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1.8. f & ~ H H O y~ Y e 6 ~ 0 n e o c o 6 ~ eC O A e P X M T C R e l e H M R 0 HFOT.1 O X H b l X COCTORHMRX. ~ a M 6 0 . ~ l eYHC'PO e BCTPeYaIOUuXCST Y 6 0 1 1 h l l b 1 ~3HAOKPI3HHhlMII an6oneRBHMRMII ~IOLIPO~IIO P a C C M O T P e H b I BOnPOCh1 IIMRI'HOCTMKM J I .I e Y e H M R OCTp b ~O x C ~ O K H ~ H ACG~ X ~ ~ I I O nI r' .Or a 6 e r a . T n p e o T o K c w e c K o r o ~ p r u~r n an o ~ r i P ~ O M A H O G~ 0 h l b 1 . H a n n o q e w x I i o n o r o r n n o a a p e l - l a n o n o r o M m T e x o n a x w e o BOI'O K P M 3 0 B . a T R K X e OCTPhlX H ~ P Y I U P I (I ~ ~ )~ o~ c~( ~ ) o P H o - K ~ . ~0~6~h I~I e~ ~Bao ~ o
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0ABTO~CK K OM n J I~e K~T M B . 2006 0O a o p ~ n e ~.n000 e (~M~AI?UHHCKO~ M H @ O ~ M ~ U I ~ O H H Oa ~r e? l i T C T B O * .
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1.1.4. 1.1.5.
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53
Aria n p a K T M s e c m i x qene5 B c e K O M ~ M I o r y T B ~ I no~pa3;[ene~br T ~ qepeBpanb~bre,a ~ a o ~ o r c c m e c r c u ae w, o T o r c c r m e c K u e II K O M ~ npyI r0%3THOJlOTH11. ~ O C K O J I ~ XKO>TaT03HbIe Y COCTORHIlFf R 3BBIICMMOCTX OT RTE10.10rEIM T ~ ~ ~ V IP O R ~T. ~ I P I Y H O T e~p~a r I H M , a TfI>Ke.XOe COCTOffHlIe Ha
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IIpMZlHHaMII. X a p a K T e p ~ b ~ e
O C O ~ ~ H H O C THak160.lee H p a C I I p O C T p a H e H H b I S KO\lRTO3FIbIX COCTORHM?~
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rax
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55
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1. AKMMOB r.A., O A I T H ~ M.M. K MHTepeC IIpaKTMseCKHX Rpaclefi K n p o 6 n e ~ eAIiEirHOCTMKM M JIe4eHMR HeOTJIOXHbIX C O C T O R H M ~OCTBeTCX ~ C T ~ ~ ~ I J IRbICOKlIM. ~ H O 3HaHM.le TOW IIp06JIeMb1 H ~ O ~ X O ~ MBpa.Iy M O n106oW CneuMa~lbHOcTM. B IIpenCTaBjreHHbIX BaILIeMy BHMMRHMIO MeTOnH4eCKMX PeKOMeHfiaUMRX aBTOPbI CAenanPi nOIIb1TK)T o606wrr~bM CMCTeM~TIT3PIpOBaTb COBpe~kleHHbIellOAXOAbI K AMarHOcTI?Ke M JIe4eHtllO He0T~10iKHb1X COCTORHI?W, BbI3BaHHbIX p~3jIMZIHbIMA~ ~ ~ O ~ I ~ B ~ H3HnOKPMHZ I R & ~ M HOW CPICTeMbI. B C B O p~a ~6 o ~ eM b I CTapa.rIIICb MCIIOnb30BaTb !'CTRHOBMBILlMeCR Ha C ~ ~ O A H E I I I I H AeIIb M~ B K ~ H H H Y ~ C K OnpaKTL1rtC % KpI$TepMM I? nOHRTMR. H ~ A ~ ~ M9 TcO R3 T,M PeKOMeHAaUZIM 6 3 1 IIOjre3HbI ~ ~ ~ He TOjrbKO CTYAeHTaM IV-VI KypCOB MeAHUMHCKMX BP30B, M3y9aIOW l I M 3II~OKpMHO~OrI~I0, H O J f 3IInOKPMHOjIOraM, TepaIIPBTaX, RHeCT ~ ~ M O J T O M~ ~peaHrmaTo.lora31, M a TaKme s p a s m ~ p y r r f xcneqxa-UbHoc~efi,lIHTepeCym4MMCR R011pOCaMll JJMarHOCTMKMI1 J7eqeFIMR HeOTJlOjrCHbIX C O C T O R H M ~3HAOKPMHHOTO ~ TeHe3a. M ~ 6I y n e ~npM3HaTe~TbHbITeM HaILIMN ~IITTaTeXRM,KOTOPbIe 3BXOTRT IlOAe.lIMTbCX C HBXI.1 C B O M M M R3rJIRAaMM M ~ 0 0 6 p a j ~ e ~ M F I ~ 1 . 1 IIO TOW n p o 6 n e ~ e .I C p ~ ~ ~ s e c~~ ~r rMe ~ ~ IKonner. ~ H M HanpasneHR Hble Ha 6onee ~ @ @ ~ K T H B HpeLUeHMe O~ 3aTpOlIYTbIX HBMI? npo6nex1, 6 y n y ~IIpEIHRTbI EtBTOpaMM C ~ J I ~ ~ O ~ ~ ~ H O C T ~ ~ O .
