with one (Fig. tibial
296) spurs
466
Mem.
Amer.
En!.
Inst.,
no.
37,
1984
\ \
\
A
Fig.
Male genitalia of Anteon wasbaueri n.
297 -
(Texas}(B:
left half removed);
A:
gonoforceps
sp.
from Speir Rch.
(laterally viewed).
1,1,2. MALE:
fully
winged;
length 1,75-2,81
mm;
black;
mandibles testaceous;
antennae brown; legs brown, with tarsi light; antennae not distally thickened; antenna I segments in following proportions: 7: 3: 3,5: 3: 4: 4: 4: 4: 6; head dull, granulated, and irregularly reticulate rugose; frons
with a complete median keel
and with
two
directed towards antenna I sockets;
occipital
carina complete;
=
3;
OOl
=
granulated;
5;
OPl
=
3.
notaul ices
Tl
=
3;
scutum shiny,
incomplete,
visible only
lateral
4:
keels around orbits POL = 6;
weakly or strongly near anterior margin of
Ol
Anteoninae
scutum;
scutellum and metanotum shiny,
propodeum reticulate rugose, and posterior surface; wing hyaline,
not pointed distal
MM!
part
(Uvalde,
holotype F!
bands;
(1,5:5,5);
inner process;
Speir Rch.
TYPICAL MATERIAL:
weakly granulated;
posterior surface without
much shorter than proximal wide,
smooth,
with a strong transversal
without dark transversal
LOCUS TYPICUS:
467
keel
distal
keels;
fore-
part of radial
gonoforceps
tibial
between dorsal
longitudinal (Fig.
297)
vein
with a
spurs 1,1,2.
Uvalde Co.,
Texas,
and 3 paratypes MM!
U.
S.
A.)
in SC;
3 paratypes
(Uvalde,
Uvalde Co.,
in OL.
HOSTS:
unknown
OISTRIBUTION: Texas),
SC!
Nearctic region:
U.
S.
A.
Speir Rch.
OU
the females of A. wasbaueri is clearly different from the funestum for its reddish prothorax, the males are a real headache. I was not able to find differences between males of A. wasbaueri and A. funestum. I fA. wasbaueri proves to be a jun ior synonym of A. funestum, then its female will be considered as a rare colour variety of AAA. funestum. NOTES:
whereas
A.
female of
The species
is named
in
honor of Mr.
Marius S.
Wasbauer.
7. An teon minimum (Fen ton)
Chelogynus minimus Fenton 1927: 13. Anteon (Chelogynus) minimus (Fenton): Krombein 1979: 1241.
FEMALE:
fully
winged;
antennae fully
length 2,00-2,50 mm;
black;
mandibles testaceous;
brown or with segments 1-2 testaceous;
abdomen brown;
legs testaceous or brown,
with
tarsi
tegulae testaceous; light;
antennal
segments in following proportions: 9: 4: 4: 3: 3: 4: 3,5: 3: 3: 5,5; head shiny, smooth, weakly punctate and alutaceous; frons with a longitudinal POL ~ 4;
median keel;
carina complete;
OL ~ 3;
OOl ~ 5,5;
pronotum shiny,
without sculpture on posterior half; scutum shi ny, notaul ices
smooth,
incomplete,
weak I y
pronotal
punctate, smooth,
with an
posterior surface;
posterior surface with 2
proportions: nearly 20
4:
2,5:
2:
prominence; lamellae;
part 4:
(2:6); 12;
bands;
keel
distal
enlarged claw group of 6
keels;
area;
(Fig.
and
median area
fore-wing
part of radial
segments
tarsus
propodeum
between dorsal
longitudinal
fore tarsal
segment 5 of front
apex with a
tegulae;
length of scutum;
with a smooth and shiny central
shorter than proximal a proximal
tubercles reaching
irregular transversal
without dark transversal
occipital
smooth and
without sculpture;
reticulate rugose,
hyaline,
TL ~ 4;
wi thout scu I pture among punctures;
reaching approximately 0,5
scutellum and metanotum shiny,
only partly rugose,
OPl ~ 4,5;
rugose on anterior half,
vein much
in following
298 with one bristle on (Fig.
lamellae;
298)
basal
with part of
row of
468
Mem. Amer. Ent.
Inst.,
no. 37,
1984
\
Fig. 298 - Chela of Anteon minimum (Fenton)(type)
Anteoninae
segment 5 shorter than distal MALE:
part;
469
tibial
spurs 1,1,2.
unknown
LOCUS TYPICUS:
Barneveld
TYPICAL MATERIAL: HOSTS:
(N.
Y.,
holotype F!
s.
U.
A.)
in WA.
unknown
DISTRIBUTION: Nearctic region: U. S. Kouchibouguac N.P. (N.B.), OT! OL!
A.:
Barneveld
(N.Y.),
WA!
CANADA:
8. An teon masoni n. sp.
FEMALE:
fully
scutellum,
winged;
metanotum,
reddish-brown;
length 2,31-2,50 mm;
abdomen
testaceous or brown;
scutum with reddish or brown spots; segments shiny,
in following proportions:
smooth,
occipital 3;
very
9:
5:
OOL = 6;
TL = 5;
6:
rugose on I y
tuberc les reachi ng
scutum,
tegu I ae;
5:
6:
5:
and
5:
lateral
pronotum shiny,
scu I pture among punctures, smooth,
6,5:
thickened; 5:
head and antennal
7;
head
without sculpture among punctures;
frons without median
OPL = 4;
with
mesopleura
somet i mes pronotum,
antennae distally
weakly punctate,
carina complete;
OL = 3;
reddish-testaceous,
propodeum and metapleura black;
keels;
punctate,
near anterior marg in;
POL = without
pronota I
scute II um and metanotum sh i ny,
without sculpture or weakly punctate and without sculpture among
punctures; scutum;
notaulices incomplete,
reaching
propodeum reticulate rugose,
approximately 0,65
with 2
longitudinal
length of
keels on posterior
surface; median area as rugose as lateral areas; fore-wing hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal
part
proximal
(4:10);
3:
11:
part
(4: 10);
19; enlarged claw
prominence;
distal
fore tarsal (Fig.
segment 5 of front
299)
apex with a group of 4
produced
into a
tibial
vein much shorter than
in following proportions:
6:
2:
with one bristle on a proximal
tarsus
lamellae;
hook;
part of radial segments (Fig.
lamellae;
299)
with
1 row of 24
segment 2 of front
tarsus
spurs 1,1,2.
MALE (not reared): fully winged; length 3,25 mm; black; mandibles testaceous; antennae black; legs testaceous, with coxae partly black and femora partly
brown;
antennae not distally
in following proportions: shiny,
11:
strongly punctate,
weakly granulated;
frons
sculpture among punctures; sculpture; transversal with
10:
10:
antennal
10:
10:
segments
11;
incomplete median keel, lateral
head
carina complete;
scutum shiny, notaul ices
POL = 7,5;
weakly punctate,
incomplete,
reticulate rugose,
reaching
between dorsal
with a
fore-wing
OL = 3,5;
without approximately
smooth,
without
track of a
and posterior surface;
tracks of 2 longitudinal keels;
visible only
keels directed
scutellum and metanotum shiny,
propodeum dull, keel
thickened;
10:
frons also with 2
occipital
TL = 5;
0,5 length of scutum;
10:
with a short
towards antenna I sockets; OPL = 5;
10:
without sculpture among punctures or very
in front of anterior ocellus; OOL = 8;
7:
hyaline,
posterior surface without dark
470
Mem.
Fig.
transversal (5:12);
bands;
Amer.
299 -
Chela of
distal
gonoforceps (Fig.
membranous process and
En!.
Ins!.,
distal
Ottawa
(Ontario,
wide,
dorsal
Canada)
part
proximal
inner pointed process;
1,1,2. LOCUS TYPICUS:
1984
vein shorter than proximal
with a very
with a
37,
Anteon masoni n. sp. (type)
part of radial 300)
no.
tibial
spurs
Anteoninae
Fig.
300 -
Male genitalia of
TYPICAL MATERIAL:
holotype F!
in TW; 1 paratype F! HOSTS: unknown DISTRIBUTION: (Ont.),
OLl
OL!
A.:
Corvall is
(Ont.),
OT!
Parke Reserve (Kam. (Oreg.),
Poughkeepsie (N.Y.),
OL! TW!
Anteon masoni n. sp. from Hull (Quebec)
and 5 paratypes FF!
5 paratypes FF!
Nearctic region:
Finland
OT!
in AM;
471
C,\NADA: Ottawa
TW!
(Ont.),
Summit of King Mt.
Co.,
Que.),
Highlands
OT!
in OT;
1 paratype F!
in OL.
OL!
(N.C.),
OT!
OL!
Rockport
(Old Chelsea,
Hull OL!
(Que.), Laurel
Que.),
OT!
(Md.),
U.
S.
OT!
4 mi. NNW Kerhonkson (Cherry town, Ulster Co.,
N.Y.),
AM!
NOTES:
the species is named in honor of Mr.
W.
R.
9. Anteon rugosiceps Kieffer
Anteon rugosiceps Kieffer 1906a: 238.
M.
Mason.
472
Mem. Amer.
Fig.
301
-
Chela of
En!.
Ins!.,
no.
37,
1984
Anteon rugosieeps Kieffer from Me Bride Springs (California)
473
Anteon i nae
Anteon rugosiceps Kieffer: Kieffer 1914b: 168. Anteon rugosiceps Kieffer: Muesebeck and Walkley 1951: 1040. Anteon rugosiceps Kieffer: Krombein 1979: 1241.
FEMALE
(not reared):
testaceous;
fully
winged;
antennae testaceous,
3,75-4,00 mm;
with hind coxae almost fully black; distally thickened; 9:
9:
9:
9:
9:
9:
antennal 11;
mandibles legs testaceous,
abdomen partly testaceous;
segments in following proportions:
head shiny,
numerous irregular keels, frons,
black;
with segments 6-10 brown;
not granulated,
antennae 12: 7:
11:
punctate and with
very numerous particularly on anterior part of
where there are numerous areolae;
frons with a complete median
keel; occipital carina complete; POL = 9; Ol = 6; OOl = 9; OPl = 7; Tl = 6; pronotum shiny, with anterior half rugose and posterior surface of pronotum shorter than scutum scutum,
(6:22);
pronotal
scutellum and metanotum shiny,
sculpture among punctures; 0,3 length of scutum; transversal
keel
part
(6:10);
reaching
propodeum reticulate rugose,
keels;
median area smooth,
distal
fore tarsal
part of radial
without
approximately
with a strong
and posterior surface;
at most with some areolae around margins; or less darkened;
weakly punctate,
notaul ices incomplete,
between dorsal
with 2 longitudinal
tubercles reaching tegulae;
smooth,
posterior surface
shiny,
without sculpture,
fore-wing with distal
half more
vein much shorter than proximal
segments in following proportions:
6:
6:
5:
13: 30;
enlarged claw (Fig. 301) with one bristle on a proximal prominence; segment 5 of front tarsus (Fig. 301) with 2 rows of 29 lamellae extending continuously to the apex; MALE:
fully
winged;
antennae brown;
legs brown,
thickened;
antennal
6,5: 7:
9;
7:
Ol = 3;
shiny,
black;
with tarsi
mandibles testaceous;
light;
antennae not distally 9:
5:
6:
7:
granulated and fully reticulate rugose;
and 2 lateral
OOl = 5;
smooth,
spurs 1,1,2.
segments in following proportions:
head dull,
with a median keel 5;
tibial
length 2,18 mm;
OPl = 4;
keels;
Tl = 4;
occipital scutum,
very weakly punctate,
7:
7:
frons
carina complete;
POL =
scutellum and metanotum
without sculpture among punctures;
notaulices incomplete, reaching at most 0,5 length of scutum; propodeum reticulate rugose, with a transversal keel between dorsal and posterior surface;
posteior surface with 2 longitudinal
as lateral distal
areas;
fore-wing hyal ine,
part of radial
gonoforceps (Fig. without distal
lOCUS TYPICUS:
302)
DISTRIBUTION:
Shasta,
holotype M!
Nearctic region:
Santa Clara Co. (Mt.
without dorsal
Santa Clara Co.
Kouchibouguac N.
P.
median area as rugose
vein much shorter than proximal
inner pointed process;
TYPICAL MATERIAL: HOSTS: unknown
keels;
without dark transversal
(N.
Siskiyou Co.,
CAl
bands;
(6:4);
membranous process and
spurs 1,1,2.
(California,
U.
S.
A.)
in CA. CANADA:
B.), OT!
(Calif.),
proximal tibial
part
u.
Onyx
Calif.),
S.
Dirleton A.:
(Cal if.), OT!
Ol!
(Ont.),
Pinehurst Ol!
OT!
OT!
Ol!
(Oreg.),
TW!
OL!
Mc Bride Springs
Washoe lake (Washoe Co.,
474
Mem.
Fig.
302 -
Amer.
Ent.
Inst.,
no.
37,
1984
Ma Ie gen i ta I i a of Anteon rugosiceps Kieffer (type)
Nev.), TE! OLl Parker Canyon Lk. (Ariz.), TW! Mt. Vernon Cn. Colo.), OT! Golden (Colo.), OT! OL! MEXICO: Ixtapan de La Sal OT!
10. Anteon puncticeps Ashmead
(Golden, (Mexico),
475
Anteoninae
Anteon puncticeps Ashmead 1893: 97. Aphelopus americanus Ashmead 1893: 100 (n. syn.); locus typicus Jacksonvi lie
(Florida,
U.S.
A.).
Chelogynus vivariensis Brad ley 1926: 7 (n. syn.); locus typicus Mc Lean Bogs Bioi.
Res.
(N.
Y.,
U.S.
A.).
Chelogynus virginiensis Fen ton 1927: 14 (n. syn.); locus typicus I"osslyn
(Virginia,
U.
S.
A.).
Chelogynus vivariensis Bradley: Muesebeck Anteon puncticeps Ashmead: Krombein 1979: Anteon (Chelogynus) virginiensis (Fenton): Anteon (Chelogynus) vivariensis (Bradley):
and Walkley
1951:
1041.
1241. Krombein
1979:
1241.
Krombein
1979:
1241.
FEMALe: fully winged; length 2,5-3,0 mm; colour variable; usually black; sometimes head reddish-testaceous, with a brown spot on ocellar triangle and behind ocelli; testaceous; black;
antennae fully
mandibles testaceous;
testaceous or brown,
mesopleura black or testaceous;
abdomen black or brown; brown;
pro;:lOdeum
metapleura black or reddish-dark;
legs testaceous or with stalk of hind femora
occasionally body fully
propodeum brown;
with segments 1-2
thorax black or reddish-dark;
antennal
testaceous,
segments
with dorsal
surface of
in following ,oroportions:
10:
5:
9:
7:
6: 6: 6: 6: 5: 8; head dull, weakly granulated, with a few irregular short keels mainly near orbits; frons with a median keel, without lateral keels, occipital carina complete; POL = 5; OL = 4; OOL = 7; OPL = 6; 7; pronotum dull, strongly transversely striate; posterior surface of pronotum shorter than scutum tegulae;
scutum,
(11:16);
posterior
scutellum and metanotum shiny,
TL
lobes of pronotum reaching smooth,
finely punctate,
part Fig.
303 -
Chela of
Anteon puncticeps Ashmead from Hc Lean Bogs Bioi. Res.
(New York)
476
Mem.
Amer.
En!.
Ins!.,
no.
37,1984
\
\\
Fig.
304 -
Chela of Anteon puncticeps Ashmead from Rosslyn variety with one row of lamellae.
(Virginia):
Anteoninae
without sculpture among punctures;
477
notaul ices
incomplete,
reaching nearly
0,50-0,66 length of scutum; propodeum reticulate rugose, transversal
keel
between
with 2 longitudinal
dorsal
keels;
and posterior surface;
with a strong
posterior surface
median area nearly as rugose as lateral
areas;
fore-wing hyal ine, without dark transversal bands; distal part of radial vein much shorter than proximal part (6:12); fore tarsal segments in
9: 3: 3: 9: 20; enlarged claw (Figs. 303, 304) with 1 bristle on a proximal prominence; segment 5 of front tarsus (Figs. 303, 304) with 1-2 rows of nearly 22 lamellae; apex with a group of 4-8 following proportions:
I ameli ae;
segment 5 of front tarsus with basa I part shorter than apica I
(8:12); segment 2 of front tarsus produced into a hook;
part
tibial
spurs
1,1,2. MALE:
fully
length 1,56-2,50 mm;
winged;
teeth brown;
antennae brown,
black;
mandibles yellow,
with
1 testaceous or with ventral
with segment
side yellow; sometimes segments 2-5 brown; tegulae testaceous; legs yellow, sometimes with basal part of hind coxae brown, always with hind clubs of femora and hind tibiae more or less brown; rugose and punctate;
frons with a
head dull,
longitudinal
granulated,
weakly
POL = 6;
median keel;
OL
= 3; OOL = 7; OPL = 4; TL = 5; occipital carina complete; scutum shiny, smooth,
finely punctate,
rugose only near anterior margin;
notaulices
incomplete, reaching mid-length of scutum; scutellum and metanotum shiny, smooth, without sculpture; propodeum reticulate rugose, with an irregular transversal keel between dorsal and posterior surface; posterior surface with 2 longitudinal straight keels; median area as rugose as lateral areas;
fore-wing hyaline,
without dark transversal
bands;
distal
part of
radial vein much shorter than proximal part (4:10); gonoforceps (Fig. C) with a wide dorsal proximal membranous process and with a distal inner pointed process; LOCUS TYPICUS:
TYPICAL MATERIAL:
spurs 1,1,2.
tibial
Arlington
(Virginia,
U.
S.
A.) of A.
and 1 para type M!
holotype M!
handwritten by Ashmead Anteon punctaticeps:
label
293
puncticeps (type
in the original
description Anteon puncticeps) in WA; holotype F! of Ch. vivariensis in IT; holotype M! of A. american us in IVA; holotype F! of Ch. virginiensis in WA. HOSTS:
unknown
DISTRIBUTION: Quadra
Is.
Nearctic region:
(B.C.),
OT!
Robson
(B.
OT!
Morley
(Alta.),
TW!
C.),
Stittsvi lie (Ont.),
OT! OT!
OT!
Funkstown
Ancaster
OT!
One Sided Lake
(Ont.),
OT!
Tillsonburg
Beach
(Madoc,
Knowlton
BM!
of King Mt.
(Old Chelsea,
(Que.),
Cap Chat
OT!
Lawrencetown
N.
C.),
S.),
Golden L.
Brighton
Braeside (Ont.), aT! OT!
Old Chelsea
(ant.),
U.
S.
A.:
OT!
OT!
aT!
P.
Mt.
aT!
(N.
Park
Crystal (Ont.),
aT!
Summit
Quyon
B.),
Ranier
OL!
Innisvi lie
Rondeau
OT!
(Que.),
(Que.),
C.),
(Alta.),
Chaffeys Locks Hull
Kouchibouguac N. OT!
WAr (B.
(ant.),
(Ont.),
Bothwell
(Que.),
C.),
Kananaskis
OL!
OT!
aT!
(B.
Cultus Lake
OT!
WAr
Dundas (Ont.),
Que.),
Is.
OT!
OT!
OT!
Cap Rouge
(Que.),
(Hal ifax Co.,
(B.
(Man.),
(Ont.),
OL!
C.),
(Ont.), OT!
(Ont.),
Ont.), OT! (Que.),
Vancouver
(B.
Mission City
(ant.),
TW!
CANADA:
Kowichan L.
OT!
(Wash.),
TW!
478
Mem. Amer. Ent.
Inst.,
no. 37,
1984
Fig. 305 - Chela of Anteon xanthothorax (Bradley)(type)
Ochoco Creek (Oreg.), TW! Corva II is (Oreg.), TW! Mc Bri de Spri ngs (Mt. Shasta, Siskiyou Co., Calif.), OT! Texas, PH! Salmon (Anderson Co., Tex.), TE! College Station (Brazos Co., Tex.), TE! Washtenaw Co. (Mich.),
Anteoninae
Ol!
lake Odessa
lake Co.,
(Mich.), Ohio),
Ridge,
(Mich.),
BM!
AM!
Tenn.),
Ol!
Columbia OT!
(Boone Co.,
Ol!
TW!
TW!
Greenville (5.
(N.
C.),
(Va.),
WA!
Island
(Md.),
TW!
Wattacoo
OT!
C.),
WAr
(D.
laurel
(Md.),
TW!
(V",.), C.),
(Md.),
TW!
WAr
WA! S.
C.),
TW! C.),
FAils Church
TW!
(Del.),
Gun
Morgan (East
Cleveland
Mc Clellanville TW!
OT!
Wayah
WA!
TW!
(Md.),
ZO!
OT!
(Va.),
(Pa.),
Cabin John
Wi Imington
BM!
(Ga.),
Ol! (N.
Spring Br.
OT!
Ol!
Chapin Sanctuary
Forsyth
Highlands
WA!
WA!
(Mich.),
NNW Mc Connelsvi lie,
Mo.),
(Pickens Co.,
Arlington
Washington
Takoma Pk.
(8 mi.
Jacksonville (Fla.),
(5. Bald
Ann Arbor
Muskingum River
(5. C.), C.),
BM!
479
Rosslyn
Plummers
IT!
Bowie (Md.),
Moorestown
(N.
J.),
TW! Westerly (R. I.), TW! Mc lean Bogs Biological Reservation (N. Y.), IT! lake Placid (N. Y.), OT! Keene Valley (Essex Co., N. Y.), OT! Poughkeepsie TW!
Shokan
(N. (N.
Y.),
TW!
Y.),
TW!
South Haven
(long
(Huntington,
N.Y.), AM!
Y.),
AM!
Island,
lewisboro
Washington
(N.
Farmingdale (N. Syracuse N.
4 mi.
Y.),
OT!
TW!
TW!
Hancock
AM! N.
Bigelow lawn
Bemus PI. (N.
(N.
Y.),
Y.),
TW!
Kalbfleisch Field Station
NNW Kerhonkson
(Westchester Co.,
H.),
Y.),
(N. Y.),
Y.),
(Mt.
(Cherry town,
AM!
Washington,
11. Anteon xanthothorax
Ulster Co.,
N.
Alpine Garden of Mt. N.
H.),
OT!
(Brad ley)
Chelogynus xanthothorax Brad ley 1926: 8. Chelogynus xanthothorax Brad ley: Muesebeck and Wa I k I ey 1951: 1041.
Anteon (Chelogynus) xanthothorax (Bradley): Krombein 1979: 1241.
FEMALE:
fully
winged;
antennae brown,
length 2,50-3,37 mm;
black;
with segments 1-3 testaceous;
mandibles testaceous;
prothorax yellow;
abdomen
sometimes brown-testaceous; legs testaceous; antennae distally thickened; antennal segments in following proportions: 11: 5: 8: 7: 7: 6: 7: 7: 7: 9; head shiny,
smooth,
puncta te on frons, median and = 7;
lateral
OPl = 6;
punctate,
weakly punctate on vertex,
wi thout scu I pture among punctures; keels;
Tl = 6,5;
occipital carina complete; pronotum shiny,
wi thout scu I pture among punctures; than scutum smooth,
(12:13);
pronotal
weakly punctate,
incomplete and weakly
smooth,
more strongly
frons
very weakly
Ol = 4;
OOl
punctate,
posterior surface of pronotum shorter
tubercles reaching tegulae;
scutum shiny,
without sculpture among punctures;
impressed,
wi thout
POL = 5;
but only on
notaulices
the anterior third;
scutellum
and metanotum shiny, smooth, without sculpture; propodeum reticulate rugose, with a strong transversal keel between dorsal and posterior surface;
posterior surface with 2 longitudinal
rugose as
lateral
the pterostigma; part
(5:10);
areas; distal
fore tarsal
keels;
fore-wing almost hyal ine,
part of radial
median area as
with a pale band near
vein much shorter than proximal
segments in following proportions:
8:
4:
4:
7:
19;
480
Mem.
Fig.
306 -
Odessa
Amer.
En!.
Inst.,
no.
37,
1984
Male genitdlia of Anteon xanthothorax
(Michigan):
the
left half is ventrally
(Bradley)
viewed;
from Lake
the right half
is
dorsally viewed.
segment 5 of front (9:
10);
tarsus
enlarged claw
prominence; lamellae;
with basal
(Fig.
305)
segment 5 of front
part sl ightly
tarsus
(Fig.
apex with a group of nearly 7
tarsus produced
into a hook;
tibial
305)
and tibiae partly
antennal
segments
head shiny, without
testaceous;
longitudinal
punctate,
segment 3 of front mandibles testaceous; light; sometimes also
antennae not distally 8:
5:
8:
7:
thickened;
8:
7:
7:
7:
without sculpture among punctures;
median keel;
part
spurs 1,1,2.
in following proportions:
smooth,
apical
proximal
with 2 rows of nearly 33
lamellae;
MALE: fully winged; length 1,56-2,75 mm; black; antennae black or brown; IpSlS brown, with tarsi femora
shorter than
with one bristle on a
occipital carina complete;
POL
6:
8;
frons
=
6;
Ol
4; OOl = 6; OPl = 4; Tl = 5; scutum, scutellum and metanotum punctate, wi thout scu I pture among punctures; notau I ices i ncomp lete, reach i ng approximately 0,5 strong transversal surface with hyal ine,
two
length of scutum; keel
longitudinal
keels;
without dark transversal
shorter than proximal
propodeum reticulate rugose,
between dorsal
part
(4:8);
and posterior surface;
median area rugose;
bands;
distal
gonoforceps
fore-wing
part of radial
(Fig.
306)
with a
posterior
with a
vein much large
Anteoninae
dorsal
proximal
process;
membranous process and with a distal
tibial
Taughannock Falls (New York,
TYPICAL MATERIAL:
holotype F!
U.S.A.).
and 1 paratype F!
in
IT.
unknown
DISTRIBUTION: C.),
OL!
Ottawa (Ont.),
Brighton (Ont.),
OT!
Nearctic region:
(B.
OT!
inner pointed
spurs 1,1,2.
LOCUS TYPICUS: HOSTS:
481
Kleanza Creek
(On!.), OT!
Chelsea, (Que.),
OT!
Que.), OT!
(Ont.), OT!
OT!
OT!
(Que.),
Kouchibouguac N. St.
P.
John's
(Que.), OT!
(N.
OT!
OT!
OL!
Stittsville (Ont.),
Innisville
B.), OT!
(On!.), OT!
St.
(Ont.), OT!
Bonaventure OT!
Is.
(Kam.
Aylmer
Andrews
OT!
U. S.
(Alta.), OT!
OT! OT!
Wakefield
Effingham
(Ont.),
Summit of King Mt.
Parke Reserve
(Newfoundland),
Robson
Waterton
Hami Iton
Fabre (Que.),
OT!
OT!
OT!
OT!
(Yukon),
C.),
Belleville (Ont.),
Chimo (Que.),
Cap Rouge
Sheldon L.
B.
(Ont.),
Marmora (Ont.),
(Ont.),
New Richmond S.), OT!
OT! Golden L.
Frankford
Chatterton
CANADA:
(Terrace,
(Que.), Co.,
(Que.),
(N.
B.),
A.:
Lowman
OT!
Que.), OT!
OT!
(Old Hull OT!
OLl
Lockeport (Idaho),
(N.
TW!
Benzie Co. (Mich.), OL! Washtenaw Co. (Mich.), OL! Iron Co. (Mich.), Ann Arbor (Mich.), OL! BM! Lake Odessa (Mich.), OL! BM! Hartford (Conn.),
OT!
Keuka Lake
Falls (Ithaca, Y.),
N.
Y.),
(Penn Yann,
IT!
Y.)(Bradley
Oneonta
(N.
Canajoharie (N. Y.), TW! Rome (N.Y.), TW! Milford Center South Haven (Long Island, N. Y.), AM! 4 mi. NNW Dryden
Ulster Co.,
(Maine),
OL!
N.
Y.),
AM!
Alpine Garden
TW!
Taughannock
TIV! TW!
(Cherry town,
(N.Y.),
OT!
(N. Y.), (N. Y.),
AM!
Millwood
1926);
lO!
Shokan
Y.),
TW!
Y.),
TW!
Kerhonkson
Y.),
N.
OLl
N.
(N.
N.
Six-Mile (Ithaca,
OL!
Westchester Co.
Brainard (Mt.
(Rensselaer Co.,
Washington,
N.
H.),
OT!
12. Anteon osborni (Fenton)
Chelogynus chlorotettixi Fenton 1918a: 202 (nomen nudum). Chelogynus osborni Fenton 1918a: 272. Chelogynus rugulosus Fenton 1927: 14 (n. syn.); locus typicus Saint John
(New Brunswick,
Canada).
Anteon flaviscapus Whittaker 1930: 67 (n. syn.); locus typicus Hollyburn
(British Columbia,
Canada).
Anteon whittakeri Muesebeck and Walkley 1951: 1041; new name for A. flaviscapus Whittaker 1930 preoccupied by A. flaviscapus Kieffer 1905.
Anteon whittakeri Muesebeck and Walkley: Krombein 1979: 1241. Anteon (Chelogynus) osborni (Fenton): Krombein 1979: 1241. Anteon (Chelogynus) rugulosus (Fenton): Krombein 1979: 1241.
FEMALE:
fully
winged;
length 2,25-3,18 mm;
occasionally reddish-light or reddish-brown;
head black or brown; sometimes with anterior half
482
Mem. Amer.
Ent.
Inst.,
no.
37,
1984
\
I
/
Fig.
307 -
Chela of Anteon osborni
I
/
/
/
/
(Fenton) (type)
Anteoninae
of frons
testaceous and with posterior half black;
testaceous;
mandibles testaceous;
testaceous,
thorax black;
with anterior half of pronotum brown;
legs testaceous,
clypeus sometimes
antennae brown or
segments 2-10 or 3-10 or 4-10 darkened; yellow,
483
with
sometimes prothorax
abdomen black or brown;
someti mes with hind coxae basa II y
black or with cl ubs of
mid femora, hind coxae, clubs of hind femora, hind tibiae brown; antennal segments in following proportions: 11: 5: 7: 7: 6: 6: 5: 5: 5: 8; head shiny,
smooth,
frons;
no sculpture among punctures;
OL
=
4;
=
OOL
striate,
finely punctate, 5;
=
OPL
5;
=
TL
6;
occipital carina complete;
pronotum shiny,
smooth only near posterior margin;
tegulae;
scutum shiny,
punctures; scutum;
smooth,
notaulices incomplete,
reaching
with an
and posterior surface;
rugose,
pronotal
weakly punctate,
scute II Um and metanotum shi ny,
propodeum reticulate rugose, dorsal
strongly punctate only on anterior half of POL = 6;
transversely
tubercles reaching
without sculpture among
approximately 0,65
smooth,
length of
without scu I pture;
irregular transversal
keel
between
posterior surface with 2 longitudinal
keels;
median area partly rugose, with a shiny and smooth central area; forewing hyal ine, without dark transversal bands; distal part of radial vein much shorter than proximal proportions: a proximal nearly 29
7:
2:
3:
8:
prominence; lamellae;
18;
part
tibial
MALE:
10);
(Fig. tarsus
segments
307)
in following
with one bristle on
(Fig.
group of 4 lamellae;
307)
with 2 rows of
segment 5 with basal
segment 2 of front tarsus produced
into a
spurs 1,1,2.
fully winged;
length 1,50-2,62 mm;
antennae testaceous or testaceous-brown; fully
fore tarsal
segment 5 of front
apex with a
part shorter than d i sta I part; hook;
(3:
enlarged claw
black;
mandibles testaceous;
sometimes segment
1 I ight;
legs
testaceous or with hind COXae partly black and femora partly brown;
antennae not distally thickened; proportions: punctate,
8:
5:
7:
7:
7:
7:
antenna I segments in following
6,5:
6,5:
7:
10;
without sculpture amongpunctures;
median keel
head shiny,
smooth,
frons with a short
in front of anterior ocellus and with a
weakly
incomplete
weak median furrow,
without lateral keels; occipital carina complete; POL = 5,5; OL = 3; OOL 7; OPL = 4; TL = 4; scutum shiny, smooth, weakly punctate, without sculpture among punctures,
weakly rugose only near anterior margin;
notaulices incomplete, reaching approximately 0,5 length of scutum; scutellum and metanotum shiny, smooth, without sculpture; propodeum reticulate rugose,
with a
posterior surface;
posterior surface with 2
strong
as rugose as
lateral
bands;
part of radial
distal
gonoforceps
(F i g.
areas;
288 C)
transversal
keel
between dorsal
longitUdinal
fore-wing hyal ine,
keels;
without dark
vein much shorter than proximal
with a
and
median
area
transversal part
(4: 10);
I arge dorsa I prox i ma I membranous process
and with a distal inner pointed process; tibial LOCUS TYPICUS: Sandusky (Ohio, U. S. A.).
spurs 1,1,2.
TYPICAL MATERIAL: holoytpe F! of Ch. osborni in WA; holotype F! of Ch. rugulosus in WA; holotype M! and 3 paratypes MM! of A. flaviscapus Whittaker in BM; 1 paratype M! of A. flaviscapus Whittaker in WA; 1 paratype M! of A. flaviscapus in LA.
484
Mem.
HOSTS:
in U.
S.
DISTRIBUTION: CANA:JA: (B.
B.
C.),
(Yukon),
Hixon
OT!
(B.
B.
C.),
OT!
Atlin
(Osoyoos,
B.
OT!
aT!
Golden L.
(Ont.),
aT!
Maynooth
Stittsville
aT!
Hollyburn
(Ont.), OT! (Ont.), aT!
aT!
OT!
P.
(Oreg.),
TIV!
Stanley
Elk Creek (Cal if.),
(3 mi. TW!
(N.
Brandon
Is.,
OT!
(Man.),
(Ont.),
aT!
aT!
(Que.),
OT!
OL!
Hull
B.),
WA!
Saint John
TW!
Corvall is
(Idaho), (Ariz.),
TW!
Somerset
Siskiyou Co., TW!
Estes Park
Ottawa
Dresden (Ont.),
(Ont.), aT!
One
(Ont.),
aT!
Fort Chimo
OT!
Parke Reserve (Kam. OT!
Chimo
Old Chelsea
(Wash.),
aT!
TIV!
(Que.),
Que.), OT!
C.),
Marmora
Niagara Glenn
(Ont.),
B.
Richter Pass
OT!
E Mc Cloud,
Portal
C.),
Frances (Ont.),
(Old Chelsea,
Ranier
LA!
Bancroft
Lac Brule (Que.),
Mt.
Chilliwack
IVA!
aT!
aT!
aT!
OT!
BM!
Innisville (Ont.), OT!
Meach Lake (Que.),
Kouchibouguac N.
C.),
Ancaster (ant.),
Chaffeys Locks
Mistassini
B.
Brighton
TW!
B.),
Ft.
OLl
(Ont.), OT!
OT!
Spring Creek
(Vancouver
(Terrace,
Cumberland Que.),
B.
c.),
(B.
Cowichan Bay
Lakelse L.
aT!
Summit of King Mt.
OT!
Galiano (B. C.), BM! Robson (B. C.), OT! C.), aT! Burnaby (B. C.), OT! Shames (Terrace,
Ninette (Man.),
(Ont.),
(Fairbanks),
(Yukon),
Sided Lake (Ont.),
Co.,
37,1984
Birch L.
Sheldon L.
(B. C.), OT!
C.),
no.
Madden Lake (Oliver,
C.),
C.), OT!
(Ont.), OT!
ALASKA:
aT!
Cowichan L. (B. c.), OT! Gagnon Road (Terrace, B. Cultus Lake (B.
Inst.,
Chlorotettix unicolor (Fitch) (Fenton 1918b).
A.
C.), OL!
BM!
Ent.
Nearctic region:
Dawson
(Terrace,
Amer.
(N.
B.),
(Oreg.), (EI
AM!
(Colo.),
(Que.),
Fredericton
(~J.
Rhododendron
Dorado Co.,
Calif.),
aT!
(Que.), OT!
WA!
TW!
(Que.),
Calif.),
TE!
Crescent City
TW!
aT!
Lawrence
(Kans.), TW! (Mich.), BM!
Benzie Co. (Mich.), OL! Iron Co. (Mich.), OL! Ann Arbor TW! Washtenaw Co. (Mich.), Sandusky (Ohio), WA! Cleveland
(S. C.), OL!
TW!
C.), OT!
(State Forest, (Md.),
lVattacoo (Pickens Co.,
Indian Gap
OT!
IV.
(Gr.
Va.),
Smoky Mt.
aT!
High Point St.
Hawksbi II
Pk.
(N.
S. C.),
Nat.
J.),
Pk.,
aLl
N.
(Shenandoah, TIV!
TW!
C.),
Va.),
Lakehurst
Highlands (N.
aT!
Cooper's Rock
aT!
(N. J.),
Laurel TW!
(R. I.), TW! Syracuse (N. Y.), TIV! South Haven (Long Island, N. Y.), AM! Ithaca (N. Y.), ZO! Sloansville (N. Y.), OT! Farmigdale (N. Y.), TIV! Bemus PI. (N. Y.), TIV! Shokan (N. Y.), TIV! Millwood (N. Y.), TIV! Canajoharie (N. Y.), TW! So. Hadley (Mass.), TIV! Alpine Garden (Mt. Washington, N. H.), OT! MEXICO: EI Saito (Durango), OT! La Ciudad (Durango), aT!
lVesterly
13. Anteon durangoi n. sp.
FEMALE:
unknown
MALE: fully winged; length 2 mm; black; mandibles testaceous; antennae and legs brown; antennae not distally thickened; antennal segments in following proportions: 7: S: 6: 6: 7: 5: 6: 6: 6: 9; head dull, granulated, with track of areolae on vertex and with numerous longitudinal striae on frons; frons without median and lateral keels; occipital
carina complete;
POL
=
5;
OL
=
3;
OOL
=
4;
OPL
=
3;
TL
=
5;
Anteoninae
Fig.
308 -
Male genitalia of n.
scutum shiny, notaul ices
sp.
Anteon ciudadi n. sp.
(type)(B):
weakly punctate,
incomplete,
faintly
without
(type)(A)
and
durangoi
right half removed.
without sculpture among punctures;
visible near anterior margin of scutum;
scutellum and metanotum shiny, reticulate rugose,
485
smooth,
without sculpture;
longitudinal keels on
propodeum
posterior surface,
with a
transversal keel between dorsal and posterior surface; fore-wing hyal ine, without dark transversal bands; distal part of radial vein shorter than proximal
part
(2,5:9);
membranous process,
gonoforceps
with a distal
(Fig.
308 B)
without dorsal
iner pointed process;
1,1,2. LOCUS TYPICUS: La Ciudad (Mexico) TYPICAL MATERIAL: holotype M! in OT.
tibial
proximal spurs
486
Mem.
HOSTS:
Amer.
En!.
Ins!.,
no.
37,
1984
unknown
DISTRIBUTION:
Nearctic region:
MEXICO:
La Ciudad
(Durango),
OT!
14. Anteon ciudadi n. sp.
FEMALE: MALE:
unknown
fully
winged;
length 2,50-2,87 mm;
antennae testaceous-brown; brown;
antennae not distally
puncta te, lateral
black;
mandibles testaceous;
testaceous or wi th coxae and femora
thickened;
antenna I segments
10: 7: 8: 9: 9: 8: 8: 8: 8:
proportions:
=
legs fu II y
10;
head shiny,
without scu I pture among punctures;
=
occipital carina complete; POL = 6; OL = 3,5; OOL = 6; OPL 5; scutum, scutellum and metanotum weakly punctate, without
sculpture among punctures;
notaul ices
incomplete,
reaching
0,3 length of scutum; propodeum reticulate rugose, keels on posterior surface; fore-wing hyal ine,
median area dull,
without dark transversal
vein shorter than proximal tibial
weakly
frons wi thout med i an and
keels;
3,5; TL
dorsal
in following
smooth,
proximal
with 2
approximately longitudinal
as rugose as
bands;
distal
lateral
(5:9); gonoforceps (Fig. 308 A)
part
membranous process,
without distal
areas;
part of radial without
inner pointed process;
spurs 1,1,2.
LOCUS TYPICUS: La Ciudad (Mexico) TYPICAL MATERIAL: holotype M! in OT; HOSTS: unknown
paratype M!
DISTRIBUTION:
Pinehurst
La Ciudad
Nearctic region:
(Durango),
U.
S.
A.:
in TW.
(Oreg.),
TW!
MEXICO:
OT!
15. Anteon menkei n. sp.
FEMALE:
unknown
MALE: fully winged; length I,BI-2,12 mm; head black, with anterior half of frons, clypeus and mandibles testaceous; antennae and legs fully testaceous;
somet i mes antennae darkened;
abdomen black;
antennae not distally
following proportions: punctate, lateral
B:
6:
6,5:
7:
thorax and propodeum black;
thickened;
7:
7:
7:
7:
without scu I pture among punctures;
keels;
occipital
carina complete;
POL
antenna I segments 7:
9;
head shiny,
frons without med i an
=
5;
OL
=
3;
OOL
=
in weakly and
6;
OPL
=
3; TL = 4; scutum shiny, smooth, weakly punctate, without sculpture among punctures; notaul ices incomplete, reaching approximately 0,5 length of scutum;
scutellum
and
metanotum shiny,
smooth,
weakly
punctate,
without sculpture among punctures; propodeum reticulate rugose, with 2 longitudinal keels on posterior surface; median area as rugose as lateral areas;
fore-wing hyal ine,
radial
vein shorter than proximal
without dark part
transversal (4:10);
bands;
distal
gonoforceps (Fig.
part of 309)
Anteoninae
Fig.
without dorsal
309 -
Male genitalia of Anteon menkei n.
proximal
membranous process,
process; tibial spurs 1,1,2. LOCUS TYPICUS: Gatlinburg (Tennessee, TYPICAL MATERIAL: holotype M! paratypes MM! in OL. HOSTS: unknown DISTRIBUTION: Nearctic region: Cleveland NOTES:
(S.
C.),
487
TW!
Wattacoo
in WA;
sp.
with a distal
(type)
inner pointed
U.S.A.). 2 paratypes MM!
U. S. A.:
Gatlinburg
(Pickens Co.,
probably this species is a male variety
S.
in TW;
(Tenn.),
C.),
TW!
2
WA!
OLl
(with frons partly
testaceous) of A. osborni. The species is named in honor of Mr. Arnold S.
Menke.
16. Anteon hirtifrons Whittaker
Anteon hirtifrons Wh i ttaker 1930: 68. Anteon hirtifrons Whittaker: Muesebeck and Walkley 1951: 1040.
488
Mem. Amer.
Anteon hirtifrons
FEMALE: MALE:
Ent.
Ins!.,
no.
37,
Krombein 1979:
Whittaker:
1984
1241.
unknown
fully
winged;
antennae testaceous, with hind coxae,
length 2,52 mm;
black;
mandibles testaceous;
wi th segments 1-3 dorsa II y
darkened;
femora and tibiae partly darkened;
legs testaceous,
antennae not distally
thickened; antenna I segments in following proportions: 13: 7: 9: 9: 9: 8: 8: 8: 8: 12; head strongly punctate and reticulate rugose, with irregular keels;
frontal
line complete;
occipital carina complete;
POL; 8;
OL ;
4;
6; TL ;
3,5; scutum, scutellum and metanotum shiny, smooth, without sculpture; notaulices incomplete, reaching approximately 0,5 length
OOL ;
of scutum;
propodeum reticulate rugose;
longitudinal
keels;
posterior surface with
median area as rugose as lateral
areas;
two
fore-wing
hyaline, without dark transversal bands; distal part of radial vein much shorter than prox i ma I part ( 5: 10); gonoforceps (F i g. 293 B) with a very large distal
inner pointed process;
LOCUS TYPICUS:
Hollyburn
TYPICAL MATERIAL: very damaged, HOSTS:
(B.
C.,
holotype M!
without body,
spurs 1,1,2.
tibial
Canada)
in BM;
1 paratype M! in LA (specimen legs and wings).
only with antennae,
unknown
DISTRIBUTION: Nearctic region: Galiano (B. C.), BM!
CANADA:
Hollyburn
NEOTROPIC SPECIES OF
(B.
C.),
LA!
BM!
ANTEON
Twenty one species are known.
KEY TO THE SPECIES
FEMALES
4 of front tarsus at most 0,5 as long as segment 310, 317) ••••••••.•••••••••••••.•••••.•.••.•••••••..... 2 Segment 4 of front tarsus slightly shorter, as long as, or longer than segment 1 (Figs. 311, 313) . . . . . . . . . . . . . . . . . . . . . . . . 4 Segment (Figs.
2
3
Posterior surface of
propodeum wi th
two longi tudinal keels ••••• 1. c1avatum Olmi and Currado Posterior surface of propodeum without longitudinal keels . . . . . 3 Scutum fully granulated; frons reticulate rugose and granulated . . • . . . • • . • • • • . • . • . . . • • • .. •• 6. mexicanum n. sp • Scutum weakly punctate, wi thout sculpture among punctures;
Anteoninae
frons
rugose,
without
sculpture among keels,
5
Posterior
surface of
propodeum without
Posterior
surface of
propodeum with
Fore-wing hyaline, Fore-wi ng wi th
6
without
1-2 dark
Pronotum and head
dark
keels . . . . 5
longitudinal
keels •• 13
bands.....
..6
bands.............
.10
transversal at
granulated •.•
sp.
longitudinal
two
transversa I
fully black,
not
oliveirai n.
• •.•••.•••••••••••••.••.•••.•• 7.
4
489
most
with mandibles . •• 7
testaceous............................................. Pronotum and head at
7
Head by
granulated,
numerous
Head
8
fully
not
part I y
red
reticulate
t estaceous •••••••••• 8
or
rugose,
sculptured by numerous areolae and
molle n.
Head
red . . . . .
pronotum fully
sp.
· . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3.
nigrorubrum n.
Head with anterior half
testaceous-whitish;
pronotum fully Scutum fully
of
of
frons
frons black
punctate and wi thout
Scutum with anterior rugose,
or partly
sp.
punctate,
region
wi th
SCUlpture among
panamense n. sp.
smooth;
irregular
posterior half
lobes;
surface of
posterior without
Posterior
surface of
surface
of
pronotum wi th
surface of
punctate,
dull,
striae •••••••••••
· . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5. jamaicanum n. 10 Posterior
vertex
reddish-dark;
or most Iy black . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9
smooth,
punctures • •••.•••••••.•••••••• 4.
11
sculptured
irregular keels •••• •••
· . . . • . . • . • . • . • . . . . • . . . . . . . . . . . 2. fully black;
not
micros n. sp.
irregular keels .. 20.
and posterior half 9
I east
two dorsal
propodeum smooth,
sp. transversal shiny,
weakly
sculpture among punctures . . . . . . . . . . . . . . . . . 11 pronotum wi thout
propodeum dull,
fully
dorsal
reticulate
lobes; rugose
posterior or with
transversal
keels . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 12
Head
without
shiny,
sculpture,
or punctate and without
noyesi n. sp. surinamense n. sp.
sculpture among punctures •.•.. 8. Head du I I, 12 Posterior
g ranu I at ed •••••••• 21. surface of
pronotum shiny,
without
sculpture,
smooth; posterior surface of propodeum wi th strong keels, not reticulate rugose.13. annulicorne Brues Posterior
surface of
surface of
pronotum dull,
propodeum reticulate
Pronotum not 14 Head
fully
two dorsal
forming
albitarse (Came ron)
lobes near posterior margin . . . . . 14
two dorsal
lobes • •••••••••••••••••••
granulated . . . . . . . . 19.
Head punctate,
without
posterior
rugose ••• ••••••••.••••••••
• •••••••••.•••••••••••••••••• 14. 13 Pronotum forming
granulated;
transversal
•• 15
bolivianum n. sp.
sculpture among punctures,
not
granulated . . . . . . . . . . . . . . . . . . . . 9. lobatum n. sp. 15 Anterior surface of pronotum nearly as long as posterior surface (Fig. 322 B); head fully except for the frontal I ine .. 10. Anterior
surface of
pronotum much
smooth,
wi thout
sculpture,
propodeale (Fenton) shorter
than posterior
490
Mem.
surface 16 Head as,
(Fig.
fully or
Amer.
En!.
Ins!.,
rugose,
longer
frons
only
with OPL
longitudinal
median
vertex
striae
occipital
rugose,
smooth,
nearly
as
long
in
the
with OPL nearly
three
wi th numerous
region
between
anterior
carina . . . . . . . . . . . . . . . . . . . . . . ..
nycteum n.
• •••••••••••••••..•••..•.•••• 1 2.
The females of
shorter,
chiriquense (Cameron)
reticulate
long as POL;
ocellus and
1984
than POL . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
•• • . •• ••••••••• •• • •• ••• • ••••• 11. times as
37,
322 A) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 16
reticulate
sl ightly
Head with
no.
sp.
A. pilicorne, plaumanni, conterminum and vivax are
unknown.
MALES
2
Posterior
surface of
propodeum with
Posterior
surface of
propodeum without
Head
fully
reticulate
rugose.18.
Head punctate and wi thout reticulate
two
longitudinal longitudinal
plaumanni n. sp.
sculpture among punctures,
rugose . . . . . . . . . . . . 15.
keels ••• 2 keels . . . . 3 not
vivax n. sp.
3
Scutum granulated •••••••••••••••••••••••••••••••••••••••••••• 4 Scutum weakly punctate, without sculpture among punctures . . . . 5
4
Gonoforceps wi thout
di stal
inner rounded pro<:css
(Fig.
329)
• •••••••••••••••••••••••••••• 1 6. pilicorne (Og I ob lin) Gonoforceps with distal inner rounded process (Fig. 318) . . .
· . . . . . . . . . . . . • • . . • • . . . . . . . . . . . 6.
5
Head with numerous
longitudinal
inner pointed process
(Fig.
6
reticulate
rugose,
sp.
gonoforceps with distal
312) . . . . . . . . . . . . . . . . . . . . . . . . . . . . ..
· • . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2. Head
mexicanum n. keels;
without
molle n. sp. longitudinal keels;
gonoforceps wi th or wi thout di stal inner pointed process (Figs 316, 330) ••••••••••••••••••••••••••••••••••••••••••••••••••.• 6 Gonoforceps wi th di stal inner pointed process (Fig. 330) . . . . . . • •••••••••••••••••••••••••••• 17. conterminum n. sp. Gonoforceps without distal inner pointed process (Fig.
. . . . . . . . . . . 5. jamaicanum n.
316) . . .
sp.
The males of A. clavatum, albitarse, lobatum, propodeale, annulicorne, chiriquense, oliveirai, micros, nigrorubrum, panamense, noyesi, surinamense, bolivianum and nycteum are unknown.
Anteoninae
491
1. Anteon clavatum Dlmi and Currado
Anteon c1avatum Dlmi and Currado 1979a: 343.
Fig.
310 -
Chela of Anteon c1avatum Dlmi
and Currado
(type)
492
Mem.
FEMALE:
fully
Amer.
winged;
Ent.
length 4,37 mm;
antenna I segments 1-2 testaceous; redd i sh;
legs testaceous,
antennae distally 17:
10:
9:
frontal
OOl = 6; fully
7:
7:
7:
8:
OPl = 6;
scutellum shiny, margin;
7:
7:
9;
segments
head fully
with
teeth
pronotal
smooth,
reticulate rugose,
tubercles reaching
weakly
punctate,
smooth,
with a
shiny;
POL = 8;
Ol = 4;
pronotum shiny,
tegulae;
scutum dull,
notaulices invisible;
reticulate rugose near posterior
without sculpture;
transversal
brown;
in following proportions:
occipital carina complete;
posterior surface with 2
surface;
with antennae brown;
reticulate rugose only on sides;
metanotum shiny,
reticulate rugose,
black,
1984
mandibles testaceous,
antennal
Tl = 3;
granulated,
37,
frons with 2 keels around orbits;
reticulate rugose;
smooth,
no.
with fore coxae and fore femora part I y
thickened;
line complete;
Inst.,
keel
propodeum shiny,
between dorsal
longitudinal
keels;
and posterior
median area as
reticulate rugose as lateral areas; fore-wing hyal ine, without dark transversal bands; distal part of radial vein much shorter than proximal part
(4:13);
pterostigma
following proportions:
one bristle located on a (Fig.
310)
3:
2,5:
4:
proximal
with some bristles;
part of segment 5 much MALE:
long and narrow;
15:
12;
fore tarsal
segments
enlarged claw
(Fig.
segment 5 of front
prominence;
apex with a
longer than apical
in
310)
group of 13 lamellae; part;
tibial
with tarsus basal
spurs 1,1,2.
unknown
lOCUS TYPICUS:
Liangtelau
TYPICAL MATERIAL:
(Brazil)
holotype F!
HOSTS: unknown DISTRIBUTION: Neotropic region:
in GE. BRAZil:
liangtelau,
GE!
2. Anteon molle n. sp.
FEMALE:
fully
winged;
length 2,12-2,62 mm;
black;
mandibles testaceous;
antennae brown; legs testaceous, with hind coxae basally brown; antennae distally thickened; antennal segments in following proportions: 10: 6: 9: 9: 9: 8: 8: 8: areolae and longitudinal
8: 10; head dull, granulated, with numerous keels; frontal I ine complete; occipital carina
complete;
=
POL = 4,5;
Ol
3;
OOl
=
7;
OPl
=
4;
Tl
=
5,5;
pronotum dull,
granulated, rugose, with transversal keels, smooth only near posterior margin; posterior surface of pronotum shorter than scutum (5:14); pronotal
tubercles reaching
smooth,
shiny,
notaul ices
tegulae;
weakly punctate,
incomplete,
scutum,
scutellum and metanotum
without sculpture among punctures;
reaching approximately 0,3
length of scutum;
propodeum reticulate rugose, with a transversal keel between dorsal and posterior surface; posterior surface without longitudinal keels; fore-wing hyal ine,
without dark transversal
bands;
distal
part of radial
vein
shorter than proximal part (4:9); fore tarsal segments in following proportions: 9: 2,5: 3,5: 5,5: 15; enlarged claw (Fig. 311) with one bristle
located on a
proximal
prominence;
segment 5 of front
tarsus
(Fig.
Anteoninae
Fig.
311)
with
311
-
1 row of 37
493
Chela of Anteon molle n.
lamellae;
apex with a
sp.
(type)
group of 5
lamellae;
tibial
spurs 1,1,2. MALE:
fully
winged;
length 1,87-2,00 mm;
black;
mandibles testaceous;
antennae brown, with segments 1-2 testaceous; legs testaceous, with coxae and clubs of femora partly brown; antennae not distally thickened; antenna I segments in following proportions: 8: 4: 5: 6: 6: 6: 5: 5: 6: 8; head shiny,
with frons sculptured by numerous
behind posterior ocelli 5,5;
OL = 2,5;
OOL = 6;
metanotum shiny, punctures; scutum; dorsal
reticulate rugose; OPL = 3;
smooth,
reaching
propodeum reticulate rugose, and posterior surface;
fore-wing hyal ine, proximal
part
scutum,
de
transversal bands;
gonoforceps
with distal
las Casas
POL =
without sculpture among
with a
(3:7);
region
scutellum and
approximately 0,25
transversal
membranous process,
tibial spurs 1,1,2. LOCUS TYPICUS: San Cristobal
carina complete;
posterior surface wi thout
without dark
vein shorter than proximal dorsal
TL = 4;
weakly punctate,
notaulices incomplete,
longitudinal keels;
occipital
keel
length of between
long i tud i na I keel s;
distal (Fig.
part of radial 312)
without
inner pointed process;
(Mexico)
494
Mem.
Fig.
Amer.
Ent.
Inst.,
no.
37,
1984
312 - Male genitalia of Anteon molle from S. Cristobal de las Casas (Mexico):
TYPICAL MATERIAL: holotype F! (4 MM, 1 F)! in OL. HOSTS: unknown
left half removed
and 9 paratypes MM!
DISTRIBUTION:
Neotropic region:
MEXICO: San Cristobal
(Chiapas),
! OT!
(Sinaloa),
OT!
OL!
Ol
COLOMBIA:
EI Cali
Palmito
(Valle del
Cauca),
Ol!
OT!
5 paratypes
in OT;
de las Casas
Concordia
(Sinaloa),
Ol!
3. Anteon nigrorubrum n. sp.
FEMALE:
fully
testaceous;
winged;
length 1,25 mm;
antennae testaceous;
and propodeum black;
head black,
prothorax red;
abdomen and
with mandibles
mesothorax,
legs testaceous;
meta thorax
antennae distally
Anteoninae
Fig.
thickened;
313 -
Anteon nigrorubrum
Chela of
antenna I segments
495
n.
sp.
in following proportions:
7:
(type)
3,5:
4:
4:
3,5:
3,5: 3,5: 4: 4: 6; head dull, fully granulated and with numerous longitudinal weak striae; frons without median and lateral keels; occipital 4;
carina complete;
pronotum dull,
hairy,
POL = 5;
OL = 2,5;
granulated and
OOL = 4,5;
posterior surface of pronotum shorter than scutum tubercles reaching smooth,
very
tegulae;
scutum,
weakly punctate,
OPL = 2,5;
with some transversal (7:10);
TL
keels;
pronotal
scutellum and metanotum shiny,
without sculpture among punctures;
notaul ices incomplete, reaching approximately 0,65 length of scutum; propodeum reticulate rugose, with a strong transversal keel between dorsal and posterior surface; posterior surface without longitudinal keels, fully
reticulate rugose,
bands;
distal
fore tarsal
segments
enlarged claw prominence; lamellae, MALE:
(Fig.
fore-wing hyaline,
in following proportions: 313)
lamellae;
with one bristle tarsus
5:
1:
1,5: 3,5:
located on a
(Fig.
313)
part shorter than apical tibial
without dark transversal
vein much shorter than proximal
segment 5 of front
with basal
group of 4
dull;
part of radial
with
part
part
(1:6);
10;
proximal 1 row of 23
(4:6);
apex with a
spurs 1,1,2.
unknown
LOCUS TYPICUS:
St.
TYPICAL MATERIAL:
Augustine
(Trinidad)
holotype F!
HOSTS: unknown DISTRIBUTION: Neotropic region:
in BM. TRINIDAD:
St.
Augustine
(St.
George),
496
Mem. Amer. En!.
Ins!.,
no. 37,
1984
BM!
Fig. 314 - Chela of Anteon panamense n. sp.
(type)
Anteoninae
497
4. Anteon panamense n. sp.
FEMALE:
fully
half of frons testaceous;
winged;
clypeus
antennae brown, reddish-dark;
testaceous;
abdomen and
8:
4:
with numerous complete; rugose,
with anterior
ventral
mandibles testaceous,
face of head
with
teeth reddish;
6 testaceous;
thorax and
except for anterior margin of pronotum which
antennae distally
proportions:
head black,
with segments 1-5 and part I y
propodeum black, dark;
length 2,06-2,12 mm;
testaceous and temples reddish-dark;
6:
6:
antennal
segments
6:
4:
head dull,
4,5:
4:
4:
8;
in following granulated and
keels on frons and vertex;
OL = 3;
OPL = 2;
OOL = 4;
is
with stalk of hind femora
thickened;
longitudinal
POL = 4,5;
legs testaceous,
occipital
TL = 4;
carina
pronotum dull,
except for a narrow region near posterior margin smooth;
posterior surface of pronotum shorter than scutum tubercles reaching
tegulae;
scutum,
(5:10);
pronotal
scutellum and metanotum shiny,
smooth, weakly punctate, without sculpture among punctures; notaul ices incomplete, reaching approximately 0,3 length of scutum; propodeum reticulate rugose,
with a
posterior surface;
posterior surface without
rugose;
hyaline,
fore-wing
radial
strong
transversal
without dark
in following (Fig.
314)
proportions: located on a
with
apical
part
1,1,2. MALE:
unknown
part 11;
(3:8);
29
holotype F!
keels,
bands;
lamellae,
and
reticulate
distal
fore tarsal
enlarged claw
prominence;
with basal lamellae;
part of
segments
(Fig.
314)
segment 5 of front
apex with a group of 4
Barro Colorado
TYPICAL MATERIAL:
2: 5:
1,5:
1 row of nearly
(4,5:5,5);
LOCU5 TYPICUS: HOSTS:
6:
proximal
between dorsal
transversal
vein much shorter than proximal
one bristle
keel
longitudinal
with
tarsus
part shorter than tibial
spurs
Island (Panama) in LE;
1 paratype F!
in OL.
unknown
DISTRIBUTION:
Neotropic region:
PAt,AMA:
Barro Colorado
Island,
LE!
OL!
5. Anteon famaicanum n. sp.
FEMALE:
fully
winged;
length 2,62 mm;
head black,
clypeus and anterior half of frons testaceous; segment
1 testaceous;
antenna I segments 6:
8;
head dull,
dull,
OL = 4;
antennae distally
in following proportions:
10:
5:
8:
with
thickened;
7:
7:
6:
5,5:
reticulate rugose and with nUmerous longitudinal
frons without median 5,5;
legs testaceous;
with mandibles,
antennae brown,
and
OOL = 5,5;
lateral OPL = 3;
keels;
occipital
TL = 5;
carina complete;
5,5: keels; POL
=
pronotum with anterior region
rugose, striate; posterior surface shiny,
smooth,
not rugose,
shorter
than scutum (7:13); pronotal tubercles reaching tegulae; scutum with anterior region near anterior margin smooth and shiny, without sculpture;
498
Mem.
Fig.
315 -
Amer.
Ent.
no.
37,
Anteon jamaicanum
Chela of
posterior half of scutum dull,
Ins!.,
punctate,
with
1984
n.
sp.
(type)
irregular striae,
notaulices visible only near anterior margin of scutum;
rugose;
scutellum and
metanotum shiny,
smooth, without sculpture; propodeum reticulate rugose, strong transversa I keel between dorsa I and posterior surface; posterior surface without longitudinal keels; fore-wing hyaline, without wi th a
dark transversal part 18;
(4:8);
bands;
fore tarsal
enlarged claw
prominence;
(Fig.
distal
part of radial
segments
in following
315)
vein shorter than proportions:
with one bristle
segment 5 of front
lamellae extending continuously
tarsus
(Fig.
located on a
315)
to the apex;
7:
proximal
2,5:
3:
9:
proximal
with 2 rows of nearly 26
tibial
spurs 1,1,2.
MALE: fully winged; length 1,56 mm; black; mandibles testaceous; antennae testaceous-dark; legs testaceous; antennae distally thickened; antennal segments in following proportions: 8: 3: 6: 5: 5: 5,5: 5,5: 5: 8; head dull, reticulate rugose and granulated; frons with a median keel; TL
occipital
=
2;
carina complete;
scutum,
scutellum
POL = 4,5;
without sculpture among punctures; margin of scutum; keel
distal
and posterior surface; fore-wing hyaline,
part of radial
gonoforceps tibial
keels; (Fig.
316)
3;
=
OOL = 4;
smooth,
near anterior
with a strong transversal
posterior surface without
without dark
vein much shorter than without distal
OPL = 2;
weakly punctate,
notaulices visible only
propodeum reticulate rugose,
between dorsal
longitudinal
OL
and metanotum shiny,
5:
transversal
proximal
part
bands;
(2:7);
inner pointed or rounded process;
spurs 1,1,2.
LOCUS TYPICUS: Hardwar Gap (Jamaica) TYPICAL MATERIAL: holotype F! and 1 paratype M!
in OT.
Anteoninae
Fig.
316 -
499
Male genital ia of Anteon jamaicanum n.
HOSTS: unknown DISTRIBUTION: Neotropic region:
JAMAICA:
sp.
Hardwar Gap,
(paratype)
OT!
6. Anteon mexicanum n. sp.
FEMALE:
fully
winged;
antennae brown,
length 2,00-3,18 mm;
coxae and clubs of femora partly brown; antennal
black;
with segments 1-3 testaceous;
mandibles testaceous;
legs testaceous,
with hind
antennae distally thickened;
segments in following proportions:
10: 5:
5:
4:
4:
4:
5:
5:
5:
6,5; head dull, granulated and reticulate rugose; sometimes head with a few areolae, mostly smooth on vertex; frons with a median keel; occipital carina complete; POL = 5,5; Ol = 3,5; OOl = 6; OPL = 4; Tl = 4; pronotum du II, granu I ated and with transversa I keel s; pronota I tubercles reaching tegulae;
posterior surface of pronotum shorter than scutum
500
Mem.
Fig.
(6:12);
317 -
scutum dull,
Amer.
Chela of
En!.
granulated;
without sculpture;
transversal without
keel
keels;
37,
1984
notaulices
indistinct,
and posterior surface;
fore-wing
faintly
visible
scutellum and metanotum shiny,
propodeum reticulate rugose,
between dorsal
longitudinal
no.
Anteon mexicanum n. sp. (type)
only near anterior margin of scutum; smooth,
Ins!.,
hyaline,
without
with
a
posterior surface dark
transversal
Anteoninae
501
318 - Male genitalia of Anteon mexicanum n. sp. from S. Cristobal
Fig.
de las Casas
(Mexico):
right
half removed.
bands; distal part of radi21 vein shorter than tarsal segments in following proportions: 5: 2:
proximal
2:
part
2,5: 9,5;
(2:6); fore enlarged claw
317) with one bristle located on a proximal prominence; segment 5 tarsus (Fig. 317) with 2 rows of 6 lamel!ae; apex with a group of 8 lamellae; tibial spurs 1,1,2. MALE: fully winged; length 1,81-2,12 mm; black; mandibles testaceous; (Fig.
of front
antennae black; thickened; 5:
5:
5:
median
legs black,
antenna I segments
7; keel;
head dull,
with
tarsi
I ight;
antennae not distally
in following proportions:
9:
granulated and reticulate rugose;
occipital carina complete;
POL = 5;
OL = 3;
5:
5:
5:
5:
5:
frons with OOL = 5;
OPL
3; TL = 3,5; scutum dull, granulated; notaul ices indistinct, faintiy visible only near anterior margin of scutum; scutellum and metanotum
502
Mem.
shiny,
smooth
transversal without
Amer.
distal
(1,5:6);
between dorsal
(Fig.
with distal
LOCUS TYPICUS:
1984
propodeum reticulate rugose,
and posterior surface;
318)
de
TYPICAL MATERIAL:
holotype F!
3 paratypes
las Casas
tibial
2 MM)!
F,
part
membranous
spurs 1,1,2.
(Mexico)
and 5 paratypes
(1
transversal
proximal
proximal
inner rounded process;
with a
posterior surface
without dark
without dorsal
paratype F! HOSTS:
37,
vein much shorter than
San Cristobal
in TW;
no.
fore-wing hyaline,
part of radial
gonoforceps
process and
I nst.,
and without sculpture;
keel
longitudinal keels;
bands;
Ent.
(1
in
F,
4 MM)!
in OT;
OL.
unknown
DISTRIBUTION: (Chiapas),
Neotropic region:
OT!
OL!
JAMAICA:
MEXICO:
Blue Mt.,
San Cristobal
OT!
PERU:
de
las Casas
Machu Picchu,
TW!
7. Anteon oliveirai n. sp.
FEMALE:
fully
winged;
antennae brown,
distally 7:
7:
thickened;
8:
with a
with
7:
7:
length 4 mm; segments
antennal
7:
10;
median keel
segments
head shiny,
and
two
black;
mandibles testaceous;
1-2 testaceous;
legs
testaceous;
in following proportions:
antennae
19:
9:
irregularly sculptured by folds;
lateral
keels around orbits;
occipital
10:
frons carina
complete; POL = 6,5; OL = 4; OOL = 6; OPL = 6,5; TL = 3; region behind ocelli weakly granulated, with irregular folds and with tracks of two oblique keels connecting posterior ocelli to occipital carina; pronotum shiny, with a strong transversal impression between anterior and posterior surface; transversal reaching
anterior part of posterior surface forming
carina,
tegulae;
shiny,
with
pronotal
posterior surface shorter than scutum
scutellum and metanotum shiny, sculpture among punctures; 0,3
irregular folds;
length of scutum;
smooth,
weakly
notaul ices incomplete,
narrow dark band beneath proportions:
11:
located on a
proximal
with
2:
3:
the pterostigma;
part 3:
(6:10);
11,5; enlarged claw
prominence;
tibial
spurs 1,1,2. unknown
LOCUS TYP I CUS:
Serra do Caraca
TYPICAL MATERIAL: HOSTS:
holotype F!
approximately
with a strong posterior surface fore-wing with a
part of radial
vein
segments in following (Fig.
319)
segment 5 of front
two rows of nearly 10 lamellae;
MALE:
distal
fore tarsal
scutum,
without
reaching
transversa I keel between dorsa I and posterior surface; without longitudinal keels, dull and reticulate rugose; shorter than proximal
raised
(7,5:19);
punctate,
propodeum reticulate rugose,
a
tubercles
apex with a
with one bristle
tarsus
group of 2
(Fig.
319)
lamellae;
(Brazi I) in TW.
unknown
DISTRIBUTION:
Neotropic region:
Minas Gerais), TW! NOTES: the species is named
BRAZIL:
Serra do Caraca
in honor of Mr.
F.M.
(5.
Oliveira.
Barbara,
Anteon i nae
Fig.
319 -
503
Anteon oliveirai n. sp. (type)
Chela of
8. Anteon noyesi n. sp.
FEMALE: black
fully
winged;
segments 7-10 brown; distally 4:
length 2,62-3,00 mm;
3,5:
thickened; 5:
7:
6:
6:
9;
punctures,
hairy,
reddish;
antennae
segments in following proportions:
head shiny,
occipital carina complete;
pronotum shiny,
with propodeum
antennae sometimes with
propodeum sometimes partly
antennal
weakly punctate,
without sculpture among punctures; orbits; = 6;
testaceous,
(mesopleura and metapleura testaceous);
smooth,
frons with only 2 lateral
POL = 5;
punctate,
Ol = 3;
7:
6:
keels around
OOl = 6;
without sculpture,
OPl = 5;
transversal
Tl
among
with posterior surface shorter than scutum
posterior surface of pronotum with 2 dorsal
10:
mainly on frons,
(4:20);
lobes;
pronotal
tubercles reaching tegulae; scutum, scutellum and metanotum shiny, smooth, wihtout sculpture; notaulices invisible; propodeum shiny, with a strong
transversal
keel
between dorsal
and posterior surface;
dorsal
surface very short, with a few areolae; posterior surface shiny, smooth, without sculpture, without areolae or keels; fore-wing with 2 dark transversal pterostigma; (4: 10);
bands, one on the basa I cell s and another beneath the distal
fore tarsal
enlarged claw
(Fig.
part of radial segments 320)
with one bristle
prominence;
segment 5 of front
than apical
part
nearly 31
(6:11);
lamellae;
MALE: unknown lOCUS TYPICUS:
St.
TYPICAL MATERIAL:
vein much shorter than proximal
in following proportions: tarsus
located on a
(Fig.
segment 5 of front
apex with a Augustine
7:
group of 6
320)
2,5:
3:
7:
part 17;
proximal
with basal
part shorter
tarsus with some rows of lamellae;
tibial
spurs 1,1,2.
(Trinidad)
holotype F!
and
1 paratype F!
in BM;
1 paratype F!
504
Mem. Amer.
En!.
Inst.,
no.
37,
1984
, \
\ I
t Fig.
in aT;
320 -
1 paratype F!
Chela of Anteon noyesi n.
TRINIDAD:
BM!
Amazonas,
NOTES:
(Arima),
J/
sp.
(type)
in TW.
HOSTS: unknown DISTRIBUTION: Neotropic region: Simla
)
OT!
BRAZIL:
SI.
Augustine
TW!
the species is named in honor of Mr. J.S.
Noyes.
(St.
George),
505
Anteoninae
Fig.
321 -
Chela of Anteon lobatum n.
sp.
(type)
506
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
9. Anteon lobatum n. sp.
FEMALE:
fully
winged;
reddish-testaceous; brown;
length 2,5-3,0 mm;
antennae testaceous,
scutum reddish-testaceous,
scutellum reddish-dark; metapleura black, reddish-brown; segments
in
head and prothorax fully
with segments 7-10 or 9-10
with sides and anterior half black;
metanotum and propodeum black;
with more or
less wide reddish
legs testaceous-dark;
following proportions:
tegulae
11:
6:
spots;
testaceous;
6:
4:
4:
6:
7:
mesopleura and abdomen antennal 7:
7:
9;
head
smooth, hairy, shiny, weakly punctate, without sculpture among punctures; frons without longitudinal median keel, with two lateral around orbits on
the frons POL = 6;
carina complete; shiny,
smooth,
pronotal
and metanotum shiny, visible only
reticulate rugose,
forming
tUbercles reaching
smooth,
lateral
sockets;
two dorsal
tegulae;
without sculpture;
transversal
keel
areas reticulate rugose;
lobes near
scutum,
scutellum
notaulices indistinct,
near anterior margin of the scutum;
with a
keels
occipital
8; OPL = 5; TL = 6; pronotum
OOL =
hairy,
between dorsal
posterior surface with 2 longitudinal
sudace; shiny;
OL = 3;
weakly punctate,
posterior margin; faintly
directed towards antennal
keels;
propodeum and posterior
median area smooth,
mesopleura and metapleura smooth;
fore-wi ng with 2 dark transversa I bands, one on the basa I cell sand another beneath the pterostigma; distal part of radial vein much shorter than proximal part (4:9); fore tarsal segments in following proportions: 3: 2: 3: 5: 14; enlarged claw (Fig. 321) with one bristle located on a proximal
prominence;
nearly 26
lamellae;
segment 5 of front
tarsus
(Fig.
apex with a group of nearly 6
321)
with 2 rows of
lamellae;
tibial
spurs
1,1,2. MALE: unknown LOCUS TYPICUS:
Gamboa
TYPICAL MATERIAL: F!
(Panama)
holotype F!
and 2 paratypes FF!
in LE;
1 paratype
in OL.
HOSTS:
unknown
DISTRIBUTION:
Neotropic region:
PANAMA:
Gamboa
(Canal
Zone),
LE!
OL!
10. Anteon propodeale (Fenton) n. comb.
Chelogynus propodealis Fenton 1927: 13.
FEMALE: tegulae, brown; 4:
6:
with a
fully winged; length 2,25 mm; black; mandibles, antennae, legs testaceous, except hind stalks of femora and hind tibiae antenna I segments
8;
in following proportions:
head very shiny and smooth,
median
longitudinal
keel
11:
5:
without sculpture,
and with 2
lateral
5:
4:
hairy;
4:
4:
4:
frons
keels around orbits;
Anteoninae
507
o
A
Fig.
322 -
Anteon chiriquense (Cameron) propodeale (Fenton) (type) (B) (laterally viewed).
Head and prothorax of females of
(type) (A)
and
occipital carina complete; POL = 5; OL = 3; OOL = 5; OPL = 3,5; TL = 3,5; pronotum hairy, very shiny and smooth, without sculpture, convex and raised (Fig.
into a
322 B);
transversal
pronotal
carina between
tubercles reaching
metanotum very shiny and smooth, invisible;
longitudinal
dorsal
keels;
scutum scutellum and
without sculpture,
propodeum with a transversal
and posterior surface; with 2
anterior and posterior half
tegulae;
hairy;
irregular keel
surface reticulate rugose;
median area shiny,
with short keels only near margins;
lateral
smooth,
notaulices
between dorsal posterior surface without areolae,
areas with
wide areolae;
fore-wing with 2 dark transversal bands, one on the basal cells and another beneath the pterostigma; distal part of radial vein much shorter than proximal 4:
1,5:
2:
proximal nearly as
4:
(2:6);
segments in following proportions:
(Fig.
323)
segment 5 of front
long as distal to
fore tarsal
enlarged claw
prominence;
continuously MALE:
part 10;
part,
with nearly
the apex;
tibial
Santarem
(Brazil)
with one bristle
tarsus 14
(Fig.
323)
located on a
with basal
lamellae extending
spurs 1,1,2.
unknown
LOCUS TYPICUS:
TYPICAL MATERIAL: HOSTS: unknown DISTRIBUTION:
holotype F!
Neotropic region:
in WA. BRAZIL:
Santarem
(Para),
WA!
part
508
Mem.
Fig.
323 -
Amer.
En!.
Ins!.,
no.
37,
1984
Anteon propodeale (Fenton)(type)
Chela of
11. Anteon chiriquense (Cameron) n. comb.
Dryinus chiriquensis Cameron 1888: 447. f'4esodryinus chiriquensis (Cameron): Kieffer 1914b: 32.
FEMALE:
fully
winged;
legs testaceous, basally black;
length 3,62-6,00 mm;
with stalks of hind femora antennae testaceous,
testaceous or brown-testaceous;
black; dark;
mandibles testaceous;
sometimes hind coxae
with segments 6-10 dark;
head hairy,
dull,
fully
abdomen
reticulate rugose;
occipital carina complete; POL = 5; OL = 3; OOl = 7; OPl = 6 or 4; Tl = 9; pronotum hairy, punctate, forming a raised transversal carina between anterior and posterior surface sometimes with an
(Fig.
322 A);
incomplete median keel;
than posterior surface;
pronotal
posterior surface of pronotum
anterior surface much shorter
tubercles reaching
tegulae;
sides of
posterior surface not rounded, but raised into longitudinal carinas; scutum, scute II um and metanotum smooth, sh i ny, weak I y puncta te, without sculpture among punctures; 0,25
length of scutum;
notaul ices short,
propodeum with a
incomplete,
transversal
keel
reaching only
between dorsal and posterior surface; dorsal surface reticulate rugose; areolae very wide, with internal surface granulated; posterior surface with 2 longitudinal keels; median and lateral areas almost fully smooth, shiny; mesopleura and metapleura shiny and smooth; fore-wing with 2 light or dark transversa I bands, one on the basa I cell s and another beneath the
Anteoninae
Fig.
pterostigma; (4: 13);
distal
fore tarsal
enlarged claw prominence; lamellae; MALE:
324 -
(Fig;
Chela of
509
Anteon chiriquense
part of radial
(Cameron) (type)
vein much shorter than proximal
segments in following proportions: 324)
8:
4:
4:
part
10: 24;
with one bristle located on a proximal
segment 5 of front
tarsus
(Fig.
apex with a group of 5 lamellae;
324) tibial
with 2 rows of nearly spurs 1,1,2.
unknown
LOCUS TYPICUS:
Volcan de Chiriqui
TYPICAL MATERIAL: HOSTS:
holotype Fl
(Panama)
in BM.
unknown
DISTRIBUTION:
Neotropic region:
PANAMA:
Volcan de Chiriqui,
BM!
BOLIVIA: 10 Km E San Antonio (Rio Mamore, Dept. Beni), AM! BRAZIL: Nova Teutonia (S.ta Catarina), OT! 12° 31' S 55° 37' W (Sinop, Mato Grosso),
TWl
18
510
Mem.
Fig.
Amer.
En!.
Inst.,
no.
37,
1984
325 - Chela of Anteon nycteum n. sp.
(type)
12. Anteon nycteum n. sp.
FEMALE:
fully
winged;
of frons and a small reddish;
length 3,37 mm;
head black,
spot between posteior ocelli
clypeus and mandibles testaceous;
testaceous,
with hind coxae basally black;
and abdomen
black;
antennae distally
with anterior region
and occipital
carina
antennae testaceous; thorax,
thickened;
propodeum,
legs petiole
antenna I segments
in
following proportions: 11: 9: 8: 8: 8: 8: 8: 8: 8: 11; head shiny, with frons fully reticulate rugose; frons without median keel, but with a median furrow connecting anterior ocellus to clypeus;
vertex smooth,
punctate,
numerous parallel
median striae connecting anterior ocellus to occipital
carina; = 3,5;
head
with
OL = 5,5;
without sculpture among punctures,
not
reticulate rugose,
two keels around orbits; OOL = 8;
OPL = 12;
with
occipital carina complete;
TL = 9;
pronotum shiny,
POL
with a
strong transversa I impression;
anterior surface of pronotum much shorter
than posterior surface
pronotum form i ng a
(8: 17);
ra i sed transversa I
carina between anterior and posterior surface; posterior surface smooth, shiny, weakly punctate, without sculpture among punctures, except for SOme transversa I str; ae near the transversal reaching
tegulae;
scutum,
cari na;
pronota I tuberc I es
scutellum and metanotum shiny,
smooth,
weakly
Anteoninae
punctate,
approximately 0,5
transversal with
two
keel
length of scutum;
between dorsal
longitudinal
511
keels;
propodeum with a
and posterior surface;
dorsal
surface reticulate rugose;
areas of posterior surface rugose;
median area smooth,
hyal ine,
bands;
without dark
transversal
shorter than proximal proportions:
11:
located on a
3:
part
4:
10:
proximal
lamellae;
tibial
22;
fore tarsal
enlarged claw
prominence;
with 2 rows of nearly 28 MALE:
(6: 14);
distal
lamellae;
strong
posterior surface lateral
shiny;
fore-wing
part of radial
vein
segments in following (Fig.
325)
segment 5 of front apex with a
with one bristle
tarsus
(Fig.
325)
group of nearly 6
spurs 1,1,2.
unknown
LOCUS TYPICUS:
12° 31'
TYPICAL MATERIAL: HOSTS:
S 55°
37'
holotype F!
W (Sinop,
Mato Grosso,
Brazil)
in TW.
unknown
DISTRIBUTION: Grosso), TW!
Neotropic region:
BRAZIL:
12° 31'
S 55° 37'
W (Sinop,
Mato
13. Anteoll annulicorne Brues
Anteon annulicornis Brues 1905: 185.
FEMALE;
fully
winged;
length 2,75 mm;
head black,
with mandibles
testaceous; antennae testaceous, with segments 2-4 brown, dark; thorax, propodeum and abdomen black; legs brown, light;
antennae distally thickened;
proportions:
11;
numerous short
5:
8;
=
OPL
forming
a
surface; reaching
5;
5:
5:
irregular keels;
keels around orbits; 5,5;
4:
=
TL
median
occipital 5;
6:
antennal 5:
5:
segments median
carina complete;
POL
smooth,
carina between
scutum shiny,
granulated;
granulated,
keel
=
7;
hairy,
and OL
=
with
two lateral 4,5;
OOL
=
without sculpture,
anterior and posterior
posterior surface shorter than scutum tegulae;
in following
head dull,
frons with a
pronotum shiny,
transversal
8;
8-10 brownwith tarsi
(7,5: 17);
pronota I tubercles
notaulices very reduced,
faintly visible near anterior margin of scutum; scutellum and metanotum shiny, amooth, without sculpture, punctate; propodeum with a strong transversal
keel
between dorsal
reticulate rugose;
and posterior surface;
posterior surface without
numerous transversal
irregular keels;
dorsal
longitudinal
fore-wing
with
surface
keels,
with
two dark transversal
bands, one on the basal cell s and another benea th the pterost i gma; distal part of radial vein shorter than proximal part (2:10); fore tarsal segments tarsus with
in following proportions:
(Fig.
326)
with basal
two rows of nearly
enlarged claw prominence;
(Fig.
tibial
326)
part
20:
8:
8,5:
22:
46;
longer than apical
segment 5 of front part
17 lamellae extending continuously with one bristle
spurs 1,1,2.
(28: 18)
and
to the apex;
located on a proximal
512
Mem.
Fig.
MALE:
326 -
Amer.
Chela of
Ent.
Inst.,
no.
37,
1984
Anteon annulicorne Brues (type)
unknown
lOCUS TYPICUS:
Mexico
TYPICAL MATERIAL:
holotype F!
HOSTS: unknown DISTRIBUTION: according
in PH.
to the morphology of the holotype,
I
think
that
A. annulicorne is a Neotropic species. The Mexican locality is unknown. The morphology is very similar to A. albitarse (Cameron).
14. Anteon albitarse (Cameron) n. comb.
Dryinus albitarsis Cameron 1888: 447. Nesodryinus albitarsis (Cameron): Kieffer 1914b: 33.
FEMALE:
fully
winged;
or partly black;
length 1,81-3,00 mm;
antennae testaceous,
segments 6-10 black;
tegulae testaceous;
hind coxae partly black; brown;
with
mid,
black;
mandibles testaceous
last segment brown or with
legs testaceous,
with all or only
hind and sometimes fore clubs of femora
mid and hind tibiae brown;
antenna I segments in following
proportions: 12: 6: 7: 5: 4: 5: 5: 5: 5: 7; head dull, granulated, smooth; frons with a longitudinal median keel and with 2 lateral keels near orbits directed towards antennal POL = 6;
Ol = 4;
OOl = 6;
OPl = 5;
sockets; Tl = 6;
occipital
carina complete;
pronotum dull,
granulated,
Anteoninae
Fig.
327 -
transversal
keels on anterior half;
pronotum forming
med i an cari na between anterior and posterior ha If;
tubercles reaching not visible;
Anteon albitarse (Cameron) (type)
Chela of
with numerous transversal
513
tegulae;
scutum dull,
smooth,
scutellum and metanotum smooth,
granulated,
shiny,
a
pronota I
notaul ices
without sculpture or
weakly punctate; propodeum reticulate rugose, with a transversal between dorsal and posterior surface; posterior surface without
keel
longitudinal keels; mesopleura and anterior half of metapleura dull and rugose; posterior half of metapleura smooth, shiny, without sculpture; fore-w i ng
wi th 2 dark
another beneath than proximal 6:
2:
3:
Bol ivia);
5:
part
16
transversa I bands,
the pterostigma; (in
(2,5:8);
(Fig.
lamellae; apex with a MALE: unknown LOCUS TYPICUS:
DISTRIBUTION:
1,5:
the basa I cell sand
3:
8:
(Fig.
327)
vein much shorter
in following 20
with one bristle
tarsus
group of nearly 17
Panajachel
TYPICAL MATERIAL: HOSTS: unkown
segments
or 5:
327)
segment 5 of front
one on
part of radial
fore tarsal
the holotype)
enlarged claw
prominence;
distal
located on
with
lamellae;
proportions:
(in a specimen from a proximal
1 row of nearly
tibial
spurs 1,1,2.
(Guatemala)
holotype F!
Neotropic region:
in BM. MEXICO:
Vista Hermosa
(Tuxtepec,
12
514
Mem.
Fig.
328 -
Amer.
Ent.
Inst.,
no.
37,
Male genitalia of Anteon vivax n.
1984
sp.
(type):
right
half
removed.
Oaxaca),
OL!
TW!
GUATEMALA:
(5t. George),
OL!
BM!
Panajachel,
COLOMBIA:
Domingo, TW! BOLIVIA: Chulumani Notes; in the borderlands between
Cal i
BM!
(Valle),
TRINIDAD: CAl
51. Augustine
ECUADOR:
Quito /
5.
(La Paz), BM! the Neotropic and Nearctic regions,
A. albitarse can be confused with A. iunestum (Nearctic). The two species are distinguished by the fore wings (hyal ine in iunestum and very dark in albitarse).
15. Anteon vivax n. sp.
FEMALE: MALE:
unknown
fully
winged;
antennae brown,
length 1,93 mm;
black;
mandibles testaceous;
with segments 1-2 testaceous;
coxae basally brown;
legs testaceous,
antennae not distally thichened;
antennal
with hind segments
Anteoninae
Fig.
515
Male genitalia of Anteon pilicorne Ogloblin from Burzaco
329 -
(Argentina) (A:
left
half removed);
B:
gonoforceps.
in following proportions: 8: 6: 7: 6: 7: 7: 7: 7: 6,5: 9; head shiny, punctate, without sculpture among punctures; frons without median and lateral =
2,5;
keels;
occipital
Tl = 4;
scutum,
carina complete;
sculpture among punctures; 0,3
length of scutum;
proximal
notaulices incomplete,
part
(3:
7);
TYPICAL MATERIAL:
de
holotype M!
bands;
OOl
=
punctate,
with 2
gonoforceps
las Casas
3;
reaching
(Fig.
5,5;
approximately areas;
part of radial 328)
without
inner pointed
(Mexico)
in OT.
unknown
DISTRIBUTIOI',: (Chiapas),
Neotropic region:
MEXICO:
San
Cristobal
OT!
16. Anteon pilicorne Ogloblin
Anteon pilicornis Ogloblin 1938: 39.
de
OPl
without
longitudinal
lateral
distal
membranous process and without distal
process; tibial spurs 1,1,2. lOCUS TYPICUS: San Cristobal HOSTS:
=
Ol
median area as rugose as
without dark transversal
vein shorter than proximal dorsal
5;
propodeum reticulate rugose,
keels on posterior surface; fore-wing hyal ine,
=
POL
scutellum and metanotum weakly
las Casas
516
Mem.
FEMALE: MALE:
Amer.
Ent.
Inst.,
no.
37,
1984
unknown
fully
winged;
antennae brown,
mm; black; mandibles testaceous;
length 2,75-3,43
with segment
1 testaceous;
legs testaceous,
and sometimes clubs of femora partly brown;
with coxae
sometimes also hind
tibiae
brown; antennae not distally thichened; antenna I segments in following proportions: 14: 6: 6: 6: 7: 7: 5: 6: 5: 9; head dull, fully reticulate rugose;
frons with a median keel
and with 2
;
5; OOL ;
7; OPL ;
5,5; TL ;
some areolae on anterior third; approximately 0,3 scu I pture; transversal without
keel
between
longitudinal
dark transversal proximal
part
occipital
notaul ices
length of scutum;
metanotum rugose;
bands;
(2:9);
membranous process
incomplete,
reaching
scutellum shiny,
smooth,
and posterior surface;
reticulate rugose;
distal without
(Fig. distal
329)
without
with a strong
posterior surface
fore-wing hyaline,
part of radial
gonoforceps and
keels near orbits
carina complete;
propodeum ret icu I ate rugose,
dorsal
keels,
lateral
POL; 8; OL 5; scutum dull, fully granulated, with
directed towards antenna I sockets;
without
vein much shorter than
without dorsal
proximal
inner pointed process,
with
a
series of papi Ilae on inner face; tibial spurs 1,1,2. LOCUS TYPICUS: General Urquiza (Buenos Aires, Argentina) TYPICAL MATERIAL: HOSTS:
DISTRIBUTION: Veado, BM!
holotype M!
in BA.
unknown Neotropic region:
Distrito Federal),
General
Urquiza
OL!
BRAZIL:
aT!
Estacao Forestal
ARGENTINA:
(Buenos Aires),
(Cabeca do
Burzaco (Buenos Aires),
BA!
17. Anteon conterminum n. sp.
FEMALE: "'·,ALE:
unknown
fully
winged;
antennae brown,
testaceous,
length 2,06
with
segment
mm; black; mandibles testaceous;
1 testaceous;
legs brown,
fore tibiae and femora partly testaceous;
with
tarsi
antennae not
distally thickened; antennal segments in following proportions: 10: 4,5: 4: 4,5: 5: 5: 5: 5: 5; antennal segment 10 missing in the only known specimen;
head dull,
granulated and fully
median complete keel antennal
sockets;
OPL; 2,5; weak I y
TL ;
two
occipital 3;
puncta te,
incomplete,
and
scutum,
lateral
carina complete;
without scu I pture among punctures;
faintly
posterior surface;
posterior surface without
without dark trasversal
shorter than proximal LOCUS TYPICUS:
tibial
part
(1 :7);
trasversal bands;
keel
distal
(Brazil)
3;
OOL ;
4;
smooth, propodeum
between dorsal keels;
part of I'adial (Fig.
towards
notau I ices
longitudinal
gonoforceps
spurs 1,1,2.
Nova Teutonia
frons with a
OL ;
visible near anterior margin of scutum; with a strong
pointed process;
POL; 5,5;
scutellum and metanotum shiny,
reticulate rugose, hyal ine,
reticulate rugose;
keels near orbits directed
and fore-wing vein much
330) with distal
inner
Anteoninae
Fig.
330 -
517
Anteon conterminum n. sp. (type): left half
Male genitalia of
removed.
TYPICAL MATERIAL: HOSTS:
holotype M!
in OT.
unknown
DISTRIBUTION: OT!
Neotropic region:
BRAZil Nova Teutonia
(Sta Catarina),
18. Anteon plaumanni n. sp.
FEMALE: MALE:
unknown
fully
winged;
length 2,0 mm;
head black;
testaceous;
thorax black or brown-reddish;
testaceous;
antennae not distally
following proportions: rugose;
frons
with
9:
5:
7:
a median
thickened;
6:
sockets;
~
Tl
OOl
weakly
~
5,
OPl
punctate,
~
2,5;
without
6:
~
2;
scutum,
transversal with 2
keel
wing hyal ine,
6:
9;
and
keels;
with 2
membranous process and
reticulate keels
POL ~ 5;
Ol
scutellum and metanotum
notaulices
in addition scutum
incomplete, with
and posterior surface;
(3:7);
in
lateral
carina com,:Jiete;
median area as rugose as
without dark trasversal
legs
segments head fully
propodeum reticulate rugose,
much shorter than proximal proximal
6:
sculpture among punctures;
between dorsal
longitudinal
antennal
keel
occipital
reticulate rugose near anterior margin; mid-length of scutum;
6:
longitudinal
directed towards antennal 3;
6:
mandibles and antennae
abdomen black;
bands;
gonoforceps
distal
posterior surface
lateral
areas;
part of radial
(Fig.331)
without distal
reaching
a strong forevein
without dorsal
inner pointed process;
518
Mem.
Fig.
331
Amer.
Ent.
Inst.,
no.
37,
Male genitalia of Anteon plaumanni n.
-
1984
sp.
(type):
right
half
removed.
tibial
spurs 1,1,2.
LOCUS TYPICUS:
Nova Teutonia
TYPICAL MATERIAL: M! in OL. HOSTS:
(Brazil)
holotype M!
in BM;
1 paratype M!
in OT;
1 paratype
unknown
DISTRIBUTION:
Neotropic region:
Nova Teutonia
(S.ta Catarina)
NOTES:
the species is named
JAMAICA:
Hardwar Gap,
OT!
OL!
BRAZIL:
BM!
in honor of Mr.
Fritz Plaumann.
19. Anteon bolivianum n. sp.
FEMALE:
fully
winged;
distally thickened; 3:
2:
5:
7:
only keel POL = 5;
7:
6:
length 2,52 mm;
antennal 9;
head dull,
is around the orbits), OL =
3,5;
fully
reddish-testaceous;
segments in following proportions:
OOL = 9;
not reticulate rugose, granulated;
OPL = 5;
occipital
TL = 6;
with posterior tubercles reaching
with a
impression;
transverse
furrow;
long,
scutum,
sculpture,
very
scutell urn
approximately 0,30
tegulae;
smooth,
pronotum
lobes separated by
and metanotum sh i ny,
weakly punctate;
smooth,
notaul ices incomplete,
length of scutum;
5: (the
carina complete;
posterior surface of pronotum nearly
with two prominent dorsal
5:
without keels
pronotum shiny,
without sculpture, as broad as
antennae 11:
twice
a median
wi thout
reaching
propodeum reticulate rugose,
with a
Anteoninae
332 - Chela of Anteon bolivianum n. sp.
Fig.
strong
transversal
surface with weakly
keel
two weak
rugose,
the basal
519
shiny;
between dorsal
and posterior surface;
longitudinal keels; fore-wing
(type)
median and
lateral
with two dark transversal
cells and another beneath
the pterostigma;
distal
posterior areas
bands,
one on
part of
part (3:9); fore tarsal segmens 7: 3: 3: 8: 21; enlarged claw (Fig. 332) with one bristle located on a proximal prominence; segment 5 of front tarsus (Fig. 332) with two rows of nearly 40 lamellae; apex with a group of nearly 17 lamellae; tibial spurs 1,1,2. radial
vein much shorter than proximal
in following proportions:
MALE:
unknown
LOCUS TYPICUS:
Mouth of Rio Baures (Bolivia)
TYPICAL MATERIAL: HOSTS:
DISTRIBUTION: Itenez,
holotype F!
in AM.
unknown Dept.
Neotropic region: Beni),
BOLIVIA:
Mouth of Rio Baures
AM!
20. Anteon micros n. sp.
(Rio
520
Mem.
Fig.
333 -
Amer.
En!.
Ins!.,
no.
37,
1984
Anteon micros n. sp. (type)
Chela of
FEMALE: fully winged; length 1,64 mm; black; mandibles testaceous; legs brown, with fore tibiae and tarsi yellow; antennae distally thickened; antennal 5;
segments
head dull,
carina complete; dull,
granulated;
POL = 5;
granulated,
more than
in following proportions:
fully
Ol =
frons 3;
7:
3:
3:
3:
3:
2,5:
without median keel;
OOl = 3;
OPl = 4;
Tl 4;
with posterior surface shorter than scutum
twice as broad
as
long
(15:4);
3:
3:
4:
occipital pronotum (4: 10)
and
posterior tubercles reaching
tegulae; scutum dull, granulated; notaul ices incomplete, reaching approximately 0,30 length nf scutum; scutellum and metanotum shiny, smooth,
without
transversal
keel
sculpture;
propodeum
between dorsal
reticulate
rugose,
and posterior surface;
with
a
strong
posterior surface
Anteoninae
without
longitudinal
bands;
distal
keels;
521
fore-wing hyaline,
part of radial
without dark
fore tarsal
segments in following proportions:
5:
claw
333)
proximal
(Fig.
with one bristle
segment 5 of front
group of 4
apex with a MALE:
tarsus
transversal
vein much shorter than proximal
(Fig.
located on a 333)
lamellae;
2:
3,5:
part
9;
(2:5);
enlarged
prominence;
two rows of nearly 8 lamellae;
with
tibial
2:
spurs 1,1,2.
unknown
LOCUS TYPICUS:
Miranda
TYPICAL MATERIAL: HOSTS:
(Venezuela)
holotype F!
in AM.
unknown
DISTRIBUTION:
Neotropic region:
VENEZUELA:
Miranda
(Caracas),
AM!
21. Anteon surinamense n. sp.
FEMALE:
fully
winged;
length 3,20 mm;
testaceous,
with posterior margin
of metanotum and med i an area of propodeum brown; thickened; 8:
8:
7:
9;
head dull,
lower face; keel
antennae d i sta II y
antenna I segments in following proportions: occipital
POL
smooth,
of pronotum nearly
with a
carina complete;
around orbits;
pronotum shiny,
granulated,
=
8;
OL
with a
=
6:
5:
OOL
transversal long,
=
10;
=
OPL
impression;
4:
keels on
frons without median keel,
4;
twice as broad as
12:
track of weak
6;
=
TL
3:
7:
the with a
9;
posterior surface
shorter than scutum
(9:18),
wi th two dorsa I transverse lobes separated by a med I an furrow; posterior tubercles reaching tegulae; scutum shiny, smooth, without sculpture; notaul ices
indistinct,
faintly
visible near anterior margin of scutum;
scutellum and metanotum shiny, a strong transversal
keel
smooth,
without sculpture;
between dorsal
surface of propodeum reticulate rugose; without
longitudinal
sculpture,
keels,
and
propodeum with
posterior surface;
dorsal
posterior surface of propodeum
with median area shiny,
surrounded by areolae and,
smooth,
near the bottom,
by
without transversal
keels; fore-wing with two dark transversal bands, one on the basal cells and another beneath the pterostigma; radial vein with distal part shorter than proximal 9:
3,5:
4:
proximal nearly 37 1,1,2. MALE:
9:
part 23;
(4: 11);
fore tarsal
enlarged claw
prominence; lamellae;
(Fig.
segments 334)
segment 5 of front apex with a
in following proportions:
with one bristle
tarsus
(Fig.
group of nearly
12
334)
located on a
with 2 rows of
lamellae;
unknown
LOCUS TYPICUS:
holotype F!
TYPICAL MATERIAL:
in AM.
holotype F!
HOSTS: unknown. DISTRIBUTION: Neotropic region: Paramaribo,
AM!
in AM. SURINAME:
Botanic Gardens of
tibial
spurs
522
Mem. Amer.
Fig.
334 -
Chela of
Ent.
Inst.,
no.
1984
Anteon surinamense n. sp. (type)
ANTEON
AUSTRALIAN SPECIES OF
Fourty
37,
two species are known.
KEY TO THE SPECIES
FEMALES
Segment
4
of
front
tarsus at
Segment
4
of
front
tarsus
longer 2 3
than
most
0,5 as
slightly
surface of
propodeum with
Posterior
surface of
propodeum without
fully
Head at
reticulate
most
than,
segment as
•• 1..2
long as,
or
1 .•.••••....•••.••••••.••••••..•...•.••.. 10
segment
Posterior Head
long as
shorter
rugose.
reticulate
1.
rugose
two
longitudinal longitudinal
keels •••• 3 keels ••••• 4
giluwense n. sp.
only
on anterior half
of
frons
or behind ocell i, wi th a wide, punctate and smooth frontal area in front of the ocell i •.•••.• 2. chelogynoides (R.C.L. Perkins) 4
Head
smooth,
reticulate
punctate,
rugose;
without
posterior
sculpture among punctures,not
surface
of
propodeum almost
523
Anteoninae
smooth,
not
reticulate
rugose,
keel s •••••••••••••••••••••••• Head
reticulate rugose;
reticulate
5
posterior
or with
strong
Scutum reticulate
rugose,
or with
wi thout
surface
of
transversal strong
propodeum keels ••••••••••.• 5
or weak
sculpture or very
Propodeum wi th a
strong
surface;
longitudinal
transversal
scutum very
keels •••••••••••
Propodeum usually without dorsal
and posterior
transversal
keel
Scutum fully
weakly
faintly
irregular punctate
or
between dorsal sculptured by
transversal
occasionally
visible among
aulicum n. irregular
sides ..
keel
between
weak
the other keels . . . . . . . . 7
5.
the
with a
rugose ••••••••••.••.••••••••••
sculptured by weak
mainly visible on
and
irregular
reticulaticeps Dodd
4.
reticulate
• ..•...••..•......•..........
Antennal
keel
strongly
a distinct
surface;
strongly
Scutum shiny,
8
longitudinal
weakly granulated • • • • • • • • • • • • • • . . • • • . . • • • • . . • • . • • • • • • . • • • 8
posterior
7
short
keels •••••••••••••••••••••••••••.••.••••••..••.•• 6
Scutum smooth, very
weak and
myrmecophilum (R.C.L. Perkins)
rugose
longitudinal
6
with 3.
sp. longi tudinal
keels
sedlaceki n. sp.
6.
1 nearly as long as segments 2+3 . . . . . . . . . . . . . .
segment
. . . . . . . . . . . . . . . . . . . . . . . . • . . . . 8. australe n. sp. Antennal segment 1 nearly as long as segments 2+3+4 . . . . . . . . . . . 9
9
Fore-wing wi thout
dark
transversal
· . . . . . . . • . . . . . . . . . . . . . • . . . . .. Fore-wing with a dark
bands •••••••••.•••••••••••••
anxium n.
7.
transversal
• • • . . • • . • • . • • • . . • . . • • • . • • . •• 41. 10 Posterior surface of propodeum with Posterior
surface of
sp.
band •••••••••.•••••••••.. rufiscapum n. sp. two
propodeum without
longi tudinal longitudinal
keels ••. 11 keels . . . . 23
11 Fore-wing hyaline, without dark transversal bands . . . . . . . . . . . . 12 Fore-wi ng wi th 1-2 dark transversa I bands . . . . . . . . . . . . . . . . . . . . 20 12 Head granulated, not reticulate rugose or reticulate rugose only behind Head at 13 Head
ocell i . . . . . . . . . . .
least
partly
9.
reticulate
reticulate rugose
parvulum (R.C.L. Perkins) rugose
on anterior half
on of
frons and vertex . . . 13 frons,
punctate
on
vertex •••••••••••••••••••••••••••••••••••••.••••••••••••••••• 14 Head
fully
reticulate
rugose
or with
numerous
longitudinal
k eel s on f ron s • • • • . . • • . • • • • • • • • • • . • • • • • • • • • • • • • • . • • . • • • . • • • . • 1 5 14 Median area of posterior surface of propodeum smooth •••••••••••
• • • • • • • • • • • • • • • • • . • • • • • • • • •• Median
area of
posterior
· . . . . . . . . . . . . . . . . . . . . . . . . . .. 15 Scutum fully
reticulate
Scutum punctate,
not
16 Scutellum smooth; anterior margin Scutellum partly
orientale n. sp. propodeum rugose ••••••••.••
11. aculeatum n.
sp. rugose . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 16
reticulate
notaulices
of
10.
surface of
scutum . . .
reticulate
rugose ••••.•••••.•••••••••••• 17
invisible or faintly
12.
visible near
corruptum n. sp.
rugose;
notaulices
reaching
0,75 length of scutum •••••••••••••••••••••••••••• • • . • • . . • . . • . • • . . • . . • . . . . . . .. 1 3. miles n. sp.
approximately
524
Mem.
Amer.
17 Frons with numerous
Ent.
Inst.,
longitudinal
no.
37,
1984
keels . . . . . . . . . . . . . . . . . . . ..
• • • • • • • • • • • • • • • • • • • • . • • • • • •• 18. curraJoi n. sp. Frons fully reticulate r u g o s e . . . . . . . . . . . . . . . . . . . . . . . 18 Head with OPL much
shorter
· • • • • • • • • • • • • • . • • • • • • • . • . • •• Head with OPL 19 Segment
4
of
longer front
4
of
front
• • • • . . • • • • . . • • • . 20 Fore-wing wi th
tarsus
longer
wi th
sp.
tarsus
shorter
. ...•.•..• dark
than
1 ••••.••••••••••
than
segment
1 •••••••••••
superbum Dodd
40.
transversa I
band beneath
the
typicum (R.C.L. Perkins)
16.
2 dark
segment
destructor (R.C.L. Perk i ns)
14.
pt eros! i gma ••••• Fore-wing
caledonianum n.
15.
than TL . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 19
.. .. .. . . . . . . . . . . . . . . . . . . . . Segment
.18
than TL . . . . . . . . . . . . . . . . ..
transversal
bands,
one
on
the basal
cell s
21
and another beneath the pterost igma . . . . . . . . . . . . . . . . . . . . . . . . . . 21 Head with frons sculptured by strong longitudinal keels, not
22
Head fully reticulate rugose . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 22 Scutum reticulate rugose . . . . 19. firmum n. sp.
reticulate
rugose . . . . . . . . . . .
Scutum punctate,
not
reticulate
............................ 23 Fore-wing
wi th
Fore-wing 24 Segment
4
1-2 dark
of
tarsus nearly
front
tarsus
wi th
1 dark
leiosomum (R.C.L. Perkins)
transveral
wi thout
5 of
rugose •••••••••••••••••••••••••
20.
hyal ine, front
bougainvillei n. sp.
17.
dark
wi th basal
bands . . . . . . . . . . . . . . . . . . . . . 24
transversal as
part
bands . . . . . . . . . . . . 25
long as
long
segment
(Fig.
363);
1;
segment
fore-wing
Segment
4
of
front
band . . . . . . . . . . . . . . . . . . . . . . . . . . . .. 21. nitidum (R.C.L. Perkins) tarsus nearly twice as long as segment
segment
5
of
front
tarsus with basal
fore wing
transversa I
wi th
•...•.•.
2 dark
transversal
•.••.•.•••••••••••
25 Head
fully
reticulate
Head
fully
or almost
26 Head with numerous
rugose fully
22.
part
(Fig.
bands.......
1
365)
.. . . . . . . .
bouceki n. sp.
or with
longitudinal
granulated,
longitudinal
short
keels,
keels . . . . . . 26
not
reticulate
rugose.28
not
reticulate
rugose
. . . . . . . . . . . . . . . . . . . . . . . . . . . . 23. mundum n.
sp.
Head fully reticulate rugose . . . . . . . . . . . . . . . . . . . . . 27 Segment 1 of front tarsus shorter than segment 4 • •••••••••••••••••••••••••••• 28
42.
Segmen t
of
front
tarsus
Segmen t
5 of
front
tarsus almost
d i st i nct
longer
5 of front
29 Scutum wi thout
sculpture,
areas very
sp
segment
rect i I inear,
4 . . . . . . . . . . . . . 28 wi thout
a
25.
bellator n. sp. distinct
curved apex
(Fig.368)
nigricorne (R.C.L. Perk i ns)
shiny and
smooth,
except
for
the
weakly granulated •••••••••••••••••••••
• • • • . • • • • • • • • • • • • • • • • • • • • • •• Scutum fully
24.
tarsus with a
•••••••••••••••••••••••••••• lateral
than
curved apex. (Fig .367) •••••••••••••••••••••••• ••••••••••••••••••••
Segment
giraulti n.
• .27
26.
fiji anum n.
sp.
granulated ••••••••••••••••••••••••••
.. .. 29
Anteoninae
525
30 Species shiny; notaulices reaching approximately 0,5 length of scutum . . . . . . . . . . . . . . . . . . . . . . 27. dubium Fouts Species dull; notaul ices shorter, reaching approximately 0,25 length of scutum ..••••.•.••• 28. coriaceum (R.C.L. Perkins)
The females of A. sucklingi, bismarckense, gloriosum, vulsum, luctuosum, hageni, haustum, corallinum, tasmanianum, permirum and involutum are unknown.
MALES
2
3
4
5 6
7
8
Propodeum wi thout a strong transversal keel between dorsal and posteri or surface . . . . . . . . . . . . . . . . . . . . . .2 Propodeum wi th a strong transversal keel between dorsa I and posterior surface . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 Posterior surface of propodeum with two longitudina keels .•
· . . . . . . . . . . . . . . . . . . . . . . . . . . . 29. sucklingi n. sp. Posterior surface of propodeum wi thout longi tudinal keel s . . . . • 3 Head smooth, weakly punctate, wi thout sculpture among punctures • • • . . . . . . • • . . . • • • . . 3. myrmecophilum (R.C.L. Perkins) Head rugose, fully reticulate rugose • • . • . . • . • . . . . . • . . . . . • . . . · . . . . . . . . . . . . . . . . . . . . . . . . . .. 5. aulicum n. sp. Posterior surface of propodeum with two longitudinal keels . . . . . . . . . . . . ... 5 Post eri or surface of propodeum wi thout longitudinal keels . . . . 15 Gonoforceps wi th a long dorsal process (Figs 356, 357, 372) . . . 6 Gonoforceps wi thout long dorsal process (Figs 373, 374, 375) .. 7 Dorsa I process of gonoforceps very long and narrow, wi th apex curved (Figs 356, 357); scutum very weakly punctate . . . . . . . . . .. · . . . . . . • . •. . . ••••. . . . . . 15. caledonianum n. sp. Dorsal process of gonoforceps shorter and broad, wi th apex not curved (Fig.372) scutum very strongly punctate; punctures on scutum very broad . . . . . . . . . . 30. bismarckense n. sp. Head fully granulated . . . . . . 31. gloriosum n. sp. Head reticulate rugose, or punctate or with numerous longitudinal keels on frons . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8 Head fully or mostly reticulate rugose or with longitudinal keels on frons . . . . . . . . . . • . . . . . . . . . . . . . . • . . . . . • • • . . . . • • . . . • • . . . 9 Head
9
punctate,
at
most
with
areolae near
occipital
carina,
on malar space and on anterior half of frons . . . • . • . . . . . . . • • . • 13 Scutum almost fully reticulate rugose or with weak irregular keels . • . . . . . • • • • • . • • . . . • • • . 32. VUlSUIll n. sp. Scutum punctate and wi thout sculpture among punctures, not . . . 10 reticulate rugose . . . . . . . . . .
526
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
10 Median area of
posterior
surface of
propodeum smooth ...•••••• 11
Median area
posterior
surface of
propodeum rugose • • • • • • • • 12
11
Head
fully
of
reticulate
rugose ••••••••..•••••••••••••••••••
• •••••••••••••••••••••••••• 33. luctuosum n. sp. Head with frons sculptured by numerous longitudinal
keel •••.••.
hageni n. sp. haustum n. sp. 14. destructor (R.C.L. Perkins)
· ••. . . . . . . . . . . . . . . •. . . . . . .
34. 35.
12 Species mostly reddish •.•. Species mostly black •••••••
13 Notaulices reaching approximately 0,5 length of scutum •••••• •. • . ••. . . . . . •. . . •. . • . . •. 36. corallinum n. sp. Notaulices very reduced, at most reaching 0,3 length of scutum • •• 14 14 Median area
of
posterior
surface of
• • • • • • • • • • • • • • • • • • • • • • • • • •• Median area of
posterior
37.
propodeum smooth ••••.
tasmanianum n. sp.
surface of
• . . . • . . . • . . . • • . • • . • • • • • . • •.
1 0.
propodeum rugose. sp.
orientale n.
15 Head punctate, without sculpture among punctures •••• • . . . . . . . . • . • • • • • • • • • . . • • . . . 21. nitidum (R.C.L. Perkins) Head ret iculate rugose or with longi tudinal keels, or granu I at ed ••••••••.•••••••••...•••••••••••..••••••••••••••••• 16 16 Head granulated, sometimes also wi th a few irregular keels •••
· . . . . . . . . • . . . • . . . . . . . . . . . . . • 8.
australe n.
Head with numerous
keels
longitudinal
on
sp.
frons
or
reticulate
rugose •••.•.•••••••••••••••.••••••••••••••••••••••••••••••••• 17
17 Head with numerous longitudinal keels on frons ••••••••••••••••• · . . . . . . . . . . . . . . . . . . . . . . . . . . 38. permirum n. sp. Head
fully
or partly
18 Scutum punctate,
reticulate
rugose ••••••••••••••••••••••• 18
sculpture among punctures •••••• • . . . . . • • • . . . . . . . • • • • . • . . . . . 21. nitidum (R.C.L. Perkins)
Scutum fully
wi thout
weakly granulated.
39. involutum n.
· . . . . . . . . . . . . . . . . . . . . . . . . ..
sp.
The males of A. dubium, coriaceum, superbum, hjianum, nigricorne, reticulaticeps, bouceki, typicum, leiosomum, parvulum, chelogynoides, bougainvillei, anxium, corruptum, giraulti, giluwense, curradoi, sedlaceki, aculeaturn, miles, firmum, mundum, bellator and rufiscapum are unknown.
1. Anteon giluwense sp.
FEMALE:
fully
light;
length 2,50 mm;
winged;
antennae black,
with segment 1 partly
antennae distally thickened;
proportions:
8:
5:
8:
6:
6:
5:
5:
5:
black;
antennal 5:
mandibles testaceous;
testaceous; 8;
legs brown,
segments
head dull,
with
tarsi
in following fully
reticulate
Anteoninae
Fig.
335 -
527
Chela of Anteon giluwense n.
sp.
(type)
rugose; frons with a complete median keel; occipital carina complete; OL = 3; OOL = 4,5; OPL = 4; TL = 3; pronotum dull, reticulate
POL
= 5;
rugose and with transversal
keels;
pronotal
tubercles reaching
posterior surface of pronotum shorter than scutum scutellum and metanotum shiny, sculpture among punctures; 0,65 length of scutum; transversal
keel
smooth,
scutum,
weakly punctate,
without
notaul ices incomplete,
reaching approximately
propodeum reticulate rugose,
between dorsal
tegulae;
(3:11);
with a strong
and posterior surface;
posterior surface
with two longitudinal keels; median aerea rugose; fore-wing hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part (3:7); fore tarsal segments in following proportions: 8:
2:
2:
2:
12; enlarged claw
proximal prominence; nearly 12 bristles; MALE:
(Fig.335)
with one bristle located on a
segment 5 of front tarsus
(Fig.
apex with a group of 10 lamellae;
335)
with 2 rows of
tibial
spurs 1,1,2.
unknown
LOCUS TYPICUS:
Mt.
TYPICAL MATERIAL:
Giluwe (New Guinea) holotype F!
HOSTS: unknown DISTRIBUTION: Australian region:
in TW. NEW GUINEA:
Mt.
Giluwe,
TW!
2. Anteon chelogynoides (R.C.L. Perkins)
Prosanteon chelogynoides R.C.L. Perkins 1905: 66. Prosanteon melanostigmus R.C.L. Perkins 1906a: 493 (n. syn.); locus typicus Mittagong (Austral ia). Anteon chelogynoides (R.C.L. Perkins): R.C.L.Perkins 1907: 50. Anteon melanostigmus (R.C.L. Perkins): Kieffer 1914b: 170. Chelogynus cognatus (R.C.L. Perkins) partim: Kieffer 1914b: 193.
528
Fig.
Mem. Amer.
336 -
Chela of
En!.
Ins!.,
no.
37,
1984
Anteon chelogynoides (R.C.L. Perkins) from Pimel ia (Western Austral ia)
FEMALE:
fully
winged;
with teeth brown;
length 2,25-3,43 mm;
antennae fully
part of segment 2 yellow;
black;
mandibles yellow,
testaceous or brown with segment 1 and
legs yellow,
with only hind coxae brown
(in
chelogynoides) or with all coxae, femora and mid and hind tibiae brown (in melanostigmus); antennae distally thickened; antennal segments in
Anteoninae
following proportions: punctate, keels;
9:
4,5:
4,5:
4:
4:
529
4:
4:
4:
4:
with frons strongly punctate and rugose; OL = 3;
POL = 5;
OOL = 4;
OPL = 4,5;
numerous areolae and transversal than scutum
(5:11)
scutum smooth,
keels,
notaulices incomplete,
reaching
head shiny,
TL = 4,5;
with
irregular
pronotum with
with posterior surface shorter
and slightly shorter than
weakly punctate,
6;
vertex
anterior surface
without sculpture among approximately 0,3-0,5
(5:6)
punctures;
length of scutum;
scutellum and metanotum smooth, weakly punctatae, without sculpture among punctures; propodeum reticulate rugose, with a transversal keel between
dorsal
longitudinal rugose; radial
and posterior surface;
keels;
fore-wing
median hyal ine,
posterior surface with or shiny,
with basal
part
numerous bristles; proximal MALE:
2:
2:
3:
part
10;
enlarged claw tibial
(Fig.
(3:6);
two
lateral
bands;
areas
distal
fore tarsal
part of
segments
segments 5 of front
tarsus
part
lamellae and
longer than apical
prominence;
smooth;
without dark transversal
vein much shorter than proximal
in following proportions:6: 336)
area dull
336)
(6:4),
with 9
with one bristle
(Fig.
located on a
spurs 1,1,2.
unknown
LOCUS TYPICUS:
Bundaberg
TYP I CAL IvlATER I AL:
(Queensland,
lectotype F!
Australia)
Pro chelogynoides in B; lectotype F!
of
Pro melanostigmus in B. in Australia Tartessus sp. (?)(R.C.L. Perkins 1906c) for the var. chelogynoldes. of
HOSTS:
DISTRIBUITION: B!
Mittagong
Australian
region:
(New South Wales),
AUSTRALIA:
B!
Wanneroo
Bundaberg (32 mls.
( Queensland),
NE of Perth,
Western Australia), 8M! Pimelia (Penberton, Western Australia), OL! Kununurra (North Western Austra I i a), OL! Mt. Bundey Road (v i a Darw in, Northern Territory),
UQ!
3. Anteon myrmecophilum (R.C.L. Perkins)
Paranteon myrmecophllus R. C. L. Perk i ns 1905: 67 Anteon myrmecophilus (R.C.L. Perkins): Kieffer 1907b: 27
FEMALE:
fully
winged;
with metanotum, not distally 7:
8:
8:
8:
length 2,74-2,93 mm;
propodeum,
thickened; 7:
7:
7:
antenna I segments
8;
yellow-reddish,
petiole and abdomen
head convex,
sometimes
reddish-brown;
antennae
in following proportions:
strongly punctate,
14:
6:
without sculpture
among punctures; frons without median and lateral keels; POL = 5; OL = 3; OOL = 7; OPL = 6; TL = 5; pronotum short, faintly visible in dorsal view;
scutum convex,
dull,
weakly punctate,
without sculpture among
punctures, with weak longitudinal keels on anterior half; notaul ices distinct, incomplete, reaching approximately 0,3 length of scutum; scutellum shiny,
smooth,
very convex,
without sculpture,with a
weak
530
Mem.
Fig.
337 -
Chela of
Amer.
Ent.
Inst.,
metanotum shiny,
propodeum with dorsal surface by a
weak
1984
(Queensland)
smooth,
surface dull,
transversal
without sculpture,
rugose,
keel
posterior margin of propodeum; lateral
37,
Anteon myrmecophilum (R.C.l. Perkins) from Bundaberg
median furrow;
no.
very convex;
separated from posterior
semicircle shaped,
not reaching
posterior surface smooth and shiny;
areas rugose and dull; dorsal surface shorter than breadth of fore-wing with a dark transversal band beneath the
metanotumj
pterostigma; part
distal
(5:7 or 4:5):
(5:2,2); tarsus basal
part of radial segment
fore tarsal (Fig.
337)
segment 3 produced
long as segment 4;
located on a MALE:
fully
tarsus much into hook;
proximal winged;
twice as
segment 5 of front
prominence;
tibial
length 2,56-2,74 mm;
lamellae at apex;
long as apical
enlarged claw
(
proximal
longer than segment 4
with numerous bristles and some
part of segment 5 nearly
nearly as
vein slightly shorter than
1 of front
Fig.
337)
part;
distal
part
with one bristle
spurs 1,1,2. brown-black;
antennae brown;
tibiae and tarsi light; tegulae testaceous; mandibles brown, with teeth light; antenna I segments in following proportions: 9: 4,5: 5: 6: 6: 6: 5: 5:
5:
6;
head shiny,
smooth,
convex,
without sculpture among punctures; Tl = 3;
scutum sh i ny,
anterior third;
smooth,
and sparse punctures,
Ol = 3;
OOl = 6;
OPl = 2,5;
with strong and sparse punctures on
posterior surface without punctures or weakly punctate,
without sculpture among punctures; reaching
with strong
POL = 6;
approximately 0,5
notaul ices distinct,
length of scutum;
shiny,
smooth,
without sculpture,
shiny,
smooth,
not reticulate rugose;
convex;
semicircle shaped separating dorsal the posterior marg i n of propodeum;
incomplete,
scutellum and metanotum
mesopleura and metapleura
propodeum with a
transversal
and posterior surface, dorsa I surface rugose,
keel
not reaching shorter than
531
Anteoninae
Fig.
338 -
Male genitalia of Anteon myrmecophilum Bundaberg
breadth of metanotum; near dorsal surface;
(Queensland):
right
(f'.C.L.
posterior surface almost smooth, lateral
without dark transversal
areas reticulate rugose;
bands,
with distal
Perkins)
from
half removed.
sl ightly rugose
fore-wing
part of radial
hyal ine,
vein shorter
than proximal part (4:7); gonoforceps (Fig. 338) without inner distal process and without dorsal proximal membranous process, narrow in dorsal
view;
tibial
LOCUS TYP I CUS:
TYP I CAL MATER I AL: HOSTS:
(Austra I i a)
lectotype F!
and 2 para I ectotypes FF!
in Australia Jpo conferta Kirk.
DISTRIBUTlm,: Sandhi II
spurs 1,1,2.
Bundaberg
Australian region:
(Queensland),
(R.C.L.
AUSTRALIA:
in B.
Perkins 1906c).
Bundaberg
OL!
4. Anteon reticuiaticeps Dodd
(Oueensland),
B!
532
Mem. Amer.
Fig.
339 -
Chela of
Ent.
Ins!.,
no.
37,
1984
Anteon reticulaticeps Dodd from Finisterre Mts. (New Guinea)
Anteon reticuiaticeps Dodd 1914a: 131.
FEMALE: fully winged; length 2,12-2,18 mm; black; mandibles testaceous, with teeth brown; antennae brown, with segments 1 or 1-2 testaceous; tegulae brown-testaceous; legs brown; sometimes tarsi testaceous; antennae distally thickened; antenna I segments in following proportions: 10: 5:
5:
4:
longitudinal 3;
TL
~
3;
4:
fully
between dorsal
vein
with one bristle
with basal
part much
frons with a
OL
~
3;
smooth,
transversal
keel
7:
transversal
proximal 1,5:
a
1,5:
part 1,5:
proximal
bands;
(4:12); 6;
fore
enlarged
prominence;
with some bristles and 2 apical
longer than distal
part;
tibial
spurs
1,1,2. MALE:
unknown
LOCUS TYP ICUS:
Nelson
TYPICAL MATERIAL:
(Austral ia)
holotype F!
HOSTS: unknown DISTRIBUTlm,: Australian
region:
~
keels and
with a
without dark
located on 339)
OOL
posterior surface without
much shorter than
(Fig.
6;
irregular longitudinal
in following proportions: tarsus
~
POL
scutellum and metanotum shiny,
fore-wing hyal ine,
segments
segment 5 of front
reticulate rugose; keels;
pronotum very short and reduced;
and posterior surface;
keels;
339)
lateral
propodeum reticulate rugose,
tarsal
lamellae,
head fully
sculptured by
part of radial (Fig.
6;
without
carina complete;
distal claw
4:
notaulices invisible;
without sculpture; longitudinal
4: 4:
occipital
scutum dull, granulated;
4:
median keel,
in AD. NEW GUINEA:
Finisterre IMs.
(Madang
Anteoninae
Fig.
Dist.),
340 -
BM!
Wau
Chimbu div., AUSTRALIA:
Chela of Anteon aulicum n.
(Morobe Distr.,
NE New Guinea),
Nelson
533
sp.
from Taui
NE New Guinea),
B!
(N Queensland),
Mt.
Kaindi
B!
(New Guinea)
Daulo Pass
(NE New Guinea),
AD!
5. Anteon aulicum n. sp.
(AsaroOL!
534
""em.
FEMALE:
fully
winged;
antennae testaceous, with
Amer.
Ent.
Inst.,
length 2,75 mm;
shiny,
in following proportions:
fully
keels;
reticulate rugose;
occipital
black;
16:
5:
POL ~ 5;
surface of pronotum shorter than scutum tegulae;
reaching
approximately 0,3
keel
rugose;
between
bristle 340)
8:
2:
2,5: 4:
located on
bands;
(3:10); 13;
a proximal
6:
length 2,18-2,31
distal
half light;
light;
antennae not distally
proportions:
antennae black;
TL ~ 5;
keels;
posterior
tubercles
notaul ices
incomplete,
without a strong
transversal
posterior surface without
transversely striate; distal
fore-wing
part of radial
segments
12: 6:
7:
8:
mm;
8:
8:
8:
8:
tarsus with basal (Fig.
340)
black;
12;
head dull,
part
with one
tarsus
(Fig.
tothe apex;
mandibles black,
with fore
with
tibiae and
antenna I segments
8:
vein
in following
segment 5 of front
legs brown,
thickened;
head
and 2 lateral
with 2 rows of nearly 16 lamellae extending continuously
tibial spurs 1,1,2. MALE: fully winged;
antennal 8;
OPL ~ 4;
enlarged claw
prominence;
5:
scutellum and metanotum
fore tarsal
(10:3);
5:
pronotal
segment 5 of front
part
legs testaceous,
keel
OOL ~ 6;
(7:18);
reticulate rugose and
longer than apical
5:
median
length of scutum;
part
5:
transversal
and posterior surface;
proximal
proportions:
5:
propodeum reticulate rugose,
keels,
mandibles testaceous;
reticulate rugose;
without dark transversal
shorter than much
scutum shiny,
dorsal
longitudinal hyal ine,
7:
reticulate rugose and with
reaching shiny,
1984
antennae distally thickened;
frons with a
carina complete;
pronotum dull,
37,
with segments 2-3 and 9-10 brown;
hind coxae proximally brown;
segments
no.
tarsi
in following
fully
reticulate
rugose; frons with a com,,,:ete median keel; occipital carina complete; ~ 6; OL ~ 3; OOL ~ 6; OPL ~ 2; TL ~ 2; scutum dull, with anterior surface fully by
reticulate rugose and with posterior third weakly sculptured
irregular striae;
sh i ny,
weak I y
notaul ices invisible;
punctate,
scutellum and metanotum smooth,
without scu I pture among punctures;
reticulate rugose,
without a strong
posterior surface;
posterior surface without
hyaline,
without dark transversal
shorter than proximal inner pointed process;
part
(3:8);
tibial
transversal bands;
keel
longitudinal
distal
gonoforceps
in B;
HOSTS:
3 paratypes MM!
propodeum
between
dorsal
keels;
part of radial
(Fig.
341)
and
fore-wing vein
without distal
spurs 1,1,2.
LOCUS TYP ICUS: Mt. Gi luwe (New Guinea) TYPICAL MATERIAL: holotype F! and 4 paratypes MM! F!
POL
in TW;
1 paratype
in OL.
unknown
DISTRIBUTION: Guinea),
B!
OL!
Gap
Tari
Australian
Mt. Giluwe, (Mt.
region: OL!
Hagen),
TW!
NEW GUINEA: Daulo Pass,
Taui OL!
(Mt.
TW!
Mt.
Kaindi,
TW!
6. Anteon sedlaceki n. sp.
FEMALE:
fully
winged;
length 3 mm;
black;
NE
Wilhelm,
mandibles testaceous;
New TW!
Anteoninae
Fig.
341
535
Male genitalia of Anteol1 aulicum n.
-
sp.
from Mt.
Giluwe
(New
Guinea)
with segment 1 proximally
antennae brown, antennae distally
thickened;
antennal
testaceous;
segments
legs testaceous;
in following proportions:
16: 6: 6: 6: 5: 5: 5: 5: 5: 9; head fully reticulate rugose, with a complete median keel;
4; OOl
=
5; OPl
=
4; Tl
=
occipital
carina complete;
5; pronotum strongly rugose,
dull; frons POL = 5,5; Ol = with numerous
transversal keels, with posterior surface shorter than scutum (5:18); pronotal tubercles reaching tegulae; scutum shiny, punctate, without sculpture among punctures, keels mainly on incomplete,
lateral
smooth,
with a
very
weak
faintly
visible among
length of scutum;
without sculpture;
transversal
irregular longitudinal
and near posterior margin;
approximately 0,5
reaching
metanotum shiny,
with numerous weak
regions
keel
between
the other keels;
notaul ices
scutellum and
propodeum reticulate rugose, dorsal
and posterior surface,
posterior surface of propodeum
without longitudinal keels; fore-wing hyal ine, without dark transversal bands; distal part of radial vein shorter than proximal part (4:10); fore tarsal
segments
(Fig.
342)
front
tarsus
in following proportions:
with
1 bristle
(Fig.
342)
located on
lOCUS TYPICUS:
(New Guinea)
HOSTS:
4:
13;
enlarged claw
prominence;
segment 5 of
14 lamellae extending
in TW.
unknown
DISTRIBUTIOi~:
TW!
Gap
holotype F!
3:
spurs 1,1,2.
tibial
Tari
2:
with 2 rows of nearly
continuously to the apex; MALE: unknown TYPICAL MATC:RIAl:
9:
a proximal
Australian
region:
NEW GUINEA:
Tari
Gap
(Mt.
Hagen),
536
Mem. Amer. Ent.
Inst.,
no. 37,
1984
Fig. 342 - Chela of Anteon sedlaceki n. sp.
NOTES:
(type)
the species is named in honor of Mr. J. Sedlacek.
7. Anteon anxium n. sp.
Anteoninae
Fig.
FEMALE:
fully
with proximal testaceous
343 -
winged;
5:
4:
3:
rugose,
3:
thickened;
4,5: 4,5: OL ;
legs black,
antennal
4:
median keel,
POL; 6;
black;
4:
7;
without
2,5;
segments
with
lateral
OOL ;
5;
fully
keels;
OPL ;
smooth,
tegulae;
scutum,
approximately 0,3
or
testaceous;
5;
reticulate rugose;
occipital TL ;
carina
4;
pronotum dull,
(3:16);
pronotal
scutellum and metanotum shiny,
without sculpture or weakly punctate;
reaching
tarsi
in following proportions:
head shiny,
with posterior surface shorter than scutum;
tubercles reaching
mandibles testaceous,
antennae brown with segment 1 testaceous,
with segments 2-3 brown;
frons with a complete;
Anteon anxium n. sp. (type)
length 2,00-2,50 mm;
half black;
antennae distally 14:
Chela of
537
length of scutum;
notaul ices
incomplete,
propodeum reticulate rugose,
wi th a strong transversa I keel between dorsa I and posterior surface; posterior surface without longitudinal keels, rugose, with transversal keels;
fore-wing
hyaline,
without dark transversal
bands;
distal
part of
538
Mem.
radial
Amer.
Ent.
Ins!.,
vein much shorter than proximal
in following proportions: basal
part
6:
2:
longer than apical
one bristle
located on
2:
3:
part
a proximal
tibial
MALE: unknown LOCUS TYPICUS:
(2:7);
fore tarsal
(8:2);
enlarged
claw
segments
tarsus with
(Fig.
343)
segment 5 of front
with
tarsus
lamellae extending continuously
to
spurs 1,1,2. Wamena
TYPICAL MATERIAL:
(New Guinea)
holotype F!
in B;
HOSTS: unknown DISTRIBUTION: Australian region: Numbu
1984
segments 5 of front
prominence;
343) with 2 rows of nearly 11
(Fig.
the apex;
part
10;
37,
no.
(Upper Chimbu V.,
NE
1 para type F!
NEW GUINEA:
New Guinea),
in OL.
Wamena
(Irian
Barat),
B!
OL!
8. Anteon australe n. sp.
FEMALE: fully
fully
winged;
coxae partly brown; partly brown; proportions:
occasionally
5:
6:
median
OOL = 6;
5:
5:
5:
TL = 3;
mandibles brown;
legs testaceous,
hind coxae, thickened;
5:
longitudinal
OPL = 3;
mm;
1 testaceous;
antennae distally
10:
frons with a = 5;
length 2,37-2,81
brown or with segment
5:
5:
keel,
hind femora and
antennal
7;
segments
head dull,
without
occipital
antennae
with hind hind tibiae in following
reticulate rugose;
lateral
keels;
carina complete;
POL = 9;
OL
pronotum dull,
reticulate rugose, with transversal keels, almost smooth on posterior surface; posterior surface shorter than anterior surface and shorter than scutum (4:16); scutum shiny, smooth, without sculpture or very weakly granulated; notaulices incomplete, reaching anterior third of scutum; scutellum and metanotum shiny, smooth, without sculpture; propodeum reticulate rugose,
with a
posterior surface;
posterior surface without
rugose;
with a
fore-wing
distal
part of radial
tarsal
segments
tarsus with basal (Fig.
MALE:
tibial
fully
tarsus
7,5:
located on 344)
with
keels,
and
reticulate
band beneath the pterostigma; proximal 2:
longer than apical
(Fig.
between dorsal
15
3:
part
3:
part
15;
(3:8);
(10:5);
a proximal
fore
segment 5 of enlarged
prominence;
lamellae and numerous
spurs 1,1,2.
winged;
antennae brown;
part
with one bristle
segment 5 of front
keel
longitudinal
dark transversal
in following proportions:
front
bristles;
transversal
vein much shorter than
claw
344)
strong
length 1,62-1,81
legs brown,
mm;
black;
mandibles testaceous;
with fore tibiae and tarsi
light;
antennae
not distally thickened; antennal segments in following proportions: 6: 4: 5: 5: 5: 5: 5: 5: 5: 7; head dull, granulated, sometimes with irregular keels on ocellar region;
frons with a short median
front of the anterior ocellus; OOL = 5;
OPL = 2;
TL = 3;
occipital
scutum,
keel
carina complete;
visible only POL
=
5;
OL
in
=
scutellum and metanotum shiny,
smooth, weak I y punctate, wi thout scu I pture among punctures; notau I ices incomplete, reaching only 0,25 length of scutum; ;:->ropodeum reticulate
3;
Anteoninae
Fig.
344 -
539
Chela of Anteon australe n.
rugose,
with a strong transversal
surface;
posterior surface without
keel
sp.
between dorsal
longitudinal
keels;
(type)
and posterior fore-wing
hyaline,
without dark transversal bands; distal part of radial vein shorter than proximal part ( 2:5); gonoforceps (Fig. 345) without distal inner pointed process; tibial spurs 1,1,2. LOCUS TYP ICUS: Dongarra (Western Austral ia) TYPICAL MATERIAL: F,
2 MM)!
HOSTS:
in TW;
holotype F! 3 paratypes
and (1
F,
1 paratype F! 2 MM)!
in BM;
3 paratypes
in OL.
unknown
DISTRIBUTION: Austral ia), (Tasmania),
Australian
BM! TW!
Yuendumu Waldheim
region:
AUSTRALIA:
Dongarra
(Northern Territory), (Tasmania),
TW!
OL!
TW!
(Western Roseberry
OL!
9. Anteon parvulum (R.C.L. Perkins)
nee Anteon pallidieornis Ashmead 1893: 97. Neoehelogynus parvulus R.C.L. Perkins 1905: 65. Neoehelogynus pallidieornis R.C.L. Perkins 1905: 65 (n. syn.); locus typieus Bundaberg (Austra I i a) nec Anteon parvulus Kieffer in Kieffer et Marshall 1905: 193. Anteon laetieornis Kieffer 1907b: 26; new name.
(1
540
Mem.
Fig.
345 -
Amer.
Ent.
Inst.,
no.
37,
1984
Male genitalia of Anteon australe n. sp. (Austral ia): left half removed.
from Yuendumu
Anteon parvulus (R.C.L. Perkins): Kieffer 1907b: 27. Chelogynus pallidicornis (R.C.L. Perkins): Kieffer 1914b: 191. Chelogynus parvulum (R.C.L. Perkins): Kieffer 1914b: 192.
FEMALE:
fully
winged;
with teeth brown; brown with
length
with segments
1-2 testaceous;
hind femora brown;
sides)
of body;
long as broad;
1,83-2,38 mm;
black;
antennae yellow or slightly body shiny,
head granulated;
tegulae and hairy
reaching
occasionally
legs yellow,
sometimes
(mainly on head and on
pronotum granulated,
scutum without sculpture or very
notaulices incomplete,
mandibles yellow,
darkened;
nearly twice as
weakly granulated;
approximately 0,25
length of scutum;
Anteoninae
Fig.
346 -
Chela of
541
Anteon parvulum (R.C.L. Perkins) from Mapleton (Australia)
scutell um and metanotum smooth;
propodeum rugose,
ma i n I y
near
metanotum, with a transversal keel between dorsal and posterior surface; posterior surface with 2 longitudinal keels; median area rugose; forewing hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal proportions:
4,5:
1,5: 2:
part;
3,5:
bristle located on a proximal 346)
with basal
fore tarsal
prominence;
(Fig.
346)
(Queensland,
lectotype F!
with one
segment 5 of front part
(3,5:5,5),
apex with a group of 10 lamellae;
Cairns
TYPICAL MATERIAL:
segments in following
enlarged claw
part shorter than apical
nearly 25 lamellae; MALE: unknown LOCUS TYPICUS:
9;
tarsus
(Fig.
with two rows of
tibial
spurs 1,1,2.
Australia)
(designated by Olmi
1982) of
N. parvulus
in B; lectotype F! (designated by Olmi 1982) of N. pallidicornis in B. HOSTS: in Australia Euleimonios sp. (R.C.L. Perkins 1906c) for parvulus;
Deltocephalus sp. luteotactella. DISTRIBUTION: Bundaberg
Australian
(Queensland),
Queensland) Roseberry
(R.C.L.
,
UQ !
Perkins 1906c) region: B!
TW
AUSTRALIA:
Eurimbula Ck.
Mapleton,
(T asman i a),
for
BR
! Mt.
pallidicornis; N. Cairns
(Queensland),
(Via Round Hi II
Hotham
(Victoria),
!
10. Anteon orientale n. sp.
Head, OL!
TW!
B!
542
Mem.
Fig.
FEMALE:
fully
testaceous;
347 -
Amer.
En!.
no.
37,
1984
Anteon orientale n. sp.
Chela of
winged;
Ins!.,
length 3,18 mm;
antennae testaceous,
black;
with dorsal
(type)
mandibles and
legs
side of segments 4-10 brown;
antennae distally thickened; antennal segments in following proportions: 14: 6: 8: 6: 6: 6: 5: 6: 7: 7; head shiny, strongly punctate, without sculpture among punctures, with anterior half of frons reticulate rugose; frons with a
longitudinal
orbits directed complete;
towards ~
POL
8;
median
~
Ol
keel
and with 2
inner side of antennal 4;
strongly
punctate,
pronotal
tubercles reaching
OOl
~
8;
OPl
~
5;
shiny,
smooth,
punctate,
tegulae;
scutum shiny,
only partly distal
transversal part
6:
longer than
apical
located on a with
2:
bands;
(3,5; 2,5:
10);
4:
part
proximal
1 row of 9
carina
pronotum shiny,
smooth,
(6:21);
strongly
notaul ices incomplete, scutellum and metanotum
without scul;:>ture among punctures;
without dark than proximal
5;
propodeum
with a strong transversal keel between dorsal and posterior surface with 2 longitudinal keels; median
area mostly smooth and shining,
proportions:
~
Tl
keels around
occipital
with posterior surface shorter than scutum
punctate, without sculpture among punctures; reaching approximately 0,3 length of scutum; reticulate rugose, posterior surface;
lateral
sockets;
15;
(9:6);
fore tarsal
segments
segment 5 of front enlarged claw
prominence;
lamellae;
rugose;
fore-wing
part of radial
in following
tarsiusi
(Fig.
347)
segment 5 of front
apex with a
group of 4
hyaline,
vein much shorter with basal
part
with one bristle
tarsus
lamellae;
(Fig. tibial
347) spurs
1,1,2. MALES:
fully
testaceous;
winged;
samet i mes
length 1,56-3,75 mm; legs brown;
black;
mandibles and
legs
antennae brown or testaceous-darkened;
Anteoninae
Fig.
Male genitalia of Anteon orientale n.
348 -
(Queensland):
antennae not distally thickened; proportion: 7: punctate,
543
4:
5:
5:
4,5: 5:
antennal 5:
sp.
from Tambourine Mts.
left half removed.
5:
5:
segments 8;
in following
head shiny,
without sculpture among punctures;
occipital
and malar space granulated and reticulate rugose;
smooth, region,
occipital
strongly
temples
carina
complete; frons with a median keel and without lateral keels; POL ~ 6; OL ~ 4; OOL ~ 4; OPL ~ 2; TL ~ 2; scutum, scutellum and metanotum shiny,
smooth,
weakly punctate,
notaulices incomplete, of scutum; between
longitudinal
and posterior surface;
keels;
hyaline,
median area dull, part
membranous process,
process; tibial spurs 1, 1, 2. LOCUS TYP ICUS: Tambourine Mts. TYPICAL MATERIAL: TW;
in BR;
holotype F!
1 paratype M!
2 paratypes MM!
HOSTS:
with
approximately 0,25
length
a strong transversal
keel
posterior surface with 2 nearly as rugose as
without dark transversal
vein much shorter than proximal without proximal
M!
reaching
propodeum reticulate rugose,
dorsal
fore-wing
without sculpture among punctures;
very reduced,
(3:7);
bands;
distal
gonoforceps
without distal
lateral
areas;
part of radial (Fig.
348)
inner pointed
(Austral ia) and 2 paratypes MM!
in AD;
1 para type F!
in UQ;
in BM;
1 paratype
2 para type I,jf;i!
in
in OL.
unknown
DISTRIBUTION: Queensland), Round Hi II
Australian region:
OL!
Head,
Gordonva!e
AUSTRALIA:
(NE Queensland),
Queensland),
UQ!
Loxton
(32 mls. N of Perth, Western Australia), TW! Mt. Giluwe, T'.\'!
Tambourine Mts. SR!
(South Austral ia),
BM!
(SE
Eurimbula Ck.
NEW GUINEA:
AD!
(via Yanchep
Baiyer R.,
OL!
544
Mem. Amer. En!.
Fig. 349 -
Inst.,
no.
37,
1984
Chela of Anteon aculeatum n. sp.
(type)
Anteoninae
545
11. An rcon aculeatum n. sp.
FEMALE:
fully
black;
antennae distally thickened;
following proportions: with
length 1,87 mm;
winged;
legs testaceous;
mandibles,
antennae and
antenna I segments
in
8: 4: 5: 4: 6: 5: 4,5: 4,5: 4,5: 7; head shiny,
vertex and posterior half of frons smooth and
without sculpture;
anterior half of frons reticulate rugose; ocellar region, region behind ocelli and regions on sides of ocelli with irregular keels; frons with a median kell; occipital carina complete; POL = 4; OL = 3; OOL = 3; OPL 2; TL = 3; ;:>ronotum strong I y rugose, with posterior surface shorter than scutum (5:10); pronotal tubercles reaching tegulae; scutum, scutellum and metanotum shiny, punctures; scutum;
smooth,
notaul ices
propodeum
between dorsal
weakly
punctate,
incomplete,
reticulate rugose,
with a strong
and posterior surface;
longitudinal keels; transversal bands;
without sculpture among
I ittle visible near anterior margin of transversal
posterior surface with
keel
two
median area rugose; fore-wing hyaline, without dark distal part of radial vein much shorter than proximal
(2:7); fore tarsal segments in following proportions: 3,5: 2,5: 2: 5: 12; enlarged claw (Fig. 349) with one bristle located on a proximal prominence; segment 5 of front tarsus (Fig. 349) with 2 rows of nearly 23 lamellae; apex with a group of nearly 19 lamellae; tibial spurs 1,1,2.
part
MALE:
unknown
LOCUS TYP ICUS:
Mt.
TYPICAL MATERIAL: HOSTS:
Glorious
(Austral ia)
holotype F!
in TW.
unknown
DISTRIBUTION: TWI
Australian
region:
AUSTRALIA:
Mt.
Glorious
(Queensland),
12. Ilnteon corruptum n. sp.
Ft:MALE:
fully
with proximal testaceous; proportions: keel
with coxae,
antennae distally
15: 6:
8:
9:
granulated on
and with 2
6:
7:
internal
lateral
sockets;
occipital
TL = 5;
pronotum shiny,
region
antennae brown,
legs testaceous,
partly brown; rugose,
length 2,81-3,00 mm;
winged;
half brown;
wi th a strong
9;
fully
head shiny,
reticulate
=
6;
OL
=
reticulate rugose,
and without sculpture;
reticulate rugose, shiny,
POL
(5:18); shiny;
pronotal
fully
frons with a
median
towards antenna I 3;
OOL
=
8;
OPL
=
4,5;
except for a short posterior surface of
tubercles reaching
notaulices invisible;
without sculpture;
transversa I keel
tibiae in following
surface of areolae;
carina complete;
near scutum smooth
metanotum smooth,
7:
side of segment
antenna I segments
keels near orbits directed
pronotum shorter than scutum scutum fully
7:
mandibles testaceous,
ventral
clubs of femora and
thickened; 7:
black; with
tegulae;
scutellum and
propodeum reticulate rugose,
between dorsa I and posterior surface;
546
Mem.
Fig.
350 -
Chela of
posterior surface with 2 as
lateral
areas;
part of radial
tarsal
segments
front
tarsus
En!.
Ins!.,
hyal ine,
keels; without
37,
1984
median
area nearly as rugose
dark transversal
vein much shorter than proximal
in following proportions:
with basal (Fig.
no.
Anteon corruptum n. sp. (type)
longitudinal
fore-wing
distal
enlarged claw
Amer.
350)
6:
3:
3:
9:
part much shorter than apical with one bristle
part 22;
bands;
(4:12);
fore
segment 5 of
part
(8:14);
located on a proximal
Anteoninae
prominence;
segment 5 of front
tarsus
547
(Fig.
350)
lamellae extending continuously to the apex; MALE:
spurs 1,1,2.
unknown
LOCUS TYPICUS:
Murray Pass
TYPICAL MATERIAL: F!
with 2 rows of nearly 31
tibial
(New Guinea)
holotype F!
in B;
1 paratype F!
in OL;
1 paratype
in TW.
HOSTS: unknown DISTRIBUTION: Australian region: NEW GUINEA: Murray Pass (W slope, New Guinea), B! Mt. Kaindi (NE New Guinea), OL! L. Sirunki, TW!
SE
13. Anteon miles n. sp.
FEMALE:
fully
winged;
length 2,87-3,12 mm;
anterior margin of clypeus reddish;
black;
mandibles testaceous;
antennae testaceous or brown,
with
segment 1 testaceous; legs testaceous, with hind coxae partly brown; antennae distally thickened; antenna I segments in following proportions: 10: 5:
10:
9:
9:
9:
with a median keel, = 5;
OL = 3;
rugose,
8:
8:
8:
without
OOL = 6;
10;
head dull,
lateral
OPL = 5;
keels;
TL = 5;
fully
reticulate rugose;
occipital carina complete;
pronotum dull,
except for a short region near scutum which
surface of pronotum shorter than scutum reaching tegulae; complete,
(5:16);
scutum fully reticulate rugose,
is smooth;
dull;
POL
reticulate
pronotal
reaching approximately 0,75 length of scutum;
frons
posterior
tubercles
notaul ices almost scutellum with
strong areolae on central region, with weak areolae laterally; metanotum reticulate rugose; propodeum reticulate rugose, with a strong transversal keel
between dorsal
longitudinal transversal part 17;
(4:13);
keels;
and posterior surface; median area rugose;
bands;
distal
fore tarsal
segment 5 of front
(7:10);
enlarged claw
prominence;
part of radial
(Fig.
351)
Angoram
DISTRIBUTION: Guinea),
B!
Kainde,
3:
part
with one bristle located on a proximal (Fig.
351) tibial
with 2 rows of 23 spurs 1,1,2.
(New Guinea)
holotype F!
Australian region:
Mt.
2,5:
part shorter than apical
lamellae; apex with a group of 6 lamellae; MALE: unknown TYPICAL MATERIAL: HOSTS: unknown
without dark
vein much shorter than proximal
segments in following proportions: 7,5: tarsus with basal
segment 5 of front tarsus
LOCUS TYPICUS:
posterior surface with 2
fore-wing hyal ine,
in B;
1 paratype F!
NEW GUINEA:
in TW.
Angoram
TW!
14. Anteon destructor (R.C.L. Perkins)
Neochelogynus destructor R.C.L. Perkins 1905: 64.
(Sepik,
NE New
7:
548
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
\
Fig.
351
-
Chela of
Anteon miles n. sp. (type)
Neochelogynus cognatus R.C.L. Perkins 1905: 64 (n. syn.); locus typicus Bundaberg (Austra I i a) •
549
Anteoninae
Neochelogynus ignotus R.C.L. Perkins 1906a: 493 (n. syn.); locus typicus Sydney (Austra I i a) • Anteon cognatus (R.C.L. Perkins): Kieffer 1907b: 25. An teon destructor (R. C. L. Perk i ns): Kieffer 1907b: 25 Chelogynus ignotus (R.C.L. Perkins): Kieffer 1914b: 192. Chelogynus destructor (R.C.L. Perkins): Kieffer 1914b: 192. Chelogynus cognatus (R.C.L. Perkins): Kieffer 1914b: 193.
FEMALE:
fully
winged;
with teeth brown;
length 2,50-3,11
sometimes only
mm;
distal
black;
mandibles yellow,
half of mandibles yellow
(destructor); antennae brown with segment 1 testaceous (ignotus); (cognatus) or segments 1-3 testaceous (destructor); tegulae testaceous; legs testaceous, sometimes with coxae
occasionally segments 1-2 testaceous brown; 5:
5:
keel
antennal 6;
segments in following proportions:
head fully
and
reticulate rugose;
with 2 l.:lteral
sockets;
POL = 7;
complete;
frons
11: 5:
with a
4:
3:
4:
longitudinal
4:
5:
median
keels near orbits directed towards antenna I
OL = 5;
OOL = 7;
OPL = 5;
pronotum re-ticulate rugose,
short,
posterior surface shorter than scutum
TL = 4;
occipital
carina
much broader than
(9:17);
pronotal
long,
with
tubercles reaching
tegulae; scutum shiny, smooth, weakly punctate, without sculpture among punctures; notaul ices very short, visible only near anterior margin of scutum;
scutellum and metanotum shiny,
among punctures; between
dor5al
longitudinal hyaline,
and posterior surface;
keels;
without dark transversal proximal
proportions:
6:
2:
4:
enlarged claw
prominence;
part 8:
20;
(Fig.
bands;
(4:10):
lateral
distal
segment 5 of front
transversa I keel
fore tarsal
2
areas;
fore-wing
part of radial
segments
segment 2 of front
352)
without sculpture
wi th a
posterior surface with
median area as rugose as
shorter than hook;
punctate,
propodeum ret i cu I ate rugose,
vein much
in following
tarsus produced
into a
with one bristle located on a proximal tarsus
(Fig.
352)
with 2 rows of 18
lamellae; apex with a group of 13 lamellae; tibial spurs 1,1,2. MALE: fully winged; length 1,43-2,68 mm; black; mandibles partly black or with
teeth brown;
testaceous; brown;
antennae brown or testaceous-dark,
tegulae testaceous;
rarely fully brown;
legs testaceous,
head reticulate rugose
reticulate rugose that head of female); keel
and
sockets;
with 2 lateral POL = 6;
complete; punctures; scutum;
OOL = 6;
smooth,
notaulices incomplete,
propodeum reticulate rugose, area as rugose as transversa I bands; part
OPL = 2;
weakly
(3:8);
longitudinal
TL = 3;
punctate,
reaching 0,3
scutellum and metanotum shiny,
posterior surface;
(less strongly
with a
median
keels near orbits directed towards antenna I
OL = 3;
scutum shiny,
frons
with segment 1
with fore coxae partly
(rarely 0,5)
smooth,
with a transversal
occipital
areas;
fore-wing
length of
without sculpture; keel
between dorsal
posterior surface with 2 longitudinal lateral
carina
without sculpture among
hyal ine,
keels;
without dark
d i sta I part of rad i a I vei n much shorter than
gonoforceps without dorsal
proximal
and
median prox i ma I
membranous process,
550
Fig.
Mem.
352 -
Amer.
En!.
Ins!.,
Chela of Anteon destructor
no.
(R.C.L.
37,
1984
Perkins)
from Bundaberg
(Queensland)
without distal process
(Fig.
inner process 354)
LOCUS TYP ICUS:
(Fig.
Bundaberg
lectotype F!
(designated by Olmi
1982)
1982) of N.
or with an tibial
indistinct rounded
spurs 1,1,2.
(Austral ia)
TYPICAL MATERIAL: by Olmi
353)
extending medially;
of N.
and 2 paralectotypes
(1
F,
1 M)!
destructor in B; lectotype F!
cognatus in 3; lectotype F!
(designated
(designated by Olmi
Anteoninae
551
B
Fig. 353 - Male genitalia of Anteon destructor (R.C.l. Perkins) from Tulley Falls (Queensland) (A: left half removed); B: gonoforceps (laterally viewed) •
1982) of N.
ignotus in B.
HOSTS: in Australia Eurinoscopus sp. (R.C.l. Perkins 1906c)(destructor) and Idiocerus sp. or ftiacropsis sp. (R.C.l. Perkins 1906c) (cognatus). DISTRIBUTION: Australian region: AUSTRALIA: Bundaberg (Queensland), B! Gordonvale (NE Queensland), SR! Tulley Falls (N Queensland), B! Mt. Glorious (Queensland), TIV! Wi Ison 's Peak (via Teviot Gap, SE Queensland), Ol! UQ! longreach (Queensland), Dl! Sydney (New South Wales), B! Cabramatta (New South Wales), BM! Yuendumu (Northern Territory), TW! Ol! Mt. Baw Baw (E Victoria), UQ! Strahan (Tasmania), Ol! TW! Bronte Park (Tasmania), TW! NEW BRITAIN: Gaulim (Gazelle Pen.), B! NEW GU INEA: Wau (Morobe Distr., NE New Guinea), B! Ambunti (Sepik R., NE New Guinea), B! Mi Ine Bay (SE New Guinea), B! Swart Val. (Irian), B! Karimui (NE New Guinea), S! Nabire (S Geelvinck Bay, NW New Guinea), B! Malgi (Mt. Giluwe, SE t<ew Guinea), Ol! lake Habbema (3250-3300 m), LEI Baiyer R., Ol! TW! Mt. Kainde, Ol! TW! Nadzab, OL! TW! Busu R. (lae), TW! Gent R. (Jimmy Valley), TW! Tari Gap (Mt Hagen), TW! SOLOMON ISLANDS: Soaika Hi II (Hutuna, Rennell Island),
15. Anteon caledonianum n. sp
CO!
552
Mem.