~ M @ H AIlarHOCTMKa ~ H HepBHbIX 60ne3~eW:PYKOBOACTBO AjIX Bpa~er'r.M ~ A 2-e, . HCnp a s n e m o e PI AononHeHHoe. - Cn6.: r s n n o ~ p a ~2000. , 664 c. 2. B a n n 6 0 . ? 1 c ~M.H. ~ n ~ a 6 e ~ o r r or ~M.: ~ . M ~ A H ~ H2000. H~, 672 c. 3. 6aJIa60.lI~1.1~ M.M. S ~ n o ~ p n ~ o n or rM.: ~ ~ Y, ~ r < s e p c yna6~ nr?rur?Hr, 1998. - 582 c. 4. F a n a 6 o n ~ 1 ?M.R., ~ ICneGa~osaE.M., I C p e ~ ~ 1 - 1 c ~B.M. afl n ~ c b ~ ) e p e ~ ~ M a n h HfiMarIrOCTAKa aR PI JIeYeHMe 3HnOKPIIHHbIX 3a60.7eB ~ H M W : ' ~ v K O B O ~ C T B O. M., 2002. - 752 C. B M I~? H T ~ H C P ~ I? RH HeOTXOXHaR ~R TepanLiR B 3 H 5. E O ~ A ~ H OB.X. AOKpMH0,TOrPII.r. - H. Hosropon, 2000. - 324 C. 6 . Boronenon H.K., r y c e ~E.M., ? K > l y p ~ aB~. n . r i ~ p K. o > ~ a : B 3 0 T. / n o 2 pen. F.B. n e ~ p o e c ~ o Ir /o B KH. EO.TSUI~R MeaxUMHCKaR 3HIlHK~3OIIeJMR.- 3-e Man. M.: COB.3HUMK.TOnegrlR, 1979. - T. 11. - C. 515-580. 7. 6 o n e a ~ k wep~aoi? i crrcTe>lbI: ~ ~ K O B O ~ C TnB. O7 ~~paqeW:B 2 T. / ITon pen. H.H. BXHO,2.P. LLI~yjlb31a~a. - 3-e 1?3n., nepepa6. 1.1 son. - M.: M e n a q a ~ a 2003. , T. 1. - 74.1 c. ~ ~ PYKOBOACTBO AAR Bpaseii: B 2 T. .' 8 . ~ O X ~ ~ HH ~I P? B H OClICTeMbI: n o n pen. H.H. BXHO,A . P . L U T ~ , T ~ M-~ 3-e M ~n3n., . nepepa6. M ,Ion. - M.: ~ ~ ~ A M u 2003. M H ~ T. , 2. - 512 '2. 9. r a a x a p a II.C., HOBHKOB A.A. Y s e 6 ~ o enoco61.1eno Knnrrnrec~ 0 fTOKClIKO.lIOrMM. i - M.: M ~ L [ ~ ~ 1979. I I H ~-, 336 C. 10.Aenon M.M., M e n b ~ M . r e ~ 1T.A., ~ 0 Q a ~ e eB.B. ~ 3~no~p111ronorMn. - M.: M e f i n q ~ ~2000. a, - 632 C . 11.Aenon M.M., Oa.!iee~ B.B. BBeneHMe B n~a6e~o:1orn10:pyKOB02CTBO Anrr ~paseW.- hI.: M3n-BOn6eperu , 1998. - 200 c. l 2 . n e ~ o nM.M., L U ~ C T ~ K O M.B. R E ~CaxapHblW n~aGe*r: PVKOBOACTBO Ann spa~leii.- M.: YHIlBepcyM na6naruaar, 2003. 