Fig.
354 -
Amer.
FEMALE:
fully
winged;
Ins!.,
no.
37,
1984
Anteon destructor (R.C.L. Perkins): variety
Male genital ia of with
Ent.
weak
rounded process.
length 2,50 mm;
antennae with segments 1-3 testaceous;
black;
mandibles testaceous;
other segments missing
in the only
known female specimen; legs testaceous, with hind coxae proximally infuscate; antennae damaged in the only known female specimen, with only first
three segments;
first
three segments
in following proportions:
9: 4,5: 6; head shiny, fully reticulate rugose; frons with a median keel and with 2 lateral keels near orbits; occipital carina complete; POL = 5; OL = 3; on I y
OOL = 2,5;
TL = 4;
pronotum shiny,
near posterior marg in;
scutum
(5: 12);
puncta te, reaching
pronotal
reticulate rugose,
tubercles reaching
tegulae;
without scu I pture among punctures; approximately 0,3
smooth
posterior surface of pronotum shorter than
length of scutum;
scutum shiny,
notau Ii ceS
weakly
i ncomp I ete,
scutellum and metanotum
shiny, smooth, weakly punctate, without sculpture among punctures; propodeum reticulate rugose, with a strong transversal keel between dorsal
and posterior surface;
posterior surface with 2
longitudinal
keels;
median area as rugose as lateral areas; fore-wing hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part
(2:8);
fore tarsal
segment 5 of front (6,5: 1,5); proximal lamellae; MALE:
segments
tarsus
enlarged claw prominence;
fully
winged;
coxae proximally
(Fig.
355)
group of 2
length
infuscate;
tarsus
lamellae;
(Fig.
tibial black;
1 testaceous;
5:
2:
3:
longer than apical
with one bristle
1,25-2,00 mm;
with segment
proportions:
part much
segment 5 of front
apex with a
antennae brown,
in following
with basal
8;
part
located on a 355)
with
1 row of 9
spurs 1,1,2. mandibles testaceous;
legs testaceous,
antennae not distally
4:
thickened;
with hind antenna I
553
An teon i nae
355 - Chela of Anteon caledonianum n. sp.
Fig.
segments shiny,
in following proportions:
reticulate rugose,
7:
4:
4:
3:
4:
4: 4:
4:
(type)
4:
6;
head
with ocellar region and a median frontal
less reticulate rugose or without areolae,
punctate,
among punctures; frons with a median keel, without lateral keels; occipital carina complete; POL = 6; OL = 3; OOL = 3; OPL = 2; TL scutum,
scutellum and metanotum shiny,
sculpture among punctures;
smooth,
weakly punctate,
notaulices incomplete,
0,3 length of scutum; propodeum reticulate rugose,
region
without sculpture
= 2; without
reaching approximately with a strong
transversal keel between dorsal and posterior surface; posterior surface with 2 longitudinal keels; meciian area as rugose as lateral areas; forewing hyal ine,
without dark transversal
bands;
distal
part of radial
vein
(2:6); gonoforceps (Fig. 356) with a long and narrow dorsal process (Fig. 357), without distal inner pointed process; tibial spurs 1,1,2.
much shorter than proximal
LOCUS TYPICUS:
Col
TYPICAL MATERIAL:
part
de la Pirogue (New Caledonia) holotYiJe F!
and 2 paratypes MM!
in B;
1 paratype
M! in OL. HOSTS: unknown DISTRIBUTION: Col
Australian
des Roussettes,
OL!
region:
Plum,
B!
NEW CALEDm
Col
B!
16. Anteon typicum (R.C.L. Perkins)
de la Pirogue,
B!
554
Mem.
Fig.
Amer.
En!.
Ins!.,
no.
37,
1984
Male genitalia of Anteon caledonianum n. sp.: A: viewed; 3: ventrally viewed (left half removed).
356 -
laterally
Neochelogynus typicus R.C.l. Perkins 1905: 61. Anteon typicus (R.C.l. Perkins): Kieffer 1907b: 27. Anteon doddi Kieffer 1911b: 232 (n. syn.); locus typicus Townsville (Australia).
Liodryinus doddi (Kieffer): Kieffer 1914b: 57. Chelogynus typicus (R.C.L. Perkins): Kieffer 1914b: 189. Anteon doddi Kieffer: Richards 1953: 54.
FEMALE:
fully
winged;
with teeth brown; distal
length 3,75-4,87 mm;
antennae testaceous,
part of 7 brown;
testaceous;
legs
11:
10: 9:
10: 9:
mandibles yellow,
sometimes also segments 1-4 brown;
testaceous,
femora and tibiae brown; 8:
black;
with segments 8-10 and sometimes
8:
with coxae brown;
antennal
8:
11;
tegulae
sometimes also clubs of
segments in following proportions:
sometimes reticulate rugose only on frons, wi thout scu I pture among punctures;
fronta I
whereas vertex line comp lete;
is punctate and frons
without
lateral keels; occipital carina complete; POL = 9; Ol = 4; OOl = 5; = 6; Tl = 5; pronotum strongly punctate and rugose, hairy, with posterior surface shorter than scutum tegulae;
scutum shiny,
punctures; scutum
smooth,
(12:22);
pronotal
weakly punctate,
notaulices incomplete,
(in doddi)
22:
head dull, fully reticulate rugose;
OPl
tubercles reaching
without sculpture among
visible only near anterior margin of
or reaching approximately 0,5 length of scutum
(typicum); scutell um and metanotum sh i ny, smooth, weak I y punctate, without sculpture among punctures; transversal
keel
between
dorsal
propodeum reticulate rugose,
and posterior surface;
with a
posterior surface
Anteoninae
555
.. '.
Fig.
Male of Anteon caledonianum n.
357 -
gonoforceps
with 2 longitudinal
keels;
median area shiny,
only near anterior margin; beneath
the pterostigma;
proximal 3:
4: 9:
part 22;
proximal
(5:13);
(Fig.
358)
Bundaberg
TYPICAL MATERIAL:
smooth,
punctate,
vein
rugose
band
much shorter than
with one bristle located on tarsus
lamellae extending continuously
1,1,2. MALE: unknown lOCUS TYP ICUS:
process of
segments in following proportions:
segment 5 of front
prominence;
dorsal
with 1 dark transversal
part of radial
tarsal
enlarged claw
of nearly 55
fore-wing
distal
fore
sp.:
(laterally viewed).
(Fig.
358)
to the apex;
10:
a
with some rows tibial
spurs
(Austral ia)
holotype F!
of N.
typicus in B; holotype F! of A.
doddi in 8M. HOSTS:
unknown
DISTRIBUTION:
Australian
Eurimbula Ck.
(via Round Hill
region:
AUSTRALIA:
Head,
Bundaberg
Queensland),
(Queensland),
Bri,!
Tamborine Mt.
(Queensland),
(Queensland),
Ol!
TW!
UQ!
Canberra,
Clare
Ol!
UQ!
(Queensland),
B!
Townsville
Mt.
Glorious
(South Australia),
TW!
17. Anteon bougainvillei n. sp.
FEMALE:
fully
winged;
antennae black; thickened; 9:
8:
8:
8;
length 3,75 mm;
legs black,
partly
black;
testaceous;
mandibles testaceous; antennae distally
antenna I segments in following proportions: antenna I segment 10 missing
in
17:
7:
12:
10: 9:
the only kmown specimen;
head shiny, rugose; frons with numerous longitudinal keels and with a median keel; vertex and region between ocellar triangle and occipital
556
Mem.
Fig.
358 -
Amer.
En!.
Ins!.,
Chela of Anteon typicum
no.
(R.C.L.
37,
1984
Perkins)
from Townsville
(Queens I and)
carina strongly sculptured by obi ique keels and with some areolae; occipital
POL = 7;
carina complete;
pronotum hairy,
dull,
OL = 5;
scutum shiny,
among punctures; length of scutum;
smooth,
strongly punctate,
transversal
surface with 2 transversal bands,
keel
between
longitudinal
keels,
one on
keels,
TL = 8; with
without sculpture
notaulices incomplete, reaching approximately 0,25 scutellum and metanotum shiny, smooth, punctate,
without sculpture among punctures; strong
OPL = 10;
transversal
(11 :20); pronotal tubercles reaching
posterior surface shorter than scutum tegulae;
OOL = 7;
strongly sculptured by
as
keels;
lateral
the basal
propodeum
dorsal
reticulate rugose,
and posterior surface;
with a
posterior
median area strongly sculptured by
areas;
fore-wing
with 2 dark
transversal
cells and another beneath the pterostigma;
fore
8: 2,5: 3: 8: 21; segment 5 of front tarsus with basal part shorter than apical part (8: 13); enlarged claw (rig. 359) with one bristle located on a proximal prominence; segment 5 front tarsus (Fig. 359) with 2 rows of nearly 27 lamellae; apex with a group of 4 lamellae; tibial spurs 1,1,2. tarsal
segments
MALE:
unknown
in following proportions:
LOCUS TYPICUS: Kokure (Bougainville I., TYPICAL MATERIAL: holotype F! in B. HOSTS:
Solomon
Islands).
unknown
DISTRIBUTION: Prince Ra.,
Australian
Bougainvi lie
region: Island),
SOLOMON
ISLANDS:
B!
18. Anteon curradoi n. sp.
Kokure
(near Crown
Anteoninae
Fig.
FEMALE:
fully
359 -
Chela of
winged;
antennae brown, brown;
legs
testacous,
with
tarsi
antennal 10;
segments
head shiny,
longitudinal 6,5;
1 testaceous;
black;
with hind coxae proximally brown;
in following proportions: occipital
TL = 5;
mandibles testaceous;
sometimes antennae fully
and fore tibiae testaceous;
reticulate rugose,
keels;
OPL = 4;
Anteon bougainvillei n. sp. (type)
length 3,12-4,18 mm;
with segment
brown,
557
sometimes
antennae distally
13:
7:
11:
10:
10:
with frons sculptured by
carina complete;
pronotum rugose,
POL = 6;
9:
9:
strong
OL = 3;
transversely striate,
legs
thickened;
OOL
with
9:
9:
558
Mem. Amer.
Fig.
Inst.,
no.
37,
1984
360 - Chela of Anteon curradoi n. sp.
posterior region smooth;
(5:20); pronotal
Ent.
(type)
posterior surface of pronotum shorter than scutum
tubercles reaching tegulae;
without sculpture among punctures;
scutum strongly punctate,
notaulices
incomplete,
reaching only
0,3 length of scutum; scutellum and metanotum shiny,smooth, punctate,
without sculpture among punctures;
with a strong
transversal
keel
between
weakly
propodeum reticulate rugose,
dorsal
and posterior surface;
posterior surface with two longitudinal keels; median area rugose; forewing hyaline,without dark transversal bands; distal part of radial vein much shorter than proximal
part
(5:11)
fore tarsal
segments in following
16; enlarged claw (Fig. 360) with one bristle located on a proximal prominence; segment 5 of front tarsus (Fig. 360) with 2 rows of nearly 14 lamellae; apex with a group of 4 lamellae; tibial spurs 1,1,2. proportions:
MALE:
7: 2: 3: 6:
unknown
LOCUS TYPICUS:
Tari
TYPICAL MATE::IAL:
Gap
(New Guinea)
holotype F!
and
1 paratype F!
in TW;
1 paratype F!
in OL. HOSTS:
unknown
DISTRIBUTION: TW!
Australian
region:
NEW GUINEA:
Tari
Gap
(Mt.
Hagen),
OL!
NOTES:
the species is named
in honor of Mr.
Italo Currado.
19. Anteon firmum n. sp.
FEMALE:
fully
winged;
length 4,56 mm;
black;
legs reddish,
with mid
559
Anteoninae
Fig.
361
-
Chela of Anteon firmum
and hind coxae blackish;
n.
sp.
(type)
mandibles and antennae black;
antennae
distally thickened; antennal segments in following proportions: 18: 9: 16: 14: 11: 12: 13: 11: 10; last antennal segment missing in the only known
560
Mem.
specimen;
head dull,
and with 2 OL = 5,5;
lateral
Amer.
fully
Ent.
Ins!.,
reticulate rugose;
keels near orbits;
OOL = 8;
no.
OPL = 11;
37,
frons
occipital
TL = 8;
1984
with a median
carina complete;
pronotum dull,
transversely striate,
except for posterior surface which
posterior surface of pronotum shorter than
(14:20);
pronotal
tubercles reaching
reticulate rugose, scutellum dull,
with notaulices faintly
reticulate rugose;
posterior area
and
latera!
reticulate rugose, posterior surface; area smooth, beneath
shiny,
with a
4:
5:
13:
(5:14);
31;
shorter than
scutum dull,
scutum
hairy,
fully
visible near anterior margin;
metanotum reticulate rugose, smooth
few
bands,
the pterostigma; part
areas
is weakly
and
without
with
sculpture;
a
propodeum
with a strong transversal keel between dorsal and posterior surface with 2 longitudinal keels; median
with 2 dark transversal proximal
tegulae;
8·,
strongly
punctate and
hairy;
keel
POL
distal
fore
irregular keels near margins; one on
part of radial
tarsal
segments
segment 5 of front apical
part
the basal
tarsus
(13:18);
fore-wing
cells ar,d another
vein much shorter than
in following proportions:
(Fig.
361)
enlarged claw
with basal
(Fig.
361)
10:
part
with one
bristle located on a proximal prominence; segment 5 of front tarsus with 2 rows of 39 lamellae; apex with a group of 7 lamellae; tibial spurs 1,1,2. MALE:
unknown
LOCUS TYPICUS:
Bulolo R
TYPICAL MATERIAL: HOSTS:
(NE New Guinea)
holotype F!
in B.
unknown
DISTRIBUTION:
Australian
region:
NEW GUINEA:
(NE New Guinea),
Bulolo R
8!
20. Anteon leiosomum (R.C.L. Perkins)
Neochelogynus leiosomus R.C.L. Perkins 1905: 62. Anteol1 leiosomus (R.C.L. Perkins): Kieffer 1907b: 26. Chelogynus leiosomus (R.C.L. Perkins): I
FEMALE:
fully
teeth brown; brown;
winged;
length 3,11
antennae yellow,
legs yellow,
punctate,
weakly
with keels on
punctate;
transversal with 2
pronotum nearly
scutum smooth,
approximately 0,5 keel
mandibles reddish,
propodeum rugose, between keels;
near anterior margin;
body shiny,
sides and on anterior surface;
dorsal
hairy;
long,
posterior surface
incomplete,
reaching
scutellum and metanotum smooth, mainly near metanotum,
and posterior surface;
median
fore-wing
notaul ices
with
tegulae yellow-
twice as broad as
weakly punctate;
length of scutum;
longitudinal
black;
with mid and hind coxae brown;
head reticulate rugose; smooth;
mm;
with segments 7-10 brown;
area
smooth,
with 2.dark
with
a
posterior surface
punctate,
transversal
rugose only bands,
one on
561
Anteoninae
Fig.
the basal rad i a I
362 -
Chela of Anteon leiosomum
cells and another beneath
Perkins)(type)
the ;:oterostigma;
vei n much shorter than proxi ma I part;
nearly as long as segment 1; chela: ,\1ALE:
(R.C.L.
fig.
362;
distal
part of
segment 4 of fron t tibial
tarsus
spurs 1,1,2.
unknown
LOCUS TYP I CUS:
Bundaberg
TYPICAL MATERIAL: HOSTS:
(Austra I i a)
holotype F!
in B.
in Australia Eurinoscopus sp.
DISTRIBUTlm,:
Australian region:
(R.C.L.
AUSTRALIA:
Perkins 1906c) 8undaberg
(Queensland),
21. Anteon nitidurn (r<'C.L. Perkins) n. comb.
Neochelogynus nitidus R.C.L. Perkins 1905: 62. nec Anteon nitidus I
FEMALE:
fully
winged;
with teeth brown;
length 2,25-4,02 mm;
yellow-reddish with segments 7-10 brown; yellow-brown,
black;
mandibles yellow,
antennae brown with segments 1-2 testaceous, tegulae yellow-brown;
or legs
sometimes with coxae dark or mid and hind femora and
tibiae brown; body shiny, hairy; head reticulate rugose; pronotum punctate, with keels on sides and on anterior surface; pronotum nearly
8!
562
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
~. . • . . .
Fig.
363 -
Chela of
Anteon nitidum (R.C.L. Perkins) from Mt. Lindesay Hawy.
twice as broad as
long,
notaulices incomplete,
(New South Wales)
with posterior surface smooth,
reaching
approximately 0,25-0,50
scutell um and metanotum smooth, mostly near metanotum, posterior surface;
with a
weak I y
punctate;
transversal
posterior surface without
keel
weakly punctate; length of scutum;
propodeum rugose,
between
longitudinal
dorsal
and
keels or with
Anteoninae
Fig.
364 -
Male genitalia of
Anteon nitidum (R.C.L. Perkins) from Tathra
(New South Wales):
only
tracks of
longitudinal
keels;
fore-wing
distal
part of radial
segments
with
keels,
1 dark
563
left half removed.
faintly visible among the
transversal
vein much shorter than
in following proportions:
5:
irregular
band beneath the pterostigma;
2,5:
3:
proximal 6:
15;
part;
fore
tarsal
enlarged claw
(Fig.
363) with one bristle located on a proximal prominence; segment 5 of front tarsus (Fig. 363) with basal part shorter than apical part (5:10), with 2 rows of nearly 70 tibial
spurs 1,
MALE:
fully
lamellae extending continuously
winged;
length 1,81-2,81
antennae brown or testaceous dark; proximally broxn Guinea; rugose,
(in holotype);
antennae not distally
proportions: dull;
7:
to the apex;
2.
1,
5:
frons
6:
5:
5:
6:
mm;
black;
fully brown thickened; 5:
5:
mandibles testaceous;
legs testaceous,
5:
8;
with or without median
with hind coxae
in some para types from New
antenna I segments
in following
head fully or partly keel,
with or without
reticulate lateral
564
,\'lem.
keels;
head
sometimes
Amer.
not
carina complete;
punctate, reaching
rugose,
=
POL
OL
=
37,
1984
punctate and
5;
=
OOL
6;
without
=
OPL
3;
punctures;
notaul ices
length of scutum;
strong transversal
keel
posterior surface without
=
TL
scutellum and melanotum shiny,
without sculpture among
a
no.
rugose,
6;
scutum,
approximately 0,3
Wilh
surface;
Inst.,
reticulate
sculpture among punctures; occipital
Ent.
3;
weakly
incomplete,
propodeurn retuculate
between dorsal
longitudinal
keels;
and posterior fore-wing
hyal ine,
without dark transversal bands; distal part of radial vein much shorter than proximal part (3:8); gonoforceps (Fig. 364) without dorsal proximal membranous process and 1,1,2. LOCUS TY!) I CUS:
without distal
Bundaberg
TYPICAL MATERIAL:
inner pointed process;
tibial
spurs
(Austra I i a)
holotype F!
N. nitidus in B; holotype F! of i'';.
of
dimidiatus in B. HOSTS:
Eurinoscopus sp.
in Australia
(".C.L.
Perkins 1906c:
under name
N. dimidiatus). DISTRIBUTIO:--J:
Australian
region:
AUSTRALIA:
Bundaberg
(Queensland),
B!
Yangaborra (NE Queensland), OL! Mt. :--Jebo (Queensland), TW! 1I1t. Tamborine (Queensland), TV/! Tathra (I~ew South Wales), OL! TIV! Mt. Lindesay
Hawy.
(3 Km S Border Gate,
(Northern Territory), :--Jew Guinea),
B!
OL!
Jimmi
rA'!
New South Wales),
NEIV GU IIJEA:
Valley -
Baiyer R.,
r.lt.
Kaindi
BR!
Yuendumu
(Morobe Clistr.,
NE
TW!
22. Anteon bouceki n. sp.
FEMALE: with
fully
winged;
teeth brown;
clubs of femora,
length 3,25-3,68 mm;
antennae and hind coxae,
tegulae
black;
mandibles testaceous,
testaceous;
legs testaceous,
mid and hind tibiae brown;
with
sometimes clubs
of fore and mid femora testaceous; antennal segments in following proportions: 11: 6: 7: 7: 6: 5: 5: 5: 6: 8; head shiny, strongly punctate,
with numerous irregular keels,
longitudinal
median keel
and with 2
towards antenna I sockets; OOL
=
5,5;
OPL
=
7;
TL
occipital
=
5,5;
also among ocell i;
lateral
carina complete;
pronotum shiny,
POL = 3;
hairy,
(20:18);
pronotal
scutellum and metanotum shiny, sculpture among punctures;
tubrcles reaching
smooth,
notaul ices
posterior surface tegulae;
weakly punctate,
incomplete,
OL = 3;
with anterior half
rugose and posterior half smooth and without sculpture; longer than scutum
frons with a
keels near orbits directed
scutum,
without
reaching
approximately
0,5 length of scutum; propodeum reticulate rugose, with a transversal keel between dorsal and posterior surface; dorsal surface with areolae much wider than areolae of posterior surface; longitudinal basal
keels;
fore-wing
cells and another beneath
the pterostigma;
vein much shorter than proximal following proportions:
8:
3:
posterior surface without
with 2 dark transversal
5:
part
19:
28;
(4: 10);
fore
bands,
distal
one on
the
part of radial
tarsal
segment 2 of front
segments
in
tarsus produced
565
Anteoninae
Fig.
into a
hook;
proximal
365 -
enlarged claw
prominence;
much shorter than
apical
Kinglake
TYPICAL M;\TERIAL: r!
HOSTS:
365)
with one bristle tarsus
(Fig.
located on a
365)
with 2 rows of nearly 31
part,
lamellae;
tibial
with basal lamellae;
part
apex
spurs 1,1,2.
(Australia)
holotype F!
1 paratype F!
in QU;
(Fig.
segment 5 of front
group of nearly 5
with a
MALE: unknown LOCUS TYPICUS:
Anteon bouceki n. sp. (type)
Chela of
1 paratype F in BM;
in CB;
paratype
in OL.
unknown
DISn,IBUTION:
Australian
region:
(Melbourne,
NOTES:
honor of Mr.
the species
is named
in
Mt.
AUSn,ALIA:
Oueensland), aU! OL! Kinglake (I'ew South Wales), B,\i!
Glorious
Victoria), Zdenek
CB!
(SC: Cabramatta
Boucek.
23. Anteon mundum n. sp.
rE!,'!ALC::
fully
proximal
third black;
winged;
length 3,81
slightly distally
thickened;
16:
8:
5:
12:
keels, OL
~
8:
8:
8:
mm;
antenna I segment 8:
9;
smooth, scutum
DOL
~
5;
OPL
~
weakly ;:>unctate, (10:20);
and rnetanotum
pronotal shiny,
5;
TL
head shiny, ~
3;
mandibles testaceous,
1 testaceous beneath;
antenna I segments
6:
without sculpture among keels; 5;
:,Iack;
in following
without sculpture arT'ong
smooth,
POL
~
5;
with anterior surface
punctures,
tegulae;
weakly punctate,
longitudinal
carina complete;
pronotum shiny,
tubercles reaching
pro;:>ortions:
with numerous
occipital
with
antennae
scutum,
shorter than scutellum
without sculpture among
566
Mem.
Amer.
En!.
Inst.,
no.
37,
\i
/lI
I
I l /
\
\I,
I
\
1
\
\ \
1
\
\
\
\
Fig.
punctures; scutum;
366 -
notaulices incomplete,
bands;
strong
part of radial
fore-wing
sp.
(type)
approximately 0,5
transversal
except for a central
is without sculpture; distal
reaching
posterior surface without
transversely striate, which
Chela of Anteon mundum n.
propodeum with a
posterior surface;
/
i /
,
I
1984
keel
length of
between dorsal
longitudinal
keels,
and
strongly
smooth area near dorsal
hyal ine,
without dark
surface
transversal
vein much shorter than proximal
part
567
Anteoninae
(3: 12);
fore tarsal
segment 5 of front enlarged claw prominence; 55
segments
in following proportions:
tarsus with basal
(Fig.
366)
with one bristle
segment 5 of front
tarsus
lamellae extending continuously
MALE:
part
6:
3:
6:
longer than apical
Fig.
Kassam
367 -
(12:9);
located on a proximal
(Fig.
366)
to the apex;
with some rows of nearly tibial
spurs
unknown
LOCUS TYPICUS:
12: 31; part
(NE New Guinea)
Chela of Anteon bellator n.
sp.
(type)
1,1,2.
568
Mem.
TYPICAL MATERIAL: HOSTS:
Amer.
En!.
holotype F!
Ins!.,
no.
37,
1984
in B.
unknown
DISTRIBUTION: Kainantu,
Australian
region:
NE New Guinea),
NEW GUINEA:
Kassam
(48 Km E of
B!
24. Anteon bellator n. sp.
FEMALE: fully
fully
thickened; 8:
8:
7:
OL = 3;
striate,
10;
length 3 mm;
abdomen
antennal
8:
longitudinal 5;
winged;
testaceous;
mandibles,
petiole black;
antennae and
without
OOL = 6;
fully retuculate rugose;
lateral
OPL = 7;
keels;
TL = 4;
occipital
(8:17);
13: 8:
pronotal
10:
10:
10:
frons with a median
carina complete;
pronotum shiny,
except for a smooth area near scutum;
pronotum shorter than scutum
legs
antennae not distally
segments in following proportions:
head shiny,
keel,
black;
testaceous;
POL
transversely
posterior surface of tubercles reaching
tegulae;
scutum shiny, smooth, weakly punctate, without sculpture among punctures or very weakly granulated; notaulices incomplete, reaching only approximately 0,25 without sculpture; keel
between dorsal
longitudinal distal
length of scutum;
scutellum and metanotum shiny,
propodeum reticulate rugose,
keels;
and posterior surface; fore-wing hyaline,
part of radial
vein
with a strong
transversal
posterior surface without
without
dark transversal
much shorter than proximal
tarsal segments in following proportions: 10: 4: 4: 7: front tarsus with basa! part shorter than apical part
part
bands;
(5:10);
fore
17; segment 5 of (7: 10); enlarged
claw (Fig. 367) with one bristle located on a proximal prominence; segment 5 of front tarsus (Fig. 367) with 2 rows of nearly 42 lamellae extending continuously MALE: unknown LOCUS TYPICUS:
U.
Watut
TYPICAL MATERIAL: HOSTS:
to the apex;
tibial
spurs 1,1,2.
(NE New Guinea)
holotype F!
in B.
unknown
DISTRIBUTION: Guinea),
Australian
region:
NEW GUlrjEA:
U.
Watut SW
(NE New
B!
25. Anteon nigricorne (R.C.L. Perkins)
Neochelogynus nigricornis R.C.L. Perkins 19Q5: 63. Anteon nigricornis (R.C.L. Perkins): Kieffer 1907b: 27 Chelogynus nigricornis (R.C.L. Perkins): Kieffer 1914b: 191.
FEMALE:
fully
winged;
wi th teeth brown;
length 2,62-2,93 mm;
antennae brown;
black;
mandibles testaceous,
tegu I ae testaceous;
legs redd i sh-
569
Anteoninae
368 -
Fig.
Anteon nigricorne (i'.C.L. Perkins) from Sandhill
Chela of
(Austral ia)
testaceous, with coxae brown; occasionally brown, with tarsi light; antennal segments in following proportions: 11: 5: 5; 5: 5: 5: 5: 5: 6;
head fully
reticulate rugose;
frons with a
median
longitudinal
5:
keel
and with 2 lateral keels near orbits directed towards antenna I sockets; occipital carina complete; POL ~ 7; OL ~ 3; OOL ~ 4; OPL ~ 4; TL ~ 4; pronotum reticulate rugose,
(8:12);
scutum
pronotal
metanotum smooth,
hairy,
shiny,
weakly
punctures;
notaulices incomplete,
of scutum;
propodeum
dorsal
reticulate
and posterior surface;
fore-wing
hyaline,
with posterior surface shorter than
tubercles reaching
without
punctate,
tegulae;
reaching only rugose,
scutum,
with
a
approximately 0,25 transversal
posterior' surface without
dark
scutellum and
without sculpture among
transversal
bands;
keel
length
between
longitudinal
distal
keels;
part of radial
(3:8); fore tarsal segments in 12; enlarged claw (Fig. 368) with one bristle located on a proximal prominence; segment 5 of front tarsus (Fig. 368) with 1 row of 21 lamellae; apex with a group of 7 lamellae; tibial spurs 1,1,2. vein much shorter than proximal
following proportions:
MALE: unknown LOCUS TYPICUS:
Bundaberg
TYPICAL MATERIAL: Olmi
1982)
HOSTS:
(Australia)
lectotype F!
and 3 paralectotypes FF!
(designated by
in B.
in Austral ia
DISTRIBUTION: Sandhi II,
part
8: 2: 2: 5:
B!
Eutettix sp. (R.C.L. Perkins 1906c).
Australian
region:
AUSTRALIA:
Bundaberg
(Queensland),
B!
OL!
26. Anteon fljianum n. sp.
FElj,ALE:
fully
tegulae and
winged;
length 1,25-1,56 mm;
legs yellow;
antenna I segments
black;
mandibles,
antennae,
in following proportions:
7:
570
Mem.
Fig.
Amer.
Ent.
Inst.,
no.
369 - Chela of Anteon fijianum n. sp.
37,
1984
from Lautoka
3,5: 2: 2: 2: 2: 2: 2: 2,5: 4;
antennae distally thickened;
granulated,
area
except for a
is without sculpture; carina complete; shiny,
( 3,5:7);
with
a
OL = 3;
pronotal
without sculpture,
incomplete,
reaching
metanotum shiny, wi th a
frons
POL = 5;
with numerous transversal
than scutum smooth,
small
smooth,
longitudinal OOL = 5; keels,
head dull,
median
OPL = 4;
keel;
occipital
TL = 5;
pronotum
with posterior surface shorter
tubercles reaching
with sides very
tegulae;
scutum shiny,
weakly granulated;
length of scutum;
without sculpture;
between
Islands)
in front of the anterior ocellus which
approximately 0,3
transversa I keel
(Fiji
notaulices
scutellum and
propodeum reticulate rugose,
dorsa I and posterior surface;
posteri or
surface wi thout long i tud i na I keel s; areol ae of dorsa I surface wider than those of posterior surface; fore-wing hyaline, without dark transversal bands;
distal
part of radial
vein much shorter than
proximal
part
(1,5:4); fore tarsal segments in following proportions: 4: 1: 2,5: 2,5: 8; enlarged claw (Fig. 369) with one bristle located on a proximal prominence; segment 5 of front tarsus (Fig. 369) with 1 row of 6 lamellae; apex with a group of 3 lamellae; tibial spurs 1,1,2. MALE:
unknown
LOCUS TYPICUS:
Lautoka
TYPICAL MATERIAL: F!
Islands) and 3 paratypes FF!
in 8M;
1 paratype
in OL.
HOSTS:
in Fiji
DISTRIBUTION: BM!
(Fiji
holotype F!
AUSTRAL IA:
Islands
Nesosteles sanguinescens Kirkaldy.
Australian Narrabri
region:
FIJI
ISLANDS:
(New South Wales),
Lautoka
8M!
27. Anteon dubium Fouts
(Viti
Levu),
DL!
Anteoninae
571
Anteon dubius Fouts 1935: 157.
FEMALE:
fully
winged;
with teeth brown; with coxae brown; granulated; broad
length 1,70 mm;
body shiny,
scutum granulated;
approximately 0,5 without
longitudinal
bands;
distal
very short,
head
twice as long as
reaching
scutellum and metanotum smooth;
fore-wing
part of radial
segment 4 of front
more than
mainly near scutellum;
keels;
legs yellow,
mainly on head;
notaulices incomplete,
length of scutum;
propodeum strongly rugose,
mandibles yellow,
tegulae yellow;
with sparse hairs,
pronotum granulated,
(3:1);
brown-black;
antennae yellow-brown;
hyal ine,
posterior surface
without dark transversal
vein much shorter than
proximal
part;
tarsus slightly shorter than segment 1 (17:22);
tibial
spurs 1,1,2. MAL:::
unknown
LOCUS TYPICUS:
Teivipakeka
TYPICAL MATERIAL:
(Marquesas
holotype F!
HOSTS: unknown DISTRIBUTION: Australian region: (Nukuhiva),
Islands)
in B. MARQUESAS
ISLANDS: Teivipakeka
B!
28. Anteon coriaceum (R.C.L. Perkins)
Neochelogynus coriaceus R.C.L. Perkins 1905: 65. Anteon coriaceus (R. C. L. Perk i ns): Kieffer 1907b: 25. Anteon apterus Dodd 1913: 181 (n. syn.); locus typicus Nelson (Austral ia)
Chelogynus coriaceus (R.C.L. Perkins): Kieffer 1914b: 192.
FEMALE: with and
fully
winged;
teeth brown; legs yellow;
length 2,01-2,12 mm;
antennae brown or yellow, legs sometimes brown;
without median and
lateral
keels;
body
black;
mandibles yellow,
weakly darkened; dull;
pronotum granulated,
surface shorter than
scutum
incomplete,
anterior third or fourth of scutum;
reaching
(6: 11 );
tegulae
head granulated;
scutum granu I ated;
frons
with posterior notau Ii ces scutellum and
metanotum sh i ny and smooth; propodeum rugose, ma i n I y near metanotum, with a transversal keel between dorsal and posterior surface; posterior surface fully
rugose,
without
longitudinal
keels;
fore-wing
hyal ine,
without dark transversal bands; distal part of radial vrin much shorter than proximal part; fore tarsal segments in following proportions: 6: 1,5: 2:
4:
10; enlarged claw
(Fig.
prominence;
segment 5 of front
than
part
apical
(4,5:5,5),
group of 6 lamellae;
tibial
370)
with one bristle located on
tarsus
(Fig.
370)
with basal
with one row of 12 lamellae; spurs 1,1,2.
a proximal
part shorter
apex with a
572
Fig.
Mem.
370 -
Amer.
En!.
Chela of ;jnteon coriaceum Creek
MALE:
Ins!.,
unknown
no.
(FLC.L.
(Queensland)
37,
1984
Perkins)
from Marlborough
Anteoninae
LOCUS TYPICUS:
Cairns
TYPICAL MATERIAL:
(Queensland,
lectotype F!
573
Australia) 1982) of N.
(designated by Olmi
coriaceus in S; hoiotype F! of A. apterus in AD. HOSTS: in Australia Euleimonios sp. (R.C.L. Perkins 1906c). DISTRIBUTION: Marlborough Cent.
Australian region:
(Queensland),
Queensland),
OLl
SR!
SR!
AUSTRALIA:
Cairns
Marlborough Creek
Nelson
(Cairns,
(Queensland),
S!
(2 Km N Marlborough,
N Queensland),
AD!
NOTES: A. apterus Dodd was described as wingless. I bel ieve that the single female specimen is damaged and the wings have been broken off.
29. Anteon sucklingi n. sp.
FEMALE:
unknown
MALE: fully winged; length 1,68-1,87 mm; black; mandibles testaceous; antennae brown with segments 1 or 1-2 testaceous; legs testaceous with hind coxae proximally brown; with all
thickened; 7:
7:
7:
antennal
9;
2;
=
OOL
5;
posterior half, incomplete,
OPL
lateral
=
2;
371
-
TL
keels;
=
2;
10: 5:
occipital
carina complete;
scutum shiny,smooth,
8:
gonoforceps
7:
7:
7:
frons
POL = 4;
OL
notaulices
length of scutum;
scutum without
weakly granulated in specimen
Male genital ia of Anteon sucklingi n. S:
also
weakly punctate on
strongly punctate near anterior margin;
reaching approximately 0,65
removed);
Wilhelm)
antennae not distally
with weak areolae or irregular keels;
sculpture among punctures in holotype;
Fig.
(specimen from Mt.
segments in following proportions:
head shiny,
without median and
=
sometimes
coxae and clubs of femora partly brown;
sp.
(type) (A:
(laterally viewed).
right
half
574
Mem.
from Mt.
Wilhelm;
sculpture; keel
Amer.
between
much shorter than proximal
with part
TYPICAL MATERIAL:
Fig.
smooth and
without
longitudinal
bands;
distal
gonoforceps
with an
keels;
fore-
part of radial
(Fig.
371)
inner subdistal
vein
without pointed
(New Guinea)
holotype M!
in B;
1 para type M!
in OL.
unknown
DISTRIBUTION: Wi Ihelm
(3:9);
1984
posterior surface with
two distinct
membranous process,
process; tibial spurs 1,1,2. LOCUS TYPICUS: Mt. Suckling HOSTS:
37,
without a distinct transversal
and posterior surface;
without dark transversal
proximal
no.
scutellum and metanotum shiny,
dorsal
hyal ine,
dorsal
Inst.,
propodeum reticulate rugose,
irregular areolae and keels, wing
Ent.
Australian
(NE New Guinea),
372 -
region:
NEW GUINEA:
Mt.
Suckling,
B!
Mt.
OL!
Male genitalia of
Anteon bismarckense n. half removed.
sp.
(type):
right
Anteoninae
575
30. Anteon bismarckense n. sp.
FEMALE: MALE:
unknown
fully
testaceous;
winged;
length 2,80 mm;
antennae brown,
propodeum black;
head black,
with mandibles
wi th segments 1-3 testaceous;
abdomen brown;
legs testaceous;
thorax and
antennae ha i ry;
antennal segments in following proportions: 13: 7: 10: 7: 8: 9: 9: 8: 9: 12; head shiny, fully reticulate rugose; frons with a complete median keel; = 4;
occipital
carina complete;
scutum smooth,
broad,
=
POL
=
8; OL
very strongly punctate;
similar to areolae;
5; OOL
=
7; OPL
=
6; TL
punctures of scutum very
notaulices incomplete approximately reaching
0,5 length of scutum; scutellum and metanotum strongly puctate; propodeum reticulate rugose, with 2 longitudinal keels on posterior surface;
median area of posterior surface rugose;
without dark proximal
transversal
part
(3:15);
bands;
distal
gonoforceps
fore-wing
part of radial
(Fig.
372)
with a
hyal ine,
vein shorter than
dorsal
process not
curved at apex; tibial spurs 1,1,2. LOCUS TYPICUS: Yalom (Bismarck Islands) TYPICAL MATERIAL:
holotype M!
HOSTS: unknown DISTRIBUTION: Australian
CO!
in CO;
region:
1 paratype M!
BISMARCK
ISLANDS:
in OL. Yalom
(New Britain),
OL!
31. An teon gioriosum n. sp.
FEMALE: MALE:
unknown
fully
winged;
length
1,25-2,00 mm;
mandibles testaceous,
with
proximal half black; antennae brown; legs brown, with tarsi light; antennae not distally thickened; antennal segments in following proportions:
6:
4:
3:
4:
4:
4:
smooth;
occipital
or with
incomplete median keel
3; OOL smooth,
4:
4:
carina complete;
4:
6;
head dull,
frons without
and
without
fully granulated,
longitudinal
lateral
keels;
median keel
POL = 5;
OL =
=
4; OPL = 2; TL = 3; scutum, scutellum and metanotum shiny, weakly punctate, without sculpture among punctures; notaulices
incomplete,
reaching
reticulate rugose, posterior surface; area shiny, hyaline,
approximately 0,3
slightly
rugose,
nearly as rugose as
without dark transversal
shorter than
length of scutum;
proximal
part
(2:6);
bands;
lateral
areas;
distal part of radial
gonoforceps
(Fig.
proximal
membranous process and without distal
process;
tibial
LOCUS TYPICUS:
propodeum
with a strong transversal keel between dorsal and posterior surface with 2 longitudinal keels; median
373)
fore-wing vein much
without dorsal
inner pointed or rounded
spurs 1,1,2. Mt.
TYPICAL MATERIAL:
Glorious
(Australia)
holotype ~'.!
in 8;
1 paratype M!
in 81-::;
1 paratype
576
Mem.
Amer.
Ent.
Ins!.,
no.
37,
1984
B Fig.
Male genitalia of Anteon gloriosurn n.
373 -
removed);
M!
in TV/;
HOSTS:
B:
paratype M!
gonoforceps
sp.
(type)(A:
left half
(laterally viewed).
in OL.
unknown
DISTRIBUTION:
Australian region:
SE Queensland), Queensland),
8!
BR!
AUSTRALIA:
2nd Palen Ck.
Mt.
Nebo
(SE
Mt.
Glorious
(Rain Forest,
Crossing from Rathdowney(SE
Queensland),
OL!
Strahan
(Tasmania),
TW!
32. Anteon vulsum n. sp.
FEMALE: unknown t.\ALE: fully winged; with
teeth brown;
testaceous,
with coxae and fore and
antennae not proportions: rugose;
distally 11:
frons
OL
=
rugose,
4;
6:
thickened;
7:
6:
with a
orbits directed 6;
length 2,06-2,25 mm;
antennae testaceous,
6:
antenna I segments 5:
6:
6:
10;
keel
towards antenna I sockets;
OOL
=
5,5;
OPL
approximately
=
mandibles testaceous,
4,5;
visj~le
legs
hind clubs of femora partly brown;
track of median
with areolae less
incomplete,
5:
black;
with segments 4-10 darkened;
=
TL
in following
head shiny,
and with 2 occipital
3,5;
reticulate keels near
carina complete;
scutum shiny,
near posterior margin;
reaching 0,5
fully
lateral
POL
reticulate
notaulices
length of scutum;
scutellum and
metanotum shiny, smooth, without sculpture; propodeum reticulate rugose, with a strong transversal keel between dorsal and posterior surface; posterior surface with 2 areolae smaller than dark
transversal
proximal
part
bands;
(2:9);
longitudinal
areolae of distal
keels;
lateral
median
areas;
part of radial
gonoforceps
(Fig.
374)
area rugose,
fore-wing
hyaline,
with without
vein much shorter than
without dorsal
proximal
577
Anteoninae
A
Fig.
Male genitalia of Anteon
374 -
B:
removed);
membranous process and
without
Mt. Suck ling
LOCUS TYP I CUS:
TYPICAL MATERIAL:
vulsum n. sp.
gonoforceps
distal
(I a tera II y
(type)(A:
right
half
viewed).
inner process;
tibial
spurs 1,1,2.
(Nevi Gu i nea)
holotype M!
and
1 paratype M!
in B;
1 paratype Iv'i!
in OL. HOSTS:
unknown
DISTRIBUTIO;":
Australian
(NE ~,ew Guinec.),
OL!
~,EW
region:
rill.
I
GUINEA:
11''.1. Suckling, B! Karimui
U"E New Guinea),
3!
33. Anteon 1uctuosum n. sp.
F::'MALE: IvIALE:
unknown
fully
testaceous;
winged;
length 2,37-2,75 !;om;
legs testaceous,
antennae not distally proportions: rugose;
frons
19:
6:
with
6:
thickened; 5:
5:
black;
mandibles and antennae
with coxae and clubs of femora partly brown; 5:
antennal 5:
5:
5:
segments 9,5;
a complete median keel
near orbits directed towards antennal
and
sockets;
in following
head fully with
two
occipital
reticulate lateral
keels
carina complete;
578
Mem.
Fig.
POL
Amer.
En!.
Ins!.,
no.
37,
Male genitalia of Anteon luctuosum n. removed.
375 -
1984
sp.
(type):
right half
= 6,5; OL = 4,5; OOL = 5; OPL = 4; TL = 3; scutum, scutellum and
metanotum shiny, punctures;
smooth,
weakly punctate,
notaulices incomplete,
scutum;
propodeum reticulate rugose,
between
dorsal
and posterior surface;
longitudinal
keels;
transversal
bands;
part tibial
(3,5:9);
with a strong
gonoforceps
part of radial (Fig.
length of
transversal
posterior surface with
median area smooth; distal
without sculpture among
reaching approximately 0,5
fore-wing hyal ine,
keel
two without dark
vein much shorter than proximal
375) without distal inner pointed process;
spurs 1,1,2.
LOCUS TYP ICUS:
Baiyer R.
TYPICAL MATERIAL: MM! in OL.
(New Guinea)
holotype M!
HOSTS: unknown DISTRIBUTION: Australian
region:
and 6 paratypes MM!
NEW GUINEA:
in TW;
Baiyer R.,
34. Anteon hageni n. sp.
TWI
3 paratypes
OLI
Anteoninae
Fig.
376 -
Male genital ia of
579
Anteon hageni n. sp. (type): right half removed.
FErviALE: MALE:
unknown
fully
winged;
antennae brown,
length 2,00-2,50 mm;
with
segmePits
coxae proximally brown;
1-2
black;
testaceous;
mandibles testaceous;
legs
testaceous,
antennae not distally thickened;
with
hind
antennal
segments in following proportions: 11: 5: 8: 7: 7: 7: 6: 7: 7: 9; head shiny, reticulate rugose and with a few or numerous longitudinal keels on frons;
frons with a median keel
and 2 lateral
directed towards antenna I sockets;
occipital
=
scutum,
3;
DOL
shiny,
=
6;
smooth,
OPL
=
3,5;
TL
=
4;
weakly punctate,
notaul ices incomplete,
smooth,
POL = 7;
OL
scutellum and metanotum
without sculpture among punctures;
reaching approximately 0,25 length of scutum;
propodeum with a strong transversa I keel surface;
keels near orbits
carina complete;
between dorsa I and posterior
posterior surface with 2 longitudinal without sculpture,
keels;
median area shiny,
except for a few short keels near margins;
fore-wing hyaline, without dark transversal bands; distal part of radial vein much shorter than proximal part (3:9); gonoforceps (Fig. 376) without dorsal
proximal
process;
spurs 1,1,2.
tibial
LOCUS TYP I CUS:
M t.
TYPICAL MATERIAL: MM!
in TW;
membranous process and without distal
Hagen
(New Gu i nea)
holotype M!
4 paratypes MM!
HOSTS: unknown DISTRIBUTION: Australian
inner
and 1 paratype M!
in Bj
5 paratypes
in OL.
region:
AUSTRALIA:
Mt.
Glorious
(Queensland),
580
Mem.
Fig.
377 -
Amer.
En!.
Ins!.,
no.
37,
II,ale genitalia of Allteon haustum n.
1984
sp.
(type):
left half
removed.
OL!
TV/!
NEW GUIf'JEA:
Mt.
Distr.,
NE I'Jew Guinea),
Valley,
TW!
Hagen
OLi
[}!
(NE
New Guinea),
Bulolo,
TW!
B!
Wau
8aiyer R.,
(Morobe
OLi
TW!
Jimmy
35. Anteon haustum n. sp.
FEr"ALE: unknown MALE: fully winged; length 1,37 mm; reddish; mandibles, antennae and legs fully testaceous; antennae not distally thickened; antenna I segments in following proportions: 8: 3,5: 4: 4: 4: 5: 4: 5: 4: 6,5; head shiny, fully reticulate rugose; frons with a median keel, without lateral keels; occipital scutum,
carina complete;
"'OL ~ 4;
OL ~ 2;
scutellum and metanotum shiny,
without sculpture among punctures;
notaul ices
approximately 0,3
length of scutum;
strong
keel
transversal
surface with 2
keels;
very
OPL ~ 2;
TL ~ 2;
weakly punctate,
incomplete,
reaching
propodeum reticulate rugose,
between dorsal
longitudinal
OOL ~ 3,5;
smooth,
and posterior surface;
with
median area slightly rugose,
nearly
rugose as lateral areas; fore-wing hyal ine, without dark transversal bands; distal part of radial vein much shorter than proximal part (2,5:7);
gonoforceps
process and
LOCUS TYP ICUS:
Mt.
TYPICAL HATERIAL: HOSTS:
(Fig.
without distal
unknown
377)
without dorsal
proximal
inner pointed process;
Glorious
(Austral ia)
holotype M!
in OL.
tibial
a
posterior
membranous spurs 1,1,2.
as
Anteoninae
Fig.
378 -
581
Anteon corallinum n. sp. (type):
Male genital ia of
left half
removed.
DISTRIBUTION: Forest,
Australian
SE Queensland),
region:
AUSTRALIA:
Mt.
Glorious
(Sclerophyll
OL!
36. Anteon corallinum n. sp.
FEMALe:; MALE:
unknown
fully
testaceous; testaceous; segments
winged;
length 1,56-2,62 mm;
black;
mandibles and clypeus
antennae fully testaceous or brown with segments 1-2 legs testaceous; antennae not distally thickened; antennal
in following proportions:
8: 4:
5:
5:
5:
5: 5: 5:
5: 7;
head
shiny, smooth, strongly punctate, without sculpture among punctures; occipital carina complete; frons without median and lateral keels; POL
4; OL = 2; OOL = 4; OPL = 1,5; TL = 2; scutum, scutellum and metanotum shiny,
smooth,
notaul ices
weakly punctate,
incomplete,
propodeum with
a
reaching
transversal
posterior surface with 2 few
irregular keels,
without
dark
than proximal
keel
as rugose as (2:5);
bands;
keels;
lateral
distal
gonoforceps
membranous process and without
length of scutum;
between dorsal
longitudinal
transversal part
without sculpture among punctures; approximately 0,5
distal
and posterior surface;
median
areas;
area shiny,
fore-wing
part of radial (Fig.
378)
with a
hyal ine,
vein much shorter
without dorsal
inner pointed process;
proxirnal
tibial
spurs
1,1,2. LOCUS TYP ICUS:
t;:1.
TYPICAL MATERIAL:
I\i!
in OL.
HOSTS:
unknown
Sinewit
(New Britain)
holotype M!
in [3;
1 paratype M!
in UQ;
1 paratype
582
Mem.
DISTRIBUTION: Pen.),
B!
Australian
AUSTRALIA:
(via Darwin,
Amer.
::Ont.
~EW
region:
I
I nst.,
no.
37,
BRITAIN:
1984
,V,t.
Sinewit
(Northern Territory),
Northern Territory),
Ol!
(Gazelle
11,t.
Bundey ?d.
UQ!
37. Anteon tasmanianurn n. sp.
FErvlALE: MALE:
unknown
fully
segment fully
winged;
length 2,75-3,37 mm;
1 testaceous;
black,
legs
partly brown;
antennae not distally
strongly punctate,
12:
8:
10:
10:
thickened; 10:
ventral
side of antenna I
hind coxae black,
10:
mid and
hind
hind femora tibiae
antenna I segments
10:
10:
10:
13;
in
head shiny,
almost reticulate rugose on anterior half of frons;
frons without median and Ol = 4;
black;
with
fore and mid femora partly black,
following proportions:
7;
testaceous,
OOl = 6,5;
lateral
OPl = 5;
keels;
occipital
Tl = 4;
scutum,
carina complete;
POL =
scutellum and metanotum
shiny, smooth, weakly punctate, without sculpture among punctures; ,ootaul ices incomplete, reaching approximately 0,3 length of scutum; propodeum ret icu I a te rugose, dorsal
Fig.
and posterior surface;
379 -
Male genitalia of
wi th a
strong
transversa I keel
posterior surface with
Anteon tasmanianum n. sp. removed.
between
two longitudinal
(type):
left half
Anteoninae
keels;
median area smooth and shiny;
transversal (6:12);
bands;
gonoforceps
spurs 1,1,2. lOCUS TYPICUS: MM!
part of radial
(Fig.
379)
without dark
vein shorter than proximal
without distal
inner pointed process;
part tibial
(Tasmania)
holotype M!
and 3 paratypes MM!
in TW;
3 paratypes
in Ol.
HOSTS: unknown DISTRIBUTION: Australian TW!
fore-wing hyaline,
distal
Mount Barrow
TYPICAL MATERIAL:
583
NEW GU INEA:
region:
Baiyer R.,
Ol!
AUSTRALIA: TW!
Mount Barrow
Jimmy Valley -
(Tasmania),
Baiyer R.,
TV/!
38. Anteon permirum n. sp.
FEMALE: MALE:
unknown
fully
winged;
antennae brown,
length 2,0-2,87 mm;
black;
with segments 1-2 testaceous;
mandibles testaceous;
legs testaceous,
with coxae
and clubs of femora partly brown; antennae not distally thickened; antennal segments in following proportions: 10: 5: 8: 6: 6: 6: 6: 6: 9;
head shiny,
Fig.
380 -
strongly sculptured by
Male genitalia of
longitudinal
keels;
6:
vertex with
Anteon permirum n. sp. (type): right half removed.
584
Mem.
areolae and 5,5;
~
OPl
keels;
4;
Tl
Arner.
occipital ~
En!.
Ins!.,
carina complete,
4; scutum shiny,
sculpture among punctures;
no.
1984
~
POL
smooth,
notaulices
37,
7; Ol
~
4;
OOl
strongly punctate,
incomplete,
reaching
~
without
approximately
0,5 length of scutum; scutellum and metanotum shiny, smooth, without sculpture; between
propodeum
dorsal
longitudinal
reticulate rugose,
and posterior surface;
keels,
1,1,2. lOCUS TYP ICUS:
a strong transversal
with numerous transversal
without dark transversal bands; distal than proximal part (3:10); gonoforceps membranous process,
with
with
keels;
fore-wing
pointed process;
Ol!
spurs
in TVI;
1 paratype
unknown
DISTRIBUTION: Mts.,
tibial
(New Guinea)
TYPICAL MATERIAL: holotype M! and 9 paratypes /VIM! tAl in B; 4 paratypes lAM! in Ol. HOSTS:
hyaline,
part of radial vein much shorter (Fig. 380) without dorsal proximal
inner subdistal
Baiyer R.
keel
posterior surface without
Australian
West New Guinea),
nV!
Kougel
R.,
NEW GUI~jEA:
region: 8!
Bulolo,
TW!
Archbold lake
Saiyer R.,
Ol!
TW!
(Central
Jimmi
Valley,
TW!
39. Anteon involutum n. sp.
FEf'0AlE: MALE: brown,
Fig.
unknown
fully
winged;
length 2 mm;
with segment 1 testaceous;
381
-
Male genital ia of
black;
mandibles testaceous;
legs testaceous,
antennae
with coxae and clubs
Anteon involutum n. sp. (type): left half removed.
585
Anteoninae
of femora partly brown; segments shiny,
reticulate rugose;
occipital reaching
POL = 5;
weakly granulated;
0,5
length of scutum;
without sculpture; between
propodeum
dorsal
9:
4:
5:
thickened;
5:
5:
frons with a median
carina complete;
scutum shiny,
keel
antennae not distally
in following proportions:
OL = 2;
5:
keel,
notaulices
antenna I
5:
5:
without
OOL = 5;
8;
head
lateral
OPL = 2;
incomplete,
scutellum and
keels;
TL = 2;
approximately
metanotum shiny,
reticulate rugose,
and posterior surface;
5:
with a strong
smooth, transversal
posterior surface without
longitudinal keels, with numerous transversal keels; fore-wing hyal ine, without dark transversal bands; distal part of radial vein much shorter than proximal
part
(3:8);
membranous process and LOCUS TYP I CUS:
1,1(.
Gi I uwe
(Fig.
381)
without dorsal
inner process;
tibial
proximal
spurs 1,1,2.
(New Gu i nea)
holotype I'/I!
TYPICAL i\lATERIAL: HOSTS:
gonoforceps
without distal in B.
unknown
DISTRI3UTI0t-l: Guinea),
Australian
region:
I'~EW
GUINEA:
h1t.
Giluwe
(Malgi,
sE
~Iew
8!
40. Anteon superbum Dodd
Anteon Anteon Anteon typicus
FC::MALE:
superbus Jodd 1913: 181. superbus Dodd: Kieffer 1914b: 171. australis Dodd 1914a: 131 (n. syn.); locus r-lel son
fully
(Austra I i a) •
winged;
length 2,4 mm;
head black;
thorax and propodeum black or brown; antennae distally
thickened;
antennal
abdomen
antennae yellow;
testaceous;
segments
legs yellow;
in following proportions:
10: 6: 10: 6: 6: 6; 7: 6: 5: 7; head dull, granulated and reticulate rugose; frontal line complete; occipital carina complete; POL = 6; OL = 4,5;
OOL = 4,5;
keels,
POL = 4;
TL = 3;
pronotum rugose,
with anterior surface approximately
(13:13);
posterior surface shorter than
and metanotum shiny, punctures; scutum;
notaul ices
propodeum
posterior surface; hyal ine,
smooth,
scutum
punctate,
incomplete,
long
(6:14);
with
transversal
bands;
transversal
scutum,
approximately 0,5 two
longitudinal
median surface as rugose as
without dark
with
as posterior surface scutellum
without sculpture among
reaching
reticulate rugose,
as
distal
lateral
length of
keels on
areas;
fore-wing
part of radial
vein
shorter than proximal part (4:8); fore tarsal segments in following proportions: 22: 7: 8: 14: 42; enlarged claw (Fig. 382) with one bristle located on with
a proximal
prominence;
two rows of nearly 20
tibial spurs 1,1,2. i,1AL>:: unknown
segment 5 of front
larnellae;
apex with a
tarsus
group of 7
(Fig.
382)
lamellae;
586
Mem. Amer.
Fig.
En!.
Ins!.,
no.
37,1984
Chela of Anteon superbum Dodd from Nelson
382 -
LOCUS TYPICUS: Kuranda (Australia) TYPICAL MATERIAL: holotype F! of A.
(Queensland)
superbus in AD; holotype F! of A.
australis in AD. HOSTS:
unknown
DISTRIBUTION: Queensland),
Australian
AD!
Nelson
region:
AUSTRALIA:
(N Queensland),
Kuranda
(Cairns,
N
AD!
41. Anteon rufiscapum Dodd
Anteon rufiscapus Dodd 1913: 180 Anteon rufiscapus Dodd: Kieffer 1914b: 171.
FEMALE:
fully
winged;
length 2,16 mm;
black;
mandibles testaceous;
Anteoninae
Fig.
antennae brown,
383 -
Chela of Anteon rufiscapum Dodd
with segment 1 yellow;
antennae distally thickened; 14: 6:
5:
3:
5:
I ine complete;
5: POL
carina complete;
5:
587
5:
4:
antennal 8;
legs brown,
(type)
with tarsi
light;
segments in following proportions:
head dull,
fully reticulate rugose;
frontal
= 8; Ol = 4; OOl = 5; OPl = 5; Tl = 3; occipital
pronotum short,
long as anterior surface;
with posterior surface approximately
posterior surface shorter than scutum
as
(5:15);
scutum, scutellum and metanotum smooth, shiny, punctate, without sculpture among punctures; notaul ices incomplete, reaching approximately 0,20 length of scutum; transversal
keel
propodeum reticulate rugose,
between
without
longitudinal
beneath
the pterostigma;
dorsal
keels;
fore-wing
radial
with a strong
and posterior surface; with a dark
vein with distal
posterior surface
transversal
band
part shorter than
proximal part (2:9); fore tarsal segments in following proportions: 56: 15: 14: 16: 63; enlarged claw (Fig. 383) with one bristle located on a proximal prominence; segment 5 of front tarsus with one row of 5 lamellae, in ilddition to some bristles; tibial spurs 1,1,2. MALE:
unknown
lOCUS TYP ICUS:
Kuranda
TYPICAL lviAT"RIAL: HOS TS:
(Austral ia)
holotype F!
in AC>.
unknown
DISTRIBUTIOr,: Queensland),
Australian region:
AUSTRALIA:
Kuranda
AD!
42. An teon gira u Iti Dodd
Anteon giraulti Dodd 1914d: 251.
(Cairns,
N
588
Mem.
Fig.
384 -
Amer.
En!.
Inst., no. 37,
1984
Chela of Anteon giraulti Dodd
(type)
Anteoninae
FEMALE:
fully
winged;
abdomen brown;
antenna I segments 8;
head shiny,
shiny,
I"OL
reticulate rugose, ~
4;
~
3,5;
is smooth;
(6:11);
scutum,
mandibles testaceous;
antennae distally 10:
except for a
is smooth;
OL
reticulate rugose,
wh i ch
black;
legs yellow;
in following proportions:
ocellar triangle which complete;
length 2,4 mm;
antennae and
589
frontal ~
OOL
3;
8:
9:
7:
wide area
I ine complete; ~
OPL
6:
2,5;
6:
6:
6:
in fr'ont of
occipital ~
TL
thickened; 6:
2,5;
carina
pronotum
except for a short area near posterior margin
posterior surface of pronotum shorter than
scutellum
and metanotum shiny,
smooth,
scu tum
without
sculpture; notaulices incomplete, faintly visible near anterior margin of scutum; propodeum reticulate rugose, with a strong transversal keel between dorsal longitudinal distal
posterior surface; fore-wing
part of radial
segments 384)
and
keels;
front
in following proportions:
tarsus
(Fig.
1\-1ALE:
without dark
vein shorter than
with one bristle located on 384)
group of nearly 30
posterior surface without
hyaline,
with
7:
proximal
3: 4:
11:
a proximal
21;
tibial
(2:10);
bands;
fore tarsal
enlarged claw
prominence;
two rows of nearly 33
lamellae;
transversal
part
(Fig.
segment 5 of
lamellae;
apex with a
spurs 1,1,2.
unknown
LOCUS TYP I CUS:
Austra I i a
TYPICAL MATERIAL: HOSTS:
holotype F!
in AD.
unknown
DISTRI8UTIO~I:
Australian
,I Queensland),
region:
AUSTRALIA:
Yungaburra
(Cairns distr.,
AD!
5. Prioranteon n. gen.
TYPE SPEC I ES: Prioranteon casalei n. sp. FEMALE (Figs 385, 386): micropterous; thoracic structure simi lar to that of
Gonatopodinae: pronotum remarkably elongate; scutum forming a stalk
between
the pro- and metanotum;
propodeum segments;
humped;
located further' distally 390)
MALE:
than
i nv i si b Ie;
with 6 segments;
inner side of enlarged claw
prominence not bearing bristles; 388,
metanotum
maxillary palpi
(Fig.387)
metathorax + labial
tibial
prominence;
chela
spurs 1,1,2.
unknown
Palaearctic,
with 3
1-2 short bristles or peg-like hairs
the proximal
without rudimentary claw;
DISTRI8UTIO~:
palpi
with a proximal
Ethiopian
HOSTS: unknown SPECIES: 3
PALAEARCT I C SPEC I ES OF PRIORANTEON
(Figs 387,
590
Mem. Amer. En!.
Fig.
385 -
Ins!.,
no. 37,
1984
Female of Prioranteon casalei n.
sp.
(type)
Anteoninae
Fig.
386 -
One species
Female of
591
l'riora"teon casalei n. sp.
(type)
is known.
Prioranteon biroi n. sp.
FEMALE:
micropterous;
scutellum;
fore
testaceous,
with
teeth brown;
segments 1-3 testaceous; testaceous; black, 10:
8:
10:
10;
punctures;
segments
occipital
frons
mandibles ';,ith 4 teeth progressing
posterior; weakly
vertex flat;
punctate,
posterior
transversal
narrower than tegulae,but nearly as reduced; long as scutum humped,
disc;
very
without median
and
smooth,
tarsi
13:
13:
11:
keel,
but
POL = 7;
with Tl =
anterior one to
without sculpture or very
very
posterior lobe directed
humped,
scute II um; 389 8);
shiny,
smooth,
tegulae and
long anterior disc,
towards
carinate;
reduced
tegulae;
hollow,
without fore
reaching scutum
rugose,
sculpture,
wings on
very
nearly
as
sides of
fore wi ngs not reachi ng transversa I furrow beh i nd metanotum
invisible;
separated from scutellum by
transversely
hind 18:
posterior margin of collar almost straight,
as broad;
(Fig.
OOl = 11;
larger from
long as posterior collar of pronotum, scutellum
5:
without sculpture among
Ol = 5;
humped and
10:
impression se;oarating disc from a posterior collar,
without a
long
scutellum
pronotum shiny,
with a
with
with coxae almost fully
proportions:
POL = 4;
as
mandibles
antennae brown,
last segment of mid and
weakly punctate,
furrow;
long
black;
with posterior collar whitish-
legs testaceous,
carina complete;
median
nearly as
testaceous;
in following
head shiny,
longitudinal
12;
clypeus
pronotum black,
tegulae testaceous;
antennal
reduced,
length 3,25-3,80 mm;
clubs of femora darkened,
brown;
a
wings very
hind wings invisible;
a
metathorax + propodeum
transversal
hollow
impression
anterior surface of metathorax + propodeum shiny,
592
Mem. Amer.
Ent.
Inst.,
no. 37,
1984
i
I
j
!
\\\. /' -.
~~/
Fig. 387 -
/
Chela of Prioranteon biroi n. sp.
(type)
593
Anteoninae
partly granulated and smooth; disc of metathorax + propodeum with a distinct longitudinal median furrow; posterior surface of propodeum dull, granulated and with some weak smooth,
shiny,
complete;
fore tarsal
enlarged claw
(Fig.
the proximal
prominence; lamellae;
long);
meso-metapleural
with one bristle
into a
small
unknown
hook;
LOCUS TYPICUS:
tibial
11:
3:
6:
13:
25;
tarsus
a group of 5
387)
(Fig.
lamellae
with 2 rows
(among which
segment 2 of front
1
tarsus produced
spurs 1,1,2.
SzSztMiklos
TYPICAL MATERIAL:
pleura
located further distally than
segment 5 of front
apex with
rudimentary claw absent;
MALE:
striae;
suture distinct and
segments in following proportions:
387)
of nearly 29 very
irregular longitudinal
without sculpture;
(Hungary)
holotype F!
and 1 paratype F!
1 paratyep F!
in BU;
in OL. HOSTS:
unknown
DISTRIBUTION: NOTES:
Palaearctic region:
the species
HUNGARY:
is named in honor of the
SzSztMiklos, late Mr.
L.
BU! Biro.
ETHIOPIAN SPECIES OF PRIORANTEON
Two species are known.
KEY TO THE SPECIES
FEMALES
Posterior
surface of
propodeum strongly
. . • • • • • • . • • . . . . • • . . • . . . . • • . . • . . • . .. -
Posterior
surface of
transversely striate .•• sp.
1. richardsi n.
propodeum granulated,
kee Is. • • • • • • • . • • • • • . • . . • • • • • • • • • . •.
2.
without
casalei n.
transversal
sp.
MALES
Unknown
In the key to the Ethiopian species, the Palaearctic species PI'. biroi shou I d be near PI'. casalei, because the posterior surface of the propodeum is granu I ated and not transverse I y str i ate. Casalei and biroi NOTES:
can be distinguished as follows:
594
{v,em.
Amer.
Ent.
Inst.,
no.
t,letathorax
+ propodeum with a distinct
disc;
with
head
reaching
frons
transversal
fully
black;
furrow behind
frons black, reaching
with
a
central
furrow behind
winus
scutellum
longitudinal
yellowish
scutellum
furrow on
shorter, (Fig.
biroi n;
+ propodeum without
Metathorax
1984
longitudinal
fore
. .. . .. . .. . . . . . . . . .. . . . .. . . . . .. . -
37,
not
tl).
389
sp.
furrow on disc;
area;
fore
wings
longer,
389 A}......
(Fig.
casalei n.
.. . . . . .
sp.
1. Prioranteon richardsi n. sp.
FEMALE:
rnicropterous;
scutellum;
fore
hind wings
testaceous with
teeth brown;
segments 1-2 and proximal pronotum black, legs brown, tibiae and B:
5:
17:
with
with tarsi
12:
wings
reduced,
clypeus
testaceous;
part of coxae,
testaceous; 10:
8:
7:
antenna I segments 7:
9;
TL =
mandibles
carina complete;
larger from anterior one to posterior; smooth,
weakly punctate,
only on
sides;
pronotum
posterior transversal narrower than tegulae, nearl y
but
as
reduced;
disc;
a
behind
posterior
humped,
nearly
as
and
shiny,
scutellum;
(Fig.
24;
distally
weakly
fore
proximal
tarsal (Fig.
and
hook;
MALE:
unknown
LOCUS TYPICUS:
tibial
388)
absent;
spurs
Van Reenen
TYPICAL MATERIAL:
12;
very
OPL
=
6;
granulated
long anterior disc;
towards
posterior collar, reaching
tegulae;
hollow,
scutum
rugose,
without sculpture,
very
nearly
as
tegulae and reduced fore wings
wings
not
reaching
invisible; a
transversal
metathorax +
transversal
hollow
posterior surface strongly striate;
with
in following 2 bristles
apex
with
a
11:
tarsus
(Fig.
3BB)
group of nearly 4
segment 2 of front
South Africa)
in 8M.
pleura strongly proportions:
located further
segment 5 of front
1,1,2.
(Natal,
holotype F!
punctate,
teeth progressing
disc from a
metanotum
segments
lamellae;
rudimentary claw
small
=
pronotum shiny,
strong and complete;
prominence;
with 2 rows of nearly 43 lamellae;
OOl
anterior surface of metathorax +
granulated;
suture distinct,
striate;
into a
proportions:
strongly
3;
with 4
separated from scutellum by
enlarged claw
than
fore
389 C);
transversely carinate;
transversely
smooth,
long as broad;
impression
12:
humped
lobe directed
humped,
meso-metapleural
=
OL
vertex flat;
impression separating
scutellum
propodeum dull,
testaceous;
sculpture among punctures,
very
propodeum
7:
smooth,
long as posterior coil ar of pronoturn, scutellum
sides of scutum
furrow
with
with
stalk of femora,
posterior margin of collar almost straight,
without a
long as scutum, on
without
4;
as
antennae brown,
in following
head shiny,
=
long
mandibles
tegulae testaceous;
trochanters,
POL
occipital
black;
half of 3 testaceous;
without sculpture among punctures; 11;
nearly as
length 4,68 mm;
anterior margin and posterior collar
distal
10:
very
invisible;
tarsus produced
3:
595
Anteoninae
('ig.
388 -
Chela of Prioranteon richardsi n.
sp.
(type)
596
Mem.
Fig.
389 -
Amer.
Inst.,
no.
37,1984
Prioranteon casalei n. sp. (A), biroi n. richardsi n. sp. (C).
Reduced fore wings of sp.
HOSTS:
Ent.
(B),
unknown
DISTRIBUTION:
Ethiopian
Natal),
8M!
NOTES:
the species
region:
is named
SOUTH AFRICA:
in honor of Mr.
Van Reenen
Owain W.
(Drakensberg,
Richards.
2. Prioranteon casalei n. sp.
FEMALE long
(Figs 385,
386):
as scutellum;
mandibles testaceous, longitudinal
segments shiny,
=
with
genae,
=
OPL
5;
pronotum shiny, with a
TL
=
very
5:
clypeus and a
abdomen reddish-brown;
tarsi
13:
8;
occipital
9:
8:
testaceous;
7:
carina complete;
humped and very
than disc;
posterior margin of collar almost straight, posterior
disc from
antennal
5:
9;
head
POL = 3,5;
mandibles with 4 vertex flat;
long anterior disc;
impression separating a
6:
weakly punctate only on
trasversal but without
5:
anterior one to posterior;
without sculpture,
with
with posterior collar and
without sculpture among punctures;
larger from smooth,
8:
and
nearly as
black;
antennae brown,
testaceous;
junctions
in following proportions:
teeth progressing sides,
length 3,25-4,00 mm;
pronotum black,
tegulae
trochanters,
weakly punctate,
9;
teeth brown;
testaceous;
margins testaceous;
legs brown,
OL
with
fore wings very reduced,
invisible;
median area on frons yellowish;
segments 1--2 partly lateral
micropterous;
hind wings
a
lobe directed towards
posterior
posterior collar, tegulae;
reaching
narrower tegulae,
scutum nearly as
Anteoninae
Fig.
390 -
Chela of
597
Prioranteon casalei n. sp. (type)
long as posterior coil ar of pronotum,
hollow,
rugose,
very reduced;
scutellum humped, shiny, smooth, without sculpture, nearly as long as scutum, nearly as broad; tegulae and reduced fore wings on sides of scutum and scutellum; scutellum humped,
(Fig.
389 A);
fore w;ngs reaching metanotum
transversal
invisible;
separated from scutellum by a
furrow
behind
metathorax + propodeum
transversal
hollow
impression
598
Mem.
transversely carinate;
Amer.
En!.
Ins!.,
no.
37,
1984
anterior surface of metathorax + propodeum shiny,
smooth, without sculpture; posterior surface dul', granulated, without transversal striae; meso-metapleural suture distinct, strong and complete; pleura dull,
granulated,
with weak transversal
segments in following proportions: 390)
with 2 bristles
prominence; lamellae; absent;
9:
2: 5:
keels;
12: 21;
located further distally than
segment 5 of front
tarsus
(Fig.
390)
tarsus produced
(Fig.
the proximal with 1 row of nearly 33
apex with a group of nearly 3 lamellae;
segment 2 of front
fore tarsal
enlarged claw
rudimentary claw
into a small
hook;
tibial
spurs 1,1,2. MALE:
unknown
LOCUS TYPICUS:
Milnerton
TYPICAL MATERIAL: FF!
(South Africa)
holotype F!
and 7 paratypes FF!
in BM;
4 paratypes
in OL.
HOSTS: unknown DISTRIBUTION: Ethiopian region:
SOUTH AFRICA:
BM! OL! NOTES: the species is named in honor of Mr.
Fig. 391
-
Milnerton
(Cape Town),
Achi lie Casale.
Female of Bocchoides bekilyensis Benoit
(holotype)
Bocchinae
599
5. SUBFAMILY BOCCHINAE RICHARDS
Bocchini Richards 1939: 189. Bocchini Muesebeck and Walkley 1951: 1039. Bocchinae Nagy 1967b: 334.
TYPE
Bocchus Ashmead 1893
GENUS:
FEMALE:
fully
brachypterous
winged (Fig.
(Fig.
424);
391)
or micropterous (Fig.
fore-wing
(in fully
452)
or
winged species)
with costal,
median and submedian cells clearly enclosed by pigmented veins (Fig. 391); maxi Ilary palpi with 6 segments; labial palpi with 3 segments; front with
tarsus chelate; 1,
2,
chela
3 or 4 teeth
three
large teeth and a
(Fig.
396 A);
rarely
(Fig.
(Fig.
405)
396);
with rudimentary claw;
rudimentary
tooth between
quadridentate mandibles with
larger from anterior one to posterior; of strong
teeth
(Fig.
405);
mandibles
quadridentate mandibles usually teeth progressing
enlarged claw usually with a row
segment 5 of front
tarsus
(Fig.
405)
only with a preapical lamella, without other lamellae; pronotal directed towards the tegulae present; occipital carina complete, incomplete;
petiole usually very
long,
with
the two posterior teeth
rarely
short;
usually tubercles rarely
antennae without
tufts of long hairs; pterostigma usually very reduced, sometimes wide; tibial spurs 1,1,1, sometimes 1,1,2. MALE: fully winged (Fig. 393) or micropterous (Fig. 454); fore-wing (in fully
winged species)
enclosed by
Fig.
with costal,
pigmented veins)(Fig.
392 -
Female of
median and submedian cells clearly 393);
maxillary palpi
with 6 segments;
Bocchoides bekilyensis Benoit (holotype)
600
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
/
Fig.
393 -
Male of Bocchoides bekilyensis Benoit
(allotype):
antennal
segments 7-10 missing.
labial
palpi
with 3 segments;
mandibles with 1, 2,
3 or 4 teeth;
tridentate or quadridentate mandibles with teeth not progressing from anterior one to posterior;
occipital
carina complete;
tibial
larger spurs
1,1,2. DISTRIBUTION: Palaearctic, Ethiopian, Oriental, Nearctic, Australian HOSTS: Issidae; Cicadellidae (exc Iud i ng I diocerinae, Macropsinae and
Typhlocybinae) GENERA: 6 SPECIES:
39
KEY TO THE GENERA OF BOCCHINAE
FEMALES
Occipital head;
carina
enlarged
incomplete,
claw with
only
visible on dorsal
one preapical
lamella
(Fig.
side
of
399) .•
Bocchinae
...........................
2
Occipital
carina
preapical
lamella;
preapical
lamella
Segment with tooth
5 of
other
enlarged Enlarged front
most
segment
tarsus with (Fig.
front
of
lamella,
claw at
never
least
with
one
lamellae
preapical
(Figs
395,
lamella, 442,
451,
sometimes 456);
claw without teeth (Figs 395,442,451,456) . . . . . . . . . 3 claw wi th numerous bristles (Fig. 395 A); segment 4 of
Enlarged
least
claw withoit
451,456);
segment
segment
Fore-wing
as
long as
segment
front
1
(Fig.
395 B) . . . . . . . .
Bocchoides Beno i t
1.
brist les or
4 of
other appendixes
tarsus
than
0,5
(Figs 442, times as
1
(Figs
442,
(Fig.
452) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ..
complete
Fore-wing with
451,
less
reduced
........................... Fore-wing 5
one
tarsus with
Bocchus Asmead
tarsus without
group
tarsus at
long as
front
enlarged
3.
• • • • • • • • . • • • • • • • • • • • • • • • • ••
4
claw never with 5 of
one preapical
405);
405) •••••••••••
5 of
Chelothelius Reinhard
2.
enlarged
(Figs 395,405) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2
front
wi th a di stal 3
at
lamellae
(Fig.
Segment
complete;
601
radial
f\1ystrophorus Forster
6.
(Figs 438,
456) •••••••••••••••••••••••• 4
449) ••••••.••••••••••••••••••••• 5
vein distinct
(Fig.
438) . . . . . . . . . . . . . . . . .
••••••••• • ••••••••• • • •••• •• 4. Mirodryinus Ponomarenko Fore-wing with no radial vein (Fig. 449) . . . . . . . . . . . . . . . . . . . . . . . • ••••••••••••••••••••••••••
Nadiimancus Moczar
5.
MALES
Temp I e s not
vis i b I e
(F i g s
393,
• • • • • • • • • • • • • • • • • • • • • • • • • •• 2
Temples distinct (Figs 402,403,440,441,454,455) . . . . . . . . . . 2 Fore-wi ng reduced (F i g. 454) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . ........................... Fore-wing
3
394) . . . . . . . . . . . . . . . . . . . . . . . . . . ..
Bocchoides Ben 0 i t
1.
compl ete
Mystrophorus Forster
6.
(Figs 402,
440) . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3
Propodeum with areolae very
small,
440) ••.••••••••••••••••••••
4.
Propodeum wi th at
I east
least
tegulae
as
large as
part
of
(Fig.
• • • . • • • • • • • • • • • • • • • • • • • • • ••
3.
less wide
than
tegulae
(Fig.
Mirodryinus Ponomarenko the areolae very
large,
at
402) . . . . . . . . . . . . . . . . . . . . . . . . . . .
Bocchus Asmead
The ma les of Chelothelius Rei nhard and Radiimancus Moczar are unknown.
1. Bocchoides Benoit
602
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
Bocchoides Benoit 1953b: 424.
Bocchoides bekilyensis Benoi t 1953, monotyp ic.
TYPE SPEC I ES: FEMALE labial
(Figs 391,392): palpi
tridentate
(Fig.
396 B);
segment 5 of front front
tarsus at
not reaching MALE
least as
palpi
visible;
394):
HOSTS:
(Fig.
lamellae (Fig.
395);
1 (Fig.
pronotal
395);
395);
segment 4 of tubercles
spurs 1,1,2. winged;
with 3 segments;
mandibles tridentate,
DISTRIBUTION:
mandibles
with numerous bristles
long as segment
fully
with 6 segments;
carina complete;
with numerous
tibial
anterior one to posterior
maxillary palpi
occipital
enlarged claw
tarsus
tegulae;
(Figs 393,
labial
fully winged;
with 3 segments;
(Fig.
maxillary palpi
occipital with
with 6 segments;
carina complete;
teeth not progressing
396 D);
tibial
temples not larger from
spurs 1,1,2.
Ethiopian
unknown
SPECIES:
1
13occhoides bekilyensis Benoi t
Fig.
394 -
Male of
Bocchoides bekilyensis Benoit (allotype): antenna I segments 7-10 missing.
603
Bocchinae
Fig.
395 -
Female of Bocchoides bekilyensis Benoit fore tarsa I segmen ts.
(holotype):
A:
chela;
B:
604
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
Bocchoides bekilyensis Benoi t 1953b: 424.
FEMALE
(Figs 391,392):
fully
winged;
clypeus and mandibles testaceous; darkened;
thorax,
testaceous,
propodeum, 14:
7:
33:
distally thickened;
14:
12:
head dull,
10:
8:
hairy,
7:
with
legs brown-
antennal
8:
black,
with segments 8-10
pet iole and abdomen black;
with coxae and clubs of femora dark;
following proportions: slightly
length 6,25-6,50 mm;
antennae testaceous,
10;
rugose,
segments
in
antennae
sculptured by short
irregular keels; occipital carina complete, reaching posterior margin of eyes; temples invisible; thorax and propodeum shiny, hairy; pronotum and scutum smooth,
without sculpture;
without sculpture;
propodeum reticulate rugose; median
area rugose;
five times as as broad
enlarged claw
for", tarsal
(Fig.
segment 5 of front
395)
tarsus
extending continuously MALE
(Figs 393,
mandibles yellow;
394):
(14:2,5);
(Fig.
and
hind
to the apex; fully
4:
14:
rugose;
mandibles tridentate,
396 D);
occipital
temples
invisible;
scutum;
scutellum granulated; without
(25:7);
tibial
yellow;
9;
legs yellow,
antennal
with
segments
1-6
intermediate tooth
in
(Fig.
visible only on anterior half of
metanotum reticulate rugose;
longitudinal
propodeum
keels on posterior surface;
long and ovoidal,
more than
fore-
three times as
long as
spurs 1,1,2. in P.
unknown
DISTRIBUTION: Ethiopian region: MADAGASCAR: 8ekily, P! NOTES: B. bekilyensis is very different from the other Bocchinae. general
spurs
(1,1,2)
are similar to tibial
tridentate mandibles the species
spurs
is included
arrangement of thi s genus prQbab I y
in
is between
The
Anteoninae or as in Dryininae. Female of Dryininae. Only for the Bocchinae. The exact Bocchinae and Dryininae.
structure of the chela is simi lar to that of
Dryininae. The pterostigma is long and narrow tibial
with
in allotype);
posterior margin of eyes;
LOCUS TYP ICUS: Beki Iy (Madagascar) TYPICAL MATERIAL: holotype F! and allotype M! HOSTS:
head black,
head and scutum reticulate
reaching
notaul ices weakly
wings with pterostigma broad
tegulae
with one rudimentary
carina complete,
lamellae
spurs 1,1,2.
length 3,12 mm;
10:
long
(11:14);
without other appendixes;
with 2-3 rows of 63
(segments 7-10 missing
10:
keels;
nearly
fore trochanters nearly twice as
abdomen brown-black;
tibiae darkened;
longitudinal
1 shorter than segment 4
tibial
winged;
antennae yellow
following proportions: 5:
reticulate rugose,
395)
two
long and narrow,
with numerous bristles,
thorax and propodeum black; femora
with pterostigma
segment
scutellum smooth,
with some areolae near scutellum;
posterior surface with
fore-wing
long as broad
(8:4);
notaul ices invisible;
metanotum smooth,
2. Chelothelius Reinhard
Bocchinae
605
A
Fig.
396 -
E
Mandibles of Bocchinae:
rudimentary tooth;
B:
from anterior one to posterior; mandible; E:
quadridentate mandible with one
C:
mandible with one tooth;
tridentate mandible with one rudimentary
Chelothelius Chelothelius Chelothelius Chelothelius
Reinhard
1863:
D:
larger
tridentate
intermediate tooth.
409.
Reinhard:
Kieffer
Reinhard:
Kieffer 1914b: 56.
in Kieffer et Marshall
Reinhard:
Berland
1928:
1905:
125.
143.
Chelothelius gryps Reinhard 1863, monotypic.
TYPE SPECIES: FEMALE
A:
tridentate mandible with teeth progressing
(Figs 397,
398):
fully
winged,
maxillary palpi
with 6 segments;
labial palpi with 3 segments: occipital carina incomplete, only visible on dorsal side of head; mandibles tridentate: enlarged claw with one preapical with
lamella and some bristles
lamellae
(Fig.
399);
pronotal
(Fig.
399);
segment 5 of front
tubercles not reaching
spurs 1,1,2. MALE: unknown DISTRIBUTION: Palaearctic HOSTS: unknown SPECIES:
2
PALAEARCTIC SPECIES OF
Two species are known.
CHELOTHELIUS
tegulae;
tarsus tibial
606
Mem.
Fig.
397 -
Amer.
Ent.
Ins!.,
no.
37,
1984
Female of Chelothelius gryps Reinhard from South France
KEY TO THE SPECIES
FEMALES
Head
fully
granulated,
scutellum dull,
wi th numerous keel s,
granulated and
•••• •••• •••••••• ••• 1. gryps Rei nhard Head shiny, granulated and with keels, wide areas
on
sides
of
ocell i
except
whi ch are
sculpture;
scutum and
sculpture,
weakly granulated,
without
berlandi
Bernard
•••••••••••••••••• • 2.
dull;
scutum and
with keels . . . . . . . .
scutellum shiny,
for
two
smooth and wi thout smooth,
almost
without
keels . . . . . . . . . . . . . . . . . . .
607
Bocchinae
MALES
Unknown
1. Chelothelius gryps Reinhard
Chelothelius gryps Rei nhard 1863: 410. Dryinus gryps (Reinhard): Dalla Torre 1898: 544. Chelotelius gryps Reinhard: Kieffer in Kieffer et Marshall 1905: 127. Chelothelius gryps Reinhard: Kieffer 1914b: 57. Chelothelius gryps Rei nhard: Ber I and 1928: 143. Chelothelius gryps Rei nhard: Bernard 1935a: 41.
FEMALE
(Figs 397,
398):
fully
winged;
length 5,0-5,4 mm;
antennae brown,
with segments 1-2 testaceous; head black, with mandibles, clypeus, a narrow frontal region near clypeus, malar space and a region around orbits reddish-testaceous;
Fig.
398 -
propectus black;
pronotum black,
with margins
Female of Chelothelius gryps Reinhard from South France
608
Mem.
Fig.
reddish;
399 -
Amer.
Ent.
Inst.,
no.
37,
1984
Chela of Chelothelius gryps Reinhard from South France
scutum,
scutellum,
metanotum,
mesopleura,
metapleura and
propodeum black; petiole black; abdomen black-brown; legs testaceous, with coxae partly black and clubs of femora brown; tegulae testaceous; antenna I segments in following proportions:
12: 7:
24:
13:
13: 8:
6:
6:
6:
9; head dull, granulated, with numerous keels, mainly longitudinal; mandibles with 3 teeth; ocellar triangle and occiput smooth; frons with incomplete median
longitudinal
carina visible only on dorsal temples almost transversal transversal
invisible;
keel;
= 5; Ol = 4; OOl = 10; occipital
POL
region of head,
pronotum shiny,
but not reaching eyes;
granulated,
with weak
keels; pronotal turcles not reaching tegulae; pronotum with a impression between anterior collar and posterior humped disc;
scutum dull,
granulated,
with numrous irregular keels or areolae;
notaulices invisible; scutellum as sculptured as scutum; metanotum fully reticulate rugose; propodeum fully reticulate rugose, with 2 incomplete longitudinal bands,
keels on posterior surface;
one on
the basal
fore wings with 2 dark
transversal
cell s and another beneath the pterost i gma;
pterostigma long and narrow;
distal
part of radial
vein
nearly as
long as
proximal part, not reaching wing margin; fore tarsal segments in following proportions: 13: 3: 5: 9: 16; segment 5 of front tarsus (Fig. 399) with some rows of nearly 26 lamellae extending continuously to the apex; enlarged claw
(Fig.
segment 3 of front tibial
spurs 1,1,2.
MALE:
unknown
lOCUS TYPICUS:
399)
with 3 proximal
tarsus produced
Bolzano
(Italy)
into a
teeth and one preapical hook,
with no very
lamella;
long bristle,
Bocchinae
TYPICAL MATERIAL: HOSTS:
probably
609
lost.
unknown
DISTRIBUTION: Goudargues
Palaearctic region:
(Gard),
OL!
ITALY:
FRANCE: Southern France,
Bolzano
P!
Ussel-
(Reinhard 1863).
2. Chelothelius berlandi Bernard
Chelothelius berlandi Bernard 1935a: 41. Chelothelius berlandi Bernard: Bernard 1935b: 53.
FEMALE:
fully
winged;
and mandibles
length 3,75 mm;
testacous;
head black,
antennae testaceous;
segment 10 distally testaceous;
thorax fully black,
(black,
petiole black;
with margins reddish);
testaceous,
with clypeus brown
with segments 6-10 brown; except for pronotum
abdomen black-brown;
with coxae and clubs of femora partly brown;
testaceous;
antennal 6:
9:
ocelli;
frons granulated and with
Fig.
9;
segments in following proportions:
10: 9:
head shiny,
400 -
smooth and
legs
tegulae
10: 5:
23:
14:
12:
without sculpture on sides of
irregular keels;
ocellar triangle rugose;
Chela of Chelothelius berlandi Bernard
(type)
610
Mem. Amer.
occipital
carina
reaching eyes;
incomplete,
Ent.
Ins!.,
smooth,
weakly
tegulae;
without keels;
granulated,
propodeum fully longitudinal basal
proportions:
long and narrow; 12:
and without
3:
5:
9:
very
16;
(Fig.
Fig.
401
400)
-
part
transversal pronotal
smooth,
(9:8),
impression
tubercles not
weakly granulated,
reticulate rugose;
posterior surface with 2 transversal
region;
fore tarsal
distal
bands,
the
vein
wing margin;
in following
tarsus produced
segment 5 of front
tarsus
lamellae extending continuously
with 7 bristles and one preapical
Female of Bocchus umber n.
incomplete
one on
part of radial
not reaching segments
segment 3 of front
long bristles;
with some rows of nearly 30 enlarged claw
dull;
the apical
longer than proximal
pterostigma
disc;
metanotum fully
fore wings with 2 dark
cells and another on
slightly
hook
reticulate rugose,
keels;
region of head,
with a
scutum and scutellum shiny,
notaulices invisible;
1984
POL = 5; Ol = 3; OOl = 8;
between anterior collar and posterior humped reaching
37,
visible only on dorsal
mandibles with 3 teeth;
pronotum shiny,
no.
sp.
to
into a
(Fig.
lamella;
(type)
400)
the apex; tibial
Bocchinae
611
spurs 1,1,2. MALE: unknown LOCUS TYP ICUS:
Plage de Saint-Raphael
TYPICAL MATERIAL: HOSTS:
(France)
in P.
unknown
DISTRIBUTION: Var),
holotype F!
Palaearctic region:
FRANCE:
Plage de Saint-Raphael
(Frejus,
P!
3. Bocchus Ashmead
Bocchus Ashmead 1893: 91. Phorbas Ashmead 1893: 90 (preoccup i ed by Phorbas Duchassa i ng and Michelotti
Fig.
402 -
1864;
syn.
proposed by Richards 1939);
type species
Male of Bocchus pedunculatus Nagy from Sarawak
612
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
Phorbas laticeps Ashmead 1893, orig. desig. Eukoebeleia R.C.L. Perkins 1905: 59 (syn. proposed by Fenton 1918b); type species Eukoebeleia mirabilis R.C.L. Perkins 1905, monotypic.
Tetradryinus Kieffer 1913c: 325 (n. syn.); type species Tetradryinus flavipes (Kieffer 1906), orig. desig. Anteonella Dodd 1913: 181 (n. syn.); type species Anteonella robusta Dodd 1913, orig. desig. Phorbasia Kieffer 1913b: 58 (syn. proposed by Richards 1939); type species Phorbasia laticeps (Ashmead 1893), orig. desig. Neoanteon Fouts 1922: 633 (syn. proposed by Nagy 1969); type species Neoanteon rubrica Fouts 1922, monotypic. Hymenodryinus Benoit 1953b: 427 (n. syn.); type species flymenodryinus seyrigi Benoit 1953b, orig. desig. Bocchus Ashmead: Nagy 1969: 321.
TYPE SPEC IES: FEMALE
Bocchus flavicollis Ashmead 1893, orig. desig.
(Figs 24,
401,
424,
425):
fully
winged
brachypterous or micropterous (Figs 424, segments;
labial
palpi
with 3 segments;
quadridentate mandibles usually
425);
(Figs 24,
401)
maxi Ilary palpi
mandibles with 1,
with a rudimentary
2,
or with 6 3 or 4 teeth;
tooth between
the two
posterior teeth (Fig. 396 A); rarely quadridentate mandibles with teeth progressing larger from anterior one to posterior; occipital carina complete; segment 5 of front tarsus with one preapical lamella, never with other lamellae (Fig. 404); enlarged claw with at least one tooth (Fig. 404);
pronotal
Fig.
tubercles reaching
403 -
tegulae;
tibial
spurs 1,1,1.
Male of Bocchus pedunculatus Nagy from Sarawak
Bocchinae
MALE
(Figs 402,
403):
fully
winged;
613
mandibles with 1, 2,
3 or 4 teeth;
quadridentate and tridentate mandibles with teeth not progressing from anterior one to posterior; wide as tegulae
(Fig.
segments;
palpi
labial
402);
larger
areolae of propodeum at least partly as
temples distinct;
with 3 segments;
cnaxillary palpi
fore wings complete;
with 6 tibial
spurs
1,1,2. DISTRIBUTION: HOSTS:
Palaearctic,
Ethiopian, Oriental,
Nearctic,
Australian
Issidae
SPECIES:
29
PALAEARCTIC SPECIES OF
BOCCHUS
Nine species are known.
KEY TO THE SPECIES
FEMALES
Species brachypterous, wi th fore-wing extending apex of propodeum ••••.•••••• 1. italicus n. sp. Species
fully
winged
(Fig.
wi thout
only
to
the
401) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 transversal bands ••••••••••••••
2
Fore-wing hyal ine,
dark
3
umbel' n. sp. 1-2 dark transversa I bands ••••••••••••••••••••• 3 Fore-wing wi th one dark transversal band •••••••••••••••••••••• 4 · . • . . • . . . . . . . . . . . . . . . . . . . . .. 2.
Fore-wi ng wi th
bands . . . . . . . . . . . . . . . . . . . . . 5 much shorter than scutellum; head black, with mandibles yellow; scutellum and propodeum black; pronotum, scutum and metanotum reddish • ••• · . . . . . . . . . . . . . . . . . . . . . . . . . .. 3. szelenyii Mo c za r Head with OOL more than twice as long as POL; scutellum nearly twice as long as metanotum; head reddish-testaceous; thorax and Fore-wing with
4
two dark
Head wi th POL not
transversal
shorter
than OOL;
propodeum black . . . . . . . . . . . . .
5
Thorax
fully
Thorax at
6
reddish ••••••••
least
8
4. slovacus Strejcek 9. parthicus Ponomarenko
partly black . . . . . . . . . . . . . • . . • . . . . . . . . . . . . . . . . • 6
5. paglianoi n. sp. reddish . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7 Head mostly reddish; frons and vertex at least partly reddish · . • . . • . . . • . . • . . • . . . . . . . . . . .. 6. scaramozzinoi n. sp. Head fully black, at most only with mandibles and and clypeus not black ••••••••••••••••••••••••••••••••••••••••••••••••••••• 8 Pronotum reddish; scutum, scutellum and metanotum black . . . . . . . . Thorax
fully black •••••••.••
Thorax partly black and
7
metanotum not
614
Mem.
Amer.
Ent.
Inst.,
7. Pronotum and
scutum reddish;
no.
37,
1984
scobiolae Nagy
scutellum and metanotum black ..•.•
8. europaeus Bernard
1v1AlES
Only known.
B. scobiolae Nagy and B. slovacus Strejcek are
the males of
I cannot propose a key
to these
two ma les because
I
have not seen
B. scobiolae.
the male of
1. Bocchus italicus n. sp.
FEMALE:
brachypterous
propodeum); occipital distally 7:
7:
carina complete; thickened;
6,5:
6:
scutum dull, pronotal dull,
(fore wings reaching
length 2,56 mm;
6:
head dull,
granulated; antennae
antenna I segmentes in following proportions:
6:
9;
POL
granulated,
=
6;
=
Ol
3;
=
OOl
10;
OPl
with notaulices complete, tegulae;
scutellum
propodeum reticulate rugose,
posterior surface;
only posterior apex of
testaceous;
frons with a median complete keel;
tubercles reaching
rugose;
fully
fore-wing
short,
11:
=
TL
7:
metanotum
longitudinal
keels on
reaching only posterior
apex of propodeum, without pterostigma; mandibles tridentate, with a rudimentary tooth between the two posterior teeth; fore tarsal segments following proportions: 46: produced
6:
into a short hook;
10:
35:
42;
segment 3 of front
enlarged claw
(Fig.
404)
with
teeth and one bristle; segment 5 of front tarsus (Fig. 404) bristles and one preapical lamella; tibial spurs 1,1,1. MALE:
in
tarsus a row of 5 with
two
unknown
lOCUS TYPICUS:
lido (Venezia,
TYP ICAl MATER IAl: HOSTS:
11:
9;
granulated;
two
not complete,
8;
posteriorly separated;
dull,
with
=
holotype F!
Italy) in CO.
unknown
OISTRIBUTION:
Palaearctic region:
ITALY:
Lido
(Venezia),
CO!
2. Bocchus umber n. sp.
FEMALE
(Figs 24,401):
testaceous; with
tergite
hairy;
winged;
almost hairless;
occipital
distinct and complete,
length 2,50-2,62 mm;
thorax and propodeum black;
1 and apex brownish;
abdomen
granulated;
fully
antennae yellow;
tegulae yellow;
head,
carina complete;
pronotum, POL = 4;
posteriorly separated;
head reddishabdomen black,
legs yellow;
body dull,
scutum and scutellum OOl = 5;
notaulices
metanotum smooth,
without
Bocch i nae
615
Ii I
I
) 1,\ I \
\
\
Fig. 404 -
sculpture;
Chela of Bocchus italicus n.
scutellum nearly
reticulate rugose,
with 2
twice as
longitudinal
sp.
long as metanotum;
(type)
propodeum
keels on posterior surface;
fore-wing
616
Mem.
Amer.
Ent.
Inst.,
no.
37,1984
,, " \"1;: - - - - - >
f
,
Fig.
I~ :.-~
,
405 -
Female of Bacchus umber n.
claw of a specimen from Sesto D:
without dark teeth;
apophysis of front
transversal
mandibles tridentate, fore tarsal
(except for distal
bands;
(type):
tarsal
A: C:
chela;
tarsus with a distal 6,1
enlarged segments;
ovoidal,
large;
tooth between the two posterior apophysis
segment 1-4 in following proportions:
apophysis):
B:
fore tarsal
segment 3.
pterostigma brown,
with a rudimentary
segment 3 of front
segment;
sp.
Fiorentino (Italy);
longer than 8:
; segment 5 of front tarsus
1,5:
1,2
(Fig.
405)
Bocchinae
with one preapical
lamella and 2 bristles;
row of 7-8 teeth and 2 bristles; MALE:
617
tibial
enlarged claw
(Fig.
405)
with a
spurs 1,1,2.
unknown
LOCUS TYP ICUS:
Bivio Castel
TYPICAL MATERIAL:
Rigone
holotype F!
(I taly)
and 1 paratype F!
in OL;
1 para type F!
in MI. HOSTS:
unknown
DISTRIBUTION: OL!
Palaearctic region:
Sesto Fiorentino (Firenze),
ITALY:
Bivio Castel
Rigone
(Perugia),
M I!
3. Bocchus szelenyii Moczar
Bocchus szelenyii Moczar 1974: 131. Bocchus szelenyii Moczar: Moczar 1983b: 206.
FEMALE:
fully
winged;
Fig.
406 -
length 3,12 mm;
head black,
with mandibles yellow;
Chela of Bocchus szelenyii Moczar (type)
618
Mem.
antennae testaceous;
black;
abdomen black; OOL = 7;
visible,
pronotum
Inst.,
scutellum shiny,
with one transversal
mandibles quadridentate, posterior teeth; 10;
punctate;
fore tarsal
enlarged claw
segment 5 of front
with some
hairy;
POL
notaulices weak,
metanotum dull,
pterostigma short and narrow;
rudimentary
406)
with a
(Fig.
406)
but
reticulate
tooth between
the two
segments 1-4 in following proportions:
(Fig.
tarsus
swollen,
carina complete;
propodeum reticulate rugose;
dark band;
with a
scutellum
petiole reddish;
near posterior margin of scutum;
not much shorter than scutellum;
fore-wing
hairy,
occipital
weakly punctate,
weakly
1984
reddish-testaceousj
head shiny,
obsolete only
smooth,
37,
propodeum black;
without sculpture;
scutum shiny,
no.
and scutum
legs testaceous;
almost complete,
rugose,
3:
En!.
metanotum reddish-testaceous;
short keels on frons, 5;
Amer.
14:
2:
row of 5 teeth and 1 long bristle;
with one preapical
lamella and 2
long
bristles; tibial spurs 1,1,1. MALE: unknown LOCUS TYP ICUS: Spring "Talyn Bi Igech" TYPICAL MATERIAL:
holotype F!
(Mongol ia)
in BU.
HOSTS: unknown DISTRIBUTION: Palaearctic region: Km E guard-house "Caganbu lag",
MONGOLIA:
Spring
"Talyn Bilgech"
Bajanchongor a imak),
(47
BU!
4. Bocchus slovacus Strejcek
Bocchus slovacus Strejcek 1964: 325. Bocchus slovacus Strejcek: Nagy 1969: 322. Bocchus slovacus Strejcek: Ponomarenko 1978: 23.
FEMALE:
fully
winged;
antennae testaceous;
length 2,48-2,75 mm;
thorax and
brown-reddish nuances; head dull, than
hairy,
twice as
long as POL;
tooth between
swollen;
twice as
rarely pronotum with
abdomen black;
occipital
scutum dull,
posteriorly separated;
long as metanotum;
narrow;
fore
407)
with a
tarsus
(Fig.
row of 6 teeth and 407)
with one apical
bristle; tibial spurs 1,1,1. MALE: fully winged; length 2,20-2,67 mm; testaceous;
tarsi
thickened,
hairy;
9:
9:
9:
8:
7:
and fore tibiae
10;
granulated,
antennal
black;
testaceous;
segments
head dull,
hairy;
granulated,
pterost i gma short
segments 1-4 in following proportions:
(Fig.
segment 5 of front
OOL more
rudimentary
metanotum and propodeum reticulate
fore-w i ngs wi th one transversa I dark band;
enlarged claw
with a
scutellum dull,
rugose;
tarsal
legs testaceous;
carina complete;
mandibles quadridentate,
the two posterior teeth;
notaulices complete, nearly
petiole testaceous;
granulated,
head reddish-testaceous;
propodeum black;
13:
3:
and
9;
1 long bristle; lamella and 2
long
mandibles brown-
antennae not distally
in following proportions:
granulated;
2:
12:
6:
8:
frons with a median complete
9:
619
Bocchinae
Fig.
keel;
=
8;
407 -
Che I a of
occipital carina complete; scutum dull,
granulated;
Bocchus slovacus Strejcek {type}
POL
=
6;
OL
=
3,5;
notaul ices complete,
OOL
=
7;
OPL
=
4;
TL
posteriorly separated;
scutellum and metanotum dull, granulated; propodeum reticulate rugose, with two longitudinal keels on posterior surface; areolae of propodeum partly broader than hyaline,
tegulae;
without dark
lanceolate;
radial
LOCUS TYPICUS:
vein curved;
Cenkov
TYPICAL MATERIAL:
abdomen
transversal
with petiole very short;
bands;
radial
pterostigma
cell
open;
long
tibial
and
spurs
fore wings narrow,
not
1,1,2.
{Czechoslovakia}
holotype F!
in PG;
3 paratypes FF
in Strejcek's
collection. HOSTS: unknown DISTRIBUTION:
Palaearctic region:
U. S. S. R.: Papanga {Russian
Romania}, PK! HUNGARY: Keleshalom, BU! CZECHOSLOVAKIA: Cenkov, PG!
BU!
Topart
{Kelebia},
BU!
Budapest,
620
Mem.
Fig.
408 -
Amer.
Chela of
En!.
Inst.,
no.
37,
1984
Bocchus paglianoi n. sp. (type)
5. Bocchus paglianoi n. sp.
FEMALE:
fully
winged;
antennae brown, black;
length 3,75 mm;
head black,
with segments 1-3 testaceous;
petiole testaceous;
abdomen black;
with mandibles yellow;
thorax and propodeum
legs testaceous,
with mid and
Bocchinae
hind coxae partly occipital
brown;
head swollen,
carina complete;
tooth between
hairy,
shiny,
mandibles quadridentate,
the two posterior teeth;
notaulices complete,
621
scutum dull,
posteriorly separated;
with posterior third rugose;
segments 1-4 in following proportions:
claw
(Fig.
408)
front
tarsus
tibial
spurs 1,1,1.
MALE:
unknown
LOCUS TYPICUS:
15:
3: 10; enlarged
2:
teeth and one long bristle;
with one preapical
Montemitro
TYPICAL MATERIAL: HOSTS:
row of 4
408)
granulated;
pterost i gma short and narrow;
fore tarsal
(Fig.
hairy,
scutellum shiny and smooth,
metanotum and propodeum reticulate rugose;
fore-wi ng with 2 transversa I dark bands; with a
granulated;
with a rudimentary
lamella and 2
segment 5 of
long bristles;
(Italy)
holotype F!
in OL.
unknown
DISTRIBUTION: NOTES:
Palaearctic region:
the species
is named
in
ITALY:
Montemitro (Campobasso),
honor of Mr.
OL!
Guido Pagliano.
6. Bocchus scaramozzinoi n. sp.
FEMALE:
fully
testaceous,
winged;
length 3,6-4,2 mm;
anterior part of frons,
head black,
with mandibles
temples and region between ocelli
and
occipital margin reddish; ventral part of head partly black and reddish; sometimes head fully reddish, except for ocellar triangle black; antennae brown,
with segments 1-2 and partly 3 testaceous;
scutum and scutellum reddish; mesopleura, tegulae basal
propectus,
testaceous;
segments
sometimes
metapleura and posterior surface of propodeum reddish; petiole testaceous;
abdomen black;
legs testaceous,
part of hind coxae and clubs of hind femora brown;
shiny,
pronotum,
metanotum and propodeum black;
in following proportions:
almost
hairless,
2 keels around orbits;
15:
8:
with a median
14:
12:
11: 9: 9: 9: 9:
longitudinal
keel
with
antennal 12;
head
on frons and with
head with numerous irregular keels among which
the surface is granulated;
only a central
region of frons
is smooth and
without areolae, keels or sculpture; occipital carina complete; ocellar triangle convex, nearly equilateral; OOL = 12; OPL = 9; TL = 12; mandibles quadridentate, shiny, keels;
with a strong pronotal
with one rudimentary
transversal
impression
tubercles reaching
tegulae;
numerous areolae or irregular keels; separated; rugose; surface;
scutellum with numerous
propodeum reticulate rugose,
pronotum hairy,
scutum shiny,
notaul ices complete, longitudinal with 2
keels;
one on basal
long and narrow,
20:
3:
4:
prox i ma I 13:
19;
with
posterior keels on posterior fore-wing
with
with 2 dark transversal
cells and another beneath the pterostigma;
part of rad i a I ve in shorter than following proportions:
irregular
hairy,
metanotum reticulate
longitudinal
median area of posterior surface almost smooth;
pterostigma very reduced, bands,
tooth;
and with numerous
(5: 7);
distal
fore tarsa I segmen ts
enlarged claw
(Fig.
409)
in
with a
622
Mem.
Fig.
Amer.
En!.
Ins!.,
no.
37,
1984
Chela of Bocchus scaramo2zinoi n.
409 -
row of 6 teeth and 2 bristles; bristles and one preapical
segment 5 of front
lamella;
tibial
tarsus
(Fig.
409)
with 2
in OL;
1 paratype F!
unknown
DISTRIBUTION:
Palaearctic region:
Sardinia), OL!
Musei
NOTES:
(type)
spurs 1,1,1.
MALE: unknown LOCUS TYPICUS: Tempio Pausania (Italy) TYPICAL MATERIAL: holotype F! and 1 paratype F! in PL. HOSTS:
sp.
the species
(Cagliari),
is named
ITALY: OL!
Tempio Pausania
(Sassari,
PL!
in honor of Mr.
Pier Luigi Scaramozzino.
7. Bocchus scobiolae Nagy
Bocchus scobiolae Nagy 1967b: 334. Bocchus scobiolae Nagy 1968: 1029.
Bocchinae
Fig. 410 -
623
Chela of Bocchus europaeus Bernard from Saint-Paul-et-Valmalle (France)
624
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
Bocchus scobiolae Nagy 1969: 322, 323.
I
haven't seen the three only known specimens of this species
females and 1 male).
I have included
Palaearctic Bacchus only on
the basis of the original
typicus: Agigea (Dobroudja, Romania).
(2
the species in the key to the
Locus
description.
I don't know where is the holotype.
8. Bacchus europaeus (Bernard)
Eukoebeleia europaea Bernard 1939: 109. Bacchus europaeus (Bernard): Strejcek 1964: 325. Bacchus europaeus (Bernard): Nagy 1969: 322. Bacchus europaea (Bernard): Abdul-Nour 1976: 277.
FEMALE: clypeus
fully
winged;
testaceous;
reddish;
pronotum and scutum reddish;
propodeum black; legs yellow; 7:
5:
6:
Bernard
5:
abdomen black,
antenna I segment
1939);
head smooth, along
sides of median keel 2 or 3 teeth;
rugose; rugose,
scutellum,
weakly brown; 1 without
hairy,
internal
weakly
occipital
metanotum and tegulae yellow-brown;
proportions: teeth
10: 5,5:
(with teeth,
granulated,
margin of eyes,
of frons;
with mandibles and
with segments 1-4 yellow-
10:
OOl more than
reticulate rugose;
carina,
long as POL;
longitudinal
keels;
on
mandibles with 1,
notaul ices distinct and complete,
scutellum with numerous
7: to
with some areolae
near occipital
carina complete;
twice as
7:
according
tridentate mandibles with a rudimentary tooth between
two posterior teeth; separated;
head black,
antenna I segments in following 7;
near clypeus,
shiny,
length 3,25 mm;
antennae reddish-brown,
the
scutum hairy, posteriorly
metanotum reticulate
scutellum nearly twice as long as metanotum; propodeum reticulate with 2 longitudinal keels on posterior surface; fore-wing with 2
dark transversa I bands;
pterost i gma short and narrow;
fore tarsa I
segments 1-4 in following proportions: 14: 2: 3: 10; segment 5 of front tarsus (Figs 410, 411) with a preapical lamella and 2 bristles; enlarged claw 1,1,1. MALE:
(Figs 410,
411)
with a row of 4-5
teeth and 1-2 bristles;
tibial
unknown
lOCUS TYPICUS:
St-Aygulf
TYPICAL MATERIAL:
(France)
holotype F!
in P.
HOSTS: in France Caliscelis sp. (Abdul-No'Jr 1976). DISTRIBUTION: Palaearctic region: FRANCE: St-Aygulf Paul-et-Valmalle (Herault),
(Var),
P!
9. Bacchus parthicus Ponomarenko
P!
Saint-
spurs
Bocchinae
B
625
A / /
/
/
/
Fig.
411
-
Bacchus eurapaeus Bernard (type): A: chela; B: fare tarsal segments.
Bacchus parthicus Ponomarenko 1979b: 166.
FEMALE:
fully
winged;
length 3,60 mm;
fully
reddish,
triangle brown,
posterior margin of scutellum brown,
abdomen brown;
antennae brown,
with occipital propodeum darkened,
with segments 1-2 yellow;
antennae
626
Mem.
Fig.
412 -
Amer.
Ent.
Ins!.,
no.
37,
1984
Chela of Bocchus parthicus Ponomarenko
(type)
weakly distally thickened; antenna I segments in following proportions: 10: 20: 15: 15: 15: 14: 13: 13: 15; mandibles quadridentate, with a rudimentary
tooth between
the two posterior teeth;
head shiny,
20:
punctate;
Bocch i nae
frons with a occipital
median complete keel
carina complete;
pronotum shiny, reaching
hairy,
tegulae;
and
= 7;
POL
smooth,
627
with 2 keels around orbits;
= 7;
OL
=
OOL
weakly punctate;
pronotum crossed by
13;
a transversal
=
OPL
pronotal
16;
TL
=
18;
tubercles
impression;
scutum
shiny, hairy, weakly punctate, with notaulices complete and posteriorly separated; scutellum and metanotum shiny, smooth, weakly punctate; propodeum reticulate rugose, strongly one on long
the basal
and
tarsal
(Fig.
1,1,2. MALE:
with 2 brown
fore-wing
cells and another beneath
narrow,
segments
front tarsus claw
with posterior surface not reticulate rugose,
transversely striate; not
lanceolate;
radial
vein
in following proportions:
(Fig.
412)
412)
with a
regularly curvilinear;
lamella and 2 bristles;
with a row of 6 teeth and
bands,
pterostigma fore
5: 8: 25: 34; segment 5 of
33:
preapical
transversal
the pterostigma;
1 long bristle;
tibial
enlarged spurs
unknown
LOCUS TYPICUS:
Tashauz
TYPICAL MATERIAL: HOSTS:
(Turkmenistan,
holotype F!
U.
S.
S.
S.
R.:
R.)
in LN.
unknown
DISTRIBUTION:
Palaearctic region:
U.
S.
Tashauz
(Turkmenistan),
LN!
ETHIOPIAN SPECIES OF
BaCCHUS
Three species are known
KEY TO THE SPECIES
FEMALES
Micropterous;
fore-wing
nearly
as
long as
• • . . • . . . . . . . . . . • • • • . • • . . . • . • . . • . .. -
Fully
2
Segment
1 of
414 B);
antennae
least -
tegulae . . . . . . . . . . . . . . .
richardsi n. sp.
winged . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2
front
twice as
Segment
of
antennae nearly
1.
as
tarsus nearly
longer and more
long
front
shorter,
1,5
times as
slender,
as broad ••.•.• tarsus nearly as
2.
MALES
3.
4
segments
(Fig. 5-9 at
bini n. sp.
long as
distally more broadened,
long as broad •••••••••••
long as
wi th 4
(Fig.
with
415 A);
segments
seyrigi (Benoi t)
5-9
628
Mem.
Amer.
En!.
Inst.,
no.
37,
1984
Unknown
1. Bocchus richardsi n. sp.
FEMALE: 3,31
15: 6:
9:
areolae, lateral 10;
micropterous,
mm;
with
fore-wing nearly
fully reddish-dark; 6:
6:
6:
with a
5:
5:
median
5:
8;
keel
keels around orbits;
occipital
antennal
as
segments
head shiny,
long as tegulae;
length
in following proportions:
granulated and with
weak
joining anterior ocellus to clypeus and with 2 POL = 5;
carina complete;
OL = 3;
pronotum shiny,
OOL = 12; hairy,
OPL = 8;
weakly
TL =
granulated,
with a transversal impression between anterior and posterior collar; pronotal tubercles reaching tegulae; scutum dull, granulated, with weak areolae and strong
longitudinal
with
longitudinal
keels;
fully
reticulate rugose;
keels;
notaulices invisible;
metanotum very short and reduced; posterior surface without
scutellum dull, propodeum
longitudinal
keels;
fore
tarsal segments in following proportions: 15: 3: 3: 11: 14; enlarged claw (Fig. 413) with a row of 9 teeth and one bristle; segment 5 of front tarsus
(Fig.
413)
with a
preapical
lamella and a
row of 8 bristles;
spurs 1,1,1.
(\
Fig.
413 -
Chela of
Bocchus richardsi n. sp. (type)
tibial
Bocchinae
MALE:
unknown
LOCUS TYPICUS:
Milnerton
TYPICAL MATERIAL: HOSTS:
(South Africa)
holotype F!
in BM;
1 paratype F!
in WA.
unknown
DISTRIBUTlmJ: BM!
629
Ethiopian region:
Cape Town,
NOTES:
SOUTH AFRICA:
Milnerton
(Cape Town),
WAr
the species
is named
in
honor of Mr.
Owain W.
Richards.
2. Bocchus bini n. sp.
FEMALE: black
fully
winged;
length 3,12-4,06 mm;
head fully reddish or fully
(with mandibles and sometimes also clypeus testaceous);
occasionally
head black with region between ocell i and occipital carina reddish; thorax fully reddish; propodeum fully reddish or fully black; occasionally partly black
and reddish;
antennae fully
testaceous or brown,
legs testaceous;
reticulate rugose;
sometimes not reticulate rugose on frons on sides of
median keel,
where the surface is granulated;
ocellar triangle equilateral; rugose,
abdomen black;
with segments 1-2
or 1-3 testaceous;
hairy,
shiny;
pronotum hairy;
notaulices
head du II,
occipital
ha i ry,
fu II y
carina complete;
scutum fully reticulate
invisible or slightly visible;
scutellum
and metanotum reticulate rugose; propodeum reticulate rugose, with 2 longitudinal keels on posterior surface; fore-wing with 2 dark transversal bands; pterostigma short and narrow; fore tarsal segments 1-4 in following proportions: preapical
13: 2:
2:
8;
5-6 teeth and one bristle; MALE:
segment 5 of front
lamella and 2 bristles; tibial
tarsus
enlarged claw
(Fig.
(Fig.
414)
414)
with one
with a row of
spurs 1,1,1.
unknpwn
LOCUS TYP ICUS:
Afgoi
TYPICAL MATERIAL: in BM;
(Somal ia)
holotype F!
1 para type F!
and 3 para types FF!
in OL;
1 paratype F!
in OT.
HOSTS: unknown DISTRIBUTION: Ethiopian region:
SOMALIA:
Afgoi
(Benadir),
OL!
Ibadan, OT! Ikenne (\'I-State), BM! GHANA: Kwadaso (Kumasi), NOTES: the species is named in honor of Mr. Ferdinando Bin.
NIGERIA: OL!
3. Bocchus seyrigi (Benoit) n. comb.
Hymenodryinus seyrigi Benoi t 1953b: 427.
FEMALE:
fully
antennal
segments 3-10 brown,
winged;
reticulate rugose,
length 4,62 mm;
hairy;
reddish,
with ocellar spot brown,
abdomen black-brown;
antennae short,
head dull,
distally broadened,
fully
with
630
Mem.
Amer.
Ent.
Ins!.,
no.
37,
1984
, >
'
\ !
,
'
~
'
,
'0\ \\ \
\
B
"
\
\
Fig.
414 -
Bocchus bini n. sp. B:
segments 5-9 nearly proportions:
18:
7:
as 15:
(paratype from
fore tarsal
long as broad; 7:
5,5:
5:
6:
6:
Ibadan,
Nigeria);
A:
chela;
segments.
antenna I segments 5:
7;
occipital
in following
carina complete;
Bocchinae
Fig.
415 -
Bocchus seyrigi
(Benoit) (type);
631
A:
fore tarsal
segments;
B:
chela.
ocellar triangle equilateral; fully
reticulate rugose;
with 2
longitudinal
scutum,
scutellum and metanotum dull,
notaulices invisible;
hairy,
propodeum reticulate rugose,
keels on posterior surface;
fore-wing
with 2 dark
632
Mem.
transversa I bands;
Amer.
12: 2:
LOCUS TYPICUS:
3:
12;
37,
Behara
1984
fore tarsa I segments 1-4
segment 5 of front
teeth and one bristle;
TYPICAL MATERIAL: HOSTS:
no.
lamella and 2 bristles;
with one preapical
with a row of 11 MALE: unknown
Inst.,
pterost i gma short and narrow;
in following proportions: 415)
En!.
tibial
tarsus
enlarged claw
(Fig.
(Fig.
415)
spurs 1,1,1.
(Madagascar)
holotype F!
1 paratype F in TV.
in P;
unknown
DISTRIBUTION: 1953b) •
Ethiopian region:
MADAGASCAR:
Behara,
P!
3ekily
(Benoit
OR I ENTAL SPEC I ES OF BOCC/JUS
Four species are known.
KEY TO THE SPECIES
FEMALES
-
Fore-wing with Fore-wing with
2 Fore-wing of A);
front
with
1 dark
transversal transversal
band •••••
2 dark
radial
vein
curved
tarsus without
enlarged
weakly
a distinct
claw with a
row of
thorax and propodeum black
some
(except
(Fig.
proximal teeth
for
Fore-wing with segment
5 of
radial
front
(Figs 419,422); 422);
head,
vein more
tarsus
enlarged
wi th
strongly a
416 A);
prominence (Fig.
2.
418);
(Fig.
di st inct proximal
claw with
segment (Fig.
5
416
head,
muluensis n.
curved
1 only
thorax and propodeum reddish
tooth
sp.
416 B);
prominence
(Figs
419,
or brown-testaceous
••..••.••••••••••.•••...••••....•.....•........•..........•..•• 3 much longer than POL; head, thorax and propodeum redddish-brown •• •••••••••••••••••......•..• 3. pedunculatus Nagy OL more than twi ce as long as POL; head, thorax and propodeum reddi sh-I ight .••.•••••••.••••••••.•••..•••• 4. rubricus (Fouts)
3 OL not -
laotianus n. sp.
the yellow mandibles) .•••
. . . • • . • • . . . . • . . . • • • • • • • . • . • . . • • • • . . . . . . . . .. -
1.
bands • • • . . • . . . . . . . . . . . • . . . . . • 2
fAALES
Only
the male of B.
pedunculatus Nagy is known.
Bacchinae
633
A
,
,
E
Radial veins of females of Bacchus muluensis n. sp. (type)(A), pedunculatus Nagy from Tagembung (Phi I ippines) (B), laatianus n. sp. (type)(C), guineensis n. sp. (type)(D), australiae n. sp. (type)(E).
Fig. 416 -
634
Mem.
Amer.
Ent.
lnst.,
no.
37,
1984
i
i~
() /
/J 1/ I /
Fig. 417 -
Chela
0
f Bocchus laotianus n.
sp.
( type)
i
Bocchinae
635
1. Bocchus laotian us n. sp.
FEMALE:
fully
winged;
length 3,43-3,87 mm;
with segments 1-2 or 1-3 yellow; legs yellow;
head dull,
hairy,
thorax,
weakly granulated,
except for the sides of the ocelli rugose;
occipital
tooth between
equilateral;
scutum shiny,
granulated,
scutellum shiny,
punctate,
twice as
with
1 dark
transversal
posteriorly separated;
reticulate rugose only near posterior margin,
long as metanotum; with 2
ocellar triangle
reticulate rugose near
notaulices distinct and complete,
nearly
with a
mandibles quadridentate,
anterior margin;
reticulate rugose,
reticulate rugose,
the two posterior teeth; hairy,
antennae brown,
where the surface is not reticulate
carina complete;
rudimentary
head black;
propodeum and abdomen black;
metanotum reticulate rugose;
longitudinal band;
propodeum
keels on posterior surface;
pterostigma short and narrow;
fore-wing
radial
vein
weakly curved (Fig. 416 C); fore tarsal segments 1-4 in following proportions: 20: 3: 3: 14; segment 5 of front tarsus (Fig. 417) with a preapical
lamella and 2 bristles;
10 teeth and MALE:
1 bristle;
enlarged claw
(Fig.
417)
with a
row of
spurs 1,1,1.
unknown
LOCUS TYPICUS:
Ban Van Eue
TYPICAL MATERIAL: HOSTS:
tibial
(Laos)
holotype F!
1 para type F!
in B;
in OL.
unknown
DISTRIBUTION:
Oriental
region:
LAOS:
Ban Van Eue
(Vientiane),
B!
OL!
2. Hocchus muluensis n. sp.
FEMALE:
fully
winged;
segments 9-10 brown; legs testaceous,
length 3,25 mm;
head black;
thorax and propodeum black;
with hind coxae brown;
head dull,
antennae yellow, abdomen hairy,
with
black-brown; reticulate
rugose; frons narrow, very converging towards the clypeus; in front of the anterior ocellus the frons is nearly as broad as each eye;ocellar triangle equilateral;
occipital carina complete;
mandibles with 4 teeth,
larger from
anterior one to posterior;
scutum reticulate rugose,
and with numerous metanotum hairy, 2
longitudinal
transversal short and 3:
12;
bristle;
longitudinal
dull,
keels;
notaul ices invisible;
weakly punctate;
keels on posterior surface;
bands; narrow;
radial
vein
fore tarsal
segment 5 of front enlarged claw
propodeum reticulate rugose, fore-wing
weakly curved
(Fig.
(Fig.
418)
418)
with a
with a
Gunong Mulu National
TYPICAL MATERIAL:
holotype F!
and
416 A); preapical
row of 7 teeth and
Park
(Sarawak,
1 paratype F!
hairy with
with 2 dark pterostigma
1 long
Malaysia)
in 8M.
14:
lamella and
bristle; tibial spurs 1,1,2. MALE: unknown LOCUS TYPICUS:
dull,
scutellum and
segments 1-4 in following proportions:
tarsus
(Fig.
progressing
3: 1
636
Mem.
Fig.
HOSTS:
418 -
Amer.
Ent.
Inst.,
no.
37,
1984
Chela of Bocchus muluensis n.
sp.
(type)
unknown
DISTRIBUTION: District,
Oriental
Sarawak),
region:
MALAYSIA:
Gunong Mulu Nat.
Park
(Baram
BM!
3. Bocchus pedunculatus Nagy
Bocchus pedunculatus Nagy 1969: 323. Bocchus rubricus Fouts 1922: Nagy 1969: 324.
FEMALE: fully winged; length 2,18-3,20 mm; reddish-brown, with antennae yellow (with segments 9-10 or only 10 brown); head shiny, smooth, weakly punctate,
sometimes with
complete;
mandibles quadridentate,
two posterior teeth; weakly
punctate,
and distinct,
transversal
OL not much
keels on frons;
longer than
POL;
carina
tooth between
scutum shiny,
without sculpture among punctures;
posteriorly separated;
occipital
with a rudimentary
the
smooth,
notaulices complete
scutellum and metanotum shiny,
smooth, weakly punctate; propodeum reticulate rugose, with 2 lateral points and with 2 longitudinal keels on posterior surface; fore-wing with 2
Bocchinae
Fig.
419 -
637
Chela of Bocchus pedunculatus Nagy from Gunong Mulu (Sarawak)
dark transversal strongly curved proportions:
bands; (Fig.
13: 3:
3:
pterostigma short and narrow;
416 B);
fore tarsal
radial
vein
segments 1-4 in following
11; segment 5 of front tarsus (Fig. 419) with a
638
Mem. Amer.
Fig.
420 -
Ent.
Inst.,
no.
37,
1984
Male of Bocchus pedunculatus Nagy from Gunong Mulu
(Sarawak): A:
ninth abdominal sternite;
B:
genitalia.
preapical lamella, 2 bristles and a proximal distinct prominence; enlarged claw (Fig. 419) with 1 tooth and 1 long bristle; tibial spurs 1,1,2. MALE (Figs 402, 403): fully winged; length 2,60-3,00 mm; head black, with mandibles and sometimes clypeus testaceous; antennae yellow, with segments 6-10 brown; thorax, propodeum and petiole black or brpwn;
legs
testaceous, with coxae blackish; head hairy, dull, fully reticulate rugose; occipital carina complete; ocellar triangle equilateral OPL = OOL; mandibles quadridentate, wite. a rudimentary tooth between the two posterior teeth; scutum dull, hairy, fully reticulate rugose; notaulices distinct and complete, posteriorly separated; scutellum and metanoturn dull, weakly punctate, hairy; propodeum reticulate rugose, with 2 longitudinal keels on posterior surface and with 2 lateral points (as in female); forewing with 2 transversal darkened bands; radial vein strongly curved; pterostigma short and narrow, lanceolate, 420; abdomen with petiole very long. LOCUS TYPICUS: Pinigisan (Philippines) TYPICAL MATERIAL: holotype M! in CO. HOSTS: unknown
hyaline;
male genitalia:
fig.
DISTRIBUTION: Oriental region: MALAYSIA: Gunong Mulu (Baram District, Sarawak), OU BM! PHILIPPINES: Pinigisan (Mantalingajan, Palawan), CO! Tagembung (Mantalingajan, Palawan)(Nagy 1969: under name Bocchus rubricus Fouts), CO!
Bocchinae
Fig. 421
-
Female of
639
Bocchus rubricus (Fouts)(type)
4. Bocchus rubricus (Fouts)
Neoanteon rubrica Fouts 1922: 634. Neoanteon rubrica Fouts: Baltazar 1966: 205. nec Bocchus rubricus (Fouts): Nagy 1969: 324 (see Bocchus pedunculatus Nagy).
FEMALE
(Fig. 421):
fully winged;
length 3,0-4,0 mm;
reddish,
with
antenna I segments 1, 2, 3, 7, 8 and part of 9 yellow; head almost hairless, shiny, smooth, without sculpture, weakly punctate; OL more than twice as long as POL; occipital carina complete; mandibles quadridentate, with a rudimentary tooth between the two posterior teeth; scutum weakly punctate, almost hairless, smooth, shiny; notaulices weak, incomplete, not visible in posterior third of scutum; scutellum and metanotum smooth, shiny,
weakly punctate;
propodeum reticulate rugose,
with 2 longitudinal
640
Fig.
Mem.
422 -
Amer.
Ent.
Inst.,
Female of Hocchus rubricus tarsal
no.
(Fouts)(type):
strongly curved; 3:
12;
bands;
fore tarsal
segment 5 of front
Mt.
points on sides;
tibial
Makiling
(Fig.
422)
prominence;
with a preapical
unknown
fore
fore-wing radial
enlarged claw
lamella, (Fig.
(Philippines)(Los Banos according in WA.
vein
15:
422)
spurs 1,1,1.
original description; Mt. Makiling on type label). TYPICAL MATERIAL: holotype F! and 1 paratype F! HOSTS:
B:
segments 1-4 in following proportions:
bristles and a distinct proximal
LOCUS TYPICUS:
chela;
pterostigma short and narrow;
tarsus
1 tooth and 1 long bristle; MALE: unknown
A:
segments.
keels on posterior surface and with 2 lateral with 2 dark transversal
37,1984
to the
4: 2 with
641
Bocchinae
DISTRIBUTION: Los Banos
Oriental
(Luzon),
region:
PHILIPPINES:
Mt.
Makiling
(Luzon),
WA!
WA!
NEARCTIC SPECIES OF
BaCCHUS
Eight species are known
KEY TO THE SPECIES
FEMALES
Species micropterous -
Species
fully
(Figs
424,
425) .•••••••••••••••••••••••••• 2
winged . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3
2 Fore-wing less reduced, reaching approximately the keel between anterior and posterior surface of propodeum; notaul ices complete, sometimes slightly visible •••••••••••••••••••••••••••• 1. mirabilis (R.C.L. Perkins - Fore-wing more reduced, reaching approximately anterior margin of propodeum (Figs 424,425); notaulices invisible . . . . . . . . . . . . . . . • • . . . • . • • • . . . . . . . • . . . . • . .. 2. testaceus n. sp. 3 Species fully testaceous . . . 3. wasbaueri n. sp. - Species at least partly black . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 4 Thorax partly black . . . . . . . . 6. flavicollis Ashmead - Thorax fully black ••••••••••••••••••••••••••••••••••••••••••••• 5 5 Scutum dull, granulated . . . . 4. arizonicus (R.C.L. Perkins) - Scutum shiny, weakly punctate, wi thout sculpture among punctures . . . • . . . . . . . . . . . . . . . . . . • . . . . 5. flavipes Kieffer
The females of B.
laticeps (Ashmead) and dubius n. sp. are unknown
MALES
Scutum dull, -
fully granulaterJ;
head granulated,
rugose . . . . . . . . . . . . . . . . . . . . . 7. laticeps (Ashmead) Scutum shiny, without sculpture; head reticulate g ranu I at ed. • • • • • • • • • • • • • • ••
The males of
8.
not
reticulate
rugose and
dubius n. sp.
B. mirabilis (R.C.L. Perkins), testaceus n. sp.,
642
Fig.
Mem. Amer.
423 -
Ent.
Inst.,
no.
37,
Chela of Bocchus mirabilis (R.C.L. (Ohio)
1984
Perkins) from Columbus
wasbaueri n. sp. flavicollis Ashmead, arizonicus (R.C.L. Perkins) and flavipes Kieffer are unknown. Probab I y B. laticeps (Ashmead) and dub ius n. sp.
are the opposite sex of the other six species.
Bocchinae
643
1. Bocchus mirabilis (R.C.L. Perkins)
Eukoebeleia mirabilis R.C.L. Perkins 1905: 59. Eukoebeleia mirabilis R.C.L. Perkins: Kieffer 1914b: 47. Phorbas mirabilis (R.C.L. Perkins): Fenton 1918b: 261. Bocchus mirabilis (R.C.L. Perkins): Richards 1939: 189. Bocchus mirabilis (R.C.L. Perkins): Nagy 1969: 322.
FEMALE:
micropterous,
with fore-wing reduced and extending beyond the
keel
between anterior and posterior surface of propodeum;
mm;
usuallyfully testaceous with antenna I segments 4-10 black or brown;
occasionally head reddish fully black;
with brown ocellar spot;
occasionally antennae brown,
pronotum and scutum usually reddish, scutellum,
head dull,
rudimentary tooth between complete;
rarely head almost
with segments 1-2 or 1-3 yellow;
rarely
metanotum and propodeum black;
tegulae testaceous;
granulated;
almost fully black;
abdomen brown-black;
the two posterior teeth;
notaulices distinct and complete,
scutellum dull,
as sculptured as scutum;
scutellum nearly twice as
legs and
mandibles quadridentate, occipital
OOL nearly three-four times as long as POL;
granulated;
length 2,50-2,92
with a
carina
scutum dull,
posteriorly separated;
metanotum reticulate rugose;
long as metanotum;
propodeum reticulate rugose,
with 2 longitudinal keels on posterior surface; fore reduced wings without pterostigma; fore tarsal segments 1-4 in following proportions: 23: 4: 6: 17; segment 5 of front tarsus (Fig. 423) with a preapical lamella and 2 bristles; enlarged claw (Fig. 423) with a row of 6-7 teeth and 1 bristle; tibial
spurs 1,1,1.
MALE:
unknown
LOCUS TYPICUS:
Ohio
(U.
S. A.)
TYPICAL MATERIAL: lectotype F! and 1 para lectotype F! in B. HOSTS: in U. S. A. Bruchomorpha oculata Newm. (Issidae) (Fenton 1918b). DISTRIBUTION: Place (Ont.),
Nearctic region: Ol!
OT!
Ont.),
OT!
U. S. A.:
Refugio
Tex.),
BY!
Ol!
B!
Ohio,
CANADA:
Grenadier
I.
(Bee Co.,
Columbus
Foremost
Centre (St. Tex.),
(Ohio),
(Alta.),
Lawrence BY!
Waco
OT! Is.
Carleton Nat.
Park,
(Mc lennan Co.,
BM!
2. Bocchus testaceus n. sp.
FEMALE
(Figs 424,
425):
micropterous,
only anterior margin of propodeum;
with fore-wing reduced and reaching
length 2,01
mm;
fully
testaceous;
head
dull, granulated; occipital carina complete; mandibles quadridentate, with a rudimentary tooth between the two posterior teth; OOL nearly four times as long as POL;
scutum shiny,
scutellum shiny,
without sculpture,
weakly granulated;
notaulices invisible;
nearly twice as long as metanotum;
644
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
I 424 - Female of Bocchus testaceus n. sp.
Fig.
metanotum reticulate rugose;
propodeum reticulate rugose,
longitudinal
keels on posterior surface;
pterostigma;
fore tarsal
enlarged claw
with 2
fore reduced wings without
segments 1-4 in following proportions:
6; segment 5 of front tarsus (Fig. 426) with a preapical bristles;
(type)
(Fig.
426)
11:
1,5: 2:
lamella and 2
with a row of 8 teeth and 2 bristles;
tibial spurs 1,1,1. MALE: unknown LOCUS TYPICUS:
Nogales (U.
TYPICAL MATERIAL: HOSTS:
S.
holotype F!
A.) in OL.
unknown
DISTRIBUTION:
Nearctic region:
U. S. A.: Nogales (Arizona), OL!
3. Bocchus wasbaueri n. sp.
FEMALE:
fully
winged;
length 3,12 mm;
fully
testaceous,
ocellar spot and antenna I segments 4-10 darkened; occipital
carina complete;
tooth between
mandibles quadridentate,
the two posterior teeth;
with brown
head dull,
granulated;
with a rudimentary
OOL nearly five times as long as
Bocchinae
Fig.
POL;
425 -
scutum dull,
separated,
645
Female of Bocchus testaceus n.
granulated;
scutellum dull,
transversa I band;
2:
3:
8;
fore-wing
tibial
with a
dark
fore tarsa I segments 1-4
segment 5 of front
lamella and 1 bristle;
row of 7 teeth and 1 bristle; MALE:
propodeum reticulate rugose,
pterost i gma short and narrow; 14:
posteriorly
metanotum reticulate rugose;
keels on posterior surface;
in following proportions: with a preapical
(type)
notaulices complete and distinct,
granulated;
scutellum nearly twice as long as metanotum; with 2 longitudinal
sp.
enlarged claw
tarsus
(Fig.
(Fig.
427)
427)
with a
spurs 1,1,1.
unknown
LOCUS TYPICUS:
Uvalde (Texas,
TYPICAL MATERIAL: HOSTS:
U.
S.
A.)
in SC.
unknown
DISTRIBUTION: NOTES:
holotype F!
Nearctic region:
the species is named
U.
S.
A.:
Uvalde (Tex.),
in honor of Mr.
4. Bacchus arizonicus
Marius S.
(R.C.L.
SC!
Wasbauer.
Perkins)
Eukoebeleia arizonica R.C.L. Perkins 1907: 44. Eukoebeleia arizonica R.C.L. Perkins: Kieffer 1914a: 47. Piwrbas arizonica (R.C.L. Perkins): Fenton 1918b: 263. Bacchus arizonica (R.C.L. Perkins): Nagy 1969: 322.
FEMALE: yellow;
fully
winged;
length 3,11-3,31
antennae brown,
mm;
head black,
with mandibles
with segments 1-2 or 1-3 testaceous;
thorax and
propodeum black; legs and tegulae testaceous; head dull, granulated, numerous areolae near clypeus, along orbits, near occipital carina; occipital carina complete; tooth between
mandibles quadridentate,
the two posterior teeth;
with
with a rudimentary
OOL nearly three times as long as
646
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
, ,
,-, I
(
(
\
'-
\
\
\
\
\
\
\
\
..i
"
\
\:/.;I--~;~)
s:;:::..- --.. . 'S. I
.-//
Fig.
426 -
Female of
Bocchus testaceus segments;
POL;
scutum dull,
granulated;
B:
n.
sp.
(type):
A:
fore tarsal
chela.
notaulices distinct and complete,
posteriorly
separated; scutellum shiny, punctate, nearly twice as long as metanotum; metanotum reticulate rugose; propodeum reticulate rugose, with 2 longitudinal band;
keels on posterior surface;
pterostigma short and narrow;
proportions:
31:
5:
6:
18;
fore-wing
fore tarsal
segment 5 of front
with 1 dark transversal segments 1-4 in following
tarsus
(Fig. 428)
with a
Bocchinae
647
"'-"- .....
\ \
Fig. 427 -
Chela of Bacchus wasbaueri n. sp.
(type)
preap ica I I ameli a and 1-2 brist les; en I arged cl aw (F i g. 428) wi th a raw of 6-8 teeth and no or 1 bristle; tibial spurs 1,1,1. MALE: unknown LOCUS TYP ICUS: Nogales (Ariz., U. S. A.) TYPICAL MATERIAL: holotype F! in B. HOSTS: in U.S.A. Bruchamorpha sp. {Issidae} (Fenton 1918b). DISTRIBUTION: Nearctic region: U. S. A.: Nogales (Ariz.), B! Uvalde
648
Mem. Amer.
Fig.
428 -
Ent.
Inst.,
no. 37,
1984
Chela of Bocchus arizonicus
(R.C.L.
Perkins)
(Texas)
from Uvalde
Bocchinae
Fig. 429 -
Chela of
649
Bocchus flavipes Kieffer from Almanor (California)
650
Mem.
(Tex.),
Amer.
En!.
Ins!.,
no.
37,
1984
OL!
5. Bocchus flavipes Kieffer
Bocchus flavipes Kieffer 1906a: 237. Tetradryinus flavipes (Kieffer): Kieffer 1913c: 325. Tetradryinus flavipes (Kieffer): Kieffer 1914b: 48. Tetradryinus flavipes (Kieffer): Richards 1939: 189. Tetradryinus flavipes (Kieffer): Muesebeck and Walkley 1951: 1035. Tetradryinus flaviceps (Kieffer): Krombein 1979: 1243.
FEMALE:
fully
winged;
antennae brown, testaceous; proportions:
length 3,37-3,56 mm;
antennae distally thickened; 10: 8:
12:
11:
rugose and with numerous with two OL = 3; strong
lateral
black;
mandibles testaceous;
with segments 1-2 or 1-5 testaceous; 9:
8:
7,5:
OOL = 10;
6:
6:
irregular keels;
keels around orbits; POL = 10;
transversal
antennal
head dull,
carina complete;
pronotum shiny,
pronotal
legs
reticulate
frons with a median keel
occipital
TL = 10;
impression;
11;
abdomen brown;
segments in following
hairy,
tubercles reaching
and
POL = 3; with a
tegulae;
scutum
sh i ny, smooth, weak I y punctate, wi thoul scu I pture among punctures, weakly rugose near anterior margin; notaulices complete, posteriorly separated; scutellum shiny, smooth, weakly punctate, without sculpture among punctures, weakly rugose near posterior margin; metanotum dull, granulated, with some keels; propodeum with broad areolae; posterior surface of propodeum with
two longitudinal
keels;
among numerous broad areo I ae;
fore-wi ng
beneath
part of radial
part
the pterostigma;
(6:9);
fore tarsal
enlarged claw front
tarsus
(Fig.
(Fig.
distal
segments in following
429)
429)
with a
small
areolae are visible
wi th 1 dark transversa I band vein shorter than proportions:
20:
4:
row of 6 teeth and 1 bristle;
with a preapical
proximal 5:
17: 25;
segment 5 of
lamella and 8 bristles;
tibial
spurs 1,1,1. MALE: unknown LOCUS TYPICU5: Ormsby Co. (Nevada, U. TYPICAL MATERIAL: holotype F! in CA. HOSTS:
S.
A.)
unknown
DISTRIBUTION: Ormsby Co.
Nearctic region:
(Nev.),
U.S. A.: Almanor
{Plumas Co.,
Calif.),OT!
CA!
6. Bacchus flavicollis Ashmead
Bocchus flavicollis Ashmead 1893: 91. Bocchus flavicollis Ashmead: Kieffer in Kieffer et Marshall 1905: 67.
Bocchinae
Fig.
430 -
Chela of
651
Bocchus flavicollis Ashmead (type)
Bocchus flavicollis Ashmead: Kieffer 1914b: 45. Bocchus flavicollis Ashmead: Muesebeck and Walkley 1951: 1039. Bocchus flavicollis Ashmead: Krombei n 1979: 1243.
FEMALE:
fully
winged;
length 2,80-2,81
mm;
head reddish,
with a broad
brown spot on frons and vertex; mandibles and clypeus testaceous; prothorax reddish; scutum reddi sh or reddi sh-brown; scutell um, metanotum, pleura and propodeum black; testaceousj
sometimes scutellum partly reddish;
abdomen brown-testaceousj
legs testaceousj
petiole
antennae missing in
652
Mem.
holotype;
head dull,
rudimentary
flat,
complete;
separated;
12:
5:
12: 9:
scutellum smooth,
transversal
9:
8:
19: 3:
4:
1984
shiny,
10;
part;
carina
pronotum dull,
pronotal
tubercles
notaulices complete,
weakly granulated; fore-wing
distal
fore tarsal
12:
16; enlarged claw
lamella;
segment 5 of front tibial
frons with a
occipital
TL = 9;
impression;
very
with a
antenna I segments in
8:
OPL = 7;
granulated;
preapical lamella and 2 bristles; MALE: unknown
posteriorly
metanotum
with 1 dark
part of radial
vein
segments in following (Fig.
430)
tarsus
with a row of 7
(Fig.
430)
with a
spurs 1,1,1.
LOCUS TYPICUS: Marquette (Michigan, U. TYPICAL MATERIAL: holotype F! in WA. HOSTS:
9:
the pterostigma;
nearly as long as proximal teeth and 1 long
9:
propodeum reticulate rugose;
band beneath
proportions:
37,
keels around orbits;
transversal
scutum dull,
reticulate rugose;
no.
mandibles quadridentate,
OL = 2,5; OOL =7;
with a strong
tegulae;
Ins!.,
the two posterior teeth;
and with 2 lateral
POL = 2;
granulated, reaching
Ent.
granulated;
tooth between
following proportions: median keel
Amer.
S.
A.)
unknown
DISTRIBUTION:
Nearctic region:
Marquette (Mich.),
CANADA:
Nerepis (N.
B.),
u.
WA!
S.
A.:
WA!
7. Bocchus laticeps (Ashmead)
Phorbas Bocchus Bocchus Bocchus
FEMALE: MALE:
laticeps laticeps laticeps laticeps
Ashmead 1893: 90. (Ashmead):
Muesebeck and Walkley 1951:
(Ashmead):
Nagy 1969: 323.
(Ashmead):
Krombein
1979:
1040.
1243.
unknown
fully
testaceous;
winged;
length 2,32-2,50 mm;
antennae brown;
thorax,
head black,
propodeum,
with mandibles
pet iole and abdomen
black; legs black; tegulae yellow; head dull, almost hairless, granulated; antennal segments in following proportions: 11: 5: 7: 7: 7: 7: 7: 7: 7: 9; POL = 6;
Ol = 3;
OOl = 8;
OPl = 3;
Tl = 5;
notaulices longer than breadth of ocelli hairless; smooth,
notaulices complete, weak I y
punctate;
scutum dull,
posteriorly separated;
smooth and
propodeum reticulate rugose,
keels on posterior surface;
the
granulated,
scutellum shiny,
metanotum rugose on anterior ha If,
weakly punctate on posterior half; longitudinal
minimum distance between
(7:3);
median surface smooth,
with 2 without
sculpture, except for regions near keels which are rugose; internal surface of areolae rugose; fore-wing hyal ine, without dark transversal bands; pterostigma hyaline, oval, with distal genital ia:
part fig.
with only posterior margin brown;
longer than proximal
431;
tibial
spurs 1,1,2.
part;
radial
vein
abdomen with petiole short;
Bocchinae
Fig.
431
-
Bocchus laticeps (Ashmead}(type): A: ninth abdominal
Male of sternitej
LOCUS TYPICUS:
DISTRIBUTION: (Tex.), OLl
B:
genitalia
(right gonoforceps removed).
Jacksonville (Florida,
TYPICAL MATERIAL: HOSTS: unknown
653
holotype M!
Nearctic region:
U. S.
A.)
in WA. U. S. A.: Jacksonville
(Fla.),
WA!
Uvalde
8. Bocchus dubius n. sp.
FEMALE: MALE:
unknown
fully
wingedj
length 2,0 mmj
antennae not distally thickened; proportions:
10: 6:
8: 8:
7:
7,5:
black,
with antennae and
legs brownj
antenna I segments in following 7:
7:
7:
9;
head dull,
granulated,
654
Fig.
Mem.
432 -
Amer.
Male genitalia of
Ent.
Inst.,
no.
37,
Bocchus dubius n.
1984
sp.
(type)(right half
removed)
reticulate rugose;
frons with a complete median keel;
occipital
carina
complete; POL = 6; OL = 4; OOL = 7; OPL = 4; TL = 4; scutum shiny, without sculpture; notaul ices complete, posteriorly separated; minimum distance between the notaulices slightly
longer than
temples
(5:4);
655
Bocchinae
scutellum shiny,
without sculpture;
reticulate rugose; mostly granulated; distal fig.
fore-wing hyaline,
part of radial
432;
propodeum
without dark transversal
vein shorter than proximal
part
(6:8);
bands;
genitalia:
t i b i a I sp u rs 1, 1 , 2.
LOCUS TYPICUS:
Green River
TYPICAL MATERIAL: HOSTS:
metanotum reticulate rugose;
posterior surface of propodeum weakly reticulate rugose,
(Wyoming,
holotype M!
U. S.
A.)
in AM.
unknown
DISTRIBUTION:
Neotropic region:
U.
S.
A.:
Green River
AUSTRALIAN SPECIES OF
(Wyoming),
AM!
BOCCHUS
Five species are known.
KEY TO THE SPECIES
FEMALES
Segment
4 of
front
tarsus
shorter
than
segment
1 ••••••••••••••• 2
Segment 4 of front tarsus longer than segment 1 ••••••• •••• '" 2 Fore-wing wi thout dark transversal bands; radial vein less
-
curved
(Fig.
416 E);
scutum reticulate
•• 3
rugose; notaulices 1. australiae n. sp. - Fore-wing wi th 2 dark transversal bands; radial vein more strongly curved (Fig. 416 D); scutum smooth, wi thout sculpture; notaulices incomplete . . . . . . . . . . 2. guineensis n. sp. 3 Head granulated, reticulate rugose only behind ocelli, near occipi tal carina . . . . . . . . . . . . . . . 3. minimus n. sp. - Head fully reticulate rugose ••••••••••••••••••••••••••••••••••• 4 4 Enlarged claw and segment 5 of front tarsus regularly curved (Fig. 436); head, thorax and propodeum black (with mandibles testaceous) . . . . . . . . . • . . . . . . . . . . 4. robustus (Dodd) - Enlarged claw and segment 5 of front tarsus less curved, partly rect i I inear (Fig. 437); head, pronotum and scutum fully or partly reddish; scutellum, metanotum and propodeum black . . . . . . . . . . • . . • . . • . . . • . . • • . . • . • . . . . . . .. 5. bouceki n. sp. complete . . . . . . . . . . . . . . . . . . . . . . .
MALES
Only the male of
B. guineensis n. sp. is known.
Mem.
656
Amer.
Ent.
Inst.,
no.
37,
1984
1. Bacchus australiae n. sp.
FEMALE:
fully
winged;
length 3,62-3,87 mm;
yellow;
antennae brown dark or
thorax,
propodeum and abdomen black;
fully
reticulate rugose;
occipital
with 2-3 rudimentary teeth; reticulate rugose;
POL
weak
with 2
longitudinal
transversal
tarsal front
band;
(Fig.
433)
distinct prominence; long bristle;
tibial
MALE: unknown LOCUS TYPICUS:
=
5;
OL
weakly
=
4;
Mt.
TYPICAL MATERIAL:
oval,
proportions:
with 1 preapical
433)
hairy,
propodeum reticulate brown 3:
fore-wing (Fig. 12;
Tibrogargan
2 bristles and
1 paratype F!
in QU.
I
433 -
1 proximal
with a row of 4 teeth and 1
(f
Fig.
fore
segment 5 of
(Australia) in CB;
with 1
416 E);
spurs 1,1,2.
holotype F!
fully
posteriorly separated;
14: 3:
lamella,
(Fig.
ha i ry,
mandibles quadridentate,
scutum dull,
punctate;
pterostigma short,
enlarged claw
with mandibles head du II,
legs testaceous;
carina comlete;
keels on posterior surface;
segments 1-4 in following tarsus
head black,
with segments 1-4 testaceous;
notaul ices distinct and complete,
scutellum and metanotum dull, rugose,
light,
Chela of Bocchus australiae n.
sp.
(type)
Bocchinae
Fig. 434 -
657
Chela of Bacchus guineensis n.
HOSTS: unknown DISTRIBUTION: Australian region:
AUSTRALIA:
Mt.
sp.
(type)
Tibrogargan
658
Mem.
(Queensland),
CB!
Amer.
Woogaroo,
Ent.
Inst.,
37,
no.
1984
QU!
2. Boccus guineensis n. sp.
FEMALE:
fully
winged;
length 3,00-3,50 mm;
head black,
with mandibles
and sometimes clypeus yellow; antennae fully testaceous; thorax, propodeum and abdomen black; legs testaceous, with hind coxae black; head shiny or dull, some transversal
without sculpture or weakly granulated,
mandibles with 3 teeth;
occipital
smooth,
shiny,
only on
anterior third of scutum;
sculpture;
except for
weak keels on frons and some areolae behind ocell i; carina complete;
without sculpture hairless;
scutellum smooth,
metanotum weakly punctate;
POL = 2;
Ol = 3;
notaulices incomplete, shiny,
scutum
visible
without
propodeum reticulate rugose,
with 2
longitudinal keels on posterior surface; fore-wing with 2 dark transversal bands; pterostigma short and narrow; radial vein strongly curved (Fig.
416 D); fore tarsal segments 1-4 in following proportions: 12: 4: 3: 9; segment 5 of front tarsus (Fig. 434) with a preapical lamella, one proximal prominence and one bristle; enlarged claw (Fig. 434) with one teeth and one long bristle; tibial spurs 1,1,1. MALE: fully winged; length 2,00-2,62 mm; black; mandibles testaceous; antennae brown or wi th segments 1-2 testaceous; abdomen brown; legs brown or testaceous,
with coxae and fore clubs of femora brown;
antennae
antennal segments in following proportions: 9: 5: 6: 6: 8: 7: 7: 7: 6: 9; head dull, granulated; frons with a median keel; occipital carina complete; POL = 3; Ol = 2,5; OOl = 6,5; OPl = 4; Tl = not distally thickened;
2,5; scutum dull, granulated, with numerous
longitudinal
reaching approximately 0,5 shiny,
weakly granulated;
longitudinal
with numerous areolae on anterior half and
keels on posterior half; length of scutum;
notaulices incomplete,
scutellum and metanotum
propodeum reticulate rugose,
keels on posterior surface;
fore-wi ng wi th a dark transversal
with distal
lOCUS TYPICUS: Baiyer R. (New Guinea) TYPICAL MATERIAL: holotype F! and 7 paratypes MM! in B;
HOSTS:
1 paratype M!
in OT;
two
band beneath the pterost i gma;
pterostigma lanceolate; radial vein curvi I inear, than proximal part; tibial spurs 1,1,2.
F!
with
median area shiny and smooth;
4 paratypes
(1
M,
in TW;
3 FF)!
part shorter
1 paratype
in Ol.
unknown
DISTRIBUTION:
Australian region:
B!
TW!
Baiyer R.,
BRITAIN:
Kerrvat,
Jimmi
Valley,
NEW GUINEA: Ol!
Wau,
Ol!
Tifalmin
(NE New Guinea),
Kassam Pass,
OT!
3. Bocchus minimus n. sp.
Ol!
NEW
80cchinae
FEMALE:
fully
and mandibles
winged; light;
length 2,81
mm;
antennae fully
659
head reddish-brown,
testaceousj
with clypeus
thorax and propodeum
reddi sh, part I y darkened on scutum and propodeum; abdomen brown; iegs testaceous; head dull, granulated, reticulate rugose only behind ocelli, near occipital POL
~
4;
OL
~
anterior third;
carina; 3;
mandibles with 3 teeth;
scutum dull,
granulated,
notaulices distinct and complete,
scutellum and metanotum shiny, reticulate rugose, fore tarsal
band;
(Fig. (Fig.
435) 435)
carina complete;
posteriorly separated;
weakly punctate;
propodeum
keels on posterior surface; pterostigma short and oval,
segments 1-4 in following proportions:
tarsus
enlarged claw MALE:
smooth,
with 2 longitudinal
with 1 weak dark transversal of front
occipital
reticulate rugose only on
with a preapical
13: 2:
2:
14;
fore-wing brown; segment 5
lamella and 2 bristles;
with a row of 14 teeth;
tibial
spurs 1,1,2.
unknown
LOCUS TYPICUS: Spargoville (Australia) TYPICAL MATERIAL: holotype F! in 8M. HOSTS: unknown DISTRIBUTION: Australian region: Western Australia),
AUSTRALIA:
Spargoville
(Coolgardie,
BM!
L L
•.......•.....
Fig. 435 -
Chela of
~ ~~ ....•..•....
Bacchus minimus
n.
....
sp.
(type)
660
Mem. Amer.
Fig. 436 -
Ent.
Inst.,
Chela of Bocchus robustus
no.
(Dodd)
37,
1984
from Townsville
(Australia)
4. Bacchus robustus (Dodd) n. comb.
Anteonella robusta Dodd 1913: 182. Anteonella bicolor Dodd 1913: 182 (n. syn.); locus typicus Townsville (Australia).