455 c. ~ ~ . & A o BM.M., ~ ~ C T ~ M.B., K O BM~~ K C H M M.A. O B ~0 e ~ e p a . T b ~ ~ qenesacr nporpaMMa aCaxap11b1fi~Ma6eTu:>IeTonrmecKMe pexoMeHAaqMlI. - M.: Me~uaccbepa,2003. - 88 C. ~ ~ . A ~ M I ? M.K). A o B ~OCTpb1e OCJIomHeHMR HHCynMHO3aBMCMMOrO caxapHoro ~ M a 6 e ~/ /a n ~ a 6 e ~ o r p a c P a s r . N! 8. - 1996. C. 17-25. ~ ~ . E @ I ? MA.C., o B C K ~ O ~ O H H.A., C K ~ Rqe6af1 A.H M np. A ~ G y n a TOPHaR I I O M O q b BHAOKPMNNOMY 6 0 j r b H 0 ~ ~/' ~ O Apen. A.C. E @ x ~ o s a. K s e ~ ~: A O ~ O B ' R1988. , - 256 C. 16.Kaaa~~ A.R., e ~ MRTKOBCKPIW B.C. C ~ ~ ~ B O Y no H IMI ~KT @ ~ K U M O H H b I M 6 0 J l e 3 ~~ ~3-e . Man., nepepa6. A AOn. - M.: M ~ A M ~ M Ha, 1985. - 320 c. 1 7 . I C a p n o ~ B.A. H e ~ p o n o r ~ l s ip: y ~ o ~ o ~ cAnsI ~ n spaseir. o M.: MMA, 1999. - 624 c. -
157
C ~ M Cfl!4TepOTypbl OK --
1 8 . K o s a . l e s c ~ aJI.M., ~ J ~ ~ K ~ R H ~ B.C. I I I Kr ~OnBo n a r > a ~ ~~i~rrieo3 1 9 . I C o ~ o s a n A.H., o~ J I n x ~ e p JI.G., ~ a ~ n o ~ a n o nA.A. HefipoTpaR. v a ~ o ~ ? o r l tCr ~n:p a s o ~ r ~ ~-~ PI., t i . 1994. - 416 c. ~ ~ . K ~ ? . c Hn o. M B . Xp~~1~1~f!CKafI i: OcTpa53 Ha~IIO~IellHEII:OBfln HenocTaTocrHocTb B npatiTmce c e ~ ~ e i i t < onpasa ro :'/ Pocc~riicxrriicelcrei?~b~fi Rpacl. - 2002. - 8; 4 . - T . 6. - C. 17-25. 21. K y n e r u o ~E.B., ICyneruo~C.E. Caxap~rbrli~ a a 6 PIe xrrpyprpruec~ KI?F ~ ~ ~ O ; I ~ B ~H M.1 K RB . o c ~ p e c e ~ b 1996. e, - 216 c. 2 2 . ~ ' I y ~ r ~ r rE.A., r r ~ o ~Kocroxaposa J . T . O c ~ p h ~oTpnnsrcar:n: e Pytiononc?Bo 3.i1sr spasefi. >I.: Menrlulr~a,1989. - 432 c. ~~.OKO~O A.13. K OAllarH0c~ill;a E 60.?e3~eiiBH>JTr)eHHIIXOPrAHOR. ~ H ~ ~ H O C Tpf?BMaTI.r~lecKE!S M K ~ M CKCTeNHbIX 3~60.rie~al1l~lfi COe,~MHITTC?J~HOI., 2000. 1'.2. - 6 0 8 c. 2 5 . n o v e p a ~ u e nF3.n. P ? . ~ ~ O ~ O X no C TnRi O r a r n o c r ~ r ~xie :Iecieiiltio nHyTpe:imrx 6 o n e a 1 ~ ~n . 3-e m j . , nepepa6. rr iron. - 31.: r03BYHMLI >I3 P O , 2001. - ,528 c. 2 6 . n o T e ~ l i r rB.B. ~ ~ ~ ~ o l ; p 1 1 ~ 0 . 7 0 r 1y1~rr7e: 6 1 - 1~ ~3-e . 1132.. nepepa6. rr Aon. - hT.: Mea:rrtir~a. 1999. - 640 r.