Anteonella robusta Dodd: Kieffer 1914b: 213. Anteonella bicolor Dodd: Kieffer 1914b: 214. Dicondylus sulciscutellum Girault 1926: 135 (n. syn.); locus typicus Townsville
(Australia).
Bocchinae
FEMALE:
fully
testaceous;
winged;
length 4,87-5,12 mm;
antennae testaceous,
propodeum and abdomen black; clubs of femora black;
661
head black,
with mandibles
with segments 4-10 or 6-10 dark;
thorax,
legs fu Ily testaceous or wi th coxae and
head swollen,
dull shiny,
fully reticulate rugose,
with a median keel between the antennal sockets; ocellar region swollen; temples distinct; occipital carina complete; POL = OL; OOL = OPL; mandibles tridentate,
with an
intermediate rudimentary tooth;
pronotum
shiny, with a transversal impression, reticulate rugose and with transversal keels; pronotal tubercles reaching tegulae; scutum shiny, reticulate rugose; posteriorly
notaul ices complete and distinct,
separated;
scutellum shiny,
median furrow composed of areolae;
smooth,
fully
composed of areolae,
weakly punctate,
with a
metanotum shiny and smooth,
reticulate rugose;
propodeum fully reticulate rugose;
transversa I band;
pterost i gma broad and ova I,
fore-wing
brown;
laterally
with 1 dark
segment 3 of front
tarsus without distal apophysis longer than segment 3; segment 4 of front tarsus longer than segment 1; segment 5 of front tarsus (Fig. 436) with an apical
lamella and 1 long proximal
a row of 16 teeth and 3 proximal MALE: unknown LOCUS TYPICUS:
Townsville
bristle;
enlarged claw
bristles;
tibial
(fig.
436)
with
spurs 1,1,1.
(Australia)
of A. robusta in AD; holotype F! of A. bicolor in AD; holotype F! of D. sulciscutellum in QU. TYPICAL MATERIAL: HOSTS:
holotype F!
in Australia Fulgorids.
DISTRIBUTION: Australian region: AUSTRALIA: AD! Blunder Cr. (Brisbane, Queensland), UQ! Australia),
WA!
Sir Graham Moore
Island
Townsville (Queensland), Anjo Peninsula (Western
(Western Australia),
QU!
OL!
5. Bocchus bouceki n. sp.
FEMALE:
fully
reddish
winged;
in holotype;
length 3,37-4,87 mm;
head black
head,
pronotum and scutum
(with clypeus and mandibles
testaceous),
prothorax reddish, scutum black (with notaulices reddish) in paratype; antennae brown with segments 1-2 testaceous or fully testaceous; scutellum, metanotum,
propodeum and abdomen black;
hind coxae brown; = 4;
occipital
rudimentary rugose,
head hairy,
carina complete;
tooth between
hairy,
shiny;
shiny,
legs testaceous,
fully
sometimes with
reticulate rugose;
mandibles quadridentate,
the two posterior teeth;
POL = 5;
OL
with a
scutum fully reticulate
notaulices complete and distinct,
posteriorly
separated; scutellum and metanotum smooth, hairy, weakly punctate; propodeum reticulate rugose, with 2 longitudinal keels on posterior surface; brown;
fore-wing fore tarsal
segment 5 of front bristle; bristles;
with 1 dark transversal tarsus
enlarged claw tibial
band;
pterostigma short,
segments 1-4 in following proportions: (Fig.
(Fig.
spurs 1,1,2.
437)
417)
with a preapical
18:
3:
3:
oval, 26;
lamella and no
with a row of 17 teeth and without
662
Mem. Amer. Ent.
Ins!.,
no. 37,
1984
Fig. 437 - Chela of Bocchus bouceki n. sp.
MALE: unknown LOCUS TYPICUS: Mt. Tibrogargan
(Australia)
(type)
Bocchinae
TYPICAL MATERIAL:
holotype F!
in CB;
663
1 paratype F!
in AD.
HOSTS: unknown DISTRIBUTION: Australian region: AUSTRALIA: Mt. Tibrogargan (Queensland), CB! Landrigan Cliffs (North Western Australia), AD! NOTES:
the species
is named
in honor of Mr.
Zdenek Boucek.
4. Mirodryinus Ponomarenko
Mirodryinus Ponomarenko 1972: 673.
Fig. 438 -
Female of Mirodryinus xerophilus
(Benoit)
from Wassiya
(Egypt)
664
Mem. Amer.
Fig. 439 -
Ins!.,
no.
Female of Mirodryinus xerophilus
TYPE SPECIES: Mirodryinus FEMALE
En!.
(Figs 438, 439):
37,
1984
(Benoit) from Wassiya
(Egypt)
ungulatus Ponomarenko 1972, orig. desig.
fully winged;
maxi Ilary palpi
with 6 segments;
labial palpi with 3 segments; occipital carina complete; mandibles quadridentate, with a rudimentary tooth bet wen the two posterior teeth; segment 4 of front tarsus less than 0,5 times as long as segment 1 (Fig. 442); segment 5 of front tarsus without lamellae, with bristles or peg-like hairs (Fig. 442); enlarged claw without teeth, lamellae, bristles or peglike hairs (Fig. 442); fore-wing with radial vein distinct (though sometimes it is very short); pronotal tubercles reaching tegulae; tibial spurs 1,1,1. MALE (Figs 440, 441): fully winged; mandibles quadridentate, with a rudimentary tooth between the two posterior teeth; maxillary palpi with 6 segments; labial palpi with 3 segments; temples distinct; propodeum with areolae very small, less wide than tegulae (Fig. 440); tibial spurs 1,1,2. DISTRIBUTION: Palaearctic HOSTS: unknown SPECIES: 5
PALAEARCT I C SPEC I ES OF
MIRODRYINUS
Five species are known.
KEY TO THE SPECIES
Bocchinae
Fig.
665
Male of Mirodryinus xerophilus
440 -
(Benoit)
from
Ikingi
Mariout
(Egypt)
FEMALES
Fore-wing distal
with
parts;
radial
vein
curved,
with distinct
proximal
and
thorax and propodeum fully black . . . . . . . . . . • . . . . • .
1. gobiensis Ponomarenko -
Fore-wing wi th proximal pa r t I y
part
radial
vein
straight,
(Figs 438,439);
only wi th a distinct
thorax and propodeum fully
or
redd i sh •••••••••••••••••••••••••••••••••• ••' ••••••••••••• 2
2 Thorax and propodeum fully reddish . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . •••••••••••••••••••••••••••••• 2. xerophilus (Benoi t) -
Thorax and propodeum partly
3 Scutum and
or black • . . . . . . . . 3 ungulatus Ponomarenko
reddish and brown
scutellum brown . . . .
3.
666
Mem.
Fig.
441
-
Male of
Amer.
En!.
Ins!.,
37,
no.
1984
Mirodryinus xerophilus (Benoit) from Ikingi Mariout (Egypt)
- Scutum and scutellum reddish •••••• ••••••••••••••••••••••••••••• 4 4 Metapleura reddish; radial vein longer . . . . . . . . . . . . . . . . . . . . . . . . . . . • . . . . . . . . . . . . • . . . . . . . . . . . . . .. 4. atlanticus n. sp. - Metapleura black; radial vein shorter (Fig. 448 A) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. 5. ponomarenkoae n. sp.
MALES
Only
the male of M.
xerophilus is known.
1. Mirodryinus gobiensis Ponomarenko
Mirodryinus gobiensis Ponomarenko 1972: 676.
FEMALE:
fully
legs fully
winged;
testaceous;
length 2,62 mm;
head,
ocellar region brown;
mandibles,
antennae and
thorax and propodeum black;
abdomen black; antennae distally thickened; antennal segments in following proportions: 7: 3: 6: 5,5: 5: 5: 5: 4,5: 5: 5; head shiny, weakly
punctate,
or lateral 4;
keels;
Ol = 2,5;
without sculpture among punctures; temples very prominent;
OOl = 10; OPL = 4;
Tl = 9;
occipita!
frons without median
carina complete;
pronotum very short,
POL
weakly
Bocchinae
667
B
Fig.
442 -
Female of Mirodryinus gobiensis Ponomarenko (type):
B: fore tarsal segments.
A: chela;
668
Mem.
transversely striate;
pronotal
and metanotum shiny, punctures; scutum;
Amer.
Ent.
Inst.,
weakly punctate,
notaulices incomplete,
longitudinal
keels;
fore-wing
pterostigma;
radial
vein curved,
segments in following proportions: 442)
1 of front
LOCUS TYP ICUS:
DISTRIBUTION:
442);
and distal part
18: 29:
hollow
long
the
parts;
(2,5:4,5);
65;
segment 5
apex with a
lamella);
(antenna-cleaner)
fore
enlarged claw
bristles or other appendixes;
tibial
Bordson-Gobi
TYPICAL MATERIAL: HOSTS: unknown
15:
(among which one very
tarsus with a proximal
a big prominence (Fig. MALE: unknown
aymag,
teeth,
82:
or
bands beneath
with 2 rows of nearly 15 bristles;
group of nearly 5 lamellae
scutellum
without transversal
vein much shorter than proximal
without subapical (Fig.
dull,
with distinct proximal
part of radial
tarsus
scutum,
without sculpture among
with a dark transversal
tarsal
442)
1984
reaching approximately 0,3 length of
distal (Fig.
37,
tubercles reaching tegulae;
smooth,
propodeum fully reticulate rugose,
of front
no.
segment
del imitated by
spurs 1,1,1.
(Mongol i a)
holotype F!
in LN
Palaearctic region:
South-East Mongolia),
MONGOLIA:
Bordson-Gobi
(South Gobi
LN!
2. Mirodryinus xerophilus (Benoit) n. comb.
Anteon xerophilus Benoit 1951a: 301. Anteon xerophilum Benoit: Olmi and Currado 1979a: 346.
FEMALE
(Figs 438,
439):
fully
petiole and abdomen black; testaceous,
winged;
length 2,68 mm;
with segments 5-10 brown;
head shiny,
smooth,
with antennae
antennae distally thickened;
antenna I segments in following proportions: 4,5: 7;
reddish,
sometimes propodeum partl y brown; 6:
4:
strongly punctate,
11: 7:
5,5:
4,5:
4:
5:
without sculpture among
punctures; mandibles quadridentate, with a rudimentary tooth between the two posterior teeth; frons without medial keel; occipital carina complete; POL = 6,5; dorsal
OL = 4;
view;
punctate,
scutum,
OOL = 7;
with a
TL = 10; pronotum
without sculpture among punctures;
propodeum reticulate rugose, fore-wing
OPL = 8;
scutellum and metanotum shiny,
track of dark spot on the basal
proximal part, six times as
without the distal
long as segment 4,
part;
weakly
notaulices invisible;
without transversal
with a broad dark transversal
invisible in
smooth,
and
longitudinal
keels;
band beneath the pterostigma and cells;
radial
vein only
segment 1 of front
with a proximal
delimitated by a big prominence (Fig. 443);
hollow
with the
tarsus nearly
(antenna-cleaner)
enlarged claw
(Fig.
443)
without bristles or teeth or lamellae; segment 5 of front tarsus (Fig. 443) with a long row of small peg-like hairs; apex with a group of 4 lamellae; tibial
spurs 1,1,1.
669
Bocchinae
B
Fig. 443 -
Female of
Mirodryinus xerophilus (Benoit) from Egypt: A: chela; B:
fore tarsal
segments.
670
Mem. Amer.
Ent.
Ins!.,
no.
-1/
/
, \
II
\I
/
\
/ /
I /
I
II
I
II 1\
I
I
)
/
I
I
1/
/'
1984
I
\
/ \
37,
'\
I
1\
I I
I
I
\
I
I \ I \
/
I
1/ "-
I
I
\ I v
II
I / /
Fig. 444 - Male genitalia of Mil'odl'yinus xel'ophilus (Benoit)(type)
Bocch i nae
MALE
(Figs 440,
441),
fully
winged;
clypeus yellow and a small sockets and eyes; testaceous; scutellum,
671
length 2,5 mm;
head black,
reddish spot on region between
mandibles yellow,
with teeth brown;
antennae fully
pronotum and propectus reddi sh-testaceous or black; metanotum,
with
antennal
mesopleura and metapleura brown-black;
scutum propodeum
black; abdomen brown; legs fully testaceous; tegulae testaceous; antenna I segments in following proportions: 6: 2: 10: 9: 8: 7: 7: 6,5: 5: 7; head sh i ny,
swoll en,
punctures; complete;
smooth,
POL = 7;
wi th sparse punctures,
OL = 4;
scutum shiny,
smooth,
sculpture among punctures; approximately 0,3 smooth,
without sculpture, almost pointed;
propodeum dull,
hairy,
with sparse punctures,
notaulices distinct,
length of scutum;
convex,
wi thout scu I pture among
= 8; OPL = 7; TL = 7; occipital carina
OOL
swollen;
weak,
without
incomplete,
reaching
scutellum and metanotum shiny, laterally viewed metanotum very
mesopleura and metapleura shiny and smooth;
granulated,
with some irregular keels or areolae,
without
a visible transversal keel between dorsal and posterior surface; dorsal surface very reduced, shorter than breadth of metanotum, as scu I ptured as posterior surface; surface;
pterostigma, part,
propodeum almost fully composed of the posterior
fore wing hyal ine, with distal
nearly
invisible;
and without dorsal
with a dark transversal
part of radial gonoforceps
proximal
TYPICAL MATERIAL:
Ikingi
Mariout,
444)
without
BM!
proximal
inner distal
dorsally viewed
process long and
(Libya)
holotype M!
in GE.
HOSTS: unknown DISTRIBUTION: Palaearctic region: EGYPT:
(Fig.
membranous process,
broad; tibial spurs 1,1,1. LOCUS TYP ICUS: Giarabub oasis
band beneath the
vein much shorter than
LIBYA:
Giarabub Oasis (Cirenaica),
Fayoum-Tamiya,
WA!
Wassiya,
GE!
WA!
3. Mirodryinus ungulatus Ponomarenko
Mirodryinus ungulatus Ponomarenko 1972: 674.
FEMALE:
fully
winged;
length 2,31
part of frons darkened;
protorax redd ish-testaceous; brown-b lack; black,
mm;
head reddish,
antennae brown,
scutum brown;
propodeum black;
pleura,
scutell um and metanotum
mesosternum and metasternum
except for a reddish spot on mesopleura;
testaceous-dark;
antennae distally thickened;
following proportions,
7:
3,5, 9,
6:
5,
with vertex and
wi th segments 1-3 testaceous;
4,5:
abdomen brown;
antennal 4,5:
4,5:
segments 4,5:
6;
legs in
head shiny,
puncta te, wi thout scu I pture among punctures; frons wi thou! med i an and lateral keels; occipital carina complete; temples very prominent; POL = 6; OL = 3;
OOL
=
9;
tubercles reaching
OPL = 7,5; tegulae;
TL
=
scutum,
10;
pronotum shiny,
short;
pronotal
scutellum and metanotum shiny,
672
Fig.
Mem.
Amer.
Ins!.,
no.
37, 1984
445 - Female of Mirodryinus ungulatus Ponomarenko (type): A: chela; B:
smooth,
weakly punctate,
incomplete,
fore tarsa I segments.
without sculpture among punctures;
reaching approximately 0,5
reticulate rugose, wing
Ent.
dull,
without transversal
with a dark transversal
without a distinct distal
length of scutum;
part;
or longitudinal
band on distal fore tarsal
half;
segments
radial
notaulices
propodeum keels;
fore-
vein straight,
in following
proportions: 85: 13: 15: 16: 41; segment 1 of front tarsus with a proximal hollow ("antenna-cleaner") distally del imitated by a big prominence (Fig.
445); enlarged claw (Fig. 445) without subapical tooth, bristles or other appendixes; segment 5 of front tarsus (Fig. 445) with 2 rows of nearly 26 peg-like lamellae; apex with a group of nearly 5 lamellae; tibial spurs 1,1,1. MALE:
unknown
LOCUS TYPICUS: Husseyn-Dey (Algeria) TYPICAI_ MATERIAL: holotype F! in LN. HOSTS:
unknown
DISTRIBUTION:
Palaearctic region:
ALGERIA:
Husseyn-Dey,
4. Mirodryinus atlanticus n. sp.
LN!
Bocchinae
Fig. 446 -
FEMALE:
Fore tarsal
fully winged;
673
segments of Mirodryinus atlanticus n.
length 2,62 mm;
head reddish;
sp.
(type)
antennae brown,
674
Mem.
Amer.
with segments 1-3 testaceous;
En!.
pronotum,
metanotum and propodeum black; abdomen brown-dark; punctate, 10;
with
occipital
legs and
keels;
dark
straight,
band on
not curved;
scutum shiny,
(Fig.
446)
lamellae; segment
=
5;
head shiny, OL
1 of front
shiny,
distal
3:
LOCUS TYPICUS:
radial oval,
3,5:
7;
without
with a proximal
hairy,
(Canary
holotype F!
9;
TL
=
with smooth,
propodeum reticulate
vein
fore-wing
brown;
fore tarsal
bristles,
segments
tarsus
in addition
lamellae,
(Fig.
with a
very short and
segment 5 of front
hollow
big prominence
Gran Tarajal
TYPICAL MATERIAL:
446)
strongly
=
OOL
weakly punctate,
hairless;
region;
2,5:
(Fig.
tarsus
3;
rugose,
with some bristles and 3 peg-I ike hairs, enlarged claw
=
keels on posterior surface;
17:
distally del imitated by a MALE: unknown
HOSTS:
and scutellum reddish;
visible only on anterior third;
pterostigma short,
in following proportions:
1984
and metap leura redd i sh;
POL
notaul ices incomplete,
longitudinal
transversal
scutum
pronotum short,
scutellum and metanotum smooth, without
37,
tegulae testaceous;
carina complete;
without sculpture;
no.
mesop leura
temples very prominent;
numerous transversal
rugose,
Ins!.,
to 4 distal
hairs;
("antenna-cleaner")
446);
tibial
spurs 1,1,1.
Islands)
in HE.
unknown
DISTRIBUTION:
Palaearctic region:
(Fuerteventura),
CANARY
ISLANDS:
Gran Tarajal
HE!
5. Mirodryinus ponomarenkoae n. sp.
Mirodryinus ungulatus Ponomarenko partim 1972: 674.
FEMALE:
fully
winged;
on ocellar region; reddish,
length 3,20 mm;
antennae missing
with anterior surface brown;
propodeum and abdomen black;
head reddish,
in holotype;
with a brown spot
pronotum reddish;
scutellum reddish;
mesopleura reddish,
scutum
metanotum,
with sutures blackish;
metapleura black; legs testaceous; head shiny, weakly alutaceous, strongly punctate on frons; occipital carina complete; frons without median keel; POL
=
=
10;
9;
OL
=
TL
=
10;
? (anterior ocellus damaged and not visible); OOL temples very prominent;
with numerous transversal punctate;
keels;
notaulices incomplete,
scutellum and metanotum shiny, rugose, dark
without
transversal
straight tarsal (Fig.
longitudinal
(Fig.
segments 447)
a proximal
scutum shiny, smooth, region;
not curved;
hollow
lamellae,
bristles,
("antenn-cleaner")
20:
hairs;
3:
=
8;
OPL
hairy, not
length of scutum; propodeum reticulate
vein
fore-wing
3: 4:
segment
with a
very short and
pterostigma short and oval,
in following proportions:
without
hairless; radial
rugose,
weakly alutaceous,
reaching only 0,3
keels on posterior surface;
band on distal
448 A),
pronotum short,
10;
brown;
fore
enlarged claw
1 of front
distally delimitated by a
tarsus with big
Bocchinae
Fig.
447 -
Fore tarsal
segments of female of Mirodryinus ponomarenkoae n. sp.
prominence (Fig.
447);
nearly
17 lamellae;
1,1,1. MALE:
unknown
675
(type)
segment 5 of front
tarsus
LOCUS TYPICUS: Western part of Mount Zamryn-Ula TYPICAL MATERIAL: holotype F!in LN. HOSTS;
Zamryn-Ula
Palaearctic region:
(Central
with 1 row of tibial
spurs
(Mongolia)
Aymag),
MONGOLIA:
Western part of Mount
LN!
the holotype of this species was designated by Ponomarenko
as para type of M. of Mrs.
447)
unknown
DISTRIBUTION: NOTES:
(Fig.
apex with a group of nearly 8 lamellae;
(1972)
ungulatus Ponomarenko. The species is named in honor
Nadezdha Ponomarenko.
5. Radiimancus Moczar
Radiimancus Moczar 1983b: 203.
TYPE SPECIES: Radiimancus olmii Moczar 1983b: 204, FEMALE labial
(Figs 449, palpi
tooth between 0,5 times as
450):
ful!y
with 3 segments;
winged;
long as segment 1 (Fig. tarsus without
segment 4 of front 451);
lamellae,
occipital
desig.
with 6 segments;
mandibles quadridentate,
the two posterior teeth;
segment 5 of front
orig.
maxi Ilary palpi
with a rudimentary tarsus
less than
carina complete;
with bristles or peg-like hairs
Mem.
676
Amer.
En!.
Ins!.,
no.
37,
1984
A~
B~ Fig. 448 - Pterostigma and radial vein of females of Mirodryinus ponomarenkoae n. sp. (type)(A) and Radiimancus olmii Moczar (type)(B)
(Fig.
451);
bristles
enlarged claw without teeth,
(Fig.
451);
radial
vein
(Fig.
MALE:
unknown
DISTRIBUTION: HOSTS:
pronotal
448 B);
lamellae,
peg-I ike hairs or
tubercles reaching tegulae;
tibial
fore-wing with no
spurs 1,1,1.
Palaearctic
unknown
SPECIES:
1
Radiimancus olmii Moczar
Radiimancus olmii Moczar 1983b: 204.
FEMALE
(Figs 449,
450):
fully winged;
anterior surface of frons,
length 2,80 mm;
head black,
clypeus and mandibles testaceous;
with
antennae
testaceous, with last 5 segments darkened; pronotum and prosternum reddish; scutum reddish, with central surface darkened; scutellum, metanotum and propodeum black; mesosternum reddish, with black spots; metasternum and metapleura black; mesopleura reddish; abdomen brown; legs testaceous;
antennae weakly distally thickened;
following proportions: quadri dentate, head shiny, occipital
8:
3:
12: 8,5:
with a rud imentary
smooth,
6:
6:
6:
6:
antennal 8;
segments in
mandibles
tooth between the two posterior teeth;
weakly punctate,
carina complete;
7:
without sculpture among punctures;
frons without median keel;
POL = 8;
OL = 4;
OOL = 10; OPL = 8; TL = 8; pronotum short, hairy, rugose, pronotal tubercles reaching tegulae; scutum shiny, weakly punctate, without sculpture among punctures;
notaulices incomplete,
reaching only 0,3 lenth
of scutum; scutellum and metanotum shiny, smooth, without sculpture; propodeum reticulate rugose, without longitudinal or transversal keels;
677
Bocchinae
Fig. 449 -
Female of Hadiimancus olmii Moczar
(type)
fore-wing with a dark transversal band on distal surface; pterostigma broad and oval; radial vein absent (Fig. 448 B); fore tarsal segments following proportions:
17: 2:
2,5:
3:
9;
in
segment 1 of front tarsus with a
proximal hollow ("antenna-cleaner") distally del imitated by a big prominence (Fig. 451); segment 5 of front tarsus (Fig. 451) with 2 rows of nearly 27 bristles extending continuously to the apex; enlarged claw (Fig. 451)
without
lamellae, bristles or hairs;
tibial
spurs 1,1,1.
MALE: unknown LOCUS TYP I CUS: Bu I gan (Mongol i a) TYPICAL MATERIAL: holotype F! in BU. HOSTS: unknown DISTRIBUTION: Palaearctic region:
MONGOLIA:
Bulgan
(10 Km SSW Somon,
678
Mem.
Fig.
Chovd
aimak),
450 -
Amer.
Ent.
Inst.,
no.
37,
1984
Female of Radiimancus olmii Moczar (type)
BU!
6. Mystrophorus Forster
Mystrophorus Forster 1856: 91. Mystrophorus Forster: Richards 1937: 119.
TYPE SPECIES: Mystrophorus FEMALE labial
(Figs 452, 453): palpi
with 3 segments;
tooth between tarsus
less than 0,5 times as
or peg-like hairs
456);
I ike hairs or bristles (Fig. tibial spurs 1,1,2. (Figs 454,
rudimentary
less wide than tegulae;
with 6 segments;
396 A);
tarsus without
with a rudimentary
segment 4 of front 456);
456);
pronotal
occipital
lamellae,
enlarged claw without teeth,
455): micropterous;
tooth between
(Fig.
long as segment 1 (Fig.
segment 5 of front (Fig.
maxillary palpi
mandibles quadridentate,
the two posterior teeth
carina complete;
MALE
formicaeformis Ruthe 1859, monotypic.
micropterous;
with bristles lamellae,
mandibles quadridentate,
the two posterior teeth;
maxi Ilary palpi
peg-
tubercles reaching tegulae; with a
areolae of propodeum
with 6 segments;
labial
palpi
with
3 segments; temples distinct. DISTRIBUTION: Palaearctic HOSTS:
Cicadellidae (excl ud i ng I diocerinae, Macropsinae,
SPECIES:
1
Mystrophorus formicaeformis Ruthe
Typhlocybinae).
Bocchinae
679
A
Fig. 451 - Female of Radiimancus oimii Moczar (type): A: fore tarsal segment 1; B: fore tarsal segments 1 (partly)-4 and chela
Mystrophorus iormicaeiormis Ruthe 1859: 120. Mystrophorus iormicaeiormis Ruthe: Ponomarenko 1978: 23.
FEMALE (Figs 452, 453): micropterous, with fore wings as long as tegulaej hind wings rudimentary, rounded; length 1,75-2,50 mm; black, with
680
Mem. Amer. Ent.
Fig. 452 -
Inst.,
no.
37,
1984
Female of Mystrophorus formicaeformis Ruthe
Bocchinae
Fig.
453 -
antennae brown
Female of
681
Mystrophorus formicaeformis Ruthe
(except for segments 1-2 testaceous),
tegulae brown-testaceous,
legs yellow
mandibles yellow,
(with hind coxae almost
clubs of femora and hind tibiae brownish);
body dull,
hairy,
fully black, except for
abdomen hairless; head without median keel between antenna I sockets; occipital carina complete; head, thorax and propodeum granulated, without striae or keels; invisible;
some keels are visible near apex of propodeum;
fore tarsal
segments 1-4 in following proportions:
notaulices
5,5:
1,2:
2,3:
2,3; en I arged cl aw (Fig. 456) without I ameli ae or brist les, wi th apex rounded; segment 5 of front tarsus (Fig. 456) without lamellae, with numerous rows of bristles or peg-like hairs. MALE
(Figs 454,
455):
micropterous,
with fore wings narrow and broadened
at apex; hind wings rudimentary, pointed; length 1,87-2,75 mm; black, with antennae brown(except for segments 1-2 testaceous), tegulae yellowbrown,
fore-wing
femora blackish, for abdomen sockets;
almost hairless;
occipital
granulated; propodeum;
yellow-brown,
legs yellow
(with coxae and clubs of
mid and hind tibiae brownish);
body dull,
head without median keel
carina complete;
head,
without striae or keels,
hairy,
between
except
antenna I
thorax and propodeum
except for some keels near apex of
notaulices invisible.
LOCUS TYPICUS:
Boden
TYPICAL MATERIAL: HOSTS: in Germany
(Germany)
not found
(not
in BM and
in Gent,
Belgium).
Deltocephalus sp. (Haupt 1938); in U. S. S. R. Diplocolenus abdominalis F. (Ponomarenko 1978). DISTRIBUTION: 1978); det.
Palaearctic region:
HUNGARY: Or Sz.
Miklos,
U.
BU!
S. Vacz
S.
R.:
Moscow
(Tudosdomb),
(Ponomarenko BU!
Hungary
(male
Mystrophorus cursor Forster type), VI! CZECHOSLOVAKIA (Ogloblin
1924b);
GERMANY:
E I steraue (Ha lie),
HA!
BM!
Boden
(Ruthe 1859);
Ha II e-a-
5., HA! Bellinchen (Oder), HA! KUpka (Th.), EB! Mallertshofer Holzb. (MUnchen), EB! OT! Saale-Ufer (Haupt 1938); Schl inzheim (Mallertshofer Holz), GV! ENGLAND: Hayling Island (Hampshire), BM! GREECE: Kerkyra Island,
BU!
682
Mem.
Fig.
454 -
FEMALE
En!.
Inst.,
Male of Mystrophorus
no.
37,
1984
formicaeformis Ruthe from Hungary
5UBFAMILY THAUMATODRYININAE N.
6.
TYPE GENUS:
Amer.
Thaumatodryinus R.C.L. Perkins 1905
(Figs 19,
457):
fully
winged;
fore-wing
with costal,
submedian cells clearly enclosed by pigmented veins also first brachial labial
palpi
SUBF.
cell
clearly enclosed;
with 3 segments;
front
(Fig.
maxillary palpi
tarsus chelate;
median and
457);
usually
with 6 segments;
chela with rudimentary
claw (Fig. 459); antennae with tufts of long hairs on segments 5-10 (last segment with two tufts) (Fig. 19); occipital carina complete; mandibles with 4 teeth progressing spurs 1,1,2.
larger from anterior one to posterior (Fig.
5);
tibial
Thaumatodry i n i nae
Fig.
MALE
Male of Mystrophorus
455 -
(Figs 28,458):
fully
winged;
683
formicaeformis Ruthe from Hungary
fore-wing
with costal,
submedian cells clearly enclosed by pigmented veins palpi
with 6 segments;
teeth progressing carina complete; pterostigma
28);
GENERA:
with 3 segments;
with metacarpus as
tibial
Palaearctic
Oriental, Nearctic, HOSTS: Flatidae SPECIES:
palpi
median and 28); 5);
occipital
long as or longer than
spurs 1,1,2.
(?)(fossil
Neotropic,
maxillary
mandibles with 4
larger from anterior one to posterior (Fig. fore-wing
(Fig.
DISTRIBUTION:
labial
(Fig.
species:
see notes),
Ethiopian,
Australian
1 17
NOTES: Brues described three species of Thaumatodryinus from Palaearctic region (Baltic amber): filicornis (Brues 1923), deletus Brues 1933 and
gracilis Brues 1933. Though the original descriptions are incomplete and the typical correct.
material
is
In the original
lost,
I think that the Brues determination
description of Th.
is
deletus in fact Brues wrote
about the presence on the I ast two antenna I segments of tufts of long hairs: this character is recognizable only in Thaumatodryinus. In living
Thaumatodryinus, however, the tufts are on antennal segments 5-10. Probably
the three Palaearctic fossi I species are belonging
extinct genus of Thaumatodryininae,
to another
perhaps Harpactosphecion
Haupt 1944
(described by Haupt for one of the three above-mentioned species,
filicornis) .
Thaumatodryinus R.C.L. Perkins
Thaumatodryinus R.C.L. Perkins 1905: 58.
684
Mem. Amer.
En!.
Ins!.,
no.
37,
1984
Ilflll III/flilltilf
Fig. 456 -
Female of Nystrophorus tormicaeiormis Ruthe: tarsal
A:
chela;
B:
fore
segments 1-4.
Thaumatodryinus R.C.L. Perk i ns: Kieffer in Kieffer et Marshall 1906: 490.
Thaumatodryinus Thauma todryin us Thaumatodryinus Thaumatodryinus
TYPE SPECIES: FEMALE
R.C.L. R.C.L. R.C.L. R.C.L.
Perk i ns: Perkins: Perkins: Perkins:
Kieffer 1914b: 16. Krombein 1952: 97. Krombein 1958: 98. Krombein 1979: 1243.
Thaumatodryinus koebelei R.C.L. Perkins 1905, orig. desig.
(Figs 19, 457):
fully winged;
fore-wing with costal,
median and
submedian cells clearly enclosed by pigmented veins (Fig. 457); usually also first brachial cell clearly enclosed; maxi Ilary palpi with 6 segments; labial palpi with 3 segments; front tarsus chelate; chela with rudimentary claw (Fig. 459); antennae with tufts of long hairs on the segments 5-10 (last segment with two tufts) (Figs 19, 471); pronotal tubercles reaching tegulae; occipital carina complete; mandibles with 4 teeth progressing
Thaumatodry i n i nae
685
/
Fig.
457 -
Female of Thaumatodryinus Guinea)
flavus n. sp. from Waris (New
686
Fig.
Mem.
458 -
Amer.
En!.
Ins!.,
no.
37,
1984
n.
sp.
from Kira-Kira
Male of Thaumatodryinus flavus
(Solomon
Islands)
larger from anterior one to posterior MALE
(Figs 28,
458):
fully
winged;
(Fig.
5);
fore-wing
tibial
submedian cells clearly enclosed by pigmented veins palpi
with 6 segments;
teeth progressing carina complete; pterostigma
(Fig.
DISTRIBUTION: fossil
labial
palpi
spurs 1,1,2.
with costal,
with 3 segments;
(Fig.
28);
larger from anterior one to posterior (Fig. fore-wing 28);
with metacarpus as
tibial
Ethiopian,
maxillary
mandibles with 4 5);
occipital
long as or longer than
spurs 1,1,2.
Oriental,
Nearctic,
Neotropic, Australian
Palaearctic species see note on page 683).
HOSTS: Flatidae GENERA: 1 SPECIES:
median and
17
ETH lOP I AN SPEC I ES OF THAUMATODRYINUS
Five species are known.
KEY TO THE SPECIES
(for
Thaumatodry i n inae
687
FEMALES
Fore and hind wings with a dark
transversal
band ••••••••••••••• 2
bands •.••.•••.•••• 3 2 Head mostly black; mesothorax, metathorax and propodeum black ••• • • • • • • • • • • • • • • • • • • • • • • • • • • •. 1. mirabilis n. sp. Fore and
-
hind wings without
dark
transversal
Head fully reddi sh-testaceous; mesothorax, metathorax and propodeum reddi sh-testaceous, wi th scutellum black ••••••.••••.••
· . . . . • . . . . . . . . • . . • • . . . . . . . . . 2. infuscatus n. sp. 3 Head wi th OPL longer than OL; thorax and propodeum partly black • . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3. dentatus Benoi t -
Head wi th OPL as
long
as or
short er
than OL;
thorax and
4 4 Occiput more strongly excavated; fore trochanters nearly as long as fore femora; posterior surface of propodeum wi th 2 complete propodeum fully
longitudinal
reddish-testaceous •••••••••••••••••.•••••••••••
keels;
notaulices more converging . . . . . . . . . . . . . . . . ..
· . . . . . . • . . . . . . . . . . . . . . . . . . .• -
Occiput
not
long as
fore
tracks
of
or weakly femora;
incomplete
townesi n.
4.
excavated; posterior
sp. trochanters nearly
fore
surface
longitudinal
of
keels;
0,5 as
propodeum only wi th notaulices
2
less
5. migratorius Beno i t
converg i ng . . . . . . . . . . . . . . . . . .
MALES
Only
the supposed male of
Th.
tOlvnesi n. sp. is known.
1. Thaumatodryinus mirabilis n. sp.
FEMALE:
fully
winged;
length 6,25-7,00 mm;
clypeus and mandibles yellow;
head black,
with malar space,
sometimes occiput partly brown-light;
antennae testaceous; pronotum and propectus yellow ; mesothorax, metathorax and propodeum black, with propodeal spiracles brown; black;
abdomen reddish;
legs testaceous,
petiole
with fore clubs of femora black,
mid and hind clubs of femora brown; antenna I segments in following proportions: 20: 7: 40: 45: 44: 45: 35: 25: 19: 15; head dull, reticulate rugose; hairy,
occipital
carina complete;
crossed by a
transversal
striae;
reticulate rugose;
transversal pronotal
OPL
notaul ices complete, metanotum smooth,
reticulate rugose;
fore-wing
part of radial
vein
tegulae;
with weak
scutum dull,
posteriorly separated; transversal
longer than
proximal
fully
scutellum
with some anterior furrows;
with a brown
much
pronotum shiny,
weakly punctate,
tubercles reaching
reticulate rugose; distal
longer than OL;
impression,
propodeum
band and with part;
radial
cell
688
Mem.
Fig.
459 -
Chela of
Amer.
Ent.
Inst.,
no.
37,
1984
Thaumatodryinus mirabilis n. sp. from Ankasoka (Madagascar)
closed;
pterostigma brown-I ight;
transversal
band;
segment
hind wings with a
1 of front
distal
tarsus nearly as
brown
long as segment 4;
Thauma todry i n i nae
689
(
Fig. 460 -
Chela of Thaumatodryinus infuscatus n.
sp.
(type)
segment 3 produced into a hook; enlarged claw with 2 subapical teeth and with 32 lamellae (Fig. 459); segment 5 of front tarsus (Fig. 459) with 2 rows of 59 lamellae; apex with a group of 5 lamellae; tibial MALE: unknown LOCUS TYPICUS: Ankasoka (Madagascar) TYPICAL MATERIAL: holotype F! and 1 paratype F! in P.
spurs 1,1,2.
690
Mem.
HOSTS:
Amer.
Ent.
Inst.,
no.
37,
1984
unknown
DISTRIBUTION:
Ethiopian region:
MADAGASCAR:
Ankasoka
(Route Lakato),
P!
2. Tha umatodryinus infuscatus n. sp.
FEMALE:
fully
winged;
length 4,68 mm;
and scutellum black;
head shiny,
carina complete;
longer than OL;
POL
granu I a ted,
with weak
impression;
pronotal
rugose; radial
tubercles reaching
scutellum
fore-wing vein much
pronotum shiny,
transversa I stri ae,
sculptured by numerous separated;
reddish-testaceous,
weakly reticulate rugose;
longitudinal
tegulae;
and metanotum shiny,
with a
brown
longer than
weakly
transversa I
scutum shiny,
fully
notaulices complete, punctate;
transversal
proximal
occipital
hairy,
crossed by a
keels;
with petiole
posteriorly
propodeum reticulate
band and with distal
part;
radial
cell
part of
closed;
pterostigma brown-I ight; hind wings with a brown distal transversal band; segment 1 of front tarsus nearly as long as segment 4; segment 3 produced into a (Fig.
hook; 460);
enlarged claw with 2 sUbapical segment 5 of front
tarsus
lamellae;apex with a group of 6 MALE:
(Fig.
lamellae;
teeth and with 32 460)
lamellae
with 2 rows of 43
tibial
spurs 1,1,2.
unknown
LOCUS TYPICUS:
St.
TYPICAL MATERIAL:
Augustin
(Madagascar)
holotype F!
in BM.
HOSTS: unknown DISTRIBUTION: Ethiopian region:
MADAGASCAR:
St.
Augustin
(Tulear),
BM!
FEMALE: fully winged; length 4,50 mm; head yellow-testaceous, with occiput partly black, ocellar region black, frons with 4 black spots
(two
3. Thaumatodryinus dentatus Benoit
Thaumatodryinus dentatus Benoit 1953b: 404.
in front of the antennal antennae testaceous, pronotum
sockets and two behind the antennal
with segments 1 and 4-10 dark;
yellow-testaceous,
testaceous,
with sides partly brown;
with anterior margin black;
between scutellum and scutum brown; and metanotum black; petiole black;
clubs of femora brown; 28:
33:
41:
incomplete, OPL
40:
28:
scutellum yellow-testaceous;
13: 9:
11;
with anterior third reddish;
legs testaceous,
head shiny, the ocell i
pronotum sh i ny,
furrow
sides of scutellum with fore and hind
antenna I segments in following proportions:
visible only behind
longer than OL;
propectus yellow; scutum yellow-
metanotum black;
propodeum black-brown,
abdomen brown-yellow;
sockets);
granulated; and on
granu I a ted,
occipital
15: 9:
carina
the sides of the vertex; ha i ry,
crossed by
a
Thaumatodryi n i nae
Fig.
461
-
Chela of
691
Thaumatodryinus dentatus n.
sp.
(type)
transversal impression; pronotal tubercles reaching tegulae; scutum shiny, hairy, granulated, with weak longitudinal striae; notaulices almost complete, sculpture;
posteriorly separated; metanotum rugose;
third almost smooth; distal closed;
part of radial
fore-wing
without dark
vein much
pterostigma brown-light;
than segment 4
(26:19);
with 2 subapical tarsus
scutellum shiny,
(Fig.
461)
transversal
longer than proximal segment
1 of front
segment 3 produced
teeth and with 36 with 2 rows of 63
apex; tibial spurs 1,1,2. MALE: unknown
smooth,
without
propodeum reticulate rugose,
into a
lamellae (Fig.
with anterior
bands and with
part;
radial
tarsus much hook; 461);
cell
longer
enlarged claw segment 5 of front
lamellae extending continuously
to the
692
Mem.
LOCUS TYPICUS:
Bekily
TYPICAL MATERIAL: HOSTS:
Amer.
Ent.
Ins!.,
37,
no.
1984
(Madagascar)
holotype F!
in P;
2 paratypes FF!
in TV.
unknown
DISTRIBUTION:
Ethiopian region:
MADAGASCAR:
Bekily,
P!
TV!
4. Thaumatodryinus townesi n. sp.
FEMALE: black;
fully
length 3,43-5,62 mm;
winged;
slender,
weakly distally thickened;
7: 5: 50: 40;
proportions:
complete; shiny,
6; OL
tubercles
smooth,
transversal
with 2 cell
smooth,
granulated,
logitudinal
keels;
=
8; OPL
tegulae;
lateral
scutum,
occipital
=
1,5; TL a
notaulices complete,
with a
carina
7; pronotum
transversal
scutellum
with
keels,
impression;
and
metanotum
very converging;
with some areolae or tracks of areolae, and posterior surface;
median
area dull,
without dark
closed and with distal
part;
weakly alutaceous,
crossed by
between dorsal
hind wings hyal ine,
=
3,5; OOL
reaching
keel
flat,
frons without median or
without sculpture;
propodeum dull, a
=
with petiole
antennae very
in front of the anterior ocellus;
without sculpture,
pronatal
shiny,
=
POL
testaceous,
segments 1-4 in following
antennal
head shiny,
occiput strongly excavated; short median furrow
fully
with segments 1-2 testaceous;
antennae brown,
part of radial
fore trochanters nearly as
smooth,
transversal long
with
posterior surface
granulated;
fore and
fore-wing
with radial
bands;
vein much
as fore femora
longer than proximal
(44:46) and nearly as
(44:46) and as fore tibiae (44:42); fore tarsal segments 16: 4: 8: 31: 41; enlarged claw (Fig. 462) with 2 subapical teeth and with 6 lamellae; segment 5 of front tarsus (Fig. 462) with a row of 9 lamellae; apex with a group of 10 lamellae; tibial spurs 1,1,2. MALE (not reared: from Sokoke Forest, Kenya): fully winged; length 2,68 long as fore coxae
in following proportions:
mm;
head black,
with mandibles testaceous;
segments 3-10 darkened; partly
testaceous
antennae testaceous,
prothorax testaceous;
thorax black,
(notaulices and region between
with
wi th scutum
the notaulices);
abdomen
brown; legs testaceous, antennae not distally thickened; antennal segments in following proportions: 6: 6: 9: 12: 11: 12: 11: 10: 8: 10; head dull, granulated and reticulate rugose; median
keel;
granulated, separated;
POL
=
weakly
6; OL
occipital
2; OOL
=
reticulate rugose;
reticulate rugose,
keels;
fore-wing hyaline,
radial
vein
longer than proximal
LOCUS TYPICUS:
Kampala
TYPICAL MATERIAL: unknown
part;
=
1; TL
=
smooth,
tibial
posteriorly
without sculpture;
transversal
transversal
frons without
4; scutum dull,
notaul ices complete,
without
without dark
carina complete;
6; OPL
scutellum and metanotum shiny,
propodeum dull,
in OL. HOSTS:
=
or
bands;
longitudinal distal
part of
spurs 1,1,2.
(Uganda)
holotype F!
and 1 paratype F!
in TW;
1 paratype F!
Thaumatodry in inae
Fig.
462 -
Chela of Thaumatodryinus
693
townesi n. sp. (type)
694
Mem. Amer.
Ent.
Inst.,
no.
37,1984
DISTRIBUTION: Ethiopian region: UGANDA: Kampala, TW! Forest, TW! SOUTH AFRICA: Tzaneen (Transvaal), TW! NOTES:
the species is named in honor of Mr.
KENYA:
Sokoke
Henry Townes.
5. Thaumatodryinus migratorius Benoit
Fig. 463 -
Chel a of Thaumatodryinus migratorius n.
Thaumatodryinus migratorius Benoi t 1953b: 404.
sp.
(type)
Thaumatodry i n i nae
FEMALE: black;
fully
winged;
ocellar region
length 3,5 mm; weakly brownish;
4-10 darkened;
antennal
22:
10: 8;
23:
20:
incomplete, OPL = OL; transversal
10:
reddish-testaceous,
with petiole
antennae testaceous,
segments in following proportions:
head shiny,
weakly punctate;
visible only behind the ocelli pronotum shiny, impression;
695
and on
weakly punctate,
pronotal
with segments 7:
occipital
5:
14:
15:
carina
the sides of the vertex;
hairy,
tubercles reaching
crossed by tegulae;
a
scutum shiny,
hairy, punctate, notaulices almost complete, posteriorly separated; scutellum and metanotum shiny, weakly punctate; propodeum reticulate rugose,
with anterior third almost smooth;
transversal proximal
bands and with distal
part;
radial cell
fore-wing
part of radial
closed;
pterostigma brown-light;
front tarsus nearly as long as segment 4;
teeth and with 21
segment 5 of front
463)
(Fig.
a group of 4 lamellae; MALE: unknown
tibial
LOCUS TYP I CUS:
(Madagascar)
Beki I y
TYPICAL MATERIAL:
segment 1 of
segment 3 produced
enlarged claw with 2 subapical tarsus
without dark
vein much longer than
lamellae
with 2 rows of 38
into a hook;
(Fig.
lamellae;
463); apex with
spurs 1,1,2.
holotype F!
in P.
HOSTS: unknown DISTRIBUTION: Ethiopian region: 'MADAGASCAR:
Bekily,
P!
THAUMATODRYINUS
OR I ENTAL SPEC I ES OF
Three species are known
KEY TO THE SPECIES
FEMALES
segment 3 less than 1,5 times as long as segment ........................... " 2. asiaticus n. sp.
Antennal -
Antennal
segment
3 more
2 Scutum with numerous
than
1,5
than POL . . . . . . . . . . . . . . . . . . . . . -
Scutum without
times as
longitudinal
longitudinal
1.
striae;
3.
segment
slightly
1 •••• 2
longer
philippinus n. sp.
striae;
long as POL .••••••••• ••••••••
long as OOL
POL more
than
malayanus n. sp.
MALES
three
times as
696
Mem.
One only male
Amer.
is known.
Ent.
It
Ins!.,
no.
37,
1984
is not possible to know
if it's a
philippinus or an asiaticus.
Fig.
464 -
Chela of
Thaumatodryinus philippinus n. sp. (type)
Thaumatodryi n i nae
697
1. Thaumatodryinus philippinus n. sp.
FEMALE:
fully
winged;
suture darkened; darkened;
length 4,80 mm;
legs yellow,
reddish-testaceous;
segment 3 more than
antennal
epistomal
with clubs of fore and hind femora 1,5 times as
long
as segment
(30: 19); head swollen, dull, granulated; frons with tracks of irregular striae; occipital carina complete; frons without median keel; POL = 9; OL = 4;
OOL = 12; OPL = 3; TL = 3; pronotum dull, hairy, granulated, strong transversal impression and with posterior tubercles reaching tegulae;
scutum granulated and with numerous
and areolae;
notaulices complete,
metanotum shiny,
granulated;
transversal
or
transversal
bands;
part 37;
(30:15);
longitudinal distal
fore tarsal
enlarged claw
23 41
lamellae; lamellae;
MALE:
(Fig.
unknown
(see key
to
Dumaguete
TYPICAL MATERIAL:
vein
without
without dark
twice as
long as proximal
in following proportions:
with 2 preapical
32:
9: 26:
5:
teeth and a row of nearly
the males) (Philippines)
holotype F!
in AM.
unknown
DISTRIBUTION: Negros
part of radial segments
hyal ine,
segment 5 of front tarsus (Fig. 464) with two rows of nearly apex with a group of 4 lamellae; tibial spurs 1,1,2.
LOCUS TYPICUS: HOSTS:
fore-wing
weak striae
scutellum and
propodeum reticulate rugose,
keels;
464)
longitudinal
posteriorly separated;
with a
Oriental
Island),
region:
PHILIPPINES:
Dumaguete
(Camp Lookout,
AM!
2. Thaumatodryinus asiaticus n. sp.
FEMALE:
fully
winged;
length 4,00-5,00 mm;
someti mes wi th brown spot on frons; testaceous;
propodeum yellow-testaceous,
yellow-testaceous;
head reddish-testaceous
antennae testaceous;
propectus
with sides partly brown;
scutum
scutell um and metanotum yellow-testaceous or reddish-
brown; propodeum fully or partly black; petiole black; abdomen reddish; legs yellow, sometimes with clubs of femora brown; head swollen, granulated,
dull,
antennal
segment 3 sl ightly
longer than segment 1 (23:20
or 24:18); occipital carina incomplete, visible only behind the ocelli and on the sides of the vertex; POL = 7; OL = 3,5; OOL = 9; OPL = 2; TL = 2,5;
propodeum shiny,
weakly
granulated,
impression;
pronotal
granulated;
notaulices complete,
almost
invisible;
reticulate rugose;
tubercles reaching
hairy,
tegulae;
crossed by a scutum dull,
posteriorly separated,
scutellum and metanotum shiny, fore-wing
with posterior parts
granulated;
without dark transversal
bands,
pterostigma brown dark and with distal
part of radial
than
segment 1 of front
as
proximal
long
part;
as segment 4
radial
cell
(24:22);
closed;
segment 3 produced
transversal
hairy, propodeum with
vein much
into a
longer
tarsus nearly hook;
enlarged
698
Mem.
Fig.
465 -
Chela of
claw with 2 subapical front
tarsus
lamellae; MALE
(Fig.
tibial
(supposed:
head testaceous,
Amer.
465)
Ent.
Inst.,
teeth and with 26
lamellae
see key
to the males):
fully
antennae slender,
proportions:
11: 6:
10:
granulated,
weakly
rugose;
OPl
2;
Tl
12:
=
4;
13:
(type)
segment 5 of
apex with a group of 5
13:
12:
abdomen and
12;
head hairy,
carina complete;
scutum hairy,
dull,
scutellum dull,
metanotum shiny,smooth,
with
legs
segments in following
10:
11:
length 2,40 mm;
antennae testaceous;
light;
antennal
occipital
posteriorly separated;
and punctate;
margin;
=
sp.
(Fig. 465);
winged;
with occiput and vertex brown; with scutum
7;
n.
spurs 1,1,2.
hairy;
=
1984
with 2 rows of 48 lamellae;
testaceous;
OOl
37,
Thaumatodryinus asiaticus
thorax and propodeum brown,
complete,
no.
POL = 7;
granulated; rugose,
dull, Ol = 5;
notaulices
reticulate rugose
some areolae near anterior
propodeum dull,fully reticulate rugose,
without transversal or
longitudinal keels; fore-wing hyaline, without dark transversal bands, hairy; pterostigma long and narrow; distal part of radial vein longer
Thaumatodry i n i nae
Fig.
than
466 -
proximal
Chela of Thaumatodryinus malayanus n.
part
LOCUS TYPICUS:
699
(19:14);
radial
Gunung Mulu Nat.
cell
open;
Park
tibial
(Sarawak,
sp.
(type)
spurs 1,1,2. Malaysia)
TYPICAL MATERIAL:
holotype F! in BM; 1 paratype F! in B; 1 para type F! 1 paratype M! in CO. HOSTS: unknown DISTRIBUTION: Oriental region: MALAYSIA: Gunung Mulu Nat. Park (Sarawak), BM! Lake Area (Bau, Sarawak), B! PHILIPPINES: Mt. Data, TW! N of Batu Batu (Tarawakan, Tawi Tawi), CO! in TW;
3.
Thaumatodryinus malayanus n. sp.
FEMALE: fully winged; length 4,60 mm; head testaceous, with ocellar triangle and a median frontal region brown; occiput and ventral side of head partly brown;
antennae brown,
with segments 1-3 testaceous and with
regions bearing tufts of hairs whitish; testaceous,
with a
lateral
propectus brown;
streak brown;
scutum,
pronotum
scutellum and metanotum
testaceous;
mesopleura and metapleura black,
testaceous; testaceous;
propodeum black, with anterior region of dorsal surface abdomen brown; legs testaceous, with cl ubs of femora brown;
antenna I segment 3 more than
1,5 times as long as segment 1 (39:20);
head swollen,
dull,
median keels;
POL = 3;
shiny,
without sculpture,
hairy,
and with a complete,
granulated;
transversal
with posterior apexes
occipital
carina complete;
OL = 1; OOL = 15; OPL = 3;
pronotum
with posterior tubercles reaching tegulae
impression;
posteriorly separated;
frons without
TL = 4;
scutum dull,
granulated;
notaulices
scutellum and metanotum smooth,
shiny,
700
Mem. Amer.
very weakly granulated; transversal or transversal part 53;
distal
fore
tarsal
enlarged claw
Ins!.,
no.
keels;
fore-wing
part of radial segments in
(Fig.
466)
with 2 preapical
segment 5 of front
apex with a group of 4
HOSTS:
hyaline,
vein much
nearly 38 lamellae;
Templer Park
TYPICAL MATERIAL:
1984
tarsus
(Selangor,
holotype F!
dull,
without
without dark
longer than proximal
following proportions:
nearly 54 lamellae; MALE: unknown LOCUS TYP ICUS:
37,
propodeum reticulate rugose,
longitudinal
bands;
(50:22);
En!.
38:
6:
13: 34:
teeth and with a row of (Fig.
466)
lamellae;
Malaya,
with
tibial
two rows of
spurs 1,1,2.
Malaysia)
in CO.
unknown
DISTRIBUTION: Malaya),
Oriental
region:
MALAYSIA:
Templer Park
(Selangor,
CO!
NEARCT I C SPEC I ES OF THAUMATODRYINUS
One species is known.
Thaumatodryinus perkinsi Krombein
Thaumatodryinus perkinsi Krombei n 1952: 99.
FEMALE:
fully
winged;
petiole black; occipital
carina
of the vertex; impression, shiny,
length 4,37-5,62 mm;
pterostigma brown incomplete,
granulated;
granulated; fore-wing
fully
yellow-testaceous,
head swollen,
shiny,
visible only behind the ocell i
with
granulated;
and on
the sides
= OL; pronotum sh i ny, ha i ry, crossed by a transversa I
OPL
pronotal
tubercles reaching
notaulices almost complete,
scutellum and metanotum shiny, rugose;
dark;
with distal
weakly punctate;
part of radial
tegulae;
scutum hairy,
posteriorly separated; propodeum reticulate
vein much
longer than
proximal part and without dark transversal bands; segment 1 of front tarsus longer than segment 4 (23:21); segment 3 produced into a hook; enlarged claw
with 2 subapical
segment 5 of front a group of 5 MALE:
lamellae;
(Fig.
tibial
teeth and with 467)
19 lamellae
(Fig.
with 2 rows of 28 lamellae;
467); apex with
spurs 1,1,2.
unknown
LOCUS TYP ICUS:
Westmoreland State Park
TYPICAL MATERIAL: HOSTS:
tarsus
holotype F!
(U.
S.
A.)
in WA.
unknown
DISTRIBUTION:
Nearctic region:
Westmoreland State Park (Oaxaca),
TE!
U.
S.
A.:
Greenville
(Westmoreland Co.,
Va.),
(S.
WAr
C.),
LE!
MEXICO:
Miltepec
Thaumatodryininae
Fig.
467 -
Chela of
701
Thaumatodryinus perkinsi n. sp.
(type)
Th. perkinsi Krombein is very similar to Th. koebelei R.C.L. Th. perkinsi, however, has pterost i gma brown dark and transversal impression of pronotum deeper. Th. perkinsi is also similar to Th. rufus Richards from Neotropic region. Th. rufus, NOTES:
Perk ins from Austral i an region.
however,
has transversa I
impression of pronotum
less deep.
702
Mem.
Amer.
En!.
Inst.,
no.
37,
1984
THAUMATODRYINUS
NEOTROPIC SPECIES OF
Six species are known.
KEY TO THE SPECIES
FEMALES
-
Not a u I ice s
c omp let e •••••••••••••••••••••••••••••••••••••••••••• 2
Notaulices
incomplete ••••••••••••••••••••.•••••••••••••••••••.. 4
2 Antennal
segment
3
. . . . . . . . . . . . . . . . . .. -
Antennal
I ittle
longer
than
2. minimus n.
segment
1 •••••••••••••••••
sp.
segment
3 more than 1,5 times as long as segment 1 . . . . 3 long as breadth of ocell i; radial cell wide (Fig. 468 B} ••••••••••••••••••••••••.•••••••••••••••••.•••• . . . . . . . . . . . . . . . . . . . 1. variegatus De Santis and Vidal Sarmiento - Head wi th POL nearl y three times as long as breadth of ocel I i ; radial cell narrow (Fig. 468 A} . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . • . . . . . . • . • • • • • • •• 6. clarus n. sp. 4 Segment 1 of front tarsus nearly as long as segment 4; head black or brown-b lack •••••••••••••••••••••••••••••••••••••••••••• . . . . . . . . . . . . . . . . . . . 3. macilentus De Santis and Vidal Sarmiento - Segment 1 of front tarsus much longer than segment 4; head fully reddi sh or reddi sh-testaceous •••••••••••••••••••••••••••••••••• 5 5 Head with OL shorter than OPL ••••••••••••••••••••••••••••••••••• . . . . .. .. .. .. .. .. ... 4. rufus Richards - Head wi th OL IOr)ger than OPL . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. 5. bruchi De Santis and Vidal Sarmi en t 0
3 Head wi th POL nearly as
MALES
Antennal
segment
head with OOL
4
less
than
less
than
three
................... -
Antennal
segment
2 Head wi th OL as
5.
4 more
1,5
times as
times as
long as
bruchi De Santis and Vidal than
1,5
segment
times as
long as
Sarmiento segment
Head wi th OL
3 .... 2
long as OPL •••••••••••••••••••••••••••••••••••••
. . . . . . . . . . . . . . . . . . . 3. macilentus De Santis and Vidal -
3;
long as OL . . . . . . . . . . . . . . .
longer
Sarmiento
than OPL ••••••••••••••••••••••••••••••••••••
4. rufus Ri chards
703
Thaumatodry i n i nae
\
\ \
\
\ \
\
\ \
\
:.:
...
Fig. 468 - Radial cells of females of Thaumatodryinus clarus n. sp. (type) (A) and variegatus De Santis and Vidal Sarmiento (type) (B)
The males of Th. n.
sp.
and minimus n.
variegatus De Santis and Vidal Sarmiento, clarus sp.
are unknown
1. Thaumatodryinus variegatus De Santis and Vidal Sarmiento
704
Mem.
Amer.
Ent.
I nst.,
no.
37,
1984
Thaumatodryinus variegatus De Santis and Vidal Sarmiento 1974: 25.
FEMALE:
fully
reddish;
antennae brown,
medial
winged;
length 7,18 mm; with joints,
region of segment 7,
head,
thorax and propodeum half of segments 5-6,
proximal
segments 8-10 testaceous;
petiole black;
abdomen reddish-testaceous; legs testaceous, with clubs of femora brown, distal half of tibiae black; antennal segment 3 more than 1,5 times as long as segment 1 (22:13); head dull, granulated; frons with a track of median keel;
occipital
POL = 5;
carina complete;
3; POL nearly as long as breadth of ocelli pronotum dull,
hairy,
with a strong
OL = 3;
(5:5);
transversal
OOL = 10;
temples
OPL
invisible;
impression,
with numerous
irregular keels; pronotal tubercles reaching tegulae; scutum shiny, granulated and with numerous irregular weak keels; notaul ices complete, posteriorly separated; reticulate rugose, wing
longitudinal
with first recurrent vein
fore-wing hyaline, vein
scutellum and metanotum shiny,
without
longer than distal
without dark transversal
longer than proximal
part
punctate;
propodeum
keels on posterior surface;
(30:12);
bands;
radial
fore-
part of discocubitus;
cell
distal open
part of radial (Fig.
468 B);
26: 3,5: 5,5: 19: 26; enlarged claw (Fig. 469) with 2 subapical teeth and with 26 lamellae; segment 5 of front tarsus (Fig. 469) with 2 rows of 53 lamellae; apex with a group of 4 lamellae; tibial spurs 1,1,2. fore tarsal
segments
MALE: unknown LOCUS TYPICUS:
in following proportions:
Dos de Mayo (Argentina)
TYP ICAL MATER IAL:
holotype F!
in PT.
HOSTS: unknown DISTRIBUTION: Neotropic region:
ARGENTINA:
Dos de Mayo (Misiones),
PT!
2. Thaumatodryinus minimus n. sp.
FEMALE:
fully
winged;
reddish-testaceous; black;
length 4,12 mm;
mesothorax;
head,
antennae and prothorax
metathorax and propodeum black;
abdomen brown-testaceous;
legs testaceous,
petiole
with tibiae and stalk of
fore femora brown; antenna I segment 3 sl i ght I y longer than segment 1 (11:9); read dull, granulated, with tracks of weak irregular striae, convex; frons without median and lateral keels; occipital carina complete; POL = 6; OL = 4; OOL = 10; OPL = 2; TL = 3; pronotum dull, granulated, with a strong scutum dull,
transversal granulated;
impression;
scutellum and metanotum dull,
distal open;
fore-wing
vein
segments
tegulae;
without sculpture among
without transversal
with a dark transversal
part of radial fore tarsal
tubercles reaching
posteriorly separated;
weakly punctate,
punctures or very reticulate rugose, keels;
pronotal
notaulices complete,
or longitudinal
band beneath the pterostigma;
longer than proximal
part
in following proportions:
(26:15); radial cell 23:
3:
7:
19:
30;
Thaumatodry i n i nae
Fig.
469 -
Chela of Thaumatodryinus variegatus De Santis and Vidal Sarmiento (type)
705
706
Mem.
Fig.
(Fig.
segment 5 of front
Ent.
470)
no.
with two subapical
tarsus
(Fig.
470)
apex with a group of 3 lamellae; MALE:
Ins!.,
37,
1984
Chela of Thaumatodryinus minimus n.
470 -
enlarged claw
Amer.
sp.
(type)
teeth and with 25
with 2 rows of nearly 44
tibial
lamellae; lamellae;
spurs 1,1,2.
unknown
LOCUS TYP I CUS: Tucuru i (Brazi I ) TYPICAL MATERIAL: holotype F! in TW HOSTS:
unknown
DISTRIBUTION:
Neotropic region:
BRAZIL:
Tucurui
(Para),
TW!
3. Thaumatodryinus macilentus De Santis and Vidal Sarmiento
Thaumatodryinus macilentus De Santis and Vidal Sarmiento 1974: 24.
FEMALE: space,
fully
winged:
clypeus and
4-10 darkened;
length 6,25-6,87 mm;
ma~dibles
reddish;
brown;
antenna I tufts: incomplete, OPL
transversal tegulae;
petiole black;
with segments mesothorax,
abdomen reddish;
legs
with clubs of femora and partly clubs of hind femora black-
carina vertex;
with malar
propectus and pronotum redd ish-testaceous;
meta thorax and propodeum black; testaceous,
head black,
antennae testaceous,
fig.
471;
head shiny,
granulated,
visible only behind the ocelli
longer than OL; impression,
scutum hairy,
pronotum sh i ny,
weakly punctate; shiny,
reaching at
and metanotum shiny,
weakly punctate;
ha i ry,
pronotal
granulated,
notaulices incomplete,
least 0,5
and on
swollen;
occipital
the sides of the
crossed by
a
tubercles reaching
with numerous irregular keels; length of scutum;
scutellum
propodeum reticulate rugose;
fore-
T ha uma todryi n i nae
Fig.
471
707
Tuft of long hairs on antenna of female of Thaumatodry inus macilentus De Santis and Vidal Sarmiento from Brazi I -
708
Mem. Amer.
Fig.
472 -
En!.
Inst.,
no.
37,
1984
Chela of Thaumatodryinus macilentus De Santis and Vidal Sarmiento from Nova Teutonia
(Brazil)
709
Thaumatodry in inae
Fig. 473 - Male genitalia of Thaumatodryinus macilentus De Santis and Vidal Sarmiento from Nova Teutonia (Brazi I): right half removed.
wing with a brown transversal distal part of radial
vein much
band
(rarely band very
longer than proximal
light)
part;
and with
radial
cell
710
Mem.
Amer.
En!.
closed;
pterostigma brown dark;
bands;
segment 1 of front
produced into a hook; lamellae
(Fig.
lamellae; MALE:
472);
Ins!.,
no.
winged;
1984
hind wings without dark transversal
tarsus nearly as
long as segment 4;
enlarged claw with 2 subapical
segment 5 of front
tarsus
apex with a group of 6 lamellae;
fully
37,
length 4,87;
(Fig.
tibial
head reddish,
segment 3
teeth and with 40
472)
with 2 rows of 67
spurs 1,1,2.
with ocellar region black;
antennae testaceous, wi th segments 3-10 dark; thorax and propodeum reddish, with scutellum and metanotum black, scutum dark; abdomen brown,
with segment
tibiae and tarsi
1 testaceous;
brown;
in following
proportions:
granulated;
smooth;
carina complete; dull,
legs testaceous,
8:
frons
7:
13: 23:
19:
OL = 2;
OOL = 7;
notaulices complete,
distance between the notaulices nearly as smooth, without
19:
without median and
POL = 7;
fully granulated;
notau Ii ces and
with mid and hind
antennae not distally thickened;
the paraps i da I furrows;
17:
18:
lateral
antennal
14:
17;
keels;
OPL = 2;
segments
head,
fully
occipital
TL = 4;
scutum
posteriorly separated;
minimum
long as distance between
the
scutell um and metanotum sh i ny,
without sculpture, finely hairy; propodeum reticulate rugose, dull, transversal or longitudinal keels; fore-wing hyaline, without dark
transversal bands; distal part of radial vein longer than proximal (36:21); genital ia: fig. 473; tibial spurs 1,1,2. LOCUS TYP ICUS:
Loreto (Argentina)
TYPICAL MATERIAL: HOSTS:
part
holotype F!
and
1 paratype F!
in PT.
unknown
DISTRIBUTION: ECUADOR:
Neotropic region:
Napo
(Muyuna,
Tena),
PANAMA: BM!
Barro Colorado
Island,
SURINAME: Sumatraweg
LE!
(Lelydorp),
OL! BRAZIL: Campsite (Mato Grosso), BM! Jatai (Goias), TW! OT! Nova Teutonia (S.ta Catarina), BM! 12° 50' S 51° 47' W (Mato Grosso), BM! OL! ARGENT INA:
Loreto
(Misiones),
PT!
4. Thaumatodryinus rufus Richards
Thaumatodryinus rufus Richards 1951: 815. Thaumatodryinus rufus Richards: Krombein 1952: 99.
FEMALE:
fully
winged;
length 3,0-4,0 mm;
reddish-testaceous,
with petiole
black, clubs of fore and hind femora brown; antennae testaceous, with segments 4-10 darkened; mesothorax, metathorax and propodeum reddish dark; antennal segment 3 more than 1,5 times as long as segment 1 (25:14); head swollen, shiny, granulated, with frontal weak longitudinal striae;
occipital
carina
incomplete,
the sides of the vertex; shiny,
hairy,
pronotal numerous
crossed by
POL = 4; a
tubercles reaching
visible only benhind the ocell i and on
OL = 2;
transversal tegulae;
irregular weak striae;
OOL = 8,5;
impression,
scutum hairy,
OPL = 3;
pronotum
weakly punctate; shiny,
notaul ices incomplete,
granulated,
reaching
at
with
least
Thaumatodry i n i nae
711
\
Fig.
474 -
Chela of Thaumatodryinus rufus Richards from St. Augustine (Trinidad)
0,5 length of scutum; scutellum and metanotum hairy, shiny, weakly punctate; propodeum reticulate rugose; fore wings with a brown transversal band and with distal part of radial vein much longer than proximal part; radial cell closed; pterostigma brown dark; hind wings without dark transversal bands; segment 1 of front tarsus longer than segment 4 (27:21); segment 3 produced into a hook; enlarged claw (Fig. 474) with 2 subapical teeth and with 29 lamellae; segment 5 of front tarsus (Fig. 474) with 2 rows of 45 lamellae; apex with a group of 5 lamellae; MALE:
tibial
fully
spurs 1,1,2.
winged;
length 2,50-2,87 mm;
head reddish-testaceous,
with
712
Mem. Amer.
Fig.
475 -
Ent.
Ins!.,
no.
37,
1984
Male genitalia of Thaumatodryinus rufus Richards from St. Augustine
(Trinidad):
right
half removed.
ocellar region black;
antennae testaceous;
with scutellum black,
metanotum and propodeum dark;
testaceous;
testaceous;
legs fully
thorax and propodeum reddish, abdomen brown-
antennae not distally thickened;
antennal 11: 10: 9: 9: 7: 10; head dull, fully granulated; frons without median and lateral keels; occipital carina complete; POL = 5; Ol = 2; OOl = 5; OPl = 1,5; Tl = 2; scutum dull, granulated; notaulices complete, posteriorly separated;minimum distance between the notaulices nearly as long as distance between the notaulices and the parapsidal furrows; scutellum and metanotum shiny, smooth, weak I y punctate, without SCu I pture among punctures; propodeum reticulate rugose, without transversal and longitudinal keels; fore-wing hyaline, without dark transversal bands; distal part of radial vein longer than proximal part (19:12); genitalia: fig. 475; tibial spurs 1,1,2. lOCUS TYPICUS: St. Augustine (Trinidad) segments in following proportions: 7:
5:
8:
14:
Thaumatodry i n i nae
Fig. 476 -
713
Chela of Thaumatodryinus bruchi De Santis and Vidal Sarmiento from Las Cesas
(Argentina)
TYPICAL MATERIAL: holotype F! in BM. HOSTS: in Trinidad Ormenis sp. (Richards 1951). DISTRIBUTION: Neotropic region: TRINIDAD: 51. Augustine (51. BM!
Curepe
(Sta.
Margarita,
Circular Rd.),
OT!
OLl
George),
OL!
714
Mem.
Amer.
En!.
Inst.,
no.
37, 1984
5. Thaumatodryinus bruchi De Santis and Vidal Sarmiento
Thaumatodryinus bruchi De Santis and Vidal Sarmiento 1974: 22.
FEMALE:
fully
winged;
length 6,56-8,62 mm;
head reddish;
antennae
testaceous, wi th segments 3-10 darkened; propectus and pronotum redd i sh; thorax (except for scutum which is reddish) and propodeum reddish-brown or black, reddish;
sometimes with
irregular brown spots;
legs testaceous,
clubs of hind femora brown-black; as
long as segment 1 (38:20);
longitudinal behind = 13;
keels on frons;
the ocelli OPL = 2;
impression,
and on
petiole black;
abdomen
with clubs of fore femora and distal
antenna I segment 3 more than
head swollen,
occipital
carina
dull,
granulated,
weakly punctate;
sh i ny,
pronotal
1,5 times
with some
incomplete, only visible
the sides of the vertex;
pronotum ha i ry,
part of
POL = 5;
crossed by
OL = 3,5;
OOL
a transversa I
tubercles reaching tegulae;
scutum
hairy, dull, reaching at least 0,5 length of scutum; scutellum shiny, smooth; metanotum with lateral keels; propodeum reticulate rugose forewing with a brown transversal much
longer than proximal
dark;
band and with distal part of radial
part;
radial
cell
hind wings without dark transversal
closed; bands;
vein
pterostigma brown segment
1 of front
(37:30); segment 3 produced into a hook; enlarged claw with 2 subapical teeth and with 29 lamellae (Fig. 476); segment 5 of front tarsus (Fig. 476) with 2 rows of 44 lamellae extending continuously to the apex; tibial spurs 1,1,2. MALE: fully winged; length 3,37 mm; head reddish-testaceous; antennae brown, with segments 1-2 testaceous; ocellar region usually dark; tarsus
longer than segment 4
prothorax reddish-testaceous; brownish;
scutellum,
propodeum testaceous; testaceous,
scutum reddish-testaceous,
metanotum,
petiole black;
darkened;
head dull,
granulated;
7: 7:
11 :16:
legs
mid and hind tibiae,
antennae not distally thickened;
segments in following proportions:
sometimes
abdomen brown-testaceous;
with mid and hind stalks of femora,
and hind tarsi
with sides
propodeum and pleura black;
mid
antennal
14: 14: 12: 12: 11: 14;
frons without median and
lateral
keels;
occipital
carina complete; POL = 7; OL = 3; OOL = 6; OPL = 1,5; TL = 4; scutum dull, granulated; notaul ices complete, posteriorly separated; scutellum and metanotum shiny, punctate, very weakly granulated; propodeum reticulate rugose, without transversal or longitudinal keels; fore-wing hyaline, without dark transversal proximal
part
bands;
LOCUS TYPICUS:
Alta Gracia
TYPICAL MATERIAL:
(Roraima),
Horco Molle
part of radial
vein
longer than
(Argentina)
holotype F!
HOSTS: unknown DISTRIBUTION: Neotropic region: Surumu
distal
(26-15); genitalia: fig. 477; tibial spurs 1,1,2. and 1 paratype F! BRAZIL:
in PT.
Encruzilhada (Bahia),
OL!
TW!
ARGENTINA:
Las Cejas
(Tucuman),
TU!
Alta Gracia
(Cordoba),
(Tucuman), PT!
TW! OL!
La Paz
TU!
(Cordoba),
715
Thaumatodry i n i nae
'/
Fig.
477 -
Male genital ia of Thaumatodryinus
Sarmiento from La Plata
PT!
La Plata,
TW!
(Argentina):
bruchi De Santis and Vidal right half removed
OU
6. Thaumatodryinus clarus n. sp.
716
Mem. Amer. En!.
Ins!., no. 37,
1984
Fig. 478 - Chela of Thaumatodryinus clarus n. sp.
(type)
Thaumatodry i n i nae
FEMALE:
fully
winged;
length 4,52 mm;
testaceous,
antennae with segment 3 more than 1,5 times as head dull, complete; times as strong dull,
granulated; POL
= 9;
OL
impression;
granulated;
=
or longitudinal
transversal
bands;
proportions:
cell:
radial fig.
= 3;
OPL
(9:3);
keels;
occipital carina
= 5;
POL nearly three
pronotum hairy,
dull,
with a
tegulae;
posteriorly separated;
fore tarsal
hyaline,
scutum
scutellum and
propodeum reticulate rugose,
fore-wing
vein with distal
468 A;
TL
posterior tubercles reaching
weakly granulated;
transversal radial
12;
notaulices complete,
metanotum shiny,
part;
OOL
with petiole black;
long as segment 1 (25: 15);
frons without median keel;
= 3;
long as breadth of ocelli
transversal
717
without
without dark
part much longer than proximal segments in following
5: 12: 28: 43; enlarged claw (Fig. 478) with 2 subapical segment 5 of front tarsus (Fig. 478)
3:
teeth and with nearly 24 lamellae;
with two rows of nearly 409 lamellae extending continuously to the apex; tibial spurs 1,1,2. MALE: unknown LOCUS TYPICUS: Yepocapa (Guatemala) TYPICAL MATERIAL: holotype F! in AM. HOSTS:
unknown
DISTRIBUTION:
Neotropic region:
Chimaltenango),
GUATEMALA:
Yepocapa
(Dep.
AM!
AUSTRAL I AN SPEC I ES OF THAUMATODRYINUS
Two species are known
KEY TO THE SPECIES
FEMALES
Fore-wing with
shorter than proximal 1. flavus n. sp. Fore-wing with first recurrent vein longer than proximal part of cubi tus (Fig. 479 A) ••••••••••••• 2. koebelei R.C.L. Perkins
part -
of
cubitus
first
(Fig.
recurrent
vein much
479 B) •••••
MALES
-
brown . . . . . . . . . . . . . . . flavus n. sp.
Head,
thorax and propodeum most Iy or 1.
fully
Head,
thorax and propodeum mostly
fully
or
testaceous ••••••••••
718
Mem.
Fig.
479 -
Nadzab,
Amer.
Ent.
Ins!.,
no.
37,
1984
Females of Thaumatodryinus koebelei R.C.L. and flavus
New Guinea) (A)
CD = first recurrent vein;
n.
sp.
Perkins
(from Waris,
DE = proximal
(from
New Guinea) (B):
part of cubitus.
1. Thaumatodryinus flavus n. sp.
FEMALE
(Figs 19, 457):
yellow-testaceous,
fully winged;
clypeus and mandibles yellow; 5-10 darkened;
length 6,30-6,87 mm;
head brown or
with ocellar spot brown and occiput partly brown: antennae testaceous,
propectus black or brown;
or with sides and partly anterior collar brown; yellow-testaceous or brown-b lack;
with segments 4-8 or
pronotum fu II y
yellow-testaceous
scutum and scutellum
metanotum brown-reddish or brown-b lack;
propodeum and pet iole brown-b I ack or black;
abdomen fu II y
redd ish or
with tergite 1 partly black;
legs yellow-testaceous or with fore clubs of
femora brown;
dull,
head swollen,
visible only behind the ocelli than OL; weakly shiny,
pronotum shiny,
granulated; granulated;
hairy,
pronotal
granulated; occipital
and on
carina
the sides of the vertex;
crossed by
a transverse
tubercles reaching tegulae;
notaul ices complete,
incomplete, OPL
longer
impression,
scutum hairy,
posteriorly separated;
scutellum
and metanotum shiny, weakly punctate; propodeum reticulate rugose; forewing with first recurrent vein much shorter than proximal part of cubitus (Fig.
479 B);
fore-wing
without dark transversal
brown dark and with distal part of radial part;
radial
cell
closed;
bands,
with pterostigma
vein much longer than proximal
segment 1 of front
tarsus
longer than segment 4
(35:32); segment 3 of front tarsus produced into a hook; enlarged claw (Fig. 480) with 2 subapical teeth and with 38-41 lamellae; segment 5 of front
tarsus
(Fig.
of 5-6 lamellae; MALE
480)
tibial
(Figs 28,458):
with 2 rows of 55-57
fully
a more or less wide frontal testaceous;
lamellae;
apex with a group
spurs 1,1,2. winged;
length 1,56-2,87 mm;
area testaceous;
occiput testaceous;
thorax and propodeum brown;
antennae brown, abdomen brown,
head brown,
with
clypeus and mandibles with segment 1 testaceous; wi th anterior region
T haumatodryi n i nae
Fig.
480 -
Chela of Thaumatodryinus
719
flavus n. sp. (type)
testaceous; legs testaceous; antennae not distally thickened; antennal segments in following proportions: 6: 6: 10: 13: 13: 13: 13: 11: 9: 10; head dull, hairy, granulated; frons without median and lateral keels occipital carina complete; POL = 5; OL = 2,5; OOL = 6; OPL = 1; TL = 3; scutum dull,hairy,
granulated;
min imum distance between
notaulices complete,
posteriorly separated;
the notau I ices of scutum shorter than distance
720
Mem.
Fig.
481
-
Islands) (A);
the notaulices and
metanotum shiny,
smooth,
distal
B:
Ins!.,
no.
1984
flavus n. sp. from Kira-Kira
furrows;
scutellum and
without sculpture among punctures;
without
transversal
posterior surface without
without dark transversal
37,
gonoforceps laterally viewed.
the parapsidal
punctate,
propodeum reticulate rugose, posterior surface; hyaline,
En!.
Male genitalia of Thaumatodryinus (Solomon
between
Amer.
bands;
keel
between dorsal
longitudinal
keels;
pterostigma lanceolate,
part almost reaching apex of radial
vein;
radial
and
fore-wing with
vein with distal
part longer than proximal part (29:13), almost reaching margin of wing; gonoforceps (Fig. 481) without proximal or distal processes, laterally viewed with a medial lOCUS TYPICUS:
TYPICAL MATERIAL: MM!
in TW;
prominence;
Tangtalau
holotype F!
6 para types
tibial
(Solomon
(1
F,
spurs 1,1,2.
Islands)
and 5 paratypes MM! 5 MM)!
in B;
4 paratypes
in Ol.
HOSTS: unknown DISTRIBUTION: Australian
region: NEW GUINEA: Wau, TW! all Jimmi Valley, all TW! Dumpu, TW! Waris (Hollandia, Western Irian), Ol! SOLOMON ISLANDS: B!
Ol!
Tangtalau
(Malaita
Island),
B!
Kira-Kira
(San Cristobal
Island),
Thaumatodryininae
Fig.
482 -
721
Chela of Thaumatodryinus koebelei R.C.L. Namatasopa
Perkins from
(New Hebrides)
2. Thaumatodryinus koebelei R.C.L. Perkins
Thaumatodryinus koebelei R.C.L. Perkins 1905: 58. Thaumatodryinus koebelei R.C.L. Perkins: Krombein 1952: 99.
FEMALE:
fully
testaceous,
winged;
length 3,47-6,00 mm;
lectotype fully
reddish-
with petiole black and antenna I segments 5-10 darkened;
other specimens head with ocellar spot brown, brown,
spots on si des of pronotum,
brown;
si des of scu tum brown ish;
transversal
in
impression
furrow between scutum and scutell um
furrow between scutell um and metanotum
722
Fig.
Mem.
483 -
Amer.
Ins!.,
no.
37,
1984
Male genital ia of Thaumatodryinus koebelei R.C.l. Nadzab
brown;
Ent.
(New Guinea):
right
propodeum brown or black abdomen brown;
femora brown;
head dull,
granulated,
Perkins from
half removed
swollen;
legs with clubs of fore
occipital
carina
incomplete,
visible only behind the ocelli and on the sides of the vertex; OPl longer than Ol; pronotum not humped, crossed by a transversal impression, hairy,
granulated;
pronotal
tubercles reaching
tegulae;
scutum dull,
hairy, granulated; notaulices incomplete (reaching at least 0,5 length of scutum) or complete (in a specimen from New Britain); scutellum and metanotum shiny,
weakly
punctate;
propodeum reticulate rugose fore-wing
with first recurrent vein longer than proximal part of cubitus (Fig. 479 A); fore-wing without dark transversal bands, with pterostigma yellow dark or brown and with distal part of radial vein much longer than proximal part; radial cell closed; segment 1 of front tarsus slightly longer than segment 4
(27:22);
with 2 subapical tarsus
(Fig.
lamellae;
482)
tibial
segment 3 produced
teeth and with 30 with 2 rows of 48
into a
lamellae lamellae;
(Fig.
hook; 482);
enlarged claw segment 5 of front
apex with a
group of 5
spurs 1,1,2.
MALE:
fully
brown
and antennal
winged;
length 2,25-3,62 mm;
testaceous,
segments 3-10 darkened;
with ocellar region
antennae not distally
I
Dry i n i nae thickened; 17:
16:
antennal
14:
12:
13;
segments in following proportions: head dull,
frons without median and 7;
=
OL
3;
=
OOL
7;
lateral
=
OPL
2;
keels;
=
TL
4;
keels;
fore-wing hyaline, vein
6:
10:
occipital
13:
carina complete;
scutum dull,
granulated;
scutellum and metanotum shiny,
propodeum reticulate rugose,
radial 1,1,2.
6:
15:
granulated and weakly reticulate rugose;
complete posteriorly separated; punctate;
723
without transversal
without dark transversal
longer than proximal
part;
genital ia:
bands; fig.
POL =
notaulices strongly
or longitudinal distal
483;
part of
tibial
spurs
LOCUS TYPICUS: Cairns (Australia) TYPICAL MATERIAL: lectotype F! in B. HOSTS:
in Australia
DISTRIBUTION: GUINEA:
Siphanta sp.
Australian
BUlolo, OL!
Lae-Zenag Road,
TW!
TW!
(R.C.L.
region: Bogia
(Om.
Busu R.
Perkins 1906c).
AUSTRALIA:
Cairns
New Guinea),
(Lae),
OL!
TW!
SUBFAMILY
Nadzab,
NEW HEBRIDES:
(Anelgaunat, Aneityum Island), BM! Namatasopa OL! NEW BRITAIN: Vunabakan (Keravat), B!
7.
(Queensland),
TW!
B!
TW!
NEW
OL!
Red Crest
(Espiritu Santo
Island),
DRYININAE KIEFFER
Dryininae partim Kieffer in Kieffer et Marshall 1906: 495. Dryininae partim Kieffer 1907b: 3. Lestodryinini partim Kieffer 1914b: 11. Dryinini partim Richards 1939: 189. Dryinini partim Muesebeck and Walkley 1951: 1034. Dryininae Richards 1953: 51. Dryininae partim Krombein 1979: 1243.
TYPE GENUS: FEMALE
Dryinus Latreille 1804
(Figs 495,
wing with costal,
I
pigmented veins palpi
529,
611,
632,
(Figs 611,
with 3 segments; front
612);
spurs 1,1,2 or 1,1,1MALE (Figs 531, 532, 613,
posterior veins
labial
occipital
fully
winged;
396 B);
with 6 segments;
fore-
lamella;
634,
palpi
segment 5 of front
680,
pronotal
705):
fully
with 3 segments;
dorsal
incomplete
(Fig.
tarsus
winged:
485);
(Fig.
tubercles present;
occipital
labial
larger from
carina complete or
mandibles with 3 teeth progressing
(Fig.
with costal,
703):
chela with rudimentary claw
long hairs;
never with one only preapical
incomplete;
678,
maxillary palpi
5);
tarsus chelate;
antennae without tufts of
with 6 segments;
662,
mandibles with 4 teeth progressing
anterior one to posterior (Fig. or absent;
659,
median and submedian cells clearly enclosed by
485)
tibial
maxi Ilary palpi carina complete or
larger from anterior one to
process of gonoforceps absent;
fore-wing
median and submedian cells clearly enclosed by pigmented
(Figs 532,613);
tibial
spurs 1,1,2.
724
Mem.
DISTRIBUTION:
Amer.
Ent.
Inst.,
no.
37,
1984
worldwide
HOSTS: Dictyopharidae,
Cixiidae, Lophopidae, Flatidae, Acanaloniidae, Issidae, Ricaniidae, Delphacidae, Tropiduchidae, Fulgoridae. GENERA:
8
SPEC I ES:
149
I<EY TO THE GENERA OF DRYININAE the fossi I genus Cretodryinus Ponomarenko)
(excluding
FEMALES
Enlarged
659, -
Enlarged
611,
claw more
than
three
660) . . . . . . . . . . . . . . . . . . . 632,
claw as
662,
long as
front
tibia
(Figs
Megadryinus Richards
or shorter than front tibia (Figs 528, 703) . . • . . . . • . . • . • • • • . • • . . • . • • . • • . . . . . . . . . . . 2
long as
678,
2 Enlarged claw very than arol ium
times as
4.
(Fig.
reduced,
nearly as
711) •••••
7.
long as or
sl ightly
longer
Perodryinus R.C.L. Perkins
reduced, much longer than arolium (Fig. 485).3 1 , 1 , 1 . . . . . . . . . . 2. Richardsidryinus Mocza r - Tibial spurs 1,1,2 •••••..••••••••.•••••••••••••.•••••••••.•.••. 4 4 Enlarged claw with one subapical tooth (Fig. 485) . . . . . . . . . . . . . . 5 -
Enlarged
3 Tib ia I
-
claw not
spu rs
Enlarged claw without teeth near apex (Figs
subapical
tooth
or with a
series of
small
669,691) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6
5 Notaulices at least partly visible . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1. Dryinus Lat rei I Ie Notaul ices invisible! . . . . . . 3. Tridryinus Kieffer 6 Notaulices at
least
partly
visible . . . . . . . . . . . . . . . . . . . . . . . . . . . . ..
. . . . • . . . . . • . . . • • . . . . . . • . • . .. 5. Alphadryinus n. g en. - Notaul ices invisible 1 . . . . . . 6. Mesodryinus Kieffer
NOTES:
the key doesn I t
inc I ude the fossi I genus Cretodryinus Ponomarenko,
because the only known specimen
is in
too bad conditions for
distinguishing generic differences.
! If the scutum
is sculptured by
longitudinal
keels,
two of the furrows
del imitated by the keels are considered notaul ices and the species is included in Dryinus Latrei lie or Alphadryinus n. gen. In some species with scutum reticulate rugose or smooth two keels at first sight can seem notaulices. I have considered, however, the notaulices invisible, the true notau I ices are furrows and not keel s.
because
Dryininae
725
MALES
I
Dryininae. The
was not able to find generic differences in males of
number of known males is too small
for drawing conclusions.
In addition
perhaps some males were associated by mistake with some genera. For the above mentioned reasons keys to males of species living
I think that actually only general
in a zoogeographic region are proposable,
as follows.
DRYININAE
KEY TO THE MALES OF THE KNOWN PALAEARCTIC
Scutum strongly as -
reticulate
rugose;
numerous areolae are as wide
ocelli •.•••••••••••••••••••••••••••••••••••••••••••••••••••• 2
Scutum granulated or weakly than
reticulate
rugose;
areolae
less wide
ocelli •••••••••••••••••••••••••••••••••••••••••••••••••••• 3
2 Posteri or
ocell i
not
areolae as wide as longer
than
touching
those of
segment
4,
less
the
occipi tal
scutum; than
carina;
antennal
three
times as
2; gonoforceps slightly shorter than penis
head wi th
3 slightly
segment
(Fig.
long as
segment
686) ••••••..••
· ............••.....•.••...•••. Mesodryinus niger (K i ef f er) -
Posterior ocell i areolae much approximately
touching
smaller
than
twice as
the
occipi tal
those of
long as
carina;
scutum;
segment
head wi th
antennal
segment
3
4, approximately four
long as segment 2; gonoforceps much shorter than penis (F i g. 684) •••••••.•••••••.••••• Mesodryinus dayi n. sp. 3 Fore wing with distal part of radial vein nearly as long as proximal part (Fig. 490 B) ••••• Dryinus tarraconensis Marshall times as
-
Fore wing wi th distal
part of radial vein much longer than 490 A) ••••••••••••••••••••••••••••••••••••• 4 4 Notaulices reaching approximately 0,5 length of scutum •••••.••• 5 - Notaulices reaching approximately 0,75 length of scutum •••••••• 7 5 Distivolsella farther from apex of penis (Fig. 491) . . . . . . . . . . . . . . . . . . . . . . . . • . . . . . . . . . . . . . . . . . . . Dryinus collaris (Linnaeus) - Distivolsella nearer apex of penis (Figs 501,618) . . . . . . . . . . . . . 6 6 Head more or less retiCUlate rugose ••••••.••.••••••••••••••••••• proximal
part
(Fig.
· . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Richardsidryinus corsicus (Ma rsha I I ) -
Head granulated,
not
reticulate
rugose ••.•••••••••••••••••••••••
· .•.........•...•.•..•.•..•..•. Dryinus koreanus (Moe za r) 7 Antennal
segment
3 nearly
twice as
long as
segment
2 . . . . . . . . . . ..
· ...•.••..•.••• , •.••.•.•....... Dryinus canariensis (Ceba I los) -
Antennal
segment
3 more
than
twice as
long as
segment
· •..........•..•..•............ Dryinus nepalensis n. sp.
2 .........
726
Mem.
NOTES: sex of
Amer.
Ent.
Ins!.,
no.
37,
1984
the male associated with D. canariensis could be also the opposite Richardsidryinus albrechti n. sp. On I y breed i ng experiments can
answer the question.
Dryinus sanderi n. sp., Nichardsidryinus maroccanus n. R. albrechti n. sp. (for the last species, however, see note
The males of sp.
and
above)
are unknown.
The fossil
Dryinus antiquus (Ponomarenko) was not included
species
into the key because the
type
(only known specimen)
is in
too bad
conditions for distinguishing differences with other males of Palaearctic
Dryininae.
KEY TO THE MALES OF THE KNOWN ETH I OP IAN
Notaulices
invisible;
scutum strongly
DRYININAE
reticulate
rugose,
with
numerous areolae as wide as
· . . . . . • . . . . . . . j\1esodryinus -
ocelly . . . . . . . . . . . . . . . . . . . . .. ethiopicus n. sp.
Notaulices visible; scutum granulated, not reticulate rugose; sometimes more or less weakly reticulate rugose, but in this case areolae much
less wide
than
ocel Ii •••••••••.•••••••••••••• 2
2 Occipital carina complete; dorsal part of occipital carina not touching eyes ••••••.••••••••••••••••••••••••••••.•••.••••.••••• 3 Occipital carina incomplete; dorsal part of occipital carina usually touching eyes ••••••••••••••••.•••••••.••••.•••••••••••• 4 3 Ninth abdominal sternite with apex rounded (Fig. 623} . . . . . . . . . . · . . . . . . . . . . . . . Richardsidryinus erraticus (Tu rn e r) - Ninth abdominal sterni te wi th apex emarginated . . . . . . . . . . . . .
· .....•....•.. Dryinus pretorianus n. 4 Head wi th OL
sp.
= OOl . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5
- Head wi th Ol more or less shorter than OOl ••••••. '" .•••••••••• 6 5 Fore-wing with radial cell closed; head with posterior ocelli not
touching
occipi tal
carina ••••••••••••••••••••••••••••••
••........•.• . Dryinus mahensis (Ki effer) -
Fore-wing with touching
radial
occipi tal
cell
open;
head with posterior ocelli
carina • . . • • • . • • • • . . • • •
· ...•....•••.. Dryinus saussurei (C eba I I os) 6 Fore-wing
wi th
radial
cell
open . . . . . . . . . .
. . . . . . • . . . . . . . Dryinus orophilus (Benoi t) -
Fore-wing with
radial
7 Frons with a median
cell
closed . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7
longitudinal
hump
from anterior ocellus
to
antennal
sockets ••••••••••••••.••••••••••••. . . . . . . . . . . . . . . Dryinus deceptor (Turner)
-
Frons
flat
in
the antennal 8 Breadth
of
front sockets
of
the anterior ocellus,
only
convex between
•..•.••••.••••.••••••••••••••••••••.••... 8
anterior ocellus
longer
than POL . . . . . . . . . . . . . . . . . . ..
.•....•••....• Lestodryinus luweli Benoi t
(see
incertae sedis nomi na)
Dryininae
- Breadth of
anterior ocellus
727
shorter
than POL . . . . . . . . . . . . . . . . . . ..
.......•• . Dryinus afer (Olmi)
KEY TO THE MALES OF THE KNOWN OR lENT AL DRYININAE
Dorsal
part
of
occipi tal
carina
touching
eyes •••••••••••••••••••
. . . . . . . . . . Dryinus indicus (Kieffer) -
Dorsal
part
of
2 Head wi th TL
occipital
shorter
carina not
than breadth
touching
of
eyes . . . . . . . . . . . . . . 2
ocel Ii •••••••••••••••••••••
· .•...•... Dryinus browni Ashmead -
Head with TL as
long as breadth
of
ocel Ii •••••••••••••••••••••••
· . . . . . . . . . Richardsidryinus pyrillae (K i ef fer)
KEY TO THE MALES OF THE KNOWN ~JEARCT I C DRYININAE
Head with breadth
of
ocelli
more
· •........ Tridryinus testaceus n. 2 Head wi th
ocell i
than
twice as
long as OPL . . . . . .
sp.
sl ight Iy broader
than OPL . . . . . . . . . . . . . . . . . . . . ..
· .......•. Perodryinus amoenus R. C • L. Perk ins
The male of Dryinus alatus check again
the
only known
(Cresson)
is known:
I,
however,
must
specimen.
KEY TO THE MALES OF THE KNOWN NEOTROP IC DRYININAE
Notaulices -
invisible •••••••••••••••••.••••••••••••••••••••••••• 2
Notaulices at
least
2 Posterior ocelli
partly
touching
visible . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3 occipital
carina . . . . . . . . . . . . . . . . . . . . . .
· . . . . . . . . . Tridryinus ruficauda R i cha rd s Posterior ocell i
not
touching
occipital
carina . . . . . . . . . . . . . . . . . .
· . . . . . . . . . Tridryinus poecilopterae Richards 3
Scutum fully granulated,
not
reticulate
rugose •.•••••.••••••••••
· .......•. Dryinus antilleanus (Evan s ) -
Scutum fully
reticulate
4 Head with OPL
less
than
rugose . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4 0,5 as
long as breadth
of
ocelli . . . . . . ..
· . . . . . . . . . Dryinus striatus (F en t on) -
Head
wi th OPL
5 Distal
part
of
sl ight Iy radial
shorter vein
than breadth
longer
than
· ....•.... Tridryinus striaticeps (K i ef fer)
of
proximal
ocell i . . . . . . . . . . 5 part . . . . . . . . . . . .
728
Mem. Amer. Ent.
Distal
part
of
radial
Ins!.,
no. 37, 1984
vein shorter than proximal
part ••••••.••••
. • • . . . • . . • • . . . . • . • • • • • . • • • • . • Dryinus surinamensis n. sp.
KEY TO THE MALES OF THE KNOWN AUSTRALIAN
DRYININAE
Notaul ices complete .•...•.... Dryinus areolatus n. sp. - Not a u I ice s i n c omp let e •••••••••••••••••••••••••.•••••.•••.••.•.• 2 2 Head granulated, not reticulate rugose ••••.••..•••••.••.••.•.••. •..•.•...•..•......••........ Dryinus pallidus (R.C.L. Perkins) - Head more or less reticulate rugose •••••••.••••••.•.•.••••..•• 3 3 Dorsal surface of propodeum approximately as long as metanotum .• .•.....••....•.••..•.•..•.•. . Dryinus aterrimus (Dodd) - Dorsal surface of propodeum longer than metanotum .•.••••••...•. 4 4 Scutellum granulated and punctate, not ret iculate rugose •.•••.•• .......•.....•.•..•.• .•••.•. . Dryinus australianus n. sp. - Scutellum granulated and reticulate rugose . . . . . . . . . . . . . . . . . . . . . 5 5 Scutum reticulate rugose, very rugose; areolae wide . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Dryinus koebelei (R.C.L. Perkins) - Scuum reticulate rugose, less rugose; areolae less wide ••••••••• . . .• .. • . . . . • , ...•..•..•..•... Dryinus venator (R.C.L. Perk i ns)
1. Dryinus Latrei lie
Dryinus Latrei lie 1804: 176. nee Dryinus Fabricius 1805: 200. Dryinus Latreille 1805: 228. Campylonyx Westwood 1835: 52 (n. syn.); type species Campylonyx ampuliciformis Westwood 1835, monotypic. Paradryinus R.C.L. Perkins 1905: 53 (n. syn.); type species Paradryinus koebelei R.C.L. Perkins 1905, orig. desig. Chlorodryinus R.C.L. Perkins 1905: 57 (n. syn.); type species Chlorodryinus pallidus R.C.L. Perkins 1905, orig. desig. Dryinus Latreille partim: Kieffer in Kieffer et Marshall 1905: 69 Campylonyx Westwood: Kieffer in Kieffer et Marshall 1905: 84. Plastodryinus Kieffer in Kieffer et Marshall 1906: 496 (syn. proposed by Moczar 1965); type species Plastodryinus szepligetii (Kieffer 1905), orig. desig.
Dryinus Latreille partim: Kieffer in Kieffer et Marshall 1906: 497. Hesperodryinus R.C.L. Perkins 1907: 40 (n. syn.); type species Hesperodryinus arizonieus R.C.L. Perkins 1907, orig. desig. Lestodryinus Kieffer 1911a: 108, new name for Dryinus Latreille (syn. proposed by Richards 1937); lype species Lestodryinus formiearius (Latreille 1804)(first species included in Dryinus
729
Dryininae
Latreille).
Lestodryinus Kieffer: Kieffer 1914b: 20. Campylonyx Westwood: Kieffer 1914b: 43. Dryinus Latrei lie: Richards 1935: 144. Dryinus Latreille: Richards 1937: 118. Dryinus Latrei lie: Richards 1939: 229. Mesodryinus Kieffer: Muesebeck and Walkley 1951: 1034. Hesperodryinus R.C.L. Perkins: Muesebeck and Walkley 1951: 1034. Hesperodryinus R. C. L. Perk i ns: Krombe i n 1979: 1242. Mesodryinus Kieffer: Krombei n 1979: 1243.
Dryinus iormicarius Latreille 1804, first included species
TYPE SPECIES: (Latreille 1805) FEMALE
(Figs 495,
segments;
labial
528,
palpi
529):
as or shorter than front subapical visible;
tooth
(rarely
pronotal
fully
tibia, two)
much
(Figs 531,
labial
palpi
532):
fully
tibial
winged;
with 6
(Fig.
longer than arol ium,
and some lamellae;
with 3 segments;
DISTRIBUTION:
maxi Ilary palpi
enlarged claw
tubercles reaching or not
complete or incomplete or absent; MALE
winged;
with 3 segments;
485)
notaulices at
tegulae;
as long
with one
occipital
least partly carina
spurs 1,1,2.
maxi Ilary palpi
with 6 segments;
for morphology see fami Iy characteristics.
worldwide
Dictyopharidae, Cixiidae, Flatidae, Acanaloniidae, Issidae, Ricaniidae, Delphacidae, Fulgoridae. HOSTS:
SPECIES: NOTES:
87.
on a different I
(Chlorodryinus for instance) were based
some synonym ized genera
have seen,
variable with
length of occipital however,
that the
carina
(complete or incomplete).
length of occipital
intermediate cases from occipital
Generic differences so are not proposable.
carina can be very
carina absent to complete.
The shape of posterior margin of
head also is variable, with intermediate cases from occiput concave to straight or convex (Hesperodryinus). In addition the posterior ocelli can be located on an front or behind,
imaginary
I ine connecting
with numerous
As the type of D.
hind margins of eyes or in
intermediate cases.
iormicarius Latreille seems lost and the original
description is unreliable, the diagnosis of Dryinus is based on classical opinions of Kieffer (1914b) and Richards (1939).
PALAEARCTIC SPECIES OF DRYINUS
Seven species are known.
KEY TO THE SPECIES
the
730
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
FEMALES
Occipital
margin
incomplete,
only dorsally visible
............................ -
Occipital
margin
complete
6.
(Fig.
2 Pronotum with weak posterior short
posterior collar
propodeum nearly as -
a
long as dorsal
long posterior collar
transversal
shorter
impression and wi th a
posterior surface of
surface;
fore-wing
hyal ine or
(Fig.
484 A,B,C);
than dorsal
posterior surface of
surface;
fore-wing
wi thout
dark
transversal
bands;
head and
sanderi n. sp.
prothorax mostly black . . . . . . 1. Fore-wing wi th dark transversal
bands;
redd ish •••••••••••••••••••••
canariensis (Ceba I los)
4 Transversal 488 A); furrow
(Fig.
488 B);
2.
head and prothorax
fully
furrow between metanotum and propodeum narrow
metanotum at
Transversal
wi th dark
bands ••••••••••.•••...••••.•••..••••••.•••.•••••••• 4
3 Fore-wing hyal ine,
-
4840);
wi th dark transversal bands •••••••••••••••••••••••••••••••••••• 3 Pronotum with a strong posterior transversal impression and with propodeum much
-
499 A) •
492) . . . . . . . . . . . . . . . . . . . . . . . . . . . 2
transversal
(Fig.
(Fig.
koreanus (Moczar)
least
five
488 A) . . . . . . . . .
3.
times as
long as breadth
this
collan's (Linnaeus)
furrow between metanotum and propodeum broad
metanotum nearly
(Fig. of
three-four
times as
(Fig.
long as breadth
of
thi s furrow (Fig. 488 B) •••.••••••••••••••••••••••••••••••••••• 5 5 Temples more prominent (Fig. 492 A); TL nearly twice as long as OL -
(Fig.
Temples (Fig.
492 A) . . . . . . . . . . . . . less prominent
(Fig.
492 B) ••••••••••••••••
The female of the fossil
4.
nepalensis n.
492 B); 5.
TL
sp.
slightly
longer
than OL
tarraconensis Marshall
species D.
antiquus (Ponomarenko) is
unknown.
MALES (excluding the fossi I Dryinus antiquus Ponomarenko)
Fore-wing with distal part of radial vein nearly as long as proximal part (Fig. 490 B) •• 5. tarraconensis Marshall -
Fore-wing proximal
with distal part
2 Notaulices -
Notaulices
(Fig.
part
of
radial
vein much
reaching approximately
0,5
reaching
0,75
3 Oistivolsella
longer
than
490 A) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2 approximately
farther
from apex
length
of
scutum . . . . . . . . . 3
length of
of penis
(Fig.
scutum . . . . . . . . 4
491) . . . . . . . . . . . ..
3. collaris (Linnaeus) -
Oistivolsella nearer apex
of penis
(Figs.
487,
494) . . . . . . . . . . . ..
Dryininae
731
. . . . . . . . . . . . . . . . . . . . . . . . . . . . 6. koreanus (Moczar) segment 3 nearly twice as long as segment 2 • . . . . . . . • . . . . • . . . . . . • . . . . . . . •. . . . . • . . . .. 2. canariensis (Ceba I los) - Antennal segment 3 more than twice as long as segment 2 . . . . . . . . . 4. nepalensis n. sp.
4 Antennal
NOTES:
the male of D.
The
fossil
sanderi n. sp. is unknown. antiquus (Ponomarenko) isn't present in the
species D.
key because the only known specimen
is in
too bad conditions for
distinguishing differences with I iving species. The male associated in the key with D. canariensis could be also the opposite sex of Richardsidryinus albrechti n.
sp.
Only breeding
experiments could answer the question.
1. Dryinus sanderi n. sp.
FEMALE: fully winged; length 3,75 mm; head black, with mandibles, clypeus and malar space testaceous; antennae fully testaceous; thorax and propodeum black,
with posterior collar of pronotum reddish-testaceous;
A
B
c Fig. sp.
(laterally viewed) of females of Dryinus nepalensis n. collaris (Linneaus) from France (B), tarraconensis Marshall from Montecatini (Italy)(C) and sanderi n. sp. (type)(D).
484 -
Pronotum
(type) (A),
732
Mem.
Fig.
485 -
abdomen black-brown;
Amer.
Chela of
Ent.
no.
37,
legs testaceous,
with coxae partly black or brown
antennae distally
in following proportions:
8:
6:
23:
14:
punctate,
frons,
where there are numerous irregular striae;
complete;
thickened;
10:
shiny,
incomplete keel,
1984
lJryinus sanderi n. sp. (type)
and with clubs of femora brown; segments
Inst.,
8:
6:
antennal
5:
4,5:
8;
head
weakly granulated and rugose mainly on anterior half of frons with a
not visible on anterior half of frons;
POL = 4;
Ol = 3;
OOl = 8;
OPl = 1,5;
median
occipital
Tl = 3;
carina
pronotum
(Fig.
484 0) with a very weak anterior transversal impression and with a strong posterior transversal impression; disc humped; posterior collar very short; pronotal
tubercles not reaching
granulated; numerous,
scutum shiny,
short,
posterior margin; of scutum; smooth;
weak,
very
tegulae;
pronotum shiny,
weakly
weakly granulated and rugose,
longitudinal
with
keels near anterior margin and near
notaulices incomplete,
reaching
scutellum and metanotum shiny,
propodeum reticulate rugose,
very
approximately 0,65
length
weakly granulated,
with dorsal
surface nearly as long as
posterior surface; posterior surface without longitudinal keels; fore-wing hyaline, without dark transversal bands; distal part of radial vein longer than proximal 17: with
3:
6:
14:
part 24;
1 row of 7
of 19 lamellae; 1,1,2. MALE:
unknown
(14:7);
fore tarsal
enlarged claw lamellae;
(Fig.
segments 485)
segment 5 of front
apex with a
in following proportions:
with a
group of nearly
tarsus
subapical (Fig.
12 lamellae;
tooth and
485) tibial
with 2 rows spurs
Dryininae
LOCUS TYPICUS: Melnik (SW Bulgaria) TYPICAL MATERIAL: holotype F! in OL. HOSTS: in Bu I gari a Hysteropterum latifrons DISTRIBUTION: Palaearctic region: BULGARIA:
Fig. 486 -
Chela of
733
Melnik, OL!
Dryinus canariensis n. sp. from Tahodio (Canary Islands)
734
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
2. Dryinus canariensis (Ceballos) n. comb.
Paradryinus canariensis Ceba Ilos 1927: 101.
FEMALE:
fully
legs reddish;
winged;
length 4,25-4,50 mm;
mesothorax,
metathorax,
head,
antennae,
prothorax,
propodeum and petiole black;
abdomen brown, with segment 1 reddish; tegulae testaceous; antennae not distally thickened; antenna I segments in following proportions: 13: 5: 20: 9:
9:
6:
6:
6:
5:
8;
head dull,
smooth,
granulated;
frons with a median
longitudinal keel, without lateral keels; occipital carina complete; POL = 4; OL = 3; posterior ocelli not touching occipital carina; OPL nearly as long as breadth of ocell i; granulated, impression
partly and
collar short,
temples prominent;
without sculpture,
with
a
strong
nearly 0,5 as
pronotum shiny,
with an
posterior transversal
long as pronotal
partly
anterior weak transversal impression;
tubercles;
posterior
pronotal
tubercles
not reaching tegulae; scutum dull, hairy, granulated and with numerous weak areolae; notaul ices incomplete, reaching approximately 0,65 length of scutum; weakly
scute II um and metanotum sh iny, granulated;
strongly transversely striate, dark transversal
smooth,
wi thout scu I pture or very
propodeum reticulate rugose,
bands,
without
two on
with posterior surface
longitudinal
the basal
keels;
pterostigma;
distal
part of radial
(13:7);
tarsal
segments in following proportions:
fore
enlarged claw
(Fig.
segment 5 of front same length; MALE
testaceous;
(Fig.
incomplete; carina;
legs brown,
with
=
granulated;
proximal
part
broadened;
winged;
tibial
5;
OL
=
3;
OOL
with ventral
light;
=
3;
scutum dull,
(12:7);
granulated;
part of radial
gonoforceps (Fig.
ninth abdominal
sternite:
black;
side of segment 10: 8:
487 A)
fig.
8:
occipital
posterior ocelli
length of scutum;
distal
part
15: 24;
antennae not distally thickened;
propodeum reticulate rugose; bands;
7:
spurs 1,1,2.
length 2,50 mm;
frons with a median keel;
temples invisible;
dark transversal
tarsi
15: 4:
tooth and with 12 lamellae;
in following proportions: 8,5: 4:
granulated;
POL
fully
with 3
with 2 rows of nearly 27 lamellae
antennae black,
reaching approximately 0,75 dull,
486)
longer tha proximal
apex with a group of 12 lamellae;
antenna I segments head dull,
vein much
with a subapical
(see note on page 731):
mandibles testaceous;
9;
486)
tarsus
fore-wing
cells and one beneath the
8:
8:
6:
6:
carina
touching occipital
notaulices incomplete,
scutellum and metanotum fore-wing
hyaline,
vein much
with distal
487 B;
tibial
without
longer than region
spurs 1,1,2.
LOCUS TYPICUS: La Cuesta (Canary Islands) TYPICAL MATERIAL: holotype F! in MD. HOSTS: unknown DISTRIBUTION: Palaearctic region: OL!
TN!
B.co de Marchar
(Tenerife) (F), Tenerife) (F),
MO! TN!
CANARY
(San Sebastian,
Tahodio
(Tenerife) (F),
ISLANDS:
B.co Santos
Gomera)(F), MD! MO!
(F),
MD!
La Cuesta
Los Calderones
(Medano,
Dryininae
735
Fig. 487 - Male genital ia of Dryinus canariensis n. sp. from Los Calderones (Canary Islands) (A: left half removed; B: ninth abdominal sternite) .
3. Dryinus collaris (Linnaeus)
Sphex collaris Linnaeus 1767: 946. Dryinus formicarius Latreille 1805: 228 (syn. proposed by Fitton, Graham,
Boucek, Fergusson,
Huddleston, Quinlan, Richards 1978);
locus typicus Southern France. Campylonyx ampuliciformis Westwood 1835: 52 (n. syn.); locus typicus Lyon (France)
Dryinus formicarius Latrei lie var. : Kieffer in Kieffer et Marshall 1905: 78.
Dryinus formicarius Latreille var. : Kieffer in Kieffer et Marshall 1905: 79.
Lestodryinus formicarius (Latreille): Kieffer 1914b: 20. Lestodryinus corsicae Kieffer 1914b: 21 (n. syn.); locus typicus Corse (France).
Lestodryinus formicarius (Latrei lie): Berland 1928: 140. Lestodryinus corsicae Kieffer: Berland 1928: 141.
736
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
488 - Metanotum of Dryinus collaris (L.) from Le Puy (France )(A) and tarraconensis Marsha II from Orleans (France) (B). In metanotum of tarraconensis the anterior half is rugose and inclined. 5 ; transversal
Fig.
furrow between metanotum and propodeum.
Campylonyx ampuliciformis Westwood: Berl and 1928: 142. Lestodryinus formicarius (Latreille): Picard 1932: 29. Lestodryinus formicarius (Latrei lie): Maneval 1935: 4. Dryinus formicarius Latrei lie: Richards 1937: 118. Dryinus formicarius Latrei lie: Richards 1939: 229. Dryinus formicarius Latrei lie: Berland 1940: 121. Dryinus formicarius Latrei lie: Abdul-Nour 1974: 1015. Dryinus formicarius Latreille: J.I'. Perkins 1976: 27. Dryinus formicarius Latrei lie: Olmi 1976: figs 10-14. Dryinus collaris (L i nnaeus): F i !ton, Graham, Boucek, Fergusson, Huddleston,
Quinlan,
Richards 1978:
127.
Dryinus formicarius Latreille: Currado and Olmi 1979b: 4. Dryinus collaris (Linnaeus): Day 1979: 60.
FEMALE: fully winged; length 5,12-6,25 mm; head reddish-testaceous, with a brown small or wide spot on frons and vertex; antennae testaceous; propectus, pronotum, scutum, scutellum, metanotum, mesopleura, mesosternum, metasternum and metapleura reddish-testaceous; sometimes brown
spots on mesosternum,
metasternum,
between mesopleura and metapleura;
mesop leura and on furrow
rarely pronotum mostly black;
scutell um and metanotum someti mes reddi sh-dark;
mesosternum,
metasternum, mesopleura and metapleura sometimes almost fully brownblack; rarely scutum, scutellum and metanotum fully black; propodeum black,
with apex reddish;
fully reddish-testaceous, head dull,
granulated;
petiole black;
ocellar region swollen;
OPL shorter than breadth of ocelli; B)
crossed by
abdomen brown-reddish;
legs
sometimes with mid and hind coxae brownish;
two transversal
collar usually smooth and
impressions,
shiny;
without sculpture;
impression with short longitudinal
occipital carina complete;
temples prominent;
keels;
pronotum
(Fig.
484
disc and posterior
posterior transversal
anterior collar and sides weakly
Dryininae
Fig.
489 -
granulated; reaching dull;
Chela of
sides with a
tegulae;
Dryinus collaris (L.) from Dornbach (Austria)
few
longitudinal
scutum shiny,
notaul ices distinct and
of scutum;
737
keels;
pronotal
tubercles not
weakly punctate and granulated,
incomplete,
reaching
posterior third of scutum with
weak
rarely
approximately 0,5
longitudinal
length
short keels;
scutellum usually flat, shiny, smooth, weakly punctate; metanotum usually weakly punctate, smooth, always without transversal furrow in posterior half (Fig.
488 A);
propodeum with transversal proximal
tarsus
segment 1 of front with subapical (Fig.
489)
reticulate rugose;
longitudinal keels;
bands and with distal
part;
enlarged claw of front
propodeum fully
two short
part of radial tarsus slightly
tooth and
11
vein much
longer than
longer than segment 4;
lamellae
with 2 rows of 27
posterior surface of
fore-wing with 2 brown
(Fig.
lamellae;
489);
segment 5
apex with a
group
Mem.
738
Amer.
Ent.
Inst.,
no.
37,
1984
A
B Fig.
490 -
Radial
(Italy)(A)
of 19 lamellae; MALE:
fully
and
veins of males of Dryinus collan"s (L.) from Rosta tarraconensis Marshall from Orleans (France)(B).
tibial
spurs 1,1,2.
winged;
length 5,00 mm;
black,
with
tegulae and
legs
testaceous; mandibles with 3 teeth; scutum dull, hairy, granulated; notaulices incomplete, reaching approximately 0,5 length of scutum; scutellum and metanotum shiny,
smooth,
weakly punctate;
propodeum fully
reticulate rugose, without longitudinal keels on posterior surface; forewing with distal part of radial vein much longer than proximal part (20:10)(Fig.
490 A),
without dark transversal
the apex of penis (Fig. LOCUS TYPICUS:
491);
tibial
bands;
distivolsella far from
spurs 1,1,2.
Spain
holotype F of S. collaris in Ll; holotype F of D. formicarius probably lost; lectotype F! (hereby designated) and 1 para lectotype F! of C. ampuliciformis in P; lectotype F! (hereby designated) of L. corsicae in MT. HOSTS: in France Cixius nervosus (L.)(Maneval 1935); in England Cixius contaminatus Fl. (Capron 1885); in Italy Issus coleoptratus (F.) (Olmi 1976); in Germany Cixius nervosus L. (Haupt 1932) (quotation of Dictyophara europaea L. in Richards 1939 valid for D. tarraconensis Marshall); in addition Tachycixius pilosus (OI.)(Thompson 1955). TYPICAL MATERIAL:
DISTRIBUTION:
Palaearctic region:
U.
S.
S.
R.
? (Ponomarenko 1978);
Dryininae
Fig.
491
HUNGARY:
-
Male genitalia of
Zelenika,
BU!
739
Dryinus collaris (L.) from Rosta (Italy)
YUGOSLAVIA:
Opatija
(Istra), BU!
Krapina
(Hrvatska), BU! GERMANY: Aix-Ia-Chapelle (Kieffer in Kieffer et Marshall 1905); AUSTRIA: Werfen (Slzb.), LE! Dornbach (Wien), P! BELGIUM (Kieffer in Kieffer et Marshall 1905); SWITZERLAND: Peney, GV! Geneve, SP! FRANCE: Mont-de-Marsan (Abdul-Nour 1974); Lyon (Rhone), P! St. Gely-duFesc (Herault), P! St. Guilhem (Herault), P! Digne (Basses-Alpes), P! Hyeres (Var), P! Le Puy, P! Toulon (Var), P! Les Guerraux (Saone-etLoire) (Berland 1928); Corse, MT! ENGLAND: Cobham (Surrey) (Richards 1939); Shere (Surrey)(Richards 1939; Capron 1885); SPAIN (Linnaeus 1767; Day 1979); ITALY: Rosta (Torino), OL! Gaibola (Bologna), OL! S. Olcese (Genova), BM! Lucca (Kieffer in Kieffer et Marshall 1905); Napoli (Kieffer
740
Mem.
Amer.
Ent.
Ins!.,
no.
37,
1984
A o Fig.
492 -
B
o
Heads
0
o
of females of Dryinus nepalensis n. sp. tarraconensis Marshall (lectotype)(B)
and
in Kieffer et Marshall the cocoon
1905); S.
is oval
elongate. According by
0
(dorsally viewed)
(type)(A)
NOTES:
o
(Fig.
to Berland
Basilio
(Mottola,
498 A),
whereas
(1940),
Taranto)(Moczar 1965).
D. tarraconensis it's
in
occasionally the female
is attracted
light traps.
4. Dryinus nepalensis n. sp.
FEMALE: fully winged; length 6,25-6,55 mm; head black, wit>, ",andibles and anterior margi n of c I ypeus testaceous; antennae brown, wi th segments 1-2 testaceous and segments 7-10 whitish; black;
legs reddish,
distally 26:
thickened;
18:
broad
13:
11:
(26:5),
complete;
10:
8:
12;
POL
=
2,5;
Ol
around disc;
pronotum
collar
long,
flat,
distinct;
=
granulated and with numerous
median complete keel;
2,5;
OOl
=
11;
OPl
=
484 A)
with a
pronotal
keels;
metanotum nearly keels;
fore-wing
Tl
=
3;
6:
50:
long as
longitudinal carina temples:
fig.
impression; tegulae;
notaulices
scutellum dull,
posterior scutum dull,
incomplete, granulated;
wi th a strong transversa I furrow near scutell um and wi th
furrow near propodeum; transversal
occipital
tubercles not reaching
transversa I furrow near propodeum;
granulated;
15:
weak anterior transversal
almost reaching posterior margin of scutum; a strong
2;
strong posterior transversal
granulated and with some areolae near margins; metanotum du II,
antennae
in following proportions:
weakly granulated and with numerous striae (Fig.
impression and with a
propodeum and abdomen
antenna I segment 4 nearly five times as
frons with a
pronotum shiny,
thorax,
tibiae and part of coxae brown;
antenna I segments
head dull,
keels on frons; 492 A;
with
propodeum dull,
posterior surface with
with
and another beneath
centra I reg ion short and
three-four times as two short
two dark transversal the pterostigma;
long as breadth of
reticulate rrgose,
distal
bands,
without
incomplete longitudinal one on the basal
part of radial
vein
cells
longer
Dryininae
Fig.
than proximal
Chel a of Dryinus nepalensis n.
493 -
part
741
(19:10);
fore tarsal
sp.
(type)
segments in following proportions:
16: 3: 6: 15: 24; segments 2 and 3 of front tarsus produced into hooks; enlarged claw (Fig. 493) with a subapical tooth and with a row of 15 lamellae;
segment 5 of front
lamellae;
apex with a group of nearly 18 lamellae;
MALE:
fully
winged;
(partly black
in
tarsus
(Fig.
length 2,00-3,56 mm;
long specimens);
segments 1-2 testaceous; partly black or brown;
carina
(not touching occipital invisible;
antennae fully
testacoeus or brown,
legs testaceous, 7:
6:
12:
incomplete;
10:
with all
10:
11:
10: 9: 7:
in very
9;
segments
head dull,
touching occipital
long specimens);
scutum dull,
reaching approximately 0,75
sometimes with a strong
antennal
frons with a median complete
scutellum and metanotum as sculptured as scutum; rugose,
with
coxae or only hind coxae
posterior ocelli
carina only
spurs 1,1,2.
mandibles testaceous
POL = 5; Ol = 2; OOl = 4;
notaul ices incomplete,
tibial
black;
granulated and weakly reticulate rugose; occipital
with 2 rows of nearly 37
antennae not distally thickened;
in following proportions: keel;
493)
transversal
keel
carina
temples
as sculptured as head; length of scutum; propodeum reticulate
between dorsal
and
742
Mem.
Fig.
494 -
Amer.
Male genital ia of
Ent.
Ins!.,
no.
37,
1984
Dryinus nepaiensis n. sp. from Godavari
(Nepal) (left half removed)
posterior surface;
fore-wing
hyal ine,
without dark transversal
bands;
distal part of radial vein longer than proximal part (13:9 or 9:5 or 14:9); gonoforceps (Fig. 494)with distivolsella located near apex of penis; tibial spurs 1,1,2. lOCUS TYPICUS:
lothar
TYPICAL MATERIAL: (1
F,
2 MM)!
HOSTS:
(Nepal)
holotype F!
and 5 paratypes MM!
in OT,
3 paratypes
in Ol.
unknown
DISTRIBUTION: Godavari
Palaearctic region:
(Katmandu), OT!
Ol!
r,EPAl:
Pulchauki
lothar (Birganj), (Katmandu),
OT!
OLi
OTt
5. Dryinus tarraconensis Marshall
Dryinus tarraconensis Marshall 1868: 204. Dryinus szepligetii Kieffer in Kieffer et Marshall 1905: 77 (n. syn.);
Dryininae
Fig.
495 -
Female of Dryinus tarraconensis Marshall from Montecatini ( Italy)
743
744
Mem.
Fig. 496 -
Amer.
Ent.
Inst., no. 37,
1984
Chela of Dryinus tarraeonensis Marshall from Orleans (France)
locus typieus Novi (Yugoslavia) (nee Budapest: see Moczar 1965). Dryinus tarraeonensis Marshall: Kieffer et Marshall 1905: 82. Plastodryinus szepligetii (Kieffer): Kieffer in Kieffer et Marshall 1906: 496.
Dryinus tarraeonensis Marsha II: Kieffer in Kieffer et Marsha II 1906: 497.
Lestodryinus tarraconensis (Marshall): Kieffer 1914b: Plastodryinus szepligetii (Kieffer): Kieffer 1914b: 43. Lestodryinus tarraconensis (Marshall): Berl and 1928: Lestodryinus gregori Hoffer 1936: 164 (syn. proposed 1965); locus typicus Pouzdranyi (Czechoslovakia). Lestodryinus bidens Haupt 1937: 70 (n. syn.); locus Bellinchen
21. 141. by Moczar
typicus
(Germany).
Lestodryinus tarraconensis Marshall: Pi Ilault 1951: 67. Dryinus szepligetii Kieffer: Moczar 1965: 380. Dryinus tarraconensis Marshall: Moczar 1965: 381. Dryinus tarraconensis Marshall: Currado and 01 mi 1979b: 4. nee Dryinus szepligetii Kieffer: Ponomarenko 1981 b: 879 (see Dryinus stantoni Ashmead and Dryinus browni Ashmead).
FEMALE (Fig. 495): fully winged; length 5,12-8,25 mm; head usually black, with frons, part of vertex (more along orbits), clypeus and mandibles yellow-reddish; sometimes also partly temples reddish; occiput
Dryininae
Fi g.
745
Ma I e gen i ta I i a of Dryinus tarraconensis Marsha II from Orl eans
497 -
(France)
always reddish; and vertex; testaceous;
rarely head reddish-testaceous with brown spot on frons
antennae brown,
with segments 1-2 and sometimes also 3
segments 9-10 or only 10 yellow;
sometimes reddish-testaceous; and sides reddish;
with posterior collar
sometimes fully reddish-testaceous;
and metanotum usually black, mesosternum,
propectus usually black,
pronotum usually black, sometimes fully
scutum,
mesopleura and metapleura usually black,
less reddish-testaceous or fully reddish;
scutellum
reddish-testaceous; sometimes more or
propodeum usually black,
sometimes with apex reddish, rarely fully reddish; petiole black; abdomen brown-reddish or black-brown; legs usually testaceous, with coxae partly brown,
sometimes also with clubs of femora brown,
testaceous; longitudinal
head dull, keels;
granulated;
vertex swollen;
rarely fully reddish-
frons and vertex with tracks of POL
= 4; Ol
= 3;
OOl = 13;
OPl = 2,5;
746
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
A
C _______B-~ Fig.
498 -
Italy)(A)
Dryinus collaris (L.) (female from Rosta, tarraconensis Marshall (female from Orleans, France)(B)
Shapes of cocoons of and
TL = 7; occipital carina complete; OPL nearly as long as breadth of ocell i; temp I es: fi g. 492 B; pronotum (F ig. 484 C) crossed by 2 transversal impressions, shiny, granulated, with strong longitudinal keels on sides; posterior collar sometimes smooth and without sculpture; anterior impression sometimes weakly tegulae;
scutum shiny,
longitudinal
pronotal hairy,
keels on posterior third;
approximately 0,5 strong
visible;
granulated,
length of scutum;
longitudinal
metanotum with a propodeum fully
keels;
deep
rarely
with numerous short
notaulices incomplete, scutellum dull,
almost smooth,
transversal
reticulate rugose;
tubercles not reaching
rugose,
reaching
with numerous and
with weak keels;
furrow on posterior half (Fig.
488 B);
posterior surface with 2 short
longitudinal keels; fore-wing with 2 brown transversal bands and with distal part of radial vein much longer than proximal part; segment 1 of front
tarsus sl ightly
longer than segment 4;
tooth and 13 lamellae (Fig. 2 rows of 38
lamellae;
496);
enlarged claw with subapical
segment 5 of front
apex with a group of at
tarsus
least 15
(Fig.
lamellae;
496)
spurs 1,1,2. MALE:
fully
winged;
length 3,37-3,87 mm;
black,
with
tibial
with mandibles yellow
Dryininae
and tarsi carina
light;
fore tibiae yellow;
incomplete;
almost complete;
scutellum and metanotum shiny,
vein
dark transversal
granulated,
scutum dull,
occipital
hairy,
granulated;
reaching posterior third or fourth of scutum,
fully reticulate rugose, part of radial
head dull,
temples not visible;
notaul ices incomplete,
747
without
longitudinal
keels;
nearly as long as proximal
bands;
distivolsella
granulated; fore-wing
part
(Fig.
rarely
propodeum with distal
490 B),
located near apex of penis
without
(Fig.
497); tibial spurs 1,1,2. lOCUS TYPICUS, Torla (Spain) TYPICAL MATERIAL; holotype F! of D. tarraconensis in BU; holotype F! of L. gregori in Hoffer coli.; holotype F! of D. szepligetii in BU; holotype F! and allotype M! of L. bidens in HA! in VI there is the type M! of Labeo rufipes Forster (never described) without local ity label: it's a male of D. tarraconensis. HOSTS; Dictyophara europaea (l.) in France (Pi Ilault 1951) and in Germany (Richards 1939; under name Dryinus formicarius latrei lie; Haupt 1937: under name Lestodryinus bidens Haupt). DISTRIBUTION;
Palaearctic region;
U.
S.
S.
? (Ponomarenko 1978);
R.
HUNGARY; Kenese, BU! Crkvenica, Ol! ROMANIA: Nagyenyed (Transilvania)(under name Plastodryinus sziladyi n. sp.), BU! CZECHOSLOVAKIA; BU!
Sutomore
FRANCE;
Orleans
(Madrid), ITALY:
BM!
Sarre
(L.)
BM!
Parma,
BO!
P!
GERMANY; SPAIN;
S. NA!
Perugia,
(Caltanissetta),
Baghdad, NOTE:
GE!
(Hoffer 1936;
BU!
(Tarragona),
(Aosta),
(Pistoia),
Cataldo
(Loiret),
Salou
(Alessandria), Terme
Pouzdranyi
(Dalmacija),
CO!
Torla
OL!
Moczar 1965);
Bell inchen
BM!
(Huesca),
BU!
Benicasim (Cuneo),
Pietra Ligure
(Savona),
Maiella
GREECE;
(Abruzzo),
Kremasti
Hills
P!
Escorial
PL!
BM!
Gavi
PL!
NA!
Novi,
HA!
(Castellon),
Benedetto Belbo GE!
YUGOSLAVIA;
(Oder),
Montecatini
Messina,
(Rhodes),
BM!
CO!
WA!
the cocoon it's ovale.
(Fig.
498 B)
is very elongate,
Notes on biology by Pillault
whereas in
D. collaris
(1951).
6. Dryinus korean us (Moczar) n. comb.
Chlorodryinus koreanus Moczar 1983a; 192.
FEMALE:
fully
winged;
length 3,84 mm;
testaceous-light,
with petiole
black; antennae distally thickened; antenna I segments in following proportions: 10: 6; 33: 18; 15; 11: 9; 7; 7; 10; head (Fig. 499 A) swollen,
dull,
granulated;
frons with a
weak median complete keel;
occipital carina incomplete, visible only behind posterior ocelli and on sides of posterior ocelli; POL = 3; OL = 4; OOL = 8; OPL = 3; temples prominent;
pronotum shiny,
striae on sides,
without sculpture,
crossed by an
with some longitudinal
anterior and a posterior transversal
S.
IRAQ;
748
Mem.
Amer.
Ent.
A
Inst.,
no.
37,
B
o
1984
0
DQll) Fig.
499 -
Heads of Dryinus koreanus
from De-sang san, Wanuma,
impression;
Korea)
(Moczar) (A:
and pallidus
New Guinea;
D:
(R.C.L.
type female;
Perkins) (B:
C:
male
female from
male from Mulgrave, Australia)
anterior collar very short;
disc very
long;
posterior collar
short; pronotal tubercles not reaching tegulae; scutum granulated, with anterior surface of median area smooth and without sculptur0; notaulices incomplete,
reaching approximately 0,65
metanotum dull,
granulated;
transversal or longitudinal
length of scutum;
propodeum reticulate rugose, keels;
dorsal surface sl ightly
scutellum and without longer than
(27:20); fore-wing hyaline, without dark transversal bands; distal part of radial vein longer than proximal part (20:8); fore tarsal segments in fOllowing proportions: 24: 3: 6: 20: 29; enlarged claw (Fig. 500) with a subapical tooth and with a row of 11 lamellae; segment 5 of front tarsus (Fig. 500) with two rows of 17 lamellae; apex with a posterior surface
group of 7 lamellae; tibial spurs 1,1,2. MALE: fully winged; length 2,72 mm; head black,
with mandibles,
anterior region of frons and malar space testaceous; 1 testaceous
(other segments missing
and propodeum black; 499 C)
dull,
carina
incomplete,
in two only known specimens);
abdomen brown;
granulated;
legs yellow-testaceous;
frons with a median complete keel;
fully visible dorsally,
clypeus,
antennae with segment head
thorax (Fig.
occipital
partly visible on sides and
Dryininae
Fig.
500 -
Chela of
Dryinus koreanus (Moczar) (type)
749
750
Fig.
Mem.
501
-
Amer.
Ent.
Inst.,
no.
Male genital ia of Dryinus koreanus (Korea) ( I eft ha I f
37,
1984
(Moczar)
from De-sang san
removed)
occipital carina dorsally touching eyes; POL = 5; Ol = 2; OOl = = 1; scutum dull, granulated; notaulices incomplete, reaching
ventrally; 3;
OPl
approximately 0,5 shiny, without
smooth,
length of scutum;
without sculpture;
transversal
scutellum dull,
propodeum dull,
or longitudinal
keels;
granulated;
metanotum
reticulate rugose,
fore-wing hyal ine,
without dark
transversal bands; distal part of radial vein longer than proximal (16:7); genitalia: fig. 501; tibial spurs 1,1,2. lOCUS TYPICUS:
Pyongyan
TYPICAL MATERIAL: HOSTS:
part
(Korea)
holotype F!,
allotype M!
and 1 paratype M!
in BU.
unknown
DISTRIBUTION:
Palaearctic region:
Pyongan),
BU!
lyong-ak san
Pyongan),
BU!
De-sang san
KOREA:
Pyongyan
(Prov.
South
(14 Km IV from Pyongyan,
Prov.
South
(12 Km NE from Pyongyan,
Prov.
South
Dryininae
Pyongan),
751
BU!
7. Dryinus antiquus (Ponomarenko) n. comb.
Laberius antiquus Ponomarenko 1981 a: 139.
FEMALE: MALE:
unknown
fully
winged;
length 3,20 mm;
black,
with
legs testaceous;
palpi
not visible; antennae not distally thickened, densely hairy; antennal hairs less short than breadth of antennal segments; antennae la-segmented; antennal
segments
in following proportions:
7;
breadth of antenna I segments
4:
4:
3;
mandibles not visible;
by short dense hairs; posterior ocelli occipital
scutum dull,
posteriorly separated;
5:
head dull,
11:
10:
10:
9:
carina;
TL = a
bands;
three basal
cells enclosed by pigmented veins;
covered
(eyes almost touching notaulices complete,
fore-wing hyaline,
transversal
7:
eyes apparently bare;
metanotum and propodeum weakly
apparently propodeum reticulate rugose;
7:
4: 4: 4: 4: 4,5: 4: 4;
apparently granulated;
scutellum,
pterostigma
11:
apparently granulated,
occipital carina complete;
not touching occipital
carina);
8:
(greatest breadth):
long and narrow;
visible;
without dark
apparently fore-wing
distal
part of radial
with
vein
longer than proximal part; tibial spurs 1,1,2. LOCUS TYPICUS: Taimyr amber (Sibiria) TYPICAL MATERIAL: HOSTS:
holotype M!
in MPI.
unknown
DISTRIBUTION:
only
known
from Taimyr amber
(Sibiria,
U.
S.
S.
R.)
(Laberius Kieffer) is not correct, because Laberius is a genus of Gonatopodinae (it's a junior synonym of Dicondylus Hal iday). As its occipital carina is complete, L. antiquus is belonging to Dryininae. The new generic name Dryinus is provisionally proposed. The only known specimen of L. antiquus is in too bad conditions to permit to associate it for a certainty to a genus of Dryininae. D. antiquus is the oldest fossil dryinid known (nearly 100 millions NOTES:
the original
generic status
years old).
ETHIOPIAN SPECIES OF
Nineteen
species are
DRYINUS
known.
KEY TO THE SPECIES
752
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
FEMALES
Posterior collar of
2
3
4
5
6 7
8
pronotum invisible . . . . . . . . . . . . . . . . . . . . . . . .. . . . . . . . . . . . . . . . . . . . . . . . . . . . 1. bisulcatus (Benoi t) Posterior collar of pronotum visible (Figs 503 B, 519, 522) ... 2 Enlarged claw much shorter than segment 5 of front tarsus and wi th subapical tooth very small (Fig. 504) . . . . . . . . . . . . . . . . . . . .. · . . . . . . . . . . . . . . . . . . . . • • . . .. 2. seyrigi Ben 0 i t Enlarged claw nearly as long as segment 5 of front tarsus and usually with subapical tooth big (Figs 506,508) . . . . . . . . . . . . . . 3 Head (dorsally viewed) with posterior ocelli placed behind the imaginary straight line joining posterior edges of eyes (Figs 505, 507) ••••.••••••••••••••••••••••••••••••••.••••••••••••••• 4 Head (dorsally viewed) wi th posterior ocell i placed in front of the imagina;"y straight line joining posterior edges of eyes or on t his lin e (F i g s 51 4, 524) •••••.•••••••••••..•••••••••••• 8 Head wi th posterior margin of vertex excavated (Fig. 505 A); head excavated; posterior ocell i touching occipi tal carina (Fig. 505 A) . . . . . . . . . . . . . . . 3. alticolus (Benoit) Head wi th posterio margin of vertex convex (Figs 505 B, 507 B); head flat; posterior ocelli not touching occipital carina (Figs 505 B, 507) ••••••••••••••••••••••••••••••••••••••••••••.•••••• 5 At I east the propodeum black . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6 Species fully testaceous or reddish-testaceous, with petiole black ••••••••••••••••••••••••••••••••••••••••••••••••••••••••• 7 Head almost fully black . . . . 7. sierranus n. sp. Head almost fully reddish •• 8. pretorianus n. sp. Posterior margin of vertex more convex (Fig. 507 B); antennal segment 3 nearly ten times as long as segment 2 . . . . . . . . . . . . . . .. · . . . . . . . • • . . • . . . . . • • . • . . . .. 4. spangleri n. sp. Posterior margin of vertex less convex (Fig. 507 A); antennal segment 3 nearly six-seven times as long as segment 2 •••••••••• . . . . . . . . . . . . . . . . . . . . . . . . . . . 5. deceptor (Turner) Species fully testaceous, with petiole black and sometimes abdomen brownish; scutum shiny, weakly punctate ••••••••••.••••• . . . . . . . . . . . . . . . . . . . . . . . . . . . 6. mahensis Ki effer Species at least partly black; scutum shiny or dull, punctate or granulated, usually with more or less short irregular keels
· •.••••••..•..•..••.. " .••.•••.•.••.••••••.•• " •.••••. " •••••. 9 9
Posterior ocelli
touching
Posterior ocelli
not
524) ••••••••••••••••• " 10 Notaulices
occipital
touching
•••••••••• "
reaching approximately
........................... Notaulices
9. 10.
(Fig.
carina
514 A) . . . . . . 10
(Figs
514 B,
•.••.••••••••.•••••••••••. 11 0,65
length of
scutum . . . . . . . .
saussurei (Ceballos)
reaching approximately
• ••••••••••••••••••••••• "
carina
occipital
0,3
length of
hova (C eba I I os)
scutum . . . . . . . . .
753
Dryininae
11 Temples prominent prominent,
but
(Figs 514 B,
in
this
524 A);
rarely
temples weakly
case head with posterior margin
of
514 B, 524 A) . • . . . . . . . . . • • • • • • • • • • • • . . 12 Temples invisible (Fig. 524 B, C); head wi th posterior margin of vertex convex (Fig. 524 B, C) . • . • • . . . • . • • • . . . . . • • • • . • • • • • . 18 vertex
excavated
(Figs
12 Fore-wing with distal as
long as proximal
Fore-wing with distal times as
part
of
part ••
11.
part
of
radial
vein nearly
three
times
basilewskyi Benoit radial
vein
less
than
three
part ••••••••••••••••••••••••••••••• 13
long as proximal
13 Scutum punctate, not granulated, wi thout keel s ••••••••••••••••• •••••••••••••••••.•••••••• 12. chamaeleo Benoi t Scutum granulated or with keels ••••••••••••••••••.•.••••.•••• 14 14 Head granulated, with irregular or regular keels . . . . • • . • • • . . • 15 Head granulated,
without
keels ••••••••••••••••••••••••••••••• 17
15 Pronotum wi th anterior impression stronger (Fig. 519 A); head mostly reddish •••••••••••• 13. schoutedeni (Benoit) Pronotum with anterior impression weaker (Fig. 519 B); head mostly black ••••••.•••.••••••••••••••••.••......•.••••••••••• 16
16 Propodeum long, wi th dorsa I surface more than twi ce as long as posterior surface ••••••••• 14. mayogo (Benoit) Propodeum short, with dorsal surface slightly longer than posterior surface ••••••••• 15. ol'ophilus (Benoit) 17 Scutum granulated, not reticulate rugose; subapical tooth of enlarged claw very near apex (Fig. 525) •••••..•••••.••••••••••• 16. modestus n. sp. Scutum granulated, with posterior half reticulate rugose; subapical tooth of enlarged claw farther from apex (Fig. S 2 6) • • • • • • • • • • • • • • . • . . • . .• 1 7. incel'tus n. sp. 18 Anterior impression of pronotum very weak (Fig. 522 B) •••••••••
18. bUl'geoni Beno it of pronotum stronger (Fig.
•••••••••••••••••••••••••, Anterior
impression
••••••••••••••••••••••••••
19.
522 A) ••••••••••
aiel' (Dlmi)
MALES
Head wi th OL
= OOL •••••••••.••••••.•.•.••••••••••••••••••....• 2
Head wi th OL more or 2
Fore-wing with
radial
less
shorter
cell
than OOL •••••••••••••••••••• 3
closed;
head with posterior ocelli
not touching occipital carina . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .. . . . . . . . . . . . . . . . . . . . . . . . . . . . 6. mahensis (Kieffer) Fore-wing with touching
radial
occipi tal
cell
• • • • • • • • •• •• • •• • • • • • • •• • • •• 3
open;
head with posterior ocelli
carina •••••••..••...••••••••••••.....••••••• 9.
saussul'ei (Ceba I los)
Fore-wing wi th
radial
cell
closed . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4
Fore-wing with
radial
cell
open . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5
754
4
Mem.
Amer.
Ent.
Ins!.,
no.
37,
1984
Frons flat in front of the anterior ocellus, convex only between the antennal sockets • • . . • • • • • • • • . . • • • • • • • • • . • . • • • • •
19. afer (Olmi) Frons wi th a
5
median
antennal
sockets •••
Vo I s e I I a
n ea r I y
as
longitudinal •••••••
shorter
from anterior
long as gonoforceps
• • • • • • • . • • • • • • • • • • • .. Volsella
hump
(Fig.
to
513 B) . . . . . . . . . . .
(Fig.
pretorianus n.
8.
513 A)
ocellus
deceptor (Turner)
5.
sp.
••••••••.
15. orophilus (Benoit)
The males of other species are unknown. The description of the male allotype of (1953b)
is not reliable.
addition
the
D. chamaeleo by Benoit
This male was not obtained by breeding;
in
locality of the holotype is different.
Dryinus bisulcatus (Benoit) n. comb.
1.
Mercetia bisulcata Benoit 1953b: 400.
FEMALE:
fully
winged;
length 5,62 mm;
head black,
with clypeus and
mandibles testaceous; antennae testaceous; propectus black; pronotum black, with sides reddish; mesothorax, metathorax and propodeum black; abdomen black; black;
legs testaceous,
antennae distally
proportions:
11: 6;
shiny,
smooth,
behind
the ocelli);
21:
weakly
with coxae and clubs of femora brown or
weakly
13:
11:
thickened; 7,5:
alutaceous;
OL
6,5:
antenna I segments
4,5:
occipital
longer than POL;
6:
6;
carina
in following
head excavated, invisible
pronotum shiny,
(no track
weakly
granulated, crossed by a transversal impression on anterior third; the pronotum so is divided into 2 collars, without disc; the posterior collar is weakly
humped;
tegulae;
pronotal
scutum shiny,
converging,
reaching
surface with 2 other wing
proximal
part;
with subapical
median keels;
spurs 1,1,2. unknown
tarsus produced
tooth and
transversal
strongly
scutellum shiny propodeum not
keels;
anterior
sides of anterior surface with 2
posterior surface with 2
segment 1 of front
with 2 rows of 41 MALE:
keels;
with 3 brown spots and with distal
segment 3 of front
not reaching
incomplete,
metanotum very reduced and short;
with numerous parallel
longitudinal
longitudinal
notaul ices
almost posterior margin of sctum;
and without sculpture; reticulate rugose,
tubercles not very prominent,
granulated;
longitudinal
keels;
fore-
vein as
long as
tarsus shorter than segment 4
(18:22);
into a
17 lamellae;
part of radial hook;
enlarged claw
segment 5 of front
lamellae extending continuously
tarsus
to the apex;
(Fig. (Fig. tibial
502) 502)
I
Dryininae
Fig.
I
502 -
LOCUS TYPICUS: HOSTS:
Chela of Dryinus bisulcatus
Behara
TYPICAL MATERIAL:
755
(Benoit)(type)
(Madagascar)
holotype Fl
in P.
unknown
DISTRIBUTION:
Ethiopian region:
MADAGASCAR:
Behara,
Pl
D. bisulcatus is species of transition from Dryininae to Gonatopodinae. For the tibial spurs (1,1,2) and the pronotal tubercles directed towards the tegulae it is belonging to Dryininae; for the absence NOTES:
of occipital
carina,
metanotum very
the clypeus ventrally rounded
reduced
it
is near Gonatopodinae.
(not bidentate)
and
the
756
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
A
B Fig. 503 - Pronotum (laterally viewed) of females of Tridryinus ampuliciformis (Turner) from Afgoi (Soma I i a )(A) and Dryinus seyrigi Benoi t (type) (B)
2. Dryinus seyrigi Benoit
Dryinus seyrigi Benoit 1953b: 407.
FEMALE:
fully
winged;
length 3,75 mm;
head black,
with front part of
vertex, clypeus and mandibles testaceous; antennae testaceous, with segments 6-9 brown; propectus black; pronotum black, with sides and posterior collar reddish;
mesothorax,
metathorax and propodeum black;
abdomen missing in holotype (only specimen known); legs testaceous, with coxae and clubs of femora brown; antennae distally weakly thickened; antennal 8;
segments
head shiny,
prominent;
in following proportions:
granulated;
pronotum
posterior transversal
(Fig.
occipital 503 B)
impresssion
10:
5:
21:
carina complete;
shiny,
granulated,
and by a
10:
9:
8:
OL = POL;
6:
6:
4,5:
temples
crossed by a
strong
weak anterior transversal
impression; disc humped; posterior collar short, distinct; anterior collar weak; pronotal tubercles not reaching tegulae; scutum dull, granulated, with short keels near posterior margin; approximately 0,5 short, without
length of scutum;
with numerous longitudinal
transversal
longitudinal
notaulices incomplete,
scutellum dull,
keels;
metanotum
propodeum reticulate rugose,
keels on posterior surface;
bands and with distal
reaching
granulated;
fore-wing
part of radial
with 3 brown
vein much
longer than
D'ry i n i nae
Fig.
504 -
Chela of
Dryinus seyrigi Benoit (type)
757
758
Mem. Amer.
Ent.
Inst.,
no.
37,
1984
A
Fig.
Heads of females of Dryinus alticolus (Benoit) (type) (A) and deceptor (Turner) from Port SI. Johns (South Africa) (B)
505 -
proximal part; segment 1 of front tarsus nearly twice as long as segment 4; segment 3 of front tarsus produced into a hook; enlarged claw (Fig. 504) with a small subapical tooth and 3 lamellae; segment 5 of front tarsus (Fig. 504) with 2 rows of 22 lamellae extending continuously to the apex; segment 5 of front tarsus apically without prominence and much longer than enlarged claw; tibial spurs 1,1,2. MALE: unknown LOCUS TYPICUS: Ranomafana (Madagascar) TYPICAL MATERIAL: holotype F! in P. HOSTS: unknown DISTRIBUTION: Ethiopian region MADAGASCAR:
Ranomafana,
3. Dryinus alticolus (Benoit) n. comb.
Lestodryinus alticola Benoit 1951c: 4.
P!
Dryininae
Fig.
FEMALE:
fully
506 -
winged;
Chela of Dryinus alticolus
length 5,0 mm;
side of head;
(Benoit) (type)
head ferruginous,
brown spot on ocell ar reg ion and part I y visible on ventral
759
on frons;
with an elongate
another brown spot is
antennae testaceous;
propectus black,
wi th a dorsal reddish spot; pronotum ferrug i nous; mesothorax, meta thorax and propodeum black; abdomen brown; legs testaceous, with hind and mid coxae partly brown;
head
(Fig.
505 A)
excavated,
dull,
granulated,
with
760
Mem.
Amer.
En!.
Ins!.,
no.
37,
1984
oa
o Fig.
507 -
Heads of females of
Johns
temples
(South Africa) (A)
0
Dryinus deceptor (Turner) from Port St. and spangleri n. sp. (type) (B)
very prominent and posterior margin of vertex excavated;
occipital
carina complete; TL nearl,' three times as long as POL; OL weakly longer than POL; posterior ocell i touching occipital carina; pronotum dull, granulated;
anterior collar and
posterior collar distinct;
anterior transversal
posterior transversal
tubercles not reaching
tegulae;
near posterior margin;
notaulices incomplete,
scutum dull,
length of scutum;
scutellum dull,
reticulate rugose,
without
wing
with 3 brown
much
longer than proximal
granulated;
longitudinal
transversal part
impression
granulated,
weak;
pronotal
with some areolae
reaching approximately 0,75 metanotum rugose;
keels on segment
propodeum
posterior surface;
bands and with distal (17:4);
very
impression strong;
fore-
part of radial
1 of front
vein
tarsus nearly
as long as segment 4; segment 3 of front tarsus produced into a hook; enlarged claw (Fig. 506) with subapical tooth and 4 lamellae; segment 5 of front of 10 MALE:
tarsus
lamellae; unknown
(Fig. tibial
506)
with 2 rows of 24
spurs 1,1,2.
lamellae;
apex with a
group
761
Dryininae
Fig.
508 -
Chela
0
f Dryinus spanglen. n.
sp.
(type)
762
Mem.
LOCUS TYPICUS:
Kilinga
TYPICAL MATERIAL: HOSTS: unknown
Amer.
Ent.
Inst.,
no.
37,
1984
(Zaire)
holotype F!
in TV;
1 paratype F!
DISTRIBUTION: Ethiopian region: ZAIRE: CAMEROON: N I Kongsamba (Beno i t 19S 1c) •
Kilinga
in P.
(Rutshuru),
TV!
4. Dryinus spangleri n. sp.
FEMALE:
fully
winged;
length 5,62 mmj
triangle and petiole black; white,
6-8 brown;
antennae distally weakly
in following
proportions:
507 B)
flat,
dull,
10:
OOL = 13;
OPL = 2j
imaginary straight of vertex convex;
57:
testaceous,
36:
24:
weak I y
antennal
14:
7:
11: 8:
frons
vis i b Ie on si des of occi put;
TL = 7;
posterior ocelli
POL = 3;
OL
posterior margin
impression;
disc humped;
longer than pronota I tubercles;
pronota I
pronotum granulated and with numerous
longitudinal
striae on sides;
longitudinal
striae on
scutum dull,
lateral region; fore-wing
granulated and with numerous
notaulices weak,
scutellum and metanotum dull,
fully reticulate rugose;
(Fig.
crossed by an anterior weak transversal
much
tubercles reaching tegulae;
segments
head
placed behind the
a posterior strong transversal long,
10;
with a median keel;
I ine joining posterior edges of eyes;
impression and by
with ocellar
with segments 4-5 partly
thickened;
weakly rugose;
pronotum du II,
posterior coil ar very
complete;
7:
granulated,
occipi ta I cari na comp I ete, = 2;
fully
antennae testaceous,
granulated;
but distinct and
propodeum shiny,
with 2 dark transversal
bands,
one on
the basal cells and another beneath the pterostigma; distal part of radial vein much longer than proximal part; fore tarsal segments in following proportions:
27:
3:
8:
22:
tooth and 12 lamellae; Clearly 33 lamellae;
33;
enlarged claw
segment 5 of front
(Fig.
tarsus
apex with a group of 23
508)
(Fig.
lamellae;
with a subapical 508) tibial
with 2 rows of spurs 1,1,2.
MALE: unknown LOCUS TYPICUS: Mombasa (Kenya) TYPICAL MATERIAL: holotype F! in WA. HOSTS:
unknown
DISTRIBUTION:
Ethiopian region:
KENYA:
Mombasa,
WAr
5. Dryinus deceptor (Turner) n. comb.
Lestodryinus deceptor Turner 1928: 147.
FEMALE:
fully
winged;
length 6,0 mm;
and abdomen reddish-brown; granulated;
occipital
head
yellow-testaceous,
(Figs 505 B,
carina complete;
507 A)
with petiole black
flat,
OPL shorter than POL;
dull, OL = POL;
Dryininae
763
Fig. 509 - Chela of Dryinus deceptor (Turner) from Port SI. Johns (South Africa)
764
Mem.
temples short;
Amer.
pronotum dull,
posterior transversal
Ent.
Ins!.,
granulated,
impression strong;
anterior transversa I impression weak; tegulae;
scutum shiny,
approximately 0,5 granulated;
granulated;
no.
with numerous keels on sides;
pronota I tuberc les not reach i ng incomplete,
longitudinal
reaching
scutellum and metanotum shiny,
propodeum weakly reticulate rugose;
propodeum without
1984
posterior collar long and visible;
notaul ices
length of scutum;
37,
keels;
posterior surface of
fore-wing
with 2 brown
transversal
bands and with distal part of radial vein much longer than proximal part; segment 1 of front tarsus shorter than segment 4 (18:22); segment 3 of front
tarsus produced
subapical
tooth and
2 rows of 34
into a
hook;
12 lamellae;
lamellae;
enlarged claw
segment 5 of front
(Fig.
509)
tarsus
apex with a group of 19 lamellae;
with
(Fig. tibial
509)
with
spurs
1,1,2. MALE: fully winged; length 2,43 mm; black; mandibles testaceous; antennae brown, with segments 1-2 testaceous; legs testaceous, with coxae, clubs of femora, mid and hind tibiae brown; antennae not distally thickened; antennal segments in following proportions: 5: 3: 11: 7: 6:
6:
5:
7;
head dull,
with a median frons
longitudinal
without median keel;
OOL = 4; dull,
granulated,
OPL = 1;
granulated,
reaching 0,5 transversal transversal
with weak tracks of areolae;
hump from anterior ocellus to antennal occipital
temples
invisible;
with weak
carina
incomplete;
scutum,
6:
sockets;
= 5,6; OL = 2;
scutellum and metanotum
tracks of areolae;
length of scutum;
POL
6:
frons
notaul ices incomplete,
propodeum reticulate rugose,
dull,
without
and longitudinal keels; fore-wing hyal ine, without dark bands; distal part of radial vein longer than proximal part;
radial cell closed; tibial spurs 1,1,2. LOCUS TYPICUS: Port SI. Johns (South Africa) TYPICAL MATERIAL:
holotype F!
and 1 paratype F!
HOSTS: unknown DISTRIBUTION: Ethiopian region:
UGANDA:
Port St.
St.
Johns
(Pondoland),
BM!
in SM.
Kampala,
TW!