2 9 . C y ~ 1 . C.A. 1 ~ N[eo.inox~breCocToZi~rin.- 3-e rl3,q.. nepcpab, I? norr. - M.: MMA, 2005. - 7.52 c. 30.GJ;laee~R.B., i v ~ e . ? ~ ~ > ~ ~ rT.A. e r ~ r~; oa r r n o ~ r e ~ ~ r r ~:ezoc.iii~rio~m .r~uTrfloc,rL /,' Pycc~iiiii ~ e , q r r u r t a c ~ r r jicvprra.1. ii -2001. Nb 24. - T. 9 C. 1088-1094. 31 .d)a;zeer? B.H., Me.;r~rr~v~crr~io T.A. ~ M ~ O T M I I ~ PYI~OBOACTHO O ~ : n:rn spaseii. - M.. 2002. - 216 c. 32. X > ~ ~ . ~P. I . /I IHI Z ( ~ ) ( ] ) C ~ ~ IJ[lTHrHOCTllKa - I U ~ ~ ~ ~BtI~TPCHHllX I ~ M R R 60nerr~ei'~: Hep. c Irex. /' - M., 1993. - 794 c. 3 3 . 3 r 1 n 0 1 i p ~ ~ i 0 ~ ~ 0/r c no^ r n p m . H. J I a e x ~ a ;TIep. c a a r n . M . : l l p a ~ r r ~1999. ~ a , - 1128 c . 3 4 . 8 ~A 0 .~ A . , ICpsf~opyqtioM.Q., Bpasesa 3.H. 1.i np. H e p ~ ~ I b l e 6one3~&1 i TIon pen. A.A. Fipoma. - IZPI~B, 1985. - 463 c. : 3.5.ADA Practice R c c o n ~ n ~ e r ~ c l a t i oHvperglycen~ic r~s: crises in paiients w i t h cliabetes m e l l i t ~ ~/s/. Diabetes Care. - 2001. 24 (1). Suppl. L. - S. 83-90. 36.Basic & clinical el~clocrinologv/ Ed.: F.S. Greenspan, J.D. B ~ x t e r .- 4 - t h ecl. - USA: Prentice-Ha11 I ~ t e r n a t i o n a l ,Inc., 1994. - 8 1 1 p. 37.Berger W., Keller U . Treatmel~t.of diabetic kctoacidosis a n d 1 non-ketotic hyperos~nolarconla. / / C l i n . Enclocrinol. ,Vetah. 1992. - Val. 6. - P. 1-22. --
'
38.Faish G.A., Fishbein H.A., Ellis S.E. The epidenliolo~yof tliabetic acitlosis: a populatioll-basecl study. / i A111 J Epidemiol. 1983. - Vol. 117. - P. 551-558. 39.Hol1nan R.C.. Herr011 C.A, S i r l ~ ~ o cP. k Epiclemiologic characteristics of ~ n o r t a l i t yfrom cliabetes with acidosis o r coma. i ; All). J . Publ. H e a l ~ l i .- 1983. -Val. 73. - P. 1169-1174. 40.Joslin's Diabetes Mellitus / Eel.: C.R. Kahn, G.S. Weir. - 13t l etl. ~ - New Delhi: B.I.\Vaverly PYT. LTD., 1998. - 1068 p. 41.Malnlberg K . and DIGAMI Study Grollp. Prospective ranelon]ized study of intensive insulin treatment on long tern1 s ~ i r v i v a l after acute !nyocarciial illfarctioll in p a t i e r ~ t swit!l dii11)etcs ~nellitlis.!i; Brit bled.J. - 1997. - Vol. 314. - P.1522-1515. 42.Marshall S.M., Walker M . , Alberti K.G.M.M. Diabetic kctoacidosis ar~tlhypergl~.caemicnon-kctotic coma. 111: I ~ ~ r . e r ~ i a t i o ~ l a Textbook on Diabetes M e l i i t ~ ~ s2nd . edition .' Ed. K.G.M.31. A!herti, P. Zinlmet, R. De Fronzo. - 1997. - 1-01. 11. - I-'. 1213-1229. 43.3IcAuiag. I-.,Deary I.J.. Frier M . Synptorns of hypoglycwc~~~ia in people wit11 diabetes . ; Diabetic medicille. - 2001. Vol. 18 (9). - P.690-903. 44.Polonsky W.H.. Aliderson B.J. eL al. 111suli!i olilissio:l in women ~ v i t IDDlI l~ . ' , Diabetes Care. - 19S:l. i . 0 1 . 17 ( i 0 ) . - . .~ P. 1178-1.185. 4.5.Princi!1les and practice of el-1r1oc:riilologyant1 ~!~~t;ibolisn!Ed. K.L. B p c k ~ r .- 3-rd pel. - P i ~ i i ~ ~ t i c l p hLi ai :p p i ~ l ~ \'Cilliams or~ & Willtil~s,2001. - 2477 p. 46.Ruder R.E.J. The danger of higli dose sodi~1111 bicirri~orintein biguanide-incl~~ced lactic acitlosis: the theory, t h e prac1-ice allti alterriative therapies. /,' Br. J . Clin. Pract. - 1.987. - \-01. 4 1 . P. 730-731. 47.Zpigel- M. Hypopartl~yroitlis~n, psei~clol~!;popi~r~i~i:~~:~oi(!i and p s e u t l o p s e u d o l i y ~ ~ o p a r a t l ~ ~ r o i ~TcxtSook iism: of el~dr!r.!-i~le surgery : Ed. 0. Clark, Q.-Y. Philaclelphia: W.B. S o ~ ~ i l t l ? r Co~npan!?,1997. - P. 523-528.