SOUTH AFRICA:
Lucia Estuary,
TW!
OL!
6. Dryinus mahensis (Kieffer) n. comb.
Mesodryinus mahensis Kieffer 1912a: 45. Labeo saxetanus Kieffer 1912a: 46 (n. syn.); locus typicus Mahe Island
(Seychelles
Islands).
Mesodryinus mahensis Kieffer: Kieffer 1914b: 31. Laberius saxetanus (Kieffer): Kieffer 1914b: 64.
FEMALE:
fully
winged;
abdomen darkened; complete, slightly
length 4,75 mm;
head shiny,
testaceous,
weakly punctate,
almost invisible on sides of occiput; longer than POL;
pronotum shiny,
with petiole black and
hairy;
occipital carina
OPL shorter than OLi
smooth,
without sculpture;
OL
Dryininae
Fig.
510 -
posterior transversal
765
Dryinus mahensis (Kieffer)(type)
Chela of
impression strong;
anterior transversal
impression
and anterior collar weakly visible; posterior collar visible; pronotal tubercles not reaching tegulae; scutum shiny, smooth, weakly punctate; notaul ices incomplete, reaching approximately 0,65 length of scutum; scutellum and metanotum shiny, smooth, weakly punctate; propodeum reticulate rugose,
without
wing
with 2 brown
much
longer than proximal
longitudinal
transversal
part;
segment
segment 4;
segment 3 of front
(Fig.
510)
with subapical
(Fig.
510)
with 2 rows of 30 lamellae;
tibial
spurs 1,1,2.
·.lALE:
fully
winged;
1 of front
tarsus produced
hook;
vein
long as
enlarged claw
segment 5 of front
apex with a head black,
fore-
part of radial
tarsus as
into a
tooth and 14 lamellae;
length 3,43 mm;
antennae yellow-brown;
keels on posterior surface;
bands and with distal
tarsus
group of 15 lamellae;
with mandibles yellow;
thorax and propodeum bl ack;
abdomen brown;
legs
yellow, with hind coxae partly black; head dull, granulated; occipital carina incomplete; temples invisible; POL; 8; OL ; 3; OOL ; 3; OPL ; -~
;
0;
scutum dull,
granulated,
with
weak
areolae on
lateral
regions;
2;
766
Mem.
Fig.
511
-
Amer.
Chela of
Ent.
Inst.,
no.
37,
1984
Dryinus sierranus n. sp. (type)
notaulices incomplete, reaching approximately 0,65 length of scutum; scutellum shiny, weakly granulated; metanotum dull, rugose; propodeum reticulate rugose; fore-wing without dark transversal bands and with distal part of radial vein much longer than proximal part; tibial spurs 1,1,2 •• LOCUS TYP ICUS: Port Glaud (Seychelles TYPICAL MATERIAL: holotype F! of M.
Islands)
mahensis in BM; holotype M! of L.
saxetanus in BM. HOSTS: unknown DISTRIBUTION: Ethiopian region:
SEYCHELLES
ISLANDS: Port Glaud
Island), BM! Mount Sebert (Mahe Island), BM! (Silhouette Island)(Kieffer 1912a).
Mare aux Cochons
7. Dryinus sierranus n. sp.
(Mahe
Dryininae
767
FEMALE: fully winged; length 8,75 mm; head black, with occiput, anterior region of frons, clypeus and mandibles reddish; antennae black, with segments 1-2 reddish, segment 4 yellow,
junction between segments 3-4 testaceous, half of segment 4 yellow,
proximal
prothorax reddish-brown-dark; mesosternum reddish-dark; metasternum black;
scutum bl ack,
scutellum,
abdomen black,
with junctions and tarsi
reddish;
apex of
segment 10 yellow;
wi th sides redd ish-brown;
metanotum,
propodeum,
pleura,
with some reddish regions;
legs black,
antennae distally weakly thickened;
antennal segments in following proportions: 10: 7: 58: 33: 24: 14: 11: 8: 8: 10; head dull, granulated, with tracks of irregular keels on frons; frons with a median keel; occipital carina complete; POL = 5; OL = 3,5; OOL = 11; OPL = 2; TL = 7; posterior ocelli placed behind the imaginary straight line joining posterior edges of eyes; posterior margin of vertex convex; pronotum crossed by strong transversal impression and with a posterior collar long and distinctly visible; pronotum dull, granulated and with numerous longitudinal keels around disc; pronotal tubercles not reaching tegulae; scutum dull, granulated, with numerous weak longitudinal striae; notaul ices incomplete, almost reaching posterior margin of scutum, numerous
but weakly visible; longitudinal striae;
fully reticulate rugose,
scutellum dull, metanotum dull,
granulated and with reticulate rugose;
with 2 incomplete longitudinal
propodeum
keels on posterior
surface; fore-wing with 2 dark transversal bands, one on the basal cells and another beneath the pterostigma; distal part of radial vein longer than proximal 10: 23:
38;
lamellae;
part;
fore tarsal
enlarged claw
(Fig:
segment 5 of front
segments in following proportions: 511)
tarsus
apex with a group of 14 lamellae; MALE:
with a subapical (Fig.
tibial
511)
28:
5:
tooth and with 20
with 2 rows of 49 lamellae;
spurs 1,1,2.
unknown
LOCUS TYP ICUS:
Freetown
TYPICAL MATERIAL: HOSTS:
(Sierra Leone)
holotype F!
in TW.
unknown
DISTRIBUTION:
Ethiopian region:
SIERRA LEONE:
Freetown,
TW!
8. Dryinus pretorianus n. sp.
FEMALE:
fully
winged;
brown spot on frons; whi t ish;
prothorax,
propodeum black;
length 6,25 mm;
antennae brown,
head reddish-testaceous,
scutum and scutell um redd ish-dark;
abdomen brown,
with a
with segments 1-2 reddish,
with segment 1 partly reddish;
with chelae and segments 2 and partly
1 of hind tarsi
10
metanotum and whitish;
legs,
antennae
distally thickened; antenna I segments in following proportions: 9: 7: 49: 29: 21: 14: 10: 9: 7: 8; head dull, flat, granulated; frons with a median keel;
POL = 4,5;
complete;
DL = 3,5.
DOL = 11; DPL = 2;
posterior margin of vertex convex;
the imaginary straight
TL = 7;
occipital
carina
posterior ocelli placed behind
line joining posterior edges of eyes;
pronotum with
768
Mem. Amer. Ent.
Inst., no. 37, 1984
Fig. 512 - Chela of Dryinus pretorianus n. sp.
(type)
Dryininae
Fig.
513 -
769
Dryinus orophilus (Benoit) from Ile-ife
Male genitalia of
(Nigeria) (A: right half removed) and pretorianus n. sp. from Pretoria (South Africa) (8: right half removed; C: ninth abdominal sternite)
a strong
transversal
dull,
hairy,
dull,
granulated,
impression and with a
granulated;
pronotal
posterior collar
tubercles not reaching
with numerous irregular weak keels;
long;
tegulae;
pronotum scutum
notaul ices
incomplete, reaching approximately 0,5 length of scutum; scutellum and metanotum as sculptured as scutum; propodeum reticulate rugose, dull, with
two median
longitudinal
keels on dorsal
surface;
fore-wing
with 2
dark transversa I bands, one on the basa I cell s and another beneath the pterostigma; distal part of radial vein longer than proximal part (15:7); fore tarsal
segments
claw
(Fig.
512)
front
tarsus
to the apex; MALE:
fully
in following proportions:
with a subapical
(Fig.
512)
tibial winged;
with 2 rows of 45
4:
10:
legs brown,
3;
occipital
scutum dull,
17:
thickened; 8:
8:
8:
black;
28;
enlarged
segment 5 of
antennal 8:
carina complete; granulated;
mandibles and antennal
with joints and tarsi
6:
6:
segments 8;
with some weak areolae on sides of frons; keel;
8:
lamellae extending continuously
length 2,37-2,62 mm;
antennae not distally 4:
3,5:
spurs 1,1,2.
segments 1-2 testaceous; proportions:
20:
tooth and with 10 lamellae;
POL = 6;
notaul ices
head dull,
granulated and
frons with an OL = 3;
incomplete,
testaceous;
in following incomplete median
OOL = 4; reaching
OPL = 1;
TL =
approximately
770
Mem.
Amer.
En!.
Inst.,
no.
37,
1984
\) 0
Fig.
514 -
Heads of females of Dryinus saussurei (Madagascar) (A)
(Ceballos)
and basilewskyi Benoit
from Beki Iy
(type) (B)
0,65 length of scutum; scutellum dull, granulated; metanotum dull, granulated and with numerous areolae near anterior margin; propodeum reticulate rugose,
without transversal
or longitudinal
keels;
fore-wing
hyaline, without dark transversal bands; distal part of radial vein longer than proximal part (9:7); distivolsella (Fig. 513 B) with apex located nearly on the same level with the apex of gonoforceps; sternite: fig. 513c; tibial spurs 1,1,2. LOCUS TYP ICUS:
Pretoria
TYPICAL MATERIAL: HOSTS:
ninth abdominal
(South Africa)
holotype F!
and 2 paratypes MM!
in TW.
unknown
DISTRIBUTION:
Ethiopian region:
SOUTH AFRICA:
Pretoria
(Transvaal),
TVY!
9. Dryinus saussurei (Ceballos)
Lestodryinus saussurei Ceballos 1935: 48. Dryinus saussurei (Ceballos): Benoit 1953b: 412.
FEMALE:
fully
winged;
mandibles yellow;
length 4,00-5,93 mm;
rarely clypeus black;
with segments 7-8 or 3-10 brown;
head black,
propectus black;
margins and posterior collar reddish;
with clypeus and
antennae testaceous,
mesothorax,
sometimes
pronotum black,
with
metathorax and
propodeum black; abdomen black-brown; legs testaceous, wi th coxae and clubs of femora partly brown; head (Fig. 514 A) shiny or du", fully or partly
granulated;
occipital
carina complete;
posteior ocell i
touching
Dryininae
Fig.
515 -
Chela of
771
Dryinus saussurei (Ceballos) from Bekily (Madagascar)
772
Mem.
occipital
carina;
Amer. En!.
OL = POL;
Ins!.,
pronotum dull,
with numerous weak striae around disc; nearly 0,5 times as
no.
37,
1984
partly or fully granulated,
posterior collar of pronotum short,
long as distance from
tegulae to pronotal
tubercles;
pronotal tubercles not reaching tegulae; scutum dull, granulated, with short keels or areolae near posterior margin; notaulices incomplete, almost reaching posterior margin of scutum; dull,
rugose;
posterior surface; proximal twice as into a claw
hook;
with
segment
515)
(Fig.
lamellae; MALE:
fore-wing
with 2
granulated;
longitudinal
with 3 brown transversal
bands
metanotum
keels on
(the two
bands partly joint) and with distal part of radial vein nearly long as proximal part (18:8); segment 3 of front tarsus produced
(Fig.
tarsus
scutellum dull,
propodeum reticulate rugose,
515)
tibial
fully
1 of front tarsus as
with subapical
tooth and
with 2 rows of 39
long as segment 4; 13
lamellae;
lamellae;
enlarged
segment 5 of front
apex with a
group of 11
spurs 1,1,2.
winged;
teeth brown;
length 2,60-2,84 mm;
antennae brown,
black;
mandibles testaceous,
with segments 1-2 testaceous;
palpi
testaceous; abdomen brown; legs testaceous, with coxae, clubs of hind femora and hind tibiae brown; antennal segments in following proportions: 8:
7:
17:
12:
13:
13:
11:
incomplete median keel, between
10:
10:
flat
the antenna I sockets;
DOL = 3; posterior ocelli scutum dull, granulated;
11;
head dull,
granulated;
frons with an
in front of the anterior ocellus and humped occipital
carina complete;
POL = 9;
OL = 3;
touching occipital carina; temples invisible; notaulices incomplete, reaching approximately
0,65 length of scutum; scutellum almost fu!ly dull, granulated, with a narrow posterior region smooth and shiny; metanotum with anterior half rugose and with posterior half shiny and smooth;
propodeum reticulate
rugose,
fore-wing hyal ine,
without
longitudinal
without dark
transversal
part slightly
longer than
segments;
labial
LOCUS TYPICUS:
palpi Rogez
TYPICAL MATERIAL:
or transversal
bands;
proximal
MD!
Behara,
part
with 3 segments;
cell open; (14:13); tibial
radial
vein
with distal
maxillary palpi
with 6
spurs 1,1,2.
(Madagascar)
holotype F!
HOSTS: unknown DISTRIBUTION: Ethiopian region: Rogez,
radial
keels;
TV!
in MD. MADAGASCAR:
Fort Dauphin,
TV!
Bekily,
TV!
P!
Mandraka,
TV!
Ranomafana
Ivondro,
P!
(Benoit 1953b).
10. Dryinus hova
(Ceballos)
n.
comb.
ft;esodryinus hova Ceballos 1936: 53. fiJesodryinus hova Ceballos: Benoit 1953b: 414.
FEMALE:
fully
winged;
mandibles testaceous;
length 5,31
mm;
propectus black;
head black,
with clypeus and
pronotum black,
with margins and
773
Dryininae
Fig.
516 -
Chela of
Dryinus hova
(Ceballos)(type)
posterior collar reddish; mesothorax, metathorax, propodeum and abdomen black; legs testaceous, with coxae and clubs of femora brown; antennae distally weakly 5:
24:
10: 9:
transversal
thickened;
7:
6:
6:
5:
antennal 7,5;
segments
head dull,
keels on anterior half of frons;
posterior ocelli edges of eyes;
located on
in following proportions:
granulated,
occipital carina complete;
the imaginary straight
posterior ocelli
9:
with numerous short
touching occipital
line joining posterior carina;
OL = POL;
pronotum dull, granulated, with anterior transversal impression weak; anterior collar weak; posterior transversal impression strong; posterior collar short,
distinct;
pronotal
tubercles not reaching
tegulae;
scutum
dull, granulated, with some strong punctures; notaulices incomplete, reaching approximately 0,3 length of scutum; scutellum dull, granulated; metanotum rugose;
propodeum reticulate rugose,
on posterior surface;
fore-wing
half of fore wings missing species); of front
without
longitudinal
transversal
bands;
in holotype (only known specimen of the
tarsus produced into a hook; lamellae;
with
tarsus
(Fig.
516)
apex with a group of 12 lamellae;
tibial
spurs
segment 5 of front
unknown
LOCUS TYP I CUS:
Bek i I Y (Madagascar)
TYPICAL MATERIAL:
holotype F!
enlarged claw
in P.
(Fig.
segment 3
516)
tooth and 8 lamellae;
2 rows of 21
keels
distal
segment 1 of front tarsus nearly as long as segment 4;
subapical 1,1,2. MALE:
with 2 brown
with
774
Mem. Amer.
Fig. 517 -
Ent.
Ins!., no. 37,
1984
Chela of Dryinus basilewskyi Benoit
HOSTS: unknown DISTRIBUTION: Ethiopian region:
MADAGASCAR:
Bekily, P!
11. Dryinus basilewskyi Benoit
(type)
Dryininae
775
Dryinus basilewskyi Benoi t 1953a: 143.
FEMALE: vertex
fully
winged:
length 7,50 mm;
head black,
testaceous;
propectus black;
coil ar redd i sh; yellow-brown; sculpture,
mesothorax,
pronotum black, head
posterior ocelli
514 B)
very near occipital
than OL;
OL = POL;
except for posterior collar very
impression strong; distinct,
tubercles
reaching
tegulae;
abdomen
shiny,
without
(4:9);
granulated;
length of scutum; fore-wing
part of radial
vein
segment 3 of front
posterior collar
weak;
tegulae to
pronotal
tubercles not
notaulices incomplete,
without
longitudinal
transversal
three times as
tarsus nearly as
tarsus produced
weak;
reaching
scutellum and metanotum dull,
with 2 brown
more than
segment 1 of front
occipital
OPL shorter
posterior transversal
long as distance from
propodeum reticulate rugose,
posterior surface;
hairy;
impression
anterior collar very
scutum dull,
carina,
with numerous striae around disc,
anterior transversal
approximately 0,65 granulated;
pronotum shiny,
nearly 0,5 times as
pronotal
(17:5);
(Fig.
except for numerous weak striae on frons and vertex;
hairless,
distal
antennae
with sides and posterior
meta thorax and propodeum black;
legs yellow-testaceous;
carina complete;
short,
with front part of
(more along orbits), clypeus and mandibles yellow;
into a
long as proximal
long as segment 4
hook;
keels on
bands and with
enlarged claw
part
(17:16); (Fig.
517)
with subapical tooth and 11 lamellae (lamellae near subapical tooth very long and simi lar to peg-I ike hairs); segment 5 of front tarsus (Fig. 517) with 2 rows of 30 lamellae; 1,1,2. MALE:
apex
with
a group of 13 lamellae;
tibial
spurs
unknown
LOCUS TYPICUS:
Thysville
TYPICAL MATERIAL: HOSTS:
(Zaire)
holotype F!
in TV.
unknown
DISTRIBUTION:
Ethiopian region:
ZAIRE:
Thysville,
TV!
12. Dryinus chamaeleo Benoit
Dryinus chamaeleo Benoit 1953b: 409.
FEMALE: frons;
fully
winged;
length 6,0 mm;
antennae testaceous;
metanotum redd ish-brown; ferruginous;
propectus,
head reddish, pronotum,
propodeum black;
with a brown spot on
scutum,
scutell um and
abdomen reddi sh-brown;
antennae distally weakly thickened;
antennal
segments
legs in
following proportions: 10: 5: 48: 27: 22: 15: 12: 10: 8: 10; head shiny, weakly punctate, with numerous longitudinal keels on median region of frons; carina;
occipital
carina complete;
OPL = OL;
OL = POL;
posterior ocelli
pronotum shiny,
not touching occipital
without sculpture,
except
776
Memo Amero En!.
Ins!.,
noo 37,
1984
Fig. 518 - Chela of Dl'yinus chamaeleo Benoit (type)
777
Dryininae
A
B
Fig.
519 -
Pronotum of females of Dryinus schoutedeni and mayogo (Benoit) (type) (B).
for the sides which show
numerous
longitudinal
impression and anterior collar weak; strong;
posterior collar
not reaching notaulices,
tegulae;
reaching
rugose; distal
fore-wing
part of radial
1 of front produced
into a
14 lamellae; lamellae;
vein
tarsus nearly hook;
without
with 2 brown
nearly
as
keels;
long
twice as
segment 5 of front
tarsus
scutellum shiny,
longitudinal
keels on
bands and with
long as proximal
(Fig. (Fig.
apex with a group of 12 lamellae;
tubercles
punctate;
metanotum punctate,
transversal
as segment 4;
enlarged claw
impression
pronotal
length of scutum;
longitudinal
propodeum reticulate rugose,
posterior surface;
disc humped;
without sculpture,
approximately 0,65
punctate and with numerous
anterior transversal
posterior transversal
long and distinct;
scutum shiny,
keels;
(Benoit) (type) (A)
part;
segment 3 of front
518)
with subapical
518) tibial
segment tarsus
tooth and
with 2 rows of 42 spurs 1,1,2.
(in P!) of D. chamaeleo by Benoit (1953b) is not rei iable, because this male specimen MALE:
unknown;
the description of the male allotype
was not obtained by breeding. LOCUS TYPICUS:
Behara
TYPICAL MATERIAL:
(Madagascar)
holotype F!
in P.
778
Mem.
Fig.
HOSTS:
520 -
Amer.
Chela of
Ent.
Inst.,
no.
37,
Dryinus schoutedeni
1984
(Benoit) (type)
unknown
DISTRIBUTION:
Ethiopian region:
1953b);
(Beno i t 1953b ) •
Bek i I Y
MADAGASCAR:
13. Dryinus schoutedeni
Lestodryinus schoutedeni
FEMALE:
fully
winged;
Benoi t
(Benoit)
1950:
n.
P!
Ivondro (Benoit
comb.
224.
length 8,43-9,37 mm;
with clypeus and mandibles light;
Behara,
head reddish or reddish-dark,
antennae black,
with segments 1-2 and
8-10 testaceous; propectus and pronotum brown-reddish or reddi sh; mesothorax, meta thorax and propodeum black; abdomen brown; legs brownblack,
with tarsi
occipital
light;
head dull,
carina complete;
granulated,
posterior ocelli
weakly reticulate rugose;
located on
the imaginary
779
Dryininae
Fig.
521
-
Chela of
Dryinus mayogo
(Benoit) (type)
straight line joining posterior edges of eyes, not touching occipital carina; OPL = OL; OL = POL; TL nearl y twice as long as POL; pronotum (Fig.
519 A)
dull,
granulated,
with numerous keels around disc;
transversal
impression strong;
anterior collar distinct;
transversal
impression very strong;
posterior collar
anterior
posterior
long;
pronotal
780
Mem.
tubercles not reaching numerous
longitudinal
approximately 0,65 dull;
tegulae;
Ins!.,
rugose;
notaul ices
part;
part of radial
segment
with subapical
1 of front
with 2 rows of 34
18
lamellae;
1984
granulated,
hairy,
incomplete,
with
reaching
scutellum as sculptured as scutum, fore-wing
vein
hook;
long
as segment 4;
enlarged claw
segment 5 of front
apex with a
transversal
nearly twice as long as
into a
lamellae;
with 2
with 2 brown
tarsus nearly as
tarsus produced
tooth and
37,
propodeum reticulate rugose,
keels on posterior surface;
segment 3 of front
no.
scutum dull,
weak keels;
bands and with distal proximal
En!.
length of scutum;
metanotum dull,
longitudinal
Amer.
group of 13
(Fig.
tarsus
lamellae;
520)
(Fig. tibial
520) spurs
1,1,2. MALE:
unknown
LOCUS TYP ICUS:
Banana
TYPICAL MATERIAL: HOSTS;
(Zaire)
holotype F!
in TV.
unknown
DISTRIBUTION: Ethiopian region: ZAIRE: Banana ZIMBABWE: Chishawasha (Salisbury), BM!
(Low-Zaire),
TV!
14. Dryinus mayogo (Benoit) n. comb.
Lestodryinus mayogo Benoit 1950: 225.
FEMALE:
fully
winged;
part of vertex
length 8,12-10 mm;
(more along orbits),
sometimes with some small pronotum black, tarsi
light;
with genae,
reddish spots along orbits;
head dull,
abdomen black-brown;
weakly reticulate rugose;
front head
propectus black;
with sides and posterior collar reddish;
metathorax and propodeum black; with
head black,
clypeus and mandibles yellow;
mesothorax, legs brown-black,
occipital
carina
complete; posterior ocelli not touching occipital carina; OPL = OL; OL shorter than POL; TL little shorter than POL (8:9); pronotum (Fig. 519 B) dull, granulated, with numerous striae around disc; anterior transversal impression very weak; anterior collar almost invisible; posterior transversal
impression very strong;
tubercles not reaching some weak 0,65
irregular keels;
length of scutum;
rugose;
tegulae;
notaulices
hairy,
incomplete,
long;
fore-wing
with 2 short
with 2 brown
pronotal
granulated,
reaching
scutell um as scu I ptured as scutum;
propodeum reticulate rugose,
posterior surface;
posterior collar
scutum very
with
metanotum
longitudinal
transversal
dull,
approximately keels on
bands and with
distal part of radial vein nearly twice as long as proximal part; 1 of front tarsus nearly as long as segment 4; segment 3 of front
segment tarsus
produced into a hook; enlarged claw (Fig. 521) with subapical tooth and 19 lamellae; segment 5 of front tarsus (Fig. 521) with 2 rows of 38 lamellae; MALE:
apex with a group of
unknown
14 lamellae;
tibial
spurs 1,1,2.
Dryininae
781
A
B
c Fig.
522 -
Pronotum of females of Dryinus afer
Benoit from Kankunda
(Zaire) (B)
LOCUS TYP I CUS: Pau lis (Za ire) TYPICAL MATE,qIAL: holotype F! HOSTS:
burgeoni
(Benoit) (type) (C).
in TV.
unknown
DISTRIBUTION: LEONE:
(Olmi)(type)(A),
and orophilus
Ethiopian region:
Freetown,
ZAIRE:
Paulis
(High-Uele),
TV!
SIERRA
TW!
15. Dryinus orophilus (Benoit) n. comb.
Lestodryinus orophilus Benoit 1950: 226. Lestodryinus orophilus Benoit: Benoit 1951c: 7.
FEMALE:
fully
winged;
length 3,75-5,31
mm;
head black,
with clypeus and
782
Mem.
Fig.
523 -
mandibles testaceous; 3,
8-10 testaceous;
Amer.
Chela of
abdomen black;
antennae fully
occipital pronotum
(Fig.
around disc,
522 C)
crossed by
1984
testaceous or brown with segments 1with margins and
metathorax and propodeum black;
longitudinal
OPl = Ol;
shiny,
37,
with coxae and clubs of femora brown;
with some
carina complete;
no.
pronotum black,
mesothorax,
legs testaceous,
granulated,
Ins!.,
Dryinus orophilus (Benoit)(type)
propectus black;
posterior collar reddish; head dull,
Ent.
keels on frons and vertex;
Ol = POL;
granulated,
Tl
longer than POL;
with numerous weak striae
a strong posterior transversal
impression and by
a weak anterior transversal impression, posterior collar short, nearly 0,5 times as long as distance from tegulae to pronotal tubercles; anterior collar weak;
pronotal
tubercles not reaching
tegulae;
scutum dull,
granulated, with some irregular short keels on posterior half; notaulices incomplete, reaching almost posterior margin of scutum; scutellum dull, granulated,
metanotum reticulate rugose;
propodeum reticulate rugose,
without longitudinal keels on posterior surface; fore-wing with 3 brown transversal bands and with distal part of radial vein less than twice as long as proximal segment 4
enlarged claw of front tarsus of 10 lamellae; MALE:
part
(21 :15);
fully
(13:9);
segment
segment 3 of front
(Fig.
523)
1 of front
with subapical
tooth and
(Fig. 523) with 2 rows of 24 tibial spurs 1,1,2.
winged;
tarsus
tarsus produced
length 2,81-3,62 mm;
11
lamellae;
head black,
longer than
into a
hook;
lamellae;
segment 5
apex with a
group
with mandibles
Dryininae
o Fig.
524 -
783
0
Heads of females of Dryinus modestus n. sp.
(Olmi) (type) (B)
and burgeoni Benoit from Kankunda
yellow;
antennae yellow-brown;
brown;
legs
yellow,
thorax and propodeum black;
with coxae partly brown;
head dull,
afer
(type) (A), (Zaire) (C).
abdomen
granulated,
weakly reticulate rugose; occipital carina incomplete; OL shorter than OOL; OPL shorter than POL; OL shorter than POL; scutum du II, as scu I ptured as head;
notaulices incomplete,
reaching approximately 0,65 length of scutum;
scutell um and metanotum as scu I ptured as scutum; rugose; radial
fore-wing vein
without dark transversal
longer than proximal
part;
propodeum ret icu I ate
bands and with distal part of
radial
cell
open;
genitalia:
fig.
513 A; tibial spurs 1,1,2. LOCUS TYPICUS: Rutshuru (Zaire) TYPICAL MATERIAL:
holotype F!
designated after the original
HOSTS; unknown DISTRIBUTION: Ethiopian region: National Park,
TV!
Lusinga
in TV;
allotype M!
invalid,
because
description. ZAIRE:
Rutshuru
(Upemba National
(Kivu),
Park),
TV!
TV!
Garamba
Kalinga-Bitshumbi
784
Mem.
Amer.
(Albert National
Park),
GHANA:
(Kumasi),
Kwadaso
OT!
Ile-ife
(W State),
BM!
ZIMBABWE:
Fig.
TV! BM!
Ent.
Pweto OL!
Chishawasha
no.
37,1984
(Mweru Lake,
BM!
Akure
Ins!.,
NIGERIA:
(W State),
(Salisbury),
Shaba)(Benoit 1951c)j
Kabba OL!
BM!
(Lokoja),
BM!
ANGOLA:
Ikenne
CA!
Vila Luso,
525 - Chela of Dryinus modestus n. sp.
Ibadan,
(W State),
(type)
CA!
Dry i n i nae
CAMERON:
Victoria,
785
CAl
16. Dryinus modestus n. sp.
FEMALE:
fully
winged;
partly clypeus yellow; black,
length 5,0 mm;
head black,
antennae testaceous;
with margins and posterior collar reddish;
and propodeum black;
abdomen black-brown;
and clubs of femora brown; occipital carina complete; OPL shorter than OL; granulated,
OL = POL;
impr"ssion strong;
without
reaching
granulated;
longitudinal
flat,
dull,
granulated;
without sculpture;
posterior collar short; scutum dull,
approximately 0,75
metanotum rugose;
carina;
pronotum shiny, pronotal
posterior transversal granulated;
length of scutum;
propodeum reticulate rugose,
keels on posterior surface;
fore-wing
with 3 brown
transversa I bands and wi th di sta I part of radi a I ve in much proximal
metathorax wi th coxae
not touching occipital
TL shorter than POL;
anterior collar weak;
incomplete,
scutellum dull,
tegulae;
pronotum
mesothorax,
legs testaceous,
524 A)
(Fig.
with anterior collar smooth and
tubercles not reaching notaul ices
head
posterior ocelli
with mandibles and
propectus black;
longer than
(11 :5); segment 1 of front tarsus longer than segment 4
part
(12:9); segment 3 of front tarsus produced into a hook; enlarged claw (Fig.
525)
with subapical
tooth and 10 lamellae;
(Fig. 525) with 2 rows of 20 tibial spurs 1,1,2. MALE:
segment 5 of front
apex with a
group of 11
tarsus
lamellae;
unknown
LOCUS TYP I CUS:
Bek i I y
TYPICAL MATERIAL: HOSTS:
lamellae;
(Madagascar)
holotype F!
in TV.
unknown
DISTRIBUTION:
Ethiopian region:
MADAGASCAR:
Bekily,
TV!
17. Dryinus incertus n. sp.
FEMALE:
fully
winged;
length 3,75 mm;
head black,
with clypeus and
mandibles reddish; propectus black; pronotum black, with sides and posterior collar reddish; mesothorax, metathorax and propodeum black; abdomen brown; granulated;
legs testaceous;
occipital
imaginary straight
antennae testaceous;
carina complete;
carina;
shorter than
POL;
pronotum shiny,
around disc;
anterior transversal
posterior transversal pronotal
head shiny, located on
line joining posterior edges of eyes;
touching occipital
distinct;
posterior ocelli
OPL shorter than OL;
impression
granulated,
OL shorter than POL;
not
TL
with numerous weak striae
impression and anterior collar weak;
strong;
tubercles not reaching
reticulate rugose on
the
posterior ocelli
posterior half;
posterior collar short, tegulae;
notaul ices
scutum dull,
incomplete,
but granulated,
reaching
786
Mem.
Fig.
526 -
Amer.
Ent.
Inst.,
no.
37,
1984
Dryinus incertus n. sp. (type)
Chela of
approximately 0,65 length of scutum; scutellum and metanotum granulated; propodeum reticulate rugose without longitudinal keels on posterior surface; radial
fore-wing
vein
much
with 3 brown
tarsus
longer than segment 4
into a
hook;
lamellae;
transversal
longer than proximal
enlarged claw
segment 5 of front
(9:7);
(Fig.
tarsus
MALE: unknown LOCUS TYPICUS:
Grahamstown
TYPICAL MATERIAL: HOSTS:
bands and with distal (7:3);
with subapical
(Fig. tibial
526)
part of
segment 1 of front
segment 3 of front
526)
apex with a group of 13 lamellae;
part
tarsus produced tooth and 11
with 2 rows of 24
lamellae;
spurs 1,1,2.
(South Africa)
holotype F!
in BM.
unknown
DISTRIBUTION:
Ethiopian region:
SOUTH AFRICA:
Grahamstown,
BM!
18. Dryinus burgeoni Benoit
Dryinus burgeoni Benoi t 1953a: 142.
FEMALE: fully winged; length 6,00-8,43 mm; head black, with clypeus and mandibles reddish; sometimes clypeus black; antennae fully testaceous or
Dryininae
Fig.
527 -
Chela of Dryinus burgeoni Benoit
black with segments 8-10 light; brown-reddish; mesothorax,
metathorax and propodeum black; with
tarsi
light;
carina complete;
OPl shorter than Ol; with
propectus black;
(type)
pronotum fully black or
occasionally black with posterior collar brownish;
reddish-brown, occipital
787
(Fig.
posterior ocelli
Ol ;
longitudinal
numerous
head
and anterior collar weak;
POL;
not
pronotum
keels on sides;
abdomen brown;
524 C)
dull,
legs
granulated;
touching occipital
(Fig.
522 B)
dull,
anterior transversal
posterior transversal
carina;
granulated, impression
impression strong;
posterior collar long and distinct; pronotal tubercles not reaching tegulae; scutum dull, granulated, fully sculptured by numerous longitudinal keels; notaulices almost complete,
reaching almost posterior margin of scutum;
scutellum and metanotum dull, reticulate rugose, wing
without
with 2 brown
curv iii near, nearly as
much
with numerous
longitudinal
transversal
bands and with distal
longer than prox i ma I part;
long as segment 4;
longitudinal
keels;
propodeum
keels on posterior surface; segment 1 of front
segment 3 of front
fore-
part of radial
tarsus produced
vein
tarsus into a
hook; enlarged claw (Fig. 527) with subapical tooth and 13 lamellae; segment 5 of front tarsus (Fig. 527) with 2 rows of 37 lamellae; apex with a group of 10 lamellae; MALE: unknown lOCUS TYP ICUS:
holotype F!
in TV.
unknown
DISTRIBUTION: CAMERON: TV!
spurs 1,1,2.
Moto (Zaire)
TYPICAL MATERIAL: HOSTS:
tibial
Moto
Ethiopian region:
Nkoemvon, (I turi),
Ol!
BM!
NIGERIA:
ZAIRE:
Ibadan,
Kankunda
BM!
Ile-Ife,
(Upemba National
TV!
19. Dryinus afer (Olmi) n. comb.
TW! Park),
788
Mem.
Fig.
528 -
Amer.
Ent.
Inst.,
no.
37,
Female of Dryinus afer
1984
(Olmi)(type)
Paradryinus afer Olmi 1975: 221.
FEMALE
(Figs 528,
529):
fully
winged;
length 5,02 mm;
head black,
with
Dryininae
Fig.
529 -
front part of vertex antennae brown,
789
Female of Dryinus afer
(more along orbits),
clypeus and mandibles yellow;
with segments 1-2 yellow;
posterior collar reddish; abdomen brown;
mesothorax,
legs brown,
(Olmi)(type)
pronotum black,
with sides and
metatorax and propodeum black;
with trochanters and part of fore coxae
yellow; head (Fig. 524 B) dull, granulated; occipital carina complete; posterior ocell i located in front of the imaginary straight line joining posterior edges of eyes; pronotum crossed by 2 transversal impressions (anterior impression very weak; posterior impression strong); pronotum (Fig.
522 A)
dull,
granulated,
and on posterior transversal nearly as
with numerous
impression;
long as distance from
tegulae to pronotal
tubercles not reaching tegulae; longitudinal
longitudinal
keels on sides
posterior collar of pronotum
scutum dull,
short keels on posterior third;
tubercles;
granulated,
with
pronotal numerous
tracks of incomplete notaulices
(reaching approximately 0,5 length of scutum) are visible; scutellum and metanotum dull, granulated and reticulate rugose; propodeum reticulate rugose,
without
longitudinal
keels on posterior surface;
fore-wing
with 2
brown transversal bands and with distal part of radial vein rectilinear, much longer than proximal part (18:8); enlarged claw (Fig. 530) with subapical
tooth and 14 lamellae;
2 rows of nearly 38 of front MALE
tarsus nearly as
(Figs 531,
532):
mandibles yellow; abdomen brown; tarsi
light;
frons flat antennal
lamellae;
segment 5 of front
long as segment 4;
fully
winged;
legs brown,
with tibiae
scutum dull,
invisible;
reticulate rugose,
530)
with
segment 1
spurs 1,1,2. head black,
(except for the hind tibiae)
fully reticulate rugose;
temples
(Fig.
with
thorax and propodeum black;
in front of the anterior ocellus, sockets;
tibial
length 2,71-3,07 mm;
antennae yellow-brown;
head dull,
tarsus
apex with a group of 13 lamellae;
occipital
carina
only convex between
and
incomplete; the
POL = 6; Ol = 3; OOl = 4; OPl = 1;
hairy;
notaul ices weakly visible,
790
Mem.
Amer.
Ent.
Ins! .,
no. 37, 1984
Dryininae
Fig.
incomplete,
-
Male of
Dryinus afer (Olmi) (allotype)
reaching approximately 0,5 length of scutum;
metanotum dull, wing without much
531
791
granulated,
punctate;
dark transversal
longer than proximal
tibial spurs 1,1,2. LOCUS TYP ICUS: Canavial
bands and with distal
part;
radial
cell
closed;
part of radial genitalia:
(Sao Tome)
TYPICAL MATERIAL:
holotype F!
HOSTS:
Ricania insularis lall. (Olmi 1975)
in Sao Tome
nISTRIBUTION:
Ethiopian reclion:
and 3 paratypes MM! SAO TOME':
OR lENT AL SPEC I ES OF
Fifteen
scutellum and
propodeum reticulate rugose;
Canavial.
DI2YINUS
species are known.
KEY TO THE SPECIES
in OL. Ol!
fig.
forevein 533;
792
Mem.
Fig.
Amer.
Ent.
Inst.,
no.
37,
1984
532 - Male of Dryinus afer (Olmi)(allotype)
FEMALES
i
\~
Posterior excavated
collar of pronotum invisible (Fig. 534 A); pronotum (Fig. 534 A) ••••••••••• 1. irregularis n. sp.
Posterior collar of more
2
or
Segment
1 of
long
segment
as
Segment
pronotum visible
less humped,
1 of
front
rarely
flat
tarsus nearl y
534 B); pronotum 534 8) •••.•••.••••••••• 2
(Fig.
(Fig.
twi ce or more
than
twi ce as
4 •••.••••••••••••••.•••••••••••..••••••••••••• 3
front
tarsus
slightly
longer
than
segment
4 or
shorter ••••••••••••••••••••••••••••••••••••••••••••••••••••••• 4
3
Posteri or ocell i black;
fore
touching
coxae
shorter
occipi tal than
carina;
head;
prothorax
fore
trochanters kiefferi (F ou t s )
less
than
three times as long as broad . . . . . 2. Posteri or ocell i not touching occipi tal carina; prothorax redd ish; fore coxae nearl y twi ce as long as head; fore trochanters nearly
five
times as
. . . . . . • . . . . . . . . . . . . . . . . . • . . . . . . .•
4
Posterior ocell i (Fig.
541
touching
long
3.
or almost
as broad •••••••••••••••••
longipes n. touching
sp.
occipi tal
carina
A) ••••••••••••••••••••• ~ •••••••••••••••••••••••••••• 5
Posterior ocelli
farther
from occipital
carina
(Fig.
541
B) . . . .
••••••• •••••••••••• ••••••••••••••••••• ••••••••••••••••••••••• 10
5
Head and pronotum fully
testaceous •••••••••••••..••••••••••••••
• • • • • • . . . . . • • . • • • • • • . . . . . • • • . . . .. 6
4.
pulcher n.
sp.
Head and pronotum mostly black . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6 Head shiny, smooth, hairless, without sculpture, very excavated
Dryininae
Fig.
8
-
Male genitalia of
Dryinus afer (Olmi)(allotype)
5.
sculpture,
shiny
or dull,
granulated,
never without usually
flat,
sometimes
or without
krombeini Ponomarenko
excavated •••••••••••••••••••
Head
f I at ••••••••••••••.•••••••••••...••.••••..••...•••••••.•• 8
Head with
longitudinal
keels
or
Head without
6.
with
excavated ••••••••.• 7
Head
tracks
• •••..••••••.••••.••••••••••..•••
9
exilis n. sp.
• • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • •• Head
keels, 7
533
793
7.
of
longitudinal
keels •••
stantoni Ashmead
longitudinal keels or tracks of longitudinal keels • ., ••••••••••••••••••.•.•• , •••••..•••••.•••.•••••••.•••..•••.• 9 Head longer, with sides of frons more converging (Fig 543 A). • •••••••••••••••••••••••••••••••• 8. trifasciatus K i ef f er
794
Mem.
Head
transverse,
Amer.
wi th
En!.
sides
Inst.,
of
frons
B) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 10 Species full:,Species at
testaceous,
least
less
long as
segment
dorsal
surface; times as
slender,
10.
surface more
antennae more long as
543
3
segment
less
twi ce as
six
long as
3 more 2 ........................ . with
segment
1 1. praeciarus n. 12.
than
indicus (Kieffer)
than
segment
fully black,
long as posterior
slender,
reddish . . . . . . . . . .
Pronotum almost
as
with
2 ......
• • • • • • • . • • • • • • • • • • • . . . . • • • • • . . .. 12 Pronotum fully
(F i g.
converging
• .12
surface nearly
antennae
eight
less
latus n. sp. .. . . . . . . . 11
times as
than
1984
with petiole black.
surface;
Propodeum wi th
9.
37,
partly black ••••••••••••••••
11 Pronotum with dorsal
posterior
no.
sp.
fulvus n. sp.
sometimes partly
reddish . . . . . . . 13
13 Metapleura wi th a narrow region smooth, shiny, hairless, wi thout sculpture ••••••••••••••• 13. lucens n. sp. Metapleura wi thout without
narrow region
smooth,
shiny,
hairless and
sculpture •••••••••••••••••••••.••••••..•••••••••••••• 14
14 Antennae wi th segments 4-5 not broader
than
other
segmen t s
cyl indri c,
(F i g.
but
flat,
much
551) . . . . . . . . . . . . . . . . . . . . . . . ..
14. browni Ashmead· Antennae
with
segments 4-5 as
cylindric as
other
segments,
not
broader than other segments . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15 15 Dorsal surface of propodeum nearly twice as long as posterior surface ••••••.••••••••••••••••••• 7. Dorsal surface of propodeum sl ightly
stantoni Ashmead
15.
lankanus n. sp.
surface •••••••••••••••••••••••••
longer
than posterior
MALES
Dorsal
part
of
occipi tal
carina
laterally
touching
eyes . . . . . . .
10. indicus (Kieffer) Dorsal
part
of
occipital
carina
laterally not touching 14. browni Ashmead
eyes . . . .
The males of the other species are unknown.
1. Dryinus irregular-is n. sp.
FEMALE: fully winged; length 5,00 mm; head fJlack, with mandibles yellow; antennae brown, with segments 1-2 and partly 3 yellow; propectus black; pronotum black, mesothorax,
with sides reddish and posterior margin
metathorax and propodeum black;
weakly reddish;
abdomen brown;
legs
795
Dryininae
A
Fig.
(laterally viewed) of females of Dryinus irregularis n. and st2ntoni Ashmead from Sandakan (Sabah) (B)
534 - Pronotum sp. (type) (A)
testaceous,
with mid and
antennae short
than
hind coxae black;
body,
with segment 3
head shiny,
reticulate rugose;
longer than segment 4;
occipital
carina complete; pronotum (Fig. 534 A) with a median transversal impression, shiny, with deep punctures; anterior surface and sides without sculpture, fully 0,5
smooth;
reticulate rugose;
length of scutum;
pronotal
tubercles not reaching
notaulices almost
invisible,
tegulae;
scutum
approximately reaching
scutellum and metanotum reticulate rugose;
propodeum reticulate rugose,
with 2
longitudinal
keels on
posterior
surface; fore-w ing without dark transversa I bands, with dista I part of radial vein nearly as long as proximal part; segment 1 of front tarsus twice as long as segment 4; segment 3 produced into a hook; enlarged claw with subapical tooth and 6 lamellae (Fig. 535); segment 5 of front tarsus
(Fig.
group of 6 MALE:
535)
with 2 rows of 3
lamellae;
(proximal)
+ 11
lamellae;
spurs 1,1,2.
unknown
LOCUS TYPICUS:
Chiangdao
TYPICAL MATERIAL: HOSTS:
tibial
(Thailand)
holotype F!
in B.
unknown
DISTRIBUTION:
Oriental
region:
THAILAND:
Chiangdao,
2. Dryinus kiefferi (Fouts)
n.
8!
comb.
apex with a
796
Mem.
Fig.
535 -
Amer.
Chela of
Ent.
Inst.,
no.
37,
1984
Dryinus irregularis n. sp. (type)
Lestodryinus kiefferi Fouts 1922: 630. Lestodryinus kiefferi Fouts: Ba I tazar 1966: 203.
FEMALE:
fully
testaceous;
winged;
propodeum black; and tarsi
length 6,87 mm;
antennae brown, light;
head black,
with mandibles
with segments 8-10 yellow;
abdomen black-reddish; head hairy,
flat,
dull,
legs brown, granulated,
thorax and with coxae black
fully sculptured by
numerous longitudinal keels; occipital carina incomplete, visible only behind ocelli and on sides of vertex, where the occipital carina is touching eyes; on sides of occiput the occipital carina is invisible; occipital carina visible on ventral face of head; temples invisible; pronotum dull, hairy, granulated, with some keels on sides; pronotal tubercles not reaching tegulae; transversal
impression strong;
posterior collar short, notaulices complete, dull,
almost invisible among keels;
reticulate rugose;
into 3 regions;
posterior
impression
weak;
longitudinal
keels;
scutellum and metanotum
propodeum reticulate rugose,
fore-wing vein
humped;
fully sculptured by numerous
keels on posterior surface; part of radial
disc not very
anterior transversal
with 2 longitudinal
posterior half of posterior surface is so divided
with 2 brown
transversal
longer than proximal part;
bands and with distal
fore coxae shorter than
head; fore trochanters less than three times as long as broad; segment of front tarsus nearly twice as long as segment 4; segment 3 of front tarsus produced into a hook;
enlarged claw
(Fig.
536)
spatulate,
with
797
Dryininae
Fig.
subapical 536)
535 -
Chela of Dryinus kiefferi
tooth and 13 very
long
with 2 rows of 39 lamellae
other lamellae)
lamellae;
(Fouts)(type)
segemnt 5 of front
(among which 5 lamellae much
extending continuously
to the apex;
tibial
tarsus
(Fig.
longer than
spurs 1.1,2.
798
Mem. Amer.
Fig.
537 -
Ent.
Inst.,
no.
37,
1984
Chela of Dryinus longipes n. sp.
MALE: unknown lOCUS TYPICUS: Mt. Makiling (Philippines) TYP ICAl MATER IAl: holotype F! in WA. HOSTS: unknown DISTRIBUTION: Oriental region: PHiliPPINES: Mt.
(type)
Makiling
(luzon),
WA!
Dryininae
NOTES:
for the morphology of the chela
of the enlarged claw very
long;
lamellae
lamellae)
Moczar.
longer than other The tibial
799
(enlarged claw spatulate;
segment 5 of front
lamellae
tarsus with some
this species is near Nichardsidryinus
spurs however are 1,1,2.
3. Dryinus longipes n. sp.
FEMALE:
fully
winged;
length 6,43 mm;
mand ib I es testaceous; darkened;
prothorax redd i sh;
abdomen black, trochanters,
head black,
antennae testaceous, mesothorax,
meta thorax and propodeum black;
with some regions testaceous;
clubs of fore femora,
with clypeus and
with segments 3 and 4 legs testaceous,
with fore
stalks of mid and hind femora,
distal
apexes of tibiae brown; antennae distally thickened; antenna I segments in following proportions: 14: 7: 49: 30: 26: 19: 15: 11: 9: 9; antenna I segment 4 nearly fifteen
times as long as broad
reticulate rugose and with keel; = 5;
occipital
irregular keels;
carina complete;
pronotum shiny,
POL = 3,5;
without sculpture on
(30:2);
frons
head shiny,
with a median complete
OL = 3;
OOL = 12;
OPL = 3;
reticulate rugose and punctate on sides and on posterior collar; with strong long;
anterior and posterior transversal
pronotal
tubercles not reaching
TL
anterior collar and on disc, impressions;
tegulae;
scutum,
pronotum
posterior collar
scutellum and
metanotum reticulate rugose; notaulices incomplete, reaching approximately 0,65 length of scutum; propodeum reticulate rugose, without transversal keels, with two incomplete longitudinal keels on posterior surface; forewing with two dark transversa I bands, one on the basa I cell s and another beneath part 28;
the pterostigma
(14:11);
segment 3 of front
and slender,
with a subapical
tarsus 7
(Fig.
lamellae;
distal
part of radial
537) tibial
MALE: unknown LOCUS TYPICUS:
vein
longer than proximal
segments in following proportions:
tarsus produced
into a
with coxae nearly twice as
trochanters nearly five 537)
;
fore tarsal
times as
hook;
fore
long as head
long as broad
(30:6);
tooth and with 12 lamellae;
with 2 rows of 25 lamellae;
46:
4:
legs very
(50:25)
Nilgira Hills
and with
HOSTS: unknown DISTRIBUTION: Oriental
region:
apex with a
group of nearly
in OT. INDIA:
(Fig.
segment 5 of front
(India)
holotype F!
18:
long
enlarged claw
spurs 1,1,2.
TYPICAL MATERIAL:
7:
Nilgira Hills
(S
India),
4. Dryinus pulcher n. sp.
Dryinus formicarius Latreille: Ponomarenko 1981b: 879.
OT!
800
Mem. Amer. Ent.
Ins!.,
no. 37,
1984
Fig. 538 - Chela of Dryinus pulcher n. sp.
(type)
FEMALE: fully winged;length 3,25-5,00 mm; head, antennae, propectus a'lG propodeum fully testaceous; mesothorax; metathorax and propodeum black; abdomen brown-testaceous; legs testaceous; head shiny, weakly granulated; occipital carina complete; posterior ocelli almost touching occipital carina;
Dryininae
Fig.
539 -
Chela of
OL slightly
longer than POL;
granu I a ted;
anterior transversa I
posterior transversal pronotal
801
Dryinus exilis n.
temples prominent;
and with some irregular keels;
(type)
pronotum shiny,
impress ion and anterior coil ar weak;
impression strong;
tubercles not reaching
sp.
tegulae;
posterior collar short, scutum dull,
notaulices incomplete,
hairy,
distinct;
granulated
reaching
802
Mem.
approximately 0,5 granulated and short
Amer.
Ent.
Ins!.,
length of scutum;
with
longitudinal
no.
37,
1984
scutellum and metanotum dull,
irregular keels;
propodeum reticulate rugose,
keels on posterior surface;
fore-wing
transversa I bands and wi th di sta I part of rad i a I vei n much proximal
part;
segment
1 of front
tarsus as
with 2
with 2 brown longer than
long as segment 4;
segment 3
of front tarsus produced into a hook; enlarged claw (Fig. 538) with subapical tooth and 9 lamellae; segment 5 of front tarsus (Fig. 538) 2 rows of 18 1,1,2. MALE:
lamellae;
apex with a
group of 6 lamellae;
tibial
with
spurs
unknown
LOCUS TYPICUS:
Labugama
TYPICAL MATERIAL: HOSTS:
(Sri
Lanka)
holotype F!
in BM;
2 paratypes FF!
in WA.
unknown
DISTRIBUTION:
Oriental
Gilimale
Dist.)(Ponomarenko 1981b:
(Rat.
Latrei lie), IVA!
region:
SRI
LANKA:
Udawattakele Sanc!.
Labugama
under name
(Kandy,
Kan.
(Colombo),
BM!
Dryinus formicarius
Dis!.),
WA!
5. Dryinus exilis n. sp.
FEMALE:
fully
winged;
length 3,75 mm;
clypeus and mandibles yellow; brown; shiny,
legs yellow, smooth,
head black,
with clubs of femora and
excavated,
with malar space,
thorax and propodeum black;
without
sculpture,
abdomen
hind coxae brown; with
temples
very
head prominent;
occipital carina complete; posterior ocell i touching occipital carina; pronotum shiny, smooth, without sculpture, with longitudinal keels on sides;
pronotum crossed by a posterior transversal
tubercles not reaching sculpture,
smooth;
weakly granulated; distance between
tegulae;
lateral
scutum shiny,
region
impression;
with central
without sculpture,
notaulices complete,
smooth;
smooth,
posteriorly separated; without sculpture;
without
lateral
the notaulices longer than breadth of ocelli
scutellum and metanotum shiny,
pronotal
region
regions
minimum (6:3);
propodeum
reticulate rugose, without longitudinal keels on posterior surface; forewing with 3 brown transversal bands and with distal part of radial vein much longer than proxi ma I part; segment 1 of front tarsus s light I y longer than segment 4; enlarged claw with subapical tooth and 12 lamellae (Fig. 539);
segment 5 of front
apex with a MALE:
group of 6
(Fig.
539)
tibial
with 2 rows of 28
lamellae;
spurs 1,1,2.
unknown
LOCUS TYPICUS:
IVapikhamthong
TYPICAL MATERIAL: HOSTS:
tarsus
lamellae;
holotype F!
(Laos) in B.
unknown
DISTRIBUTION:
Oriental
region:
LAOS:
Wapikhamthong
(Wapi),
6. Dryinus krombeini Ponomarenko
B!
Dryininae
Fig.
540 -
Chela of
803
Dryinus krombeini Ponomarenko from Culion I. (Philippines)
Dryinus krombeini Ponomarenko 1981 b: 879.
FEMALE: vertex,
fully
winged;
malar space,
length 4,37-6,00 mm;
head black,
clypeus aC'd mandibles yellow;
thorax and propodeum black,
with front part of
antennae testaceous;
sometimes with posterior collar of pronotum
reddish; abdomen brown-reddish; legs reddish-testaceous, with coxae, clubs of femora and tibiae brownish; head excavated, dull, granulated, with some
longitudinal
keels on frons;
occipital
carina complete;
posterior
ocell i almost touching occipital carina; pronotum dull, granulated, with numerous keels around disc; pronotal tubercles not reaching tegulae; scutum granulated, areolae on central
with numerous areolae on region
lateral
near posterior margin;
regions
lateral
and with some
regions usually
804
Mem.
Fig.
541
-
Amer.
Ent.
and fulvus
with areolae on posterior half; metanotum granulated;
37,
part of radial tarsus slightly
n.
sp.
1984
(type) (8)
notaulice almost complete;
propodeum reticulate rugose,
keels on posterior surface; front
no.
Heads of females of Dryinus stantoni Ashmead from Sandakan (Sabah) (A)
distal
Ins!.,
fore-wing
vein much
with 3 brown
longitudinal
transversal
longer than proximal
longer than segment 4;
scutellum and
without part;
bands;
segment 1 of
enlarged claw with subapical
tooth and 13 lamellae (Fig. 540); segment 5 of front tarsus (Fig. 540) 2 rows of 31 lamellae; apex with a group of 11 lamellae; tibial spurs 1,1,2. MALE:
unknown
LOCUS TYPICUS:
Udawattakele
TYPICAL MATERIAL: HOSTS:
with
(Sri
holotype F!
Lanka)
in WA.
unknown
DISTRIBUTION:
Oriental
region:
VIET NAM:
Phan Rang,
B!
LAOS:
Ban Van
Eue (Vientiane), OL! B! Wapikhamthong (Wapi), B! THAILAND: Mae Sa (Chiang Mai), B! Konthathan waterfall area (Doi Suthep-Pui natn. Park), CO! PHILIPPINES: Dis!.), WA!
Culion
Island,
OL!
SRI
LANKA:
Udawattakele
(Kan.
Dryininae
Fig.
805
542 - Chela of Dryinus stan toni Ashmead from Forest Camp (Sabah)
806
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
7. Dryinus stan toni Ashmead
Dryinus stan toni Ashmead 1904: 134. Prodryinus stantoni (Ashmead): Fouts 1922: 629. Paradryinus javanus R.C.L. Perkins 1912: 11 (n. syn.); locus typicus Pekalongan (Java). Lestodryinus luzonicus Kieffer 1913a: 189 (n. syn.); locus typicus Mt.
Maki I ing
(Phi I ippine
Islands).
Lestodryinus stan toni (Ashmead): Fouts 1922: 629. Neodryinus stantoni (Ashmead): Baltazar 1966: 204. Dryinus szepligettii Kieffer partim: Ponomarenko 1981 b: 879.
FEMALE:
fully
winged;
length 3,12-5,62 mm;
head black,
with malar space,
clypeus and mandibles yellow; antennae testaceous, sometimes brownish; rarely with last segment whitish; pronotum black, with sides and posterior collar reddish;
sometimes posterior collar black;
meta thorax black; A)
dull,
abdomen brown-reddish;
granulated,
complete;
with some longitudinal
posterior ocell i
touching occipital
mesothorax and
legs testaceous; keels;
located very near occipital
carina);
head
occipital
(Fig.
541
carina
carina
(but not
pronotum crossed by 2 transversal
impressions
(anterior impression weak; posterior impression strong); pronotum (Fig. B) du II, ha i ry, granu I a ted; pronota I tuberc I es not reach i ng tegu I ae; scutum
dull,
granulated,
hairy;
notaul ices
incomplete,
534
reaching
approx i matel yO, 5 length of scutum; scute II um and metanotum as scu I ptured as scutum; propodeum rugose, without longitudinal keels on posterior surface; radial
fore-w i ng
vein
much
with 3 brown
slightly
longer than segment 4;
lamellae
(Fig.
lamellae;
transversa I bands and with di sta I part of
longer than proximal
542);
part;
segment
1 of front
enlarged claw with subapical
segment 5 of front
tarsus
apex with a group of 14 lamellae;
(Fig.
tibial
542)
tarsus
tooth and 11
with 2 rows of 27
spurs 1,1,2.
MALE: unknown LOCUS TYPICUS: Manila (Philippines) TYPICAL ,'vlATERIAL: lectotype F! and 1 para lectotype F!
(designated by
javanus in B; holotype F! of D. stantoni in WA; holotype F! of L. luzonicus in WA. HOSTS: in Java Thanatodictya sp. (Dictyopharidae) (R.C.L. Perkins 1912). PARASITES: in Java probably Cheiloneurus javanus R.C.L. Perkins (R.C.L.
Olmi
1982) of P.
Perkins 1912). DISTRIBUTION: Oriental region: INDONESIA: Pekalongan (Java), B! PHILIPPINES: Manila (Luzon), WA! La Trinidad (Luzon), B! Mt. Polis (Luzon),
MD!
Nueva Vizcaya
(Imugan,
Luzon),
B!
Mt.
Makiling
(Luzon),
WA! Zamboanga (Mindanao), B! Wa-wa Dam (Rizal, Mt. Montalban), B! Pinigisan (Mantalingajan, Palawan), CO! Tagembung (Mantalingajan, Palawan), TAIWAN:
CO!
THAILAND:
Klapan,
8!
Chiangdao
VIET NAM:
(Chiangmai),
Pleiku,
B!
Kontum
OL!
B!
Banna,
(Pleiku),
B!
B!
M'Drak
Dryininae
Fig.
543 -
Heads of females of JJryinus
Wapikhamthong
(Ban Me Thuot),
B!
LAOS:
Sayaboury,
Sion,
B!
MALAYSIA:
8M!
Forest Camp
(Kalabakan,
(Sabah),
Kanneliya Jungle (Induruwa Jungle,
OL!
8!
Phou-kow-kuei,
Port Dickson
Sabah),
B!
Singkor Sabah),
(Gal.
Dist.),
Rat.
from
Ban Van Eue (Vientiane),
B!
Vientiane),
trifasciatus Kieffer from
(Laos)(A) and latus n. sp. (Phi I ippines) (B).
(Phou-kow-kuei),
Sandakan
807
Dist.),
WA!
(Negeri SRI
Lahugala
WA!
B! 8!
LANKA:
(Amp.
HONG KONG:
Ban Van
Heue
De Tha Ngone
Sambi lan,
Gunong Mulu B!
Irawan Riv.
(Sarawak), Kandy,
Dist.),
Taipokau,
WA!
(Gi
Malaya), OL!
B~1!
BM! Gilimale
B!
8. JJryinus trifasciatus Kieffer
Dryinus trifasciatus Kieffer 1906b: 335.
FEMALE:
fully
winged;
length 3,75-4,37 mm;
head black,
with clypeus and
mandibles reddish; antennae testaceous; pronotum black, with sides and posterior collar reddish; mesothorax, meta thorax and propodeum black; abdomen brown, legs fully testaceous or with clubs of fore femora and hind coxae brown;
head
(Fig.
543 A)
flat,
shiny,
smooth,
weakly
808
Fig.
Mem. Amer.
544 -
Ins!.,
no.
37,
1984
Chela of Dryinus trifasciatus Kieffer from Wapikhamthong
granulated; ocelli
Ent.
sides of frons converging;
touching occipital
carina;
occipital
carina complete;
impression,
with anterior collar smooth and without sculpture; pronotal
granulated;
notaul ices more or less visible,
propodeum
surface; radial
tubercles not reaching
reticulate
rugose,
544);
without
with 3 brown
extending continuously MALE: unknown
part;
tarsus
to the apex;
Matheran
region:
MALAYSIA:
(Kalabakan,
(Maharashtra),
BU!
tibial
LAOS:
part of
tarsus sl ightly
tooth and 6 lamellae
with 2 rows of 37 lamellae
spurs 1,1,2.
Wapikhamthong
Sabah),
Cheramgode (Nilgiri
Pinigisan
544)
and 1 para lectotype F!
HOSTS: unknown DISTRIBUTION: Oriental
PHILIPPINES:
segment 1 of front
(Fig.
posterior
(India)
TYPICAL MATERIAL: lectotype F! Moczar 1965) in BU.
Forest Camp
keels on
bands and with distal
enlarged claw with subapical
segment 5 of front
LOCUS TYPICUS:
disc weakly
scutum dull,
reaching approximately 0,5
longitudinal
transversal
longer than proximal
longer than segment 4; (Fig.
tegulae;
scutell um and metanotum as scu I ptured as scutum;
fore-wing
vein
posterior
pronotum with a posterior transversal
granulated;
length of scutum;
(Laos)
(Mantalingajan,
B!
Hills,
(Wapi),
INDIA: S.
Palawan),
(designated by
OLl
Matheran
India),
TW!
CO!
9. Dryinus latus n. sp.
FEMALE:
fully
winged;
length 2,68-3,12 mm;
head black,
with clypeus and
Dryininae
Fig.
545 -
Chela of Dryinus latus n. sp.
809
(type)
810
Mem.
mandibles yellow;
antennae fully
and 10 testaceous;
Amer.
Ent.
Ins!.,
no.
37,
1984
testaceous or brown with segments 1-2
pronotum black,
with sides and posterior collar
redd i sh; mesothorax, meta thorax and propodeum black; abdomen brown; legs fully testaceous or brown with tarsi light; head (Fig. 543 B) dull, weakly excavated,
granulated;
touching occipital around disc;
carina;
pronotal
occipital
carina complete;
pronotum shiny,
granulated,
tubercles not reaching
tegulae;
posterior ocelli
with numerous keels scutum dull,
hairy,
granulated, with tracks of keels on posterior half; notaul ices incomplete, reaching approximately 0,5 length of scutum; scutellum as sculptured as scutum;
metanotum rugose;
longitudinal bands and
propodeum reticulate rugose,
keels on posterior surface; with distal
part of radial
fore-wing
vein much
without
with 3 brown
transversal
longer than proximal
part; segment 1 of front tarsus sl ightly longer than segment 4; enlarged claw (Fig. 545) with subapical tooth and 9 lamellae; segment 5 of front tarsus
(Fig.
545)
with 2 rows of 14 lamellae;
lamellae; tibial MALE: unknown
spurs 1,1,2.
LOCUS TYP ICUS:
Zamboanga
TYPICAL MATERIAL: FF! in OL. HOSTS:
apex with a group of 7
(Phi I ippines)
holotype F!
and 2 paratypes FF!
in B;
2 paratypes
unknown
DISTRIBUTION: Irawan Riv.
Oriental
(Mt.
Wapikhamthong
region:
Beaufort,
(Wapi),
B!
PHILIPPINES:
Palawan), VIET NAM:
OL!
Zamboanga
THAILAND:
Dalat,
(Mindanao),
Banna,
B!
B!
LAOS:
OL!
10. Dryinus indicus C
Nesodryinus indicus Kieffer 1914c: 311. Mesodryinus indicus (Kieffer): Pruthi and Mani 1942: 425. Chlorodryinus pallidus R.C.L. Perkins: Pruthi and Mani 1942: 425. i\,'esodryinus indicus Kieffer: Kurian 1954a: 256. Chlorodryinus pallidus R.C.L. Perkins: Kurian 1954a: 257. Mesodryinus indicus Kieffer: Kelner-Pillault 1958: 146.
FEMALE:
fully
antennal
segments
8:
11;
winged;
length 3,56-5,00 mm;
in following proportions:
testaceous, 11:
5:
41:
proportions of antenna I segments 2-3 different
specimens
(in small
granulated;
on sides of vertex; without sculpture,
impression; distinct;
specimens 5:22;
occipital carina POL
13:
10:
10:
and big
head dull,
visible only behind the ocelli
and
= 3; OL = 3; OOL = 7; OPL = 1,5; pronotum shiny,
smooth,
anterior collar
21:
in small
in big specimens 5:41);
incomplete,
crossed by a posterior strong
anterior transversal
scutum shiny,
with petiole black; 25:
weak;
impression weak; pranatal
strongly punctate;
tubercles
transversal
posterior collar short, not
reaching
notaulices reaching 0,5
tegulae;
length of scutum
Dryininae
Fig.
546 -
Chela of
(posterior half of scutum punctate;
longitudinal
Dryinus indicus (Kieffer)(type)
not visible,
metanotum rugose,
811
du II;
hidden by pin);
scutellum shiny,
propodeum reticu I a te rugose,
keels on posterior surface;
fore-wing
without dark
wi thout transversal
812
Mem.
Fig.
547 -
Amer.
Male genitalia of
En!.
Ins!.,
no.
37,1984
Dryinus indicus (Kieffer) from Pusa (India): left half removed.
bands,
with distal
segment front
1 of front
tarsus produced
subapical
tooth and 9
2 rows of 16 1,1,2. MALE:
part of radial tarsus
fully
lamellae; winged;
vein much
longer than proximal
longer than segment 4
into a
hook;
lamellae;
(15: 12);
enlarged claw
segment 5 of front
apex with a group of length 2,20-3,60 mm;
head with anterior part of frons,
testaceous;
antennae not distally
546)
with
tarsus
(Fig.
546)
lamellae;
tibial
spurs
black;
testaceous;
thickened;
12
(Fig.
part;
segment 3 of with
legs and antennae
clypeus and mandibles antennal
segments
in following
Dryininae
Fig.
548 -
Chela of
Dryinus praeclarus n. sp. (type)
813
814
Mem.
proportions:
7:
7:
19:
Amer.
14:
En!.
14:
14:
Ins!.,
14:
frons with a complete median keel; part
laterally reaching
eyes;
scutellum and metanotum dull, transversal
transversal
bands;
or
HOSTS:
in
10:
11;
head dull,
carina
OL = 4;
granulated;
granulated;
incomplete,
OOL = 4;
with dorsal
OPL = 1,5;
holotype F!
propodeum reticulate rugose,
keels;
part of radial
(17:10); genitalia: fig. 547; tibial LOCUS TYPICUS: Pusa (India) TYPICAL MATERIAL:
1984
granulated, with posterior third rugose; approximately 0,65 length of scutum;
longitudinal
distal
37,
occipital
POL = 7;
temples invisible; scutum dull, notaul ices incomplete, reaching without
12:
no.
fore-wing vein
hyal ine,
without dark
longer than proximal
part
spurs 1,1,2.
and 4 paratypes FF!
in P.
Phromnia marginella (Kieffer 1914b); Pyrilla sp. (I(urian
India
1954a) . DISTRIBUTION:
Oriental
region:
Pusa
(Bihar),
P!
ND!
11. Dryinus praeclarus n. sp.
FEMALE:
fully
winged;
antennae slender, proportions: weakly
10:
length 7 mm;
distally
5:
45:
granulated;
30:
and on with a
shiny,
occipital
flat,
10:
8:
10;
in
and without sculpture;
carina
incomplete,
POL =2;
following
head flat,
shiny,
frons with a
visible behind the ocell i
OL = 4,5;
OOL = 8,5;
transversal
posterior collar very short;
pronotum
without sculpture,
keels on
impression;
incomplete,
weak
visible only
and metanotum sh i ny,
punctate,
tubercles not reaching tracks of areolae;
and
except for
tegulae;
near anterior mar"in of scutum;
propodeum reticulate rugose,
longitudinal
dorsal
scutum
notaulices weakly
without scu I pture among
weakly alutaceous; keels;
OPL = 2,5;
impression
sides and for some short keels on posterior
pronotal
granulated and with
visible,
a
11:
posterior
longitudinal
with
with petiole black;
antenna I segment
weak
transversal dull,
15:
sides of posterior ocelli;
pronotum
some
26:
temples long
median complete keel;
testaceous,
thickened;
scutellum
punctures or
without transversal
surface of propodeum more than
twice as
or long as
posterior surface (27: 10); fore-wing hyal ine, without dark transversal bands; distal part of radial vein longer than proximal part (26:9); fore tarsal (Fig. tarsus
segments 548)
(Fig.
lamellae;
in following proportions:
with a 548)
tibial
MALE: unknown LOCUS TYPICUS:
subapical
tooth and
with 2 rows of 21
12
32:
3,5:
lamellae;
lamellae;
5:
20:
26;
enlarged claw
segment 5 of front
apex with a group of 7
spurs 1,1,2. Los Banos
TYPICAL MATERIAL:
(Philippines)
holotype F!
HOSTS: unknown DISTRIBUTION: Oriental
region:
in TW. PHILIPPINES:
Los Banos
(Luzon),
TW!
Dryininae
815
/r~
I
!
/cf '
cf=
t
\ \
Fig.
549 -
Chela of
Dryinus fulvus n. sp. (type)
12. Dryinus fulvus n. sp.
FEMALE: vertex, reddish,
fully
winged;
length 5,62 mm;
clypeus and mandibles yellow; with brown
spots on
head black,
with front part of
antennae testaceous;
anterior weak
collar;
pronotum
propectus reddish;
816
Mem.
Fig.
mesothorax; testaceous,
Ent.
Chela of
Inst.,
no.
(Fig.
keels;
541
B)
occipital
dull,
around disc;
pronotal
1984
abdomen brown-reddish;
granulated,
carina complete;
carina pronotum dull,
37,
Dryinus lucens n. sp. (type)
metathorax and propodeum black; head
longitudinal occipital
550 -
Amer.
with a few
posterior ocelli
weakly granulated,
tubercles not reaching
legs
weak not touching
with numerous keels
tegulae;
scutum dull,
hairy,
Dryininae
granulated; 0,5
notaul ices almost
length of scutum;
reticulate rugose, transversal proximal
of 8
incomplete,
reaching
longitudinal
keels;
fore-wing
part of radial
approximately
granulated;
vein
propodeum
with 3 brown longer than
longer than segment 4;
tarsus slightly
549) with subapical tooth and 9 lamellae; segment 5 549) with 2 rows of 21 lamellae; apex with a group tibial spurs 1,1,2.
(Fig.
tarsus
(Fig.
lamellae;
MALE:
without
segment 1 of front
enlarged claw of front
invisible,
scutellum and metanotum dull,
bands and with distal
part;
817
unknown
LOCUS TYP ICUS:
Kontum
TYPICAL MATERIAL: HOSTS:
(Viet Nam) in B.
holotype F!
unknown
DISTRIBUTION:
Oriental
region:
VIET NAM:
Kontum
(Plateau),
B!
13. Dryinus lucens n. sp.
FEMALE:
fully
winged;
mandibles reddish; abdomen
reddish;
length 6,25 mm;
legs reddish;
carina complete;
impression strong);
numerous
keels
around
smooth,
regions punctate; of scutum;
di sc;
notaul ices
reticulate rugose,
without
with 3 brown
much
longer than proximal
550);
Ranau
TYPICAL MATERIAL: HOSTS:
tarsus
group of 15
MALE: unknown LOCUS TYPICUS:
part;
enlarged claw
segment 5 of front
apex with a
incomplete,
bands,
reaching
(Sabah,
part of radial
550)
tibial
forevein
tarsus slightly
tooth and
length
propodeum
keels on posterior surface;
segment 1 of front (Fig.
weak; with
approximately 0,5
metanotum dull,
with distal
with subapical
lamellae;
holotype F!
without sculpture,
not reach i ng tegu I ae; without sculpture; lateral
weakly punctate;
transversal
pronotum
tuberc I es
region
longitudinal
wing
than segment 4;
pronota I
occipital
carina;
(anterior impression very
pronotum shiny,
with central
scutellum shiny,
reticulate rugose;
not touching occipital
impressions
posterior
with clypeus and
thorax and propodeum black;
head dull,
posterior ocelli
crossed by two transversal
scutum shiny,
head black,
antennae testaceous;
longer
12 lamellae (Fig.
with 2 rows of 28
lamellae;
spurs 1,1,2.
Malaysia) in
B.
unknown
DISTRIBUTION:
Oriental
region:
MALAYSIA:
Ranau
(Sabah),
B!
14. Dryinus browni Ashmead
Dryinus browni Ashmead 1905: 109. Paradryinus terryi R.C.L. Perkins 1912: 10 (n. s)-n.); locus typicus Hong Kong
(China).
818
Mem.
Amer.
Ent.
Ins!.,
no.
37,
1984
5
3
4
Fig.
551
-
Dryinus browni Ashmead: antenna I segments 3-6.
Female of
Lestodryinus browni (Ashmead): Kieffer 1914b: 24. Paradryinus terryi R. c. L. Perk i ns: Ki effer 1914b: 35. Dryinus szepligettii Kieffer partim: Ponomarenko 1981 b: 879.
FEMALE:
fully
winged;
mandibles yellow; yellow;
antennae brown,
propodeum black; light;
length 6,25-7,68 mm;
antennal
head black,
with clypeus and
sometimes also malar space and front part of vertex with
abdomen
last 2-4 segments whitish;
brown;
thorax and
legs testaceous or brown,
with
tarsi
much broader than other segments
segments 4-5 flat,
(F i g.
551}; head dull, granulated, with some longitudinal keels on sides of frons; occipital carina complete; posterior ocell i not touching occipital carina;: pronotum dull, granulated, hairy, with numerous keels around disc; pronotal tubercles not reaching tegulae; pronotum crossed by two
transversal
impressions {anterior impression very weak;
posterior
impression strong}; scutum hairy, dull, granulated, rugose on posterior third; notaulices complete, posteriorly separated; scutellum as sculptured as scutum;
propodeum reticulate rugose,
posterior surface; distal
fore-w i ng
part of radial
wi th
vein much
without
three brown longer than
longitudinal
keels on
transversa I bands and with proximal
part;
segment
1 of
front tarsus slightly longer than segment 4; enlarged claw (Fig. 552) with subapical tooth and 14 lamellae; segment 5 of front tarsus (Fig. 552) with 2 rows of 40 1,1,2.
lamellae;
apex with a
group of 19 lamellae;
tibial
spurs
Dryininae
Fig.
MALE:
552 -
fully
Dryinus browni
Chela of
winged;
antennae yellow-brown;
brown;
legs testaceous,
=
head black,
with hind coxae partly brown; 9;
OL
with mandibles
thorax and propodeum black;
granulated and reticulate rugose; POL
Ashmead from De Tha Ngone (Laos)
length 3,25-3,75 mm;
yellow;
antenna I sockets;
819
=
frons with a median keel 4;
OOL
=
5;
OPL
=
2;
abdomen
head dull, between the
TL = 3;
occipital
Mem.
820
Amer.
Ent.
Inst.,
no.
37,
1984
\
\
\ \
I I /
Fig.
553 -
Male genitalia of Dryinus browni Ashmead from Kalutara Lanka):
right
half removed.
(Sri
Dryininae
carina complete;
temples
reticulate rugose; shiny,
scutum dull,
notaulices complete,
granulated;
longitudinal
invisible;
metanotum rugose;
bands and with distal
part of radial
(15:9);
553;
lOCUS TYPICUS:
fig.
Manila
TYPICAL MATERIAL: terryi in B. HOSTS:
PHiliPPINES: GV!
(Gi Sion,
Oriental Manila
Vientiane),
(Kanneliya Jungle,
scutellum
propodeum reticulate rugose, fore-wing
vein much
without dark
without
transversal
longer than proximal
part
spurs 1,1,2.
of D.
browni in WA; holotype F! of P;
(=Pyrops) dohrni (Fulgoridae).
region:
(luzon),
Pinigisan
weakly
(Philippines)
holotype F!
in Sri lanka Zanna
DISTRIBUTION: Island,
tibial
granulated,
posteriorly separated;
keels on posterior surface;
genitalia:
821
CHINA:
Ta Hian
(Hainan
Id.),
WA!
los Banos
(luzon),
(Mantalingajan,
Palawan),
CO!
B!
Cal.
SRI
Ol!
lANKA:
Kalutara,
BM!
lAOS:
Ol!
B!
TW! WA!
Bilaran
De Tha Ngone
Hiniduma
under name Dryinus
Dist.)(Ponomarenko 1981b:
szepligettii Kieffer), WA! MALAYSIA: Gn. Mulu (Sarawak), BM! HONG KONG, B!
15. Dryinus lankanus n. sp.
Dryinus trifasciatus Kieffer: Ponomarenko 1981b: 879.
FEMALE: fully winged; length 3,60-4,88 mm; head black, with clypeus, malar space, mandibles and part of ventral side testaceous; antennae testaceous with segments 9-10 light or brown thorax and propodeum black, of pronotum
testaceous or reddish;
testaceous or black, antennal 7:
4:
with tarsi
10: 8:
6:
5:
5:
4:
abdomen black or brown;
light;
segments 4-5 not flat;
21,5:
with segments 9-10 testaceous,
with posterior collar of pronotum and sides antennal
antenna I segments in following proportions: 5;
head flat,
irregular keels on frons and vertex;
frons
dull,
not reaching
tegulae;
around disc;
scutum dull,
impression;
pronotum dull,
=
10: OPl = 1,5; Tl impression and by a
disc humped;
granulated,
granulated,
keels near posterior margin;
granulated and with
with a complete median keel;
occipital carina complete; POL = 5; Ol = 4; OOl pronotum crossed by a weak anterior transversal strong posterior transversal
legs
segments cylindric;
pronotal
tubercles
reticulate rugose or with weakly
irregular visible
scutell um and metanotum du II,
granulated;
propodeum reticulate rugose,
longitudinal
keels;
fore-wing
with 3 dark transversal
bands beneath the
pterostigma and on
the basal
cells;
distal
vein much longer than proximal following proportions: 28: 5: 9: subapical
5;
with numerous striae
notaul ices almost complete,
near posterior marg i n of scutum;
=
without transversal or
radial
cell
open;
part of radial
part (10:5); fore tarsal segments in 21: 33; enlarged claw (Fig. 554) with a
tooth and with a row of 11
lamellae;
segment 5 of front
tarsus
822
Fig.
Mem. Amer.
Ent.
Inst.,
no.
37,
1984
554 - Chela of Dryinus lankanus n. sp. from Kandy
(Sri
lanka)
(Fig. 554) with two rows of nearly 31 lamellae; apex with a group of nearly 15 lamellae; tibial spurs 1,1,2. MALE: unknown lOCUS TYPICUS: Kibissa (Sri lanka) TYPICAL MATERIAL: holotype F! and 9 paratypes FF! in WA; 1 paratype F! in Ol. HOSTS: unknown DISTRIBUTION: Oriental region: SRI lANKA: Kibissa (W of Sigiriya, Mate Dist.), WA! Angunakolapelessa (Mon. Dist.), Ol! WA! Kandy (Kan. Dist.), WA! Parayanalankulam (25 mi. NW Medawaehehiya, Vav. Dist.), WA! Hasalaka (Kan. Dist.), WA! Ekgal Aru Palatupana tank (Ham. Dist.), WA!
(Sanet. Jungle, Amp.
NEARCTIC SPECIES OF
DRYINUS
Dist.), WA!
823
Dryininae
Four species are known.
KEY TO THE SPECIES
FEMALES
Segment
1 of
enlarged Segment 4; 2
1 of
enlarged
tarsus nearly
front
tarsus at
claw not
Head with TL
twi ce as
long as
segment
4;
longer
most
1,5
times as
long as
segment
spatulate •••••••••••••••••••••••••••••••• 3
propectus and pronotum 1. mexicanus (R.C.L. Perkins) Head with TL shorter than POL; head (except for yellow mandibles), propectus and pronotum black ••••••••••••••••••••••• . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 2. crawfordi (Krombei n) Head (dorsally viewed) wi th posterior ocell i located in front of the imaginary straight line joining posterior edges of eyes (Fig. 557 A) •••••••••••••• 3. ameMcanus (Ashmead) Head (dorsally viewed) wi th posterior ocell i located behind the imaginary straight I ine joining posterior edges of eyes (Fig. 557 B) ••••••••••••••••••••••••• 4. alatus (Cresson) fully
3
front
claw spatulate ••••••••••••••••••••••••••••••••••••••• 2
than POL;
head,
reddish-testaceous . . . . . . .
MALES
Only the male of
D. alatus
(Cresson)
1. Dryinus mexicanus
Paradryinus mexicanus
FEMALE:
fully
winged;
abdomen reddish-brown; very prominent;
Perkins)
head,
mesothorax,
comb.
antennae,
propectus and
meta thorax and propodeum black;
legs testaceous-reddish;
occipital carina complete;
pronotum dull,
n.
Perkins 1907: 39.
length 4,68 mm;
pronotum reddish-testaceous; reticulate rugose;
R.C.L.
(R.C.L.
is known.
head swollen,
dull,
POL shorter than OL;
crossed by a strong transversal
temples
posterior
impression, granulated; anterior transversal impression weak; disc humped; posterior collar very visible; anterior collar weak; pronotal tubercles not reaching tegulae; on
lateral
scutum
dull,
regions and near posterior margin;
granulated,
reticulate rugose
notaulices almost complete,
824
Mem.
Fig.
reaching
555 -
Amer.
Ent.
Ins!.,
no.
Chela of Dryinus mexicanus
almost posterior margin of scutum;
37,
1984
(R.C.L.
Perkins)(type)
scutellum and metanotum dull,
granulated; propodeum reticulate rugose, without longitudinal keels on posterior surface; fore-wing with 2 brown transversal bands and with distal front hook;
part of radial
vein much
longer than
proximal
tarsus nearly twice as long as segment 4; enlarged claw
lamellae; (among
(Fig.
555)
with subapical
segment 5 of front tarsus
which 6 lamellae much
continuously to the apex; MALE:unknown LOCUS TYPICUS:
tibial
(Fig.
555)
part;
tooth and with 5 very
holotype F!
HOSTS: unknown DISTRIBUTION: Nearctic region:
longer than other lamellae) spurs 1,1,2.
in WA. MEXICO:
long
with 3 rows of 36 lamellae
Patzcuaro (Mexico)
TYPICAL MATERIAL:
segment 1 of
segment 3 produced into a
Patzcuaro,
WA!
extending
Dryininae
------------------Fig.
556 -
Chela of
825
-
Dryinus crawfordi (Krombein) (type)
2. Dryinus crawfordi (Krombein) n. comb.
Mesodryinus crawfordi Krombein 1962: 3.
FEMALE:
fully
winged;
antennae brown, black;
abdomen brown-reddish;
head dull,
granulated,
of vertex;
occipital
granulated, distinct; notaul ices
carina complete;
posterior
granulated,
incomplete,
notaulices reticulate; without
10 yellow;
legs testaceous,
with mandibles yellow;
thorax and propodeum
with coxae partly black;
OPL shorter than OL;
anterior transversal
impression weak;
disc not very swollen;
scutum dull,
head black,
with numerous areolae or short keels on front part
not very humped;
collar invisible;
rugose,
length 4,37 mm;
with segments 1-3 and
pronotal
posterior collar short,
tubercles not reaching
with numerous areolae on
lateral
reaching posterior margin of scutum; scutellum and metanotum dull,
longitudinal
pronotum dull,
impression and anterior
furrow of
granulated;
keels on posterior surface;
tegulae;
regions; reticulate
fore wings with 2
826
Mem. Amer.
Ent.
Inst.,
no.
37,
1984
o
Fi g.
557 -
Heads of fema les of lJryinus american us
0
(Ashmead) (type) (A)
and
alatus (Cresson) from Nogales (Arizona) (B).
brown
transversal
than proximal segment 4; subapical
bands and with distal part of radial
part;
segment 1 of front
segment 3 produced into a hook; tooth and 9 lamellae;
3 rows of 45
lamellae (among
tarsus
lamellae much
to the apex;
LOCUS TYP ICUS: Plummers Island (Maryland, TYPICAL MATERIAL: holotype F! in WA. HOSTS:
enlarged claw
segment 5 of front
which 5
lamellae) extending continuously MALE: unknown
vein much
tarsus nearly twice as
tibial U.
S.
longer
long as
(Fig. (Fig.
556) 556)
with with
longer than other spurs 1,1,2.
A.)
unknown
DISTRIBUTION: Tompkins Co.
Nearctic region: (S.
Lansing,
N.
U. Y.),
S.
A.:
Plummers
Island
(Md.)
WA!
OLl
3. Dryinus american us (Ashmead)
Mystrophorus american us Ashmead 1887b: 128. Dryinus americanus (Ashmead): Ashmead 1893: 87. Prodryinus american us (Ashmead): Kieffer 1907b: 9. Mesodryinus americanus (Ashmead): Muesebeck and Walkley 1951: 1034.
FEMALE: black
fully
winged;
length 5,0 mm;
and abdomen brown-black;
head
reddish-testaceous, (Fig.
557 A)
shiny,
with petiole weakly
granulated, weakly rugose on frons; occipital carina incomplete, only visible behind the ocell i and on sides of vertex; posterior ocell i almost touching occipital
carina;
OL sl ightly
longer than POL;
vertex flat;
827
Dryininae
Fig.
558 -
pronotum shiny, disc not very
Chela of
granulated,
humped;
Dryinus american us
crossed by 2 weak
anterior collar invisible;
(Ashmead) (type)
transversal
impressions;
posterior collar
invisible,
because the posterior transversal impression is just on posterior margin of pronotum; pronotal tubercles not reaching tegulae; scutum shiny,
828
Mem.
granulated,
Amer.
Ent.
with weak areolae on
not very visible,
reaching
I nst.,
lateral
no.
regions;
approximately 0,65
and metanotum as scu I ptured as scutum, propodeum reticul a te rugose;
37,
1984
notaul ices incomplete,
length of scutum;
scutellum
but wi thout weak areol ae;
posterior surface wi th transversa I keel sand
with 2 longitudinal keels; the posterior surface so is divided into 3 regions; fore wings with 3 brown transversal bands and with distal part of radial
vein slightly
tarsus nearly as into a
hook;
lamellae;
enlarged claw
segment 5 of front
extending continuously MALE:
longer than proximal
long as segment 4; (Fig.
558)
tarsus
part;
segment 1 of front
segment 3 of front with subapical
(Fig.
to the apex;
tibial
558)
tarsus produced
tooth and 8
with 2 rows of 35
lamellae
spurs 1,1,2.
unknown
LOCUS TYP ICUS: Jacksonvi lie (Florida, U. S. A.) TYPICAL MATERIAL: holotype F! and 1 paratype F! HOSTS: unknown DISTRIBUTION: Nearctic region:
U. S.
A.:
in WA.
Jacksonville (Fla.),
WA!
4. Dryinus alatus (Cresson)
Gonatopus alatus Cresson 1872: 193. Dryinus alatus (Cresson): Patton 1879: 65. Dryinus alatus (Cresson): Ashmead 1893: 86. Hesperodryinus amphiscepae R.C.L. Perkins 1907: 41 (n. syn.); locus typicus Nogales (Arizona, U. S. A.). Hesperodryinus arizonicus R.C.L. Perkins 1907: 42 (n. syn.); locus typicus Nogales (Arizona, U. S. A.). Hesperodryinus audax R. C. L. Perk ins 1907: 43 (n. syn.); locus typicus Nogales (Arizona, U. S. A.). Prodryinus alatus (Cresson): Kieffer 1907b: 9. Mesodryinus alatus (Cresson): Kieffer 1914b: 32. Hesperodryinus amphiscepae R. C. L. Perk i ns: Kieffer 1914b: 39. Hesperodryinus arizonicus R.C.L. Perkins: Kieffer 1914b: 40. Hesperodryinus audax R. C. L. Perk ins: Kieffer 1914b: 41. Mesodryinus alatus (Cresson): Muesebeck and Walkley 1951: 1034. Hesperodryinus amphiscepae R.C.L. Perkins: Muesebeck and Walkley 1951:
1034.
Hesperodryinus arizonicus R.C.L. Perkins: Muesebeck and Walkley 1951:
1035.
Hesperodryinus audax R.C.L. Perkins: Muesebeck and Walkley 1951: 1035.
Hesperodryinus amphiscepae R.C.L. Perkins: Krombein 1979: 1242. Hesperodryinus arizonicus R. C. L. Perk i ns: Krombei n 1979: 1243. Hesperodryinus audax R. C. L. Perk i ns: Krombei n 1979: 1243. Mesodryinus alatus (Cresson): Krombein 1979: 1243.
Dryininae
Fig.
559 -
Chela of
829
Dryinus alatus {Cresson} from Nogales {Arizona}
FEMALE: fully winged; length 5,00-7,87 mm; yellow-testaceous, with petiole black; sometimes with black spots on head, scutum, scutellum, propodeum and abdomen; sometimes in addition frons almost fully black, propectus black, mesothorax, meta thorax and propodeum almost fully black; head {Fig. 557 B} dull, granulated; occipital carina incomplete, only visible behind the ocell i and on sides of vertex; pronotum as sculptured as head,
830
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
A
c Fig.
Varieties of enlarged claws of Dryinus alatus
560 -
from Nogales
(Arizona);
B:
from Ann Arbor (Michigan); Co. (Michigan)
with a strong posterior transversal transversal
impression;
posterior collar and a
impression and a
the pronotum so has a central less visible anterior collar;
as head or sculptured by more or less numerous notaulices complete,
posteriorly separated;
as sculptured as scutum;
prcpodeum shiny,
with a series of transversal anterior surface smooth, median
series
from
parallel
(Cresson):
C:
weak anterior humped disc,
a
scutum as sculptured
longitudinal
keels,
shiny;
scutellum and metanotum shiny, more or less reticulate rugose,
weak keels on posterior surface;
with more or less numerous areolae forming a
anterior margin
A:
from Midland
to posterior surface;
occasionally
831
Dryininae
anterior surface with strong keels; bands,
with distal
part of radial
trochanters more than twi ce as
fore wings with 2 brown vein
transversal
longer than proximal
part;
fore
(15: 6); segment 4 of front
long as broad
tarsus nearly twice as long as segment 1 or sl ightly shorter than segment 1 (33:41); segment 3 of front tarsus produced into a hook; enlarged claw (Fig. 559) with subapical tooth (big or small) and 5-11 lamellae or pegI ike hairs,
without series of teeth;
subapical
tooth more or less far from
560); segment 5 of front tarsus (Fig. 559) with 2 rows of 18-30 lamellae; apex with a group of nearly 20 lamellae; tibial spurs 1,1,2. MALE: fully winged; length 2,85-4,00 mm; head black; with mandibles the apex
reddish;
(Fig.
antennae brown;
legs brown, i ncomp lete;
with tarsi
thorax and propodeum black;
light;
head dull,
abdomen brown;
reticulate rugose;
OPL shorter than POL and than OL;
scutum,
occipital
carina
scute II um and
metanotum dull, as reticulate rugose as head; notaulices almost invisible; propodeum reticulate rugose, with areolae broader than areolae of head; fore wings without dark transversal longer than proximal part; tibial LOCUS TYPICUS: Texas (U. S. A.) TYPICAL MATERIAL:
alatus in WA;
bands,
with distal part of radial
alatus in AE; 1 para type F! of G.
holotype F of G.
and 2 paralectotypes
lectotype F!
vein
spurs 1,1,2.
(1
F,
1 M)!
(designated
amphiscepae in B; 1 paratype F! of H. amphiscepae in BM; lectotype F! (designated by Olmi 1982) of H. arizonicus in B; holotype F! of H. audax in B. HOSTS: in U.S.A. Acanalonia (=Amphiscepa) bivittata (Say) (Acanaloniidae) and Mistharnophantia sonorana Kirkaldy (Flatidae) (R.C.L. Perkins 1907). by Olmi
1982) of H.
DISTRIBUTION: A.: Co.,
Nogales Tex.),
Midland Co. Wattacoo
Nearctic region:
(Ariz.), TE!
OLl
B!
Waukesha Co.
(Mich.),
(Pickens Co.,
WA! S.
CANADA:
BM!
Texas,
(Wis.),
SI.
Catharines
AE!
WA!
WA!
Ann Arbor
Columbia
(Mo.),
C.),
Cleveland
TW!
OL!
WA! (S.
(ant.),
aT!
College Station (Mich.),
Forsyth
C.),
TW!
U.
S.
(Brazos
TW!
(Ga.),
OL! OT!
Moorestown
OLl (N.
J.), OL! TW! Horicon Lake (Lakehurst, Ocean Co., N. J.), AM! Westfield (N. J.), OL! Kalbfleisch Field Station (Huntington, N. Y.), AM! Shokan (N. Y.),
TW!
NEOTROP I C SPEC I ES OF
Nineteen species,
including a fossil
DRYINUS
species,
are known.
KEY TO THE SPECIES (excluding
the fossi I species Dryinus palaeodominicanus Currado and Olmi)
FEMALES
832
2
Mem.
Occipital
carina
Occipital
carina
Amer.
Ent.
no.
37,
invisible ••••••••••••
1.
complete
or
Thorax and propodeum fu I I y Head with POL
longer
constans n. sp.
fu I I y
t estaceous •••••••••• 3
black ••••••••.•••••••••••• 4
or fully
than OL;
1984
incomplete ••••••••••.•••••••••••• 2
or a I most
Thorax and propodeum mostly
3
Inst.,
occipital
carina
complete •••••••••
••••••••••••. •••. •••. . ••••••••. •. . . . " 2. citricolus n. sp. Head with POL almost 0,5 as long as OL; occipital carina incomplete •.•.•.•••••••••••••••••••••• 3. flavoniger n. sp. 4
Scutum fully
sculptured by more
or
less numerous
longitudinal
keels •••••••••••••••••••••••••••••••••••••••••••••••••••••••••• 5 Scutum without keels, fully reticulate rugose or partly g ranu I at ed •••••••••••••••••••••••••••••••••••••••••••••••••••• 16
5
Pronotum fully
granulated
keels on posterior
,wi thout
transerversal
keel s,
except
impression;
for
some
occipital
4. wellingensis n.
comp let e ••.•••••••••••••••••••••
short
carina
sp.
Pronotum differently sculptured, with numerous irregular keels; occipital carina incomplete •••••.••.••••••••••••••••••••••••••• 6
6
Posterior ocelli more distant from occipital carina (Fig. 566 A); head with OPL nearly as long as POL (Fig. 566 A) . . . . . . . . . . ..
. . . . . . . . . . . . . . . . . . . . . •. . •. . . . . . . . " Posteri or ocell i OPL
shorter
nearer occipi tal
than POL
(Fig.
Segment Segment
8
Prothorax mostly b l a c k . . . . . . . . . . . . .
6.
Prothorax
7.
front front
tarsus tarsus
(Fig.
566 B);
sp.
head wi th
566 B) . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7
7
1 of of
surinamensis n.
5.
carina
shorter than segment 4 ••••••••••••••• 8 slightly or much longer than segment
4 ••••••••••••••••••.•••.••••••••••••••••••••••••••••••••••••••• 9
9
Antennae broad;
fully less
slender,
prothorax
Antennae more long
fully
slender,
as broad;
10 Dorsal
or mostly
wi th
segment
wi th
segment
fully
striatus (Fenton) argentinus n. sp.
4 nearly
testaceous..
prothorax
surface of
reddish..
4 nearly
or partly
twice as
long
as
caraibicus n. sp.
8.
five-nine
times as
reddish . . . . . . . . . . . . . . 10
propodeum sculptured by numerous
longitudinal
keel s ••••••••••••••••••••••••••••••••••••••••••••••••••••••••• 11 11
Dorsal surface of propodeum fully reticulate rugose . . . . . . . . . . . 13 Head with OL nearly two-three times as long as POL; head fully redd ish. • . • • • • • • • • • • • • • • • • • • • • • • • • • • 9. napensis n. sp. Head with OL as long as or slightly longer than POL; head partly black and
12 Pronotum
testaceous •••••••••••••••••••••••••••••••••••••••••• 12 (laterally
viewed)
with disc nearly
as anterior collar . . . . . . . . . . . . . . . . . . . Pronotum
(laterally
viewed)
with disc
anterior collar . . . . . . . . . . . . . . . . . . . . . . 13 Fore
tibiae nearly
Fore 14 Head
tibiae less than twice as fully reddish-testaceous;
by
longitudinal
twice as
keels;
long as
less 10.
fore
three
times as
long
kimseyae n. sp.
18.
than
twice as
long as
opacifrons n. sp. trochanters . . . . . . . . . . 14
long as fore trochanters ••••••• 15 scutum more strongly sculptured
notaulices weakly
• • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • ••
1 1.
visible among keels
grandis (Og I ob lin)
Dryininae
Head partly black and keels;
833
testaceous;
notaulices clearly
distinct
scutum wi th weak longitudinal among kee Is ••••••••••
alvarenganus n • sp. sinopensis n. sp. 14. belizensis n. sp.
...................................
12.
15 Head part Iy black •••••••.••••••••••• Head
fully
13.
reddish-testaceous . . . . . .
16 Scutum granulated, wi th areolae on lateral posterior half of median region ••••.
regions and
on
15. antilleanus (Evans) Scutum fully
17 Antennal
reticulate
4
segment
less
rugose •••••.•••••••••.••••..•••••••••• 17 slender,
near I Y
twi ce as
8. Antennal
long as broad ••
caraibicus n.
sp.
4 more slender, nearly five times as
segment
long as
broad ••••••••••••••••••••••••••••••••••••••••••••••••••••••••• 18
18 Posterior ocelli
more distant
from occipital
• • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • • •• Posteri or ocell i
nearer
occipi tal
16.
carina
carina
(Fig.
580 A)
teutoniae n. sp. 580 B)
(Fig.
17. catarinae n. sp.
MALES
Frons wi thout
2
median keel . . . . . . . . . . . .
Frons with a
complete or
Frons with a
complete median keel;
5.
surinamensis n. sp.
incomplete median keel . . . . . . . . . . . . . . . . 2 notaul ices
reaching
approximately 0,5 length of scutum; scutum granulated, not reticulate rugose . . . . . . . . . . . . . . . . . . . 15. antilleanus (Evans) Frons wi th an half
of
incomplete median keel
frons:
notaulices complete.
scutum reticulate
onlv visible on posterior posteriorlv
ruCJose . . . . . . . . . . . . .
6.
separated:
striatus (Fenton)
The males of other species are unknown.
D. palaeodominicanus n. sp., fo=sil species from Dominica, isn't present
in
through
the amber,
the key because the only known so that
its morphology
specimen
is weakly
visible
isn't very distinct.
1. Dryinus constans n. sp.
FEMALE:
fully
winged;
broad median frontal 9-10 testaceous;
head testaceous,
antennae brown,
thorax and propodeum black,
pronotum reddish; testaceous,
length 6,87 mm;
stripe black;
abdomen black,
with posterior margin of
with some regions testaceous;
with hind coxae partly black;
antenna I segments
with vertex and a
with segments 1-2 and
in following proportions:
legs
antennae distally thickened; 16: 6:
44:
24:
20:
20:
14:
14:
834
Mem.
Fig.
561
-
Amer.
En!.
Ins!.,
no.
37,
1984
Chela of Dryinus cons tans n.
sp.
(type)
12: 15; head dull, granulated and with numerous weak irregular striae; frons with a complete median keel; occipital carina invisible; POL = 4; OL = 2,5; OOL = 8,5; pronotum with a strong anterior transversal impression and with a strong posterior transversal
impression;
disc humped;
posterior
835
Dryininae
A
Fig.
562 -
and
collar
Dryinus striatus (Fenton) (type) (A) and citricolus n. sp. from Manati (Puerto Rico) (8)
Pronotum of females of
invisible;
pronotal
by numerous parallel separated;
tubercles not reaching
longitudinal
keels;
scutellum and metanotum dull,
tegulae;
scutum sculptured
notaul ices complete, reticulate rugose,
posteriorly with 2
incomplete longitudinal keels on posterior surface; fore wings with two broad dark transversa I bands, one on the basa I cell s and another beneath the pterostigma; distal part of radial vein longer than proximal part (21:15); fore tarsal segments in following proportions: 21: 4,5: 11: 20:
36;
proportions of fore tibiae and trochanters: 37:23; enlarged claw (Fig. 561) with a subapical tooth and 20 lamellae; segment 5 of front tarsus (Fig.
561) with two rows of nearly 24 lamellae; apex with a group of nearly 12 lamellae; tibial spurs 1,1,2. MALE:
unknown
LOCUS TYP I CUS:
Represa do Rio Grande
TYPICAL MATERIAL: HOSTS:
holotype F!
(Braz i I)
in TW.
unknown
DISTRIBUTION: Neotropic region: (Guanabara), TW!
BRAZIL:
Represa do Rio Grande
2. Dryinus citricolus n. sp.
FEMALE: fully winged; length 3,12-3,50 mm; fully testaceous, with petiole black; head dull, granulated; occipital carina complete; OPL ; OL; OL shorter than
POL;
pronotum
(Fig.
562 B)
dull,
granulated,
not humped,
836
Mem.
Fig.
563 -
Amer.
Ent.
Inst.,
no.
37,
Chela of Dryinus citricolus n.
with 2 weak anterior and posterior transversal posterior collar weakly visible; scutum shiny,
granulated;
I ength of scutum; shiny,
1984
pronotal
sp.
(type)
impressions;
anterior and
tubercles not reaching
tegulae;
notaulices incomplete,reaching approximately 0,5
scutell um and metanotum du II,
not reticulate rugose or weakly
granu I a ted;
reticulate rugose;
propodeum
areolae more
visibi Ie on anterior surface; posterior surface smooth, without sculpture and without longitudinal keels; fore wings with 2 brown transversal bands and with distal front hook;
part of radial
enlarged claw
segment 5 of front
(Fig.
tarsus
group of 9 lamellae; MALE:
vein
longer than proximal
tarsus nearly twice as long as segment 4;
tibial
563) (Fig.
with subapical 563)
part;
segment 1 of
segment 3 produced tooth and without
with 1 row of 10 lamellae;
into a
lamellae; apex with a
spurs 1,1,2.
unknown
LOCUS TYPICUS: Lajas (Puerto Rico) TYPICAL MATERIAL: holotype F! and 1 paratype F! 1 paratype F!
in WA;
in OL.
HOSTS: unknown DISTRIBUTION: Neotropic region:
PUERTO RICO:
Lajas,
WA!
Manati,
OL!
Dryininae
Fig.
564 -
Chela of
Dryinus flavoniger n. sp. (type)
837
838
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
3. Dryinus flavoniger n. sp.
FEMALE: fully winged; length 4,56-5,00 mm; testaceous, with a central black spot on propodeum; petiole black; internal side of stalks of hind femora black; antennae distally thickened; antennal segments in following proportions: 8: 6: 28: 15: 12: 8: 6,5: 5,5: 5: 7; head flat, dull, granulated; incomplete,
frons without median and visible behind the ocelli
occipital carina OPl = 3;
keels;
invisible behind the temples;
pronotum dull,
POL
as sculptured as head,
anterior and posterior transversal pronotal
lateral
and on sides,
tubercles not reaching
impressions;
tegulae;
occipital
carina
but not reaching eyes;
=
3;
=
Ol
5,5;
weakly humped, posterior collar
scutum,
=
OOl with
7;
weak
invisible;
scutellum and metanotum
dull, as sculptured as head; notaulices incomplete, weakly visible, reaching 0,65 length of scutum; propodeum reticulate rugose, with two longitudinal
median keels joining metanotum
to petiole;
fore wings with
two
dark transversa I bands, one on the basa I cell s and another beneath the pterostigma; distal part of radial vein longer than proximal part (27:8); fore tarsal
segments in following proportions:
2 and 3 of front tarsus produced into hooks; a subapical 564)
tooth and with 10 lamellae;
with 1 row of 15 lamellae;
21:
4:
5,5:
18: 27;
enlarged claw
(Fig.
segment 5 of front tarsus
apex with a
group of nearly
segments 564)
14 lamellae;
tibial spurs 1,1,2. MALE: unknown lOCUS TYPICUS: Faz. Taperinha (Para, TYPICAL MATERIAL: holotype F! in OT; HOSTS: unknown DISTRIBUTION: Neotropic region: Encruzilhada
(Bahia),
Fig.
565 -
Brazil) 1 paratype F!
BRAZil:
Faz.
in OL.
Taperinha
(Para),
OL!
Chela of
with
(Fig.
Dryinus wellingensis n. sp. (type)
OT!
839
Dryininae
4. Dryinus wellingensis n. sp.
FEMALE:
fully
testaceous; black,
length 5,62 mm;
winged;
with tarsi
ligth;
following proportions:
black;
mandibles and clypeus
with segments 1-3 and 10 testaceous;
antennae brown,
antennae distally thickened;
antennal
legs
segments in
11: 6: 22: 11: 11: 9: 8: 7,5: 6,5: 9; antenna I (11 :3); head dull, flat,
segment 4 nearly four times as long as broad granulated,
with numerous areolae on frons and vertex around eyes;
OPL
=
=
1,5; TL
occipital
on posterior transversal
carina complete;
impression;
transversa I impression weak;
= 4;
POL
pronotum weakly
= 4;
frons
= 11; 3; pronotum dull, granulated, with numerous short keels
with a median keel;
OL
humped,
OOL
with anterior
posterior transversa I impression stronger;
posterior collar almost invisible;
pronotal
tubercles reaching tegulae;
scutum dull, granulated, with numerous longitudinal parallel keels; notaulices weakly visible among keels; scutellum and metanotum dull, granulated;
propodeum with dorsal surface sculptured by
longitudinal
parallel keels; posterior surface reticulate rugose and without longitudinal keels; propodeum without transversal keels; fore wings with two dark transversal
bands,
one on
the basal
cells and another beneath
the
(16:7); 19: 2: 5: 13: 19; segments 2 and 3 of front tarsus produced into hooks; enlarged claw (Fig. 565) with a subapical tooth and with 10 lamellae; segment 5 of front tarsus (Fig. 565) with 2 rows of nearly 36 lamellae extending continuously to the apex; tibial spurs 1,1,2. pterostigma;
fore tarsal
MALE:
distal
vein
longer than proximal
part
segments in following proportions:
unknown
LOCUS TYPICUS:
Middlesex
TYPICAL MATERIAL: HOSTS:
(British Honduras)
holotype F!
in OT.
unknown
DISTRIBUTION: NOTE:
part of radial
Neotropic region:
the species is named
in
BRITISH HONDURAS: honor of Mr.
E.C.
Middlesex,
OT!
Welling.
5. Dryin us surinamensis n. sp.
FEMALE: vertex
fully
reddish,
with
testaceous, testaceous;
length 6,87-10,12 mm;
head black,
clypeus and mandibles yellow;
irregular black spots on vertex;
sometimes head
occiput yellow;
meta thorax ,
propodeum and abdomen black;
light or testaceous with brown spots on coxae;
shiny or dull,
granulated,
with ocellar region swollen; the ocelli
with front part of antennae
with segment 3 brown; propectus black, with dorsal region pronotum black, with margins and posterior collar reddish;
mesothorax, tarsi
winged;
(more along orbits),
with numerous occipital
and on sides of vertex;
longitudinal
carina
legs brown
head
wi th
(Fig. 566 A)
keels;
vertex flat,
incomplete, only visible behind
pronotum shiny,
with numerous keels
840
Fig.
Mem. Amer. Ent.
566 -
Ins!., no. 37,
1984
Heads of females of Dryinus surinamensis n. striatus (Fenton) (type) (B).
sp.
(type)(A)
and
Dryininae
Fig. 567 -
around disc;
keels;
impression and anterior collar weak;
impression strong;
tubercles not reaching
longitudinal of keels;
Dryinus surinamensis n. sp. (type)
anterior transversal
posterior transversal pronotal
Chela of
841
tegulae;
notaulices,complete,
posterior collar short, scutum shiny, but almost
scutellum and metanotum punctate;
distinct;
with numerous
invisible among furrows
propodeum reticulate rugose,
842
Mem.
Fig.
568 -
Amer.
Ent.
no.
37,
1984
Male genitalia of Dryinus surinamensis n. (Brazi I):
with 2 short brown
Ins!.,
longitudinal
transversal
than proximal
keels on posterior surface;
bands and with distal
part
proportions: 70:35;
(25:18);
with subapical
567)
with 2 rows of 35 lamellae;
Itapetinca
vein much longer
trochanters in following
tarsus nearly as long as segment 4;
tarsus produced
567)
from
fore wings with 2
part of radial
fore tibiae and
segment 1 of front
segments 2 and 3 of front
sp.
right half removed.
tooth and 26 lamellae;
into hooks;
enlarged claw
segment 5 of front
tarsus
apex with a group of 20 lamellae;
(Fig. (Fig. tibial
spurs 1,1,2. MALE: fully winged; length 3,50-4,25 mm; black; clypeus and mandibles testaceous; antennae brown, wi th segments 1-2 testaceous; legs testaceous,
Dryininae
Fig.
with hind coxae, thickened; 10: 9:
9:
569 -
Dryinus striatus (Fenton)(type)
hind femora and hind tibiae brown;
antennal 10;
Chela of
843
antennae not distally
segments in following proportions: 7:
head dull,
fully reticulate rugose;
frontal
7:
20:
11:
line absent;
12:
12:
844
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
/ ./ 1 - - - - - 1 / ___ I
/
/
/
Fig.
570 -
Male genitalia of Dryinus striatus (Brazil):
(Fenton)
left half removed.
from Encruzilhada
Dryininae
occipital
carina
of head,
on
incomplete,
not visible on
sides and on ventral
side;
head directed towards the eyes and 4;
OOL = 5,5;
dull,
OPL = 3;
complete,
temples,
occipital
laterally
temples invisible;
reticulate rugose;
apparently
845
visible on dorsal
carina on dorsal
touching eyes;
scutum,
side
side of
POL = 7;
OL =
scutellum and metanotum
notaulices almost invisible near posterior margin,
incomplete, but reaching
posteriorly separated;
almost posterior margin of scutum;
propodeum;
transversal or longitudinal keels; pterostigma; distal part of radial
dull,
reticulate rugose,
if
without
fore wings weakly darkened beneath the vein shorter than proximal part (15:18);
genitalia: fig. 568; tibial spurs 1,1,2. LOCUS TYPICUS: Charlesburg (Suriname) TYPICAL MATERIAL: (1
F,
2 MM)!
HOSTS:
holotype F!
in TW;
in LE;
3 paratypes
(1
1 paratype F!
M,
Neotropic region:
Raleigh Falls
(Coppename River),
ECUADOR:
Ongota,
TW!
BRAZIL:
SURINAME: OL!
Lestodryinus striatus
Itapetinca
FEMALE:
fully
winged;
(more along orbits),
with a
median frontal specimen);
mesothorax,
brown-reddish;
legs testaceous;
incomplete,
13;
strong
a
collar short, long
head
(Fig.
with 2
transversal
long as proximal
proportions: fore
25:
5:
part 12:
37:
carina;
56;
an
tegulae,
anterior
weakly
fore tarsal
visible among
keels;
part of radial
segments
twice
fully propodeum
keels on posterior surface; longitudinal
posterior
nearly
metanotum granu la ted;
dorsal
fore wings vein nearly
in following
segments 2 and 3 of fore tarsi
tibiae and trochanters
pronotum pronotum
scutum shiny,
notaul ices complete,
bands and with distal (19:20);
carina POL =
anterior collar visible;
tubercles not reaching
longitudinal
abdomen
occipital
transversa I impress ion and by
keels;
(only
granulated and
sides without sculpture;
disc humped;
pronotal
longitudinal
holotype
and on sides of vertex;
surface reticulate rugose or sculptured by with 2 brown
dull,
keels on frons;
scutell um as scu I ptured as scutum;
reticulate rugose,
hooks;
566 B)
as distance between tegulae and tubercles;
sculptured by
sometimes face in
with sides and posterior
posterior ocell i touching occipital
impression;
distinct;
with front part of
antennae missing
longitudinal
posterior strong
trasversal
comb.
head black,
pronotum black,
wih numerous keels around disc;
crossed by
OL!
meta thorax and propodeum black;
with some
COL =
n.
LE!
WA!
2.
only visible behind the ocelli
1,5; OL = 3; shiny,
1927:
stripe testaceous;
propectus black;
reticulate rugose,
(Fenton)
TW!
(Loreto),
clypeus and mandibles yellow;
collar redd i sh;
as
3 paratypes
(Paramaribo),
San Antonio
(Bahia),
length 6,0-7,5 mm;
vertex known
Fenton
Charlesburg
PERU:
6. Dryinus striatus
keel s;
in WA;
in OL.
unknown
DISTRIBUTION:
as
2 FF)!
in following proportions:
produced
44:28;
into
enlarged
846
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
claw (Fig. 569) with subapical tooth and 11 lamellae; segment 5 of front tarsus (Fig. 569) with 2 rows of 27 lamellae; apex with a group of 28 lamellae; MALE:
tibial
fully
spurs 1,1,2.
winged;
length 4,44 mm;
and hind coxae black;
black;
hind femora brown;
antenna I segments in following proportions:
mandibles testaceous,
with mid
antennae not distally thickened; 9:
8:
23:
12:
12:
12:
12:
10: 9:
10; head dull, reticulate rugose; frontal I ine incomplete, only visible on posterior half of frons; occipital carina incomplete, invisible on temples; POL = 7; Ol = 4,5; OOl = 4,5; OPl = 1; scutum, scutellum and metanotum reticulate rugose, dull; notaulices complete, posteriorly separated, weakly visible near posterior margin of scutum; a transversal
strong keel
surface reticulate rugose; of radial tibial
vein
nearly as
and posterior surface;
fore wings with distal long as proximal
part
with
posterior
half darkened;
distal part
(17: 17); genital ia:
fig.
570;
38'
W 40
spurs 1,1,2.
lOCUS TYPICUS:
Santarem
TYPICAL MATERIAL: HOSTS: unknown DISTRIBUTION: 33'
propodeum reticulate rugose,
between dorsal
(Brazil)
holotype F!
in WA.
Neotropic region:
S (Amazonas),
TW!
BRAZil:
Encruzi Ihada
Santarem
(Bahia),
(Para),
TW!
WA!
71
0
Ol!
7. Dryinus argentinus n. sp.
FEMALE:
fully
winged;
length 4,37-5,62 mm;
head
testaceous,
with vertex
and posterior ha If of frons black; antennae brown-testaceous; prothorax reddish, sometimes with two black spots on sides; mesothorax, metathorax and propodeum black; brown;
abdomen brown;
antennae distally thickened;
proportions:
10: 7:
32:
15:
14:
with numerous longitudinal occipital
10: 9:
striae;
legs testaceous,
antennal 9: 7:
wi th coxae part I y
segments in following 10;
head flat,
granulated and
frons with a median complete keel;
carina
incomplete,
visible behind the ocelli
posterior ocelli;
POL = 2,5;
Ol = 2,5;
and on sides of
OOl = 10; Tl = 2;
pronotum shiny,
with two strong transversal impressions; anterior collar distinct and long; posterior collar invisible; pronotum with numerous longitudinal keels around disc; pronotal tubercles not reaching tegulae; scutum dull, sculptured by numerous parallel longitudinal keels; notaul ices invisible among longitudinal
keels;
scutellum sculptured by numerous parallel
longitudinal
keels; metanotum dull, granulated; propodeum with dorsal surface sculptured by numerous parallel longitudinal keels; posterior surface reticulate rugose and with two incomplete longitudinal without transversal
keels between dorsal
with two dark transversal beneath the pterostigma; proximal
part
fore tarsal
(14:16);
bands, one on distal
keels;
propodeum
and posterior surface; the basal
part of radial
vein
fore wings
cells and another little shorter than
proportions of fore tibiae and trochanters:
segments in following
proportions:
18: 4:
9:
23:
38;
30:22;
enlarged
Dryininae
Fig. 571
claw
(Fig.
front
tarsus
571)
continuously MALE:
-
Chela of
Dryinus argentinus n. sp.
with a subapical
(Fig.
571)
tibial
(type)
tooth and with 7 lamellae;
with 2 rows of nearly 45
to the apex;
847
segment 5 of
lamellae extending
spurs 1,1,2.
unknown
LOCUS TYP ICUS:
Horco Molle (Argentina)
TYPICAL MATERIAL:
holotype F!
and 5 paratypes FF!
in TW;
3 paratypes
848
Mem. Amer.
Fig.
FF!
in AM;
572 -
Ent.
Inst.,
no.
37,
1984
Chela of Dryinus caraibicus n.
5 paratypes FF!
sp.
(type)
in Ol.
HOSTS: unknown DISTRIBUTION: Neotropic region: ARGENTINA: Horco Molle (Tucuman), TW! BRAZil: Caruaru, OL! Silva Jardin (Rio de Janeiro) TW! Sinop (Mato Grosso),
TW!
Represa do Rio Grande (Guanabara),
Ol!
BOLIVIA:
Rio
Ichilo
Dryininae
Fig.
573 -
Chela of Dryinus napensis n.
849
sp.
(type)
(Santa Cruz), AM! 2 Km N mouth of Rio Chapare (Rio Mamore, Cruz), AM! OLl ECUADOR: Napo and Coca Rivers, TW!
Dept.
Santa
8. Dryinus caraibicus n. sp.
FEMALE:
fully winged; length 4,56-4,62 mm; head testaceous, with region of vertex near occipital carina dark; antennae testaceous-dark; prothorax testaceous; mesothorax, meta thorax and propodeum black; abdomen browntestaceous, with fore and mid coxae almost fully black, hind coxae apically black, clubs of femora brown; antennae distally thickened; antennal segments in following proportions: 14: 7: 22: 7: 7: 8: 7: 6: 6: 9; antenna I segment 4 nearly twice as long as broad; head dull, very weakly reticulate rugose or sculptured by irregular keels; frons with a median keel; occipital carina incomplete, only visible behind the ocelli and on sides of ocelli; occipital carina not visible on temples; POL = 3; OL = 2,5; OOL = 9; OPL 2; TL = 0; pronotum dull, broader than long (22:17); pronotum as sculptured as head, with some keels around disc, with an anterior strong transversa I impress ion; disc very humped; posterior transversa I impression strong; posterior collar invisible; pronotal tubercles reaching tegulae; scutum dull, sculptured by numerous paC'allel longitudinal keels; sometimes (in Bolivian specimen) keels weakly visible and sculpture partly reticulate rugose; notaul ices complete, posteriorly separated; scutum punctate, dull, with some irregular striae near anterior margin; metanotum without sculpture, smooth; propodeum reticulate rugose, dull; fore wings with two dark transversa I bands, one on the basa I cell s and another beneath the pterostigma; distal part of radial vein longer than proximal part (17:11); fore tibiae and trochanters in following proportions: 21 :10; fore tarsal segments in following proportions: 18: 3: 6: 14: 24; enlarged claw (Fig. 572) with a subapical tooth and with 6 lamellae; segment 5 of front tarsus
850
Mem.
(Fig.
572)
lamellae;
with
Amer.
Ent.
Inst.,
1 row of 19 lamellae;
segments 2 and 3 of front
no.
37,
1984
apex with a
group of nearly 10
tarsus produced
into hooks;
tibial
spurs 1,1,2. MALE:
unknown
LOCUS TYPICUS:
Simia
TYPICAL MATERIAL:
(Trinidad)
holotype F!
1 para type F!
in OT;
1 para type F!
in TW;
in AM. HOSTS:
unknown
DISTRIBUTION: 31'
S 55°
37'
Neotropic region: W (Sinop,
(Rio Mamore,
Dept.
TRINIDAD:
Mato Grosso),
Seni),
TW!
Simia
(Arima),
SOL IVIA:
OTt
BRAZIL:
12°
10 Km E San Antonio
AM!
9. Dryinus napensis n. sp.
FEMALE:
fully
winged;
margins of clypeus testaceous; partly
thorax,
testaceous;
propodeum,
abdomen
tarsus nearly seven
in following proportions:
excavated"
dull,
sculptured by
visible among ocell i;
frons
visible behind the ocelli OOL =
13;
OPL =
head reddish,
antennae brown, and
1;
11:
numerous
with a median
52:
pronotal
15:
11:
longitudinal keel;
9:
antennal
8:
7:
parallel
occipital
9;
head
keels,
carina
also
incomplete,
POL = 1;
OL = 3;
with a strong anterior transversal
impression and a strong posterior transversal invisible;
trochanters
long as broad;
14:
and on sides of posterior ocelli;
pronotum humped,
fore
antennae distally thickened;
times as 8:
with mandibles and
with segments 1-2
legs black;
hind trochanters testaceous;
segment 4 of front segments
length 6,87 mm;
testaceous;
tubercles not reaching
impression;
tegulae;
posterior collar
pronotum with numerous
keels around disc; scutum and scutellum dull, sculptured by numerous longitudinal parallel keels; notaul ices not clearly visible among keels; metanotum dull, dorsal
weakly sculptured by
surface sculptured by
surface reticulate rugose and with propodeum wi thout
two dark
another beneath
the pterostigma;
proximal
part
(24:12);
fore tarsal
longitudinal
two complete
transversa I keel s between
fore wings with
37:18;
irregular keels;
numerous
transversal
bands,
distal
fore tibiae and
segments
segments 2 and 3 of front
keels;
longitudinal
keels;
posterior
dorsa I and posterior surface; one on
the basal
part of radial trochanters
vein
cells and
longer than
in following proportions:
in following proportions: tarsus produced
propodeum with
parallel
25:
into hooks;
5:
11:
23:
42;
enlarged claw
(Fig.
573) with 2 subapical teeth and with 22 lamellae; segment 5 of front tarsus (Fig. 573) with 2 rows of nearly 50 lamellae extending continuously to the apex;
tibial
spurs 1,1,2.
MALE: unknown LOCUS TYPICUS:
Limoncocha
TYPICAL MATERIAL: HOSTS:
(Ecuador)
holotype F!
in OT.
unknown
DISTRIBUTION:
Neotropic region:
ECUADOR:
Limoncocha
(Napo),
OTt
Dryininae
Fig.
574 -
Chela of
851
Dryinus opaciirons n. sp. (type)
10. Dryinu5 opaciirons n. sp.
FEMALE:
fully
space brown frons;
winged;
and a
length 5,00-5,62 mm;
antennae testaceous,
pronotum
head testaceous,
with malar
broad black spot on vertex and on posterior half of
testaceous,
with
somet imes darkened;
two lateral
propectus testaceous;
black or brown spots;
pronotal
margins testaceous; mesothorax, metathorax and propodeum black; abdomen brown-testaceous; legs testaceous; antennae distally thickened; antenna I segments
in following proportions:
12:
5:
34:
15:
12:
12:
10:
9:
11;
8:
head
dull, flat, granulated, with some weak longitudinal keels on frons; frontal line complete; occipital carina incomplete, invisible on sides of head, only visible on dorsal
side of head,
where the occipital
carina
is touching
posterior edges of eyes; POL = 2; OL = 3; OOL = 10; OPL = 1; temples invisible; pronotum shiny, with two strong transversal impressions; anterior collar
long and distinct;
pronotum collar;
posterior collar invisible;
with numerous keels around disc,
pronotal
tubercles not reaching
disc very humped;
less than
tegulae;
twice as
scutum,
long
as
scutellum and
metanotum dull, sculptured by numerous parallel longitudinal keels; notaul ices weakly visible among keels; propodeum with dorsal surface sculptured by
numerous parallel
surface reticulate rugose; longitudinal basal
keels and with posterior
fore-wing
with
two dark
nearly as with
tarsus
long as proximal
part
in following proportions: two subapical
(Fig.
574)
(16:16); 26:
4:
teeth and with 8
spurs 1,1,2.
distal
radial
cell
9:
38;
24:
lamellae;
with 2 rows of nearly 40
to the apex; tibial MALE: unknown
two incomplete
transversal
cells and another beneath the pterostigma;
segments 574)
keels;
longitudinal
posterior surface also with
bands,
one on
part of radial
open;
fore
the vein
tarsal
enlarged claw
(Fig.
segment 5 of front
lamellae extending continuously
852
Mem.
Fig.
lOCUS TYPICUS:
575 -
Amer.
Ent.
no.
37,
1984
Dryinus grandis (Ogloblin)(type)
Chela of
Jacareacanga
TYPICAL MATERIAL:
Ins!.,
(Brazil) and 1 para type F!
holotype F!
in TW;
1 para type F!
in
Ol. HOSTS:
unknown
DISTRIBUTION: Teutonia
Neotropic species:
(Santa Catarina),
TW!
BRAZil:
Jacareacanga
(Para),
TW!
Nova
Ol!
11. Dryinus grandis (Og lob lin) n. comb.
Nesodryinus grandis Ogloblin 1938: 35.
FEMALE:
fully
winged;
length 8,12 mm;
thorax and propodeum black, pronotum reddish; trochanters, tarsi
abdomen brown-reddish;
femora
testaceous,
antennae distally weakly
following proportions:
carina
16: 8: 50: 20:
OPl =
longitudinal
incomplete,
1,5; Tl
transversa I
=
keels;
impression and by distinct;
longer than broad;
thickened;
with coxae,
legs black,
with
antenna I segments in
18: 16: 14: 12: 10: 14; antennal
frons
with a
head dull,
humped,
granulated and
median keel;
POL = 2,5;
temples;
2,5; pronotum shiny,
legs testaceous,
mid and hind
long as broad;
reaching only
posterior collar short, pronotum
fore
and tibiae partly brown;
segment 4 nearly five times as with numerous
head and antennae testaceous;
except f0r posterior margin and sides of
crossed by a
a strong posterior transversa I pronotal
occipital
Ol = 2,5;
anterior
impression;
tubercles not reaching
scutum shiny,
OOl = 14;
strong
tegulae;
strongly sculptured by numerous
longitudinal keels; notaul ices invisible among keels; scutellum and metanotum dull, reticulate rugose,; propodeum reticulate rugose with 2
Dryininae
Fig.
576 -
Chela of Dryinus alvarenganus n.
853
sp.
(type)
854
Mem.
Fig.
incomplete
577 -
longitudinal
transversa I bands, pterostigma;
Amer.
segments
Inst.,
37,
keels on posterior surface;
one on
1984
sp.
(type)
fore-wing
with 2 dark
the basa I cell s and another beneath the
distal part of radial
vein
longer than proximal
in following proportions:
in following proportions: 40:
with a subapical
no.
Chela of Dryinus sinopensis n.
fore tibiae and trochanters 575)
Ent.
4:
60:30;
part
(15:8);
fore tarsal
11: 27: 45; enlarged claw (Fig. segment 5 of front tarsus
tooth and 16 lamellae;
Dryininae
Fig.
(Fig.
575)
578 -
Chela of
Dryinus belizensis
with 2 rows of nearly 40 lamellae;
20 lamellae;
tibial
spurs 1,1,2.
855
n.
sp.
(type)
apex with a group of nearly
856
Mem. Amer.
Fig.
579 -
Che-Ia of
En!.
Ins!.,
no.
37,
1984
Dryinus antillean us (Evans) (type)
MALE: unknown LOCUS TYPICUS: Jujuy (Argentina) TYPICAL MATERIAL: holotype F! in BA. HOSTS: unknown DISTRIBUTION: Neotropic region: ARGENTINA:
Jujuy,
BA!
Dryininae
Fig.
857
Heads of females of Dryinus teutoniae n.
580 -
catarinae n. sp.
sp.
(type) (A)
and
(type) (B).
12. Dryinus alvarenganus n. sp.
FEMALE:
fully
winged;
antennae testaceous;
length 6,25 mm;
propectus black;
head
testaceous,
for sides of pronotum testaceous;
abdomen brown;
distally thickened,
very slender;
antennal
proportions:
60:
12: 5:
irregularly striate;
25:
frons
21:
14:
with vertex black;
thorax and propodeum black,
10: 9:
legs testaceous;
except antennae
segments in following 7:
9;
with a median keel;
head dull,
occipital
granulated and
carina
incomplete,
only visible behind the ocelli and on sides of ocellar triangle; temples invisible; POL = 3; OL = 4; OOL = 10; OPL = 2,5; pronotum with a strong anterior transversal impression; reaching complete, scutum;
tegulae;
posterior collar short;
scutum dull,
posteriorly separated, scutellum punctate,
reticulate rugose;
with numerous
pronotal
tubercles not
longitudinal
with areolae along margins;
fore-wing
cells and another beneath
keels;
notaulices
weakly distinct near posterior margin of
propodeum reticulate rugose,
on posterior surface; basal
impression and with a strong posterior transversal
disc humped;
metanotum
with two
longitudinal
with two dark transversal the pterostigma;
bands,
keels
one on
distal part of radial
longer than proximal part (25:15); fore tarsal segments in following proportions: 32: 5: 10: 29: 45; enlarged claw (Fig. 576) with a subapical tooth and with 25
lamellae;
segment 5 of front
rows of nearly 33
lamellae;
apex with a group of nearly 16 lamellae;
proportions of fore tibiae and trochanters: MALE:
60:28;
(Fig.
tibial
576)
Represa do Rio Grande
TYPICAL MATERIAL:
holotype F!
HOSTS: unknown DISTRIBUTION: Neotropic region: (Guanabara),
(Brazil)
in TW. BRAZIL:
Represa do Rio Grande
TW!
the species is named
in honor of Mr.
M.
with two
spurs 1,1,2.
unknown
LOCUS TYPICUS:
NOTES:
tarsus
Alvarenga.
the
vein
858
Mem.
Amer.
Ent.
Inst.,
no.
37,
1984
13. Dryinus sinopensis n. sp.
FEMALE:
fully
spots on si des;
head testaceous,
propectus testaceous;
mesothorax,
legs testaceous; proportions:
length 5 mm;
winged;
antennae testaceous;
pronotum
metathorax and propodeum black;
antennae distally thickened;
12: 7:
33:
with vertex black;
testaceous,
16:
12:
11:
10: 9:
antennal
8:
12,5;
wi th
two
abdomen
and
segments in following
head dull,
granulated,
with numerous longitudinal keels on vertex and frons; frontal I ine complete; occipital carina incomplete, only visible behind the ocell i and on sides of ocellar triangle;
temples
pronotum shiny, transversal invisible;
invisible;
impressions; pronotal
2;
anterior collar
=
OL
long;
tubercles not reaching
sculptured by numerous parallel among keels;
=
POL
3;
with numerous striae around disc,
metanotum dull,
longitudinal
=
disc humped;
tegulae;
granulated;
OOL
=
10; OPL
crossed by
1;
two strong
posterior collar
scutum and scutellum
keels;
notaul ices not distinct
propodeum fully reticulate rugose,
with two longitudinal keels on posterior surface; fore-wing with two dark transversal bands, one on the basal cells and another beneath the pterostigma; (16:15); tarsal
distal
fore
part of radial
vein slightly
segments
in following proportions:
segment 4 more than five times as (Fig.
577)
tarsus
577)
with 2 rows of nearly 35
to the apex; tibial MALE: unknown LOCUS TYPICUS:
3,5:
30:20;
part fore
9: 22: 36; antennal (16:3);
enlarged claw segment 5 of front
lamellae extending continuously
spurs 1,1,2.
12° 31'
TYPICAL MATERIAL: HOSTS:
27:
long as broad
tooth and with 9 lamellae;
with a subapical
(Fig.
longer than proximal
tibiae and trochanters in following proportions:
S 55° 37'
holotype F!
W (Sinop,
Mato Grosso,
Brazil)
in TW.
unknown
DISTRIBUTION: Grosso),
Neotropic region:
BRAZIL:
12° 31'
S,
55° 37'
W (Sinop,
Malo
TW!
14. Dryinus belizensis n. sp.
FEMALE: brown,
fully
winged;
length 5,62 mm;
with segments 1-2 testaceous;
meta thorax and propodeum black;
head reddish-testaceous;
prothorax reddish;
abdomen and
antennae
mesothorax,
legs redd i sh-testaceous;
antennae distally thickened; antennal segments in following proportions: 13: 7: 38: 20: 16: 13: 12: 10: 9: 12; antennal segment 4 nearly six times as long as broad;
head dull,
the sides of ocelli; and on POL
=
flat,
occipital
the sides of ocelli; 2;
OL
=
2,5;
OOl
=
granulated,
carina
occipital 12; OPL
with some
incomplete,
=
carina 1,5;
longitudinal
keels on
visible behind the ocelli
invisible behind the temples;
pronotum humped,
shiny,
with
numerous keels around disc,
with strong anterior and posterior transversal
impressions;
invisible;
posterior collar
pronotal
tubercles not reaching
Dryininae
Fig.
tegulae; clearly
581
-
Chela of
Dryinus teutoniae n. sp. (type)
scutum sculptured by numerous visible among keels;
859
longitudinal
keels;
notaulices not
scutellum sculptured by numerous
longitudinal
keels; metanotum smooth, dull, granulated; propodeum reticulate rugose; posterior surface of propodeum with two incomplete longitudinal keels; forewing with two dark transversal beneath part
the pterostigma;
(19:17);
fore tarsal
fore
subapical
one on the basal vein
in following proportions:
tarsus produced into hooks;
tooth and with
with 2 rows of nearly 51
13 lamellae;
29:
4:
10: 24:
enlarged claw
segment 5 of front
lamellae extending continuously
spurs 1,1,2. MALE: unknown LOCUS TYPICUS:
Middlesex
TYPICAL MATERIAL: HOSTS:
(Belize)
holotype F!
in OT.
unknown
DISTRIBUTION:
cells and another
longer than proximal
tibiae and trochanters in following proportions:
segments
and 3 of front
bands,
distal part of radial
Neotropic region:
BELIZE:
Milddlesex,
OT!
40;
(Fig. tarsus
34:23;
segments 2 578)
with a
(Fig.
to the apex;
578) tibial
860
Mem. Amer. En!.
Ins!., no. 37,1984
Fig. 582 - Chela of Dryinus catarinae n. sp.
15. Dryinus antillean us (Evans) n. comb.
(type)
Dryininae
861
Mesodryinus antilleanus Evans 1969: 11.
FEMALE;
fully
winged;
mandibles testaceous;
length 3,60-4,00 mm; antennae testaceous;
with sides and posterior collar reddish; propodeum black; thickened; (11:1,5);
abdomen brown-reddish;
head dull,
hairy,
pronotum du II,
transversal
POL
granu I a ted,
impression and by
posterior collar short, pronotal
granulated,
carina complete; a
distinct;
tubercles not reaching
=
region;
pronotum black,
metathorax and antennae distally
10: 6:
23:
11:
9:
9;
times as long as broad
with some 4,5;
OL
=
longitudinal
3; OOL
=
keels on
7,5; OPL
=
1;
crossed by a posterior strong
weak anterior transversal
impression;
anterior collar less visible; tegulae;
scutum dull,
numerous areolae on posterior half of lateral margin of central
with clypeus and
legs testaceous;
antenna I segment 4 nearly seven
occipital
TL = 4;
mesothorax,
antenna I segments in following proportions;
8: 6: 5: 7; frons;
head black,
propectus black;
disc humped;
granulated,
with
regions and near posterior
notaul ices incomplete,
distinctly visible on
anterior half of scutellum, weakly visible on posterior half, because hidden among areolae; scutellum and metanotum dull, granulated; propodeum reticulate rugose, surface very
without
inclined;
longitudinal
fore-wing
distal
part of radial
tarsal
segments in following proportions:
front
vein much
keels on posterior surface;
with 2 brown transversal longer than proximal
tarsus produced into a hook;
18: 2:
5:
enlarged claw
part
13: (Fig.
(13:7);
fore
segment 3 of
579) with a
(Fig. 579) with 2 rows of 40 lamellae extending continuously to the apex; fore tibiae and trochanters in following proportions: 31:18; tibial spurs 1,1,2. MALE: fully winged; length 3,00 mm; head black, with mandibles yellow; antennae brown; abdomen black; legs yellow, wi th fore and hi nd coxae partly black; antennal segments in following proportions: 6: 4,5: 12: 9: 10: 10: 8: 8: 7: 7; head dull, granulated; occipital carina incomplete; POL = 5,5; OL = 3; OOL = 5; OPL = 1; TL = 1,5; frontal line complete; scutum dull, granulated; notaulices incomplete, extending to beyond 0,65 length of scutum; scutellum dull, granulated, except for a narrow region near metanotum which is shiny, smooth and without sculpture; metanotum smooth and without sculpture; propodeum reticulate rugose, without longitudinal keel s on posterior surface; fore-w i ng wi thout dark transversa I bands and with distal part of radial vein much longer than proximal part (11 :6); tibial spurs 1,1,2. LOCUS TYP ICUS: Mouth of Layou River (Dominica) TYPICAL MATERIAL: holotype F! and allotype M! in WA; 1 paratype F! in CM. HOSTS: unknown DISTRIBUTION: Neotropic region: DOMINICA: Mouth of Layou River, WA! South Chiltern, CM! subapical
tooth and 13 lamellae;
segment 5 of front
19;
posterior
bands and with
tarsus
862
Mem. Amer.
Fig.
583 -
En!.
Inst.,
no.
37,
1984
Chela of Dryinus kimseyae n.
sp.
(type)
16. Dryinus teutoniae n. sp.
FEMALE fully winged; length 4,50 mm; head black, with mandibles brown; antennae testaceous; thorax and propodeum black; abdomen black-brown; legs testaceous, with coxae partly brown and clubs of hind femora black; head (Fig. 580 A) dull, hairy, convex, reticulate rugose; occipital carina complete; posterior ocell i not touching occipital carina; OPl = Ol; Ol
Dryininae
Fig.
584 -
863
Female of Dryinus palaeodominicanus Currado and Olmi
(type)
shorter than POL; pronotum dull, granulated, crossed by a weak anterior transversa I impression; anterior coil ar weak; posterior transversa I impression strong; posterior collar short, distinct; scutellum fully reticulate rugose; notaulices weakly visible, incomplete, reaching approximately 0,5 length of scutum; scutell um and metanotum ret icu I ate rugose; propodeum reticulate rugose, without longitudinal keels on posterior surface; fore-wing without dark transversal bands and with distal part of radial vein longer than proximal part (15:7); segment 1 of front tarsus longer than segment 4 (20:12); segments 2 and 3 produced into hooks; enlarged claw (Fig. 581) with subapical tooth and 16 lamellae; segment 5 of front tarsus (Fig. 581) with 2 rows of 37 lamellae extending continuously to the apex; tibial spurs 1,1,2. MALE: unknown LOCUS TYPICUS: Nova Teutonia (Brazil) TYPICAL MATERIAL: holotype F! in BM. HOSTS: unknown DISTRIBUTION: Neotropic region:
BRAZIL:
Nova Teutonia
(S.ta Catarina),
864
Mem.
Amer.
En!.
Inst.,
no.
37,
1984
BM!
17. Dryinus catarinae n. sp.
FEMALE:
fully
winged;
length 4,50 mm;
head black,
with clypeus and
mandibles reddish; antennae testaceous; propectus black; pronotum black, with posterior collar reddish; mesothorax, metathorax and propodeum black; abdomen brown; B)
shiny,
carina complete; than OL;
legs testaceous,
granulated,
with coxae partly brown;
with some longitudinal
posterior ocelli
OL shorter than
POL;
head
keels on frons;
not touching occipital pronotum dull,
carina;
granulated;
(Fig.
580
occipital POL shorter
anterior
transversa I impress ion and anterior coil ar weak; posterior transversa I impression strong; posterior collar short, distinct; pronotal tubercles not reaching tegulae; scutum shiny, fully reticulate rugose; notaulices weakly visible, incomplete, reaching approximately 0,5 length of scutum; scutellum and metanotum shiny, granulated; propodeum reticulate rugose, without longitudinal keels on posterior surface; fore-wing with 2 brown transversal bands and with distal part of radial vein longer than proximal part (13:8); segment 1 of front tarsus longer than segment 4 (18:12); segments 2 and 3 of front tarsus produced into hooks; enlarged claw (Fig. 582) with subapical tooth and 16 lamellae; segment 5 of front tarsus (Fig. 582) with 2 rows of 42 lamellae extending continuously to the apex; tibial spurs 1,1,2. MALE: unknown LOCUS TYPICUS: Nova Teutonia (Brazil) TYPICAL MATERIAL: holotype F! in BM. HOSTS: unknown DISTRIBUTlmj: Neotropic region: BRAZIL: Nova Teutonia (S.ta Catarina), BM!
18. Dryinus kimseyae n. sp.
FEMALE: spot on
fully
winged;
ventral
length 10,8 mm;
head black,
with a broad yellow
side near mouth-parts and with anterior region of frons
near clypeus testaceous; clypeus testaceous with a brown spot; mandibles testaceous, with teeth brown; antennae brown-reddish, with segment 1 yellow;
thorax and propodeum black,
testaceous; brown;
abdomen brown;
antennae distally
proportions:
24:
11: 90:
nearly nine times as
weakly 33:
keel;
occipital
carina
30:
thickened; 21:
long as broad
with numerous areolae and
with posterior margin of pronotum
legs testaceous, 16:
15:
(35:4);
irregulae keels;
incomplete;
POL
= 4;
with anterior tibiae partly antennal 12:
17;
segments in following antenna I segment 4
head flat, frons OL
=
dull,
granulated,
with a complete median 4;
OOL
=
17;
OPL
=
2;
Dryininae
Fig.
Dryinus palaeodominicanus Currado and Olmi (type)
585 -
occipital
carina not visible behind the eyes;
transversal collar
(in
strong;
numerous weakly
impression, lateral
with disc nearly
view:
50:14);
posterior collar short,
reaching
longitudinal tegulae;
scutell um dull,
distinct;
keels,
keels; basal
cells
keels;
pronotum shiny,
hairy,
fore-wing
is very
with
posterior tubercles not
longitudinal
keels;
notaul ices
except for the posterior half not metanotum dull,
granulated;
surface sculptured by numerous strong
posterior surface with
transversal
long as anterior impression
but complete and posteriorly separated;
reticulate rugose,
with dorsal
times as
keels on collar and disc;
scutum with numerous
visible among
pronotum crossed by a strong
three
a posterior transversal
reticulate rugose and granulated; shiny,
865
two with
longitudinal
propodeum
longitudinal
keels and with numerous
two dark transversal
and another beneath the pterostigma;
bands,
distal
one on
the
part of radial
vein longer than proximal part (43:22); radial cell open; fore tarsal segments in following proportions: 50: 7: 14: 45: 72; enlarged claw (Fig. 583) front
with a tarsus
subapical (Fig.
583)
tooth and a with
row of nearly 28
two rows of nearly 45
lamellae; lamellae;
segment 5 of apex with a
866
Mem. Amer.
group of nearly 25 MALE:
lamellae;
Ent.
Ins!.,
tibial
no.
37,
1984
spurs 1,1,2.
unknown
LOCUS TYP I CUS:
Barro Co lorado
TYPICAL MATERIAL: HOSTS:
I.
holotype F!
(Panama)
in DA.
unknown
DISTRIBUTION:
Neotropic region:
PANAMA:
Barro Colorado
I.
(Canal
Zone),
DA! NOTE:
the species
is named in honor of Mrs.
Lynn S.
Kimsey.
19. Dryinus palaeodominicanus Currado and Dlmi
Dryinus palaeodominicanus Currado and Olmi 1983: 330.
FEMALE
(Figs 584,
585):
brown-testaceous and weakly
thickened;
fully winged;
legs
light;
length 6 mm;
black,
antennae 10-segmented,
densely hairy;
with mandibles
slender,
antenna I hairs very short,
distally
much shorter
than breadth of segments, weakly visible through the amber; antennal segments in following proportions (length): 12: 7: 39: 19: 18: 13: 8: 8: 6,5:
6,5;
aantennal
segments in following proportions
(breadth):
3:
2:
2:
2,5: 3: 3: 3: 2,5: 2: 2; antennae more than three times as long as head (length of head in dorsal view from occipital margin behind the ocelli to apex of mandibles):
137:40;
antennal
sockets touching margins of clypeus;
clypeus with ventral margin strongly bidentate; mandibles with 4 teeth progressing larger from anterior one to posterior; eyes apparently bare, except for some
long hairs;
of eye to base of mandibles; head,
where
it
malar space with a furrow joining occipital
is directed towards
carina
distinct only on
the upper corners of eyes;
carina not visible on sides of head and on ventral
lower corner dorsal
side of
occipital
side because of the
thickness of the amber; temples not prominent, absent; eyes normally protruding; POL; 5; OL ; 3; DOL; 9; OPL ; 1; frons with a complete median
keel;
vertex and frons apparently rugose,
pronotum crossed by an
anterior weak
anterior collar and the disc; strong posterior transversal
disc humped;
posterior collar very short,
(3:11);
pronotum fully with
granulated;
reticulate rugose; impression between
the
pronotum also crossed by a
impression between
collar; (36:40),
transversal
the disc and the posterior
much shorter than anterior collar pronotum slightly shorter than
posterior angles produced
into
lobes directed towards
head the
tegulae, but not reaching tegulae; sides of pronotum contracted where they meet the scutum; posterior transversal impression and posterior collar with a
distinct dorsal
median carina,
granulated and reticulate rugose, reaching
approximately 0,65
apparent I y scutell um,
scutum shorter than pronotum with
notaul ices incomplete,
lenth of scutum;
as scu I ptured as scutum,
scutellum and metanotum
shorter than scutum;
metanotum and pronotum usua I for
(22:36),
distinct,
shape of scutum,
Dryininae; propodeum not
Dryininae
Fig.
easily
586 -
867
Dryinus paJaeodominicanus Currado and Olmi (type)
Chela of
distinct through
the amber,
because of the thickness of the amber
and because the abdomen
is hiding
dorsal
reticulate rugose;
surface apparently
long as posterior' surface; on basal
cells and on distal
venation of the (30:4);
radial
proximal basal
part
closed;
(50:8);
very slender,
legs
in proportions:
(tarsal
dorsal
surface apparently as
two broad dark transversal
half of wings;
distal
radial
fore-wing
part of radial
vein s-shaped; (costal,
without dark bands,
wing
(tarsal
with
with
bands
the normal
Dryininae; pterostigma narrow, much longer than broad cell
cells clearly enclosed
wings hyaline,
21
fore-wing
the posterior surface of propodeum;
with shape usual 35
segment 1):
segment 4):
34
(coxae): 3
(tarsal
(tarsal
segments 2 and 3 of front
median
with
longer than the usual
and submedian cells);
three
hind
for Dryininae; foreDryininae; segments of the fore
with shape usual for
(trochanters):
segment 2):
segment 5):
trochanters nearly four times as fore tronchanters with a
40
vein much
fore-wing
8
34
long as broad
long proximal
62
(tarsal
56
24
fore
(greatest breadth):
into hooks;
(tibiae):
segment 3):
(enlarged claw);
slender stalk,
tarsus produced
(femora):
40:
9,5;
distally broadened; rudimentary
claw
868
Mem.
apparently present
31
lamellae;
central (tarsal
(34:8),
(tarsal
proportions:
25 (coxae):
segment 1): 11
(Fig.
1984
30
segment 2):
segment 5):
maxillary palpi
tibial
labial
43
49
(tibiae):
segment 3):
for
(femora):
spurs 1,1,2; segment very for
28 5
(tarsal
legs in following
15 (tarsal
length and breadth of the wings usual
morphology of the body usual
row
legs in following
(femora):
74 (tibiae): segment 3): abdomen
apparently with 6 segments;
last
much
lamellae located around a
segments of hind
segment 2):
586)
with 2-3 rows of nearly
10 (tarsal
10 (trochanters):
19 (tarsal
(Fig.
tooth and with a
586)
segments of mid
(trochanters):
apparently with 3 segments;
MALE:
37,
enlarged claw
tarsus
segment 5);
(tarsal
distinct petiole:
5
15 (tarsal
8
shape,
distinct);
lamellae;
14 (coxae):
segment 1):
segment 4):
no.
apex with a group of nearly 30
segment 4): (tarsal
Inst.,
with a strong subapical
segment 5 of front
hollow without
proportions:
Ent.
(but weakly
longer than arolium of 16 lamellae;
Amer.
31
7 (tarsal
without a
labial
palpi
long and slender;
Dryininae; shape and
Dryininae.
unknown
LOCUS TYP I CUS:
Dom i n i ca amber.
TYPICAL MATERIAL: HOSTS:
DISTRIBUT ION: NOTES:
holotype F!
in TR.
unknown Neotropic region:
only known from Dominica amber.
Dryinus. It Dryinus. The only species of Dryinus known from
this species shows morphologic characteristics of living
is belonging to the genus
D. antillean us (Evans), clearly distinct. D. palaeodominicanus is the only fossi I Dryinid known from Dominica
Dominica is
amber and from
the Neotropic region.
DRYINUS
AUSTRAL I AN SPEC I ES OF
Twenty
three species are known.
KEY TO THE SPECIES
FEMALES
Fore wings without
dark
........................ Fore wings with dark 2
Propodeum not (transversal
bands •••••••••••••••••••••••
1. pallidus (R.C.L. Perk i ns)
transversal
reticulate or
transversal
rugose,
longitudinal)
bands ••••••••••••••••••••••••• 2 with numerous parallel
keels at
least
on anterior half;
sometimes with weak areolae among keels •••••••••••••••••••••••• 3 Propodeum reticulate 3
rugose at
least
Propodeum with numerous parallel
on anterior half •••••••••• 5
longitudinal
keels .••••••••••••
Dry i n i nae
869
2. speciosus (8odd) Propodeum with numerous parallel
4
Pronotum wi th posterior nearly
three
species
times as
transversal
transversal
long as pronotal
reddish-testaceous,
tubercles
6
subtilis n.
darkened •..•••••..••••.
3.
Pronotum with posterior
transversal
7
carina
incomplete ••••••••••••••••••••
carina
complete .••••..•••
excava t ed ••••••••• swol len ••••••.••••
Occipital
carina
sides
of
• ••• 7 6.
ocellar
triangle;
...................•... fully fully
7.
the posterior ocelli;
head
pseudophanes (R.C.L. Perkins)
carina visible behind the
.•••.••.••••.. 6 . ••••••••••.•• 8
montanus n. sp.
only visible behind
granulated ••••••••••••• Occipital
5.
nearly
species mostly
Occipital Head
589 A);
sp.
impression narrow,
Occipital Head
(Fig.
wi th petiole black and abdomen
as long as pronotal tubercles (Fig. 589 B); b I a ck • • • • • • . • • • • . • • • • •• 4. pacificus n. sp.
5
keels •••••..••••• 4
impression very broad,
the posterior ocell i head punctate,
not
and
on
the
granulated
insularis (Dodd)
8
Head Head
or mostly reddish • • • . . • • • • . . • • • • • . • • • . . . • • • . • • • • . • • 9 or mostly black •••••••••••••••.•••..••..•••.•.•••• 10
9
rugose, without longitudinal 8. bouceki n. sp. Scutum reddish, rarely black, not reticulate rugose, with numerous longitudinal keels ••..••••.•••••.••••.••• . . . • • . . • • • • . . . . • . . . . • . . 9. koebelei (R.C.L.Perkins) Frons and vertex almost fully sculptured by numerous strong longitudinal irregular keels •••.••••.••••••••••..••••..•••..• 11 Frons and vertex without longitudinal keels or with a few keels on a narrow surface •••••••••..••••.•••••.••••.••••••••••••••• 12 Posterior ocelli touching occipital carina ••••••••••••••••••••• •••••••••••••••••••• 10. guineensis n. sp. Posterior ocelli not touching occipital carina •••••• •.••....•••...••.•••.. 11. gigas (R.C.L. Perkins) Metapleura wi th a narrow region smooth, shiny and hairless, without sculpture; pronotum, sometimes also head and scutum, fully smooth, shiny, hairless, without sculpture, sometimes only with Some keels on sides of pronotum .••••••••••••••••••• 13 Metapleura without region smooth, shiny and hairless; pronotum, head and scutum usually not smooth, shiny and hairless ••••••• 15 Head ex cava ted. • • • • • •• 15. montanus n. sp. Head flat •••••••••••••••••••••••••••••••••••••••••••••••••••• 14 Scutum shiny, smooth, without sculpture •••••••••••••••••••••••• •••••••••••••••••••••• 13. aterrimus (Dodd) Scutum dull, with numerous longitudinal keels •••••••••••••••••• • • • • • • • . • • • • • • • • • • • • •• 14. biaki n. sp. Scutum almost fully reticulate rugose •••••••••••••••••••••••• 16 Scutum not reticulate rugose or with a few areolae (sometimes
Scutum black,
fully
reticulate
keels ••••••••••••••••••
10
11
12
13 14
15
870
Mem.
on I y
n ea r
Amer.
Inst.,
frons and wertex not
punctate;
no.
37,
1984
ma rg in) •••••••••••••••••••••••••••••••••• 1 8
po s t e rio r
16 Pronotum,
Ent.
pronotum wi th
15. punctatus n.
· . . . . . . . . . . . . . . . . . . . .. Frons and vertex as
reticulate
rugose;
f ron s
smoo t h ,
weak keels around disc ••
reticulate
sp.
rugose as
scutum •••••••••••••• 17
17 Pronotum as reticulate rugose as scutum ••••••••••••••••• • • • • • • • • • • • • • • • • • • • • ••
16.
areolatus n.
sp.
Pronoium with keels around disc, not as reticulate scutum •••••••••••••••• 17. leptias (R.C.L. Perkins) 18 Scutum without
keels
or with
very
short
keels near
rugose as scutellum
• •••••••••••••.•••.••••••.•••••••••.••••.•.•••••••••••••••••• 19 Scutum almost
fully
19 Posterior ocel I i
sculptured by
touching
• • • • • • . • . . . • • • . . • . . • •. Posterior
ocelli
not
1 8.
20 Pronotum smooth,
19.
shiny,
carina;
TL
keels ••••••••• 20 shorter
than POL
qua te i n . sp.
touching
POL. • . • • . • • • • • • • • • • • ••
longitudinal
occipital
occipital
carina;
TL
longer
than
dahmsi n. sp.
almost
hairless,
wi thout
sculpture or
wi th a few keel s around di sc ••••••••••••••••••••••••••••••••• 21 Pronotum not smooth and shiny, sculptured by numerous keels around d i s c . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .
• ••••• 22
21 OPL = POL; posterior collar of pronotum yellow.... • • • • • • . • . . • • • . . • . • • • •. 20. glaber n. sp.
• •••••••
Posterior POL;
ocell i
posterior
very
near occipital
collar of
carina;
OPL
shorter
than
pronotum black • . • . . • • • . . • • • • • • . • • • • • . •
• .•.••.•.•..•..••••••• 21. australianus n. sp. 22 Frons and vertex reticulate rugose •••••••••••••.•••••••••••••••
. . . . . . . . . . . . . . . . . . . . . . 22. venator (R.C.L. Perkins) Frons and vert ex not
ret i cu I ate
rugose,
punctate and wi th
some
keels •••••••••••••••••••••••••••••••••••....•••.••••••••••.•• 23
23 Posteri or ocell i farther from the occipi tal carina (Fig. 598 B); frons and vertex smooth, hai rl ess, finely punctate, wi th weak
keel s
numerous
connect ing
ocell i
to
longitudinal
keels,
punctate ••••••••••••••••
• • • . • • • • • • • • • • • • • . . . •• Posterior ocelli and vertex connecting
nearer
1 2.
frons;
bismarcki n.
occipital
scutellum wi thout sp.
carina
(Fig.
598 A)
smooth, hairless, weakly punctate, without ocelli to frons; scutellum with numerous
longitudinal
kels •••••
23.
papuanus n.
frons keels
sp.
MALES
Notaulices complete ••• Notaulices